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In regions where both species coexist, they are divided by landscape gradients such as the age of forest, elevation, and land openness. Grizzly bears tend to favor old forests with high productivity, higher elevations and more open habitats compared with black bears. However, a bear shot in autumn 1986 in Michigan was thought by some to be a grizzly×black bear hybrid, due to its unusually large size and its proportionately larger braincase and skull, but DNA testing was unable to determine whether it was a large American black bear or a grizzly bear.