红岩
The Red Crag Formation is representative of a temperate, shallow nearshore environment, perhaps at the mouth of a large river indicated by conifer pollen and small terrestrial vertebrate remains. Lydekker, along with "O. citoniensis", identified several other whales from the formation, for example the dubious baleen whale "Balaenoptera sibbaldina", the sperm whale "Hoplocetus", the shark toothed dolphin "Squalodon antverpiensis", as well as some species that exist in modern day such as the northern bottlenose whale ("Hyperoodon ampullatus"), the strap-toothed whale ("Mesoplodon layardii"), and the long-finned pilot whale ("Globicephala melas"). Shark teeth and skate coprolite were also found here.