奧塞內爾·查利斯·馬什
In the subsequent years of the 19th century, the London specimen was subject to a number of analyses by other researchers of historical importance, including Thomas Huxley, Othniel Charles Marsh, Wilhelm Dames, and Bronislav Petronievics. Much of this early study was relevant to controversy surrounding the possible taxonomic divisions of "Archaeopteryx" specimens.