Shantar Islands
The brown bear is found in the Ussuri krai, Sakhalin, the Amur Oblast, northward to the Shantar Islands, Iturup Island, northeastern China, the Korean peninsula, Hokkaidō and Kunashiri Island. Until the 13th century, bears inhabited the islands of Rebun and Rishiri, having crossed the La Pérouse Strait to reach them. They were also present on Honshu during the last glacial period, but were possibly driven to extinction either by competing with Asian black bears or by habitat loss due to climate change. There are three genetic groups, distinct for at least 3 million years which reached to Hokkaido via Honshu at different times.