Presidente Prudente
"Pepesuchus" is known from the holotype MN 7005-V, which consists of nearly complete skeleton and skull, and from the paratype MCT 1788-R, a nearly complete skull and lower jaw likely from the same locality of the holotype. The specimens were recovered from the Presidente Prudente Formation of the Bauru Group of the northern Paraná Basin, between the cities of Pirapozinho and Presidente Prudente. In 2019, new fossil material from the Adamantina Formation was described.