تشارلز موريس
Apart from Charles Sanders Peirce (1839–1914) and Charles W. Morris (1903–1979), early pioneers of biosemiotics were Jakob von Uexküll (1864–1944), Heini Hediger (1908–1992), Giorgio Prodi (1928–1987), Marcel Florkin (1900–1979) and Friedrich S. Rothschild (1899–1995); the founding fathers of the contemporary interdiscipline were Thomas Sebeok (1920–2001) and Thure von Uexküll (1908–2004).