Hrib
Chemical analysis of pigments and chromogens showed that "Suillus" was more closely related to "Gomphidius" and "Rhizopogon" than to other boletes, and hence "Suillus bovinus" and its allies were transferred from the Boletaceae to the newly circumscribed family Suillaceae in 1997. Molecular studies have reinforced how distantly related these fungi are from "Boletus edulis" and its allies.