岡山大学
He was lecturer in parasitology in Kyoto University (1927-1943), parasitologist at the Naval Institute of Tropical Hygiene in Macassar (then "Celebes" – now Sulawesi, Indonesia) with the Japanese Navy (1943-1944), and special consultant of the Malaria Survey Detachment of the US Army (1946-1950). He became Professor of parasitology in Okayama University Medical School (1950), was a visiting professor at the University of Hawaii (1962-1966) and a Graduate Professor of Biology at Tulane University (1968-1969).