Duvalius

Duvalius  er en slekt av billene i delgruppen (underfamilien) skyggeløpere (Trechinae) i familiegruppen løpebiller. 


== Utseende ==
Små til middelsstore (ca. 5 millimeter), spinkle, blanke, rødbrunlige løpebiller med lange antenner. De har reduserte øyne og er blinde.


== Levevis ==
Artene lever i huler, der de trolig ernærer seg av andre smådyr.


== Utbredelse ==
Slekten er utbredt i hulesystemer i Frankrike, Italia, på Balkan, i Lilleasia og Kaukasus.


== Systematisk inndeling ==
Systematikken følger Carabidae of the World. 

ordenen biller, Coleoptera Linnaeus, 1758
underordenen Adephaga Schellenberg, 1806
overfamilien løpebiller og vannkalver, Caraboidea Latreille, 1802
familien   løpebiller, Carabidae Latreille, 1802
underfamilie skyggeløpere Trechinae Bonelli, 1810
stammen Trechini Bonelli, 1810
delgruppe Trechina Bonelli, 1810
Slekten Duvalius Delarouzee, 1859
Underslekten Biharotrechus Bokor, 1922
Duvalius abnormis Knirsch, 1913
Duvalius bedelensis Janak & P. Moravac, 1989
Duvalius bekchievi Gueorgiev & Kostova, 2011
Duvalius beshkovi Coiffait, 1970
Duvalius biokovensis Holdhaus, 1911
Duvalius breitianus Knirsch, 1912
Duvalius brevipilosus Knirsch, 1927
Duvalius budae Kenderessy, 1879
Duvalius cicioarae Jeannel, 1930
Duvalius cognatus J. Frivaldszky, 1879
Duvalius coiffaiti Decou, 1967
Duvalius comes Scheibel, 1934
Duvalius droveniki Magrini, 1998
Duvalius durmitorensis Apfelbeck, 1904
Duvalius edithae Janak & P. Moravec, 1989
Duvalius golesensis Winkler, 1926
Duvalius hegeduesii J. Frivaldszky, 1880
Duvalius herculis J. Frivaldszky, 1889
Duvalius hickeri Knirsch, 1913
Duvalius hurkai Janak & P. Moravec, 1989
Duvalius kanabei Csiki, 1940
Duvalius knirschi Janak & P. Monavec, 1989
Duvalius laevigatus Bokor, 1913
Duvalius maglicensis Winkler, 1933
Duvalius mallaszii Csiki, 1901
Duvalius mandibularis Jeannel, 1930
Duvalius mariannae Knirsch, 1924
Duvalius meschniggi Meixner, 1928
Duvalius milleri J. Frivaldszky, 1862
Duvalius nannus Jeannel, 1931
Duvalius oltenicus Jeannel, 1928
Duvalius paroecus J. Frivaldszky, 1865
Duvalius peristericus J. Muller, 1914
Duvalius pilifer Ganglbauer, 1891
Duvalius redtenbacheri I. Frivaldszky von Frivald & J. Frivaldszky, 1857 
Duvalius roseni Jeannel, 1929
Duvalius scarisoarae Knirsch, 1913
Duvalius semecensis Winkler, 1926
Duvalius speiseri Ganglbauer, 1892
Duvalius spiessi Jeannel & Mallasz, 1929
Duvalius spinifer Jeannel, 1928
Duvalius stilleri Reitter, 1913
Duvalius sziladyi Csiki, 1904
Duvalius transcarpathicus Shilenkov & Rizun, 1989
Duvalius trescavicensis Ganglbauer, 1891
Duvalius voitestii Jeannel, 1930
Duvalius wercharatskii Rizun & Yanitskiy, 1994
Duvalius winneguthi Apfelbeck, 1907
Underslekten Duvalius (i snever forstand)
Duvalius adelphus Magrini, Petrioli, Degiovanni & Bastianini, 2008
Duvalius aliciae Magrini, Petrioli, Degiovanni & Bastianini, 2008
Duvalius andreinii (Gestro, 1907)
Duvalius andreuccii Magrini & Vanni, 1984
Duvalius annamariae Vanni & Magrini, 1989
Duvalius antonellae Casale, Giachino, Vailati & Vigna Taglianti, 1996
Duvalius apuanus Dodero, 1917
Duvalius aroaniae Casale, Giachino, Vailati & Vigna Taglianti, 1996
Duvalius auberti Grenier, 1864
Duvalius auruncus Vigna Taglianti & Magrini, 2008
Duvalius ausonicus Vigna Taglianti & Magrini, 2008
Duvalius avoni Vigna Taglianti & Magrini, 2008
Duvalius balazuci Bruneau de Mire, 1948
Duvalius baldensis Putzeys, 1870
Duvalius balearicus Henrot, 1964
Duvalius bastianini Magrini, 1998
Duvalius battonii Straneo, 1959
Duvalius bensae Castro, 1892
Duvalius bertagnii Magrini, 1998
Duvalius berthae (Jeannel, 1910)
Duvalius bianchii Jeannel, 1928
Duvalius bischoffi Maschnigg, 1936
Duvalius boldorii Jeannel, 1926
Duvalius bonadonius Giordan & Raffaldi, 1982
Duvalius bonzanoi Casale & Vigna Taglianti, 1990
Duvalius boschi Jeannel, 1929
Duvalius brandisi Ganglbauer, 1896
Duvalius brandmayri Vigna Taglianti & Magrini, 2008
Duvalius breiti Ganglbauer, 1900
Duvalius brucki (Piccioli, 1870)
Duvalius brujasi Sainte-Claire Deville, 1901
Duvalius cadurcus Jeannel, 1955
Duvalius cailloli Sainte-Claire Deville, 1902
Duvalius calandrii Casale & Vigna Taglianti, 1990
Duvalius canevae Gestro, 1885
Duvalius caoduroi Pace, 1986
Duvalius carantii Sella, 1874
Duvalius carchinii Vigna Taglianti, Magrini & Vanni, 1993
Duvalius casalei Sciaky, 1992
Duvalius casellii Gestro, 1898
Duvalius cerrutii Sbordoni & Di Domenico, 1967
Duvalius cirocchii Magrini & Vanni, 1986
Duvalius clairi Abeille de Perrin, 1881
Duvalius comottii Casale, Giachino, Vailati & Vigna Taglianti, 1996
Duvalius consortii Magrini & Degiovanni, 2006
Duvalius convexicollis Peyerimhoff, 1904
Duvalius cornilloni Giordan & Raffaldi, 1998
Duvalius curtii Giordan & Raffaldi, 1983
Duvalius dattii Vigna Taglianti & Magrini, 2008
Duvalius degiovannii Magrini & Vanni, 1985
Duvalius delphinensis Abeille de Perrin, 1869
Duvalius diaphanus Rottenberg, 1874
Duvalius didonnai Magrini, Vanni & Inguscio, 1996
Duvalius diniensis Peyerimhoff, 1904
Duvalius doderoi Gestro, 1885
Duvalius dolops Jeannel, 1937
Duvalius doriae Fairmaire, 1859
Duvalius dragacevensis S.Curcic, Pavicevic & B.Curcic, 2001
Duvalius exaratus Schaum, 1860
Duvalius fodori Scheibel, 1937
Duvalius franchettii Luigioni, 1926
Duvalius franziniorum Vigna Taglianti & Magrini, 2008
Duvalius fuchsi Scheibel, 1937
Duvalius fulvii Casale, Giachino, Vailati & Vigna Taglianti, 1996
Duvalius gabriellavernae Casale, Giachino, Vailati & Vigna Taglianti, 1999
Duvalius genesti Casale & Vigna Taglianti, 1984
Duvalius gentilei Gestro, 1885
Duvalius georgii J. Muller, 1922
Duvalius gestroi Dodero, 1900
Duvalius ghidinii Gestro & Dodero, 1909
Duvalius giachinoi Casale & Vigna Taglianti, 1990
Duvalius graecus Casale, Giachino, Vailati & Vigna Taglianti, 1996
Duvalius guareschii Moscandini, 1950
Duvalius hartigi Magrini, Bavieri & Vigna Taglianti, 2006
Duvalius innamoratorum Magrini & Cirocchi, 2002
Duvalius iolandae Magrini & Vanni, 1986
Duvalius iuliae Casale, Giachino, Vailati & Vigna Taglianli, 1996
Duvalius iulianae Vigna Taglianti & Casale, 1973
Duvalius jureceki Dodeno, 1917
Duvalius krueperi Schaum, 1862
Duvalius kyllenicus Scheibel, 1937
Duvalius laneyriei J. Ochs, 1948
Duvalius lantosquensis Abeille da Perrin, 1881
Duvalius latellai Vigna Taglianti & Magrini, 2008
Duvalius laurentii Magrini & Bastianini, 2003
Duvalius lemairei Giordan & Raffaldi, 1982
Duvalius leonhardi Reitter, 1901
Duvalius lepinensis Cenuti, 1950
Duvalius lespesii Fairmaire, 1867
Duvalius longhii Comolli, 1837
Duvalius lucarellii Casale & Vigna Taglianti, 1990
Duvalius lydiae Vigna Taglianti & Magrini, 2008
Duvalius magdelainei Jeannel, 1914
Duvalius magistrettianus Schatzmayr, 1940
Duvalius maglianoi Giordan & Raffaldi, 1983
Duvalius marii Vanni, Magrini & Pennisi, 1991
Duvalius matocqi Giordan, 1987
Duvalius meixneri Kreissl, 1993
Duvalius merisioi Casale, Giachino, Vailati & Vigna Taglianti, 1996
Duvalius milenae Casale, 1983
Duvalius minozzii Dodero, 1917
Duvalius mirei Deuve, 2001
Duvalius mixanigi Daffner, 1993
Duvalius moczarskyi J. Muller, 1917
Duvalius montisageli Jeannel, 1947
Duvalius montisoetae Casale, 1987
Duvalius morisii Vigna Taglianti & Casale, 1973
Duvalius muelleri Winkler, 1933
Duvalius muriauxi Jeannel, 1957
Duvalius nambinensis Boldori, 1935
Duvalius nardii Vigna Taglianti & Magrini, 2008
Duvalius occitanus Casale & Vigna Taglianti, 1993
Duvalius ochsi Dodero, 1921
Duvalius oertzeni L. Miller, 1884
Duvalius olympiadicus Scheibel, 1937
Duvalius oscus A. Franzini & C. Franzini, 1984
Duvalius parnassianus Deuve, 2001
Duvalius paulinae (Fagniez, 1922)
Duvalius pecoudi Jeannel, 1937
Duvalius pennisii Magrini & Vanni, 1984
Duvalius perrinae Giordan, 1989
Duvalius pominii Schatzmayr, 1943
Duvalius raffaldii Curti, 1981
Duvalius ramorinii (Gestro, 1887)
Duvalius raymondi Delarouzee, 1859
Duvalius regiszogui Maschnigg, 1936
Duvalius roberti Abeille de Perrin, 1903
Duvalius rossii Magrini & Vanni, 1991
Duvalius ruffoi Magistretti, 1956
Duvalius sardous (Dodero, 1917)
Duvalius sbordonii Vigna Taglianti, Genest & Sciaky, 1980
Duvalius sclanoi Magrini & Vanni, 1996
Duvalius sicardi Fagniez, 1923
Duvalius siculus (Baudi di Selva, 1882)
Duvalius silvestrii (Gestro, 1896)
Duvalius simoni (Abeille de Perrin, 1881)
Duvalius smolikanus Casale, Giachino, Vailati & Vigna Taglianti, 1996
Duvalius spaethi (Ganglbauer, 1904)
Duvalius springeri (J. Muller, 1921)
Duvalius starivlahi B.Gueorguiev, S.Curcic & B.Curcic, 2000
Duvalius straneoi Jeannel, 1931
Duvalius strupii Scheibel, 1937
Duvalius sturanyi (Apfelbeck, 1904)
Duvalius taygetanus Casale, 1979
Duvalius turcati Giordan & Raffaldi, 1983
Duvalius vallombrosus (C. & F. Rasetti, 1920)
Duvalius vannii Magrini & Sclano, 1998
Duvalius vermionensis (Casale, 1983)
Duvalius vignai Casale, 1983
Duvalius villiersi Giordan & Raffaldi, 1983
Duvalius virginiae Magnini, Vanni & Cirocchi, 1996
Duvalius volscus A. Franzini & G. Franzini, 1984
Duvalius voraginis Jeannel & J. Ochs, 1938
Duvalius waillyi Giordan & Raffaldi, 1982
Duvalius weiratheri Scheibel, 1937
Duvalius wichmanni Jeannel, 1929
Duvalius wingelmuelleri (Ganglbauer, 1904)
Duvalius winklerianus Jeannel, 1926
Duvalius zaimisi Jeannel, 1929
Underslekten Euduvalius Jeannel, 1928
Duvalius erichsonii Schaufuss, 1864
Duvalius lucidus J. Muller, 1903
Duvalius maglajensis Apfelbeck, 1908
Duvalius novaki J. Muller, 1911
Duvalius petrochilosi Coiffait, 1965
Duvalius reiseri Ganglbauer, 1891
Duvalius ruffoanus Casale, Giachino, Vailati & Vigna Taglianti, 1996
Duvalius zhalovi Gueorgiev, 2005
Underslekten Hungarotrechus Bokor, 1922
Duvalius babicola Knirsch, 1925
Duvalius bokori Csiki, 1910
Duvalius bucurensis Janak, 1993
Duvalius centenarius Knirsch, 1925
Duvalius corpulentus Weise, 1875
Duvalius delamarei Decou, 1967
Duvalius deubelianus Csiki, 1903
Duvalius dieneri Csiki, 1910
Duvalius gaali Mallasz, 1928
Duvalius ganglbauerianus Knirsch, 1913
Duvalius godeanus Janak, 1993
Duvalius goemoeriensis Bokor, 1922
Duvalius gracilis Petri, 1912
Duvalius hungaricus Csiki, 1903
Duvalius kimakowiczi Ganglbauer, 1891
Duvalius merkli J. Frivaldszky, 1877
Duvalius microphthalmus L. Miller, 1859
Duvalius onaci Moldovan, 1993
Duvalius poporogui Decou, 1973
Duvalius proceroides Jeannel, 1926
Duvalius procerus Putzeys, 1847
Duvalius richardi Knirsch, 1925
Duvalius roubali Jeannel, 1926
Duvalius ruthenus Reitter, 1878
Duvalius subterraneus (L. Miller, 1868)
Duvalius szaboi Csiki, 1914
Underslekten Neoduvalius G. Muller, 1913
Duvalius bolei Pralmer, 1963
Duvalius bradycephalus Jeannel, 1928
Duvalius cuniculinus Knirsch, 1929
Duvalius cvijici (Jeannel, 1924)
Duvalius eurydice Schaufuss, 1881
Duvalius humerosus Knirsch, 1926
Duvalius klimeschi Winkler, 1914
Duvalius kodrici Scheibel, 1938
Duvalius langhofferi Csiki, 1913
Duvalius neumanni J. Muller, 1911
Duvalius opermanni Scheibel, 1933
Duvalius petraeus Knirsch, 1927
Duvalius reitteri Miller, 1881
Duvalius schatzmayri J. Muller, 1912
Duvalius styx Apfelbeck, 1904
Duvalius vranensis Breit, 1904
Underslekten Paraduvalius Knirsch, 1924
Duvalius balcanicus (J. Frivaldszky, 1879)
Duvalius beroni V.B. Gueorguiev, 1971
Duvalius bulgaricus Knirsch, 1924
Duvalius bureschi Jeannel, 1928
Duvalius deltshevi V.B. Gueorguiev, 1965
Duvalius garevi Casale & Genest, 1991
Duvalius germanae Casale & Vigna Taglianti, 1990
Duvalius glabellus G. Etonti & M. Etonti, 1984
Duvalius hanae Hurka, 1990
Duvalius joakimovi Gueorgiev, 2005
Duvalius joannidisi Casale, Giachino & M. Elonti, 1990
Duvalius karelhurkai Farkac, 1990
Duvalius kotelensis Genest, 1978
Duvalius legrandi Genest, 1983
Duvalius marani Knirsch, 1930
Duvalius papasoffi Mandl, 1942
Duvalius petrovi Gueorgiev, 2005
Duvalius philippensis Jeannel, 1937
Duvalius pirinensis Gueorgiev, 2005
Duvalius pretneri V.B. Gueorguiev, 1971
Duvalius pruinosus Jeannel, 1937
Duvalius rajtchevi Genest & Juberthie, 1983
Duvalius regisborisi Buresch, 1926
Duvalius stankovitchi Jeannel, 1924
Duvalius sydowi Jeannel, 1930
Duvalius winkleri Jeannel, 1924
Duvalius zivkovi (Knirsch, 1925)
Underslekten Platyduvalius Jeannel, 1928
Duvalius gogalai Pretnen, 1963
Duvalius macedonicus J. Muller, 1917
Underslekten Trechopsis Peyerimhoff, 1908
Duvalius ferreresi Lagar Mascare, 1976
Underslekten Typhloduvalius Hurka & Pulpan, 1980
Duvalius gebhardti Bokor, 1926


=== Liste over arter ikke plassert til underslekt ===
Duvalius abyssimus Reboleira & Ortuño, 2014
Duvalius alexeevi Belousov, 1992
Duvalius antoniae Reitter, 1892
Duvalius armeniacus Casale, 1979
Duvalius arnoldii Jeannel, 1962
Duvalius baborensis Bruneau de Mire, 1955
Duvalius bicikensis Perrault, 1971
Duvalius bodoanus Reitter, 1913
Duvalius bortesii Casale & Vigna Taglianti, 1984
Duvalius bruschii Vigna Taglianti, 1999
Duvalius dvoraki P. Moravec, 1986
Duvalius gusevi Belousov, 1989
Duvalius hetschkoi (Reitter, 1911)
Duvalius huberi Casale, 2011
Duvalius huetheri Jeannel, 1934
Duvalius iblis Payerimhoff, 1910
Duvalius iljukhini Dalzhanski & Ljovuschkin, 1985
Duvalius jurjurae Peyerimhoff, 1909
Duvalius koeni Muilwijk & Felix, 2008
Duvalius kryshanovskii Jeannel, 1962
Duvalius kurnakovi Jeannel, 1960
Duvalius lapiei Peyerimhoff, 1908
Duvalius lencinai Mateu & Ortuno, 2006
Duvalius martensi Casale, 1983
Duvalius martinae Jeanne, 1996
Duvalius megrel Belousov, 1992
Duvalius miroshnikovi Belousov & Zamotajlov, 1995
Duvalius mohammadzadehi Muilwijk & Felix, 2008
Duvalius ovtshinnikovi Belousov, 1992
Duvalius panoecus J. Frivaldszky, 1865
Duvalius phoenicius Vigna Taglianti, 1973
Duvalius putshkovi Belousov & Kabak, 1999
Duvalius saueri Farkac & P. Moravec, 1993
Duvalius sokolovi Ljovuschkin, 1963
Duvalius stepanavanensis Iablokoff-Khnzorian, 1963
Duvalius stopicensis (Jeannel, 1924)
Duvalius vartashensis Belousov, 1989
Duvalius yatsenkokhmelevskyi Iablokoff-Khnzorian, 1960
Duvalius zlatiborensis (Curcic, Brajkovic & Curcic, 2005)


== Referanser ==


== Eksterne lenker ==
Bilde av Duvalius delphinensis
(en) Duvalius i Encyclopedia of Life 
(en) Duvalius i Global Biodiversity Information Facility 
(en) Duvalius hos Fauna Europaea 
(en) Duvalius hos NCBI 
Duvalius – detaljert informasjon på Wikispecies