Athous

Athous er en slekt av biller som hører til familien smellere (Elateridae).


== Utseende ==
Små til middelsstore (5 – 20 millimeter), slanke, blanke, fint hårete, brune til svarte smellere. Hodet er kort og bredt, med små fasettøyne. Antennene er 11-leddete, vanligvis ganske lange, mer elelr mindre sagtakkede. Brystskjoldet (pronotum) er rundet firkantet, gjerne noe lengre enn bredt. Sidekantene er tydelig markerte, og bakhjørnene er litt tilbaketrukket. Dekkvingene er langsmale, omtrent så brede som brystskjoldet, som oftest med tydelige lengdestriper.
Larvene er sylindriske, glatte med en voksaktige overflate og spredte hår, gjerne rød- eller gulaktige. Hodet er noe flatt med fremoverstrakte kjever. Det bakerste leddet har vanligvis et par kraftige utvekster av varierende form. Beina er ganske korte.


== Levevis ==
Denne slekten har larver som lever i jorda, der de gjerne eter planterøtter.


== Skadedyr og nyttedyr ==
Noen arter i slekten er brysomme skadedyr i jord- og hagebruk fordi larvene (kjølmark) spiser røtter.


== Utbredelse ==
Slekten har en holarktisk utbredelse, med særlig mange arter i Sør-Europa og Tyrkia.


== Systematisk inndeling ==
Noen arter som tradisjonelt var plassert her blir nå regnet til slekten Hemicrepidius.

Ordenen biller, Coleoptera
Underordenen Polyphaga
Overfamilien smellere, Elateroidea
Familien smellere, Elateridae
Underfamilien tannsmellere (Dendrometrinae) Gistel, 1856 – slanke arter
Stammen Dendrometrini Gistel, 1856
Understammen Athoina Candéze, 1860
Slekten Athous Eschscholtz, 1829
Underslekten Alcimathous Reiiter, 1905
Athous mokrzeckii Lomnicki, 1923
Athous sacheri Kiesenwetter, 1858
Underslekten Athous (i snever forstand)
Athous cingulatus L. Miller, 1881
Athous densatus Reitter, 1905
Hasselsmeller, Athous haemorrhoidalis (Mal:Fabricius, 1801)
Athous ineptus Candèze, 1860
Athous limoniiformis Candèze, 1865
Athous nadoraz Mertlik & Dusanek, 2006
Athous nigricornis Brisout, 1856
Athous rubrotestaceus Desbrochers de Loges, 1871
Athous vittatoides Reitter, 1905
Løvsmeller, Athous vittatus (Fabricius, 1792)
Underslekten Euplathous Reitter, 1905
Athous canus Dufour, 1843
Athous dejeanii (Laporte, 1840)
Athous frigidus Mulsant & Guillebeau, 1855
Athous iablokoffi Leseigneur, 1972
Athous mandibularis (Dufour, 1843)
Athous proximus Hampe, 1864
Athous villardi Carret, 1904
Underslekten Haplathous Reitter, 1905
Athous abdurachmanovi Dolin in Dolin & Penev, 2004
Athous alpestris Orlov, 1994
Athous angulifrons Reitter, 1905
Athous austriacus Desbrochers des Loges, 1873
Athous barthei Leseigneur, 1958
Athous belloi Guglielmi & Platia, 1985
Athous brachati Zeising & Brunne, 2005
Athous brevicornis Desbrochers des Loges, 1871
Athous cachecticus Candèze, 1860
Athous carpathicus Reitter, 1905
Athous cavifrons L. Redtenbacher, 1858
Athous circassiensis Reitter, 1905
Athous circumductus (Ménétriés, 1832)
Athous corcyreus Reitter, 1905
Athous corsicus Reiche, 1861
Athous crassicornis Candèze, 1860
Athous csikii Platia, 2001
Athous curtus Dolin in Dolin & Penev, 2004
Athous daghestanicus Reitter, 1890
Athous delmastroi Platia & Gudenzi, 1998
Athous dilaticornis Reitter, 1905
Athous dilutus (Illiger, 1807)
Athous emaciatus Candèze, 1860
Athous evae Platia & Nemeth, 2011
Athous euxinus Buysson, 1912
Athous filicornis (Dufour, 1851)
Athous flavipennis Candèze, 1860
Athous hilfi Reitter, 1912
Athous ilniczkyi Platia & Nemeth, 2011
Athous incognitus Platia, 1988
Athous iristonicus Dolin, 1971
Athous kerkyranus Reitter, 1905
Athous kobachidzei Dolin & Chantladze, 1982
Athous korabicus Csiki, 1940
Athous korsosi Platia & Nemeth, 2011
Athous laevistriatus (Dufour, 1851)
Athous latior Orlov, 1994
Athous lepontinus Schwarz, 1900
Athous leprieuri Desbrochers des Loges, 1870
Athous lgockii Dolin, 1983
Athous longicornis Candèze, 1865
Athous magnanii Guglielmi & Platia, 1985
Athous marginicollis Reitter, 1890
Athous melanoderes Mulsant & Guillebeau, 1855
Athous mollis Reitter, 1889
Athous monilicornis Schwarz, 1897
Athous oromii Platia & Gudenzi, 2005
Athous osellai Guglielmi & Platia, 1985
Athous pacei Guglielmi & Platia, 1985
Athous panellai Platia & Schimmel, 1991
Athous patoni Dolin in Dolin & Penev, 2004
Athous pfefferi Roubal, 1932
Athous protoracicus Platia & Schimmel, 1991
Athous putschkovi Dolin & Penev, 2004
Athous pyrenaeus Candèze, 1865
Athous reflexicollis Dolin & Penev, 2004
Athous reynosae C. N. F. Brisout de Barneville, 1866
Athous ruteri Chassain, 1985
Athous settei Guglielmi & Platia, 1985
Athous siteki Platia, 2006
Athous sosybius Reitter, 1905
Athous spalatrensis Reitter, 1894
Busksmeller, Athous subfuscus (Müller, 1764)
Athous talamellii Platia & Gudenzi, 1998
Athous tauricola Reitter, 1905
Athous tauricus Candèze, 1860
Athous transylvanicus Platia, 2001
Athous utschderensis Reitter, 1890
Athous tschukini Reitter, 1910
Athous turcicus Reitter, 1905
Athous villiger Mulsant & Guillebeau, 1855
Athous villosulus Desbrochers des Loges, 1875
Athous vivesi Platia & Gudenzi, 2005
Athous zebei Bach, 1852
Underslekten Neonomopleus Schenkling, 1927
Athous alcantarensis De la Rosa et al., 2011
Athous chiarae Platia, 2010
Athous chloroticus Candèze, 1865
Athous cobosi Platia, 2003
Athous discors Reitter, 1904
Athous elongatus Brisout, 1866
Athous escorialensis Mulsant & Guillebeau, 1856
Athous fernandensis Platia, 2010
Athous kiesenwetteri (Schaufuss, 1862)
Athous koschwitzi Platia, 2003
Athous longissimus Reitter, 1904
Athous lopezcoloni Platia & Gudenzi, 2005
Athous martinae Platia, 2003
Athous martinezi Reitter, 1904
Athous masoae Platia, 2003
Athous procerus (Illiger, 1807)
Athous recticollis Graells, 1858
Athous schimmeli Platia, 2003
Athous schurmanni Platia & Serrano, 2002
Athous sofiae Platia, 2010
Athous suarezi Platia & Gudenzi, 2005
Athous tenuis Brisout, 1866
Althous toribioi Zapata de la Vega & Sanchez-Ruiz, 2005
Underslekten Orthathous Reitter, 1905
Athous abkhazianus Platia & Nemeth, 2011
Athous acutangulus Fairmaire, 1866
Athous albanicus Csiki, 1940
Athous alticola Platia, 2006
Athous anatolicus Platia, 1989
Athous angustulus Candéze, 1860
Athous apfelbecki Reitter, 1905
Athous artvinensis Platia, Yildirim & Kesdek, 2007
Athous astrabadensis Faust, 1877
Athous asturiensis Platia, 2006
Athous audisioi Guglielmi & Platia, 1985
Athous azoricus Platia & Gudenzi, 2002
Athous balcanicus Reitter, 1905
Athous barriesi Platia & Gudenzi, 1996
Athous benedikti Platia, 2003
Athous bicolor (Goeze, 1777)
Athous binaghii Platia, 1984
Athous bolivari Reitter, 1904
Athous bolognai Guglielmi & Platia, 1985
Athous brevipennis Schwarz, 1897
Athous brisouti Sanchez-Ruiz, 1996
Athous bulgaricus Platia, 2001
Athous cagatayae Kabalak & Sert, 2012
Athous campyloides Newman, 1833
Athous cantabricus Schaufuss, 1862
Athous carpathophilus Reitter, 1905
Athous catalonicus Platia, 2006
Athous cavazzutii Platia, 2003
Athous caviformis Reitter, 1905
Athous cavulus Reitter, 1905
Athous cavus (Germar, 1817)
Athous cervicolor Heyden, 1880
Athous chamboveti Mulsant & Godart, 1868
Athous codinai Platia, 2006
Athous conradi Platia, 2006
Athous croaticus Platia & Gudenzi, 2002
Athous curtulus Desbrochers des Loges, 1873
Athous daccordii Guglielmi & Platia, 1985
Athous dalmatinus Platia, 2005
Athous dasycerus Buysson, 1890
Athous debilis Reiche, 1869
Athous demirsoyi Platia, Kabalak & Sert, 2007
Athous desbrochersi Platia, 2006
Athous difficilis (Dufour, 1843)
Athous distinctithorax Desbrochers des Loges, 1873
Athous divaricatus Platia, 2006
Athous dorgaliensis Buysson, 1912
Athous durazzoi Platia, 1985
Athous dusaneki Platia, 2003
Athous eckerleini Platia & Gudenzi, 2000
Athous edirnensis Platia & Gudenzi, 2000
Athous epirus Stierlin, 1875
Athous escolai Platia, 2006
Athous espanoli Platia, 2006
Athous espinamensis Platia, 2006
Athous fodorjenoi Platia & Nemeth, 2011
Athous fragariae Platia & Kovanci, 2005
Athous freudei Platia, 1989
Athous frontalis Platia & Schimmel, 1991
Athous fuentei Platia, 2006
Athous gagliardii Platia, 1988
Athous galiberti Buysson, 1918
Athous gallicus Platia & Gudenzi, 2000
Athous ganglbaueri Schwarz, 1897
Athous gerezianus Reitter, 1905
Athous gianassoi Platia & Gudenzi, 1996
Athous giustoi Platia, 2006
Athous gobanzi Reitter, 1905
Athous gonzalesi Platia, 2006
Athous goriciensis Reitter, 1905
Athous gottwaldi Lohse, 1978
Athous gracacensis Platia, 2005
Athous graecus Platia, 1989
Athous guadalupensis Platia, 2006
Athous gudenzii Guglielmi & Platia, 1985
Athous hajeki Platia & Gudenzi, 2005
Athous harmodius Reitter, 1905
Athous herbigradus Mulsant & Guillebeau, 1855
Athous hercegovinensis Platia & Nemeth, 2011
Athous hetzeli Platia, 2004
Athous holtzi Reitter, 1905
Athous ibericus Platia, 2006
Athous insularis Desbrochers des Loges, 1869
Athous iranicus Platia, 2004
Athous jakupiciola Platia & Nemeth, 2011
Athous jejunus Kiesenwetter, 1858
Athous judicariensis Schwarz, 1900
Athous kabalaki Platia, 2010
Athous kasovskyi Platia & Gudenzi, 2007
Athous kaszabi Dolin, 1986
Athous kovancii Platia, 2003
Athous kozufensis Platia, 2010
Athous kruegeri Reitter, 1905
Athous lambeleti Leseigneur, 2004
Athous lasoni Platia, 2011
Athous lassallei Platia & Gudenzi, 1996
Athous leonhardi Reitter, 1905
Athous leseigneuri Platia, 2006
Athous lomnickii Reitter, 1905
Athous luigionii Platia, 1988
Athous lusitanus Platia, 2006
Athous maljenensis Platia, 2013
Athous malkinorum Platia, 2006
Athous malmusii Platia & Gudenzi, 2000
Athous margheritae Guglielmi & Platia, 1985
Athous mauroi Platia, 2010
Athous melonii Platia, 1984
Athous mendesi Giuseppe & Serrano, 2002
Athous mertliki Platia & Gudenzi, 2002
Athous meuseli Reitter, 1905
Athous mingrelicus Reitter, 1890
Athous monguzzii Platia & Gudenzi, 2007
Athous nadari Buysson, 1904
Athous naseri J. Müller, 1916
Athous nigror Platia, 2006
Athous novaki Penecke, 1907
Athous obsoletus (Illiger, 1807)
Athous olbiensis Mulsant & Guillebeau, 1856
Athous olcesei Buysson, 1905
Athous paflagonensis Platia & Gudenzi, 1998
Athous paganettii Platia, 2006
Athous pallidus Platia & Gudenzi, 2002
Athous pedemontanus Platia, 1988
Athous penevi Platia, 2001
Athous perezarcasi Platia, 2006
Athous picipennis Reitter, 1905
Athous plagipennis Reitter, 1905
Athous podlussanoi Platia & Nemeth, 2011
Athous pomboi Platia & Borges, 2003
Athous ponticus Platia & Gudenzi, 2007
Athous propinquus Buysson, 1890
Athous prouzai Platia, 2005
Athous putativus Platia, 2006
Athous reitteri Platia, 2006
Athous ruffoi Guglielmi & Platia, 1985
Athous sabatinellii Guglielmi & Platia, 1985
Athous samai Guglielmi & Platia, 1985
Athous sameki Platia, 2003
Athous sandanskiensis Platia, 2010
Athous senaci Buysson, 1890
Athous serbicus Reitter, 1905
Athous serranoi Platia, 2006
Athous silicensis Laibner, 1975
Athous singularis Reitter, 1905
Athous sinuatocollis Desbrochers des Loges, 1869
Athous slavonicus Platia & Nemeth, 2011
Athous snizeki Platia, 2004
Athous solitarius Platia, 2010
Athous stoimenovae Platia, 2001
Athous subtruncatoides Platia, 2006
Athous subtruncatus Mulsant & Guillebeau, 1856
Athous subvirgatus Daniel, 1904
Athous svihlai Platia & Gudenzi, 1998
Athous szombathyi Schenkling, 1927
Athous tadzhikistanicus Platia, 2010
Athous tekkirazicus Platia, 2003
Athous telciuensis Platia, 2013
Athous tomentosus Mulsant & Guillebeau, 1855
Athous tribertii Guglielmi & Platia, 1985
Athous vasconicus Platia, 2006
Athous vavrai Platia, 2006
Athous vicinus Desbrochers des Loges, 1873
Athous vomeroi Platia, 1988
Athous warchalowskii Platia & Tarnawski, 1998
Athous wewalkai Platia, 1989
Athous zanettii Guglielmi & Platia, 1985
Athous zappiorum Platia, 1985
Athous zbuzeki Platia & Gudenzi, 2007
Athous ziegleri Zeising & Brunne, 2005
Underslekten Pleurathous Reitter, 1905
Athous circassicus Reitter, 1888
Athous godarti Mulsant & Guillebeau, 1853
Athous rosinae Reitter, 1899
Athous uncicollis Perris, 1864
Underslekten Roznerathous Nemeth & Platia, 2014
Athous szalokii Nemeth & Platia, 2014
Ikke plassert til underslekt
Athous alnicola Gistel, 1857
Athous dimidiatus Gistel, 1857
Athous fausti Reitter, 1890
Athous francoisi Fleutiaux, 1928
Athous humeralis (Fischer von Waldheim, 1824)
Athous kubani Schimmel, 1998
Athous opacus Fleutiaux, 1934
Athous parallelopipedus Brullé, 1832
Athous roralis Gistel, 1857
Athous rufithorax Miwa, 1930
Athous shirozui Ôhira, 1966
Athous strictus (Fischer von Waldheim, 1824)
Athous temperatus Miwa, 1930
Athous weigeli Schimmel, 1998


=== Liste over ikke-europeiske, uplasserte arter ===
Athous acanthus (Say, 1839)
Athous agriotoides Fall, 1907
Athous appalachius Van Dyke, 1932
Athous arizonicus Van Dyke, 1932
Athous aterrimus Fall, 1910
Athous axillaris Horn, 1871
Athous bedeli Fleutiaux, 1928
Athous birmanicus Fleutiaux, 1942
Athous brevis Fleutiaus, 1928
Athous brightwelli (Kirby, 1837)
Athous chapaensis Fleutiaux, 1928
Athous coomani Fleutiaux, 1928
Athous coquerelli Reitter, 1908
Athous cribratus LeConte, 1876
Athous cucullatus (Say, 1825)
Athous demangei Fleutiaux, 1918
Athous diplogrammus Orlov, 1994
Athous discrepans Reitter, 1908
Athous ebeninus Fleutiaux, 1918
Athous equestris (LeConte, 1853)
Athous essigi Van Dyke, 1932
Athous excavatus (Motschulsky, 1859)
Athous farallonicus Van Dyke, 1951
Athous fossularis (LeConte, 1853)
Athous fueguensis Golbach & Aranda, 1991
Athous funestus Champion, 1896
Athous imitans Fall, 1910
Athous lecontei (Candèze, 1889)
Athous limbatus LeConte, 1866
Athous malaisei Fleutiaux, 1942
Athous massiei Fleutiaux, 1928
Athous minutus Fleutiaux, 1928
Athous neacanthus Becker, 1974
Athous nigropilis Motschulsky, 1859
Athous nodieri Fleutiaux, 1918
Athous olgae Iablokov-Khnzorian, 1961
Athous opilinus Candèze, 1860
Athous ornatipennis (LeConte, 1863)
Athous orvus Becker, 1974
Athous paradisus Knull, 1934
Athous perroti Fleutiaux, 1940
Athous phylander Tottenham, 1948
Athous polygenus (Fall, 1910)
Athous posticus (Melsheimer, 1844)
Athous productus (Randall, 1838)
Athous rufifrons (Randall, 1838)
Athous rufipennis Van Dyke, 1932
Athous rufotestaceous Fall, 1907
Athous sanguinicollis Frivaldsky, 1892
Athous scapularis (Say, 1839)
Athous scissus LeConte, 1857
Athous striatus Fleutiaux, 1940
Athous vanmeeri Chassain, 2007
Athous vitalisi Fleutiaux, 1918
Athous vittiger LeConte, 1853


== Referanser ==


== Kilder ==
Silfverberg, H. 1992. Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae (Liste over Nordens biller). Helsinki.
Zahradník, J. og Chvála, M. 1991. Teknologisk forlags store bok om insekter. N.W. Damm. Om smellere, side 282-287.


== Eksterne lenker ==
Victor Hanssen (1966). Biller 23. Smældere og pragtbiller (dansk). 74 (1 utg.). København: G. E. C. Gads forlag. s. 53–61.  [Bestemmelsesnøkler]
Bilder av mange typeeksemplarer i slekten
(en) Athous i Encyclopedia of Life 
(en) Athous i Global Biodiversity Information Facility 
(no) Athous hos Artsdatabanken 
(sv) Athous hos Dyntaxa 
(en) Athous hos Fauna Europaea 
(en) Athous hos Fossilworks 
(en) Athous hos NCBI 
(en) Kategori:Athous – bilder, video eller lyd på Wikimedia Commons 
Athous – detaljert informasjon på Wikispecies