Drusilla (bille)

Drusilla er en slekt av biller som hører til den store underfamilien småkortvinger (Aleocharinae) i familien kortvinger (Staphylinidae). De er spesialiserte på å leve sammen med maur.


== Utseende ==
Små til middelsstore (gjerne 3-5 millimeter), brunlige eller rødlige kortvinger, kroppen lite hårete bortsett fra hårpenslene på bakkroppen. Antennene er forholdsvis lange og kraftige. Hodet er forholdsvis lite og rundt. Pronotum er lengre enn bredt, gjerne med en bred midtfure. Dekkvingene er forholdsvis korte. Beina er forholdsvis lange og tynne.


== Levevis ==
Drusilla-artene har spesialisert seg på å leve sammen med maur. Disse artene lever rundt maurtuene  der de dels jakter på maurene, dels eter maurenes matrester.


== Systematisk inndeling / norske arter ==

Ordenen biller, Coleoptera
Underordenen Polyphaga
Overfamilien åtsel- og rovbiller, Staphylinoidea
Familien kortvinger (Staphylinidae) Latreille, 1802
Underfamilien småkortvinger (Aleocharinae) Fleming, 1821
Stammen Lomechusini
Slekten Drusilla Leach, 1819 (synonym Astilbus Dillwyn, 1829)
Drusilla adjacens (Cameron, 1939)
Drusilla aeneicollis (Fauvel, 1899)
Drusilla aequatorialis (Bernhauer, 1927)
Drusilla aerea (Scheerpeltz, 1934)
Drusilla affinis (Cameron, 1939)
Drusilla aino Nakane, 1963
Drusilla alutacea Reitter, 1901
Drusilla anceps Assing, 2005
Drusilla andrewesi (Cameron, 1939)
Drusilla angularis (Fauvel, 1899)
Drusilla ashei Eldredge, 2012
Drusilla assamensis (Cameron, 1939)
Drusilla attenuata Kistner, 1997
Drusilla baliensis Pace, 1987
Drusilla bartolozzii Pace, 2003
Drusilla bernhaueri (Scheerpeltz, 1934)
Drusilla besucheti Assing, 2005
Drusilla bicarinicollis Pace, 2008
Drusilla bicolor (Cameron, 1950)
Drusilla bifida Assing, 2016
Drusilla bilobata Pace, 2014
Drusilla bispinosa Assing, 2019
Drusilla bituberculata (Motschulsky, 1858)
Drusilla borneensis (Bernhauer, 1915)
Drusilla borneoapicalis Pace, 2014
Drusilla borneoclara Pace, 2014
Drusilla borneoruficollis Pace, 2014
Drusilla borneostricta Pace, 2014
Drusilla brevipennis (Cameron, 1939)
Drusilla breviuter Assing, 2019
Drusilla bruneiensis Pace, 2008
Drusilla bruneiorum Pace, 2008
Drusilla bryanti (Cameron, 1950)
Drusilla bulbata Assing, 2008
Drusilla bulbosa Pace, 2014
Drusilla burgeoniana (Bernhauer, 1943)
Drusilla butuanensis (Bernhauer, 1916)
Drusilla caelata Pace, 1987
Drusilla caelaticollis Pace, 2008
Drusilla calicis Pace, 2012
Drusilla cameroni (Scheerpeltz, 1934)
Drusilla canaliculata (Fabricius, 1787) - eneste art i Norge
Drusilla caputserpentis Pace, 2014
Drusilla carinithorax (Bernhauer, 1915)
Drusilla celebensis (Cameron, 1942)
Drusilla cenwanglaomontis Pace, 2012
Drusilla cernens Assing, 2005
Drusilla congoensis (Kistner, 2003)
Drusilla convexifrons (Fairmaire, 1873)
Drusilla cretica Assing, 2005
Drusilla cupulifer (Fauvel, 1906)
Drusilla damingensis Pace, 2012
Drusilla danumensis Pace, 2014
Drusilla dendrolasii Maruyama & Kishimoto, 2002
Drusilla denigrata Assing, 2005
Drusilla denticollis (Bernhauer, 1933)
Drusilla dharmapuriensis Pace, 2001
Drusilla dimidiata Pace, 1987
Drusilla divergens Pace, 2014
Drusilla dorylophila (Bernhauer, 1929)
Drusilla drescheri (Cameron, 1939)
Drusilla dumogaicola Pace, 2004
Drusilla dusunorum Pace, 1993
Drusilla endopilosella Pace, 2000
Drusilla endorica (Saulcy, 1865)
Drusilla erichsoni (Peyron, 1857)
Drusilla feae (Cameron, 1939)
Drusilla fissata (Bernhauer, 1915)
Drusilla flagellata Assing, 2015
Drusilla flava (Cameron, 1939)
Drusilla flavicornis (Motschulsky, 1858)
Drusilla foeda Pace, 2008
Drusilla fontis Pace, 2008
Drusilla foreli (Wasmann, 1890)
Drusilla fossulicollis Pace, 2008
Drusilla fossulifera Pace, 1987
Drusilla frater (Cameron, 1950)
Drusilla gabonensis Pace, 2012
Drusilla ganglbaueri (Bernhauer, 1906)
Drusilla gibberella Pace, 1998
Drusilla gilva Pace, 2012
Drusilla gracilicornis (Bernhauer, 1902)
Drusilla gracilis (Hochhuth, 1849)
Drusilla granulata (Bernhauer, 1927)
Drusilla granulicollis (Cameron, 1939)
Drusilla guineensis (Bernhauer, 1932)
Drusilla haematica (Fauvel, 1899)
Drusilla heydeni (Eppelsheim, 1887)
Drusilla hipponensis (Fauvel, 1898)
Drusilla impressicollis (Kraatz, 1857)
Drusilla imurai Maruyama & Kishimoto, 2002
Drusilla incola (Fauvel, 1904)
Drusilla inflatae Maruyama, Yek, Hashim & Ito, 2003
Drusilla intermedia (Cameron, 1933)
Drusilla italica (Bernhauer, 1903)
Drusilla jacobsoni (Cameron, 1936)
Drusilla javana (Cameron, 1939)
Drusilla kabyliana (Fagel, 1958)
Drusilla kadooriorum Pace, 1998
Drusilla kamerunensis (Bernhauer, 1912)
Drusilla kathmanduensis Pace, 1992
Drusilla katiki Pace, 1993
Drusilla keiichii Maruyama, 2004
Drusilla khamhengi Pace, 1986
Drusilla kinabaluensis Pace, 1987
Drusilla kisumuensis Pace, 1996
Drusilla laevicauda (Bernhauer, 1903)
Drusilla lateremaculata Pace, 2016
Drusilla lativentris Assing, 2017
Drusilla lethierryi (Fauvel, 1898)
Drusilla liangicola Pace, 1993
Drusilla limata Assing, 2005
Drusilla lineatocollis (Kraatz, 1859)
Drusilla longatheca Pace, 2004
Drusilla lucens (Cameron, 1950)
Drusilla lucida (Cameron, 1950)
Drusilla luzonica (Bernhauer, 1915)
Drusilla lydica Assing, 2007
Drusilla magniceps (Cameron, 1939)
Drusilla mahnerti Pace, 1996
Drusilla malaisei (Scheerpeltz, 1965)
Drusilla malindensis Pace, 1991
Drusilla manenekarum Pace, 1993
Drusilla manubrifera Pace, 2010
Drusilla memnonia (Märkel, 1844)
Drusilla meridiana (Fauvel, 1900)
Drusilla mixta (Cameron, 1939)
Drusilla montanella (Bernhauer, 1936)
Drusilla monticola (Cameron, 1943)
Drusilla nearctica Eldredge, 2012
Drusilla necaerea Pace, 2008
Drusilla nepalensis Pace, 1992
Drusilla nigra (Cameron, 1925)
Drusilla nigrescens (Motschulsky, 1858)
Drusilla nomurai Maruyama, 2000
Drusilla obliqua (Bernhauer, 1916)
Drusilla obliquoides Pace, 2003
Drusilla occipitalis Pace, 1987
Drusilla ocularis (Fenyes, 1914)
Drusilla opacicollis (Cameron, 1920)
Drusilla opacipennis (Bernhauer, 1902)
Drusilla operosa Pace, 1987
Drusilla orientalis (Cameron, 1939)
Drusilla orissaensis Pace, 2011
Drusilla pallidicornis Assing, 2005
Drusilla papuana (Cameron, 1937)
Drusilla pendleburyi (Cameron, 1950)
Drusilla penicillata  Assing, 2019
Drusilla perakensis Pace, 2013
Drusilla perdensa Pace, 2004
Drusilla perforans Assing, 2015
Drusilla persica Assing, 2005
Drusilla persulcata Pace, 2004
Drusilla pervisa Pace, 1987
Drusilla philippina (Bernhauer, 1915)
Drusilla planaticollis (Kraatz, 1857)
Drusilla plena Pace, 2004
Drusilla plicipennis (Bernhauer, 1915)
Drusilla profunda Pace, 2014
Drusilla proxima (Cameron, 1939)
Drusilla pudukkottaiensis Pace, 2001
Drusilla puetzi Assing, 2011
Drusilla punctatissima (Kraatz, 1857)
Drusilla raffrayi (Fairmaire, 1871)
Drusilla rantepaoicola Pace, 1993
Drusilla recta Assing, 2005
Drusilla refugita Pace, 2000
Drusilla rougemonti Pace, 1987
Drusilla rougemontiana Pace, 2014
Drusilla ruficornis (Cameron, 1939)
Drusilla rufiventris (Cameron, 1950)
Drusilla rufobrunnea (Cameron, 1939)
Drusilla rufocaelata Pace, 2008
Drusilla rufoflava (Cameron, 1950)
Drusilla sabahensis Pace, 2014
Drusilla sabahorum Pace, 2014
Drusilla schawalleri Kistner, 1994
Drusilla sculpticollides Hlavàč, Newton & Maruyama, 2011
Drusilla semicircularis (Bernhauer, 1929)
Drusilla semicyanea (Cameron, 1930)
Drusilla semimonticola Pace, 2008
Drusilla semiobliqua Pace, 2005
Drusilla semipunctata (Bernhauer, 1929)
Drusilla semirubra (Cameron, 1950)
Drusilla serrulae Pace, 2014
Drusilla shavrini  Assing, 2019
Drusilla singalorum Pace, 1988
Drusilla sinicola Pace, 2012
Drusilla sinuosa Assing, 2005
Drusilla smetanai Pace, 1992
Drusilla sparsa (Sharp, 1874)
Drusilla spiniventris  Assing, 2019
Drusilla spissatheca Pace, 2008
Drusilla stevensi (Cameron, 1939)
Drusilla strigicollis (Cameron, 1928)
Drusilla subsplendens Assing, 2011
Drusilla suturalis (Erichson, 1843)
Drusilla takashii Maruyama, 2004
Drusilla taygetana Assing, 2005
Drusilla terrestris Pace, 2008
Drusilla thai Pace, 2012
Drusilla thaifuscicollis Pace, 2012
Drusilla theryi (Fauvel, 1898)
Drusilla tigrata Pace, 1992
Drusilla todolarum Pace, 1993
Drusilla trigonocephala (Cameron, 1939)
Drusilla trina Pace, 2014
Drusilla tronqueti Pace, 1988
Drusilla tuberculata (Bernhauer, 1916)
Drusilla ulusegamicola Pace, 2015
Drusilla umranicola Pace, 2005
Drusilla umtaliensis Pace, 1996
Drusilla unicolor (Cameron, 1950)
Drusilla veluticollis (Cameron, 1928)
Drusilla villosa (Peyerimhoff, 1909)
Drusilla wallacei (Kistner & McNairn, 1991)
Drusilla watanabei Maruyama & Kishimoto, 2002
Drusilla yunnanensis Pace, 1993
Drusilla zhejiangensis Pace, 1998
Drusilla zimbabwensis Pace, 1996
Drusilla zyrasoides Dvořàk, 1988


== Referanser ==

Silfverberg, H. 1992. Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae (Liste over Nordens biller). Helsinki.


== Eksterne lenker ==
(en) Drusilla (bille) i Encyclopedia of Life 
(en) Drusilla (bille) i Global Biodiversity Information Facility 
(no) Drusilla (bille) hos Artsdatabanken 
(sv) Drusilla (bille) hos Dyntaxa 
(en) Drusilla (bille) hos ITIS 
(en) Drusilla (bille) hos NCBI 
(en) Kategori:Drusilla (Staphylinidae) – bilder, video eller lyd på Wikimedia Commons 
Drusilla – detaljert informasjon på Wikispecies