Ataenius

Ataenius er en artsrik slekt av biller som hører til familien skarabider (Scarabaeidae).


== Utseende ==
Små (1,5 -7 millimeter), avlange, blanke, mørke gjødselbiller. Hodet er stort og bredt, noe hvelvet, gjerne innbuktet foran.


== Levevis ==
Disse billene har larver som utvikler seg i jorda, der de etter på gressrøtter. De kan gjøre en del skade på golfbaner og lignende 


== Utbredelse ==
Slekten er utbredt i alle verdensdeler bortsett fra Antarktis.


== Systematisk inndeling ==

Ordenen biller, Coleoptera
Underordenen Polyphaga
Overfamilien skarabider, Scarabaeoidea
Familien skarabider, (Scarabaeidae) Latreille, 1806
Underfamilien Gjødselbiller, Aphodiinae Leach, 1815
Stammen Eupariini Schmidt, 1910
Slekten Ataenius Harold, 1867


=== Liste over artene ===
Ataenius abancay Stebnicka, 2005
Ataenius abditus (Haldeman, 1848)
Ataenius abdominalis Petrovitz, 1973
Ataenius aciculus Hinton, 1937
Ataenius acutulus Schmidt, 1913
Ataenius aequalis Harold, 1880
Ataenius aequatorialis Petrovitz, 1961
Ataenius africanus Endrödi, 1967
Ataenius alternatus (Melsheimer, 1845)
Ataenius ambaritae Stebnicka, 1988
Ataenius angulatus Petrovitz, 1964
Ataenius annanus Stebnicka & Howden, 1997
Ataenius aphodiiformis Bordat & Théry, 2012
Ataenius apicalis Hinton, 1937
Ataenius arenosus Harold, 1868
Ataenius argillaceus Schmidt, 1916
Ataenius arrowi Hinton, 1936
Ataenius asper Petrovitz, 1975
Ataenius atramentarius (Erichson, 1847)
Ataenius attenuator Harold, 1874
Ataenius australasiae Boheman, 1858
Ataenius australis Harold, 1875
Ataenius balthasari Petrovitz, 1973
Ataenius barberi Cartwright, 1974
Ataenius basiceps Lea, 1923
Ataenius beattyi Chapin, 1940
Ataenius benarabyensis Stebnicka & Howden, 1997
Ataenius bicolor Petrovitz, 1963
Ataenius biroi Endrödi, 1951
Ataenius bispinulosus Schmidt, 1911
Ataenius blapoides Balthasar, 1947
Ataenius bolivarensis Stebnicka, 2007
Ataenius bolivari Stebnicka, 2001
Ataenius borgmeieri Hinton, 1936
Ataenius borjae Stebnicka, 2005
Ataenius brevicollis (Wollaston, 1854)
Ataenius brevis Fall, 1930
Ataenius brevitarsis Petrovitz, 1964
Ataenius brouni Sharp, 1876
Ataenius brunneus Schmidt, 1922
Ataenius buenavistae Stebnicka, 2001
Ataenius caesoides Fairmaire, 1899
Ataenius caicarae Stebnicka, 2005
Ataenius californicus Horn, 1887
Ataenius canoasus Stebnicka, 2007
Ataenius carinatipennis Petrovitz, 1973
Ataenius carinator Harold, 1874
Ataenius cartago Stebnicka, 2007
Ataenius cartwrighti Chalumeau & Gruner, 1974
Ataenius castaniellus Bates, 1887
Ataenius catarinaensis Stebnicka, 2007
Ataenius catenulatus (Erichson, 1847)
Ataenius cavagnaroi Cartwright, 1970
Ataenius ceylonensis Schmidt, 1912
Ataenius chapini Hinton, 1937
Ataenius chilensis (Solier, 1851)
Ataenius chinacotae Stebnicka, 2007
Ataenius circulusbrumalis Petrovitz, 1964
Ataenius clavatus Schmidt, 1916
Ataenius clitellarius Petrovitz, 1973
Ataenius cochabambae Stebnicka, 2005
Ataenius cognatus (LeConte, 1858)
Ataenius columbicus Harold, 1880
Ataenius communis Hinton, 1936
Ataenius complicatus Harold, 1869
Ataenius confertus Fall, 1909
Ataenius convexus Robinson, 1940
Ataenius coriarius Schmidt, 1912
Ataenius corporaali Boucomont, 1924
Ataenius corrosus Chapin, 1940
Ataenius costulifer Balthasar, 1951
Ataenius crenaticollis Petrovitz, 1973
Ataenius crenatipennis MacLeay, 1871
Ataenius crenator Harold, 1876
Ataenius crenatostriatus (Blanchard, 1851)
Ataenius crenulatus Schmidt, 1910
Ataenius cribrithorax Bates, 1887
Ataenius cristobalensis Cook & Peck, 2000
Ataenius cucutae Stebnicka, 2007
Ataenius cylindrus Horn, 1871
Ataenius dentatus Koshantshikov, 1916
Ataenius depilis Petrovitz, 1976
Ataenius deserti Blackburn, 1894
Ataenius desertus Horn, 1871
Ataenius ecruensis Stebnicka & Lago, 2005
Ataenius edungalbae Stebnicka & Howden, 1997
Ataenius elegans Harold, 1868
Ataenius elisaensis Stebnicka, 2002
Ataenius elongatulus MacLeay, 1888
Ataenius elongatus (Palisot de Beauvois, 1811)
Ataenius endroedyyoungai Endrödi, 1973
Ataenius erinaceus Petrovitz, 1961
Ataenius eringundae Stebnicka & Howden, 1997
Ataenius erratus Fall, 1930
Ataenius eurynotus Lea, 1923
Ataenius excisicollis Petrovitz, 1961
Ataenius exiguus Brown, 1932
Ataenius fattigi Cartwright, 1948
Ataenius floreanae Cook & Peck, 2000
Ataenius forgiei Bordat & Théry, 2012
Ataenius forsteri Balthasar, 1960
Ataenius freyi Petrovitz, 1961
Ataenius gammonensis Stebnicka & Howden, 1997
Ataenius garamas Peyerimhoff, 1929
Ataenius gascoyneensis Stebnicka & Howden, 1997
Ataenius gibbus Blackburn, 1904
Ataenius gilesi Stebnicka & Howden, 1997
Ataenius glabriventris Schmidt, 1911
Ataenius glaseri Cartwright, 1974
Ataenius gracilis (Melsheimer, 1845)
Ataenius granchacoensis Balthasar, 1939
Ataenius granocostatus Schmidt, 1912
Ataenius griffini Cartwright, 1974
Ataenius gruneri Chalumeau, 1979
Ataenius guanacastae Stebnicka, 2005
Ataenius guayasi Stebnicka, 2001
Ataenius gungareei Stebnicka & Howden, 1997
Ataenius guriensis Stebnicka, 2005
Ataenius heinekeni (Wollaston, 1854)
Ataenius hesperius Cartwright, 1974
Ataenius hirsutus Horn, 1871
Ataenius hispaniolae Chalumeau, 1982
Ataenius hispidus Harold, 1867
Ataenius holopubescens Hinton, 1938
Ataenius horticola Harold, 1869
Ataenius howdeni Chalumeau, 1978
Ataenius hrubanti Balthasar, 1967
Ataenius huanus Stebnicka, 2007
Ataenius humptydooensis Stebnicka & Howden, 1997
Ataenius hygrophilus Paulian, 1947
Ataenius icanus Balthasar, 1941
Ataenius illaetabilis Lea, 1923
Ataenius imbricatoides Schmidt, 1909
Ataenius imbricatus (Melsheimer, 1845)
Ataenius imparilis Blackburn, 1904
Ataenius impiger Schmidt, 1916
Ataenius impressus (Petrovitz, 1963)
Ataenius incanus Balthasar, 1951
Ataenius indutus Petrovitz, 1961
Ataenius inquisitus Horn, 1887
Ataenius insculptus Horn, 1887
Ataenius insolitus Schmidt, 1909
Ataenius insulae Chalumeau & Gruner, 1974
Ataenius insularis Lea, 1923
Ataenius integer Harold, 1868
Ataenius integricollis Lea, 1923
Ataenius intermedius Bates, 1887
Ataenius iquitosae Stebnicka, 2007
Ataenius isabelae Franz, 1985
Ataenius jamaicensis Chapin, 1940
Ataenius jardinensis Stebnicka, 2002
Ataenius jelineki Chalumeau, 1982
Ataenius kapalgaensis Stebnicka & Howden, 1997
Ataenius klapperichi Howden, 1978
Ataenius kochi Balthasar, 1941
Ataenius koebelei Blackburn, 1904
Ataenius koelleri Balthasar, 1963
Ataenius koghianus Paulian, 1991
Ataenius lamarensis Stebnicka, 2007
Ataenius lamotteiroyi Bordat, 1992
Ataenius lanei Petrovitz, 1973
Ataenius languidus Schmidt, 1911
Ataenius latus Petrovitz, 1963
Ataenius lenkoi Petrovitz, 1973
Ataenius liogaster Bates, 1887
Ataenius lobatus Horn, 1871
Ataenius londrinae Stebnicka, 2007
Ataenius longiclavus Petrovitz, 1970
Ataenius lucanus Horn, 1871
Ataenius luctuosus (Burmeister, 1877)
Ataenius luteomargo Chapin, 1940
Ataenius madagassicus Bordat, 1990
Ataenius maghribinicus Baraud, 1985
Ataenius martinezi Petrovitz, 1973
Ataenius miamii Cartwright, 1934
Ataenius michelii Chalumeau, 1978
Ataenius microtrichopterus Lea, 1923
Ataenius millstreamae Stebnicka & Howden, 1997
Ataenius montanus Schmidt, 1911
Ataenius monteithi Paulian, 1991
Ataenius monticola Paulian, 1940
Ataenius montreuili Bordat & Théry, 2012
Ataenius morator Harold, 1869
Ataenius murchisoni Stebnicka & Howden, 1997
Ataenius nakpandurii Endrödi, 1973
Ataenius nanus (DeGeer, 1774)
Ataenius napoensis Stebnicka, 2007
Ataenius nigricans Paulian, 1933
Ataenius noques Stebnicka, 2007
Ataenius noronhai Stebnicka, 2007
Ataenius nudus Blackburn, 1904
Ataenius nugator Harold, 1880
Ataenius oaxacaensis Stebnicka & Lago, 2005
Ataenius ocumarensis Stebnicka, 2001
Ataenius oklahomensis Brown, 1930
Ataenius onkonensis Stebnicka, 2005
Ataenius opacipennis Schmidt, 1910
Ataenius opatrinus Harold, 1867
Ataenius opatroides (Balthasar, 1947)
Ataenius ovatulus Horn, 1871
Ataenius pacificus (Sharp, 1879)
Ataenius palmaritoensis Stebnicka, 2007
Ataenius palmerstoni Blackburn, 1891
Ataenius palustris (Montrouzier, 1860)
Ataenius panamensis Hinton, 1936
Ataenius parallelipennis Petrovitz, 1973
Ataenius parvus Lea, 1923
Ataenius pearlensis Stebnicka, 2006
Ataenius peregianensis Stebnicka & Howden, 1997
Ataenius peregrinator Harold, 1877
Ataenius pereirai Petrovitz, 1970
Ataenius perforatus Harold, 1867
Ataenius pertuga Balthasar, 1961
Ataenius petrovitzi Balthasar, 1960
Ataenius picinus Harold, 1867
Ataenius platensis (Blanchard, 1847)
Ataenius plaumanni Petrovitz, 1973
Ataenius polyglyptus Bates, 1887
Ataenius pseudimparilis Stebnicka & Howden, 1997
Ataenius pseudocarinator Balthasar, 1947
Ataenius pseudoclavatus Stebnicka, 2005
Ataenius pseudocommunis Stebnicka, 2001
Ataenius pseudostercorator Stebnicka, 2003
Ataenius pseudousingeri Galante, Stebnicka & Verdú, 2003
Ataenius punctatohirsutus Schmidt, 1909
Ataenius puncticollis LeConte, 1868
Ataenius punctipennis Harold, 1868
Ataenius purator Harold, 1868
Ataenius quintanaroo Stebnicka, 2006
Ataenius raccurti Chalumeau, 1978
Ataenius rakovici Bordat & Théry, 2012
Ataenius raslani Bordat & Théry, 2012
Ataenius raucus Schmidt, 1908
Ataenius robustus Horn, 1871
Ataenius rosinae Balthasar, 1967
Ataenius rubrotessellatus (Blanchard, 1843)
Ataenius sabinoi Cartwright, 1974
Ataenius saltae Stebnicka, 2007
Ataenius santarosae Stebnicka, 2007
Ataenius saramari Cartwright, 1939
Ataenius saulensis Stebnicka, 2006
Ataenius scabrelloides Petrovitz, 1962
Ataenius scabrellus Schmidt, 1909
Ataenius scalptifrons Bates, 1887
Ataenius schmidti Stebnicka, 2003
Ataenius sculptilis Harold, 1868
Ataenius sculptor Harold, 1868
Ataenius scutellaris Harold, 1867
Ataenius seaforthensis Stebnicka & Howden, 1997
Ataenius semicoecus MacLeay, 1888
Ataenius semicornutus MacLeay, 1871
Ataenius semipilosus Van Dyke, 1928
Ataenius setiger Bates, 1887
Ataenius setosus Schmidt, 1909
Ataenius seydeli Endrödi, 1964
Ataenius siminasus Petrovitz, 1973
Ataenius simplicipes Mulsant & Rey, 1870
Ataenius skelleyi Stebnicka, 2007
Ataenius sparsicollis Blackburn, 1904
Ataenius speculator Blackburn, 1891
Ataenius spinipennis Lea, 1923
Ataenius spretulus (Haldeman, 1848)
Ataenius stebnickae Bordat & Théry, 2012
Ataenius steinheili Harold, 1874
Ataenius stephani Cartwright, 1974
Ataenius stercorator (Fabricius, 1775)
Ataenius striatocrenatus Fairmaire, 1889
Ataenius strigatus (Say, 1823)
Ataenius strigicaudus Bates, 1887
Ataenius strigifrons Schmidt, 1920
Ataenius suineata Deloya, 2012
Ataenius synnotensis Stebnicka & Howden, 1997
Ataenius talpoides Stebnicka, 2001
Ataenius tambopatae Stebnicka, 2001
Ataenius tarumensis Stebnicka, 2007
Ataenius temperei Chalumeau & Gruner, 1974
Ataenius terminalis (Chevrolat, 1864)
Ataenius texanus Harold, 1874
Ataenius tindalensis Stebnicka & Howden, 1997
Ataenius tomentosus Petrovitz, 1969
Ataenius torridus Blackburn, 1892
Ataenius tovarensis Stebnicka, 2001
Ataenius tuberculatus Schmidt, 1911
Ataenius uriarrae Stebnicka & Howden, 1997
Ataenius usingeri Hinton, 1937
Ataenius utahensis Cartwright, 1974
Ataenius vandykei Cartwright, 1974
Ataenius variopunctatus Schmidt, 1922
Ataenius versicolor Schmidt, 1916
Ataenius vethianus Schmidt, 1909
Ataenius vinacoensis Stebnicka, 2006
Ataenius walkeri Blackburn, 1904
Ataenius waltherhorni Balthasar, 1938
Ataenius warisensis Stebnicka, 1998
Ataenius wenzelii Horn, 1887
Ataenius windjanae Stebnicka & Howden, 1997
Ataenius yungasus Stebnicka, 2001


== Referanser ==


== Kilder ==
Generic Guide to New World Scarab Beetles - Ataenius [2]
Z. Stebnicka og P.K. Lago (1995) The New World species of Ataenius Harold, 1867. V. Revision of the A. strigatus group (Scarabaeidae: Aphodiinae: Eupariini). Insecta Mundi 19: 55-83 [3]
biolib.cz - Oversikt over stammer og slekter


== Eksterne lenker ==
(en) Ataenius i Encyclopedia of Life 
(en) Ataenius i Global Biodiversity Information Facility 
(en) Ataenius hos Fauna Europaea 
(en) Ataenius hos Fossilworks 
(en) Ataenius hos ITIS 
(en) Ataenius hos NCBI 
(en) Kategori:Ataenius – bilder, video eller lyd på Wikimedia Commons 
Ataenius – detaljert informasjon på Wikispecies