Cyclocephala

Cyclocephala er en artsrik slekt av biller som hører til gruppen hornbiller i familien skarabider (Scarabaeidae). 


== Utseende ==
Middelsstore (7-25 millimeter), avlange, blanke, brungule, rødlige eller mørke biller. Mange arter har mørke fargetegninger, de kan også være ensfarget mørke. De er blant de minste hornbillene og mangler horn. Et særtrekk er at de vifteformede antennene bare har to «vifteblader».


== Levevis ==
De voksne billene er nattaktive og kommer til lys. I alle fall noen arter besøker blomster (for eksempel på planter i myrkonglefamilien) og kan være viktige pollinatorer. Larvene er kjent bare for et fåtall av artene, disse lever i jorda og eter på gressrøtter.


== Utbredelse ==
Artene er utbredt i Nord-Amerika, der de kan være meget tallrike, men de aller fleste lever i Sør- og Mellom-Amerika.


== Systematisk inndeling ==
Avgrensingen av slekten er noe uklar, spesielt er den ofte vanskelig å skille fra slekten Aspidolea.

Ordenen biller, Coleoptera
Underordenen Polyphaga
Overfamilien skarabider (Scarabaeoidea)
Familien skarabider, Scarabaeidae Latreille, 1806
Underfamilien hornbiller, Dynastinae MacLeay, 1819
Stammen Cyclocephalini
Slekten Cyclocephala Latreille, 1829 – nordlige arter:
Cyclocephala borealis Arrow, 1911
Cyclocephala hirta LeConte, 1861
Cyclocephala longula LeConte, 1863
Cyclocephala lunulata Burmeister, 1847
Cyclocephala lurida Bland, 1863
Cyclocephala melanocephala (Fabricius, 1775)
Cyclocephala miamiensis Howden & Endrödi, 1966
Cyclocephala pasadenae (Casey, 1915)
Cyclocephala puberula LeConte, 1863


=== Liste over arter utbredt sør for USA ===
Cyclocephala abrelata Ratcliffe & Cave, 1992
Cyclocephala acoma Ratcliffe, 2008
Cyclocephala acuta Arrow, 1902
Cyclocephala aequatoria Endrödi, 1963
Cyclocephala affinis Endrödi, 1966
Cyclocephala alazonia Ratcliffe, 2003
Cyclocephala alexi Ratcliffe & Delgado, 1990
Cyclocephala almitana Dechambre, 1992
Cyclocephala altamontana Dechambre, 1999
Cyclocephala alutacea Höhne, 1923
Cyclocephala alvarengai Dechambre, 1980
Cyclocephala amazona (Linnaeus, 1767)
Cyclocephala amblyopsis Bates, 1888
Cyclocephala ampliata Bates, 1888
Cyclocephala amplitarsis Ratcliffe, 1992
Cyclocephala angularis (Knoch, 1801)
Cyclocephala anibali Joly, 2009
Cyclocephala antoinei Dupuis, 2018
Cyclocephala aravaipensis Ratcliffe, 1992
Cyclocephala arenosa Howden & Endrödi, 1966
Cyclocephala arnaudi Dechambre, 1980
Cyclocephala arrowiana Martínez, 1967
Cyclocephala atricapilla Mannerheim, 1829
Cyclocephala atriceps (Casey, 1915)
Cyclocephala atricolor Chapin, 1932
Cyclocephala atripes Bates, 1888
Cyclocephala aulustjaorum Hiekema, 2017
Cyclocephala barrerai Martínez, 1969
Cyclocephala batesi Delgado-Castilla & Castaneda, 1994
Cyclocephala bella Endrödi, 1969
Cyclocephala berti Delgado-Castillo, 1992
Cyclocephala bicolor Laporte de Castelnau, 1840
Cyclocephala bicolorata Endrödi, 1964
Cyclocephala bimaculata Dechambre, 1999
Cyclocephala binotata Dechambre, 1999
Cyclocephala bleuzeni Dechambre, 1995
Cyclocephala boliviana Dechambre, 1992
Cyclocephala bollei Dechambre & Endrödi, 1984
Cyclocephala borburatae Endrödi, 1980
Cyclocephala boucheri Dechambre, 1997
Cyclocephala boulardi Dechambre, 1979
Cyclocephala brasiliana Endrödi, 1966
Cyclocephala brevipennis Endrödi, 1985
Cyclocephala brevis Höhne, 1923
Cyclocephala brittoni Endrödi, 1964
Cyclocephala burmeisteri Endrödi, 1964
Cyclocephala caelestis Delgado & Ratcliffe, 1990
Cyclocephala camachicola Ohaus, 1910
Cyclocephala capitata Höhne, 1923
Cyclocephala carbonaria Arrow, 1911
Cyclocephala cardini Chapin, 1935
Cyclocephala carinatipennis Martínez & Morón, 1984
Cyclocephala carlsoni Ratcliffe, 2008
Cyclocephala cartwrighti Endrödi, 1964
Cyclocephala casanova Ratcliffe & Cave, 2009
Cyclocephala castanea (Olivier, 1789)
Cyclocephala castaniella Bates, 1888
Cyclocephala caussaneli Dechambre, 1999
Cyclocephala cearae Höhne, 1923
Cyclocephala celata Dechambre, 1980
Cyclocephala cerea Burmeister, 1847
Cyclocephala chalumeaui Martínez, 1978
Cyclocephala chera Ratcliffe, 2008
Cyclocephala chiquita Ratcliffe, 2008
Cyclocephala colasi Endrödi, 1964
Cyclocephala collaris Burmeister, 1847
Cyclocephala comata Bates, 1888
Cyclocephala compacta Ratcliffe, 2008
Cyclocephala complanata Burmeister, 1847
Cyclocephala concolor Burmeister, 1847
Cyclocephala confusa Endrödi, 1966
Cyclocephala conspicua Sharp, 1877
Cyclocephala contraria Kirsch, 1873
Cyclocephala coriacea Dechambre, 1992
Cyclocephala couturieri Dechambre, 1999
Cyclocephala crassa Endrödi, 1967
Cyclocephala crepuscularis Martínez, 1954
Cyclocephala cribrata Burmeister, 1847
Cyclocephala cubana Chapin, 1932
Cyclocephala cucullata Ratcliffe & Cave, 2020
Cyclocephala curta Bates, 1888
Cyclocephala dalensi Ponchel, 2009
Cyclocephala danforthi Chapin, 1935
Cyclocephala deceptor (Casey, 1915)
Cyclocephala dechambrei Dupuis, 2018
Cyclocephala decorella Endrödi, 1966
Cyclocephala defecta Endrödi, 1970
Cyclocephala deltoides Ratcliffe, 1992
Cyclocephala dichroa Dechambre, 1992
Cyclocephala dicula Dupuis, 2021
Cyclocephala dilatata Prell, 1934
Cyclocephala diluta Erichson, 1847
Cyclocephala discicollis Arrow, 1902
Cyclocephala discolor (Herbst, 1792)
Cyclocephala dispar (Herbst, 1790)
Cyclocephala distincta Burmeister, 1847
Cyclocephala divaricata Joly, 2005
Cyclocephala dolichotarsa Ratcliffe & Cave, 2008
Cyclocephala dominicana Endrödi, 1985
Cyclocephala duodecimpunctata Endrödi, 1966
Cyclocephala dupuisi Ratcliffe, 2014
Cyclocephala durantonorum Dechambre, 1999
Cyclocephala dyscinetoides Dechambre, 1999
Cyclocephala emarginata Endrödi, 1966
Cyclocephala endroedii Martínez, 1965
Cyclocephala endroedy-youngai Endrödi, 1964
Cyclocephala englemani (Ratcliffe, 1977)
Cyclocephala enigma Ratcliffe, 2003
Cyclocephala epistomalis Bates, 1888
Cyclocephala ergastuli Dechambre, 1997
Cyclocephala erotylina Arrow, 1914
Cyclocephala everardoi Grossi, Santos & Almeida, 2016
Cyclocephala falsa (Arrow, 1911)
Cyclocephala fankhaeneli Endrödi, 1964
Cyclocephala fasciolata Bates, 1888
Cyclocephala ferruginea (Fabricius, 1801)
Cyclocephala figurata Burmeister, 1847
Cyclocephala flavipennis Arrow, 1914
Cyclocephala flavoscutellaris Höhne, 1923
Cyclocephala flora Arrow, 1911
Cyclocephala forcipulata Howden & Endrödi, 1966
Cyclocephala forsteri Endrödi, 1963
Cyclocephala freudei Endrödi, 1963
Cyclocephala freyi Endrödi, 1964
Cyclocephala frontalis Chevrolat, 1844
Cyclocephala fulgurata Burmeister, 1847
Cyclocephala fulvipennis Burmeister, 1847
Cyclocephala fusiformis Chapin, 1932
Cyclocephala gabaldoni Martínez & Martínez, 1981
Cyclocephala genieri Joly, 2010
Cyclocephala gigantea Dupuis, 1999
Cyclocephala goetzi Endrödi, 1966
Cyclocephala gravis Bates, 1888
Cyclocephala gregaria Heyne & Taschenberg, 1908
Cyclocephala guaguarum Dechambre & Endrödi, 1984
Cyclocephala guianae Endrödi, 1969
Cyclocephala guttata Bates, 1888
Cyclocephala guycolasi Dechambre, 1992
Cyclocephala halffteriana Martínez, 1968
Cyclocephala hardyi Endrödi, 1975
Cyclocephala hartmannorum Maly, 2006
Cyclocephala helavai Endrödi, 1975
Cyclocephala herteli Endrödi, 1964
Cyclocephala hiekei Endrödi, 1964
Cyclocephala hielkemaorum Ratcliffe, 2008
Cyclocephala hirsuta Höhne, 1923
Cyclocephala histrionica Burmeister, 1847
Cyclocephala holmbergi Martínez, 1968
Cyclocephala howdenannae Endrödi, 1975
Cyclocephala huamilule Romero-López & Morón, 2017
Cyclocephala husingi Endrödi, 1964
Cyclocephala iani Ratcliffe, 1992
Cyclocephala immaculata (Olivier, 1789)
Cyclocephala inca Endrödi, 1966
Cyclocephala insulicola Arrow, 1937
Cyclocephala isabellina Höhne, 1923
Cyclocephala isthmiensis Ratcliffe, 1992
Cyclocephala italoi Dupuis, 1999
Cyclocephala jalapensis Casey, 1915
Cyclocephala jauffreti Dechambre, 1979
Cyclocephala kahanoffae Martínez, 1975
Cyclocephala kaszabi Endrödi, 1964
Cyclocephala kechua (Martínez, 1957)
Cyclocephala knobelae (Brown, 1934)
Cyclocephala krombeini Endrödi, 1979
Cyclocephala kuntzeniana Höhne, 1923
Cyclocephala labidion Ratcliffe, 2003
Cyclocephala lachaumei Dechambre, 1992
Cyclocephala laevis Arrow, 1937
Cyclocephala lamarcki Dechambre, 1999
Cyclocephala laminata Burmeister, 1847
Cyclocephala larssoni Endrödi, 1964
Cyclocephala latericia Höhne, 1923
Cyclocephala latipennis Arrow, 1911
Cyclocephala latreillei Dechambre, 1999
Cyclocephala lecourti Dechambre, 1992
Cyclocephala letiranti Young, 1992
Cyclocephala lichyi Dechambre, 1980
Cyclocephala ligyrina Bates, 1888
Cyclocephala lineata Dupuis, 2008
Cyclocephala lineigera Höhne, 1923
Cyclocephala liomorpha Arrow, 1911
Cyclocephala literata Burmeister, 1847
Cyclocephala lizeri Martínez, 1964
Cyclocephala longa Endrödi, 1963
Cyclocephala longicollis Burmeister, 1847
Cyclocephala longimana Dechambre, 1980
Cyclocephala longitarsis Dechambre, 1999
Cyclocephala lutea Endrödi, 1966
Cyclocephala machadoi Grossi, 2016
Cyclocephala macrophylla Erichson, 1847
Cyclocephala maculata Burmeister, 1847
Cyclocephala maculiventris Höhne, 1923
Cyclocephala mafaffa Burmeister, 1847
Cyclocephala magdalenae Young & Le Tirant, 2005
Cyclocephala malleri Martínez, 1968
Cyclocephala malyi Dupuis, 2014
Cyclocephala mannheimsi Endrödi, 1964
Cyclocephala marginalis Kirsch, 1871
Cyclocephala marginicollis Arrow, 1902
Cyclocephala marianista Dechambre & Endrödi, 1984
Cyclocephala marqueti Dechambre, 1997
Cyclocephala marquezi Delgado & Mora-Aguilar, 2020
Cyclocephala martinezi Endrödi, 1964
Cyclocephala marylizae Ratcliffe, 2003
Cyclocephala mathani Dechambre, 1982
Cyclocephala mechae Martínez
Cyclocephala mecynotarsis Höhne, 1923
Cyclocephala megalophylla Endrödi, 1966
Cyclocephala meinanderi Endrödi, 1964
Cyclocephala melanae Bates, 1888
Cyclocephala melanopoda Ratcliffe, 2008
Cyclocephala melolonthida Ratcliffe & Cave, 2002
Cyclocephala mesophylla Mora-Aguilar & Delgado, 2012
Cyclocephala metrica Steinheil, 1872
Cyclocephala minuchae Joly, 2003
Cyclocephala minuta Burmeister, 1847
Cyclocephala modesta Burmeister, 1847
Cyclocephala molesta Endrödi, 1969
Cyclocephala monacha Ratcliffe, 2008
Cyclocephala monzoni Ratcliffe & Cave, 2009
Cyclocephala moreti Dechambre, 1992
Cyclocephala morphoidina Prell, 1937
Cyclocephala multiplex Casey, 1915
Cyclocephala munda Kirsch, 1870
Cyclocephala mustacha Ratcliffe, 2003
Cyclocephala mutata Harold, 1869
Cyclocephala nana Dechambre, 1999
Cyclocephala nicolasi Dupuis, 2018
Cyclocephala nigerrima Bates, 1888
Cyclocephala nigra (Endrödi, 1979)
Cyclocephala nigricollis Burmeister, 1847
Cyclocephala nigritarsis Ratcliffe, 1992
Cyclocephala nigrobasalis Höhne, 1923
Cyclocephala nigropicta Dechambre & Endrödi, 1980
Cyclocephala niguasa Dechambre & Endrödi, 1984
Cyclocephala nike Ratcliffe, 1992
Cyclocephala nodanotherwon Ratcliffe, 1992 - artsnavnet tyder på en viss frustrasjon over tallet på arter i denne slekten
Cyclocephala notata (Illiger, 1806)
Cyclocephala obscura Endrödi, 1966
Cyclocephala occipitalis Fairmaire, 1892
Cyclocephala ocellata Burmeister, 1847
Cyclocephala ochracea Prell, 1937
Cyclocephala octopunctata Burmeister, 1847
Cyclocephala ohausiana Höhne, 1923
Cyclocephala olivieri Arrow, 1911
Cyclocephala ovulum Bates, 1888
Cyclocephala pan Ratcliffe, 1992
Cyclocephala panthera Dechambre, 1979
Cyclocephala paraflora Martínez, 1978
Cyclocephala paraguayensis Arrow, 1903
Cyclocephala parallela (Casey, 1915)
Cyclocephala pardolocarnoi Dechambre, 1995
Cyclocephala perconfusa Dechambre, 1992
Cyclocephala pereirai (Martínez, 1960)
Cyclocephala perforata Arrow, 1913
Cyclocephala perplexa Ratcliffe, 2008
Cyclocephala peruana Endrödi, 1966
Cyclocephala pichinchana Dechambre, 1992
Cyclocephala picipes (Olivier, 1789)
Cyclocephala picopijola Ratcliffe & Cave, 2006
Cyclocephala picta Burmeister, 1847
Cyclocephala pilosa Dupuis, 2006
Cyclocephala pilosicollis Saylor, 1936
Cyclocephala pokornyi Dupuis, 2014
Cyclocephala pompanoni Dechambre, 1979
Cyclocephala poncheli Dechambre & Duranton, 2005
Cyclocephala porioni Dechambre, 1979
Cyclocephala prelli Endrödi, 1967
Cyclocephala prolongata Arrow, 1902
Cyclocephala proxima Dechambre, 1997
Cyclocephala pseudoconfusa Ratcliffe, 1992
Cyclocephala pseudomelanocephala Dupuis, 1996
Cyclocephala pugnax Arrow, 1914
Cyclocephala pulchra Dechambre, 1999
Cyclocephala puncticollis Endrödi, 1966
Cyclocephala putrida Burmeister, 1847
Cyclocephala pygidialis Joly, 2000
Cyclocephala pygidiata Dupuis, 1999
Cyclocephala quadripunctata Höhne, 1923
Cyclocephala quatuordecimpunctata Mannerheim, 1829
Cyclocephala quercina Burmeister, 1847
Cyclocephala quisqueya Joly, 1998
Cyclocephala rangelana Chapin, 1935
Cyclocephala ratcliffei Endrödi, 1977
Cyclocephala recta Dupuis, 2008
Cyclocephala regularis Casey, 1915
Cyclocephala rogerpauli Moore, Branham & Cave, 2018
Cyclocephala rogezi Dechambre, 1992
Cyclocephala rondoniana Ratcliffe, 1992
Cyclocephala rorulenta Höhne, 1923
Cyclocephala rotundipenis Dupuis, 2009
Cyclocephala rubescens Bates, 1891
Cyclocephala rufa Endrödi, 1967
Cyclocephala rufescens Endrödi, 1967
Cyclocephala ruficollis Burmeister, 1847
Cyclocephala rufonigra Demay, 1838
Cyclocephala rufovaria Arrow, 1911
Cyclocephala rustica (Olivier, 1789)
Cyclocephala saltini Ratcliffe, 2008
Cyclocephala sanguinicollis Burmeister, 1847
Cyclocephala santaritae Ratcliffe, 1992
Cyclocephala sarahae Ratcliffe, 1992
Cyclocephala sardadebiae Dechambre & Duranton, 2005
Cyclocephala sarpedon Ratcliffe, 1992
Cyclocephala scarabaeina (Gyllenhal, 1817)
Cyclocephala schmitzorum Ratcliffe, 1992
Cyclocephala sculpta Dupuis, 2021
Cyclocephala seditiosa LeConte, 1863
Cyclocephala setosa Burmeister, 1847
Cyclocephala sexpunctata Laporte de Castelnau, 1840
Cyclocephala signaticollis Burmeister, 1847
Cyclocephala signatoides Höhne, 1923
Cyclocephala similis Dechambre, 1980
Cyclocephala simillima Dechambre, 1999
Cyclocephala simulatrix Höhne, 1923
Cyclocephala sinaloae Howden & Endrödi, 1966
Cyclocephala sinuosa Höhne, 1923
Cyclocephala sororia Bates, 1888
Cyclocephala spangleri Joly, 2000
Cyclocephala sparsa Arrow, 1902
Cyclocephala spilopyga Erichson, 1847
Cyclocephala stictica Burmeister, 1847
Cyclocephala stockwelli Ratcliffe, 2003
Cyclocephala striata Endrödi, 1963
Cyclocephala subsignata Burmeister, 1847
Cyclocephala supernana Dechambre, 1999
Cyclocephala suturalis Ohaus, 1911
Cyclocephala sylviae Dechambre, 1995
Cyclocephala tarsalis Dechambre, 1979
Cyclocephala testacea Burmeister, 1847
Cyclocephala tetrica Burmeister, 1847
Cyclocephala tidula Dechambre, 1999
Cyclocephala toulgoeti Dechambre, 1992
Cyclocephala tridentata (Fabricius, 1801)
Cyclocephala tronchonii Martínez, 1975
Cyclocephala tucumana Brethes, 1904
Cyclocephala tutilina Burmeister, 1847
Cyclocephala tylifera Höhne, 1923
Cyclocephala unamas Ratcliffe, 2003
Cyclocephala undata (Olivier, 1789)
Cyclocephala unidentata Endrödi, 1980
Cyclocephala variabilis Burmeister, 1847
Cyclocephala varians Burmeister, 1847
Cyclocephala variipenis Dechambre, 1999
Cyclocephala variolosa Burmeister, 1847
Cyclocephala verticalis Burmeister, 1847
Cyclocephala vestita Höhne, 1923
Cyclocephala vidanoi Dechambre, 1992
Cyclocephala vidua Endrödi, 1966
Cyclocephala villosa Blanchard, 1846
Cyclocephala vincentiae Arrow, 1900
Cyclocephala vinosa Arrow, 1937
Cyclocephala virgo Dechambre, 1999
Cyclocephala viridis Dechambre, 1982
Cyclocephala vittoscutellaris Prell, 1937
Cyclocephala wandae Hardy, 1974
Cyclocephala warneri Ratcliffe, 1992
Cyclocephala weidneri Endrödi, 1964
Cyclocephala williami Ratcliffe, 1992
Cyclocephala zodion Ratcliffe, 1992
Cyclocephala zurstrasseni Endrödi, 1964


== Kilder ==
Generic Guide to New World Scarab Beetles - Cyclocephala [1]


== Eksterne lenker ==
(en) Cyclocephala i Encyclopedia of Life 
(en) Cyclocephala i Global Biodiversity Information Facility 
(en) Cyclocephala hos Fossilworks 
(en) Cyclocephala hos ITIS 
(en) Cyclocephala hos NCBI 
(en) Kategori:Cyclocephala – bilder, video eller lyd på Wikimedia Commons 
Cyclocephala – detaljert informasjon på Wikispecies 
Bildegalleri over nordamerikanske arter
Dynastidae du monde- sjekkliste over artene Arkivert 5. august 2013 hos Wayback Machine.