Scarites

Scarites er en slekt av billene i delgruppen (underfamilien) graveløpere (Scaritinae) i familiegruppen løpebiller. 


== Utseende ==
Store (opptil 50 millimeter), langstrakte, blanke, oftest svarte løpebiller. Hodet er stort og mer eller mindre rektangulært, med forstørrede, svært kraftige mandibler (overkjever). Fasettøynene er små og rundede, antennene trådformede og nokså korte. Pronotum er mye bredere enn langt, bakover kraftig innsnevret slik at det danner seg en markert "stilk" mellom pronotum og dekkvingene, scutellum sitter på denne. Dekkvingene er parallellsidige med dype punktstriper. Beina er korte og kraftige.


== Levevis ==
Som andre Scaritinae lever disse billene hovedsakelig underjordisk. De graver godt, og lever som rovdyr.


== Utbredelse ==
Denne artsrike slekten er representert i alle verdensdeler bortsett fra Antarktis, men mangler i Nord-Europa.


== Systematisk inndeling ==

Oversikten har med de europeiske artene.   Systematikken følger Carabidae of the World.

ordenen biller, Coleoptera Linnaeus, 1758
underordenen Adephaga Schellenberg, 1806
overfamilien løpebiller og vannkalver, Caraboidea Latreille, 1802
familien   løpebiller, Carabidae Latreille, 1802
underfamilie graveløpere Scaritinae Bonelli, 1810
stammen Scaritini Bonelli, 1810
slekten Scarites Fabricius 1775 – ikke i Norge
underslekt Orientolobus Dostal, 1996
Scarites (Orientolobus) beesoni (Andrewes, 1929)
Scarites (Orientolobus) birmanicus (Bates, 1892)
Scarites (Orientolobus) lucidus (Chaudoir, 1880)
Scarites (Orientolobus) modestus (Chaudoir, 1880)
Scarites (Orientolobus) sexualis Banninger, 1938
Scarites (Orientolobus) stygicus (Chaudoir, 1880)
Scarites (Orientolobus) vilhenai (Basilewsky, 1955)
Scarites (Orientolobus) wittei (Banninger, 1933)
underslekt Parallelomorphus Motschulsky, 1849
Scarites (Parallelomorphus) acutidens Chaudoir, 1855
Scarites (Parallelomorphus) aterrimus A. Morawitz, 1863
Scarites (Parallelomorphus) cultripalpis Quedenfeldt, 1883
Scarites (Parallelomorphus) discoidalis Banninger, 1938
Scarites (Parallelomorphus) inconspicuus Chaudoir, 1855
Scarites (Parallelomorphus) indus Olivier, 1795
Scarites (Parallelomorphus) laevigatus Fabricius, 1792
Scarites (Parallelomorphus) mandli Jedlicka, 1963
Scarites (Parallelomorphus) nitidulus Klug, 1862
Scarites (Parallelomorphus) punctum Wiedemann, 1823
Scarites (Parallelomorphus) quadripunctatus Dejean, 1825
Scarites (Parallelomorphus) semicircularis W.S. MacLeay, 1825
Scarites (Parallelomorphus) subcylindricus Chaudoir, 1843
Scarites (Parallelomorphus) subnitens Chaudoir, 1855
Scarites (Parallelomorphus) terricola Bonelli, 1813
underslekt Scallophorites Motschulsky, 1858
Scarites (Scallophorites) abbreviatus Dejean, 1825 - finnes på Madeira
Scarites (Scallophorites) afghanus Jedlicka, 1967
Scarites (Scallophorites) basiplicatus Heyden, 1884
Scarites (Scallophorites) bucida (Pallas, 1778) - det sørøstlige Europa
Scarites (Scallophorites) buparius (Förster, 1771) - Sør-Europa, dessuten funnet i Estland
Scarites (Scallophorites) cycloderus Chaudoir, 1880
Scarites (Scallophorites) cyclops Crotch, 1871 - Den iberiske halvøya
Scarites (Scallophorites) guineensis Dejean, 1831
Scarites (Scallophorites) impressus Fabricius, 1801 - Spania og Sardinia
Scarites (Scallophorites) kabakovi Dostal, 1997
Scarites (Scallophorites) klapperichi Banninger, 1956
Scarites (Scallophorites) nitens Andrewes, 1929
Scarites (Scallophorites) saxicola Bonelli, 1813 - Hellas
Scarites (Scallophorites) striatus Dejean, 1825
Scarites (Scallophorites) turkestanicus Heyden, 1884
underslekt Scarites (i snever forstand)
Scarites (Scarites) aestuans Klug, 1855
Scarites (Scarites) angulifrons Chaudoir, 1880
Scarites (Scarites) angustesulcatus Baehr, 2002
Scarites (Scarites) angustus (Chaudoir, 1855)
Scarites (Scarites) anomalus Andrewes, 1930
Scarites (Scarites) anthracinus Dejean, 1831
Scarites (Scarites) atronitens Fairmaire, 1887
Scarites (Scarites) baenningeri Emden, 1932
Scarites (Scarites) barbarus Dejean, 1825
Scarites (Scarites) batesi Andrewes, 1929
Scarites (Scarites) bengalensis Dejean, 1826
Scarites (Scarites) biangulatus Fairmaire, 1898
Scarites (Scarites) bokalensis Banninger, 1932
Scarites (Scarites) bottegoi Banninger, 1937
Scarites (Scarites) boucardi Chaudoir, 1880
Scarites (Scarites) bruchi Banninger, 1932
Scarites (Scarites) cayennensis Dejean, 1825
Scarites (Scarites) ceylonicus Chaudoir, 1880
Scarites (Scarites) colossus Csiki, 1927
Scarites (Scarites) comoricus Alluaud, 1932
Scarites (Scarites) corbetti Andrewes, 1929
Scarites (Scarites) cormoides Andrewes, 1929
Scarites (Scarites) costipennis Peringuey, 1896
Scarites (Scarites) crassus Andrewes, 1929
Scarites (Scarites) cylindriformis Banninger, 1933
Scarites (Scarites) cylindronotus Faldermann, 1836
Scarites (Scarites) defletus Banninger, 1933
Scarites (Scarites) denticulatus Chaudoir, 1880
Scarites (Scarites) derogatus Andrewes, 1929
Scarites (Scarites) distinguendus Chaudoir, 1855
Scarites (Scarites) doguereaui Gory, 1833
Scarites (Scarites) dubiosus Andrewes, 1929
Scarites (Scarites) dyschromus Chaudoir, 1855
Scarites (Scarites) edentatus Banninger, 1932
Scarites (Scarites) emarginatus Herbst, 1806
Scarites (Scarites) epaphius Chaudoir, 1880
Scarites (Scarites) estriatus Fairmaire, 1887
Scarites (Scarites) eurytus Fischer von Waldheim, 1828 - spredt i Sør-Europa, også i Estland
Scarites (Scarites) exaratus Dejean, 1825
Scarites (Scarites) excavatus Kirby, 1818
Scarites (Scarites) fairmairei Banninger, 1933
Scarites (Scarites) fatuus Karsch, 1881
Scarites (Scarites) feanus Banninger, 1937
Scarites (Scarites) ferus Banninger, 1933
Scarites (Scarites) fletcheri Andrewes, 1929
Scarites (Scarites) granulatus Andrewes, 1929
Scarites (Scarites) heterogrammus Perty, 1830
Scarites (Scarites) hypsipus Alluaud, 1917
Scarites (Scarites) illustris Chaudoir, 1880
Scarites (Scarites) inaequalis Fairmaire, 1893
Scarites (Scarites) limitaneus Andrewes, 1932
Scarites (Scarites) linearis Boheman, 1848
Scarites (Scarites) liopterus Chaudoir, 1880
Scarites (Scarites) liostracus Alluaud, 1930
Scarites (Scarites) lissopterus Chaudoir, 1880
Scarites (Scarites) lomaensis Basilewsky, 1972
Scarites (Scarites) longiusculus Chaudoir, 1880
Scarites (Scarites) lubricipennis Minowa, 1932
Scarites (Scarites) lunicollis Banninger, 1933
Scarites (Scarites) madagascariensis Dejean, 1831
Scarites (Scarites) malangensis Quedenfeldt, 1883
Scarites (Scarites) mandarinus Banninger, 1928
Scarites (Scarites) mandibularis Andrewes, 1929
Scarites (Scarites) marinus Nichols, 1986
Scarites (Scarites) mayumbensis Banninger, 1933
Scarites (Scarites) melanarius Dejean, 1831
Scarites (Scarites) migiurtinus G.Muller, 1944
Scarites (Scarites) minowai Habu, 1947
Scarites (Scarites) natalensis Boheman, 1848
Scarites (Scarites) nigrita Boheman, 1848
Scarites (Scarites) nitidiceps Baehr, 2002
Scarites (Scarites) ocalensis Nichols, 1986
Scarites (Scarites) orthomus Chaudoir, 1855
Scarites (Scarites) paraguayensis Banninger, 1928
Scarites (Scarites) parallelus Dejean, 1825
Scarites (Scarites) passaloides Quedenfeldt, 1883
Scarites (Scarites) patroclus Murray, 1857
Scarites (Scarites) patruelis Leconte, 1845
Scarites (Scarites) perplexus Dejean, 1825
Scarites (Scarites) pinguis Andrewes, 1929
Scarites (Scarites) politus Bonelli, 1813
Scarites (Scarites) polyphemus Herbst, 1806
Scarites (Scarites) praedator Chaudoir, 1880
Scarites (Scarites) procerus Dejean, 1825
Scarites (Scarites) productus Banninger, 1933
Scarites (Scarites) quadratus Fabricius, 1801
Scarites (Scarites) quadriceps Chaudoir, 1843
Scarites (Scarites) quadricostis Chaudoir, 1880
Scarites (Scarites) raptor Andrewes, 1932
Scarites (Scarites) reductus Banninger, 1933
Scarites (Scarites) retusus Andrewes, 1929
Scarites (Scarites) richteri Chaudoir, 1847
Scarites (Scarites) rugiceps Wiedemann, 1823
Scarites (Scarites) rugosus Wiedemann, 1821
Scarites (Scarites) salinus Dejean, 1825 - Sørøst-Europa
Scarites (Scarites) scaevus Andrewes, 1929
Scarites (Scarites) selene Schmidt-Gobel, 1846
Scarites (Scarites) senegalensis Dejean, 1825
Scarites (Scarites) similis Chaudoir, 1880
Scarites (Scarites) simogonus Chaudoir, 1880
Scarites (Scarites) stenodes Andrewes, 1929
Scarites (Scarites) stenops Bousquet & Skelley, 2010
Scarites (Scarites) strigifrons Baehr, 2002
Scarites (Scarites) subpatroclus Basilewsky, 1972
Scarites (Scarites) subrugatus Chaudoir, 1880
Scarites (Scarites) subterraneus Fabricius, 1775
Scarites (Scarites) sulcatus Olivier, 1795
Scarites (Scarites) tauropus Andrewes, 1929
Scarites (Scarites) tenebricosus Dejean, 1831
Scarites (Scarites) texanus Chaudoir, 1880
Scarites (Scarites) timorensis Banninger, 1949
Scarites (Scarites) trachydermon Andrewes, 1936
Scarites (Scarites) unicus Minowa, 1932
Scarites (Scarites) urbanus Minowa, 1932
Scarites (Scarites) zambo Steinheil, 1875
underslekt Taeniolobus Chaudoir, 1855
Scarites (Taeniolobus) aberrans Banninger, 1941
Scarites (Taeniolobus) bonariensis (Chaudoir, 1880)
Scarites (Taeniolobus) carinatus Dejean, 1825
Scarites (Taeniolobus) convexipennis Fairmaire, 1868
Scarites (Taeniolobus) convexiusculus (Chaudoir, 1880)
Scarites (Taeniolobus) corvinus Dejean, 1831 (Taeniolobus)
Scarites (Taeniolobus) cubanus (Banninger, 1937)
Scarites (Taeniolobus) deplanatus (Banninger, 1937)
Scarites (Taeniolobus) dubius Banninger, 1941
Scarites (Taeniolobus) ecuadorensis Banninger, 1941
Scarites (Taeniolobus) furcatus Banninger, 1941
Scarites (Taeniolobus) gratus (Chaudoir, 1855)
Scarites (Taeniolobus) guerini (Chaudoir, 1855)
Scarites (Taeniolobus) holcocranius (Chaudoir, 1880)
Scarites (Taeniolobus) interpositus (Banninger, 1933)
Scarites (Taeniolobus) lebasii (Chaudoir, 1855)
Scarites (Taeniolobus) meridionalis Banninger, 1941
Scarites (Taeniolobus) multisetosus Banninger, 1941
Scarites (Taeniolobus) oberthueri Banninger, 1938
Scarites (Taeniolobus) obliteratus Banninger, 1941
Scarites (Taeniolobus) planatus Dejean, 1831
Scarites (Taeniolobus) planiusculus (Chaudoir, 1855)
Scarites (Taeniolobus) pronotalis Banninger, 1941
Scarites (Taeniolobus) reichei (Chaudoir, 1880)
Scarites (Taeniolobus) rugatus (Chaudoir, 1880)
Scarites (Taeniolobus) rugicollis Dejean, 1825
Scarites (Taeniolobus) schubarti Banninger, 1939
Scarites (Taeniolobus) seriepunctatus (Banninger, 1933)
Scarites (Taeniolobus) setosus Banninger, 1941
Scarites (Taeniolobus) silvestris Castelnau, 1835
Scarites (Taeniolobus) spectabilis (Chaudoir, 1880)
Scarites (Taeniolobus) subcostatus (Chaudoir, 1880)
Scarites (Taeniolobus) subsulcatus Dejean, 1831
Scarites (Taeniolobus) sulciceps (Chaudoir, 1855)
Scarites (Taeniolobus) sulcifrons (Chaudoir, 1855)
Scarites (Taeniolobus) thiemei (Banninger, 1933)
Scarites (Taeniolobus) vilcanotanus (Banninger, 1932)
Scarites (Taeniolobus) zikani Banninger, 1941


== Referanser ==


== Eksterne lenker ==
(en) Scarites i Encyclopedia of Life 
(en) Scarites i Global Biodiversity Information Facility 
(en) Scarites hos Fauna Europaea 
(en) Scarites hos ITIS 
(en) Scarites hos NCBI 
(en) Kategori:Scarites – bilder, video eller lyd på Wikimedia Commons 
Scarites – detaljert informasjon på Wikispecies 
Scarites – Database Carabidae of the World – engelskspråklig
Scarites - Fauna Europaea - engelskspråklig
Norsk Entomologisk forening