Stantonia sirena Braet & Quicke 2004, sp. n.

Stantonia sirena sp. n. (figures 5, 14, 19, 27, 52, 53, 55, 56) Etymology. From the name of the area from where the holotype was collected. Type material. HOLOTYPE, ” (INBIO): Costa Rica: Prov. Punt., Est. Sirena, 0–100 m, P. N. Corcovado, March–June 1991 (Malaise trap; L-S-270500, 508300). PARATYPES: Costa Rica: ” and „, topotypic (INBIO). Limon Province: ”, Sector Cerro Cocori, Fca. de E. Rojas, 150 m, E. Rojas, October 1991 (L-N-286000, 567500) (INBIO); 2””, idem but: ‘ August 1991 ’ (INBIO); ”, idem but: ‘ 14 October– 11 December 1992 (Malaise trap)’ (INBIO); ”, idem but: ‘ 10–30 September 1992 ’ (INBIO); ”, idem but: ‘ 12 April–19 May 1992 ’ (INBIO); ”, idem but: ‘ March 1992 ’ (INBIO); „, idem but: ‘ August 1991 ’ (INBIO); „, idem but: ‘ March 1992 (Malaise trap)’ (INBIO); ”, Est. Hitoy Cerere, 100 m, R. Cerere, Res. Biol. Hitoy, Cerere, June 1992 (G. Carballo, L-N-184200, 643300) (INBIO); ”, idem but: ‘ 5–19 March 1992 ’ (INBIO); 2””, idem but: ‘ September 1992 ’ (INBIO); ”, idem but: ‘ 19–29 April 1992 ’ (R. Guzman) (INBIO); „, idem but: ‘ 7–26 January 1992 ’ (INBIO); 2„„, 4 km NE Bribri, September–November 1989 (50 m; P. Hanson) (UWIM); ” and „, idem but: ‘ April–June 1990 ’(UWIM); ”, P. N. Tortuguero, Est. 4-esquinas, June–August 1989 (Solano, 0 m) (UWIM). Puntarenas: 2””, P. N. Corcovado, Estac. Sirena, October–December 1990 (50 m; P. Hanson) (UWIM); 2””, road to Rincon, 24 km W Pan-Amer. Hwy, March–May 1989 (200 m; Hanson and Gauld) (UWIM); ” and 2„„, Golfo Dulce, 10 km W Piedras Blancas, June–August 1989 (100 m; P. Hanson) (UWIM); ” and „, Golfo Dulce, Reserva Forestal Golfo Dulce, 3 km SW Rincon, December 1992 (10 m, primary forest; P. Hanson) (UWIM); ”, idem but: ‘ January 1992 ’ (UWIM); ”, idem but: ‘ April 1993 (Malaise)’ (UWIM); 2”” and „, idem but: ‘ June 1991 ’ (UWIM); 4””, idem but: ‘ October 1991 ’ (UWIM); ”, Golfo Dulce, Reserva Forestal Golfo Dulce, 5 km W Piedras Blancas, November–December 1992 (100 m; P. Hanson) (UWIM); ”, idem but: ‘ 15 km W. Piedras Blancas, November 1990 ’ (UWIM); ”, idem but: ‘ 24 km W Piedras Blancas, June–August 1989 (200 m)’ (UWIM); 4„„, idem but: ‘ July 1992 (P. Hanson)’ (TAMU); ” and „, idem but: ‘ December 1992 ’ (TAMU); ”, idem but: ‘ November 1990 (Malaise, primary forest)’ (UWIM); ” and „, idem but: ‘ February 1993 ’ (UWIM); 2„„, idem but: ‘ March 1993 ’ (UWIM). Panama (AEI): Darien: ”, 1967 (C. A. Triplehorn). Barro Colo. [rado] Is. [land] (USNM): ” and „, CZ September 1947 – January 1948, zetek 5328 (Berl[ese] Funnel). Holotype, ”, length of body 4.4 mm, of fore wing 3.7 mm. Head. Antenna with 39 segments, 1.37 times longer than fore wing, length of third segment 1.5 times fourth, length of third, fourth and penultimate segments 3, 2 and 1 times their maximal width, respectively, length of apical segment twice its maximal width; maxillary palp 1.43 times height of head; eyes setose and protuberant in frontal view; length of eye in dorsal view 2.4 times temple; temple directly narrowed posteriorly, punctate, shiny; OOL: diameter of ocellus: POL~ 6.0:3.0:2.0; frons smooth medially, shiny (figure 5); vertex flattened near stemmaticum, smooth and shiny; face shortened, weakly convex (quasi flat) in lateral view, shiny, sparsely punctate with long setae; clypeus weakly convex and smooth; gena in frontal view very short; length of malar space one times basal width of mandible; malar suture absent; occipital flange rounded and very wide (figure 53). Mesosoma. Length of mesosoma 1.26 times its maximal height; side of pronotum smooth posteriorly near tegula and near fore coxa, antero-medially finely crenulate and remainder smooth and shiny, posteriorly setose; mesopleuron smooth and shiny; precoxal sulcus largely punctate and hardly reaching anterior margin of mesopleuron; metapleuron finely punctate, its ventral margin protruding; metapleural flange absent (figure 27); mesoscutum sparsely punctate; scutellar sulcus with small carinae (figure 19); notauli punctate; mesopleuron, scutum, metapleuron, propodeum highly setose; scutellum convex, shiny and setose; propodeum medially convex, smooth, shiny and finely punctate. Wings. Fore wing: r: SR1z3-SR~9:47; 2-SR: r-m~2.0:1.0; second submarginal cell distinctly petiolate (figure 14); subbasal cell setose anteriorly; vein cu-a sub interstitial; vein 3-CU1 twice longer than vein CU1b. Hind wing: 2-M coloured; membrane weakly setose basally. Legs. Hind coxa smooth and shiny; lengths of hind femur, tibia and basitarsus 5, 10 and 7.5 times their widths, respectively; legs setose; tibial spurs 0.53 and 0.3 times basitarsus; hind tibia with three apical teeth. Metasoma. First tergite 3.5 times longer than apically wide, finely coriaceous, flattened in lateral view (figure 55); spiracles weakly protruding (figure 56); second tergite shiny, medially finely punctate, smooth elsewhere; T2 1.3 times longer than T3; second suture smooth and curved (figure 52); ovipositor sheath 0.7 times fore wing length. Colour. Black. Tegula and humeral plate yellowish; maxillary palps, mid coxa, mid trochanter and mid trochantellus, hind trochanter, S1–S3 whitish; dorsal stripe on mid and fore femur, frons, vertex and pronotum dorsally brownish; fore legs (except telotarsi), membrane of fore wing slightly infuscate; antenna, scape and pedicellus, stemmaticum, mesopleuron, mesoscutum, scutellum, propodeum, mesopleuron, tergites, ovipositor sheath, mid tibia (one-third distal), mid tarsus, hind coxa (except basally), hind trochantellus, hind femur (except basal quarter), hind tibia except a small basal annulus, hind tibial spurs, fore telotarsus, hind tarsus blackish. Variation. Upper part of head (frons, vertex and upper part of temples) sometimes brownish to blackish, body sometimes completely blackish except for tegula and whitish yellow hind trochanter, fore and mid coxa. Face sometimes pale yellowish to yellow, basal third of hind coxa and basal fifth of hind femur sometimes yellowish to black. Pronotum sometimes whitish yellow to blackish; S1–S3 are always whitish. Number of antennal segments varies between 37 and 39. Specimens that have a whitish pronotum also have the hind leg with relatively larger white patch. Remarks. This species is closely related to S. scotti, but it may be distinguished by the shorter T1 and the colour pattern of the mesosoma and mid coxa.

Topic: Botany and Geology in Latin America and Caribbean
Subfield: Ecology, Evolution, Behavior and Systematics
Field: Agricultural and Biological Sciences
Domain: Life Sciences