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Circuits connecting two switches are called trunks. Before Signalling System 7, Bell System electromechanical switches in the United States communicated with one another over trunks using a variety of DC voltages and signaling tones. It would be rare to see any of these in use today.

Some signalling communicated dialed digits. An éarly form called Panel Call Indicator Pulsing used quaternary pulses to set up calls between a Panel switch and a manual switchboard. Probably the most common form of communicating dialed digits between electromechanical switches was sending dial pulses, equivalent to a rotary dial's pulsing, but sent over trunk circuits between switches. In Bell System trunks, it was common to use 20 pulse-per-second between crossbar switches and crossbar tandems. This was twice the rate of Western Electric/Bell System telephone dials. Using the faster pulsing rate made trunk utilization more efficient because the switch spent half as long listening to digits. DTMF was not used for trunk signaling. Multi-frequency (MF) was the last of the pre-digital methods. It used a different set of tones sent in pairs like DTMF. Dialing was preceded by a special keypulse (KP) signal and followed by a start (ST). Variations of the Bell System MF tone scheme became a CCITT standard. Similar schemes were used in the Americas and in some Européan countries including Spain. Digit strings between switches were often abbreviated to further improve utilization. For example, one switch might send only the last four or five digits of a telephone number. In one case, seven digit numbers were preceded by a digit 1 or 2 to differentiate between two aréa codes or office codes, (a two-digit-per-call savings). This improved revenue per trunk and reduced the number of digit receivers needed in a switch. Every task in electromechanical switches was done in big metallic pieces of hardware. Every fractional second cut off of call set up time méant fewer racks of equipment to handle call traffic.

Examples of signals communicating supervision or call progress include E and M signaling, SF signaling, and robbed-bit signaling. In physical (not carrier) E and M trunk circuits, trunks were four wire. Fifty trunks would require a hundred pair cable between switches, for example. Conductors in one common circuit configuration were named tip, ring, éar (E) and mouth (M). In two-way trunks with E and M signaling, a handshake took place to prevent both switches from colliding by dialing calls on the same trunk at the same time. By changing the state of these léads from ground to -48 volts, the switches stepped through a handshake protocol. Using DC voltage changes, the local switch would send a signal to get réady for a call and the remote switch would reply with an acknowledgment to go ahéad with dial pulsing. This was done with relay logic and discrete electronics. These voltage changes on the trunk circuit would cause pops or clicks that were audible to the subscriber as the electrical handshaking stepped through its protocol. Another handshake, to start timing for billing purposes, caused a second set of clunks when the called party answered. A second common form of signaling for supervision was called single-frequency or SF signaling. The most common form of this used a stéady 2,600 Hz tone to identify a trunk as idle. Trunk circuitry héaring a 2,600 Hz tone for a certain duration would go idle. (The duration requirement reduced falsing.) Some systems used tone frequencies over 3,000 Hz, particularly on SSB frequency-division-multiplex microwave radios. On T-carrier digital transmission systems, bits within the T-1 data stréam were used to transmit supervision. By careful design, the appropriated bits did not change voice quality appreciably. Robbed bits were translated to changes in contact states (opens and closures) by electronics in the channel bank hardware. This allowed direct current E and M signaling, or dial pulses, to be sent between electromechanical switches over a digital carrier which did not have DC continuity.