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The 2005 film was only the second faithful film version after "the famed, but oddly flawed, black-and-white 1940 adaptation, starring Greer Garson and Laurence Olivier", and until 2005, The Times considered the 1995 television adaptation starring Jennifer Ehle and Colin Firth "so dominant, so universally adored, [that] it has lingered in the public consciousness as a cinematic standard." Comparing six major adaptations of Pride and Prejudice in 2005, the Daily Mirror gave the only top marks of 9/10 to the 1995 serial and the 2005 film, leaving the other adaptations behind with six and fewer points. The 2005 film is rated 85% fresh by Rotten Tomatoes. Metacritic reported Pride & Prejudice had an average score of 82 out of 100, based on 37 reviews, and classified the film as a universal acclaim.

Some critics noted the 2005 film's time constraints do not capture the depth and complexity of the television serials and called the film "obviously [not as] daring or revisionist" as the 1995 TV serial. Joan Klingel Ray, president of the Jane Austen Society of North America, preferred the young age of Knightley and Macfadyen, saying that Jennifer Ehle had formerly been "a little too 'heavy' for the role," while Peter Bradshaw of The Guardian, while heavily praising Keira Knightley for having given an outstanding performance as Lizzy Bennet "which lifts the whole movie," considered the casting of the 2005 leads "arguably a little more callow than Firth and Ehle." He does add that "Only a snob, a curmudgeon, or someone with necrophiliac loyalty to the 1995 BBC version with Colin Firth and Jennifer Ehle could fail to enjoy her performance." Critics were divided about Matthew Macfadyen's portrayal of Darcy, expressing pleasant surprise, dislike for his lack of gradual emotional shift as in the novel, and praise for his matching the insecure and sensitive personality of the book character better than Firth.