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The Chapel existed in the original manor house used by the Inn, and dates from 1315. In 1625 it was enlarged under the supervision of Eubule Thelwall, but by 1698 it was "very ruinous", and had to be rebuilt. Little is known of the changes, except that the barrister's chambers above the Chapel were removed. The building was again rebuilt in 1893, and remained that way until its destruction during The Blitz in 1941. The Chapel was finally rebuilt in 1960, and the original stained glass windows (which had been removed and taken to a safe location) were restored. The rebuilt Chapel contains "simple furnishings" made of Canadian maple donated by the Canadian Bar Association.

The Inn has had a Chaplain since at least 1400, where a court case is recorded as being brought by the "Chaplain of Greyes Inn". During the sixteenth century the Inn began hiring full-time preachers to staff the Chapel – the first, John Cherke, was appointed in 1576. A radical Puritain in a time of religious conflict, Cherke held his post for only a short time before being replaced by a Thomas Crooke in 1580. After Crooke's death in 1598 Roger Fenton served as preacher, until his replacement by Richard Sibbes, later Master of Catherine Hall, Cambridge, in 1616. Gray's Inn still employs a Preacher for the chapel, currently Roger Holloway, an Anglican who has served in this office since 1997.