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Bus services within Eastbourne have been provided by Stagecoach Group since November 2008, when the company acquired the former Eastbourne Buses   and subsequently the independent company Cavendish Motor Services.  The services operate under the name Stagecoach in Eastbourne.

Eastbourne Buses was the main operator of bus services in the borough until November 2008: it was one of the few surviving municipal bus companies. Following complaints about the poor service provided by independent operators, the County Borough of Eastbourne in 1903 became the first local authority in the world authorised to run motor buses. This long history was a source of pride for the council, and the company Eastbourne Buses, which was formed after bus deregulation and remained part-owned by the Borough Council.

As well as local journeys within the town, Stagecoach also runs routes to Polegate, Hailsham, Tunbridge Wells, Uckfield and East Grinstead at various frequencies, while the two routes to Hastings via Bexhill are run by Stagecoach South East from Hastings.  The other main operator into Eastbourne is Brighton & Hove, owned by Go-Ahead Group, which runs frequent services seven days a week from Brighton via Seaford and Newhaven.  Limited numbers of additional buses are run by the Cuckmere Community Bus service, and a regular National Express coach service operates daily from London's Victoria Coach Station.