Jabarkan Pendidikan dari Shalihuddin Djalal Tandjung

S1 Fakultas Biologi, Universitas Gadjah Mada, Yogyakarta (1968)

S2 bidang Ekologi, Dept. of Life Science, Indiana State University, Terre Haute, Amerika Serikat (1977)

S3 bidang Environmental Science and Pollution Ecology, the Louis Calder Conservation and Ecology Study Center, Dept. of Biology, Fordham University, New York, Amerika Serikat (1982)