Given a context passage, generate a question from the passage such that its answer is shortest continous span from the passage.

Example Input: Precursor cells have been shown to be affected by oxidative stress, in vivo and vitro, but little is known about the expression of antioxidant mechanisms in neuronal/glial differentiation. We have characterized the expression of Cu/Zn superoxide dismutase (Cu/Zn SOD), one of the main antioxidant proteins involved in the breakdown of superoxide, in the immature rat dorsolateral subventricular zone (SVZ), rostral migratory stream (RMS) and hippocampal subgranular zone (SGZ). Progenitor cells were identified immunohistochemically on cryostat sections by 5'Bromodeoxyuridine (BrdU) incorporation and expressing cells were further characterized using double labeling for progenitor markers. In the SVZ, only a subpopulation of BrdU+ cells, mostly found in the medial SVZ, expressed Cu/Zn SOD. These cells were mostly nestin+ and some were also vimentin+. In contrast, in the lateral SVZ few Cu/Zn SOD+/BrdU+ cells were found. These were primarily nestin+, vimentin-, showed some PSA-NCAM expression, but only a few were NG2+. In the RMS and SGZ virtually all BrdU+ progenitors were Cu/Zn SOD+ and expressed nestin and vimentin. Some RMS cells were also PSA-NCAM+. These findings show a heterogeneous expression of Cu/Zn SOD in restricted cell types in the germinative zones and suggest a role for antioxidant Cu/Zn SOD in progenitor cells of the immature rat brain.
Example Output: Which intermediate filament (IF) protein can be used as a non-specific marker of the neuronal precursor cells of the subventricular zone?

Example Input: Mosqueiro is an island near the south bank of the Para River in the Brazilian state of Para, roughly north of downtown Belem.
Example Output: What body of water was Mosqueiro next to?

Example Input: The most widely accepted explanation of how the solar system formed is called the nebular hypothesis. According to this hypothesis, the Sun and the planets of our solar system formed about 4.6 billion years ago from the collapse of a giant cloud of gas and dust, called a nebula. The nebula was drawn together by gravity, which released gravitational potential energy. As small particles of dust and gas smashed together to create larger ones, they released kinetic energy. As the nebula collapsed, the gravity at the center increased and the cloud started to spin because of its angular momentum. As it collapsed further, the spinning got faster, much as an ice skater spins faster when he pulls his arms to his sides during a spin. Much of the clouds mass migrated to its center but the rest of the material flattened out in an enormous disk. The disk contained hydrogen and helium, along with heavier elements and even simple organic molecules. As gravity pulled matter into the center of the disk, the density and pressure at the center became intense. When the pressure in the center of the disk was high enough, nuclear fusion began. A star was bornthe Sun. The burning star stopped the disk from collapsing further. Meanwhile, the outer parts of the disk were cooling off. Matter condensed from the cloud and small pieces of dust started clumping together. These clumps collided and combined with other clumps. Larger clumps, called An artists painting of a protoplanetary disk. planetesimals, attracted smaller clumps with their gravity. Gravity at the center of the disk attracted heavier particles, such as rock and metal and lighter particles remained further out in the disk. Eventually, the planetesimals formed protoplanets, which grew to become the planets and moons that we find in our solar system today. Because of the gravitational sorting of material, the inner planets Mercury, Venus, Earth, and Mars formed from dense rock and metal. The outer planets Jupiter, Saturn, Uranus and Neptune condensed farther from the Sun from lighter materials such as hydrogen, helium, water, ammonia, and methane. Out by Jupiter and beyond, where its very cold, these materials form solid particles. The nebular hypothesis was designed to explain some of the basic features of the solar system: The orbits of the planets lie in nearly the same plane with the Sun at the center The planets revolve in the same direction The planets mostly rotate in the same direction The axes of rotation of the planets are mostly nearly perpendicular to the orbital plane The oldest moon rocks are 4.5 billion years Click image to the left or use the URL below. URL:
Example Output:
the inner planets are made primarily of