Given a context passage, generate a question from the passage such that its answer is shortest continous span from the passage.

[Q]: Fly! is a flight simulator for PC and Apple Macintosh computers developed by Terminal Reality and published by Gathering of Developers.
[A]: By whom was Fly! developed?


[Q]: A Vampyre Story is a 2008 point-and-click adventure game developed by Autumn Moon Entertainment for Windows, published by Crimson Cow.
[A]: Which company developed the videogame A Vampyre Story?


[Q]: Viroids, subviral pathogens of plants, are composed of a single-stranded circular RNA of 246-399 nucleotides. Within the 27 viroids sequenced, avocado sunblotch, peach latent mosaic and chrysanthemum chlorotic mottle viroids (ASBVd, PLMVd and CChMVd, respectively) can form hammerhead structures in both of their polarity strands. These ribozymes mediate self-cleavage of the oligomeric RNAs generated in the replication through a rolling circle mechanism, whose two other steps are catalyzed by an RNA polymerase and an RNA ligase. ASBVd, and presumably PLMVd and CChMVd, replicate and accumulate in the chloroplast, whereas typical viroids replicate and accumulate in the nucleus. PLMVd and CChMVd do not adopt a rod-like or quasi rod-like secondary structure as typical viroids do but have a highly branched conformation. A pathogenicity determinant has been mapped in a defined region of the CChMVd molecule.
[A]:
Which are the smallest known subviral pathogens of plants?