Table:
text: Australian Dung Beetle Project
Further dung beetle research
Species of introduced dung beetle that have established in Australia
Species Country_of_origin Total_released First_release Last_release Areas_of_release Areas_established Pasture_type_
Bubas bison France, Spain 1613 April 1983 1996 (not by CSIRO) WA NSW, SA, WA Winter rainfall
Copris elphenor Klug South Africa 2287 January 1977 May 1983 QLD QLD Summer rainfall
Copris hispanus Linnaeus Spain 294 October 1983 June 1994 WA WA Winter rainfall
Euoniticellus africanus Harold South Africa 49009 October 1971 February 1984 NSW, QLD, SA, Tas, Vic, WA NSW, QLD Summer rainfall
Euoniticellus fulvus Goeze France, Turkey 76944 March 1978 February 1983 NSW, SA, Tas, Vic, WA NSW, SA, Tas, Vic, WA Winter rainfall

text: Australian Dung Beetle Project
Further dung beetle research
Species of introduced dung beetle that have established in Australia
Species Country_of_origin Total_released First_release Last_release Areas_of_release Areas_established Pasture_type_
Euoniticellus intermedius Reiche South Africa 248637 November 1971 February 1984 ACT, NSW, NT, QLD, SA, Vic, WA NSW, NT, QLD, SA, Vic, WA Summer rainfall
Euoniticellus pallipes Fabricius Iran, Turkey 46642 March 1977 September 1982 NSW, SA, WA NSW, SA, Vic, WA Winter rainfall
Geotrupes spiniger Marsham France 12082 April 1979 December 1983 ACT, NSW, Tas, Vic ACT, NSW, SA, Tas, Vic Winter rainfall
Liatongus militaris Castelanu South Africa (via Hawaii) 70450 January 1968 November 1979 NSW, NT, QLD, WA NSW, NT, QLD Summer rainfall
Onitis alexis Fabricius South Africa 186441 August 1972 February 1984 NSW, NT, QLD, WA NSW, NT, QLD, SA, Vic, WA Rainfall, summer rainfall, winter rainfall

text: Australian Dung Beetle Project
Further dung beetle research
Species of introduced dung beetle that have established in Australia
Species Country_of_origin Total_released First_release Last_release Areas_of_release Areas_established Pasture_type_
Onitis aygalus Fabricius South Africa 18682 January 1977 January 1982 NSW, SA, WA NSW, SA, Vic, WA Winter rainfall
Onitis caffer Boheman South Africa 8738 October 1979 April 1984 WA QLD, NSW, SA Summer rainfall, winter rainfall
Onitis pecuarius Lansberge South Africa 11395 November 1976 May 1979 NSW, QLD NSW, QLD Summer rainfall
Onitis vanderkelleni Lansberge Kenya, Rwanda, Zaire 10852 October 1974 February 1982 NSW, QLD QLD Summer rainfall
Onitis viridulus Bohemann South Africa 8008 September 1976 July 1980 NSW, NT, QLD NSW, NT, QLD, WA Summer rainfall
Onthophagus binodis Thunberg South Africa 173018 October 1971 February 1982 NSW, QLD, SA, Tas, Vic, WA, Norfolk Islands NSW, QLD, SA, Tas, Vic, WA, Norfolk Islands Winter rainfall

text: Australian Dung Beetle Project
Further dung beetle research
Species of introduced dung beetle that have established in Australia
Species Country_of_origin Total_released First_release Last_release Areas_of_release Areas_established Pasture_type_
Onthophagus gazella Fabricius South Africa 420415 February 1968 February 1984 ACT, NSW, NT, QLD, SA, Tas, Vic, WA, Norfolk Islands ACT, NSW, NT, QLD, SA, WA, Norfolk Islands Summer rainfall
Onthophagus nigiventris d`Orbigny East Africa 29960 May 1975 March 1983 NSW NSW Summer rainfall
Onthophagus obliquus Nigeria, Senegal, Zaire 9300 January 1976 November 1977 QLD, NT QLD Summer rainfall
Onthophagus sagittarius Sri Lanka (via Hawaii) 9075 January 1968 March 1977 NSW, NT, QLD, WA NSW, NT Summer rainfall
Onthophagus taurus Schreber Spain, Greece, Italy, Turkey 164499 February 1975 January 1984 NSW, SA, Tas, Vic, WA NSW, SA, Tas, Vic, WA Winter rainfall
Sisyphus rubrus Paschalidis South Africa 85933 March 1973 February 1980 NSW, QLD, WA NSW, QLD Summer rainfall
Sisyphus spinipes South Africa 36125 March 1972 December 1978 NSW, NT, QLD, WA NSW, QLD Summer rainfall

Question: Which beetle released the largest amount from the Australian Dung Beetle Project?