Question: Where was the author of On the Origin of Species educated?

full_context: Document 1: James Mallet: He became professor of biological diversity at the Department of Biology, University College London. He was co-director of the Centre for Ecology and Evolution, a centre of excellence in research and teaching formed by University College London, the Institute of Zoology (Zoological Society of London), Natural History Museum, Imperial College, Queen Mary, Royal Holloway and Kew Gardens. In 2013 he was distinguished lecturer on Organismic and Evolutionary Biology at Harvard University. His research has included work on the species concept central to evolutionary biology, along with hybridization and the process of speciation.
Document 2: Alfred Nalepa: He studied natural sciences at the University of Vienna, and from 1886 was associated with the "Lehrerbildungsanstalt" in Linz. In 1892 he returned to Vienna, where he was appointed professor of natural history at the "Elisabethgymnasium".
Document 3: Robert Jameson: As Regius Professor of Natural History at the University of Edinburgh for fifty years, Jameson is notable for his advanced scholarship, his superb museum collection and for his tuition of Charles Darwin. Jameson was not at his best in the lecture theatre however, and, for the first half of his career, he grappled with his predecessor John Walker's perverse "Neptunian" geological theories. Darwin attended Robert Jameson's natural history course at the University of Edinburgh in his teenage years, learning about stratigraphic geology and assisting with the collections of the Museum of Edinburgh University, then one of the largest in Europe. At Jameson's Wernerian Natural History Association, the young Charles Darwin saw John James Audubon give a demonstration of his method of using wires to prop up birds to draw or paint them in natural positions. Robert Jameson was the great-uncle of Sir Leander Starr Jameson, Bt, KCMG, CB, British colonial official and inspiration for the "Jameson Raid".
Document 4: Evolution: In the mid-19th century, Charles Darwin formulated the scientific theory of evolution by natural selection, published in his book On the Origin of Species (1859). Evolution by natural selection is a process first demonstrated by the observation that often, more offspring are produced than can possibly survive. This is followed by three observable facts about living organisms: 1) traits vary among individuals with respect to morphology, physiology, and behaviour (phenotypic variation), 2) different traits confer different rates of survival and reproduction (differential fitness), and 3) traits can be passed from generation to generation (heritability of fitness). Thus, in successive generations members of a population are replaced by progeny of parents better adapted to survive and reproduce in the biophysical environment in which natural selection takes place.
Document 5: On the Origin of Species: On the Origin of Species was first published on Thursday 24 November 1859, priced at fifteen shillings with a first printing of 1250 copies. The book had been offered to booksellers at Murray's autumn sale on Tuesday 22 November, and all available copies had been taken up immediately. In total, 1,250 copies were printed but after deducting presentation and review copies, and five for Stationers' Hall copyright, around 1,170 copies were available for sale. Significantly, 500 were taken by Mudie's Library, ensuring that the book promptly reached a large number of subscribers to the library. The second edition of 3,000 copies was quickly brought out on 7 January 1860, and incorporated numerous corrections as well as a response to religious objections by the addition of a new epigraph on page ii, a quotation from Charles Kingsley, and the phrase "by the Creator" added to the closing sentence. During Darwin's lifetime the book went through six editions, with cumulative changes and revisions to deal with counter-arguments raised. The third edition came out in 1861, with a number of sentences rewritten or added and an introductory appendix, An Historical Sketch of the Recent Progress of Opinion on the Origin of Species, while the fourth in 1866 had further revisions. The fifth edition, published on 10 February 1869, incorporated more changes and for the first time included the phrase "survival of the fittest", which had been coined by the philosopher Herbert Spencer in his Principles of Biology (1864).
Document 6: On the Origin of Species: Evolution had less obvious applications to anatomy and morphology, and at first had little impact on the research of the anatomist Thomas Henry Huxley. Despite this, Huxley strongly supported Darwin on evolution; though he called for experiments to show whether natural selection could form new species, and questioned if Darwin's gradualism was sufficient without sudden leaps to cause speciation. Huxley wanted science to be secular, without religious interference, and his article in the April 1860 Westminster Review promoted scientific naturalism over natural theology, praising Darwin for "extending the domination of Science over regions of thought into which she has, as yet, hardly penetrated" and coining the term "Darwinism" as part of his efforts to secularise and professionalise science. Huxley gained influence, and initiated the X Club, which used the journal Nature to promote evolution and naturalism, shaping much of late Victorian science. Later, the German morphologist Ernst Haeckel would convince Huxley that comparative anatomy and palaeontology could be used to reconstruct evolutionary genealogies.
Document 7: History of biology: The emerging discipline of geology also brought natural history and natural philosophy closer together; the establishment of the stratigraphic column linked the spatial distribution of organisms to their temporal distribution, a key precursor to concepts of evolution. Georges Cuvier and others made great strides in comparative anatomy and paleontology in the late 1790s and early 19th century. In a series of lectures and papers that made detailed comparisons between living mammals and fossil remains Cuvier was able to establish that the fossils were remains of species that had become extinct—rather than being remains of species still alive elsewhere in the world, as had been widely believed. Fossils discovered and described by Gideon Mantell, William Buckland, Mary Anning, and Richard Owen among others helped establish that there had been an 'age of reptiles' that had preceded even the prehistoric mammals. These discoveries captured the public imagination and focused attention on the history of life on earth. Most of these geologists held to catastrophism, but Charles Lyell's influential Principles of Geology (1830) popularised Hutton's uniformitarianism, a theory that explained the geological past and present on equal terms.
Document 8: On the Origin of Species: On the Origin of Species, published on 24 November 1859, is a work of scientific literature by Charles Darwin which is considered to be the foundation of evolutionary biology. Darwin's book introduced the scientific theory that populations evolve over the course of generations through a process of natural selection. It presented a body of evidence that the diversity of life arose by common descent through a branching pattern of evolution. Darwin included evidence that he had gathered on the Beagle expedition in the 1830s and his subsequent findings from research, correspondence, and experimentation.
Document 9: History of science: Geology did not undergo systematic restructuring during the Scientific Revolution, but individual theorists made important contributions. Robert Hooke, for example, formulated a theory of earthquakes, and Nicholas Steno developed the theory of superposition and argued that fossils were the remains of once-living creatures. Beginning with Thomas Burnet's Sacred Theory of the Earth in 1681, natural philosophers began to explore the idea that the Earth had changed over time. Burnet and his contemporaries interpreted Earth's past in terms of events described in the Bible, but their work laid the intellectual foundations for secular interpretations of Earth history.
Document 10: James Hutton: James Hutton FRSE (/ ˈhʌtən /; 3 June 1726 -- 26 March 1797) was a Scottish geologist, physician, chemical manufacturer, naturalist, and experimental agriculturalist. He originated the theory of uniformitarianism -- a fundamental principle of geology -- that explains the features of the Earth's crust by means of natural processes over geologic time. Hutton's work established geology as a science, and as a result he is referred to as the ``Father of Modern Geology ''.
Document 11: The Descent of Man, and Selection in Relation to Sex: The Descent of Man, and Selection in Relation to Sex is a book by English naturalist Charles Darwin, first published in 1871, which applies evolutionary theory to human evolution, and details his theory of sexual selection, a form of biological adaptation distinct from, yet interconnected with, natural selection. The book discusses many related issues, including evolutionary psychology, evolutionary ethics, differences between human races, differences between sexes, the dominant role of women in mate choice, and the relevance of the evolutionary theory to society.
Document 12: On the Origin of Species: In December 1831, he joined the Beagle expedition as a gentleman naturalist and geologist. He read Charles Lyell's Principles of Geology and from the first stop ashore, at St. Jago, found Lyell's uniformitarianism a key to the geological history of landscapes. Darwin discovered fossils resembling huge armadillos, and noted the geographical distribution of modern species in hope of finding their "centre of creation". The three Fuegian missionaries the expedition returned to Tierra del Fuego were friendly and civilised, yet to Darwin their relatives on the island seemed "miserable, degraded savages", and he no longer saw an unbridgeable gap between humans and animals. As the Beagle neared England in 1836, he noted that species might not be fixed.
Document 13: On the Origin of Species: On the Origin of Species The title page of the 1859 edition of On the Origin of Species Author Charles Darwin Country United Kingdom of Great Britain and Ireland Language English Subject Natural selection Evolutionary biology Published 24 November 1859 (John Murray) Media type Print (Hardback) Pages 502 OCLC 352242 Preceded by On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection Followed by Fertilisation of Orchids
Document 14: Common descent: Universal common descent through an evolutionary process was first proposed by the English naturalist Charles Darwin in On the Origin of Species (1859), which concluded: ``There is grandeur in this view of life, with its several powers, having been originally breathed into a few forms or into one; and that, whilst this planet has gone cycling on according to the fixed law of gravity, from so simple a beginning endless forms most beautiful and most wonderful have been, and are being, evolved. ''
Document 15: On the Origin of Species: On the Origin of Species (or, more completely, On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life), published on 24 November 1859, is a work of scientific literature by Charles Darwin which is considered to be the foundation of evolutionary biology. Darwin's book introduced the scientific theory that populations evolve over the course of generations through a process of natural selection. It presented a body of evidence that the diversity of life arose by common descent through a branching pattern of evolution. Darwin included evidence that he had gathered on the "Beagle" expedition in the 1830s and his subsequent findings from research, correspondence, and experimentation.
ground_truth: University of Edinburgh
sub_queries: who wrote the book the origin of species Answer: Charles Darwin
Charles Darwin >> educated at Answer: University of Edinburgh