Question: At what university did the man who figured out the evolutionary mechanism called natural selection, receive his education?

full_context: Document 1: Survival of the fittest: Herbert Spencer first used the phrase, after reading Charles Darwin's On the Origin of Species, in his Principles of Biology (1864), in which he drew parallels between his own economic theories and Darwin's biological ones: ``This survival of the fittest, which I have here sought to express in mechanical terms, is that which Mr. Darwin has called 'natural selection', or the preservation of favoured races in the struggle for life. ''
Document 2: Natural selection: Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in heritable traits of a population over time. Charles Darwin popularised the term ``natural selection '', and compared it with artificial selection.
Document 3: On the Origin of Species: In December 1831, he joined the Beagle expedition as a gentleman naturalist and geologist. He read Charles Lyell's Principles of Geology and from the first stop ashore, at St. Jago, found Lyell's uniformitarianism a key to the geological history of landscapes. Darwin discovered fossils resembling huge armadillos, and noted the geographical distribution of modern species in hope of finding their "centre of creation". The three Fuegian missionaries the expedition returned to Tierra del Fuego were friendly and civilised, yet to Darwin their relatives on the island seemed "miserable, degraded savages", and he no longer saw an unbridgeable gap between humans and animals. As the Beagle neared England in 1836, he noted that species might not be fixed.
Document 4: On the Origin of Species: On the Origin of Species The title page of the 1859 edition of On the Origin of Species Author Charles Darwin Country United Kingdom of Great Britain and Ireland Language English Subject Natural selection Evolutionary biology Published 24 November 1859 (John Murray) Media type Print (Hardback) Pages 502 OCLC 352242 Preceded by On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection Followed by Fertilisation of Orchids
Document 5: History of science: Geology did not undergo systematic restructuring during the Scientific Revolution, but individual theorists made important contributions. Robert Hooke, for example, formulated a theory of earthquakes, and Nicholas Steno developed the theory of superposition and argued that fossils were the remains of once-living creatures. Beginning with Thomas Burnet's Sacred Theory of the Earth in 1681, natural philosophers began to explore the idea that the Earth had changed over time. Burnet and his contemporaries interpreted Earth's past in terms of events described in the Bible, but their work laid the intellectual foundations for secular interpretations of Earth history.
Document 6: On the Origin of Species: Evolution had less obvious applications to anatomy and morphology, and at first had little impact on the research of the anatomist Thomas Henry Huxley. Despite this, Huxley strongly supported Darwin on evolution; though he called for experiments to show whether natural selection could form new species, and questioned if Darwin's gradualism was sufficient without sudden leaps to cause speciation. Huxley wanted science to be secular, without religious interference, and his article in the April 1860 Westminster Review promoted scientific naturalism over natural theology, praising Darwin for "extending the domination of Science over regions of thought into which she has, as yet, hardly penetrated" and coining the term "Darwinism" as part of his efforts to secularise and professionalise science. Huxley gained influence, and initiated the X Club, which used the journal Nature to promote evolution and naturalism, shaping much of late Victorian science. Later, the German morphologist Ernst Haeckel would convince Huxley that comparative anatomy and palaeontology could be used to reconstruct evolutionary genealogies.
Document 7: On the Origin of Species: Chapter I covers animal husbandry and plant breeding, going back to ancient Egypt. Darwin discusses contemporary opinions on the origins of different breeds under cultivation to argue that many have been produced from common ancestors by selective breeding. As an illustration of artificial selection, he describes fancy pigeon breeding, noting that "[t]he diversity of the breeds is something astonishing", yet all were descended from one species of rock pigeon. Darwin saw two distinct kinds of variation: (1) rare abrupt changes he called "sports" or "monstrosities" (example: ancon sheep with short legs), and (2) ubiquitous small differences (example: slightly shorter or longer bill of pigeons). Both types of hereditary changes can be used by breeders. However, for Darwin the small changes were most important in evolution.
Document 8: Robert Jameson: As Regius Professor of Natural History at the University of Edinburgh for fifty years, Jameson is notable for his advanced scholarship, his superb museum collection and for his tuition of Charles Darwin. Jameson was not at his best in the lecture theatre however, and, for the first half of his career, he grappled with his predecessor John Walker's perverse "Neptunian" geological theories. Darwin attended Robert Jameson's natural history course at the University of Edinburgh in his teenage years, learning about stratigraphic geology and assisting with the collections of the Museum of Edinburgh University, then one of the largest in Europe. At Jameson's Wernerian Natural History Association, the young Charles Darwin saw John James Audubon give a demonstration of his method of using wires to prop up birds to draw or paint them in natural positions. Robert Jameson was the great-uncle of Sir Leander Starr Jameson, Bt, KCMG, CB, British colonial official and inspiration for the "Jameson Raid".
Document 9: Natural selection: Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the heritable traits characteristic of a population over generations. Charles Darwin popularised the term ``natural selection '', contrasting it with artificial selection, which is intentional, whereas natural selection is not.
Document 10: On the Origin of Species: Later chapters provide evidence that evolution has occurred, supporting the idea of branching, adaptive evolution without directly proving that selection is the mechanism. Darwin presents supporting facts drawn from many disciplines, showing that his theory could explain a myriad of observations from many fields of natural history that were inexplicable under the alternate concept that species had been individually created. The structure of Darwin's argument showed the influence of John Herschel, whose philosophy of science maintained that a mechanism could be called a vera causa (true cause) if three things could be demonstrated: its existence in nature, its ability to produce the effects of interest, and its ability to explain a wide range of observations.
Document 11: Darwinism: Darwinism is a theory of biological evolution developed by the English naturalist Charles Darwin (1809 -- 1882) and others, stating that all species of organisms arise and develop through the natural selection of small, inherited variations that increase the individual's ability to compete, survive, and reproduce. Also called Darwinian theory, it originally included the broad concepts of transmutation of species or of evolution which gained general scientific acceptance after Darwin published On the Origin of Species in 1859, including concepts which predated Darwin's theories. It subsequently referred to the specific concepts of natural selection, the Weismann barrier, or the central dogma of molecular biology. Though the term usually refers strictly to biological evolution, creationists have appropriated it to refer to the origin of life, and it has even been applied to concepts of cosmic evolution, both of which have no connection to Darwin's work. It is therefore considered the belief and acceptance of Darwin's and of his predecessors' work -- in place of other theories, including divine design and extraterrestrial origins.
Document 12: James Hutton: James Hutton FRSE (/ ˈhʌtən /; 3 June 1726 -- 26 March 1797) was a Scottish geologist, physician, chemical manufacturer, naturalist, and experimental agriculturalist. He originated the theory of uniformitarianism -- a fundamental principle of geology -- that explains the features of the Earth's crust by means of natural processes over geologic time. Hutton's work established geology as a science, and as a result he is referred to as the ``Father of Modern Geology ''.
Document 13: History of biology: The emerging discipline of geology also brought natural history and natural philosophy closer together; the establishment of the stratigraphic column linked the spatial distribution of organisms to their temporal distribution, a key precursor to concepts of evolution. Georges Cuvier and others made great strides in comparative anatomy and paleontology in the late 1790s and early 19th century. In a series of lectures and papers that made detailed comparisons between living mammals and fossil remains Cuvier was able to establish that the fossils were remains of species that had become extinct—rather than being remains of species still alive elsewhere in the world, as had been widely believed. Fossils discovered and described by Gideon Mantell, William Buckland, Mary Anning, and Richard Owen among others helped establish that there had been an 'age of reptiles' that had preceded even the prehistoric mammals. These discoveries captured the public imagination and focused attention on the history of life on earth. Most of these geologists held to catastrophism, but Charles Lyell's influential Principles of Geology (1830) popularised Hutton's uniformitarianism, a theory that explained the geological past and present on equal terms.
Document 14: Evolution: In the mid-19th century, Charles Darwin formulated the scientific theory of evolution by natural selection, published in his book On the Origin of Species (1859). Evolution by natural selection is a process first demonstrated by the observation that often, more offspring are produced than can possibly survive. This is followed by three observable facts about living organisms: 1) traits vary among individuals with respect to morphology, physiology, and behaviour (phenotypic variation), 2) different traits confer different rates of survival and reproduction (differential fitness), and 3) traits can be passed from generation to generation (heritability of fitness). Thus, in successive generations members of a population are replaced by progeny of parents better adapted to survive and reproduce in the biophysical environment in which natural selection takes place.
Document 15: Alfred Nalepa: He studied natural sciences at the University of Vienna, and from 1886 was associated with the "Lehrerbildungsanstalt" in Linz. In 1892 he returned to Vienna, where he was appointed professor of natural history at the "Elisabethgymnasium".
ground_truth: University of Edinburgh
sub_queries: who figured out the mechanism of evolution known as natural selection Answer: Charles Darwin
Charles Darwin >> educated at Answer: University of Edinburgh