Two cladograms proposed for the genus "Panthera". The upper cladogram is based on a 2006 phylogenetic study by Warren E. Johnson (of the National Cancer Institute) and colleagues, and a 2009 study by Lars Werdelin and colleagues. The lower cladogram is based on a 2010 study by Brian W. Davis (of the Texas A&M University) and colleagues and a 2011 study by Ji H. Mazák (of the Shanghai Science and Technology Museum) and colleagues.