Original sentence: (2008), haplogroup D is older than haplogroup E, the exact difference in age between the two clades is disputed.(2008), in which he was leading contributor, he is described as saying that:.Although, according to Shi et al.The relative ages of DE and its two branches, haplogroup D and haplogroup E, are also considered relevant to the likelihood of Asian or African origins
Edited sentence: (2008), in which he was leading contributor, he is described as saying that....The relative age of DE and its two branches D (Asian) and E (African) is also considered relevant to the likelihood of Asian or African origins