## 3 Results and Discussion
### The Concentration of Biogenic Amines in Lentil Wholemal Samples  
Biogenic amine concentration in lentil samples is tabulated in Table 4. Tryptamine (TRIP), cadaverine (CAD), histamine (HIS), and tyramine (TYR) were not detected in any lentil groups. Phenylethylamine (PHE) was found only in SMFLR24 samples. In most of the cases, fermented samples contained higher concentrations of other detected biogenic amines--chiefly, putrescine (PUT), spermidine (SPRMD), and spermine (SPRM)--in comparison to the respective sample control groups. In this regard, total BA concentration in L and LR SMF and SSF sample groups were significantly higher in than the respective control sample groups. All factors and their interactions were statistically significant on PHE concentration in lentil samples (see supplementary file: Table S6). Moreover, all factors and the growing conditions * type of fermentation interaction were statistically significant for PUT concentration. Almost all factors and the interactions thereof, except type of fermentation, were statistically significant for SPRMD concentration. The type of fermentation, the growing conditions * type of fermentation interaction, and the growing conditions * fermentation  
\begin{table}
\begin{tabular}{l l l l l l l l l l l} \hline \hline \multirow{2}{*}{**Lentil Samples**} & \multirow{2}{*}{**TRIP**} & \multirow{2}{*}{**PHE**} & \multirow{2}{*}{**PUT**} & \multirow{2}{*}{**CAD**} & \multirow{2}{*}{**HIS**} & \multirow{2}{*}{**TYR**} & \multirow{2}{*}{**SPRMD**} & \multirow{2}{*}{**SPRM**} & \multicolumn{2}{c}{**Total BA**} \\  & & & & & & & & & & & **Content** \\ \hline \multicolumn{10}{c}{**mg/kg**} \\ \multirow{5}{*}{LR} & C\({}_{\text{LB}}\)SM & nd & nd & nd & nd & nd & nd & nd & 75.8 \(\pm\) 5.06 \({}^{\text{\,\text{\text{\text{\  
duration interaction were statistically significant for SPRM concentration in lentil samples. The correlation analysis (see supplementary file: Table S5) discovered that only PUT and SPRM possessed strong and weak positive correlations with LAB viable counts (r = 0.761, \(p\) < 0.001 and r = 0.331, \(p\) = 0.049, respectively), while only PUT showed a significant strong positive correlation with the pH of the lentilts (r = 0.719, \(p\) < 0.001). Strong and weak negative correlations of PUT with arginine and glutamine (r = \(-\)0.686, \(p\) < 0.001 and r = \(-\)0.341, \(p\) = 0.042, respectively) were established. Moreover, strong positive correlations of PUT with glutamic and aspartic acids, valine, phenylalanine, leucine/isoleucine, and lysine (r = 0.734, \(p\) < 0.001; r = 0.881, \(p\) < 0.001; r = 0.614, \(p\) < 0.001; r = 0.639, \(p\) < 0.001; r = 0.697, \(p\) < 0.001; and r = 0.583, \(p\) < 0.001, respectively), and moderate positive correlations with threonine, glycine, and proline (r = 0.594, \(p\) < 0.001; r = 0.349, \(p\) = 0.037; and r = 0.468, \(p\) = 0.004, respectively) were found. SPRMD showed moderate positive correlations with GABA and tyrosine (r = 0.424, \(p\) = 0.010; and r = 0.534, \(p\) < 0.001, respectively). Also, a weak positive correlation of SPRMD with methionine (r = 0.356, \(p\) = 0.033) was observed. Strong positive correlations were found between SPRM and alanine, proline, glycine, phenylalanine, and histidine (r = 0.787, \(p\) < 0.001; r = 0.604, \(p\) < 0.001; r = 0.644, \(p\) < 0.001; r = 0.610, \(p\) < 0.001; and r = 0.668, \(p\) < 0.001, respectively). Furthermore, moderate positive correlations between SPRM and glutamic acid, serine, threonine, GABA, leucine/isoleucine, tyrosine, valine, and lysine (r = 0.461, \(p\) = 0.005; r = 0.430, \(p\) = 0.009; r = 0.563, \(p\) < 0.001; r = 0.566, \(p\) < 0.001; r = 0.425, \(p\) = 0.010; r = 0.466, \(p\) = 0.004; r = 0.520, \(p\) = 0.001; and r = 0.575, \(p\) < 0.001, respectively) were established. Yet, a weak positive correlation between SPRM and glutamine (r = 0.354, \(p\) = 0.034) was observed.