); [EMAIL_ADDRESS] (Y.H)  
[PERSON]  
1 College of Agronomy, Henan Agricultural University, Zhengzhou 450002, China; [EMAIL_ADDRESS] (J.Z. ); [EMAIL_ADDRESS] (C.Y. ); [EMAIL_ADDRESS] (X.L. ); [EMAIL_ADDRESS] (X.L. ); [EMAIL_ADDRESS] (Y.H)  
[PERSON]  
1 College of Agronomy, Henan Agricultural University, Zhengzhou 450002, China; [EMAIL_ADDRESS] (J.Z. ); [EMAIL_ADDRESS] (C.Y. ); [EMAIL_ADDRESS] (X.L. ); [EMAIL_ADDRESS] (X.L. ); [EMAIL_ADDRESS] (Y.H)  
[PERSON]  
1 College of Agronomy, Henan Agricultural University, Zhengzhou 450002, China; [EMAIL_ADDRESS] (J.Z. ); [EMAIL_ADDRESS] (C.Y. ); [EMAIL_ADDRESS] (X.L. ); [EMAIL_ADDRESS] (X.L. ); [EMAIL_ADDRESS] (Y.H)  
[PERSON]  
1 College of Agronomy, Henan Agricultural University, Zhengzhou 450002, China; [EMAIL_ADDRESS] (J.Z. ); [EMAIL_ADDRESS] (C.Y. ); [EMAIL_ADDRESS] (X.L. ); [EMAIL_ADDRESS] (X.L. ); [EMAIL_ADDRESS] (Y.H.) [PERSON]  
1 College of Agronomy, Henan Agricultural University, Zhengzhou 450002, China; [EMAIL_ADDRESS] (J.Z. ); [EMAIL_ADDRESS] (C.Y. ); 1883891693@163.  
herb in China. In recent decades, many chemical and pharmacological studies have been done on fresh, dried, and steamed tuberous root of _R. glutinosa_. Many natural compounds, including phenylethanol glycosides (PhGs), iridoid glycosides (IGs), ionone glycosides, polysaccharides, amino acids, and other components, have been found in the herb [1; 2; 3]. Many studies show that _R. glutinosa_ and its active compounds possess wide pharmacological actions on the immune system, the cardiovascular system, the blood system, the endocrine system, and the nervous system [4]. _R. glutinosa_ is also thought of as a tonic brain medicine and is commonly used in the prevention and treatment of some central-nervous-system-related diseases, such as depression, cognitive impairment, and anti-anxiety [5]. Extensive research has revealed that the contents of effective components in different organs and tissues of medicinal plants are different. HPLC analysis has revealed that the production of chlorogenic acid, caffe acid, ferulic acid, luteoloside, and quercetin is reduced after flowering [6]. Phytochemical analysis demonstrates the localization of tanshinones in the peeled periderm [7] and hydrophilic phenolic acids in the phloem and xylem [8], respectively. The content of major phenolic compounds varies significantly among the olive cultivars and changes during fruit development and maturation, with some compounds showing specificity for certain cultivars [9]. Studies on _R. glutinosa_ active compounds accumulation have mainly focused on producing area and harvesting time. For example, the concentration of catalpol in _R. glutinosa_ tuberous root is higher in the genuine producing areas [10], and total iridoid glycosides content gradually increases with tuberous root development [11]. There are great differences in acteoside content in _R. glutinosa_ tuberous roots and leaves among the different cultivars. [PERSON] and Qinhuai are high-acteoside cultivars with a higher content of acteoside in tuberous root [12]. A transverse section of _R. glutinosa_ has obvious radial striation from the genuine producing area, which was identified as a sign of genuine medicinal material by the generation of pharmacists. Therefore, the existence of radial striation might be involved in the quality of medicinal materials. In our previous research, we found that the content of catalpol and acteoside of _R. glutinosa_ at different development stages varied with different change trends, and there was a significant difference in the component accumulation between radial striation (RS) and non-radial striation (nRS) of tuberous root [13]. However, the molecular mechanism of _R. glutinosa_ radial striation and non-radial striation quality formation is still unclear. Catalpol is an iridoid glycoside, a main active compound of _R.