## 4 Discussion
### Conflict between Morphological and Molecular Inferences  
Neither maximum-likelihood nor Bayesian multilocus phylogenetic approaches supported the two putative genera based on paramere morphology. However, there appear to be state specific clades distinguished by forceps morphology. All clades have unique male forceps morphology, highlighting the trait's utility in distinguishing morphospecies, but in some cases paramere morphology is necessary to distinguish species. At broader scales, across clades, sole reliance on paramere morphology might lead to misleading species assignment. Some of the \(G\). sp. 1 and \(G\). sp. 2 specimens showed similar paramere morphology despite being phylogenetically distant. Moreover, the lengths of the parameres of \(A\). sp. 1 and \(A\). sp. 2 are closer to each other than to other morphospecies, but the two species are not closely related according to the molecular phylogeny. This echoes the argument made by [PERSON] et al. [113] working with another family, the Pygidicranidae; even among closely related congeners, genital characters may mislead. The generic assignments here are not all necessarily valid. [PERSON] [60] revised the systematics of the Anisolabididae genera and generic assignments based on this system would yield the following changes; \(A\) sp. 1 would become _Gonolabis_ sp. ; _Anisolabis_ sp. 2 would become _Apolabis_ sp. ; _G. forcipata_ remains unknown, keying to either _Mongolabis_ or _Anisolabella_, but lacking the snout-like paramere structure of _Mongolabis_ nor the continuous outer paramere margin of _Anisolabella_; \(G\) nr. _gilesi_ would become _Anisolabella_ sp. 1; \(G\). sp. 1 would become _Mongolabis_ sp. ; \(G\). sp. 2 would remain unknown for similar reasons to \(G\). _forcipata_; and \(G\). sp. 3 would become _Anisolabella_ sp. 2. In such a case, the _Anisolabella_ become polyphyletic, and the conjecture that similar paramere structure does not reflect phylogeny remains, regardless of the nomenclature used. This is unsurprising given [PERSON]'s heavy reliance on paramere morphology for distinguishing the genera of the Anisolabidinae subfamily [60]. Given these issues, the specimens presented here should be incorporated, along with their putative identifications, into a fuller cladistic and phylogenetic analysis. The phylogenetic signal of insect genitalia has been subject to some debate. Polyandry has been found in confamilial species _Euborellia plebeja_ ([PERSON]) [114; 115; 116; 117] and _Euborellia brunneri_ ([PERSON]) [118], so it is likely that genital morphology is subject to sexual selection. However, the precise function of the parameres is unknown [41] so it is difficult to make definitive statements about the effect of sexual selection on their morphology. Structures of the male forceps often play specific roles in male-male competition [38] so they should also be under strong sexual selection for morphological evolution. [PERSON] [119] and [PERSON] and [PERSON] [120] both cited [PERSON] [121] in arguing that rates of morphological evolution (which may be accelerated by sexual selection) in insect male genitalia relative to the rate of speciation should be high enough to swamp phylogenetic inertia and that there should therefore be little correlation between morphological similarity in male genitalia and phylogenetic distance. The present study is equivocal on the matter, as the SA clade species show similar paramere morphology, while the WA clade do not. [PERSON] and [PERSON] [122] explicitly tested the phylogenetic signal of male insect genitalia and found that the genitalia are composite structures, and some sub-components may be conserved whereas others may be free to diversify. In the present study, the parameres were used for generic identification, but other characters such as the length and asymmetry of the virgae and the length of the penis lobes may also prove useful in describing the phylogeny of the morphospecies as they have already in describing the family-level phylogeny of the order [41].