sentence: However , the genetic mechanism of GMCs in milled grains remains largely unknown.Previously , we used two sets of backcrossed inbred lines ( BILs ) derived from the same donor , and two elite new varieties in Southwestern China , Ce258 and Zhongguangxiang1 ( ZGX1 ) as recipients , to assess the genetic background and the genotypic by environment ( G × E ) effects of GMC traits in rice milled grains using QTL mapping ( Xu et al ., 2015 ) .