sentence: In particular , previous molecular phylogenetic studies using chloroplast and nuclear loci indicated that Jaltomata is close to both Solanum and Capsicum , however the relationship among these three genera has varied depending upon the specific loci used in phylogenetic reconstruction ( Bohs and Olmstead 1997 ; Olmstead et al. 1999 ; Walsh and Hoot 2001 ; Olmstead et al. 2008 ; S ä rkinen et al. 2013 ) .