sentence: In other plant species , several MYB transcription factors are associated with lignin synthesis , including PtrMYB3 , PtrMYB20 , EgMYB2 , PvMYB4 , PtMYB1 , PtMYB4 , PtMYB8 , OsMYB46 , and ZmMYB ( Patzlaff et al ., 2003 ; Bomal et al ., 2008 ; McCarthy et al ., 2010 ; Rahantamalala et al ., 2010 ; Zhong et al ., 2011 ; Shen et al ., 2012 ) .