sentence: The Doa10 complex also localizes to the INM and has nuclear substrates such as the soluble transcriptional repressor Mat2 α ( Deng and Hochstrasser , 2006 , Swanson et al ., 2001 ) and the membrane proteins Asi2 ( Boban et al ., 2014 ) and Ubc6 ( Walter et al ., 2001 ) .