sentence: The highest copy numbers were found in R. belli transformed with pRAM18dRGA ( 13.3 to 28.1 ) , followed by R. parkeri ( 9.9 ) , R. montanensis ( 7.5 ) , and R. monacensis ( 5.5 ) ( 15 ) , while plasmid copy numbers in R. conorii ( 1 – 3 ) ( 16 ) and R. prowazekii ( 0.86 ) were relatively low ( 17 ) .