This ability has evolved independently twice as it seems -- in the flanged bombardier beetles ( Paussinae ) which are among the most ancient ground beetles , and in the typical bombardier beetles ( Brachininae ) which are part of a more `` modern '' lineage .
This ability has evolved twice -- in the flanged bombardier beetles ( Paussinae ) which are among the most ancient ground beetles , as well as in the typical bombardier beetles ( Brachininae ) which are part of a more `` modern '' lineage . ''