Proposal for a Council Directive adapting to technical and scientific progress Directive 92/43/EEC on the conservation of natural habitats and of wild fauna and flora
Date: 1997-05-28
Institutions: European Commission
Subjects: protected area, protected species, protection of animal life, protection of plant life, wildlife

COMMISSION OF THE EUROPEAN COMMUNITIES 

Brussels,  28.05.1997 
COM(97)  195  final 

Proposal  for  a 

COUNCIL  DIRECTIVE 

adapting to technical and  scientific 

progress Directive 92/43/EEC on the conservation  of natural habitats and of 

wild fauna  and  flora 

(presented by  the  Commission) 

EXPLANATORY  MEMORANDUM 

Subject: 

Proposal  for  a  Council  Directive  (EC)  adapting  to  technical  and  scientific 
progress Directive 92/43/EEC on the conservation of natural  habitats and of 
wild fauna and flora 

1. 

2. 

3. 

4. 

Council  Directive 92/43/EEC  of 21 May  1992 on the conservation  of  natural  habitats  and 
of  wild  fauna  and  flora  (the  "Habitats  Directive")  provides  for  the  creation,  in  several 
stages, of a network of protected natural habitats called Natura 2000. 

Under  the  terms  of  Article  2(2)  of  Directive  92/43/EEC,  measures  taken  pursuant  to  the 
Directive  must  be  designed  to  maintain  or  restore,  at  favourable  conservation  status, 
natural habitats and species of wild fauna and flora of Community  interest. 

Pursuant to Article 4 of Directive 92/43/EEC, each Member  State is required, on the  basis 
of the criteria set out in Annex III (Stage  1) and relevant scientific  information,  to  propose 
a list of sites indicating which natural habitat types in Annex I and which species in Annex 
II that are native to its territory the sites host. 

Annexes  I  and  II  to  Directive  92/43/EEC  may  be  amended  in  accordance  with  the  first 
paragraph  of  Article  19.  The  amendments  necessary  for  adaptation  to  technical  and 
scientific  progress  must  be  adopted  by  the  Council  acting  by  qualified  majority  on  a 
proposal from  the Commission. 

5.  When Austria, Finland and Sweden acceded to the Union, Annexes I and II were  amended 

(OJNo LI,  1.1.1995, p. 135). 

6. 

However,  several  Member  States  and  the  Scientific  Working  Group  set  up  by  the 
"Habitats" Committee (established by the Directive) wanted Annexes I and II to  Directive 
92/43/EEC  to be adapted  to technical  and  scientific  progress.  Adaptation  was  requested 
to cover: 

- 

- 

the  description  of  the  habitat  types  of  Community  interest  in  the  Boreal  region  to 
better  define  the  additions  made  to  the  original  lists  when  Sweden  and  Finland 
acceded. 

the scientific  nomenclature of several species, which has changed  since the  original 
annexes were drawn up. 

All  up-to-date  scientific  information  on  natural  habitat  types  and  the  relevant  species  of 
Community  interest indicates that it is necessary to adapt Annexes I and II. 

As  regards  Annex  I,  the  process  of  updating  the  classification  of  European  habitats, 

carried  out  on  the  basis  of  the  Corine  biotopes  project, 1  led  to  the  production  of  an 
Interpretation  Manual of European Union Habitats.  This Manual  includes the new Natura 
2000  codes  which  identify  each  natural  habitat  type.  The  first  EUR  12  version  of  the 
Manual  was  published  by  the  Commission  in  April  1995  and  contained  updated 
information  for twelve Member  States.  A second  EUR  15 version  was published  in April 
1996,  incorporating  the  new  priority  habitats  of  Community  interest  which  were  added 
when the three new Member States acceded. 

i  Corine biotopes - Technical Handbook, Volume 1, pp. 73-109; Corine/Biotopes/89-2.2,  19 May 1988, 

partially updated  14 February 1989. 

7. 

* 

- 

- 

- 

- 

- 

- 

- 

- 

* 

- 

At  the  "Habitats"  Committee  meeting  of  13  September  1996, Member  States'  delegates 
unanimously approved the following  amendments: 

Annex I: 

to replace the reference  to Corine biotopes for  classifying  habitat types with  a reference  to 
the  new  Interpretation  Manual  of  European  Union  Habitats,  EUR  15  version  of  April 
1996; 

to  replace  the  1989  Corine  code  with  the Natura  2000  code,  now  the  standard  scientific 
reference; 

to describe  new types of Boreal habitats, including  seven priority  habitats,  in line with the 
opinion of the Scientific  Working Group of the Habitats Committee. 

to  remove  the  habitat  sub-types  (e.g.  "grey  dunes")  which  are  now  described  in  the 
Interpretation  Manual, EUR  15 version; 

to take  account of the recent scientific  identification  of the habitat type Turloughs  outside 
Ireland;  to  therefore  delete  the  word  "(Ireland)"  from  the  habitat  type  "Turloughs 
(Ireland)"; 

to  reclassify  the habitat "*Taxus baccata woods" in keeping with  scientific  advice  so that 
formations  in  the  British  Isles  will  be  placed  in  the  sub-category  "Forests  of  temperate 
Europe" rather than "Mediterranean  mountainous  forests"; 

to  replace  a  number  of titles  causing  habitat  types  to  be  confused  by  more  explicit  titles 
from  the Interpretation Manual, EUR  15 version; 

to  replace  the  common  names  of  species  under  each  title  by  their  Latin  names,  so  as  to 
conform  to scientific  usage. 

Annex II: 

to adapt the names of species not correctly  identified  when the Annexes were drawn up, to 
the taxonomy and nomenclature currently  in force  in the scientific  community; 

- 

to add certain Nordic species identified jointly by scientists in Sweden and Finland. 

Member  States  are  not  expected  to  experience  any  additional  difficulty  in  complying  with  the 
amendments to this Directive. 

8. 

In conformity  with  the principle  of subsidiarity  and  with the principle  of  proportionality 
it  turns  out  to  be  necessary  that  the  present  proposal  is  carried  out  at  the  Community 
level  because  it concerns  the  simple  adaptation  to technical  and  scientific  progress  of  a 
basic act which emanates from  the Community. 

*** 

The Commission  proposes to the Council to adopt the attached  proposal for a Council  Directive. 

Proposal for a Council Directive (EC) adapting to technical and  scientific 

progress Directive 92/43/EEC on the conservation of natural habitats and of 

wild fauna and flora 

THE COUNCIL OF THE EUROPEAN UNION, 

Having regard to the Treaty establishing the European Community, 

Having  regard  to  Council  Directive  92/43/EEC  of  21  May  1992  on the  conservation  of 

natural  habitats  and  of  wild  fauna  and  flora,1  and  in  particular  the  first  paragraph  of 

Article  19 thereof, 

Having regard to the proposal from the Commission, 

Whereas  certain  natural  habitat  types  and  species  in  Annexes  I  and  II  to  Directive 

92/43/EEC should be adapted to technical and scientific progress; 

Whereas the Interpretation Manual of European Union Habitats (EUR  15 version of April 

1996)  includes  the  new  Natura  2000  codes  which  identify  each  natural  habitat  type; 

whereas  the  reference  to  the  Corine  code  in Annex  I to  Directive  92/43/EEC  should  be 

replaced by a reference  to the Natura 2000 code; 

HAS ADOPTED THIS DIRECTIVE: 

Article 1 

Annexes  I  and  II to  Directive  92/43/EEC  shall  be  replaced  by  the  text  in  the  Annex  to 

this Directive. 

1  OJ No L 206, 22.7.1992, p. 7, amended by the Act of Accession of Austria, Finland and Sweden. 

Arttekl 

Member States shall bring into force the laws, regulations and administrative provisions 

necessary  to  comply  with  this  Directive  by  31 December  1997.  They  shall  forthwith 

inform the Commission thereof. 

The  provisions  adopted  pursuant  to  this  paragraph  shall  contain  a  reference  to  this 

Directive  or  shall  be  accompanied  by  such  reference  at  the  time  of  their  official 

publication.  The procedure for such reference shall be adopted by the Member States. 

Article 3 

This Directive shall enter into force on the twentieth day following that of its publication 

in the Official  Journal of the European Communities. 

Article 4 

This Directive is addressed to the Member States. 

Done at...,... 

For the Council 

The President 

ANNEX 

"ANNEX  I 

NATURAL HABITAT TYPES OF COMMUNITY INTEREST 
WHOSE CONSERVATION REQUIRES THE DESIGNATION OF SPECIAL AREAS OF CONSERVATION 

Interpretation 

Guidance on the interpretation  of habitat types is given in the 'Interpretation  Manual of European 
Union  Habitats'  as  approved  by  the  committee  set  up  in  Article  20  ('Habitats  Committee')  and 
published by the European Commission1. 
The code corresponds to the NATURA 2000 code. 
The sign '*' indicates priority habitat types. 

1. COASTAL  AND HALOPHYTIC  HABITATS 

11. Open  sea  and  tidal  areas 
1110 
1120 
1130 
1140 
1150 
1160 
1170 
1180 

Sandbanks which are slightly covered by sea water all the time 
• * Posidonia beds (Posidonion oceanicaé) 
Estuaries 
Mudflats and sandflats not covered by seawater at low tide 
* Coastal lagoons 
Large shallow inlets and bays 
Reefs 
Submarine structures made by leaking gases 

12. Sea  cliffs  and  shingle  or stony  beaches 
1210 
1220 
1230 
1240 
1250 

Annual vegetation of drift lines 
Perennial vegetation of stony banks 
Vegetated sea cliffs of the Atlantic and Baltic Coasts 
Vegetated sea cliffs of the Mediterranean coasts with endemic Limonium spp. 
Vegetated sea cliffs with endemic flora of the Macaronesian coasts 

13. Atlantic and  continental  salt  marshes  and  salt  meadows 
1310 
1320 
1330 
1340 

Salicornia and other annuals colonizing mud and sand 
Spartina swards (Spartinion maritimi) 
Atlantic salt meadows (Glauco-Puccinellietalia maritimae) 
* Inland salt meadows 

14. Mediterranean  and  thermo-Atlantic salt  marshes  and  salt  meadows 
1410 
1420 
1430 

Mediterranean salt meadows (Juncetalia maritimi) 
Mediterranean and thermo-Atlantic halophilous scrubs (Sarcocorneteajruticosi) 
Halo-nitrophilous scrubs (Pegano-Salsoleted) 

1 "Interpretation manual of European Union habitats, version EUR 15" adopted by the Habitats Committee on 25 April 
1996, European Commission, DGXI. 

15. Salt and gypsum inland steppes 
1510 
1520 
1530 

* Mediterranean salt steppes (Limonietalia) 
* Iberian gypsum vegetation (Gypsophiletalid) 
* Pannonic salt steppes and salt marshes 

16. Boreal Baltic archipelago, coastal and landupheaval areas 
1610 

Baltic esker islands with sandy, rocky and shingle beach vegetation and sublittoral 
vegetation 
Boreal Baltic islets and small islands 
* Boreal Baltic coastal meadows 
Boreal Baltic sandy beaches with perennial vegetation 
Boreal Baltic narrow inlets 

1620 
1630 
1640 
1650 

f 

2. COASTAL SAND DUNES AND INLAND DUNES 

21. Sea dunes of the Atlantic, North Sea and Baltic coasts 
2110 
2120 
2130 
2140 
2150 
2160 
2170 
2180 
2190 
21A0 

Embryonic shifting  dunes 
Shifting dunes along the shoreline with Ammophila  arenaria ('white dunes') 
* Fixed coastal dunes with herbaceous vegetation ('grey dunes') 
* Decalcified fixed dunes with Empetrum nigrum 
* Atlantic decalcified fixed dunes {Calluno-Ulicetea) 
Dunes with Hippophaë rhamnoides 
Dunes with Salix repens ssp. argentea (Salicion arenariae)i 
Wooded dunes of the Atlantic, Continental and Boreal region 
Humid dune slacks 
Machairs (* in Ireland) 

22. Sea dunes of the Mediterranean coast 
2210 
2220 
2230 
2240 
2250 
2260 
2270 

Crucianellion maritimae fixed beach dunes 
Dunes with Euphorbia terracina 
Malcolmietalia  dune grasslands 
Brachypodietalia  dune grasslands with annuals 
* Coastal dunes with Juniperus spp. 
Cisto-Lavenduletalia dune sclerophyllous scrubs 
* Wooded dunes with Pinuspinea  and/or  Pinuspinaster 

23. Inland dunes, old and decalcified 
2310 
2320 
2330 
2340 

Dry sand heaths with Calluna and Genista 
Dry sand heaths with Calluna and Empetrum  nigrum 
Inland dunes with open Corynephorus and Agrostis  grasslands 
* Pannonic inland dunes 

t 

3. FRESHWATER HABITATS 

31. Standing water 
3110 

Oligotrophic waters containing very few minerals of sandy plains 
(Littorelletalia uniflorae) 
Oligothrophic waters containing very few minerals generally on sandy soils of the West 
Mediterranean, with Isoetes spp. 
Oligotrophic to mesotrophic standing waters with vegetation of the Littorelletea 
uniflorae and/or of the Isoëto-Nanojuncetea 
Hard oligo-mesotrophic waters with benthic vegetation of Char a spp. 
Natural eutrophic lakes with Magnopotamion or Hydrocharition  - type vegetation 
Natural dystrophic lakes and ponds 
* Mediterranean temporary ponds 
*Turloughs 

3120 

3130 

3140 
3150 
3160 
3170 
3180 

32.  Running  water  -  sections  of  water  courses  with  natural  or  semi-natural  dynamics 
(minor, average and major beds) where the water  quality  shows  no 
significant  deterioration 

3210 
3220 
3230 
3240 
3250 
3260 

3270 
3280 

3290 

Fennoscandian natural rivers 
Alpine rivers and the herbaceous vegetation along their banks 
Alpine rivers and their ligneous vegetation with Myricaria  germanica 
Alpine rivers and their ligneous vegetation with Salix elaeagnos 
Constantly flowing Mediterranean rivers with Glaucium flavum 
Water courses of plain to montane levels with the Ranunculion fluitantis and 
Callitricho-Batrachion vegetation 
Rivers with muddy banks with Chenopodion rubri p.p. and Bidention p.p. vegetation 
Constantly flowing  Mediterranean rivers with Paspalo-Agrostidion  species and 
hanging curtains of  Salix and  Populus alba 
Intermittently flowing Mediterranean rivers of the  Paspalo-Agrostidion 

4. TEMPERATE HEATH AND SCRUB 

4010 
4020 
4030 
4040 
4050 
4060 
4070 

4080 
4090 

Northern Atlantic wet heaths with Erica tetralix 
* Temperate Atlantic wet heaths with Erica ciliaris and Erica tetralix 
European dry heaths 
* Dry Atlantic coastal heaths with Erica vagans 
* Endemic macaronesian heaths 
Alpine and Boreal heaths 
* Bushes with Pinus mugo and Rhododendron hirsutum (Mugo-Rhododendretum 
hirsuti) 
Sub-Arctic Salix spp. scrub 
Endemic oro-Mediterranean heaths with gorse 

5. SCLEROPHYLLOUS  SCRUB  (MATORRAL) 

51.  Sub-Mediterranean  and  temperate  scrub 
5110 

Stable xerothermophilous formations with Buxus sempervirens on rock slopes 
(Berberidion p.p.) 
Mountain Cytisus purgans formations 
Juniperus communis formations on heaths or calcareous grasslands 
* Cistus palhinhae formations on maritime wet heaths 

5120 
5130 
5140 

52. Mediterranean  arborescent  matorral 
5210 
5220 
5230 

Arborescent matorral with Juniperus spp. 
* Arborescent matorral with Zyziphus 
* Arborescent matorral with Laurus nobilis 

53. Thermo-Mediterranean  and  pre-steppe  brush 
5310 
5320 
5330 

Laurus nobilis thickets 
Low formations of Euphorbia close to cliffs 
Thermo-Mediterranean and pre-desert scrub 

54.  Phrygana 
5410 
5420 
5430 

West Mediterranean clifftop phryganas (Astragalo-Plantaginetum subulatae) 
Sarcopoterium spinosum phryganas 
Endemic phryganas of the Euphorbio- Verbascion 

10 

6.  N A T U R AL  A ND  S E M I - N A T U R AL  G R A S S L A ND  F O R M A T I O NS 

61.  Natural  grasslands 
6110 
6120 
6130 
6140 
6150 
6160 
6170 
6180 

* Rupicolous calcareous or basophilic grasslands of the Alysso-Sedion albi 
* Xeric sand calcareous grasslands 
Calaminarian grasslands of the  Violetalia calaminariae 
. Siliceous Pyrenean Festuca eskia grasslands 
Siliceous alpine and boreal grasslands 
Oro-Iberian Festuca indigesta grasslands 
Alpine and subalpine calcareous grasslands 
Macaronesian mesophile grasslands 

62.  Semi-natural  dry  grasslands  and  scrubland  faciès 
6210 

Semi-natural dry grasslands and scrubland facies on calcareous substrates (Festuco-
Brometalia) (* important orchid sites) 
* Pseudo-steppe with grasses and annuals of the Thero-Brachypodietea 
* Species-rich Nor dus grasslands, on silicious substrates in mountain areas (and 
submountain areas in Continental Europe) 
* Sub-Pannonic steppic grasslands 
* Pannonic loess steppic grasslands 
* Pannonic sand steppes 
* Fennoscandian lowland species-rich dry to mesic grasslands 
* Nordic alvar and precambrian calcareous flatrocks 

6220 
6230 

6240 
6250 
6260 
6270 
6280 

63.  Sclerophillous  grazed  forests  (dehesas) 
Dehesas with evergreen Quercus spp. 
6310 

64.  Semi-natural  tall-herb  humid  meadows 
6410 
6420 
6430 
6440 
6450 

Molinia meadows on calcareous, peaty or clayey-silt-ladensoils (Molinion caeruleae) 
Mediterranean tall humid grasslands of the Molinio-Holoschoenion 
Hydrophilous tall herb fringe communities of plains and of the montane to alpine levels 
Alluvial meadows of river valleys of the Cnidion dubii 
Northern boreal alluvial meadows 

65.  Mesophile  grasslands 
6510 
6520 
6530 

Lowland hay meadows (Alopecurus pratensis, Sanguisorba officinalis) 
Mountain hay meadows 
* Fennoscandian wooded meadows 

<\ 

7. RAISED  BOGS AND MIRES  AND  FENS 

71.  Sphagnum  acid  bogs 
7110 
7120 
7130 
7140 
7150 
7160 

* Active raised bogs 
Degraded raised bogs still capable of natural regeneration 
Blanket bogs (* if active bog) 
Transition mires and quaking bogs 
Depressions on peat substrates of the Rhynchosporion 
Fennoscandian mineral-rich springs and springfens 

72.  Calcareous  fens 
7210 
7220 
7230 
7240 

* Calcareous fens with Cladium mariscus and species of the Caricion davallianae 
* Petrifying springs with tufa formation (Cratoneurion) 
Alkaline fens 
* Alpine pioneer formations of the Caricion bicoloris-atrofuscae 

73. Boreal  mires 
7310 
7320 

* Aapa mires 
* Palsa mires 

8. ROCKY  HABITATS  AND  CAVES 

81.  Scree 
8110 

8120 

8130 
8140 
8150 
8160 

Siliceous  scree  of  the  montane  to  snow  levels  (Androsacetalia  alpinae  and 
Galeopsietalia ladani) 
Calcareous  and  calcshist  screes  of  the  montane  to  alpine  levels (Thlaspietea 
rotundifolii) 
Western Mediterranean and thermophilous scree 
Eastern Mediterranean screes 
Medio-European upland siliceous screes 
* Medio-European calcareous scree of hill and montane levels 

82. Rocky  slopes with  chasmophytic  vegetation 
8210 
8220 
8230 

Calcareous rocky slopes with chasmophytic vegetation 
Siliceous rocky slopes with chasmophytic vegetation 
Siliceous  rock  with  pioneer  vegetation  of  the  Sedo-Scleranthion or  of  the 
Sedo albi-Veronicion dillenii 
* Limestone pavements 

8240 

83.  Other  rocky  habitats 
8310 
8320 
8330 
8340 

Caves not open to the public 
Fields of lava and natural excavations 
Submerged or partially submerged sea caves 
Permanent glaciers 

U 

9. FORESTS 
(Sub)natural woodland vegetation  comprising native species forming forests  of tall trees, with 
typical  undergrowth,  and  meeting  the  following  criteria:  rare  or  residual,  and/or  hosting 
species of Community  interest 

90.  Forests  of  Boreal  Europe 
9010 
9020 

* Western Taïga 
*  Fennoscandian  hemiboreal  natural  old  broad-leaved  deciduous  forests  (Quercus, 
Tilia, Acer, Fraxinus or Ulmus) rich in epiphytes 
* Natural forests of primary succession stages of landupheaval  coast 
Nordic subalpine/subarctic forests with Bêtula pubescens  ssp. czerepanovii 
Fennoscandian herb-rich forests with Picea abies 
Coniferous  forests on, or connected to,-glaciofluvial  eskers 
Fennoscandian wooded pastures 
* Fennoscandian deciduous swamp woods 

91. Forests of Temperate Europe 
9110 
9120 

Luzulo-Fagetum  beech  forests 
Atlantic  acidophilous  beech  forests  with  Ilex  and  sometimes  also  Taxus  in  the 
shrublayer (Quercion robori-petraeae or Hici-Fagenion) 
Asperulo-Fagetum  beech  forests 
Medio-European subalpine beech woods with Acer and Rumex arifolius 
Medio-European limestone beech forests of the  Cephalanthero-Fagion 
Sub-Atlantic and medio-European oak or oak-hornbeam forests of the Carpinion betuli 
Galio-Carpinetum oak-hornbeam  forests 
* Tilio-Acerion forests of slopes, screes and ravines 
Old acidophilous oak woods with Quercus robur on sandy plains 
Old sessile oak woods with Ilex and Blechnum in the British Isles 
Thermophilous Fraxinus angustifolia woods 
* Caledonian  forest 
* Bog woodland 
* Alluvial forests with Alnus glutinosa and Fraxinus excelsior (Alno-Padion, Alnion 
incanae, Salicion albae) 
Riparian mixed forests of Quercus robur, Ulmus laevis and Ulmus minor, Fraxinus 
excelsior or Fraxinus angustifolia, along the great rivers (Ulmenion minoris) 
* Pannonic woods with Quercus petraea  and Carpinus betulus 
* Pannonian woods with Quercus pubescens 
* Euro-Siberian steppic woods with Quercus spp. 
* Taxus baccata woods of the British Isles 

9030 
9040 
9050 
9060 
9070 
9080 

9130 
9140 
9150 
9160 
9170 
9180 
9190 
91A0 
91B0 
91 CO 
91D0 
91E0 

91F0 

91 GO 
91 HO 
9110 
91 JO 

92.  Mediterranean  deciduous  forests 
9210 
9220 
9230 
9240 
9250 
9260 
9270 

* Apeninne beech forests with Taxus and Ilex 
* Apennine beech forests with Abies alba and beech forests with Abies nebrodensis 
Galicio-Portuguese oak woods with Quercus robur and Quercus pyrenaica 
Quercus faginea  and Quercus canariensis Iberian woods 
Quercus trojana woods 
Castanea sativa woods 
Hellenic beech forests with Abies  borisii-regis 

<1 

9280 
9290 
92A0 
92B0 

92C0 
92D0 

Quercus frainetto woods 
Cupressus forests (Acero-Cupression) 
Salix alba and Populus alba galleries 
Riparian formations on intermittent Mediterranean water courses with Rhododendron 
ponticum, Salix and others 
Platanus orientalis and Liquidambar orientalis woods (Platanion orientalis) 
Southern riparian galleries and thickets (Nerio-Tamaricetea and Securinegion 
tinctoriae) 

93. Mediterranean  sclerophyllous  forests 
9310 
9320 
9330 
9340 
9350 
9360 
9370 
9380 

Aegean Quercus brachyphylla woods 
Olea and Ceratonia forests 
Quercus suber forests 
Quercus ilex and Quercus rotundifolia forests 
Quercus macrolepis forests 
* Macaronesian laurel forests (Laurus,  Ocotea) 
* Palm groves of Phoenix 
Forests of Hex aquifolium 

94. Temperate mountainous coniferous forests 
9410 
9420 
9430 

Acidophilous Picea forests of the montane to alpine levels (Vaccinio-Piceetea) 
Alpine Larix decidua and/or Pinus cembra forests 
Subalpine and montane Pinus uncinata forests (* if oh gypsum or limestone) 

95. Mediterranean and Macaronesian mountainous coniferous forests 
9510 
9520 
9530 
9540 
9550 
9560 
9570 
9580 

* Southern Apennine Abies alba forests 
Abies pinsapo forests 
* (Sub-) Mediterranean pine forests with endemic black pines 
Mediterranean pine forests with endemic Mesogean pines 
Canarian endemic pine forests 
* Endemic forests with Juniperus spp. 
* Tetraclinis articulata forests 
* Mediterranean Taxus baccata woods 

n 

ANNEX II 

ANIMAL AND PLANT SPECIES OF COMMUNITY INTEREST WHOSE 
CONSERVATION REQUIRES THE DESIGNATION OF SPECIAL AREAS OF 
CONSERVATION 

Interpretation 

(a) 

(b) 

Annex II follows  on from Annex I for  the establishment  of a consistent network of special 
areas of conservation. 
The species listed in this Annex are indicated : 

(c) 

by the name of the species or subspecies, or 
by  the  all  the  species  belonging  to  a  higher  taxon  or  to  a  designated  part  of  that 
taxon. 

The  abbreviation  'spp.'  after  the  name  of  a  family  or  genus  designates  all  the  species 
belonging to that family or genus. 
Symbols 
An asterisk (*) before the name of a species indicates that it is a priority species. 
Most species listed in this Annex are also listed in Annex IV. 
Where a species appears in this Annex but does not appear in either Annex IV or Annex V, 
the  species  name  is  followed  by  the  symbol  (o);  where  a  species  which  appears  in  this 
Annex also appears in Annex V but does not appear  in Annex IV,  its name is followed by 
the symbol (V). 

(a) ANIMALS 

VERTEBRATES 

MAMMALS 
INSECTIVORA 
Talpidae 

Galemys pyrenaicus 

CHIROPTERA 

Rhinolophidae 

Rhinolophus blasii 
Rhinolophus euryale 
Rhinolophus  ferrumequinum 
Rhinolophus hipposideros 
Rhinolophus mehelyi 

Vespertilionidae 

Barbastella barbastellus 
Miniopterus schreibersi 
Myotis bechsteini 
Myotis blythii 
Myotis capaccinii 
Myotis dasycneme 

K 

Myotis emarginatus 
Myotis myotis 

RODENTIA 

Sciuridae 

* Pteromys volans (Sciuropterus russicus) 
Spermophilus citellus (Citellus citellus) 

Castoridae 

Castor fiber (except the Finnish and Swedish populations) 

Microtidae 

Microtus cabrerae 
* Microtus oeconomus arenicola 

CARNIVORA 

Canidae 

*  Alopex lagopus 
* Canis lupus  (Spanish  populations:  only  those  south  of  the  Duero;  Greek 
populations:  only  south  of  the  39th  parallel;  Finnish  populations 
excepted). 

Ursidae 

* Ursus arctos (except the Finnish and Swedish populations) 

Mustelidae 

* Gulo gulo 
Lutra lutra 
Mustela lutreola 

Felidae 

Lynx lynx (except the Finnish populations) 
* Lynx pardinus 

Phocidae 

Halichoerus grypus (V) 
* Monachus monachus 
Phoca hispida bottnica (o) 
* Phoca hispida saimensis 
Phoca vitulina (V) 

ARTIODACTYLA 

Cervidae 

*  Cervus elaphus corsicanus 
Rangifer tarandus fennicus (o) 

Bovidae 

Capra aegagrus (natural populations) 
* Capra pyrenaica pyrenaica 
Ovis gmelini musimon (Ovis ammon musimon) (natural populations - Corsica 
and Sardinia) 
* Rupicapra pyrenaica ornata (Rupicapra rupicapra ornata) 
Rupicapra rupicapra balcanica 

CETACEA 

Tursiops truncatus 
Phocoena phocoena 

a 

REPTILES 
CHELONIA (TESTUDINES) 
Tesîudinidae 

Testudo hermanni 
Testudo graeca 
Testudo marginata 

Cheloniidae 

* Caretta caretta 

Emydidae 

Emys orbicularis 
Mauremys caspica 
Mauremys leprosa 

SAURIA 

Lacertidae 

Lacerta bonnali (Lacerta monticola) 
Lacerta monticola 
Lacerta schreiberi 
Gallotia galloti insulanagae 
* Gallotia simonyi 
Podarcis lilfordi 
Podarcis pityusensis 

Scincidae 

Chalcides simonyi (Chalcides occidentalis) 

Gekkonidae 

Phyllodactylus europaeus 

OPHIDIA (SERPENTES) 
Colubridae 

Elaphe quatuorlineata 
Elaphe situla 

Viperidae 

* Macrovipera schweizeri (Vipera lebetina schweizeri) 
Vipera ursinii 

AMPHIBIANS 
CAUDATA 

Salamandridae 

Chioglossa lusitanica 
Mertensiella luschani (Salamandra luschani) 
* Salamandra atra aurorae 
Salamandrina terdigitata 
Triturus carnifex (Triturus cristatus carnifex) 
Triturus cristatus (Triturus cristatus cristatus) 
Triturus dobrogicus (Triturus cristatus dobrogicus) 
Triturus karelinii (Triturus cristatus karelinii) 

Proteidae 

Proteus anguinus 

Plethodontidae 

Hydromantes (Speleomantes)  ambrosii 
Hydromantes (Speleomantes)  flavus 

n 

Hydromantes (Speleomantes)  genei 
Hydromantes (Speleomantes)  imperialis 
Hydromantes (Speleomantes)  strinatii 
Hydromantes (Speleomantes)  supramontes 

ANURA 

Discoglossidae 

Bombina bombina 
Bombina variegata 
Discoglossus galganoi  (including Discoglossus "jeanneae") 
Discoglossus montalentii 
Discoglossus sardus 
* Alytes muletensis 

Ranidae 

Rana latastei 

Pelobatidae 

* Pelobates fiiscus insubricus 

FISHES 
PETROMYZONIFORMES 

Petromyzonidae 

Eudontomyzon spp. (o) 
Lampetra fluviatilis  (V) (except the Finnish and Swedish populations) 
Lampetra planeri (o) (except the Finnish and Swedish populations) 
Lethenteron zanandreai (V) 
Petromyzon marinus (o) (except the Swedish populations) 

ACIPENSERIFORMES 

Acipenseridae 

* Acipenser naccarii 
* Acipenser sturio 

CLUPEIFORMES 
Clupeidae 

Alosa spp. (V) 

SALMONIFORMES 

Salmonidae 

Hucho hucho (natural populations) (V) 
Salmo salar (only in fresh water) (V) (except the Finnish populations) 
Sahno marmoratus (o) 
Salmo macrostigma (o) 

Coregonidae 

* Coregonus oxyrhynchus (anadromous populations in certain sectors of the 
North Sea) 

CYPRINIFORMES 
Cyprinidae 

Alburnus albidus (o) (Alburnus vulturius) 
Anaecypris hispanica 
Aspius aspius (o) (except the Finnish populations) 
Barbus plebejus (V) 
Barbus meridionalis (V) 
Barbus comiza (V) 

f& 

Chalcalburnus chalcoides (o) 
Chondrostoma soetta (o) 
Chondrostoma polylepis (o) (including C. willkommi) 
Chondrostoma genei (o) 
Chondrostoma lusitanicum (o) 
Chondrostoma toxostoma (o) 
Gobio albipinnatus (o) 
Gobio uranoscopus (o) 
Iberocypris palaciosi (o) 
*Ladigesocypris ghigii (o) 
Leuciscus lucumonis (o) 
Leuciscus souffia (o) 
Phoxinellus spp. (o) 
Rutilus pigus (o) 
Rutilus rubilio (o) 
Rutilus arcasii (o) 
Rutilus macrolepidotus (o) 
Rutilus lenMningii (o) 
Rutilus frisii meidingeri (o) 
Rutilus alburnoides (o) 
Rhodeus sericeus amarus (o) 
Scardinius graecus (o) 

Cobitidae 

Cobitis trichonica (o) 
Cobitis taenia (o) (except the Finnish populations) 
Misguraus fossilis  (o) 
Sabanejewia aurata (o) 
Sabanejewia larvata (o) (Cobitis larvata and Cobitis conspersa) 

SILURIFORMES 
Siluridae 

Silurus aristotelis (V) 

ATHERINIFORMES 

Cyprinodontidae 

Aphanius iberus (o) 
Aphanius fasciatus (o) 
* Valencia hispanica 
* Valencia letourneuxi (Valencia hispanica) 

PERCIFORMES 
Percidae 

Gymnocephalus schraetzer (V) 
Zingel spp. [(o) except Zingel asper and Zingel zingel (V)] 

Gobiidae 

Pomatoschistus canestrini (o) 
Knipowitschia (Padogobius) panizzae (o) 
Padogobius nigricans (o) 

SCORPAENIFORMES 
Cottidae 

Cottus petiti (o) 
Cottus gobio (o) (except the Finnish populations) 

fS 

INVERTEBRATES 

ARTHROPODS 
CRUSTACEA 

Decapoda 

Austropotamobius pallipes (V) 

INSECTA 

Coleoptera 

Agathidium pulchellum (o) 
Boros schneideri (o) 
Buprestis splendens 
*Carabus menetriesi pacholei (o) 
* Carabus olympiae 
Cerambyx cerdo 
Corticaria planula (o) 
Cucujus cinnaberinus 
Dytiscus latissimus 
Graphoderus bilineatus 
Limoniscus violaceus (o) 
Lucanus cervus (o) 
Macroplea pubipennis (o) 
Mesosa myops (o) 
Morimus funereus (o) 
*Osmoderma eremita 
Oxyporus mannerheimii (o) 
Pytho kolwensis (o) 
•Rosalia alpina 
Stephanopachys linearis (o) 
Stephanopachys substriatus (o) 
Xyletinus tremulicola (o) 

Hemiptera 

Aradus angularis (o) 

Lepidoptera 

Agriades glandon aquilo (o) 
*Callimorpha (Euplagia, Panaxia) quadripunctaria (o) 
Clossiana improba (o) 
Coenonympha oedippus 
Erebia calcaria 
Erebia christi 
Erebia medusa polaris (o) 
Eriogaster catax 
Euphydryas (Eurodryas, Hypodryas) aurinia (o) 
Graellsia isabellae (V) 
Hesperia comma catena (o) 
Hypodryas maturna 
Lycaena dispar 
Maculinea nausithous 
Maculinea teleius 

«ào 

Melanargia arge 
Papilio hospiton 
Plebicula golgus 
Xestia borealis (o) 
Xestia brunneopicta (o) 

Mantodea 

Apteromantis aptera 

Odonata 

Coenagrion hylas (o) 
Coenagrion mercuriale (o) 
Cordulegaster trinacriae 
Gomphus graslinii 
I^uconhinia pectoralis 
Lindenia tetraphylla 
Macromia splendens 
Ophiogomphus cecilia 
Oxygastra curtisii 

Orthoptera 

Baetica ustulata 

ARACHNIDA 

Pseudoscorpiones 

Anthrenochernes stellae (o) 

MOLLUSCS 
GASTROPODA 

Caseolus calculus 
Caseolus commixta 
Caseolus sphaerula 
Discula leacockiana 
Discula tabellata 
Discus guerinianus 
Elona quimperiana 
Geomalacus maculosus 
Geomitra moniziana 
•Helicopsis striata austriaca (o) 
Idiomela (Helix) subplicata 
Leiostyla abbreviata 
Leiostyla cassida 
Leiostyla corneocostata 
Leiostyla gibba 
Leiostyla lamellosa 
Vertigo angustior (o) 
Vertigo genesii (o) 
Vertigo geyeri  (o) 
Vertigo moulinsiana (o) 

BIVALVIA 

Unionoida 

Margaritifera  durrovensis (Margaritifera  margaritifera) (V) 
Margaritifera  margaritifera (V) 

*1 

Unio crassus 

(b) PLANTS 

PTERIDOPHYTA 
ASPLENIACEAE 

Asplenium jahandiezii (Litard.) Rouy 

BLECHNACEAE 

Woodwardia radicans (L.) Sm. 

DICKSONIACEAE 

Culcita macrocarpa C. Presl 

DRYOPTERIDACEAE 

Diplazium sibiricum (Turcz. ex Kunze) Kurata 
* Dryopteris corleyi Fraser-Jenk. 
Dryopteris fragans (L.) Schott 

HYMENOPHYLLACEAE 

Trichomanes speciosum Willd. 

ISOETACEAE 

Isoetes boryana Durieu 
Isoetes malinverniana Ces. & De Not. 

MARSILEACEAE 

Marsilea batardae Lauiiert 
Marsilea quadrifolia L. 
Marsilea strigosa Willd. 

OPHIOGLOSSACEAE 

Botrychium simplex Hitchc. 
Ophioglossum polyphyllum A. Braun 

GYMNOSPERMAE 
PINACEAE 

* Abies nebrodensis (Lojac.) Mattei 

ANGIOSPERMAE 
ALISMATACEAE 

* Alisma wahlenbergii (Holmberg) Juz. 
Caldesia parnassifolia  (L.) Pari. 
Luronium natans (L.) Raf. 

AMARYLLIDACEAE 

Leucojum nicaeense Ard. 
Narcissus asturiensis (Jordan) Pugsley 
Narcissus calcicola Mendonça 
Narcissus cyclamineus DC. 
Narcissus fernandesii G. Pedro 
Narcissus humilis (Cav.) Traub 
* Narcissus nevadensis Pugsley 
Narcissus pseudonarcissus L. subsp. nobilis (Haw.) A. Femandes 
Narcissus scaberulus Henriq. 

22_ 

Narcissus triandrus L. subsp. capax (Salisb.) D. A. Webb. 
Narcissus viridiflorus  Schousboe 

BORAGINACEAE 

* Anchusa crispa Viv. 
* Lithodora nitida (H. Ern) R. Femandes 
Myosotis lusitanica Schuster 
Myosotis rehsteineri Wartm. 
Myosotis retusifolia R.  Afonso 
Omphalodes kuzinskyanae Willk. 
* Omphalodes littoralis Lehm. 
Solenanthus albanicus (Degen & al.) Degen & Baldacci 
* Symphytum cycladense Pawl. 

CAMPANULACEAE 

Asyneuma giganteum (Boiss.) Bornm. 
* Campanula sabatia De Not. 
Jasione crispa (Pourret) Samp, subsp. serpentinica Pinto da Silva 
Jasione lusitanica A. DC. 

CARYOPHYLLACEAE 

Arenaria ciliata L.  ssp. pseudofrigida Ostenf. & O.C. Dahl 
Arenaria humifusa Wahlenberg 
* Arenaria nevadensis Boiss.  & Reuter 
Arenaria provincialis Chater & Halliday 
Dianthus arenarius L. subsp. arenarius 
Dianthus cintranus Boiss. & Reuter subsp. cintranus Boiss. & Reuter 
Dianthus marizii (Samp.) Samp. 
Dianthus rupicola Biv. 
* Gypsophila papulosa P. Porta 
Herniaria algarvica Chaudhri 
* Herniaria latifolia Lapeyr.  subsp. litardierei Garnis 
Herniaria lusitanica (Chaudhri) subsp. berlengiana Chaudhri 
Herniaria maritima Link 
Moehringia laterifolia (L.) Fenzl. 
Moehringia tommasinii Marches. 
Petrocoptis grandiflora Rothm. 
Petrocoptis montsicciana O. Bolos & Rivas Mart. 
Petrocoptis pseudoviscosa Fernandez Casas 
Silène furcata Rafm.  ssp. angustiflora  (Rupr.) Walters 
* Silène hicesiae Brullo & Signorello 
Silène hifacensis Rouy ex Willk. 
* Silène holzmanii Heldr. ex Boiss. 
Silène longicilia (Brot.) Otth. 
Silène mariana Pau 
* Silène orphanidis Boiss. 
* Silène rothmaleri Pinto da Silva 
* Silène velutina Pourret ex Loisel. 

CHENOPODIACEAE 

* Bassia (Kochia) saxicola (Guss.) A.  J. Scott 
* Salicornia veneta Pignatti & Lausi 

23> 

CISTACEAE 

Cistus palhinhae Ingram 
Halimium verticillatum (Brot.) Sennen 
Helianthemum alypoides Losa & Rivas Goday 
Helianthemum caput-felis Boiss. 
* Tuberaria major (Willk.) Pinto da Silva & Rozeira 

COMPOSITAE 

* Anthémis glaberrima (Rech. f.) Greuter 
Artemisia campestris L. subsp. bottnica A.N.  Lundstrom ex Kindb. 
* Artemisia granatensis Boiss. 
* Artemisia laciniata Willd. 
Artemisia oelandica (Besser) Komaror 
* Artemisia pancicii (Janka) Ronn. 
* Aster pyrenaeus Desf. ex DC 
* Aster sorrentinii (Tod) Lojac. 
* Carduus myriacanthus Salzm. ex DC. 
* Centaurea alba L. subsp. heldreichii (Halacsy) Dostal 
* Centaurea alba L. subsp. princeps (Boiss. & Heldr.) Gugler 
* Centaurea attica Nyman subsp. megarensis (Halacsy & Hayek) Dostal 
* Centaurea balearica J. D. Rodriguez 
* Centaurea borjae Valdes-Berm. & Rivas Goday 
* Centaurea citricolor Font Quer 
Centaurea corymbosa Pourret 
Centaurea gadorensis G. Blanca 
* Centaurea horrida Badaro 
* Centaurea kalambakensis Freyn & Sint. 
Centaurea kartschiana Scop. 
* Centaurea lactiflora Halacsy 
Centaurea micrantha Hoffmanns.  & Link subsp. herminii (Rouy) Dostal 
* Centaurea niederi Heldr. 
* Centaurea peucedanifolia Boiss. & Orph. 
* Centaurea pinnata Pau 
Centaurea pulvinata (G. Blanca) G. Blanca 
Centaurea rothmalerana (Arènes) Dostal 
Centaurea vicentina Mariz 
* Crépis crocifolia Boiss. & Heldr. 
Crépis granatensis (Willk.) B. Blanca & M. Cueto 
Crépis tectorum L.  subsp. nigrescens 
Erigeron frigidus Boiss. ex DC. 
Hymenostemma pseudanthemis (Kunze) Willd. 
* Jurinea cyanoides (L.) Reichenb. 
* Jurinea fontqueri Cuatrec. 
* Lamyropsis microcephala (Moris) Dittrich & Greuter 
Leontodon microcephalus (Boiss. ex DC.) Boiss. 
Leontodon boryi Boiss. 
* Leontodon siculus (Guss.) Finch & Sell 
Leuzea longifolia Hoffmanns.  & Link 
Ligularia sibirica (L.) Cass. 
Santolina impressa Hoffmanns.  & Link 

^<r 

Santolina semidentata Hoffmanns.  & Link 
* Senecio elodes Boiss. ex DC. 
Senecio jacobea L. subsp. gotlandicus (Neuman) Sterner 
Senecio nevadensis Boiss. & Reuter 

CONVOLVULACEAE 

* Convolvulus argyrothamnus Greuter 
* Convolvulus fernandesii Pinto da Silva & Teles 

CRUCIFERAE 

Alyssum pyrenaicum Lapeyr. 
Arabis sadina (Samp.) P. Cout. 
* Biscutella neustriaca Bonnet 
Biscutella vincentina (Samp.) Rothm. 
Boleum asperum (Pers.) Des vaux 
Brassica glabrescens Poldini 
Brassica insularis Moris 
* Brassica macrocarpa Guss. 
Braya linearis Rouy 
* Coincya rupestris Rouy 
* Coronopus navasii Pau 
Diplotaxis ibicensis (Pau) Gomez-Campo 
* Diplotaxis siettiana Maire 
Diplotaxis vicentina (P. Cout.) Rothm. 
Draba cacuminum Elis Ekman 
Draba cinerea Adams 
Erucastrum palustre (Pirona) Vis. 
* Iberis arbuscula Runemark 
Iberis procumbens Lange subsp. microcarpa Franco & Pinto da Silva 
* Jonopsidium acaule (Desf.) Reichenb. 
Jonopsidium savianum (Caruel) Ball ex Arcang. 
Rhynchosinapis erucastrum (L.) Dandy ex Clapham subsp. cintrana (Coutinho) 

Franco & P.  Silva (Coincya cintrana (P. Cout.) Pinto da Silva) 

Sisymbrium cavanillesianum Valdes & Castroviejo 
Sisymbrium supinum L. 

CYPERACEAE 

Carex holostoma Drejer 
* Carex panormitana Guss. 
Eleocnaris carniolica Koch 

DIOSCOREACEAE 

* Borderea chouardii (Gaussen) Heslot 

DROSERACEAE 

Aldrovanda vesiculosa L. 

EUPHORBIACEAE 

* Euphorbia margalidiana Kuhbier & Lewejohann 
Eurphorbia transtagana Boiss. 

GENTIANACEAE 

* Centaurium rigualii Esteve 
* Centaurium somedanum Lainz 
Gentiana ligustica R. de Vilm. & Chopinet 
Gentianella angelica (Pugsley) E. F. Warburg 

2S 

GERANIACEAE 

* Erodium astragaloides Boiss. & Reuter 
Erodium paularense Fernandez-Gonzalez & Izco 
* Erodium rupicola Boiss. 

GLOBULARIACEAE 

•Globularia stygia Orph. ex Boiss. 

GRAMINEAE 

Arctagrostis latifolia (R. Br.) Griseb. 
Arctophila fulva (Trin.) N. J. Anderson 
Avenula hackelii (Henriq.) Holub 
Bromus grossus Desf.  ex DC. 
Calamagrostis chalybaea (Laest.) Fries 
Cinna latifolia (Trev.) Griseb. 
Coleanthus subtilis (Tratt.) Seidl 
Festuca brigantina (Markgr.-Dannenb.) Markgr.-Dannenb. 
Festuca duriotagana Franco & R.  Afonso 
Festuca elegans Boiss. 
Festuca henriquesii Hack. 
Festuca summilusitanica Franco & R.  Afonso 
Gaudinia hispanica Stace & Turin 
Holcus setiglumis Boiss. & Reuter subsp. duriensis Pinto da Silva 
Micropyropsis tuberosa Romero - Zarco & Cabezudo 
Pseudarrhenatherum pallens (Link) J. Holub 
Puccinellia phryganodes (Trin.) Scribner  +  Merr. 
Puccinellia pungens (Pau) Paunero 
* Stipa austroitalica Martinovsky 
* Stipa bavarica Martinovsky & H. Scholz 
* Stipa styriaca Martinovsky 
* Stipa veneta Moraldo 
Trisetum subalpestre (Hartman) Neuman 

GROSSULARIACEAE 

* Ribes sardoum Martelli 

HIPPURIDACEAE 

Hippuris tetraphylla L. Fil. 

HYPERICACEAE 

* Hypericum aciferum (Greuter) N.K.B.  Robson 

JUNCACEAE 

Juncus valvatus Link 
Luzula arctica Blytt 

LABIATAE 

Dracocephalum austriacum L. 
* Micromeria taygetea P. H.  Davis 
Nepeta dirphya (Boiss.) Heldr. ex Halacsy 
* Nepeta sphaciotica P. H.  Davis 
Origanum dictamnus L. 
Sideritis incana subsp. glauca (Cav.) Malagarriga 
Sideritis javalambrensis Pau 
Sideritis serrata Cav. ex Lag. 
Teucrium lepicephalum Pau 

2fc 

Teucrium turredanum Losa & Rivas Goday 
* Thymus camphoratus Hoffmanns.  & Link 
Thymus carnosus Boiss. 
* Thymus lotocephalus G. Lôpez & R. Morales (Thymus cephalotos L.) 

LEGUMINOSAE 

Anthyllis hystrix Cardona, Contandr. & E. Sierra 
* Astragalus algarbiensis Coss. ex Bunge 
* Astragalus aquilanus Anzalone 
Astragalus centralpinus Braun-Blanquet 
* Astragalus maritimus Moris 
Astragalus tremolsianus Pau 
•Astragalus verrucosus Moris 
*Cytisus aeolicus Guss. ex Lindl. 
Genista dorycnifolia Font Quer 
Genista holopetala (Fleischm. ex Koch) Baldacci 
Melilotus segetalis (Brot.) Ser. subsp. fallax Franco 
•Ononis hackelii Lange 
Trifolium saxatile All. 
*Vicia bifoliolata J.D. Rodriguez 

LENTIBULARIACEAE 

Pinguicula nevadensis (Lindb.) Casper 

LILIACEAE 

Allium grosii Font Quer 
•Androcymbium rechingeri Greuter 
•Asphodelus bento-rainhae P. Silva 
Hyacinthoides vicentina (Hoffmans.  & Link) Rothm. 
•Muscari gussonei (Pari.) Tod. 

LINACEAE 

*Linum muelleri Moris (Linum maritimum muelleri) 

LYTHRACEAE 

*Lythrum flexuosum Lag. 

MALVACEAE 

Kosteletzkya pentacarpos (L.) Ledeb. 

NAJADACEAE 

Najas flexilis  (Willd.) Rostk.  & W.L. Schmidt 
Najas tenuissima (A. Braun) Magnus 

ORCHIDACEAE 

Calypso bulbosa L. 
*Cepnalanthera cucullata Boiss. & Heldr. 
Cypripedium calceolus L. 
Gymnigritella runei Teppner & Klein 
Liparis loeselii (L.) Rich. 
*Ophrys lunulata Pari. 
Platanthera obtusata (Pursh) subsp. oligantha (Turez.) Hulten 

PAEONIACEAE 

Paeonia cambessedesii (Willk.) Willk. 
Paeonia parnassica Tzanoudakis 
Paeonia clusii F.C.  Stem subsp. rhodia (Steam) Tzanoudakis 

2f 

PALMAE 

Phoenix theophrasti Greuter 

PAPAVERACEAE 

Corydalis gotlandica Lidén 
Papaver laestadianum (Nordh.) Nordh. 
Papaver radicatum Rottb. subsp. hyperboreum Nordh. 

PLANTAGINACEAE 

Plantago algarbiensis Sampaio (Plantago bracteosa (Willk.) G.  Sampaio) 
Plantago almogravensis Franco 

PLUMBAGINACEAE 

Armeria berlengensis Daveau 
•Armeria helodes Martini & Pold 
Armeria neglecta Girard 
Armeria pseudarmeria (Murray) Mansfeld 
•Armeria rouyana Daveau 
Armeria soleirolii (Duby) Godron 
Armeria velutina Welw. ex Boiss. & Reuter 
Limonium dodartii (Girard) O. Kuntze subsp. lusitanicum (Daveau) Franco 
*Limonium insulare (Beg. & Landi) Arrig. & Diana 
Limonium lanceolatum (Hoffmans.  & Link) Franco 
Limonium multiflorum Erben 
•Limonium pseudolaetum Arrig. & Diana 
* Limonium strictissimum (Salzmann) Arrig. 

POLYGONACEAE 

Persicaria foliosa (H. Lindb.) Kitag. 
Polygonum praelongum Coode & Cullen 
Rumex rupestris Le Gall 

PRIMULACEAE 

Androsace mathildae Levier 
Androsace pyrenaica Lam. 
•Primula apennina Widmer 
Primula nutans Georgi 
Primula palinuri Petagna 
Primula scandinavica Bruun 
Soldanella villosa Darracq. 

RANUNCULACEAE 

•Aconitum corsicum Gayer (Aconitum napellus subsp. corsicum) 
Adonis distorta Ten. 
Aquilegia bertolonii Schott 
Aquilegia kitaibelii Schott 
•Aquilegia pyrenaica D.C. subsp. cazorlensis (Heywood) Galiano 
•Consolida samia P.H.  Davis 
Pulsatilla patens (L.) Miller 
Ranunculus lapponicus L. 
Pulsatilla vulgaris Hill,  subsp. gotlandica (Johanss.) Zaemelis & Paegle 
•Ranunculus weyleri Mares 

RESEDACEAE 

•Reseda decursiva Forssk. 

2& 

ROSACEAE 

Agrimonia pilosa Ledebour 
Potentilla delphinensis Gren. & Godron 
Sorbus teodorii Liljefors 

RUBIACEAE 

•Galium litorale Guss. 
•Galium viridiflorum Boiss. & Reuter 

SALICACEAE 

Salix salvifolia Brot. subsp. australis Franco 

SANTALACEAE 

Thesium ebracteatum Hayne 

SAXIFRAGACEAE 

Saxifraga berica (Beguinot) D.A.  Webb 
Saxifraga florulenta Moretti 
Saxifraga hirculus L. 
Saxifraga osloensis Knaben 
Saxifraga tombeanensis Boiss. ex Engl. 

SCROPHULARIACEAE 

Antinhinum charidemi Lange 
Chaenorrhinum serpyllifolium (Lange) Lange 
subsp. lusitanicum R. Femandes 

•Euphrasia genargentea (Feoli) Diana 
Euphrasia marchesettii Wettst. ex Marches. 
Linaria algarviana Chav. 
Linaria coutinhoi Valdés 
•Linaria ficalhoana Rouy 
Linaria flava (Poiret) Desf. 
•Linaria hellenica Turrill 
•Linaria ricardoi Cout. 
•Linaria tursica B. Valdes & Cabezudo 
Linaria tonzigii Lona 
Odontites granatensis Boiss. 
Verbascum litigiosum Samp. 
Veronica micrantha Hoffmanns.  & Link 
•Veronica oetaea L.-A.  Gustavsson 

SOLANACEAE 

•Atropa baetica Willk. 

THYMELAEACEAE 

Daphne petraea Leybold 
•Daphne rodriguezii Texidor 

ULMACEAE 

Zelkova abelicea (Lam.) Boiss. 

UMBELLIFERAE 

•  Angelica heterocarpa Lloyd 
Angelica palustris (Besser) Hoffm. 
•  Apium bermejoi Llorens 
Apium repens (Jacq.) Lag. 
Athamanta cortiana Ferrarini 
•  Bupleurum capillare Boiss. & Heldr. 

24 

•  Bupleurum kakiskalae Greuter 
EryngiumalpinumL. 
•  Eryngium viviparum Gay 
•  Laserpitium longiradium Boiss. 
•  Naufraga  balearica Constans & Cannon 
•  Oenanthe conioides Lange 
Petagnia saniculifolia Guss. 
Rouya polygama (Desf.) Coincy 
•  Seseli intricatum Boiss. 
Thorella verticUlatinundata (Thore) Briq. 

VALERIANACEAE 

Centranthus trinervis (Viv.) Beguinot 

VIOLACEAE 

•  Viola hispida Lam. 
Viola jaubertiana Mares & Vigineix 
Viola rupestris F.W. Schmidt 

subsp. relicta Jalas 

Lower plants 

BRYOPHYTA 
Bruchia vogesiaca Schwaegr. (o) 
Bryhnia novae-angliae (Sull & Lesq.) Grout (o) 
•Bryoerythrophyllum  campylocarpum  (C.  Mull.)  Crum.  (Bryœrythrophyllum  machadoanum 
(Sergio) M. O. Hill)) (o) 
Buxbaumia viridis (Moug.) Moug. & Nestl. (o) 
Cephalozia macounii (Aust.) Aust. (o) 
Cynodontium suecicum (H. Am. & C. Jens.) I. Hag. (o) 
Dichelyma capillaceum (Dicks) Myr. (o) 
Dicranum viride (Sull. & Lesq.) Lindb. (o) 
Distichophyllum carinatum Dix. & Nich. (o) 
Drepanocladus (Hamatocaulis) verrucosus (Mitt.) Warnst. (o) 
Encalypta mutica  (I. Hagen)  (o) 
Hamatocaulis lapponicus (Norrl.) Hedenas (o) 
Herzogiella turfacea (Lindb.) I. Wats, (o) 
Hygrohypnum montanum (Lindb.) Broth, (o) 
Jungermannia handelii (Schiffh.)  Amak. (o) 
Mannia triandra (Scop.) Grolle (o) 
•  Marsupella profunda  Lindb. (o) 
Meesia longiseta Hedw. (o) 
Nothothylas orbicularis (Schwein.) Sull. (o) 
Orthothecium lapponicum (Schimp.) C. Hartm. (o) 
Orthotrichum rogeri Brid. (o) 
Petalophyllum ralfsii (Wils.) Nees & Gott. (o) 
Plagiomnium drummondii (Bruch & Schimp.) T. Kop.  (o) 
Riccia breidleri Jur. (o) 
Riella helicophylla (Bory & Mont.) Mont, (o) 
Scapania massolongi (K. Mull.) K. Mull, (o) 
Sphagnum pylaisii Brid. (o) 

Tayloria rudolphiana (Garov) B. & S. (o) 
Tortella rigens (N. Alberts) (o) 

SPECIES FOR MACARONESIA 

PTERIDOPHYTA 
HYMENOPHYLLACEAE 

Hymenophyllum maderensis Gibby & Lovis 

DRYOPTERIDACEAE 

• Polystichum drepanum (Sw.) C. Presl. 

ISOETACEAE 

Isoetes azorica Durieu & Paiva ex Milde 

MARSILEACEAE 

•  Marsilea azorica Launert & Paiva 

ANGIOSPERMAE 
ASCLEPIADACEAE 

Caralluma burchardii N. E. Brown 
•Ceropegia chrysantha Svent. 

BORAGINACEAE 

Echium candicans L. til. 
•  Echium gentianoides Webb & Coincy 
Myosotis azorica H. C. Watson 
Myosotis maritima Hochst. in Seub. 

CAMPANULACEAE 

•  Azorina vidalii (H. C. Watson) Feer 
Musschia aurea (L. f.) DC. 
•  Musschia wollastonii Lowe 

CAPRIFOLIACEAE 

•  Sambucus palmensis Link 

CARYOPHYLLACEAE 

Spergularia azorica (Kindb.) Lebel 

CELASTRACEAE 

Maytenus umbellata (R. Br.) Mabb. 

CHENOPODIACEAE 
Beta patula Ait. 

CISTACEAE 

Cistus chinamadensis Banares & Romero 
•  Helianthemum bystropogophyllum Svent. 

COMPOSITAE 

Andryala crithmifolia Ait. 
•  Argyranthemum lidii Humphries 
Argyranthemum thalassophylum (Svent.) Hump. 
Argyranthemum winterii (Svent.) Humphries 
•  Atractylis arbuscula Svent. & Michaelis 
Atractylis preauxiana Schultz. 
Calendula maderensis DC. 
Cheirolophus duranii (Burchard) Holub 

3>1 

Cheirolophus ghomerytus (Svent.) Holub 
Cheirolophus junonianus (Svent.) Holub 
Cheirolophus massonianus (Lowe) Hansen & Sund. 
Cirsium latifolium Lowe 
Helichrysum gossypinum Webb 
Helichrysum oligocephala (Svent. & Bzamw.) 
•  Lactuca watsoniana Trel. 
•  Onopordum nogalesii Svent. 
•  Onorpordum carduelinum Bolle 
•  Pericallis hadrosoma Svent. 
Phagnalon benettii Lowe 
Stemmacantha cynaroides (Chr. Son. in Buch) Ditt 
Sventenia bupleuroides Font Quer 
•  Tanacetum ptarmiciflorum Webb & Berth 

CONVOLVULACEAE 

•  Convolvulus caput-medusae Lowe 
•  Convolvulus lopez-socasii Svent. 
•  Convolvulus massonii A. Dietr. 

CRASSULACEAE 

Aeonium gomeraense Praeger 
Aeonium saundersii Bolle 
Aichryson dumosum (Lowe) Praeg. 
Monanthes wildpretii Banares & Scholz 
Sedum brissemoretii Raymond-Hamet 

CRUCIFERAE 

•  Crambe arborea Webb ex Christ 
Crambe laevigata DC. ex Christ 
•  Crambe sventenii R. Petters ex Bramwell & Sund. 
•  Parolinia schizogynoides Svent. 
Sinapidendron rupestre (Ait.) Lowe 

CYPERACEAE 

Carex malato-belizii Raymond 

DIPSACACEAE 

Scabiosa nitens Roemer & J. A.  Schultes 

ERICACEAE 

Erica scoparia L.  subsp. azorica (Hochst.) D. A. Webb 

EUPHORBIACEAE 

•  Euphorbia handiensis Burchard 
Euphorbia lambii Svent. 
Euphorbia stygiana H.  C. Watson 

GERANIACEAE 

•  Geranium maderense P. F.  Yeo 

GRAMINEAE 

Deschampsia maderensis (Haeck.  & Bom.) Buschm. 
Phalaris maderensis (Menezes) Menezes 

GLOBULARIACEAE 

•  Globularia ascanii D.  Bramwell & Kunkel 
•  Globularia sarcophylla Svent. 

3^ 

LABIATAE 

•  Sideritis cystosiphon Svent. 
•Sideritis discolor (Webb ex de Noe) Bolle 
Sideritis infernalis Bolle 
Sideritis marmorea Bolle 
Teucrium abutiloides L'Hér. 
Teucrium betonicum L'Hér. 

LEGUMINOSAE 

•  Anagyris latifolia Brouss. ex. Willd. 
Anthyllis lemanniana Lowe 
•  Dorycnium spectabile Webb & Berthel 
•  Lotus azoricus P. W. Ball 
Lotus callis-viridis D. Bramwell & D. H.  Davis 
•  Lotus kunkelii (E. Chueca) D. Bramwell & al. 
•  Teline rosmarinifolia Webb & Berthel. 
•  Teline salsoloides Arco & Acebes. 
Vicia dennesiana H.  C. Watson 

LILIACEAE 

•  Androcymbium psammophilum Svent. 
Scilla maderensis Menezes 
Semele maderensis Costa 

LORANTHACEAE 

Arceuthobium azoricum Wiens & Hawksw. 

MYRICACEAE 

•  Myrica rivas-martinezii Santos. 

OLEACEAE 

Jasminum azoricum L. 
Picconia azorica (Turin) Knobl. 

ORCHIDACEAE 

Goodyera macrophylla Lowe 

PITTOSPORACEAE 

•  Pittosporum coriaceum Dryand. ex. Ait. 

PLANTAGINACEAE 

Plantago malato-belizii Lawalree 

PLUMBAGINACEAE 

•  Limonium arborescens (Brouss.) Kuntze 
Limonium dendroides Svent. 
•  Limonium spectabile (Svent.) Kunkel & Sunding 
•  Limonium sventenii Santos & Fernandez Galvan 

POLYGONACEAE 

Rumex azoricus Rech. fil. 

RHAMNACEAE 

Frangula azorica Tutin 

ROSACEAE 

•  Bencomia brachystachya Svent. 
Bencomia sphaerocarpa Svent. 
•  Chamaemeles coriacea Lindl. 
Dendriopoterium pulidoi Svent. 
Marcetella maderensis (Bom.) Svent. 

32 

Prunus lusitanica L. 

subsp. azorica (Mouillef.) Franco 

Sorbus maderensis (Lowe) Dode 

SANTALACEAE 

Kunkeliella subsucculenta Kammer 

SCROPHULARIACEAE 

•  Euphrasia azorica H.C. Watson 
Euphrasia grandiflora Hochst. in Seub. 
•  Isoplexis chalcantha Svent. & O'Shanahan 
Isoplexis isabelliana (Webb & Berthel.) Masferrer 
Odontites holliana (Lowe) Benth. 
Sibthorpia peregrina L. 

SOLANACEAE 

•  Solanum lidii Sunding 

UMBELLIFERAE 

Ammi trifoliatum (H. C. Watson) Trelease 
Bupleurum handiense (Bolle) Kunkel 
Chaerophyllum azoricum Trelease 
Ferula latipinna Santos 
Melanoselinum decipiens (Schrader & Wendl.) Hoffm. 
Monizia edulis Lowe 
Oenanthe divaricata (R. Br.) Mabb. 
Sanicula azorica Guthnick ex Seub. 

VIOLACEAE 

Viola paradoxa Lowe 

Lower plants 

BRYOPHYTA 

•  Echinodium spinosum (Mitt.) Jur. (o) 
•  Thamnobryum fernandesii Sergio (o) 

" 

3«f 

ISSN 0254-1475 

COM(97)  195  final 

DOCUMENTS 

EN 

14  15  03 

Catalogue  number  :  CB-CO-97-211-EN-C 

ISBN  92-78-19665-7 

Office  for  Official  Publications of the European  Communities 

L-2985  Luxembourg 

3: