 The first known fossil salamanders are Kokartus honorarius from the Middle Jurassic of Kyrgyzstan and two species of the apparently neotenic, aquatic Marmorerpeton from England of a similar date. They looked superficially like robust modern salamanders but lacked a number of anatomical features that developed later. Karaurus sharovi from the Upper Jurassic of Kazakhstan resembled modern mole salamanders in morphology and probably had a similar burrowing lifestyle.
