The scapula (shoulder blade) was fused with the coracoid, forming a single bone without any visible demarcation between the two original elements. The C. nasicornis holotype was found with an articulated left fore limb including an incomplete manus (hand). Although disarticulated during preparation, a cast had been made of the fossil beforehand to document the original relative positions of the bones. Carpal bones were not known from any specimen, leading some authors to suggest that they were lost in the genus. In a 2016 paper, Matthew Carrano and Jonah Choiniere suggested that one or more cartilaginous (not bony) carpals were probably present, as indicated by a gap present between the forearm bones and the metacarpals, as well as by the surface texture within this gap seen in the cast. In contrast to most more-derived theropods, which showed only three digits on each manus (digits I–III), Ceratosaurus retained four digits, with digit IV reduced in size. The first and fourth metacarpals were short, while the second was slightly longer than the third. The metacarpus and especially the first phalanges were proportionally very short, unlike in most other basal theropods. Only the first phalanges of digits II, III, and IV are preserved in the holotype; the total number of phalanges and unguals (claw bones) is unknown. The anatomy of metacarpal I indicates that phalanges had originally been present on this digit, as well. The pes (foot) consisted of three weight-bearing digits, numbered II–IV. Digit I, which in theropods is usually reduced to a dewclaw that does not touch the ground, is not preserved in the holotype. Marsh, in his original 1884 description, assumed that this digit was lost in Ceratosaurus, but Charles Gilmore, in his 1920 monograph, noted an attachment area on the second metatarsal demonstrating the presence of this digit.Uniquely among theropods, Ceratosaurus possessed small, elongated, and irregularly formed osteoderms (skin bones) along the midline of its body. Such osteoderms have been found above the neural spines of cervical vertebrae 4 and 5, as well as caudal vertebrae 4 to 10, and probably formed a continuous row that might have extended from the base of the skull to most of the tail. As suggested by Gilmore in 1920, their position in the rock matrix likely reflects their exact position in the living animal. The osteoderms above the tail were found separated from the neural spines by 25 millimetres (0.98 in) to 38 millimetres (1.5 in), possibly accounting for skin and muscles present in between, while those of the neck were much closer to the neural spines. Apart from the body midline, the skin contained additional osteoderms, as indicated by a 58 millimetres (2.3 in) by 70 mm (2.8 in) large, roughly quadrangular plate found together with the holotype; the position of this plate on the body is unknown. Specimen UMNH VP 5278 was also found with a number of osteoderms, which have been described as amorphous in shape. Although most of these ossicles were found at most 5 m  apart from the skeleton, they were not directly associated with any vertebrae, unlike in the C. nasicornis holotype, so their original position on the body cannot be inferred from this specimen.

Question: Ceratosaurus had two metacarpals that were considered short, which were they?
Answer: first and fourth