Sentence:  With the elucidation of the mysteries of "Oenothera", and as the mutation theory came under increasing attack, some proponents of a <WSD>multivalenced</WSD> view of mutation found it increasingly difficult to speak clearly about mutations on several different levels without seeming fuzzy-headed latecomers to seemingly discarded de Vriesian views. 
Meaning of the word marked with <WSD> tags:  Having three or more homologous chromosomes during meiosis