Sentence:  The term "forest carnivores" denotes a smaller group of four species - the marten, <WSD>fisher</WSD>, lynx, and wolverine - and is only marginally descriptive, inasmuch as it excludes many carnivores that live in forests, and includes the wolverine, which can thrive in the complete absence of trees. 
Meaning of the word marked with <WSD> tags:  A North American marten, "Martes pennanti", that has thick brown fur.