Sentence:  The possible influence of various sensory factors in migration was tested by Dodson and Leggett (1974) who compared the behaviour of ultrasonically tagged control shad with ones which had been blinded, or had the olfactory system occluded (<WSD>anosmic</WSD> fish), or had been subjected to both these operations. ... Both blind and blind/<WSD>anosmic</WSD> fish orientated into the tidal current and altered their swimming speed in relation to the speed of the tide as did intact fish. 
Meaning of the word marked with <WSD> tags:  Having anosmia; lacking a sense of smell.