Sentence:  In 1558 the beagle had become well patronised by royalty and was painted by court painters, so that we know his type to have been already well established, a small hound with long, drooping ears, short pudgy body and <WSD>throaty</WSD> neck. 
Meaning of the word marked with <WSD> tags:  (of livestock or dogs) Having a dewlap or excess skin hanging under the neck.