Sentence:  Would she be able to appreciate the subtlety of their advances, if  they made any; those little twitches and twists of the lips, those narrowings and rollings of the eye, those <WSD>wrinklings</WSD> of the speculative temple, that shrugging of an eye-brow at play? 
Meaning of the word marked with <WSD> tags:  The act of wrinkling.