Sentence:  As early as 1913 Ghoreyeb and Karsner demonstrated with <WSD>perfusion</WSD> studies in dogs that bronchial artery flow would remain constant at a certain low level when pressure was maintained in the pulmonary artery and vein , but that increases in bronchial artery flow would occur in response to a relative drop in pulmonary artery pressure . 
Meaning of the word marked with <WSD> tags:  pumping a liquid into an organ or tissue (especially by way of blood vessels)