Sentence:  In tests with the Lincoln and Virginia varieties of cowpea, Crowley (1959) found that, in plants infected with bean southern mosaic virus before blossoming, the virus was present in nearly 100% of the <WSD>testae</WSD> and endosperms of seeds of both varieties, but could not be detected in the embryos. 
Meaning of the word marked with <WSD> tags:  A seed coat.