Sentence:  In the wild state the flowers are single—that is to say, only the outer florets are strap-shaped, and usually of a rosy-lilac tint, with yellowish disc florets; but under cultivation, all the florets have become ligulate or <WSD>quilled</WSD> […] 
Meaning of the word marked with <WSD> tags:  Having long, narrow petals or florets.