Sentence:  What ... are the steps from ... the sort of inarticulate <WSD>pushmi-pullyu</WSD> comprehensions the bee has and the dim sort of <WSD>pushmi-pullyu</WSD> comprehension that mediates responses to behavior releasers, to articulate, well-differentiated, and uncommitted human beliefs and desires ...? 
Meaning of the word marked with <WSD> tags:  in teleosemantics, being a representation with both indicative and imperative content.