Sentence:  ["Helianthus"] "frondosus", ... stem smooth below, leaves lance ovate, remotely and acutely serrate, <WSD>scabrous</WSD> above, paler and sub-<WSD>scabrous</WSD> beneath, 3-nerved: peduncles <WSD>scabrous</WSD>; petiole ciliate; calyx squarose, undulate, leafy, ciliate: rays 8-flowered. 
Meaning of the word marked with <WSD> tags:  Covered with scales or scabs; "hence", very coarse or rough.