Sentence:  The primary, secondary, and <WSD>tertiary</WSD> wing feathers (remiges) of adult mourning doves are shed and replaced yearly in an orderly sequence, so flight is possible during molt. ... Little is known about the shedding sequence of the secondary or <WSD>tertiary</WSD> feathers or the related coverts. If it is similar to that of passerines, however, loss of the outermost <WSD>tertiary</WSD> feather occurs coincident with loss of the fifth or sixth primarynb.... The molt then would continue sequentially until the last or innermost tertiary was shed and replaced. 
Meaning of the word marked with <WSD> tags:  Of quills: growing on the innermost joint of a bird's wing; tertial.