Sentence:  The American Registry of Pathology <WSD>operates</WSD> as a cooperative enterprise in medical research and education between the Armed Forces Institute of Pathology and the civilian medical profession on a national and international basis , under such conditions as may be agreed upon between the National Research Council and The Surgeons General of the Army , Navy , and Air Force . 
Meaning of the word marked with <WSD> tags:  perform as expected when applied