He received his Ph.D. in Microbiology and Molecular Genetics from Harvard University, where he studied various aspects of the natural history of reoviruses. He also received his M.D. from Harvard Medical School and completed a residency in Clinical Pathology at the Brigham and Womens Hospital in Boston, MA. Work in his new lab in Madison is focused on understanding how the structure of virus particles and proteins relates to the mechanisms of virus entry into cells.