His research, however, predominantly focuses on determining the specific contributions of the hippocampus to long-term memory. He has published several papers demonstrating that the hippocampus, contrary to common memory consolidation views, has a lasting contribution in the organization of long-term memory. He has also findings suggesting that long-term memories can become increasingly resistant to brain damage, particularly extensive hippocampal damage, through a reinstatement process. His research involves animal models of amnesia and a range of neuroscience techniques including, permanent brain lesions, reversible brain lesions, and immunohistochemistry.