He his primarily interested in the determinism of the spatial distributions of animal species, from fine (movement models) to large scale (distribution in the landscape) in relation to environmental features (e.g. climate change) and other species. His work lies on a strong theoretical and methodological basis using the concept of ecological niche, leading to applications for animal population management, as well as basic developments at the intersection of evolutionary ecology and behavioral ecology. His main study species are large vertebrates, such as lynx in Norway, caribou in Québec, and wood stork and sea turtles in Florida.