He is a physician-scientist who divides his time between his cancer research laboratory and the clinic. Dr. Amatruda earned his medical degree from Washington University School of Medicine in St. Louis. He completed his residency in internal medicine at Brigham and Women's Hospital in Boston and fellowships in hematology and oncology at Massachusetts General Hospital, Brigham and Women's Hospital and the Dana-Farber Cancer Institute; in molecular genetics at Consglio Nazionale delle Ricerche in Italy; and in pediatric hematology at Children's Hospital of Boston.