He received his Bachelor’s and Master’s Degrees in Biology from the University of Konstanz (Germany). He completed his Ph.D. with Dr. George Somero at Oregon State University, and he did postdoctoral work with Dr. Somero at Stanford University and Dr. Dietmar Kültz at the University of California, Davis. Working mainly with intertidal invertebrates, he and his students integrate across levels of biological organization, from the subcellular to the whole-organism, to study physiological responses to temperature, osmotic, pH, hypoxic, heavy metal, and endocrine stressors. For the Instructor