He studied Pharmacy at the University of Athens, Greece and carried out his D.Phil. thesis on cortical interneurons at the MRC Anatomical Neuropharmacology Unit of the University of Oxford, UK. As a postdoctoral fellow in the laboratory of Gord Fishell at New York University Medical Center, USA, he worked on the synaptic integration of interneurons in the developing cortical circuit and on neurodevelopmental disorders. He is a recipient of a NARSAD award and an ERC Starting Grant.