He received a Ph.D. in Computational Biology from the University of Pennsylvania where he developed computational models of cell signaling with Scott Diamond and Ravi Radhakrishnan. He completed his postdoctoral training at Harvard Medical School under the mentorship of Galit Lahav, where he studied the role of p53 dynamics in the DNA damage response. As an independent investigator, his group seeks to understand how individual cells make fate decisions and how groups of interacting cells give rise to emergent properties. Their work focuses primarily on early differentiation decisions in human embryonic stem cells and cell cycle arrest decisions in the context of cancer.