He graduated summa cum laude from Yale University with a B.S./M.S. in Molecular Biophysics and Biochemistry. He received his medical doctorate from WCMC and his Ph.D. in immunology from The Rockefeller University. He is an active gastroenterologist and mucosal immunologist focused on basic research to understand the causes and to develop new therapies for inflammatory bowel disease (IBD). The major focus of his research is to define cellular and molecular regulation of innate intestinal barrier immunity and to evaluate the functional implications of IBD-associated microbiota.