He also served as a research fellow at Mount Allison University (Canada), Linköping University (Sweden) and Matsuyama University (Japan). He trained as a biochemist and has a PhD in Theoretical Biochemistry from the University of Lisbon. He has teaching experience in systems biology and metabolic regulation. His research interests lie in the quantitative biology of redox regulation and oxidative stress, and also in the development of a redox therapy program based on the delivery of reactive oxygen species as active pharmaceutical ingredients. He has published over 60 papers that gathered more than 4000 citations (h-index: 28).