He received his Ph.D. in molecular genetics in the City University of New York, New York, in 1997, and had his postdoctoral training in the areas of mammalian cell cycle regulation and proteosome-dependent protein degradation regulation in Dr. Michele Pagano’s laboratory at New York University, New York, New York, from 1997-1998 and of neuronal differentiation and survival regulation in Dr. Lloyd Greene’s laboratory at Columbia University, New York, New York, from 1998-2002. Before joining the WSU, he was anAssistant Professor at the Penn State College of Medicine, Pennsylvania, U.S.A., from 2006-2002, andaResearch Associate at Columbia University, New York, New York, from 2002-2006.