He received his B.S. degree (honors), York University, Ph.D. degree, University of Toronto, and completed his Postdoctoral studies at UCSD and Cornell University. His areas of expertise are membrane trafficking, endolysosomal trafficking, phagocytosis, phagosome maturation, macrophages, yeast, phosphoinositides, and organelle function. His research program aims to elucidate the molecular mechanisms that establish, maintain, or change organelle identity, that is, the biochemical, biophysical, and functional properties of an organelle. He is particularly interested in the regulation and function of phosphoinositide lipid signals and GTPases in mediating organelle identity and function. He uses three model organelles: the yeast vacuole, phagosome maturation, and lysosomal tubulation in macrophages.