Prehistoric Europe

Prehistoric Europe refers to Europe before the start of written records, beginning in the Lower Paleolithic. As history progresses, considerable regional unevenness in cultural development emerges and grows. The region of the eastern Mediterranean is, due to its geographic proximity, greatly influenced and inspired by the classical Middle Eastern civilizations, and adopts and develops the earliest systems of communal organization and writing. The Histories of Herodotus (from around 440 BC) is the oldest known European text that seeks to systematically record traditions, public affairs and notable events.
Widely dispersed, isolated finds of individual fossils of bone fragments (Atapuerca, Mauer mandible), stone artifacts or assemblages suggest that during the Lower Paleolithic, spanning from 3 million until 300,000 years ago, palaeo-human presence was rare and typically separated by thousands of years. The karstic region of the Atapuerca Mountains in Spain represents the currently earliest known and reliably dated location of residence for more than a single generation and a group of individuals.
Homo neanderthalensis emerged in Eurasia between 600,000 and 350,000 years ago as the earliest body of European people that left behind a substantial tradition, a set of evaluable historic data through a rich fossil record in Europe's limestone caves and a patchwork of occupation sites over large areas. These include Mousterian cultural assemblages. Modern humans arrived in Mediterranean Europe during the Upper Paleolithic between 45,000 and 43,000 years ago, and both species occupied a common habitat for several thousand years. Research has so far produced no universally accepted conclusive explanation as to what caused the Neanderthal's extinction between 40,000 and 28,000 years ago.
Homo sapiens later populated the entire continent during the Mesolithic, and advanced north, following the retreating ice sheets of the last glacial maximum that spanned between 26,500 and 19,000 years ago. A 2015 publication on ancient European DNA collected from Spain to Russia concluded that the original hunter-gatherer population had assimilated a wave of "farmers" who had arrived from the Near East during the Neolithic about 8,000 years ago.
The Mesolithic era site Lepenski Vir in modern-day Serbia, the earliest documented sedentary community of Europe with permanent buildings, as well as monumental art, precedes by many centuries sites previously considered to be the oldest known. The community's year-round access to a food surplus prior to the introduction of agriculture was the basis for the sedentary lifestyle. However, the earliest record for the adoption of elements of farming can be found in Starčevo, a community with close cultural ties.
Belovode and Pločnik, also in Serbia, is currently the oldest reliably dated copper smelting site in Europe (around 7,000 years ago). It is attributed to the Vinča culture, which on the contrary provides no links to the initiation of or a transition to the Chalcolithic or Copper Age.
The process of smelting bronze is an imported technology with debated origins and history of geographic cultural profusion. It was established in Europe about 3200 BC in the Aegean and production was centered around Cyprus, the primary source of copper for the Mediterranean for many centuries.
The introduction of metallurgy, which initiated unprecedented technological progress, has also been linked with the establishment of social stratification, the distinction between rich and poor, and use of precious metals as the means to fundamentally control the dynamics of culture and society.
The European Iron Age culture also originates in the East through the absorption of the technological principles obtained from the Hittites about 1200 BC, finally arriving in Northern Europe by 500 BC.
During the Iron Age, Central, Western and most of Eastern Europe gradually entered the actual historical period. Greek maritime colonization and Roman terrestrial conquest form the basis for the diffusion of literacy in large areas to this day. This tradition continued in an altered form and context for the most remote regions (Greenland and Eastern Balts, 13th century) via the universal body of Christian texts, including the incorporation of East Slavic peoples and Russia into the Orthodox cultural sphere. Latin and ancient Greek languages continued to be the primary and best way to communicate and express ideas in liberal arts education and the sciences all over Europe until the early modern period.
The climatic record of the Paleolithic is characterised by the Pleistocene pattern of cyclic warmer and colder periods, including eight major cycles and numerous shorter episodes. The northern maximum of human occupation fluctuated in response to the changing conditions, and successful settlement required constant adaption capabilities and problem solving. Most of Scandinavia, the North European Plain and Russia remained off limits for occupation during the Paleolithic and Mesolithic. Populations were sparse and small throughout the Palaeolithic.
Associated evidence, such as stone tools, artifacts and settlement localities, numbers more than fossilised remains of the hominin occupants themselves. The simplest pebble tools with a few flakes struck off to create an edge were found in Dmanisi, Georgia, and in Spain at sites in the Guadix-Baza basin and near Atapuerca. The Oldowan tool discoveries, called Mode 1-type assemblages are gradually replaced by a more complex tradition that included a range of hand axes and flake tools, the Acheulean, Mode 2-type assemblages. Both types of tool sets are attributed to Homo erectus, the earliest and for a very long time the only human in Europe and more likely to be found in the southern part of the continent. However, the Acheulean fossil record also links to the emergence of Homo heidelbergensis, particularly its specific lithic tools and handaxes. The presence of Homo heidelbergensis is documented since 600,000 BC in numerous sites in Germany, Great Britain and northern France.
Although palaeoanthropologists generally agree that Homo erectus and Homo heidelbergensis immigrated to Europe, debates remain about migration routes and the chronology.
The fact that Homo neanderthalensis is found only in a contiguous range in Eurasia and the general acceptance of the Out of Africa hypothesis both suggest that the species has evolved locally. Again, consensus prevails on the matter, but widely debated are origin and evolution patterns.
The Neanderthal fossil record ranges from Western Europe to the Altai Mountains in Central Asia and the Ural Mountains in the North to the Levant in the South. Unlike its predecessors, they were biologically and culturally adapted to survival in cold environments and successfully extended their range to the glacial environments of Central Europe and the Russian plains. The great number and, in some cases, exceptional state of preservation of Neanderthal fossils and cultural assemblages enables researchers to provide a detailed and accurate data on behaviour and culture. Neanderthals are associated with the Mousterian culture (Mode 3), stone tools that first appeared approximately 160,000 years ago.
Homo sapiens arrived in Europe around 46,000 and 43,000 years ago via the Levant and entered the continent through the Danubian corridor, as the fossils at the sites of Bacho Kiro cave and Peștera cu Oase suggest. With an approximate age of 46,000 years, the Homo sapiens fossils found in Bacho Kiro cave consist of a pair of fragmented mandibles including at least one molar This site yielded the oldest known ornaments in Europe, Radiocarbon dated to over 43,000 years ago.
The fossils' genetic structure indicates a recent Neanderthal ancestry and the discovery of a fragment of a skull in Israel in 2008 support the notion that humans interbred with Neanderthals in the Levant.
After the slow processes of the previous hundreds of thousands of years, a turbulent period of Neanderthal–Homo sapiens coexistence demonstrated that cultural evolution had replaced biological evolution as the primary force of adaptation and change in human societies.
Generally small and widely dispersed fossil sites suggest that Neanderthals lived in less numerous and more socially isolated groups than Homo sapiens. Tools and Levallois points are remarkably sophisticated from the outset, but they have a slow rate of variability, and general technological inertia is noticeable during the entire fossil period. Artifacts are of utilitarian nature, and symbolic behavioral traits are undocumented before the arrival of modern humans. The Aurignacian culture, introduced by modern humans, is characterized by cut bone or antler points, fine flint blades and bladelets struck from prepared cores, rather than using crude flakes. The oldest examples and subsequent widespread tradition of prehistoric art originate from the Aurignacian.
After more than 100,000 years of uniformity, around 45,000 years ago, the Neanderthal fossil record changed abruptly. The Mousterian had quickly become more versatile and was named the Chatelperronian culture, which signifies the diffusion of Aurignacian elements into Neanderthal culture. Although debated, the fact proved that Neanderthals had, to some extent, adopted the culture of modern Homo sapiens. However, the Neanderthal fossil record completely vanished after 40,000 years BC. Whether Neanderthals were also successful in diffusing their genetic heritage into Europe's future population or they simply went extinct and, if so, what caused the extinction cannot conclusively be answered.
Around 32,000 years ago, the Gravettian culture appeared in the Crimean Mountains (southern Ukraine). By 24,000 BC, the Solutrean and Gravettian cultures were present in Southwestern Europe. Gravettian technology and culture have been theorised to have come with migrations of people from the Middle East, Anatolia and the Balkans, and might be linked with the transitional cultures mentioned earlier since their techniques have some similarities and are both very different from Aurignacian ones, but this issue is very obscure. The Gravettian also appeared in the Caucasus and Zagros Mountains but soon disappeared from southwestern Europe, with the notable exception of the Mediterranean coasts of Iberia.
The Solutrean culture, extended from northern Spain to southeastern France, includes not only a stone technology but also the first significant development of cave painting and the use of the needle and possibly that of the bow and arrow. The more widespread Gravettian culture is no less advanced, at least in artistic terms: sculpture (mainly venuses) is the most outstanding form of creative expression of such peoples.
Around 19,000 BC, Europe witnesses the appearance of a new culture, known as Magdalenian, possibly rooted in the old Aurignacian one, which soon superseded the Solutrean area and also the Gravettian of Central Europe. However, in Mediterranean Iberia, Italy, the Balkans and Anatolia, Epigravettian cultures continued to evolve locally.
With the Magdalenian culture, the Paleolithic development in Europe reaches its peak and this is reflected in art, owing to previous traditions of paintings and sculpture.
Around 12,500 BC, the Würm Glacial Age ended. Slowly, through the following millennia, temperatures and sea levels rose, changing the environment of prehistoric people. Ireland and Great Britain became islands, and Scandinavia became separated from the main part of the European Peninsula. (They had all once been connected by a now-submerged region of the continental shelf known as Doggerland.) Nevertheless, the Magdalenian culture persisted until 10,000 BC, when it quickly evolved into two microlith cultures: Azilian, in Spain and southern France, and Sauveterrian, in northern France and Central Europe. Despite some differences, both cultures shared several traits: the creation of very small stone tools called microliths and the scarcity of figurative art, which seems to have vanished almost completely, which was replaced by abstract decoration of tools.
In the late phase of the epi-Paleolithic period, the Sauveterrean culture evolved into the so-called Tardenoisian and strongly influenced its southern neighbour, clearly replacing it in Mediterranean Spain and Portugal. The recession of the glaciers allowed human colonisation in Northern Europe for the first time. The Maglemosian culture, derived from the Sauveterre-Tardenois culture but with a strong personality, colonised Denmark and the nearby regions, including parts of Britain.
A transition period in the development of human technology between the Paleolithic and the Neolithic, the Balkan Mesolithic began around 15,000 years ago. In Western Europe, the Early Mesolithic, or Azilian, began about 14,000 years ago, in the Franco-Cantabrian region of northern Spain and southern France. In other parts of Europe, the Mesolithic began by 11,500 years ago (the beginning Holocene) and ended with the introduction of farming, which, depending on the region, occurred 8,500 to 5,500 years ago.
In areas with limited glacial impact, the term "Epipaleolithic" is sometimes preferred for the period. Regions that experienced greater environmental effects as the Last Glacial Period ended had a much more apparent Mesolithic era that lasted millennia. In Northern Europe, societies were able to live well on rich food supplies from the marshlands, which had been created by the warmer climate. Such conditions produced distinctive human behaviours that are preserved in the material record, such as the Maglemosian and Azilian cultures. Such conditions delayed the coming of the Neolithic to as late as 5,500 years ago in Northern Europe.
As what Vere Gordon Childe termed the "Neolithic Package" (including agriculture, herding, polished stone axes, timber longhouses and pottery) spread into Europe, the Mesolithic way of life was marginalised and eventually disappeared. Controversy over the means of that dispersal is discussed below in the Neolithic section. A "Ceramic Mesolithic" can be distinguished between 7,200 and 5,850 years ago and ranged from Southern to Northern Europe.
The European Neolithic is assumed to have arrived from the Near East via Asia Minor, the Mediterranean and the Caucasus. There has been a long discussion between migrationists, who claim that the Near Eastern farmers almost totally displaced the European native hunter-gatherers, and diffusionists, who claim that the process was slow enough to have occurred mostly through cultural transmission. A relationship has been suggested between the spread of agriculture and the diffusion of Indo-European languages, with several models of migrations trying to establish a relationship, like the Anatolian hypothesis, which sets the origin of Indo-European agricultural terminology in Anatolia.
Apparently related with the Anatolian culture of Hacilar, the Greek region of Thessaly was the first place in Europe known to have acquired agriculture, cattle-herding and pottery. The early stages are known as pre-Sesklo culture. The Thessalian Neolithic culture soon evolved into the more coherent Sesklo culture (6000 BC), which was the origin of the main branches of Neolithic expansion in Europe. The Karanovo culture on the territory of modern day Bulgaria, was another early Neolithic culture (Karanovo I-III ca. 62nd to 55th centuries BC) which was part of the Danube civilization and it is considered the largest and most important of the Azmak River Valley agrarian settlements. The Karanovo I is considered a continuation of Near Eastern settlement type. The Starčevo culture is dating to the period between c. 6200 and 4500 BCE. It originates in the spread of the Neolithic package of peoples and technological innovations including farming and ceramics from Anatolia. The Starčevo culture marks its spread to the inland Balkan peninsula as the Cardial ware culture did along the Adriatic coastline. It forms part of the wider Starčevo–Körös–Criş culture. Practically all of the Balkan Peninsula was colonized in the 6th millennium from there. The expansion, reaching the easternmost Tardenoisian outposts of the upper Tisza, gave birth to the Proto-Linear Pottery culture, a significant modification of the Balkan Neolithic that was the origin of one of the most important branches of European Neolithic: the Danubian group of cultures. In parallel, the coasts of the Adriatic and of southern Italy witnessed the expansion of another Neolithic current with less clear origins. Settling initially in Dalmatia, the bearers of the Cardium pottery culture may have come from Thessaly (some of the pre-Sesklo settlements show related traits) or even from Lebanon (Byblos). They were sailors, fishermen and sheep and goat herders, and the archaeological findings show that they mixed with natives in most places. Other early Neolithic cultures can be found in Ukraine and Southern Russia, where the Epigravettian locals assimilated cultural influxes from beyond the Caucasus (e.g. the Dniepr-Donets culture and related cultures) and in Andalusia (Spain), where the rare Neolithic of La Almagra Pottery appeared without known origins very early (c. 7800 BC).
This phase, starting 7000 years ago was marked by the consolidation of the Neolithic expansion towards western and northern Europe but also by the rise of new cultures in the Balkans, notably the Dimini (Thessaly) and related Vinca (Serbia and Romania) and Karanovo cultures (Bulgaria and nearby areas). Meanwhile, the Proto-Linear Pottery culture gave birth to two very dynamic branches: the Western and Eastern Linear Pottery Cultures. The western branch expanded quickly, assimilating Germany, the Czech Republic, Poland and even large parts of western Ukraine, historical Moldavia, the lowlands of Romania, and regions of France, Belgium and the Netherlands, all in less than 1000 years. With this expansion came diversification and a number of local Danubian cultures started forming at the end of the 5th millennium. In the Mediterranean, the Cardium pottery fishermen showed no less dynamism and colonised or assimilated all of Italy and the Mediterranean regions of France and Spain. Even in the Atlantic, some groups among the native hunter-gatherers started the slow incorporation of the new technologies. Among them, the most noticeable regions seem to be southwestern Iberia, which was influenced by the Mediterranean but especially by the Andalusian Neolithic, which soon developed the first Megalithic burials (dolmens), and the area around Denmark (Ertebölle culture), influenced by the Danubian complex.
This period occupied the first half of the 6th millennium BC. The tendencies of the previous period consolidated and so there was a fully-formed Neolithic Europe, with five main cultural regions:
Also known as "Copper Age", the European Chalcolithic was a time of significant changes, the first of which was the invention of copper metallurgy. This is first attested in the Vinca culture in the 6th millennium BC. The Balkans became a major centre for copper extraction and metallurgical production in the 5th millennium BC. Copper artefacts were traded across the region, eventually reaching eastwards across the steppes of eastern Europe as far as the Khavalynsk culture. The 5th millennium BC also saw the appearance of economic stratification and the rise of ruling elites in the Balkan region, most notably in the Varna culture (c. 4500 BC) in Bulgaria, which developed the first known gold metallurgy in the world.
The economy of the Chalcolithic was no longer that of peasant communities and tribes, since some materials began to be produced in specific locations and distributed to wide regions. Mining of metal and stone was particularly developed in some areas, along with the processing of those materials into valuable goods.
Early Chalcolithic, 5500-4000 BC
From 5500 BC onwards, Eastern Europe was apparently infiltrated by people originating from beyond the Volga, creating a plural complex known as Sredny Stog culture, which substituted the previous Dnieper-Donets culture in Ukraine, pushing the natives to migrate northwest to the Baltic and to Denmark, where they mixed with the natives (TRBK A and C). The emergence of the Sredny Stog culture may be correlated with the expansion of Indo-European languages, according to the Kurgan hypothesis. Near the end of the period, around 4000 BC, another westward migration of supposed Indo-European speakers left many traces in the lower Danube area (culture of Cernavodă I) in what seems to have been an invasion.
Solnitsata ("The Saltworks"), a prehistoric town located in present-day Bulgaria, is believed by archaeologists to be the oldest town in Europe - a fortified stone settlement - citadelle, inner and outer city with pottery production site and the site of a salt production facility approximately six millennia ago; it flourished ca 4700–4200 BC.
Meanwhile, the Danubian Lengyel culture absorbed its northern neighbours in the Czech Republic and Poland for some centuries, only to recede in the second half of the period. The hierarchical model of the Varna culture seems to have been replicated later in the Tiszan region with the Bodrogkeresztur culture. Labour specialisation, economic stratification and possibly the risk of invasion may have been the reasons behind this development.
In the western Danubian region (the Rhine and Seine basins), the Michelsberg culture displaced its predecessor, the Rössen culture. Meanwhile, in the Mediterranean basin, several cultures (most notably the Chasséen culture in southeastern France and the Lagozza culture in northern Italy) converged into a functional union of which the most significant characteristic was the distribution network of honey-coloured silex. Despite the unity, the signs of conflicts are clear, as many skeletons show violent injuries. This was the time and area of Ötzi, the famous man found in the Alps.
This period extends through the first half of the 4th millennium BC. During this period the Cucuteni-Trypillia culture in Ukraine experienced a massive expansion, building the largest settlements in the world at the time, described as the first cities in the world by some scholars. The earliest known evidence for wheeled vehicles, in the form of wheeled models, also comes from Cucuteni-Trypillia sites, dated to c. 3900 BC.
In the Danubian region the powerful Baden culture emerged circa 3500 BC, extending more or less across the region of Austria-Hungary. The rest of the Balkans was profoundly restructured after the invasions of the previous period, with the Coțofeni culture in the central Balkans showing pronounced eastern (or presumably Indo-European) traits. The new Ezero culture in Bulgaria (3300 BC), shows the first evidence of pseudo-bronze (or arsenical bronze), as does the Baden culture and the Cycladic culture (in the Aegean) after 2800 BC.
In Eastern Europe, the Yamnaya culture took over southern Russia and Ukraine. In western Europe, the only sign of unity came from the Megalithic super-culture, which extended from southern Sweden to southern Spain, including large parts of southern Germany as well. However, the Mediterranean and Danubian groupings of the previous period appear to have fragmented into many smaller pieces, some of them apparently backward in technological matters. From 2800 BC, the Danubian Seine-Oise-Marne culture pushed directly or indirectly southwards and destroyed most of the rich Megalithic culture of western France. After 2600 BC, several phenomena prefigured the changes of the upcoming period:
Large towns with stone walls appeared in two different areas of the Iberian Peninsula: one in the Portuguese region of Estremadura (culture of Vila Nova de Sao Pedro), strongly embedded in the Atlantic Megalithic culture; the other near Almería (southeastern Spain), centred around the large town of Los Millares, of Mediterranean character, probably affected by eastern cultural influxes (tholoi). Despite the many differences, both civilisations seem to have had friendly contact and to have maintained productive exchanges. In the area of Dordogne (Aquitaine, France), a new unexpected culture of bowmen appears: the Artenac culture soon takes control of western and even northern France and Belgium. In Poland and nearby regions, the putative Indo-Europeans reorganised and reconsolidated with the culture of the Globular Amphoras. Nevertheless, the influence of many centuries in direct contact with the still-powerful Danubian peoples had greatly modified their culture.
Use of Bronze begins in the Aegean around 3200 BC. From 2500 BC the new Catacomb culture, whose origins were obscure but were also Indo-Europeans, displaced the Yamna peoples in the regions north and east of the Black Sea, confining them to their original area east of the Volga. Around 2400 BC, the Corded Ware culture replaced their predecessors and expanded to Danubian and Nordic areas of western Germany. One related branch invaded Denmark and southern Sweden (Scandinavian Single Grave culture), and the mid-Danubian basin, though showing more continuity, had clear traits of new Indo-European elites (Vučedol culture). Simultaneously, in the West, the Artenac peoples reached Belgium. With the partial exception of Vučedol, the Danubian cultures, which had been so buoyant just a few centuries ago, were wiped off the map of Europe. The rest of the period was the story of a mysterious phenomenon: the Beaker people, which seemed to be of a mercantile character and to have preferred being buried according to a very specific, almost invariable, ritual. Nevertheless, out of their original area of western Central Europe, they appeared only within local cultures and so they never invaded and assimilated but went to live among those peoples and kept their way of life, which is why they are believed to be merchants.
The rest of Europe remained mostly unchanged and apparently peaceful. In 2300 BC, the first Beaker Pottery appeared in Bohemia and expanded in many directions but particularly westward, along the Rhone and the seas, reaching the culture of Vila Nova (Portugal) and Catalonia (Spain) as their limits. Simultaneously but unrelatedly, in 2200 BC in the Aegean region, the Cycladic culture decayed and was substituted by the new palatine phase of the Minoan culture of Crete.
The second phase of Beaker Pottery, from 2100 BC onwards, is marked by the displacement of the centre of the phenomenon to Portugal, within the culture of Vila Nova. The new centre's influence reached to all of southern and western France but was absent in southern and western Iberia, with the notable exception of Los Millares. After 1900 BC, the centre of the Beaker Pottery returned to Bohemia, and in Iberia, a decentralisation of the phenomenon occurred, with centres in Portugal but also in Los Millares and Ciempozuelos.
Though the use of bronze started much earlier in the Aegean area (c. 3.200 BC), c. 2300 BC can be considered typical for the start of the Bronze Age in Europe in general.
Derivations of the sudden expansion were the Sea Peoples, who attacked Egypt unsuccessfully for some time, including the Philistines (Pelasgians?) and the Dorians, most likely Hellenised members of the group that ended invading Greek itself and destroying the might of Mycene and later Troy.
Simultaneously, around then, the culture of Vila Nova de Sao Pedro, which lasted 1300 years in its urban form, vanishes into a less spectacular one but finally with bronze. The centre of gravity of the Atlantic cultures (the Atlantic Bronze Age complex) was now displaced towards Great Britain. Also about then, the Villanovan culture, the possible precursor of the Etruscan civilisation, appeared in central Italy, possibly with an Aegean origin.
Though the use of iron was known to the Aegean peoples about 1100 BC, it failed to reach Central Europe before 800 BC, when it gave way to the Hallstatt culture, an Iron Age evolution of the Urnfield culture. Around then, the Phoenicians, benefitting from the disappearance of the Greek maritime power (Greek Dark Ages) founded their first colony at the entrance of the Atlantic Ocean, in Gadir (modern Cádiz), most likely as a merchant outpost to convey the many mineral resources of Iberia and the British Isles.
Nevertheless, from the 7th century BC onwards, the Greeks recovered their power and started their own colonial expansion, founding Massalia (modern Marseilles) and the Iberian outpost of Emporion (modern Empúries). That occurred only after the Iberians could reconquer Catalonia and the Ebro valley from the Celts, separating physically the Iberian Celts from their continental neighbours.
The second phase of the European Iron Age was defined particularly by the Celtic La Tène culture, which started around 400 BC, followed by a large expansion of them into the Balkans, the British Isles, where they assimilated druidism, and other regions of France and Italy.
The decline of Celtic power under the expansive pressure of Germanic tribes (originally from Scandinavia and Lower Germany) and the forming of the Roman Empire during the 1st century BC was also that of the end of prehistory, properly speaking; though many regions of Europe remained illiterate and therefore out of reach of written history for many centuries, the boundary must be placed somewhere, and that date, near the start of the calendar, seems to be quite convenient. The remaining is regional prehistory, or, in most cases, protohistory, but no longer European prehistory, as a whole.
The genetic history of Europe has been inferred by observing the patterns of genetic diversity across the continent and in the surrounding areas. Use has been made of both classical genetics and molecular genetics. Analysis of the DNA of the modern population of Europe has mainly been used but use has also been made of ancient DNA.
This analysis has shown that modern man entered Europe from the Near East before the Last Glacial Maximum but retreated to refuges in southern Europe in this cold period. Subsequently, people spread out over the whole continent, with subsequent limited migration from the Near East and Asia.
According to a study in 2017, the early farmers belonged predominantly to the paternal Haplogroup G-M201. The maternal haplogroup N1a was also frequent in the farmers.
Evidence from genome analysis of ancient human remains suggests that the modern native populations of Europe largely descend from three distinct lineages: Mesolithic hunter-gatherers, derivative of the Cro-Magnon population of Europe, Early European Farmers (EEF) introduced to Europe during the Neolithic Revolution, and Ancient North Eurasians which expanded to Europe in the context of the Indo-European expansion. The Early European Farmers migrated from Anatolia to the Balkans in large numbers during the 7th millennium BC. During the Chalcolithic and early Bronze Age, the EEF-derived cultures of Europe were overwhelmed by successive invasions of Western Steppe Herders (WSHs) from the Pontic–Caspian steppe, who carried about 60% Eastern Hunter-Gatherer (EHG) and 40% Caucasus Hunter-Gatherer (CHG) admixture. These invasions led to EEF paternal DNA lineages in Europe being almost entirely replaced with EHG/WSH paternal DNA (mainly R1b and R1a). EEF maternal DNA (mainly haplogroup N) also declined, being supplanted by steppe lineages, suggesting the migrations involved both males and females from the steppe. EEF mtDNA, however, remained frequent, suggesting admixture between WSH males and EEF females.
The written linguistic record in Europe first begins with the Mycenaean record of early Greek in the Late Bronze Age. Unattested languages spoken in Europe in the Bronze and Iron Ages are the object of reconstruction in historical linguistics, in the case of Europe predominantly Indo-European linguistics.
Indo-European is assumed to have spread from the Pontic steppe at the very beginning of the Bronze Age, reaching Western Europe contemporary with the Beaker culture, after about 5,000 years ago.
Various pre-Indo-European substrates have been postulated, but remain speculative; the "Pelasgian" and "Tyrsenian" substrates of the Mediterranean world, an "Old European" (which may itself have been an early form of Indo-European), a "Vasconic" substrate ancestral to the modern Basque language, or a more widespread presence of early Finno-Ugric languages in northern Europe. An early presence of Indo-European throughout Europe has also been suggested ("Paleolithic continuity theory").
Donald Ringe emphasizes the "great linguistic diversity" which would generally have been predominant in any area inhabited by small-scale, tribal pre-state societies.
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Paleolithic sanctuaries: