The Treaty of Dunkirk was signed by France and the United Kingdom on 4 March 1947, during the aftermath of World War II and the start of the Cold War, as a Treaty of Alliance and Mutual Assistance in the event of possible attacks by Germany or the Soviet Union. In March 1948, this alliance was expanded in the Treaty of Brussels to include the Benelux countries, forming the Brussels Treaty Organization, commonly known as the Western Union. Talks for a wider military alliance, which could include North America, also began that month in the United States, where their foreign policy under the Truman Doctrine promoted international solidarity against actions they saw as communist aggression, such as the February 1948 coup d'état in Czechoslovakia. These talks resulted in the signature of the North Atlantic Treaty on 4 April 1949 by the member states of the Western Union plus the United States, Canada, Portugal, Italy, Norway, Denmark, and Iceland. Canadian diplomat Lester B. Pearson was a key author and drafter of the treaty.
Please give me a comma separated list of the countries that signed the North Atlantic Treaty in April 1949 given the text below
France, United Kingdom, Belgium, Luxembourg , Netherlands, United States, Canada, Portugal, Italy, Norway, Denmark, Iceland