page_title: Nuphar carlquistii
section_title: History and classification
hierarchical_section_title: Nuphar carlquistii / History and classification
caption_attribution_description: Nuphar advena, Nymphaeaceae
context_page_description: Nuphar carlquistii is an extinct species of flowering plant in the family Nymphaeaceae related to the modern spatterdock, Nuphar advena. The species is known from fossil seeds and fruits found in the early Eocene deposits of northern Washington state, United States, and a similar aged formation in British Columbia, Canada.
context_section_description: Nuphar carlquistii fossils have been identified from two locations in Western North America, the 49 million year old Klondike Mountain Formation near Republic, Washington and at the Thomas Ranch locality near Princeton, British Columbia. Permineralized seeds identified as the extinct Nymphaeaceae genus Allenbya has been identified from the related Okanagan Highlands Princeton chert site in south central British Columbia.
  Ages for the Okanagan Highland locations are, in general, Early Eocene, with the sites that have current uranium-lead or argon–argon radiometric dates being of Ypresian age, while the undated sites or those given older dates being possibly slightly younger and Lutetian in age.
  The specimens were studied by paleobotanists Melanie DeVore, Witt Taylor, and Kathleen Pigg with the type description for N. carlquistii being published in a 2015 International Journal of Plant Sciences article.  They chose the specific name carlquistii, in honor of Sherwin Carlquist in recognition of the work he had done with Nymphaeaceae anatomy and with plant biology.  In the type description, the species name was applied to fruits and seeds specifically, while fossils of associated but not attached tepals, stigmatic disks and rhizome sections were discussed as Nuphar cf carlquistii.  One of the rhizome segments, UWBM 57225A showing root attachment points, had previously been published with an informal identification as banana fruit of the genus Ensete.  The specimen was figured in a 1996 article with a caption of "Ensete sp. fruits with seeds, UWBM 57225A, loc. A0307"  though no discussion of the fossil was made in the article text.  The fossils display a combination of features, such as a barrel shaped fruit that has vertical furrows, with the stigmatic disk being occasionally to fully crenulated, that are seen in Nuphar Section Astylus, and are noted to be quite similar to the living Nuphar advena.
caption: Nuphar advena flower and leaves
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