page_title: Ground beetle
section_title: Basal ground beetles
hierarchical_section_title: Ground beetle / Evolution and systematics / Subfamilies and selected genera / Basal ground beetles
caption_attribution_description: Clivina fossor
context_page_description: Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it is one of the 10 most speciose animal families.
context_section_description: Carabinae Latreille, 1802 – including Agoninae and Callistinae
  Altagonum
  Aplothorax (monotypic genus)
  Calosoma - including Callisthenes
  Carabus
  Cychrus
  Homothes
  Fortagonum
  Laemostenus
  Notagonum
  Coptolabrus
  Cicindelinae – tiger beetles (roughly 2,600 species; sometimes included in the Carabidae)
  Cicindinae
  Elaphrinae Latreille, 1802
  Blethisa
  Diacheila Motschulsky, 1844
  Elaphrus
  Hiletinae
  Loricerinae Bonelli, 1810
  Loricera
  Migadopinae
  Nebriinae (includes Notiophilinae, often included in Carabinae)
  Leistus
  Nebria
  Notiophilus
  Pelophila Dejean, 1821
  Nippononebria
  Nototylinae
  Omophroninae Bonelli, 1810 – round sand beetles
  Omophron
  Paussinae – ant nest beetles, flanged bombardier beetles
  Promecognathinae
  Scaritinae Bonelli, 1810 – pedunculate ground beetles
  Clivina
  Dyschirius Bonelli, 1810
  Scarites
  Siagoninae Bonelli, 1810
caption: Clivina fossor (Scaritinae: Clivinini)
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