A New Lobelia from Mexico, with Additional New Combinations in World Campanulaceae

Lobelia guerrerensis is a new species endemic to pine, pine-oak, and mixed deciduous forests of the Sierra Madre del Sur in Guerrero, Mexico. Though collected several times over the past 60 years, the specimens were consistently misidentified as the narrow-leaved phase of L. laxiflora. Detailed phenetic analyses, however, have showed them to be morphologically quite distinct. In addition, several new combinations needed for a forthcoming checklist of world Campanulaceae are published. Work is drawing to a close on a checklist of Campanulaceae s.1. currently recognized in the world's taxonomic literature. Four precursor papers effecting requisite nomenclatural changes (Lammers, 1995, 1998a, 1999) and describing recently detected species (Lammers, 1998b) have already appeared. This is the fifth and (I hope) final installment in that series. It should be noted that some of the names validated here stem from recent studies conducted under my direction by two students, Michael James Eakes of the University of Illinois and Kari Ellen Morris of the University of Illinois-Chicago. It will not be possible to publish their complete results prior to publication of the checklist. Therefore, nomenclatural innovations dictated by their studies are extracted and published here in advance, so that the resulting names will be available for use in the checklist and elsewhere. Actions undertaken here are of three types: (1) description of a new species of Lobelia L. from Guerrero, Mexico (with a change in rank for a related taxon); (2) new combinations in Campanula L., Centropogon C. Presl, and Lobelia, dictated by the principle of priority; and (3) new combinations for species and subspecies of Codonopsis Wallich, Cyanea Gaudichaud, Cyclocodon Griffith, and Delissea Gaudichaud, which do not have a name at that rank in the genus or species to which they are assigned. NEW SPECIES Unpublished morphometric studies in Lobelia sect. Homochilus A. DC. by Eakes (cf. Hamlin, 1995; Eakes & Lammers, 1996) disclosed the existence of a discrete group of specimens from the Mexican state of Guerrero, which did not conform to any previously recognized taxon. These specimens had been collected over the past 60 years by different botanists and widely distributed to the world's herbaria. In every instance, however, they were identified as the narrow-leaved phase of L. laxiflora Kunth, a species widespread in Mexico and Central America (see below). It was only with detailed phenetic analyses that the distinctness of these plants was revealed. Because these plants were as distinct morphologically as any species in the section, they are here described as a new species: Lobelia guerrerensis Eakes & Lammers, sp. nov. TYPE: Mexico. Guerrero: municipio de Atoyac de Alvarez, a 6 km al SO de Puerto del Gallo, bosque mes6filo de montafia, 2320 m, 29 Mar. 1983, Soto & Martinez 5136 (holotype, MEXU). Figure 1. A Lobelia laxiflora foliis cernuis cum marginibus integris vel subintegris et minute revolutis, corollae tubo flavo vel luteo-aurantiaco et lobis rubris vel aurantiacis, tubo antherarum dense villoso a basi ad summum tecto cum trichomatibus isabellinis 2-4 mm longis, antheris ventralibus apice caespitosis cum trichomatibus isabellinis 1.5-2 mm longis, capsulis 9-11 mm diametro, et seminibus 0.70.8 mm longis 0.3-0.4 mm latis differt. Stems 1-2 m tall, woody or suffruticose, branched or unbranched, erect or ascending, glabrous. Leaves sessile, glabrous; lamina linear or lanceolate, 7-18 cm long, 0.3-1.1 cm wide, cernuous, chartaceous; margin entire or sometimes with a few distant minute callose teeth, minutely revolute; apex narrowly acuminate; base cuneate or attenuate. Flowers solitary in the axils of the upper leaves; pedicels 60-130 mm long, ca. 1 mm diam., straight or slightly incurved, glabrous, bibracteolate in the lower half; bracteoles 0.5-3 mm long, linear. Hypanthium depressed hemispheric, 4NovoN 9: 381-389. 1999. This content downloaded from 157.55.39.35 on Mon, 29 Aug 2016 04:45:53 UTC All use subject to http://about.jstor.org/terms