Question: What different species of birds live mainly in the desert uplands and cliffs of the inner canyon?
About the article "Grand Canyon", section "Upper Sonoran and Transition".

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Vegetation in the Paria Canyon-Vermilion Cliffs Wilderness can be divided into three different zones of riparian vegetation communities, which run parallel to each side of the river.
Plants found closest to the water include cattails, common reeds, sedges, rushes, and horsetails.
Drier soil farther from the water makes a good environment for woody vegetation which has survived the repeated flooding, such as coyote willow, cottonwood, ash, tamarisk, and seep-willow.
This zone is referred to as the "regeneration site" because some of the trees that grow here survive to produce the next generation of mature canopy.
Beyond the regeneration site trees grow strong and tall.
Dominant vegetation includes cottonwood, willow, ash, and tamarisk.
This zone is the fringe between riparian and desert uplands, often sharing vegetation like rabbitbrush, Indian ricegrass, arrowweed, and sand dropseed.
Other vegetation found in the Wilderness include pinyon pine, juniper, boxelder, various species of cactus, and other desert flora. 
The Upper Sonoran Life Zone includes most of the inner canyon and South Rim at elevations from .
This zone is generally dominated by blackbrush, sagebrush, and pinyon-juniper woodlands.
Elevations of are in the Mojave Desert Scrub community of the Upper Sonoran.
This community is dominated by the four-winged saltbush and creosote bush; other important plants include Utah agave, narrowleaf mesquite, ratany, catclaw, and various cacti species.
Approximately 30 bird species breed primarily in the desert uplands and cliffs of the inner canyon.
Virtually all bird species present breed in other suitable habitats throughout the Sonoran and Mohave deserts.
The abundance of bats, swifts, and riparian birds provides ample food for peregrines, and suitable eyrie sites are plentiful along the steep canyon walls.
Also, several critically endangered California Condors that were re-introduced to the Colorado Plateau on the Arizona Strip, have made the eastern part of the Park their home.
The conifer forests provide habitat for 52 mammal species.
Porcupines, shrews, red squirrels, tassel eared Kaibab and Abert's squirrels, black bear, mule deer, and elk are found at the park's higher elevations on the Kaibab Plateau.
Above the desert scrub and up to is a pinyon pine forest and one seed juniper woodland.
Within this woodland one can find big sagebrush, snakeweed, Mormon tea, Utah agave, banana and narrowleaf Yucca, winterfat, Indian ricegrass, dropseed, and needlegrass.
There are a variety of snakes and lizards here, but one species of reptile, the mountain short-horned Lizard, is a particularly abundant inhabitant of the piñon-juniper and ponderosa pine forests.
Ponderosa pine forests grow at elevations between , on both North and South rims in the Transition life zone.
The South Rim includes species such as gray fox, mule deer, bighorn sheep, rock squirrels, pinyon pine and Utah juniper.
Additional species such as Gambel oak, New Mexico locust, mountain mahogany, elderberry, creeping mahonia, and fescue have been identified in these forests.
The Utah tiger salamander and the Great Basin spadefoot toad are two amphibians that are common in the rim forests.
Of the approximately 90 bird species that breed in the coniferous forests, 51 are summer residents and at least 15 of these are known to be neotropical migrants.
The Lower Sonoran life zone spans from the Colorado River up to 3500 feet.
Along the Colorado River and its perennial tributaries, a riparian community exists.
Coyote willow, arrowweed, seep-willow, western honey mesquite, catclaw acacia, and exotic tamarisk (saltcedar) are the predominant species.
Hanging gardens, seeps and springs often contain rare plants such as the white-flowering western redbud, stream orchid, and McDougall's flaveria.
Endangered fish in the river include the humpback chub and the razorback sucker.
The three most common amphibians in these riparian communities are the canyon tree frog, red-spotted toad, and Woodhouse’s Rocky Mountain toad.
Leopard frogs are very rare in the Colorado River corridor, and are known to exist at only a few sites.
There are 33 crustacean species found in the Colorado River and its tributaries within Grand Canyon National Park.
Of these 33, 16 are considered true zooplankton organisms.
Only 48 bird species regularly nest along the river, while others use the river as a migration corridor or as overwintering habitat.
The bald eagle is one species that uses the river corridor as winter habitat.
River otters may have disappeared from the park in late 20th century, and muskrats are extremely rare.
Beavers cut willows, cottonwoods, and shrubs for food, and can significantly affect the riparian vegetation.
Other rodents, such as antelope squirrels and pocket mice, are mostly omnivorous, using many different vegetation types.
Grand Canyon bats typically roost in desert uplands, but forage on the abundance of insects along the river and its tributaries.
In addition to bats, coyotes, ringtails, and spotted skunks are the most numerous riparian predators and prey on invertebrates, rodents, and reptiles.
Raccoons, weasels, bobcats, gray foxes, and mountain lions are also present, but are much more rare.
Mule deer and desert bighorn sheep are the ungulates that frequent the river corridor.
Since the removal of 500 feral burros in the early 1980s, bighorn sheep numbers have rebounded.
Mule deer are generally not permanent residents along the river, but travel down from the rim when food and water resources there become scarce.
The insect species commonly found in the river corridor and tributaries are midges, caddis flies, mayflies, stoneflies, black flies, mites, beetles, butterflies, moths, and fire ants.
Numerous species of spiders and several species of scorpions including the bark scorpion and the giant desert hairy scorpion inhabit the riparian zone.
Eleven aquatic and 26 terrestrial species of mollusks have been identified in and around Grand Canyon National Park.
Of the aquatic species, two are bivalves (clams) and nine are gastropods (snails).
Twenty-six species of terrestrial gastropods have been identified, primarily land snails and slugs.
There are approximately 47 reptile species in Grand Canyon National Park.
Ten are considered common along the river corridor and include lizards and snakes.
Lizard density tends to be highest along the stretch of land between the water's edge and the beginning of the upland desert community.
The two largest lizards in the Canyon are gila monsters and chuckwallas.
Many snake species, which are not directly dependent on surface water, may be found both within the inner gorge and the Colorado River corridor.
Six rattlesnake species have been recorded in the park.
Above the river corridor a desert scrub community, composed of North American desert flora, thrives.
Typical warm desert species such as creosote bush, white bursage, brittlebush, catclaw acacia, ocotillo, mariola, western honey mesquite, four-wing saltbush, big sagebrush, blackbrush and rubber rabbitbrush grow in this community.
The mammalian fauna in the woodland scrub community consists of 50 species, mostly rodents and bats.
Three of the five Park woodrat species live in the desert scrub community.
Except for the western (desert) banded gecko, which seems to be distributed only near water along the Colorado River, all of the reptiles found near the river also appear in the uplands, but in lower densities.
The desert gopher tortoise, a threatened species, inhabits the desert scrublands in the western end of the park.
Some of the common insects found at elevations above 2,000 feet are orange paper wasps, honey bees, black flies, tarantula hawks, stink bugs, beetles, black ants, and monarch and swallowtail butterflies.
Solpugids, wood spiders, garden spiders, black widow spiders and tarantulas can be found in the desert scrub and higher elevations. 
Mammals found in the Wilderness include mule deer, bobcat, fox, mountain lion, porcupine, beaver, coyote, jack rabbit, cottontail rabbit, ground squirrels, kangaroo rat, and various other rodent species.
Desert bighorn sheep were successfully reintroduced to Paria Canyon in the 1980s and are usually found in the cliffs and crags of the lower canyon. 
The cliffs consist mainly of beds of Namurian shale and sandstone, with the oldest rocks being found at the bottom of the cliffs.
It is possible to see 300 million year-old river channels cutting through, forming unconformities at the base of the cliffs.
There are an estimated 30,000 birds living on the cliffs, representing more than 20 species.
These include Atlantic Puffins, which live in large colonies at isolated parts of the cliffs and on the small Goat Island.
Also present are hawks, gulls, guillemots, shags, ravens and choughs.