Vessel end-walls scalariform. Root anatomy . Root xylem with vessels. Vessel end-walls scalariform. Reproductive type, pollination .
Unisexual flowers present .
Plants dioecious, or polygamodioecious . Inflorescence, floral, fruit and seed morphology . Flowers minute, or small. Placentation when apocarpous or one-carpelled marginal, or basal. Endosperm formation nuclear.