Vessel end-walls scalariform, or scalariform and simple. Vessels without vestured pits. Primary medullary rays mixed wide and narrow. Reproductive type, pollination . Fertile flowers functionally male and functionally female .
Unisexual flowers present .
Inflorescence, floral, fruit and seed morphology . Free hypanthium absent. Androecium 5, or 10, or 15. Pollen shed as single grains. Carpels reduced in number relative to the perianth to isomerous with the perianth.