Xylem with vessels. Vessel end-walls simple. Vessels without vestured pits. Wood storied, or partially storied. Reproductive type, pollination .
Unisexual flowers present .
Plants monoecious, or dioecious, or polygamomonoecious. Inflorescence, floral, fruit and seed morphology . The terminal inflorescence unit cymose . Hypogynous disk absent . Androecium exclusively of fertile stamens.