Xylem without vessels. Root anatomy . Root xylem with vessels. Vessel end-walls scalariform. Reproductive type, pollination .
Unisexual flowers present .
Plants dioecious, or polygamomonoecious. Inflorescence, floral, fruit and seed morphology . The terminal inflorescence unit racemose. Perigone tube absent. Endothecium developing fibrous thickenings.