This model suggests that metabolic processes play some role in overall salt regulation, in contradict Scholander's findings. Although Werner and Stelzer do not offer a complete explanation of the ultrafiltration mechanism, they offer a plausible model that can be tested experimentally. Unlike the red mangrove, the black mangrove secretes excess salt through specialized glands on the surface of leaves in addition to carrying out root ultrafiltration. Salt gland of A. germinans. From Dschida et.
Salt glands are microscopic and occur in epidermal depressions on the upper leaf surface.
Evidence suggests that these salt glands, unlike the ultrafilter of R. mangle, rely on metabolic processes to function. Based on analysis of the application of specific metabolic inhibitors, Dschida et. Their analysis also suggests that mitochondria provide the energy source for sustaining secretion. Peels of adaxial leaf surfaces were floated on treatment solutions containing various concentrations of NaCl, and exposed to different temperatures. Volume of secretion varied directly with temperature, and varied indirectly with NaCl concentration.