Secondary thickening developing from a conventional cambial ring. Included' phloem absent. Vessel end-walls simple. Vessels without vestured pits. Reproductive type, pollination .
Unisexual flowers present .
Gynoecium of male flowers vestigial, or absent. Inflorescence, floral, fruit and seed morphology . The terminal inflorescence unit cymose. Free hypanthium absent. Endothecium developing fibrous thickenings.