Internal phloem present . Secondary thickening developing from a conventional cambial ring. Xylem without fibre tracheids. Wood parenchyma apotracheal, or paratracheal. Reproductive type, pollination .
Unisexual flowers present .
Plants dioecious, or polygamomonoecious. Inflorescence, floral, fruit and seed morphology . The terminal inflorescence unit cymose, or racemose. Free hypanthium present . Petals punctate, shortly clawed.