Vessel end-walls scalariform. Vessels without vestured pits. Wood parenchyma apotracheal. Reproductive type, pollination . Fertile flowers functionally male, or functionally female .
Unisexual flowers present .
Inflorescence, floral, fruit and seed morphology . Free hypanthium absent. Hypogynous disk absent . Androecium 4, or 6. Androecium exclusively of fertile stamens.