They move on all fours when foraging. They are active during the day, starting off at dawn. They may rest during the day in between foraging sprees. Although they forage on the ground, all climb trees for safety when threatened by predators, and sleep in trees at night. They break up into harems to forage during the peak of the fruiting season.
Small groups are quieter, while larger bands are very noisy.
The dominant male decides where the group should forage, and directs them from behind. But if there is danger, he moves to the front. Some harems may include 2 or more subordinate males. These are duller in colour and are slimmer. The presence of the dominant appears to suppress the maturity of sons.