Overlapping runways but generally never share home range systems. Will share common thoroughfare system in historically infested areas. Avoid each other unless during highly charged breeding cycle and weaning process. Only social when breeding. Systems are typically made up of shallow feeding tunnels that run downward into deep nesting tunnel systems that can be few feet underneath the shallow systems.
Upper systems can leave heaved turf.
Infested areas will have above ground conical shaped mounds created by the excavated tunnel soil and debris. Mounds seem to follow a line, unlike a gopher's random mounding pattern and crescent shaped mounds. Spend about half their time sleeping in their deep tunnel system nests. Although moles are typically referred to as an insectivore, they are more of an omnivore. They eat methodically on earthworms and soft-bodied insects but will consume vegetative matter as well as other rodents and even birds.