In other work the polarity of force generation and the intrinisic sliding velocities have been established for sub-sets of the flagellar motor proteins. For light microscopy, the technique of caged-ATP photolysis allowed use to witness the immediate and transitory events relating to reactivation. The importance of the basal flagellar structures for regular beating was demonstrated by micromanipulation. A lengthy study of the structure and motility of eel spermatozoa was undertaken. Currently efforts are being made to measure sperm energy metabolism using a probe for inorganic phosphate.
Work continues also on the kinematics of avian spermatozoa.
Morphology of nexin links in relation to sliding in the sperm flagellum . Cell Motility and the Cytoskelton . Direct evidence for tension development between flagellar doublet microtubules . The propagation of a zone of activation along groups of flagellar doublet microtubules. Studies on the eel sperm flagellum.