At other times, active submission involves a group of subordinate wolves surrounding the dominant wolf with their noses up against it. Sometimes the pack will howl. Various facial muscles, eyes, ears and the nose are extremely important when wolves are expressing their feelings. Bared teeth, an open mouth, ears erect and pointed forward indicate a threat by a dominant wolf. Wolves are also very territorial animals and do not readily share it with wolves who are not members of their pack.
Wolves communicate and mark their territories by scent.
They often do this by urinating near the edges of their territory, and on stumps, rocks and logs that are within their territory. Most of this is done by the dominant wolves, usually the alpha male.