The living lemurs of Madagascar are an excellent example of primates that use signals of sound, sight, and smell about equally well. The ring-tailed lemur has two specialized glandular areas, one on the forearm and one in the armpit. The lemur then faces his opponent and arches his colorful tail over his head, quivering it violently so that the scent is disseminated into the air around him. Both the slow loris and the capuchin monkey have stereo-typed gestures of anointing their feet and hands with urine. The skunk group has further specialized these into defensive devices, but like all other anal glands, their original evolution related to their role as a social organ.
Some male deer rub themselves with urine increasing their threat signal during the rut.
The dog with the fireplug is an even more familiar example of urine scenting. In general, the position of social glands is related to the animal's particular life history. Mammals that use runways or burrows more often than not have glands located on the sides to swipe the vegetation as they pass. Deer have lands on the hind, legs that leave a scent on the trail behind them. Almost all glandular areas have elongate tufts of hair to disseminate the odor.