Cranium heavily sclerotized, glossy, yellowish or reddish-brown. Antennae 3-segmented, terminal segment minute, penultimate segment with apical sensory area. Frontoclypeal suture distinct. Galea and lacinia distinctly separate. Hypopharynx without oncyli.
Spiracles cribriform, lacking closing apparatus, concavities of respiratory plates oriented ventrad.
Terga of abdominal segments 3-7 each with 4 dorsal annulets. Anal opening V or Y-shaped, not surrounded by fleshy lobes. Phylogeny of the families of Scarabaeoidea based on characters of the hindwing articulation, hindwing base and wing venation. A revision of the genus Pleocoma . Feeding habits of Pleocoma larvae in coniferous forests of western Oregon.