After the fraction do two homologous fragments join so that one chromosome without a centromere forms, while the other chromosome has two. The chromosome without a centromere is lost. During anaphase do the two centromeres of the other chromosome segregate to opposite poles, thus causing a break of the in-between domain. Since the exact place of the breaking is subjected to chance, will one of the daughter cells get an elongated, the other one a shortened fragment. In the elongated one is the additional domain inverted.
Translocations occur as a result of the pairing of non-homologous chromosomes during meiosis.
The classic example is Oenothera . In contrast to many other animal and plant species is it impossible to correlate the linkage groups and the chromosomes of Oenothera . During meiosis do the homologous maternal and paternal domains pair. Normally are these domains and homologous chromosomes identical. This is different here.