All range from late Oligocene to middle Miocene in age. Hill features have been further subdivided into cone karst, mogotes, and river coastal rampart. Evergreen and semi-evergreen seasonal forests are present on the first two. These are sharp-pointed or oval hills. In cone karst, the hills are surrounded by limestone whereas mogotes are surrounded by blanket deposits.
Soils are well-drained, rocky, steep and shallow.
Pollen and seed dissemination agents are unknown. No information is available concerning seed biology. A total of seven mature individuals were observed in an area of approximately 3000 square meters. Seedlings have been observed in the past. The soil is rocky and shallow.