Wolves mark the outside border of their territories twice as much as inside the territory. A wolf's nose is 14 times the size of a human's, their sensitivity to smells is 100 times greater than the human sense. Wolves will communicate social position in the pack based on their body posture. High ranking animals have high tail positions and carry their body tall, showing dominant status. Lower ranking wolves carry their tails lower, often lowering their body while pawing at the higher ranking wolves.
Wolves use several types of vocalizations for different purposes.
Wolves will howl any time of the day, but are most often heard in the evening hours during their higher activity periods. Wolves will howl when they have a need to defend territory or reunite pack members whether there is a full moon or not.