The cushion marks the point of attachment of abscised leaves. Parichnos scars mark the position of parichnos tissue within the leaf and stem.
Parichnos tissue is parenchyma with lots of air space between cells.
Paralycopodites has leaf cushions that are intermediate in shape between Lepidodendron and Lepidophloios . Bothrodendron leaf scars are faint or absent in external impressions. Leaf cushions in arborescent lycopsids. In anatomical cross section, the stem of Lepidodendron displays an exarch siphonostele surrounded by secondary xylem in ranks. The evolution of secondary growth in the arborescent lycopsids is another synapomorphy for this clade.