Mesophytic, or xerophytic. Lamina margins of the leaflets serrate to dentate. Leaves without a persistent basal meristem. Leaf anatomy . Urticating hairs present.
Minor leaf veins without phloem transfer cells.
Stem anatomy . Vessel end-walls scalariform and simple. Reproductive type, pollination . Inflorescence, floral, fruit and seed morphology . Androecium 8, or 10.