They have Multicellular Apical Meristems and Lateral Branching . They have an Actinostele like most other roots. Some species have symbiotic relationships with soil microbes that fix Nitrogen. The most famous of these is the legume-Rhizobium symbiosis which has been profoundly important for agriculture. Acacia koa has this kind of symbiosis which doubtless has significant ecological importance.
Most species have mychorrizal associations with soil fungi .
These are very important for the absorption of water and minerals from soil. Large tropical trees may have root buttresses at the base. These help to support the large aerial portions of the tree. Vines have roots which cling to aerial structures. The Stems of Dicots are very similar to those of the Coniferophyta.