Nutrient dynamics and toxicology of aquatic ecosystems. Systematics and phylogenetic reconstruction using morphological and molecular approaches.
Primary research focuses on systematics, evolution and development of amphibians and reptiles.
Sexual selection and evolution in freshwater fishes. Genetic and biochemical processes that regulate the differentiation of specialized cells in a spatial pattern, in filaments of nitrogen-fixing cyanobacteria. Characterization and application of symbioses of nitrogen-fixing cyanobacteria and plants of agricultural importance. Computer-aided analysis of genomic DNA sequences. Sequencing the genome of the common soil bacterium Agrobacterium tumefaciens, using genomics and other techniques to understand the overall biology of the bacterium, including how it causes disease in plants and animals.