It's worth noting that this problem has never been addressed. The challenge has set me thinking about how selection interacts with such self-ordered properties. This job is hardly begun, but several points are clear. Eventually, mutation overwhelms selection and disperses an adapting population away from optimal genotypes. But a second limitation on selection seems to be emerging.
Natural selection is a kind of combinatorial optimiation process.
Under strong selection, a population will at least climb to a local peak. Worse, this appears to be a general tendency in any combinatorial optimization process. As the entities under selection become more complex, the optima that can be reached become progressively more mediocre. What in the post-Darwin world this might imply has me even more deeply puzzled. Embracing him, and moving on.