Secondary thickening developing from a conventional cambial ring. Reproductive type, pollination . Fertile flowers functionally male, or functionally female, or functionally male and functionally female . Plants monoecious, or dioecious. Inflorescence, floral, fruit and seed morphology .
Inflorescences bracteate spikes or catkins of mostly sessile flowers.
Free hypanthium absent. Androecium in male flowers, exclusively of fertile stamens. Pollen shed as single grains. Gynoecium 3 carpelled, or 5 carpelled. Carpels isomerous with the perianth, or increased in number relative to the perianth.