Reproductive characters Fertilization is external. Epidermal lining is absent from the anterior cloacal chamber of females. Evaginations are absent from the dorsolateral walls of the male cloacal tube. Anterior ventral glands are present in the cloacae of females. No spermathecae are present in the female cloacal chamber.
Glands secreting into the dorsal walls of the female cloaca are absent.
Anterior ventral glands are present in male cloacae. Posterior ventral glands are absent from male cloacae. Kingsbury's glands are absent from male cloacae. Dorsal pelvic glands are absent in males. Lateral pelvic glands are absent in males.