Lamina margins entire, or crenate to serrate. Leaf anatomy . Stem anatomy . Secondary thickening developing from a conventional cambial ring. Reproductive type, pollination .
Fertile flowers hermaphrodite .
Plants hermaphrodite . Inflorescence, floral, fruit and seed morphology . The terminal inflorescence unit cymose. Inflorescences loosely or closely cymose, the pedicels often long. Free hypanthium absent .