Traveling 9-10 hours a day. Group size ranges from small to as large as 236 members, but the average size is 9-58 members. Groups contain several adult males, 2-3 times as many adult females, and an equal number of offspring. Dominance network highly dependent on material ranks. Adolescent males leave the group to form new groups while females stay with the original group throughout their lives.
Male dominance is directly related to reproductive success.
Males are aggressive toward other males and females. The females benefit from having relations with a high ranking male. Females also receive protection along matrilineal lines. Vocalization highly important to group. Highly intelligent and lively, may become bad-tempered as an adult.