Other hosts have been reported, but not recently confirmed. North and east. Middle East, south Russia, Uzbekistan, and Japan. South-central and southern, particularly Spain, southern Russia, Portugal, southern France, Italy, Greece, and Bulgaria. In Israel the female lays 300 or more eggs at a rate of more than 35 per day.
Many young larvae die, but mortality among older larvae is very low.
Pupae of the first generation occur in the fruit or in a fold of a leaf. Overwintering pupae of second or later generations occur beneath bark in fissures of the stock. There are two generations in central Europe with a third generation further south. In Israel, up to four generations may occur. All moths of the same generation do not emerge at the same time.