Males have turquoise and red genitals. Live in both forest and savannah, often moving between the two. Variable-sized bands of young males. Mating groups usually composed of one male and group of related females.
Females are dominant, although males are twice as large.
When predators approach, males may act as decoys, jumping on bushes, while females and juveniles hide. Males sexually mature at 4.5 years, Females sexually mature at 3 years. No estrus swellings. In females, parental relationships last until adulthood. Males driven from group at or near sexual maturity.