Dominance network highly dependent on material ranks. Adolescent males leave the group to form new groups while females stay with the original group throughout their lives. Male dominance is directly related to reproductive success. Males are aggressive toward other males and females. The females benefit from having relations with a high ranking male.
Females also receive protection along matrilineal lines.
Vocalization highly important to group. Highly intelligent and lively, may become bad-tempered as an adult. Studies have been done to test the memory and leaming capabilities. Mating occurs in September and October. Males sexually mature at the age of 4 and a half years and females at he age of 3 and a half years.