PPL announced last year that it had produced the world's first cloned pigs, but the birth of the new piglets is important because it represents the first time piglets were produced from cells that had been genetically modified. PPL estimated that clinical trials involving xenotransplantation with pig organs could start in four or five years. With the birth of Dolly in July 1996, PPL, along with the Roslin Institute, became the first company to clone an adult mammal. PPL later became the first company to achieve gene knock-out in livestock. PPL Therapeutics is based in Scotland, but the bulk of its work involving xenografts is conducted in Blacksburg, Virginia.
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