That is in some doubt now, but we are still pretty sure of the upper bounds of their mass. The Strong Force acts on hadrons, through the exchange of pion particles. Hadrons, as we have seen, are made up of quarks held together by the Color force, which is an exchange of gluons . It is what holds the nucleus of an atom together. The Electromagnetic Force , which we've discussed before, acts on charged particles through the exchange of photons .
Gravity operates on any particle with mass, and is mediated by the graviton .