Primary vascular tissue centrifugal. Secondary thickening absent, or developing from a conventional cambial ring. Xylem with fibre tracheids. Vessel end-walls simple. Reproductive type, pollination .
Fertile flowers hermaphrodite .
Unisexual flowers absent . Inflorescence, floral, fruit and seed morphology . The terminal inflorescence unit cymose. Floral receptacle markedly hollowed. Petals clawed, or sessile.