The males in the group compete for rank. High rank is obtained through fighting or by forming alliances with other males. High ranking males have first access to females in season. Subordinate males may also achieve mating success by ingratiating themselves with females by protecting them and their young. Although males and females are promiscuous, a male and female may form a long-lasting friendship.
Males are usually larger and have some sort of mane.
One young, rarely twins. No specific breeding season. The dominant male decides when and where to forage. Females are in front as they move, but it is the dominant male that directs them from behind. Females with young stay closer to the dominant male, subordinate females hang out with the subordinate males.