Males sexually mature at 4.5 years, Females sexually mature at 3 years. No estrus swellings. In females, parental relationships last until adulthood. Males driven from group at or near sexual maturity. Juvenile males often harrass adult males during copulation.
Groups are adaptable, climbing in forest, terrestrial in savannah.
Females have not estrus swelling, and can have postconception estrus. Females are dominant. At predator s approach, females often pick up the nearest infant, not necessarily their own, before hiding. Barbara Smuts et.