The mesophyll crystals druses. Stem anatomy . Secondary thickening developing from a conventional cambial ring. Vessel end-walls scalariform. Reproductive type, pollination .
Fertile flowers hermaphrodite .
Unisexual flowers absent . Inflorescence, floral, fruit and seed morphology . Androecial members free of the perianth. Androecium exclusively of fertile stamens. Stamens 1 .