Internal phloem absent. Included' phloem absent. Vessel end-walls simple, or scalariform and simple. Vessels without vestured pits. Reproductive type, pollination .
Fertile flowers hermaphrodite .
Unisexual flowers absent . Inflorescence, floral, fruit and seed morphology . Sometimes the gynoecium acyclic . Flowers tetracyclic to polycyclic. Endothecium developing fibrous thickenings.