The cortex containing cristarque cells, or without cristarque cells. Secondary thickening developing from a conventional cambial ring. Vessel end-walls simple. Vessels without vestured pits. Reproductive type, pollination .
Fertile flowers hermaphrodite .
Inflorescence, floral, fruit and seed morphology . The terminal inflorescence unit cymose. Inflorescences cincinni or dichasia, sometimes pseudoracemose. Free hypanthium absent. Calyx lobes markedly longer than the tube.