The two that fly project their calls upward, and flying conspecifics land at or near the calls. The one that cannot fly does not call, though it retains a courtship song. Sound production by female mole crickets is somewhat a mystery. On rare occasions, when in contact with other mole crickets, females have been heard making sounds as they rub their wings together. S.M. Ulagaraj tape recorded and analyzed such sounds of a female tawny mole cricket.
Females seem to produce sounds during aggression or defense.
Two aspects of hearing in mole crickets are puzzling. In other crickets, the tympana are not evident in the early instars and are fully developed only in the adult. Dave Yager, in a pilot study, established that some southern mole cricket nymphs had hearing similar to, but generally less sensitive than, the adult. The function and exact nature of precocious hearing in mole crickets are unexplored. The specimens he studied had flown to lights.