Males sexually mature at 5 yrs. Females sexually mature at 4 yrs. No estrus swellings. Infant birth wt. Infants carried by males for 4 mo., transferred to females for nursing.
Males also share food, groom, and play with offspring.
Siblings carry infants. Extensive paternal and sibling care of infants. Terborgh, J. 1983. Five new world primates. Princeton University Press.