Again a constraint on optimisation. I guess that won't need 2-6 million years. They have produced homogeneous mice in the laboratory in a couple of years, for tens of genes. Thus genetic variance in male trait is maintained at equilibrium by recurrent mutations, which females select against. Again the optimum is to eliminate mutation, but this cannot normally be done for any trait.
Male traits are an indicator of viability.
Females choose 'handicapped' males with expensive traits, because only healthy males can afford the cost of making these traits. The optimum is for all males to be equally healthy, but this can't be achieved if health is determined by something that changes fast, like parasite genotypes, because of the high population size and rapid generation time of parasites. Yes indeed, finally I see something that points to a random chance every individual has because that is what I think plays an important role as well, just a lot of chance, extra genetic chance. If chance plays indeed an important role than we can stop trying to argue and define rules for the female's behaviour at the arena. That's a pity.