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These unusual cells are among the most powerful biological tools available. These properties suggest novel applications in biotechnology and health care. The relationship of ES cells to the various projects within the group is represented schematically below. Their ability to colonise the germ line has proven very useful in allowing us to use gene targeting to develop animal models of human genetic disease. At the same time, their in vitro differentiation suggests how the same cells can then be used to treat the modelled disease.