This species emerges late, usually 1-2 hr after sunset in summer, and at sunset in winter. A second peak of activity occurs at about 10 pm. Bats return to the day roost by about 1 hr before sunrise. Leaf-nosed bats do not hibernate. They move within the roost to find suitable temperatures, and undergo brief periods of reduced body temperature.
Individuals occupy different roosts in summer and winter.
Probably some individuals migrate to Mexico in winter. Groups of up to 500 have been observed in California. Both sexes may roost together in the non-breeding season, but roost separately during spring and summer. Copulation occurs from September to November. Delayed implantation is followed by births from May to June.