Pollen in the Upper Cretaceaous and other plant parts in the Oligocene. Inflorescence a solitary flower or in cymes. The calyx sometimes connate at the base, the corolla in two whorls, the filaments of the stamens strap- shaped and the carpels of the gynoecium are free not fused. Berry-like or follicle, small and oily with endosperm. Habit is generally shrubs or trees.
Leaves are simple, entire and alternate.