Metanotal groove usually distinct. Pretarsal claws of hind legs not pectinate. Unlike Ponerinae, the second gastral segment does not embody a secondary girdling constriction or stridulatory organs. Females generally like conspecific workers, but usually somewhat to much larger and with the mesosoma relatively well-developed. They have relatively well-developed eyes and ocelli, and complexly subdivided mesosomas.
Males always alate, with well-developed eyes and ocelli.
Antennal scapes usually short, except in species of Polyergus. Mesosoma like that of females. Petiole usually nodiform, without definite spines or teeth. Cerci usually present. The rest of body similar to workers.