Annales de la Soci t Entomologique de Belgique. Conspicuous polymorphism is seen in the workers of almost all species.
Large workers have relatively wide heads.
Mandibles large and strong, with many denticles, which are frequently worn-down. Eyes of medium size, situated at the midlength of head viewed laterally. Antennae 12-segmented, not forming a club. Frontal lobes short but conspicuous, concealing at least parts of the antennal insertions. Frontal carinae short, barely extending beyond antennal insertions.