Stem anatomy . Secondary thickening developing from a conventional cambial ring. Xylem with tracheids, or without tracheids. Vessel end-walls simple. Reproductive type, pollination .
Fertile flowers hermaphrodite .
Inflorescence, floral, fruit and seed morphology . The terminal inflorescence unit cymose. Inflorescences 'boragoid' cincinni . Free hypanthium absent. Androecium exclusively of fertile stamens.