Results of the 12 crosses show that both the non-recombinant F1 female backcrosses and the recombinant F1 male backcrosses show a decline in fitness relative to parentals. This implies that coadapted gene complexes may extend beyond single chromosomes, which may make outbreeding depression more difficult to purge than has often been assumed.
Future work is directed at further characterizing the extent of coadaptation.
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