Leaf blades with pinnate secondary veins and reticulate tertiary veins. Anthers not poricidal. Leaves with small, often caducous stipules. Fruits dehiscent, seeds not winged. Leaves without stipules.
Fruits indehiscent, or, if dehiscent, the seeds with wings.
Fruits conspicuously ridged. Fruits not conspicuously ridged. Plants with abundant, white exudate. Plants without exudate. Seeds with white hairs.