Root xylem with vessels. Vessel end-walls scalariform. Reproductive type, pollination . Plants dioecious, or polygamodioecious . Inflorescence, floral, fruit and seed morphology .
Flowers minute, or small.
Placentation when apocarpous or one-carpelled marginal, or basal. Endosperm formation nuclear. Gynoecia of adjoining flowers generally combining to form a multiple fruit . Fruit 1 seeded. Endosperm oily, or not oily.