Secondary thickening developing from a conventional cambial ring. Xylem with tracheids. Vessel end-walls scalariform . Primary medullary rays tending to be mixed wide and narrow. Reproductive type, pollination .
Fertile flowers hermaphrodite .
Unisexual flowers absent . Inflorescence, floral, fruit and seed morphology . Flowers aggregated in 'inflorescences'. The terminal inflorescence unit cymose, or racemose. Floral receptacle with neither androphore nor gynophore .