Fruit, embryo and seedling . Pericarp free . Abaxial leaf blade epidermis . Guard-cells overlapping to flush with the interstomatals. Intercostal silica bodies absent.
Large prickles present costally.
Costal short-cells conspicuously in long rows. Transverse section of leaf blade, physiology . Lamina mid-zone in transverse section open. PCR sheath extensions absent. Mesophyll with radiate chlorenchyma.