Plants hermaphrodite, or dioecious, or gynodioecious. Gynoecium of male flowers vestigial. Inflorescence, floral, fruit and seed morphology . The terminal inflorescence unit cymose. Free hypanthium absent.
Calyx lobes markedly longer than the tube.
Androecium exclusively of fertile stamens. Seeds copiously endospermic. Micropyle not zigzag . Geography, cytology .