Vessels with vestured pits. Reproductive type, pollination . Inflorescence, floral, fruit and seed morphology . The terminal inflorescence unit cymose. Free hypanthium absent.
Corolla lobes markedly longer than the tube.
Corolla contorted . Carpels reduced in number relative to the perianth, or isomerous with the perianth, or increased in number relative to the perianth. Placentation basal to axile, or axile, or axile to apical. Capsules when dehiscent, loculicidal . Seeds with starch.