The snout is short and toothed. The carpus and manus are separate. The metacarpals are separate and with digits. The pelvic girdle elements are free rather than co-ossified. The metatarsals are fused only at the proximal ends.
Gastral ribs are present, as are several advanced avian traits.
The tail is short, and a pygostyle is present. The shoulder joint permits raising of the wing above the level of the vertebral column. The second digit of the manus and the ulna support flight feathers. The wing seems to have been capable of folding. The manus is shorter than the forearm or the humerus.