These pairs are tightly associated with each other. These complexes apear to have two parts and are called bivalents. During this stage sections of chromosome are exchanged between homologs. This stage ends with the breakdown of the nuclear envelope. A spindle forms.
Individual microtubules connect to the kinetochore of each chromosome at the centromere.
Homologs separate and are pulled to opposite ends of the cell. Each pair of homologs segregate independently of one another. Begins when the chromosomes stop moving. During this stage chromosomes may disperse to form chromatin and the nuclear envelope reforms. Telophase in pollen formation is coupled with cytokinesis.