The sporophyll, at maturity, becomes a cone scale. The cone scale starts out as a sporophyll with either spores or seed developing on its upper surface. A cluster of sporophylls at the tip of a branch is termed a strobilus. In pines and firs these strobili are woody and produce seeds. Cork contains suberin , a waxy substance that is water proof.
Cork is synonymous with periderm .
The cork cambium is a synonym for phellogen . It is a cambium that forms in the cortex of the young stem and the pericycle of the young root. It then divides, as a typical cambium does, on both sides of the cambium. On the outer side about 4-5 layers of cells are formed. On the inside only one layer of cells is formed.