Carpels reduced in number relative to the perianth to increased in number relative to the perianth. Gynoecium very shortly stylate, or non-stylate. Placentation finally axile. Outer integument not contributing to the micropyle. Polar nuclei fusing prior to fertilization.
Capsules basally loculicidal .
Embryo well differentiated. Embryo slightly curved, or straight. Physiology, biochemistry . Iridoids not detected. Ellagic acid absent.