Floral nectaries present. Inflorescence, floral, fruit and seed morphology . The terminal inflorescence unit cymose. Inflorescences cincinni or corymbs . Floral receptacle not markedly hollowed.
Corolla lobes markedly longer than the tube.
Endothecium developing fibrous thickenings. Anther epidermis persistent. The initial microspore tetrads tetrahedral. Anther wall initially with more than one middle layer. Carpels isomerous with the perianth.