Endothecium developing fibrous thickenings. Carpels reduced in number relative to the perianth to isomerous with the perianth. Placentation axile to apical. Outer integument not contributing to the micropyle. Endosperm formation nuclear.
Capsules sometimes circumscissile .
Embryo well differentiated. Physiology, biochemistry . Iridoids not detected. Geography, cytology . Species about 55.