Vessels without vestured pits. Reproductive type, pollination . Inflorescence, floral, fruit and seed morphology . The terminal inflorescence unit cymose. Free hypanthium absent .
Calyx lobes markedly longer than the tube.
The androecial bundles when bundled, alternating with the corolla members, or opposite the corolla members. Androecium exclusively of fertile stamens, or including staminodes. Endothecium developing fibrous thickenings. Anther epidermis persistent. The initial microspore tetrads tetrahedral.