Ovary 3 locular. Polar nuclei fusing prior to fertilization. Endosperm formation helobial. Mericarps in Trichoryne , 3. Fruit usually a capsule .
Capsules denticidal, or septicidal and loculicidal, or loculicidal.
Seedling collar not conspicuous. Seedling cataphylls absent. First leaf centric, or dorsiventral. Primary root ephemeral. Physiology, biochemistry .