Androecium exclusively of fertile stamens. The initial microspore tetrads tetrahedral, or isobilateral. Carpels reduced in number relative to the perianth to isomerous with the perianth. Polar nuclei fusing prior to fertilization. Endosperm formation nuclear.
Capsules loculicidal .
Seeds endospermic . Physiology, biochemistry . Iridoids not detected. Geography, cytology . Temperate to tropical.