Seed dispersal takes place in October. Late spring leaves and summer leaves persist through the winter. Decreases in soil moisture and amounts of precipitation have a significant impact on the phenology of this species. Generally there is a gradual buildup of root reserves from a seasonal low in March to floral bud development in August. Immediately afterwards a prolonged decline occurs which lasts until September.
Data is inconsistent on the buildup of reserves through initial flower opening and fruit set.
He indicates that dates for full bloom and seed dissemination are somewhat predicatable.