Gynoecium of male flowers pistillodial, or vestigial, or absent. Inflorescence, floral, fruit and seed morphology . Inflorescences terminal, or axillary, or cauliflorous. Floral receptacle with neither androphore nor gynophore . Free hypanthium absent.
Calyx lobes markedly longer than the tube.
Androecium of male flowers exclusively of fertile stamens. Endothecium developing fibrous thickenings. Anther epidermis persistent. The initial microspore tetrads tetrahedral, or isobilateral, or decussate. Carpels reduced in number relative to the perianth, or isomerous with the perianth.