Unisexual flowers absent . Inflorescence, floral, fruit and seed morphology . Inflorescences usually cincinni . Floral receptacle with neither androphore nor gynophore . Hypogynous disk absent.
Calyx lobes markedly longer than the tube.
Petals shortly clawed. Androecium exclusively of fertile stamens. Endothecium developing fibrous thickenings. The initial microspore tetrads tetrahedral, or isobilateral. Tapetum glandular, or amoeboid.