They have crowns as the sign of ruling over someone. The second of John's vision of God's Throne resolves this when Jesus Christ appeared there. They encircled the throne and the living creatures and the elders . There are not any resurrected human beings. Therefore, they cannot be their princes.
Angels are there, then they are their rulers, so they are from angels.
We may distinguish them better at the time of the presentation of human being before God the Father . The Bible knows angels as rulers. We call them archangels and believe that they are before God's Throne. One from them is Gabriel. He gives this testimony about himself.