Vessel end-walls simple. Wood parenchyma scanty paratracheal. Reproductive type, pollination . Inflorescence, floral, fruit and seed morphology . Free hypanthium absent.
Calyx lobes markedly shorter than the tube to markedly longer than the tube.
The androecial bundles when bundled, opposite the corolla members. Androecium exclusively of fertile stamens, or including staminodes. Endothecium developing fibrous thickenings. Anther wall initially with one middle layer. Tapetum amoeboid, or glandular.