Anther epidermis persistent. The initial microspore tetrads tetrahedral, or isobilateral, or decussate, or T-shaped. Carpels reduced in number relative to the perianth. Polar nuclei fusing prior to fertilization. Endosperm formation cellular.
Capsules loculicidal .
Seeds thinly endospermic, or non-endospermic. Seeds with amyloid . Physiology, biochemistry . Ellagic acid absent. Aluminium accumulation not found.