Aquatic larvae feed on littoral, benthic, and planktonic arthropods. During breeding migrations, individuals are sometimes found under surface objects such as rocks and logs. Postmetamorphic juveniles retreat to small-mammal burrows after spending a few hours or days in mud cracks near water or tunnels constructed in soft soil.
Aquatic larvae seek cover in turbid water, clumps of vegetation, and other submerged debris.
Rainfall is important to the formation and maintenance of breeding ponds. Most surface migrations and other movements by adults are associated with sustained rainfall, especially at night. In some localities, dispersal of postmetamorphic juveniles from breeding ponds is not associated with rainfall. In such cases, desiccation can cause significant mortality. Prime habitat in California is annual grassland, but seasonal ponds or vernal pools are crucial to breeding.