Correlation with faunas assigned to the Uintan and Duchesnean mammal ages in North America and to the later Robiacian and Headonian mammal ages in Eurpoe is more tenuous. It has a short, vertical, deepened symphysis with both an inferior and strongly developed superior transverse torus. The P3 through M2 on both sides of the jaw have smooth enamel crowns with faint or no buccal cingulids.
All major cusps are about equal in height.
The M1 has a small and anterior placed paraconid near the tooth midline but has no hypoconulid. The trigonid of M1 is narrower transversely than the talonid. The metaconid and protoconid are about equal in size and height and the small paraconid is anteromedial. The trigonid forms a broad and low transvrse lophid with only a small trigonid basin. Distally, there is a small posterior closure of the talonid basin between the massive hypoconic and the more marginally placed but equally high entoconid.