Epicalyx absent . Androecium exclusively of fertile stamens. Carpels reduced in number relative to the perianth to increased in number relative to the perianth. Polar nuclei fusing prior to fertilization. Endosperm formation cellular, or nuclear.
Capsules loculicidal .
Physiology, biochemistry . Ursolic acid absent . Geography, cytology . Holarctic and Paleotropical. Temperate to sub-tropical.