Secondary thickening developing from a conventional cambial ring. Wood parenchyma absent. Reproductive type, pollination . Inflorescence, floral, fruit and seed morphology . Free hypanthium absent.
Calyx lobes markedly longer than the tube.
Pollen shed as single grains. Gynoecium 3 carpelled . Carpels reduced in number relative to the perianth. Placentation free central . Geography, cytology .