A pair of nuclei migrate to the tip of the ascus mother cell and fuse to form a diploid nucleus. Then meiosis occurs followed by four successive mitotic divisions creating eight ascospores per ascus. They reported observing eight nuclei per ascospore, presumably resulting from additional mitosis. As they also reported, hyphae contain porate septa. Typically there are 10-15 nuclei per compartment, but sometimes as many as 50.
Anastomosis is highly prevalent.
Nuclear pairs are sometimes observed in the undifferentiated hyphae. The authors theorized a life-cycle where plasmogamy between strain variants creates a heterokaryotic mycelium, which they referred to as secondary mycelium. It then is theorized to form heterokaryotic sclerotia which can either revert back to secondary mycelium or lead to ascocarp formation. The assumption that heterokaryotic mycelium has a role in the life-cycle is contradicted by much evidence. The unusual mycelium taken from the meld was difficult to grow, which is probably why it was not studied more thoroughly.