Anther epidermis persistent. Carpels isomerous with the perianth. Ovary 3 locular . Outer integument contributing to the micropyle, or not contributing to the micropyle. Polar nuclei fusing prior to fertilization.
Capsules apically loculicidal.
Fruit many seeded. Embryo well differentiated. Geography, cytology . Holarctic, Paleotropical, Neotropical, Cape, and Antarctic. Temperate to tropical.