Most surface migrations and other movements by adults are associated with sustained rainfall, especially at night. In some localities, dispersal of postmetamorphic juveniles from breeding ponds is not associated with rainfall. In such cases, desiccation can cause significant mortality. Prime habitat in California is annual grass, but seasonal ponds, or vernal pools are crucial to breeding. Permanent ponds or reservoirs sometimes used as well.
Adults exist in subterranean refugia most of the year.
Before and after breeding, they emerge at night during rains. During breeding, some diurnal activity occurs. In late spring or early summer, postmetamorphic juveniles disperse from breeding sites at night. In California, the first rains of November usually initiate adult migration to breeding ponds. They usually stay at the ponds a few days, but some individuals may remain up to several weeks after breeding is completed.