The other four are found in the chromosomes of the nucleus, including BRCA-1, commonly known as the breast cancer gene. The results in Nature indicate those groups should remain together, in contrast to previously published molecular results. The rodents, including the guinea pig, belong together. And rabbits probably do go with rodents. On the other hand, a group known as Archonta should be split apart according to Springer et al.
Bats are somewhere completely different.
They're closer to pigs and cows than rodents and primates. If this is starting to sound a bit confusing, the biologists have a very simple explanation for the divergence and resulting evolutionary tree. The relationships which evolved closely parallel the movement of continental land masses during geologic time. That's why Springer named one group Afrotheria for its African origins and another Laurasiatheria after the land mass which gave rise to North America, Europe and Asia. One in the southern hemisphere and the other in the northern hemisphere.