Anther epidermis persistent. Outer integument contributing to the micropyle. Polar nuclei fusing prior to fertilization. Synergids sometimes with filiform apparatus. Endosperm formation nuclear, or helobial.
Capsules sometimes denticidal, or poricidal.
Seeds copiously endospermic. Testa encrusted with phytomelan . Physiology, biochemistry . Ellagic acid absent. Geography, cytology .