The thickenings spiral. The initial microspore tetrads tetrahedral, or isobilateral. Ovary 1 locular, or 3 locular. Polar nuclei fusing prior to fertilization. Endosperm formation helobial.
Capsules usually loculicidal.
Endosperm not oily. Seeds with starch. Physiology, biochemistry . Cynogenic constituents tyrosine-derived. Ellagic acid absent.