Vessel end-walls simple. Reproductive type, pollination . Inflorescence, floral, fruit and seed morphology . Free hypanthium absent. Hypogynous disk absent.
Calyx lobes markedly longer than the tube.
Androecium exclusively of fertile stamens. Endothecium not developing fibrous thickenings. Anther epidermis persistent. Anther wall initially with one middle layer. Pollen shed as single grains.