Vessels with vestured pits. Reproductive type, pollination . Inflorescence, floral, fruit and seed morphology . Flowers solitary, or aggregated in 'inflorescences'. Hypogynous disk absent .
Calyx lobes markedly longer than the tube.
Corolla lobes about the same length as the tube, or markedly longer than the tube. Androecium exclusively of fertile stamens. Microsporogenesis successive, or simultaneous. The initial microspore tetrads linear. Gynoecium 2 carpelled.