Coracle appears simultaneously in the epidermis , hindgut and foregut .
As the germ band retracts, expression increases in these tissues and also appears in the tracheal branches and salivary glands . Coracle is not expressed within the CNS although it is expressed within some peripheral nervous system sensory neural cells such as the scolopidia of the pentascolopidial chordotonal organ. Once established, this pattern of expression is maintained throughout embryogenesis. Coracle is closely associated with the cell membrane in all expressing cells. Larval Imaginal disc cells show intense staining in regions of cell contact just below the apical epithelial surface, just as was observed in embryonic epithelia.