DOI: 10.5061/dryad.cvdncjth4
title: Supporting information for phylogenetic analysis and imaging of Agriodontosaurus

description: The Lepidosauria is the most species-rich group of land-dwelling
 vertebrates. The group includes around 12,000 species of lizards and
 snakes (Squamata) and one species of Rhynchocephalia, the tuatara
 Sphenodon punctatus from New Zealand. Squamates owe their success to their
 generally small size, but also to their highly mobile skull that enables
 them to manipulate large prey. These key features of lizard and snake
 skulls are not seen in Sphenodon, which makes it important to understand
 the nature of their common ancestor. Lepidosaurs originated in the
 Triassic 252–201 million years ago, but confusion has arisen because of
 incomplete fossils, many of which are generalized lepidosauromorphs,
 neither squamates nor rhynchocephalians. Here we report a reasonably
 complete skull and skeleton of a definitive rhynchocephalian from the
 Middle Triassic (Anisian) Helsby Sandstone Formation of Devon, UK that is
 around 3–7 million years older than the oldest currently known lepidosaur.
 The new species shows, as predicted, a non-mobile skull but an open lower
 temporal bar and no large palatine teeth, and it seems to have been a
 specialized feeder on insects. This specimen helps us understand the
 initial diversification of Lepidosauria as part of the Triassic
 Revolution, when modern-style terrestrial ecosystems emerged.
description: # Supporting information for phylogenetic analysis and imaging of
 Agriodontosaurus Dataset DOI:
 [10.5061/dryad.cvdncjth4](https://doi.org/10.5061/dryad.cvdncjth4) ##
 Description of the data and file structure Anatomical and phylogenetic
 analysis of a small rhynchocephalian fossil from the Helsby Sandstone
 Formation of Devon, UK. ### Files and variables #### File:
 OSFBeastie_mb_revised.txt **Description:**  MrBayes command file for
 phylogenetic analysis of the new specimen in context of modern and fossil
 Lepidosauria. #### File:
 Character_List_(modified_from_talanda_et_al_2022).txt **Description:**
 Modified character list for MrBayes file; including new taxa and a new
 character (ch383) #### File: OSFBeastie_CT21.tnt **Description:** tnt
 command file for phylogenetic analysis of the new specimen in context of
 modern and fossil Lepidosauria. #### File:
 Complete_Specimen_(Bristol_scan).stl **Description:** single object 3D
 file of the segmented Bristol scan of Agriodontosaurus #### File:
 Complete_Skull_Model_(ESRF_scan).stl **Description:** single object 3D
 file of the segmented ESRF scan of Agriodontosaurus #### Individual STL
 files: **Description:** individually segmented stl files for each skull
 element of Agriodontosaurus ##### File: L_Frontal(ESRF_scan).stl #####
 File: LDe_Teeth(ESRF_scan).stl ##### File: L_Jugal(ESRF_scan).stl #####
 File: LMx_Teeth(ESRF_scan).stl ##### File: L_Mandible(ESRF_scan).stl 
 ##### File: L_Parietal(ESRF_scan).stl ##### File: L_Maxilla(ESRF_scan).stl
 ##### File: RDe_Teeth(ESRF_scan).stl ##### File: RMx_Teeth(ESRF_scan).stl
 ##### File: L_Prefrontal(ESRF_scan).stl ##### File:
 L_Postfrontal(ESRF_scan).stl ##### File: L_Squamosal(ESRF_scan).stl #####
 File: L_Postorbital(ESRF_scan).stl ##### File: L_Quadrate(ESRF_scan).stl
 ##### File: R_Frontal(ESRF_scan).stl ##### File: R_Parietal(ESRF_scan).stl
 ##### File: R_Maxilla(ESRF_scan).stl ##### File: R_Mandible(ESRF_scan).stl
 ##### File: R_Prefrontal(ESRF_scan).stl ##### File:
 R_Postorbital(ESRF_scan).stl ##### File: R_Postfrontal(ESRF_scan).stl 
 ##### File: R_Quadrate(ESRF_scan).stl ##### File:
 VomerPalatinePterygoid(ESRF_scan).stl ## Code/software Both phylogenetic
 files are text files and can be viewed in any text viewer. The analyses
 will run in any versions of the free software MrBayes and tnt
 respectively.  STL files may be viewed in a pdf viewer or 3D programs such
 as Blender/3DSlicer/MeshLab etc