DOI: 10.5281/zenodo.15142274
title: Table 7 in Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam

description: Table 7. Examples of integrative taxonomy in other species of the genus Tylototriton. *partial 16S rRNA and COI, and complete tRNA Leu, ND1, ND2, tRNA Ile, tRNA Gln, tRNA Met, tRNA Trp, tRNA Ala, tRNA Asn, tRNA Cys, and tRNA Tyr.
SourceGene usedSpecies at stakeGenetic var. (%)Morphological variation and conclusionsZhang et al. (2007)partial cyt bT. shanjing × T. verrucosusmean 1.2; range 0.4-2.6Morphology not discussed. Conspecificity. Stuart et al. (2010), Nishikawa et al. (2013a) and Nishikawa et al. (2015) point out that only a single, unvouchered sample of T. verrucosus was included, and cautiously defended the need for further examinations before taxonomic conclusions.Khatiwada et al. (2015)complete ND2 & partial cyt bT. shanjing × T. verrucosus1.9Size; head proportions; grooves on tail base; coloration. "The topotypic T. verrucosus were deeply nested within T. shanjing."Khatiwada et al. (2015)complete ND2 & partial cyt bT. pulcherrimus × T. verrucosus; T. pulcherrimus × T. shanjing2.1; 2.8Not further discussed. Treated as separated species.Nishikawa et al. (2013a)partial ND2, POMC & Rag 1T. shanjing (Jingdong and Nu Jiang, Yunnan, China)mean 1.4; range 0.2-2.1Morphology not discussed. Treated as intraspecific variation.Nishikawa et al. (2013a)partial ND2, POMC & Rag 1T. shanjing (from above) × T. pulcherrimusmean 2.6; range 2.5-2.8Morphology not discussed. Considered conspecific due to small genetic differences.Nishikawa et al. (2015)partial ND2T. yangi × T. daweishanensismean 0.4; range 0.2-0.5Coloration. Considered conspecific due to small genetic differences.Khatiwada et al. (2015)complete ND2 & partial cyt bT. yangi × T. daweishanensismean 0.7Not discussed. Treated as separate species.Phimmachak et al. (2015a)*T. anguliceps × T. pulcherrimusmean 3.1; range 2.8-3.4Morphology not discussed. Treated as separate species and used as example for low genetic divergence within species of the genus.Phimmachak et al. (2015a)*T. podichthys (description) × T. pulcherrimusmean 2.9; range 2.5-3.4Morphology not discussed. The new species formed a unique clade within an unresolved polytomy containing T. verrucosus, T. shanjing, and T. pulcherrimus.Phimmachak et al. (2015a)*T. podichthys (description) × T. shanjingmean 3.4; range 2.1-4.6Ridge on midline of crown; coloration. Treated as separate species.Le et al. (2015)partial ND2T. podichthys (Xam Neua, Laos) × T. shanjing0.4When the paper was published, T. podichthys was not yet described. The authors referred to this population as T. verrucosus from Laos, which formed a clade with T. shanjing, T. pulcherrimus, and T. verrucosus from the type locality.Phimmachak et al. (2015a)*T. podichthys (description) × T. verrucosusmean 3.1; range 2.1-4.4Ridge on midline of crown; coloration; skin on cranial crest; orientation of parotoids. Separate species.Shen et al. (2012)complete ND2T. broadoridgus (description) × T. wenxianensismean 3.9; range 3.8-4Dorsal ridge; height of tail; presence of genital papillae; form of rib warts. Treated as separate species.Shen et al. (2012)complete ND2T. broadoridgus (description) × T. dabienicusmean 3.5; range 3.4-3.5Inferred morphological differences. Separate species.Nishikawa et al. (2013b)partial ND2T. broadoridgus × T. dabienicusmean 3.3; range 3.3-3.4Morphology not examined. Treated as separate species and used as example for low genetic divergence within species of the genus.Nishikawa et al. (2013a)partial ND2T. broadoridgus × T. dabienicus3.3Morphology not examined. The authors suspect conspecificity.Khatiwada et al. (2015)complete ND2 & partial cyt bT. broadoridgus × T. dabienicus3.4Not discussed. Treated as separate species.Stuart et al. (2010)* (except COI)T. notialis (description) × T. hainanensisrange 3.7-3.8Form of rib warts. Treated as separate species.Nishikawa et al. (2013b)partial ND2T. notialis (type from Laos) × T. notialis (Nghe An, Vietnam)2.8Coloration. Considered conspecific.Nishikawa et al. (2013b)partial ND2T. ziegleri (description)mean 1.7; range 0.1-2.8Treated as intraspecific variation.Nishikawa et al. (2013b)partial ND2T. asperrimus (China) × T. asperrimus (Thuong Tien, Hoa Binh, Vietnam)mean 2.7; range 0.1-3.4Morphology not examined. Considered conspecific by Nishikawa et al. (2013b); separate species, Tylototriton pasmansi, this study.Qian et al. (2017)ND1, ND2 & cyt bT. ziegleri x T. vietnamensis2.3Known morphological differences based on Nishikawa et al. (2013b): ridges on head; skin; vertebral ridge; form of rib warts; tail size and height. Treated as separate species.Qian et al. (2017)ND1, ND2 & cyt bT. anhuiensis (description) × T. broadoridgus3.2Dorsal ridge width; head proportions. Treated as separate species.Grismer et al. (2018)ND2T. ngarsuensis (description) × T. shanorumrange 3.0-3.4Size; head length; rib nodules; dorsal ridge; parotoid position; coloration. Treated as separate species.
description: Published as part of Bernardes, Marta, Le, Minh Duc, Nguyen, Truong Quang, Pham, Cuong The, Pham, Anh Van, Nguyen, Tao Thien, Roedder, Dennis, Bonkowski, Michael & Ziegler, Thomas, 2020, Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam, pp. 121 in ZooKeys 935 on page 121, DOI: 10.3897/zookeys.935.37138