DOI: 10.5281/zenodo.17081079
title: TABLE 3 in A new limestone-associated species of wolf snake in the Lycodon fasciatus complex (Squamata: Serpentes: Colubridae) from central Vietnam

description: TABLE 3. Comparison of morphological characters of Lycodon calcarophilus sp. nov. with those of the Lycodon fasciatus group. Characters highlighted in bold are considered diagnostic (with respect to Lycodon calcarophilus sp. nov.). Numeric values provided for SVL refer to the maximum SVL attained by each species.
SpeciesMax SVL (mm)TaL/TLVENSCBBTBDSR keeled?CPLOR-EDorsal ground colorColor of light crossbandsVentral patternDistributionsSourcescalcarophilus sp. nov.5050.19–0.22214–21983–8819–2510–15KeeledEntireYesBlackWhite anteriorly, slight brownish-grey posteriorlyBandedVietnamThis studybutleri6480.19–0.24216–22881–9728–3714–25KeeledEntireYesGrey or blackWhite or greyBanded & spottedThailand, Peninsular MalaysiaGrismer et al. (2014); Nguyen et al. (2024a)davidi3080.21224998438KeeledEntireNoDark brownBrownish-greyBanded & spottedLaosVogel et al. (2012)cavernicolus4060.20–0.21232–24592–11336–4529–41KeeledEntireYesLight brownWhite or light brownPlainThailand; Peninsular MalaysiaGrismer et al. (2014); Vogel et al. (2024)fasciatus7060.19–0.22197–21374–9028–4310–20KeeledEntireYesBlack or dark brownWhite or light brown anteriorly, variable posteriorly, usually light brown or reddish brownBanded & spottedChina, Myanmar, India, Bhutan, Thailand, Laos, VietnamNguyen et al. (2024a); Vogel et al. (2024)gammiei11100.19–0.25205–228101–11242–5816–23KeeledEntireNoBlackYellow or olive yellowBandedIndia, Bhutan, ChinaMistry et al. (2007); Chettri & Bhupathy (2009); Shu et al. (2024)gongshan7980.21–0.27203–21679–9626–469–16KeeledEntireYesBlackWhite anteriorly, variable posteriorly, usually orange-brown or reddish-brownVariable (banded or banded & spotted)China, MyanmarVogel & Luo (2011); Huang et al. (2021); Nguyen et al. (2024a)latifasciatus6800.21–0.23206–21690–9629–3613–21KeeledEntireYes (No)Dark brown or blackOrange or light brownBanded & spottedChina, MyanmarNguyen et al. (2025)liuchengchaoi6760.16–0.22200–22868–8118–547–16KeeledDivided or EntireYesBlackCream anteriorly, yellow, orange or light brown posteriorlyBandedChinaZhang et al. (2011); Nguyen et al. (2024a)multifasciatus5700. 23–0.26227–237106–11751–7225–35KeeledEntireNoDark brownLight brownPlainJapanMaki (1931); Vogel et al. (2009); Nguyen et al. (2024a)multizonatus5050. 16–0.18191–20256–7555–6211–19KeeledDividedYesBlackOrange or reddishbrownVariable (plain or banded)ChinaLei et al. (2014); Nguyen et al. (2024a)obvelatus4470.19199763113SmoothEntireYesBlackLight orange or salmonBanded & spottedChinaWang et al. (2021); Nguyen et al. (2024a)pictus5430.18–0.23209–21882–9125–328–16SmoothEntireYes or NoBlack or dark brownWhite or cream anteriorly, light brown posteriorlyBandedVietnam, ChinaJanssen et al. (2019, 2020); Xu et al. (2023); Nguyen et al. (2024a)ruhstrati abditus7620.19–0.25210–23090–11517–479–23KeeledEntireNoDark brownWhite anteriorly, white or grey-brown posteriorlyPlain & spottedChina, Vietnam, LaosThis studyruhstrati ruhstrati8100.21–0.25211–22899–11433–4614–25KeeledEntireNoBlack or dark brownGrey-brownPlain & spottedTaiwan IslandThis studyserratus4800. 24198846626SmoothDividedYesDark brownOlive brown (tawny olive)Banded & spottedChinaWang et al. (2021)sidiki6500.19–0.24195–21277–9125–3310–16KeeledEntireYesBlackLight greyBandedIndonesia (Sumatra)Wostl et al. (2017); Nguyen et al. (2024a)synaptor3950.19201–20368–6930–319KeeledEntireNoBlack or dark brownWhiteBandedChinaVogel & David (2010)yunnanensis5500.18–0.21186–20267–8020–367–12KeeledEntireYesBlack or dark brownWhiteBanded & spottedChina, VietnamVogel et al. (2024)
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description: Published as part of Vogel, Gernot, Bragin, Andrey M., Poyarkov, Nikolay A. & Nguyen, Tan Van, 2025, A new limestone-associated species of wolf snake in the Lycodon fasciatus complex (Squamata: Serpentes: Colubridae) from central Vietnam, pp. 10-11 in Zootaxa 5679 (1) on page 10, DOI: 10.11646/zootaxa.5679.1.1, http://zenodo.org/record/16983543