DOI: 10.5061/dryad.7722q
title: Data from: Morphology and phylogenetic relationships of fossil snake mackerels and cutlassfishes (Trichiuroidea) from the Eocene (Ypresian) London Clay Formation

description: ‘Gempylids’ (snake mackerels) and trichiurids (cutlassfishes) are pelagic
 fishes characterised by slender to eel-like bodies, deep-sea predatory
 ecologies, and large fang-like teeth. Several hypotheses of relationships
 between these groups have been proposed, but a consensus remains elusive.
 Fossils attributed to ‘gempylids’ and trichiurids consist almost
 exclusively of highly compressed body fossils and isolated teeth and
 otoliths. We use micro-computed tomography to redescribe two
 three-dimensional crania, historically assigned to †Eutrichiurides
 winkleri and †Progempylus edwardsi, as well as an isolated braincase
 (NHMUK PV OR 41318). All from the London Clay Formation (Eocene:
 Ypresian), these specimens represent some of the oldest fossils identified
 as trichiuroids. We find that †Eutrichiurides winkleri does not show
 diagnostic characters of †Eutrichiurides, and it is assigned to a new
 genus. In order to investigate the placement of these fossils relative to
 extant lineages, we combine existing morphological character sets for
 ‘gempylids’ and trichiurids along with published mitogenomic data in order
 to investigate the placement of these fossils relative to extant lineages.
 Our analyses recover a monophyletic Trichiuridae nested within a
 paraphyletic ‘Gempylidae’. The taxon formerly known as †Eutrichiurides
 winkleri is considered Trichiuroidea incertae sedis, while †Progempylus
 edwardsi and NHMUK PV OR 41318 are recovered within the ‘gempylid’ grade.
 Using previously published descriptions and character optimisations from
 our phylogenetic analyses we suggest possible placements for laterally
 compressed body fossils historically associated with Trichiuroidea
 (†Argestichthys, †Abadzekia, †Chelifichthys, †Anenchelum, †Eutrichiurides,
 †Musculopedunculus).
description: Macroynis casieri CT
 dataThis zip file contains
 the CT data for Macroynis casieri NHMUK PV 26904. Includes the .mcs file,
 which can be opened by the program Mimics, a vol and vgi file which can be
 opened in programs including the open source program Drishti in addition
 to segmentation programs like VGStudioMax, Aviso and Mimics, and PLY files
 for the segmented specimen which can be opened in open source programmes
 like Meshlab, Blender and others.Macroynis-casieri.zipProgempylus edwardsi CT dataThis zip file contains the CT data for Progempylus edwardsi NHMUK OR 32388. Includes the .mcs file, which can be opened by the program Mimics, a vol and vgi file which can be opened in programs including the open source program Drishti in addition to segmentation programs like VGStudioMax, Aviso and Mimics, and PLY files for the segmented specimen which can be opened in open source programmes like Meshlab, Blender and others.Progempylus-edwardsii.zipSupplementary Information S1 - Scan Parameters Radiography/CTAppendix S1: Scan parameters for radiography and computed tomography for taxa used in this study.Supplementary Information S1 - Scan Parameters Radiography:CT.xlsxSupplementary Information S2 – Character ListAppendix S2: Trichiuroid character list used in this study.Supplementary Information S3 - Morphological Character MatrixAppendix S3: Trichiuroid morphological character matrix used in this study. Character numbers refer to characters outlined in Supplementary Information S2.Supplementary Information S5 - Extant onlyAppendix S5: Maximum clade credibility tree from the Bayesian analysis of the combined morphology and molecular dataset (mitochondrial data from Miya et al. 2013) for extant trichiuroids. Bayesian posterior probabilities are indicated on nodes.Supplementary Information S4 - Optimisations MorphologyAppendix S4: Trichiuroid optimisations from morphological parsimony analysis. This pdf file is output from MacClade. A key is included.Supplementary Information S4 - Optimisations AA6.pdf