DOI: 10.5281/zenodo.15135592
title: Table 2 in Helen's twins in the Balkans: discovery of two new Paraptychoptera Tonnoir, 1919 species closely related to P. helena Peus, 1958, with systematic revision of the " lacustris " group (Diptera, Ptychopteridae)

description: Table 2. Morphological characters of male Paraptychoptera specimens used in the phylogenetic analyses.
1Antennal segments: (0), segment 3 equal with segments 4+5; (1) segment 3 shorter than segments 4+52Wing R2+3+4: oblique straight (0); curved or angled in the middle (1)3Wing: R2+3+4 length: (0)> 2 × length of rm; (1) <2 × length of rm4Male auxiliary sexual organ on abdominal sternite III: only cluster of setae (0); cluster of setae and distal lobes (1)5Male auxiliary sexual organ, distal lobe: absent or weakly developed (0), well developed, ventrally projected outer lip depressed in the middle (1)6Male auxiliary sexual organ, distal lobe: outer lip truncate or straight: (0) absent; (1) present7Median sclerotized strip of the auxiliary sexual organ with a transverse ornamentation: (0) absent; (1) present8Epandrial clasper: well sclerotized with squared basal lobe and complex ventral lobe (0); poorly sclerotized without basal lobe (1)9Epandrial clasper configuration: (0), simple cylindrical; (1), cylindrical with additional subterminal ventral lobes10Epandrial clasper configuration: (0), simple cylindrical; (1), cylindrical with basal lobes11Epandrial clasper apex: rounded (0), pointed and curved (1)12Epandrial subapical process: absent (0); present (1)13Epandrial subapical process bilobate: absent (0); present (1)14Epandrial subapical process bilobate, inferior arm chitinous and curved: (0) absent; (1) present15Epandrial subapical process bilobate, superior arm laterally compressed, lobate, inferior arm truncate: (0) absent; (1) present16Epandrial subapical process bilobate, both arms chitinous and curved: (0) absent; (1) present17Epandrial subapical process bilobate, superior arm beak-shaped, inferior arm triangular: (0) absent; (1) present18Epandrial subapical process with chitinous process having curved apex: (0) absent; (1) present19Epandrial subapical process with a rounded apex, and strong basal thorn, longer than subapical process: (0) absent; (1) present20Epanadrial subapical process with rounded aped and strong basal thorns, shorter than subapical process: (0) absent; (1) present21Hypoproct: reduced to paired rectangular plates in subepandrial membrane (0); hypoproct well developed, triangular or tongue like (1)22Hypoproct short triangular lobe with rounded apex: (0) absent; (1) present23Hypoproct long, tongue like process: (0) absent (1) present24Hypoproct long, tongue like process, apex covered with dense hear: (0) absent (1) present25Hypoproct long, tongue like process, apex glabrous and bilobate: (0) absent (1) present26Hypoproct long, tongue like process, apex pointed, harpoon-shaped: (0) absent (1) present27Hypoproct long, tongue like process, apex truncate or slightly depressed: (0) absent (1) present28Gonostylus, shape of anterior lobule: (0) scythe-like; (1), lobe-like with rostrum29Gonostylus, shape of medial lobule: lobe like (0); scythe-like (1)30Gonostylus, apical stylus of gonostylus apex with strong spines: absent (0); present (1)31Gonostylus, apical stylus or gonostylus: simple (0); inflated (1)32Gonostylus, secondary lobe of apical stylus: present (0); absent (1)33Hypandrium apex terminal division spade like, without trichoid sensilla: (0) present; (1) absent;34Hypandrium apex terminal division wide spade like, with trichoid sensilla (0) absent; (1) present35Hypandrium apex terminal division long lobe like process, with rounded apex: (0) absent; (1) present36Hypandrium apex terminal division long lobe like process with bilobate apex: (0) absent; (1) present37Hypandrium eversible sac extended anteriorly nearly to margin: (0), absent; (1), present38Hypandrium basal scale: (0), absent; (1) present39Hypandrium basal scale chitinous, hat shape: (0), absent; (1) present40Hypandrium basal scale chitinous, hat shape, with medial lobe (0), absent; (1) present41hypandrium: basal scale lobe like, well developed (0), absent; (1) present42Hypandrium basal scale lobe like, weakly developed (0), absent; (1) present43Aedeagus: ejaculatory apodeme size: (0), larger, than sperm sac; (1), subequal44Aedeagus subapical plate wide spatulate: (0) present; (1) absent45Aedeagus subapical plate narrow, apex rounded or pointed: (0) present; (1) absent46Aedeagus subapical plate narrow, apex depressed: (0) present; (1) absent47Paramere apical lobes filiform process: (0) present; (1) absent48Paramere apical lobes with a notch in the middle: (0) present; (1) absent49Paramere apical lobes tip pointed: (0) present; (1) absent50Paramere apical lobes tip rounded: (0) present; (1) absent51Paramere lateral arms reduced: (0) present; (1) absent52Paramere lateral arms long, beak shaped: (0) present; (1) absent53Paramere lateral arms short, truncate: (0) present; (1) absent
description: Published as part of Keresztes, Lujza, Kappert, Juergen, Henning, Maria & Toeroek, Edina, 2021, Helen's twins in the Balkans: discovery of two new Paraptychoptera Tonnoir, 1919 species closely related to P. helena Peus, 1958, with systematic revision of the " lacustris " group (Diptera, Ptychopteridae), pp. 63 in ZooKeys 1071 on page 63, DOI: 10.3897/zookeys.1071.58598