DOI: 10.5281/zenodo.15160383
title: TABLE 1 in Description of a new species of the earthworm genus Moniligaster Perrier, 1872 (Clitellata, Moniligastridae) from the Nilgiri Biosphere Reserve of Tamil Nadu State, India

description: TABLE 1. Comparison of the characters of M. gatesi Ahmed & Narayanan, sp. nov., with other species of the ‘ gravelyi ’ group.
CharacterM. gravelyi Stephenson, 19151,2M. troyi Jamieson, 19773M. bahli Narayanan & Julka,20214M. blakemorei Narayanan & Julka, 20214,5M. keralensis Narayanan & Julka, 20214,5M. julkai Narayanan & Paliwal, 20225M. gatesi Ahmed & Narayanan, sp. nov.Length130–174 mm60–63 mm75–134 mm81–136 mm58–155 mm156–238 mm87–160 mmDiameter5–7 mm3.5–3.7 mm4.5–7 mm3.5–7 mm3–6 mm6.5–8.5 mm5–7 mmSegments139–189123–153135–200183–278109–201237–342116–181Spermathecal pore locationbetween bc setal lines, median to setae cin c setal linebetween bc, median to setae cclose to cd setal linesaligned between cd setal linesaligned at cd setal linesbetween bc setal lines, nearer to cClitellumannular, on segments ¼9– ½14 (= ¾4)annular, on segments ½9–14 (=5½)annular, on segments 10–13 (=4)annular, on segments 10–13 (=4)annular, on segments 10–13 (=4)annular, on segments 10–½14 (=4½)annular, on segments ⅓9–14 (=5⅓)Secondary male aperture or male pore locationaligned between bc setal lines, nearer to bon evaginated long cylindrical pseudopenes, at bc setal lines, lateral to blateral to seta b linesslightly lateral to b setal lineslightly lateral to b setal lineslateral to b setal linebetween bc setal lines, nearer to bFemale pore locationat b setal line?minute, at b setal lineminute, at b setal lineminute, slightly median to b setal lineminute, at b setal line?b setal lineSpermathecaatrial gland single, on each side; duct of the gland c. 1/7 the length of common atrial duct; ampulla ovoidatrial glands paired on each side; duct of each gland c. 1/4 the length of common atrial duct; ampulla elongate-ovoidatrial glands paired on each side; duct of each gland c. 1/5 the length of common atrial duct; ampulla ovoidatrial glands paired on each side; duct of each gland c. four times the length of common atrial duct; ampulla ovoidalatrial glands paired on each side; duct of each gland about as long as common atrial duct; ampulla elongate-ovoidatrial glands paired on each side; duct of each gland c. five times the length of common atrial duct; ampulla ovoidalatrial glands paired on each side; duct of the gland c. 1/6 the length of common atrial duct; ampulla pear-shapedProstatesstrap-shapedclub-shapedstrap-shapedbluntly club-shapedtubulartubularstrap-shapedProstate capsuletubular with deeply incised margins and highly nodulated surface(?)tubular with smooth margins and a few nodulations at ental endclub-shaped with smooth marginstubular with smooth marginstubular with smooth marginstubular, surface with shallow nodulations, margins nodulatedLength of prostatic ductjust over 1/4 the length of gland plus ductc.1/3 the length of gland plus ductc. 1/7 the length of gland plus ductc. 1/3 the length of gland plus ductc. 1/9 the length of gland plus ductc. 1/3 the length of gland plus ductc. 1/8 the length of gland plus ductVas deferenssegments 9–10, 11, enters prostate sub-entallysegment 9, enters prostate sub-ectallysegments 9–12(–16), enters prostate sub-entallysegments 9–11(– 12), enters prostate near entallysegments 9–10, enters prostate sub-ectallysegments 9–12(–13), enters prostate sub-ectallysegments 9–10, enters prostate sub-entallyGizzards4–5 (in segments 12–18)3 (in segments 13–15)3–4 (in segments 13–18)2 (in segments 12–16)3–6(in segments 14–20)2–3(in segments 12–16)4–5 (in segments 13–18)Intestinal originin segment 25±1in segment 27in segment 25±1in segment 27±1in segment 28±1in segment 27in segment 26±1Ovisacsextending to segments 13–14extending to segment 13extending to segment 20extending to segment 13extending to segments 12–16extending to segments 13–14extending to segments 13–16
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Data from: 1 Stephenson (1915); 2 Narayanan et al. (2023b); 3 Jamieson (1977); 4 Narayanan et al. (2021a); 5 Narayanan et al. (2022).
description: Published as part of Narayanan, S. Prasanth, Ahmed, Shakoor & Thomas, A. P., 2025, Description of a new species of the earthworm genus Moniligaster Perrier, 1872 (Clitellata, Moniligastridae) from the Nilgiri Biosphere Reserve of Tamil Nadu State, India, pp. 148-149 in Zootaxa 5589 (1) on page 148, DOI: 10.11646/zootaxa.5589.1.12, http://zenodo.org/record/14896843