DOI: 10.5061/dryad.z8w9ghxrz
title: Data from: Long-fuse evolution of carnivoran skeletal phenomes through the Cenozoic

description: Climatic change is hypothesized to promote phenotypic diversification.
 While neontological analyses are often used to test this hypothesis,
 extant data only capture time-averaged signals of surviving lineages. More
 nuanced tests require paired and longitudinal climatic and organismal
 data. Here, we developed the most comprehensive phenomic dataset of
 pan-carnivorans to test hypotheses that Cenozoic climatic change
 influenced the evolution of the cranial, appendicular, and axial skeleton.
 We found support that a hierarchical progression of ecological
 diversification across the Cenozoic significantly influenced the
 establishment of modern carnivorans. Specifically, extinctions during the
 Eocene-Oligocene Transition released crown carnivorans from a constrained
 adaptive zone to interfamilial skeletal diversification. Intrafamilial
 skeletal diversification did not occur for another 20 million years until
 after the Mid-Miocene Climate Transition. Our work demonstrates the
 essential role of macroevolutionary data from the fossil record for
 revealing how major global climatic events steered the evolutionary
 trajectories of modern skeletal phenomes.
description: # Data from: Long-fuse evolution of carnivoran skeletal phenomes through
 the Cenozoic Dataset DOI:
 [10.5061/dryad.z8w9ghxrz](10.5061/dryad.z8w9ghxrz) ## Description of the
 data and file structure tree_prune_final_2502015.tre contains the pruned
 phylogeny extant_data_250205.csv and extinct_data_250215.csv contain raw
 trait data. All traits measured in mm. Empty cells indicate that trait
 could not be measured. 
 
 Clade = clade association for each specimen. Either
 Carnivora, Carnivoramorpha, or Pan-carnivora (Hyaenodonta or Oxyaenodonta)
 Family = the family in which the specimen is a part of Species =
 specimen's species name Status = whether the specimen is extinct or
 extant Catalog = museum catalog number for each specimen 
 
 Geomean = geometric mean of all traits   Cranial traits CBL =
 condylobasal length POC = postorbital constriction breadth ZB = zygomatic
 breadth MB = mastoid breadth AFH = anterior facial height PFH = posterior
 facial height CVH = cranial vault height PW = palate width BCL =
 basicranial length cran_MRL = molar row length cran_M1L = first molar
 length cran_M1W = first molar width   mandibular traits MAT = moment arm
 of temporalis MAM = moment arm of masseter MAM2 = moment arm of masseter
 MOL = molar out-lever COL = canine out-lever MW = mandibular width AMW =
 anterior mandibular width man_MRL = molar row length man_M1L = first molar
 length man_M1W = first molar width   vertebral traits C3 = third cervical
 vertebrae; C5 = fifth cervical vertebrae; T1 = first thoracic vertebrae;
 midT = middle thoracic vertebrae; diaT = diaphragmatic thoracic vertebrae;
 lastT = last thoracic vertebrae; L1 = first lumbar vertebrae; midL =
 middle lumbar vertebrae; lastL = last lumbar vertebrae VW = vertebral
 width VH = vertebral height CL = centrum length CW = centrum width
 (posterior) CH = centrum height (posterior) ZL = inter-zygapophyseal
 length LW = mid-lamina width NSH = neural spine height TPDV = transverse
 process/diapophysis dorsoventral angle PZA = pre-zygapophysis joint angle
 NSA = neural spine angle TPAP = transverse process/diapophysis
 anteroposterior angle sacral_L = sacral length sacral_H = sacral height
 sacral_W = sacral width scap_L = scapula length scap_W = scapula width
 hum_L = humerus length hum_D = humerus mid-shaft width hum_prox = humerus
 proximal width hum_dist = humerus distal width ul_L = ulna length ul_D =
 ulna mid-shaft width ul_OL = ulnar olecranon length rad_L = radius length
 rad_D = radius mid-shaft width MC3L = 3rd metacarpel length MC3W = 3rd
 metacarpel width PPL = Manus proximal phalanx length PPW = Manus proximal
 phalanx width pel_L = pelvis length fem_L = femur length fem_D = Femur
 mid-shaft width fem_EB = Femur distal width fem_GT = Height of the greater
 trochanter of the femur tib_L = tibia length tib_D = tibia mid-shaft width
 tib_prox = tibia proximal width tib_dist = tibia distal width fib_L =
 fibula length cal_L = Calcaneus length MT3L = 3rd metatarsal length MT3D =
 3rd metatarsal width