DOI: 10.5281/zenodo.16898347
title: APPENDIX I in From Serendipity to Systematics: Tracing the Trails of Cockroach Research in India

description: APPENDIX I. Type locality data and additional species records as per the updated taxonomic changes (CSF database) to the existing extant cockroach species checklist from India by Gupta & Chandra (2019).
Systematic AccountType locality as in original description (and its present-day location/name)Voucher specimen depositorySuperfamily BlaberoideaFamily BlaberidaeSub-Family BlaberinaeGenus Blaberus Serville, 18311Blaberus giganteus (Linnaeus, 1758)AmericaBMNHSub-Family DiplopterinaeGenus Diploptera Saussure, 18642Diploptera punctata (Eschscholtz, 1822)North-Central Pacific, HawaiiZMMUSub-Family EpilamprinaeGenus Aptera Saussure, 18643Aptera brindlei Anisyutkin, 2017 *Poona (=Pune), Maharashtra, IndiaMMUMGenus Calolampra Saussure, 18934Calolampra irrorata (Fabricius, 1775)Nova Hollandia (=New Holland, Australia)ZMUCGenus Indoapterolampra Anisyutkin, 20145Indoapterolampra rugosiuscula Anisyutkin, 2014 $Coimbatore, Tamil Nadu, IndiaMMUMGenus Opisthoplatia Brunner von Wattenwyl, 18656Opisthoplatia orientalis (Burmeister, 1838)ChinaNRGenus Princisola Gurney & Roth, 19767Princisola pulchra (Shelford, 1910) $Nighiris, Coonor, Tamil Nadu, IndiaMNHNGenus Rhicnoda Brunner von Wattenwyl, 18938Rhicnoda natatrix Shelford, 1907SarawakUMOGenus Haanina Hebard, 19299Haanina cariniceps (Bolívar, 1897) *Tamil Nadu, IndiaMNHN10Haanina maindroni (Shelford, 1910) *Malabar (=Kerala), IndiaMNHN11Haanina major (Saussure, 1895)JavaNR12Haanina patinifera (Bolívar, 1897) $Maduŕe (=Madurai), IndiaMNHNGenus Morphna Shelford, 191013Morphna amplipennis (Walker, 1868) *Silhet (Sylhet, Assam), IndiaBMNH14Morphna auriculata (Brunner von Wattenwyl, 1865) *Bengale (=West Bengal), IndiaNR15Morphna decolyi (Bolívar, 1897) $Anamalai Hills, Tamil Nadu, IndiaMNHN16Morphna indica Anisyutkin & Yushkova, 2017 $Cardamon Hills, Kerala, IndiaMHNG17Morphna lucida Anisyutkin, 2014 *Devala, Nilgiri hills, Tamil Nadu, IndiaMMUM18Morphna plana (Brunner von Wattenwyl, 1865)Ceylon (Sri Lanka)NMWGenus Rhabdoblatta Kirby, 190319Rhabdoblatta excelsa (Navas, 1904) $Kurseong, Darjiling, West Bengal, IndiaNR20Rhabdoblatta lineaticollis (Bolívar, 1897) $IndiaMNHN21Rhabdoblatta melanosoma (Saussure, 1869) *Les Indes Orientales (=Tamil Nadu, India)MHNG22Rhabdoblatta ridleyi (Kirby, 1903)SingaporeBMNH23Rhabdoblatta simlansis (Baijal & Kapoor, 1996) *Himachal Pradesh, IndiaNR24Rhabdoblatta subsparsa (Walker, 1868)Ceylon (Sri Lanka)ANSPGenus Stictolampra Hanitsch, 193025Stictolampra buqueti (Serville, 1839)JavaNR26Stictolampra plicata (Navas, 1904) $Kurseong, Himalayas (West Bengal), IndiaNRGenus Phlebonotus Saussure, 186227Phlebonotus anomalus Saussure, 1863 $Les Indes Orientales. Neelgheries. Pondicheri (=Puducherry), IndiaNR28Phlebonotus pallens (Serville, 1831) ##Indes Orientales (=Tamil Nadu)MNHNGenus Thorax Saussure, 186229Thorax porcellana (Saussure, 1862) $Indes (=India)/ Les Indes orientales. Neelgheries (Tamil Nadu, India)MHNGSub-Family CalolamprodinaeGenus Calolamprodes Bey-Bienko, 196930Calolamprodes (Calolamprodes) characterosa (Walker, 1868) $Bengale (=West Bengal), IndiaBMNHSub-Family RhabdoblattellinaeGenus Rhabdoblattella Anisyutkin, 199931Rhabdoblattella alexeevi Anisyutkin & Yushkova, 2017 *Cardamon Hills, Muttapatti near Munnar, Kerala, IndiaMHNG32Rhabdoblattella euptera Anisyutkin & Yushkova, 2017 *Cardamon Hills, Valara Fall, 46 km SW of Munnar, Kerala, IndiaMHNGSubfamily OxyhaloinaeGenus Nauphoeta Burmeister, 183833Nauphoeta cinérea (Olivier, 1789)FranceZMUCSubfamily PanchlorinaeGenus Panchlora Burmeister, 183834Panchlora nivea (Linnaeus, 1758)AmericaBMNHSubfamily PanesthiinaeGenus Caeparia Stål, 187735Caeparia saussurei (Wood-Mason, 1876) *Sikkim, IndiaMHNGGenus Microdina Kirby, 190336Microdina forceps (Saussure, 1869) *Pondicheri (=Puducherry), Indes OrientalesBMNHGenus Miopanesthia Saussure, 189537Miopanesthia bigibbosa Saussure, 1895 $India Orientalis, Sikkim, Khasi Hills, IndiaBMNH38Miopanesthia discoidalis Saussure, 1895 **Java. India, Pondicheri (=Puducherry)BMNHGenus Panesthia Serville, 183139Panesthia angustipennis angustipennis (Illiger, 1801)Sumatra40Panesthia angustipennis cognata Bey-Bienko, 1969Yunnan, ChinaZILR41Panesthia birmanica Brunner von Wattenwyl, 1893Mt Mooleyit, Tanasserilli [Burma] (=Myanmar)NMW42Panesthia cribrata Saussure, 1864New Holland, USAMHNG43Panesthia flavipennis Wood-Mason, 1876 $Naga hills, Bhramputra Valley, Dikrang valley, IndiaBMNH44Panesthia lucanoides (Butler, 1882) *South IndiaBMNH45Panesthia missimensis Roth, 1979Morobe District, Mt Missim, Papua New GuineaMCZ46Panesthia monstruosa Wood-Mason, 1876 *Southern IndiaBMNH47Panesthia morsu s (Butler, 1882) *South IndiaBMNH48Panesthia morosa Kirby, 1903 *֎Anamalai Hills, Tamil Nadu, IndiaBMNH49Panesthia necrophoroides Walker, 1868Tenasserim, Bruma (=Myanmar)BMNH50Panesthia paramonstruosa Roth, 1979 *8 miles NE from Munnar, IndiaCAS51Panesthia plagiata Walker, 1859Ceylon (Sri Lanka)BMNH52Panesthia pygmaea Princis, 1951 *Port Blair, Andaman Islands, IndiaZMUC53Panesthia quinquedentata Kirby, 1903 *Nilgiris, IndiaBMNH54Panesthia regalis Walker, 1868 $Silhet (Assam), IndiaBMNH55Panesthia serrata Saussure, 1895 *Sink (=?), IndiaMHNG56Panesthia stellata Saussure, 1895 $Sikkim, India orientalis, IndiaMHNG?57Panesthia wallacei Wood-Mason, 1876Sinkep Island, near SingaporeNRGenus Salganea Stål, 187758Salganea biglumis (Saussure, 1895) $Sikkim, IndiaMus. Calcutta59Salganea cavagnaroi Roth, 1979 *NW of Pykara, Tamil Nadu, IndiaCAS60Salganea erythronota Bolívar, 1897 *Maduré (=Madurai), IndiaMNHN61Salganea incerta (Brunner, 1893)Mt Mooleyit, Tenasserim, BurmaNMW(=Myanmar)62Salganea indica Princis, 1953 *Anamalai Hills, IndiaNMM,63Salganea kodaikanalensis Roth, 1979 $Kodaikanal, Palni Hills (=Palani Hills), south India (=Tamil Nadu), IndiaANSP64Salganea morio (Burmeister, 1838)New Guinea and New Holland (=Australia)Berlin Muesum65Salganea passaloides Walker, 1868Ceylon (Sri Lanka)MBNH66Salganea raggei Roth, 1979Mt Angka, ThailandMCZ67Salganea rehni Roth, 1979 $Inde méridionale, Attakatti Shola am Iber Hill (=Tamil Nadu), IndiaANSPSubfamily ParanauphoetinaeGenus Paranauphoeta Brunner von Wattenwyl, 186568Paranauphoeta affinis Shelford, 1906BhutanMNHN69Paranauphoeta circumdata (De Haan, 1842)IndonesiaNR70Paranauphoeta indica Saussure & Zehntner, 1895 $Assam, IndiaNR71Paranauphoeta limbata Saussure, 1869 *Les Indes Orientales (=Tamil Nadu), IndiaNRSubfamily PerisphaerinaeGenus Perisphaerus Serville, 183172Perisphaerus altus Walker, 1868 $Silhet (Assam), IndiaBMNH73Perisphaerus armadillo Serville, 1831JavaMNHN74Perisphaerus cotesianus (Saussure & Zehntner, 1895) *Assam, IndiaZSIC75Perisphaerus flavicornis (Burmeister, 1838)JavaNRGenus Pseudoglomeris Brunner von Wattenwyl, 189376Pseudoglomeris glomeris (Saussure, 1863)Ceylon (Sri Lanka)BMNH77Pseudoglomeris terranea (Walker, 1871) *Bombay (=Mumbai), IndiaBMNH78Pseudoglomeris guerini (Saussure & Zehntner, 1895) *India Orientalis (=West Bengal), IndiaMHNG79Pseudoglomeris humbertiana (Saussure, 1863)Ceylon (Sri Lanka)MHNG80Pseudoglomeris sericea (Saussure, 1863) *Les Indes orientales. Pondichery (=Puducherry), IndiaMHNGSubfamily PycnoscelinaeGenus Pycnoscelus Scudder, 186281Pycnoscelus indicu s (Fabricius, 1775) *India Orientali (=West Bengal), IndiaZMUC82Pycnoscelus nigra (Brunner von Wattenwyl, 1865)Birma (=Myanmar)NR83Pycnoscelus surinamensis (Linnaeus, 1758)SurinamiBMNH84Pycnoscelus tenebrigera (Walker, 1868) $Hindostan (=Hindustan/India)BMNHGenus Stilpnoblatta Saussure & Zehntner, 189585Stilpnoblatta opaca (Walker, 1868)Ceylon (Sri Lanka)BMNHFamily PseudophyllodromiidaeSubfamily AnaplectoidinaeGenus Anaplectella Hanitsch, 192886Anaplectella indica Bey-Bienko, 1969 $Haidarabad (=Hyderabad), South IndiaZILR87Anaplectella subrotundata (Walker, 1871) *Bombay (=Mumbai), IndiaBMNHGenus Malaccina Hebard, 192988Malaccina pallidula (Bolívar, 1897) $Kodaikanal, Tamil Nadu, IndiaMNHNSubfamily PseudophyllodromiinaeGenus Allacta Saussure & Zehntner, 189589Allacta crassivenosa Bolívar, 1897 *Kodaikanal, Tamil Nadu, IndiaMNHN90Allacta diluta Saussure, 1863Ceylon (Sri Lanka)MHNG91Allacta figurata (Walker, 1871) $Bombay (=Mumbai), IndiaBMNH92Allacta jcenpro Senraj, Packiam, Prabakaran, Lucanas, and Jaiswal, 2021*K.K. Nagar, Chennai, Tamil Nadu, IndiaZSI, SRC93Allacta kalakadensis Prabakaran & Senraj, 2019 *Thirukurungudi Range, Valaiyathu odai, Tamil Nadu, IndiaZSI, SRC94Allacta kollimalai Senraj, Packiam, Prabakaran, Lucanas, and Jaiswal, 2021*Valappurnadu View Point, Namakkal, Tamil Nadu, IndiaZSI, SRC95Allacta vellimalai Senraj, Packiam, Prabakaran, Lucanas, and Jaiswal, 2021*Vellimalai Forest, Megamalai Range, Theni, Tamil Nadu, IndiaZSI, SRCGenus Balta Tepper, 189396Balta acutiventris (Chopard, 1924) $Barkuda Island, Chilika Lake, Orrisa (=Odisha), IndiaNR97Balta chopardi Princis, 1969 *Barkuda Island, Chilika Lake, Orrisa (=Odisha), IndiaNR98Balta reticulata (Fabricius, 1798) *India Orientali (=Transquebar), IndiaZMUCGenus Ceratinoptera Brunner von Wattenwyl, 186599Ceratinoptera dispar Chopard, 1924 *֎Barkuda Island, Chilika Lake, Orrisa (=Odisha), IndiaProbably IMCGenus Lupparia Walker, 1868100Lupparia ferruginea (Brunner von Wattenwyl, 1865) *Indes Orientales (=Tiruchirappalli, TN)NR101Lupparia unguiculata (Bolívar, 1897) *Maduŕe (=Madurai), IndiaMNHNGenus Plectoptera Saussure, 1864102Plectoptera insularis Princis, 1951West Indies?ZMUCGenus Pseudectobia Saussure, 1869103Pseudectobia luneli (Saussure, 1868) *India Orientalis (=West Bengal)NRGenus Supella Shelford, 1911104Supella (Supella) longipalpa (Fabricius, 1798) ***India Orientali (=Transqubar, TN)ZMUCFamily BlattellidaeGenus Blattellina Princis, 1951105Blattellina indica Princis, 1951 *Northeast India, Santal Parganas (= Jharkhand)ZMUCGenus Episymploce Bey-Bienko, 1950106Episymploce annulifera (Walker, 1871) *Bombay (=Mumbai), IndiaBMNH107Episymploce brevipes (Walker, 1871) *Bombay (=Mumbai), IndiaBMNH108Episymploce himalayica (Brunner von Wattenwyl, 1865) *Himalaya (=Sikkim), IndiaNMW109Episymploce kuluana Roth, 1987 *Kulu, Himachal Pradesh, IndiaMCZ110Episymploce suknana Roth, 1987 *IndiaBPBM111Episymploce telephoroides (Walker, 1871) *Bombay (=Mumbai), IndiaBMNHGenus Eublattella Princis, 1950112Eublattella latistriga (Walker, 1868)ChinaBMNHGenus Haplosymploce Hanitsch, 1933113Haplosymploce andamanica (Princis, 1951) *Port Blair, Andaman Islands, IndiaZMUCGenus Hemipterisca Bey-Bienko, 1950114Hemipterisca submarginata (Walker, 1871) *Cahar, Assam, IndiaBMNHGenus Hemithyrsocera Saussure, 1893115Hemithyrsocera alagara Roth, 1995 *Alagar hills, Madurai, IndiaMCZ116Hemithyrsocera ignobilis Shelford, 1906 *Khasia (=Khasi) Hills, Meghalaya, IndiaUMO117Hemithyrsocera lycoides (Walker, 1871) *Malabar (=Kerala), IndiaBMNH118Hemithyrsocera nathani (Roth, 1995) *Annamalai Hill, Tamil Nadu, IndiaZMA119Hemithyrsocera palliata (Fabricius, 1798) ***Tranquebariae (=Transqubar, TN), IndiaZMUC120Hemithyrsocera suspecta Bolívar, 1897 *Tamil Nadu, IndiaMNHN121Hemithyrsocera vittata (Brunner von Wattenwyl, 1865)CambodiaBMNHGenus Jacobsonina Hebard, 1929122Jacobsonina platysoma (Walker, 1868)Swan River, AustraliaBMNHGenus Lobopterella Princis, 1957123Lobopterella dimidiatipes (Bolívar, 1890)Causip, PhillipinesMNHNGenus Neoloboptera Princis, 1953124Neoloboptera chakrabortyi Roth, 1995 *Alagar hills, Madurai, TN, IndiaMCZ125Neoloboptera indica (Brunner von Wattenwyl, 1865) *Indes (=India)NRGenus Parcoblatta Hebard, 1917126Parcoblatta pennsylvanica (De Geer, 1773)America BorealisMCZGenus Symploce Hebard, 1916127Symploce indica (Brunner Von Wattenwyl, 1865)MalaccaNMW128Symploce fulvastra (Chopard, 1924) *֎Barkuda Island, Chilika Lake, Orrisa (=Odisha), IndiaNRGenus Symplocodes Hebard, 1929129Symplocodes impar Bey-Bienko, 1969 *104 km south of Madras (=Chennai), Tamil Nadu, IndiaZILRGenus Pholeosilpha Chopard, 1958130Pholeosilpha induta (Bolívar, 1897) *Maduŕe (=Madurai) Tamil Nadu, IndiaNRSubfamily BlattellinaeGenus Blattella Caudell, 1903131Blattella asahina i Mizukubo, 1981Okinawa Island, JapanKU132Blattella biligata (Walker, 1868)Ceylon (Sri Lanka) + Bombay (=Mumbai), IndiaBMNH133Blattella bisignata (Brunner von Wattenwyl, 1893)Bhamo, MyanmarNMWA134Blattella germanica (Linnaeus, 1767)DenmarkLSUK135Blattella humbertiana (Saussure, 1863)Ceylon (Sri Lanka)BMNH136Blattella karnyi (Princis, 1969)Fuhosho, TaiwanNR137Blattella lituricollis (Walker, 1868)Amoy China (Now-Kinmen Islands)BMNH138Blattella nipponica Asahina, 1963Tanegashima, JapanNIH139Blattella padmanabhani Roth, 1995 *Alagar Hill, Madurai, Tamil Nadu, IndiaMCZ140Blattella teressensis Roth, 1985 *Terressa Island, Nicobar Islands, IndiaZMUC141Blattella vaga Hebard, 1935Phoenix, ArizonaANSPFamily EctobiidaeSubfamily EctobiinaeGenus Ectobius Stephens, 1835142Ectobius (Ectobius) indicus Bei-Bienko, 1938 *North West IndiaBMNH143Ectobius (Ectobius) jarringi (Hanitsch, 1937*Tragbal, Kashmir, IndiaZILSSuperfamily BlattoideaFamily BlattidaeSubfamily BlattinaeGenus Archiblatta Snellen van Vollenhoven, 1862144Archiblatta phalangium (Saussure, 1864) *India Orientalis (=West Bengal)NRGenus Blatta Linnaeus, 1758145Blatta orientalis Linnaeus, 1758America, OrienteLSUKGenus Cartoblatta Shelford, 1910146Cartoblatta speciosa (Shelford, 1910) *Sangli, Maharashtra, IndiaUMOGenus Dorylaea Stål, 1877147Dorylaea semimarginalis (Hanitsch, 1915)Kuching, Sarawak, MalaysiaSMMGenus Validiblatta Luo & Wang, 2025148Validiblatta brunnea Burmeister, 1838Chile and Demerary (South America)BMNH149Validiblatta australasiae (Fabricius, 1775)Unspecified primary typeBMNHGenus Homalosilpha Stål, 1874150Homalosilpha decorata (Serville, 1839)Sarawak, MalaysiaUMO151Homalosilpha ustulata (Burmeister, 1838)JavaMHNGGenus Neostylopyga Shelford, 1911152Neostylopyga maindroni (Shelford, 1911) *Nilghiris, Coonoor (town in Tamil Nadu), IndiaMNHN153Neostylopyga ornata (Brunner von Wattenwyl, 1865) $Indes Orientales (=Tiruchirappalli, TN)NR154Neostylopyga parallela (Bolivar, 1897) *Maduŕe (=Madurai), Tamil Nadu, India155Neostylopyga picea (Brunner von Wattenwyl, 1865) *Ile de Sambelong (Southern Group of Nicobar Islands), IndiaMNI156Neostylopyga rhombifolia (Stoll, 1813) ***Bombay (=Mumbai), IndiaBMNH157Neostylopyga sexpustulata (Walker, 1871) $Bombay (=Mumbai), IndiaBMNHGenus Periplaneta Burmeister, 1838158Periplaneta affinis Saussure, 1869 *πLes Indes Orientales (=Tamil Nadu), IndiaMHNG159Periplaneta americana (Linnaeus, 1758)AmericaBMNH160Periplaneta atemeleta Princis, 1966 *Indien, Merkara (=Madikeri), Karnataka, IndiaMSG161Periplaneta ebneri Princis, 1951 *Tranquebar, Indien (Transquebar, India)ZMUC162Periplaneta lateralis (Walker, 1868)EgyptBMNH163Periplaneta monochroma (Walker, 1871) *Bombay (=Mumbai), IndiaBMNHGenus Tenumembrana Luo & Wang, 2025164Tenumembrana ceylonica (Karny, 1908)Ceylon (=Sri Lanka)NMWSubfamily HebardininaeGenus Hebardina Bey-Bienko, 1938165Hebardina concinna (Haan, 1842)JavaMusée de BruxellesSubfamily PolyzosteriinaeGenus Melanozosteria Stål, 1874166Melanozosteria castanea (Brunner von Wattenwyl, 1865)Sidney, AustraliaBMNHGenus Polyzosteria Burmeister, 1838167Polyzosteria indica (Saussure, 1863) *֎Les Indes Orientales. Pondichery (=Puducherry), IndiaMHNGGenus Platyzosteria Brunner von Wattenwyl, 1865168Platyzosteria melanaria (Erichson, 1842) *Bombay (=Mumbai), IndiaBMNHSuperfamily CorydioideaFamily CorydiidaeSubfamily CorydiinaeGenus Ergaula Walker, 1868169Ergaula nepalensis (Saussure, 1893) $Sikkim, IndiaSMNHGenus Eucorydia Hebard, 1929170Eucorydia aenea aenea (Brunner von Wattenwyl, 1865) *Indes Orientales (=Tiruchirappalli) Tamil Nadu, IndiaBMNH171Eucorydia aenea plagiata (Walker, 1868) *Hindostan (=India)BMNH172Eucorydia ornata (Saussure, 1864) *Bombay (=Mumbai), IndiaBMNH173Eucorydia westwoodi (Gerstaecker, 1861) $Assam, IndiaNRGenus Polyphaga Brullé, 1835174Polyphaga aegyptiaca (Linnaeus, 1758)EgyptUUZM175Polyphaga indica Walker, 1868 $Hindostan (=India)BMNH176Polyphaga saussurei (Dohrn, 1888)Turcomannia (=Turkmenia)BMNHGenus Therea Billberg, 1820175Therea bernhardti Fritzsche, 2009 $Thiruvannamalai, Tamil Nadu, IndiaNZSI178Therea defranceschii Grandcolas, 1993 $Indes Orientales, Trichinopoli (=Tiruchirappalli), Tamil Nadu, IndiaMNHN179Therea hyperguttata Grandcolas, 1993 *Inde (Bellary) (=Ballari), Karnataka, IndiaMNHN180Therea irreperta Fritzsche, 2009 *IndiaUMO181Therea nuptialis (Gerstaecker, 1861) $Les Indes Orientales (=Puducherry), IndiaBMNH182Therea olegrandjeani Fritzsche & Zompro, 2008 *Gooty, Andra Pradesh, IndiaNZSI183Therea petiveriana (Linnaeus, 1758) $IndiaBMNH184Therea regularis (Grandcolas, 1993) $Coromandel (=Tamil Nadu coast), IndiaMNHNSubfamily EuthyrrhaphinaeGenus Euthyrrhapha Burmeister, 1838185Euthyrrhapha pacifica (Coquebert de Montbret, 1804) ֎BrazilBMNHFamily NocticolidaeSubfamily LatindiinaeGenus Ipolatta Karny, 1914186Ipolatta paradoxa Karny, 1914 *Khasi hills, Assam (=Meghalaya), IndiaNMWSubfamily NocticolinaeGenus Alluaudellina Chopard, 1932187Alluaudellina himalayensis (Gravely, 1910) *Kurseong, Darjeeling, IndiaIMCGenus Typhloblatta Chopard, 1924188Typhloblatta caeca (Chopard, 1921) $Rupmath Cave, Jaintia Hills, Sylhet, Assam, IndiaNR
APPENDIX I. Type locality data and additional species records as per the updated taxonomic changes (CSF database) to the existing extant cockroach species checklist from India by Gupta & Chandra (2019).
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Note: ֎ = Newly added to Indian checklist (Gupta and Chandra, 2019). * = Species known only from original description and not been collected since then. π = Type specimen lost
** = Type locality uncertain, *** = Species having type locality in India and having wide range distribution globally, ## = Species described from and restricted to India and Sri Lanka; $ = Species collected are reported based on specimens after original description.
description: Published as part of Shabnam, A., Morey, Rashmi & Dinesh, K. P., 2025, From Serendipity to Systematics: Tracing the Trails of Cockroach Research in India, pp. 531-537 in Zootaxa 5661 (4) on page 531, DOI: 10.11646/zootaxa.5661.4.3, http://zenodo.org/record/16605792