DOI: 10.48579/pro/ogxugt
title: Replication data for: Fig 10.8, Top right, GPDs, clan size and clan type vs male ARS, 17 Dec 2025

description: FILE DESCRIPTION Keywords: GPDs, clan size, clan type, male ARS File name = Fig 10.8, Top right, GPDs, clan size and clan type vs male ARS, 17 Dec 2025 OLDFile name = GPDs, Clan TYPE vs Male ARS, 23 Sept 2021 Today I have created this file by starting with File name = GPDs, Male ARS vs clan size, 05 Sept 2021 Today I add variable = clan type In this file, clantype = 0.5 = Half-male clan, for which &gt;=1 are controlled by single male Clantype = 1 = One male clan Clantype = 2 = Multi-male clan Clantype = 0 = Male that lived alone, with no other females in his territory, who did not DISAPPEAR shortly after mating season If a GPD male mated and then disappeared in early April, I did not assign a clantype This file has been checked TWO TIMES for clan type on 23 Sept 2021. Notice that I have only ONE case of Clantype = 0. Almost no solitary GPD males. Other data were checked TWO TIME earlier. See below. Note that multi-male clans were VERY RARE at RMC and VCC, but were much more common at PEFO. All in this file from RMC and VCC. Microsats = microsatellites This file includes ONLY THOSE GPD MALES THAT COPULATED WITH &gt;=1 FEMALE-- OR MALES KNOWN TO SIRE OFFSPRING FROM MICROSATS--i.e., I have EXCLUDED GPD males for which I have no evidence of copulation from either observations or microsats Note: All GPD data are from RMC and VCC, with NO DATA from PEFO--because I do not have any information via microsats about siring of GPD offspring at PEFO OLDFile name = GPDs, Male ARS vs clan size, 05 Sept 2021 I started this file from File name = GPDs, Male RS for cops,babies,yrls, 01 April 2021 Details below for GPDs, Male RS for cops,babies,yrls, 01 April 2021 and related files My plan is to use only data from VCNP's RMC and VCC, because I hope to have microsat data on paternities from those colonies. In this plan, I will NOT include info from PEFO, from which I have no information on paternities from microsats I will keep info from PEFO on Sexpartners for now If a GPD male had &gt;=2 half clans, I used the LARGER of the two half-clans for clan size for the WTPD male. In 2013, for example, GPD male-8 had one half-clan of size 12 (8A) and two half clans of size 4 (8B and 8C). For GPD-male-8 for this file, I used clan size = 12 If TWO GPD males shared the same multi-male clan, I used the same clan size for both males. In 2013, for example, I used the same clan size of 20 for both male-0 and male-41. Multi-male clans were rare for GPDs. For 0-41 in 2013, I do not use clan size--because smaller clans with large territory almost certainly existed For 44 in 2013, I use smaller clan (44a) for 44 (clansize = 4), because I new so little about clan-44B For 28 in 2013, I no not use any information. Smaller clans within large territory seem likely. For 49 in 2017, I use clan size = 4 (no females in his other half-clan of clan size = 5) Clan sizes checked TWO TIMES on 05 Sept 2021 Other data were checked two times for earlier files. I DO use GPD male-31 in 2018. We observed only 2 cops, but he sired babies from 5 litters. Note on 23 Sept 2021: For clan size for males that disappeared, I used clan size in May for the clan where the disappearing male would have been. Note on 23 Sept 2021:Clan sizes are for MAY of each year OLDFile name = GPDs, Male ARS for cops, PEFO and VCNP, 11 April 2021 I created this file by combining data from the following two EXCEL files: File name = GPDs, Male RS for cops,babies,yrls, 01 April 2021 [and] File name = GPDs, Male ARS and LRS for cops, 13 Jan 2021 Details from both files immediately below OLD File name = GPDs, Male RS for cops,babies,yrls, 01 April 2021 I created this file from File name = GPDs, Male ARS-LRS summaries, 31 March 2021. See notes below. OKtouse = 0 = we did not see many of the copulations--probably &lt;50% OKtouse = 1 = we documented most of the copulations--probably &gt; 50% I checked all the data on Age and Sexpartners on the hard-copy summary sheets ONE TIME ONLY--- I found only 1 error, so I did not check a second time This file has been checked TWO TIMES on 02 April 2021 In particular, I checked the new data for Sexpartners, OK to use, and Age on 02 April 2021 For GPDs from VCC of VCNP, I use a suffix of "V" for every GPD marker and eartag-- to reduce possible confusion with GPDS from RMC of VCNP All data in this file are from GPD males that copulated with &gt;=1 female All data from males that were not observed to copulate are EXCLUDED from this file For several males listed in this file that copulated with &gt;=1 female, we have NO EVIDENCE FROM MICROSATS that those males sired &gt;=1 emergent juvenile. Copulating males that evidently did not sire any offspring include the following: For 2013: 28 (123) For 2014: 108 (C47), 25 (439), 32 (593) For 2015: 20#2 (815), 107 (C39) For 2016: 37#2 (C99) For 2017: 24 (203) For 2018: 32 (79), 6 (395). 14 (55), 47 (435), 17 (O235) Date = 31 March 2021 This new file checked TWO TIMES for Weanlings and Yearlings on 31 March-02 April 2021 Weanlings = cumulative number of babies sired by each male per year Yearlings = cumulative number of sired babies sired that survived for &gt;=9 months after weaning. OLDFile name = GPDs, Male ARS-LRS summaries, 31 March 2021 I made this file directly from file = GPDs, Male ARS-LRS for cops, babies, yrls, 30 March 2021 Details immediately below for GPDs, Male ARS-LRS for cops, babies, yrls, 30 March 2021 Name of file = GPDs, Male ARS-LRS for cops, babies, yrls, 30 March 2021 No information on survivorship of juveniles for 2016 (last year at VCNP's RMC) or 2018 (last year at VCNP's VCC) Number in parenthesis after number of babies or yearlings for a male = number of babies or yearlings ESTIMATED or INFERRED. I only made estimates or inferences that seemed reasonable. For example, if microsats showed that the single male that copulated with the mother sired 2 of 3 babies, I inferred paternity to that male for the third baby as well. Similarly, if microsats showed for a litter of 6 that one of two copulating males sired 2 babies and that the other of two copulating ales sired 2 babies, then I inferred paternity for 3 babies for both males. If a female copulated with 2 males, for example, and 1 of the copulating males sired three of four babies and we have NO INFORMATION from microsats for the fourth babies, then I inferred that the male that sired three of the babies (from microsats) also sired the fourth baby as well. I only made this inference if the one male sired AT LEAST HALF of babies. So, if a female copulated with two males and weaned five babies, for example, and if one of the copulating males sired two babies and we have NO INFORMATION from microsats for the other three babies, I have NOT inferred paternities for any of those 3 babies. For this file I have made no attempt to include information for body mass or age of the copulating males. Information on age and body mass vs male ARS and LRS is summarized in other chapters. This file has been checked TWO TIMES on 29-30 March 2021. I have excluded some cases for which paternity makes no sense. Dognumber does not equal integer--e.g., 15.5, 33.2, 58.5--means that GPD was added after I created an earlier, numbered file. File name = GPDs, Male ARS and LRS for cops, 13 Jan 2021 Date = 12 Jan 2021 This file is IDENTICAL to file = GPDs, MCs vs male body mass, ARS, 06 Jan 2021 I have created this identical file so that I can keep it with other ARS files See notes below from that file Note on 08 Feb 2021. These data are all from PEFO from 1989 through 1995 i.e., no data from RMC or VCC of VCNP Sexpartners = number of different females with whom a male copulated Weanlings = number of babies sired by a male, as determined from either cops or microsatellites Yearlings = number of sired emergent babies sired that survived for &gt;=9 months after weaning OLDFile name = GPDs, MCs vs male body mass, male cops, 08 Jan 2021 I derived this file from file = GPDs, MCs vs female body mass, ARS, 06 Jan 2021, which I checked ONE TIME ONLY on 05 Jan 2021 OLDFile name = GPDs, MCs vs male body mass, male cops, 08 Jan 2021 has been checked TWO TIMES ON 08 Jan 2021 A GPD male will have the SAME EARTAGS in different years, but will have a different Nyanzol marker in different years For most analyses, I will consider data from the same GPD male in different years to be statistically independent Oktouse = 1 means that we think we observed most or all copulations by a GPD male in a particular year Oktouse = 0 means that we think we DID NOT OBSERVE most or all copulations by a GPD male in a particular year I only use AGE if I was certain--or almost certain--about a GPD male's exact age Weanlings = all babies weaned by all females with whom a GPD male copulated. SOME of the babies might have been sired by the copulating male, but OTHER offspring might have been sired by other males that copulated with the GPD female Yearlings = all yearlings reared by all females with whom a GPD male copulated. SOME of the yrls might have been sired by the copulating male, but OTHER yrls might have been sired by other males that copulated with the GPD female SEXPARTNERS = minimal number of GPD females with whom GPD male copulated, with or without MCs by the copulating male Note GPD male with Dognum = 63.5--so that I could insert additional data for GPD male-07 (B17, C68) in 1993. Note also that GPD male-34 and GPD male 34#2 are DIFFERENT GPD MALES, and they have different Dognums and different Eartags These data used for Figure 10.8 in Hoogland 2026. This file ready for longterm storage on 14 Feb 2025. No-male clan had no resident sexually mature male in May Half-male clan shared a single sexually mature male in May with another clan One-male clan had a single resident sexually mature male in May Multi-male clan had &gt;=2 sexually mature males in May For more information about clan types, see Chapter 10 of Hoogland 2025 Marchwt = male body mass in February, March, or early April--just before during, or just after mating season Junewt = male body mass in late May or June, approximately when weanlings were first emerging from their natal burrows yrl = yearling No information for variable = "Dognumber" in this EXCEL file. Notice the several notes from several earlier versions of this EXCEL file. Data for Fig 10.8 for GPDs are all accurate. Cops = copulations File name = Fig 10.8, Top right, GPDs, clan size and clan type vs male ARS, 17 Dec 2025