DOI: 10.5061/dryad.fbg79cp8p
title: Data for: Direct and indirect ecosystem responses to vehicle compaction of soft sediments

description: In northeastern Pacific tidal flats, a native bioturbator, ghost shrimp
 Neotrypaea californiensis, engineers soft-sediment habitat and interacts
 antagonistically with bivalve shellfish. Vehicle compaction has been used
 in pest control of ghost shrimp, but this disturbance lacks quantitative
 evidence of its efficacy and environmental impacts. Through three large
 (~10 ha) experiments in Grays Harbor, Washington, USA, we tested the
 direct and indirect impacts of compaction by a tracked vehicle
 (MarshMaster) on ghost shrimp density, sediment conditions, and infauna.
 We also examined how oyster survival (cultch seeded with Pacific oysters,
 Magallana gigas) and waterbird usage of tidal flats responded
 post-compaction. In Experiment 1, compaction occurred on 13 Aug 2022 and
 19 May 2023, and sampling occurred seasonally over three years, including
 immediately post-compaction. In Experiment 2, compaction occurred on 2 Aug
 2023, and sampling occurred seasonally over one year. Due to minimal
 treatment effects at this site, a third experiment was set up on 0.16-ha
 beds that were compacted with 0, 1, 3, or 5 compaction passes on 24 Jul
 2024 and followed for a year. Compaction pushed &lt;20% of ghost
 shrimp to the surface, where they were vulnerable to predation and damage,
 yet did not significantly reduce subsurface densities within 1-2 days.
 Rather, declines in shrimp density and shifts to smaller size classes
 appeared at later sample timepoints and were more pronounced with more
 compaction passes. All compaction experiments resulted in firmer sediment
 for at least a year, even in the experiment where shrimp densities were
 unaffected by a single compaction pass. Where compaction briefly reduced
 shrimp densities below 50 m-2, sediment increased in mud and organic
 content and infauna increased in abundance, suggesting that these changes
 were mediated through reduced bioturbation rather than a direct impact of
 compaction. Similarly, multivariate responses of infauna appeared only in
 the experiments where compaction reduced shrimp densities. Habitat use by
 waterbirds was more influenced by tidal stage than by compaction;
 statistically, only dunlin (Calidris alpina) foraged more on compacted
 than on reference beds. Finally, although survival of outplanted oyster
 seed improved with compaction at one site, it remained too low (~34-40%
 yr-1) for viable farming.
description: # Data for: Direct and indirect ecosystem responses to vehicle compaction
 of soft sediments Dataset DOI:
 [10.5061/dryad.fbg79cp8p](https://doi.org/10.5061/dryad.fbg79cp8p) ##
 Description of the data and file structure Intertidal flats of sandy
 soft-sediment dominated by ghost shrimp (*Neotrypaea californiensis*) in
 Grays Harbor, Washington (USA) were compacted with a tracked vehicle
 (MarshMaster MM 2LX, 1 pound per square inch ground pressure). Because the
 tracks of the MarshMaster have a gap between them, successive rounds
 overlapped so that all surface experienced ground pressure. In Experiment
 1 (Damon Point), five 2-ha beds were established along an elevation
 contour of 0.2-0.6 m relative to mean lower low water (MLLW), and two beds
 (9 and 11) were compacted on 13 Aug 2022 and recompacted on 19 May 2023.
 In Experiment 2 (Westport Single), five 2-ha beds were established along
 an elevation contour of 0.4-0.5 m MLLW, and two beds (2 and 4) were
 compacted on 2 Aug 2023. In Experiment 3 (Westport Multiple), nine 0.16-ha
 beds were established along an elevation contour of 0.6 m MLLW; three were
 uncompacted (Reference) beds, and two each were compacted with 1x, 3x, and
 5x passes of the MarshMaster. Six stations (subsamples) per bed were used
 to record responses on the 2-ha beds established in Experiment 1 and 2,
 and four stations per bed were used in Experiment 3. For each
 post-compaction sampling event, these subsamples were measured for: a)
 ghost shrimp density by coring (five times to 70 cm within a 1 m^2^ area,
 totaling 0.0625 m^2^); b) penetrability of the sediments based on depth of
 penetration of a standard metal rod; c) organic content of sediment by
 collecting a small core (5.4 cm depth, 30 ml), drying, and ashing at 500
 C; d) mud content of sediment by dry-sieving the ashed sediment and
 determining the fraction that passed through a 63-micrometer screen; e)
 abundance and composition of invertebrate macro-infauna in cores (10 cm
 diameter, 10 cm depth) sieved to 500 micrometers. Additionally, at
 stations in Experiment 1 and 2, seeded cultch (small Pacific oysters,
 *Magallana gigas*, on a shell) were tethered on the ground and collected
 at intervals to determine oyster survival. Experiment 1 also was evaluated
 for bird use at low tide during spring migration (6 days in April 2023 and
 7 days in April 2024) by photographs from remote cameras. For all files,
 the "Notes" variable is entered as "none" when no
 notes were recorded. NA refers to data not available. ### Files and
 variables #### File: BirdCamR.csv **Description:** Counts of birds by
 taxon within the field of view of a Go-Pro Hero3+ camera attached to a
 pole 1 m above the tidal flat, deployed before the water ebbed and
 retrieved as the water flooded. All cameras were deployed at a large
 experimental site 0.2-0.6 m above mean lower low water at Damon Point,
 Grays Harbor, Washington (Experiment 1). ##### Variables * Date: Date when
 camera was deployed * GPSN: WGS84 geoposition of camera in north latitude
 * GPSW: WGS84 geoposition of camera, negative sign for west longitude *
 Camera: Number identifying specific camera * Bed : Number identifying the
 2-ha Bed in which the camera was deployed. Beds 9 and 11 experienced
 vehicle compaction. Beds 8, 10, and 12 were reference beds. * Stage: Tidal
 stage - Ebb was defined as the time at which the water’s edge passed the
 camera (when the frame appeared fully dry), and flood as the time at which
 it passed the bamboo stake 7 m from the camera (when the frame was fully
 flooded). Low water was defined as the timepoint halfway between the ebb
 and flood. The tidal stages were divided into six periods: 24-10 minutes
 before the ebb, 8 minutes before to 6 minutes after the ebb, 8-22 minutes
 after the ebb, 16 minutes around low water, 24-10 minutes before the
 flood, and 8 minutes before to 6 minutes after the flood. * NumberImages:
 Number of photographs at 2-minute intervals used at that Stage *
 DunlinNear: Number of Dunlin (*Calidris alpina*) closer than 7 m in the
 field of view * DunlinFar: Number of Dunlin (*Calidris alpina*) farther
 than 7 m but not beyond 14 m * DowitcherNear: Number of Dowitcher
 (*Limnodromus spp.*) closer than 7 m in the field of view * DowitcherFar:
 Number of Dowitcher (*Limnodromus spp.*) farther than 7 m but not beyond
 14 m * GullNear: Number of Gulls (*Larus* spp.) closer than 7 m in the
 field of view * GullFar: Number of Gulls (*Larus* spp.) farther than 7 m
 but not beyond 14 m * CrowNear: Number of Crows (*Corvus brachyrhychos*)
 closer than 7 m in the field of view * CrowFar: Number of Crows (*Corvus
 brachyrhychos*) farther than 7 m but not beyond 14 m * UnidNear:
 Unidentifiable birds closer than 7 m in the field of view * UnidFar:
 Unidentifiable birds farther than 7 m but not beyond 14 m * Notes:
 Description of factors that could influence data collection #### File:
 infauna.csv **Description:** Counts of macroinfauna by taxon in cores (10
 cm diameter, 10 cm depth) collected from Grays Harbor, Washington, in
 Experiment 1 (Damon Point, 0.2-0.6 m above mean lower low water),
 Experiment 2 (Westport, 0.4-0.5 m above mean lower low water), and
 Experiment 3 (Westport, 0.6 m above mean lower low water, identified as
 Westport Multiple). Treatments were applied at the scale of aquaculture
 beds, which were 2 ha in size in Experiment 1 and 2, but 0.16 ha in size
 in Experiment 3. Vehicle compaction occurred on Beds 9 and 11 at Damon
 Point, on Beds 2 and 4 at Westport, and in a gradient of compaction
 intensity (1x, 3x, 5x) at Westport Multiple. Infauna were collected on a
 500-micron mesh sieve. ##### Variables * Site: Experimental site *
 SampleCode: Code denoting the Bed and position on the bed where the core
 was collected * Bed: Code denoting the Bed * DateSampled: Date used to
 designate one timepoint of sampling, which could span several consecutive
 days * DateSampled_true: Actual date when core was collected *
 DaysPostComp: Core collection date relative to the experimental setup when
 vehicle compaction occurred * DateProcessed: Date when sample was sorted
 in the lab and infaunal taxa were identified and counted * GPSN: WGS84
 geoposition of core in north latitude * GPSW: WGS84 geoposition of core,
 negative sign for west longitude * Clinocardium_nuttallii: Species count
 (bivalve clam = cockle) * Cryptomya_californica: Species count (bivalve
 clam) * Macoma_spp: Count by genus (bivalve clam) * Mya_arenaria: Species
 count (bivalve clam) * Mytilus: Count by genus (bivalve mussel) *
 Nutricola: Count by genus (bivalve clam) * Nuttalia: Count by species
 (*Nuttallia obscurata*, bivalve clam) * Ruditapes_philippinarum: Count by
 species (bivalve clam) * Protothaca_staminea_(Leukoma): Count by species
 (bivalve clam) * Tellina_modesta: Count by species (bivalve clam) *
 Tellina_nuculoides: Count by species (bivalve clam) * Allorchestes: Count
 by genus (amphipod) * Ampithoie: Count by genus (Ampithoe, amphipod) *
 Calanoida: Count by order (copepod, crustacean) * Caprellidae: Count by
 family (amphipod) * Cumacean: Count by order (Cumacea, crustacean) *
 Crangon_fransiscorum: Count of *Crangon franciscorum* (shrimp, Decapod
 crustacean) * Clausiidae: Count by family (copepod, crustacean) *
 Eogammarus: Count by genus (amphipod) * Eohaustorius_estuarius: Count by
 species (amphipod) * Grandidierella: Count by genus (amphipod) *
 Hemigrapsus_sp: Count by genus (crab, Decapod crustacean) *
 Monocorophium_spp: Count by genus (amphipod) * Neotrypaea_californiensis:
 Count by species (Thallassinid shrimp) * Ostracoda: Count by class
 (crustacean) * Paracorophium: Count by genus (amphipod) * Pinnixa_spp.:
 Count by genus (crab, Decapod crustacean) * Tanaidacea: Count by order
 (crustacean) * Urothoe_spp.: Count by genus (amphipod) * Capitellidae:
 Count by family (polychaete, annelid) * Cirratulidae: Count by family
 (polychaete, annelid) * Hesionidae: Count by family (polychaete, annelid)
 * Hirudinea: Count by class (leeches, annelid) * Glyceridae_Goniadidae:
 Count by families (polychaete, annelid) * Magelonidae: Count by family
 (polychaete, annelid) * Maldanidae: Count by family (polychaete, annelid)
 * Nemertean: Count by phylum * Nephtyidae: Count by family (polychaete,
 annelid) * Nereididae: Count by family (polychaete, annelid) *
 Oligochaete: Count by order (annelid) * Opheliidae: Count by family
 (polychaete, annelid) * Orbiniidae: Count by family (polychaete, annelid)
 * Oweniidae: Count by family (polychaete, annelid) * Paraonidae: Count by
 family (polychaete, annelid) * Phoronidae: Count by phylum *
 Phyllodocidae: Count by family (polychaete, annelid) * Pilargidae: Count
 by family (polychaete, annelid) * Polynoidae: Count by family (polychaete,
 annelid) * Platyhelminthes: Count by phylum (flatworms) * Sabellidae:
 Count by family (polychaete, annelid) * Spionidae: Count by family
 (polychaete, annelid) * Syllidae: Count by family (polychaete, annelid) *
 Terebellidae: Count by family (polychaete, annelid) * Barnacle: Count by
 subclass (Cirripedia, crustaceans) * Cnidaria: Count by phylum * Hydrozoa:
 Count by class (within Cnidaria) * Nudibranch: Count by order
 (Nudibranchia, gastropod) * Sea_urchin: Count by class (Echinoidea,
 echinoderm) * Snail: Count by class (Gastropoda) * Nematoda: Count by
 phylum * Unknown_amphipoda: Count by class * Unknown_bivalve: Count by
 class * Unknown_polychaete: Count by class * Decapod_zoea: Count by order
 * Harpacticoida: Count by order * Foraminifera: Count by phylum *
 Chironomidae: Count by family (fly larvae) * Cicadellidae: Count by family
 (leafhoppers) * Chrysomelidae: Count by family (beetles) * Hemiptera:
 Count by family (bugs) * Hymenoptera: Count by family (wasps) *
 Isotomidae: Count by family (springtails) * Talitridae: Count by family
 (amphipod) * Notes: Collection notes * ID'd_by: Person doing
 identification #### File: Penetrometer.csv **Description:** Penetration
 depth into the sediment of a standard rod to determine sediment
 penetrability on intertidal flats in Grays Harbor, Washington, in
 Experiment 1 (Damon Point, 0.2-0.6 m above mean lower low water) and
 Experiment 2 (Westport, 0.4-0.5 m above mean lower low water). The
 penetrometer was a device consisting of a 159 cm tall metal rod with a
 movable 2.25 kg weight encircling an additional section of rod between two
 plates. For each subsample, the rod was placed vertically on the mudflat,
 the weight was dropped 3 times, and the distance measured from the plate
 to the sediment. The final measurement was subtracted from 159 to give the
 depth of penetration into the sediment in centimeters, a measure of bed
 softness. ##### Variables * Site : Experimental site * DateSampled_true:
 Actual date when penetrometer was used * DateSampled: Date used to
 designate one timepoint of sampling, which could span several consecutive
 days * DaysPostComp: Penetrometer use date relative to the experimental
 setup when vehicle compaction occurred * Bed: Code denoting bed * Sample:
 Code denoting the Bed and position on the bed where the penetrometer was
 used * Treatment: Treatment applied to 2-ha bed, either Compacted by
 vehicle compaction, or Reference * Elevation: For Westport Single only,
 refers to whether samples were well into the bioturbated area
 ("High") or at the lower edge near firmer sediment
 ("Low") * GPSN: WGS84 geoposition of penetrometer sample in
 north latitude * GPSW: WGS84 geoposition of penetrometer sample, negative
 sign for west longitude * Penetr1: Rod height above the sediment after
 first drop of weight (cm) * Penetr2: Rod height above the sediment after
 second drop of weight (cm) * Penetr3: Rod height above the sediment after
 third drop of weight (cm) * Notes: Observations that could affect
 penetrometer data #### File: Oyster_outplants2.GPS.csv
 **Description:** Counts of Pacific oysters (*Magallana gigas)* on cultch
 ("mother shell"). Cultch were tethered to posts inserted on
 tidal flats in Grays Harbor, Washington, at multiple stations on each bed
 in Experiment 1 (Damon Point, 0.2-0.6 m above mean lower low water) and
 Experiment 2 (Westport, 0.4-0.5 m above mean lower low water).
 Pre-outplant samples are from cultch having the same source as outplanted
 cultch, therefore representative of initial counts. When cultch were
 collected, each was counted for remaining live oysters (and in some cases
 for obvious dead oysters with open or missing top valves). ##### Variables
 * DateSampled: Date when cultch was counted for oysters * Site:
 Experimental site * DaysPostComp: Number of days after vehicle compaction
 treatment * Bed: Code denoting bed * SampleCode: Code denoting the Bed and
 position on the bed where the cultch was tethered * NGPS: WGS84
 geoposition of tethered cultch in north latitude * WGPS: WGS84 geoposition
 of tethered cultch, negative sign for west longitude * Live: Count of live
 oysters * Dead: Count of dead oysters * Total_count: Count of live and
 dead oysters * Live_shell_ht.mm: Dimension of live oyster shell in mm from
 hinge to lip * Notes: Comments regarding cultch deployment or measurements
 * Group: Separates three outplant experiments using different sets of
 cultch #### File: PacificCultch.csv **Description:** Counts of Pacific
 oysters (*Magallana gigas*) that were planted on 22 July 2023 by a
 commercial operator in a 1-acre (0.4 ha) area on a bed compacted by
 vehicle compaction on 12 and 13 August 2022 and on 13 May 2023. Quadrats
 were distributed along two transects separated by at least 20 m and
 running from NE to SW corner of the planted area. The planted area was
 within 46.9971 and 46.9977 degrees N, and within 124.1322 and 124.1329
 degrees W (WGS84 datum). ##### Variables * photo_id: Number assigned to
 this photograph by the camera used * site: This commercial planting
 occurred only at Damon Point * date: Date when photograph was taken *
 area_m2: Area in the photograph used for counting (quadrat area) *
 oyster_count: Count of visible apparently live oysters per quadrat, only
 counted once these were large enough on cultch to be seen in photographs *
 cluster_count: Count of visible cultch or clusters per quadrat *
 algae_presence: Proportional cover of green macroalgae * holes: Count of
 surface holes in the sediment per quadrat #### File:
 WP_multiplepass_shrimpcores.csv **Description:** Samples collected from
 Grays Harbor, Washington, in Experiment 3 (Westport Multiple, 0.6 m above
 mean lower low water). Treatments were applied to plots that were 0.16 ha
 in size. Vehicle compaction occurred in a gradient of compaction intensity
 on 24 Jul 2024 by driving over the area with a MarshMaster MM2LX tracked
 vehicle. Treatments included 0 (reference), 1, 3, and 5 passes of vehicle
 compaction. The samples in this dataset are burrowing shrimp densities
 (*Neotrypaea californiensis*) in 70-cm deep cores and sediment
 penetrability measured with the same tool as described for the
 Penetrometer.csv dataset. ##### Variables * Site: This dataset only
 includes one site, Westport Multiple * Sample: Code denoting Plot and the
 placement on the Plot where the sample was collected * Plot: Code denoting
 plot, which is a 40x40 m area of consistent treatment * Treatment:
 reference (no compaction passes) or 1x, 3x, or 5x representing the number
 of passes with the tracked vehicle on 24 Jul 2024 * surface.or.core:
 "surface" if shrimp were counted in the top 2 cm of sediment in
 a 0.25 m^2^ quadrat within an hour of vehicle compaction; "core"
 if shrimp were counted in a 5-core sample, which entailed quintuplet cores
 within 1 m^2^, collected with a clam gun (stainless steel, 12.8 cm
 diameter and 36 cm length with 50-cm handle), inserted twice to reach the
 target depth of 70 cm * quadrat.size.m: area of quadrat in m^2^ *
 DateSampled_true: Actual date of sampling * DateSampled: Date used to
 designate one timepoint of sampling, which could span several consecutive
 days * DaysPostComp: Number of days after vehicle compaction treatment *
 GPSN: WGS84 geoposition of sample in north latitude * GPSW: WGS84
 geoposition of sample, negative sign for west longitude * Holes: Count of
 surface holes per quadrat, including holes at least 5 mm in diameter *
 Vegetation: % Cover or notes on seagrass or macroalgae * L.Neo: Count of
 large *Neotrypaea californiensis*, carapace length &gt;17.42 mm, per
 sample * M.Neo: Count of medium *Neotrypaea californiensis*, carapace
 length 12.5-17.42 mm, per sample * S.Neo: Count of small *Neotrypaea
 californiensis*, carapace length 8.29-12.49 mm, per sample * XS.Neo: Count
 of extra-small *Neotrypaea californiensis*, carapace length 4-8.28 mm, per
 sample * R.Neo: Count of recruit *Neotrypaea californiensis*, carapace
 length &lt;4 mm, per sample * Penetrometer1: Rod height above the sediment
 after first drop of weight (cm) * Penetrometer2: Rod height above the
 sediment after second drop of weight (cm) * Penetrometer3: Rod height
 above the sediment after third drop of weight (cm) * Notes: Observations
 that could affect shrimp counts or penetrometer data #### File:
 sediment.csv **Description:** Sediment collected from Grays Harbor,
 Washington, in Experiment 1 (Damon Point, 0.2-0.6 m above mean lower low
 water), Experiment 2 (Westport, 0.4-0.5 m above mean lower low water), and
 Experiment 3 (Westport, 0.6 m above mean lower low water, identified as
 Multiple). Treatments were applied at the scale of aquaculture beds, which
 were 2 ha in size in Experiment 1 and 2, but 0.16 ha in size in Experiment
 3. Vehicle compaction with a MarshMaster MM2LX occurred on Beds 9 and 11
 at Damon Point, on Beds 2 and 4 at Westport, and in a gradient of
 compaction intensity (1x, 3x, 5x) at Westport Multiple. At east sample
 timepoint at each station, one sediment sample (30 mL) was collected with
 a syringe corer to a depth of 5.4 cm. Sediment was fully dried and then
 processed for organic content and the mineral sediment thereafter
 dry-sieved for particle size. Sediment was dried at 45°C for at least 72
 hours, homogenized and weighed, held in a furnace at 500°C for 3 hours,
 and reweighed, to determine dry mass and organic content. Ashed sediment
 was dry-sieved through a sieve series (Wentworth scale, 63 micrometers to
 2000 micrometers) on a sieve shaker (Ro-Tap, W.S. Tyler, Mentor, OH, USA).
 Sediment smaller than 63 micrometers was classed as “mud” (silt and clay).
 ##### Variables * Site: Experimental site * SampleCode: Code denoting Bed
 and the placement on the Bed where the sample was collected * Bed: Code
 denoting Bed * Treatment: Treatment applied to 2-ha bed, either Compacted
 by vehicle compaction, or Reference; for Multiple experiment applied to
 0.16-ha plots, reference (no compaction passes) or 1x, 3x, or 5x
 representing the number of passes with the tracked vehicle * DateSampled:
 Actual date of sampling * DaysPostComp: Number of days after vehicle
 compaction treatment * GPSN: WGS84 geoposition of sampling in north
 latitude * GPSW: WGS84 geoposition of sample, negative sign for west
 longitude * crucible.wt: weight (g) of empty crucible * cruc.sed.wt:
 weight (g) of crucible with dry sediment * cruc.ashed.wt: weight (g) of
 crucible with sediment that had been ashed at 500 C for 3 hr *
 crucible.wt.2: weight (g) of empty crucible to be used with a second
 portion of sediment * cruc.sed.wt.2: weight (g) of second crucible with a
 second portion of dry sediment * cruc.ashed.wt.2: weight (g) of second
 crucible with a second portion of dry sediment after ashing at 500 C for 3
 hr * boat: weight (g) of weigh-boat, 0 if tared * boat.2mm: weight (g) of
 ashed sediment caught on 2-mm sieve including any boat weight * boat.1mm:
 weight (g) of ashed sediment caught on 1-mm sieve including any boat
 weight * boat.05mm: weight (g) of ashed sediment caught on 0.5-mm sieve
 including any boat weight * boat.025mm: weight (g) of ashed sediment
 caught on 0.25-mm sieve including any boat weight * boat.0125mm: weight
 (g) of ashed sediment caught on 0.125-mm sieve including any boat weight *
 boat.0063mm: weight (g) of ashed sediment caught on 0.063-mm sieve
 including any boat weight * boat.fines: weight (g) of ashed sediment
 passing through 0.063-mm sieve including any boat weight * Notes:
 Observations that could influence interpretation of sediment mass *
 Exclude: "Y" if concerns about processing this sample mean it
 should be censored from analysis #### File: Shrimp.csv
 **Description:** Burrowing shrimp (*Neotrypaea californiensis*) samples
 from Grays Harbor, Washington, in Experiment 1 (Damon Point, 0.2-0.6 m
 above mean lower low water) and Experiment 2 (Westport, 0.4-0.5 m above
 mean lower low water). Treatments were applied at the scale of aquaculture
 beds (2 ha). Vehicle compaction with a MarshMaster MM2LX occurred on Beds
 9 and 11 at Damon Point, and on Beds 2 and 4 at Westport. Shrimp were
 counted in 5-core samples, which entailed quintuplet cores within 1 m^2^,
 collected with a clam gun (stainless steel, 12.8 cm diameter and 36 cm
 length with 50-cm handle), inserted twice to reach the target depth of 70
 cm. ##### Variables * Site: Experimental site * DateSampled: Date used to
 designate one timepoint of sampling, which could span several consecutive
 days * DateSampled_true: Actual date of sampling * DaysPostComp: Number of
 days after vehicle compaction treatment * Bed: Code denoting Bed *
 Bed.description: Code denoting Bed and the placement on the Bed where the
 sample was collected * Treatment: Treatment applied to 2-ha bed, either
 Compacted by vehicle compaction, or Reference * GPSN: WGS84 geoposition of
 sample in north latitude * GPSW: WGS84 geoposition of sample, negative
 sign for west longitude * holes.per.025.sq.m: Count of surface holes per
 quadrat, including holes at least 5 mm in diameter * L.Neo: Count of large
 *Neotrypaea californiensis*, carapace length &gt;17.42 mm, per sample *
 M.Neo: Count of medium *Neotrypaea californiensis*, carapace length
 12.5-17.42 mm, per sample * S.Neo: Count of small *Neotrypaea
 californiensis*, carapace length 8.29-12.49 mm, per sample * XS.Neo: Count
 of extra-small *Neotrypaea californiensis*, carapace length 4-8.28 mm, per
 sample * R.Neo: Count of recruit *Neotrypaea californiensis*, carapace
 length &lt;4 mm, per sample * TotalShrimp: Sum of large, medium, small,
 and extra-small shrimp counts * Shrimp_m2: Total shrimp scaled to count
 per square meter * core_number: Number of cores per sample, usually 5
 (quintuplet) * Notes: Observations that could affect shrimp counts ##
 Code/software All data files are uploaded as comma-separated values.