DOI: 10.48579/pro/qfcpba
title: Replication data for: Fig 9.14, GPDs, clan size vs male ARS, 05 Dec 2025

description: FILE INFORMATION Keywords: GPDs, Clansize, male ARS File name = Fig 9.14, GPDs, clan size vs male ARS, 05 Dec 2025 OLD File name = GPDs, Male ARS vs clan size, 05 Sept 2021 I started this file from File name = GPDs, Male RS for cops,babies,yrls, 01 April 2021 Details below for GPDs, Male RS for cops,babies,yrls, 01 April 2021 and related files I used microsatellite data from only VCNP's RMC and VCC, because I had microsat data on paternities from those colonies. I did NOT include info from PEFO, from which I have no good information on paternites from microsats I will keep info from PEFO on Sexpartners for now If a GPD male had &gt;=2 half clans, I used the LARGER of the two half-clans for clan size for the GPD male. In 2013, for example, GPD male-8 had one half-clan of size 12 (8A) and two half clans of size 4 (8B and 8C)). For GPD-male-8 for this file, I used clan size = 12 If TWO GPD males shared the same multi-male clan, I used the same clan size for both males. In 2013, for example, I used the same clan size of 20 for both male-0 and male-41. Multi-male clans were rare for GPDs. For 0-41 in 2013, I do not use clan size--because smaller clans with large territory almost certainly existed For 44 in 2013, I use smaller clan (44a) for 44 (clansize = 4), because I new so little about clan-44B For 28 in 2013, I no not use any information. Smaller clans within large territory seem likely. For 49 in 2017, I use clan size = 4 (no females in his other half-clan of clan size = 5) Clan sizes checked TWO TIMES on 05 Sept 2021 Other data were checked two times for earlier files. I DO use GPD male-31 in 2018. We observed only 2 cops, but he sired babies from 5 litters! Date = 11 April 2021 OLDFile name = GPD, Male ARS for cops, PEFO and VCNP, 11 April 2021 I created this file by combining data from the following two EXCEL files: File name = GPDs, Male RS for cops,babies,yrls, 01 April 2021 [and] File name = GPDs, Male ARS and LRS for cops, 13 Jan 2021 Details from both files immediately below Date = 02 April 2021 OLD File name = GPDs, Male ARS for cops,babies,yrls, 01 April 2021 I created this file from File name = GPDs, Male ARS-LRS summaries, 31 March 2021. See notes below. OKtouse = 0 = we did not see many of the copulations--probably &lt;50% OKtouse = 1 = we documented most of the copulations--probably &gt; 50% I checked all the data on Age and Sexpartners on the hard-copy summary sheets ONE TIME ONLY--- I found only 1 error, so I did not check a second time This file has been checked TWO TIMES on 02 April 2021 In particular, I checked the new data for Sexpartners, OK to use, and Age on 02 April 2021 For GPDs from VCC of VCNP, I use a suffix of "V" for every GPD marker and eartag-- to reduce possible confusion with GPDS from RMC of VCNP All data in this file are from GPD males that copulated with &gt;=1 female All data from males that were not observed to copulate are EXCLUDED from this file For several males listed in this file that copulated with &gt;=1 female, we have NO EVIDENCE FROM MICROSATS that those males sired &gt;=1 emergent juvenile. Copulating males that evidently did not sire any offspring include the following: For 2013: 28 (123) For 2014: 108 (C47), 25 (439), 32 (593) For 2015: 20#2 (815), 107 (C39) For 2016: 37#2 (C99) For 2017: 24 (203) For 2018: 32 (79), 6 (395). 14 (55), 47 (435), 17 (O235) Date = 31 March 2021 This new file checked TWO TIMES for Babies and Yearlings on 31 March-02 April 2021 Babies = cumulative number of babies sired by each male per year Yearlings = cumulative number of babies sired by each male per year File name = GPDs, Male ARS-LRS summaries, 31 March 2021 I made this file directly from file = GPDs, Male ARS-LRS for cops, babies, yrls, 30 March 2021 Details immediately below for GPDs, Male ARS-LRS for cops, babies, yrls, 30 March 2021 Name of file = GPDs, Male ARS-LRS for cops, babies, yrls, 30 March 2021 No information on survivorship of juveniles for 2016 (last year at VCNP's RMC) or 2018 (last year at VCNP's VCC) Number in parentheses after number of babies or yearlings for a male = number of babies or yearlings ESTIMATED or INFERRED. I only made estimates or inferences that seemed reasonable. For example, if microsats showed that the single male that copulated with the mother sired 2 of 3 babies, I inferred paternity to that male for the third baby as well. Similarly, if microsats showed for a litter of 6 that one of two copulating males sired 2 babies and that the other of two copulating ales sired 2 babies, then I inferred paternity for 3 babies for both males. If a female copulated with 2 males, for example, and 1 of the copulating males sired three of four babies and we have NO INFORMATION from microsats for the fourth babies, then I inferred that the male that sired three of the babies (from microsats) also sired the fourth baby as well. I only made this inference if the one male sired AT LEAST HALF of babies. So, if a female copulated with two males and weaned five babies, for example, and if one of the copulating males sired two babies and we have NO INFORMATION from microsats for the other three babies, I have NOT inferred paternities for any of those 3 babies. For this file I have made no attempt to include information for body mass or age of the copulating males. Information on age and body mass vs male ARS and LRS is summarized in other chapters. This file has been checked TWO TIMES on 29-30 March 2021. I have excluded some cases for which paternity makes no sense. Dognumber does not equal integer--e.g., 15.5, 33.2, 58.5--means that GPD was added after I created an earlier, numbered file. File name = GPDs, Male ARS and LRS for cops, 13 Jan 2021 Date = 12 Jan 2021 This file is IDENTICAL to file = GPDs, MCs vs male body mass, ARS, 06 Jan 2021 I have created this identical file so that I can keep it with other ARS files See notes below from that file Note on 08 Feb 2021. These data are all from PEFO from 1989 through 1995 i.e., no data from RMC or VCC of VCNP Sired weanlings = Number of sired babies weaned by females with whom male copulated Note, however, that the male did NOT necessarily sire all juveniles listed as babies Some of those babies were sired by other males For paternities, see information from Keller Each male had unique eartags. If we had data from the same male in different years, then that male might have 1, 2, 3, or even 4 "Dognumbers" This file has now been checked TWO TIMES MCs = total number of MCs given by male for all females with whom he copulated PEFO = Petrified Forest National Park in Arizona RMC = Redondo Meadow Colony at Valles Caldera National Preserve in New Mexico VCC = Visitor Center Colony at Valles Caldera National Preserve in New Mexico Note: this EXCEL file does NOT contain information on GPD clantypes. Data for GPD clantypes are in file that begins with "Fig 18.8, GPDs,,," Date = 08 Jan 2021 File name = GPDs, MCs vs male body mass, male cops, 08 Jan 2021 I derived this file from file = GPDs, MCs vs female body mass, ARS, 06 Jan 2021, which I checked ONE TIME ONLY on 05 Jan 2021 File name = GPDs, MCs vs male body mass, male cops, 08 Jan 2021 has been checked TWO TIMES ON 08 Jan 2021 A GPD male will have the SAME EARTAGS in different years, but will have a different Dognum in different years For most analyses, I will consider data from the same GPD male in different years to be statistically independent Oktouse = 1 means that we think we observed most or all copulations by a GPD male in a particular year Oktouse = 0 means that we think we DID NOT OBSERVE most or all copulations by a GPD male in a particular year I only use AGE if I was certain--or almost certain--about a GPD male's exact age Babies = all babies weaned by all females with whom a GPD male copulated. SOME of the babies might have been sired by the copulating male, but OTHER offspring might have been sired by other males that copulated with the GPD female Yrls = all yrls reared by all females with whom a GPD male copulated. SOME of the yrls might have been sired by the copulating male, but OTHER yrls might have been sired by other males that copulated with the GPD female SEXPARTNERS = minimal number of GPD females with whom GPD male copulated, with or without MCs by the copulating male Note GPD male with Dognum = 63.5--so that I could insert additional data for GPD male-07 (B17, C68) in 1993. Note also that GPD male-34 and GPD male 34#2 are DIFFERENT GPD MALES, and they have different Dognums and different Eartags Microsats = microsatellites These GPD data used for Figure 9.14 in Hoogland 2026. This file ready for longterm storage on 06 Feb 2025. File name = Fig 9.14, GPDs, clan size vs male ARS, 05 Dec 2025