DOI: 10.5281/zenodo.15159942
title: Table 1 in Phylogenetic insights into the terrestrial snails Helicoidei (Gastropoda: Stylommatophora) with special emphasis on the Camaenidae

description: Table 1. Helicoidei specimens used in this study.
Vaught family/ subfamilyMolluscaBase family/ subfamilySpeciesCollection/locationCollectorGenBank accessionBradybaenidae: BradybaeninaeCamaenidae: BradybaeninaeAcusta despecta chinensis (Sowerby, 1839) aJapanS. ChibaAY841337, PP069321Bradybaenidae: BradybaeninaeCamaenidae: BradybaeninaeAinohelix editha (A. Adams, 1868) aShimamaki, Hokkaido, JapanS. ChibaAY841338, PP069323Bradybaenidae: BradybaeninaeCamaenidae: BradybaeninaeBradybaena similaris (Férussac, 1821) aSri LankaP. KarunaratneMN022676, PP069329Bradybaenidae: BradybaeninaeCamaenidae: BradybaeninaeCathaica pyrrhozona (Philippi, 1845)bDezhou City, ChinaZ. ZhangPP069372Camaenidae: CamaeninaeCamaenidae: BradybaeninaeChosenelix impotens (Pilsbry & Y. Hirase, 1908) comb. nov. bTianmenshan, ChinaG. ZhangPP069378Bradybaenidae: BradybaeninaeCamaenidae: BradybaeninaeEuhadra amaliae (Kobelt, 1875) aOsaka City, JapanP. CallomonAY014140, PP069345Bradybaenidae: BradybaeninaeCamaenidae: BradybaeninaeEzohelix gainesi (Pilsbry, 1900) aSapporro, Hokkaido, JapanS. ChibaAY841339, PP069347Not ApplicableCamaenidae: BradybaeninaeJiaoliaous tchefouensis (Crosse & Debeaux, 1863) bKunyushan, ChinaG. ZhangPP069379Camaenidae: CamaeninaeCamaenidae: BradybaeninaeLandouria radleyi (Jousseaume, 1894) bHorton plain, Sri LankaF. NaggsPP069359Bradybaenidae: BradybaeninaeCamaenidae: BradybaeninaeManchurohelix twenhuaensis (Ping & Yen, 1933) bJinlin, Jilin Province, ChinaG. ZhangPP069337Camaenidae: Camaeninae Bradybaenidae: BradybaeninaeCamaenidae: Bradybaeninae Camaenidae: BradybaeninaeMandarina ponderosa (Pilsbry, 1901) a Metodontia yantaiensis (Crosse & Debeaux, 1863) bHahajima, Bonin Islands, Japan Xuecheng, Zaozhaung, Shandong Province, ChinaS. Chiba & A. Davison G. ZhangAY841320, PP069361 PP069389Bradybaenidae: BradybaeninaeCamaenidae: BradybaeninaeNesiohelix bipyramidalis Kuroda & Emura, 1943aRyukyu, JapanS. ChibaAY841341, PP069369Camaenidae: CamaeninaeCamaenidae: BradybaeninaeNipponochloritis bracteatus (Pilsbry, 1902) aSendai, JapanS. ChibaAY841319, PP069370Bradybaenidae: BradybaeninaeCamaenidae: BradybaeninaeParaegista takahidei Kuroda & Azuma, 1951aHokkaido, JapanS. ChibaAY841340, PP069371Bradybaenidae: BradybaeninaeCamaenidae: BradybaeninaePlectotropis vulgivaga (Schumacher & Boettger, 1890) [= Aegista vulgivaga]aOsaka City, JapanP. CallomonAY014139, PP069322Bradybaenidae: BradybaeninaeCamaenidae: BradybaeninaePliocathaica buvigneri (Deshayes, 1873) bHuanxian, Qingyang, Anhui Province, ChinaUnknownPP069363cBradybaenidae: BradybaeninaeCamaenidae: BradybaeninaePseudiberus zenois (Gredler, 1882) bJinan, ChinaG. ZhangPP069326Bradybaenidae: BradybaeninaeCamaenidae: BradybaeninaePseudobuliminus dongyiicus bLushan, ChinaG. ZhangPP069358Bradybaenidae: BradybaeninaeCamaenidae: BradybaeninaeTrishoplita hachijoensis (Pilsbry, 1902) aNiijima Island, Izu Islands, JapanS. ChibaAY841345, PP069387Bradybaenidae: BradybaeninaeCamaenidae: BradybaeninaeVirginhelix virgo (Pilsbry, 1927) bGuichuan Road, Beijing City, ChinaG. ZhangPP069360Camaenidae: CamaeninaeCamaenidae: BradybaeninaeConiglobus mercatorius (Pfeiffer, 1854) aKikai Island, Ryukyu, JapanS. ChibaAY841324, PP069340Camaenidae: CamaeninaeCamaenidae: BradybaeninaeSatsuma japonica (Pfeiffer, 1847) aOsaka City, JapanP. CallomonMN022674, PP069381Camaenidae: CamaeninaeCamaenidae: CamaeninaeAmphidromus sp. aUnknownD. ReidAY841317 & AY841318, PP069332cCamaenidae: CamaeninaeCamaenidae: CamaeninaeBeddomea albizonata (Reeve, 1849) bHomadola, Sri LankaD. RaheemPP069328Camaenidae: CamaeninaeCamaenidae: CamaeninaeTrichelix diminuta (Pilsbry & Y. Hirase, 1905) [= Moellendorfia diminuta]aRyukyu, JapanS. ChibaAY841329 & AY841330, PP069367cCamaenidae: CamaeninaeCamaenidae: HadrinaeSphaerospira fraseri (Gray in Griffith & Pigeon, 1833)aBrisbane, Queensland, AustraliaJ. StanisicAY841325, PP069382Bradybaenidae: HelicostylinaeCamaenidae: HelicostylinaeChloraea intorta (Sowerby, 1840) aBohol Island, PhilippinesS. ChibaAY841344, PP069336Bradybaenidae: HelicostylinaeCamaenidae: HelicostylinaeHelicostyla lignaria (Pfeiffer, 1842) aBohol Island, PhilippinesS. ChibaAY841342 & AY841343, PP069351cHelminthoglyptidae: CepolinaeCepolidae: CepolinaeHemitrochus streatori (Pilsbry, 1889) [= Cepolis streatori]aGrand CaymanS. ChibaAY841346, PP069334Helicidae: HelicigoninaeHelicidae: AriantinaeArianta arbustorum (L., 1758)aDeepdale, Derbyshire, UKC. WadeAY014136, PP069327Helicidae: HelicigoninaeHelicidae: AriantinaeHelicigona lapicida (L., 1758)aDeepdale, Derbyshire, UKC. WadeAY014137, PP069352Helicidae: HelicigoninaeHelicidae: AriantinaeIsognomostoma isognomostomos (Schröter, 1784) bPoľana Mountain, SlovakiaJ. GregoPP069357Helicidae: HelicinaeHelicidae: HelicinaeAllognathus minoricensis (Mittre, 1842) bLluc, MajorcaC. Wade & B. WadePP069325Helicidae: HelicinaeHelicidae: HelicinaeCantareus apertus (Born, 1778) aSicilyA. DavisonAY014129, PP069331Helicidae: HelicinaeHelicidae: HelicinaeCaucasotachea vindobonensis (C. Pfeiffer, 1828) [= Cepaea vindobonensis]bKatlanovo, North MacedoniaE. Stojoska & J. GregoPP069342Helicidae: HelicinaeHelicidae: HelicinaeCepaea hortensis (Müller, 1774) aMarlborough Downs, Wiltshire, UKA. DavisonAY014131, PP069335Helicidae: HelicinaeHelicidae: HelicinaeCepaea nemoralis (L., 1758)aMarlborough Downs, Wiltshire, UKA. DavisonAY014130, PP069338Helicidae: HelicinaeHelicidae: HelicinaeCornu aspersum (Müller, 1774) [= Helix aspersa]aKettering, Northamptonshire, UKC. WadeAY014128, PP069353Helicidae: HelicinaeHelicidae: HelicinaeEobania vermiculata (Müller, 1774) bMajorca, SpainUnkownPP069343Helicidae: HelicinaeHelicidae: HelicinaeEremina desertorum (Forskål, 1775) aUnknownUnknownAY841335, PP069344Helicidae: HelicinaeHelicidae: HelicinaeHelix lucorum L., 1758aUnknownUnknownAY841334, PP069319Helicidae: HelicinaeHelicidae: HelicinaeHelix lutescens Rossmässler, 1837bPlešivec, SlovakiaJ. GregoPP069354Helicidae: HelicinaeHelicidae: HelicinaeHelix pomatia L., 1758aPulpit Down, Buckinghamshire, UKP. MordanAY841333, PP069355Helicidae: HelicinaeHelicidae: HelicinaeMacularia sylvatica (Draparnaud, 1801) [= Cepaea sylvatica]bSwitzerlandUnknownPP069341Helicidae: HelicinaeHelicidae: HelicinaeOtala lactea (Müller, 1774) aUnknownUnknownAY841336, PP069320Helicidae: HelicinaeHelicidae: HelicinaeTheba pisana (Müller, 1774) aSicilyA. DavisonAY014134 & AY014135, PP069383Helicidae: HelicinaeHelicidae: MurellinaeMarmorana scabriuscula (Deshayes, 1830) aSicilyA. DavisonAY014132 & AY014133, PP069362cHelicidae: HelicodontinaeHelicodontidae: HelicodontinaeHelicodonta obvoluta (Müller, 1774) bWest Wood, Winchester, Hampshire, UKJ. S. Gallichan, A. H. Wood & J. ChatfieldPP069349Hygromiidae: Helicellidae?Geomitridae: GeomitrinaeCochlicella acuta (Müller, 1774) aPorthcurnick, Cornwall, UKE. BailesAY014126, PP069339Hygromiidae: TrichiinaeGeomitridae: GeomitrinaePonentina subvirescens (Bellamy, 1839) bMaritime turf on serpentine headland, Kynance, Cornwall, UKG. A. HolyoakPP069376Hygromiidae: Helicellidae?Geomitridae: HelicellinaeCernuella virgata (Da Costa, 1778)aPorthcurnick, Cornwall, UKE. BailesAY014127, PP069333Hygromiidae: HygromiinaeHygromiidae: HygromiinaeMonachoides vicinus (Rossmässler, 1842) bSlovakiaJ. SteffekPP069366Hygromiidae: HygromiinaeHygromiidae: HygromiinaePerforatella bidentata (Gmelin, 1791) bBank of Lupcica river, Starohorske Vrchy Mountain, SlovakiaUnknownPP069373Hygromiidae: HygromiinaeHygromiidae: TrochulininaeEuomphalia strigella (Draparnaud, 1801) bSlovakiaUnkownPP069346Hygromiidae: Helicellidae?Hygromiidae: TrochulininaeMonacha cantiana (Montagu, 1803) aPulpit Down, Buckinghamshire, UKP. MordanAY841331 & AY841332, PP069365cHygromiidae: TrichiinaeHygromiidae: TrochulininaeTrochulus striolatus (Pfeiffer, 1828) [= Trichia striolata]aDeepdale, Derbyshire, UKC. WadeMN022675, PP069386Camaenidae: CamaeninaePleurodontidae: LucerninaeThelidomus aspera (Férussac, 1821) aWindsor, JamaicaS. ChibaAY841321, PP069384Polygyridae: PolygyrinaePolygyridae: PolygyrinaePolygyra troostiana I. Lea, 1838bCedars of Lebanon State Park, Wilson County, TN, USAJ. Slapcinsky & B. ColesPP069377Polygyridae: AshmunelinaePolygyridae: PolygyrinaeTriodopsis alleni (Wetherby, 1883) aWilliams Creek, IA, USAR. CameronAY841316, PP069385Polygyridae: AshmunelinaePolygyridae: TriodopsinaeAllogona townsendiana (I. Lea, 1838) bDecker Creek at Matlock, Brady Road, Mason County, WA, USAD. TaylorPP069324Polygyridae: PolygyrinaePolygyridae: TriodopsinaeInflectarius rugeli (Shuttleworth, 1852) bPolk County, TN, USAJ. Slapcinsky & B. ColesPP069356Polygyridae: PolygyrinaePolygyridae: TriodopsinaeMesodon thyroides (Say, 1816) aYork Co. PA, USAF. ThompsonAY841315, PP069364Polygyridae: AshmunelinaePolygyridae: TriodopsinaeVespericola columbiana (Lea, 1838) aEugene, OR, USAD. TaylorAY014120, PP069388Helminthoglyptidae: HelminthoglyptinaeXanthonychidae (Helminthoglyptinae): HelminthoglyptinaeHelminthoglypta diabloensis (J. G. Cooper, 1869) bSonoma Mountain, CA, USAT. LawsonPP069350Helminthoglyptidae: HelminthoglyptinaeXanthonychidae: MonadeniinaeMonadenia fidelis (Gray, 1834) aOregon, USAD. TaylorMN022677, PP069368Camaenidae: CamaeninaeSagdidae: PolydontinaeHispaniolana crispata (Férussac, 1821) [= Polydontes undulata]aDominican RepublicG. SealAY014121, PP069375Sagdidae: SagdinaeSagdoidea: SagdinaeSagda sp. aWindsor, JamaicaS. ChibaAY841347, PP069380Camaenidae: CamaeninaeSagdoidea: ZachrysiidaeZachrysia auricoma (Férussac, 1821) aNr. Dorado, Puerto RicoA. DavisonAY841326, PP069390Camaenidae: CamaeninaeSolaropsidae,: CaracolinaeCaracolus marginellus (Gmelin, 1791) bUnknownG. M. BarkerPP069330cCamaenidae: CamaeninaeSolaropsidae: CaracolinaeCaracolus carocollus (L., 1758)bYunque, Puerto RicoA. DavisonPP069374Outgroup: Haplotrematoidea: HaplotrematidaeHaplotrema vancouverense (Lea, 1839)bEugene, OR, USAD. TaylorAY014090, PP069348
aSpecimens published by Wade et al. (2007) for an ~1460 bp fragment of the 5.8S–ITS2–LSU rRNA (using primers LSU 1–5) and resequenced here for an ~4000 bp fragment of the 5.8S–ITS2–LSU rRNA (using primers LSU 1–12).
bEntirely new specimens sequenced here for the ~4000 bp fragment of the 5.8S–ITS2–LSU rRNA (using the LSU 1–12 primers).
cGenBank accession numbers for which the ITS2 fragment is incomplete.
description: Published as part of Zhang, Guoyi, Naggs, Fred, Andrus, Peter S. & Wade, Christopher M., 2025, Phylogenetic insights into the terrestrial snails Helicoidei (Gastropoda: Stylommatophora) with special emphasis on the Camaenidae, pp. 3-6 in Zoological Journal of the Linnean Society 203 on page 3, DOI: 10.1093/zoolinnean/zlae027, http://zenodo.org/record/14803321