DOI: 10.5281/zenodo.18342286
title: APPENDIX I in Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species

description: APPENDIX I. New generic placements proposed for the species included in Goniocotes and Goniodes by Price et al. (2003), following this revision. The genera of species listed here as “ incerta sedis ” are in quotation marks to indicate that these generic associations are not supported by us. The generic placement of lice parasitizing hosts in the Megapodiidae follows Mey (1997), who synonymised Aurinirmus Tendeiro, 1983 under Homocerus Kéler, 1940, transferring the species to the latter genus, except for Aurinirmus keleri Tendeiro, 1983, which is here placed in Weelahia Mey, 1997.
GenusSubgenusSpeciesArchigoniodes Conci, 1946Archigoniodes Conci, 1946Archigoniodes (Archigoniodes) wilsoni (Clay, 1938)Clayarchigoniodes Conci, 1952Archigoniodes (Clayarchigoniodes) fimbriatus (Neumann, 1913)Archigoniodes (Clayarchigoniodes) hopkinsi (Clay, 1940)Archigoniodes (Clayarchigoniodes) inaequalis (Tendeiro, 1988)Archigoniodes (Clayarchigoniodes) klockenhoffi (Tendeiro, 1988)Archigoniodes (Clayarchigoniodes) meyi (Tendeiro, 1988)Archigoniodes (Clayarchigoniodes) numidae (Mjöberg, 1910)Archigoniodes (Clayarchigoniodes) plumiferae (Tendeiro, 1988)Archigoniodes (Clayarchigoniodes) reichenowii (Tendeiro, 1988)Archigoniodes (Clayarchigoniodes) schoutedenii (Tendeiro, 1988)Astrocotes Kéler, 1940Astrocotes astrocephalus (Burmeister, 1838)Astrocotes keleri (Clay, 1940)Astrocotes lootensi (Tendeiro, 1960)Astrocotes retractus (Le Souëf, 1902)Astrocotes souefi (Clay, 1940)Astrodes Kéler, 1940Astrodes coronatus (Giebel, 1874)Claygoniodes Conci, 1946Claygoniodes temporalis (Kéler, 1940)Complectodes new genusComplectodes emersoni (Tendeiro, 1965)Complectodes graecus (Liu, 1994)Dictyocotes Kéler, 1940Dictyocotes crossoptiloni (Liu, 1990)Dictyocotes furcatus Gustafsson et al., 2020Dictyocotes haplogonus (Nitzsch [in Giebel], 1866)Dictyocotes rolandi (Gustafsson et al., 2021)Dictyocotes tetraophasis (Liu, 1990)Euligoniodes Mey, 1997Euligoniodes ocrea (Piaget, 1880)Goniocotes Burmeister, 1838Goniocotes afer Tendeiro, 1989Goniocotes afropavo (Clay, 1938)Goniocotes albidus Giebel, 1874Goniocotes cacumentalis Gustafsson et al., 2024bGoniocotes caoi Gustafsson et al., 2024bGoniocotes castaneicollis Tendeiro, 1965Goniocotes chrysocephalus Giebel, 1874Goniocotes clayae Tendeiro, 1954Goniocotes congolensis Tendeiro, 1989Goniocotes coxatus Piaget, 1885Goniocotes crassicauda Tendeiro, 1989Goniocotes creber Kellogg, 1896bGoniocotes diasi Tendeiro, 1954Goniocotes diplogonus Nitzsch [in Giebel], 1866Goniocotes eurygaster Piaget, 1885Goniocotes gallinae (De Geer, 1778)Goniocotes gregarius (Nitzsch [in Giebel], 1866)Goniocotes ictiorhynchi Tendeiro, 1989Goniocotes ignitus Eichler, 1947Goniocotes jirufti Ansari, 1947Goniocotes keleri Tendeiro, 1989Goniocotes keniensis Tendeiro, 1989Goniocotes kivuensis Tendeiro, 1960Goniocotes maculatus Taschenberg, 1882Goniocotes megalocephalus Uchida, 1916Goniocotes microcephalus Uchida, 1917Goniocotes microthorax (Stephens, 1829)Goniocotes nahanii Tendeiro, 1989Goniocotes obscurus Giebel, 1874Goniocotes pallidiflavus Piaget, 1890Goniocotes pallidomaculatus Piaget, 1880Goniocotes plumiferae Tendeiro, 1989Goniocotes pternistis Tendeiro, 1965Goniocotes pucherani Tendeiro, 1989Goniocotes punctatus Neumann, 1891Goniocotes pusillus (Nitzsch [in Giebel], 1866)Goniocotes rectangulatus Kéler, 1940Goniocotes rotundiceps Piaget, 1880Goniocotes sarissa Gustafsson et al., 2024bGoniocotes schraderi Tendeiro, 1989Goniocotes shelleyii Tendeiro, 1989Goniocotes vulturini Tendeiro, 1989Goniodes Nitzsch, 1818Goniodes claytonbushorum new speciesGoniodes pavonis (Linnaeus, 1758)Gonotyles Kéler, 1940Gonotyles Kéler, 1940Gonotyles (Gonotyles) cervinicornis (Giebel, 1874)Gonotyles (Gonotyles) diardi (Clay, 1940)Syrmatichares new subgenusGonotyles (Syrmatichares) humiae (Clay, 1940)Gonotyles (Syrmatichares) intermedius (Neumann, 1913)Gonotyles (Syrmatichares) selenautes new speciesGonotyles (Syrmatichares) sinensis (Emerson & Stojanovich, 1964)Incerta sedisGonotyles bambusicolus (Liu, 1989)Gonotyles longus (Rudow, 1869)Gonotyles lophurus (Liu, 1989)Gonotyles ocellatus (Rudow, 1869)Homocerus Kéler, 1940Homocerus aepypodius (Tendeiro, 1983)Homocerus arfakianus (Tendeiro, 1983)Homocerus crassipes (Piaget, 1888)Homocerus curtiprothorax Mey, 1982Homocerus macrocephalus (Taschenberg, 1882)Homocerus talegallae (Tendeiro, 1983)Kelerigoniodes Conci, 1946Kelerigoniodes processus (Kellogg & Paine, 1914)Leipoiella Mey, 1986Leipoiella australis (Emerson & Price, 1986)Lobicrotaphus Mey, 1997Lobicrotaphus discogaster (Taschenberg, 1882)Majiodes new genusMajiodes crossoptilon (Clay, 1940)Majiodes dolani (Eichler, 1937)Maleoicus Mey, 1997Maleoicus biordinatus (Clay, 1940)Maleoicus confusio (Clay, 1940)Maleoicus minimus (Tendeiro, 1980)Maleoicus minor (Piaget, 1880)Maleophilus Mey, 1997Maleophilus major (Piaget, 1880)Margaritenes Kéler, 1940Margaritenes eurygaster (Piaget, 1885)Megatheliella Mey, 1986Megatheliella leipoae (Emerson & Price, 1984)Oulocrepis Kéler, 1940Oulocrepis ammoperdix (Clay, 1940)Oulocrepis bituberculatus (Rudow, 1869)Oulocrepis bonasus (Emerson, 1948)Oulocrepis centrocerci (Simon, 1938)Oulocrepis chrysolophi (Clay, 1940)Oulocrepis colchici (Denny, 1842)Oulocrepis corpulentus (Kellogg & Mann, 1912)Oulocrepis costatus (Kéler, 1940)Oulocrepis cupido (Rudow, 1870)Oulocrepis dissimilis (Denny, 1842)Oulocrepis ithaginis (Clay, 1940)Oulocrepis leucurus (Emerson, 1952)Oulocrepis merriamanus (Packard, 1873)Oulocrepis montschadskyi (Eichler & Vasjukova, 1981)Oulocrepis nebraskensis (Carriker, 1946)Oulocrepis oreophilus (Clay 1940)Oulocrepis sectus (Kellogg & Paine, 1914)Oulocrepis stefani (Clay & Hopkins, 1955)Oulocrepis stylophorus new speciesOulocrepis tetraonis (Linnaeus, 1761)Oulocrepis tibetanus (Eichler, 1950)Pavoniocotes Gustafsson et al., 2023Pavoniocotes mayuri (Lakshminarayana & Emerson, 1971)Pavoniocotes parviceps (Piaget, 1880)Rhopaloncus new genusRhopaloncus lepidoceros new speciesRhopaloncus megaceros (Kellogg & Paine, 1914)Sikongia new genusSikongia tetraophasis (Chou & Liu, 1986)Solenodes Kéler, 1940Solenodes antennatus (Clay, 1940)Solenodes assimilis (Piaget, 1880)Solenodes capitatus Kéler, 1940Solenodes dispar (Burmeister, 1838)Solenodes moucheti (Tendeiro, 1960)Solenodes ningxiaensis new speciesSolenodes ortygis (Denny, 1842)Solenodes scleroptilus (Bedford, 1929)Solenodes securiger (Nitzsch [in Giebel], 1866)Stenocrotaphus Kéler, 1940Stenocrotaphus agelastes (Clay, 1940)Stenocrotaphus bifurcus Tendeiro, 1988Stenocrotaphus gigas (Taschenberg, 1879)Stenocrotaphus gutterae Tendeiro, 1988Stenocrotaphus isogenos (Nitzsch [in Giebel], 1866)Stenocrotaphus phasidus Tendeiro, 1988Stenocrotaphus zairensis Tendeiro, 1988Sulciferodes new genusSulciferodes meinertzhageni (Clay, 1940)Tragoniodes new genusTragoniodes gallowayi new speciesTragoniodes spinicornis (Nitzsch [in Giebel], 1866)Tragoniodes tragopan (Clay, 1940)Tragoniodes yinhe new speciesWeelahia Mey, 1997Weelahia fissus (Rudow, 1869)Weelahia keleri (Tendeiro, 1983)Genus incerta sedis“ Goniocotes” chapini (Clay, 1938)“ Goniodes” chloropus Emerson & Elbel, 1957“ Goniodes” curvicornis Nitzsch [in Giebel], 1866“ Goniodes” indicus (Kellogg & Paine, 1914)“ Goniodes” columbianus Carriker, 1946“ Goniodes” ovoidalis Kéler, 1940“ Goniodes” pictus Emerson, 1950“ Goniodes” rhynchortyx (Carriker, 1956)“ Goniodes” squamatus Emerson, 1950“ Goniodes” submamillatus Emerson, 1950
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APPENDIX I. (Continued)
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APPENDIX I. (Continued)
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APPENDIX I. (Continued)
description: Published as part of Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling & Zou, Fasheng, 2025, Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species, pp. 96-99 in Zootaxa 5731 (1) on page 96, DOI: 10.11646/zootaxa.5731.1.1, http://zenodo.org/record/18019316