DOI: 10.5281/zenodo.15389058
title: TABLE 2 in The World Catalog of Biting Midges (Diptera: Ceratopogonidae): Additional Updates and Errata

description: TABLE 2. Numbers of extant and fossil species of Ceratopogonidae according to subgenus and genus and summarized cumulatively. The classification follows Borkent (2024).
SubfamilyTribeGenus (* = fossil taxon)SubgenusSubgenusGenusNomina dubiaExtant speciesFossil SpeciesExtant speciesFossil SpeciesTotal SpeciesLebanoculicoidinae* Lebanoculicoides44Leptoconopinae* Archiaustroconops1414Austroconops2810* Baskintoconops11* Fossileptoconops11* Jordanoconops11Leptoconops1571917613Brachyconops2Holoconops79Leptoconops5817Megaconops1Palaeoconops2Proleptoconops9Styloconops8* Minyohelea99Atriculicoidinae* Atriculicoides1212* Burmahelea11ForcipomyiinaeDasyheleiniDasyhelea6391465340ForcipomyiiniAtrichopogon519352213Forcipomyia116731119926Atopopogon2Baliohelea1Bassoforcipomyia2Blantonia2Caloforcipomyia35Collessohelea2Dycea11Euprojoannisia1015Forcipohelea1Forcipomyia33013Gampsohelea2Herakleohelea1Ixodehelea1Japygahelea1Kattangomyia2Lasiohelea1833Lepidohelea864Metaforcipomyia36Microhelea105Nicothohelea1Oreinohelea1Oxyria1Panhelea2Pedilohelea19Phytohelea261Pterobosca23Rhinohelea2Rhynchoforcipomyia8Saliohelea14Schineromyia1Schizoforcipomyia13Sinicohelea1Synthyridomyia251Thyridomyia47Trichohelea583Trithicomyia1Typhonomyia1Warmkea14unplaced51CeratopogoninaeCulicoidiniCulicoides137354142745Amossovia12Anilomyia20Avaritia106Beltranmyia51Chairopogon1Cotocripus6Culicoides67Diphaomyia25Drymodesmyia26Fastus12Glaphiromyia3Groganomyia1Haemophoructus16Haematomyidium41Hoffmania85Jilinocoides6Macfiella2Marksomyia6Mataemyia19Meijerehelea12Monoculicoides25Nullicella1Oecacta152Pontoculicoides14Psychophaena2Remmia8Selfia8Sensiculicoides38Silvaticulicoides13Sinocoides9Synhelea32Trithecoides64Tokunagahelea3Wirthomyia8in species groups348miscellaneous13154Paradasyhelea1111WashingtonheleiniWashingtonhelea11* Nelohelea11CeratopogoniniAfrohelea11Afrostilobezzia22Agilihelea11Allohelea6363Alluaudomyia20612073Ankylohelea11Austrohelea99Baeodasymyia516Baeohelea11Boreohelea111Borkenthelea44Bothahelea33Bothamia11* Brachycretacea11Brachypogon204102141Brachypogon1125Isohelea855Sarissohelea5unplaced2Cacaohelea44Calcarhelea11Camptopterohelea516Capehelea11Ceratoculicoides91101Ceratohelea22Ceratopogon442064Congohelea11Downeshelea6060* Eohelea88Fanthamia1414* Fossihelea22* Gedanohelea66Heterohelea11Isthmohelea11Kolenohelea1919Leptohelea11Luciamyia11Macrurohelea2020Metacanthohelea11* Mantohelea33* Monogedania11Monohelea9999Nannohelea426Neohelea11Notiohelea22Notoceratopogon44* Palaeobrachypogon66Parastilobezzia11* Peronehelea33Pseudostilobezzia33Rhynchohelea11Schizohelea9110Schizonyxhelea1616Serromyia38846Sinhalohelea33Stilobezzia35210362Acanthohelea1766Debenhamia2Eukraiohelea14Stilobezzia s. str.1601unplaced03Stiloculicoides66* Wirthohelea11ParabezziiniBahiahelea11Diaphanobezzia44Fittkauhelea11Parabezzia4545Spinellihelea11Yungahelea11NeurobezziiniAtyphohelea112Ceratopalpomyia112Meunierohelea1910Neurobezzia33EchinoheleiniEchinohelea2727Echinohelea26Echinoideshelea1HeteromyiiniClinohelea39391Ceratobezzia1Clinohelea s. str.38Heteromyia13114Metahelea213Neurohelea11Pellucidomyia99Physohelea213Tetrabezzia77HebetuliniHebetula2222JohannsenomyiiniAnebomyia99Austrosphaeromias44Calyptopogon66Crispomyia22Dibezzia55Groganhelea11Jenkinshelea2020Johannsenomyia2828Lanatomyia33Lanehelea22Mackerrasomyia88Macropeza2222Mallochohelea44145Neobezzia88Neosphaeromias44Nilobezzia7575Niphanohelea11Probezzia2828Sphaeromiini s. str.Alloimyia11Chelohelea11Homohelea1919Indobezzia22Leehelea99Sphaerohelea11Sphaeromias3131Xenohelea1414PalpomyiiniAmerohelea1414Bezzia33023322Clastrieromyia44Pachyhelea22Palpomyia274112852Phaenobezzia341352Wannohelea11StenoxeniniParyphoconus4242Stenoxenus2121Unplaced genera* Adelohelea33* Alautunmyia11* Archiculicoides33* Gerontodacus44* Heleageron22* Protoculicoides22suprageneric40Total63463036650189Total Number of Extant Genera—107Total Number of Fossil Genera—23Total Number of Genera—130
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description: Published as part of Borkent, Art, Evenhuis, Neal L. & Dominiak, Patrycja, 2025, The World Catalog of Biting Midges (Diptera: Ceratopogonidae): Additional Updates and Errata, pp. 251-256 in Zootaxa 5631 (2) on page 251, DOI: 10.11646/zootaxa.5631.2.2, http://zenodo.org/record/15370558