DOI: 10.6078/d10411
title: Phylogeny and classification of the genus-group taxa of Loxandrina (Coleoptera, Carabidae, Abacetini)

description: Bayesian and parsimony phylogenetic analyses of combined and partitioned
 datasets of molecular (partial sequences of 28S, wg, COI, and CAD) and
 morphological (51 characters of adults) data for exemplar taxa of five
 outgroup and 76 ingroup abacetine carabids resulted in a monophyletic
 Loxandrina Erwin &amp; Sims, 1984 that is split into Australian
 and American clades. The genus Loxandrus LeConte, 1853 as previously
 delimited is not monophyletic relative to numerous genus-level taxa in
 Abacetini Chaudoir, 1873 and is restricted to a subgenus of North American
 species. A reclassification and nomenclatural changes for the subtribe
 that are consistent with the phylogeny are provided. Three genera are
 removed from Loxandrina: Aulacopodus Britton, 1940 moved to Pterostichini
 Bonelli, 1810; Cosmodiscus Sloane, 1907 and Tiferonia Darlington, 1962
 moved to Abacetina. Based on the phylogenetic relationships and
 nomenclatural priority only four genera are recognized in Loxandrina:
 Cerabilia Laporte, 1867, Zeodera Laporte, 1867, Pediomorphus Chaudoir,
 1878, and Oxycrepis Reiche, 1843. All other previously recognized genera
 are treated as subgenera. The classification change created eight
 secondary homonyms that are resolved by the proposal of the following:
 Oxycrepis gebi, replacement name for O. balli (Straneo, 1993); O. amatona,
 replacement name for O. matoana (Straneo, 1993); O. xiproma, replacement
 name for O. proxima (Straneo, 1993); O. rasutulis, replacement name for O.
 suturalis (Straneo, 1993); O. laevinota, replacement name for O.
 laevicollis (Bates, 1871); O. arvulap, replacement name for O. parvula
 (Straneo, 1951); O. noaffine, replacement name for O. affinis (Straneo,
 1991); O. alutona, replacement name for O. notula (Tschitschérine, 1901).
 An overview of the morphological characteristics and diagnostic features
 of Loxandrina taxa is provided. A key and habitus images are provided for
 identification of genera and subgenera. The possible historical
 biogeography of the group is discussed in light of their phylogenetic
 relationships and past geological events.
description: Morphological observations and standard PCR and Sanger for
 sequences.
description: There are a series of files that that include data matrices,
 analysis script files, and output files. These are arranged in zip
 archived folders, each being a self-contained set of input-script-output
 files for Bayesian and Parsimony analyses. Also the "final"
 renamed Nexus file as produced by Mesquite and the classification are
 included.    
 80_OTUs.zip   20.36 MB: <strong>All input and output files
 for parallel MrBayes analysis for combined data for 80
 OTUs.</strong> 
 80_OTUs_no_morph.zip   20.35 MB: <strong>All input and
 output files for parallel MrBayes analysis for DNA sequence data only for
 80 OTUs.</strong> 
 81_OTUs.zip   21.02 MB: <strong>All input and output files
 for parallel MrBayes analysis for combined data and 81
 OTUs.</strong> 
 GBLOCKS_parameters.JPG   42.95 kB: <strong>Image of
 parameter dialog box from GBlocks run in Mesquite.</strong>
 Lox-MrModelTest.zip   202.25
 kB: <strong>All MrModelTest input and output files for each
 locus.</strong> 
 Loxandrina_2019.6.nameschanged.nex   426.49
 kB: <strong>Final Mesquite generated nexus file for all data and
 taxa.</strong> 
 TNT-2019.12.17-Run.1.zip   2.50 MB: <strong>All TNT input
 and output files for parallel analysis run for morphology data for 81
 OTS.</strong>