Fulgurofusus marshalli Harasewych 2011, new species
Creators: name: Harasewych, M. G.
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Neogastropoda
Columbariidae
Fulgurofusus
Fulgurofusus marshalli

description: Fulgurofusus marshalli new species.   
(Figures 117, 119–132)  
 Diagnosis. Small species with strongly fusiform shell, tall, sharply stepped spire, shoulder with weak peripheral keel high on whorl, at or above midpoint between successive sutures. Protoconch conical, with 3 rounded whorls. Spiral sculpture of numerous cords, threads, adjacent cords along periphery producing incised furrow. Axial sculpture of strong, rounded ribs extending from suture to siphonal canal, most prominent along shoulder, forming nodules at intersection with peripheral keel on early whorls, more rounded on later whorls. Anterior carina with strong spiral cord. Aperture weakly angular along outer lip, rounded along parietal region and columella. Siphonal canal comparatively short (<½ shell length) stout, tapering distally.   
 Description. Shell (Figures 120–129) of small to moderate size (to 43.3 mm), stout, with tall, stepped, conical spire (spire angle 35–38˚), narrow, ovate aperture, comparatively short, axial siphonal canal. Outer shell layers of early whorls, including protoconch, generally eroded. Protoconch (Figures 131–132) very tall, uniformly conical, with rounded whorls increasing in diameter from 435 µm to 1.8 mm in ~3 whorls. First whorl deflected from coiling axis of shell by ~40˚. Transition to teleoconch marked by onset of a weak shoulder followed almost immediately by thin, spiral threads and later by a weak peripheral keel (Figure 130, pk). Teleoconch of up to 6 whorls. Early whorls angular, later whorls more rounded. Suture (Figure 130, s) adpressed onto spiral cord along anterior carina (Figure 130, ac) of previous whorl. Spiral sculpture of 8–12 strong, narrow, equally spaced cords between suture and peripheral keel. Peripheral keel weak, comprised of two closely opposed cords forming an incised furrow between them. Furrow reduced by 4 th postnuclear whorl, disappearing as two cords fuse by 5 th postnuclear whorl. Spiral sculpture of: 6–9 cords and threads between peripheral keel and anterior carina; 4–5 between anterior carina and siphonal canal; 9–11 progressively diminishing cords along proximal ⅓ of siphonal canal. Axial sculpture of 9–14 rounded ribs, slightly narrower than intervening spaces, extending from suture to siphonal canal, most pronounced near periphery and anterior carina. Aperture narrowly ovate, slightly more rounded in larger specimens, deflected from shell axis by 21–25˚. Outer lip thin, smooth, evenly glazed, not furrowed beneath peripheral keel or anterior carina (Figure 130). Inner lip with thin, porcellaneous glaze along parietal region, columella, axial portion of siphonal canal. Siphonal canal axial, tapering distally. Shell surface generally worn or eroded. Periostracum, operculum, radula and anatomical features unknown, as all specimens dead collected or crabbed.   
 Etymology. This new species honors Bruce Marshall for his many contributions to our knowledge and understanding of the molluscan fauna of New Zealand and of the deep sea.   
 Type locality. (Figure 117,) Bounty Plateau [48°00.0’S, 180°00.0’E], 280 m, 13 Mar 1979, RV TANGAROA (NIWA I671).   
 Type material. Holotype, NIWA 67594, 2 paratypes, NMNZ M.297047, from the type locality. Additional paratypes: 1 NMNZ M.297048, 1, NIWA I669; 1, USNM 1146129, Bounty Plateau [47°49.0’S, 179°45.7’E], 355 m, 13 Mar. 1979, RV TANGAROA; 1 NMNZ M.297045, Bounty Plateau [48°10.0’S, 179°45.0’E], 457 m, 14 Mar. 1979, RV TANGAROA; 1 NMNZ M.297050, 1, NIWA I682, Bounty Plateau [48°20.0’S, 179°49.5’E], 450 m, 15 Mar. 1979, RV TANGAROA.   
 Distribution (Figures). Fulgurofusus marshalli is known only from the Bounty Plateau. This species has been reported from depths ranging from 280 to 457 m, with a mean station depth [n = 4] of 385.5 m, and a confirmed bathymetric range of 280– 457 m.   
 Remarks. The shell of Fulgurofusus marshalli differs from that of comparably sized F. maxwelli in having: a more evenly conical protoconch; a less prominent peripheral keel that is obscured by far more prominent axial ribs, which are the dominant sculptural elements; a less pronounced anterior carina; a slightly narrower spire angle; and a more stepped spire, in which the peripheral carina is situated midway between the sutures of successive whorls. The strong axial sculpture of F. marshalli is reminiscent of Fulgurofusus (Peristarium) aurora, a species that inhabits roughly comparables depths in the Straights of Florida, and which differs in having stronger, rounder and more numerous axial ribs and stronger spiral cords, a proportionally smaller aperture, and a far stouter siphonal canal. Four species of Peristarium have been reported from off the Southeastern United States, yet none overlap in their bathymetric ranges. As in the western Atlantic, the strongest axial sculpture occurs in the shallowest dwelling species.   
 Fulgurofusus marshalli is known only from the Bounty Plateau at depths ranging from 280 to 457 m. The broader geographic range of F. maxwelli includes the Bounty Plateau, where its confirmed bathymetric range extends from 516 to 917 m.
description: Published as part of Harasewych, M. G., 2011, The Living Columbariinae (Gastropoda: Neogastropoda: Turbinellidae) of New Zealand, pp. 1-33 in Zootaxa 2744 (1) on page 28, DOI: 10.11646/zootaxa.2744.1.1, http://zenodo.org/record/5291694