Landouria tonywhitteni Nurinsiyah & Neiber & Hausdorf 2019, sp. nov.
Creators: name: Nurinsiyah, Ayu Savitri
name: Neiber, Marco T.
name: Hausdorf, Bernhard
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Camaenidae
Landouria
Landouria tonywhitteni

description: Landouria tonywhitteni sp. nov.
urn:lsid:zoobank.org:act: 59260748-3438-411F-82B4-14A7847AFA58
Figs 39, 52, 60, 63; Tables 3–4
Diagnosis
Landouria tonywhitteni sp. nov. is characterized by having a strongly depressed conical, brownishcorneous, sharply keeled shell and a penis that does not become distinctly broader towards is proximal end, the lack of a coecum at the proximal end of the penis and the presence of a node at the base of the flagellum.
Etymology
Landouria tonywhitteni sp. nov. is named in honour of the late British naturalist Tony Whitten (1953– 2017), who dedicated his life to the conservation of karst habitats and especially the snails that inhabit these habitats.
Material examined
Holotype
INDONESIA • holotype (det. anat.); Central Java, Grobogan, Taruman, near Sendang Sedayu; 7°00′52″ S, 110°51′29″ E; 20 m a.s.l.; 16 Aug. 2014; A.S. Nurinsiyah, F.L.H. Irsyad, F.J. Sari and E. Nurlaela leg.; measurements: D = 17.6 mm, H = 7.6 mm; MZB 19214.
Paratypes
INDONESIA – Central Java • 9 spec.; same data as for holotype; MZB 19217 • 5 spec.; same data as for holotype; ZMH 133434 • 9 spec. (det. anat.); Grobogan, Taruman, near Sendang Sedayu; 7°00′52″ S, 110°51′29″ E; 23 m a.s.l.; MZB 19215 • 1 spec.; same data as for preceding; MZB 21268 • 5 spec.; same data as for preceding; ZMH 133436.
Description
SHELL (Fig. 60; Tables 3–4). Depressed conical, with 5.75–6 hardly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct spiral lines, umbilical side with indistinct incised spiral lines; with radial scaly processes; tuberculate around umbilicus; brownishcorneous; body whorl sharply keeled; aperture rhombic; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 32–41% of shell diameter, hardly obscured by columellar edge.
GENITALIA (Figs 39, 63; Table 4). Atrium short; penis long, consisting of a long, cylindrical distal part covered by an additional sheath, which is connected by muscular tissue to proximal part of epiphallus and a dilated proximal part containing a short penial papilla; without penis coecum; penial retractor runs from diaphragm to distal part of long, narrow epiphallus; flagellum short, broadest at its base, where it is equipped with a globular node, tapering towards its proximal end, not distinctly crenated; vas deferens narrow and long; vagina long; peduncle of bursa copulatrix subdivided into a slightly broader, cylindrical distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.
Remarks
Landouria nodifera sp. nov., L. tonywhitteni sp. nov., L. pacitanensis sp. nov., L. sewuensis sp. nov. and L. intumescens share strongly depressed, brownish-corneous, sharply keeled shells. These species cannot be reliably distinguished by shell characters. Landouria tonywhitteni sp. nov. shares a node at the base of the flagellum with L. nodifera sp. nov., from which it differs in the lack of a coecum at the proximal end of the penis and in having a penis that does not become broader towards its proximal end.
Distribution
Landouria tonywhitteni sp. nov. is endemic to Sukolilo Karst (Fig. 52), where it occurs sympatrically with L. sukoliloensis sp. nov.
description: Published as part of Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, pp. 1-73 in European Journal of Taxonomy 526 on pages 45-47, DOI: 10.5852/ejt.2019.526, http://zenodo.org/record/3152217