Anthomyzidae Czerny
Creators: name: Roháćek, Jindřich
name: Barber, Kevin N.
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Anthomyzidae

description: Key to identincation of Nearctic genera of Anthomyzidae
1 f 1 lacking ctenidial spine posteroventrally, with only a row of long nne setae. .......... 2
– f 1 with distinct (although sometimes short) posteroventral ctenidial spine (Figs 7, 51) in addition to a row of long setae. ............................................................................... 3
2(1) Wing abbreviated and narrowed or with incomplete venation (at least dm-cu absent, see ROHÁĆEK & BARBER 2005: Figs 183–185); maximum eye diameter distinctly oblique. .............................................................................. Stiphrosoma Czerny, 1928, in part
– Wing normal, with complete venation; maximum eye diameter nearly vertical. .......... ...................................................................................................... undescribed genus
3(1) Wing with distinct longitudinal infuscation, especially along R 4+5 (Figs 49, 85). ...... 4
– Wing hyaline or with only indistinct infumation, never with discrete contrasting markings (Figs 11, 34). ............................................................................................... 5
4(3) Head markedly longer than high, depressed and anteriorly angular; eye elongate, with maximum diameter oblique to almost horizontal (Fig. 47); frontal triangle small and narrow, hardly reaching midpoint of frons; f 1 with ctenidial spine small (Fig. 51); gonostylus simple, with 2 denticles on apex (Fig. 59); transandrium with distinctively forked caudal process having its arms projecting and dilated anteroventrally (Fig. 55); postgonite slightly sinuous and without setulae (Fig. 56); spermatheca with eccentric duct insertion (Fig. 61) and spermathecal duct very long. ............................................. ............................ Ischnomyia Loew, 1863 [only I. albicosta (Walker, 1849)] (p. 41)
– Head slightly shorter than high, anteriorly rounded; eye broad, maximum diameter of eye nearly vertical (Fig. 89); frontal triangle large and broad, reaching at least to anterior nfth of frons; f 1 with ctenidial spine long (Fig. 87); gonostylus with attenuated apex but without denticles (Fig. 91); transandrium with caudal process simple (Fig. 94); postgonite sickle-shaped, with distinct anteroproximal setula (Fig. 92); spermatheca with normal duct insertion (Fig. 99) and spermathecal duct short (Fig. 105). .............. ........... Arganthomyza Roháček, 2009, in part [A. vittipennis (Walker, 1857)] (p. 66)
5(3) Maximum diameter of eye nearly vertical (Fig. 35); 3 strong ors, anterior ors subequal to or slightly shorter than middle seta and with additional microsetula in front of anterior ors; subvibrissa reduced (Fig. 35); all femora distinctly variegated brown and yellow; anal nssure very small, narrowly rounded triangular and medandrium abruptly narrowed dorsally (Fig. 36); internal structures of female genital chamber weak, annular sclerite reduced, almost indistinct (Fig. 45). ............................................................ ....... Fungomyza Roháček, 1999 [only F. buccata Roháček & Barber, 2004)] (p. 30)
– Maximum eye diameter more or less oblique; 1–3 strong ors, if 3 ors then the anterior distinctly shorter; 0–2 microsetulae in front of anterior ors; subvibrissa more or less distinct (longer than anterior peristomal setula); femora usually unicolourous (yellow or ochreous-orange), sometimes partly ochreous-brown; anal nssure larger and medandrium different; internal structures of female genital chamber usually well sclerotized and annular sclerite well developed. ........................................................................... 6
6(5) A single long ors (see VOCKEROTH 1987: Fig. 75.2), at most with a small microsetula anterior to this (no more than one-eighth length of ors); frontal triangle large and shining black. .............................................................................................................. 7
– At least 2 ors long (if two, the anterior at least one-third length of posterior ors and with additional short anterior microsetula present) (Figs 1, 6; ROHÁĆEK & BARBER 2011: Fig. 11); frontal triangle rarely shining black. .................................................................... 8
7(6) Fore legs unicolourous yellow to pale brown; silvery white tomentum above occipital foramen in a single medial patch; 1st antennal nagellomere nearly orbicular, with apex rounded; longest aristal hairs shorter than half width of 1st nagellomere; postgena with pale ventral border connuent with pale gena; palpus entirely pale; female abdomen entirely brown or sometimes slightly pale lateral to midline; T7 not divided medially. ............................. Mumetopia Melander, 1913 [only M. occipitalis Melander, 1913]
– t 1 and basal tarsal segments black with apical segments white; no white tomentum above occipital foramen; 1st antennal nagellomere longer than wide, apically obliquely rounded; longest aristal hairs as long as width of 1st nagellomere; postgena entirely dark, contrasting with pale gena; palpus apically darkened; female T2–T5 usually contrastingly pale with medial brown stripe; T7 longitudinally medially divided. ............................. ......................... undescribed genus [only Mumetopia nigrimana (Coquillett, 1900)]
8(6) Wing generally shorter than body length (sometimes much shorter), cross-vein r-m arising in basal third of dm cell; arista distinctly (shortly to long) pectinate (ROHÁĆEK & BARBER 2005: Figs 1, 32, 42); prs usually small or absent; female T7+S7 dorsomedially divided or unpigmented (ROHÁĆEK & BARBER 2005: Figs 13, 27); annular sclerite in female genital chamber strongly transversely compressed (ROHÁĆEK & ABER 2005: Figs 31, 179); spermathecae pyriform with entire surface strongly spinose (ROHÁĆEK & BARBER 2005: Figs 17, 46, 80). ........ Stiphrosoma Czerny, 1928, in part
– Wing generally longer than body length; cross-vein r-m arising beyond basal third towards the middle of dm cell; arista ciliate (Figs 6, 35, 395; ROHÁĆEK & BARBER 2011: 12); prs present and usually long; female T7+S7 dorsomedially undivided, but if unpigmented, then spermathecae not spinose on entire surface and annular sclerite not transversely compressed. ............................................................................................ 9
9(8) Contrasting silvery grey tomentose stripes present between frontal triangle and posterior part of orbits and join broadly across the front of the frontal triangle leaving only the ocellar triangle shiny; nlum of distiphallus slender and formed by two ribbon-shaped sclerites (see ROHÁĆEK 2006a: Figs 502, 505, as Paranthomyza caricis); spermatheca spherical to subspherical with only sparse tubercles or spines and without striations or apical invagination (see ROHÁĆEK 2006a: Fig 510). ......... Carexomyza Roháček, 2009
– Contrasting silvery grey tomentose stripes between frontal triangle and posterior part of orbits either absent, or if present then short and not joining anteriorly; nlum of distiphallus of various composition; spermatheca of various shape and vestiture usually with at least incomplete striations (Figs 117, 215, 288, 480), but if without striations then with apical invagination (Figs 267, 399, 540) or irregularly pyriform to spindleshaped (ROHÁĆEK & BARBER 2011: Figs 26, 119). ..................................................... 10
10(9) Body shiny black, sparsely tomentose (ROHÁĆEK & BARBER 2011: Fig. 10); two strong ors with additional microsetula anterior to these no stronger than the medial frontal microsetulae; face of female darker than that of male (ROHÁĆEK & BARBER 2011: Figs 8, 9); male gonostylus deeply bilobed (ROHÁĆEK & BARBER 2011: Figs 13, 15); nlum of distiphallus formed by two very slender ribbon-shaped sclerites, apically simple (ROHÁĆEK & BARBER 2011: Fig. 18); female T7+S7 dorsomedially divided but S8 not divided (ROHÁĆEK & BARBER 2011: Figs 23, 24); spermatheca elongately spindle-shaped with distal end rugged and provided with small blunt processes, never apically invaginated (ROHÁĆEK & BARBER 2011: Figs 26, 119). ........... Quametopia Roháček & Barber, 2011
– Body ranging from yellow to shiny brown or black, with sparse to dense grey tomentum; 2 or 3 strong ors, if only 2 strong ors then additional microsetula anterior to these may be as strong as or stronger than the medial frontal microsetulae; face of female never darker than that of male but the reverse may occur; male gonostylus never bilobed; nlum of distiphallus of various construction but if composed of 2 (partly fused) sclerites they are thicker and terminate in distinctly dilated apex (Figs 155, 174); female T7+S7 not divided dorsally but S8 longitudinally divided (Figs 79, 178, 253, 393, 475); spermatheca distally simple, often apically invaginated. ................................ 11
11(9) Only 2 strong ors with additional microsetula anterior to these usually weaker (only very rarely stronger) than medial frontal microsetulae; contrasting silvery grey tomentose stripes present between frontal triangle and posterior part of orbits; if lower part of thoracic pleurae yellow and contrasting with the dark notum (A. duplex, Figs 146, 147) then gonostylus narrowing toward apex (Fig. 156) and female with S7 fused with T7 to form syntergosternum (Fig. 159); thorax never with dense grey tomentum; males always with short thickened ventral setae in distal part of f 3; nlum of distiphallus formed by two thicker, more or less fused sclerites, apically dilated and complex and saccus at most with small rounded tubercles ((Figs 75, 135, 174); spermathecal ducts short (Figs 80, 124, 199). ....................................... Arganthomyza Roháček, 2009, in part (p. 56)
– Two or 3 strong ors, if only 2 strong ors then additional microsetula anterior to these always stronger than medial frontal microsetulae; contrasting silvery grey tomentose stripes usually absent between frontal triangle and posterior part of orbits, these stripes present only when at least lower half of thoracic pleurae is yellow and contrasting with brown notum (A. macra group, Figs 205, 206) where gonostylus is broad and spatulate (Figs 208, 231) and the female S7 is separate from T7 (Fig. 238); thorax often with dense grey microtomentum; males with or without modined ventral setae in distal part of f 3; nlum of distiphallus formed by single sclerite, apically simple and saccus with (usually robust) spines (Figs 212, 285, 390, 465, 556); spermathecal ducts very long (3 times or more as long as those in Fig. 199). ........... Anthomyza Fallén, 1810 (p. 105)
description: Published as part of Roháćek, Jindřich & Barber, Kevin N., 2016, Nearctic Anthomyzidae: a monograph of Anthomyza and allied genera (Diptera), pp. 1-412 in Acta Entomologica Musei Nationalis Pragae (suppl.) (suppl.) 56 on pages 26-28, DOI: 10.5281/zenodo.4272829