Achilia jeanneli Sabella & Cuccodoro & Kurbatov 2020, n. sp.
Creators: name: Sabella, Giorgio
name: Cuccodoro, Giulio
name: Kurbatov, Sergey A.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Achilia
Achilia jeanneli

description: Achilia jeanneli n. sp.
Figs 3, 17, 24, 32, 45-46, 59-62, 83
Holotype: MHNG (# MHNG-ENTO-81519); 1 ♂; CENTRAL CHILE: Región Araucanía: Malleco prov.: Nahuelbuta National Park, Piedra del Aquila; 37° 48’S 73° 02’W; 1300 m; 24.XII.1992; site 31b, sifting of moss on rock and tree trunks and vegetational debris; D. Burckhardt.
Paratypes (70 ex.): CENTRAL CHILE: Región Araucanía: Malleco prov.: MHNG (# MHNG-ENTO- 81520-29); 7 ♂ and 3 ♀; same data as holotype. – MHNS; 1 ♂ and 1 ♀; same data. – MHNG (# MHNG- ENTO-81530-73); 9 ♂ and 35 ♀; Nahuelbuta National Park, Piedra del Aquila; 1450 m; 15.XII.1990; site 10a, sifting of vegetational debris; M. Agosti & D. Burckhardt. – FMNH (FMHD #2002-045); 2 ♀; Nahuelbuta National Park, road to Piedra del Aquila; 37° 49.29’S 73° 01.90’W; 1360 m; 06-24.XII.2002; site 1055, Nothofagus dombey & pumilio, large Araucaria and bamboo, shrub understory; flight intercept traps; M. Thayer, A. Newton, A. Solodovnikov, D. J. Clarke & M. Chani. – FMNH (FMHD #2002- 096); 2 ♀; Nahuelbuta National Park, road to Piedra del Águila; 37° 49.7’S 73° 01.9’W; 1360 m; 25.XII.2002; Nothofagus and Araucaria, berlese, leaf & log litter; A. Solodovnikov. – MHNG (# MHNG- ENTO-81574-79); 3 ♂ and 3 ♀; Nahuelbuta National Park; 1100 m; 14-17.XII.1990; site 9a, forest litter; M. Agosti & D. Burckhardt. – NHMW; 1 ♂ and 2 ♀; Cordillera Nahuelbuta; H. Franz. – FMNH (FMHD #96- 224); 1 ♀; Nahuelbuta National Park, 4.5 km W Los Portones entrance; 37° 49.25’S 72° 59.82’W; 1300 m; 21.XII.1996; site 975, Nothofagus spp. emergent Araucaria aracauna, Chusquea understory, berlese, leaf & log litter; A. Newton & M. Thayer.
Description: Body 1.60-1.70 mm long, brown with reddish elytra, sometimes also head and pronotum reddish; antennae and legs reddish; palpi yellowish. Pronotum with disc slightly more convex than in A. grandiceps; posterior portion of lateral margins subparallel. First abdominal tergite with diverging basal striae extending to about one-fourth of paratergal length, and separated at base by about one-third of tergal width. Male: Head as in Figs 59-62, with occipital region and basal half of frons raised as two triangular protuberances, longer than wide, and separated by U-shaped median notch; lateral arms of notch thickly pubescent apically; anterior portion of frons deeply excavated; median apophysis curved and directed backwards, originating from broad median sub-basal clypeal hump, tip in dorsal view horseshoe-shaped with pubescent sides; median sub-basal clypeal hump with sides bearing only some sparse setae. Antennae (Fig. 17) with scape longer than wide, slightly swollen and slightly flattened; pedicel wider than long, slightly excavated on medial surface and with broadened lateral margin; antennomere III about as long as wide, antennomeres IV-VIII wider than long and slightly swollen; antennomere IX transverse with protruding mesal margin pointed at middle, antennomere X wider than long with denticulate margins and protruding mesal margin pointed at middle; antennomere XI elongate, with denticulate margins, shorter than VII-X combined. Metaventrite entirely covered by dense pubescence formed by long converging bristles, raised at middle for two- thirds of its distal portion, this surface entirely divided by broad median sulcus. Protrochanters with ventral margin forming median spine rounded or apically truncate, with long bristle at base of spine (Fig. 24); mesotrochanters (Fig. 32) with ventral margin forming spine on basal third; mesofemora (Fig. 32) with ventral margin similar to those of A. grandiceps, covered by broad, short and thick setae on basal third; mesotibiae with medial margin bearing recurved setae (Fig. 45), or not (Fig. 46), and forming short apical spine. Aedeagus (Fig. 3) 0.43-0.44 mm long; similar to that of A. grandiceps except lateral sclerites shorter, distinctly pointed and apically bent outwards.
Female: Similar to male except head not modified with frons slightly depressed behind scarcely impressed frontal sulcus.
Collecting data: Collected in December in different types of forest, at elevations ranging from 1110 m to 1450 m. All specimens came from sifted samples of leaf and log litter, sometimes with moss and vegetational debris included. Two females were collected by flight intercept traps.
Distribution: The species is known only from central Chile (Fig. 83: fuchsia star) in Nahuelbuta National Park (Malleco province).
Etymology: The species is dedicated to the French entomologist R. Jeannel.
Comments: Within the A. grandiceps group, the males of A. jeanneli n. sp. are easily distinguished from the other species by the shape of the head (Figs 59-62) and antennae (Fig. 17). The females of this species are very similar to those of A. grandiceps, from which they can be distinguished by their larger body size (1.6-1.7 mm for A. jeanneli versus 1.35-1.55 mm for A. grandiceps), and by the slightly swollen antennal scape of A. jeanneli n. sp.
description: Published as part of Sabella, Giorgio, Cuccodoro, Giulio & Kurbatov, Sergey A., 2020, A revision of the Chilean Brachyglutini - Part 6. Revision of Achilia Reitter, 1890: A. grandiceps, A. valdiviensis, and A. bicornis species groups (Coleoptera: Staphylinidae: Pselaphinae), pp. 129-156 in Revue suisse de Zoologie 127 (1) on page 135, DOI: 10.35929/RSZ.0013, http://zenodo.org/record/5743391