Thienemanniella
Creators: name: Fu, Yue
name: Saether, Ole
name: Wang, Xinhua
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Chironomidae
Thienemanniella

description: Key to known males of Thienemanniella
1. Eyes bare, antennal apex distinctly rounded................................................................................................................. 2
- Eyes pubescent or hairy, antennal apex notched or slightly rounded ........................................................................... 7
2. Inferior volsella absent ....................................................................................................... T. flavescens Edwards, 1929
- Inferior volsella present ................................................................................................................................................ 3
3 Antenna with 9–10 flagellomeres ................................................................................................................................ 4
- Antenna with 11–13 flagellomeres ............................................................................................................................... 5
4. Antenna with 9 flagellomeres, inferior volsella rounded .................................. T. togamijika Sasa et Okazawa, 1992
- Antenna with 10 flagellomeres, inferior volsella hook-shaped ................................................. T. okigrata Sasa, 1990
5. Inferior volsella rectangular and projecting................................................................................ T. partita Schlee, 1968
- Inferior volsella nearly triangular and along the inner margin of gonocoxite ............................................................. 6
6. Antenna with 12 or 13 flagellomeres, scutellum black in male (Tokunaga, 1936) ...... T. nipponica (Tokunaga, 1936)
- Antenna with 11 or 12 flagellomeres, scutellum yellow in male (Tokunaga, 1936) T. flaviscutella (Tokunaga, 1936)
7. Inferior volsella absent ................................................................................................................................................. 8
- Inferior volsella present ............................................................................................................................................. 10
8. Superior volsella hook-shaped; transverse sternapodeme without oral projection; phallapodeme straight and short .. ....................................................................................................................................................... T. hainanensis sp. n.
- Superior volsella triangular, transverse sternapodeme with oral projections; phallapodeme more or less curved and relatively long ............................................................................................................................................................... 9
9. Antenna with 11 flagellomeres, AR 0.72–0.90; third palpomere with digitiform lobe ......... T. similis (Malloch) 1915
- Antenna with 12 flagellomeres, AR 0.25–0.28; third palpomere without digitiform lobe.................... T. absens sp. n.
10. Inferior volsella narrow, digitiform to spatulate, sometimes with basal apodeme; or broad and low ........................ 11
- Inferior volsella well developed, rectangular to triangular......................................................................................... 20
11. Superior volsella completely fused with the inner of gonocoxite ........................... T. medialis Sublette & Sasa, 1994
- Superior volsella present, well developed or partly fused with the inner of gonocoxite............................................ 12
12. Inferior volsella digitiform with an apodeme, bare, phallapodeme straight ............................................................... 13
– Inferior volsella not digitiform, most often carrying thin glandular setae, phallapodeme curved ............................. 14
13. Superior volsella with a flat rim along the gonocoxite narrowly and completely fused .. T. antennata Freeman, 1953
- Superior volsella mostly triangular and fused only anteriomedially ............ T. lobapodema Hestenes & Saether, 2000
14. Inferior volsella placed basally or medially on gonocoxite........................................................................................ 15
- Inferior volsella very low and placed caudally on gonocoxite ................................................................................... 18
15. AR 0.52–0.58, inferior volsella very broad and rounded ............................................................................................... ........................................................................................................................... T. akagiquarta Kikuchi & Sasa, 1994
- AR 0.20–0.30, inferior volsella digitiform or narrow................................................................................................. 16
16. Gonocoxite with large concavity for gonostylus, tergite IX with many long setae........................................................ ......................................................................................................................... T. cavata Fu, Hestenes & Saether, 2010
- Gonocoxite without such concavity for gonostylus, tergite IX without many long setae .......................................... 17
17. Antenna with 8 flagellomeres; tergite IX with a small, flexible, triangular hump-like extension; apex of phallapo- deme bent 90° caudally. ............................................................................................................ T. fuga Lehmann, 1979
- Antenna with 10 flagellomeres; tergite IX without hump-like extension; apex of phallapodeme slightly curved ........ .............................................................................................................................................. T. lineola Freeman, 1953
18. Antenna with 9 flagellomeres; gonostylus expanded; superior volsella more or less digitiform and projecting........... ...................................................................................................................................... T. nagaramaculata Sasa, 1989
- Antenna with 10 flagellomeres; gonostylus not expanded; superior volsella narrow and not projecting .................. 19
19. AR 0.26, tergite IX partly semitransparent, gonocoxite without a long dark apodeme ................................................. .............................................................................................................................. T. yakysetea Sasa et Suzuki, 2000 a
- AR 0.40–0.66, tergite IX not partly semitransparent, gonocoxite with a long apodeme ............................................... .............................................................................................................................. T. tusimufegea Sasa et Suzuki, 1999
20. Superior volsella distinct from inferior volsella ......................................................................................................... 21
- Superior volsella partly overlapping or joined with inferior volsella......................................................................... 33
21. Inferior volsella bare, with apodeme; transverse sternapodeme narrow and very thin, without oral projection............ ...................................................................................................................................... T. trivittata Goetghebuer, 1935
- Inferior volsella with thin glandular setae, without apodeme; transverse sternapodeme wide and thick, with oral pro- jection.......................................................................................................................................................................... 22
22. Inferior volsella low and placed caudally on gonocoxite ........................................................................................... 23
- Inferior volsella high and placed basally or medially on gonocoxite ......................................................................... 30
23. Third palpomere with small apical projection; gonostylus with 2 megasetae T. taurocapita Hestenes & Saether, 2000
– Third palpomere without small apical projection; gonostylus with 1 megaseta......................................................... 24
24. Superior volsella large, triangular and close to inferior volsella, conspicuously projecting ..................................... 25
- Superior volsella small, triangular or narrow and at a distance from inferior volsella, not projecting ...................... 27
25. AR about 0.30; inferior volsella with rounded corner, posterior margin of superior volsella flat ................................ ..................................................................................................................................................... T. safi Lehmann 1981
- AR about 0.50; inferior volsella caudally hooked, posterior margin of superior volsella oblique............................. 26
26. Tergite IX partly semitransparent; phallapodeme slightly curved, gonostylus narrowed near apex ............................. ............................................................................................................................ T. ginzanquerea Sasa et Suzuki, 2001
- Tergite IX not partly semitransparent; phallapodeme distinctly curved distinctly; gonostylus equally wide to apex .. ..................................................................................................................... T. oyabedilata Sasa, Kawai et Ueno, 1988
27. All trochanters of legs with dorsal keel ........................................................................................... T. liae Paggi 2007
- Trochanters without keel or only fore trochanter with dorsal keel............................................................................. 28
28. Superior volsella joined medially; inferior volsella largely typical triangular ................................... T. sichuana sp. n.
- Superior volsella separate medially; inferior volsella not largely triangular.............................................................. 29
29. Posterior of inferior volsella hooked; tergite IX not partly semitransparent .................................................................. ........................................................................................................... T. ginzanquinta (Sasa et Suzuki, 1998) comb. n.
- Posterior of inferior volsella rounded; tergite IX partly semitransparent............................... T. spreta (Roback, 1962)
30. Inferior volsella rectangular........................................................................................................................................ 31
– Inferior volsella largely triangular .............................................................................................................................. 32
31. Antenna with 10 flagellomeres; superior volsella conspicuously projecting ........ T. boltoni Hestenes & Saether, 2000
- Antenna with 12–13 flagellomeres; superior volsella not projecting .................................. T. minuscule Brundin 1949
32. Superior volsella triangular and projecting; phallapodeme without projection, placed below sternapodeme .............. .......................................................................................................................................................... T. wuyiensis sp. n.
- Superior volsella narrow and not projecting; phallapodeme with projection for joint with sternapodeme placed pre- lateral ................................................................................................................................................. T. triangula sp. n.
33. Antenna with 10 flagellomeres, apical part of superior volsella hooked.................................................................... 34
– Antenna with 11–13 flagellomeres, apical part of superior volsella rounded or flat.................................................. 35
34. AR 0.30; superior volsella long triangular and projecting; tergite IX not partly semitransparent ................................ ...................................................................................................................................... T. chuzeduodecima Sasa, 1984
- AR 0.46; superior volsella narrow and not projecting; tergite IX partly semitransparent.............................................. ............................................................................................................................... T. tusimuefea Sasa et Suzuki, 1999
35. Inferior volsella triangular .......................................................................................................................................... 36
- Inferior volsella nearly rectangular............................................................................................................................. 37
36. Antenna with 11 flagellomeres; AR 0.35, superior volsella projecting, triangular ........................................................ .......................................................................................................................... T. caspersi Saether et Ferrington, 2003
- Antenna with 13 flagellomeres; AR 0.27, superior volsella fused (not distinctly visible)............................................. .................................................................................................................... T. ogasaquindecima Sasa et Suzuki, 1997
37. Third palpomere with small apical projection; phallapodeme expanded medially, superior volsella nearly rectangular .......................................................................................................... T. tiunovae Makarchenko et Makarchenko 2006
- Third palpomere without apical projection; phallapodeme not expanded medially, superior volsella nearly triangular or rounded ................................................................................................................................................................... 38
38. Superior volsella separated medially, triangular, apical part nearly right-angled T. tonewquerea Sasa et Tanaka, 2002
- Superior volsella joined medially, rounded or narrowed, apical part with rounded corner........................................ 39
39. Antenna with 10 or 11 flagellomeres ......................................................................................................................... 40
- Antenna with 12 or 13 flagellomeres.......................................................................................................................... 41
40. AR 0.30–0.37; superior volsella small and not projecting; phallapodeme long and strongly curved; apex of gonosty- lus straight ......................................................................................................................... T. clavicornis Kieffer, 1911
- AR 0.60–0.76; superior volsella large and projecting; phallapodeme short and slightly curved; apex of gonostylus slightly hooked ..................................................................................................................... T. obscura Brundin, 1947
41. Superior volsella small and not projecting ................................................................................................................. 42
- Superior volsella large rounded and conspicuously projecting .................................................................................. 45
42. Anal vein of wing surpassing cubital fork; inferior volsella almost flat posteriorly .................................................. 43
- Anal vein of wing not reaching cubital fork; inferior volsella slightly oblique posteriorly ...................................... 44
43. Antenna with 13 flagellomeres; AR more than 0.50 ......................................................... T. acuticornis Kieffer, 1912
- Antenna with 12 flagellomeres; AR 0.30 ................................................... T. ogasaquardecima Sasa et Suzuki, 1997
44. Ultimate flagellomere of antenna longer than combined length of preceding eight flagellomeres; AR about 0.85 ...... ........................................................................................................................................ T. majuscula (Edwards, 1924)
- Ultimate flagellomere of antenna shorter than combined length of preceding eight flagellomeres; AR about 0.65 ..... ............................................................................................................................................... T. lutea (Edwards, 1924)
45. Body color very pale, AR more than 0.60 ............................................................ T. gotopallida Sasa et Suzuki, 2000
- Body color relatively dark, AR usually about 0.50 .................................................................................................... 46
46. Fore trochanter with keel, inferior volsella angle-like near apex .............................................. T. xena (Roback, 1957)
- Fore trochanter without keel, inferior volsella rounded near apex ....................................... T. vittata (Edwards, 1924)
description: Published as part of Fu, Yue, Saether, Ole & Wang, Xinhua, 2010, Thienemanniella Kieffer from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae), pp. 1-42 in Zootaxa 2431 on pages 32-34, DOI: 10.5281/zenodo.194711