Daedalma dognini subsp. dognini Pyrcz
Creators: name: Pyrcz, Tomasz W.
name: Greeney, Harold F.
name: Willmott, Keith R.
name: Wojtusiak, Janusz
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Nymphalidae
Daedalma
Daedalma dognini
Daedalma dognini dognini pyrcz

description: Daedalma dognini dognini Pyrcz   
(Figs. 4G, 4H, 11A, 14D)  
 Material examined: PERU: HOLOTYPE male: Cuzco, Ollantaytambo-Alfamayo, Qda. San Luís, Carrizales, 3100–3150 m, 20.V.2003, T. Pyrcz leg., red, rectangular label saying: Holotype, MZUJ (to be deposited in MUSM); PARATYPE female: same data but 3200–3250 m, TWP.   
 Diagnosis: Recognised from the only sympatric congener, D. boliviana, by the uniform brown upperside; very similar to the Colombian species D. drusilla, except for the slightly lighter upperside, wider wings and a reddish suffusion of the FWV.   
 Description: MALE (Fig. 4G): Head: eyes blackish brown, hairy; labial palpi covered with dense, short, brown hair; antennae extending to 2/5 length of costa, orange, shaft slender, club twice as thick and flattened. Thorax: blackish brown; legs light beige. Abdomen: blackish brown. Wings: FW length 27 mm; outer margin sharply produced at vein M1; fringes yellowish. HW with two tail-like extensions along veins Cu1 and Cu2, former slightly longer; fringes very short, brown, except at apex where yellowish and slightly longer. FWD: uniform medium brown, with a very faint, barely noticeably lighter postdiscal area reflecting shape of reddish patch from underside. HWD: uniform medium brown. FWV: ground colour medium brown; a faint reddish suffusion in postbasal area, and a red-brown, irregular patch in discal cell; a wide red-light brown area covering most of postdiscal to submarginal area, reddish suffusion fading away towards subapical area giving place to light brown, and some white scaling along costa, with a deep notch pointing distally along its basal edge between veins M1 and M3; subapical area chocolate brown with some silver and magenta scaling forming a zigzagging submarginal line extending as far as vein M3; three minute subapical ocelli in R4-R5, R5-M1 and M1-M2. HWV: ground colour chocolate brown with some red-brown and milky white scaling forming a complex, nondescript mosaic typical of genus; two milky white submarginal patches in M2-M3 and M3-Cu1, latter noticeably bigger and reminiscent in shape and position of that of D. drusilla. Genitalia (Fig: 11A): Uncus stout, longer than dorsum of tegumen; gnathos 2/3 length of uncus, gradually thinner from a wide base to a fine apical part; saccus deep, about length of uncus; pedunculus prominent, especially compared to D. inconspicua; valvae wide and solid in basal half, gradually narrowing towards apex, terminated with a rather short hook curved upwards; aedeagus about length of valvae, smooth, slightly arched in middle.   
 FEMALE (Fig. 4H): FW length 29 mm; FWD and HWD lighter than in male, with lighter patches showing more conspicuously from underside; FWV reddish suffusion lighter and much fainter, concentrated along basal edge of light orange-brown postmedian area. HWV colour pattern lighter, otherwise similar to male. Genitalia (Fig. 14D): Sinus vaginalis wide and shallow. Marginal parts of V-shaped batten on lamella postvaginalis consisting of heavily sclerotized slats that narrow slightly laterally. Papillae anales posteriorly close to each other. Apophyses posteriores as short triangular plates sharply pointing anteriorly. Posterior wall of lamella postvaginalis concave, with delicate wrinkles. Ductus bursae curved, with few pronounced wrinkles, twice length of antrum. Posteriorly to well developed colliculum, another ring-like cuticular thickening on wall of antrum. Hump in middle of lamella postvaginalis very small, triangular. Bursa copulatrix with two ribbon-like and narrow, parallel signa consisting of minute teeth.   
 Etymology: This species is dedicated to Paul Dognin, the French naturalist who, by the end of 19th century, had described several species and subspecies of Pronophilina, including Daedalma (inconspicua) palacio.   
 Remarks: This species superficially resembles the northern Andean D. drusilla and D. inconspicua, particularly in the uniform dorsal brown pattern. However, their ventral patterns, showing some distinctive characters such as the wide reddish FW patch (conspicuous in the nominate subspecies), and the male genitalia, are sufficiently distinct to treat D. dognini as a species. Furthermore, any closer association of D. dognini with D. inconspicua would be unjustified from a biogeographical standpoint, considering that their ranges are widely disjunct. D. dognini has no known close relatives in central Peru. D. adamsi, which occurs at similar elevations in central Peru, has very different male genitalia, which rules out any closer affinity (see under D. adamsi). Available elevational data (3100–3250 m) indicate that D. dognini is an uppermost forest species occurring near the treeline.
description: Published as part of Pyrcz, Tomasz W., Greeney, Harold F., Willmott, Keith R. & Wojtusiak, Janusz, 2011, 2898, pp. 1-68 in Zootaxa 2898 on page 26