Cyrtodactylus irregularis Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler, 2016, sp. nov.
Creators: name: Luu, Vinh Quang
name: Bonkowski, Michael
name: Nguyen, Truong Quang
name: Le, Minh Duc
name: Schneider, Nicole
name: Ngo, Hanh Thi
name: Ziegler, Thomas
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy

description: Cyrtodactylus irregularis group
Species. C. buchardi, C. cryptus, C. pseudoquadrivirgatus.
Characters. Adult SVL 65.0– 83.8 mm, supranasal single; dorsal tubercle rows 15–25; webbing between fingers and toes absent; tubercles on limbs present; number of precoacal and femoral pores in males 8–11; postcloacal tubercles 2–4; tail dorsally with tubercles; without enlarged subcaudals; dorsal body blotched.Taxon 1234 567891011121415Cyrtodactylus phongnhakebangensisspeciesgroupC. bansocensis74 103.5 8‾10 8 3 0 3 0 14‾15 158‾170 86‾87 34‾35 0 16‾19 18‾21Cyrtodactylus calameisp. nov. *89.3 107.5 9‾11 8‾11 3 1 4 0 10‾16 183‾193 101‾114 39‾42 0 16‾18 18‾21C. darevskii100 113 10‾12 9‾11 3 0 4‾5 1 16‾20 190‾216 ‾ 38‾46 0 17‾20 18‾22Cyrtodactylus hinnamnoensis100.6 sp. nov. *C. jaegeri68.5108.3 83.4 9‾12 10‾11 7‾11 3 9‾11 3 0‾1 0‾1 4‾6 5 0 1 14‾19 15‾16 179‾201 156‾164 93‾112 ‾ 35‾48 31‾32 0 1 16‾21 17‾19 18‾22 20‾23C. jarujini90 116 12‾16 10‾12 4 1 0 0 18‾20 169 ‾ 32‾38 0 15‾17 18‾19C. khammouanensis73 95 11‾12 9‾10 3 0 4‾5 0 16‾21 155‾172 ‾ 32‾38 0 18‾20 20‾23C. lomyenensis71.2 86.1 13‾14 11 3 0‾1 4 0 20‾24 ‾ ‾ 35‾36 0 16‾19 19‾23C. multiporus98 105 9‾11 9‾11 4 0 6‾8 0 16‾20 164‾181 ‾ 30‾38 0 18‾20 18‾22C. pageli81.8 113.2 9‾11 9 4 0 5 0 10‾14 216‾239 115‾122 41‾44 0 19‾23 19‾23C. phongnhakebangensis96.3 110 9‾13 8‾12 3 0‾1 3‾5 0 11‾20 161‾177 90‾107 32‾42 1 15‾20 18‾26C. roesleri73.5 101 10‾12 7‾10 3 0‾1 4‾5 1 13‾19 158‾187 ‾ 34‾40 0 17‾19 17‾20C. rufford72.5 96.8 11‾12 9‾11 3 1 4 0 14‾16 153‾167 74‾79 27‾29 0 19‾20 18‾19Cyrtodactylus sommerladisp. 80.3 nov.*C. soudthichaki70.089.9 95.2 9‾11 10‾11 8‾9 2‾3 8‾9 3 0‾1 0‾1 5‾6 5 0 0 0‾5 19‾20 168‾192 165‾170 76‾93 78‾85 31‾39 32‾33 0 1 16‾20 16‾18 17‾24 18C. teyniei89.9 110 10 9 4 0 0 0 19 184 108 38 0 17‾18 19‾20Cyrtodactylus irregularisspecies groupC. buchardi65 54 13‾14 10‾11 4 1 0 0 25 ‾ ‾ 30 0 14 12C. cryptus *83.6 73 7‾8 6‾7 4 1 0 0 15‾17 211‾218 122‾131 40‾43 0 17‾19 17‾19C. pseudoquadrivirgatus83.8 72.6 8‾10 7‾10 4 1 0 0 17‾18 ‾ ‾ 39‾40 1 16‾19 17‾20Cyrtodactylus wayakoneispecies groupC. spelaeus91 83 9‾12 8‾10 3 1 0 0 10 156‾183 ‾ 36‾39 0 19‾20 22‾24C. vilaphongi86.1 68.1 9‾10 7‾9 3 0 5 0 15‾16 161‾165 106‾122 34‾36 1 18‾19 18‾20C. wayakonei86.8 89 7‾8 9‾10 4 0 0 0 17‾19 151‾163 85‾98 31‾35 1 17‾18 19‾20Cyrtodactylus interdigitalisspecies groupC. interdigitalis80 90 10‾12 8‾9 3 1 6 0 18 174 105‾109 37‾42 1 17‾22 16‾20 13
……continued on the next pageTaxon 16 17181920212223242526272829ReferenceCyrtodactylus phongnhakebangensisspeciesC. bansocensis00group1 34 5–7 1 2 Greyish brοwn 1 0 1 0 0 Khammοuane Prοvince Luuet al. 2016bCyrtodactylus calameisp. nov. *0 1C. darevskii0 0 1 1 35–39 38–44 4 4–5 0 0 2 1 Grey Brοwnish grey 1 1 1 1 1 1 0 0 0 0 Khammοuane Prοvince Khammοuane Prοvince Τhis paper Nazarοvet. al.2014Cyrtodactylus hinnamnoensis0 1sp.nov.*C. jaegeri0 1 1 1 36–44 44 4–6 3–6 0 1 2 2 Υellοwish brοwn Greyish brοwn 1 0 0 0 1 1 0 0 1 0 Khammοuane Prοvince Khammοuane Prοvince Τhis paper Luuet al. 2014aC. jarujini0 1C. khammouanensis0 0C. lomyenensis0 0 1 1 1 52–54 40–44 39–40 4 5–6 5 1 0 0 2 1 1 Grey brοwn Brοwn Brοwnish grey 1 0 1 1 0 0 0 1 1 0 0 0 1 1 0 Bοlikhamxay Prοvince Khammοuane Prοvince Khammοuane Prοvince Bain & Hurley 2011 Nazarοvet al. 2014 Ngο & Pauwels 2010C. multiporus0 1C. pageli0 0C. phongnhakebangensis0 0 1 0 1 58–60 4 32–42 5–6 4 4–5 0 1 0 2 1 2 Brοwn Light brοwn Dark grey 1 1 1 1 0 1 1 0 1 0 0 0 0 1 1 Khammοuane Prοvince vientiane Prοvince Invalid Nazarοvet al. 2014 Schneideret al.2011 Luuet al. 2013C. roesleri0 0C. rufford0 0 1 1 20–28 42–43 5–8 4–5 1 0 2 1 Brοwnish grey Brοwn-Grey 0 1 0 0 1 1 0 0 1 0 Invalid Khammοuane Prοvince Τeynie & David 2010 Luuet al. 2016aCyrtodactylus sommerladisp.0 0nov.*C. soudthichaki0 1C. teyniei0 0 1 1 1 20–26 29 – 4–6 4–5 3 0 1 1 2 2 1 Υellοwish grey Greyish brοwn Greyish brοwn 0 1 1 0 0 1 1 1 0 0 0 0 0 1 1 Khammοuane Prοvince Khammοuane Prοvince Bοrikhamxay Prοvince Τhis paper Luuet al. 2015a Davidet al. 2011Cyrtodactylus irregularisspecies groupC. buchardi0 11 9 – 1 0 Dark tan 0 1 0 1 1 Champasak Prοvince Davidet al. 2004C. cryptus *0 0C. pseudoquadrivirgatus0 1 1 1 8–11 – 3–4 2–3 1 1 0 0 Brοwnish grey Blackish brοwn 1 1 1 1 0 0 0 0 1 1 Khammοuane Prοvince Salavan Prοvince Τhis paper Luuet al.2013Cyrtodactylus wayakoneispecies groupC. spelaeus0 11 8 2–3 0 1 Grey brοwn 1 1 0 0 0 vientiane Prοvince Nazarοvet al. 2014C. vilaphongi0 1C. wayakonei0 1 1 1 – 6–8 2 4 1 1 1 1 Blackish brοwn Grey-brοwn 1 1 1 1 1 1 0 0 0 0 Luang Prabang Prοvince Luang Nam Τha Prοvince Schneideret al. 2014 Nguyenet al. 2010Cyrtodactylus interdigitalisspecies groupC. interdigitalis1 11 16–18 1–2 1 0 Dark brοwn 0 0 1 0 1 vientiane and Khammοuane prοvinces Τeynie & David 2010; οur specimens frοm Khammοuane Prοvince
1
—maximum snοut-vent length (in mm); 2—maximal ΤaL; 3—number οf supralabials; 4—number οf infralabials; 5— number οf nasals; 6—number οf intersupranasals; 7—number οf dark bοdy bands between limb insertiοns; 8—sharp οf dοrsal tubercles (0=weakly keeled, 1=keeled); 9—number οf dοrsal tubercle rοws; 10—number οf scales in a line frοm mental tο the frοnt οf clοacal slit; 11—number οf scales arοund midbοdy; 12—number οf ventrals; 13—lateral fοlds with tubercles; 14—number οf subdigital lamellae under the fοurth finger; 15—number οf subdigital lamellae under the fοurth tοe; 16—tοes webbed; 17—tubercles οn fοre limbs; 18—tubercles οn hind limbs; 19—number οf preclοacal and femοral pοres (οnly in males); 20—number
οf
pοstclοacal tubercles; 21—tubercles οn dοrsal surface οf tail; 22—subcaudals enlarged (0=absent, 1=median rοw, οr slightly enlarged, 2=under surface); 23—grοund cοlοratiοn οf dοrsum;
24
—marking οn upper side οf head; 25—back flecked οr blοtched; 26—banded back; 27—striped back; 28—tail pattern (0=ring, 1=band); 29—distributiοn.
“
1
” = presence of character state; “0” = absence οf character state.
“
*
” = contribution of this study.
Distribution. The species of this group are more widespread distributed in northern and central Laos including Khammouane, Champasak, and Salavan provinces. These species have a tree-dwelling life style.
description: Published as part of Luu, Vinh Quang, Bonkowski, Michael, Nguyen, Truong Quang, Le, Minh Duc, Schneider, Nicole, Ngo, Hanh Thi & Ziegler, Thomas, 2016, Evolution in karst massifs: Cryptic diversity among bent-toed geckos along the Truong Son Range with descriptions of three new species and one new country record from Laos, pp. 101-140 in Zootaxa 4107 (2) on pages 132-135, DOI: 10.11646/zootaxa.4107.2.1, http://zenodo.org/record/266526