Eminespina burma Chen & Zhang & Shi 2021, sp. nov.
Creators: name: Chen, Xin-Yu
name: Zhang, Hua-Chuan
name: Shi, Xiaoxiao
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Alienoptera
Alienopteridae
Eminespina
Eminespina burma

description: Eminespina burma sp. nov.
(Figs 1-5)
urn:lsid:zoobank.org:act: B03B0F38-33E9-417E-BAA4-68E9A74EEE33
Etymology. The specific name is a Latin word “ Burma ” referring to the country from which the material originates.
Type material. Holotype: female, in Burmese amber, CSCLRCAB100340.
Diagnosis. As for the genus.
Description. Female, flattened body complete (Fig. 1). Body and appendages reddish brown, length (from anterior margin of head to tip of abdomen, exclusive of cerci) 7.4 mm; maximum width of head 2.2 mm.
Hypognathous head (Fig. 2A, C; h) strongly transversal, with a frontal visible ocellus (other lateral ocelli invisible), compound eyes (Fig. 2A, C; ce) strongly convex (hemispherical), diameter 0.68 mm, with large near-circular base and extremely small ommatidia. Chewing mouthparts standard; labrum well developed, free; clypeolabral suture not recognizable. Mandibles not visible. Maxillae long, visible four-segmented palps, maxillary palp (Fig. 2B; mp) length 1.8 mm; distal palpomere longest, widened distally, distal end truncate; 5th segment narrow, conical. Antennae (Fig. 2C; a) long and slender, slightly shorter than half of body length, filiform, multi-segmented, first 15 antennomeres (total 20) covered with numerous short, thick setae (each segment bearing up to 20 setae). Neck (Fig. 2A; n) broad, width equal to one-third of head. Anterior margin of subgenae with distinct triangular crest (Fig. 2C; tc) below each compound eye. Sulcus or sutura between antennal sockets not visible. Epicranial sutures not visible; occipital and parietal ridges not recognizable.
Pronotum cylindrical, evidently longer than wide, posterior and anterior widths equivalent (i.e., not constricted); indistinct transverse furrow (supracoxal furrow, Fig. 2A, C; sf) separates posterior one-third of pronotum from longer anterior part; three distinct stout spines distributed anterior quarter of pronotum from longer posterior part. Mesonotum and metanotum not visible. Legs moderately long, similar in their armature of spines and general shape, length slightly increasing from prothorax to metathorax. Fore femur (Figs 3C, 4A; ff) without spurs, profemoral brush absent, 0.6 mm long and 0.17 mm wide; fore tibia with two apical spurs, 0.5 mm long and ca. 0.08 mm wide; tarsus 0.33 mm long. Mid femur (Fig. 3C; mf) 1.6 mm long and 0.6 mm wide; mid tibia with two apical spurs, 1.7 mm long and 0.2 mm wide; mid tarsi 1.7 mm long. Hind femur (Figs 3C; hf & 3F) with small femoral spur (Fig. 3E), 0.5 mm long; hind tibia 2.2 mm long (0.04 mm wide) also with two long apical spurs (0.3 mm long) and three shorter spurs. Tarsus (Figs 3A, 4B & 4E) five-segmented, Ta1-3 of all legs with two pad-like euplantulae on ventral side, Ta4 with only one euplantula with shallow groove in proximal third; size of tarsi decreases from Ta1 to Ta4, small Ta4 distinctly protruding; elongate and slender apical Ta5 strongly bent upward. Arolium present, pretarsus with paired asymmetrical claws (Figs 3D & 3A; cl).
Short and stout abdomen longer than wide and displays only seven visible sternites (Fig. 4F). Cerci (Fig. 4C) distinctly developed, 0.7 mm long, conical, strongly setose, about as long as distance between their bases, arise above the subgenital plate; composed 13 cercomeres (Fig. 4C; Ce1–13); Ce1–7 wider than long, oblong, Ce8-10 elongate and slender, Ce11-12 equal length and width, and elongate and slender distal cercomere (Ce13). The large terminal sternite (subgenital plate) of the female is triangular and apically pointed. Two valves of the ovipositors (Fig. 4D) are visible at least.
Forewings (Figs 5A, 5B; fw) fully developed (hardened tegmina), 5.7 mm long and 2 mm wide, forming scale-like pseudoelytra with leathery cuticular structure and bunky present; veins not recognizable in forewings. Hindwing (Figs 5A, 5B, 5C; hw) fully developed, 5 mm long, about 2 mm wide, transparent with thick veins ending before wing margin and numerous cross-veins forming a fenestrate structure. Hindwing apex approximately reaches abdominal apex; veins C, Sc, J, &A invisible or indistinct; three cross veins between R1 and R2 at least, Rs bifurcate in distal one-third, R3 straight, simple; MA and MP fused basally in proximal one-third; MA bifurcate in distal onefifth, MA1, MA2, and MP straight, simple, slightly sinuated; pterostigma distinct (Fig. 5C; p), sharp not obvious.
Male and morphs. Unknown.
description: Published as part of Chen, Xin-Yu, Zhang, Hua-Chuan & Shi, Xiaoxiao, 2021, A new species and genus of Alienopteridae (Blattodea) from mid-Cretaceous amber of northern Myanmar, pp. 580-586 in Zootaxa 4941 (4) on pages 581-583, DOI: 10.11646/zootaxa.4941.4.7, http://zenodo.org/record/4595746