Scolytocis howdeni Lopes-Andrade 2008, sp. nov.
Creators: name: Lopes-Andrade, Cristiano
Publisher: name: Zenodo
Year: 2008
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Ciidae
Scolytocis
Scolytocis howdeni

description: Scolytocis howdeni Lopes-Andrade sp. nov. (Figs 39–41, 58, 64B, 67B, 72, 80)   
Etymology The species is named after Henry F. Howden, who collected six specimens of the type series.  
Diagnosis  
 The species belongs to the danielssoni species-group, and so it is distinguishable from the other Scolytocis species by the biconcave prosternum with a narrow longitudinal carina at midline, the rugose border along the posterior pronotal margin and the comparatively large tibiae. It differs from the other species in the group mainly by the close pronotal punctation, with punctures separated by a distance of one puncture width or less.   
Description  
 Holotype. (Figs 39–41) Measurements in mm: TL 2.00; PL 0.79; PW 0.89; EL 1.21; EW 0.95; GD 0.89. Ratios: PL/PW 0.88; EL/EW 1.28; EL/PL 1.53; GD/EW 0.94; TL/EW 2.11. Body dark brown, head and pronotum slightly darker than elytra; basal antennomeres, mouthparts and tarsi yellowish brown; dorsal and ventral surfaces almost entirely glabrous. Head with dorsum coarsely punctate; punctures irregular, with interstice finely microreticulate. Each antenna (Fig. 64B, paratype) with nine antennomeres (FL 0.098mm; CL 0.160mm; CL/FL 1.64); length of the antennomeres (in mm) as follows: 0.118; 0.055; 0.053; 0.015; 0.015; 0.015; 0.050; 0.045; 0.065. Eyes with greatest width 1.33X the basal width of scutellum. Pronotum coarsely punctate; punctures irregular, limits distinct, usually separate by a distance of one puncture width or less, but not coalescent; interstice between punctures finely microreticulate; anterolateral margins straight; posterolateral angles broadly rounded; posterior margin with a conspicuous rugose border along it. Scutellum rugose, basal width 0.13X the EW. Elytra with punctation slightly coarser, denser and more irregular than that of pronotum; interstice between punctures smooth; elytral apex truncate, with a small concavity when seen from above; cuticular globules absent. Each hindwing (Fig. 58) with apical area bearing one vague, incomplete pigmented line near the anterior margin. Prosternum biconcave, with a narrow longitudinal carina at midline; surface microreticulate to striated. Metaventrite glabrous, surface microreticulate to striated; discrimen indistinct; either side bearing large obsolescent punctures. Abdominal ventrites glabrous; surface microreticulate. Each metatibia (Fig. 67B) near 3.0X as long as broad; outer apical and outer lateral edges forming an obtuse angle that is broadly rounded; outer edge with spines (around 20) regularly distributed and more close to each other at apical third (outer apical edge), then getting sparser. Male genitalia (in a paratype). (Fig. 72) Apex of tegmen (sclerotized apical half) subtriangular, base membranous; median lobe membranous, elongate, near 1.2X longer than tegmen.   
Type series  
 Holotype. (CNCI) Mexico: / 3 mi. N.W. San Cristobal L.C., Chis. Mex. V. 29 1969 H.F. Howden / Scolytocis howdeni Lopes-Andrade HOLOTYPUS [printed on red paper]/. Paratypes. Mexico: 4 specimens (2 CNCI, 2 LAPC), same data as holotype; 1 specimen (ANIC) / 6.4mi. S. Tenango de Doria Hidalgo MEX. July 24, 1969 / S. & J. Peck Collectors / Ganoderma sp. / J. F. Lawrence Lot [printed] 3079 [handwritten]/. All the paratypes with an additional label / Scolytocis howdeni Lopes-Andrade PARATYPUS [printed on yellow paper]/   
Variation  
Measurements in mm (n = 6, including the holotype): TL 1.84–2.00 (1.92 ± 0.07); PL 0.68–0.79 (0.70 ± 0.04); PW 0.79–0.89 (0.85 ± 0.04); EL 1.16–1.26 (1.20 ± 0.04); EW 0.84–0.95 (0.92 ± 0.04); GD 0.84–0.89 (0.86 ± 0.03). Ratios: PL/PW 0.76–0.88 (0.83 ± 0.04); EL/EW 1.22–1.38 (1.31 ± 0.06); EL/PL 1.53–1.85 (1.72 ± 0.11); GD/EW 0.89–1.00 (0.93 ± 0.04); TL/EW 1.94–2.19 (2.09 ± 0.08).  
Distribution Known from Chiapas and Hidalgo, Mexico (Fig. 80).  
 Host fungi Ganoderma sp. (Ganodermataceae).   
Comments  
 It resembles Scol. indecisus sp. nov. in the coarse and dense punctation of elytra and pronotum, but is easily distinguished by the features mentioned in the diagnosis.
description: Published as part of Lopes-Andrade, Cristiano, 2008, An essay on the tribe Xylographellini (Coleoptera: Tenebrionoidea: Ciidae), pp. 1-110 in Zootaxa 1832 (1) on pages 21-22, DOI: 10.11646/zootaxa.1832.1.1, http://zenodo.org/record/5126979