Clivina apexplana Balkenohl 2021, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina apexplana

description: <i>Clivina apexplana</i> sp. nov. urn:lsid:zoobank.org:act: 1C881377-546B-402C-A10B-D83B785B2481 (Figs 30, 50, 70, 97, 114) ETYMOLOGY. The name refers to the flattened intervals at the apex of the elytron. TYPE MATERIAL. Holotype: • &amp;male;, with labels and data: white, black printed “S- Sulawesi, 20 km NE Bantmurung 30.iii.1999, at light Becvar &amp; Zabransky (CMBB). Paratypes: • 1 &amp;male;, 1 &amp;female;, same data as holotype (CMBB); • 1 spec., N SULAWESI 1992 W Bitauna, Kotamo-bagu-Gorontalo (18) leg. Jäch 23.iv. (NHMW); • 2 specs., INDONESIA, SE-Sulawesi Rawa Aopa Nat. Park Aopa vill. 8–10.ii.1994, leg. M. Strba &amp; I. Jenis (NHMW); • 2 specs., Indonesia – SULAWESI S.O. ins. BUTON – WAKARUMBA 3–7.ii.1994 M. Štrba &amp; I. Jeniš leg. (CMBB); • 1 spec., same data but (NHMW); • 1 spec., INDONESIA, SE-Sulawesi Buton Isl. Wakarmba 3–7.ii.1994, leg. M. Strba &amp; I. Jenis (NHMW); • 2 &amp;male;&amp;male;, 3 &amp;female;&amp;female;, INDONESIEN, S-SULAWESI ENREKANG, SADANG RIVER, 46 m S 03°34.58.4” E 119°45.56.5” leg. A. PUCHNER, 27.vii.2015 / <i>trachys</i> Andr. det. DR. A. DOSTAL 2015 (CADW); • 1 spec., Toekan Besi Ins. Kalidupa (MFNB). DIAGNOSIS. Amedium sized species with smooth lateral margin of the pronotum and elytron, with isodiametric reticulation on interval eight of the elytron, and nearly right angled lobe of the mentum with slightly convex anterior margin, and with the median tooth much less projecting asthe lobe. Asimilar combination of characters is present in <i>C. agona</i>. <i>Clivina agona</i> differs from <i>C. apexplana</i> sp. nov. mainly by the size, the distinct lateral group of punctures in the basal half of the pronotum, and the acute angled lobe of the mentum. In addition, in <i>C. apexplana</i> sp. nov. the intervals of the elytron are flattened at the apex. DESCRIPTION. <i>Measurements</i>. Holotype: Body length 7.98 mm, width 2.36 mm; ratio length/width of pronotum 0.91; ratio length/width of elytra 1.86. Other material: Body length 7.32–8.45 mm (: = 7.87 mm *), width 2.23–2.57 mm (: = 2.38 mm *), ratiolength/width of pronotum 0.9–0.91 (: = 0.91*), ratiolength/widthofelytra 1.82–1.88 (: = 1.85*); (*n = 10). <i>Colour</i>: Glossy. Piceous. Apex of elytron fuscous. Intermediate and hind legs dark fuscous, antennae and tarsomeres fuscous. <i>Head</i>: Athird narrower than pronotum. Clypeus slightly excised anteriorly; wing prominent, as protruding as clypeus, separated from clypeus by distinct notch; supraantennal plate convex; clypeus laterally with carina, wing slightly reflexed margined, supraantennal plate reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by distinct notch. Clypeus slightly transverse, convex and glossy in posterior four fifths, in anterior fifth with slight isodiametric reticulation, separated from frons by distinct furrow. Frons moderately convex, with small but distinct impression at middle, with blunt lateral carina at posterior eyelevel. Clypeus and frons smooth, with minute scattered punctures, separated from wing and supraantennal plate by deep moderately wide furrow continuing posteriorly aswide supraorbital furrow with two supraorbital setigerous punctures at posterior eye level. Neck constriction developed as small step, with row of rough punctures. Eye projecting, elongated in dorsal view. Gena covering eye by about a quarter in ventral view. Antenna reaching over seta of pronotum, antennomeres not flattened, four to ten of moderate length (ratio L/ W 1.73 –1.83). Labrum like a broad flat triangle, with isodiametric reticulation, seven setose. Mentum (Fig. 50) with distinct isodiametrically reticulated surface, with lobe somewhat concavely hollowed out, shape of lobe like a skewed trapezium, with slightly convex lateral margin, with convex anterior margin and nearly rectangular lateral anterior angle, carinate laterally and anteriorly, with medial angle obtuse and indistinct, with short carina at middle, median tooth wide, somewhat less protruding anteriorly as lobe, straight anteriorly with small pointed tooth at middle. <i>Pronotum</i> (Fig. 70): Disk moderately convex in lateral and frontal view. Wider than long. Reflexed lateral margin smooth, attenuate and indistinctly convex in anterior half, widest in posterior third; anterior angle slightly projecting anteriorly; posterior angle marked by small distinct sharp tooth, tooth not protruding laterally over convexity of lateral margin. Lateral channel narrow, slightly widened posteriorly, reticulated. Median line narrow, line-like; anterior transverse line longitudinally punctured, joining median line, just joining basal constriction. Surface smooth, with some minute punctures which are denser laterally, with some transverse wrinkles and longitudinal wrinkles at middle, with distinct isodiametric reticulation in basal quarter. <i>Elytron</i>: Disk indistinctly convex in anterior third in lateral view, convex on frontal view. Nearly twice as long as wide, nearly regularly convex with maximum width at middle. Reflexed lateral margin smooth. Scutellar striole short; setigerous tubercle at base of first stria, with distinct tubercle at base of third interval. Striae moderately deep anteriorly, becoming less deep to middle and posteriorly, punctuate-striate, ground of striae with reticulation, one to three free at base, fourto seven joining at humerus, one running up to apex, joining with seven, two ending free at apex, three and four, five and six joining apically, striae somewhat widened apically. Intervals flattened apically, eights with short carina at humerus and carina in apical quarter. Third interval with four setigerous punctures, first situated at second stria near base, the other three approaching third stria. Surface of intervals glossy, interval eight with distinct isodiametric reticulation, interval seven with isodiametric reticulation at apex in prolongation of striae two to six, without reticulation at humerus. <i>Hind wings</i>: Fully developed. <i>Lower surface:</i> Proepisternum covered in lateral half with distinct isodiametric reticulation, coriaceous towards middle, with some transverse wrinkles. Terminal sternite of abdomen developed apically as narrow flat transverse fold. <i>Legs</i>: Legs covered with longitudinal reticulation. Protibia with three spines of moderate length, not sulcate dorsally, movable spur sable-like. <i>Male genitalia</i> (Fig. 97): Median lobe long, slender, moderately arcuate, distinctly arcuate at base, apex elongated, moderately broad, attenuated apically, apex slightly distorted, ventral rim distinct, base with apophysis. Endophallus at top of oroficium with band of small bristles. Ventral paramere moderately wide, slightly arcuate, lateral apophysis slender, apex moderately slender, with two long and two small setae at apex. Dorsal paramere slender, sinuate, at apex with four long setae. <i>Female coxostylus and laterotergite</i> (Fig. 114): Coxostylus slender, dorsally with thirteen long to medium sized and one distinctly elongated and robust nematiform setae in basal half, ventrally with three long setae with numerous smaller setae, with one SSO. Laterotergite with four setae of moderate length and two minute setae basally and one isolated seta towards coxostylus. VARIATION. On the frons of the head the small impression at middle is indistinct in few of the paratypes. In one male paratype from S-Sulawesi, Sadang River, the first setigerous puncture on the interval three of the elytron is located at middle of the interval. DISTRIBUTION. Known from Sulawesi including the Islands of Buton and Besi.
description: Published as part of <i>Balkenohl, Michael, 2021, Belgian Journal of Entomology, pp. 1-83 in Belgian Journal of Entomology 115</i> on pages 44-46, DOI: 10.5281/zenodo.5645953