Ipsvicia Tillyard 1919
Creators: name: Lambkin, Kevin J.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Ipsvicia

description: Ipsvicia Tillyard, 1919
Ipsvicia Tillyard, 1919a: 878.
Type species. Ipsvicia jonesi Tillyard, 1919, by original designation.
Emended diagnosis. Tegmen c. 3 times longer than wide; surface sculpture complex, comprising numerous variably shaped patches of fine tubercles each encircled by a smooth, very narrow margin, the patches set in a coarser tuberculate/punctate groundmass, a little finer apically; costal margin distinctly convex; basal cell short and broad, not extending to midpoint of PCu; R evenly convex, parallel with costal margin, the basal R cell therefore of about equal width for entire length; two r-m; CuA strongly angulate at m-cua; base of 1A slightly upcurved.
Description. Tegmen c. 14–22 mm long; marginal membrane only slightly widened postero-apically; costal fracture fairly upright; bSc variable convex; dSc very weakly developed, running along middle of costal space; R with 4–6 mostly simple post 1r-m branches; M+CuA stem a little variable in length, extending just beyond ½ length of PCu; apical section of CuA directed posteriorly, straight or slightly convex; clavus extending to c. 0.6 tegmen length; 1A about as long as PCu+1A. Hind wing: size variable in proportion to variable size of the individual, maximum length c. 13.8 mm, maximum width c. 5.9 mm in measurable specimen; R, M, CuA and CuP very closely approximated basally, CuA and CuP apparently without a common stem; R, M and CuA then more or less parallel, directed towards apical margin, gradually diverging apically, apex of CuA upcurved and forming an apical loop with R and M enclosing a small network of terminal short branches and crossveins; a long, backwardly oblique m-cua (the single cua-cup illustrated by Tillyard (1923) and Evans (1963) not detected); CuP simple, very long, sinuous, its apex broadly separated from that of CuA and PCu; both PCu and 1A with thickened bases, both straight basally and sinuous apically, CuP and PCu approximating each other at their mid-points; anal area noticeably crinkled; marginal membrane well developed.
Notes. The hind wing venation is noteworthy for the close alignment and apical looping of R, M and CuA, and the long, simple and sinuous CuP, PCu and 1A. The other known hind wings of the Scytinopteroidea are those of several species of Scytinoptera Handlirsch, 1904, illustrated by Becker-Migdisova (1948, figs. 2, 33, 39–43). They apparently lack a marginal membrane, and their venation is much simpler than that of Ipsvicia, with M and CuA directed postero-apically rather than apically, R, M and CuA without apical looping, CuA and CuP with a common stem, CuP, PCu and 1A almost straight rather than sinuous, and without the long, backwardly oblique m-cua. Indeed, as noted by Evans (1956), the venation of Ipsvicia was unlike any of the Palaeozoic or Mesozoic homopteran hind wings known at that time (see also those illustrated by Becker-Migdisova 1961), and it differs from that of the extant families of Cicadomorpha (see figures in Kramer 1950). It might be observed, however, that the form of R, M and CuA bears some resemblance to that of Heteroptera: Nepomorpha (Popov 1971, plate 52).
description: Published as part of Lambkin, Kevin J., 2020, Revision of the Ipsviciidae of the Late Triassic of Queensland (Hemiptera Cicadomorpha: Scytinopteroidea), pp. 503-520 in Zootaxa 4860 (4) on pages 506-507, DOI: 10.11646/zootaxa.4860.4.2, http://zenodo.org/record/4414254