Nyctiophylax (Paranyctiophylax) anryutah Oláh & Johanson 2010, new species
Creators: name: Oláh, János
name: Johanson, Kjell Arne
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Polycentropodidae
Nyctiophylax
Nyctiophylax anryutah

description: Nyctiophylax (Paranyctiophylax) anryutah, new species   
Figs 19–22  
 This species belongs in the Nyctiophylax (Paranyctiophylax) abaya species group and is most similar to N. archemoros Malicky, from which it differs by having long, crossing, spine-like dorsal paraproctal processes present. In N. archemoros the dorsal paraproctal processes are absent. Similar, but not crossing structures were originally described for other species of this species group as parameres of the phallic organ.   
 Male. Body uniformly pale brown. Maxillary palp formula II-I-IV-III-V, segment III originates subapically from mesal face of segment II. Forewing length 4.2 mm; brown, with hypertrophied longitudinal R vein and 2 hyaline spots: small, rounded spot below discoidal cell around crossvein r-m in subradial cell; and large triangular spot along crossvein m-cu and base of fork M; discoidal cell and median cell closed; apical forks 2, 3, 4, and 5 present; A1, A2, and A3 looping. Hind wing discoidal cell closed, median cell open, apical forks 2 and 5 present.   
Spur formula 3,4,4.  
Male genitalia. Sternite IX robust, subtriangular in lateral view, with well-developed concavity on each apicodorsal corner at fulcrum. Tergite IX indiscernible, possibly fused to paraproctal body. Paraproctal body with strongly developed dorsal and ventral paraproctal processes. Segment X membranous, weakly visible, partially covered with minute microtrichae. Cerci setose, subtriangular in lateral view; long digitate in dorsal view; shorter than gonopods and ventral paraproctal processes. Paraproctal complex well developed, highly sclerotized. Dorsal paraproctal processes forming pair of large, spine-like structures crossing dorsally, arching dorsad in lateral view; pair of large ventral paraproctal processes directed posterad, each with rounded apex and with slightly ventrad-curving subapical setose hump dorsally. Gonopods with large, quadrangular “elbow” ventrobasally, continuing into almost filiform apical half in lateral view; in ventral view directed mesad, geniculate at meeting point of large “elbow” and filiform apical half. Phallic apparatus located dorsally in genitalia, fixed, guided by paraproctal processes and segment X; tube forming sclerotized phallotheca continuing into matrix of membranous apical half. Weakly discernible phallotremal sclerites present at apical ending of phallotheca and embedded in phallic membrane.  
 Holotype male: INDONESIA (Sumatra): Bukit Barisan Selatan National Park, Barisan Mts., Way Pitias, 800 m, 17–19.vi.2008, light trap [J. Oláh, Jr.]— 1 male (OPC, in alcohol).   
 Distribution: Indonesia (Sumatra).   
 Etymology: Anryutah, from Sanskrit, meaning “crossness,” referring to the crossing dorsal paraproctal processes in the genitalia.
description: Published as part of Oláh, János & Johanson, Kjell Arne, 2010, Fifteen new Trichoptera (Insecta) species from Sumatra, Indonesia, pp. 1-35 in Zootaxa 2618 (1) on pages 14-15, DOI: 10.11646/zootaxa.2618.1.1, http://zenodo.org/record/5304087