Anemesia infuscata Zonstein 2018, sp. nov.
Creators: name: Zonstein, Sergei
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Cyrtaucheniidae
Anemesia
Anemesia infuscata

description: Anemesia infuscata sp. nov.
urn:lsid:zoobank.org:act: 84CDD144-26EB-4DFF-8BE2-F878AA5C7471 Figs 14, 26, 38–41, 45–47, 61–62, 76–77, 87–89, 103, 115–116, 131–132, 186–187, 220–223, 258– 265, 289–290, 324–329, 346–353, 365–367, 373
Diagnosis
Anemesia infuscata sp. nov. differs from A. incana in a considerably narrower clypeus and noticeably larger eyes both in males and females (Figs 61–62, 76–77; cf. Figs 59, 73). From the most similarlooking A. infumata sp. nov., males of A. infuscata sp. nov. differ by a less tapering embolus which arises closer to the prolateral side of the palpal organ (Figs 220–223; cf. Figs 217–219), while the conspecific females differ in their longer cone spermathecae ending with certainly dilated receptacles (Figs 258–265; cf. Figs 255–257).
Etymology
The specific epithet is derived from the Latin adjective ‘ infuscatus ’, meaning ‘infuscate’, referring to the entirely darkened (blackened) colouration of these spiders. The gender is feminine.
Material examined
Holotype
TAJIKISTAN: Ƌ, foothills of Hissar Mts, Varzob Canyon, 7 km N of Dushanbe, vicinity of Harangon, 1060 m, 38°40′04′′ N, 68°46′56′′ E, 27 May 2002, S. Zonstein leg. (SMNH).
Paratypes (19 ƋƋ, 37 ♀♀)
TAJIKISTAN: 1 Ƌ, 2 ♀♀, same collecting data as for the holotype (SMNH); 1 ♀, same area, environs of Harangon 5 km N of Dushanbe, 1100 m, 38°39.9′ N, 68°47.7′ E, 3 May 2015, S. Zonstein leg. (SMNH); 2 ƋƋ, 6 ♀♀, Sanglok Mts, 3–5 km W Sebiston, 1300–2200 m (38°15′ N, 69°17′ E), 2 May 1990, S. Zonstein leg. (SMNH); 3 ƋƋ, 2 ♀♀, same collection data as for preceding but 7 May 1991, S. Ovchinnikov and S. Zonstein leg. (SMNH); 2 ƋƋ, 5 ♀♀, same collection data as for preceding but 16–18 May 2002, S. Zonstein leg. (SMNH); 4 ♀♀, same collection data as for preceding but 29 Apr. 2015, S. Zonstein leg. (SMNH); 6 ƋƋ, 8 ♀♀, Hazratisho Mts, Sangdara Gorge, 1650 m, 38°22′ N, 70°09′ E, 19–25 May 2002, S. Zonstein leg. (SMNH); 5 ƋƋ, same area, Iokunj Gorge, 1600–2000 m, 38°23′ N, 70°10′ E, 23–26 May 2002, S. Zonstein leg. (SMNH); 3 ♀♀, Kuhitek Mts, 7 km NE of Hovaling, 1850 m, 38°21.6′ N, 70°03.9′ E, 28 Apr. 2015, S. Zonstein leg. (SMNH, ZMMU); 3 ♀♀, Mazar Mts, Sultanmazar 16 km NNE of Hovaling, 1850 m, 38°28.3′ N, 70°04.1′ E, 28 Apr. 2015, S. Zonstein leg. (SMNH); 3 ♀♀, same area, Darai-Mukhtor 5 km NW of Hovaling, 1600 m, 38°23.6′ N, 69°57.9′ E, 28 Apr. 2015, S. Zonstein leg. (SMNH).
Other material examined (8 ♀♀, 2 ♀♀ subad.)
TAJIKISTAN: 1 ♀, Rangentau Mts, Fahrabad Pass, 1200–1300 m, 38°20′ N, 68°43′ E, 3 Apr. 1967, E. Andreeva leg. (MIZW); 1 ♀ subad., Hazratisho Mts, 25 km E of Muminabad, 24 May 1966, E.Andreeva leg. (MIZW); 1 ♀ subad., same collection data as for preceding but 13 km NE of Muminabad, 13 Jun. 1966, E. Andreeva leg. (MIZW); 7 ♀♀, Kuhitek Mts, 1 km E of Hovaling, 1600–1700 m, 38°20′ N, 69°59′ E, 11 Oct. 1987, S. Zonstein and S. Ovchinnikov leg. (SMNH, ZMMU).
Description
Male (holotype)
HABITUS. See Fig. 14.
MEASUREMENTS. TBL 17.80, CL 7.17, CW 6.28, LL 0.80, LW 1.24, SL 3.63, SW 3.33.
COLOUR. Carapace, palps and legs predominantly intensive dark reddish brown; anterior edge and lateral margins of carapace, chelicerae and femora even darker; whole eye tubercle brownish black; sternum, labium, maxillae and leg coxae light coffee brownish coloured, metatarsi and tarsi III–IV brownish orange; abdomen yellowish brown with dark chocolate brown pattern consisting of very broad median longitudinal stripe crossed medially and posteriorly with six wide procurved fasciae, and numerous small and partially fused spots located between them; ventral abdominal surface with darker horse-shoe spot covering genital area; book-lungs paler, light yellowish brown; spinnerets even paler, light yellow.
PROSOMA. Clypeus and eye tubercle as in Fig. 61. Eye diameters and interdistances: AME 0.20(0.29), 0.30, 0.27, 0.22, AME–AME 0.23(0.13), ALE–AME 0.23(0.19), ALE–PLE 0.14, PLE–PME 0.02, PME–PME 0.49. Cheliceral rastellum composed of 15–20 stout bristles lacking tips and located in front of fang base and on low mound. Cheliceral furrow with 6 promarginal teeth and 9–10 smaller retromarginal teeth each. Sternum, labium and maxillae as shown in Fig. 103. Sternal sigilla small,
posterior pair broadly oval and located distantly from sternal margin. Each maxilla with 11–12 cuspules confined to probasal maxillary edge.
SPINATION. Palp: femur d3, pd1; patella p1; tibia d1, p3, r1, v1+ca 20 stout bristles; tarsus d5–6. Leg I: femur d4, pd2; tibia p2, v6–7+m; metatarsus v5. Leg II: femur d4, pd2, rd1; patella p1; tibia p3, v8–9; metatarsus p1, v8. Leg III: femur d4, pd3, rd3–4; patella p2, r1; tibia d1, p3, r3, v7; metatarsus pd5, p5, r3, v9–11; tarsus p3. Leg IV: femur d4, p3, r3; patella r1; tibia r3, v9; metatarsus pd1, p2, r3, v13–15; tarsus pv9–10, r2. Patella I, tarsi I and II aspinose.
LEGS. Tibia and metatarsus I as shown in Fig. 132. Scopula entire and distal on metatarsi I and II, very narrowly divided on tarsi I and II, widely divided on tarsi III, absent on tarsi IV. Trichobothria: 2 rows of 9–11 each on tibiae; 10–12 on metatarsi; 16 on tarsi I and II; 13 on tarsi III and IV, 9–10 on cymbium. PTC I–II with 7 teeth on each margin; PTC III–IV with 7 teeth on outer and 6 teeth on inner margins.
PALP. Tibia, cymbium and palpal organ as in Figs 187, 220–221, 223. Tibia cylindrical with numerous thick ventral bristles (Fig. 187). Embolus tapering, slightly curved, embolic tip with small membranous keel (Figs 220–221, 223).
SPINNERETS. See Fig. 289. PMS: length 0.51, diameter 0.22. PLS: maximal diameter 0.53; length of basal, medial and apical segments 0.90, 0.65, 0.83; total length 2.38; apical segment shortly digitiform.
LEG MEASUREMENTS. Ƌ(♀)
Female (paratype, Varzob canyon)
HABITUS. See Fig. 26.
MEASUREMENTS. TBL 22.30, CL 8.55, CW 7.32, LL 1.12, LW 1.82, SL 4.70, SW 4.11.
COLOUR. Similar to that of male, but carapace and leg more intensive reddish coloured; dark brown dorsal abdominal pattern is even better developed; ventral brownish yellow surface of abdomen with numerous dense and tiny brown spots located irregularly between last pair of book lings to spinnerets.
PROSOMA. Clypeus and eye tubercle as in Fig. 76. Eye diameters and interdistances: AME 0.21(0.30), ALE 0.38, PLE 0.28, PME 0.19, AME–AME 0.24(0.19), ALE–AME 0.25(0.20), ALE–PLE 0.21, PLE– PME 0.15, PME–PME 0.56. Cheliceral rastellum composed of numerous spikes located in front of fang base and on low mound. Each cheliceral furrow with 7–8 promarginal teeth and 9–13 smaller retrolateral teeth (as in Fig. 87). Sternum, labium and maxillae as shown in Fig. 115. Sternal sigilla small, posterior pair broadly oval and located distantly from sternal margin. Maxillae with 10–11 cuspules each.
LEGS. Scopula entire and distal on metatarsi I and II, entire on palpal tarsus, very narrowly divided on tarsi I and II, elsewhere absent. Trichobothria: 2 rows of 8–9 each on tibiae; 15–18 on metatarsi; 12–17 on tarsi. Palpal claw with 4 promarginal teeth. PTC I–II with 5–6 teeth on outer and 4–5 teeth on inner margins; PTC III with 4–5 and 2–3; PTC IV with 3–4 and 1–2 teeth, respectively.
SPINATION. Palp: femur pd1; patella p1; tibia p3, v12–14; tarsus v3. Leg I: femur pd1; tibia p3, v6–7; metatarsus v5–7. Leg II: femur pd1; patella p1; tibia p3, v8–9; metatarsus v9–10. Leg III: femur r2; patella p3; tibia p2, v7; metatarsus d1, p3, r3, v9–10; tarsus p2. Leg IV: tibia v7–8; metatarsus p4, v11–12; tarsus pv7–8. All femora with 3–5 long bristles located medially, femora III and IV also with prodorsal and retrodorsal rows of bristles, with 2–3 bristles in each row; patellae I and IV, and tarsi I and II aspinose.
SPERMATHECAE. Entire and mushroom-shaped with wide tapering stalks and rounded receptacles (Fig. 259).
SPINNERETS. See Fig. 290. PMS: length 0.79, diameter 0.48. PLS: maximal diameter 1.02; length of basal, medial and apical segments 1.52, 0.75, 0.97; total length 3.24; apical segment triangular to shortly digitiform.
Variation
The length of the carapace varies from 6.48 to 6.97 in males and from 6.89 to 9.47 in females. The colour varies narrowly from somewhat lighter and more reddish to an even darker brown than that of the holotype. The thoracic fovea are sometimes slightly recurved. Variations in shape, arrangement and structure are shown for the eyes (Figs 61–62, 76–77), the cheliceral furrow (Figs 87–89), the sternum (Figs 115–116), the male tibia and the metatarsus I (Figs 131–132), the male palpal tibia and the cymbium (Figs 186–187), the palpal organ (Figs 220–223), and the spermathecae (Figs 258–265). The number of maxillary cuspules ranges from 10 to 14.
Habitat
Anemesia infuscata sp. nov. seems to be the most mesophilic congener inhabiting shrubby and forested slopes between 1000 and 2300 m (Figs 324–329).
Distribution
Tajikistan. See Fig. 373.
Notes
Some illustrations of Anemesia infuscata sp. nov. are based on the additional paratype specimens; these include the images showing variations (see above) and photos of live spiders presented in Figs 38–41 and 45–47.
description: Published as part of Zonstein, Sergei, 2018, A revision of the spider genus Anemesia (Araneae, Cyrtaucheniidae), pp. 1-100 in European Journal of Taxonomy 485 on pages 44-47, DOI: 10.5852/ejt.2018.485, http://zenodo.org/record/3829992