Schismatothele hacaritama Valencia-Cuéllar & Perafán & Guerrero & Guadanucci 2019, sp. nov.
Creators: name: Valencia-Cuéllar, Dayana
name: Perafán, Carlos
name: Guerrero, Roberto J.
name: Guadanucci, José Paulo Leite
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Theraphosidae
Schismatothele
Schismatothele hacaritama

description: Schismatothele hacaritama Perafán, Valencia-Cuéllar & Guadanucci sp. nov.
(Figs 9–16)
Type material. COLOMBIA: Holotype male, ICN-Ar 8057, Ocaña, Norte de Santander, 1200 masl, IV-2013, F. Sánchez leg.
Etymology. The specific epithet is a noun in apposition in honor of the indigenous people Hacaritama, who inhabited the territory where this species is distributed.
Diagnosis. Males of S. hacaritama sp. nov. differ from those of other species by the shape of palpal bulb bearing several triangular teeth on the ventral face (Figs 9–11), a ventral distal keel, a pronounced keel on the distally translucent dorsal edge, and a small dorsal keel (Figs 9–12). Thick spines on the retrolateral palpal tibia are separated in two groups; the proximal group arranged on a mound in a transverse row (Fig. 15). Cymbium with a noticeable distal protrusion/bulge on retrolateral lobe (Figs 14–15). Tibial spur on I with retrolateral branch long and thick, very widened and flattened distally (Fig. 16). Females are unknown.
Description. Holotype male (ICN-Ar 8057). Color (in alcohol): carapace and legs brown, abdomen light brown; carapace bordered by numerous golden setae, abdomen and legs covered by golden setae. Total length:18.1. Chelicerae basal segment: length 2.2. Carapace elongated: length 9.8, width 7.5. Abdomen: length: 9.2. Clypeus absent. Eye tubercle slightly elevated, sub-rectangular: length 1.2, width 1.7. Anterior eye row procurved, posterior straight. Eyes and interdistances: AME 0.39, ALE 0.43, PME 0.22, PLE 0.43, AME–AME 0.14, AME–ALE 0.07, ALE–ALE 0.11, PME–PME 0.77, PME–PLE 0.05, PLE–PLE 1.21, AME–PME 0.10, ALE–PLE 0.05. Thoracic fovea procurved, deep: width 1.4. Chelicerae basal segment with 11 well-developed teeth on furrow promargin, and group of 10 small teeth on proximal area of each furrow. Intercheliceral tumescence absent. Maxillae with at least 150 cuspules, located on anterior inner corner. Labium sub-quadrate: length 0.8, width 1.7; with ca. 300 cuspules. Labiosternal junction narrow medially with two well-developed lateral mounds. Sternum oval: length 4.0, width 3.8; with three pairs of oval sigilla separated by their diameter from edge. Superior tarsal claws without teeth. Tarsal scopulae: I entire, II entire with few scattered conical setae on longitudinal midline, III narrowly divided by longitudinal band of conical setae, IV divided. Metatarsal scopulae extent: I and II almost entire length, III on distal half, IV on distal 1/4. Clavate tarsal trichobothria in two rows, each with 10 trichae, interspersed with filiform trichobothria of different sizes. Tarsus IV slightly cracked.
Palpal bulb piriform with long embolus bearing several triangular teeth on ventral face (Figs 9–11), a ventral distal keel, a pronounced keel on dorsal edge distally translucent, and a small dorsal keel (Figs 9–12). Paraembolic apophysis absent. Cymbium with two asymmetric lobes, retrolateral larger and wider, prolateral lobe elongated and laterally flattened; retrolateral lobe with noticeable distal retrolateral protrusion/bulge (Figs 14–15). Palpal tibia swollen, with seven retrolateral spines separated in two groups, one apical longitudinal row with four spines, and the other with three spines arranged in transverse row on mound (Fig. 15). Tibial spur on I (Fig. 16): prolateroventral spur with two well-developed branches, on same mound and very close together; retrolateral branch bigger, slightly curved, long and thick, very widened and flattened distally, distal edge with small subtriangular spine at distal end and internal subapical spine at proximal end; prolateral branch small, subtriangular, with a spine spine of similar size. Metatarsus I bends touching distal end of retrolateral branch.
Legs formula 4123. Palpal and legs segments lengths (femur/ patella/ tibia/ metatarsus/ tarsus/ total): palp: 6.2/ 3.0/ 4.2/ –/ 2.7/ 16.1; leg I: 8.5/ 5.0/ 5.8/ 5.9/ 4.6/ 29.8; leg II: 7.8/ 4.1/ 5.2/ 5.6/ 4.0/ 26.7; leg III: 6.3/ 3.4/ 3.6/ 5.2/ 3.4/ 21.9; leg IV: 8.5/ 4.0/ 6.0/ 8.5/ 4.0/ 31.0.
Spination (proximal to distal): cymbium and tarsi without spines. Palp: femur (d) 0, (v) 0, (p) 0-0-d1, (r) 0; patella 0; tibia (d) 0, (v) 0, (p) 0-0-1, (r) 7 megaspines. Leg I: femur (d) 0, (v) 0, (p) 0-0-d1, (r) 0; patella (d) 0, (v) 0-0-1, (p) 0-1-0, (r) 0; tibia (d) 0, (v) 1-1-0, (p) 0-1-0, (r) 1-1-0; metatarsus (d) 0, (v) 0-1-ap1, (p) 0, (r) 0. Leg II: femur (d) 0, (v) 0, (p) 0-0-d2, (r) 0; patella 0-0-1; tibia (d) 0, (v) 1-1-ap3, (p) 0-1-1, (r) 1-1-0; metatarsus (d) 0, (v) 1-0-ap1, (p) 0, (r) 1-0-ap1. Leg III: femur (d) 0, (v) 0, (p) 0-0-d1, (r) 0-0-d1; patella (d) 0, (v) 0, (p) 0-0-1, (r) 0; tibia (d) 0, (v) 1-1-ap3, (p) 0-2-0, (r) 0-1-1; metatarsus (d) 0, (v) 2-1-ap2, (p) 1-2-ap2, (r) 0-1-1,ap1. Leg IV: femur (d) 0, (v) 0, (p) 0, (r) 0-0-d1; patella 0; tibia (d) 0, (v) 1-2-ap3, (p) 0-0-1, (r) 1-0-1; metatarsus (d) 0, (v) 1-3-ap1, (p) 1-1-1,ap1, (r) 0-1-1.
Distribution and natural history. Known only from its type locality, on Colombian Eastern Cordillera, Ocaña, Norte de Santander, over 1200 masl (Fig. 37). These spiders inhabit a relatively humid environment and a mean temperature of 22 °C.
description: Published as part of Valencia-Cuéllar, Dayana, Perafán, Carlos, Guerrero, Roberto J. & Guadanucci, José Paulo Leite, 2019, Schismatothelinae spiders (Araneae, Mygalomorphae, Theraphosidae) from Colombia: four new species and an approach to their diversity, pp. 548-562 in Zootaxa 4545 (4) on pages 554-556, DOI: 10.11646/zootaxa.4545.4.6, http://zenodo.org/record/2618325