Cymbionotum (Procoscinia) fernandezi , Ball and Shpeley 2005, n. sp.
Creators: name: Ball, George E.
name: Shpeley, Danny
Publisher: name: Zenodo
Year: 2005
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Cymbionotum
Cymbionotum fernandezi

description: Cymbionotum (Procoscinia) fernandezi, Ball and Shpeley, n. sp.   
(Figs. 1B, 2A, 2E, 3F, 12A–I, 13 A–J, 14A, 15A–E, 16)  
 Type material. Thirty eight specimens (20 males and 17 females). HOLOTYPE male, card­mounted, labeled: " COLOMBIA Bolivar/ Zambrano bosque seco/ Lote Papayo cultivo/ Gmelina arborea. Luz/ 10–15. V. 1999 /Col. H.M.­M.A.­M.G./ No. 967597". A microvial, pinned beneath the whole specimen, contains its genitalia (UNCB). PARATYPES, 4 labeled same as holotype: male, point­mounted, head and prothorax missing (broken and parts lost, following for habitus picture, Fig. 1B); female, pointmounted, reproductive tract removed, in microvial pinned beneath whole specimen; female, card­mounted, with left elytron removed, and attached to card; and female, cardmounted, disassembled for SEM photographs of mouthparts and ovipositor sclerites). Eighteen males, 15 females, labeled: " COLOMBIA Bolivar Zam­/brano. Hda. Monterrey For­/estal BsT 9¼42'N, 74¼53'W/ 20m. May 1999 "; "Lote Papayo Interior de/ cultivo Gmelina arborea / trampa de luz,/ H. Moreno y M. çvila/ No. 967543" With the agreement of H. Moreno, the paratypes were distributed as follows: BMNH, 2; CASC, 2; CNIF, 7; IAVH, 7; MNHP, 2; MRAC, 2; UASM, 2; UNCB, 11; and UASM, 2.   
 Specific epithet. This Latinized eponym, genitive case, is based on the surname of Prof. Fernando Fernandez, who loaned the type specimens to us, and with whom we have cooperated in extending knowledge of Colombian Carabidae.   
 Type locality. Colombia: Bolivar: Zambrano.   
 Recognition. Compared to C. negrei, adults of C. fernandezi are on average, larger, both in length and width, with wider vertex, wider head in relation to pronotum, and wider pronotal base (Table 12). The phalli (Figs. 15D, F) of these two species are similar in form, but that of C. fernandezi (Fig. 15B) is broader, with the preapical portion of the shaft bent more gradually ventrad, and ventral surface generally dark (cf. Fig. 15F).   
 Description. With recognition features as noted above, and other details specified in the subgeneric description, and illustrations, as follows: habitus, Fig. 1B; integumentary setal length, Fig. 2E; density of elytral punctation, Fig. 3F; labrum and epipharynx, Figs. 12I and J, respectively; mandibles, Figs. 12A–H; maxillae, Figs. 12K–L; labium, Figs. 13A–C; pronotum, dorsal aspect, Fig. 14A; hind wings, Fig. 15A; middle tibia and tarsomere 5, Figs. 13D and E, respectively; phallus and endophallic armature, Fig. 15B; gonocoxa of ovipositor, Figs. 13F–J; internal genitalia of female, Fig. 15E.   
 Habitat, habits, and seasonal occurrence. All specimens were collected in May, at an ultraviolet light, within a plantation of an exotic tree ( Gmelina arborea Roxb., white teak, family Verbenaceae), a species indigenous in the Oriental Region and southern China. We infer that C. fernandezi is mesophilic, occupying disturbed areas in the tropical rain forest zone (Table 17, Fig 24), at less than 200 m altitude with adults active in midyear.   
 Geographical distribution (Fig. 16). This species is known only from the Rio Magdalena valley, Colombia.   
 Relationships and chorological affinities. Based on shared morphological features, geographical distribution and allopatric ranges, C. fernandezi and C. negrei are postulated to be adelphotaxa.   
 Material examined. We have seen the type series, only. For details, see " Type material", above.
description: Published as part of Ball, George E. & Shpeley, Danny, 2005, Taxonomic review of the Tribe Melaenini (Coleoptera: Carabidae), with observations on morphological, ecological and chorological evolution, pp. 1-120 in Zootaxa 1099 (1) on pages 1-120, DOI: 10.11646/zootaxa.1099.1.1, http://zenodo.org/record/5051674