Paracymbiomma Rodrigues & Cizauskas & Rheims 2018, gen. nov.
Creators: name: Rodrigues, Bruno V. B.
name: Cizauskas, Igor
name: Rheims, Cristina A.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Gnaphosidae
Paracymbiomma

description: Paracymbiomma gen. nov.
Etymology. The generic name is a contraction of paracymbium and Lygromma, since the paracymbium is one of the diagnostic structures of the genus. Gender is neuter.
Type species. Paracymbiomma angelim (original designation).
Diagnosis. Species of Paracymbiomma gen.nov. are distinguished from those of other Prodidomidae genera by the combination of the following characters: male palp with paracymbium and RTA single (Figs 45, 54, 61), embolus long and filiform with curved tip (Figs 43, 47, 49, 61), conductor sclerotized (Figs 53, 55, 57) and median apophysis membranous with sclerotized tip (Figs 53, 49, 61, 69–74); female epigyne with posteromedian atrium and sclerotized margins (Figs. 51, 59, 63, 67), vulva with copulatory ducts long, strongly convoluted laterally with proximal part (closest to copulatory openings) translucent (Figs 52, 68); secondary spermathecae elongated (Figs 60, 64).
Description. Total length males 1.86–3.9 and females 2.25–4.15. Carapace and legs pale yellow, and abdomen weakly pigmented. Carapace longer than wide, narrower at cephalic region (Fig. 7). Fovea present as a narrow dark longitudinal line on posterior half of carapace (Figs 1–7). Eyes variable in number and size (Figs 13–18). Chilum present, entire (Fig. 8). Chelicerae relatively long (0.27–0.9) with a convex boss (Fig. 8); promargin with 1–4 teeth, retromargin with 3–6 teeth; one large promarginal escort setae at base of fang (Fig. 31); comb of whisker setae at cheliceral promargin near the base of fang on all females collected in caves (Fig. 20). Endites slightly convergent anteriorly (Fig. 9), with dense scopulae on internal margin; serrula present in single row (Fig. 21); labium longer than wide (Fig. 9). Sternum shield shaped, anteriorly straight, longer than wide, rebordered anteriorly and laterally; posterior region strongly protruding between coxae IV with numerous long and erect setae (Figs 10, 22, 33); precoxal triangles present (Fig. 10). Pedicel with ventral sclerite triangular. Leg formula 4123. Spination variable. All tarsi scopulate (Figs 23, 25). Trichobothria present on dorsal tibia, metatarsi and tarsi, arranged in two rows on tarsi (Fig. 25) and one on metatarsi (Fig. 34); bothrium with 5–9 deep ridges (Fig. 26). Tarsal organ capsulate with oval opening, located dorsally on tarsi. Two pectinate claws; tarsi I–II with long toothed claws, with 6–13 teeth (Figs. 27, 35) tarsi III–IV with short toothed claws, with 3–4 teeth (Fig. 36). Claw tufts composed of slightly widened tenent setae (Fig. 27); presence of classic claw tuft clasper (Figs 27, 35, 36) Abdomen oval, longer than wide, covered with sheet-shaped translucent scales (Figs 29, 30); sparse tactile hairs with several barbs present; dorsum of abdomen anteriorly with long, dark, posteriorly curved setae; postepigastric invaginations present (Figs 12, 37). Six spinnerets (Figs 11, 38, 39); ALS elongated (0.6–0.13), separated from each other by one diameter or more (Fig. 11), piriform gland spigots elongated with few associated setae (Fig. 40), male with one major ampullate gland spigot (Fig. 40) and female with two; PLS and PMS conical and short (Figs. 41, 42), PLS larger than PMS (Figs 11, 38); PLS in male with one minor ampular gland spigot and four aciniform gland spigots (Fig. 41), females apparently with one minor ampular gland spigot, five aciniform gland spigots and two cylindrical gland spigots; PMS in male with one minor ampular gland spigot and two aciniform gland spigots, females apparently with one minor ampular gland spigot, four aciniform gland spigots and two cylindrical gland spigots (Fig. 42). Female pedipalp with one pectinate claw with 6–8 teeth (Fig. 28). Male palp: tibia generally short, except in P. caecus sp. nov., in which it is more elongated (Figs 65, 66, 74), RTA single, directed apically (Figs 54, 56); cymbium oval with apical scopula (Figs 62, 66), paracymbium present (Figs 43, 45, 54, 61); bulb rounded (Figs 53, 55, 69–72) or ovoid (Figs 61, 65, 73, 74); embolus long with curved tip and wide base, inserted between 9–12 o’clock position (Figs 55, 71), or 3 o’clock position (Figs 61, 74); conductor sclerotized, with curved tip, inserted
prolaterally (Figs 47, 55, 70); median apopohysis inserted distally and with sclerotized tip(Figs 69–74). Epigyne: posterior margin of epigynal plate anteriorly curved (Figs 51, 67) or truncate (Fig. 59); conspicuous posteromedian atrium with well-defined margins (Figs 59, 63). Vulva: secondary spermathecae elongated (Figs 60, 64); fertilization ducts antero-lateral.
Distribution. Brazil: states of Pará (exclusively in ferruginous caves), Paraíba, Pernambuco, Bahia and São Paulo (Fig. 75).
Composition. Six species: Paracymbiomma angelim sp. nov.; P. doisirmaos sp. nov.; P. pauferrense sp. nov.; P. carajas sp. nov.; P. bocaina sp. nov.; P. caecus sp. nov. (Figs 1–6).
description: Published as part of Rodrigues, Bruno V. B., Cizauskas, Igor & Rheims, Cristina A., 2018, Description of Paracymbiomma gen. nov., a new genus of prodidomid spiders from the Neotropical region (Araneae: Prodidomidae) including a new troglobite species, pp. 301-331 in Zootaxa 4514 (3) on pages 303-304, DOI: 10.11646/zootaxa.4514.3.1, http://zenodo.org/record/2608011