Empis (Coptophlebia) hyalea Melander 1946
Creators: name: Daugeron, Christophe
name: Grootaert, Patrick
Publisher: name: Zenodo
Year: 2005
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Empididae
Empis
Empis hyalea

description: EMPIS (COPTOPHLEBIA) HYALEA CLADE   
 Monophyly: The monophyly of this clade is supported by one character: male fore tibia with at least one characteristic anterodorsal bristle on apical half [character 7(1)].   
 Distribution (Fig. 23): This group of species is typically South-east Asian, recorded from China   
(Kwantung), Indonesia (Java), Myanmar, Singapore, Thailand and Vietnam.  
 Remarks: Even if E. (C.) leptargyra Frey is not included in the phylogenetic analysis and not taxonomically reviewed, a preliminary morphological study of this species allows inclusion in the E. (C.) hyalea clade as well as in the key given below.   
 EMPIS (COPTOPHLEBIA) HYALEA MELANDER, 1946   
(FIG. 17)  
 References: Melander (1946): 113, fig. 6.   
 Type material studied: Holotype: male, China, east Kwantung, Tsin Leong San, 5.vi.1936, J.L.Gressit (according to Melander, 1946: 114), A.L.Melander Collection 1961 (USNM). The red label ‘ Holotype, Coptophlebia (= Empis) hyalea Melander, male, teste   
 KEY TO SPECIES OF THE EMPIS (COPTOPHLEBIA) ATRATATA CLADE (MALES ONLY)   
 1. Wing brown, epandrium with 4 strong apical bristles............................................. E. (C.) velutinella Frey (Myanmar) Wing clear, epandrium with 1 strong apical bristle (Daugeron et al., 2002: figs 1, 2)...................................................2   
 2. Black species; hind tibia and tarsus covered with numerous long bristles especially dorsally; hind tarsomeres strongly swollen................................................................................................. E. (C.) atratata Daugeron & Grootaert (Thailand) Brown species; hind tibia and tarsus only with shorter bristles dorsally; hind tarsomeres slender.................................................................................................................................. E. (C.) kosametensis Daugeron & Grootaert (Thailand)  KEY TO SPECIES OF THE EMPIS (COPTOPHLEBIA) HYALEA CLADE (MALES ONLY) 1. Third flagellomere distinctly aristiform.......................................................................................................................... 2 Third flagellomere shorter, styliform..................................................................  E. (C.) inferiseta sp. nov.  (Vietnam) 2. Fore tibia with 1 distinct to spine-like dorsal or anterodorsal bristle near middle, sometimes with an additional distinct dorsal or anterodorsal bristle on apical half.......................................................................................................... 3 Fore tibia with 2 spine-like anterodorsal bristles near middle, or with a single regular dorsal or anterodorsal row of bristles.............................................................................................................................................................................. 103. Cercus without a minute posteroventral projection apically......................................................................................... 4 Cercus with a minute posteroventral rounded projection apically (Figs 19B, 20; Daugeron & Grootaert, 2003a: fig. 12)................................................................................................................................................................................. 84. Fore tibia not distinctly deformed, somewhat S-shaped in frontal view...................................................................... 5 Fore tibia distinctly deformed.......................................................................................................................................... 7 5. Wing brown................................................................................................. E. (C.) thiasotes Melander (Indonesia: Java) Wing hyaline...................................................................................................................................................................... 6 6. First and basal half of second mid tarsomeres yellowish to whitish......................... E. (C.) hyalea Melander (China) Mid tarsus brownish.................................................................................................. E. (C.) leptargyra Frey (Myanmar) 7. Labrum rather short, as long as head height, dorsal bristle on fore tibia spine-like, wing clear ......................................................................................... E. (C.) pseudospinotibialis Daugeron & Grootaert (Thailand) Labrum distinctly longer than head height, dorsal bristle on fore tibia not spine-like, wing rather brownish.........................................................................................................................  E. (C.) vitisalutatoris sp. nov.  (Singapore) 8. Last four fore tarsomeres strongly modified, considerably swollen (Fig. 19A).... E. (C.) ostentator Melander (China) Last four fore tarsomeres not modified or only somewhat swollen............................................................................... 9 9. Fore tibia only narrowed at base, with 1 distinct anterodorsal bristle at middle and a second at apical quarter, mid tarsus swollen, halter blackish.................................................................................... E. (C.) sinensis Melander (China) Fore tibia distinctly deformed, with 1 anterodorsal spine at middle and a second less strong, shorter at apical quarter, mid tarsus not swollen, halter pale ................................... E. (C.) spinotibialis Daugeron & Grootaert (Thailand) 10. Fore tibia with a single regular dorsal row of bristles longer than tibia depth, epandrium with a very long ventral bristle at base (Daugeron & Grootaert, 2003a: fig. 3)............... E. (C.) lamruensis Daugeron & Grootaert (Thailand) Fore tibia with spine-like anterodorsal bristles not distinctly longer than tibia depth, epandrium without a very long ventral bristle at base..................................................................................................................................................... 11 11. Last four fore tarsomeres swollen, somewhat rounded, fore tibia with an anterodorsal row of c. 5–6 strong bristles apically ending in 2 spines bristles, epandrium with c. 10 strong bristles at tip (Daugeron & Grootaert, 2003a: fig. 8).................................................................................................. E. (C.) pakensis Daugeron & Grootaert (Thailand) Last four fore tarsomeres slender, fore tibia with an anterodorsal row of 8–10 short spine-like bristles not apically ending in 2 spines; epandrium with c. 4 strong bristles at tip (Daugeron & Grootaert, 2003a: fig. 14)............................................................................................................................... E. (C.) thapensis Daugeron & Grootaert (Thailand)  
W.J. Turner, 1981’ was added to the holotype. Paratypes: 2 males, same data but 1.vi.1936; 1 female, same data but 7.vi.1936 (USNM).
description: Published as part of Daugeron, Christophe & Grootaert, Patrick, 2005, Phylogenetic systematics of the Empis (Coptophlebia) hyalea-group (Insecta: Diptera: Empididae), pp. 339-391 in Zoological Journal of the Linnean Society 145 (3) on pages 361-362, DOI: 10.1111/j.1096-3642.2005.00189.x, http://zenodo.org/record/5435260