Mallinella tuberculata Dankittipakul & Jocqué & Singtripop 2012, sp. nov.
Creators: name: Dankittipakul, Pakawin
name: Jocqué, Rudy
name: Singtripop, Tippawan
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Zodariidae
Mallinella
Mallinella tuberculata

description: Mallinella tuberculata sp. nov.   
(Figs 104–105, 112–113, 115, 117–138)  
 Material examined. Holotype: ♂,  THAILAND, Chiang Mai Province:  Chomthong District, Doi Inthanon NP, Doi Inthanon, dry deciduous dipterocarp forest, 510 m, pitfall trap, 25 April – 23 May 2000, leg. P. Dankittipakul (MHNG, TH–4A). Paratypes: 18♂, 10♀, same data as holotype (MHNG TH–4A, TNHM).  THAILAND, Chiang Mai Province  : 1♂, Chiang Mai City, Doi Suthep-Pui NP, Doi Kham, dry deciduous dipterocarp forest, 310 m, leaf litter samples, 20 August 2002, leg. M. & P. Dankittipakul (MHNG, TH–4B); 1♀, Doi Suthep-Pui NP, Mae Hia, southwest of Chiang Mai city, dry deciduous dipterocarp forest, 350 m, 23–30 June 1987 (MHNG, TH–4C); 1♀, 16–23 June 1987 (MHNG); 1♀, 7–13 July 1987 (MHNG, TH–4D); 2♂, 1♀, 9–16 June 1987 (THNM); 1♂, 2–9 June 1987 (MHNG); 1♂, 19–30 July 1987 (MHNG); 2♀, 19 October to 19 November 1986 (MHNG); 1♀, 14–19 July 1987 (TNHM); 1♀, 1–8 July1987 (MHNG); 1♂, 16–23 June1987 (TNHM).  Lamphun Province:  1♂, Doi Khunthan NP, 1,100 m, 24 September 1990 (TNHM). All latter leg. P.J. Schwendinger.   
 Diagnosis. Males of M. tuberculata sp. nov. can be distinguished from all known species of the tuberculata - group by a complicate apex of the embolus (Fig. 134); they are similar to those of M. atromarginata sp. nov. in having an oblique mesal ridge and a triangular mesal tooth on the TA. Females can be separated from those of M. atromarginata sp. nov. by the shape of the spermathecae which are slightly excavated on their outer margins in anterior view (Fig. 136).   
 Etymology. The specific epithet (tuberculata, an adjective derived of the Latin noun tuberculum = small boss) refers to the dorsal tubercle on the RTA.   
 Description. Male  (holotype). Total length 7.37; prosoma 3.63 long, 2.97 wide; opisthosoma 3.74 long, 2.75 wide. Eye sizes and interdistances: AME 0.20, ALE 0.16, PME 0.16, PLE 0.15, AME–AME 0.12, AME–ALE 0.14, PME–PME 0.16, PME–PLE 0.26; MOQ: 0.50 long, 0.38 anterior width, 0.44 posterior width. Leg measurements: I 10.01 (2.75, 3.08, 2.20, 1.98), II 9.79 (2.53, 3.08, 2.42, 1.76), III 9.9 (2.64, 2.97, 2.64, 1.65), IV 11.79 (2.75, 3.85, 3.63, 1.56).   
 Coloration (Fig. 104). Carapace reddish brown, slightly darker along margin. Chelicerae brown. Sternum yellowish. Legs bi-colored: coxae, trochanters and proximal part of femora whitish, distal part of femora and other leg segments yellowish brown. Dorsum of opisthosoma dark sepia. Dorsal pattern: first pair represented by paired round spots situated close together; second pair by transverse median band; from third pair onwards fused, forming pale folium-like patch. Dorsal scutum longitudinal, edge sharply defined.   
 Palp (Figs 120–121, 130–134). RTA broad at base, subterminally abruptly tapered, apex bluntly pointed; dorsal tubercle blunt. Cymbial fold slightly less than half-length of cymbium. Tegular spine triangular, sharply pointed. Membranous area on tegulum with digitiform mesal hump. TA terminally with blade-like apico-prolateral process; apico-prolateral fold triangular, short, apex blunt; apico-retrolateral arm strongly pronounced, directed dorsad; oblique mesal ridge slightly elevated, situated anterior to small, sharply pointed mesal tooth. Embolus originating at 270°, with deep longitudinal groove over entire length; mesal ridge translucent, terminally forming semitransparent triangular flange; lateral ridge with sharply pointed subapical spike and terminally blunt apical process in ventral view.   
 Female (paratype, MHNG TH–4A). Total length 8.8; prosoma 4.18 long, 2.75 wide; opisthosoma 4.62 long, 3.03 wide. Eye sizes and interdistances: AME 0.18, ALE 0.15, PME 0.15, PLE 0.15, AME–AME 0.14, AME–ALE 0.26, PME–PME 0.18, PME–PLE 0.36; MOQ: 0.54 long, 0.48 anterior width, 0.50 posterior width. Leg measurements: I 9.13 (2.64, 2.75, 2.31, 1.43), II 8.107 (2.59, 2.75, 1.65, 1.11), III 9.17 (2.66, 2.92, 2.15, 1.43), IV 9.83 (2.75, 2.83, 2.64, 1.60).   
 Coloration (Fig. 105). Carapace orange-brown anteriorly, yellowish brown posteriorly. Chelicerae brown. Sternum yellowish. Legs yellowish. Dorsum of opisthosoma dark sepia. Dorsal pattern: as in male.   
 Genitalia (Figs 115, 117–119, 135–138). Epigynal plate V-shaped, anterior median incision deep, almost reaching posterior margin. Insemination ducts short and divergent, triangular in anterior view. Spermathecae tubiform, rounded terminally, outer margin slightly excavated.   
 Natural history. Most of the type specimens were collected from dry dipterocarp forests at low altitude. Only one male was collected from a forest located above 1,000 m but extension of dry deciduous forest to higher slope is common at Doi Khunthan NP.   
 Distribution. Northern Thailand, Provinces of Chiang Mai and Lamphun.
description: Published as part of Dankittipakul, Pakawin, Jocqué, Rudy & Singtripop, Tippawan, 2012, Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from Southeast Asia (Araneae, Zodariidae) 3369, pp. 1-327 in Zootaxa 3369 (1) on pages 33-37, DOI: 10.11646/zootaxa.3369.1.1, http://zenodo.org/record/5252563