Ceratophila (Ceratophila)
Creators: name: Tang, William
name: Skelley, Paul
name: Pérez-Farrera, Miguel Angel
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Cycadopsida
Cycadales
Zamiaceae
Ceratophila

description: Key to adult Ceratophila
Because of variable and overlapping morphological character states with some species, it is desirable, and in some cases necessary, to have a series of specimens representing color variations and both sexes to properly identify a
species. In the following key, text in brackets before the species name are the Mexican states of occurrence and the known host species of Ceratozamia.
1. Pronotum not explanate laterally, surface convexly curved to lateral carinae; lateral carinae in lateral view distinctly thicker anteriorly, anterior thickness 2× posterior thickness (Fig. 1D); pronotal disc lacking longitudinal groove extending anteriorly from lateral basal pore in margin (Fig. 1B); metatibia triangularly dilated toward apex (Fig. 1F)......................................................................................... [subgenus Ceratophila, new subgenus] ... 2
- Pronotum explanate laterally, surface broadly concave near lateral carinae; lateral carina weakly thickening anteriorly, anterior thickness <1.5× that at base (Fig. 1E); pronotal disc with longitudinal groove extending anteriorly from lateral basal pore in margin (Fig. 6D); metatibia not triangularly dilated toward apex (Fig. 1G)............... [Vovidesa, new subgenus] … 5
2(1). Head with broad medial transverse ridge anterior of a distinct transverse basal groove in males (Fig. 2D), ridge and groove weak in females, ridge interrupting supraocular line in both male and female; head broad, width = 0.74–0.75× pronotal width; anterior clypeal margin emarginate; pronotal hypomeron densely punctate (Fig. 2E); [Veracruz— C. euryphyllidia].................................................................................... C. (C.) chemnicki, new species
- Base of head at most with indistinct shallow transverse groove, supraocular line complete (Fig. 3D); head narrower, width = 0.66–0.68× pronotal width; anterior clypeal margin truncate or weakly convex; pronotal hypomeron with punctures sparse and minute or lacking (Fig. 3E).............................................................................. 3
3(2). Dark elytral markings (when present) laterally covering most of disc (Fig. 4A); pronotum generally more elongate, pronotal length/pronotal width (PL/PW) = 0.83–0.90; [Chiapas— C. alvarezii, C. mirandae, C. norstogii, C. vovidesii]......................................................................................... C. (C.) picipennis, new species
- Dark elytral markings (when present) only along suture and laterally (Fig. 5A); pronotum generally shorter, PL/PW = 0.80– 0.84................................................................................................ 4
4(3). Color of pronotum and elytra similar (Fig. 5A); pronotum generally more quadrate; male genitalia with penile struts relatively long, length of penile strut relative to median lobe 2:1; setae at apex of parameres with greatest length <dorsal width of parameres (Fig. 5F); [Veracruz— C. tenuis]......................................... C. (C.) sanchezae, new species
- Pronotum reddish brown contrasting with light brown of elytra (Fig. 3A); pronotum generally more trapezoidal, narrowing anteriorly; male genitalia with penile struts shorter, length of penile struts relative to median lobe 3:2; setae at apex of parameres with greatest length ± dorsal width of parameres (Fig. 3H); [Oaxaca— C. mixeorum]...................................................................................................... C. (C.) gregoryi, new species
5(1). Male meso- and metatibiae each with medial subapical emargination (Fig. 1G); female abdominal segments weakly alutaceous, punctures and setae visible except on lateral quarters.................................................... 6
- Male tibiae lacking medial subapical emargination; female abdominal segments strongly alutaceous (Fig. 6B), punctures obscured across middle, setae reduced [Chiapas— C. alvarezii, C. mirandae, C. norstogi, C. vovidesii]............................................................................................... C. (V.) chiapensis, new species
6(5). Submentum of male and female with sparse patch of setae-bearing punctures, male with long setae projecting anteriorly (Fig. 7E); male genitalia more dorsoventrally compressed, median lobe nearly cylindrical; [Veracruz— C. mixeorum]........................................................................................ C. (V.) mixeorum, new species
- Submentum of male and female with dense patch of setae-bearing punctures, male with long setae projecting laterally (Figs. 8 E–F); male genitalia laterally compressed, median lobe laterally flattened and distinctly curved in lateral view; [Veracruz— C. tenuis]....................................................................... C. (V.) vazquezi, new species
description: Published as part of Tang, William, Skelley, Paul & Pérez-Farrera, Miguel Angel, 2018, Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae), pp. 151-178 in Zootaxa 4508 (2) on pages 155-156, DOI: 10.11646/zootaxa.4508.2.1, http://zenodo.org/record/2606934