Neoperrona zoltanorum Harzhauser & Landau & Janssen 2022, nov. sp.
Creators: name: Harzhauser, Mathias
name: Landau, Bernard
name: Janssen, Ronald
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Neogastropoda
Clavatulidae
Neoperrona
Neoperrona zoltanorum

description: Neoperrona zoltanorum nov. sp.   
 Figs 30D 1 –D 3, E 1 –E 3, 4H, 5, 7   
 Perrona taurinensis (Bellardi, 1877) — Kovács & Vicián 2021: 146, pl. 3, figs 19–20 [non Neoperrona taurinensis (Bellardi, 1877)].   
 Type material. Holotype: NHMW 2019 /0151/0108a, SL: 51.9 mm, MD: 19.0 mm, Letkés (Hungary), figs 30D 1 – D 2, 4H. Paratype: NHMW 2019 /0151/0108b, SL: 49.2 mm, MD: 19.6 mm, Letkés (Hungary), figs 30E 1 –E 2. SMF 351871 /4, Letkés (Hungary).   
 Type stratum. Fossil-rich marly sand with coral blocks and andesite boulders of the Sámsonháza Formation (Császár 1997).   
 Type locality. Letkés at the western part of the Börzsöny Mts. (Hungary); see Kovács & Vicián (2013) for a map and description.   
 Age. Middle Miocene, early Badenian (= Langhian).   
 Etymology. In honor of Zoltán Kovács and Zoltán Vicián (Budapest), in recognition for their contributions on the mollusc fauna from Letkés.   
 Diagnosis. Neoperrona species of moderately large size, solid, moderately slender fusiform, with high spire, early whorls with smooth subsutural cord, canaliculated mid-portion, beaded suprasutural cord. Later whorls with broad collar, narrow ramp, tubercular shoulder, relatively strong spiral sculpture at mid-whorl and base on last whorl. Moderately long siphonal canal.   
 Description. Shell moderately large, solid, moderately slender fusiform with high spire; apical angle ~35°. Protoconch not preserved. Teleoconch of at least 11 whorls. Early teleoconch whorls flat-sided with tripartite sculpture. Weak, faintly beaded subsutural spiral cord, beaded suprasutural cord forming periphery, separated by a canaliculated mid-portion. On fourth to fifth whorl subsutural cord becomes swollen to form broad, smooth, rounded, sharply delimited collar. Central cord remains narrow, sharply depressed, almost canaliculate. Suprasutural beads weaken on sixth whorl leaving prominent broad, smooth to vaguely tubercular cord above suture. Suture narrowly impressed, linear. Last whorl about ~50% of total height; with broad subsutural collar. Subsutural ramp narrow, concave. Shoulder delimited by row of axially elongated, opisthocline tubercles, convex below, moderately strongly constricted at base. Shoulder weak, siphonal fasciole swollen, strongly twisted, with prominent growth increments. Sculpture of relatively prominent irregular spirals below shoulder, most strongly developed at mid-whorl and over fasciole, weaker over base. Aperture moderately wide, ovoid. Outer lip thin, smooth within. Anal sinus deep, moderately wide, asymmetrically U-shaped, with apex mid-ramp; siphonal canal moderately short. Columellar and parietal callus thickened, sharply delimited, forming broad callus rim and narrow pseudumbilical chink.   
 Discussion. The sculpture of the early teleoconch whorls excludes a placement in Perrona as suggested by Kovács & Vicián (2021), who identified this species as Perrona taurinensis (Bellardi, 1877), a species originally described from the Langhian of Villa Forzano in the Colli Torinesi (Italy).The Italian species differs in its prominently beaded suprasutural spiral cord on the early whorls and the less protruding shoulder (see Bellardi 1877: 187, pl. 6, fig. 10). ‘ Perrona’ taurinensis sensu Zunino & Pavia, 2009 (non Bellardi, 1877) develops spiral cords in the concave interspace of the spire whorls and lacks nodes or tubercles along the shoulder (see Zunino & Pavia 2009: 359, pl. 2, fig. 5). ‘ Perrona’ taurinensis Bellardi, 1877, based on the specimens from the Serravallian of Turkey illustrated in Landau et al. (2013), differs from N. zoltanorum nov. sp. in its shorter spire, the wider apical angle, and the weaker concavity of the spire whorls (see Landau et al. 2013: 291, pl. 50, fig. 1). Note that the name Pleurotoma taurinensis was proposed by Karl Mayer (1873) in a letter to Bellardi. The first published description, however, was provided by Bellardi (1877: 187). Contrary to Ferrero Mortara et al. (1981), the authorship has to be passed to Bellardi (1877) and not to Mayer (1873). Due to this misunderstanding, Ferrero Mortara et al. (1981) did not identify the specimen illustrated by Bellardi (1877) as syntype of Neoperrona taurinensis. ‘ Clavatula ’ inedita (Bellardi, 1877), from the Burdigalian of Italy and the Serravallian of Turkey, has a similar profile of the last whorl, but differs in its high last whorl and longer siphonal canal (see Ferrero Mortara et al. 1981, 80, pl. 15, fig. 8; Landau et al. 2013: 291, pl. 49, figs 10–11).   
 Paleoenvironment. The occurrence at Letkés (Hungary) suggests a preference for inner neritic environments in the vicinity of corals (Kovács & Vicián 2013).   
 Distribution in Central Paratethys. Badenian (middle Miocene): Pannonian Basin: Letkés (Hungary) (Strausz 1966; Kovács & Vicián 2021), Bükk Mountains: Borsodbóta (Hungary) (Csepreghy-Meznerics 1972).
description: Published as part of Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, pp. 1-172 in Zootaxa 5123 (1) on pages 99-100, DOI: 10.11646/zootaxa.5123.1.1, http://zenodo.org/record/6399827