Glyphiulus basalis Golovatch & Geoffroy & Mauriès & Spiegel 2007, n. sp.
Creators: name: Golovatch, Sergei I.
name: Geoffroy, Jean-Jacques
name: Mauriès, Jean-Paul
name: Spiegel, Didier Van Den
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Spirostreptida
Cambalopsidae
Glyphiulus
Glyphiulus basalis

description: Glyphiulus basalis n. sp.   
(Figs 27-29)  
 TYPE MATERIAL. — China. Sichuan Prov., Xin Long County, Gan Chuan Cave, Nos 414, 417 and 421, 6.VIII.1999, leg. J. & B. Lips, holotype ♂ (MNHN GA 043); paratypes 1 ♂, 3 ♀♀ (MNHN GA 043); 1 ♀ (ZMUM); 1 ♀ (ZMUC); 1 ♀ (SEM).   
ETYMOLOGY. — To emphasize the presumed basal position of this species within the group.  
DIAGNOSIS. — Differs primarily by the retained telopodites of male legs 1, coupled with the large size, the crests on the collum and the spines at claw base both strongly reduced, the crests on metaterga reduced but the ozoporiferous tubercles high, the troglomorphic traits (complete depigmentation, the lack of ocelli, the evidently elongated antennae and legs), the low, but evident, transverse pleural ridge behind the gonopod opening on male segment 7.  
DESCRIPTION  
Length of ♂, 30-32 mm, width 1.8 mm; of ♀, 23-42 and 1.5-2.2 mm, respectively. Length of holotype about 30 mm, width 1.3 mm. Coloration usually uniformly yellowish, sometimes dark red-brown due to heavy sclerotization.  
Holotype with 48p+3a+T; ♂ paratype with 45p+4+T; ♀ paratypes with 40-59p+4-1a+T.  
Antennae much more elongate than usual (Figs 27A; 29A). Ocelli absent. Gnathochilarium with (n = 1) or without (n = 2) division of mentum (Fig. 29B).  
Crests on collum, dorsad of a well-developed lateral carina, nearly completely obliterated, pattern visible only as (4)5+m+5(4) extremely faint undulations at caudal margin (Fig. 27A, B). Carinotaxy of subsequent metaterga typical (Fig. 27D; 28D), but only two crests present below rather strikingly elongated ozoporiferous tubercles; all other crests distinctly underdeveloped (Figs 27A, D; 28A, C). From segment 7 or 8 on, median crest clearly doubled anteriorly (Fig. 27D). Midbody segments round in crosssection (Fig. 28C). A low, but evident, transverse pleural ridge present behind gonopod opening on male segment 7. Epiproct narrower caudally, hypoproct more narrowly emarginate at caudal margin (Fig. 28B).  
Legs elongate (Figs 27C; 28C; 29F), longer than midbody height; claws strongly elongate, with a very small spine at base (Fig. 28D). Male legs 1 unusual in being fully leg-like, only very modestly reduced:a coxa, a 3-segmented telopodite and a claw supplied with a spine at base are all very clearly expressed (Fig. 29C); male legs 2 and 3 typical (Fig. 29D, E), but penes devoid of strong setae distolaterally.  
Anterior gonopods with evident, subunciform protuberances at median corners of coxosternum (Fig. 29G). Posterior gonopods with a flagelliform and micropilose distal process (Fig. 29H).  
REMARKS  
Based on morphological evidence alone, this species seems to be a troglobite.
description: Published as part of Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group, pp. 7-49 in Zoosystema 29 (1) on pages 39-40, DOI: 10.5281/zenodo.5392079