Glyphiulus percostulifer Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel 2007, n. sp.
Creators: name: Golovatch, Sergei I.
name: Geoffroy, Jean-Jacques
name: Mauriès, Jean-Paul
name: Spiegel, Didier Van Den
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Spirostreptida
Cambalopsidae
Glyphiulus
Glyphiulus percostulifer

description: <i>Glyphiulus percostulifer</i> n. sp. (Figs 20; 21) TYPE MATERIAL. — <b>Laos.</b> Khammouan Prov., Ban Nakok (= Nakhok), Tham Thê, 104.49°E, 17.90°N, cave, hand collection, 11.II.1998, leg. A. Bedos &amp; L. Deharveng (LAO-016), holotype &amp;male; (MNHN GA 051); paratypes 1 &amp;male;, 1 &amp;female;, 1 subad. &amp;male; (MNHN GA 051); 1 &amp;female; (ZMUM / SEM). ETYMOLOGY. — To emphasize the very close resemblance to <i>G. costulifer</i> n. sp., but with more strongly developed tergal crests. DIAGNOSIS. — Differs from congeners in the small size, the completely unpigmented tegument and ocelli, coupled with certain details of leg and gonopod structure (see also key below). DESCRIPTION Holotype with 34p+2a+T, about 14 mm long and 1.0 mm wide; adult paratypes with 36p+2a+T, about 14 mm long and 1.0 mm wide (&amp;male;), with 40p+1a+T or 38p+1a-T, about 17 mm long and 1.0 mm wide (&amp;female;); subadult &amp;male; with 31p+2a+T. Coloration uniformly pallid, ocelli not visible (Fig. 20A). Postcollar constriction, dorsal crests and carinotaxy pattern on postcollar terga (Fig. 20 A-E), segment’s texture and cross-section (Fig. 20 B-D), paraprocts and hypoproct (Fig. 20F), ventral flaps of male segment 7, male legs 2 and 3 (Fig. 21D, E), penes (Fig. 21D) generally as in <i>G. costulifer</i> n. sp., but antennae elongate, antennomeres 5-7 richer in sensilla (Fig. 21A), gnathochilarium (mentum divided, n = 2) (Fig. 21B) slightly even more oligotrichous; collum (Fig. 20A) with a nearly typical carinotaxy formula (I-IV+5a+pc+ma+pc+5a+IV-I); all tergal crests (Fig. 20 A-E) on subsequent body segments higher, subdivided (2/2+I/i+3/3+I/i+2/2), ozoporiferous cones about as high as wide; epiproct (Fig. 20E, F) upturned and broadly rounded in caudal part, with one median and two paramedian crests; legs slightly longer than body diameter (Fig. 20B), always with an accessory filament/spine at base of claw, which is usually slightly more than half as long as claw (Fig. 21 D-F); male leg 1 with normal, 5-segmented telopodites and large, paramedian, coxal processes (Fig. 21C); each anterior gonopod plate with a high and subsecuriform distomedial process (Fig. 21G); posterior gonopods with distally slightly plumose flagella (Fig. 21H). REMARKS Based on several troglomorphic traits, such as the completely unpigmented body and ocelli, as well as the elongated antennae and legs, this species seems to be troglobitic, although the small size may also be evidence of its being an inhabitant of the MSS (“milieu souterrain superficiel”).
description: Published as part of <i>Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group, pp. 417-456 in Zoosystema 29 (3)</i> on pages 441-442, DOI: 10.5281/zenodo.5393630