Sceliphron caementarium
Creators: name: Ceccolini, Filippo
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Sphecidae
Sceliphron
Sceliphron caementarium

description: Sceliphron caementarium (Drury, 1773)
Sphex caementaria Drury, 1773
Sphex caementarium Drury, 1773
Sphex caementarius Drury, 1773
Sceliphron cementarium (Drury, 1770) (incorrect date)
Pelopeus caementarius (Drury, 1773)
Pelopoeus caementarius (Drury, 1773)
Sphex flavo-maculata De Geer, 1773
Sphex flavomaculata De Geer, 1773
Sphex flavomaculatus De Geer, 1773
Sceliphron caementarium flavomaculatum (De Geer, 1773)
Sphex lunata Fabricius, 1775
Sphex lunatus Fabricius, 1775
Pelopoeus lunatus (Fabricius, 1775)
Pelopeus caementarius lunatus (Fabricius, 1775)
Sceliphron caementarium lunatum (Fabricius, 1775)
Sphex flavipes Fabricius, 1781
Pelopoeus flavipes (Fabricius, 1781)
Pelopeus caementarius flavipes (Fabricius, 1781)
Sceliphron caementarium flavipes (Fabricius, 1781)
Sphex flavipunctata Christ, 1791
Sphex flavipunctatus Christ, 1791
Sceliphron caementarium flavipunctatum (Christ, 1791)
Sphex affinis Fabricius, 1793
Pelopoeus affinis (Fabricius, 1793)
Sceliphron affine (Fabricius, 1793)
Sceliphron caementarium affinis (Fabricius, 1793)
Pelopaeus lunatus Latreille, 1805
Pelopoeus architectus Lepeletier, 1845
Pelopeus caementarius architectus (Lepeletier, 1845)
Sceliphron caementarium architectus (Lepeletier, 1845)
Pelopoeus servillei Lepeletier, 1845
Sceliphron servillei (Lepeletier, 1845)
Sceliphron caementarium servillei (Lepeletier, 1845)
Pelopoeus solieri Lepeletier, 1845
Sceliphron solieri (Lepeletier, 1845)
Pelopoeus canadensis Smith, 1856
Sceliphron caementarium canadensis (Smith, 1856)
Pelopoeus nigriventris A. Costa, 1864
Sceliphron nigriventre (A. Costa, 1864)
Sceliphron caementarium nigriventre (A. Costa, 1864)
Pelopeus tahitensis de Saussure, 1867
Pelopoeus tahitensis de Saussure, 1867
Sceliphron tahitensis (de Saussure, 1867)
Sphex economica Curtiss, 1938
Sphex economicus Curtiss, 1938
Original distribution
Sceliphron caementarium is essentially a Nearctic species (Porter 1926; van der Vecht & van Breugel 1968; Hanson & Menke 1995; Amarante 2002; Horta Vega et al. 2007; Demetriou et al. 2022). Besides the states reported by Drury (1770), in its original range the species is known in southern Canada, several states of USA, Mexico, Belize, Nicaragua, and Costa Rica (see Pulawski 2022).
Allochthonous distribution in the world
The attendance of anthropic environments makes S. caementarium a species that can easily be accidentally transported by humans through long-distance dispersal (LDD) (see Butikofer et al. 2018; Cianferoni et al. 2021).
It was accidentally introduced several times in various parts of the world and even the locus typicus (Drury 1770) could be beyond the limits of its original range (see below for further information). The first certain record out of the native range was due to de Saussure (1867) from Tahiti, although he not recognize the wasp as allochthonous (see below for further information). The first precise record quoted as allochthonous species was by Cameron in Blackburn & Cameron (1886) from the Hawaii islands, where the species was already considered common at the time. However, van der Vecht & van Breugel (1968) reported some specimens collected in the island of Madeira in 1825. This record is currently the oldest known from outside the native range, excluding the locus typicus.
Nowadays S. caementarium is widespread in most of the world.
Literature data on the distribution out of the native range
ISLANDS OF THE ATLANTIC OCEAN
Some archipelagos in the Atlantic Ocean are far from the mainland and from each other. For this reason each archipelago is treated separately. A list of all literature records is given below for them, with indication of the oldest interception year. The records are listed geographically and archipelagos are arranged from west to east.
BERMUDA ISLANDS. Bermuda is an archipelago of seven main islands and about 170 additional (named) islets and rocks; anyway, all islands are very close to each other and the main islands are even connected by bridges (www.britannica.com/place/Bermuda). Moreover, the scarce records in the literature did not specify exactly in which islands S. caementarium was reported. For these raisons, the following reference data are considered for the entire archipelago without a specific list of the islands in which S. caementarium was intercepted.
Bermuda: Ogilvie (1928); van der Vecht & van Breugel (1968).
Oldest record: 1925 (van der Vecht & van Breugel 1968).
MADEIRA ARCHIPELAGO. Madeira: Saunders (1903); van der Vecht & van Breugel (1968); Lomholdt (1975); Leclercq (1976) and Leclercq & Claparède (1978) (same data); Erlandsson (1978); Smit (2000).
Remarks. Saunders (1903) named S. tubifex (Latreille, 1809) (= S. madraspatanum tubifex) four specimens (one male and three females) of Sceliphron caught by Wollaston in the island of Madeira but he misidentified the material, which was assigned to S. caementarium by Lomholdt (1975). Indeed, Saunders (1903) reported that all specimens “have the pedicel of the abdomen black” unlike S. madraspatanum tubifex but like S. caementarium (see Pagliano & Negrisolo 2005; Bitsch et al. 2020).
Oldest record: 1825 (van der Vecht & van Breugel 1968).
CARIBBEAN ISLANDS
Due to their geographical position, it is not easy to establish which Caribbean islands are part of its native range. However, except the small islands along the Central American coast, it is not likely they are included in the original distribution of the species. Although only a detailed genetic analysis on samples from several islands can reveal the origin of colonization of Caribbean islands by S. caementarium, it seems due to several independent introductions. Although the species was described from some islands of the Antilles (Drury 1770) and Fabricius (1775) reported Antigua in the distribution of the species citing Drury, its occurrence in those islands could be due to old accidental introductions. Effectively, the currently known records in the area are scattered and Wolcott (1951) reported that “It is apparently a recent introduction from continental America” and “It was first noted in Puerto Rico on the walls of a house in Santurce in September 1924 ”. Moreover, no specific records are from the largest countries of the Caribbean Sea such as Cuba or Haiti (cf. Genaro 2006; Pulawski 2022) and this suggests that there has not been a progressive natural colonization.
Some old works reported the species from Barbados (Porter 1926; Cheesman 1928; Williams 1932) and Cuba (Cheesman 1928; Williams 1932) without specific locality; anyway, no precise data exist about the occurrence of S. caementarium in these islands. In recent times these countries have been cited among those in which S. caementarium is distributed by Díaz-Calafat (2020) for Barbados and Anagha et al. (2021) for Cuba. However, they simply quoted aforementioned papers. Anyway, in the checklist of apoid wasps of Cuba, Genaro (2006) did not list the species and in Pulawski (2022) neither Barbados nor Cuba are included for S. caementarium.
All literature records for each Caribbean island are given below, with indication of the oldest interception year. The records are listed geographically with archipelagos arranged from north to south and for each islands are listed in alphabetical order.
LEEWARD ISLANDS. Antigua: Drury (1770). Guadeloupe: Lepeletier de Saint Fargeau (1845); van der Vecht & van Breugel (1968). Montserrat: van der Vecht & van Breugel (1968). Saint Croix: van der Vecht & van Breugel (1968). Saint Kitts: Drury (1770). Saint Thomas: van der Vecht & van Breugel (1968).
Oldest record: ante 1770 (Drury from Antigua and Saint Kitts).
GREATER ANTILLES: Caja de Muertos: Medina Gaud & Martorell (1974). Hispaniola (Dominican Republic): (Palisot de Beauvois 1806). Isla de la Juventud: Holland (1917). Jamaica (Drury 1770). Puerto Rico: Wolcott (1951); van der Vecht & van Breugel (1968).
Oldest record: ante 1770 (Drury from Jamaica).
WINDWARD ISLANDS. Martinique: Cheesman (1929); van der Vecht & van Breugel (1968). Mustique: van der Vecht & van Breugel (1968). Saint Vincent: van der Vecht & van Breugel (1968).
Remarks. Van der Vecht & van Breugel (1968) reported eight specimens from Martinique: one of them, located in the British Museum, is clearly the specimen cited by Cheesman (1929) since it was collected by Cheesman herself and the collecting date coincides, whilst the other ones are placed in the Museum of Comparative Zoology of Harvard University. One of them is not dated, but it was collected during the Virginia Expedition, whilst six specimens, labelled simply “ Martinique ”, were reported by authors as “old specimens”. Also specimens from Saint Vincent and Mustique were not dated.
Oldest record: 1925 (Cheesman 1929).
In addition to aforementioned records, Costa (1866) reported two specimens from Antilles in Museo Zoologico di Napoli and Berland (1953) cited S. caementarium for French Antilles without naming the specific islands.
SOUTH AMERICA
For South America S. caementarium was first reported by van der Vecht & van Breugel (1968) based on some specimens collected in Peru in 1954 and 1955. For many years no further records were provided from the continent and likely the species was not established there. Only in recent years the species has been reported in some countries and currently it seems to be naturalized also in that continent. Fernández (1995) and Amarante (2002) reported the presence of this mud-dauber wasp in Colombia, without specific localities, but Fernández & Castro-Huertas (2014) questioned its occurrence in that country and herein it is not included in the following list. Porter (1926) said that the species was “reported by Cameron from Brazil ”, but no documentation of this record exists and this statement could be a mistake. Therefore, also Brazil is not included there.
The following list includes all records for each South American country, reporting the oldest interception year for each one. The Galápagos Islands (belonging to Ecuador) are considered separately. The records are listed geographically and South America countries are arranged from north to south; for each state, the regions are listed in alphabetical order.
GALÁPAGOS ISLANDS. Santa Cruz: Causton & Sevilla (2007).
Remarks. The occurrence reported by Causton & Sevilla (2007) had not a precise date, but the authors listed S. caementarium among the species introduced in the archipelago in 2005–2006.
Oldest record: 2005/2006 (Causton & Sevilla 2007).
PERU. Arequipa: Ceccolini (2021). Ica: Barrera-Medina & Sepúlveda-Osorio (2014); Ceccolini (2021). Lima department: Ceccolini (2021). Lima province: Ceccolini (2021). Piura: van der Vecht & van Breugel (1968); Ceccolini (2021).
Remarks. Rasmussen & Asenjo (2009) reported the presence of S. caementarium in Lima without locality. Most likely the record refers to the whole department, but in the paper the homonym province is not considered as independent territory, therefore it is not possible to ascertain to which of the two areas the authors referred. Moreover, they quoted the department of Piura indicating between brackets the locality Sullana, the same one reported by van der Vecht & van Breugel (1968).
Oldest record: 1954 (van der Vecht & van Breugel 1968).
CHILE. Arica y Parinacota: Barrera-Medina & Sepúlveda-Osorio (2014). Santiago Metropolitan: BarreraMedina & Sepúlveda-Osorio (2014).
Oldest record: 2013 (Barrera-Medina & Sepúlveda-Osorio 2014).
OCEANIA
Oceania includes a large number of islands, many of which are very far from each other; for this reason each archipelago is treated separately. A list of all literature records for each island is given below, with indication of the oldest interception year. The records are listed geographically and the archipelagos are arranged from east to west and from north to south and for each one the islands are listed in alphabetical order. The list includes the Pacific archipelago of Volcano Islands, although the other records from Japan (to which they administratively belong) are reported under the distribution in Asia (see below). Although it is not possible to establish how many times humans introduced S. caementarium in these territories, clearly this mud-dauber arrived many times in the Pacific islands as a result of human activity.
VOLCANO ISLANDS. Iwô-Jima: Terayama (2004); Takahashi & Shimizu (2007).
Oldest record: 1976 (Takahashi & Shimizu 2007 from Iwô-Jima).
HAWAIIAN ISLANDS. Subsequently to Cameron in Blackburn & Cameron (1886) many works (even if not recent) documented occurrence of this wasp in this archipelago, where it appears widespread. For example, Perkins (1899) referred to this mud dauber as “Very abundant all over the islands, but not extending to great altitudes on the mountains”, whilst Swezey (1906) wrote: “No adults were seen but the nests of this muddauber wasp were common, and they always contained larvae or pupae, which shows that this species is continuing the habit of hibernation which it had in the States, where it hibernates as larvae in the cells of its mud-nest”.
A list of all literature records for each island of the archipelago is given below (the islands are listed alphabetically). No records without precise island is quoted, e.g., in addition to those mentioned above, van der Vecht & van Breugel (1968) who referred to “a series from these islands” preserved in the British Museum of London. The list includes also the Midway Atoll that is the only part in the Hawaiian Archipelago that is not included of the state of Hawaii (it is an Insular area of United States considered unorganized unincorporated territory) and where evidently S. caementarium, recorded in 1959 (Suehiro 1960), was introduced with a cargo for the American military air base which was in operation in the island from 1941 to 1993.
Hawaii: Yoshimoto (1960). Kauai: Alfken (1904); Yoshimoto (1960). Lehua: Alfken (1904). Sand Island (in Midway Atoll): Suehiro (1960). Molokai: Alfken (1904); Yoshimoto (1960). Niihau: Yoshimoto (1960). Ohau: Alfken (1904); Bridwell (1917); Kohl (1908); Swezey (1940); Yoshimoto (1960); Suman (1964).
Remarks. The record by Bridwell (1917) has been obtained indirectly. In the short note of Bridwell (1917) about an unidentified species of Chrysididae is written: “Mr. Bridwell reported finding cocoons and adults of this insect in Sceliphron nests in a road culvert at Diamond Head”. Subsequently Rohwer (1922) described this cuckoo wasp as Chrysis (Pentachrysis) extraniens through six specimens sent him by P.H. Timberlake, one of them labeled “Ex Sceliphron cells,” “Diamond Head, Oahu, May, 1919”. Moreover, Timberlake accompanied the specimens with a letter in which referred to them as follow: “I also send six specimens of a Chrysidid which was first taken here in 1914, and is now fairly common in Oahu. […] It is known to paratisize the common Sceliphorn caementarium.” (Rohwer 1922). Even if Bridwell (1917) did not specify the species of Sceliphron, it is clear that it is the same as the one mentioned by Timberlake. Moreover, Sceliphron caementarium is the only species of the genus occurring in the Hawaiian islands (see Pulawski 2022).
Oldest record: 1896 (Alfken 1904 from Hawaii, Lehua and Molokai).
MARSHALL ISLANDS. Kwajalein: Krombein (1949).
Oldest record: 1946 (Krombein 1949 from Kwajalein).
MARIANA ISLANDS. Guam: Krombein (1949). Saipan: Krombein (1949). Tinian: Krombein (1949).
Oldest record: 1946 (Krombein 1949 from Saipan).
MARQUESAS ISLANDS. Fatu Hiva: Cheesman (1928); Williams (1932). Hiva Oa: Cheesman (1928); Williams (1932). Nuku Hiva: Cheesman (1928); Berland (1945); Ramage et al. (2015). Ua Huka: Williams (1932).
Remarks. Records by van der Vecht & van Breugel (1968) are not considered because they observed the specimens already mentioned by Cheesman (1928).
Oldest record: 1924 (Ramage et al. 2015 from Nuku Hiva).
TUAMOTU ISLANDS. Fakahina: Ramage et al. (2015). Makatea: Ramage et al. (2015).
Oldest record: 2007 (Ramage et al. 2015 from Makatea).
SOCIETY ISLANDS. Bora Bora: Cheesman (1928). Moorea: Ramage et al. (2015). Raiatea: Cheesman (1928); Ramage et al. (2015). Tahiti: de Saussure (1867); Cheesman (1928); Curtiss (1938); van der Vecht & van Breugel (1968); Leclercq & Claparède (1978); Ramage et al. (2015).
Remarks. From Tahiti de Saussure (1867) described Pelopeus [sic] tahitensis, subsequently synonymized with Sceliphron caementarium by Kohl (1918), and Curtiss (1938) described Sphex economica [sic], subsequently synonymized with S. caementarium by Menke in Bohart and Menke (1976).
Oldest record: ante 1867 (de Saussure 1867 from Tahiti).
GAMBIER ISLANDS. Mangareva: van der Vecht & van Breugel (1968); Ramage et al. (2015).
Oldest record: 1905 (van der Vecht & van Breugel 1968 and Ramage et al. 2015).
AUSTRAL ISLANDS. Rurutu: Ramage et al. (2015).
Oldest record: 1962 (Ramage et al. 2015).
SOUTHERN COOK ISLANDS. Aitutaki: Krauss (1961); van der Vecht & van Breugel (1968) (reported under Samoa). Rarotonga: Kuhlmann (2006); McCormack (2007).
Oldest record: 1958 (van der Vecht & van Breugel 1968 from Aitutaki).
SAMOAN ISLANDS. 1946 (Lever 1946); Kami & Miller (1998).
Remarks. No precise localities are available, but Kami & Miller (1998) cited the species as present in “ Samoa islands” and “ Western Samoa ”.
Oldest record: 1946 (Lever 1946).
TERRITORY OF THE WALLIS AND FUTUNA ISLANDS. Wallis: van der Vecht & van Breugel (1968); Ramage et al. (2015).
Oldest record: 1958 (van der Vecht & van Breugel 1968 and Ramage et al. 2015).
FIJI ISLANDS. Viti Levu: Lever (1940); Kumar & Kumar (2017).
Oldest record: 1939 (Lever 1940).
ARCHIPELAGO OF NEW CALEDONIA. New Caledonia: van der Vecht & van Breugel (1968); Callan (1990); Ramage et al. (2015).
Oldest record: 1948 (Ramage et al. 2015).
AUSTRALIA. Northern Territory: Naumann (1983). Queensland: van der Vecht & van Breugel (1968); Riek (1970).
Oldest record: 1956 (van der Vecht & van Breugel 1968).
NEW ZEALAND. Te Ika-a-Mâui (= North Island): Early & Townsend (1983); Harris (1992). Te Waipounamu (= South Island): Harris (1997).
Remarks. Early & Townsend (1983) reported that one specimen was found in a ship’s hold in 1974 and another one was found in 1978 in a container, without specifying the localities, but likely these specimens were collected before they could get to the mainland. They expected that “it seems only a matter of time before one or more species [of Sceliphron] (probably the cosmopolitan tramp S. caementarium, or the Australian/Pacific S. laetum) become established in New Zealand ” (Early & Townsend 1983). Anyway, some years later Harris (1994) reported that “No Sceliphron is currently established in New Zealand ” and nowadays no evidence seem to support permanent populations of S. caementarium there.
Oldest record: 1982 (Early & Townsend 1983 from North Island).
ASIA
From Asia S. caementarium was first mentioned from “ India ” at the end of the nineteenth century as Pelopoeus solieri or S. solieri (Maindron 1878; Cameron 1889; Bingham 1897), but without specific localities and the first precise record for this country was reported only more than a century later (Gurule et al. 2020). Many records in the continent are from Japan, where the species seems to have been introduced after the Second World War (van der Vecht & van Breugel 1968).
A list of all literature records for each Asian country is given below, reporting the oldest interception year for each one. The records are listed geographically and the countries are arranged from east to west; for each state the regions are listed in alphabetical order.
JAPAN. Chűbu: Haneda et al. (2004, 2006); Suda (2004); Negoro et al. (2005); Ohkusa (2007). Kantô: van der Vecht & van Breugel (1968); Nambu (1972, 1975, 2006); Dollfuss (2016); Terayama (2021).
Remarks. Danilov (2020) cited specimens from Japan housed in the Institute of Systematics and Ecology of Animals of Novosibirsk, Russia, but the only reported datum is “Honshu” (the largest main island of Japan), without more precise information.
Van der Vecht & van Breugel (1968) reported second-hand information about established population in Kantô region (Tokyo and Yokohama); moreover, they said that the life-history of S. caementarium in Japan “was studied by Dr. Toshitomo Shida, 1952”.
Oldest record: 1952 (van der Vecht & van Breugel 1968).
SOUTH KOREA. Chungcheongnam: Kim et al. (2014). Gangwon: Kim et al. (2014). Gyeonggi: Kim et al. (2014).
Oldest record: 2011 (Kim et al. 2014).
CHINA. Sichuan: Hua (2006).
Remarks. In a list of Chinese insects, Hua (2006) reported simply “ Sichuan ” beside the name of S. caementarium without other information. Therefore, the date of the interception in the country is unknown.
Oldest record: ante 2006 (Hua 2006).
BANGLADESH. Dakha: Begum & Bose (1976); Begum et al. (1989, 1995).
Remarks. Begum & Bose (1976) listed the species (as S. Solieri Smith) among the solitary wasps of Dacca city and its suburbs but without year of collection.
Oldest record: ante 1976 (Begum & Bose 1976).
INDIA. Maharashtra: Gurule et al. (2020).
Remarks. Sceliphron caementarium was reported several times from India, but only Gurule et al. (2020) provided a precise locality. In addition to the aforementioned works (Maindron 1878; Cameron 1889; Bingham 1897), also Anagha & Girish Kumar (2020) and Anagha et al. (2021) quoted the species in the list of Indian sphecids, but reporting “no specific locality”. Actually there are other records of S. caementarium with precise locality from India since Chatterjee (2015) and Kumari & Priya (2022) reported it in West Bengal and Bihar regions respectively. Anyway the photos included in their papers show some nests and an adult specimen with large yellow body parts (including the entire petiole and part of the femurs) incompatible with S. caementarium. I believe the occurrence of S. caementarium in Bihar and West Bengal needs confirmation and in my opinion the species cannot be considered present in these regions.
The record of Gurule et al. (2020) had not a precise date, but the authors reported a time range from July 2015 to February 2016.
Oldest record: 2015/2016 (Gurule et al. 2020).
IRAN. Fars: Fallahzadeh et al. (2006, 2009).
Oldest record: 2004 (Fallahzadeh et al. 2006, 2009).
AFRICA
For Africa, the species was recorded only once by Dollfuss (2016) from Erithrea (without precise locality); this record dates back to 1968 and, after this interception, the species was not documented.
For a recent study on impacts of mud-daubers in the archeological site of Dandara, Egypt, Abd-Elkareem & Fouad (2022) collected several specimens identified by the laboratories of the Department of Plant Protection at the Faculty of Agriculture, Sohag University, as S. caementarium. No adult wasp photos are available in the paper, but the abundance of the species in the territory suggests that it is not the result of a recent introduction. It cannot be excluded that the mud-dauber analyzed in the paper is a native species, for example the common S. spirifex (Linnaeus, 1758), widespread in Egypt (Pulawski 2022). In the doubt that there was a misidentification, the occurrence of S. caementarium in Egypt needs confirmation.
In the Czech BioLib portal (Zicha 1999 –2022) South Africa is listed among the countries where the species is introduced, but no written records are available from that country.
EUROPE
Outside the original range, the continent where S. caementarium has the greatest distribution is Europe, where seven species of Sceliphron occur: four of them are native (Barbier 2013)—a further species, S. aterrimum (Marquet, 1875) from France, seems to be doubtful—and three are allochthonous (see Leclercq 1974; van der Vecht 1984; Ćetković et al. 2011).
The European islands of the Atlantic Ocean have already been discussed above. In the mainland the oldest occurrences of this mud-dauber wasp are from the Czech Republic (in 1942) according to Bogusch & Macek (2005) and from France (in 1945) according to Berland (1946). These records represent proofs of unsuccessful introductions, since the species was not observed till 1970’s, when it was recorded again in France (Leclercq 1974, 1978; Leclercq & Claparède 1978). Subsequently, S. caementarium became an invasive species and in the last decades it was widespread in large part of Europe.
A list of all literature records for each European country is given below, with indication of the oldest interception year. The records are listed geographically and the countries are arranged from west to east; for each country the regions are listed in alphabetical order.
PORTUGAL. Leiria: Cross et al. (2021). Lisbon: Leclercq (1976); Cross et al. (2021). Viseu: Cross et al. (2021).
Oldest record: 1975 (Leclercq 1976).
SPAIN. Balearic Islands: Diaz-Calafat (2020) (Mallorca island).
Oldest record: 2019 (Diaz-Calafat 2020).
FRANCE. Auvergne-Rhône-Alpes: Hamon et al. (1994); Bitsch et al. (1997); Schmid-Egger (2005); Bitsch & Barbier (2006); Bitsch (2010); van der Smissen (2010); Dollfuss (2016); Grangier (2017). Bourgogne-FrancheComté: Cretin (2015); Carminati et al. (2019). Centre-Val de Loire: Chapelin-Viscardi & Larivière (2014). Corsica: Hamon et al. (1989); Pagliano et al. (2000); Canovai et al. (2000); Bitsch & Barbier (2006). Grand Est: Callot & Brua (2013). Île-de-France: Berland (1946). Nouvelle-Aquitaine: Bitsch (2010). Occitanie: Leclercq 1976; Leclercq & Claparède (1978); Hamon et al. (1989, 1994); Carrière (2003); Magdalou (2006); Bitsch & Barbier (2006); Bitsch (2010); van der Smissen (2010); Dollfuss (2016); Magdalou (2017); Bitsch et al. (2020). Provence-Alpes-Côte d’Azur: Leclercq (1974); Haeseler (1976); Piek (1985, 1986, 1989); Hamon et al. (1989, 1994); Bitsch & Barbier (2006); van der Smissen (2010); Schmid-Egger (2011); Dollfuss (2016); Schmid‐Egger et al. (2018).
Oldest record: 1945 (Berland 1946) in mainland; 1986 (Hamon et al. 1989; Pagliano et al. 2000) in Corsica.
LUXEMBOURG. Esch-sur-Alzette: Schneider & Pelles (1998).
Oldest record: 1987 (Schneider & Pelles 1988).
BELGIUM. Flanders: Ravoet et al. (2017).
Oldest record: 2015 (Ravoet et al. 2017).
NETHERLANDS. Limburg: Schmitz (2015).
Remarks. Smit & Wijngaard (2010) listed S. camentarium among the species in a checklist of Sphecinae of Netherlands. The first precise locality is reported by Schmitz (2015).
Oldest record: 2010 (Schmitz 2015).
GERMANY. Hesse: Tischendorf et al. (2011); Burger (2015). Rhineland-Palatinate: Stalling (2002); Frommer (2012); Burger (2015) (all works reported that the species was found near Kaiserslauten, maybe through the same record; Burger (2015) is the only to specify the year of the record).
Remarks. Krombein (1979) cited the species as present in Germany but without locality and year of introduction.
Oldest record: ante 1979 (Krombein 1979); 1990 with precise year reported (Burger 2015).
SWITZERLAND. Basel-Landschaft: CSCF (2022). Basel-Stadt: CSCF (2022). Fribourg: CSCF (2022). Geneva: Vernier (2003); Boillat (2012); CSCF (2022). Graubünden: CSCF (2022). Jura: CSCF (2022). Lucerne: CSCF (2022). Neuchâtel: CSCF (2022). St. Gallen: CSCF (2022). Ticino: Schmid-Egger (2005); CSCF (2022). Vaud: Vernier (2003); CSCF (2022). Valais: CSCF (2022). Zürich: CSCF (2022).
Oldest record: 2000 (Vernier 2003).
AUSTRIA. Carinthia: Gusenleitner (2002). Lower Austria: Zettel et al. (2014).
Oldest record: 2002 (Gusenleitner 2002).
ITALY. Abruzzo: Schmid‐Egger et al. (2018); Bogusch (2022). Aosta Valley: Ceccolini & Paggetti (2013). Apulia: Turrisi & Altadonna (2017). Basilicata: Ceccolini & Paggetti (2012a). Campania: Turrisi & Altadonna (2017). Emilia-Romagna: Grillenzoni & Pesarini (1998); Campadelli et al. (1999); Rosa (2006); Pavesi (2008). Friuli-Venezia Giulia: Turrisi & Altadonna (2017). Liguria: Pagliano (1995); Pagliano & Negrisolo (2005). Lombardy: Polidori et al. (2007); Pavesi (2008); Chatenoud et al. (2012); Schmid‐Egger et al. (2018). Marche: Pagliano (2009). Molise: Ceccolini & Paggetti (2012a). Tuscany: Pagliano (1992); Strumia (1996); Pagliano & Negrisolo (2005); Ceccolini & Paggetti (2012a) (Elba island); Strumia et al. (2012) (Montecristo island); Bogusch (2022). Piedmont: Pagliano (2009); Schmid-Egger (2011); Dollfuss (2016). Sardinia: Cillo & Bazzato (2013); Dollfuss (2016). Sicily: Turrisi & Altadonna (2017). Trentino-Alto Adige: Hellrigl (2004 a,b, 2012). Umbria: Ceccolini & Paggetti (2012a,b). Veneto: Pagliano & Negrisolo (2005); Hellrigl (2006); Schmid‐Egger et al. (2018).
Remarks. Turrisi & Altadonna (2017) reported the distribution of the species in Italy listing all regions in which it is present, beyond their original record from Sicily. In the list Apulia, Campania and Friuli-Venezia Giulia are reported, but these are the only data from literature and specific records lack for these regions.
Oldest record: 1990 (Pagliano 1992) in mainland; 2013 in Sardinia (Cillo & Bazzato 2013); 2013 in Sicily (Turrisi & Altadonna 2017); 2010 in Tuscan Archipelago (Ceccolini & Paggetti 2012a).
MALTA. Central Region: (Malta island): Cassar & Mifsud (2020). South Eastern Region (Malta island): Cassar & Mifsud (2020).
Oldest record: 2018 (Cassar & Mifsud 2020).
SLOVENIA. Coastal–Karst: Gogala (2011); Bogusch (2022).
Oldest record: 1996 (Gogala 2011).
CROATIA. Dubrovnik-Neretva: Dollfuss (2016). Istria: Gusenleitner (1996, 2002); Pagliano & Negrisolo (2005); Józan (2009, 2014); Dollfuss (2016); Bogusch (2022). Primorje-Gorski Kotar: Józan (2009) (Krk island); Dollfuss (2016) (Cres island, Rab island). Šibenik-Knin: Dollfuss (2016). Zadar: Józan (2009, 2014); Bogusch (2022).
Oldest record: 1996 (Gusenleitner 1996).
SERBIA. Southern and Eastern Serbia: (Demetriou et al. 2022). Vojvodina: (Demetriou et al. 2022).
Oldest record: 2018 (Demetriou et al. 2022).
BOSNIA AND HERZEGOVINA. Herzegovina-Neretva: Rovoet et al. (2017).
Oldest record: 2011 (Rovoet et al. 2017).
MONTENEGRO. Bar: (Demetriou et al. 2022). Budva: (Demetriou et al. 2022). Herceg Novi: (Demetriou et al. 2022). Kotor: (Demetriou et al. 2022).
Oldest record: 2019 (Demetriou et al. 2022).
ALBANIA. Durrës: (Demetriou et al. 2022). Elbasan: Bogusch (2022). Vlorë (Demetriou et al. 2022).
Oldest record: 2019 (Demetriou et al. 2022).
GREECE. Attica: (Demetriou et al. 2022). Ionian Islands: (Demetriou et al. 2022) (Corfu island, Ithaca island). South Aegean: (Demetriou et al. 2022) (Patmo island). Thessaly: (Demetriou et al. 2022).
Oldest record: 2016 in mainland (Demetriou et al. 2022); 2011 in Ionian Islands (Demetriou et al. 2022 from Corfu); 2021 in South Aegean (Demetriou et al. 2022).
CZECH REPUBLIC. Central Bohemian: Bogusch & Macek (2005). South Moravian: Popelka (2015).
Oldest record: 1942 (Bogusch & Macek 2005).
SLOVAKIA. Bratislava: (Demetriou et al. 2022). Nitra: (Demetriou et al. 2022). Trenčín: (Demetriou et al. 2022). Trnava: (Demetriou et al. 2022).
Oldest record: 2019 (Demetriou et al. 2022).
HUNGARY. Central Hungary: Vas & Józan (2014); Hírek (2015).
Oldest record: 2010 (Vas & Józan 2014).
POLAND. Lesser Poland: (Demetriou et al. 2022). Subcarpathian Voivodeship: Kosibowicz (2021).
Oldest record: 2021 (Kosibowicz 2021; Demetriou et al. 2022).
ROMANIA. Bihor: Gagiu (2012).
Oldest record: 2012 (Gagiu 2012).
BULGARIA. Sofia City: Gradinarov (2017); Bogusch (2022). Varna: Gradinarov (2017).
Oldest record: 2012 (Gradinarov 2017)
UKRAINE. Odessa: Antropov (1993).
Oldest record: 1991 (Antropov 1993).
AUTONOMOUS REPUBLIC OF CRIMEA (disputed territory). Crimea: Shorenko (2007, 2009, 2017, 2018, 2019, 2020); Shorenko & Konovalov (2010); Protsenko et al. (2012); Danilov (2017).
Oldest record: 2007 (Shorenko 2007).
RUSSIA. Krasnodar: Popov (2022).
Oldest record: 2021 (Popov 2022).
NEW RECORDS
Material and methods
The material examined consists of collected or photographed specimens from websites (see abbreviations below). All listed records were recognized or confirmed by the author. Sceliphron caementarium is recognizable through photo thanks to its aforementioned colour pattern (see Introduction). For each site, the following information is given (when available): locality, coordinates, altitude, date of collecting, number of specimens, collector or photographer, repository or source. Some data has been duplicated from the primary web source also in the platform GBIF— Global Biodiversity Information Facility—(iNaturalist contributors, iNaturalist 2022) but only the primary source is reported. Continents or archipelagos are listed geographically and for each country or archipelago the regions or the islands are listed in alphabetical order. For each region the lower administrative division is underscored. Each island is underscored regardless of its administrative status. Geographical coordinates are in decimal degrees (datum WGS84). The uncertainty (in meters) of data is indicated according to the point-radius method (Wieczorek et al. 2004).
Abbreviations used under material examined are the following:
BOLD = barcode of Life Data System (www.barcodinglife.org)
CEP = collection Eugenio Pacieri, c/o MZUF
CFC = collection Filippo Ceccolini, Rassina (Arezzo), Italy
CFG = collection Francesca Graziani, Florence, Italy
CLP = collection Lucia Pizzocaro, Edolo (Brescia), Italy
CPC = collection Piero Cosimi, c/o MZUF
IN = iNaturalist (www.inaturalist.org)
FAP = Forum Acta Plantarum (https://www.actaplantarum.org)
FAR = Forum Aracnofilia—Associazione Italiana di Aracnologia (https://forum.aracnofilia.org)
FEI = Forum Entomologi Italiani (www.entomologiitaliani.net)
FNM = Forum Natura Mediterraneo (www.naturamediterraneo.com)
GMI = Galérie du Monde des insectes (www.galerie-insecte.org)
MZUF = Museo di Storia Naturale dell’Università degli Studi di Firenze, Sezione di Zoologia “La Specola”, Florence, Italy
OB = observation.org (https://observation.org).
UBIN = Ukrainian Biodiversity Information Network (www.ukrbin.com)
UCBL1 = Université Claude Bernard Lyon 1
WNL = waarneming.nl (www.waarneming.nl)
leg. = legit/legerunt un. = uncertainty
Material examined
ISLANDS OF THE ATLANTIC OCEAN
BERMUDA. Hamilton Parish: Blue Hole Park, 32.346613°N - 64.708578°E (un. = 40 m), 30.VII.2021, 2 specimens, photo by Luke Foster (IN); Shelly Bay Park, 32.334167°N - 64.740386°E (un. = 10 m), 15.VIII.2021, 1 specimen, photo by Miguel A Mejias (IN). Smith’s Parish: Spittal Pond, 32.311443°N - 64.725665°E (un. = 101 m), 29.VII.2021, 1 specimen, photo by Luke Foster (IN).
AZORES. Terceira island: Angra do Heroísmo, 38.662801°N - 27.212863°E (un. = 9 m), 12.VI.2022, 1 specimen, photo by Abr„o Leite (IN).
MADEIRA ARCHIPELAGO.Madeira: Santana, near S„o Jorge, 32.8334°N - 16.9004°E (un.= 25 m), 7.VIII.2019, 1 specimen, photo by Danny Klaessens (OB); Faj„ da Rocha do Navio, 32.8135°N - 16.8720°E (un. = 10 m), 1 specimen, photo by Jord Ingenbleek (OB); Câmara de Lobos, 32.6825°N - 16.9952°E (un. = 81 m), 29.VII.2022, 1 specimen, photo by Stan van der Laan (OB); Funchal, Pico do Ariero, 32.7116°N - 16.9635°E (un. = 29 m), 14.VIII.2021, 1 specimen, photo by “robrob1975” (OB); Funchal, near Jardim Botânico da Madeira, 32.661823°N - 16.895337°E (un. not recorded), 15.IX.2017, 1 specimen, photo by Thorsten Stegmann (IN); idem, Santo Antonio, Jardim da Quinta da Achada, 32.655272°N - 16.91796°E (un. = 31 m), 29.V.2022, 1 specimen, photo by Kevin Arbuckle (IN); idem, S„o Martinho, Jardim Municipal do Funchal, 32.6481°N - 16.9112°E (un. = 6 m), 23.VIII.2022, 1 specimen, photo by “Tove” (OB); idem, 32.6478°N - 16.9115°E (un. = 209 m), 5.IX.2021, 1 specimen, photo by “lloirs” (IN); idem, Jardins do Lido, 32.637072°N - 16.933139°E (un. = 10 m), 24.VII.2021, 1 specimen, photo by Peter Lindberg Jannerup (IN); idem, 32.636845°N - 16.932496°E (un. not recorded), 21.VII.2021, 1 specimen, photo by Peter Lindberg Jannerup (IN); idem, R. Nova do Vale da Ajuda, 32.636302°N - 16.94199°E (un. = 5 m), 28.VII.2019, 2 specimens, photo by “jkt” (IN); idem, near Promenade do Lido, 32.6370°N - 16.9327°E (un. = 10 m), 6.IX.2019, 1 specimen, photo by Pietsje Fleurbaaij (OB); idem, Jardins do Avenida do Mar, 32.6468°N - 16.9089°E (un. = 10 m), 26.VI.2018, 1 specimen, photo by Hans Vanhulle (OB); idem, Parque de Santa Catarina, 32.6457°N - 16.9139°E (un. = 25 m), 22.IX.2022, 1 specimen, photo by “Nessa & Nono” (OB); between Pico Ruivo and Achada do Teixeira, 32.761235°N - 16.935721°E (un. = 50 m), 25.VII.2021, 1 specimen, photo by “miwi2020” (IN); Machico, Caniçal, near Museu da Baleia da Madeira, 32.736182° - 16.740713°E (un. = 6 m), 21.VIII.2020, 1 specimen, photo by Tim Leilich (IN); Calheta, Jardim do Mar, 32.7388°N - 17.2115°E (un. not recorded), 25.VII.2021, 1 specimen, photo by Thierry Madigou (GMI).
CANARY ISLANDS. Tenerife: Santa Cruz de Tenerife, Plaza de la Iglesia, 28.464761°N - 16.248704°E (un. = 1 m), 1.X.2019, 1 specimen, photo by “gabohq” (IN).
CARIBBEAN ISLANDS
CAYMAN ISLANDS. Grand Cayman: North Side, Breakers, 19.313911°N - 81.188754°E (un. = 4 m), 11.VIII.2018, 1 specimen, photo by “wyattherp” (IN); Bodden Town, Savannah, Governor Gore’s Bird Sanctuary, 19.277699°N - 81.308823°E (un. = 100 m), 30.III.2019, 1 specimen (Fig. 1), photo by Steven Mlodinow (IN); George Town, South Sound Road, 19.273173°N - 81.365605°E (un. = 30 m), 13.VI.2020, 1 specimen, photo by “caymankatie” (IN).
SOUTH AMERICA
PERU. La Libertad: Trujillo, Trujillo, Plazuela Antonio Raimondi, - 8.109664°N - 79.032379°E (un. = 31 m), 23.XI.2021, 1 specimen, photo by Jose Laca (IN); idem, Distrito de Víctor Larco Herrera, Parque Las Orquídeas, - 8.132983°N - 79.051053°E (un. = 4 m), 31.X.2021, 1 specimen, photo by Rob Westerduijn (IN). Lima (department): Huarochirí, Surco, - 11.882832°N - 76.438295°E (un. = 44 m), 15.IV.2022, 1 specimen, photo by Brandon Ortiz Arenas (IN); idem, near Ayas, - 11.868601°N - 76.424568°E (un. = 123 m), 15.IV.2022, 1 specimen, photo by Brandon Ortiz Arenas (IN); Lima (province): San Borja, Parque Pallardelli, - 12.085633°N - 76.99543°E (un. = 8 m), 10.IV.2022, 1 specimen, photo by Antonio W. Salas (IN).
CHILE. Santiago Metropolitan: Cordillera, Las Condes, - 33.43422°N - 70.503462°E (un. not recorded), 11.I.2021, 1 specimen, photo by Orlando Montes (IN); Santiago, Santiago de Chile, Av. Pedro Bannen, - 33.416892°N - 70.623323°E (un not recorded), 2.I.2020, 1 specimen, photo by Orlando Montes (IN); Maipú, René Olivares Becerra, - 33.531316°N - 70.772089°E (un. not recorded), 2.I.2022, 1 specimen, photo by Pablo Moreno Vallejos (IN); idem, Parque Municipal El Trapiche de Peñaflor, - 33.595769°N - 70.894617°E (un. = 4 m), 29.XII.2020, 1 specimen, photo by Germán Ortiz (IN).
ARGENTINA. La Pampa: Utracán, General Acha, - 37.368794°N - 64.599867°E (un = 2.85 km), 20.II.2022, 1 specimen (Fig. 2), photo by Daniel Oscar Molina (IN).
OCEANIA
HAWAIIAN ISLANDS. Kauai island: Kapaʻa, 22.056652°N - 159.322542°E (un not recorded), 22.V.2022, 1 specimen, photo by “jcripe” (IN); Po’ipu, 21.874437°N - 159.453857°E (un. = 7 m), 3.X.2018, 1 specimen, photo by “sea-kangaroo” (IN); Eleele, 21.899053°N - 159.584062°E (un. = 5 m), 12.I.2020, 1 specimen, photo by Tony Palmer (IN); Oahu island: Koʻolauloa, Pűpűkea, 21.651029°N - 158.061877°E (un. = 5 m), 12.X.2018, 1 specimen, photo by Harvey Perkins (IN); Waianae, 21.454047°N - 158.19246°E (un. = 65 m), 10.V.2018, 1 specimen, photo by “mcclelland” (IN); Royal Kunia, 21.401165°N - 158.023114°E (un. = 5 m), 30.XI.2020, 1 specimen, photo by “rkkam” (IN); Ko Olina, Aliinui Dr, 21.345961°N - 158.125917°E (un. = 5 m), 9.V.2018, 1 specimen, photo by Alexander Moerman (IN); Ewa Beach, 21.313148°N - 158.008999°E (un = 2.93 km), 21.X.2021, 1 specimen, photo by Nadia Tubbs (IN); Waimanalo, 21.376619°N - 157.705837°E (un. = 31 m), 3.II.2020, 1 specimen, photo by Zoe Ferraris (IN); Ala Moana, 21.2886°N - 157.8419°E (un. = 25 m), 6.V.2022, 1 specimen, photo by Arjan Loeve (OB); Maui island: Kapalua, 21.006157°N - 156.655431°E (un. = 56 m), 13.XII.2017, 1 specimen, photo by “sea-kangaroo” (IN); idem, Napili Bay, 20.994085°N - 156.668276°E (un. = 7 m), 20.V.2022, 2 specimens, photo by “sea-kangaroo” (IN); Pa’ia, 20.929296°N - 156.367064°E (un. = 18 m), 15.VIII.2016, 1 specimen, photo by “sea-kangaroo” (IN); Kula, Haleakalâ, 20.713673°N - 156.251737°E (un. = 526 m), 3.I.2019, 1 specimen, photo by Monica Ventrice (IN); Wailea-Makena, Maluaka Beach, 20.648378°N - 156.442413°E (un. = 8 m), 9.XI.2020, 1 specimen, photo by John Starmer (IN); Hawaii island: Kawaihae, Hapuna Beach, 20.005742°N - 155.824463°E (un. = 65 m), 14.X.2019, 1 specimen, photo by “mbarger” (IN); Kailua-Kona, West Hawai’i Explorations Academy, 19.715309°N - 156.036802°E (un. = 50 m), 17.II.2022, 1 specimen, photo by “kbkash” (IN); Holualoa, 19.61262°N - 155.975159°E (un. = 8 m), 24.X.2016, 1 specimen, photo by Steve Wells (IN); idem, 19.603751° - 155.974971°E (un. = 206 m), 4.III.2017, 1 specimen, photo by Joelle Belmonte (IN); Keauhou, 19.559045°N - 155.96283°E (un. = 5 m), 18.IV.2018, 1 specimen, photo by Cory Silas Sheffield (IN); Puʻuhonua o Hônaunau National Historical Park, 19.420932°N - 155.911605°E (un. not recorded), 11.I.2020, 1 specimen, photo by Lori Brandt Dudzik (IN); idem, 19.420867°N - 155.911552°E (un. not recorded), 16.IV.2022, 1 specimen, photo by Nadya Kandel (IN); idem, 19.420798°N - 155.911703°E (un. not recorded), 28.V.2022, 1 specimen, photo by Phil Calvert (IN).
SOCIETY ISLANDS. Moorea island: Piha’ena, - 17.492°N - 149.83°E (un not recorded), 6.II.2007, 1 specimen, leg. Anne Duplouy, UCBL1, photo on BOLD; near Marae Ti’i-rua, - 17.533°N - 149.839°E (un. not recorded), 12.I.2006, 1 specimen, leg. Marjorie Saillan, UCBL1, photo on BOLD.
NEW CALEDONIA. North Province: Pouembout, near Camping de Tipenga, - 21.134688°N 164.914701°E (un. = 122 m), 30.III.2019, 1 specimen, photo by “juju98” (IN).
AUSTRALIA. Queensland: Moreton Bay Region, Griffin, - 27.2638°N 153.0431°E (un. not recorded), 20.III.2022, 1 specimen, photo by Paul G. Schrijvershof (IN).
ASIA
JAPAN. Chűgoku: Hiroshima, Onomichi, Nishikubocho, 34.415182°N 133.204819°E (un. = 7.56 km), 9.VIII.2020, 1 specimen, photo by “lentom” (IN). Kansai: Hyôgo, Kôbe, 34.690083°N 135.195511°E (un. = 8.51 km), 28.VI.2014, 1 specimen, photo by “harumkoh” (IN); Himeji, Tofumachi, 34.826629°N 134.689878°E (un. = 194 m), 3.VII.2017, 1 specimen, photo by Yaron Budowski (IN). Kantô: Tokyo, Shinagawa, 35.601486°N 139.75296°E (un. = 1.95 km), 27.VI.2017, 1 specimen, photo by Steven Kurniawidjaja (IN).
SOUTH KOREA. Gyeonggi: Yangju, Deokgye-dong, 37.817614°N 127.056305°E (un. = 8 m), 1.VII.2017, 1 specimen, photo by “whaichi” (IN); Uijeongbu, 37.741467°N 127.054353°E (un. not recorded), 27.V.2016, 1 specimen, photo by “whaichi” (IN); idem, 37.736016°N 127.052684°E (un. = 17 m), 27.V.2016, 2 specimens, photo by “whaichi” (IN); Daegu: Dong, Geomdan-dong, 35.916566°N 128.630915°E (un. = 5 m), 8.VII.2017, 1 specimen, photo by “bio96” (IN); idem, 35.916375°N 128.630511°E (un. = 4 m), 21.VI.2018, 1 specimen, photo by “bio96” (IN); idem, 35.9147°N 128.630475°E (un. = 4 m), 31.VII.2018, 1 specimen, photo by “bio96” (IN); idem, 35.916339°N 128.63045°E (un. = 4 m), 30.VII.2019, 1 specimen, photo by “bio96” (IN); idem, 35.916098°N 128.630212°E (un. = 9 m), 8.VIII.2019, 1 specimen (Fig. 3), photo by “bio96” (IN); Buk, Duryu Park, 35.843842°N 128.561106°E (un. = 10 m), 28.VIII.2021, 1 specimen, photo by “bio96” (IN). North Gyeongsang: Gyeongsang, Gapje-dong, 35.825678°N 128.763429°E (un. = 300 m), 26.IX.2015, 1 specimen, photo by “ichman” (IN).
OMAN. Muscat governorate: Muscat, between 3011 Way and 3015 Way, 23.609396°N 58.553457°E (uncertainty = 29 m), 27.V.2020, 1 specimen, photo by “suvijohn” (Fig. 4).
EUROPE
PORTUGAL. Aveiro: Estarreja, Salreu, 40.728358°N - 8.587711°E (un. = 1900 m), 4.VIII.2020, 1 specimen, photo by Luís Gordinho (IN); idem, 40.723185°N - 8.569851°E (un. = 393 m), 21.VIII.2016, 1 specimen, photo by Gonçalo Andrade Hagland (IN); idem, 40.7346°N - 8.5868°E (un. = 10 m), 23.VIII.2017, 1 specimen, photo by René Beekvelt (OB); Aveiro, Marinha da Noeirinha, 40.646342°N - 8.654607°E (un. not recorded), 31.V.2020, 1 specimen, photo by Duarte Frade (IN); idem, Escola Superior De Saúde Da Universidade De Aveiro, 40.623322°N - 8.656796°E (un. = 20 m), 7.VI.2022, 1 specimen, photo by Charcos Companhia (IN). Coimbra: Coimbra, Parque Verde do Mondego, 40.201559°N - 8.425581°E (un. = 10 m), 10.IX.2017, 1 specimen, Pablo de la Fuente Brun (IN). Faro: Lagos, Bar„o de S„o Jo„o, 37.1420347°N - 8.7623525°E (un not recorded), 68 m, 13.IX.2021, 1 specimen, photo by Alain Roujas (GMI); Faro, Faro, 37.015097°N - 7.922149°E (un. = 4 m), 17.IX.2022, 1 specimen, photo by Duarte Frade (IN). Leiria: Pombal, R. da L„mpada, 39.985485°N - 8.581843°E (un. = 8 m), 15.IX.2019, 1 specimen, photo by Hugo Gaspar (IN); near Pombal, 39.902752°N - 8.665588°E (un. = 2 m), 24.V.2022, 1 specimen, photo by José M. Martins (IN); Leiria, Amor, 39.802823°N - 8.860834°E (un. = 4210 m), 24.VIII.2020, 1 specimen, photo by Vasco Duarte (IN); Caranguejeira, 39.771251°N - 8.720378°E (un. = 22 m), 27.VIII.2020, 1 specimen, photo by “zeferreira”; Caldas da Rainha, R. do Talefe, 39.404717°N - 9.160597°E (un. = 65 m), 21.VIII.2020, 1 specimen, photo by Barbara Fonseca (IN); Porto de Mós, near Mendiga, 39.4828°N - 8.8067°E (un. = 10 m), 21.VII.2016, 1 specimen, photo by Francisco Barros (OB); Óbidos, Gaeiras, 39.368022°N - 9.132597°E (un. = 242 m), 4.VII.2022, 1 specimen, photo by Anaís Fidalgo (IN). Lisbon: Cadaval, Cercal, 39.2104°N - 8.9969°E (un. not recorded), 1.IX.2010, 1 specimen, photo by Francisco Barros (OB); Loures, Forte do Mosqueiro, 38.898778°N - 9.18255°E (un. = 188 m), 18.IX.2021, 1 specimen, photo by Marta Veiga Santos (IN); S„o Jo„o da Talha, 38.829422°N - 9.089421°E (un. = 4 m), 10.VII.2021, 1 specimen, photo by Sonia Ferreirai (IN); Vila Franca de Xira, Reserva Natural do Estuário do Tejo, 38.847144°N - 8.971694°E (un. = 161 m), 8.VIII.2021, 1 specimen, photo by Tiago Carrapiço (IN); Lourinh„, Estr. da Cezaredas, 39.297508°N - 9.244251°E (un. not recorded), 20.IX.2020, 1 specimen, photo by Octávio Mateus (IN); Alenquer, Pereiro de Palhacana, 39.043151°N - 9.126182°E (un. = 58 m), 20.IX.2020, 1 specimen, photo by Paulo Caseirito (IN); Sintra, Parque Urbano Felício Loureiro, 38.7542°N - 9.2588°E (un. not recorded), 31.V.2010, 1 specimen, photo by “Cidadania Queluz” (OB); Lisboa, Rua José da Purificaç„o Chaves, 38.745318°N - 9.190359°E (un. = 24 m), 11.VI.2022, 1 specimen, photo by “analopesfsl” (IN); Parque Forestal de Monsanto, 38.739509°N - 9.182081°E (un. = 85 m), 22.VII.2021, 1 specimen, photo by “pedrodn” (IN); idem, 38.733769°N - 9.178111°E (un. = 75 m), 27.VII.2021, 1 specimen, photo by Tiago Carrapiço (IN); idem, IDMEC—Instituto de Engenharia Mecânica, 38.737412°N - 9.138327°E (un. = 26 m), 26.V.2021, 1 specimen, photo by “vproenca” (IN); idem, Jardim dos Montes Claros, 38.716192°N - 9.197869°E (un. not recorded), 5.VII.2021, 1 specimen, photo by Diogo Sebasti„o (IN); Oeiras, Bairro Comendador Joaquim Matias, 38.694985°N - 9.299924°E (un. = 498 m), 9.VII.2018, 1 specimen, photo by Filipe Ribeiro (IN). Porto: Bai„o, Parque de Lazer, 41.16517°N - 8.024447°E (un. = 50 m), 31.VIII.2021, 1 specimen, photo by Daniel Ferreira; Penafiel, near Parque Merendas de Capela, 41.111386°N - 8.354492°E (un. = 16 m), 23.VIII.2021, 1 specimen, photo by “j_naturalist” (IN); idem, Lagares, Rua de Quintandona, 41.133736°N - 8.379442°E (un. = 8 m), 23.VIII.2021, 1 specimen, photo by “j_naturalist” (IN); Gondomar, Moinhos de Jancido, 41.095878°N - 8.489383°E (un. = 4 m), 16.IX.2022, 1 specimen, photo by Jo„o Lima (IN). Santarém: Alpiarça, Reserva Natural do Cavalo do Sorraia, 39.23597°N - 8.5824°E (un. = 15 m), 25.VI.2022, 1 specimen (Fig. 5), photo by Antonio Goncalves (IN). Setúbal: Alcochete, Passil, 38.7170°N - 8.8871°E (un. = 10 m), 1.VI.2017, 1 specimen, photo by Francisco Barros (OB); Almada, near Estádio Municipal José Martins Vieira, 38.664894°N - 9.169441°E (un. = 15 m), 7.VIII.2019, 1 specimen, photo by Luís Lourenço (IN); idem, 38.6629°N - 9.169244°E (un. = 15 m), 1 specimen, photo by Luís Lourenço (IN); R. José Carlos de Melo, 38.646237°N - 9.186716°E (un. = 135 m), 13.VIII.2019, 1 specimen, photo by Luís Lourenço (IN); idem, 38.646312°N - 9.186672°E (un. = 20 m), 23.VIII.2019, 1 specimen, photo by Luís Lourenço (IN); R. Gen. Humberto Delgado, 38.64403°N - 9.173273°E (un. not recorded), 11.VIII.2020, 1 specimen, photo by Luís Lourenço (IN); R. do Cabeço Verde, 38.604247°N - 9.191939°E (un. = 39 m), 19.IX.2021, 1 specimen, photo by Tiago Carrapiço (IN); Charneca da Caparica, Rua de S„o Jo„o, 38.603923°N - 9.196052°E (un. = 18 m), 25.VIII.2022, 1 specimen, photo by Sergio Chozas (IN); Seixal, Av. Mud Juvenil, 38.645351°N - 9.093085°E (un. not recorded), 27.VII.2022, 1 specimen, photo by Joaquim Pedroso (IN); idem, Largo Alfredo Jose Almeida Lima, 38.626691°N - 9.081355°E (un. = 1690 m), 12.V.2021, 1 specimen, photo by Ana Jorge da Costa (IN); idem, Santa Marta do Pinhal, R. António Garibaldi, 38.628165°N - 9.152884°E (un. = 4 m), 1.VIII.2021, 2 specimens, photo by Daniel Raposo (IN); Barreiro, Lago da Mata da Machada, 38.60586°N - 9.025485°E (un. = 60 m), 26.VI.2021, 1 specimen, photo by Tiago Carrapiço (IN); Sesimbra, Espaço Interpretativo da Lagoa Pequena, 38.525398°N - 9.142877°E (un. not recorded), 10.VII.2020, 1 specimen, photo by Luís Lourenço (IN); idem, 38.525337°N - 9.142924°E (un. not recorded), 2.VI.2019, 1 specimen, photo by Luís Gordinho (IN): idem, 38.525221°N - 9.143028°E (un. = 15 m), 10.VI.2017, 1 specimen, photo by Daniel Raposo (IN); idem, 38.522335°N - 9.143838°E (un. = 187 m), 2.VII.2021, 1 specimen, photo by Pedro Castro (IN). Viseu: Castro Daire, Farejinhas, R. da Costa, 40.907946°N - 7.898012°E (un. = 27 m), 11.VI.2021, 1 specimen, photo by Jo„o Ponte (IN).
SPAIN. Andalusia: Cádiz, Santa Margarita, 36.203659°N - 5.335262°E (un. not recorded), 22.VII.2020, 1 specimen, photo by Juan Manuel Palmero Sánchez (IN). Aragon: Huesca, Valle de Hecho, Embún, 42.626214°N - 0.720776°E (un. = 32 m), 3.VIII.2022, 1 specimen, photo by Ana Pujante (IN). Balearic Islands: Mallorca island, Port de Pollença, 39.90278°N 3.079283°E (un. not recorded), 18.VI.2022, 1 specimen, photo by “jarvo” (IN). Catalonia: Girona, Bolvir, 42.417999°N 1.879894°E (un. = 1070 m), 3.X.2021, 1 specimen, photo by Eduard Marques (IN); Ribes de Freser, 42.318697°N 2.186978°E (un. not recorded), 18.VI.2022, 1 specimen, photo by “aurearamon” (IN); Cadaqués, 42.285737°N 3.295795°E (un. = 5 m), 8.IX.2019, 1 specimen, photo by “jofree” (IN); idem, 42.285066°N 3.296179°E (un. not recorded), 25.VIII.2021, 1 specimen, photo by Jeremy Barker (IN); idem, Carrer de Cala Montjoi, 42.267355°N 3.018481°E (un. = 65 m), 12.VI.2020, 1 specimen, photo by Bernat Espigulé (IN);Alt Empordà, Figueres, 42.266289°N 2.951822°E (un. = 443 m), 30.VIII.2020, 1 specimen, photo by Enric Capalleras (IN); idem, 42.259926°N 2.951789°E (un. = 55 m), 29.VI.2022, 1 specimen, photo by “bixaurin” (IN); Vila-sacra, Calle Capitán de Creus, 42.267463°N 3.018597°E (un. = 5 m), 27.V.2022, 1 specimen, photo by “jofree” (IN); Saus, Camallera i Llampaies, Camallera, 42.122514°N 2.961382°E (un. not recorded), 31.VII.2022, 1 specimen, photo by Jacobo Krauel (IN); L’Escala, Montgó, 42.1130°N 3.1713 E (un. = 13 m), 2.VIII.2022, 1 specimen, photo by “Glenn S” (OB); Pals, 41.9701°N 3.1462°E (un. = 25 m), 16.VII.2022, 1 specimen, photo by “Bakker” (OB); Girona, Carrer de Bescanó, 41.993005°N 2.796381°E (un. = 33 m), 27.VII.2019, 1 specimen, photo by Pavel Vernigorov (IN); idem, Palamós, 41.861058°N 3.144738°E (un. not recorded), 13.VII.2021, 1 specimen, photo by Valentin de Mazancourt (IN); idem, San Felíu de Buxalleu, 41.745182°N 2.609786°E (un. not recorded), 11.IX.2019, 1 specimen, photo by Paco Aguayo (IN); Blanes, 41.6768°N 2.8052°E (un. = 204 m), 30.VII.2022, 1 specimen, photo by “peter VdJ” (OB); Barcelona, Tordera, Carrer de la Mare de Déu de la Mercè, 41.693638°N 2.728083°E (un. = 65 m), 27.VI.2020, 1 specimen, photo by Jordi Pages (IN); Santa Susanna, 41.635535°N 2.698969°E (un. = 9 m), 9.VIII.2019, 1 specimen, photo by “ant-toniscla” (IN); idem, 41.633741°N 2.698125°E (un. = 55 m), 11.VIII.2020, 1 specimen, photo by“anttoniscla” (IN); idem, 41.634868°N 2.704805°E (un. = 20 m), 3.VIII.2019, 1 specimen, photo by “ant-toniscla” (IN); Terrassa, Carrer de Wagner, 41.562138°N 2.023197°E (un. = 5 m), 10.IX.2021, 1 specimen, photo by Sergi Massanes (IN); Polinyà, Carrer Major, 41.555663°N 2.154439°E (un. = 26 m), 22.VIII.2020, 1 specimen, photo by “Blanca” (IN); Badalona, Riera de Canyadó, 41.460527°N 2.250761°E (un. = 409 m), 28.VI.2020, 1 specimen, photo by “eev” (IN); Barcelona, La Salut, Parc Ģell, 41.414444°N 2.152222°E (un. not recorded), 15.VII.2019, 1 specimen, photo by “neurojimbo” (IN); idem, L’Hospitalet de Llobregat, Carrer Can Trias, 41.354843°N 2.121089°E (un. = 65 m), 12.VI.2021, 1 specimen, photo by “ramonhoms” (IN). Comunidad Valenciana: València, València, Parque de la Rambleta, 39.446945°N - 0.393187°E (un. = 4 m), 11.VIII.2021, 1 specimen (Fig. 6), photo by David Fhuerta (IN); Alicante, Alicante city, Sant Antoni, 38.349413°N - 0.479411°E (un. = 31 m), 24.VIII.2021, 1 specimen, photo by Alexandra Laking (IN). Navarra: Navarra, Roncal, 42.8079°N - 0.9551°E (un. = 5 m), 24.VII.2021, 8 specimens, photo by Ana Eguaras (OB); Aoiz, 42.7926°N - 1.3622°E (un. = 25 m), 9.IX.2020, 1 specimen, photo by Santiago Serrano (OB); Eģés, Mutilva, 42.8038°N - 1.6113°E (un. = 25 m), 24.VIII.2020, 1 specimen, photo by Santiago Serrano (OB).
FRANCE. Auvergne-Rhône-Alpes: Puy-de-Dôme, Saint-Germain-Lembron, 45.4645°N 3.2432°E (un. = 25 m), 5.VIII.2020, 1 specimen, photo by Lennard Barendrecht (OB); Allier, Moulins, 46.543502°N 3.3836°E (un. = 10 m), 15.VIII.2021, 1 specimen, photo by “anbalemans” (IN); Ain, Arbigny, 46.470072°N 4.962536°E (un. not recorded), 4.VIII.2020, 1 specimen, photo by “nonomay” (IN); idem, Attignat, 46.288208°N 5.157924°E (un. not recorded), 29.VII.2019, 1 specimen, photo by “brebrebre” (IN); Saint-Jean-le-Vieux, 46.0278510°N 5.3870584°E (un. not recorded), 258 m, 19.VIII.2012, 1 specimen, photo by Yves Dubois (GMI); Loire, Riorges, 46.034482°N 4.037153°E (un. = 20 m), 2.VIII.2020, 1 specimen, photo by “La Dame du Clair” (IN); Le Coteau, 46.0298°N 4.0881°E (un. = 100 m), 18.VI.2019, 1 specimen, photo by Anita van der Landen (OB); Groslée-Saint-Benoît, 45.6555980°N 5.6086707°E (un. not recorded), 23.VIII. 2008, 300 m, 1 specimen, photo by Gerard Cuchet (GMI); Sorbiers, 45.50499°N 4.428501°E (un. = 98 m), 11.VI.2022, 1 specimen, photo by “poye” (IN); La Talaudière, 45.475946°N 4.435155°E (un. = 15 m), 16.IX.2014, 2 specimens, photo by Mickaël Villemagne (IN); Saint-Chamond, 45.463248°N 4.514801°E (un. = 188 m), 10.VIII.2008, 1 specimen, photo by “ahmedm” (IN); Maclas, Goëly, 45.378351°N 4.682059°E (un. not recorded), 20.IX.2013, 1 specimen, photo by Pascal Dubois; Rhône, Liergues, Porte des Pierres Dorées, 45.97004°N 4.671172°E (un. not recorded), 8.VI.2022, 1 specimen, photo by Julien Lepage (IN); L’Arbresle, 45.8349591°N 4.6149445°E (un. not recorded), 18.VII.2011, 1 specimen, photo by “Sylvestre II” (GMI); Dardilly, 45.824727°N 4.742665°E (un. = 63 m), 29.VI.2021, 1 specimen, photo by “hugodonata” (IN); Lyon, Jardin des Pendarts, 45.772299°N 4.829894°E (un. = 263 m), 7.VII.2020, 1 specimen, photo by “matthieudepl” (IN); idem, near Hôpital Pierre Wertheimer, 45.748112°N 4.89819°E (un. = 6 m), 29.VIII.2022, 1 specimen, photo by Bernard Bal (IN); idem, between Pont Pasteur and Pont de la Mulatiére, 45.7315°N 4.8184°E (un. = 16 m), 5.VIII.2021, 1 specimen, photo by Joeri Van Beeck (OB); idem, Rue Jacques Monod, 45.730428°N 4.828062°E (un. = 14 m), 11.IX.2021, 1 specimen, photo by “chjp” (IN); idem, Av. Leclerc, 45.738881°N 4.825333°E (un. not recorded), 28.VI.2022, 1 specimen, photo by Julien Lepage (IN); Villeurbanne, 45.772269°N 4.895001°E (un. = 20 m), 17.VIII.2018, 1 specimen, photo by “s2lyon” (IN); idem, Parc de Gerland, 45.724328°N 4.822945°E (un. = 4 m), 4.VII.2022, 1 specimen, photo by “chjp” (IN); Millery, 45.627555°N 4.784145°E (un. = 65 m), 29.VIII.2021, 1 specimen, photo by “aimji” (IN); Haute-Savoie, Évian-les-Bains, Place du Port, 46.4017°N 6.5960°E (un. = 25 m), 25.VII.2021, 1 specimen, photo by Jan Plaggenmarsch (OB); Cruseilles, 46.032935°N 6.110762°E (un. = 7 m), 2.VII.2022, 1 specimen, photo by “barassi” (IN); Sevrier, Lac d’Annecy, 45.870572°N 6.144°E (un. = 4 m), 30.VII.2022, 1 specimen, photo by “chjp” (IN); Savoie, Aix-les-Bains, Jardin Vagabond, 45.70792°N 5.89105°E (un. = 148 m), 24.VII.2022, 1 specimen, photo by “seriousdom” (IN); Le Bourget-du-Lac, 45.645866°N 5.875667°E (un. = 244 m), 4.IX.2021, 1 specimen, photo by “williamdomenge9” (IN); Nances, Lac d’Aiguebelette, 45.5739°N 5.8015°E (un. = 10 m), 2.VIII.2015, 1 specimen, photo by Pierre Bonmariage (OB); Isère, Saint-Pierred’Entremont, 45.3985°N 5.8418°E (un. = 17 m), 6.VII.2011, 1 specimen, photo by Els en Ben Prins (OB); Sassenage, 45.213687°N 5.657812°E (un. = 349 m), 21.VI.2020, 1 specimen, photo by “loicos1638” (IN); Haute-Loire, Retournac, 45.2093°N 4.0031°E (un. = 10 m), 5.VIII.2018, 1 specimen, photo by Sam Gobin (OB); Ardèche, Balazuc, 44.5120°N 4.3733°E (un. = 10 m), 12.IX.2019, 1 specimen, photo by Tom Vermeire (OB); Saint-Péray, 44.945219°N 4.837013°E (un. 61 m), 20.VIII.2020, photo by “charles_5” (IN); Privas, 44.73579°N 4.597406°E (un. not recorded), 13.VIII.2020, 1 specimen, photo by Ralph Martin (IN); Vals-les-Bains, 44.6572°N 4.3679°E (un. = 15 m), 18.VII.2022, 1 specimen, photo by “illenoy” (OB); Largentière, 44.519222°N 4.310748°E (un. not recorded), 12.VIII.2018, 1 specimen, photo by Eric Verna (IN); Sampzon, 44.4228°N 4.3226°E (un. = 4 m), 23.VI.2022, 1 specimen, photo by “Robert P” (OB); idem, 44.4112°N 4.3481°E (un. = 5 m), 26.VII.2021, 1 specimen, photo by “kvangoidsenhoven” (OB); idem, 44.4228°N 4.3226°E (un. = 4 m), 23.VI.2022, 1 specimen, photo by “Robert P” (OB); Vallon-Pont-d’Arc, 44.4029°N 4.3867°E (un. = 5 m), 31.VIII.2021, 1 specimen, photo by An Defloor (OB); Les Assions, 44.428588°N 4.163003°E (un. = 4 m), 15.VIII.2022, 1 specimen, photo by “nattchen” (IN); Beaulieu, 44.3610°N 4.2317°E (un. = 8 m), 19.VII.2022, 1 specimen, photo by Manu Van Haute (IN); Saint-Paul-le-Jeune, 44.3427°N 4.1497°E (un. = 8 m), 29.VII.2021, 1 specimen, photo by Jeroen Maes (OB); Saint-Martin-d’Ardèche, Rue d’Andronne, 44.302524°N 4.563305°E (un. = 9 m), 18.VIII.2021, 1 specimen, photo by “siemi” (IN); Drôme, Bourg-lès-Valence, 44.95°N 4.8833°E (un. not recorded), 2.VII.2005, 1 specimen with nests, photo by “Alain.M” (GMI); Chantemerle-lès-Grignan, 44.412045°N 4.850487°E (un. 24 m), 20.VIII.2020, 1 specimen, photo by Laurent Vallotton (IN); Dieulefit, 44.525789°N 5.06709°E (un. = 19 m), 19.VI.2022, 1 specimen, photo by “liamnell” (IN); Beaufort-sur-Gervanne, La Tuilerie, 44.7812°N 5.1426°E (un. = 5 m), 18.VIII.2019, 1 specimen, photo by J. Wielink (OB); Pierrelongue, 44.2453°N 5.2193°E (un. = 10 m), 10.VIII.2018, 1 specimen, photo by Pierre Bonmariage (OB). Bourgogne-Franche-Comté: Doubs, Besançon, 47.2307189°N 6.0236247°E (un. not recorded), 242 m, 22.IX.2013, 1 specimen, photo by Luc Bettinelli (GMI); Broye, between Poisot and Chapey, 46.8711°N 4.3376°E (un. = 25 m), 14.VI.2022, photo by Lydia Devriese (OB); Nièvre, Luzy, La Gagére, 46.8169°N 4.0567°E (un. = 25 m), 1.VIII.2021, 1 specimen, photo by Jac Hamers (OB); Saône-et-Loire, Épinac, 46.996103°N 4.516159°E (un. 77 m), 21.VII.2021, 1 specimen, photo by “vioplo” (IN); Épervans, 46.734206°N 4.903107 E (un. not recorded), 3.VII.2022, 1 specimen, photo by “chenneviere” (IN); La Guiche, Les Janins, 46.540057°N 4.470563°E (un. = 10 m), 1.VII.2019, 1 specimen, photo by “piksou139” (IN); Mâcon, 46.3688°N 4.8439°E (un. not recorded), 2.VII.2010, 7 specimens, photo by Luc Verhelst (OB); idem, 46.3688°N 4.8436°E (un. not recorded), 11.VII.2010, 1 specimen, photo by Luc Verhelst (OB). Corsica: Haute-Corse, Calvi, 42.566°N 8.7571°E (un. not recorded), 1.VIII.2005, 1 specimen photographed but reported as some individuals, photo by “Lauzias” (GMI); Lucciana, 42.5278°N 9.4419°E (un. = 23 m), 25.VI.2022, 1 specimen, photo by Ruud Boom (OB); near Calacuccia, 42.328753°N 9.026411°E (un. = 977 m), 23.VIII.2019, 1 specimen, photo by “alexblanke” (IN); Corse-du-Sud, Ota, 42.25799°N 8.745154°E (un. 100 m), 25.VII.2020, 1 specimen, photo by Emily Dent (IN); Sollacarò, 41.7458°N 8.8661°E (un. = 100 m), 21.VIII.2019, 3 specimens, photo by Herman Blockx (OB); Zonza, Camping California, 41.665798°N 9.378532°E (un. not recorded), 16.VII.2022, 1 specimen, photo by Stefano Gellato (IN); Bonifacio, 41.389635°N 9.157481°E (un. 1420 m), 15.VII.2020, 1 specimen, photo by “antoinedl” (IN); idem, 41.3859°N 9.1551°E (un. not recorded), 25.VI.2022, 1 specimen, photo by “Nest” (OB); Plage de Cala Sciumara, 41.3679°N 9.1983°E (un. = 25 m), 27.VIII.2022, 12 specimens, photo by Patrick Gros (OB). Grand Est: Metz, Metz, Place de la Comédie, 49.121435°N 6.173007°E (un. = 37 m), 25.VII.2022, 1 specimen, photo by “leafyh” (IN); Bas-Rhin, Scharrachbergheim-Irmstett, Rue de la Féodalité, 48.595975°N 7.491157°E (un. = 24 m), 6.IX.2020, 1 specimen, photo by “tritscherfj” (IN); Strasbourg, quai de Bateliers, 48.579533°N 7.750564°E (un. = 3 m), 29.VIII.2022, 2 specimens, photo by Valentin de Mazancourt (IN); idem, Rue de Port du Rhin, 48.576338°N 7.787895° (un. not recorded), 17.VI.2021, 1 specimen, photo by Lucas Scholtes (IN); idem, quai du Woerthel, 48.5799°N 7.7400°E (un.= 40 m), 27.VIII.2021, 1 specimen, photo by “viennaverbeke” (OB); idem, Port du Rhin, Rue de la Musau, 48.561338°N 7.77919°E (un. = 4 m), 29.VII.2022, 1 specimen, photo by Simon Diemer (IN); Illkirch-Graffenstaden, 48.5320096°N 7.7145995°E (un.not recorded), 10.VI.2017, 1specimen,photo by Jean-Sébastien Carteron(GMI);Ebersmunster, Rue de Fossé, 48.309837°N 7.525702°E (un. not recorded), 31.VII.2022, 1 specimen, photo by Jean Gaget (IN); Haut-Rhin, Colmar, Rue de la Poissonnerie, 48.0741°N 7.3575°E (un. = 5 m), 30.VII.2022, 1 specimen, photo by Jonas Joos (OB); Fessenheim, 47.913563°N 7.528166°E (un. not recorded), 27.VIII.2019, 1 specimen, photo by “binbin68” (IN); Kembs, 47.6931°N 7.5007°E (un. = 32 m), 28.VIII.2021, 1 specimen, photo by Thomas Ryckmans (OB). Hauts-de-France: Nord, Saultain, 50.336203°N 3.57929°E (un. = 8 m), 30.VII.2018, 1 specimen, photo by Pierre Meignan (IN). Île-de-France: Paris, 4th arrondissement of Paris, Square Albert-Schweitzer, 48.853667°N 2.358283°E (un. not recorded), 16.VIII.2020, 1 specimen, photo by “thadeus72” (IN); idem, 5th arrondissement of Paris, RER Saint-Michel – Notre-Dame, 48.8532°N 2.3464°E (un. = 200 m), 27.VII.2022, 1 specimen, photo by Niels Zonneveld (OB); idem, 7th arrondissement of Paris, Rue de Grenelle, 48.8581°N 2.3142°E (un. = 6 m), 24.VIII.2022, 1 specimen, photo by “Egon” (OB). Nouvelle-Aquitaine: Pyrénées-Atlantiques, Andrein, 43.390006°N - 0.899617°E (un. not recorded), 21.VIII.2020, 1 specimen, photo by Misia Clive (IN); Mendive, 43.047192°N - 1.074498°E (un. = 4 m), 5.VIII.2019, 1 specimen, photo by Adrien Nouailhane (IN); Larrau, 43.0191°N - 0.9618°E (un. = 10 m), 19.IX.2018, 1 specimen, photo by Rolf Van Steenkiste (OB); Landes, Gastes, 44.3146°N - 1.1764°E (un. = 25 m), 2.VIII.2022, 1 specimen, photo by V. Engelbos (OB); Vielle-Saint-Girons, 43.95°N - 1.3°E (un. not recorded), 5 m, 18.VIII.2010, 1 specimen, photo by “Phil” (GMI); Saint-Paul-lès-Dax, 43.726848°N - 1.06779°E (un. not recorded), 20.VII.2022, 1 specimen, photo by “tomtyne” (IN); Ondres, 43.5687°N - 1.4586°E (un. not recorded), 17.VIII.2021, 1 specimen, photo by “Ocmer” (OB); Saint-Martin-de-Seignanx, Route Océane, 43.544675°N - 1.387303 °E (un. = 5 m), 25.VII.2021, 1 specimen, photo by Peter Guy (IN); Corrèze, Beaulieu-sur-Dordogne, 44.975968°N 1.838067°E (un. = 6 m), 23.VIII.2020, 1 specimen, photo by “lnjoo” (IN); Gironda, Saint-Seurin-sur-l’Isle, 45.0166°N - 0.0032°E (un. not recorded), 3.VIII.2022, 1 specimen, photo by “Silke” (OB); Dordogne, Nanteuil-Auriac-de-Bourzac, 45.3686°N 0.2592°E (un. = 25 m), 5.VIII.2021, 1 specimen, photo by Stijn Cinjaere (OB); Brantôme-en-Périgord, 45.3638°N 0.6528°E (un. not recorded), 15.VIII.2021, 1 specimen, photo by Martijn Back (OB); Eyvirat, La Jugie, 45.326554°N 0.723756°E (un. = 5 m), 25.VIII.2019, 1 specimen, photo by “marol31” (IN); Montpon-Ménestérol, 45.0552063°N 0.1724092°E (un. not recorded), 14.VIII.2017, 1 specimen, photo by “Orchallan” (GMI); Saint-Léon-sur-Vézère, La Rebeyrolle, 45.0014°N 1.0740°E (un. not recorded), 14.VI.2021, 1 specimen, photo by Els van Steenbergen (OB); Montignac, Rue Tournon, 45.0656°N 1.1638°E (un. not recorded), 7.VII.2021, 1 specimen, photo by “Pieter” (OB); Les Eyzies-de-Tayac-Sireuil, La Combe, 44.9133°N 1.0156°E (un. not recorded), 10.VIII.2021, 1 specimen, photo by “Kev B” (OB); Sarlat-la-Canéda, 44.9016°N 1.1826°E (un. = 25 m), 2.X.2021, 1 specimen, photo by Peter De Kraker (OB); Siorac-en-Périgord, 44.8256°N 0.9900°E (un. = 5 m), 16.VII.2022, 1 specimen, photo by Remko Wijling (OB); Haute-Vienne, Saint-Victurnien, near Avenue du Pont, 45.8753°N 1.0131°E (un. not recorded), 7.VIII.2022, 1 specimen, photo by Ingrid van den Berg (OB); Glandon, locality La Petite Valade, 45.4660°N 1.2028°E (un. = 10 m), 23.V.2021, 1 specimen, photo by “Benoît” (OB); Lot-et-Garonne, Sainte-Livrade-sur-Lot, 44.3974°N 0.5820°E (un. = 6 m), 20.VIII.2022, 1 specimen, photo by “joris” (OB); Courbiac, 44.3768416°N 1.0184888°E (un. not recorded), 151 m, 27.VI.2012, 1 specimen, photo by Brigitte Ulmer (GMI); Charente, Angoulême, Rue de la Fontaine du Lizier, 45.654044°N 0.159007°E (un. = 5 m), 1 specimen, photo by “gerbi” (IN); Barbezières, 45.90509°N - 0.088962°E (un. = 2 m), 23.VIII.2018, 1 specimen, photo by Paul Lewis (IN); Brossac, 45.3300°N - 0.0572°E (un. = 19 m), 30.VI.2021, 1 specimen, photo by Nicky Verlinden (OB); Deux-Sèvres, Échiré, Rue de Marécheaux, 46.388054 N - 0.408611°E (un. not recorded), 22.VIII.2019, 1 specimen, photo by “gbo” (IN). Occitanie: Lot, Bélaye, 44.455914°N 1.211028°E (un. = 6 m), 24.VII.2022, 1 specimen, photo by “nykfel”; Lozère, Meyrueis, 44.1930°N 3.4047°E (un. = 25 m), 15.VI.2017, 1 specimen, photo by Peter De Kraker (OB); idem, near Salvinsac, 44.1966°N 3.4487°E (un. = 17 m), 28.VII.2020, 1 specimen, photo by “sander01” (OB); Vebron, 44.240478 N 3.5785°E (un. = 4 m), 25.VII.2022, 1 specimen, photo by Ivo Walsmit (IN); Bédouès-Cocurès, 44.3451°N 3.6099°E (un. = 25 m), 6.VIII.2020, 1 specimen, photo by Sam Gobin (OB); Vialas, 44.3247°N 3.9119°E (un. = 25 m), 21.VI.2021, 1 specimen, photo by Hannes Stevens (OB); Aveyron, Capdenac-Gare, 44.567977°N 2.04493°E (un. = 17 m), 26.VIII.2020, 1 specimen, photo by “teoch” (IN); Salvagnac-Cajarc, 44.4389°N 1.8760°E (un. = 50 m), 12.VIII.2016, 1 specimen, photo by Harm Alberts (OB); Saint-Rome-de-Cernon, 44.0114°N 2.9658°E (un. not recorded), 17.VII.2021, 1 specimen, photo by Marc Rombouts (OB); idem, Raspaillac, 43.9990°N 2.9710°E (un. = 34 m), 17.VII.2021, 1 specimen, photo by Kris Bracke (OB); Gard, Aujac, 44.3393°N 4.0406°E (un. = 100 m), 1 specimen, photo by Ruud and Betty van Middelkoop (OB); Courry, near Grotte de la Cocalière, 44.310246°N 4.167777°E (un. = 8 m), 13.IX.2020, 1 specimen, photo by “lantinantoine” (IN); Saint-Florent-sur-Auzonnet, 44.2382°N 4.1202°E (un. = 5 m), 5.VIII.2022, 2 specimens, photo by Jean-François Meersman (OB); Cornillon, near Saint-Gély, 44.2072°N 4.4889°E (un. = 49 m), 4.VIII.2022, 1 specimen, photo by “funnie” (OB); Rousson, 44.1948°N 4.1422°E (un. = 25 m), 22.VII.2022, 1 specimen, photo by Henk Wallays (OB); Générargues, 44.0730°N 3.9791°E (un. not recorded), 25.VII.2022, 1 specimen, photo by Elise Bakker (OB); Anduze, Chemin de Recoulin, 44.068783°N 3.971745°E (un. = 8 m), 27.VIII.2020, 1 specimen, photo by Katrin Simon (IN); Saint-Hippolyte-de-Montaigu, Rue des Garrigues, 44.032546°N 4.495753°E (un. = 5 m), 1.VIII.2019, 1 specimen, photo by Barbara Cox (IN); Argilliers, Impasse des Cros, 43.979353°N 4.494111°E (un. = 5 m), 2.VIII.2021, 1 specimen, photo by “tommaso78” (IN); Sardan, 43.8667°N 4.05°E (un. not recorded), 6.VIII.2021, 1 specimen, photo by Liliane Delattre (GMI); Nages-et-Solorgues, 43.789777°N 4.251392°E (un. = 4 m), 29.VII.2021, 1 specimen, photo by Pierre Kerner (IN); Bouillargues, Rue du Mistral, 43.800188°N 4.421449°E (un. 107 m), 5.VI.2021, 1 specimen, photo by “loicalegot” (IN); Uzès, 44.0094°N 4.4258°E (un. = 25 m), 26.VII.2020, 1 specimen, photo by Stijn Cinjaere (OB); Aigues-Mortes, 43.5873°N 4.2093°E (un. = 10 m), 12.VI.2017, 1 specimen, photo by Arno van Stipdonk (OB); Bouches-du-Rhône, Saint-Martin-de-Crau, Chemin de la Rispaulide, 43.648613°N 4.756122°E (un. = 65 m), 1 specimen, photo by “majipi” (IN); Fontvieille, Chemin du Grand Clos, 43.716457°N 4.69542°E (un. = 4 m), 18.VIII.2018, 1 specimen, photo by “tonydanza” (IN); Arles, 43.681309°N 4.630534°E (un. = 310 m), 30.VI.2021, 1 specimen, photo by Benoît Segerer (IN); Réserve Naturelle des Marais du Vigueirat, 43.5147°N 4.7876°E (un. = 25 m), 9.VIII.2017, 1 specimen, photo by Philippe Stroot (OB); Saintes-Maries-de-la-Mer, 43.45026°N 4.419657°E (un. = 31 m), 10.VIII.2020, 1 specimen, photo by Andrew Lamb (IN); Gémenos, 43.3°N 5.6333°E (un. not recorded), 5.VII. 2013, 130 m, 1 specimen, photo by Stéphane Bonhomme (GMI); Gémenos, Av. du Pic de Bertagne, 43.281642°N 5.616379°E (un. not recorded), 18.VII.2022, 1 specimen, photo by Yohann Flavier (IN); Hérault, Brissac, 43.8787°N 3.7004°E (un. = 1000 m), 24.VII.2015, 10 specimens, photo by Eric Walravens (OB); Notre-Dame-de-Londres, 43.83864°N 3.75701°E (un. = 4 m), 8.IX.2020, 1 specimen, photo by Pierre-Henri Fabre (IN); Viols-le-Fort, 43.742805°N 3.709007°E (un. = 22 m), 1.VIII.2020, 1 specimen, photo by David Renoult (IN); Restinclières, 43.727049°N 4.032393°E (un. = 8 m), 21.VIII.2021, 1 specimen, photo by Jérémie Février (IN); Teyran, 43.697979°N 3.926785°E (un. not recorded), 2.VIII.2021, 1 specimen, photo by Julia Wittmann (IN); Lunel, 43.6833°N 4.1333°E (un. not recorded), 50 m, 22.VII.2010, 3 specimens, photo by Dominique Martiré (GMI); Lunel-Viel, 43.6722093°N 4.0899586°E (un. not recorded), 4.VIII.2017, 1 specimen, photo by Jean-Jacques Porteneuve (GMI); Montpellier, Vert-Bois, 43.638584°N 3.864005°E (un. = 2 m), 2.IX.2019, 1 specimen, photo by “osoandino” (IN); idem, Boutonnet, Rue Edouard Marsal, 43.62436°N 3.867999°E (un. not recorded), 1.VII.2022, 1 specimen, photo by “gaetanbrn” (IN); Lattes, 43.562798°N 3.908225°E (un. = 2 m), 8.VIII.2014, 1 specimen, photo by Simon Popy (IN); Fabrègues, 43.550282°N 3.80771°E (un. not recorded), 31.VIII.2019, 1 specimen, photo by Benoit Nabholz (IN); Mèze, 43.42617°N 3.603649°E (un. = 36 m), 19.VI.2022, 1 specimen, photo by Mathias Prat (IN); Frontignan, 43.461724°N 3.820937°E (un. = 31 m), 9.IX.2018, 1 specimen, photo by Benoit Nabholz (IN); Colombiers, 43.3140°N 3.1427°E (un. not recorded), 9.VII.2010, 1 specimen, photo by Luc Verhelst (OB); Aude, Bages, Etang de Saint-Paul, 43.1073°N 2.9758°E (un. = 10 m), 28.VIII.2018, 1 specimen, photo by Frans van Antwerpen (OB); Gruissan, Port Rive Gauche, 43.108654°N 3.099415°E (un. = 8 m), 9.X.2021, 1 specimen, photo by “milanpestalozzi” (IN); Cabrespine, 43.356207°N 2.461591°E (un. = 754 m), 17.VII.2018, 1 specimen, photo by “marketaz” (IN); Raissac-sur-Lampy, 43.267814°N 2.165766°E (un. = 122 m), 4.VIII.2017, 1 specimen, photo by Eric Verna (IN); Puilaurens, Aigues-Bonnes locality, 42.7950°N 2.3342°E (un. = 10 m), 12.VII.2015, 1 specimen, photo by Ruud and Betty van Middelkoop (OB); Tarn, Marnaves, 44.0975°N 1.8892°E (un. = 10 m), 26.VIII.2015, 1 specimen, photo by Harm Alberts (OB); Cuq-Toulza, 43.570743°N 1.897606°E (un. = 15 m), 23.VIII.2019, 1 specimen, photo by“pcaze81” (IN); idem, 43.573236°N 1.895443°E (un.= 2 m), 1.VIII.2021, 1 specimen, photo by “pcaze81” (IN); Cambounet-sur-le-Sor, 43.580276°N 2.134332°E (un. = 18 m), 5.VI.2022, 1 specimen, photo by “pcaze81” (IN); Haute-Garonne, Blagnac, Rue Dieudonné Costes, 43.6309°N 1.3804°E (un. = 33 m), 2.VIII.2021, 1 specimen, photo by “Anna C.” (OB); Toulouse, Négreneys, 43.6193°N 1.443929°E (un. = 43 m), 10.VIII.2020, 1 specimen, photo by Ivan Paz-Vinas (IN); Mons, 43.613201°N 1.573325°E (un. = 1 m), 17.VII.2022, 1 specimen, photo by “MiKL_Nicolas” (IN); Saint-Orens-de-Gameville, 43.5494665°N 1.5260243°E (un. not recorded), 27.VI.2017, 1 specimen, photo by Hélène Laviron (GMI); Cugnaux, Rue de la Marne, 43.547256°N 1.348371°E (un. = 6 m), 26.VIII.2013, 1 specimen, photo by “raboul” (IN); Goyrans, 43.490681°N 1.422697°E (un. = 1150 m), 12.IX.2020, 1 specimen, photo by Ivan Paz-Vinas (IN); Villefranchede-Lauragais, 43.4°N 1.7333°E (un. not recorded), 28.VI. 2013, 200 m, 1 specimen, photo by Genevieve Buxin (GMI); Gers, Condom, 43.9748°N 0.3691°E (un. = 34 m), 11.VIII.2012, 1 specimen, photo by Ben Sleeman (OB); Cologne, 43.721125°N 0.976344°E (un. = 420 m), 13.VIII.2019, 1 specimen, photo by “daquitaine” (IN); Ariège, Montjoie-en-Couserans, Griou locality, 43.0374°N 1.1671°E (un. = 10 m), 13.VII.2013, 1 specimen and nests, photo by Toine Rommers (OB); Prat-Bonrepaux, 43.0333°N 1.0167°E (un. not recorded), 300 m, 28.VIII.2013, 1 specimen, photo by Jean Maurette (GMI); Rimont, 42.994346°N 1.280031°E (un. 84 m), 28.IX.2020, 1 specimen, photo by “michael_t” (IN); Pyrénées-Orientales, Caudiès-de-Fenouillèdes, 42.8117°N 2.3783°E (un. = 10 m), 1.VII.2019, 1 specimen, photo by D. Moonen (OB); Argelès-sur-Mer, 42.5671°N 3.0285°E (un. = 7 m), 29.VII.2022, 1 specimen, photo by Eline Kooij (OB); Saint-André, 42.552°N 2.9713°E (un. not recorded), 35 m, 14.VI.2010, 1 specimen, photo by “Canigou” (GMI); Banyuls-sur-Mer, 42.4782°N 3.1440°E (un. = 18 m), 9.VII.2020, 1 specimen, photo by Philippe Deflorenne (OB); Saint-Cyprien, Chemin de las Clottes, 42.615653°N 3.018197°E (un. = 10 m), 29.VI.2019, 1 specimen, photo by “franccomtois” (IN). Provence-Alpes-Côte d’Azur: Bouches-du-Rhône, Saintes-Maries-de-la-Mer, 43.4521°N 4.4291°E (un. not recorded), 5.X.2008, 2 m, 1 specimen, photo by Dominique Bataille (GMI); Bouc-Bel-Air, 43.4535°N 5.4086°E (un. = 10 m), 24.VI.2011, 1 specimen, photo by Maarten Schurmans (OB); Roquevaire, 43.3471°N 5.6090°E (un. = 10 m), 9.VIII.2019, 1 specimen, photo by René van Bruggen (OB); Martigues, near La Couronne, 43.3331°N 5.0706°E (un. = 25 m), 29.VII.2020, 1 specimen, photo by Pieter Bruijsten (OB); Var, Aiguines, 43.75218°N 6.248139°E (un. = 1570 m), 2.VIII.1984, 1 specimen, photo by Lupoli Roland (IN); Carnoules, 43.299774°N 6.167135°E (un. = 98 m), 12.VI.2022, 1 specimen, photo by Robin Leyret (IN); Trans-en-Provence, Rue Nationale, 43.5032°N 6.4868°E (un. = 25 m), 2.VII.2020, 1 specimen, photo by Fabien Bonte (OB); Le Luc, 43.370972°N 6.292785°E (un. not recorded), 31.VII.2022, 1 specimen, photo by Mathis Garro (IN); Ramatuelle, near L’Escalet, 43.2069°N 6.6424°E (un. = 10 m), 22.VII.2011, 1 specimen, photo by Erwin Walstra (OB); Hyères, 43.1167°N 6.1167°E (un. not recorded), 100 m, 23.VIII.2005, 1 specimen, photo by Jean-Philippe Mathieu (GMI); La Seyne-sur-Mer, 43.0923276°N 5.8670405°E (un. not recorded), 15 m, 17.VIII.2019, 1 specimen, photo by “Liboupat” (GMI); Marseille, 43.2310360°N 5.4394314°E (un. not recorded), 139 m, 2.VII.2019, 1 specimen, photo by Bernard Ginesy (GMI); Alpes-de-Haute-Provence, Digneles-Bains, 44.096488°N 6.238709°E (un. not recorded), 9.VI.2022, 1 specimen, photo by Louis Vivier (IN); Sisteron, 44.172867°N 5.928798°E (un. = 13 m), 12.VI.2022, 1 specimen, photo by Raphaël Colombo (IN); Alpes-Maritimes,Beuil, 44.1059°N 6.9844°E (un. = 34 m), 20.VII.2021, 1 specimen, photo by Pierre Bonmariage (OB); Saint-André-de-la-Roche, Chemin de Rimiez, 43.73907°N 7.281978°E (un. = 9 m), 12.VII.2022, 1 specimen, photo by “thf06” (IN); Nice, Av. du Marechal Lyautey, 43.720942°N 7.28605°E (un. = 8 m), 27.VI.2020, 2 specimens, photo by “bernardjulien06000” (IN); idem, 43.713979°N 7.282605°E (un. = 31 m), 6.VI.2017, 1 specimen, photo by Fabien Piednoir (IN); Cannes, Jetée Albert Edouard, 43.55°N 7.0167°E (un. not recorded), 9.VIII.2011, 1 specimen, photo by Jean-Claude Jamoulle (GMI); idem, Lérins Islands, Île Saint-Honorat, 43.506145°N 7.045561°E (un. = 119 m), 1 specimen, photo by “raptorr575” (IN). Vaucluse, Vaisonla-Romaine, 44.2392°N 5.0879°E (un. = 5 m), 15.VIII.2022, 1 specimen, photo by “Bram” (OB); Malaucène, 44.1712°N 5.1274°E (un. = 100 m), 8.IX.2012, 1 specimen, photo by Alrik Hoven (OB); Bédoin, Rue de Mont-Ventoux, 44.1341°N 5.3017°E (un. = 272 m), 16.VII.2015, 1 specimen, photo by Jan Vanhoutte (OB); Orange, 44.1277°N 4.8321°E (un. = 8 m), 28.VII.2022, 1 specimen, photo by “Kristof” (OB); Loriol-du-Comtat, Chemin de Barriot, 44.072359°N 5.014302°E (un. = 65 m), 7.VIII.2022, 1 specimen, photo by “jibbito” (IN); Avignon, Jardin du Rocher des Doms, 43.9528°N 4.8068°E (un. = 9 m), 27.VI.2022, 1 specimen, photo by Frank Berbers (OB).
BELGIUM. Wallonia: Liège, Parc Naturel Hautes Fagnes-Eifel, 50.58601°N 6.128632°E (un. not recorded), 12.VII.2020, 1 specimen, photo by Chris Van Bladel (IN).
NETHERLANDS. Flevoland: Lelystad, Lelystad, 52.5094°N 5.4308°E (un. = 10 m), 17.VIII.2018, 1 specimen, photo by Jeroen de Rond (WNL). Limburg: Venlo, Venlo, 51.3764°N 6.1277°E (un. = 10 m), 12.IX.2020, 1 specimen, photo by Fons Verheyde (WNL); Roerdalen, Montfort, 51.1300°N 5.9250°E (un. = 100 m), 13.VIII.2010, 1 specimen, photo by Huub Schmitz (WNL). North Brabant: Maashorst, Uden, 51.6532°N 5.6213°E (un. = 25 m), 25.VII.2021, 1 specimen, photo by Stefan van Schaik (WNL). North Holland: Amsterdam, Amsterdam, Buiksloterweg, 52.38768°N 4.914453°E (un. = 65 m), 25.IX.2021, 1 specimen, photo by Hannah Van Dijck (IN). Overijssel: Oldenzaal, Oldenzaal, 52.2864°N 6.9238°E (un. = 100 m), 15.VII.2018, 1 specimen, photo by Martin Bonte (WNL); Hof van Twente, Goor, 52.2362°N 6.5760°E (un. = 15 m), 6.IX.2021, 1 specimen, photo by André Mensink (WNL).
GERMANY. Baden-Württemberg: Mannheim,Feudenheim, 49.48649°N 8.531681°E (un.= 1.41km), 2.VIII.2022, 1 specimen, photo by Klaus Bohn (IN); Rastatt, Rastatt, Ottersdorfer Str., 48.856863°N 8.20022°E (un. = 8 m), 18.VII.2020, 1 specimen, photo by “hobu” (IN); idem, near bridge on Richard-Wagner-Ring, 48.8696°N 8.1996°E (un. = 16 m), 26.VII.2022, 6 specimens, photo by “landsmann” (OB); Gaggenau, 48.8053°N 8.3183°E (un. not recorded), 28.VIII.2022, 1 specimen, photo by “Schmeiser91” (OB); Emmendingen, 48.121614°N 7.851431°E (un. = 6.07 km), 20.VI.2022, 1 specimen, photo by Franz Aiple (IN); Ortenaukreis, Offenburg, Durbacher Str., 48.485704°N 7.977818°E (un. = 40 m), 24.VII.2021, 1 specimen (Fig. 7), photo by Corinna Herr (IN); Europa-Park, 48.264469°N 7.721184°E (un. not recorded), 18.VIII.2020, 1 specimen, photo by “snoutch” (IN); Freiburg im Breisgau, Appenweier, 48.540982°N 7.973553°E (un. = 28 m), 15.VIII.2022, 1 specimen, photo by Micha Baum (IN); Freiburg im Breisgau, 48.007786°N 7.855605°E (un. = 35 m), 10.VIII.2022, 1 specimen, photo by “kaka_po” (IN); idem, 48.003665°N 7.853374°E (un. = 15 m), 24.VIII.2020, 1 specimen, photo by “felixf” (IN); idem, 47.990022°N 7.826584°E (un. = 44 m), 31.VIII.2019, 1 specimen, photo by “fredstr” (IN); idem, 47.991089°N 7.864861°E (un. = 65 m), 31.VIII.2019, 1 specimen, photo by “fredstr” (IN); idem, 47.96963°N 7.816799°E (un. = 15 m), 20.VI.2020, 1 specimen, photo by Corinna Herr (IN); L̂rrach, Kandern, 47.713603°N 7.662183°E (un. = 24 m), 11.IX.2022, 1 specimen, photo by Wesley Shrader (IN); idem, Grenzach-Wyhlen, 47.5554°N 7.6579°E (un. = 5 m), 16.VII.2022, 1 specimen, photo by “susumumpitz” (OB); Bad Säckingen, 47.551178°N 7.950228°E (un. = 4 m), 28.VIII.2022, 1 specimen, photo by Luca Mehlhorn (IN); Breisgau-Hochschwarzwald, Neuenburg am Rhein, 47.8145°N 7.5588°E (un. = 14 m), 7.VIII.2022, 1 specimen, photo by “Roy” (OB). North Rhine-Westphalia: Essen, Essen, Krupp Park, 51.4630°N 6.9876°E (un. = 10 m), 3.VIII.2022, 1 specimen, photo by Wilfried van de Sand (OB). Schleswig-Holstein: Kiel, Kiel, 54.328209°N 10.145628°E (un. = 488 m), 27.VI.2006, 1 specimen, photo by David Foster (IN).
SWITZERLAND. Basel-Landschaft: Liestal, Frenkendorf, 47.511059°N 7.723458°E (un. = 9 m), 2.IX.2021, 1 specimen, photo by “ullukai” (IN); idem, 47.505612°N 7.713487°E (un. = 13 m), 17.VIII.2022, 1 specimen, photo by “perowu” (IN). Basel-Stadt: Basel, Aeschenvorstadt, 47.5530°N 7.5930°E (un. = 5 m), 7.VIII.2022, 1 specimen, photo by Arthur van Ooijen (OB); idem, Scḩtzenmattpark, 47.552853°N 7.574655°E (un. = 214 m), 6.VI.2022, 1 specimen, photo by Kalyta Kateryna (IN); idem, St Louis Grenze, 47.576388°N 7.572989 °E (un. = 132 m), 17.VIII.2020, 1 specimen, photo by “laurenthoff” (IN); idem, Feldbergstrasse, 47.567062°N 7.59557°E (un. not recorded), 22.V.2021, 1 specimen, photo by Samuel B̧ttler (IN); idem, 47.566722°N 7.595614°E (un. = 220 m), 27.V.2020, 1 specimen, photo by Jens Oldenburg (IN); idem, 47.566806°N 7.59609°E (un. = 17 m), 10.VII.2021, 1 specimen, photo by Jens Oldenburg (IN). Bern: Bernese Jura, Schelten, 47.334355°N 7.552258°E (un. = 8 m), 1.VII.2022, 1 specimen, photo by David Scinto (IN). Fribourg: Broye, Estavayer, 46.851322°N 6.848478°E (un. = 3 m), 10.VIII.2021, 1 specimen, photo by Markus Greiner (IN). Geneva: Versoix, Route de Sauverny, 46.285022°N 6.161161°E (un. = 6 m), 26.V.2020, 1 specimen, photo by “morgs” (IN); Meyrin, 46.236312°N 6.074099°E (un. = 2 m), 6.VIII.2021, 1 specimen, photo by “Victor B” (IN); Geneva, Jardin botanique de Genève, 46.227622°N 6.147064°E (un. = 30 m), 20.V.2019, 1 specimen, photo by “manuelramalho” (IN); idem, Parc Mon Repos, 46.219125°N 6.150714°E (un. = 5 m), 23.VII.2020, 1 specimen, photo by “timbr” (IN); idem, Eaux-Vives, 46.199958°N 6.159185°E (un. = 1 m), 31.VII.2015, 1 specimen, photo by Tommy Andriollo (IN); idem, 46.199626°N 6.158548°E (un. = 4 m), 31.VII.2014, 1 specimen, photo by Tommy Andriollo (IN); Jussy, 46.22606°N 6.252539°E (un. = 13 m), 25.III.2022, 1 specimen, photo by Tommy Andriollo (IN). Graubünden: Moesa, near San Bernardino, 46.4729°N 9.1852°E (un. = 25 m), 6.VII.2019, 1 specimen, photo by Johan van ‘t Bosch (OB). Neuchâtel: Cortaillod, Chemin du Bugnon, 46.9392°N 6.8446°E (un. = 5 m), 11.VIII.2022, 1 specimen, photo by “RalfTB” (OB). Ticino: Locarno, Muralto, Via S. Gottardo, 46.173612°N 8.80367°E (un. = 6 m), 26.VII.2022, 1 specimen, photo by “whoosh” (IN). Vaud: Broye-Vully, Valbroye, 46.76637°N 6.88648°E (un. = 17 m), 31.VII.2021, photo by Patrick Favre (IN); Ouest lausannois, Ecublens, 46.523467°N 6.567271°E (un. = 2 m), 7.IX.2019, 1 specimen, photo by Nicolas Zwahlen (IN); Lausanne, Lausanne, Le Mont-sur-Lausanne, 46.556618°N 6.636915°E (un. = 145 m), 20.V.2022, 1 specimen, photo by Patrick Favre (IN); idem, Musée et jardins botaniques cantonaux, 46.513589°N 6.622853°E (un. = not recorded), 17.VI.2022, 1 specimen, photo by Joan Casanelles-Abella (IN); idem, 46.513467°N 6.622903°E (un. = 4 m), 21.VI.2022, 1 specimen, photo by “timbr” (IN); idem, Avenue de l’Elysée, 46.509938°N 6.634183°E (un. = 26 m), 23.VIII.2022, 1 specimen, photo by “henrik_bfg” (IN); Lavaux-Oron, La Croix-sur-Lutry, 46.514639°N 6.703204°E (un. = 33 m), 10.VII.2019, 1 specimen, photo by “dgel” (IN); idem, 46.514506°N 6.703137°E (un. = 7 m), 14.VI.2022, 1 specimen, photo by “dgel” (IN); Bourg-en-Lavaux, 46.486971°N 6.724829°E (un. = 182 m), 15.VIII.2021, 1 specimen, photo by “lagartinha” (IN); Aigle, Aigle, 46.315172°N 6.968414°E (un. = 5 m), 21.VIII.2018, 1 specimen, photo by Nicolas Zwahlen (IN); Nyon, Founex, 46.334355°N 6.19313°E (un. = 6 m), 8.VII.2020, 1 specimen, photo by Dao Nguyen and James Hardcastle (IN); Réserve Naturelle “En Bucley”, 46.393128°N 6.139149°E (un. = 244 m), 4.VII.2020, 1 specimen, photo by Tom Brooks (IN).
AUSTRIA. Burgenland: Gols, Gols, 47.8976 °N 16.9087°E (un. not recorded), 5.VI.2022, 1 specimen, photo by Gilbert Hafner (OB); Rust, Rust, 47.800803°N 16.679618°E (un. = 7 m), 5.VII.2021, 3 specimens, photo by “frank007” (IN). Carinthia: Spittal an der Drau, M̂rtschach, 46.9086°N 12.9105°E (un. = 4 m), 2.VIII.2022, 1 specimen, photo by Herman Bouman (OB); Klagenfurt am Ŵrthersee, Europaschutzgebiet Lendspitz— Maiernigg, 46.611897°N 14.247835°E (un. = 168 m), 2.VII.2022, 1 specimen, photo by Peter Holub (IN). Lower Austria: Bernhardsthal, Bernhardsthal, 48.690484°N 16.875544°E (un. = 20 m), 16.VIII.2022, 1 specimen, photo by “kienberg” (IN); M̂dling, Perchtoldsdorf, 48.11552°N 16.264128°E (un. not recorded), 25.VIII.2021, 1 specimen, photo by Michael Knapp (IN); Laxenburg, 48.068243°N 16.357747°E (un. not recorded), 21.IX.2021, 1 specimen, photo by “eliegaget” (IN); idem, 48.0676°N 16.356819°E (un. = 65 m), 7.IX.2021, 1 specimen, photo by “seakmus” (IN); idem, Schlossplatz, 48.06754°N 16.356862°E (un. = 5 m), 20.IX.2022, 1 specimen, photo by “seakmus” (IN); Baden, Traiskirchen, 48.024087°N 16.280007°E (un. = 2 m), 5.VII.2021, 1 specimen, photo by Norbert Sauberer (IN); idem, 48.004327°N 16.270952°E (un. = 4 m), 1.X.2021, 1 specimen, photo by Martin A. Prinz (IN); idem, 48.004376°N 16.271485°E (un. = 2 m), 29.VII.2022, 1 specimen, photo by Martin A. Prinz (IN); idem, Schafflerhof-Siedlung, 47.991687°N 16.279729°E (un. = 19 m), 9.VIII.2022, 1 specimen, photo by “hype_zz666” (IN); Tribuswinkel, 48.000711°N 16.276246°E (un. = 4 m), 4.X.2021, 1 specimen, photo by Martin A. Prinz (IN); idem, 47.996937°N 16.302262°E (un. = 2 m), 20.VIII.2021, 1 specimen, photo by Martin A. Prinz (IN); idem, 48.001184°N 16.260363°E (un. = 8 m), 15.IX.2021, 1 specimen, photo by Martin A. Prinz (IN); idem, 48.007168°N 16.261296°E (un. = 2 m), 4 specimens, photo by “karl_a_berger_2511” (IN); Leobersdorf, 47.92837°N 16.216487°E (un. = 10 m), 12.VIII.2022, 1 specimen, photo by “yorelle” (IN). Styria: Graz, Graz, Grabenstrasse, 47.089293°N 15.432071° E (un. = 47 m), 31.VIII.2021, 1 specimen, photo by “birgit_” (IN); idem, near Schlossberg, 47.075917°N 15.450295°E (un. = 22 m), 17.X.2019, 1 specimen, photo by David Krainer (IN); idem, Glacisstrasse, 47.069378°N 15.447267°E (un. = 4 m), 26.VIII.2022, 1 specimen, photo by Margret Huss (IN); idem, Innere Stadt, 47.064767°N 15.435117°E (un. not recorded), 6.X.2021, 1 specimen, photo by “benikabuto” (IN); idem, Skatepark Augarten, 47.062003°N 15.435469°E (un. = 102 m), 21.VII.2021, 1 specimen, photo by “nikahause” (IN); idem, Sternäckerweg, 47.03936°N 15.464592°E (un. = 54 m), 1.IX.2021, 1 specimen, photo by “salsar” (IN); Şdoststeiermark, Poppendorf, 46.874348°N 15.85693°E (un. = 15 m), 5.VIII.2020, 1 specimen, photo by Sepp Schmid (IN); idem, 46.874816°N 15.857114°E (un. = 15 m), 23.VIII.2021, 1 specimen, photo by Sepp Schmid (IN); idem, 46.874493°N 15.8569°E (un. not recorded), 28.VII.2022, 1 specimen, photo by Sepp Schmid (IN); idem, 46.873958°N 15.856563°E (un. not recorded), 1.VIII.2022, 1 specimen, photo by Sepp Schmid (IN); idem, 46.874397°N 15.856842°E (un. not recorded), 11.VIII.2022, 1 specimen, photo by Sepp Schmid (IN); idem, 46.874693°N 15.856645°E (un. not recorded), 30.VIII.2022, 1 specimen, photo by Sepp Schmid (IN); idem, 46.874708°N 15.856655°E (un. not recorded), 11.IX.2022, 1 specimen, photo by Sepp Schmid (IN); idem, 46.874637°N 15.85668°E (un. not recorded), 14.IX.2022, 1 specimen, photo by Sepp Schmid (IN). Vienna: near Kleinhausgasse, 48.295208°N 16.384117°E (un. not recorded), 15.VIII.2022, 1 specimen, photo by “akirk9” (IN); Ziegelofenweg, 48.289325°N 16.391686°E (un. = 31 m), 20.VII.2021, 1 specimen, photo by Leonid Rasran (IN); Nussdorf, 48.274277°N 16.358066°E (un. = 61 m), 14.VIII.2021, 1 specimen, photo by Konrad Zobel (IN); idem, 48.272427°N 16.352508°E (un. = 85 m), 20.VIII.2022, 1 specimen, photo by Konrad Zobel (IN); Universität f̧r Bodenkultur Wien, 48.236876°N 16.33598°E (un. = 26 m), 7.IX.2021, 1 specimen, photo by “alsbach” (IN); Breitenlee, 48.239221°N 16.488953°E (un. = 151 m), 23.VII.2022, 1 specimen, photo by Stephan Manhalter (IN); Scĥffelgasse, 48.232536°N 16.326256° E (un. = 109 m), 8.X.2021, 1 specimen, photo by “marionouk” (IN); Währinger Park, 48.232474°N 16.349837°E (un. = 4 m), 24.VI.2021, 1 specimen, photo by Lorin Timaeus (IN); Waldspielplaz, 48.227339 N 16.37839°E (un. not recorded), 3.IX.2021, 1 specimen, photo by Gertraud Seiser (IN); Kaiserm̧hlen, 48.229648°N 16.427657°E (un. not recorded), 22.VIII.2021, 1 specimen, photo by Nikola Szucsich (IN); Sonnenblumengasse, 48.2251°N 16.438557°E (un. = 24 m), 2.X.2021, 1 specimen, photo by “garten444” (IN); Roterdstrasse, 48.219405°N 16.29962°E (un. = 20 m), 12.VIII.2022, 1 specimen, photo by “moidl67” (IN); Alservorstadt, 48.219196°N 16.353067°E (un. not recorded), 11.VIII.2022, 1 specimen, photo by “isadoraisadorable” (IN); Aspern, 48.215902°N 16.493916°E (un. = 6 m), 6.VI.2020, 1 specimen, photo by “cornetto” (IN); Stadion, 48.211087°N 16.418089°E (un. = 20 m), 29.VI.2022, 1 specimen, photo by “bobbyschauer” (IN); Stadlau, 48.203438°N 16.444828°E (un. = 256 m), 7.VIII.2021, 1 specimen, photo by Fernando Useros López (IN); idem, 48.205553°N 16.443761°E (un. = 116 m), 19.VII.2022, 1 specimen, photo by “yat70” (IN); near Museum f̧r angewandte Kunst, 48.207588°N 16.382843°E (un. not recorded), 2.IX.2022, 1 specimen, photo by “isadoraisadorable” (IN); Gumpendorfer Strasse, 48.200907°N 16.361856°E (un. not recorded), 25.VII.2022, 1 specimen, photo by Nikola Szucsich (IN); Botanischer Garten, 48.1901°N 16.3824°E (un. = 5 m), 15.VII.2022, 1 specimen, photo by Jolien Morren (OB); idem, 48.193705°N 16.383575°E (un. = 5 m), 10.VIII.2022, 1 specimen, photo by “felix_s98” (IN); near Friedenspagode, 48.189397°N 16.454814°E (un. not recorded), 4.VII.2021, 1 specimen, photo by Gertraud Seiser (IN); Gumperdorfer Strasse, 48.199874°N 16.361192°E (un. not recorded), 13.VIII.2021, 1 specimen, photo by Nikola Szucsich (IN); Haupteingang Botanischer Garten der Universität, 48.193624°N 16.383417 E (un. = 49 m), 10.VIII.2021, 1 specimen, photo by “loxod0nta” (IN); idem, 48.19138°N 16.383307°E (un. = 8 m), 30.IX.2021, 1 specimen, photo by Lorin Timaeus (IN); Kolschitzkygasse, 48.18824°N 16.372061°E (un. = 14 m), 27.VIII.2021, 1 specimen, photo by Piero Visconti (IN); Rosa-Janku-Park, 48.191°N 16.353669°E (un. = 2 m), 6.VIII.2020, 1 specimen, photo by “drosophila” (IN); Grete-Salzer-Park, 48.176101°N 16.332142°E (un. = 2 m), 4.IX.2021, 1 specimen, photo by “hederahelix” (IN); Ḩtteldorf, 48.202928°N 16.238791°E (un. = 50 m), 23.VII.2018, 1 specimen, photo by “cpu” (IN); Grete Salzer Park, 48.176101°N 16.332142°E (un. = 2 m), 4.IX.2021, 1 specimen, photo by “hederahelix” (IN); idem, 48.176113°N 16.332171°E (un. = 4 m), 6.VIII.2022, 1 specimen, photo by “hederahelix” (IN); Liesing, 48.133626°N 16.294543°E (un. = 4 m), 28.VII.2021, 1 specimen, photo by “cpu” (IN); near Gross-Enzersdorf, 48.200139°N 16.536611°E (un. not recorded), 25.IX.2021, 1 specimen, photo by Barnaby Walters (IN); Modeschule der Stadt Wien “Schloss Hetzendorf”, 48.168274°N 16.309168°E (un. = 82 m), 15.VIII.2022, 1 specimen, photo by “petra_l” (IN).
ITALY. Abruzzo: L’Aquila, L’Aquila, Bazzano, 42.336857°N 13.453525°E (un. = 232 m), 29.V.2020, 1 specimen, photo by Emanuele Santarelli (IN); Monticchio, 42.320183°N 13.465872°E (un. = 77 m), 21.VII.2018, 1 specimen, photo by Emanuele Santarelli (IN); Campo di Giove, surroundings Valico della Forchetta, 1250 m, on Eryngium sp., 23.VIII.2006, 8 specimens, leg. Eugenio Pacieri, 6 in CEP, 2 in CPC; Chieti, Torino di Sangro, Borgata Marina, 42.234595°N 14.542037°E (un. not recorded), 9.IX.2020, 1 specimen, photo by Rino Iubatti (IN). Aosta Valley: Aosta, Sarre, 45.7363°N 7.2546°E (un. = 100 m), 7.VIII.2018, 1 specimen, photo by Cor Zonneveld (OB);Aosta, via Caduti del Lavoro, 45.735725°N 7.330155°E (un. = 10 m), 14.VII.2022, 1 specimen, photo by “toomes” (IN). Apulia: Foggia,Vieste, between Mandriono and Monticello, 41.901835°N 16.098943°E (un. = 4 m), 12.VII.2022, 1 specimen, photo by “srollins” (IN); idem, near Grotta Sfondata, 41.813695°N 16.195092°E (un. = 186 m), 7.VII.2019, 1 specimen, photo by “silvio_lioce1” (IN); Bari, Poggiorsini, 40.939524°N 16.29193°E (un. = 8 m), 17.IX.2021, 1 specimen, photo by Cristiano Liuzzi (IN); Altamura, 40.832158°N 16.547955°E (un. = 10 m), 9.VIII.2021, 1 specimen, photo by Alessandro G. (IN); Gravina in Puglia, Selva di Gravina, 40.771133°N 16.48196°E (un. = 8 m), 19.IX.2021, 1 specimen, photo by Alessandro G. (IN); Gioia del Colle, 40.827006°N 16.945637°E (un. = 2700 m), 17.VI.2018, 1 specimen, photo by Nicola Nitti (IN); Taranto, Martina Franca, near Carpari, 40.726256°N 17.28704°E (un. = 61 m), 5.VIII.2011, 1 specimen, photo by Natalie Raeber (IN); Pulsano, Montedarena-Bosco Caggione, Marina di Leporano, 40.370547°N 17.317417°E (un. = 12 m), 1 specimen, photo by “gianfrs” (IN); idem, 40.366595°N 17.320445°E (un.= 22 m), 25.VII.2021, 1 specimen, photo by “gianfrs” (IN);Lecce, Porto Cesareo,Torre Lapillo, 40.285329°N 17.832698°E (un. = 2 m), 21.VIII.2020, 1 specimen, photo by Valentino Traversa (IN); Copertino, between Leverano and Copertino, 40.277665°N 18.008241°E (un. = 2 m), 30.VII.2020, 1 specimen, photo by Valentino Traversa (IN); idem, 40.270645°N 18.048904°E (un. = 1.28 km), 18.VIII.2020, 1 specimen, photo by “francesco167”(IN); Otranto, Porto Badisco, 40.081426°N 18.482349°E (un. = 4 m), 30.VI.2017, 1 specimen, photo by Mattia Menchetti (IN). Campania: Caserta, Capriati a Volturno, Oasi Naturalistica Le Mortine, 41.466715°N 14.092922°E (un. = 8 m), 23.VI.2011, 1 specimen, photo by Daniele Ritella (IN); idem, 41.468656°N 14.091778°E (un. = 8 m), 1 specimen, 22.VIII.2013, photo by Daniele Ritella (IN); Alvignano, 41.237278°N 14.33883°E (un. not recorded), 6.VI.2022, 1 specimen, photo by Alberto Di Giorgio (IN); Napoli, Napoli, Colli Aminei, 40.871953°N 14.236687°E (un. = 2100 m), 19.VI.2020, 1 specimen, photo by “sasymegna01” (IN); Pompei, ancient Roman ruins of Pompeii, 40.7522°N 14.4929°E (un. = 16 m), 29. VII.2021, 1 specimen, photo by Remco Hofland (OB); idem, 40.751293°N 14.485337°E (un. = 47 m), 25.VIII.2022, 1 specimen, photo by “virginielf” (IN); idem, 40.750772°N 14.484228°E (un. not recorded), 23.VIII.2021, 1 specimen, photo by “bemma” (IN); idem, 40.752244°N 14.487583°E (un. = 80 m), 31.VII.2022, 1 specimen, photo by Alexandre Mora (IN); idem, 40.751003°N 14.490245°E (un. = 9 m), 18.IX.2022, 1 specimen, photo by Joao Brito (IN); Avellino, Avellino, via Fontanatetta, 40.923408°N 14.819073°E (un. = 20 m), 21.VIII.2021, 1 specimen, photo by “vincedreux” (IN); Sturno, 41.02047°N 15.113146°E (un. = 65 m), 29.VIII.2020, 1 specimen, photo by Michele Sisto (IN); Salerno, Pontecagnano Faiano, Biblioteca del Centro di ricerca Orticoltura e Florovivaismo, 40.648711°N 14.890351°E (un. = 20 m), 19.VII.2022, 1 specimen, photo by Alessandro Mattedi (IN). Emilia-Romagna: Parma, Parma, near via Jones Melvin, 44.81767°N 10.305577°E (un. not recorded), 29.V.2018, 1 specimen, photo by “michidisperso” (IN); Modena, Galleria Estense, 44.648701°N 10.921596°E (un. = 87 m), 23.VII.2021, 1 specimen, photo by “daviferra” (IN); Ferrara, Ferrara, Francolino, Via del Luccio, 44.901996°N 11.661336°E (un. = 184 m), 26.V.2017, 1 specimen, photo by Carla Corazza (IN); idem, 44.901835°N 11.661055°E (un. not recorded), 8.VI.2022, 1 specimen, photo by Carla Corazza (IN); idem, Crociarola, 44.875448°N 11.668605°E (un. not recorded), 21.VI.2020, 1 specimen, photo by Carla Corazza (IN); idem, Ferrara, near via Darsena, 44.83432°N 11.606208°E (un. not recorded), 26.VII.2017, 1 specimen, photo by Nicola Baraldi (IN); Copparo, via Ferrara, 44.89422°N 11.822552°E (un. = 6 m), 14.V.2018, 1 specimen, photo by Nicola Baraldi (IN); Ostellato, Rovereto, 44.763719°N 11.864192°E (un. = 3.1 km), 27.VII.2022, 1 specimen, photo by Anna Luongo (IN); Voghiera, Montesanto, 44.732024°N 11.714658°E (un. = 359 m), 2.VI.2022, 1 specimen, photo by Valentina Buono (IN); Bologna, Molinella, 44.612264°N 11.667087°E (un not recorded), 30.IV.2021, 1 specimen, photo by Roberto Casadei (IN); Calderara di Reno, Lippo, 44.534386°N 11.300797° (un. not recorded), 1 specimen, photo by Claudio Danesi (IN); Ravenna, Conselice, 44.519169°N 11.83597°E (un. not recorded), 12.VIII.2021, 1 specimen, photo by Roberto Casadei (IN); idem, 44.519192°N 11.835868°E (un. not recorded), 20.VI.2022, 1 specimen, photo by Roberto Casadei (IN); Ravenna, Lido di Dante, 44.389298°N 12.295821°E (un. = 289 m), 8.VIII.2019, 1 specimen, photo by Valentina Buono (IN); Bastia—Borgo Pasna, 44.265626°N 12.167117°E (un. = 1070 m), 1 specimen, photo by Danio Miserocchi (IN); Forlì-Cesena, Meldola, 44.109342°N 12.047774°E (un. = 232 m), 20.VII.2020, 1 specimen, photo by “daniela83fc” (IN); Cesena, Monte Grande, 44.094166°N 12.125924°E (un. not recorded), 15.VI.2019, 1 specimen, photo by Claudio Danesi (IN); Gambettola, 44.112032°N 12.337531°E (un. = 21 m), 14.VII.2016, 1 specimen, photo by Eddi Bisulli (IN); Rimini, Santarcangelo di Romagna, Case Paglierani, 44.06058°N 12.432024°E (un. not recorded), 15.VII.2021, 1 specimen, photo by Francesco Ghinelli (IN). Friuli-Venezia Giulia: Pordenone, Sequals, near Lestans, 46.170085°N 12.888914°E (un. = 4 m), 2.VII.2022, a specimen, photo by Mirco Peschiutta (IN). Udine, Ronchis, 45.812695°N 13.00603°E (un. not recorded), 26.VI.2016, 1 specimen, photo by Marcello Consolo (IN); Udine, 27.VI.2020, 1 specimen, photo by Michele Armellini (FEI); Trieste, Duino-Aurisina, Medeazza, 7.VIII.2014, 1 specimen, photo by Lucio Morin (FEI); Trieste, Basovizza, 45.642774°N 13.865443°E (un. = 4 m), 14.VI.2022, 2 specimens, photo by Felix Puff (IN). Latium: Rieti, Rieti, Terminillo, Pian de’ Rosce, 1014 m, 3.VII.2010, 1 specimen, leg. Eugenio Pacieri, CEP; idem, Lungovelino Bellagamba, 42.399787°N 12.861505°E (un. = 50 m), 6.VIII.2018, 5 specimens, photo by Francesca Graziani (IN); Castel Sant’Angelo, Canetra, 438 m, 2.VII.2018, 3 specimens, leg. Eugenio Pacieri, CEP; Roma, Rocca di Papa, 41.777067°N 12.702199°E (un. = 15 m), 19.VII.2020, 1 specimen, photo by Luca Odorico (IN); Campagnano di Roma, contrada Selva Grossa, Via Campagnanese, 12-18.VII.2010, 1 specimen, leg. Eugenio Pacieri, CEP; Sacrofano, Via Prato della Chiesa, surroundings riding school “Le Piane”, 42.121389°N 12.441667°E [obtained from label coordinates 42°07’17.0”N 12°26’30.0”E] (un. not recorded), 290 m, 30.VII.2011, 2 specimens, leg. Eugenio Pacieri, CEP; Carpineto Romano, 41.586955°N 13.119668°E (un. = 2 m), 10, VII.2016, 2 specimens, photo by Dino Biancolini (IN); Castel Gandolfo, Lago Albano, 31.VII.2016, 1 specimen, photo by Roberto Vanzini (FEI); Fiumicino, Maccarese, 11.VI.2014, 1 specimen, photo by Maurizio Gigli (FEI); Roma, Trionfale, Riserva naturale dell’Insugherata, 12.IX.2010, 10 specimens, leg. Eugenio Pacieri, CEP; idem, Aniene valley, crossroads to Anticoli Corrado, 3.IX.2006, 2 specimens, leg. Eugenio Pacieri, CEP; idem, Castel Porziano, 25.VII.2013, 2 specimens, photo by Paolo Maltzeff (FEI); idem, Via Appia Nuova, 41.858513°N 12.5364°E (un = 2 km), 15.VIII.2022, 1 specimen, photo by Masked Lynx (IN); idem, Castel Romano, 41.713623°N 12.420061°E (un. = 2 m), 31.VII.2021, 3 specimens, photo by “a_ calabrese” (IN); idem, Lido di Ostia, 41.741203°N 12.250364°E (un. = 8 m), 24.VIII.2022, 1 specimen, photo by “beethoven242” (IN); idem, 41.731355°N 12.31018°E (un. not recorded), 3.IX.2006, 1 specimen, photo by Sarah Gregg (IN); idem, Castel Fusano, 41.705948°N 12.339626°E (un. = 14 m), 6.VIII.2022, 1 specimen, photo by “king_maf” (IN); Frosinone, Piglio, 41.824082°N 13.166177°E (un. = 6 m), 7.VII.2019, 1 specimen photographed but reported as many individuals, photo by Luca Tringali (IN); idem, 41.829921°N 13.178952°E (un. = 49 m), 29.VI.2020, 1 specimen photographed but reported as some individuals, photo by Luca Tringali (IN); Latina, Sabaudia, Selva di Circe, 41.344895°N 13.074611° (un. = 1.05 km), 26.VI.2022, 1 specimen, photo by Ronald Werson (IN). Liguria: Savona, Villanova d’Albenga, 44.0452°N 8.1132°E (un. = 10 m), 6.VIII.2014, 1 specimen, photo by Herrald Damen (OB); Altare, 44.337268°N 8.341539°E (un. = 18 m), 22.VII.2019, 1 specimen, photo by “mich8” (IN); idem, 44.337149°N 8.343636°E (un. = 132 m), 8.IX.2019, 1 specimen, photo by “mich8” (IN); Cairo Montenotte, 44.415006°N 8.308041°E (un. = 4 m), 2.VII.2020, 1 specimen, photo by Elan Zucchetti (IN); La Spezia, Deiva Marina, 44.2157 N 9.5188°E (un. = 25 m), 15.IX.2021, 1 specimen, photo by “foppe” (OB); Sarzana, VI.2010, 1 specimen, photo by “Nemo Nemus” (FNM). Lombardy: Sondrio, Sorico, Riserva Naturale Pian di Spagna e Lago di Mezzola, 46.179081°N 9.428489°E (un. = 326 m), 27.VIII.2020, 1 specimen, photo by Federico Del Barba (IN); Como, Sorico, near Peschiera Lake, 46.211938°N 9.422359°E (un. = 275 m), 2.VIII.2015, 1 specimen, photo by Gaia Bazzi (IN); Valsolda, 46.039997°N 9.046628°E (un. = 15 m), 3.VII.2019, 3 specimens, photo by Harm Alberts (IN, OB); Bellagio, 45.9792°N 9.2528°E (un. = 10 m), 11.VII.2015, 1 specimen, photo by Ronsse Etienne (OB); Alta Valle Intelvi, Pellio Intelvi, 45.97314°N 9.043677°E (un. = 507 m), 18.VII.2020, 2 specimens, photo by Mirko Tomasi (IN); Lenno,Lungolago Guido Delmati, 45.9709°N 9.1965°E (un. not recorded), 15.VII.2022, 1 specimen, photo by “BabsD” (OB); Valmorea, 45.81856°N 8.921053°E (un. = 184 m), 9.VII.2021, 1 specimen, photo by Jeroen Van Akkeren (IN); Castelnuovo Bozzente, 45.763023°N 8.939609°E (un. not recorded), 13.VIII.2021, 1 specimen, photo by Paola Casale (IN); Merone, Lago di Pusiano, 45.791327°N 9.254912°E (un. not recorded), 5.IX.2021, 1 specimen, photo by “lionello” (IN); Bergamo, San Pellegrino Terme, 45.853056°N 9.663056 (un. not recorded), 19.VIII.2022, 2 specimens, photo by “theshowmustgoon” (IN); idem, 45.852778°N 9.663333°E (un.not recorded), 28.VI.2022, 1 specimen,photo by “theshowmustgoon” (IN);Sedrina, near Lisso, 45.777864°N 9.616102°E (un. = 2 m), 22.V.2022, 1 specimen, photo by Nicola Zanetti (IN); idem, near Mediglio, 45.763859°N 9.622409°E (un. = 18 m), 19.VII.2022, 1 specimen, photo by Nicola Zanetti (IN); Villa d’Adda, Oasi dell’Alberone, 45.72694°N 9.451184°E (un. = 191 m), 4.VIII.2020, 1 specimen, photo by Giuliana Pirotta (IN); Ranica, Via Borgosale, 45.733151°N 9.714134°E (un. = 15 m), 24.VI.2018, 1 specimen, photo by Giovanni Perico (IN); Alzano Lombardo, via Peppino Ribolla, 45.731031°N 9.728892°E (un. = 2 m), 6.VI.2020, 1 specimen, photo by Stefano Aguzzi (IN); Villa di Serio, Parco Carrara, 45.721854°N 9.737613°E (un. = 18 m), 26.VI.2022, 1 specimen, photo by “margherita_bg” (IN); Scanzorosciate, 45.711489°N 9.73931°E (un. = 190 m), 20.V.2021, 1 specimen, photo by “gibe_a_m” (IN); Bergamo, via Giuseppe Verdi, 45.6982°N 9.671344°E (un. = 35 m), 19.VII.2020, 1 specimen, photo by “filippo_” (IN); idem, via Sora Gennaro, 45.695625°N 9.671542°E (un.= 65 m), 24.VII.2020, 1 specimen, photo by “dennis04” (IN);Vigolo, 45.716711°N 10.021673°E (un. = 2100 m), 14.IX.2019, 1 specimen, photo by Gabriele La Grasta (IN); Vaprio d’Adda, 45.574532°N 9.535283°E (un. = 31 m), 13.VI.2021, 1 specimen, photo by “samuele2010” (IN); Romano, between Cascina Corsa and Cascina Marinoni, 45.521648°N 9.72222°E (un. not recorded), 1.VII.2018, 1 specimen, photo by “fakko” (IN); Albano Sant’Alessandro, 27.IV.2009, 1 specimen, photo by Andrea Pesenti (FEI); Brescia, Darfo Boario Terme, Lago Moro, 45.880732°N 10.161913°E (un. = 22 m), 2.VII.2021, 1 specimen, photo by “donnasara” (IN); Idro, Lake Idro, 45.7527°N 10.4941°E (un. = 100 m), 6.VII.2011, 1 specimen, photo by Johan Neegers (OB); Zone, 16.VIII.2013, 1 specimen, leg. Lucia Pizzocaro, CLP; Iseo, Ciochet, 45.653136°N 10.029384°E (un. = 281 m), 14.VI.2021, 1 specimen, photo by Gabriele La Grasta (IN); idem, 45.653396°N 10.031103°E (un. = 248 m), 22.VII.2021, 1 specimen, photo by Gabriele La Grasta (IN); Riserva Naturale delle Torbiere del Sebino, 45.646132°N 10.036745°E (un. = 194 m), 1.VII.2021, 1 specimen, photo by Gabriele La Grasta (IN); idem, 45.64389°N 10.029217°E (un. = 183 m), 1.VII.2021, 1 specimen, photo by Gabriele La Grasta (IN); idem, 45.637491°N 10.034903°E (un. = 2 m), 7.VI.2022, 1 specimen, photo by Gabriele La Grasta (IN); Brescia, via S. Gottardo, 45.541225°N 10.246095°E (un. = 10 m), 6.VI.2022, 1 specimen, photo by “mattia01” (IN); Iseo, 45.6709°N 10.0859°E (un. = 5 m), 16.VIII.2021, 1 specimen, photo by Bruno De Geest (OB); Monticelli Brusati, 45.617921°N 10.093605°E (un. = 15 m), 21.VI.2020, 1 specimen, photo by Marco Bologna (IN); Limone sul Garda, 45.823402°N 10.801781°E (un. = 82 m), 22.VIII.2021, 1 specimen, photo by “trullatrine” (IN); Tignale, 45.727219°N 10.717106°E (un. = 6 m), 17.IX.2021, 1 specimen, photo by “vickyfrank” (IN); Botticino, 45.554099°N 10.303988°E (un. = 996 m), 22.VI.2022, 1 specimen, photo by Gabriele La Grasta (IN); Manerba del Garda, 45.534153°N 10.555966°E (un. not recorded), 6.VIII.2019, 1 specimen, photo by Guenter Moeller (IN); Moniga del Garda, 45.523117°N 10.541088°E (un. not recorded), 1.VIII.2019, 1 specimen, photo by Davide Grillo (IN); Sirmione, 45.4699°N 10.5966°E (un. = 10 m), 19.VIII.2008, 1 specimen, photo by Jan Willem Dekker (OB); idem, 45.4930°N 10.6072°E (un. = 25 m), 14.VI.2019, 1 specimen, photo by Frank Berbers (OB); Desenzano del Garda, 45.4673°N 10.5457°E (un. = 64 m), 17.VIII.2022, 1 specimen, photo by “brecht” (OB); idem, Oasi San Francesco del Garda, 45.464287°N 10.590501°E (un. = 15 m), 24.VII.2021, 1 specimen, photo by “radrat” (IN); Varese, Inarzo, Palude Brabbia, 45.787042°N 8.725064°E (un. not recorded), 4.VIII.2022, 1 specimen, photo by “bethanp” (IN); Saronno, Via Padre Giuliani, 45.620258°N 9.020659°E (un. = 68 m), 26.VI.2020, 1 specimen, photo by “bracchettogiallo” (IN); Albizzate, 5.VIII.2012, 1 specimen, photo by Emanuele Zenga (FEI); Monza and Brianza, Villasanta, 45.606792°N 9.303537°E (un. = 6 m), 11.VI.2018, 1 specimen, photo by Andrea Quadrio (IN); Monza, Parco di Monza, 45.597612°N 9.2873°E (un. = 31 m), 7.VI.2019, 1 specimen, photo by Augusto Crippa (IN); idem, Via Sele, 45.577841°N 9.235737°E (un. not recorded), 14.VI.2022, 1 specimen, photo by Fernando Oceani (IN); Lesmo, 3.VI.2017, 1 specimen, photo by Giuseppe Vago (FEI); Lecco, Varenna, 46.0094°N 9.2834°E (un. = 20 m), 9.VIII.2022, 1 specimen, photo by Strandjutter Koen (OB); Bosisio Parini, 45.801727°N 9.294369°E (un. not recorded), 19.VII.2022, 1 specimen, photo by “keymi” (IN); Lecco, Monte San Martino, 27.VII.2014, 1 specimen, photo by Michele Butta (FEI); Milano, Bollate, Oasi WWF Il Caloggio, 45.551294°N 9.102408°E (un. = 9 m), 5.VIII.2022, 1 specimen, photo by “donzelli” (IN); Sesto San Giovanni, 45.546128°N 9.205981°E (un. = 8 m), 29.IX.2016, 1 specimen, photo by Massimo Urso (IN); idem, Via Maggiore Giuseppe Galliano, 45.537932°N 9.185713°E (un. = 196 m), 1.VI.2020, 1 specimen, photo by Tina Pannullo (IN); Milano San Felice, 45.45773°N 9.314367°E (un. = 258 m), 24.VI.2018, 1 specimen, photo by “geoclaudia” (IN); San Donato Milanese, Via Martiri di Cefalonia, 45.42113°N 9.271928°E (un. = 30 m), 30.VII.2020, 1 specimen, photo by “pavlaki1968” (IN); Lainate, 45.571535°N 9.047111°E (un. = 602 m), 6.VI.2022, 1 specimen, photo by “umbertuccio” (IN); Milano, Via Guido Ucelli di Nemi, 45.436745°N 9.26327°E (un. = 4 m), 10.VII.2017, 1 specimen, photo by “geoclaudia” (IN); idem, Primaticcio, 45.456336°N 9.129572°E (un. = 15 m), 23.V.2022, 1 specimen, photo by Ergor Gavrilov (IN); Turbigo, 20.VIII.2021, 1 specimen, photo by Carlo Galliani (FEI); Novate Milanese, 26.VI.2020, 1 specimen, photo by Ezio Sacchi (FEI); Paderno Dugnano, 12.VI.2019, 1 specimen, photo by Carlo Galliani (FEI); Pavia, Mortara, Via Sabotino, 45.260117°N 8.730037°E (un. not recorded), 23.VI.2020, 1 specimen, photo by Giacomo Boffa (IN); Pavia, Darsena del Naviglio Pavese di Porta S. Vito, 45.193004°N 9.157377° (un. = 8 m), 5.VII.2021, 1 specimen, photo by Alessandro G. (IN); Frascarolo, Cascina Belvedere, 45.053303°N 8.648378°E (un. = 43 m), 5.VIII.2021, 1 specimen, photo by “pierabionda” (IN); idem, 45.053406°N 8.648843°E (un. not recorded), 30.V.2022, 1 specimen, photo by Leonardo Mantovani (IN); Travacò Siccomario, Boschi, 45.149307°N 9.186219°E (un. = 747 m), 1.VIII.2022, 1 specimen, photo by Edoardo Razzetti (IN); Cremona, Genivolta, 45.333003°N 9.877046°E (un. = 699 m), 8.IX.2020, 1 specimen, photo by “theworldofpaolo” (IN); Cremona, between via Riva Fredda and via Francesco Sforza, 45.126023°N 10.053457°E (un. = 2 m), 13.VI.2017, 1 specimen, photo by “damighez” (IN); Crema, VI.2008, 1 specimen, photo by Franco Giordana (FAP); Motta Baluffi, Solarolo Monasterolo, 45.058022°N 10.234783°E (un. = 686 m), 27.VI.2020, 1 specimen, photo by “fedelf” (IN); San Giovanni in Croce, 45.079436°N 10.371963°E (un not recorded), 24.VII.2016, 2 specimens, photo by “sofia-b” (IN); Rivarolo del Re ed Uniti, Rivarolo del Re, 45.03164°N 10.475779°E (un. = 17 m), 1.X.2016, 1 specimen, photo by Daniele Fuochi (IN); Mantova, Canneto Sull’Oglio, Via Alessandro Volta, 45.146791°N 10.383881°E (un. = 22 m), 26.V.2017, 1 specimen, photo by “sofia-b” (IN); Acquanegra sul chiese, 45.138275°N 10.436048°E (un. not recorded), 1.VII.2019, 1 specimen, photo by Francesco Cecere (IN); Calvatone, Vicolo Castaldi, 45.128513°N 10.441616°E (un. = 20 m), 8.VI.2017, 1 specimen, photo by “lella” (IN); Marmirolo, Riserva naturale Bosco Fontana, 45.200291°N 10.747276°E (un. = 31 m), 8.VIII.2019, 1 specimen, photo by Andrea Angiari (IN); Mantova, viale Te, 45.147324°N 10.788619°E (un. not recorded), 17.VI.2018, 1 specimen, photo by Marco Munari (IN). Marche: Pesaro-Urbino, Fano, 2.VII.2019, 1 specimen, photo by Giovanna Olivieri (FEI); Ancona, Jesi, Riserva naturale di Ripa Bianca, 12.VI.2011, 1 specimen, photo by “mbondini” (FNM); Macerata, Camerino, Campus Universitario, 43.1468°N 13.066772°E (un. = 35 m), 6.VI.2022, 1 specimen, photo by “fede02” (IN). Piedmont: Verbano-Cusio-Ossola, near Alpe Rusa, 45.90989°N 8.385093°E (un. = 727 m), 7.VII.2021, 1 specimen, photo by Philipp Hoenle (IN); Stresa, Piazza Guglielmo Marconi, 45.884388°N 8.540675°E (un. = 63 m), 4.VIII.2020, 1 specimen, photo by “julesfromarvs” (IN); Novara, Armeno, 45.8238°N 8.4487°E (un. not recorded), 28.VII.2010, 1 specimen, photo by Jaap van der Linden (OB); idem, 45.820555°N 8.439722°E (un. not recorded), 17.VIII.2015, 1 specimen, photo by “martinbishop” (IN); Orta San Giulio, 45.7950°N 8.4130°E (un. = 4 m), 22.VII.2022, 2 specimens, photo by Christian Deutsch (OB); Pella, west shore of Lake Orta, 8.IX.2018, 1 specimen, photo by “vladim” (FNM); Gozzano, Buccione, 45.763768°N 8.426488°E (un. not recorded), 25.VIII.2021, 2 specimens, photo by “cattanea” (IN); idem, 45.765039°N 8.42823°E (un. = 122 m), 14.VII.2022, 1 specimen, photo by Gianmaria Marchese (IN); Biella, Pray, Flecchia, 45.671603°N 8.19207°E (un. = 8 m), 17.VII.2016, 1 specimen, photo by Gabriele Vaudano (IN); idem, 45.671596°N 8.192166°E (un. = 2 m), 15.VI.2020, 1 specimen, photo by Gabriele Vaudano (IN); Torino, Candia Canavese, 45.32679°N 7.898313°E (un. = 36 m), 31.VII.2019, 1 specimen, photo by “giorege1959” (IN),Torino, Via Spagna, 45.099972°N 7.589954°E (un. = 15 m), 22.VI.2011, 1 specimen, photo by “daveoshaw” (IN); idem, via Galliate, 45.084167°N 7.632222°E (un. not recorded), 3.VIII.2021, 1 specimen, photo by Enrico Valen (IN); idem, Corso Brescia, 45.082422°N 7.689872°E (un. = 13 m), 2.VII.2022, 1 specimen, photo by Stefano Gellato (IN); idem, Corso Chieri, 45.062575°N 7.738667°E (un. = 5 m), 11.V.2020, 1 specimen, photo by “chiarapo” (IN); idem, Madonna del Pilone, 45.089524°N 7.734084°E (un. = 19 m), 26.V.2022, 1 specimen, photo by Lorenzo Dotti (IN); idem, without precise locality in the city, 10.IX.2019, 1 specimen, photo by Martino Romano; Grugliasco, Strada della Pronda, 45.065927°N 7.613119°E (un. = 9 m), 1 specimen, photo by “pepik” (IN); Moncalieri, Corso Trieste, 45.01243°N 7.67039°E (un. = 3 m), 27.VI.2020, 1 specimen, photo by Enrico Castello (IN); Piobesi Torinese, 44.927222°N 7.608611°E (un. not recorded), 15.VI.2020, 1 specimen, photo by Roberto Ostellino (IN); Carmagnola, 44.885582°N 7.683397°E (un. = 4 m), 21.VI.2019, 1 specimen, photo by “mich8” (IN); Villar Perosa, 44.913067°N 7.261174°E (un. = 5 m), 21.VII.2020, 1 specimen, photo by Davide Berton (IN); Angrogna, Martel, 44.837569°N 7.240146°E (un. = 31 m), 25.VII.2020, 1 specimen, photo by Elio Giacone (IN); Cuneo, Racconigi, 44.789834°N 7.661665°E (un. = 3160 m), 13.VII.2021, 1 specimen, photo by Elio Giacone (IN);Alba, Belmonte, 44.691162°N 8.036529°E (un. = 8 m), 20.VI.2021, 1 specimen, photo by Roberto Ghiglia (IN); Monforte d’Alba, 44.5855°N 7.9692°E (un. = 5 m), 14.IX.2021, 1 specimen, photo by “Vera O” (OB); Sant’Albano Stura, Oasi Naturalistica “La Madonnina”, 44.499416°N 7.700678°E (un. = 97 m), 27.VIII.2022, 1 specimen, photo by Elio Giacone (IN); Canosio, near Lago Nero, 44.4117°N 7.0633°E (un. = 16 m), 19.VII.2021, 1 specimen, photo by Kurt Geeraerts (OB); Lesegno, 44.401954°N 7.967972°E (un. = 45 m), 7.VII.2022, 1 specimen, photo by Elan Zucchetti (IN); Limone Piemonte, 44.2196°N 7.5686°E (un. = 100 m), 11.VII.2012, 1 specimen, photo by Pietsje Fleurbaaij (OB); Valdieri, Pian del Valasco, 44.204303°N 7.265169°E [obtained from label coordinates 4896028N 361389E], 1450 m, 9.IX.2013, 1 specimen, leg. Margherita Norbiato, MZUF; Asti, Viarigi, 44.9793°N 8.3614°E (un. = 4 m), 14.VIII.2022, 1 specimen, photo by “Oetel” (OB); Dusino San Michele, 44.923421°N 7.973141°E (un. not recorded), 16.VI.2019, 1 specimen, photo by Stefano Bertoloni (IN); Asti, Bricco Roasio, 44.943304°N 8.168087°E (un. = 186 m), 21.VI.2020, 1 specimen, photo by Salvatore infanti (IN); Alessandria, Ponti, 44.6433°N 8.3616°E (un. not recorded), 10.VIII.2022, 1 specimen, photo by “Brecht” (IN). Sardinia: Sassari, Olbia, 40.889765°N 9.613326°E (un. = 61 m), 2.IX.2020, 1 specimen, photo by Joachim Louis (IN). Sicily: Trapani, Trapani Saltworks, 37.9626°N 12.5079°E (un. = 100 m), 16.VI.2018, 1 specimen, photo by Yves Pieters (OB). Trentino-Alto Adige: Bolzano, Varna, 46.73184°N 11.649502°E (un. not recorded), 8.VIII.2022, 1 specimen, photo by “plissnrutscha2” (IN); Merano, 46.671354°N 11.152518°E (un. = 3.05 km), 19.VIII.2022, 1 specimen, photo by Carlo Zoico (IN); idem, Via Enrico Toti, 46.6657°N 11.1530°E (un. = 65 m), 3 specimens, photo by Andreas Clausen (OB); Tesimo, 46.556873°N 11.157646°E (un. = 4 m), 19.VI.2021, 1 specimen, photo by Elia Guariento (IN); Bolzano, Gries-San Quirino, 46.505446°N 11.344107°E (un. not recorded), 22.VI.2021, 1 specimen, photo by David Gruber (IN); idem, near Via Trieste, 46.494159°N 11.346295°E (un. = 10 m), 30.VII.2020, 1 specimen, photo by Elia Guariento (IN); Trento, Trento, Gardolo, 46.107592°N 11.110213°E (un. = 10 m), 7.VIII.2022, 1 specimen, photo by Claudio Peterlongo (IN); idem, Vela, 46.089236°N 11.09948°E (un. = 488 m), 6.VI.2021, 1 specimen, photo by Luigi Cannizzo (IN); Caldonazzo, Caldonazzo Lake, 46.0102°N 11.2647°E (un. = 5 m), 24.VII.2021, 1 specimen, photo by Hessel Rasch (OB); Arco, 45.9253°N 10.8904°E (un. = 23 m), 15.VIII.2022, 1 specimen, photo by Jana Petermann (OB); idem, 45.8758°N 10.8696°E (un. = 5 m), 1.VIII.2022, 1 specimen, photo by Roos Eichhorn (OB); idem, Linfano, 45.874722 N 10.870277°E (un. not recorded), 19.VII.2019, 1 specimen, photo by “gitteahlers” (IN); Riva del Garda, 45.898086°N 10.855539°E (un. = 6 m), 20.VIII.2022, 1 specimen, photo by Daniel Iversen (IN). Tuscany: Livorno, Livorno, 5.VII.2011, 1 specimen, photo by Enrico Lucarelli (FEI); Castagneto Carducci, Paradù Tuscany EcoResort, 43.152285°N 10.543094°E (un. = 65 m), 6.VI.2022, 1 specimen, photo by “jonpfei” (IN); Elba island, Poggio, 42.785816°N 10.185022°E (un. = 2 m), 23.VI.2022, 1 specimen, photo by Roberto Barsaglini (IN); Porto Azzurro, 42.758482°N 10.385327°E (un. = 2 m), 1.VII.2022, 1 specimen, photo by Roberto Barsaglini (IN); Lucca, Bagni di Lucca, Scesta, 44.036817°N 10.663085°E (un. not recorded), 24.VII.2021, 1 specimen, photo by “pippocalzelunghe” (IN); Viareggio, Torre del Lago Puccini, 43.8319°N 10.3072°E (un. = 10 m), 2.IX.2013, 1 specimen, photo by “Nick W” (OB); Pistoia, Serravalle Pistoiese, La Magione, 43.89045°N 10.823221°E (un. = 14 m), 28.VII.2015, 1 specimen, photo by Mattia Menchetti (IN); Prato, Prato, Figline, Bardena torrent, 120 m, 3.VII.2007, 1 specimen, leg. F. Terzani & S. Rocchi, MZUF; Firenze, Firenze, via della Covacchia, 43.82392°N 11.23298°E (un. = 15 m), 155 m, 13.V.2000, 1 specimen, leg. Francesca Graziani, CFG; idem, 43.82385°N 11.232948°E (un. = 8 m), 26.VI.2019, 1 specimen, photo by Francesca Graziani (IN); idem,Novoli, near the confluence between torrents Terzolle and Mugnone, 43.790247°N 11.232785°E (un. = 31 m), 6.VIII.2020, 1 specimen, photo by Nicola Addelfio (IN); idem, via Buonsignori, 43.788519°N 11.234528°E (un. = 122 m), 1 specimen, photo by Nicola Addelfio (IN); idem, Ponte a Mensola, 43.78111°N 11.30920°E (un. = 90 m), 13.VIII.2016, leg. Filippo Ceccolini, Fabio Cianferoni, Saverio Rocchi & Fabio Terzani, CFC; idem, Settignano, lake of Azienda Tre Pini, 43.78350°N 11.33405°E (un. = 50 m), 13.VIII.2016, leg. Filippo Ceccolini, Fabio Cianferoni, Saverio Rocchi & Fabio Terzani, CFC; Bagno a Ripoli, Grassina, VI.2012, 1 specimen, leg. Lucia Pizzocaro, CLP; Pontassieve, between Molino del Piano and Rufina, 43.8139°N 11.4489°E (un. = 60 m), 23.VI.2022, 1 specimen, photo by “willemdie” (OB); Reggello, near Via Di Camerino, 43.670076°N 11.526992°E (un. = 28 m), 22.VI.2021, 1 specimen, photo by “p-france” (IN); Arezzo, Pratovecchio Stia, Pratovecchio, 420 m, VIII.2019, 4 specimens, legit Margherita Norbiato, MZUF; Chiusi della Verna, 43.697439°N 11.925008°E (un. not recorded), 6.IX.2021, 1 specimen, photo by Boris Mittermeier (IN); Chitignano, La Fatica, 43.65349°N 11.86979°E (un. = 90 m), 25.VII.2000, 1 specimen, leg. Filippo Ceccolini, CFC; Castel Focognano, Rassina, 43.64996°N 11.83420°E (un. = 20 m), 15.VI.2012, 1 specimen, leg. Filippo Ceccolini, CFC; idem, 43.649846°N 11.833957°E (un. = 3 m), 8.VII.2020, 2 specimens, photo by Filippo Ceccolini (IN); idem, 43.649902°N 11.834139°E (un. = 2 m), 27.VIII.2021, 1 specimen, photo by Filippo Ceccolini (IN); idem, 43.65002°N 11.834124°E (un. = 1 m), 16.VIII.2022, 1 specimen, photo by Filippo Ceccolini (IN); Bibbiena, via della Segheria, 43.696486°N 11.801012°E (un. = 2 m), 18.VI.2022, 6 specimens, photo by Filippo Ceccolini (IN); Arezzo, via Alessandro Volta, 43.471099°N 11.85505°E (un. = 4 m), 1.VI.2017, 1 specimen, photo by Mattia Menchetti (IN); Viciomaggio, Via M. Mannelli, 43.435845°N 11.772527°E (un. = 5 m), 14.VII.2020, 1 specimen, photo by “profstocchi” (IN); Lucignano, Santa Maria, 43.2725°N 11.7345°E (un. = 25 m), 17.VI.2016, 1 specimen, photo by “Albert:)” (OB); Siena, Monteriggioni, 43.3897°N 11.2237°E (un. not recorded), 2.VIII.2022, 1 specimen, photo by “Ekkel” (OB); Grosseto, Bagnoli, 42.87061°N 11.550539°E (un. = 2 m), 30.VI.2021, 1 specimen, photo by Luca Passalacqua (IN); Pitigliano, 42.6326°N 11.6487°E (un. = 10 m), 2.VIII.2013, 1 specimen, photo by Willem Deleus (OB); Capalbio, Oasi WWF Lago di Burano, 42.400316°N 11.367038°E (un. = 2 m), 13.VIII.2022, 1 specimen, photo by Luca Passalacqua (IN); Giglio island, Isola del Giglio, Giglio Campese, 42.365628°N 10.879812°E (un. = 103 m), 14.VI.2021, 1 specimen, photo by Valentin Moser (IN). Umbria: Perugia, Monte Santa Maria Tiberina, 43.4375°N 12.1585°E (un. = 8 m), 28.VIII.2021, 1 specimen, photo by Jeremy Bruggeman (OB); Tuoro sul Trasimeno, near Montecchio, 43.1941°N 12.0381°E (un. not recorded), 16.VIII.2022, 1 specimen, photo by “Wimtje” (OB); Magione, Monte del Lago, 43.149973°N 12.170778°E (un. = 3 m), 3.VII.2016, 1 specimen, photo by Roberto Sindaco (IN); idem, 43.1447°N 12.1610°E (un. = 5 m), 26.VII.2022, 1 specimen, photo by Jennifer Huizing (OB); idem, San Feliciano, 43.1165°N 12.1675°E (un. not recorded), 21.VII.2022, 1 specimen, photo by “Klara” (OB); Foligno, near Via Tagliamento, 42.9618°N 12.7098°E (un. = 25 m), 17.VIII.2020, 1 specimen, photo by Frank Coulier (OB); idem, near Statale Flaminia, 42.9708°N 12.7165°E (un. = 25 m), 15.VIII.2020, 1 specimen, photo by Frank Coulier (OB); Preci, 42.8869°N 13.0146°E (un. not recorded), 13.VIII.2008, 1 specimen and nest, photo by Michel Schreurs (OB); Terni, San Venanzo, near Rotecastello, 42.8749°N 12.3021°E (un. = 5 m), 6.VIII.2022, 1 specimen, photo by “De Pie” (OB). Veneto: Belluno, Belluno, Val Tibolla, 46.079231°N 12.246563°E (un. not recorded), 12.VII.2017, 1 specimen, photo by Gianfranco Dramis (IN); Feltre, Farra, VII.2013, 1 specimen, photo by Nicola Dal Zotto (FEI); Treviso, San Zenone degli Ezzelini, Oasi San Daniele, 45.805304°N 11.812399°E (un. = 186 m), 12.IX.2020, 1 specimen, photo by Fiorella Rigoni (IN); Moriago della Battaglia, Isola dei Morti, VII.2011, 1 specimen, photo by “Pinog” (FNM), Treviso, VI.2013, 1 specimen, photo by Pino Perino (FAP); Vicenza,Valdagno, near Spelaccia, 45.644923°N 11.272975°E (un.= 61m), 2.VII.2022, 1 specimen, photo by Marco Vicariotto (IN);Valdagno, via Dorando Pietri, 45.641716°N 11.307579°E (un. = 61 m), 27.VI.2018, 1 specimen, photo by Jessica Peruzzo (IN); idem, bike path near Agno torrent, 45.638302°N 11.309218°E (un. not recorded), 21.IX.2019, 1 specimen, photo by Jessica Peruzzo (IN); idem, via Terragli, 45.633461°N 11.308705°E (un. not recorded), 14.VI.2020, 1 specimen, photo by Marco Vicariotto (IN); idem, 45.63319°N 11.308322°E (un. not recorded), 20.VII.2017, 1 specimen, photo by Jessica Peruzzo (IN); Cornedo Vicentino, 45.615062°N 11.337639°E (un. = 2690 m), 31.VIII.2018, 1 specimen, photo by Jessica Peruzzo (IN); Trissino, via della Stampa, 45.557671°N 11.376051°E (un. not recorded), 7.IX.2019, 1 specimen, photo by Jessica Peruzzo (IN); Vicenza, viale Arnaldo Fusinato, 45.53901°N 11.536785°E (un. = 244 m), 23.I.2018, 1 specimen, photo by Marco Vicariotto (IN); idem, Parco Retrone, 45.531709°N 11.525978°E (un. = 357 m), 1 specimen, photo by Marco Vicariotto (IN); Bosco del Quarelo, 45.522218°N 11.521402°E (un. = 61 m), 1 specimen, photo by Marco Vicariotto (IN); Arcugnano, Lago di Fimon, 45.465255°N 11.54273°E (un.= 1540 m), 16.VII.2019, 1 specimen, photo by Marco Vicariotto (IN);Val Liona, Villa del Ferro, 45.384287°N 11.46992°E (un. = 8 m), 10.VII.2019, 1 specimen, photo by Marco Vicariotto (IN); Orgiano, 45.355502°N 11.41135° E (un. = 488 m), 10.VII.2019, 1 specimen, photo by Marco Vicariotto (IN); Dueville, 12.VIII.2014, 1 specimen, photo by Antonio Carraro (FEI); Arcugnano, Fimon, VIII.2013, 1 specimen, photo by Davide Tomasi (FAP); Verona, San Mauro di Saline, 45.5799°N 11.1115°E (u. = 5 m), 21.IX.2022, 1 specimen, photo by “weston” (OB); Garda, locality San Vigilio, 45.5746°N 10.6752°E (un. = 25 m), 27.VIII.2020, 1 specimen, photo by Joey Bom (OB); Peschiera del Garda, 45.4534°N 10.7007°E (un. = 12 m), 6.VI.2022, 1 specimen, photo by “Jordan” (OB); idem, 45.4444°N 10.6738°E (un. = 900 m), 15.VIII.2021, 1 specimen, photo by “karlien” (OB); idem, San Benedetto, 45.4448°N 10.6799°E (un. = 5 m), 12.VIII.2021, 1 specimen, photo by “niek” (OB); Verona, near Ponte della Vittoria, 45.4418°N 10.9910°E (un. = 10 m), 18.VIII.2011, 6 specimens, photo by Edwin Kerssens (OB); idem, via Monte Santo, 45.457174°N 10.984569°E (un. = 4 m), 23.VI.2020, 1 specimen, photo by “micdallo” (IN); idem, Santa Lucia, 45.383728°N 10.767263°E (un. = 122 m), 21.VIII.2019, 1 specimen, photo by Andrea Angiari (IN); near Verona, 45.4060°N 11.0639°E (un. not recorded), 18.VIII.2021, 1 specimen, photo by “ron” (OB); Colognola ai Colli, 45.433753°N 11.187621°E (un. = 22 m), 16.VI.2022, 1 specimen, photo by Gianfranco Ferrari (IN); Illasi, 45.4674°N 11.1958°E (un. not recorded), 18.VIII.2021, 1 specimen, photo by “joop” (OB); Lazise, 45.4919°N 10.7286°E (un. = 4 m), 6.VIII.2021, 1 specimen, photo by Petra Terpstra (OB); Valeggio sul Mincio, 45.34734°N 10.722085°E (un. = 31 m), 3.X.2019, 1 specimen, photo by Andrea Angiari (IN), idem, 45.3536°N 10.7244°E (un. = 6 m), 1.VIII.2021, 1 specimen, photo by Robin Drenth (OB), idem, Salionze, 45.3966°N 10.7179°E (un. = 5 m), 21.VIII.2017, 1 specimen, photo by Wouter Van Gasse (OB); Villafranca di Verona, 45.355251°N 10.847427°E (un. = 3040 m), 11.IX.2021, 1 specimen, photo by “polvere” (IN); Venezia, Caorle, 45.605066°N 12.880037°E (un. = 55 m), 5.VIII.2022, 1 specimen, photo by Jan Ruzicka (IN); idem, 45.597335°N 12.880065°E (un. = 1.4 km), 3.VIII.2022, 1 specimen, photo by Jan Ruzicka (IN); idem, Duna Verde, 45.5703°N 12.8174°E (un. = 5 m), 13.VIII.2017, 1 specimen, photo by Wouter Van Gasse (OB); Venezia, San Servolo, 45.41968°N 12.355674°E (un. = 40 m), 26.VII.2015, 1 specimen, photo by “ilkkat” (IN); idem, near Lungomare Gabriele D’Annunzio, 45.418808°N 12.383556°E (un. not recorded), 23.VIII.2022, 1 specimen, photo by Dan Wrench (IN); idem, near Casa Museo Andrich, 45.49824°N 12.412358°E (un. = 488 m), 4.VI.2022, 1 specimen, photo by “phobos35” (IN); idem, Riserva naturale Ca’ Roman di Pellestrina, 45.24194°N 12.292425°E (un. = 2 m), 4.VIII.2022, 1 specimen, photo by “cccrll” (IN); San Michele al Tagliamento, Bibione, 27.VII.2015, 1 specimen, photo by Alex Hasenkampf (FEI); Burano, 45.4841°N 12.4198°E (un. = 100 m), 27.VII.2015, 1 specimen, photo by Bep van Pelt-Verkuil (OB); Rovigo, Rosolina, Adige mouth, 45.1535°N 12.3283°E (un. = 10 m), 27.VII.2018, 5 specimens, photo by N. Kroese (OB); Giacciano con Baruchella, III-VII.2012, 3 specimens, photo by Umberto Devincenzo (FAR).
REPUBLIC OF SAN MARINO. San Marino: Borgo Maggiore, Valdragone, 43.939451°N 12.456753°E (un. not recorded), 15.VIII.2021, 1 specimen (Fig. 8), photo by Gertraud Seiser (IN).
SLOVENIA. Central Slovenia: Ljubljana, Ljubljana, Poljanska cesta, 46.0495°N 14.5170°E (un. not recorded), 18.VIII.2022, 1 specimne, photo by Joost Sesink (IN); idem, Gornji trg, 46.046621°N 14.507711°E (un. = 10 m), 28.VI.2021, 1 specimen, photo by Juan Cepeda Espinosa (IN). Coastal–Karst: Piran, Sečovlje, Sečovlje Saltworks, 45.4909°N 13.5948°E (un. = 100 m), 8.VIII.2007, 1 specimen, photo by Jan Westgeest (OB); idem, 45.4919°N 13.6085°E (un. not recorded), 2.VIII.2021, 1 specimen, photo by Joris De Bremme (OB); idem, 45.4890°N 13.6046°E (un. = 17 m), 2.VIII.2022, 1 specimen, photo by “Jarne” (OB). Drava: Selnica ob Dravi, Selnica ob Dravi, 46.5522°N 15.5173°E (un. = 30 m), 14.VIII.2022, 2 specimens, photo by “RangerD” (OB); Ptuj, Ptuj, 46.4172°N 15.8669°E (un.not recorded), 26.VII.2021, 1 specimen, photo by Robbe Van de Wiele (OB). Gorizia: Bovec, Bovec, 46.338517°N 13.55444°E (un. = 35 m), 24.IX.2021, 1 specimen, photo by “kazup” (IN); Kanal ob Soči, Kanal, 46.0847°N 13.6315°E (un. = 10 m), 20.VIII.2017, 1 specimen, photo by Maico Weites (OB). Lower Sava: Brežice, Brežice, 45.927308°N 15.596458°E (un. = 8 m), 13.VI.2022, 1 specimen, photo by “nikelaos” (IN). Mura: Radenci, Živalski Vrt Sikalu Zoo, 46.634727°N 16.027908°E (un. = 39 m), 2.VIII.2020, 1 specimen, photo by “kontra” (IN).
CROATIA. Dubrovnik-Neretva: Korčula island, Korčula, 42.961262°N 17.136013°E (un. = 199 m), 13.VIII.2021, 1 specimen, photo by “nadine_klar” (IN); Mljet island, Malo Lake, 42.779281°N 17.349636°E (un. not recorded), 1.IX.2021, 1 specimen, photo by “earlchesro” (IN); idem, 42.78305°N 17.350833°E (un. = 4 m), 19.VI.2021, 1 specimen, photo by “michaelct” (IN); Okuklje, 42.727119°N 17.668496°E (un. = 2500 m), 7.VIII.2016, 1 specimen, photo by Nuno Veríssimo P. (IN); Koločep island, Donje Čelo, 42.676131°N 18.003156°E (un. not recorded), 3.VI.2022, 1 specimen, photo by “bamsenaattori” (IN); Dubrovnik, near Dubrovačka Sinagoga, 42.64142°N 18.109939°E (un. = 35 m), 4.VI.2022, 1 specimen, photo by “spencerchau” (IN); Trsteno, near Arboretum, 42.7121°N 17.9771°E (un. = 25 m), 11.VIII.2020, 1 specimen, photo byTibor Mikuska (OB); Konavle, Cavtat, 42.58448°N 18.223829°E (un. = 61 m), 9.VIII.2021, 1 specimen, photo by Elena Ilina (IN); idem, 42.576131°N 18.222792 E (un. = 80 m), 8.VIII.2020, 1 specimen, photo by “mjdapifer” (IN). Istria: Lupoglav, Dolenja Vas, 45.3357°N 14.1339°E (un. not recorded), 2.VIII.2022, 1 specimen, photo by “koen vdl” (OB); Poreč, Poreč, Ul. Karla Huguesa, 45.222741°N 13.603645°E (un. = 35 m), 13.IX.2021, 1 specimen, photo by Nina Išić (IN); idem, 45.222324°N 13.603405°E (un. = 4 m), 17.VII.2022, 1 specimen, photo by Nina Išić (IN); idem, Ul. Nikola Tesla, 45.2343°N 13.6003°E (un. = 18 m), 29.VII.2021, 1 specimen, photo by “Kevin” (OB); Labin, Rabac, 45.076637°N 14.169104°E (un. = 18 m), 30.VII.2022, 1 specimen, photo by “mioplax” (IN); Barban, Orihi, 45.068738°N 13.943949°E (un. = 70 m), 2.VI.2018, 1 specimen, photo by “Kienberg” (IN); Marčana, Krnica, 44.9747°N 14.0183°E (un. = 71 m), 20.VII.2022, 1 specimen, photo by Andrew Dingley (OB); Vodnjan, near Vodnjan, 44.9730°N 13.8398°E (un. = 5 m), 13.VIII.2022, 1 specimen, photo by Elbert jan Achterberg (OB); Pula, near Jadreški, 44.864521°N 13.89931°E (un. = 9 m), 22.VII.2022, 1 specimen, photo by Karlo Jušić (IN); Medulin, Vinkuran, 44.836399°N 13.873763°E (un. not recorded), 9.IX.2019, 1 specimen, photo by “amazingalex1010” (IN); near Uvala Debeljak, 44.771337°N 13.918005°E (un. not recorded), 16.VII.2019, 1 specimen, photo by “alessandra” (IN); Ližnjan, Ližnjan, Ul. Istarskog Egzodusa, 44.8275°N 13.9561°E (un. not recorded), 9.VIII.2021, 2 specimens, photo by “GuidoV” (OB). Karlovac: Generalski Stol, between Zvečaj and Gornje Bukovlje, 45.3940°N 15.4361°E (un. = 17 m), 23.VII.2021, 1 specimen, photo by Tibor Mikuska (OB). Krapina-Zagorje: Bedekovčina, Lug Poznanovečki, 46.052451°N 16.017643°E (un. = 1570 m), 14.IX.2020, 1 specimen, photo by Magnus Bunker (IN). LikaSenj: Senj, Senj, Jablanačka ulica, 44.984562°N 14.913125°E (un. = 3 m), 22.VIII.2022, 1 specimen, photo by Emanuel Kern (IN); idem, Krkča ulica, 44.9849°N 14.9083°E (un. = 8 m), 15.IX.2022, 1 specimen, photo by Joris Bolder (OB); Zakosa, 44.9469°N 14.9253°E (un. not recorded), 12.VIII.2022, 1 specimen, photo by “Sylvia” (OB). Osijek-Baranja: Osijek, near Sarvaš, 45.5319°N 18.8462°E (un. = 50 m), 25.VI.2020, 1 specimen, photo by Tibor Mikuska (OB). Primorje-Gorski Kotar: Opatija, Opatija, Viktora Cara Emina, 45.337403°N 14.309895°E (un. = 7 m), 29.VII.2021, 1 specimen, photo by “racn123” (IN); Krk island, Porat, 45.1231°N 14.4946°E (un. not recorded), 2.VIII.2022, 1 specimen, photo by “Andrea” (OB); Vrbnik, 45.076759°N 14.677755°E (un. = 5 m), 4.IX.2022, 1 specimen, photo by “metternich” (IN); near Krk town, 45.043049°N 14.568123°E (un. = 697 m), 4.IX.2021, 1 specimen, photo by Gernot J. Bergthaler (IN); idem, 45.034867°N 14.558463°E (un. = 9 m), 4.IX.2021, 2 specimens, photo by Johannes Schlagbauer (IN); Krk town, near Ul. Krčkih iseljenica, 45.027259°N 14.577609°E (un. = 5 m), 17.VIII.2013, 2 specimens, leg. Filippo Ceccolini, CFC; idem, 45.0270°N 14.5774°E (un. = 25 m), 22.VIII.2018, 4 specimens, photo by Jan Soors (OB); idem, Ul. Vinogradska, 45.029187°N 14.569151°E (un. = 53 m), 18.X.2022, 1 specimen, photo by “croatianiguana” (IN); Punat, 45.01869°N 14.629986°E (un. = 20 m), 3.IX.2022, 1 specimen, photo by Michaela Michal (IN); idem, 45.0157°N 14.6295°E (un. not recorded), 30.VII.2022, 1 specimen, photo by Gernot J. Bergthaler (OB); Cres island, near Martinšćica, 44.816572°N 14.344442°E (un. not recorded), 20.VIII.2021, 1 specimen, photo by “mue-wie” (IN); near Osor, 44.6924°N 14.4025°E (un. = 3 m), 6.IX.2021, 1 specimen, photo by Anke Oertel (OB); idem, 44.69531°N 14.397608 E (un. = 56 m), 2.IX.2022, 1 specimen, photo by Julian Kokott (IN). Šibenik-Knin: Vodice, about 4 km north to Vodice, 43.814276°N 15.766283°E (un. = 4 m), 4.IX.2021, 1 specimen, photo by Sebastian Ćat (IN); Šibenik, Krka National Park, 43.8054°N 15.9640°E (un. not recorded), 29.VII.2022, 1 specimen, photo by Eric Stienen (OB); idem, 43.8054°N 15.9636°E (un. = 222 m), 12.VIII.2022, 1 specimen, photo by “Harold” (OB); Vodice, 43.759813°N 15.76836°E (un. = 8 m), 12.VIII.2021, 1 specimen, photo by Magdalena Magali (IN); Drniš, Skradinski Buk, 43.806332°N 15.963438°E (un. = 98 m), 1.VIII.2021, 1 specimen, photo by “alexxa14” (IN); Primošten, Vezac, 43.582591°N 16.001751°E (un. = 102 m), 13.VI.2022, 1 specimen, photo by “erniefriedo22” (IN). Split-Dalmatia: Seget, Seget Vranjica, 43.51026°N 16.19301°E (un. = 180 m), 25 m, 5.VIII.2016, 1 specimen, leg. Francesca Graziani, CFG; idem, Obala Mira Barešića, 43.509875°N 16.184823°E (un. = 21 m), 27.VII.2022, 1 specimen, photo by “siggees” (IN); Čiovo island, Trogir, 43.515831°N 16.256724°E (un. not recorded), 14.VII.2022, 1 specimen, photo by T. Abe Lloyd (IN); idem, 43.5136°N 16.2452°E (un. not recorded), 14.VIII.2022, 1 specimen, photo by “Naico” (OB); Okrug Donji, 43.498101°N 16.202106°E (un. = 463 m), 5.VIII.2016, 1 specimen, photo by Francesca Graziani (IN); Split, Varoš, 43.505376°N 16.416578°E (un. = 26 m), 13.VIII.2021, 1 specimen, photo by Anton Gjeldum (IN); Dugopolje, near Dugopolje, 43.58728°N 16.608376°E (un. = 92 m), 25.IX.2021, 1 specimen, photo by Anton Gjeldum (IN); Brač island, Dol, 43.351058°N 16.623944°E (un. not recorded), 14.VIII.2022, 1 specimen, photo by Nikola Szucsich (IN); Murvica, 43.267224°N 16.595091°E (un. = 278 m), 5.VII.2019, 1 specimen, photo by “rozi-mari” (IN); Sveti Klement island, Palmižana, 43.158739°N 16.391654°E (un. = 21 m), 28.VII.2020, 1 specimen, photo by “smectit” (IN); Hvar island, Ivan Dolac, 43.128395°N 16.642164°E (un. = 35 m), 11.VII.2022, 1 specimen, photo by “hugogelso” (IN); Zavala, 43.122339°N 16.686345°E (un. not recorded), 14.VIII.2022, 1 specimen, photo by Albert Cardona (IN); about 3 km E to Plaža Dubac, 43.153772°N 16.884153°E (un. = 4 m), 1.VIII.2017, 1 specimen, photo by Oliver Drescher (IN); Vis island, Vis, 43.0592°N 16.1870°E (un. not recorded), 11.VIII.2022, 1 specimen, photo by “Gielen” (OB); idem, 43.059578°N 16.186238°E (un. not recorded), 8.VIII.2022, 1 specimen, photo by Steven Schulting (IN); idem, 43.0581°N 16.1991°E (un. = 10 m), 6.VIII.2012, 1 specimen, photo by Joeri Claes (OB); Tučepi, Tučepi, 43.263203°N 17.058672°E (un. = 8 m), 12.VIII.2021, 1 specimen, photo by “wotsuka” (IN). Varaždin: Varaždin, Varaždin, 46.29288°N 16.35695°E (un. not recorded), 23.VII.2020, 1 specimen, photo by Ivica Pakrac (IN). Zadar: Pag island, 44.5144°N 14.9157°E (un. = 25 m), 23.VII.2020, 1 specimen, photo by Tibor Mikuska (OB); Zverinac island, Zverinac, 44.160487°N 14.915828°E (un. = 8 m), 17.VII.2020, 1 specimen, photo by Martin Scheuch (IN); Ugljan island, Preko, 44.0803°N 15.1840°E (un. = 1000 m), 5.VIII.2019, 6 specimens, photo by Lo Troisfontaine (OB); Pašman island, Ždrelac, 44.012889°N 15.271827°E (un. = 4 m), 1.VIII.2014, 1 specimen (Fig. 9), photo by Marino Linić (IN); Zadar, near Zadar city, 44.1311°N 15.2547°E (un. = 25 m), 27.VIII.2021, 1 specimen, photo by Willem Venhuizen (OB); Sukošan, Sukošan city, 44.0448°N 15.3244°E (un. = 5 m), 5.VIII.2022, 1 specimen, photo by “PS1971” (OB); Novigrad, 44.185832°N 15.541294°E (un = 5 m), 6. VII.2021, 1 specimen, photo by Niklas Wahlberg (IN); Sveti Filip i Jakov, Donje Raštane, 44.020335°N 15.408935°E (un. = 5 m), 20.VII.2022, 1 specimen, photo by “jannieto” (IN); Benkovac, Šopot, 44.023139°N 15.594217°E (un. = 3 m), 21.VIII.2021, 1 specimen, photo by “anke1980” (IN); Pakoštane, Lake Vrana, 43.933824°N 15.509728°E (un. = 95 m), 29.VII.2018, 1 specimen, photo by George Turner (IN). Zagreb: Zagreb city, Štrokinec ul., 45.81°N 15.873056°E (un. not recorded), 17.VI.2022, 1 specimen, photo by Danijel Ostović (IN).
SERBIA. Belgrade: Zemun, Batajnica, 44.905173°N 20.259036°E (un. = 183 m), 3.VIII.2022, 1 specimen, 1 specimen, photo by Viktor Kovačević (IN).
BOSNIA AND HERZEGOVINA. Republika Srpska: Doboj, Nikole Pašića, 44.730951°N 18.091718°E (un. = 5 m), 29.VII.2022, 1 specimen, photo by “harispl” (IN).
ALBANIA. Durrës: Durrës, Durrës, 41.317223°N 19.490833°E (un. = not recorded), 2.VII.2019, 1 specimen, photo by “ariani”(IN); idem, 41.316944°N 19.491112°E (un. not recorded), 31.VII.2019, 1 specimen, photo by “ariani”(IN).
GREECE. Ionian Islands: Corfu island, Benítses, 39.542235°N 19.914353°E (un. = 1.78 km), 21.VII.2022, 1 specimen, photo by “milodoingstuff” (IN).
POLAND. Lesser Poland: Gorlice, near Ropki, 49.449858°N 21.129147°E (un. = 488 m), 23.VIII.2022, 1 specimen, photo by “greenwin” (IN).
CZECH REPUBLIC. Moravian-Silesian Region: Frýdek-Místek, Čeladná, 49.528629°N 18.33469°E (un. = 98 m), 22.VII.2021, 1 specimen, photo by “radek333” (IN). Zlín: Uherské Hradiště, Uherské Hradiště, Města Mayen, 49.070529°N 17.472731°E (un. = 6 m), 27.VI.2022, 1 specimen, photo by Matyáš Adam (IN).
SLOVAKIA. Bratislava: Bratislava, Bratislava, Okres Bratislava I, Trnavské mýto, 48.1584°N 17.1269°E (un. = 25 m), 19.IX.2020, 1 specimen, photo by Lendert Van Laer (OB). Košice: Košice, Košice, Hlavné námestie, 48.72145°N 21.257313°E (un. = 13 m), 3.VIII.2022, 1 specimen, photo by “milesmwb” (IN). Nitra: Nitra, Nové Sady, 48.41462°N 17.9837°E (un. = 12 m), 31.VIII.2022, 1 specimen, photo by “rsl” (IN). Trenčín: Partizánske, Veľké Kršteňany, 48.658475°N 18.400348°E (un. = 2.4 km), 24.VI.2022, 1 specimen, photo by “marekh” (IN).
HUNGARY. Central Hungary: Pest, Pilisborosjenő, 47.598137°N 18.975453°E (un. = 4 m), 7.IX.2021, 1 specimen, photo by Gábor Keresztes (IN); Gödöllő, near Gödöllő, 47.610095°N 19.419978°E (un. = 61 m), 16.VIII.2020, 1 specimen, photo by Attila Steiner (IN); Gödöllői Dombvidék Tájvédelmi Körzet, 47.573897°N 19.455222°E (un. = 4 m), 23.VI.2020, 1 specimen, photo by Attila Steiner (IN); Budapest, Budapest, Ẑldfa utca, 47.58345°N 19.059431°E (un. = 32 m), 11.VI.2022, 1 specimen, photo by “pokelophil” (IN); idem, Filatorigát, 47.552126°N 19.045272°E (un. not recorded), 23.VII.2019, 1 specimen, photo by “vykolakas” (IN); idem, Lake Naplás, 47.507921°N 19.245721°E (un. = 22 m), 20.VIII.2020, 1 specimen, photo by Székely Áron (IN); idem, Erzsébetváros, 47.500025°N 19.062382°E (un. = 4 m), 11.VII.2018, 1 specimen, photo by “ikomposzt” (IN); idem, 47.499945°N 19.062314°E (un. = 4 m), 11.V.2018, 1 specimen, photo by “ikomposzt” (IN); idem, Németv̂lgyi út, 47.486846°N 19.00858°E (un. = 43 m), 8.IX.2020, 1 specimen, photo by “jamesatkins” (IN); idem, Kőérberek, 47.445473°N 18.997909°E (un. = 665 m), 14.VI.2022, 1 specimen, photo by Louisa Z. Riedel (IN). Northern Great Plain: Jász-Nagykun-Szolnok, Szolnok, 47.1814°N 20.1679°E (un. = 5 m), 25.VII.2022, 1 specimen, photo by Johny Berloznik (OB); Heves, Poroszló, 47.651339°N 20.660228°E (un. not recorded), 27.VIII.2019, 1 specimen, photo by Boris Berlijn (IN); Hajdú-Bihar, near Hortobágy, 47.6775°N 21.2372°E (un. = 25 m), 14.VIII.2021, 1 specimen, photo by Leo Vaes (OB); near Debrecen, 47.530181°N 21.798345°E (un. = 8 m), 13.VI.2021, 1 specimen, photo by “iaszabo” (IN). Northern Hungary: Nógrád, Nagyoroszi, 47.991993°N 19.107876°E (un. = 34 m), 23.VIII.2021, 1 specimen, photo by “mrpua” (IN); Borsod-Abaúj-Zemplén, near B̧kkzsérc, 47.9591°N 20.5319°E (un. = 25 m), 15.VI.2022, 1 specimen, photo by Jan Katsman (OB). Southern Transdanubia: Somogy, Siófok, 46.8841°N 17.9581°E (un. not recorded), 27.VI.2022, 1 specimen, photo by “Tamás” (OB); Somogyvár, 46.594767°N 17.637946°E (un. = 95 m), 30.VII.2020, 1 specimen, photo by Zsófia Varga-Szilay (IN); Baranya, Pécs, Rákóczi út, 46.073539°N 18.230906°E (un. = 5 m), 13.VII.2020, 1 specimen, photo by “ter24” (IN); Harkány, Béke utca, 45.849289°N 18.249805°E (un. = 35 m), 2.VIII.2020, 1 specimen, photo by “antalzsuzsa” (IN); Western Transdanubia: Zala, Keszthely, 46.7626°N 17.2502°E (un. not recorded), 15.VIII.2022, 1 specimen, photo by Bart Lemey (OB).
ROMANIA. Bihor: Marghita, Cheț, 47.4140°N 22.3358°E (un = 7 m), 28.VI.2022, 1 specimen, photo by “martijn” (OB). Bucharest: Bucharest, Muzeul Național al Satului ‘Dimitrie Gusti’, 44.472379°N 26.076565°E (un. = 122 m), 5.VIII.2021, 1 specimen, photo by “dmtrsav” (IN): idem, near Parc Dendrologic Agronomie, 44.470708°N 26.062348°E (un. not recorded), 10.VI.2018, 1 specimen, photo by Adrian Ruicanescu posted by “danamihaimileazachi” (IN); idem, Bulevardul Uverturii, 44.444028°N 26.019528°E (un. not recorded), 30.VI.2020, 1 specimen, photo by Emanuel C. (IN). Cluj: Cluj-Napoca, Cluj-Napoca, Gheorgheni, 46.770613°N 23.610308°E (un = 22 m), 13.VI.2020, 1 specimen, photo by Doroteea Ispas (IN); idem, Parcul Central, 46.768613°N 23.578097°E (un. not recorded), 27.VII.2022, 1 specimen, photo by Hugo Sergej Perttersson (IN); idem, Bună Ziua, 46.748322°N 23.599639°E (un. = 754 m), 11.VII.2020, 1 specimen, photo by Adrian Ruicanescu (IN). Dâmbovița: GăeȘti, GăeȘti, 44.716103°N 25.314865°E (un. = 21 m), 1 specimen, photo by “caandrei” (IN). Olt: ScorniceȘti, Jitaru, 44.543907°N 24.545367°E (un. = 20 m), 16.VIII.2022, 1 specimen, photo by Silviu Petrovan (IN).
BULGARIA. Burgas: Nesebăr, Nesebăr, 42.660173°N 27.720559°E (un. = 1.82 km), 10.VIII.2022, 1 specimen, photo by Galin Ivanov (IN); Ravda, 42.644315°N 27.678139°E (un. = 1.31 km), 19.VIII.2022, 1 specimen, photo by Galin Ivanov (IN). Kărdžali: Kărdžali, Kărdžali, ul. Bulair, 41.6359°N 25.3562°E (un. = 25 m), 30.V.2022, 1 specimen, photo by Bert Mertens (OB). Ruse: Ruse, Ruse, ul. Alexandrovska, 43.849365°N 25.953712°E (un. = 12 m), 15.VII.2018, 1 specimen, photo by Cesar Pollo (IN). Sofia: Botevgrad, 42.904094°N 23.808236°E (un. not recorded), 29.VI.2022, 2 specimens photographed but reported about a dozen individuals, photo by Dimitǎr Boevski (IN); Kostinbrod, Balsha, 42.856942°N 23.273839°E (un. = 977 m), 24.VII.2020, 1 specimen, photo by Marti Lubenov (IN).
UKRAINE. Dnipropetrovska: Dnipro, Dnipro city, 48.442868°N 35.074419°E (un. not recorded), 14.VIII.2021, 1 specimen, photo by Vika Roi (UBIN); Novooleksandrivka, 48.352286°N 35.017377°E (un not recorded), 30.VII.2021, 1 specimen, photo by Vika Roi (UBIN). Mykolaïv: Mykolaïv, Mykolaïv city, 11.VII.2016, 1 specimen, photo by “Efarilis” (UBIN). Zakarpatska: Vynohradiv, Vynohradiv city, 48.139609°N 22.993729°E (un. not recorded), 14.VI.2022, 1 specimen, photo by Vasyl Hleba (UBIN).
AUTONOMOUS REPUBLIC OF CRIMEA (disputed territory). Crimea: Simferopol’s’kyi, Hvardiis’ke, 45.113799°N 34.013503°E (un = 22 m), 21.VII.2020, 1 specimen, photo by Veronika Krasnobaeva (IN); Alušta, Alušta, 44.690375°N 34.415462°E (un. = 9 m), 12.VIII.2022, 1 specimen, photo by Maxim Shumskik (IN).
RUSSIA. Krasnodar: Dinskiy, Krasnosel’skoye, 45.26838°N 39.164452°E (un. = 30 m), 16.VIII.2022, 1 specimen, photo by “nikiriki23” (IN); Krasnodar, Krasnodar, Western District, 45.025059°N 38.903922°E (un. = 2 m), 9.VIII.2019, 1 specimen (Fig. 10), photo by Vitaly Lugachev (IN).
description: Published as part of Ceccolini, Filippo, 2023, The worldwide occurrence of Sceliphron caementarium (Drury, 1773) outside its native range, with new records (Hymenoptera: Sphecidae), pp. 1-63 in Zootaxa 5242 (1) on pages 5-38, DOI: 10.11646/zootaxa.5242.1.1, http://zenodo.org/record/7642314