Selenocosmiinae Simon 1889
Creators: name: West, Rick C.
name: Nunn, Steven C.
name: Hogg, Stephen
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Theraphosidae

description: Selenocosmiinae Simon 1889
Selenocosmiini Simon 1889: 204 (sub Selenocosmiae); Simon 1892: 147 (sub Selenocosmieae); Pocock 1895a: 168 (sub Selenocosmiidae); Raven 1985: 118.
Phlogiini Simon 1892: 144 (sub Phlogieae). (synonymised by Simon 1903: 953).
Poecilotheriini Simon 1889: 204 (sub Poecilotheriae); Simon 1892: 144 (sub Poecilotherieae); Schmidt 1995:1–2 (subfamilial elevation); Raven 2005: 16; West et al. 2008: 38. Newly replaced into synonymy of Selenocosmiinae.
Chilobrachini trib. nov. West, Nunn & Hogg
Phlogiellini trib. nov. West, Nunn & Hogg
Diagnosis Maxilla with stridulation organ formed by an oval patch of modified setae (often bacillate) on prolateral face (Figs 8, 9, 47, 49a, 50a, 51a, 52a), originating medially (not ventrally from the oral fringe), opposing modified setae (strikers) in multiple rows on the retrolateral cheliceral surface (Figs 10, 12, 48, 50b, 51b, 52b): maxillary lyra reduced or absent in some genera, cheliceral strikers retained, sternal sigilla largely removed from sternal margins (Fig. 5), many labial cuspules (>200, except Poecilotheria>50) (Fig. 4), leg spines only found distally on metatarsi, tibial apophyses on leg I in mature males absent.
Genera included: Chilobrachys Karsch 1891, Coremiocnemis Simon 1892, Haplocosmia Schmidt & von Wirth 1996, Lyrognathus Pocock 1895, Orphnaecus Simon 1892, Phlogiellus Pocock 1897, Poecilotheria Simon 1885, Psednocnemis gen. nov. West, Nunn & Hogg, Selenocosmia Ausserer 1871, Selenotholus Hogg 1902 and Selenotypus Pocock 1895.
Remarks: West et al. (2008) considered Poecilotheria as its own subfamily Poecilotherinae, presumably following Schmidt (1995). However, we do not recognize Poecilotherinae as a valid subfamily as it lacks any significant synapomorphies to define it from other subfamilies, particularly from Selenocosmiinae (but only synapomorphies of generic significance). Further, dividing the two groups would leave Selenocosmiinae lacking any synapomorphies, something Schmidt (1995) apparently did not consider. With the two groups synonymized, herein, Selenocosmiinae can again be defined by synapomorphies discovered herein (but subject to further testing through the addition of outgroups), and previously (Raven 1985). During the course of gathering data for cladistic analyses, the authors noted a distinct lack of labial cuspules in Poecilotheria species compared to all other Selenocosmiinae and have, herein, adjusted the Selenocosmiinae diagnosis to account for this trait. Additionally, the authors consider the presence of multiple rows of cheliceral strikers a serial homologue for Selenocosmiinae (see Character discussion), but only when Poecilotheriini is included.
description: Published as part of West, Rick C., Nunn, Steven C. & Hogg, Stephen, 2012, A new tarantula genus, Psednocnemis, from West Malaysia (Araneae: Theraphosidae), with cladistic analyses and biogeography of Selenocosmiinae Simon 1889, pp. 1-43 in Zootaxa 3299 on pages 12-13, DOI: 10.5281/zenodo.280919