Fannia tsachilas Grisales & De Carvalho 2019, sp. nov.
Creators: name: Grisales, Diana
name: De Carvalho, Claudio J. B.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Fanniidae
Fannia
Fannia tsachilas

description: Fannia tsachilas sp. nov.
(Figs.1M, 4 J–K, 5M, 7E, 8M)
Diagnosis. Fore tarsus with tarsomeres 3–5 flattened, tarsomere 1 and 2 yellowish on part of v surface.
Description. Holotype male (Fig. 1M)
Head: eye slightly setulose. Frontal vitta velvety black, pollinose. 12 fr. Ocellar triangle black; oc proclinate and developed, 3 less developed setulae on ocellar area. Fronto–orbital plate silver and pollinose. Parafacial bare, silver pollinose. Fronto–genal suture and gena silver pollinose, gena with long setae. Scape black, 2 setae reaching pedicel. Pedicel black pollinose with a dorsal seta developed and surrounded minor setae. Postpedicel black, golden silver pilose, 2 times the length of pedicel. Arista black, with short pubescence, rays shorter than width of base of arista. Palpus black and filiform.
Thorax: black. Scutum without vittae. Acr 2:2–3 arranged in rows. Dc 2:3, long, posterior most postsutural pair developed. Pprn 4, developed. Pra 2. Pro–epimeron setulose.
Wing: brownish, with upper third between vein C and half of cell r 2+3 darker. Calypters whitish. Haltere brown with stem and base lighter in color.
Legs: black, pulvilli brownish and tarsi black, except for fore tarsomere 1 and 2 with part of them yellowish on v surface. Fore femur setulose on posterior surface with 1 row of long pv and d. Fore tibia with 1 strong pre–apical d; 1 apical pv; 1 apical p; 1 apical v. Fore tarsomere 1 and 2 yellowish on part of v surface; tarsomeres 3–5 flattened (Fig. 4J). Mid femur with 2 rows of long pv, hooked apices on apical third and 1 row of developed av decreasing to the apex. Mid tibia on ventral surface with pronounced constriction on basal half area, densely setulose on apical half, 1 v; 2 median ad; 1 d; 2 p; 1 apical pv; 1 pd. Hind coxa bare on posterior margin. Hind femur on ventral surface with pre–apical protuberance; av, v and p surfaces setuloses, av setae longer and curved, increasing in size from base to apex (Fig. 4K). Hind tibia with 1 median d and 1 pre–apical; 1 median a; 1 median av and 1 apical (Fig. 4K).
Abdomen: elongate, black, weakly greyish pollinose with developed setae. Sternite 1 densely setulose. Sternite 5 (Fig. 5M). Terminalia (Fig. 7E): epandrium wider than long, with apical hook extensions; cercal plate pointed apically, base fused almost completely; surstylus strongly articulated with epandrium, pointed apically; bacilliform process spiral and short near epandrium extensions.
Female: unknown.
Biology. Label information indicates that this species occurs in the Andean highlands in Ecuador, from 2500 m in the western foothills. Nothing else is known about its habits or biology.
Comments. Fannia tsachilas sp. nov. is morphologically similar to Fannia iguaque, F. chingaza and F. sutagao described by Grisales, Wolff and Carvalho (2012), due to the fore tarsomeres 3–5 flattened, but differs in the hind femur and the terminalia.
Etymology. Name in apposition. The species epithet refers to the Tsachilás indigenous people, from Ecuador.
Type material. Holotype male (MZSP). ECUADOR: N. Latacunga / Ecuador, 3300m./ XII.1970 / L. E. Peña Leg / MZSP. Paratypes. ECUADOR. Latacunga: N. Latacunga / Ecuador, 3300m./ XII.1970 / L. E. Peña Leg / MZSP (6 males MZSP, 5 males DZUP); Saraguro: 2500m. Ecuador/ XI. 1970 / L. E. Peña col./ MZSP (5 males MZSP, 5 males DZUP); Quito: Ascázubi, Pifo/ N. E. Quito, Ecuador/ 2500 – 2800 m. XII–1970 / L. E. Peña col./ MZSP (1 male, DZUP); Santo. Domingo de los Colorados: Ecuador/ 2600 m. XII–1970 / L. E. Peña col./ MZSP (1 male, MZSP); Otavalo: N. Perucho / Otavalo 2000 m./ Ecuador. I. 1971 / L. E. Peña col./ MZSP (1 male, MZSP).
The holotype has no abdomen because it was taken to dissect the terminalia and it was fixed under the holotype.
Distribution. Ecuador: provinces of Cotopaxi, Loja, Santo Domingo de los Tsáchilas and Imbabúra (Andean region) (Fig. 8M).
description: Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2019, Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America, pp. 330-360 in Zootaxa 4551 (3) on pages 350-351, DOI: 10.11646/zootaxa.4551.3.4, http://zenodo.org/record/2622947