Nanoleon wangae Hu, Lu & Liu 2019, sp. nov.
Creators: name: Lu, Xiumei
name: Hu, Jiahui
name: Wang, Bo
name: Zhang, Weiwei
name: Ohl, Michael
name: Liu, Xingyue
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Neuroptera
Myrmeleontidae
Nanoleon
Nanoleon wangae

description: Nanoleon wangae Hu, Lu & Liu sp. nov.
(Figs 8–10)
Diagnosis. As for the genus.
Material. Holotype, NIGP 168260, amber piece preserving a partly preserved adult female of Nanoleon wangae. It is polished in the form of a flattened subequilateral triangular cabochon, with length × width about 43.0 × 32.4 mm, height about 42.8 mm. Paratype, NIGP 168261, amber piece preserving a nearly complete adult female of Nanoleon wangae and a beetle. It is polished in the form of a flattened subelliptical cabochon, with length × width about 28.5 mm × 21.0 mm, height about 3.87 mm.
Etymology. The new species is dedicated to Prof Xinli Wang, who has made great contributions to the taxonomy of Myrmeleontidae from China, and also helped the last author of this paper in various ways for many years.
Description. Body length 7.36–8.44 mm; head 0.62–0.90 mm long and 1.45 mm wide; distance between compound eyes 0.41 mm; diameter of compound eyes 0.60–0.78 mm; antenna length 1.73 mm; prothorax 0.74 mm long, 0.34 mm wide; forewing 10.17–10.97 mm long and 3.12–3.95 mm wide; hind wing 9.90–10.09 mm long and 2.33–2.36 mm wide; abdomen 5.09–5.73 mm long and 0.5–1.19 mm wide.
Head orthognathous, subtriangular. Compound eyes large, semi-globular; diameter of compound eye slightly longer than distance between compound eyes. Antenna slightly shorter than combined length of head and thorax, dilated at distal 1/4. Mouthparts with parts of maxillae and labium discernible; maxilla with galea slightly broadened and rounded distad; maxillary and labial palpi, each with terminal segment tapering distad.
Prothorax short and slender; meso- and metathorax robust. Wings transparent, immaculate.
Forewing elongated, gradually broadened distad. Costal space rather narrow, about 2.5 times as wide as subcostal space, but distinctly widened from pterostigmal area, with 34 veinlets, most of which are simple. ScA absent; ScP and RA fused together at distal 1/4 of wing, slightly bending and running to wing margin posteriad wing apex; subcostal crossveins absent. One presectorial crossvein present. Five crossveins in RA space present. Hypostigmal cell long. RP þ MA diverging from RA well distad wing base, nearly at proximal 1/3 of wing, with seven pectinate branches and most of them forked distally; infra radial cell short; two short series of gradate crossveins present. MA leaving a short oblique stem near wing base; diverging point of MA from RP þ MA not far beyond diverging point of RP þ MA from RA, with only one crossvein on radial space between these two diverging points. MP1 long, straight, distally trifurcated; base of MP2 (oblique vein) untraceable. CuA and CuP diverging near wing base; MP 2 þ CuA pectinately branched from midpoint, with seven branches; posterior two branches of MP 2 þ CuA1 bifurcated distally; CuA2 slightly shorter than first branch of MP 2 þ CuA1, medially forked. CuP with a short stem discernible, distally fused with A1, with six simple branches; A2 bifurcated, arcuately curved posteriad; A3 simple, strongly arched toward wing margin.
Hind wing elongated, much narrower than forewing, with wing base strongly narrowed, and with wing apex slightly narrowed and feebly excavated along distal margin. Costal space widened from pterostigmal area, with 23 veinlets, most of which are simple. ScP and RA fused together at distal 1/5 of wing, running to wing margin slightly posteriad wing apex, with a few marginally forked veinlets; subcostal crossveins absent. RP þ MA diverging from RA near proximal 1/3 of wing; five crossveins present in RA space; one presectorial crossvein present. MA trifurcated distally. MP1 with only a marginal fork, MP2 pectinately branched into 6–7 single branches; seven crossveins present between MP1 and MP2. CuA stem running parallel to posterior margin, pectinately branched into three simple branches; CuP and anal veins fused together, leaving three short simple branches.
Legs relatively short and stout, densely setose; femur slightly longer than tibia, without sensory hair; tibia with sparsely spaced long spinous setae, tibial spurs present, almost straight; tarsus much thinner and shorter than tibia, 5-segmented; tarsomere 1 long, but slightly shorter than combinational length of remaining tarsomeres; tarsomere 2–4 short, almost equal in length; paired pretarsal claws present, slenderly elongate, and curved distally, seemingly not opposable on last tarsomere.
Abdomen slenderly elongated, almost 2/3 × body length. Female genitalia preserved; tergum 8 much shorter than tergum 7; gonocoxites 8 not discernible; tergum 9 strongly broadened ventrad in lateral view; gonocoxites 9 small, with arcuate ventral view, bearing long setae; ectoprocts nearly semicircular in lateral view, with large ovoid callus cerci.
description: Published as part of Lu, Xiumei, Hu, Jiahui, Wang, Bo, Zhang, Weiwei, Ohl, Michael & Liu, Xingyue, 2019, New antlions (Insecta: Neuroptera: Myrmeleontidae) from the mid-Cretaceous of Myanmar and their phylogenetic implications, pp. 1215-1232 in Journal of Systematic Palaeontology 17 (14) on pages 1223-1224, DOI: 10.1080/14772019.2018.1517132, http://zenodo.org/record/15148813