Prodilis joanna Gordon and Hanley 2017, new species
Creators: name: Gordon, Robert D.
name: Hanley, Guy A.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Coccinellidae
Prodilis
Prodilis joanna

description: 47. Prodilis joanna Gordon and Hanley, new species   
 Description. Male holotype. Length 1.4 mm, width 1.4 mm; body nearly round, elytron with side rounded, wider than pronotal base, widest at middle of elytra. Dorsal surface entirely shiny. Color yellowish brown (Fig. 581); head black except narrow apex of frons yellowish brown (Fig. 583); pronotum medially dark brown, lateral 1/3 yellowish brown; antenna, mouthparts, epipleuron, legs yellow; abdomen brownish yellow except median portion of basal 3 ventrites brown. Head punctures large, separated by a diameter or less; pronotal punctures much smaller than on head, separated by less than 3 times a diameter; elytral punctures slightly larger than on pronotum, separated by 1 to 3 times a diameter; prosternal, mesosternal punctures large, separated by a diameter or less; metasternal punctures as large as on mesosternum, separated by less than a diameter except more widely spaced medially; abdomen with punctures on ventrites 1–2 large, separated by less than 3 times a diameter, punctures on remaining ventrites smaller, separated by about a diameter. Head with frons not widened from vertex to clypeus, lateral margins parallel, 1.4 times width of eye measured at vertex; eye canthus short; apical maxillary palpomere short, widened from base to apex. Pronotum widest at middle, reflexed lateral margin wide, equal in width from base to apex. Epipleuron flat, wide in basal ½, as wide as pronotal hypomeron, without feeble depressions for reception of femoral apices. Prosternum longer than wide, longer than short mesosternum, base arcuate medially, lateral carina slender, extended to apex of procoxa. Postcoxal line on ventrite 1 short, rounded, extended slightly beyond middle of ventrite (Fig. 582) Apex of ventrite 5 truncate. Genitalia with long phallobase, basal lobe longer than paramere, slender, lateral margin gently curved to bluntly rounded apex, apex not emarginate; paramere nearly straight, sinuate in lateral view, without marginal serrations (Fig. 584); sipho lost.   
 Female. Similar to male except female genitalia with spermathecal capsule, short, slightly curved medially.   
 Variation. Width 1.3–1.4 mm. Lateral pronotal margin sometimes entirely black or with reduced pale margin.   
 Type material. Holotype male; COSTA RICA: Heredia, Est. Biol. La Selva. 50–150m, 10 o 26' N 84 o 01W, Prov. Alas, INBio–OET, FOT/41/01–40, Tachigalia costaricensis, 28 Diciembre 1999. (USNM). Paratypes; 2, same data as holotype. (USNM).   
 Remarks. Prodilis joanna is a small, distinctive species characterized by a unique dorsal color pattern, a black male head, widely spaced elytral punctures, and the male genitalia.
description: Published as part of Gordon, Robert D. & Hanley, Guy A., 2017, South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae), pp. 1-158 in Insecta Mundi 2017 (601) on page 90, DOI: 10.5281/zenodo.5170031