Sticholotis quadriguttata
Creators: name: Tomaszewska, Wioletta
name: Łączyński, Piotr
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Coccinellidae
Sticholotis
Sticholotis quadriguttata

description: Sticholotis quadriguttata (Gorham)   
(Figs. 2, 56–67)  
 Orcus quadriguttatus Gorham, 1895: 689.   
 Sticholotis quadriguttata: Łączyṅski & Tomaszewska, 2009: 609.   
 Diagnosis. This species is easily distinguished among species of Sticholotis from Myanmar by its small body and two large, brown maculae on each elytron.   
 Redescription. Female. Length 2.1 mm; TL/EW = 1.15; PL/PW = 0.47; EL/EW = 0.85; GD/TL = 0.5.   
Body (Figs. 2, 56) rounded, strongly convex; pronotal margins very narrow, visible from above; elytral margins comparatively explanate, entirely visible from above. Pronotum, scutellum and background of elytra brownish black; head and labrum dark reddish brown. Each elytron with two large, round, reddish brown spots along mid line of disk, narrowly connected to each other; elytral and pronotal lateral margins reddish brown. Punctures on pronotum 1.5–2.5 diameters apart, rather fine and moderately deep; punctures on elytra slightly coarser and deeper than those on pronotum, 1.0–2.0 diameters apart; surfaces between elytral and pronotal punctures polished and shiny; dorsum apparently glabrous. Ventral surface dark reddish brown with meso- and metaventrite, especially along middle and anterior part of abdominal ventrite I, infuscate. Ventral mouthparts and antennae yellowish brown; legs slightly darker.  
Head (Figs. 57, 59) flat medially, finely punctate and moderately densely setose. Clypeus weakly arcuate anteriorly, scarcely reflexed at anterior edge. Eyes moderately large, rather coarsely faceted, dorsally separated by about 3 times eye width; interocular distance nearly 0.65 times head width; inner margins of eyes slightly rounded, convergent anteriorly. Maxilla (Figs. 58, 59) with terminal palpomere about 2.3 times longer than wide, parallel-sided along basal half of its length, strongly obliquely truncate towards apex; labial terminal palpomere narrowed and acuminate, distinctly narrower than penultimate palpomere. Antenna (Fig. 58) 11-segmented with narrow 3-segmented club.  
Prothorax (Figs. 57, 62) about 0.9 times base of elytra; pronotum with groove extending parallel along base, curving anteriorly near hind angles; pronotal hypomeron and prosternum smooth; anterior lobe of prosternum distinctly bordered with anterior edge emarginate medially; prosternal process (Figs. 59, 60) subtruncate at apex, without distinct lateral carinae, moderately coarsely and rather sparsely punctate, punctures with moderately long setae. Mesoventral intercoxal process (Fig. 61) about 1.35 times mesocoxal diameter. Metaventrite with complete discrimen, moderately coarsely and sparsely punctate; postcoxal lines curved and complete. Elytral epipleuron broad with maximum width at metaventrite, narrowing posteriorly but complete to apex, without distinct foveae. Wings well-developed.  
Abdomen (Figs. 63, 64) with 5 ventrites; ventrite I along mid line about 3.0 times longer than ventrite II; postcoxal line of first ventrite curved posteriorly and laterally, closely paralleling posterior margin, incomplete laterally; postcoxal disk microreticulate and sparsely punctate (Figs. 63, 65); ventrite V triangularly produced posteriorly; abdominal segment VIII in female as in Fig. 66.  
Female genitalia as in Fig. 67. Proctiger (T10) well developed, somewhat triangular in shape; ovipositor sclerotized, coxites triangularly oval, entirely separated; styli absent; spermatheca not observed.  
Male not known.  
 Material examined. Types. Lectotype (here designated), female: “ Rangoon Birmania, Fea V. 1885 / Typus / quadriguttatus Gorh. / Orcus quadriguttatus / O. quadriguttatus Gorh. typus!/ Syntypus, Orcus quadriguttatus Gorham, 1895 / Museo Civico di Genova 1895” (MCSN).   
 Note. The designation of the lectotype for O. quadriguttatus Gorham, 1895, is made to fix the taxonomic status of this species.
description: Published as part of Tomaszewska, Wioletta & Łączyński, Piotr, 2010, A contribution to Sticholotis Crotch from Myanmar (Coleoptera: Coccinellidae: Sticholotidini), pp. 51-68 in Zootaxa 2658 (1) on page 65, DOI: 10.11646/zootaxa.2658.1.3, http://zenodo.org/record/5302384