The Froggattimyia-Anagonia Genus Group (Diptera: Tachinidae)
Creators: name: Colless, Donald H.
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy

description: Genus Anagonia    
 Anagonia Brauer & Bergenstamm, 1891:348. Type species Anagonia spylosioides Brauer & Bergenstamm, 1891 (= Masicera rufifacies Macquart, 1847) by original designation and monotypy.   
 Acephana Townsend, 1916:153. Type species Masicera rubrifrons Macquart, 1847 (= Masicera rufifacies Macquart, 1847) by original designation.   
 Opsophana Townsend, 1916:153. Type species Masicera rufifacies Macquart, 1847, by original designation.   
All synonymy by Crosskey, 1966:95.  
The diagnosis is given in the foregoing key.  
  Identification of species of Anagonia    
 Females of most species are unrecognized or cannot at present be accurately identified, although a few bear distinctive attributes that link them to the male. For males, the variability of most attributes makes it very difficult to produce a sufficiently accurate, conventional key for their identification. Some characters, too, need a little experience; e.g., whether an eye is haired or not, or whether the tibiae are paler than the femora; and some are often badly preserved. I have tried to catch any leakage over such couplets by multiple entries and the key given below will normally give the correct identification. However, complete confidence will always require the examination of the terminalia, which is required in any case for a few species.The male of A. grisea is unknown and excluded from this key (see p. 192).   
  Key to males of Anagonia    
1 Eye profusely haired, or if moderately so, hairiness widely  
distributed; ocellar bristles inconspicuous or absent; tibiae often  
paler than femora........................................................................................................................... 2  
—— Eye hairs absent or very sparse and inconspicuous and/or ocellar setae well developed...................................................................................................................... 6  
 2 Tergite 4 with 1 or more stout discal bristles; intrapostalar bristles finely differentiated on one or both sides............................................... A. lasiophthalma Malloch   
—— Tergite 4 without discal bristles; intrapostalar usually not developed............................................... 3  
3 Abdominal tergite 3 with submedian pair of marginal bristles finely developed or absent....................................................................................................................... 4  
—— Tergite 3 with submedian marginals well developed.................................................................... 5  
 4 Presutural intra-alar bristle very fine or absent on both sides; upper occiput with mainly dark scales......................................................... A. perplexa sp. nov. (in part)   
 —— Presutural intra-alar bristle well developed on one, usually both, sides; upper occiput with mainly pale scales........................................ A. loripes sp. nov. (in part)   
5 Terminalia as in Figs 23–26; cerci in lateral view diagonally  
truncate at apex and longer than vertical height of syntergo-  
sternite 6–8; eye conspicuously hairy in specimens from cooler  
 climates........................................................................................................ A. rufifacies Macquart   
—— Terminalia as in Figs 30, 31; cerci in lateral view almost mammili-  
form, with rounded apex, and no longer than vertical height of  
 syntergosternite 6–8; eye often only moderately haired.................. A. conformis sp. nov. (in part)   
 6 Foretarsi with all or most segments conspicuously pale brown...................... A. tillyardi Malloch   
—— Foretarsomeres dark brown or black............................................................................................. 7  
7 Ocellar bristles strongly developed; legs dark, tibiae concolorous  
with femora; tergite 3 with strongly developed pair of submedian  
marginal bristles............................................................................................................................. 8  
—— Without this combination of attributes; ocellar bristles at most weakly developed.......................................................................................................................... 9  
 8 Presutural intra-alar bristles usually undifferentiated; intrapostalar absent; tergite 5 with small discal bristles........................................................... A. opaca Malloch   
 —— Presutural intra-alar bristles and intrapostalar bristle clearly differentiated; tergite 5 without discal bristles.................................................... A. zentae sp. nov.   
 9 Abdomen in glancing posterior view intensely silver-pollinose with black spots around hair bases; thorax usually lacking presutural median vitta; (hindtibia with pd1 clearly as long as or longer than sdd)....................................................................................................................... A. major Malloch   
—— Abdomen in posterior view with brownish sublateral patches; thorax usually with distinct presutural median vitta.................................................................... 10  
10 Abdominal tergite 3 with submedian marginal bristles clearly differentiated, longer than adjacent bristles.......................................................................................... 11  
—— Tergite 3 with submedian marginals scarcely or not at all differentiated or extremely short.......................................................................................................... 16  
11 Presutural intra-alar bristle very fine or absent on both sides...................................................... 12  
—— Presutural intra-alar well developed on at least one side............................................................. 13  
 12 Tibiae slightly, but distinctly, paler than femora; usually only 2 postocellar bristles; frons/head width less than 0.2............................ A. angustifrons sp. nov. (in part)   
 —— Tibiae dark, concolorous with femora; usually more than 2 postocellar bristles; ratio frons/head width greater than 0.2;..................................... A. similis sp. nov.   
 13 Hindtibia with Pd1 bristle clearly as long as or longer than Sdd................................................. 25   
 —— Hindtibia with Pd1 scarcely as long as, or clearly shorter than Sdd............................................ 14   
14 Intrapostalar bristle well differentiated on at least one side, usually both.............................................................................................................................................. 15  
—— Intrapostalar bristle weakly or not at all differentiated.................................................................. 5  
 15 Tibiae dark, concolorous with femora; terminalia as in Figs 69–71............. A. lateralis Macquart   
 —— Tibiae at least a little paler than femora; terminalia as in Figs 47–49;............................................................................................... A. dayi sp. nov. (in part)   
16 Presutural intra-alar bristles well developed on at least one side (usually both)............................................................................................................................... 17  
—— Presutural intra-alar bristles very fine or absent on both sides.................................................... 30  
17 Upper occiput with hairs all or mostly pale................................................................................. 18  
—— Upper occiput with hairs all or mostly dark................................................................................ 21  
 18 Intrapostalar bristle well differentiated on at least one side........................... A. commoni sp. nov.   
—— Intrapostalar bristle at most very weakly differentiated.............................................................. 19  
 19 Hindtibia with Pd1 clearly longer than Sdd; tibiae dark, concolorous with femora; apical ad spine on foretibia often very small or vestigial...................................................................................... A. anguliventris Malloch (in part)   
 —— Hindtibia with Pd1 shorter than Sdd; tibiae usually at least slightly paler than femora; ad spine on foretibia normal.......................................................................... 20   
 20 Tibiae brown, at least slightly paler than femora; eye at least slightly haired;................................................................................................... A. loripes sp. nov. (in part)   
 —— Tibiae black, concolorous with femora; eye scarcely or not at all haired.................................................................................................... A. norrisi sp. nov. (in part)   
 21 Foretibia with ad spine more or less vestigial........................... A. anguliventris Malloch (in part)   
 —— Foretibia with ad spine clearly developed................................................................................... 22   
22 Intrapostalar bristle not differentiated.......................................................................................... 23  
—— Intrapostalar bristle at least weakly differentiated....................................................................... 28  
 23 Hindtibia with Pd1 bristle clearly longer then Sdd; (scutellum   
usually with apical bristles straight or slightly downcurved, parallel  
or divergent)................................................................................................................................. 24  
 —— Hindtibia with Pd1 bristle scarcely as long as, or (usually) distinctly shorter than Sdd; (apical scutellar bristles upcurved or at least directed upwards)......................................................................................................................... 25   
 24 Terminalia as in Figs 35, 36.......................................................................... A. propinqua sp. nov.   
 —— Terminalia as in Figs 32–34........................................................................... A. scutellata Malloch   
 25 Tibiae usually paler than femora; eye usually at least slightly haired............................................................................................... A. conformis sp. nov. (in part)   
—— Tibiae dark, concolorous with femora; eye scarcely or not at all haired........................................................................................................................................... 26  
26 Apical scutellar bristles upcurved................................................................................................ 27  
 —— Apical scutellar bristles more or less horizontal; terminalia as in Figs 61–63........................................................................................................ A. latistylus sp. nov.   
 27 Terminalia as in Figs 66–68................................................................... A. uptoni sp. nov. (in part)   
 —— Terminalia as in Figs 58–60.................................................................. A. norrisi sp. nov. (in part)   
 28 Hindtibia with Pd1 bristle clearly longer than Sdd......................................... A. crosskeyi sp. nov.   
 —— Hindtibia with Pd1 no longer than Sdd..................................................... A. dayi sp. nov. (in part)   
 29 Upper occiput with mainly pale scales................................................. A. norrisi sp. nov. (in part)   
—— Upper occiput with mainly dark scales........................................................................................ 30  
30 Tibiae at least slightly, but distinctly, paler than femora............................................................. 31  
—— Tibiae dark, concolorous with femora......................................................................................... 32  
 31 Terminalia as in Figs 64, 65; ratio Frw/Hdw 0.1–0.2.................. A. angustifrons sp. nov. (in part)   
 —— Terminalia as in Fig. 78; ratio Frw/Hdw 0.2–0.3.............................. A. perplexa sp. nov. (in part)   
 32 Intrapostalar bristle small but distinct; terminalia as in Fig. 76.................. A. teratostylus sp. nov.   
—— Intrapostalar bristle not differentiated; apical scutellar bristles upcurved or at least directed upwards.............................................................................................. 33  
 33 Surstylus lacking anteriorly-directed apical hook or spine (Fig. 78)............................................................................................... A. perplexa sp. nov. (in part)   
—— Surstylus with a minute apically-directed or anteromediallydirected hook or spine (Figs. 79, 80)........................................................................................... 34  
 34 Surstylus 1.5 times or less than length of syncercus (Figs 66, 67, 79).................................................................................................... A. uptoni sp. nov. (in part)   
—— Surstylus 2 or more times as long as syncercus (Figs 56, 57, 77, 80)................................................................................................................................................ 35  
 35 Posterior margin of surstylus with distinct angle (Figs 66, 67, 77)...................... A. minor sp. nov.   
 —— Posterior margin of surstylus more evenly rounded (Fig. 80)............................ A. errator sp. nov.
description: Published as part of Colless, Donald H., 2012, The Froggattimyia-Anagonia Genus Group (Diptera: Tachinidae), pp. 167-211 in Records of the Australian Museum 64 (3) on pages 183-186, DOI: 10.3853/j.0067-1975.64.2012.1590, http://zenodo.org/record/4684127