Kumatoeides anomala Gómez-Zurita 2018, sp. nov.
Creators: name: Gómez-Zurita, Jesús
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Chrysomelidae
Kumatoeides
Kumatoeides anomala

description: Kumatoeides anomala sp. nov.
(Figs 1f, 2a, 3a, 4a, 5e)
Holotype: Male, New Caledonia (N), Mt. Mandjélia, rainforest, -20.40439 164.51798, 550–600 m, 19.xi.2008, beating, leg. M. Wanat, HOLOTYPE Kumatoeides anomala sp. nov. J. Gómez-Zurita det. [red label] (MNHN). Paratype: Female, New Caledonia (N), -20.40439 164.51798, 550–600 m, Mt Mandjélia, beating, rainforest, 19.xi.2008, leg. M. Wanat PARATYPE Kumatoeides anomala sp. nov. J. Gómez-Zurita det. [red label] (MNHW).
Other material examined: (1) one female, New Caledonia (N), -20.40007 164.52788, Mt Mandjélia, 700– 780 m, montane rainforest, 20.xi.2008, leg. M. Wanat, Kumatoeides anomala m. J. Gómez-Zurita det. 2018 (MNHW); (2) two females, New Caledonia (N), -20.40007 164.52788, Mt Mandjélia, 700–780 m, montane rainforest, 20.xi.2008, leg. M. Wanat, Kumatoeides anomala m. J. Gómez-Zurita det. 2018 (MNHW); (3) one male, New Caledonia (N), 21°09'S 165°19'E, Aoupinié, 420–530 m, road to sawmill, 7.ii.2004, leg. M. Wanat, Kumatoeides anomala m. J. Gómez-Zurita det. 2018 (MNHW).
Head, pronotum, scutellum, ventral thoracic parts and trochanters dark brown, with slight bronze metallic luster on head, pronotum and scutellum; elytra hazelnut brown with suture, punctures and inner border of epipleura dark brown; labrum, palpi, antennae and legs yellowish orange (Fig. 1f). Length: 2.77 mm; width: 1.39 mm.
Head hypognathous at rest; frons and vertex regularly convex, microsculptured, sulcate medially and with scattered strong, deep punctation; sulcus above eyes imprecise; glabrous except for long seta above eye. Clypeus nearly as long as wide at apex, microsculptured, punctate. Labrum transverse, smooth, weakly emarginate at apex. Apical maxillary palpomere spindle-shaped, obliquely cut shortly at apex. Eyes large, bulging, entire. Antennae slender, reaching beyond middle of elytra; scape nearly cylindrical, slightly bent posteriorly, twice as long as thick, strongly narrowed at base; pedicel cup-shaped, half as long as scape; antennomeres 3–10 club-shaped, 3–6 glossy with sparse long setae, 7–11 slightly incrassate, finely granulose, clothed with denser, shorter hairs; antennomeres 3, 4 and 6 subequal, longer than pedicel; antennomeres 5 and 8–10 subequal, longer than sixth and shorter than seventh; eleventh antennomere longest, pointed, obliquely cut at apex internally. Pronotum transverse (1.44x as wide at base than long at middle), convex with stronger slopes at sides and toward anterior angles; anterior border nearly straight, slightly produced forward medially, unmargined, narrower than posterior border, weakly bisinuate, with very short median lobe before scutellum, finely margined; anterior angles obtuse, feebly callous, with setigerous pore anteriorly at middle; posterior angles obtuse, very slightly salient laterally; sides feebly sinuate before base, widest after middle, with explanate margin; surface glabrous, shiny, very finely microsculptured, slightly irregular with transverse depression at anterior half, and with dense, strong deep roundish punctures larger than distance separating them, sometimes confluent. Hypomera smooth, very finely shagreened, with few scattered punctures near basal border; anterior border very feebly convex. Prosternum very narrow before procoxae, with anterior border concave and finely margined; prosternal process relatively wide, about 0.8x as wide as transverse diameter of procoxae, slightly widened at apex to enclose procoxae posteriorly, finely rugose-punctate at both ends, glossy and punctate at middle, with disheveled long, fine translucent hairs. Mesoventral process as wide as prosternal process, shortly expanded before convex apical border, glossy, with scattered fine punctures and very short fine hairs. Metaventrite slightly longer than mesoventrite, smooth on disc, with fine transverse impressions near basal border and deep narrow median notch at apical border, sides finely microreticulate with sparse punctures and short fine translucent hairs. Scutellum arched, as long as wide at base, round at apex, finely microreticulate. Elytra long, as wide at base as base of pronotum, gently curved at sides, broadest at middle and regularly curved at apex; elytra with very weak transverse impression at basal 1/3; humeri strongly marked, unpunctured; eight regular striae on apical half of each elytron, with only striae 6–8 reaching humeral area more or less regularly; striae on apical half of elytra impressed with longitudinally cojoined slit-like small punctures or dot-like, deeper punctures in outer rows; punctation in basal half of elytra, including regular outer striae deep, strong, dense and generally confused; punctures of scutellar stria confused with surrounding punctation; short subhumeral, curved stria of large, deep puntures joining premarginal stria before middle of elytron; intervals convex in regularly striated part of elytra; intervals very finely microreticulate. Femora strongly widened medially; anterior and hind tibiae nearly as long as their corresponding femur and mesotibiae slightly shorter; tarsi much shorter than corresponding tibia, with first pro- and mesotarsomeres expanded; claws appendiculate. Abdominal ventrites finely shagreened with sparse fine punctures and short appressed translucent hairs; last visible ventrite weakly emarginate at apex, with small curcular median impression before apical border. Penis (Fig. 2a) narrow, slender, strongly curved ventrally at base and weakly bent toward apex; more or less parallel-sided except at level of operculum in dorso-ventral view, slightly broadened before converging to apex; dorsal flap of operculum short triangular; apical end of penis expanded as subrectangular tooth with blunt angles.
Females (Fig. 4a): The females of K. anomala sp. nov. lack modified tarsomeres in pro- and mesotarsi, have more convex elytra with raised costae in their lateral declivities, and the punctures on pronotum and in basal area of elytra are stronger and tighter; moreover, at least their head, pronotum, scutellum, suture and all ventral surfaces (as in the paratype) and also the elytra (in all other females examined) are dark metallic bronze. Spermatheca (Fig. 5e) weakly sigmoidal; cornu bulbous, shorter than wide at base, connected to nodulus, more than twice as long as cornu, narrowed at middle, at level with insertion of spermathecal gland, wider at distal half compared to bulbous basal half, with spermathecal duct inserted at base, opposite to cornu orientation.
Diagnosis: The defining trait for this species, absent in all other known species of the genus is the lack of ordered punctation at the base of elytra. Apart from this trait, obvious both in males and females, it can be distinguished from K. metallica sp. nov., the most similar species, and also found in Mt. Mandjélia, by the lack of intense green metallic shine on dorsal surfaces in males.
Derivatio nominis: The name derives from the Greek adjective (fem.) όνώµαλoς, meaning anomalous, unusual, in reference to the scrambling of punctures at the base of elytra, which deviates from the norm for the genus.
Distribution: The few known specimens of this species were found in two distant localities at mid elevations in the mountains of Mandjélia and L'Aoupinié (Fig. 3a).
description: Published as part of Gómez-Zurita, Jesús, 2018, Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia, pp. 89-115 in Zootaxa 4521 (1) on pages 93-94, DOI: 10.11646/zootaxa.4521.1.4, http://zenodo.org/record/2609701