Toechorychus vilhenus Tedesco & Aguiar 2013, sp. nov.
Creators: name: Tedesco, Anazélia M.
name: Aguiar, Alexandre P.
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Toechorychus
Toechorychus vilhenus

description: Toechorychus vilhenus Tedesco, sp. nov.   
(Figs 12, 99, 138, 172)  
Description. Holotype FEMALE. Fore wing length 6.79 mm.  
 Head. Mandible 1.46 as long as basal width, moderately pilose; ventral margin projected as crest; dorsal tooth distinctly longer than ventral tooth. Clypeus 1.67 as wide as high, subquadrate, entirely finely strigulate; apex 1.41 as long as base, truncate; apical margin sharp, medially slightly concave. Supraclypeal area entirely strigulate, also densely punctate, moderately pilose, medially slightly prominent; between antennal foramens with V-shaped carina, medially interrupted; radicle foveolate. Antenna with 25 flagellomeres; white band starting at flagellomere 4, reaching flagellomere 12; flagellum slender; subapical flagellomeres slightly flattened. Supra-antennal area without distinct median carina, medially with small rounded elevation, near antennal sockets striolate; dorsal half medially strigulate. Paraocular area smooth. Vertex with very coarse punctures around ocelli, without sulcus; gena and vertex behind ocelli smooth, densely pilose; gena in lateral view at level of dorsal portion of occipital carina narrow, ventrally wide; occipital carina faint, dorsally uniformly arched, ventrally slightly projected as crest, reaching hypostomal carina at mandible base or nearly so; hypostomal carina slightly projected as crest; malar space 1.14 as long as basal width of mandible.   
 Mesosoma. Mesosoma in dorsal view 2.04 as long as wide; in lateral view, mesosoma middle width 0.19 mm. Pronotum centrally smooth, latero-ventrally rugulose behind collar, margin near mesopleuron not corrugated; pronotal swelling finely rugulose; collar dorso-laterally rounded, distinctly swollen, anteriorly punctulate; epomia absent. Lobes dorsally punctulate, laterally markedly corrugated or covered with sparse coarse punctures; notauli faintly impressed, posteriorly parallel throughout. Scutoscutellar groove faintly corrugated; scutellum 1.00 as long as wide, smooth; scutellar carina incomplete, restricted approximately to anterior 0.3; postscutellum 0.32 as long as wide; hind margin of metanotum with two short lateral carinae extending towards transverse furrow at base of propodeum. Subalar ridge somewhat rounded; mesopleuron dorsally strigate, include over hypoepimeron, ventrally strigulate; mesopleural groove corrugated; epicnemial carina stout, dorsally straight; dorsal end of epicnemial carina reaching 0.2 of posterior margin of pronotum, behind epicnemial carina without rounded confused-rugulose area; sternaulus complete, reaching base of mid coxa, moderately sinuate, posteriorly almost faint; faintly corrugated; scrobe deeply impressed, forming pit. Mesothoracic venter strigulate; median portion of posterior transverse carina of mesothoracic venter convex, regularly shaped, without projections. Transverse furrow at base of propodeum laterally 0.09 as long as anterior portion of propodeum, shallow and wide, uniformly corrugated; propodeum 1.31 as long as wide medially; anterior margin of propodeum without teeth; anterior portion of propodeum finely strigulate or punctulate, medially without longitudinal carinae; lateral longitudinal carina of propodeum represented by short curved carina; posterior portion of propodeum entirely markedly strigate, medially without distinctly stout longitudinal carina; anterior transverse carina of propodeum complete, faint, medially slightly arched forwards; posterior transverse carina absent; propodeal spiracle 1.36 as long as wide; pleural carina of propodeum absent, apparently present because of sculpture patterns of lower division of metapleuron and propodeum; lower division of metapleuron faintly strigulate, densely pilose, juxtacoxal carina absent.   
 Wings. Fore wing: vein 2+3Rs slightly concave; ramellus absent; vein 1M+Rs entirely sinuous; vein 1cu-a posteriorly markedly curved, arising far from vein 1M+Rs base, angle with vein M+Cu about 90°; vein 2Cu 0.73 as long as vein 2cu-a; vein 4Rs 1.10 as long as vein 4M, straight; cell 1+2Rs 0.80 as high as pterostigma, rectangular; vein 2M approximately as long as vein 3M; vein 3M distinct, 0.50 as long as vein 2M. Hind wing: vein 1Cu 1.09 as long as vein cu-a; vein 2-1A reaching 0.79 of distance to posterior margin.   
 Legs. Tibia with very sparse short bristles; hind coxa globose.   
 Metasoma. T1 0.52 as long as hind femur, without basolateral tooth; minutely coriarious; spiracle at anterior 0.54, slightly prominent; postpetiole dorso-anteriorly flat; T2–8 minutely coriarious; sparsely pilose; T2 1.25 as long as wide at apex; apex of T2 1.92 as wide as base. Dorsal valve of ovipositor with nodus indistinct; apex of lower valve with series of teeth; notch absent.   
 Color. Head and mesosoma pale yellow with black marks, metasoma orange (206,091,028). Head: pale yellow (238,221,131); mandible teeth, supra-antennal area medially, interocellar area, vertex, and occiput, black; scape ventrally yellow, dorsally brown; flagellum ventrally brown, dorsally entirely whitish at f5–7, partially at f4 and f8–9. Mesosoma: mostly pale yellow; pronotum black, except for lateral of collar and pronotal swelling, pale yellow; mesoscutum black, except longitudinal lanceolate mark at entire length of central lobe; mesopleuron pale yellow with two black marks: anteriorly around hypoepimeron, and ventral half of epicnecium; remaining of mesosoma pale yellow, except for the following black marks: scutoscutellar groove, both axillary trough of mesonotum and metanotum, tegula apically, carinal triangle, and anterior portion of propodeum; posterior portion of propodeum with three longitudinal stripes, basally black, apically orange. Legs: all legs entirely orange (211,089,024), yellowish towards apex, except for all t5, dark brown (086,067,053). Metasoma: orange; S2–8 medially white.   
 Comments. Very similar to T. fluminensis sp. nov., from which it can be readily differentiated by lacking epomia (vs. with very stout epomia, restricted to space between posterior margin of collar and pronotal swelling); and metasoma entirely orange (vs. striped; Fig. 187). Also similar to T. amapaeus sp. nov., from which it can be separated by having mesopleuron ventrally and sternaulus, yellow (vs. with rounded black spot just behind epicnemial carina, and along posterior half of sternaulus; Fig. 13); hind coxa dorsally yellow (vs. entirely orange; Fig. 168); T1 with basolateral tooth, as in Fig. 53 (vs. without, as in Fig. 54).   
 Male. Unknown.   
 Distribution. Known only from Brazil (Fig. 233).   
 Biology. Unknown.   
 Material examined. Holotype ♀, BRAZIL: Rondônia, Vilhena, XI.1973, M.Alvarenga leg., Toechorychus, Townes det. (AEIC). Complete, in good shape, except for mesoscutum somewhat damaged by pin.
description: Published as part of Tedesco, Anazélia M. & Aguiar, Alexandre P., 2013,  Phylogeny and Revision of  Toechorychus  Townes (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of thirty-five new species , pp. 1-138 in Zootaxa 3633 (1) on pages 71-72, DOI: 10.11646/zootaxa.3633.1.1, http://zenodo.org/record/5261931