Campsicnemus gladiator Evenhuis, new species
Creators: name: Evenhuis, Neal L.
Publisher: name: Zenodo
Year: 2009
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Dolichopodidae
Campsicnemus
Campsicnemus gladiator

description: Campsicnemus gladiator Evenhuis, new species
(Figs. 2–3, 11)
Diagnosis. Easily distinguished from the other members of the gladiator subgroup by the lack of a conspicuous modified spur apically on the male mid basitarsus (present in various shapes in C. limnobates, C. scurra, and C. uncleremus) and the apical insertion of IIt2 onto IIt1 (insertion is subapical in the other species).
Description. Male. Body length: 2.5–2.8 mm. Wing length: 2.2–2.5 mm.
Head. Black, face dark brown to black with reddish highlights, brown near clypeus; oc and vt black, about one-third length of antennal arista; front, occiput, and vertex black with blue highlights; face constricted at middle, almost holoptic, eyes separated below antennae by width of 1–2 ommatidia; palp small, dark brown; proboscis brown, extending below eye in lateral view; antennal segments black to brown; scape length ca. 2.5 x length of pedicel; postpedicel long, arrow-shaped, thin apically, length about 3.5 x width; arista slightly longer than head height.
Thorax: Mesoscutum and scutellum, brown, dark brown dorsomedially on mesonotum, with a few metallic greenish highlights laterally; pleura yellow except brown anepimeron; thoracic setae black: 1 + 3 dc; 2 np; 2 ph; 1 pa; 1 + 1 sc; ac absent; halter stem white, knob dark brownish black.
Legs: CI yellowish white, with apical tuft of three strong black setae, approximate basally, almost fused apically so as to appear as a single strong seta; CII and CIII brown, slightly paler than surrounding pleura; FI and FII yellowish brown, FIII yellowish brown basally, becoming darker brown on apical one-fourth; remainder of legs brown; FI and remainder of foreleg unmodified, without MSSC; FII slightly bowed with strong chetae basally, setae thinner toward apex, with subapical brown patch (sensory area?) harboring two small setae; FIII slightly bowed, middle of FIII bearing very long curved, sinuous, and wrinkled setae on ventral surface (MSSC); TiII slightly bowed, thin basally, swollen on apical half, with long setae in middle on dorsal surface (MSSC), smaller black setae and hairs along entire ventral surface. TiIII with row of stiff long hairs basoventrally, dorsal surface with small hairs and only two long setae (one subbasally; the other near middle) (MSSC). IIt1 (Fig. 3) short, about three-fourths length of IIt2, with single strong black, lateroapical seta (MSSC); IIt2 inserted apically on IIt1. Remainder of leg segments without MSSC.
Wing: pale smoky throughout; posterior crossvein perpendicular to vein M.
Abdomen. Brown with short black hairs dorsally on each tergite, a few longer hairs laterally; tergal interstices whitish; sternites paler brown than tergites. Hypopygium brown with paler brown cerci, not dissected.
Female. As in male except for lack of MSSC; postpedicel subconical, length subequal to width; legs normal, without modifications; FI and FII brown.
Types. Holotype ɗ (BPBM Type 16,633) [BPBM1013183] and 5ɗ [BPBM1013184] paratypes from FRENCH POLYNESIA: Marquesas: UA POU: Paeoa River, 1150–1500 ft [350–457 m], 17 Aug 2001, pools/ cascades, R.A. Englund. Other paratypes: FRENCH POLYNESIA: Marquesas: U A P OU: 24ɗ [BPBM1013185], 19Ψ [BPBM1013186], Upper Paeoa River, 1200 ft [366 m], 12 Aug 2001, skating on pools, D.A. Polhemus; 5ɗ [BPBM1013187], 11Ψ [BPBM1013188], Middle Paeoa River at road, 650 ft [198 m], 9°24'01"S, 140°03'30"W, skating on stream pools, 19 Aug 2001, D.A. Polhemus; 3ɗ [BPBM1013189], 2Ψ [BPBM1013190], Pouaoa River cascade, 400 ft [122 m] (see Fig. 11), 9°25'15"S, 140°05'27"W, skating on stream pools, 18 Aug 2001, D.A. Polhemus; Upper Anakooma River, 1100–1300 ft [335–396 m], 9°23'36"S, 140°03'49"W, skating on stream pools, 19 Aug 2001, D.A. Polhemus. Holotype in BPBM. Paratypes in BPBM and USNM.
Etymology. The specific epithet derives from the Latin gladiator = “swordsman”; referring to an anthropomorphic similarity of this species possessing a spike-like cluster of setae on the fore coxae of males as if it were ready to go into battle like a gladiator.
description: Published as part of Evenhuis, Neal L., 2009, Review of Campsicnemus (Diptera: Dolichopodidae) of the Marquesas, French Polynesia, with description of four new species groups, pp. 25-48 in Zootaxa 2004 on pages 29-30, DOI: 10.5281/zenodo.185641