Lophophaena kaonashii, n. sp.
Creators: name: Trubovitz, Sarah
name: Renaudie, Johan
name: Lazarus, David
name: Noble, Paula
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Chromista
Radiozoa
Polycystina
Nassellaria
Plagiacanthidae
Lophophaena
Lophophaena kaonashii

description: Lophophaena kaonashii n. sp.   
Plate 34, Figs. 1A – 8B.  
urn:lsid:zoobank.org:act: 8C9A5395-F8AD-43EE-BE0B-64C2173A99D2  
 Arachnocorys? sp 2/3 partim., Trubovitz et al., 2020, supplementary data 7.   
 Diagnosis. Lophophaena with a heavily-silicified, spherical cephalis that has 1 or 2 conical horns unrelated to the apical spine, and a shouldered thorax with large pores around the top and no appendages.   
 Description. The cephalis of this species is spherical in shape and is more heavily silicified than the thorax. Well-preserved specimens show strongly-framed pores on the cephalis and many small bumps on the nodes of pores, which appear to have served as the bases of tiny, needle-like spines. 1, or rarely 2, prominent horns are present near the apex of the cephalis, but these are not related to the apical spine, which merges with the shell wall immediately above the collar structure and is so muted that it is difficult to detect. The thorax has a ring of large pores around the top where it connects to the cephalis. It reaches its widest point at the shoulders, and then tapers downward slightly. The lateral and dorsal spines merge with the thorax wall and do not form appendages. Pores on the thorax are small and tend to be more closely spaced than those on the cephalis.   
 Remarks. This species is structured similarly to Lithomelissa dybbuki (Pl. 19, Figs. 1–7), but differs in heaving a more heavily silicified cephalis, with large pores around the base. In addition, the apical spine of L. kaonashii does not pass through the center of the cephalis, disqualifying it from being in Lithomelissa.   
 Material examined. 16 specimens from samples 321-1337A-31X-6W, 4–6cm (Middle Miocene), 321-1337D-30H- 3, 103–104cm (Middle Miocene), 321-1337D-26H-3W, 142–144cm (Late Miocene), 321-1337D-23H- 6, 134– 137cm (Late Miocene).   
 Holotype. Pl. 34, Figs. 5A–B; sample 321-1337D-23H- 6, 134– 137cm; ECO-147; W22-4.   
 Paratypes. (1) Pl. 34, figs. 9A–B; sample 321-1337D-26H- 3, 142–144cm; ECO-148; G8-4. (2) Pl. 34, figs. 4A–B; sample 321-1337A-31X-6, 4–6cm; ECO-151; O11-4. (3) Pl. 34, figs. 8A–B; sample 321-1337A-31X-6, 4–6cm; ECO-151; T 25-1. (4) Pl. 34, fig. 7; sample 321-1337A-31X-6, 4–6cm; ECO-151; U32-2. (5) [not figured] sample 321-1337D-26H- 3, 142–144cm; ECO-148; Z8-1.   
 Measurements. Cephalis height 34–41 (38)μm; cephalis width 30–40 (36)μm; neck width 25–34 (30)μm; width at shoulders 40–55 (48)μm. Based on 14 specimens.   
 Etymology. Named for the spirit Kaonashi (“No-Face”) in Hayao Miyazaki's animated film, Spirited Away.   
 Range. Middle—Late Miocene in the EEP, lower boundary not determined (Table 1).
description: Published as part of Trubovitz, Sarah, Renaudie, Johan, Lazarus, David & Noble, Paula, 2022, Late Neogene Lophophaenidae (Nassellaria, Radiolaria) from the eastern equatorial Pacific, pp. 1-158 in Zootaxa 5160 (1) on pages 60-61, DOI: 10.11646/zootaxa.5160.1.1, http://zenodo.org/record/10544058