Tylotrypes Bezzi 1914
Creators: name: Korneyev, V. A.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Pyrgotidae
Tylotrypes

description: <i>Tylotrypes</i> Bezzi, 1914 (fig. 10) Type species: <i>Tylotrypes immsi</i> Bezzi, 1914. D i a g n o s i s. Robust, densely setulose dark brown flies with darkened wings (fig, 10, <i>1, 2, 4, 13, 14</i>); head without subocular sclerite; face with (fig. 10, <i>5</i>) or without carina (fig. 10, <i>15</i>); wing with straight and entire vein Sc, straight R 2+3, and without a spurious vein, costa more or less thinned posterior of R 4+5 apex; femora without ventral rows of shortened and thickened setae, usually densely setose; femoral organ in females usually lacking, except in <i>T. fuscipes</i> (Wulp); scutellum heavily setulose; synsternite 1+2 wide, with setulose posterior part; epandrium setulose dorsally, with moderately short surstyli (medial surstylus without prensisetae); hypandrium with two posterior lobes forming a phallic guide between surstyli (fig. 10, <i>10</i>, phg); oviscape with long medioventral sclerotized lobe, but without hook- or nail-like unpaired sclerotized process (fig. 10, <i>6, mvl</i>), often with calluses on ventral surface (fig. 10, <i>2, 4, 6, 14, opc</i>); aculeus with basal part thickened dorsoventrally, but neither Y- or T-shaped (fig. 10, <i>7, 8</i>); three sclerotized sausage-like spermathecae (fig. 10, <i>9</i>), one of them smaller and shorter. <i>Tylotrypes</i> shares the absence of the subocular sclerite, and straight veins Sc with <i>Pyrgota</i> Wiedemann and <i>Tephritopyrgota</i> Hendel and apparently is related to them, differing by having very long medioventral lobe of the oviscape apex instead of a short hook and also by the shape of aculeus with its basal part thickened dorso-ventrally and triangular in profile (fig. 10, <i>7, 8</i>, a unique character state not occurring elsewhere, possibly a synapomorphy of <i>Tylotrypes</i>). R e m a r k s. I have examined about two dozen specimens from Nepal, Thailand, Malaysia, Mainland China, Taiwan, and Indonesia (Java, Borneo), which represent six to eight species of the genus <i>Tylotrypes</i> similar or identical to species described by Shi (1994; 1996) from China (Yunnan) as <i>Apyrgota</i> species. As the type material is unavailable at this moment and illustrations and descriptions are incomplete or based on males alone, identification of that material with Chinese nominal species is pending until a forthcoming revision of <i>Tylotrypes</i>. Judging from the photographs of the type specimen of <i>Apyrgota longa</i> Shi kindly provided by Dr. Li-hua Wong (Beijing) it must be with certainty transferred from <i>Apyrgota</i> to <i>Tylotrypes</i>, which now contains seven nominal species: <i>Tylotrypes breviventris</i> (Shi, 1996) (<i>Apyrgota</i>) (China: Yunnan), <b>comb. n.;</b> <i>Tylotrypes fura</i> (Shi, 1996) (<i>Apyrgota</i>) (China: Yunnan), <b>comb. n.;</b> <i>Tylotrypes fuscipes</i> (Wulp, 1885) (<i>Campylocera</i>) (Indonesia: Java), <b>comb. n.;</b> <i>Tylotrypes immsi</i> Bezzi, 1914 (India); <i>Tylotrypes jiangleensis</i> (Shi, 1994) (<i>Apyrgota</i>) (China: Fujian), <b>comb. n.;</b> <i>Tylotrypes longa</i> (Shi, 1996) (<i>Apyrgota</i>) (China: Yunnan), <b>comb. n.,</b> and <i>Tylotrypes hispida</i> (Kim &amp; Han, 2001), <b>comb. n.</b> (<i>Adapsilia</i>) (South Korea). These taxonomic acts are based either on the material examined (to be listed in the forthcoming publication) or on the comparison of the diagnosis of the genus with original descriptions. Some of these species were described from males and females alone, and can belong to the same species; I do not synonymize them here as it needs study of additional material and will be subject to a taxonomic revision (Korneyev, in preparation). This paper is partly resulting from the study supported by the Belgian Federal Office for Scientific, Technical and Cultural Affairs Research Fellowship and conducted in the Royal Museum for Central Africa, Tervuren, Belgium in 2005–2006; most type specimens were photographed there. My thanks are due to Ben Brugge (RMNH), Robert Copeland (NMKE), Allen L. Norrbom (USDA SEL, Washington, D. C., U. S. A., material from USNM), David Notton (BMNH), Thomas Pape (ZMUC), Peter Sehnal (NHMW), and Chen W. Young (CMNH), who kindly put at my disposal important material, both type and undetermined, or provided necessary data on the specimens deposited in collections under their care. Chris Raper kindly read the early proof of this paper. I thank Robert Copeland and an anonymous reviewer for their criticism and valuable comments.
description: Published as part of <i>Korneyev, V. A., 2015, Pyrgotid Flies Assigned To Apyrgota. Iii. Species Of Afropyrgota Gen. N. And Tylotrypes (Diptera, Pyrgotidae), pp. 25-40 in Vestnik Zoologii 49 (1)</i> on pages 38-40, DOI: 10.1515/vzoo-2015-0003, http://zenodo.org/record/6404027
description: {"references": ["Bezzi, M. XXI. - Indian Pyrgotinae (Diptera) // The Annals and Magazine of Natural History. Ser. 8. - 1914. - 14. - P. 153 - 163.", "Shi, Y-s. Pyrgotidae / Eds. W. Xue, C. Chao. Flies of China. -. Shenyang: Liaoning Science and Technology Press, 1996. - Vol. 1. - P. 575 - 595. - Chinese."]}