Clivina acutimentum Balkenohl 2021, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina acutimentum

description: <i>Clivina acutimentum</i> sp. nov. urn:lsid:zoobank.org:act: 99384C09-2B9C-4F01-86A6-495DE04A49B9 (Figs 27, 47, 67, 94, 111) ETYMOLOGY. The name refers to the mentum with its acute lobes and is composed of the Latin noun “mentum” and the Latin adverb “ <i>acutatus</i>, a, um”. TYPE MATERIAL. Holotype: • &amp;male;, NW Thailand, 30.iv.1992, MAE HONG SON 350 m, S. Bilý leg. (NHMB). Paratypes: N-VIETNAM. • 1 &amp;female;, Vietnam Tam Dao 10–25.vi.1992 V. Seidl / <i>Cl. castanea</i> WESTW. Balkenohl iv.96 / COLL. Wrase Berlin (CWGP); • 2 specs., Annam Phuc-Son Nov. Dez. H. Fruhstorfer (MFNB). LAOS. • 1 &amp;male;, LAOS, PHONGSALI PROV. MEUNG KHOA DISTR. BAN HOUAYWANG 10–17.vi.2010 LG. SASA KAMPHILAVONG (CADW); • 1 spec., N-Laos Khan River, at light 5 km E Luang Prabang, leg. K.W. Anton, 21.iv.1999 (CMBB); • 1 spec., LAOS B. Na Gnao le 1.ii.1918, R. Vitalis de Salvaza / <i>Clivina castanea</i> West. det. R. Vitalis de Salvaza / I.R.Sc.N.B 16.493 (RBINS); • 3 specs., LAOS Pak Pha le 4–ii.1918, R. Vitalis de Salvaza / <i>Clivina castanea</i> West. det. R. Vitalis de Salvaza / I.R.Sc.N.B 16.493 (RBINS); • 5 specs., LAOS, SEKONG prov. ca. 12 km S Sekong TAD FAEK waterfalls (at light) 15°14.7’N, 196°45.1’E, 188 Ji&amp;rcaron;í Hájek leg., 8–12.v.2010 / <i>Clivina castanea</i> s.l. Westwood 1837 P. Bulirsch det. 2010 (NHMP). THAILAND. • 1 &amp;male;, THAILAND 1988 CHANG MAI 18–25.vi. ex coll W ZELENKA / <i>Clivina castanea</i> Westw. det. Alex. Dostal 1993 (CADW); • 1 &amp;male;, same data as holotype (CMBB); • 5 specs., THAIL. Juli 1990 160 km N W Bangkok 90 m leg. Thielen / 10 km W Han-Kha Lichtfang (NHMW); • 2 specs., Thailand, 9.xi. 1992, 12 km SW Pak Chong 130 km NO Bangkok Mischobstplantage, 400 m, Lichtfang. Thielen leg. (CMBB); • 1 &amp;male;, 3 &amp;female;&amp;female;, 1 spec., THAILAND, 230 km NW Bangkok, W. Thielen leg. vii. 1988, 95 m NN (CMBB); • 1 &amp;male;, same data (NHMW); • 2 specs., Thailand Han-Kha, 60 km n. Suphon-Buri, 1.i.1990, leg. Thielen (CMBB); • 1 spec., Thailand 55 km w. Utahi-Thani ii.1992, leg. W. Thielen (CMBB); • 1 &amp;male;, 1 &amp;female;, 2 specs., Thailand 3 km w. Ban Rai 170 km nw Bangkok, xi.1991, leg. Thielen <i>/ Clivina castane</i> a WESTW. det. Balkenohl’ 95 (CMBB); • 1 &amp;male;, 1 &amp;female;, 3 specs., THAIL., vii. 1990, 150 km NW Bangkok 90 m, leg. Thielen / 10 km W Han-Kha Lichtfang (NHMW); • 2 &amp;male;&amp;male;, NO-Thailand Khon Kaen 21.ii.1981, lux leg. S. Saowakontha (MFNB); • 1 &amp;male;, same data but 26.i.1981 (MFNB); • 4 specs., same data but 23.xi.1980 (MFNB); • 1 spec., same data but 25.ii.1981 (MFNB); • 1 spec. same data but 25.5.1980 (MFNB); • 1 spec., same data but 29.viii.1980 (MFNB); • 11 specs., Nordost-Thailand Khon Kaen Gelände mit Sekundärwald in der Nähe / eines Teiches, am Licht 10.ix.1978, leg. H.J. Bremer (MFNB); • 1 &amp;male;, 1 &amp;male;, 4 specs., THAILAND: Amphoe Chom Thong, Ban Huai Muang 4.i.1989 250 m, leg. TRAUTNER &amp; GEIGENMÜLLER (SMNS); • 3 specs., same data (CMBB); • 2 specs., C-THAILAND: Lan-Sak 20 km, W Uthai-Thani ca. 80 m, vii.1986, leg. W. Thielen (SMNS); • 1 spec., Thailand, 25 km NW Uthai Thani, v.1989, leg. Thielen / <i>Clivina castanea</i> WESTW. (CMBB); • 1 &amp;male;, 1 &amp;female;, 5 specs., THAILAND, vi. 1990, 25 km NW LanSak 65 km, NW Uthai Thani leg. Thielen / collection NATURKUNDE-MUSEUM ERFURT / <i>Clivina castanea</i> WESTW. det. M. Balkenohl x.97 (NKME); • 2 spec., same data but (CMBB); • 1 spec., THAILAND: Tap Tan, 20 km WNW Uthai Thani, 260 km NW Bangkok. W. THIELEN leg. 1–3.i.1979 (CMBB); • 2 &amp;male;&amp;male;, Thailand, Tap Tan Uthai Thani 200 m, i.1979 leg. Thielen / <i>Clivina castanea</i> Westw. det. Alex. Dostal 1982 (CADW); • 1 spec., C-THAILAND; Krok-Phra 10 km S Nakhon-Sawan ca. 50 m vii.1986, leg. W. THIELEN (SMNS); • 2 specs., THAILAND 25 km NW Lan Sak, x.1989, W. Thielen (CWGP); • 2 specs., THAILAND 230 km NW Bangkok W. Thielen leg., vii. 1988, 95 m NN (SMNS); • 27 specs., NW THAILAND, 24–27.iv.1991 CHOM THONG LEG PACHOLATKO / <i>CLIVINA (Leucocara) castanea</i> Westwood, 1837, P. Bulirsch det. 2016 (NHMB); • 3 specs., same data as before but without determination label (CMBB); • 3 specs., NE Thailand, 23–27.iv.1991, Chom Thong S. Bilý leg. / <i>Clivina castanea</i> WESTW. det. Balkenohl, 2007 (NHMB); • 13 specs., NW THAILAND Chom Thong 24.–27.iv.1991, J. Horák leg. / <i>Clivina castanea</i> W. det. Alex. Dostal 1997 (CADW); 1 spec., THAILAND, 5.iv.1991, KANCHANABURI, leg. PACHOLÁTKO (NHMB); • 1 spec., Thailand, Kanchanaburi, 31.iii.1994, S. Biliý leg. / <i>CLIVINA castanea</i> Westwood P. Bulirsch det. 1999 (NHMP); • 1 &amp;male;, 3 &amp;female;&amp;female;, 1 spec., THAILAND, S 40 km NE OF BETONG, LAKE BANG LANG (KG.TO) 12–23.vi. 2007, 350 m Z. LINEK leg. (CADW); • 1 &amp;female;, THAILAND PAI env. 1– 14.v.2001, J. Hromádka leg. (CADW); • 1 &amp;male;, 2 &amp;female;&amp;female;, THAI, 29.iv.1993, PAI City Pacholatko &amp; Dembicky leg. (CADW); • 1 spec., Thailand – nord Pai 22–29.v.1996, A. Kudrna jr. leg. (CMBB); • 1 spec., Siam / Soc.Ent.Belg. Coll. PUTZEYS / <i>Clivina agona</i> Putz. dét. J. Putzeys (RBINS); • 1 &amp;male; [immature], THAILAND Prov. Ayutthaya Umg. 26.i.1995 Weide leg. A. Weigel / <i>Clivina castanea</i> WESTW. det. Balkenohl, ix.96 (NKME). CAMBODIA. • 1 &amp;male;, 1 &amp;female;, Coll. I.R.Sc.N.B. Cambodia – 8 km north of Sre Noi (road to Along Vaeng) Light trap 29.v.2003, leg. J. Constant &amp; K. Smets (RBINS). DIAGNOSIS. Amedium sized species with smooth lateral margin of the pronotum and elytron, and isodiametric reticulation on intervals five to eight of the elytron. Distinguished from the other species with isodiametric reticulation on intervals five to eight of the elytron by the shape of the lobe of the mentum which is distinctly triangle-like acute antero-laterally. The other species with a similar acute lobes is <i>C. agumbea</i> sp. nov., but in that species the anterior margin of the lobe is slightly convex, the lateral margins of the pronotum are attenuating anteriorly and are not spherical as it is in <i>C. acutimentum</i> sp. nov. In addition, <i>C. agumbea</i> is a much bigger species. DESCRIPTION. <i>Measurements</i>. Holotype: Body length 8.53 mm, width 2.51 mm; ratio length/width of pronotum 0.89; ratio length/width of elytra 1.89. Other material: Body length 7.75–9.11 mm (: = 8.52 mm *), width 2.31–2.68 mm (: = 2.51 mm *), ratiolength/width of pronotum 0.89–0.93 (: = 0.91*), ratiolength/widthofelytra 1.88–1.94 (: = 1.9*); (*n = 10). <i>Colour</i>: Glossy to partly semi-opaque. Piceous. Intermediate and hind legs dark fuscous, antenna fuscous. <i>Head</i>: Athird narrower than pronotum. Clypeus slightly excised anteriorly; wing prominent, as protruding as clypeus, separated from clypeus by distinct notch; supraantennal plate moderately convex; clypeus laterally with carina, wing amargined, supraantennal plate reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by distinct notch. Clypeus transverse, slightly convex and glossy, separated from frons by moderately deep furrow. Frons moderately convex, with slight pore at middle, with lateral carina at posterior eye-level. Clypeus and frons smooth, with minute scattered punctures, separated from wing and supraantennal plate by deep moderately wide furrow continuing posteriorly aswide supraorbital furrow with two supraorbital setigerous punctures at posterior eye level. Neck constriction developed as small step, with row of small punctures. Eye hemispherical projecting. Gena small. Antenna reaching over posterior seta of pronotum, antennomeres four to ten oblongoelongate (ratio L/W around 2.2). Labrum indistinctly convex anteriorly, obtusely pointed at middle, with indistinct reticulation, seven setose. Mentum (Fig. 47) with longitudinally reticulated surface, with lobe flattened, shape of lobe like a conspicuously askew trapezium, with slightly concave anterior margin and distinctly acute-angled lateral anterior angle, all margins carinate, with medial angle obtuse, epilobe small, developed at medial angle only, with complete carina at middle, median tooth wide, slightly less protruding anteriorly as lobe, obtuse angled anteriorly. <i>Pronotum</i> (Fig. 67): Disk somewhat flattened in lateral view, moderately convex in frontal view. Slightly wider than long. Reflexed lateral margin smooth, indistinctly convex in anterior third, widest in posterior third; anterior angle slightly not projecting; posterior angle marked by distinct tooth, tooth not protruding laterally over convexity of lateral margin. Lateral channel moderately wide, reticulated, stepped from disk. Median line distinct; anterior transverse line punctured, joining median line, finely joining basal constriction. Surface smooth, with some transverse wrinkles, with distinct isodiametric reticulation at base. <i>Elytron:</i> Disk flattened in anterior half in lateral view, moderately convex in frontal view. Nearly twice as long as wide, indistinctly dilated in apical half with maximum width behind middle. Reflexed lateral margin smooth. Scutellar striole distinct; setigerous tubercle at base of first stria, with distinct tubercle at base of third interval. Striae moderately deep, punctuatestriate, one to three free at base, four to seven joining at humerus, one joining with seven at apex, two ending free at apex, three and four, and five and six deep and joining apically. Intervals moderately convex, eights with short indistinct carina at humerus, carina at apex distinct but relatively short. Third interval with four setigerous punctures, first situated at second stria near base, the other three approaching third stria. Intervals on disk with isodiametric reticulation of decreasing intensity, most distinct on interval eight, very faint on interval four (best visible along striae), intervals three to one glossy. Reticulation more distinct in apical quarter. <i>Hind wings</i>: Fully developed. <i>Lower surface</i>: Proepisternum with transverse wrinkles, in lateral half with distinct isodiametric reticulation, with short rough longitudinal scars towards middle. Terminal sternite of abdomen developed apically as deep narrow transverse fold. <i>Legs</i>: Femur covered with isodiametric and tibiae with irregular reticulation. Protibia with three spines of moderate length, carinate dorsally, movable spur distinctly arcuate in apical third. <i>Male genitalia</i> (Fig. 94): Median lobe long, slender, of moderate diameter, regularly arcuate, apex developed as long slightly sinuate stick-like spatula, rounded at apex, ventral rim distinct, base with small apophysis, on the surface with large group of microtrichia at middle. Endophallus apically with small group of short bristles. Ventral paramere moderately wide, regularly arcuate, lateral apophysis medium sized, apex slender, with two long setae at apex and one small one preapically. Dorsal paramere regularly curved, at apex with two long and one smaller setae and one small seta preapically. <i>Female coxostylus and laterotergite</i> (Fig. 111): Coxostylus moderately broad and moderately curved, dorsally with eight long to medium sized and one distinctly elongated and robust nematiform setae in basal half, ventrally with three long and numerous smaller setae, with one SSO. Laterotergite with five long and one small setae basally and one isolated seta towards coxostylus. VARIATION. On the disk of the elytron the reticulation varies to a certain degree. In most of the specimens it is present on intervals five to eight and most distinctly developed on interval eight. It is always more distinctly present along the striae. In some specimens traces are only visible on the inner intervals. Usually, the distinctness of the reticulation decreases considerably towards middle but there are also two specimens with very indistinct reticulation on the inner intervals (one to four). The short longitudinal scars and wrinkles on the proepisternum vary slightly in their number and deepness. DISTRIBUTION. Known from North East and Central Thailand, Cambodia as well as from North Vietnam.
description: Published as part of <i>Balkenohl, Michael, 2021, Belgian Journal of Entomology, pp. 1-83 in Belgian Journal of Entomology 115</i> on pages 37-40, DOI: 10.5281/zenodo.5645953
description: {"references": ["WESTWOOD J. O., 1837. - A collection of insects collected at Manilla by Mr. Cuming. Proceedings of the Zoological Society, 5: 127 - 130."]}