Trichoclinocera stigmatica Saigusa & Sinclair, 2016, sp. nov.
Creators: name: Saigusa, Toyohei
name: Sinclair, Bradley J.
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Empididae
Trichoclinocera
Trichoclinocera stigmatica

description: Trichoclinocera stigmatica sp. nov.
(Figs 8 C, 9G, 10, 15A–C, 16G, 17M, N, 18M, N)
Type material. HOLOTYPE ♂, labelled: “(OITA)/Kyûsuikei/ Kujûsan/ 28.v.1972 / T. Saigusa”; [HOLOTYPE]/ Trichoclinocera / stigmatica / Saigusa et/ Sinclair, 2016 [red label]” (KUMF). PARATYPES: JAPAN. KYUSHU. Fukuoka: Fukuoka-shi, Nogochi, 7.vi., 17.viii.1997, 24.iv.1998, BJS (4 ♂, 5 ♀, CNC); Magaribuchi (Nogochi), Sawara, 5.x.1961, TS (11 ♂, 4 ♀, KUMF, CNC). Kumamoto: Hae, Itsuki village, 25.viii.1997, BJS (1 ♂, 2 ♀, CNC); Hirasawado, Gokanoshô, 3.viii.1975, TS (10 ♂, 2 ♀, KUMF); Nakamichi-bashi, 400 m, Izumi-mura, 24.v., 25.viii.1997, BJS (26 ♂, 22 ♀, CNC); Izumi-mura, Momiki, 20.viii.1999, BJS (2 ♂, 2 ♀, CNC); Shiratoriyama, Izumi-mura, 12–14.v.1983, K. Ôhara (1 ♂, KUMF). Miyazaki: Seki-no-O, 28.iv.1963, S. Takagi (2 ♂, HUS). Oita: Kyûsuikei, Kujûsan, 21.v.1962, TS (42 ♂, 74 ♀, KUMF, BMNH, CNC); same locality, 19.v.1963, TS (9 ♂, 19 ♀, KUMF); same data as holotype (108 ♂, 55 ♀, KUMF).
Recognition. Distinguished from other Japanese species by its distinct pterostigma, presence of antero- and posterodorsal bristles on mid and hind femora, and face with bright blue pruinescence.
Description. Wing length 4.8–5.4 mm. Male. Clothed in blue pruinescence except vertex of head, scutum, and dorsum of abdomen brown. Head rounded; compound eyes large, rounded. Face with blue pruinescence, with broad, median brown patch on carina and narrow dorsal stripe; face narrowest just below antennae, slightly wider than antennal sockets, lower margin of face with deep cleft and distinct carina; gena one-fifth as wide as height of eye. Ocellar bristles longest among cranial bristles; occiput with 2 pairs of bristles posterior to ocellar triangle; postocular with row of short bristles along margin of eye, upper third darker and more stout than lower bristles. Arista-like stylus long and slender, tapered apically. Palpus clavate, length greater than half height of eye.
Pleura, postpronotal lobe, notopleuron, postalar, prescutellar depression, and scutellum with bright blue pruinescence; posterolateral corner of scutum near postalar ridge bluish; scutum faintly bivittate. Thoracic setae long, some ca. as long as width of eye; acrostichal setulae lacking; 6–8 pairs dc; 1 pprn; 1 presut spal; 3 npl; 2 psut spal; 1 pal; 1 pair sctl, lacking setulae on disc; several setulae on postpronotal lobe and notopleural depression. Laterotergite with patch of pale setae; prothoracic episternum with long pale setae; katepisternum with pale setulae near middle, metepisternum with pale setulae at posterodorsal margin.
Wing narrow, infuscate (Fig. 16 G); pterostigma narrow, distinct, elliptical. Basal costal seta ca. subequal to posterior dc. Row of setulae from base of R to tip of R1; cell dm produced apically; R4 and R5 curved, parallel except extreme apex divergent. Halter brown with pale base.
Legs greyish, generally darker towards tips, with coxae and trochanters clothed in blue pruinescence. Coxae with long pale, marginal setae. Fore femur (Figs 17 M, N) with 2 stout, oblique anterior setae on apical third; widely spaced row of 3–5 anterodorsal setae on apical two-thirds, less than half width of femur; row of 6–8 stout anteroventral setae on apical half, short and long setae intermixed, some longer than half width of femur; basal half with row of ca. 9–12 very short, stout setae and 4–6 stout, sub-basal setae, some longer than half width of femur; posteroventral margin with 4–5 stout, widely spaced setae on apical half, less stout apically; basal half with row 9– 13 short, slender setae and 2–3 stout, sub-basal setae some longer than basal width of femur. Fore tibia with biserial row of short, stout setae beneath. Foreleg tarsomeres with white, dense ventral pubescence, length ca. half width of tarsomeres. Mid and hind femora with widely spaced pairs of antero- and posteroventral; widely spaced anterodorsal bristles on apical three-quarters; widely spaced posterodorsal bristles on apical half. Base of hind tibia with pair of divergent dorsal setae (1 posterodorsal and 1 anterodorsal); anteroventral margin lacking bristles; 1 dorsal and 1 ventral preapical bristles present. Tarsus length longer than corresponding fore tibia, shorter than mid and hind tibiae. Empodium pulvilliform, slightly shorter than claw, longer than half length of tarsomere.
Hypandrium quadrate, broad, shorter than epandrium (Figs 8 C, 9G). Phallus not extended beyond apex of clasping cercus, strongly arched at mid-length; apex with anterior, slender, sickle-shaped extension enclosing membrane, posterior similar to ventral process; ejaculatory apodeme nearly straight at anterior third. Surstylus narrow, short, inconspicuous with membranous posteroapical margin; anterior margin with several short teeth-like projections. Clasping cercus broadest in middle, expanded also subapically, strongly tapered to rounded apex; bearing peg-like setae on inner dorsal face.
Female. Similar to male except as follows: basal anteroventral bristles of fore femur absent; posteroventral basal setae less pronounced. Fore tibiae lacking biserial row of erect setae beneath (Figs 18 M, N). Posterior marginal setae of tergum 7 fairly strong, slightly denser than in takagii; sternum 7 weakly emarginated posteriorly, with midventral length 3/4 as long as lateral length, bearing fairly strong hind marginal setae evenly distributed, posterolateral extension of sternum 7 with rather strong isolated seta directed posteroventrally; tergum 8 with posterior marginal setae fairly long and dense dorsally, moderately long laterally; sternum 8 weakly emarginated posteriorly, densely clothed with minute pile-like setulae, sparsely clothed with minute setulae on lateral surface; tergite 10 bearing some 50 apically curved setae evenly distributed except basal marginal area, 2 apical setae only slightly stronger than others on disc; sternite 10 with short setae mixed with 2–3 longer setae 1.5 times as long as sternum (Figs 15 A–C).
Distribution. This species is restricted to the southern island of Kyushu (Fig. 10). Adults are collected from May to October.
Etymology. In reference to the distinct pterostigma, which is weakly differentiated in other Japanese species of Trichoclinocera.
description: Published as part of Saigusa, Toyohei & Sinclair, Bradley J., 2016, Revision of the Japanese species of Trichoclinocera Collin (Diptera: Empididae: Clinocerinae), pp. 201-229 in Zootaxa 4103 (3) on pages 217-218, DOI: 10.11646/zootaxa.4103.3.1, http://zenodo.org/record/255748