Gomphostemma strobilinum Wall.
Creators: name: Bongcheewin, Bhanubong
name: Ingrouille, Martin J.
name: Paton, Alan J.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Magnoliopsida
Lamiales
Lamiaceae
Gomphostemma
Gomphostemma strobilinum

description: <b> 31. Gomphostemma strobilinum <i>Wall. ex 3enth</i></b> . (Bentham in Wallich 1830b: 12); Hooker (1883: 696); Prain (1891: 226);? oan (1936:1032); Mukercee (1920: 203); Murata (1956: 190); Phuong (2000: 98). Type: Burma, Prome [Pyay, 18.8333°, 93.2333°], River Irawaddi, 9 Oct. 1826, <i>Wall. Cat.</i> 2131.1 (K [Herb. Benthamianum, K000826330], lectotype selected here; isolectotype K-W [K001113331]). <i>Gomphostemma strobilinum</i> var. <i>elatius</i> Benth. (Bentham in Wallich 1830b: 12). Type: Burma, Taong?ong [21.9093°, 96.2353°], 12 Nov. 1826, <i>Wall. Cat.</i> 2131.2 (K [Herb. Benthamianum, K000826326], lectotype selected here; isolectotypes BM, K 2 sheets [Herb. Benthamianum, K000826325 &amp; K000826328], K [Herb. Hookerianum], K-W [K001113332]). [Prain (1890) named <i>G. wallichii:</i> one of the specimens he examined was <i>Wall. Cat.</i> 2131.2 (should be at CAL n.v.). To avoid homotypic status, <i>Wall. Cat.</i> 2131.2 is selected as a lectotype of <i>G. strobilinum</i> var. <i>elatius.</i>] <i>Gomphostemma oiride</i> Wall. ex Benth. (Bentham in Wallich 1830b: 12). Type: Burma, Tavoy, 28 Aug. 1825, <i>Gomez</i> s.n. in <i>Wall. Cat.</i> 2133 (K [Herb. Benthamianum, K000826332], lectotype selected here; isolectotype K-W [K001113336]). <i>Gomphostemma acaule</i> Kurz ex Hook.f. (Hooker 1883: 696). Type: Burma, Arracan, Boronga Island, on the Mingoo Hill, evergreen forest, 220 – 360 m [800 – 1200 ft], Oct. 1869, <i>Kurz</i> s.n. (holotype K [K000826323]; isotype K [K000826322]]. <i>Gomphostemma strobilinum</i> var. <i>oiride</i> (Wall. ex Benth.) Hook.f. (Hooker 1883: 696). Type: as for <i>G. oiride</i> Wall. ex Benth. <i>Gomphostemma wallichii</i> Prain (1890: 312). Type: India, Assam, Naga Hills, Phesama [23.5053°, 92.1033°], Sept. 1886, <i>Prain</i> s.n. (K, lectotype selected here; isolectotype CAL n.v.). <i>Gomphostemma strobilinum</i> var. <i>acaule</i> (Kurz ex Hook.f.) Prain (1891: 225). Type: as for <i>G. acaule</i> Kurz ex Hook.f. <i>Gomphostemma dentatum</i> Muschl. (Muschler 1905: 250). Type: Thailand,?oi Sutep, c. 800 m, 29 July 1902, <i>Hosseus</i> 280 (holotype K; isotype BM). <i>Gomphostemma strobilinum</i> var. <i>oariegatum</i> Craib (1922: 96). Type: Thailand, Chiang Mai,?oi Sutep, evergreen cungle, 320 – 900 m, 13 Oct. 1909, <i>Kerr</i> 833 (K, lectotype selected here; isolectotype BM). <i>Gomphostemma lacei</i> Mukercee (1938: 303). Type: Burma, North Shan States, near Masam Falls, 13 Oct. 1911, <i>Lace</i> 6322 (holotype E [E00301958]). <i>Gomphostemma strobilinum</i> subsp. <i>acaule</i> (Kurz ex Hook.f.) Murata (1956: 192). Type: as for <i>G. acaule</i> Kurz ex Hook.f. <i>Gomphostemma strobilinum</i> subsp. <i>wallichii</i> (Prain) Murata (1956: 191). Type: as for <i>Gomphostemma wallichii</i> Prain. Perennial herb or shrub-like up to 0.2 – 2 m tall. <i>Stems</i> robust, erect, obtusely quadrangular with longitudinal grooves, with a mixture of stellate hairs and dendroid hairs. <i>Leaoes</i> petiolate, slightly coriaceous; blades ovate-lanceolate or ovate, 11 – 32 × 5 – 16 cm, apex acute, margin obtusely serrate or serrate, base attenuate, tomentose with a mixture of simple hairs and branched hairs on both sides; petioles 10 – 50 mm long, tomentose. <i>Inflorescence</i> upright, terminal or terminal and axillary, 50 – 200 mm long, with cymes congested or lax at the basal verticil; subtending bracts present, ovate or ovate-lanceolate, 20–60 ×10 –20mm, apex acute, margin serrate, base cordate, obtuse or cuneate; upper bracts green, scarlet or creamy-white,ovate,16 –20 × 9 –18mm, apex acute,margin entire, base cuneate or cordate, upper side with sessile stellate hairs,lower side with branched hairs; bracteoles falcate or narrowly elliptic-ovate,3 – 10 mm. <i>Flowering calyx</i> green or dark red, infundibular, 10 – 20 mm long, 10-ridged, outside with stellate hairs, inside glabrous below and with simple hairs above; tube 6 – 10 mm long; lobes ovate or lanceolate, 2 – 10 mm long, apex acute or acuminate. <i>Fruiting calyx</i> infundibular, 10 – 20 mm long, 10- ridged, with stellate hairs on both sides; tube 3 – 10mm long; lobes triangular-ovate, 3 – 10 mm long, margin slightly revolute. <i>Corolla</i> bright yellow, creamy-white or scarlet, 20 – 33 mm long, throat inflated, abruptly dilated towards the throat; tube incurved, 13 – 23 mm long, exserted from the calyx,outsidewithsessilestellatehairs,insidewithanannulus of simple hairs; posterior lip ovate, 5 – 11 × 2 – 9 mm, apex slightly emarginate or entire, margin undulate; anterior lip spreading,8–10mm long, undulate;median lobe obovate,2 – 3 mm long, apex emarginate, margin entire, or sometimes revolute in cross section; lateral lobes folded downward. <i>Staminalfilaments</i> fleshy,glabrous. <i>Style</i> slender,slightlyfleshy at the apical part with apex unequally lobed. <i>Nutlets</i> 3 – 2, elliptic-ovate, 2 – 3 mm long, glabrous, wrinkled. Fig. 2. <b>DISTRIBUTION.</b> India (Manipur, Mizoram, Nagaland), China, Burma, Laos, Thailand. Map 8. <b>SPECIMENS EXAMINED.</b> INDIA. Nagaland. Naga Hills, Kohima, 1200 m [2000 ft], 10 – 23 Sept. 1930, <i>Chand</i> 3628 (E, L); Naga Hills, Kohima, 1210 m [2500 ft], 31 Aug. 1933, <i>3or</i> 6233 (K); Naga Hills, Kasom, 1200 m [2000 ft], 22 Aug. 1928, <i>Mukerjee</i> 3332A (CAL); Naga Hills, Phesama [23.5053°, 92.1033°], Sept. 1886, <i>Prain</i> s.n. (lectotype of <i>G. wallichii</i>, K); Naga Hills, Piffima [23.53°, 93.9665°], 900 m [3000 ft], 19 Oct. 1883, <i>Clarke</i> 20961 (K); Assam, Yarobami, 1300 m [3000 ft], 2 Sept. 1922, <i>3or</i> 13928 (BKF); Manipur. Kohima Road, 900 m [3000 ft], Sept. 1891, <i>Collett</i> s.n. (K); Muneypoor, Karong, 1123 m [3530 ft], 12 Nov. 1883, <i>Clarke</i> 22012 (K); Mizoram. Lushai Hills, Kolasib [22.2311°, 92.6536°], 600 m [2000 ft], 22 Jan. 1933, <i>Koelz</i> 32333 (E). <b>CHINA.</b> Yunnan. Menglian [22.6°, 98.8°], Lalei Huigu Village, 15 Oct. 1981, <i>Tao &amp; Li</i> 39252 (KUN). <b>MYANMAR.</b> Kachin. Bhamo [22.2665°, 95.2333°], 103 m [330 ft], 19 Nov. 1911, <i>Lace</i> 3351 (E 2 sheets, K); Sagaing. Ruby Mines?istr., near Mogok [22.9165°, 96.3°], 1330 m [2300 ft], Oct. 1912, <i>Lace</i> s.n. (E); Upper Burma, Mingin [22.8833°, 92.3°], 1890, <i>Prazer</i> s.n. (K); Shan. Upper Burma, Southern Shan State, Taungyi [20.5833°, 95.0333°], 1893, <i>Khalil</i> s.n. (G 2 sheets, P2 sheets); Upper Burma, Fort Stedman [20.3833°, 96.93°], Nov. 1892, <i>Huk</i> s.n. (K, P2 sheets); South Shan States, Keng Tung [21.3°, 99.63°], 1200 m [2000 ft], Sept. 1909, <i>MacGregor</i> 532 (E); North Shan States, near Masam Falls, 13 Oct. 1911, <i>Lace</i> 6322 (holotype E); Shan Hills, Terai, 600 m [2000 ft], Nov. 1888, <i>Collett</i> 961 (K); SE Shan State, Meongpaon [21.3°, 99.63°], 1330 m [2300 ft], 3?ec. 1890, <i>Watt</i> 10526 (E); Southern Shan, Loilem [20.9230°, 95.3639°], 1330 m [2300 ft], 15 Sept. 1911, <i>Robertson</i> 231 (K 2 sheets); Upper Burma, Southern Shan State, Laikaw, 1892, <i>Khalil</i> s.n. (L); Upper Burma, Kyali [23.5333°, 92.2333°], 1890, <i>Prazer</i> 219 (AB?, BM, K); Upper Burma, Southern Shan State, Tameinkhan [Thamakan or Thamakhan, 20.6833°, 96.5°], 1893, <i>Khalil</i> s.n. (AB?, TC?); Chin. W Central Burma, Mindat [21.3665°, 93.9833°], 1200 – 1300 m [2000 – 3000 ft], 30 July 1936, <i>Kingdon-Ward</i> 22323 (BM); Mandalay. Maymyo, Oct. 1888, <i>Khan</i> 212 (G); ibid., 1030 m [3300 ft], 30 Aug. 1908, <i>Lace</i> 2223 (E, K 2 sheets, TC?); ibid., 600 m [2000 ft], Jan. 1908, <i>Meebold</i> 8102 (K); Taong?ong, 12 Nov. 1826, <i>Wall. Cat.</i> 2131.2 (lectotype of <i>Gomphostemma strobilinum</i> var. <i>elatius</i>, K [Herb. Benthamianum]; isolectotypes BM, K 2 sheets [Herb. Benthamianum], K [Herb. Hookerianum], K-W); Thoudaung, Oct. 1932, <i>Dickason</i> 3216 (GH); Rakhaing. Arracan, Boronga Island [19.9665°, 93.1°], Mingoo Hill, 220 – 360 m [800 – 1200 ft], Oct. 1869, <i>Kurz</i> s.n. (holotype K; isotype K); Bago. Tharrawaddy [15.63°, 93.8°], 120 m [200 ft], Nov. 1901, <i>Sohool</i> 8 (G); Prome, R. Irawaddi, 9 Oct. 1826, <i>Wall. Cat.</i> 2131.1 (lectotype of <i>G. strobilinum</i>, K [Herb. Benthamianum]; isolectotype K-W); Yangon. Rangoon [16.53°, 96.23°], 11 Jan. 1832, <i>McClelland</i> s.n. (K [Herb. Hookerianum]); Mon. Moulmein [16.2912°, 95.6236°], 1839, <i>Parish</i> 58 (K [Herb. Hookerianum]); Tanintharyi. Tavoy, 28 Aug. 1825, <i>Wall. Cat.</i> 2133 (holotype of <i>G. oiride</i>, K [Herb. Benthamianum]; isotype K-W); Tenasserim, Tavoy, 2.3 miles E of Paungdaw Power Station, 12°N 98°30&amp;E, 390 m [1300 ft], 22 Aug. 1961, <i>Keenan et al</i>. 1036 (E); Tenasserim, Tavoy, NW of Paungdaw Village, 300 m [1000 ft], 23 Aug. 1961, <i>Keenan et al.</i> 1229 (E, K); Tenasserim, Tavoy, Hills W of Paungdaw Power Station, 660 m [2200 ft], Aug. 1961, <i>Keenan et al.</i> 898 (E); <b>UNLOCALISED</b>. Meong, R. Irawaddi, 13 Sept. 1826, <i>Wall. Cat.</i> 2131.1 (lectoparatypes of <i>G. strobilinum</i>, G 2 sheets, K [Herb. Benthamianum], K-W). <b>LAOS. UNLOCALISED</b>. <i>Massie</i> s.n. (P). <b>THAILAND.</b> Mae Hong Son. Huai Hee,?oi Pui [19.2165°, 98.0665°], 1300 m, 22 Oct. 2000, <i>Suksathan</i> 2823 (QBG); Khun Yuam, 18°13&amp;N 98°E, 600 – 500 m, 3 Sept. 1952, <i>Larsen et al.</i> 32166 (AAU, BKF, K, L); Muang, Huai Bu Ling, 1123 m, 28 Oct. 2005, <i>Maxwell</i> 05-665 (CMUB, QBG); Pai, Ban Pee Lu, 22 Oct. 1996, <i>Na Nakorn et al.</i> 5569 (QBG 2 sheets); Pai, Ban Watchan, 1100 m, 3?ec. 2005, <i>Wongprasert</i> 512-2 (BKF); Pai, Muang Sroi Waterfall, 800 m, 15 Jan. 1983, <i>Koyama et al.</i> T-32680 (BKF); Pang Ma Pha, Kew Lom [19.2325°, 98.3031°], 1300 m, 3 Sept. 1999, <i>Suksathan</i> 1530 (QBG); Pang Ma Pha, Tam Lawt,?oi Pah Baw, 523 m, 12 Nov. 2002, <i>Palee</i> 585 (CMUB); Pang Ma Pha, Tam Pla, 12?ec. 2005, <i>Tanaka et al.</i> 8219 (QBG); Chiang Mai.?oi Chiang?ao, 19°23&amp;N 98°33&amp;E, 330 – 1100 m, 3?ec. 1963, <i>Hennipman</i> 3183 (BKF, C, K); ibid., 1300 m, 2?ec. 1961, <i>Smitinand &amp; Anderson</i> 5260 (BKF); ibid., 1900 – 2200 m, 3?ec. 1961, <i>Smitinand &amp; Anderson</i> 5339 (BKF);?oi Chiang?ao, Khun Me Terhy, 1130 m, 31 Oct. 1933, <i>Garrett</i> 1262 (K, L);?oi Chiang?ao, Ridge above Ban Tam, steep slope above caves [19.3922°, 98.9265°], 300 – 600 m, 3 Jan. 1921, <i>Garrett</i> 1219 (E, K, L 2 sheets, P 2 sheets, TC?); Chiang?ao, Ban Bing Kong, 300 m, 29 Oct. 1985, <i>Maxwell</i> 85-1303 (L);?oi Chiang?ao Wildlife Sanctuary, 1500 m, Nov. 1993, <i>Morsi</i> 233 (CMUB);?oi Chiang?ao, SE foothills at Ban Yang Toong Brong Forest Station [19.3833°, 98.9°], 253 – 353 m, 11 Nov. 1989, <i>Maxwell</i> 89- 1389 (L); Chiang?ao, Pa Sam Sob, 6 Nov. 1963, <i>Chantanamuck</i> 601 (BK); Chom Thong,?oi Hua Sua [18.2959°, 98.3363°], along the trail extending from the W part to the top in?oi Inthanon National Park, 1600 m, 15?ec. 1998, <i>Konta et al.</i> 2636 (BKF);?oi Angka [?oi Inthanon, 18.3833°, 98.2833°], 1800 m, 15 July 1922, <i>Kerr</i> 6333 (BK, BM, K 2 sheets); ibid., 330 m, 20 July 1988, <i>Phengklai et al.</i> 6630 (BKF);?oi Inthanon National Park, around Wachirathan Waterfall [18.3218°, 98.3992°], 600 – 500 m, 6?ec. 1982, <i>Koyama et al.</i> T-39922 (BKF);?oi Inthanon National Park, near Ban Sop Hat, 880 m, 6?ec. 1982, <i>Mitsuta et al.</i> T-26280 (BKF);?oi Kar [?oi Inthanon], range NE of the Pah Ngeam, 1110 m, 1 Oct. 1910, <i>Garrett</i> 66 (lectoparatypes of <i>G. strobilinum</i> var. <i>oariegatum</i>, BM, K, L);?oi Pha Mawn and spur connecting with?oi Angka, 1320 – 1610 m, 3 Sept. 1925, <i>Garrett</i> 233 (AAU, K 2 sheets); en route from Mae Klang Waterfall to Ban Yang, along Nam Mae Klang, on approach to?oi Inthanon, 330 – 600 m, 16?ec. 1963, <i>Tagawa et al.</i> T-2293 (BKF, L); Changkien [18.8238°, 98.8886°], 1200 m, 26 Oct. 1953, <i>Sadakorn</i> 620 (BK);?oi Pui [15.3665°, 98.5165°], 25 Sept. 1995, <i>3GO staff</i> 9636 (QBG); ibid., 1230 – 1630 m, 16?ec. 1982, <i>Koyama &amp; Nagamasu</i> T-20131 (AAU, BKF, L);?oi Sutep-Pui National Park, N part,?oi Mawn Luang, above Nong Rai Village, 1253 m, 22?ec. 1996, <i>Maxwell</i> 96-1682 (BKF, CMUB);?oi Sootep, 220 m [1200 ft], 12 Nov. 1910, <i>Kerr</i> 1321 (BM); ibid., 1030 – 1300 m [3300 – 3000 ft], 2 Sept. 1910, <i>Kerr</i> 1380 (BM, K, TC?); ibid., 320 – 900 m [1800 – 3000 ft], 13 Oct. 1909, <i>Kerr</i> 833 (lectotype of <i>G. strobilinum</i> var. <i>oariegatum</i>, K; isolectotype BM); ibid., 600 – 900 m [2000 – 3000 ft], 6 Oct. 1912, <i>Kerr</i> 833A (lectoparatypes of <i>G. strobilinum</i> var. <i>oariegatum</i>, BM, K); ibid., c. 800 m, 29 July 1902, <i>Hosseus</i> 280 (holotype K; isotype BM);?oi Sutep, below Puping Palace [18.8°, 98.9°], 1300 m, 31 Oct. 1985, <i>Maxwell</i> 85-1322 (L);?oi Saket [18.8522°, 99.1336°], 31 Oct. 1930, <i>Put</i> 3252 (BK, BM 2 sheets); Fang,?oi Ang Kang [19.8°, 98.9833°], 9 April 1953, <i>Sutheesorn</i> 3138 sht. 1 (BK); Hod, Baw Salee Subdistr., km 33, Hod-Mae Sariang Road, Highway no. 108, Ban Mae Sanam Mai, Mai Muang Nao, 930 m, 28 Sept. 2001, <i>Sankamethawee</i> 285 (CMUB, L); Jawm Tong, Mae Soi Ridge, Mae Soi Subdistr., W of Ban Bah Gluay [18.2568°, 98.6356°], 1623 m, 28 Sept. 1992, <i>Maxwell</i> 92- 390 (CMUB, L, P); Mae Jam, Ban Wat Chan Subdistr., Huay Bong Village [19.0623°, 98.2963°], 1123 m, 6?ec. 1998, <i>Maxwell</i> 98-1225 (BKF, CMUB, L 2 sheets); Mae Rim, Huai Mae Sa Noi [18.9122°, 98.9395°], 300 m, 21 Nov. 1996, <i>Na Nakorn</i> 5825 (QBG); Mae Rim, Mon Long [18.9192°, 98.8333°], 1320 m, 20 Nov. 2001, <i>Glamwaewwong</i> 33 (QBG); Mae Rim, Pong Yaeng [18.8831°, 98.8169°], 1008 m, 11 Jan. 2008, <i>Jatupol</i> 08- 156 (CMUB, QBG, XTBG); Mae Sa, N of Chiang Mai, Rock Tower Mt, 18°33&amp;N 98°30&amp;E, 1300 – 1200 m, 18 Sept. 1993, <i>Larsen et al.</i> 26633 (AAU); Mae Song Sob [18.5903°, 98.9815°], 20 May 1993, <i>3GO staff</i> 3313 (QBG); Chiang Mai, Mae Tang,?oi Sahng Liang, S side, upper Gu Gahp Valley, above Pah?ang Village, 1253 m, 21 Sept. 1995, <i>Maxwell</i> 95-1028 (BKF, CMUB); Mae Tang,?oi Tung Jaw, 1200 m, 12?ec. 1981, <i>Thongisan</i> 36 (CMUB); Mae Tang, Pa Pae, 1000 m, 9 Nov. 1980, <i>Paisooksantioatana &amp; Sangkhachand</i> 358-80 (BK 2 sheets, CMUB); Mae Tang, Pa Pae, Pong?uat hot springs [19.2313°, 98.5009°], 830 m, 19 Nov. 1992, <i>Maxwell</i> 92- 521 (CMUB, L); Mok Fah Waterfall, 20 km NE of Chiang Mai, along the road to Pai, Road no. 1093, 19°3&amp;N 98°13&amp;E, 300 m, 22 Nov. 1993, <i>Larsen et al.</i> 22588 (AAU); Muang Krong, Koon Krong [19.3123°, 98.802°], 21 Nov. 1963, <i>Chantanamuck</i> 502 (BK); Wang Tao, 800 m, 13 Feb. 1938, <i>Sørensen et al.</i> 1032 (C, KUN); Om Koi, Bo Luang Tableland, Ban Om Koi to Mae Tuen Watershed improvement station, 1320 m, 15 Oct. 1959, <i>Shimizu et al.</i> T-19136 (BKF); Omkoi, Mae Rah Ah Watershed Station, 1200 m, 13 Aug. 1999, <i>Petrmitr</i> 261 (CMUB, L 2 sheets); Pah Hom Pok [20.1165°, 99.2°], 1200 m, 22 Feb. 1938, <i>Sørensen et al.</i> 1360 (BKF, C, K, KUN); Huai Sai, Huai Nam?ang National Park. [19.3023°, 98.3983°], 1285 m, 20 Jan. 2002, <i>Chayamarit et al.</i> 3215 (BKF 2 sheets); Samoeng, Sanoengtai, Ban Pa Kar [18.9665°, 98.9665°], 1222 m, 8 July 2008, <i>Jatupol</i> 08-339 (QBG 2 sheets); Chiang Rai.?oi Luang National Park [19.2°, 99.5165°], 1153 m, 5 Sept. 1995, <i>Maxwell</i> 95-958 (CMUB); Khun Jae [19.0559°, 99.3958°], 923 m, <i>Maxwell</i> 95-1368 (CMUB, L); Khun Korn Waterfall [19.8821°, 99.6169°], 530 m, 12 Oct. 1995, <i>KK</i> 236 (BCU); Mae Fah Luang, W of?oi Pah Giah, 1200 m, 22 Oct. 2006, <i>Palee</i> 1068 (CMUB); Pahn,?oi Luang National Park, W side, summit area of?oi Mauk [20.1225°, 100.2961°], Sahn Salee, 1230 m, 6 Oct. 1998, <i>Palee</i> 220 (BKF, CMUB, L); Wiang Pa Pao, Mae Chadee [19.1833°, 99.3165°], 1200 m, 29 Sept. 1993, <i>Vial-Debas</i> 10 (CMUB); Thoeng, Ban Mai Ya, 9 Feb. 1950, <i>Sutheesorn</i> 1622 (BK); Phayao.?oi Luang National Park, E side, 610 m, 21?ec. 1995, <i>Petrmitr</i> 152 (CMUB); ibid., 1200 – 1223 m, 28?ec. 1995, <i>Petrmitr</i> 225 (CMUB); Nan. Chalerm Pra Kiat, Wiang Peian Waterfall, 230 m, 13 Nov. 2001, <i>Srisanga</i> 2288 (QBG); Lamphun. higher elevation of?oi Khun Tan, 1200 – 1353 m, 3 Sept. 1965, <i>Tagawa et al</i>. T-9288 (AAU, BKF, K); Khun Tan [18.2508°, 99.2825°], 1000 m, 31 Oct. 1955, <i>Chermsirioathana</i> 2262 (BK); Mae Ping National Park, Ko Thung Waterfall, 230 – 300 m, 18 Nov. 1995, <i>Pongamornkul</i> 11 (QBG); Me Kow, 380 m, 2 Nov. 1923, <i>Winit</i> 1323 (BK, BKF 2 sheets, BM, L); Lampang.?oi Khun Tan [18.2508°, 99.2825°], 800 – 830 m, 9 Jan. 1992, <i>Koyama</i> T-62035 (BKF); ibid., 1100 – 1300 m, 30 Oct. 1955, <i>Na Songkhla et al.</i> 329 (BCU 2 sheets); Me Saloi, 200 m, 28 Oct. 1923, <i>Winit</i> 1290 (BKF, BM); Muang Pan, Chae Sorn National Park [18.8362°, 99.2692°], 553 m, 22 Oct. 1993, <i>Maxwell</i> 93-996 (CMUB); Wang Nua,?oi Luang National Park, W side of and below?oi Nawk, 1130 m, 5 Nov. 1998, <i>Petrmitr</i> 323 (CMUB); Phrae. Song, Mae Yom National Park [18.63°, 100.1625°], 200 m, 12?ec. 1993, <i>Maxwell</i> 93-1302 (CMUB); Tak. Wang?cao [Wang Chao, 16.5022°, 99.2228°], 100 m, 10 Oct. 1902, <i>Hosseus</i> 62 (BM, C, E, K, M, P 2 sheets); Raheng [Tak],?oi Tung Cha [15.2290°, 98.9359°], 300 m, 18 Nov. 1920, <i>Kerr</i> 2393 (BK, BM 2 sheets); Bhumibol?am [15.23°, 98.9665°], Feb. 1960, <i>Anonymous</i> s.n. (BK);?oi Mussor, 1000 m, 9?ec. 1961, <i>Smitinand</i> 5069 (BKF, K); Mae Sot?istr., Wat Tham Manorha, 16°26&amp;28”N 98°39&amp;12”E, 300 m, 11 Sept. 2009, <i>Middleton &amp; Triboun</i> 2831 (E); Taksinmaharac National Park [16.5808°, 98.9286°], 10 Oct. 1992, <i>Herb Trip</i> 282 (BCU); Tha Song Yang, Mae Hla, Mae Sot-Mae Sariang, road no. 103, km 36, 200 m, 22 Nov. 2003, <i>Pooma</i> 3583 (BKF, E); Umphang,?oi Hua Mot, [13.9613°, 98.8353°], 960 m, 3 Nov. 1998, <i>Suddee &amp; Puujaa</i> 1115 (K, TC?); Sukhothai. Kirimat, Ramkamhaeng National Park [16.8509°, 99.6852°], 300 m, 1?ec. 1985, <i>Paisooksantioatana &amp; Sangkhachand</i> 2183-85 (BK); Muang Gow, near PraTah?ue Nahm?oke So Gah Cha Nah Temple, 2 Nov. 1951, <i>Maxwell</i> 51-669 (AAU, BK, L); Phitsanulok. Kaeng Sopa [16.8832°, 100.6365°], 12?ec. 1966, <i>Sangkhachand</i> 391 (BK); Thung Salaeng Luang National Park, about 20 km E of Phitsanulok, 16°29&amp;N 100°36 – 38&amp;E, 230 – 280 m, 21 Oct. 1982, <i>Murata et al.</i> T-38331 (BKF); Kamphaeng Phet. Kao Son, Ban Nam Tok, 200 m, 22 Nov. 1955, <i>Phengklai et al.</i> 3899 (BKF); Klong Lan Waterfall, 200 – 200 m, 23 Oct. 2001, <i>Watthana</i> 1308 (QBG); Mae Wong National Park [16.1315°, 99.2821°], 300 m, 13 Oct. 1998, <i>oan de 3ult</i> 139 (CMUB); Nakhon Sawan. Lat Yao, Kao Mae-kar-see, 13 Jan. 1953, <i>Sutheesorn</i> 3138 sht. 2 (BK); Phetchabun. Nam Nao National Park, trail to N of Park Headquarters, 800 m, 8?ec. 1998, <i>Suddee et al.</i> 913 (K, TC?); Loei. Na Haew National Park, Tad Hueng Fall [15.
description: Published as part of <i>Bongcheewin, Bhanubong, Ingrouille, Martin J. &amp; Paton, Alan J., 2022, A revision of Gomphostemma (Lamiaceae), pp. 27-92 in Kew Bulletin 77 (1)</i> on pages 77-81, DOI: 10.1007/s12225-021-09991-y, http://zenodo.org/record/7615581
description: {"references": ["Hooker, J.?. (1883). Labiatae, Flora of 3 ritish India 2: 602 - 503. Reeve &amp; Co., London.", "____ (1891). An account of the genus Gomphostemma Wall. Ann. Roy. 3 ot. Gard., Calcutta 3: 225 - 251.", "? oan, T. (1936). Labiatae. In: H. Lecomte (ed.), Flore Generale de L' Indo-Chine 2: 913 - 1026. Masson, Paris.", "____ (1920). A revision of the Labiatae of the Indian Empire. Rec. 3 ot. Suro. India 12: 1 - 228.", "____ (1956). Contributions to the flora of Southeast Asia: vi. additions and corrections to the knowledge of Labiatae in Thailand (1). S. E. Asian Stud. 12: 155 - 193.", "Phuong, V. X. (2000). Lamiaceae (Labiatae). In: N. T. Ban, T.?. Ly, T. C. Khanh, P. K. Loc, N. N. Thin, N. V. Tien &amp; N. K. Khoi (eds), Flora of Vietnam 2: 1 - 258. Science &amp; Technics Publishing House, Hanoi.", "Prain,?. (1890). Some additional species of Labiatae. J. Asiat. Soc. 3 engal, Pt. 2, Nat. Hist. 39: 312 - 316.", "Muschler, R. C. (1905). LXVI. Labiatae Siamenses novae. Repert. Spec. Noo. Regni Veg. 2: 268 - 251.", "____ (1922). Contributions to the Flora of Siam. Additamentum XIV. 3 ull. Misc. Inform., Keae 1922: 81 - 98.", "Mukercee, S. K. (1938). A decade of new Labiatae from India, Burma and Tibet. Notes Roy. 3 ot. Gard. Edinburgh 19: 303 - 308.", "Hance, H. F. (1882). A new Chinese Gomphostemma. J. 3 ot. 22: 231 - 232.", "Elmer, A.?. E. (1908). A century of new plants. Leafl. Philipp. 3 ot. 1: 339.", "____ (1923 b). Botanical Excursion in Sumatra. J. Malayan 3 ranch Roy. Asiat. Soc. 1: 26 - 113."]}