Asymphyloptera tama Ramos-Pastrana & Córdoba-Suarez & Sinclair 2023, sp. nov.
Creators: name: Ramos-Pastrana, Yardany
name: Córdoba-Suarez, Eric
name: Sinclair, Bradley J.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Empididae
Asymphyloptera
Asymphyloptera tama

description: Asymphyloptera tama sp. nov.
(Figs 8–14, 15–16)
Diagnosis. Postpedicel with white setae, medial setae shorter than lateral and dorsal setae; 3 dorsocentral setae, anterior dorsocentral setae offset; phallus slightly arched medially, apex pointed, hook-shaped, with membrane expanded posteriorly.
Description. Holotype male. Head (Fig. 8). Dark brown. Postpedicel ovoid with white setae scattered, medial setae shorter than lateral and dorsal setae; slender apical extension arising subapically (Fig. 9). Ocellar setae stout, shorter than length of postpedicel extension. Labrum equal in length to clypeus; palpus brown, subequal in length to labrum, tapered to slender apex (Fig. 8).
Pleura dark brown (Fig. 8). Chaetotaxy stout; 1 postpronotal seta short, very slender; 2 notopleural setae, lower seta shorter and more slender; 3 dorsocentral setae, anterior dorsocentral seta offset; 2 apical scutellar setae, subequal in length to dorsocentral setae, lacking lateral scutellar setae (Fig. 10). Legs (Fig. 8). Coxae, trochanters and base of femora yellowish, remaining segments brown; all femora with anteroventral and posteroventral rows of pale and slender setae, length about ½ width of femur; hind femur with fewer and shorter setae; all tibiae with row of setae ventrally, setae on fore tibia stronger and shorter. Wing length 1.7 mm (Fig. 11); crossvein h distinct; base of wing with slender setulae along posterior margin. Halter brown.
Male terminalia (Figs 12–13). Cercus with anterior lobe slightly produced; posterior lobe long, slender. Hypandrium prolonged as broad, paired postgonites; apical margin of postgonite acute, not extended anteriorly. Epandrium narrow, with elongate setae on apical half. Surstylus shorter than cercus, slender, arched, tapered to pointed apex; extending beyond phallus. Phallus slightly arched medially, apex pointed, hook-shaped, with membrane expanded posteriorly. Ejaculatory apodeme narrow, not expanded.
FEMALE. Similar to male. Terminalia not dissected (Fig. 14).
Type material. HOLOTYPE ♁: COLOMBIA, Huila, Garzón, Vda.[Vereda] Las Mercedes, Parque Natural Regional Cerro Páramo Miraflores, 02º08′06″N / 75º30′04″W, 2074 m [eters], 24. Nov [XI]–8. Dic [XII].2022, Y. Ramos-Pastrana Leg. (1 ♁ LEUA-59560) (photographed specimen) / Captura con trampa Malaise en bh-MB [Bosque húmedo montano bajo] (Vegetación secundaria),. PARATYPES: same data as Holotype (1 ♁ LEUA-59561); (1 ♁ LEUA-59562); (1 ♁ LEUA-59563); (1 ♁ LEUA-59564); (1 ♁ LEUA-59565); (1 ♁ LEUA-59566); (1 ♁ LEUA-59567); (1 ♀ LEUA-59568) (photographed specimen); (1 ♀ LEUA-59569); (1 ♀ LEUA-59570); (1 ♀ LEUA-59571); (1 ♀ LEUA-59572); (1 ♀ LEUA-59573); (1 ♀ LEUA-59574); (1 ♀ LEUA-59575); (1 ♀ LEUA-59576); (1 ♀ LEUA-59577); (1 ♀ LEUA-59578); (1 ♀ LEUA-59579); (1 ♀ LEUA-59580); (1 ♀ LEUA-59581); (1 ♀ LEUA-59582); (1 ♀ LEUA-59583); (1 ♀ LEUA-59584); (1 ♀ LEUA-59585); (1 ♀ LEUA-59586); (1 ♀ LEUA-59587); (1 ♀ LEUA-59588); (1 ♀ LEUA-59589); (1 ♀ LEUA-59590).
Etymology. The specific epithet refers to the indigenous people “Tama” that inhabit the type locality and is a noun in apposition.
Geographical distribution. Colombia (Huila, Garzón) (Fig. 15).
Habitat. See under A. miraflorensis sp. nov.
Taxonomic notes. Asymphyloptera tama sp. nov. runs to A. cajanuma in couplet 5 of the key presented by Sinclair (2015). It differs from A. cajanuma by having a sclerotized pointed hook-shaped apex of the phallus (Figs 12–13) [versus sclerotized portion of apex of phallus slender with apex truncated in A. cajanuma (Sinclair 2015, fig. 8)]. Additionally, based on male specimen and due to the shape of the cercus, epandrium and surstylus A. tama sp. nov. is also related to A. dominica (Sinclair 2015, fig. 13). It differs by the slightly arched phallus, with apex of phallus lacking jagged margin (Figs 12–13) [versus phallus strongly arched posteriorly, apex of phallus with jagged lateral margin in A. dominica (Sinclair 2015, fig. 13)].
Key to males of New World species of Asymphyloptera [adapted from Sinclair (2015)]
(male unknown for A. discrepans)
1 Hypandrium strongly produced posteriorly, with expanded, flattened ejaculatory apodeme [see Sinclair (2015, fig. 9)].................................................................................... A. chilensis Sinclair, 2015
– Hypandrium rounded, not strongly produced posteriorly, with slender, rod-like ejaculatory apodeme [see Sinclair (2015, figs 8, 11, 13–16)].......................................................................................... 2
2 Surstylus digitiform, with rounded tip; phallus with apex arched anteriorly [see Sinclair (2015, fig. 11)].............................................................................................. A. chiricahua Sinclair, 2015
- Surstylus slender and tapered toward narrow apex; phallus either gradually arched posteriorly or apex sharply bent anteriorly [see Sinclair (2015, figs 8, 13–16)]....................................................................... 3
3 Phallus straight, not arched; apex of phallus distinctly bent anteriorly [see Sinclair (2015, figs 14, 16)]................. 4
– Phallus slightly to strongly arched posteriorly; apex arched posteriorly or expanded laterally [see Sinclair (2015, figs 8, 13, 15)]............................................................................................... 5
4 Apex of phallus strongly bent anteriorly; apex of postgonite sharply pointed [see Sinclair (2015, fig. 16)].............................................................................................. A. mexicana Sinclair, 2015
- Apex of phallus expanded anteriorly; apex of postgonite rounded to slightly pointed [see Sinclair (2015, fig. 14)]......................................................................................... A. havasu Sinclair, 2015
5 Phallus slightly arched................................................................................. 6
- Phallus strongly arched posteriorly....................................................................... 7
6 Sclerotized apex of phallus slender with apex truncated [see Sinclair (2015, fig. 8)].......... A. cajanuma Sinclair, 2015
- Sclerotized apex of phallus pointed, hook-shaped (Figs 12–13).................................... A. tama sp. nov.
7 Apex of phallus with jagged lateral margin [see Sinclair (2015, fig. 13)]..................... A. dominica Sinclair, 2015
- Apex of phallus lacking jagged margin [Figs 5–6, and see Sinclair (2015, fig. 15)]................................. 8
8 Cercus with anterior lobe slightly produced and posterior lobe long, slender and apex rounded [see Sinclair (2015, fig. 15)]; phallus with apex posterodorsally-directed [see Sinclair (2015, fig. 15)]......................... A. lutea Sinclair, 2015
- Cercus with anterior lobe not produced and narrow posterior lobe (Figs 5–6); phallus with apex ventrally-directed........................................................................................ A. miraflorensis sp. nov.
description: Published as part of Ramos-Pastrana, Yardany, Córdoba-Suarez, Eric & Sinclair, Bradley J., 2023, Two new species of Asymphyloptera Collin (Diptera: Empididae: Clinocerinae) from Colombia, with an updated key to males of New World species, pp. 437-444 in Zootaxa 5374 (3) on pages 440-443, DOI: 10.11646/zootaxa.5374.3.8, http://zenodo.org/record/10150959