Chileanthicus dentatus Kejval 2009, comb. nov.
Creators: name: Kejval, Zbyněk
Publisher: name: Zenodo
Year: 2009
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Anthicidae
Chileanthicus
Chileanthicus dentatus

description: Chileanthicus dentatus (Dajoz, 1980) comb. nov.   
(Figs 58–62, 222)  
 Pseudoleptaleus dentatus Dajoz, 1980: 192, fig. 1 (see Remarks).   
 Type locality. W-Madagascar, environs of Morondava, Belo and Tsiribihina villages.   
 Redescription (male, FMNH). Head largely dark brown to brown black, with paler, rufous brown neck and surroundings of antennal pits; pronotum brown black, with paler, rufous brown collar and basal margin; elytra brown black, with rufous brown suture, lateral margins and two, paired, rufous to rufous brown bands (set off by silvery setation), the anterior band wide, oblique and somewhat sinuous, the posterior band rather straight and transverse; legs rufous brown, distal swollen portion of metafemora darkened, brown black; antennae and palpi rufous brown.   
Head 1.2 times as long as wide, widely rounded posteriorly; tempora subparallel, posterior temporal angles rounded but distinct. Eyes small to medium-sized, moderately convex. Dorsal surface matte, very densely, finely punctured (Fig. 58); punctation simple, evenly developed, including median line. Setation pale to slightly brownish, evenly short and fine, appressed, with inconspicuous, sparsely scattered, erect setae. Antennae rather strong, distinctly enlarged in terminal third; antennomere III 2.1 times as long as wide, distinctly longer than IV; antennomere X 1.3 times, XI 1.9 times as long as wide.  
Pronotum robust, 1.3 times as long as wide, moderately wider than head; widely rounded anteriorly; pronotal disc moderately convex, with indication of shallow median longitudinal impression, its lateral margins forming rounded edge, lateral outlines nearly straightly narrowing posteriad in dorsal view. Surface matte, very densely, finely punctured, including lateral sides; dorsal punctation and setation as on head.  
Elytra 1.6 times as long as wide, narrowed and conjointly rounded apically, rather convex, but somewhat flattened mesally along slightly raised sutural margins. Surface slightly glossy, densely and finely punctured; basal punctation moderately sparser than on head (Fig. 58), punctures at places arranged in transverse/oblique rows, and thus surface may appear finely, transversely corrugated (mainly in basal third near suture, best visible in dorso-caudal view). Setation similar to that on head, mostly pale with slight brownish tinge, rather silvery on pale elytral bands; short erect setae somewhat more numerous than on head.  
Metafemora conspicuously swollen/clavate, unidentate, subapical process prominent, facing inner side of tibia (Fig. 59); metatibiae distinctly arcuate, with apical margin moderately produced on inner side. Setation rather uniformly short, fine and appressed.  
Abdominal sternum VII and tergum VII simple, with posterior margin evenly rounded apically. Sternite VIII (Figs 60, 61); paired prongs simple, sharply curved ventrad, their distal narrowed portion, except fine setation, with about six, short and thick, peg-like setae ventro-laterally. Tergite VIII with posterior margin somewhat unevenly rounded.  
Aedeagus (Fig. 62); apical portion of tegmen short, 0.5 times as long as basal-piece, simple, moderately bent in lateral view, nearly evenly narrowing towards bluntly pointed apex; median lobe of aedeagus membranous, bilobed and with peculiar, lamelliform structure apically.  
Female. Externally identical to male, with nearly simple metafemora (less distinct apical protrusion).  
Body length (♂ ♀). 3.0– 5.1 mm.  
 Variation. Rather variable in size, colouration and body proportions. In dark coloured specimens, head and pronotum unicoloured brown black, paler marking elytra less distinct, and all femora darkened, brown black distally. In pale coloured specimens, head and pronotum pale rufous brown, paler marking of elytra conspicuous (bands merging with extensively paler suture and lateral margins), and legs rufous brown. In some specimens, especially smaller ones, head less widely rounded posteriorly and pronotum as wide as to slightly narrower than head.   
 Type material. Holotype and a paratype (MNHN, not examined, see Remarks).   
 Specimens examined. 37 ♂♂, 39 ♀♀, Madagascar, 48 km ENE Morondava, 20°04‘S, 44°39‘E, 30 m, 4.- 10.i.1991, tropical dry forest, litter, D. M. Olson leg. (FMNH); 1 ♂, 5 ♀♀, Madagascar, Morondava, Kirindi forest, ii.-iii.1990, M. Butterweck leg. (SMNS, ZKDC).   
 Differential diagnosis. C. dentatus nearly shares the same robust body form of C. cursor sp. nov. It can be easily distinguished by the matte, very densely and finely punctured body surface (cf. Figs 54, 58), including the lateral sides of the pronotum, by the generally shorter, appressed body setation, and by numerous details of the male characters, e.g. presence of peg-like setae on sternite VIII.   
 Distribution. Madagascar (Toliara).   
 Remarks. Dajoz (1980) gave the MNHN as the depository of the types of P. dentatus. I have not examined these specimens (they were not found during my visit to MNHN), but I neverthless have no doubts about identity of P. dentatus, as it is a very conspicuous species, figured in the original description, and the examined specimens were taken from near the type locality.   
 Telnov (2007) redefined large genus Pseudoleptaleus to include only several species from New Guinea and Australia, and mentioned that: “all species from the African continent need to be transferred to a new groups...”
description: Published as part of Kejval, Zbyněk, 2009, Taxonomic revision of the genus Chileanthicus Werner (Coleoptera: Anthicidae) 2180, pp. 1-82 in Zootaxa 2180 (1) on pages 28-29, DOI: 10.11646/zootaxa.2180.1.1, http://zenodo.org/record/5313537