Cardiodactylus contrarius Robillard & Gorochov & Poulain & Suhardjono 2014, n. sp.
Creators: name: Robillard, Tony
name: Gorochov, Andrej V.
name: Poulain, Simon
name: Suhardjono, Yayuk R.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Orthoptera
Haglotettigoniidae
Cardiodactylus
Cardiodactylus contrarius

description: Cardiodactylus contrarius Gorochov, n. sp.
(Figs 4, 5C, 18)
Type material. Male holotype: Indonesia. Sulawesi Tengah Province, Sulawesi I., Lore Lindu National Park, ~ 75 km SE of Palu City, environs of Wuasa Vill. (near eastern edge of park), ~ 1000 m, secondary forest, on large leaf of grassy plant, at night, 7–12.II.2011, A. Gorochov (ZIN). Paratypes (2♀): same data as for holotype, 1♀ (ZIN), 1♀ (MNHN-EO-ENSIF3653).
Type locality. Indonesia, Sulawesi I., Lore Lindu National Park, environs of Wuasa village.
Other material examined. Indonesia. Sulawesi Tengah Province, Sulawesi I., Lore Lindu National Park, ~ 45 km SSE of Palu City, environs of Tomado Vill. on Lindu Lake, ~ 1000 m, disturbed primary forest, on leaf of bush, at night, 13–17.II.2011, A. Gorochov, 1♀ (ZIN).
Etymology. The species is named in the connection with its contrasting (contrarius in Latin) coloration.
Distribution. Indonesia, north-west of Sulawesi Island.
Diagnosis. The new species is most similar to C. oeroe (also from Sulawesi), but it is distinguished from the latter species by the frons under rostral apex lighter, majority of darkened parts of male tegmina darker (including Sc-R area dark brown; in C. oeroe, this area is reddish), middle part of costal area dark with the proximal halves of Sc branches and crossveins between distal halves of Sc branches light (in C. oeroe, this part is with a wide light band along costal edge), tegminal mirror narrower, Sc-R area in female tegmina dark brown (not reddish) and with all crossveins or some of them clearly lighter, and epiphallus with the proximal corners of dorsal ridges widely rounded (not roundly angular) in profile.
Description. Male (holotype): Coloration very characteristic: head light brown with labrum, clypeus and frons under rostal apex yellowish (Fig. 18A), with dorsum brown and having a few lines yellowish (Fig. 18B), with apex of apical segment of all palpi and middle and distal parts of antennal flagellum brown, and with a pair of small spots near dorsolateral corners of clypeus dark brown; pronotum with lateral lobes brown but having narrow stripe along each ventral edge yellowish (Fig. 18C), and with disk yellowish and having rather numerous brown and light brown marks on its median 2/3; tegmina dark brown with several very large whitish spots (apical area of dorsal field almost completely whitish; Fig. 18D), with R and stock of Sc as well as narrow areas between them and around diagonal vein light brown, with 3–4 whitish stripes along proximal half of some branches of Sc, and with a few whitish crossveins between distal halves of these branches (Fig. 18E); exposed part of hind wings dark grayish brown with light brown venation; legs reddish brown but somewhat lighter than brown parts of head and of pronotum, with apical part of hind femora and middle segment of all tarsi dark brown, and with hind tibiae slightly darkened (grayish brown); rest of body slightly lighter than most part of legs (intermediate between brown and light brown) but with dark brown tergites having sparse lighter marks, and with uniformly light brown cerci. Body structure similar to that of C. reticulatus, however tegminal mirror 1.9 times as long as wide, Sc comb in tegmina with 7 branches (Fig. 18E), and epiphallus of genitalia somewhat narrower in middle part and with proximal corners of dorsal ridges widely rounded in profile (Fig. 18H–K).
Female: General appearance similar to that of male, but with following differences: head in one specimen with narrow brown stripe along clypeal suture; tegminal dorsal field with 9–10 brown and moderately convex longitudinal veins, majority of membranes (excepting basal and lateral areas) light brown, large light spots less numerous, majority of longitudinal veins brown, and almost all crossveins whitish (Fig. 18F); tegminal lateral field (Fig. 18G) with Sc branches somewhat more straight, with comb of Sc having 4–5 branches, and with distal spot and rest of venation from whitish to light brown (including crossveins between Sc and R; but in one female, crossveins in distal half of Sc-R area brown); abdominal apex almost as in female of C. retuculatus but with ovipositor comparatively longer (hind femur 1.2 times as long as ovipositor).
Variation. Female from Tomado village may correspond to another species. It is darker than the other specimens and C. oeroe, and ovipositor is slightly shorter (hind femur 1.3 times as long as ovipositor).
Measurements. Length in mm. Body: male 16.5, female 18–18.5; body with wings: male 21, female 22–23; pronotum: male 2.4, female 2.6–2.7; tegmina: male 12, female 13–14; hind femora: male 13.5, female 14–15; ovipositor 12–12.5.
Habitat and life history traits. Unknown.
Behavior. Unknown.
description: Published as part of Robillard, Tony, Gorochov, Andrej V., Poulain, Simon & Suhardjono, Yayuk R., 2014, Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species, pp. 1-104 in Zootaxa 3854 (1) on pages 29-31, DOI: 10.11646/zootaxa.3854.1.1, http://zenodo.org/record/4929336