Philisca robinson Soto & Ramírez 2012, new species
Creators: name: Soto, Eduardo M.
name: Ramírez, Martín J.
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Anyphaenidae
Philisca
Philisca robinson

description: Philisca robinson new species   
Figs 21–23, 32c, d, 35c  
 Type material: Holotype male and female paratype from Chile, Región V (Valparaíso), Archip. Juan Fernández, Is. Robinson Crusoe, área de Plazoleta El Yunque, S33.65276º W78.84403º (GPS, trayecto +- 25m), elev. 299m (barómetro), 16–20.II.2011, bosque de luma, M. Ramírez, E. Soto, J. Pizarro (MHNS, temporary preparations, male: EMS-00156; female: EMS-00155); same locality, 1 male and 1 female paratypes, 23–28.I.1992, S. Marshall (AMNH; male EMS-00078, female EMS-00068).   
 Etymology: The specific name is a noun in apposition taken from the type locality, which is also the name of the main character in the novel Robinson Crusoe, by Daniel Defoe.   
 Diagnosis: It resembles other species of the ornata group by the modified male chelicerae and the epigyne with smooth, heart shaped median field close to the epigastric furrow (Figs 35c, 32c, d), and is very similar to P. pizarroi by the contrasted coloration. It can be distinguished from the latter species by having a pattern of dark spots in the thoracic area (Figs 21b, 22b) and a darker venter of the opistosoma (Figs 21c, 22c). The females have the accessory bulbs closer to the spermathecae (Fig. 32d).   
 Description: Female (paratype MHNS EMS-00155): Total length 6.92. Prosoma length 2.67, width 2.07, wider between legs II–III. Tibia/metatarsus length: I, 1.50/1.23; II, 1.33/1.27; III, 1.13/1.40; IV, 1.67/2.03. Chelicerae with three teeth (middle tooth larger) on promargin, three on retromargin (basal tooth larger). Sternum length 1.40, width 1.10. Spines: leg I, femur d 1–1–1, p 2ap; tibia v 2–2–p1; metatarsus v 2bas. II, femur d 1–1–1, p d1ap; tibia v 2–2–2; metatarsus v 2bas, p d1. III, femur d 1–1–1, p and r d1ap; patella r d1; tibia v p1–p1–r1–2, p and r d1–1; metatarsus v 2–2–2, d 0–p1–2, p and r d1–1–1. IV, femur d 1–1–1, p and r d1ap; patella r d1; tibia v p1–2–2, p 1–1, r 1–1–1; metatarsus v 2–2–2, d 0–p1–2, p and r d1–1–1. Color in ethanol (Fig. 21): the entire specimen has a striped appearance. Prosoma and legs reddish brown, almost uniform, both with dark brown spots. Sternum brown, paler in the center. Opisthosoma yellowish brown, venter with dark brown marking with general“w” form (Fig. 21c). Opisthosoma length 3.99, epigastrium–spiracle 2.17, spiracle–spinnerets 0.37. Epigyne small, sclerotized, translucent (Fig. 32c, d). MF smooth, heart-shaped, close to the epigastric furrow. Anterior pouch wide, with short lumen. Ducts of AB long, with conspicuous gland ducts.   
 Male (holotype): Total length 5.59. Prosoma length 2.40, width 1.83. Length of tibia/metatarsus: I, 1.60/1.40; II, 1.47/1.43; III, 1.07/1.30; IV, 1.53/1.83. Chelicerae modified (internal superior margins projecting) (Fig. 35c), with three teeth (middle tooth larger) on promargin, three on retromargin. Sternum length 1.28, width 1.03. Spines: leg I, femur d 1–1–1, p 2ap; tibia v 2–2–2; metatarsus v 2bas, p d1. II, femur d 1–1–1, p d1ap; tibia v 2–2–2, p 0– v1 or 0; metatarsus v 2–p1–0, p 0–d1–0. III, femur d 1–1–1, p and r d1ap; patella r d1; tibia v p1–2–2, p and r d1– 1; metatarsus v 2–2–2, d 0–p1–2, p and r d1–1–1. IV, femur d 1–1–1, p and r d1ap; patella r d1; tibia v p1–2–2 or 2–2–2, p 1–1, r 1–1–1; metatarsus v 2–2–2, d 0–p1–2, p and r d1–1–1. Opisthosoma length 2.87, spiracle– epigastrium 1.73, spiracle–spinnerets 0.30. Color in ethanol as for female described above (Fig. 22). Palp (Fig. 23): Tibia width/length 0.48, Embolus normal, thin and cylindrical (Fig. 23). MA long, with few small thin branches at base (Fig. 23f, h). PMA long and wide, pPMA simple. Conductor normal and well developed, canal vestigial, limited to tip, ending in longitudinally ridged peak, on rounded projection directed backward.   
All adults and immatures have a group of thick setae on the base of ALS.  
 Variability: Spines in males: I, tibia, v 2–2–p1; metatarsus v 2bas–p1, p 0. III, metatarsus v 2–0–2. Spines in females: I, tibia, v 2–2–0. III, tibia v p1–2–2; metatarsus v 2–p1–2. IV, metatarsus v 2–p1–r1–2, p d1–2–2.   
 Distribution: Known only from the Juan Fernández Archipelago, Is. Robinson Crusoe, Chile.   
  Other material examined: CHILE: Región V (Valparaíso):  Archip. Juan Fernández: Is. Robinson Crusoe: same locality as holotype, S33.65276º W78.84403º (GPS, trayecto +- 25m), elev. 299m (barómetro), 16–20.II.2011, bosque de luma, M. Ramírez, E. Soto, J. Pizarro, 1 females, 2 males and 2 immatures (MHNS), 1 female and 3 immatures (MACN-Ar 27960), 1 male and 2 immatures (MACN-Ar 28460), 1 female (MACN-Ar 28089), 1 female (MACN-Ar 28088), 1 male (MACN-Ar 27961), pitfall, 3 males (MACN-Ar 27958), pitfall, 1 female and 1 immature (MACN-Ar 27962), pitfall, 2 females and 1 immature (MACN-Ar 27964), 1 male and 4 immatures (UC), 2 immatures (MACN-Ar 27959), pitfall, 2 immatures (MACN-Ar 27963); Sector Salsipuedes, S33.63087° W78.84386° (GPS, área +- 100m), elev. 482m (barómetro), 21.II.2011, bosque de canelo, maqui y luma, M. Ramírez, E. Soto, J. Pizarro, 2 females and 31 immatures (UC); same locality as holotype, 23–29.I.1992, S. Marshall, 1 female and 1 male (AMNH); same locality, 200–250m Camote side, 1–28.IV.1962, B. Malkin, 3 females (AMNH); same locality, site 2, pans, 23–29.I.1992, S. Marshall, 1 female and 7 males (AMNH); same locality, wet areas nr. Plazoleta, pans, 24– 28.I.1992, S. Marshall, 2 females and 6 males (AMNH); Plazoleta El Yunque, pans in wet areas, 23–29.I.1992, S. Marshall, 14 males (AMNH); same locality, pans nr. Plazoleta campsite, 23–29.I.1992, S. Marshall, 10 males (AMNH); same locality, site 1, open forest, pans, 24–28.I.1992, S. Marshall, 1 female and 1 male (AMNH); same locality, upper part; El Yunque trail, ferns, dung, 23–28.I.1992, S. Marshall, 1 female and 1 male (AMNH); same locality, slopes of El Yunque, fern covered, pan traps, 28.I.1992, S. Marshall, 2 females and 3 males (AMNH); same locality, site 1, open forest, dung traps, 23–28.I.1992, S. Marshall, 1 female and 1 male (AMNH); same locality, dung traps, 23– 29.I.1992, S. Marshall, 2 males (AMNH); Sendero, luego de Mirador Selkirk, ladera W, S33.63749º W78.85155º (GPS, trayecto +- 50m), elev. 460–539m (barómetro), 18–20.II.2011, bosque de luma con helechos arborescentes, M. Ramírez, E. Soto, J. Pizarro, 1 immature (MACN-Ar 28091), 1 immature (MACN-Ar 28090).
description: Published as part of Soto, Eduardo M. & Ramírez, Martín J., 2012, Revision and phylogenetic analysis of the spider genus Philisca Simon (Araneae: Anyphaenidae, Amaurobioidinae) 3443, pp. 1-65 in Zootaxa 3443 (1) on pages 37-38, DOI: 10.11646/zootaxa.3443.1.1, http://zenodo.org/record/5254614