Abida secale subsp. brongersmai Gittenberger 1973
Creators: name: Kokshoorn, Bas
name: Gittenberger, Edmund
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Chondrinidae
Abida
Abida secale
Abida secale brongersmai gittenberger, 1973

description: Abida secale brongersmai Gittenberger, 1973 (pl. 4 figs A–E)   
 Abida secale brongersmai Gittenberger, 1973: 115.   
Type locality: Spain, Lérida, road to Alás, near Torres, right side of the river Segre, east of La Seu d’Urgell.  
Holotype: National Museum of Natural History (RMNH), Leiden 54778.  
Distribution.—See fig. 2.  
Description (after Gittenberger 1973).—Shell slender cylindro-conical to fusiform, with 9¼–11¼ weakly to moderately inflated whorls, sculptured with regularly placed, fine axial ribs. Initial teleoconch whorls often without sculpture. Body whorl obliquely flattened, slightly narrowed, with a distinct keel. Palatalis superior accompanied by a slight to moderately prominent indentation, infrapalatalis with conspicuous external indentation just behind apertural lip. Palatal side of the aperture vertical or slightly leaning backwards (in lateral view). Umbilicus moderately wide, circular. Shell height, 8.0– 11.2 mm; width, 2.8–3.3 mm.  
Aperture roundish. Apertural lip strongly thickened, continuous across the parietal side, often protruding up to 1 mm. Angularis relatively high near the aperture but quickly decreasing in prominence, reaching past the front end of the parietalis, not or more or less clearly connected with spiralis. Subangularis well developed, not connected to angularis. Parietalis and infraparietalis well developed. Several weak to strong folds may be present on the apertural lip. Columellaris ending abruptly on the columella, clearly broader than the other lamellae there. Infracolumellaris, reaching further beyond the columella, often connected with a lamella on the apertural lip. Infrapalatalis, palatalis inferior and palatalis superior indented, reaching up to the edge of the aperture. At the anterodorsal palatal center a suturalis, an equally long or longer suprapalatalis and often a short basalis are present.  
Genetic barcode. —GenBank accession numbers EU395348, EU395357 – EU395367.  
 Notes.—Both shell sculpture and apertural protrusion vary in this conspicuous subspecies. This variation is partly correlated with altitude (Kokshoorn 2008: 99–114). In the easternmost part of its present range A. s. brongersmai occurs close to A. s. margaridae, but intermediate populations are not known. This might be due to the introduction of A. s. brongersmai in the area north of Pedra (SE of Bellver de Cerdanya). In that area, limestone rocks have been used for the construction of walls, creating an artificial habitat for A. secale. Maybe the rocks originated from the region where A. s. brongersmai occurs.
description: Published as part of Kokshoorn, Bas & Gittenberger, Edmund, 2010, Chondrinidae taxonomy revisited: New synonymies, new taxa, and a checklist of species and subspecies (Mollusca: Gastropoda: Pulmonata) 2539, pp. 1-62 in Zootaxa 2539 (1) on pages 16-17, DOI: 10.11646/zootaxa.2539.1.1, http://zenodo.org/record/10094211