Aleochara (Emplenota) fucicola Sharp 1874
Creators: name: Yamamoto, Shûhei
name: Maruyama, Munetoshi
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Aleochara
Aleochara fucicola

description: Aleochara (Emplenota) fucicola Sharp, 1874
(Figs. 1, 7–22, 93–94, 97, 101)
Aleochara fucicola Sharp, 1874: 9 (original description; type locality: “Amakusa and Iwosima, near Nagasaki” [Amakusa-shi, Kumamoto-ken (or Iô-jima, Nagasaki-shi, Nagasaki-ken), Kyûshû, Japan]); Lewis, 1879: 6 (catalogue of Japanese Coleoptera); Schönfeldt, 1887: 66 (catalogue of Japanese Coleoptera); Sawada, 1972: 36 (maxilla, illustrated), 37 (description of lacinia), 39 (chaetotaxy of labial pulpus), 40 (comments on pores on labial palpus), 40 (comments on paramerite); Shibata, 1985: 321 (description in Japanese; Coleoptera of Japan), pl. 56 (habitus of specimen); Naomi, 1989: 280 (checklist of Japanese Staphylinidae); Li & Wang, 1993: 154 (catalogue of soil beetles of Anhui Province, China).
Aleochara (Emplenota) fucicola Sharp, 1874; Fenyes, 1921: 415 (catalogue of world species of Aleocharinae); Scheerpeltz, 1925: 447 (catalogue of Palaearctic species of Staphylinidae); Bernhauer & Scheerpeltz, 1926: 796 (catalogue of world species of Staphylinidae); Adachi 1957: 34 (catalogue of Japanese species of Staphylinidae); Nakane, 1963: 100 (description in Japanese; Coleoptera of Japan), pl. 50 (habitus of specimen); Sawada, 1971: 309 (redescription); Sawada, 1972: table (character states, listed); Welch, 1997: 9 (comments on carina on mesoventrite); Maus & Ashe, 1998a (online) (checklist of subgenus of world; phylogenetic relationship); Cho & Ahn, 2001: 14, 30 (checklist of Korean species of Silphidae and Staphylinidae), 157 (pl. 2) (habitus of specimen); de Rougemont, 2001: 84 (record from Hong Kong); Park & Ahn, 2004: 195 (key to littoral species of Korean Aleochara; redescription); Smetana, 2004: 356 (catalogue of Palaearctic Aleocharinae); Yamazaki, 2008: 152 (dipteran host record); Frank & Ahn, 2011: 19 (checklist of coastal Staphylinidae of world); Yamazaki, 2012: 32 (ecological research).
Emplenota fucicola (Sharp, 1874); Assing, 1995: 220 (diagnostic key to Palaearctic species of Emplenota), 223 (redescription; lectotype designation); PaŠnik, 2001: 231 (record from North Korea); Dauphin, 2005: 48 (record from Gironde, France).
Homalota variolosa Weise, 1877: 89 (original description; type locality: “Hagi” [Yamaguchi-ken, western end of Honshû, Japan]); Lewis, 1879: 6 (catalogue of Japanese Coleoptera); Schönfeldt, 1887: 66 (catalogue of Japanese Coleoptera); Assing, 1995: 223 (synonymized with A. fucicola; lectotype designation); Ahn et al., 2000: 244 (as synonym of A. fucicola); Smetana, 2004: 356 (catalogue; as synonym of A. fucicola); Park & Ahn, 2004: 196 (as synonym of A. fucicola); Frank & Ahn, 2011: 19 (checklist; as synonym of A. fucicola).
Aleochara (Emplenota) variolosa (Weise, 1877); Fenyes, 1921: 415 (catalogue of world species of Aleocharinae); Scheerpeltz, 1925: 447 (catalogue of Palaearctic species of Staphylinidae); Bernhauer & Scheerpeltz, 1926: 797 (catalogue of world species of Staphylinidae); Adachi 1957: 34 (catalogue of Japanese species of Staphylinidae).
Aleochara variolosa (Weise, 1877); Li & Chen, 1990: 19 (catalogue of northeast Chinese species of Staphylinidae); Li, 1993: 46 (catalogue of northeast Chinese species of Staphylinidae); Li & Wang, 1993: 154 (catalogue of soil beetles of Anhui Province, China).
Type specimens. A. fucicola: [Lectotype]: Ƥ, “ Japan [YRLS]// Japan./G. Lewis//Sharp Coll./1905-313// Aleochara / fucicola Ƥ[HW]/ type D. S. [HW]// Lectotypus /design. Assing, 1994//LECTO-/ TYPE [PRL]” [Paralectotypes]: 2 3, “ Type [RRL]// Japan [YRLS]// Japan./G. Lewis//Sharp Coll./1905-313// Aleochara / fucicola Sharp ”; 1 Ƥ, 1 sex?, “ Japan [YRLS]// Japan./G. Lewis//Sharp Coll./1905-313//Para-/lecto-/ type [BRL]”. Designated by Assing (1995).
H. variolosa: Not examined.
Non-type specimens. JAPAN: [Honshû]: 1 3, 1 Ƥ, Kosode, Ube-chô, Kuji-shi, Iwate-ken (40.168N, 141.848E), 28 VI 2010, Ôhara-M., Kobayashi-N., Yamamoto-H. (40°10ʹ0 5ʺN, 141°50ʹ54ʺE; collected from seaweed (Sargassum sp., Laminaria sp.) and eel grass (Zostera sp.) on cobble beach; collections of HUM: HN-10- MO-043/CO); 1 3, 1 Ƥ, Raga, Tanohata-mura, Shimohei-gun, Iwate-ken (39.939N, 141.940E *), 30 V 1998, Watanabe-T. (cWat); 2 3, 3 Ƥ, Omoto, Iwaizumi-chô, Shimohei-gun, Iwate-ken (39.850N, 141.974E), 27 VI 2010, Ôhara-M. (39°50ʹ59ʺN, 141°58ʹ27ʺE; sandy beach; HUM: HN-10-MO-042/SA); 2 3, 1 Ƥ, Hamaogi-kaigan, Hamaogi, Kamogawa-shi, Chiba-ken (35.118N, 140.144E), 24 IV 1990, Takeda-T. (KUM); 1 Ƥ, Shinmaikokaigan, Yawata, Futtsu-shi, Chiba-ken (35.250N, 139.866E), 11 VI 1995, Emoto-K. (cWat); 2 Ƥ, river mouth of Obitsu-gawa, Kuzuma, Kisarazu-shi, Chiba-ken (35.412N, 139.906E *), 19 II 2007, Ono-H. (cOno); 1 3, 2 Ƥ, same data, but 2 III 2007; 5 3, 8 Ƥ, same data, but 13 III 2007; 1 3, 2 Ƥ, river mouth of Obitsu-gawa, Kuroto, Kisarazu-shi, Chiba-ken (35.411N, 139.900E), 27 V 1983, Nemoto-K. (under drifted woods; KUM); 4 3, 9 Ƥ, same place, but 25 IV 1987, Haga-K. (from decaying seaweed; KUM); 3 Ƥ, Yakenga-hama (Hachijô-jima), Ôkagô, Hachijô-machi, Tôkyô-to (33.100N, 139.770E), 26 VI 2011, Hayama-T. (33d06'02N, 139d46'11E; shingle beach, composed of lava; cHayam); 3 3, 1 Ƥ, same data, but 27 VI 2011; 1 Ƥ, Kaneda beach, Kaneda, Minamishitauramachi, Miura-shi, Kanagawa-ken (35.167N, 139.660E), 19 III 2001, Watanabe-T. (cWat); 1 Ƥ, Bishamon, Minamishitaura-machi, Miura-shi, Kanagawa-ken (35.142N, 139.650E *), 21 IV 1994, Ohgi-K. (cWat); 2 3, Jôgashima (Jôga-shima), Misaki-machi, Miura-shi, Kanagawa-ken (35.132N, 139.621E *), 10 IV 2003, Watanabe- T. (cWat); 3 Ƥ, Mito beach, Hasse-machi, Miura-shi, Kanagawa-ken (35.177N, 139.620E), 28 V 2001, Watanabe- T. (cWat); 1 3, 2 Ƥ, same data, but 29 V 2007; 1 Ƥ, same data, but 14 V 2009; 1 Ƥ, same data, but 12 VI 2009; 1 Ƥ, Arasaki, Nagai, Yokosuka-shi, Kanagawa-ken (35.194N, 139.600E *), 12 VI 2009, Watanabe-T. (cWat); 1 Ƥ, Enoshima (Eno-shima), Fujisawa-shi, Kanagawa-ken (35.300N, 139.485E *), 11 VI 2001, Watanabe-T. (cWat); 1 Ƥ, Hatsushima (Hatsu-shima), Atami-shi, Shizuoka-ken (35.041N, 139.167E *), 12 XI 2000, Kurihara-T. (shore reef on intertidal zone; EEEU); 39 3, 59 Ƥ, Tsumeki-zaki, Shimoda-shi, Shizuoka-ken (34.659N, 138.987E), 21 IV 2003, Maruyama-M. (cMar); 5 3, 7 Ƥ, Ogi, Noto-machi, Fugeshi-gun, Ishikawa-ken (N37.301, 137.234E), 1 V 1961, Hayashi-Y. (cHayas); 1 Ƥ, Ôshima (Ô-shima), Kushimoto-chô, Higashimuro-gun, Wakayama-ken (33.468N, 135.800E *), 30 IV 1959, Kimura-Y. (cHayas); 1 3, Isonoura, Wakayama-shi, Wakayama-ken (34.259N, 135.093E), 21 II 1981, Itô-T. (cHayas); 1 Ƥ, Miyama (Tomoga-shima), Wakayama-shi, Wakayama-ken (34.280N, 135.014E *), 16 VI 2007, Kawakami-Y. (cKaw); 1 3, 2 Ƥ, Nada-kaigan (Awaji-shima), Nada, Minamiawaji-shi, Hyôgo-ken (34.207N, 134.812E *), 1 IV 1977, Kaneda (KUM); 1 Ƥ, Nakayagi, Akashi-shi, Hyôgo-ken (34.670N, 134.931E), 9 V 1999, Kawakami-Y. (cKaw); 1 Ƥ, river mouth of Chigusa-gawa, Onzaki, Akô-shi, Hyôgo-ken (34.731N, 134.393E), 15 VI 1999, Kawakami-Y. (cKaw); 1 Ƥ, Kugui, Higashi-ku, Okayama-shi, Okayama-ken (34.584N, 134.086E *), 31 III 2003, Fujitani-Y. (under seaweed; KUM); 2 3, 2 Ƥ, same data, but 5 V 2003; 1 3, 4 Ƥ, same data, but 9 V 2003 (under seaweed); 4 3, 3 Ƥ, same data, but 28 V 2003 (under seaweed); 1 3, 1Ƥ, Kusumihana, Ôbatake, Kurashiki-shi, Okayama-ken (34.430N, 133.822E *), 8 VI 2003, Fujitani-Y. (KUM); 7 3, Okidomari, Tako, Shimane-chô, Matsue-shi, Shimane-ken (35.604N, 133.096E *), 3 VI 2008, Hayama-T. (from seaweed; cHayam); 1 Ƥ, Konami-kaigan, Nonami, Shimane-chô, Matsue-shi, Shimane-ken (35.588N, 133.098E), 19 VII 2010, Yamamoto-S. (from seaweed on small sandy beach; cYam); 9 3, 9 Ƥ, Kakanokuketo, Shimane-chô, Matsue-shi, Shimane-ken (35.578N, 133.050E), 1 IV 2009, Hayama-T. (shingle beach; cHayam); 4 Ƥ, same data, but 9 VI 2009; 3 3, 3 Ƥ, same data, but 17 VII 2010, Yamamoto-S. (from rotten seaweed on shingle beach; cYam); 2 3, 5 Ƥ, same data, but 18 VII 2010; 1 Ƥ, same data, but 19 VII 2010; 1 3, 39 Ƥ, Koura-kaigan, Koura, Kashimachô, Matue-shi, Shimane-ken (35.521N, 132.975E), 8–10 VI 2009, Hayama-T. (sandy beach; FIT; cHayam); 1 3, 41 Ƥ, same data, but 18–20 VI 2009; 1 Ƥ, same data, but 20–22 VI 2009; 1 3, 5 Ƥ, same data, but 3–5 VII 2009; 18 Ƥ, same data, but 16–18 VII 2009; 5 3, 6 Ƥ, same data, but 17 VII 2010, Yamamoto-S. (under flotsam on sandy beach; cYam); 3 3, 1 Ƥ, same data, but 18 VII 2010; 1 Ƥ, Sakaura-kaigan, Sakaura-chô, Izumo-shi, Shimane-ken (35.508N, 132.861E), 4 VII 2008, Hayama-T. (shingle beach with shore reef; cHayam); 1 3, same data, but 20 IV 2009; 4 Ƥ, same data, but 15 V 2009 (from seaweed); 1 Ƥ, river mouth of Sakaura-gawa, Sakaura-chô, Izumo-shi, Shimane-ken (35.508N, 132.861E), 30 IV 2008, Hayama-T. (cHayam); 1 3, same data, but 31 VIII 2008; 1 3, same data, but 24 III 2009; 2 3, 2 Ƥ, river mouth of Mitsu-gawa, Mitsu-chô, Izumo-shi, Shimane-ken (35.498N, 132.826E), 30 IV 2008, Hayama-T. (cHayam); 4 3, 8 Ƥ, Owashi-hama, Nakayama, Taisha-chô, Izumo-shi, Shimane-ken (35.433N, 132.634E), 24 III 2009, Hayama-T. (shingle beach; cHayam); 3 3, 5 Ƥ, same data, but 10 IV 2009;; 1 Ƥ, Akaishihana, Hinomisaki, Taisha-chô, Izumo-shi, Shimane-ken (35.411N, 132.650E *), 2 IV 2009, Hayama-T. (cHayam); 8 3, 11 Ƥ, same data, but 2–8 V 2009 (FIT); 1 Ƥ, Kuchitaki, Taki-chô, Izumo-shi, Shimaneken (35.271N, 132.581E), 18 IV 2009, Hayama-T. (cHayam); 2 Ƥ, Kiami-chô, Masuda-shi, Shimane-ken (34.676N, 131.750E *), 15 VI 2008, Hayama-T. (cHayam); 1 Ƥ, same data, but 10 V 2009. [Shikoku]: 1 3, Komatsubara, Matsubara, Higashikagawa-shi, Kagawa-ken (34.254N, 134.377E), 5 IV 2008, Fujimoto-H. (KUM); 1 3, 4 Ƥ, Ariake-hama, Muromoto-chô, Kanonji-shi, Kagawa-ken (34.139N, 133.642E), 9 IV 2011, Fujimoto-H. (sandy beach; KUM); 20 3, 11 Ƥ, Iwagi (Akahone-jima), Kamijima-chô, Ochi-gun, Ehime-ken (34.235N, 133.159E *), 2 V 2009, Satô-Y. (EEEU); 1 3, same data, but 7 V 2010, Senda-Y.; 1 3, Shimoda (sandy and pebbly beach near Hirano-gyokô fishing port), Shimanto-shi, Kôchi-ken (32.948N, 132.995E), 21 IV 2012, Yamamoto-S. (from decaying seaweed on sandy beach; cYam); 2 Ƥ, Murotomisaki, Murotomisaki-chô, Muroto-shi, Kôchi-ken (33.245N, 134.177E *), Kan-T. (EEEU). [Kyûshû]: 3 3, 4 Ƥ, Kuroga-hama, Ôguro, Saganoseki, Ôita-shi, Ôita-ken (33.258N, 131.898E), 20 VI 2012, Yamamoto-S. (under seaweed and flotsam during daytime on cobble beach; cYam); 3 3, 1 Ƥ, Kin (Tsu-shima), Kamitsushima-chô, Tsushima-shi, Nagasaki-ken (34.568N, 129.469E), 5 V 2009, Yamamoto-S. (from decaying seaweed on sandy beach with Aleochara (Triochara) trisulcata; cYam); 2 3, Kazusa-chô, Minamishimabara-shi, Nagasaki-ken (32.625N, 130.164E *), 8 II 1978, Imasaka-S. (KUM); 1 Ƥ, Kuchinotsu-chô, Minamishimabara-shi, Nagasaki-ken (32.610N, 130.190E *), 18 II 1979, Imasaka-S. (KUM); 1 Ƥ, Nomo-machi (Nomo-zaki), Nagasaki-shi, Nagasaki-ken (32.580N, 129.754E *), 3 VI 1987, Yahiro-K. (KUM); 24 3, 39 Ƥ, Tomioka (Amakusa-syotô), Reihoku-machi, Amakusa-gun, Kumamoto-ken (32.519N, 130.037E), 3 IV 1977, Naomi-S.I. (KUM); 2 3, Sato (Kamikoshiki-jima), Sato-machi, Satsumasendai-shi, Kagoshima-ken (31.844N, 129.919E), 11–12 IV 1973, Furuki-Y. (EEEU).
Redescription. Body (Fig. 1): medium size; robust, narrowly subparallel; surface of head, pronotum and elytra covered with numerous hairs; somewhat densely and numerous distinct punctures on dorsal surface (head, pronotum and elytra). Colour (Fig. 1): blackish brown in ground colour, but slightly paler in elytra; legs, especially tarsal segments, brown to reddish brown; maxillary and labial palpi reddish brown to yellowish brown; antennae dark brown but reddish brown around basal segements. Head: longitudinal impunctured area along midline on dorsal surface not clearly swollen (not carinate); entire surface roughly covered with setae except for impunctured area. Antennae (Fig. 8): moderately thick and robust; segment I, 2.3 times as long as broad; segment II clearly shorter than I; segment III slightly longer than II but shorter than I; each segment of IV to VI as long as width; segments VII to X moderately broader than length; segment XI subconical with rounded apical margin, 1.4 times as long as broad; relative length (width) of segments from basal to apical: 9(4): 5.5(3): 6(3.5): 3.5(3.5): 4(4): 4(4): 3.5(4.5): 3.5(5): 3.5(5): 3.5(5.5): 7.5(5.5). Thorax: pronotum slightly wider than long (PW/PL =1.23), a little broader than head (PW/HW =1.34); dorsal surface mat with hexagonal reticulations, and with somewhat shallow, but distinct punctures. Metaventrite (Fig. 7), about 1.8 times as long as mesoventrite. Inter coxal process of mesoventrite pointed, with short carina, about 0.4 times as long as mesoventrite. Inter coxal process of metaventrite (Fig. 7) broad, rather short, one third as long as mesocoxal cavity, rounded apically. Legs: relative lengths of tarsomeres from basal to apical: 5: 3: 3: 3: 11 in foretarsus, 7: 4.5: 4.5: 5: 13 in midtarsus, 8: 7: 6: 6: 15.5 in hindtarsus.
[Male]: tergite VIII (Fig. 14) slightly rounded toward apex, with around 6 macrosetae. Sternite VIII (Fig. 16) with about 4 macrosetae and around 10 thin macrosetae; posterior margin weakly produced medially. Median lobe of aedeagus (Figs. 18–19) elongated, moderately narrowed apically, elongated pyriform in ventral view (Fig. 19); a pair of subapico-ventral projections gently curved in lateral view (Fig. 18); basal swelling of aedeagus (bs), circularly rounded in lateral view (Fig. 18); apical lobe of median lobe clearly isosceles shape in ventral aspect (Figs. 19–20); flagellum slightly shorter than the whole length of median lobe (Figs. 18–19).
[Female]: posterior margin of tergite VIII (Fig. 15) rounded, with around 7 macrosetae. Sternite VIII (Fig. 17) with 3 large macrosetae and around 4 thinner macrosetae; posterior margin slightly pointed. Spermatheca (Fig. 22): spermathecal head more or less as long as apical portion of spermathecal stem; basal portion of spermathecal stem (sb) slightly narrowing toward base, lacking any bent around middle and usually at base; sclerotized portion of spermathecal stem erect, connected with (sb) almost horizontally; each part of spermatheca except for membraneous portion of spermathecal duct (sm) entirely and moderately sclerotized; (sm) moderate in length.
Measurements (male: n=10): BL, 3.58–4.59 (4.02±0.32); FBL, 1.16–2.03 (1.83±0.16); HL, 0.43–0.60 (0.51±0.06); HW, 0.46–0.66 (0.59±0.06); AL, 0.79–1.27 (1.08±0.15); PL, 0.48–0.73 (0.64±0.07); PW, 0.50–0.90 (0.77±0.12); EL, 0.54–0.83 (0.67±0.09); EW, 0.77–1.21 (1.02±0.13); HTL, 0.45–0.73 (0.61±0.08).
Measurements (female: n=10): BL, 3.36–5.18 (4.23±0.55); FBL, 1.63–2.12 (1.90±0.16); HL, 0.47–0.57 (0.53±0.03); HW, 0.49–0.65 (0.60±0.05); AL, 0.87–1.23 (1.08±0.12); PL, 0.56–0.73 (0.66±0.05); PW, 0.66–1.04 (0.83±0.11); EL, 0.58–0.80 (0.69±0.08); EW, 0.86–1.18 (1.06±0.11); HTL, 0.48–0.71 (0.61±0.08).
Diagnosis. This species can be distinguished from the other species of Emplenota by a combination of the following character states: normally medium body sized; dorsal surface of forebody with numerous distinct punctures (Fig. 1); impunctured area of dorsal surface of head scarcely swollen (almost flattened); mesoventrite with short carina about 0.41 times as long as mesoventrite (Fig. 7). [Male]: sternite VIII (Fig. 16) somewhat weakly pointed toward apex; subapico-ventral projections on median lobe gently curved in lateral view (Fig. 18); basal swelling of median lobe (bs), circularly rounded in lateral view (Fig. 18); median lobe elongated pyriform (Fig. 19) and apical lobe isosceles shape in ventral aspect (Fig. 20). [Female]: basal portion of spermathecal stem (sb) (Fig. 22) simple, and gently or moderately curved without bent at middle and base, with sclerotized and erect sclerotized portion of spermathecal stem (ss).
Confirmed distribution by present study. [JAPAN]: Honshû, Shikoku, Kyûshû, Hachijô-jima, Awaji-shima, Tsu-shima, Amakusa-shotô, Kamikoshiki-jima. See, Fig. 101 for Japanese distribution.
Other localities in literature. [SOUTH KOREA]: Chungnam Province, Gyeongnam Province, Jeonnam Province, Jeju Province (Park & Ahn, 2004); [NORTH KOREA]: Hamgyong Province (PaŠnik, 2001); [CHINA]: Heilongjiang (Li & Chen, 1990, 1993: both doubtful records), Anhui (Li & Wang, 1993), Hong Kong (de Rougemont, 2001); [RUSSIA]: Far East (Ahn et al., 2000: no original citation; locality doubtful); [FRANCE]: Région Aquitaine (Dauphin, 2005).
Remarks. This species was originally described “under seaweed at Amakusa and Iwosima, near Nagasaki”, (Kyûshû, Japan: Sharp, 1874) and later redescribed by Sawada (1971), Assing (1995), and Park and Ahn (2004). Sawada (1971) emphasised the chaetotaxy of mouth parts, while Assing (1995) paid special attention to the setal condition of the dorsal surface of pronotum. Park and Ahn (2004) provided a short diagnosis with figures.
Ahn et al. (2000) shortly redescribed Aleochara fucicola, but Park and Ahn (2004) regarded it as a misidentification of A. puetzi (Assing, 1995) and redescribed it again. However, according to their illustrations of the aedeagus, it is part of a new species being described in the present paper, A. segregata, although the figure of the spermatheca in Ahn et al. (2000) seems to belong to A. fucicola.
The distribution of A. fucicola is very wide. Although records have been restricted in East Asia for a long time, Dauphin (2005) recorded the species from north-western France (Région Aquitaine, Gironde). This amazing record seems accurate, as Dr. V. Assing identified the specimens. Dauphin (2005) also mentioned that the records might have been unintentional invasions from other regions, and special attention needs to be paid to this taxon with regard to its distribution.
The records from mainland China adopted in Li and Chen (1990, 1993) are highly doubtful, because this species’ record locality (Heilongjiang) is inland.
Assing (1995) synonymised Homalota variolosa Weise, 1877 with A. fucicola, and designated a lectotype for the type series of each species.
Yamazaki (2008) revealed one of the host species of A. fucicola as Fucellia apicalis Kertész, 1908 (Diptera, Anthomyiidae). Yamazaki (2012) reported that the kelp fly, Coelopa frigida (Fabricius, 1805) (Diptera, Coelopidae) was an additional fly host. The article also noted the parasitized months (April, November, December) by A. fucicola.
description: Published as part of Yamamoto, Shûhei & Maruyama, Munetoshi, 2012, Revision of the Seashore-dwelling Subgenera Emplenota Casey and Triochara Bernhauer (Coleoptera: Staphylinidae: genus Aleochara) from Japan, pp. 1-52 in Zootaxa 3517 on pages 11-16, DOI: 10.5281/zenodo.282595