Mortoniella (Mortoniella) quadrispina Blahnik & Holzenthal 2017, new species
Creators: name: Blahnik, Roger J.
name: Holzenthal, Ralph W.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Magnoliopsida
Gentianales
Apocynaceae
Mortoniella
Mortoniella quadrispina

description: Mortoniella (Mortoniella) quadrispina, new species   
Fig. 54  
 This species is apparently very closely related to M. auricularis, n. sp., differing primarily in the shape of the apex of the dorsal phallic spine. Mortoniella quadrispina has two apicolateral projections, as in M. auricularis, but also has the ventral lobe sharply angulate, rather than rounded, thus producing 4 spine-like projections on the apex of the dorsal phallic spine (1 apicodorsal, 2 lateral, and 1 apicoventral).   
 Adult —Length of forewing: male 3.5 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0:4:4. Overall color (in alcohol) medium brown.   
 Male genitalia —Ventral process of segment VI laterally compressed, ventrally projecting, large, subtriangular, length about equal to width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short, lateral margins subparallel, apicomesal margin with U-shaped emargination, extending about 1/3 length of tergum, apicolateral lobes tapering, acute apically, relatively short and narrow as viewed laterally; apicomesal lobes visible in lateral view, rounded apically; ventrolateral lobes distinct, projecting. Inferior appendages without dorsolateral projections, apicoventral projections paired, narrow, nearly linear, moderately elongate, acute apically. Mesal pockets of inferior appendage with apical processes large, prominent, posterodorsally curved. Paramere appendage moderately elongate (shorter than dorsal phallic spine), nearly uniformly narrow, apex acute. Dorsal phallic spine compressed apically; as viewed laterally, with acute apicoventral projection, preapically (or apically) with pair of small acute lateral projections, apicomesally with additional small acute projection; spine, in dorsal view, nearly uniform in width, apicolateral projections evident. Phallicata with pair of small rounded basoventral projections, and somewhat indistinct, rounded, apicoventral lobes. Endophallic membrane short, with pair of very enlarged, curved, sclerotized spines (possibly phallotremal complex); ventromesal spine absent.   
 Holotype male (alcohol)— ECUADOR: Pastaza: Puyo (27 km. N), Estación Fluvia Metrica, 4.ii.1976, Spangler, et al. (UMSP000097062) (NMNH).   
 Etymology —This species is named M. quadrispina for the four acute projections on the apex of the dorsal phallic spine.
description: Published as part of Blahnik, Roger J. & Holzenthal, Ralph W., 2017, Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *, pp. 1-251 in Insecta Mundi 2017 (602) on pages 69-70, DOI: 10.5281/zenodo.5170203