Klyveria Burckhardt, Serbina and Malenovsky 2024, gen. nov.
Creators: name: Aléné Geonho Cho, Daniel Burckhardt Liliya Š. Serbina Igor Malenovský Dalva L. Queiroz Désirée C.
name: Percy, Diana M.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Liviidae
Klyveria

description: Klyveria Burckhardt, Serbina and Malenovský gen. nov.
Zoobank registration: urn:lsid:zoobank.org:act: 568C5A77- 9010-4F66-B83E-575317BA95A5.
Type species: Paurocephala crassiflagellata Burckhardt, 1996, by present designation.
Diagnosis: Adult. Head, in lateral view, deflexed 45–90° from longitudinal axis of body (Fig. 1H); in dorsal view slightly wider than thorax, very short and wide. Vertex rhomboidal, moderately to strongly transverse (Fig. 2B); covered in finely granular microsculpture, sometimes much reduced on disc; passing smoothly into genae anteriorly; coronal suture fully developed; genae weakly produced ventrally but not enlarged into processes; frons small trapezoidal (Fig. 3G); median ocellus visible in perpendicular view to vertex; compound eyes, in dorsal view, subspherical, collared. Clypeus pear-shaped, large, flattened ventrally, hardly visible in lateral view as it is hidden by the genae. Antenna 10-segmented, distinctly longer than head width; flagellum with simple setae; segment 3 thickened, longest, about as long as segments 4–5 together; segments 3, 4, 6, 8, and 9 bearing each a subapical rhinarium lacking marginal spines. Thorax moderately slender; dorsal outline, in lateral view, weakly curved. Pronotum, in dorsal view, weakly curved posteriad laterally; propleurites narrowly subrectangular, divided by perpendicular suture into subequal epimeron and episternum. Metapostnotum with blunt tubercle. Mesosternum (Fig. 5C) narrower than head, forming transverse band more than three times as wide as long laterally; anterior margin weakly concave with median hump; pleurosternal suture not visible; basisternum triangular; katepisternum small antero-laterally, not bent dorsad laterally; angle between arms of precoxale right or acute. Pro- and mesotibiae cylindrical. Metacoxa with subacute horn-shaped meracanthus. Metafemur with the three ventral sense organs in medial position (Fig. 6K); apex with a group of stout, long setae. Metatibia longer than metafemur, hardly widened apically; bearing 12 or 13 densely spaced apical, weakly sclerotized spurs, without posterior peg-likw or thorn-like setae (Fig. 7I). Both metatarsal segments relatively short, subequal in length. Forewing oval or oblong-oval, widest in apical third; 2.2–2.3 times as long as wide, membranous; vein C + Sc weakly, evenly convex, slender, distinctly delimited from cell; costal break developed, close to apex of vein R 1; pterostigma narrow or wide, entirely membranous; nodal line developed; veins R and M + Cu subequal; vein Rs weakly convex relative to costal margin or sinuous; vein M longer than M 1 + 2; vein Cu 1a weakly or strongly curved towards anal margin; veins M 1 + 2 and M 3 + 4 perpendicular to wing margin apically; anal break adjacent to apex of vein Cu 1b; surface spinules fine, spaced, present in all cells. Hindwing slightly shorter than forewing; with two or three costal setae proximal and five to eight setae distal to costal break; vein R + M + Cu bifurcating into R and M + Cu. Abdominal base with a sclerotized area on either side covered in spines. Male proctiger simple (Fig. 9A). Aedeagus with simple proximal portion bearing many weak folds subapically; distal portion moderately to strongly expanded basally, apex subdivided from stem (Fig. 9G). Female subgenital plate bearing apical process.
Last instar immature. Antenna 10 segments; bearing sectasetae or lanceolate setae on antennal flagellum. Mid- and hindlegs without massive peg-like setae. Tarsal arolium small, pedicel with lateral wings (Fig. 9L). Dorsal body surface bearing minute clavate setae. Precaudal abdominal tergites lacking densely spaced simple setae or sectasetae. Anus in terminal position; with additional pore fields developed.
Etymology: Dedicated to the US entomologist Frederick Detlev Klyver for his exemplary contributions to psyllid taxonomy.
Comments: Monophyly strongly supported in all analyses (Figs 10, 11; Supporting Information, File S4). Klyveria currently includes two described species: K. crassiflagellata (Burckhardt 1996: 79), comb. nov. (from Paurocephala; Diclidophlebia, Burckhardt and Mifsud 2003: 13) and K. setinervis (Burckhardt 1996: 78), comb. nov. (from Paurocephala; Diclidophlebia, Burckhardt and Mifsud 2003: 14). Their distribution and host plants are summarized in Table 3 and Supporting Information, File S3.
description: Published as part of Aléné Geonho Cho, Daniel Burckhardt Liliya Š. Serbina Igor Malenovský Dalva L. Queiroz Désirée C. & Percy, Diana M., 2024, Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data, pp. 387-421 in Zoological Journal of the Linnean Society 201 (2) on pages 409-410, DOI: 10.1093/zoolinnean/zlad128, http://zenodo.org/record/13219879