Cambonilla securicula Jocqué & Jocque & Stock & Rin & Henrard 2019, gen. et sp. nov.
Creators: name: Jocqué, Rudy
name: Jocque, Merlijn
name: Stock, Willem
name: Rin, Naroeun
name: Henrard, Arnaud
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Zodariidae
Cambonilla
Cambonilla securicula

description: Cambonilla securicula Jocqué gen. et sp. nov.
urn:lsid:zoobank.org:act: 00D14C53-6330-42BA-A2F1-63C760959CF3
Figs 2, 3 B–F, 4A, 5, 6, 7, 11
Diagnosis
The male of Cambonilla securicula sp. nov. is recognized by the presence of small lateral scuta behind the epigastric fold and by the male palp in which both the MA and the C are hatchet shaped. The female has a remarkable, small, modified spine on the tibia of the palp and a characteristic epigyne in which the central plate has a concave anterior and a straight posterior margin.
Etymology
The species name ‘ securicula ’ meaning hatchet, is a noun in apposition referring to the shape of both the MA and the C.
Material examined
Holotype CAMBODIA • ♂; Kratie Province, Base Camp; 13.00986° N, 106.0640° E; 16 May 2018; M. Jocque & W. Stock leg.; Herpetology pitfall in forest; RBINS IG 33.889/001.
Paratypes CAMBODIA • 3 ♂♂; same data as for holotype; RBINS IG 33.889/002 (=CA3) • 2 ♂♂, 1 ♀; 17 May 2018; same data as preceding; RBINS IG 33.889/003 • 4 ♂♂ 2 ♀♀; 13 May 2015; same data as preceding; RBINS IG 33.889/004 • 4 ♂♂; Kratie Province, Base Camp 4; 13.0198° N, 105.9268° E; 5 May 2018; M. Jocque and W. Stock leg.; pitfall set 1; RBINS IG 33.889/005 • 1 ♂; Kratie Province; 13.00986° N, 106.06395° E; 10 May 2018; M. Jocque and W. Stock leg; pitfall set 3; RBINS IG 33.889/006.
Other material
CAMBODIA • 1 ♂, 1 ♀; Kratie Province, Base Camp; 13.00986° N, 106.0650° E; 6–11 May 2018; M. Jocque and W. Stock leg.; forest, by hand; RBINS IG 33.889/010 • 2 ♂♂; Kratie Province, Base Camp 1; 12.61788° N, 106.0 0 296° E; 10 Apr. 2018; M. Jocque and W. Stock leg.; flooded forest on sand, Malaise trap, by hand; RBINS IG 33.889/011.
LAOS – Vientiane Province • 1 ♀; Vientiane, Ban Don Ma Khay; 18.08384° N, 102.1713° E; 13 Jun. 2013; P. Jäger leg.; 186 m, secondary forest, at night by hand; SMF – Champasak Province • 1 ♀; Muang Bachieng, That Paxuam; 15.17653° N, 105.9225° E; 3.Aug. 2016; P. Jäger and S. Munnich leg.; 197 m,secondary forest with single primary trees, by night, by hand; SMF – Khammouan Province • 1 ♀; Ban Tathot; 17.62361° N, 105.1466° E; 18–19 Fev. 2003; P. Jäger leg.; ca 180 asl, village and neighbourhood, shrubs and rocks, night and evening; SMF.
Description
Male holotype
TOTAL LENGTH. 6.96. Carapace: length 3.48, width 2.70, height 2.00.
COLOUR (Fig. 5 A–D). Carapace uniform dark chestnut brown; chelicerae dark brown with orange extremity and white tip; chilum medium brown; sternum medium brown; labium and endites medium brown, paler towards anterior white tip; legs: Cx and Tr medium brown, Fe dark brown, P and T greyish brown, Mt and t pale brown.Abdomen: dorsum with pale reniform patches fused in front and at posterior end, overlaid by reddish orange, translucent oval scutum; further backward with five transverse bands and pale patch above spinnerets on dark background; venter sclerotized in front of epigastric furrow and yellow sclerotized area on either side behind epigastric furrow; remainder marbled with sepia on pale background; spinnerets pale brown.
CARAPACE GRANULATED. With sparse cover of thin, pale, branched setae.
EYES. ALE 0.16; AME 0.20; PLE 0.13; PME 0.13; ALE-AME 0.10; AME-AME 0.07; PLE-PME 0.12; PME-PME 0.12. Clypeus 0.66 high.
CHILUM. 0.21 high, 0.43 wide, with slightly concave inferior side, centre protruding.
STERNUM (Fig. 5B). Shield shaped, granular; 1.42 wide, 1.49 long. Posterior tip shallowly indented; with rounded depressions in front of coxae; no precoxal sclerites.
LEGS. Trochanters III and IV with anterior ventral indentation; dorsal hinged hair on base of T I and II. Cx and ventral side of F granular.
LEG MEASUREMENTS.
FePTMttTotalI2.310.841.751.891.618.40II2.030.841.401.751.407.42III2.030.841.261.610.986.72IV2.590.911.962.871.8910.22
SPINATION.
FPTMtIpl1d2-v1-1 -2disp3 dw4IIpl1d3pl2v1-1 -2disp5 dw4IIIpl1d3rl1pl2rl1pl3d3rl3 v1-2 -2disp6 dw4IVpl1d4rl1pl2rl1pl4d3rl4 v2-2 -2disp8 dw4
ABDOMEN. With PVS in ill-defined group and dispersed thick setae in posterior half (Fig. 5D); with short tubular extension around pedicel, its anterior margin ventrally indented (Fig. 5B).
MALE PALP (Figs 4 C–F, 5E–F, 7A–B). T with strong, slightly sinuous, downcurved, blunt RTA; cymbial fold as long as half the cymbium’s length; embolus with large base, originating at an angle of 270° to longitudinal axis, halfway with short, flat bifurcation, distal end corkscrew shaped; MA hatchet-shaped with short ventral tooth at base; C with short, sharp, flat, distal protrusion, pointing prolaterad.
Female (RBINS IG 33.889/003)
TOTAL LENGTH. 7.31. Carapace: length 3.05, width 2.06, height 1.42.
COLOUR (Fig. 6 A–C). Carapace uniform dark brown; chelicerae medium brown with pale tip; chilum medium brown. Endites and labium medium brown with pale frontal margin. Sternum uniform medium brown. Legs I: Cx medium brown, Tr pale brown, F dark brown, remainder yellowish orange; palp medium brown except T yellowish orange. Abdomen: dorsum with two ill-defined reniform spots touching in front, without scutum overlay, followed by small, faint, pale patches and large patch in front of spinnerets; venter with sclerotized band in front of epigastric furrow, without tubular extension, remainder marbled with sepia on pale background, spinnerets pale yellow.
CARAPACE. Rugose with sparse cover of branched setae.
EYES. ALE 0.18; AME 0.18; PLE 0.15; PME 0.13; ALE-AME 0.15; AME-AME 0.07; PLE-PME 0.23; PME-PME 0.08. Clypeus 0.51 high.
CHILUM. A single sclerite 0.12 high, 0.33 wide, with central protrusion.
STERNUM (Fig. 6B). Shield shaped; 1.28 wide, 1.35 long. Posterior tip rounded; with rounded depressions in front of coxae; no precoxal sclerites.
LEGS. Tr III and IV posteriorly indented. Hinged hair on T I and II.
SPINATION.
FPTMtIpl1d3-v1-1 -1-2disp3 dw4IIpl1d3pl2v1-1 -1-2disp5 dw4IIIpl1d3rl1pl2rl1pl4d3rl3 v1-2 -2disp5 dw4IVpl1d3rl1pl2rl1pl4d3rl4 v1-2 -2disp8 dw4
LEG MEASUREMENTS.
FePTMttTotalI1.890.771.471.401.266.79II1.610.771.121.331.055.88III1.680.700.911.611.055.95IV2.100.911.752.451.408.61
FEMALE PALP. Tarsus with toothed claw turned inward over 30°, eight dispersed spines; tibia with short, sinuous prolateral spine (Fig. 6F).
ABDOMEN. Venter without PVS or thick setae (Fig. 6C).
EPIGYNE (Figs 6 D-E, 9D). With wide rectangular plate, concave in front, posterior margin convex and thickened in the middle. Copulatory ducts with few whorls, ending in poorly delimited spermathecae near posterior end.
Variation
The abdominal venter of the female specimen from Ban Tathot (17.62361 °N, 105.1466 °E) is not marbled but has three dark, faint longitudinal lines.
Distribution
Known from lowland forest in Cambodia and Laos; all localities are along the Mekong river (Fig. 11).
description: Published as part of Jocqué, Rudy, Jocque, Merlijn, Stock, Willem, Rin, Naroeun & Henrard, Arnaud, 2019, The new Southeast Asian genus Cambonilla gen. nov. (Zodariidae, Araneae): ' bis repetita placent', pp. 1-24 in European Journal of Taxonomy 501 on pages 12-17, DOI: 10.5852/ejt.2019.501, http://zenodo.org/record/2578299