Antrops diversipennis KITS & MARSHALL 2013, sp. nov.
Creators: name: KITS, JOEL H.
name: MARSHALL, STEPHEN A.
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Sphaeroceridae
Antrops
Antrops diversipennis

description: Antrops diversipennis sp. nov.   
(Figs. 34, 37, 201)  
 Description. Head black, face, clypeus, prementum, and maxillary palp dark brown. Covered with microtomentum except large spots lateral to ocelli and ventral half of face. Antenna orange to dark brown. Ocellar bristles about 1 ocellus width anterior to median ocellus. Anterior genal bristle about 0.3X length of vibrissa, subvibrissal bristle barely strengthened, about 0.25X length of vibrissa.   
Thorax black, covered with microtomentum except proepisternum, ventromedial patch on anepisternum, and small patch behind fore coxa. Halter whitish, reduced in brachypterous specimens. Acrostichal bristles in 2 rows. Presutural dorsocentral bristle sometimes barely differentiated from surrounding setae.  
Legs black with orange at joints, coxae 1 and 3, trochanters, and femur 1 orange. Tarsi dark brown. Tibia 2 with row of anterodorsal, 1 anteroventral, 1 posteroventral, and 5 subapical bristles. Tibia 3 with 1 strong anteroventral, 3 ventroapical bristles.  
 Wing pale brown, with whitish spots on crossveins r-m, bm-cu, dm-cu, and 2 on vein R 4+5 distal to r-m. Wing length variable, from less than 0.25X length of abdomen to slightly longer than abdomen.   
Abdomen with tergites and sternites black, heavily sclerotized in both sexes, covered with microtomentum. Pleural setae short.  
Male postabdomen: Sternite 5 with lateral margins rounded, posterior margin deeply emarginated, anterior apodeme slightly narrower and about 0.75X length of external portion (Fig. 37). Surstylus scooped, fairly narrow. Pregonite small, distinct. Postgonite with lobes well-separated, anterior lobe broad and rounded, posterior lobe narrower, somewhat pointed. Basiphallus with very broad, short epiphallus, narrow, long preepiphallus. Distiphallus with spinose dorsal tube, flanked by a single spike-like sclerite, with ventral lateral clear sacs (Fig. 34).  
Female postabdomen: Tergites 6–7 and sternites 6–7 slightly wider than long, covered with microtomentum, weakly sclerotized medially except sternite 6. Tergites with 3 posterior strips, sternites with 2 posterior strips. Tergite 8 approximately square, with anterior margin shallowly emarginated, covered with microtomentum except anterior margin. Epiproct with long anterior arms, 1 pair of setae, distal portion with microtomentum. Cerci thin. Sclerites of sternite 8 subtriangular, distal two-thirds covered with microtomentum, nearly meeting tergite 8 along lateral margins. Hypoproct subrectangular, with an unsclerotized hole medially near anterior margin, distal half covered with microtomentum. Spermathecae barrel-shaped, slightly longer than wide, with long apical invagination, sclerotized duct about 1.2X as long as spermathecae.  
 Type material. Holotype ♂: ECUADOR: Napo: Lago Papallacta, nr., 3400 m, forest above lake, pans/ dung, 4–8 Nov 1999, S.A. Marshall, debu00135643 (QCAZ). Paratypes: same data as holotype (10 ♂, 5 ♀, DEBU, QCAZ); Lago Papallacta, 3400 m, elfin forest, leaf litter, 6 Nov 1999, R. Anderson (3 ♂, 2 ♀, DEBU); Quito – Baeza road, 4000 m, open páramo, pan traps, 10–18 Feb 1983, L. Masner (1 ♂, 2 ♀, DEBU); as above but low páramo, 18–23 Feb 1983 (1 ♂, 2 ♀, DEBU); Pichincha: Latacunga, 45km NNE, 3700 m, shrub grass páramo, carrion traps, 19–25 Jul 1985, S. & J. Peck (1 ♀, DEBU); Quito – Baeza pass, 6 km W, pans/dung, 4–8 Nov 1999, S.A. Marshall (2 ♀, DEBU).   
 Comments. The species name refers to the variable wing length within this species. While there are many brachypterous or apterous species of Archiborborinae, and a few individuals of some normally macropterous species have been found with somewhat reduced wings, the range of variation in wing length in this species is unique. Without field observations, it is unclear whether or not macropterous individuals can fly. Longer-winged individuals resemble A. aurantifemur, but the microtomentum on the scutellum and scutum easily identifies this species.
description: Published as part of KITS, JOEL H. & MARSHALL, STEPHEN A., 2013, 
  Generic classification of the Archiborborinae (Diptera: Sphaeroceridae), with a revision of  Antrops  Enderlein,  Coloantrops  gen. nov.,  Maculantrops  gen. nov.,  Photoantrops  gen. nov., and  Poecilantrops  gen. nov.  , pp. 1-113 in Zootaxa 3704 (1) on pages 27-28, DOI: 10.11646/zootaxa.3704.1.1, http://zenodo.org/record/6507447