Scydmaenozila latipennis Lea, comb. n.
Creators: name: Jałoszyński, Paweł
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Scydmaenozila
Scydmaenozila latipennis

description: Scydmaenozila latipennis (Lea) comb. n.
(Figs. 73, 78–79, 83–84, 91–92, 195 b)
Phagonophana latipennis Lea, 1910: 188.
Syndicus latipennis (Lea); implied new combination ex Phagonophana, which was synonymized with Syndicus by Franz, 1971b.
Horaeomorphus latipennis (Lea); Franz, 1975: 146, Fig. 124.
Type material studied. Lectotype (here designated): AUSTRALIA: ♂: seven labels (Figs. 78–79): " latipennis / Lea TYPE / Bridgetown" [thick mounting card on which originally four syntypes were mounted; white, handwritten]; " latipennis / Lea, Type / Bridgetown" [white, handwritten and printed]; "13794 / Phagonophana / latipennis Lea / W. Australia / TYPE " [white, handwritten, " TYPE " in red ink], " SAMA Database / No. 25- 037011" [white, printed], " SAMA Digital Image / 14. 8. 2013 " [green, printed and handwritten], " PHAGONOPHANA / latipennis Lea, 1910 / det. P. Jałoszyński, 2014 / LECTOTYPUS " [white, printed], " SCYDMAENOZILA / latipennis (Lea, 1910) / det. P. Jałoszyński, 2014" [white, printed] (SAM). Paralectotypes (7 exx.): 2 ♀♀ originally mounted on one card with the lectotype, remounted during present study and labeled with white, printed copies of the lectotype labels (SAM); 1 ♂, originally mounted on one card with the lectotype, remounted by Franz, three labels: "Bridgetown" [white, handwritten], " Horaeomorphus / latipennis / Lea / det. H. Franz" [white, handwritten and printed], "S. Aust. Museum / specimen" [orange, printed] (SAM); 1 ♀, four labels: " Australia / 1910–172." [white, printed, first line underlined in red], "Co-type" [circular, white with yellow margin, printed], " Phagonophana / latipennis Lea / W. Aust. / Co-type Lea" [white, handwritten and printed], "♀" [white, printed] (NHM); 1 ♂, 1 ♀ (mounted on separate cards but on one pin): six labels: "Bridgetown / W.A. Lea" [white, printed], " Phagonophana / latipennis Lea / W. Australia / Co-type" [white, handwritten, "Co-type" in red ink], "On permanent loan from / MACLEAY MUSEUM / University of Sydney" [white, printed], " PARATYPE " [blue, printed], " ANIC Database No. / 25 062435 " [white, printed], "Aust. Nat. / Ins. Coll." [green, printed] (ANIC); 1 ♀, seven labels: "WESTERN AUSTRALIA " [white, handwritten], "E. W. Ferguson / Collection" [white, printed], " Phagonophana / latipennis Lea / W. Australia / Co-type" [white, handwritten, "Co-type" in red ink], " PARATYPE " [blue, printed], " PARATYPE " [grey, printed], " ANIC Database No. / 25 062434 " [white, printed], " ANIC / specimen" [green, printed] (ANIC). All paralectotypes additionally with two white, printed labels: " PHAGONOPHANA / latipennis Lea, 1910 / det. P. Jałoszyński, 2014 / PARALECTOTYPUS ", and " SCYDMAENOZILA / latipennis (Lea, 1910) / det. P. Jałoszyński, 2014".
Additional material (4 exx.). 2 ♂♂, Western Australia, Millbrook, W Millstream Road - Millbrook Nature Reserve, 26.iv.2001, " JVAP #31 pitfall" (ANIC, cPJ); 1 ♂, 1 ♀, Western Australia, side track off Pemberton Northcliffe Road, sifted litter, 11.x. 2010, 112 m, leg. K. Will (ANIC, cPJ).
Revised diagnosis. Male and female: body uniformly brown. Male: aedeagus stout, in ventral view with distinctly emarginate apex.
Redescription. Body of male (Fig. 73) strongly convex, elongate and slender, with long appendages, BL 2.80–2.95 mm (mean 2.87 mm); cuticle glossy, uniformly dark brown with slightly reddish hue, vestiture reddishbrown.
Head (Fig. 73) broadest at eyes, HL 0.48–0.50 mm (mean 0.50 mm), HW 0.55–0.60 mm (mean 0.58 mm); tempora longer than eyes, strongly curved posteromesally; vertex weakly convex; frons subtrapezoidal. Eyes large and strongly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine, separated by spaces 1.5– 2 x as wide as diameters of punctures; setae on frons and anterior portion of vertex short, sparse and suberect, tempora and posterior portion of vertex densely covered with long bristles directed posteriorly and posterolaterally. Antennae (Fig. 73) slender, AnL 1.75–1.90 mm (mean 1.83 mm); all antennomeres distinctly elongate, XI slightly shorter than IX–X together, 2.5x as long as broad, pointed.
Pronotum (Fig. 73) in dorsal view strongly elongate, broadest near anterior third, PL 0.78–0.85 mm, (mean 0.80 mm), PW 0.58–0.63 mm (mean 0.60 mm); anterior and lateral margins in anterior half rounded; sides slightly concave in posterior third; posterior margin distinctly bisinuate; base of pronotum with two pairs of large and deep pits, additionally with a pair of shallow but distinct impressions located dorsolaterally. Punctures on pronotal disc fine and inconspicuous; entire dorsal and lateral surface of pronotum covered with moderately dense bristles, dorsally with admixture of thin setae.
Elytra (Fig. 73) slightly rhomboidal, broadest in or slightly behind middle, EL 1.50–1.63 mm (mean 1.57 mm), EW 1.13–1.23 mm (mean 1.17 mm), EI 1.30–1.40 (mean 1.35); basal impressions short but distinct; elytral apices separately rounded. Punctures on elytral disc fine but slightly more distinct than those on pronotum, separated by spaces 2– 3 x as wide as diameters of punctures; setae long, curved, sparse and erect. Hind wings well developed, about twice as long as elytra.
Legs (Fig. 73) long and slender, without modifications.
Aedeagus (Figs. 83–84, 91–92) elongate but relatively stout; AeL 0.53 mm; in ventral view apex not demarcated, with distinctly emarginate apical margin; internal armature complicated, with submedian complex of elongate and darkly sclerotized structures and subapical finely denticulate lateral lobes of internal sac; parameres long and slender, apices barely noticeably broadened.
Female. Similar to male, on average slightly larger, but body lengths and proportions of body parts overlap with those measured in males, distinguished by antennomere VIII about as long as broad and antennomeres IX–X slightly transverse. BL 2.85–3.05 mm (mean 2.95 mm); HL 0.48–0.50 mm (mean 0.50 mm), HW 0.58–0.60 mm (mean 0.58 mm), AnL 1.58–1.75 mm (mean 1.69 mm); PL 0.78–0.85 mm (mean 0.81 mm), PW 0.58–0.63 mm (mean 0.60 mm); EL 1.55–1.70 mm (mean 1.64 mm), EW 1.13–1.23 mm (mean 1.18 mm), EI 1.38–1.42 (mean 1.39).
Distribution (Fig. 195 b). South-western Australia: Western Australia.
Remarks. Franz (1975) studied four specimens originally mounted on a single card, on which one male was annotated as "TY" (Fig. 78). Franz treated this male as a holotype and remaining specimens as paratypes, but studied the aedeagus only in a single remounted " paratype " male (Franz 1975). In fact, Lea (1910) clearly stated that he collected "numerous specimens obtained under a log on sandy ground in the company of ants", and in his description none of them is treated as a type. Therefore, four specimens preserved at SAM, one at NHM and three at ANIC, all identifiable as coming from the original type series, are syntypes. The male originally mounted together with three other specimens on one card and annotated as "TY" is here designated as lectotype, to provide a name-bearing type for Phagonophana latipennis. Three more specimens labeled as Phagonophana latipennis with the same hand but not belonging to the type series (collected in different localities) (ANIC) belong to other species: two are designated here as the holotype and paratype of Scydmaenozila inexpectata sp. n. (from the Ferguson Collection), and one is a Syndicus (from the Ropes Collection).
Scydmaenozila latipennis is similar to Scydmaenozila mallacootana sp. n. and Scydmaenozila inexpectata sp. n.; it clearly differs from these species in the broader elytra in relation to the pronotum (PW/EW 0.48–0.56 vs. 0.60 vs. 0.58–0.61, respectively), and in the structures of the aedeagus.
description: Published as part of Jałoszyński, Paweł, 2014, ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae), pp. 1-76 in Zootaxa 3828 (1) on pages 33-36, DOI: 10.11646/zootaxa.3828.1.1, http://zenodo.org/record/286513