Schraderiellus Rider 1998
Creators: name: Roell, Talita
name: Campos, Luiz Alexandre
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Pentatomidae
Schraderiellus

description: Schraderiellus Rider, 1998
Schraderia Ruckes, 1959; Schrader, 1960: 505; Rolston, 1992: 04–10
Schraderiellus Rider, 1998: 505; Campos & Grazia, 2006: 151–161
Type species: Schraderia hughesae Ruckes, 1959, by original designation.
Description. Body oval, general color dark brown to black, some species with yellow spots; punctures confluent, sometimes forming wrinkles. Female larger than male.
Head triangular, wider than long, declivous. Mandibular plates slightly longer than clypeus, apices acute and declivous, margins reflexed and sinuate midway between base and apex, ending subapically in a very small toothlike projection; margin sinuous between the projection and the apex of each mandibular plate. Wrinkles from posterior margin of head to apex of clypeus and mandibular plates, subparallel on clypeus and diagonal on mandibular plates. Eyes large, ocelli yellow to red. Antennae five-segmented, setose, dark brown with light bands between the segments; segment Iexceeding the apex of the head and slightly wider than the remaining segments, these progressively longer from II to V; proportion: I≥II<III<IV<V. Antennal tubercles each with lateral spine visible from above. Bucculae subparallel, evanescent posteriorly, elevated anteriorly in an acute tooth-like projection. Labium long, reaching urosternites VI to VII; first segment the shortest, reaching prosternum, second segment the longest attaining or surpassing the mesocoxae; insertion of first segment before the middle of bucculae; proportion: I<II>III<IV.
Pronotum trapezoidal, about twice as wide as long, anterior half declivous; anterior margin concave mesially, with a submarginal sulcus. Punctures confluent forming shallow wrinkles. Cicatrices flat with less punctures. Anterolateral angles of pronotum each with a small acute projection laterally directed, slightly exceeding the lateral margin of eyes. Anterolateral margins rectilinear; posterior margin sinuate. Humeri obtuse, not produced. Posterolateral angles obtuse. Scutellum longer than wide at base, reaching connexival segment VI; punctures denser and sometimes confluent near lateral margins. Foveae at basal angles black and shallow. Corium as long as scutellum, apical margin sinuate reaching connexival segment VI; punctures sparse. Membrane reaching or slightly surpassing apex of abdomen, with ten to twelve subparallel veins not forming cells. Prosternum sulcate, meso- and metasternum flat, not punctured, with median longitudinal carina. Mesopleural evaporatorium on posterior margin of mesopleurae; metapleural evaporatorium occupying half the width of metapleura. Peritreme spout, reaching more than half of metapleural evaporatorium. Tibiae sulcate laterally and dorsally, dorsal surface of apical tarsomere of hind legs largely depressed in females, convex in males (Fig. 4).
Connexivum exposed, with sparse punctures. Spiracles black, circular. Trichobothria lateral to imaginary line tangent to spiracles; trichobothrial tubercles yellowish.
Male. Pygophore subglobular; exposed surface brown to black. Posterolateral angles divergent or subparallel, concolor with pygophore or yellowish, posteriorly depressed, apices obtuse, inner surface yellowish (Fig. 5, pa). Base of posterolateral angles, at the lateral limits of ventral rim, bearing 1+1 projections with a tuft of setae (Fig. 5, p 2). Dorsal and ventral rims moderately sinuate, median third concave (Fig. 5, dr, vr). Inferior layer of ventral rim U-shaped (Fig. 5, il). Surface between inferior and superior layers of ventral rim excavated and wrinkled. Segment Xsubcylindrical surpassing posteriorly the inferior layer of ventral rim, with fringe of setae at lateral and apical margins; basal half membranous and apical half sclerotized, the latter visible from below; anal opening apical, genital opening as a ventral longitudinal crevice on apical half of segment X; ventral portion sclerotized (Fig. 5, X). Parameres lateral to segment X, subparallel to posterolateral angles (Fig. 5, par). Phalloteca globose; vesica sinuous covering ductus seminis distalis throughout its length (Fig. 7, ph, v, ds).
Female. Gonocoxites VIII oval, as wide as long, posterior margin subrectilinear to sinuous, covering the base of laterotergites IX; surface irregular with a shallow lateral depression. Posterolateral angles laminar over the base of laterotergites VIII; sutural margins juxtaposed (Fig. 8, gcVIII). Laterotergites VIII triangular, wider than long, with spiracle at basal angle (Fig. 8, laVIII). Gonocoxites IX diamond shaped, longitudinally sulcate (Fig. 8, gcIX). Laterotergites IX digitiform directed to each other and touching at apex, depressed along median margin (Fig. 8, laIX). Segment Xhidden by laterotergites IX, weakly sclerotized (Fig. 8, X). Gonapophyses IX partly exposed, bent posteriorly, with lateral thickenings following the arms of gonocoxites IX (Fig. 9, gpIX). Ring sclerites placed laterally to the foldings of gonapophyses IX (Fig. 9, rs). Thickening of vaginal intima folded in the center; ventrally subtriangular, laterally subcylindrical. Orificium receptaculi apical in the thickening of vaginal intima (Fig. 9, vi). Basal half of outer duct of vesicular area narrow, not dilatable, apical half evenly dilatable (Fig. 9, od). Medium and inner ducts uniformly wide throughout the length (Fig. 9, md, id). Pars intermedialis three times longer than capsula seminalis, basal third narrow, half the width of apical 2/3 (Fig. 9, pi). Capsula seminalis globose with three processes of variable length, each process as long as or longer than pars intermedialis (Fig. 9, cs). Annular flanges convergent.
Distribution: Nicaragua, Costa Rica, Panama, Colombia, Ecuador.
description: Published as part of Roell, Talita & Campos, Luiz Alexandre, 2018, Phylogeny and revision of Schraderiellus Rider, 1998 with the description of five new species (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini), pp. 47-67 in Zootaxa 4508 (1) on pages 50-55, DOI: 10.11646/zootaxa.4508.1.3, http://zenodo.org/record/4983222