Anyphaena quadrata Rivera-Quiroz & Álvarez-Padilla 2023, sp. nov.
Creators: name: Rivera-Quiroz, F. Andrés
name: Álvarez-Padilla, Fernando
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Anyphaenidae
Anyphaena
Anyphaena quadrata

description: Anyphaena quadrata sp. nov.
urn:lsid:zoobank.org:act: DE0EFD57-6E7A-4854-9B6D-2C919B2AF7D7
Figs 34–35, 52
Differential diagnosis
Males of A. quadrata sp. nov. are differentiated from all those of the pectorosa and pacifica groups by the ventral tegular projection distal section being broad and rectangular, the median apophysis also rectangular, longer than wide (Figs 34E–F, 35A–D). Coxa III tubercle bifurcated, anterior ramification blunt, posterior ramification spine-shaped (Fig. 35E–F). Similar tegular apophyses are found in males of A. zuyelenae (Durán-Barrón et al. 2016: figs 12–14, 16) and A. accentuata (Platnick 1974: figs 63, 70; Dondale & Redner 1982: fig. 336; Brescovit 1997: figs 1–6), but they differ by the hook-shaped median apophysis, the RTA shape and coxal ventral surfaces.
Etymology
The species epithet is derived from the Latin ‘ quadrata ’ (‘square’), referring to the distinctive broad semi-squared shape of the ventral tegular projection.
Material examined
Holotype MEXICO • ♂; Veracruz, Calcahualco, Xamaticpac, Plot I; 19.14172° N, 97.20597° W; alt. 1710 m; 2–11 Oct. 2013; Aracnolab team leg.; oak and tropical wet forest fragment; LUP; CNAN-T01535.
Paratype MEXICO • 1 ♂; Veracruz, Calcahualco, Xamaticpac, Plot II 19.12614° N, 97.06708° W; alt. 1700 m; 2–11 Oct. 2013; Aracnolab team leg.; oak and tropical wet forest fragment; BEAT; CNAN-T01566.
Additional material
MEXICO • 1 ♂; same collection data as for paratype; BEAT; ANYM054 • 1 ♂; same collection data as for preceding; ANYM055.
Description
Male
Total length 4.5. Carapace yellow, with two darker bands over ocular quadrangle, around fovea and clypeus. Sternum surface white, intercoxal triangles present on all legs. Labium yellow, white at tip, longer than wide. Endites white, rectangular, broader at tip (Fig. 34A–C). Chelicerae yellow, paturon without pattern (Fig. 34D), promargin with four teeth, retromargin with eight to nine denticles. Abdomen hirsute, dorsal surface dark yellow, anterior border white, lateral sides with same dark pattern on top and diffusing ventrally, ventral surface white. Tracheal spiracle at center of abdomen (Fig. 34A, C). Distal surface of median apophysis with spine. Prolateral apophysis present. Tibia slightly longer than wide. Ventral branch of median tibial apophysis displaced towards proximal border of tibia (Figs 34E–F, 35A–B, D). Coxa II with short basal tubercle, coxa IV with bent spur (Fig. 35 F-F). Cephalothorax length 2.24, thoracic width 1.76, cephalic width 0.77. Clypeus height 0.07. Eye diameters: AME 0.06, ALE 0.1, PME 0.1, PLE 0.11. Eye interdistances: AME–AME 0.04, AME–ALE 0.02, ALE–PLE 0.09, PME– PME 0.15, PME–PLE 0.07. Femur lengths: I 2.29, II 2.05, III 1.71, IV 2.2. Leg spination: femur I d1- 1-1, p0-2-1, r0-0-1. Tibia I v2-2-0, p1-1-1, r1-1-1. Metatarsus I v2-2-0, p1-1-1, r1-1-1. Femur II d1-1-1, p0-1-1, r0-1-1. Tibia II v2-2-0, p1-1-1, r1-1-1. Metatarsus II v2-2-0, p1-1-1, r1-1-1. Femur III d1-1-1, p0-1-1, r0-1-1. Tibia III v2-2-2, p0-1-1, r0-1-1. Metatarsus III v2-2-2, p1-1-2, r1-1-2. Femur IV d1-1-1, p0-0-1, r0-0-1. Tibia IV v2-2-2, p0-1-1, r0-1-1. Metatarsus IV v2-2-2, p1-1-2, r1-1-2.
Female
Unknown.
Variation
Males (N=4): total length 4.63 (± 0.15), cephalothorax length 2.24 (± 0.04), thoracic width 1.76 (± 0.04), cephalic width 0.81 (± 0.04), femur I 2.37 (± 0.05).
Distribution
Only known from the type locality (Fig. 52).
Natural history
Most specimens collected by beating and direct searching over vegetation.
description: Published as part of Rivera-Quiroz, F. Andrés & Álvarez-Padilla, Fernando, 2023, Integration or minimalism: twenty-one new species of ghost spiders (Anyphaenidae: Anyphaena) from Mexico, pp. 1-94 in European Journal of Taxonomy 865 on pages 56-58, DOI: 10.5852/ejt.2023.865.2097, http://zenodo.org/record/7867448