Ligyrus (Ligyrus) paranaensis López-García & Deloya 2022, new combination
Creators: name: López-García, Margarita M.
name: Deloya, Cuauhtémoc
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Dynastidae
Ligyrus
Ligyrus paranaensis

description: Ligyrus (Ligyrus) paranaensis (López-García & Deloya, 2019) new combination
(Figs. 9E, 15P, 18H, 26E; 36)
Tomarus paranaensis López-García & Deloya, 2019: 133. Original combination.
Male holotype (USNM) “ BRASIL / Halik 1966 / Collection // Halik / det. 1965 // Ligyrus / gibbosus / DeGeer // Ponta Grossa / Parana / 3.II 1965 / Moses / 26438 // Tomarus paranaensis / López-García & Deloya / HOLOTYPE ”
Description. Habitus as in Fig. 26E. Length 15.5–16.1 mm; humeral width 8.7–9.1 mm. Color dark reddish brown. Head: Frons coarsely punctate, area between eyes nearly impunctate. Frontoclypeal region with a carina, narrowly interrupted at middle. Clypeus slightly narrowed towards apex, base 2 times as wide as apex. Clypeal teeth triangular, separated by a tooth diameter. Mandible with 2 apical, acute teeth and a basal, widely rounded lobe. Maxilla subrectangular; galea with 3 dorsal and 2 ventral teeth. Apex of labrum bilobed. Interocular distance 5 times an eye width. Ocular canthus acute, with ventral setae. Antennal club long, 1.5 times longer than antennomeres 2–7. Pronotum: Surface with small, dense punctures. Apical tubercle and fovea absent. Elytra: First interval punctate, punctures similar to the other intervals. Pygidial surface without rugosity (Fig. 9E). Punctures small, evenly distributed. Strongly convex in lateral view. Apex regularly rounded. Legs: Protibia tridentate without a denticle; basal tooth slightly distant from others. Protarsus subcylindrical, not enlarged, inner claw entire. Metatibia strongly narrowed before apex. Apex of metatibia entire, with 17–19 spinules. Female genitalia: Subcoxite subrectangular (2 times wider than long). Coxite subquadrate (as long as wide); surface flat. Subcoxite wider and subequal in length to coxite. Male genitalia: Spiculum gastrale thin, apex truncate. Parameres with a medial tooth on each side; apex widened but ending in an acute, triangular point (Figs. 15P, 18H). Internal sac with lamellar spiny belt long (5 times longer than copulatory lamellae), accessory lamella thin and folded.
Diagnosis. Ligyrus paranaensis and L. burmeisteri are similar in their external morphology, but the metatibia is strongly narrowed before apex and has 17–19 spinules on the apical margin in L. paranaensis, while it has the sides nearly parallel and the apical margin with 25–27 spinules in L. burmeisteri. The parameres have the apex acute in L. paranaensis (Figs. 15P, 18G) and rounded to subquadrate in L. burmeisteri (Fig. 15K; see also fig. 18 in NeitaMoreno and Ratcliffe 2017).
Distribution. It is known only from the type locality, Ponta Grossa (Paraná, Brazil).
Locality records (Fig. 36). 3 examined specimens from USNM and MN/UFRJ. BRAZIL (3). Paraná (3): Ponta Grossa.
Natural history. Specimens known are from the Atlantic Forest biome and they were collected in February.
description: Published as part of López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, pp. 1-119 in Zootaxa 5211 (1) on page 48, DOI: 10.11646/zootaxa.5211.1.1, http://zenodo.org/record/7345007