Landouria pakidulan Nurinsiyah & Neiber & Hausdorf 2019, sp. nov.
Creators: name: Nurinsiyah, Ayu Savitri
name: Neiber, Marco T.
name: Hausdorf, Bernhard
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Camaenidae
Landouria
Landouria pakidulan

description: Landouria pakidulan sp. nov.
urn:lsid:zoobank.org:act: D059D3B1-85D9-4760-B9F2-ABC24A7735ED
Figs 46, 54, 70, 85; Tables 3–4
Landouria ciliocincta – van Benthem Jutting 1941: 310 (in part); 1950: 466 (in part). — Dharma 2015: 8 (in part). (not Möllendorff, 1897)
Diagnosis
Landouria pakidulan sp. nov. is characterized by a whitish, depressed conical shell with a reddish-brown band at the angular periphery and a wide umbilicus, a short penis with a large bulge at the proximal end and a moderately long, almost smooth flagellum, which tapers abruptly near its proximal end.
Etymology
Landouria pakidulan sp. nov. (from the Sundanese and Javanese word ʻpakidulanʼ meaning ʻsouthernʼ) is named after its occurrence in southern Java.
Material examined
Holotype
INDONESIA • holotype (det. anat.) (voucher for L. ciliocincta in van Benthem Jutting 1941, 1950); Central Java, Nusa Kambangan Island, Nature Monument Gligir; 7°44′ S, 108°53′ E; 15.–16 Feb. 1927; P.F. Franck leg.; D = 12.5 mm, H = 6.9 mm; ZMA 417011 a.
Paratypes
INDONESIA – West Java • 2 spec.; Pangandaran, Green Canyon; 7°44′ S, 108°27′ E; MZB 14067 • 1 spec.; Pangandaran Nature Reserve; 7°42′ S, 108°39′ E; MZB 6162. – Central Java • 2 spec. (vouchers for L. ciliocincta in van Benthem Jutting 1941, 1950); same data as for holotype; ZMA 417011.
Description
SHELL (Fig. 85; Tables 3–4). Depressed conical, with 5.25–5.5 slightly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes not observed; tuberculate around umbilicus; whitish with a reddish-brown band at periphery; body whorl angular; aperture oval; upper insertion of peristome slightly descending; peristome expanded, reflexed and thickened; umbilicus eccentric, wide, comprising 24–30% of shell diameter, hardly obscured by columellar edge.
GENITALIA (Figs 46, 70; Table 4). Atrium short; penis short, almost cylindrical, with a large bulge at proximal end, without penis coecum; membranaceous sheath around distal part of penis; penial retractor not observed; epiphallus cylindrical, short; flagellum moderately long, broadest in its middle part, abruptly tapering near its proximal end, almost smooth, without node; vas deferens narrow and long; vagina long; oviduct short, broad, proximally tapering; peduncle of bursa copulatrix subdivided into a broad, abruptly tapering distal part and a narrower, longer proximal part. Right ommatophoral retractor runs between penis and vagina.
Remarks
Landouria pakidulan sp. nov. shares with L. menorehensis sp. nov. and L. ciliocincta the whitish shell with a brown band at the periphery. Landouria pakidulan sp. nov. differs from L. menorehensis sp. nov. in the more depressed shell (D/H 1.70–1.86 vs 1.31–1.51 in L. menorehensis sp. nov.) with an angular body whorl and a wider umbilicus that comprises 24–30% of the shell diameter (vs 16–24% in L. menorehensis sp. nov.). The holotype and only anatomically examined specimen of Landouria pakidulan sp. nov. (Figs 70, 85) differs from L. menorehensis sp. nov. (Figs 71, 88) in the triangular dilation at the proximal end of the penis. More specimens must be examined to corroborate that this is a species specific character and not an individual aberration. Landouria pakidulan sp. nov. differs from L. ciliocincta in the less depressed shell (D/H 1.70–1.86 vs 1.86–2.12 in L. ciliocincta), the shorter penis and epiphallus, and the longer flagellum.
Distribution
Landouria pakidulan sp. nov. is known from southern Java from Pangandaran in West Java and Nusa Kambangan Island in Central Java (Fig. 54).
description: Published as part of Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, pp. 1-73 in European Journal of Taxonomy 526 on pages 62-64, DOI: 10.5852/ejt.2019.526, http://zenodo.org/record/3152217