Clivina schillhammeri Balkenohl 2021, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina schillhammeri

description: <i>Clivina schillhammeri</i> sp. nov. urn:lsid:zoobank.org:act: D0FD6935-0961-4DBE-B9FB-49124FD3486F (Figs 31, 51, 52, 71, 83, 98, 115) ETYMOLOGY. The species is dedicated to Dr. Harald Schillhammer (NHMW) who collected numerous specimens of the species in Laos and in Myanmar and who supported my work over many years. TYPE MATERIAL. Holotype: • &amp;male;, with labels and data: white, black printed “N- LAOS: Prov. leg. Nam Tha Muang Sing, at light 9–13.vi. 1996, 600 m leg. Schillhammer (18) (NHMW). Paratypes: KOREA. • 1 &amp;female;, KOREA: Prov. south Phenan: Pyongyang 4.vi.1970 / Hung. Zool. Exp. I. in Korea. No. 92., leg. Dr. S. Mahunka et Dr. H. Steinmann / <i>Clivina castanea</i> Westw. det. Alex. Dostal 1989 (CADW). JAPAN. • 8 &amp;male;&amp;male;, 8 &amp;female;&amp;female;, Saeki, Oita Prev. 24.vii.1948 Kiushiu. Japan (CADW); • 1 &amp;male;, 1 &amp;female;, same data but 5.viii.1948 (CADW); • 2 &amp;male;&amp;male;, 4 &amp;female;&amp;female;, same data but 11.viii.1948 (CADW); • 5 &amp;male;&amp;male;, 12 &amp;female;&amp;female;, same data but 28.viii.1948 (CADW); • 1 &amp;male;, Kobe, Hyogo. vii.1949 Honshin, Japan (CADW); • 3 &amp;male;&amp;male;, 1 &amp;female;, Fujioka-chô, Tochigi Pref. / Wartarase-yusuichi / 8.ix.2007 T. Suda leg. (CSMT); • 1 &amp;female;, handwritten in black ink: Japon. Lewis / <i>Clivina parryi</i> Putz det. J. Putzeys / printed: Soc. Ent. Belge Coll. PUTZEYS (RBINS); • 1 &amp;male;, 1 &amp;female;, handwritten in black ink and printed: Matsuyama Ehime Pref. 10.vi.1955 / R. I. Sc. N. B. I. G. 22.024 (RBINS); • 1 spec., same data but 30.vi.1954 (RBINS). TAIWAN. • 1 &amp;male;, 1 &amp;female;, TAIWAN, Pingtung County Kenting Liu-fu Hotel N21°57.388 E 120°47.017, 02.viii.2008, at light H. Mendel, MVL Barclay &amp; R. Ewers BMNH(E)2008–85 (BMNH); • 1 spec., TAIWAN, Lanyu Island (Taitung County) Yung Hsin Farm N 22°01.703 E 121°34.642, 29–30.vii.2008, at light H. Mendel &amp; MVL Barclay BMNH(E)2008-85 (BMNH); • 1 &amp;male;, S. Formosa Kagi 20–viii.1907 H. Sauter S.V. No. 11474 (MFNB); • 1 spec., TAIWAN vii.1990 Pingtung Hsien Suchung-Shi / 44 leg. M.L. Jang (NHMW); • 1 spec., Kosempo Formosa H. Sauter 22.v.1912 / <i>Clivina castanea</i> West. det. Ing. Jedli <b>&amp;Ccaron;</b> ka / ex Coll. Jedli <b>&amp;Ccaron;</b> ka Nat. Mus. Prague (NHMP). HONG KONG. • 1 &amp;male;, Hong Kong, China coll: Paul Aston 2–9.x.1912 Pui O, Lantau. H.K. PT in Marsh / <i>Clivina castanea</i> Westwood 1837 Determined D. Fedorenko (CMBB); • 2 &amp;female;&amp;female;, HONGKONG, 1983 Tai Po Kau, 10.viii. downstream leg. Dudgeon (NHMW). CHINA. • 1 &amp;male;, Kiautschau China / <i>C. castanea</i> / COLLECTIO KAREL KULT COLL. A. DOSTAL, 1999 (CADW); • 1 spec., Kiautschau China /? <i>moerens</i> Putz. Det. Fleischer (NHMP); • 1 &amp;female;, China, Chinkiang 32°12’N 119°30’E, 4.i.1919 Eigin Suenson leg. (CADW); • 2 &amp;male;&amp;male;, China, Soochow 31°21’N 120°40’E, 6.iv.1919 Eigin Suenson leg. (CADW); • 2 &amp;female;&amp;female;, China, Shanghai 31°13’N 121°25’E 6.v.1937 Eigin Suenson leg. (CADW); • 1 spec., Zi-ka-wei 16.vi.20 / O. PIEL coll. (CADW); • 1 &amp;male;, 1 &amp;female;, China, Hangchow 30°18’N 120°07’E, 18.x.1920, Eigin Suenson leg. (CADW); • 1 &amp;male;, China, Canton, Pak wan Asi, 15.vi.1910 Mell S. V. (MFNB); • 1 spec., China Canton /? <i>moerens</i> Putz. Det. Fleischer (NHMP); • 1 spec., Tsangen /? <i>moerens</i> Putz. Det. Fleischer (NHMP); • 1 spec., Shaowu – Fukien (500m) J. Klapperich / <i>Clivina castanea</i> West. Dr. Maran det. (NHMP); • 3 spec., KUATUN FUKIEN China meridion. Klapperich / <i>castanea</i> Westw. det. Ing. Jedli <b>&amp;Ccaron;</b> ka / ex Coll. Jedli <b>&amp;Ccaron;</b> ka Nat. Mus. Prague (NHMP); • 1 &amp;female;, Südchina Wuhan 15.viii.1984 Katschok leg. / <i>Clivina castanea</i> Westw. det. Balkenohl 1999 (SMNS); • 1 &amp;male;, CHINA – Guangdong Chebaling Nat. Reserve E. Shaoguan, 300–500 m 25–27.vi.1996 Chr. Häuser leg (SMNS); • 6 &amp;male;&amp;male;, 3 &amp;female;&amp;female;, China, Buong Kä n. of. Foochow – 27°N 119°E, 26.iv.1935 Eigin Suenson leg. (CADW); • 1 &amp;male;, China, Kushan 26°4’N 119°25’E 8.iv.1935 Eigin Suenson leg. (CADW); • 1 spec., KATUN, FUKIEN China 4.iv.1946, (TSCHUNG SEN.) (CADW); • 2 &amp;male;&amp;male;, 1 &amp;female;, China, Kuliang 26°12’N 119°28’E 11.iv.1935 Eigin Suenson leg. (CADW); • 1 &amp;female;, SO China: Fujian Wuyi Gebirge (ca 125 km N Yvanping 200– 400 m 7–22.iv.1988 leg. Görgner &amp; Kleinfeld (CWGP); • 1 &amp;female;, China, Nanping 26°50’N 118°05’E 17.v.1935 Eigin Suenson leg. (CADW); • 1 spec., Kiang-Si T‘ en- gan (MFNB); • 1 spec., same data and ex Coll. Jedli <b>&amp;Ccaron;</b> ka Nat. Mus. Prague (NHMP); • 1 &amp;male;, CHINA, N GUANGXI reg. MIAOERSHAN, S slope, 25–26.vi.1997, 1300–2000 m, Bolm lgt / <i>CLIVINA (Leucocara) castanea</i> Westwood, 1837 P. Bulirsch det. 2016 (NHMB); • 1 spec., Chine centrale SI ‘ Shan 25.v.1931 R.P. Marga / R. Mus. Hist. Nat. Belg.I.G.13.336 / <i>Clivina</i> sp. det. S.W.Nichols 1984 <i>/ Clivina castanea</i> s.l. Westwood, 1837 P. Bulirsch det. 2011 (RBINS); • 1 spec., CHINA, ZHEJIANG Prov., West Tianmu Shan (Mts) reserve, border of secondary mixed forest nr entrance of reserve; at light 30°18´34.5“N, 119°28´46“E, 310 m, 23.vi.–6.vii.2017 J. Hájek &amp; J., R&amp;uring;zi&amp;ccaron;ka leg. (NHMP); • 1 &amp;female;, Kweichou China, Plason (CADW), • 1 &amp;male;, CHINA / Sichuan 103.20el/29.30 NW Mt. Emei 500–1200 m 4–19.v.1989 S. &amp; J. Kolibé&amp;ccaron; leg. / Freiwilliger Museumsverein Basel 1989 (NHMB); • 2 &amp;male;&amp;male;, 1 &amp;female;, 10 specs., CHINA (Yunnan) Lincang Pref., Mekong Valley, small creek cleft 38 km SSE Lincang. 854 m 23°33’13.2”N / 100°09’56.8”E (field edge in small cleft, in loamy soil) 11.ix.2009 D.W. Wrase [44] / Coll. Wrase Berlin (CWGP, CMBB); • 1 &amp;female;, same data but (NKME); • 1 spec., China, Prov. Yunnan Gbg. B. Mengtze / <i>Clivina castanea</i> Westw. H.E. Andrewes det. / ex Coll. Jedli <b>&amp;Ccaron;</b> ka Nat. Mus. Prague (NHMP); • 1 spec., CHINA: YUNNAN Prov. Tongbiguan vill. 24–26.vi.2016 24°36.7’N, 97°39.4’E, 1340 m at light in village near river J. Hájek &amp; J., R&amp;uring;zi&amp;ccaron;ka leg. / <i>CLIVINA castanea</i> Westwood, 1837, P. Bulirsch det. 2018 (NHMP); • 1 spec., Tsushima Rost 1919 / ex Coll. Jedli <b>&amp;Ccaron;</b> ka Nat. Mus. Prague (NHMP); • 3 historic specs. with black framed and printed labels, one without abdomen: CHINA (NHMB); • 1 historic spec. with double framed and handwritten label: Lilong. China (NHMB); • 1 &amp;male;, Vallis flumin. Soling-ho, Yun. / <i>castanea</i> Westw. det. Ing. Jedli&amp;ccaron;ka / A 1975 / WIEN/COLLECTIO Dr. R. WEBER; 1994 (CADW); • 1 spec., Tchenkiang / Lou bou 10.vii.1923 / O. PIEL, coll. (CADW); • 1 spec., Kiang-si Kiu-kiang (MFNB); • 1 spec., Kiang-si Chang-tsin-cheng (MFNB); • 1 spec., Tientsin 16, IX. 29 / <i>castanea</i> Westw. det. Ing. Jedli&amp;ccaron;ka (NHMP). VIETNAM. NORTH VIETNAM. • 1 &amp;male;, Pa-kha 1914 / Indo Chine Coll. Dussault (NHMB); • 1 spec., VIETNAM SaPa – 1530 m 25.v–9.vi.1991 Strnad Jan leg. (CMBB); • 1 spec., N-VIETNAM 1991 75 km NW Hanoi Tam Dao 16–23. v. leg. J. Strnad (NHMW); • 1 &amp;female;, Annam Phuc Son (RBINS); • 2 specs., Annam Phuc son Nov. Dez. H. Fruhstorfer (MFNB); • 2 &amp;male;&amp;male;, Annam Phuc-Son Nov. Dez. H. Fruhstorfer (MFNB); • 1 spec., N-Vietnam Vinh Phu pr. Tam Dao 1050–1200 m 21°27.18N 105°38.58E 2–6.vi.1999, leg. Ahrens (CJSR); • 3 specs., Coll. I.R.Sc.N.B. VIETNAM: Phia Den 08.vii..2010, Light Trap I.G. 31.668 leg. J. Constant &amp; P. Limbourg (RBINS); • 1 spec. Hanoi 1923/25 / Indo Chine Coll. Dussault / <i>Clivina castanea</i> WESTW. det. Balkenohl xi.1991 (NHMB); • 1 &amp;male;, 10 specs., N-Vietnam Lao Chai pr. Bac Ha env. 980–1000 m 22°22,05N 104°17.32E 28–30.v.1999, leg. Ahrens (CJSR); • 1 &amp;female;, Tonkin Hanoi / <i>castanea</i> Westw. det. Ing. Jedli&amp;ccaron;ka / <i>castanea</i> Hoppe [sic] det. K. Kult / COLLECTIO KAREL KULT COLL. A. DOSTAL, 1999 (CADW); • 2 &amp;male;&amp;male;, TONKIN Hoa-Binh, vii.1940, leg. A. de Cooman (CADW); • 1 spec., same data but iii.1939 (CADW); • 2 specs., same data but iii.1931 (CADW); • 1 &amp;male;, TONKIN Hoa-Binh leg A. de Cooman / No 3 (CADW); • 1 spec., Hoah Binh Tonkin / <i>Clivina castanea</i> WESTW. det. M. Baehr’ 05 (CMBM); • 1 spec., Vietnam, Hoa binh 4–7.vi.1986 Ha son binh prov. Jan Horák leg. / <i>CLIVINA castanea</i> group P. Bulirsch det. 2009 (NHMP); • 1 spec., VIETNAM-N, Tuyen Quang pr. SE-E env. of Na Hang 22°17’30”- 22°30’N 105°26’ –28’E, 200–700 m L. Dembický leg., 1–12.v.2010 / <i>CLIVINA (Leucocara</i>) <i>castanea</i> Westwood, 1837 P. Bulirsch det. 2016 (NHMB); • 1 &amp;male;, N VIETNAM (Tonkin) pr. Hoang Lien Son SAPA 11–15.v.1990 Vít. Kubán leg. (CADW); • 1 &amp;female;, Kansou mer. Hoei-sien. (MFNB); • 2 specs., VIETNAM N, Cuc Phuong 2–11.v.1991 Strand Jan leg. (NHMB); • 1 spec., N-VIETNAM Ninh Binh Cuc Phuong NP, 270 m N 20°17’57’’, E 105°40’05’’, 04–09.v.2017. A. Weigel KL / collection NATURKUNDE-MUSEUM ERFURT / <i>CLIVINA (Leucocara) castanea</i> Westwood, 1837 P. Bulirsch det. 2018 (NKME); • 1 spec, same data but 25.iv. 2012, 190 m, N20°14’24’’, E105°42’53’’ leg. A. Weigel Li.-Fa. / <i>CLIVINA (Leucocara) castanea</i> Westwood, 1837 P. Bulirsch det. 2014 (NKME); • 2 specs., N-VIETNAM Cao Bang Pr. vic. Vin Den, Nui Pia Oac Nature Res., 6–10.v.2013, 22°33’53’’N, 105°52’53’’E 900–1300 m leg. A. Weigel / collection NATURKUNDE-MUSEUM ERFURT / <i>CLIVINA (Leucocara) castanea</i> Westwood, 1837 P. Bulirsch det. 2014 (NKME); • 1 spec., N-VIETNAM Thai Nguyen Pr. vic. Ngoc Thanh, vic. Me Linh (IEBR station), 12.v.2012, 21°22‘3’’N, 105°42’44’’E 60–80 m leg. A. Weigel / collection NATURKUNDE-MUSEUM ERFURT / <i>CLIVINA (Leucocara) castanea</i> Westwood, 1837 P. Bulirsch det. 2014 (NKME); • 3 specs., VIETNAM, N (Na Hang) 160 km NNW Hanoi NE env. of Na Hang 28.v.–10.vi.1996, LF, 150–200 m, NN, leg. A. Napolov &amp; I. Roma / collection NATURKUNDE-MUSEUM ERFURT / <i>CLIVINA (Leucocara) castanea</i> Westwood, 1837 P. Bulirsch det. 2012 (NKME); • 1 &amp;female;, (305) Annam-Vinh 1910 / Indo Chine Coll. Dussault (NHMB); • 1 spec., VIETNAM: Me linh Light Trap 20–24.viii.2010, leg J. Constant &amp; P. Limbourg I.G.31.668 (RBINS); • 1 &amp;female;, same data as before (CMBM). SOUTH VIETNAM. • 1 &amp;male; [immature], VIETNAM: Buon-loi 40 km Nof Ankhe Prof. Gialai Kontum 15.vi.1985 (SMNS); • 2 specs., S-VIETNAM 40km NW An Khe Buon Luoi, 620– 750 m / 14°10’N 108°30’E 28.iii.–12.iv.1995 Pacholatko &amp; Dembicky (NHMW); • 1 &amp;male;, 4 specs., S-VIETNAM Nam Cat Tien Nat. Park 1–15.v.1994 Pacholatko &amp; Dembicky (NHMW); • 1 spec., VIETNAM Buon-loi 40 km Nof Ankhe Prov. Gia-lai Kontum 15.vi.1985 (CMBB). THAILAND. • 1 &amp;female;, THAILAND bor. CHIANG DAO env. 17–19.vi.1995 leg. SNIZEK M. (CMBB); • 2 &amp;female;&amp;female;, NO-THAILAND Nan – Pha Khab 11-16.v.1993 leg. Pacholatko (NHMW); • 5 &amp;female;&amp;female;, NO – THAILAND 1993 NAN – PHA 11-16. v. leg PACHOLATKO (CADW); • 1 &amp;male;, O – THAILAND 1990 Ko Chang (11) am Licht leg. Jäch 8– 12.xii. (NHMW); • 1 spec. THAIL. 9.xii.1990 Ko Chang NP, leg. Forster (NHMW); • 1 spec., E Thailand, 5–13.v.1998, Chanthaburi Dist. Khao Soi Dao, J. Fyman lgt (NHMP); • 1 spec. Nord-Thailand Chieng Mai Dr. Jaratpun Sa-nguansermsri leg ix.1981 (MFNB); • 1 &amp;male;, NW THAILAND, Pai, 500 m, 17– 19.iv.2004, leg. W. SCHAWALLER (SMNS); • 1 &amp;female;, NW-THAILAND, CW Mae Hong Son Pai Thapai Hot Springs 25.iv.2004, R. Grimm (SMNS); • 1 &amp;male;, THAILAND PAI env. 1– 14.v.2001, J. Hromádka leg. (CADW); • 1 &amp;male;, 1 &amp;female;, NW Thailand MAE HONG SON 1992 Nupa-ah 7– 9.v, Strnad Jan leg. (CADW); • 1 &amp;male;, THAILAND: Changwat Mae Hong Son. Ban Huai Kha 5.i. 1989, 300 m leg. TRAUTNER &amp; GEIGENMÜLLER (SMNS); • 1 spec., Thai, 26–28.v.1991, PALONG 750 m 19°55’N 99°06’E Vít Kubá&amp;ncaron; leg. / Thailand ’91 “Thanon Thong Chai” D. Král &amp; V. Kubá&amp;ncaron; (NHMB); • 1 spec., Coll. I.R.Sc.N.B. THAILAND (loei) Na-Haeo (field res stat) 15–19.v.2003 Light trap, leg. J. Constant, K. Smets &amp; P. Grootaert (RBINS); • 2 specs., THAI, 9–13.iv.1991, THIMONGHTA 350 m, 15°02’N 98°35’E Vít Kubá&amp;ncaron; leg. / Thailand ’91 “Thanon Thong Chai” D. Král &amp; V. Kubá&amp;ncaron; (NHMB); • 1 spec., Thai 18–24.iv.1991 LANSANG nat. park 16°48’N 98°57’E leg. D. Král 500 m / Thailand’91 “Thanon Thong Chai” D. Král &amp; V. Kubá&amp;ncaron; / <i>Clivina castanea</i> WESTW. det. Balkenohl, 2007 (NHMB); • 1 &amp;male;, THAI 20.v.1993, NAKHON THAI Pacholatko &amp; Dembicky leg. (CADW); • 1 &amp;male;, 1 &amp;female;, 1 spec., THAILAND, S 40 km NE OF BETONG, LAKE BANG LANG (KG.TO) 12–23.vi. 2007, 350 m Z. LINEK leg. (CADW); • 1 spec., S-THAILAND Taku Pa, 20.xi.1996, leg. J. REJSEK (SMNS). LAOS. • 1 &amp;female;, same data as holotype (NHMW); • 12 specs., LAOS, Phongsaly prov., PHONGSALY env., 6–17.v.2004,~ 1500 m, 21°41’N 102°06–08’E, M. Brancucci leg. / <i>Clivina</i> (<i>Leucocara</i>) <i>castanea</i> Westwood, 1837 P. Bulirsch det. 2016 (NHMB); • 1 &amp;female;, LAOS, PHONGSALI PROV. MEUNG KHOA DISTR. BAN HOUAYWANG 10–17.vi.2010 leg. SASA KAMPHILAVONG (CADW); • 1 &amp;female;, LAOS-Houaphan prov. 38 km Sof Sam Neua Saleui 9–22.v.2009 Martinu leg. 1350–1900 m (CADW); • 3 &amp;female;&amp;female;, NE-LAOS Hua Phan prov. Ban Saleui, Phou Pan Mt. 20°12’N 104°01’E 23.iv.–15.v.2008, 1500–1900 m, leg. C. Holzschuh (CADW); • 1 &amp;female;, same data but 11.iv.–15.v.2012, 1300–1900 m / collection NATURKUNDE-MUSEUM ERFURT / <i>Clivina</i> (<i>Leucocara</i>) <i>castanea</i> Westwood, 1837 P. Bulirsch det. 2013 (NKME); • 1 &amp;male;, 6 specs., same data but 01–31.v.2011, leg. C. Holzschuh and local collector (NKME); • 4 specs., LAOS: N-VIENTIANE Prov. Vang-Vieng, 300 m N 18°55’23”, E 102°26’55” 10–15.v. &amp; 01–06.vi.2001, JI&amp;Rcaron;I KOLIBÁ&amp;Ccaron; leg. (NHMB); LAOS, Vientiane prov. LAO PAKO env. 200 m 55 km NE Vientiane 19–22.v.2004, J. Bezdĕk leg. / <i>CLIVINA castanea</i> group P.Bulirsch det. 2009 [sic] (NHMP); • 1 &amp;male;, 1 &amp;female;, LAOS, Xieng Khouang Prov., 19°37-8’N 103°20’E, Phonsavan (30 km NE); Phou Sane Mt. 1400–1700 m 10–30.v.2009, D. Hauck leg. (NHMB); • 4 specs., LAOS, Khammouan prov. 17°57 -59’N/ 105°13 -16’E, Nakai- Nam Hheun NPA, BAN NAVANG env., 600–750 m, 18–21.v.2012, / NHMB Basel, Expedition Laos 2012: M. Brancucci, M. Geiser, K. Phanthavong, S. Xayalath / <i>Clivina</i> (<i>Leucocara</i>) <i>castanea</i> Westwood, 1837, P. Bulirsch det. 2016 (NHMB); • 3 specs., LAOS, Bokeo prov., 5 km W Ban Toup, Bokeo Nature reserve, 500–700 m, 20°27 –28’ N/ 100°45’ E, 4–18.v.2011 / NHMB Basel, Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala, &amp; E. Vongphachan / <i>Clivina</i> (<i>Leucocara</i>) <i>castanea</i> Westwood, 1837 P. Bulirsch det. 2016 (NHMB); • 2 specs., LAOS Nam Tha le 9.iv.1918, R. Vitalis de Salvaza / <i>Clivina castanea</i> West. det. R. Vitalis de Salvaza / R.I.Sc.N.B 16.493 (RBINS); • 6 specs., same data but Pak Pha 4.ii.1918 (RBINS); • 1 spec., LAOS Pak Pha 4.ii.1918, R.V.de Salvaza / <i>Clivina castanea</i> Westw., H.E. Andrewes det. / Ex Misc Coleoptera 142 Tribes of the Carabidae Hope Ent. Coll. Oxford (OUMNH); • 1 spec., LAOS Ehalan le 20.xi.1917, R. Vitalis de Salvaza / <i>Clivina castanea</i> West. det. R. Vitalis de Salvaza / R.I.Sc.N.B 16.493 (RBINS); • 1 spec., LAOS Ban Hou le 7.x.1918, R. Vitalis de Salvaza / <i>Clivina castanea</i> West. det. R. Vitalis de Salvaza / R.I.Sc.N.B 16.493 (RBINS); • 1 spec., LAOS Muong Pek le 15.xii.1918, R. Vitalis de Salvaza / <i>Clivina castanea</i> West. det. R. Vitalis de Salvaza / R.I.Sc.N.B 16.493 (RBINS); • 1 spec., LAOS B. Honei Soni le 30.i.1918, R. Vitalis de Salvaza / <i>Clivina castanea</i> West. det. R. Vitalis de Salvaza / R.I.Sc.N.B 16.493 (RBINS); • 1 spec., LAOS Natung le 29.iii.1918, R. Vitalis de Salvaza / <i>Clivina castanea</i> West. det. R. Vitalis de Salvaza / R.I.Sc.N.B 16.493 (RBINS); • 4 specs., LAOS-C: KHAMMOUAN pr., Ban Khoun Ngeun. ~ 200 m; 18°07’N 104°29’E; Pacholátoko leg., 24–29.iv.2001 / <i>Clivina</i> (<i>Leucocara</i>) <i>castanea</i> Westwood, 1837 P. Bulirsch, det. 2016 (NHMB); • 2 specs., LAOS, Salavan prov., 15°42’N / 106°24’E, SALAVAN town, 200 m, 23–24.v.2012, M. Geiser leg. / Collection Naturhistorisches Museum Basel / <i>Clivina</i> (<i>Leucocara</i>) <i>castanea</i> Westwood, 1837, P. Bulirsch det. 2016 (NHMB); • 2 specs., LAOS centr., Kammouan prov. NAKAI env. 4–8.v.1998, Route No 8, alt. 560+ 20 m N 17°42.8, E 105°08.9 (GPS), M. Strba &amp; R. Hergovits (CWGP); • 3 specs., LAOS, Salavan prov., 16°08’N / 106°40 -43’E, Xe Xap NPA, ca. 15 km NE of Ta-oy, BAN DOUB env. 400–1000 m, 25–31.v.2012, / NHMB Basel, Expedition Laos 2012: M. Brancucci, M. Geiser, K. Phanthavong, S. Xayalath / <i>Clivina</i> (<i>Leucocara</i>) <i>castanea</i> Westwood, 1837 P. Bulirsch det. 2016 (NHMB); • 3 specs., LAOS-N (Oudonxai), 1–9.v.2002, ~ 1100 m, 20°45’N 102°09’E, OUDOM XAI (17 km NEE). Vít Kubá&amp;ncaron; leg. / Collection Naturhistorisches Museum Basel / <i>Clivina</i> (<i>Leucocara</i>) <i>castanea</i> Westwood, 1837 P. Bulirsch det. 2016 (NHMB); • 1 spec., LAOS-N(Louangphra-bang), 1–21.v.2002, 19°35’N 101°58’E, THONG KHAN, ~ 750 m, Vít Kubá&amp;ncaron; leg. / Collection Naturhistorisches Museum Basel / <i>Clivina</i> (<i>Leucocara</i>) <i>castanea</i> Westwood, 1837 P. Bulirsch det. 2016 (NHMB); • 1 spec., LAOS, Champasak Prov., 15°02’N 106°31 -34’E, Ban Nong Panouan env. 770–800 m 10– 17.vi.2010, M. Geiser &amp; D. Hauk leg. / coffee plantations, secondary and degrades primary forest; hand collecting, sweeping, beating, coll. at flowering trees and in small water puddles, light trapping (NHMB); • 1 spec., LAO, 26–27.v.2003, Bolikhamxai prov., PAKKADING, ~ 300 m, 18°20’N 104°00’E, Vít Kubá&amp;ncaron; leg. / Collection Naturhistorisches Museum Basel / <i>Clivina</i> (<i>Leucocara</i>) <i>castanea</i> Westwood, 1837 P. Bulirsch det. 2016 (NHMB); •
description: Published as part of <i>Balkenohl, Michael, 2021, Belgian Journal of Entomology, pp. 1-83 in Belgian Journal of Entomology 115</i> on pages 46-53, DOI: 10.5281/zenodo.5645953
description: {"references": ["WESTWOOD J. O., 1837. - A collection of insects collected at Manilla by Mr. Cuming. Proceedings of the Zoological Society, 5: 127 - 130.", "BALKENOHL M., 1999. - New Clivinini from the Oriental region. 2. Clivina rugosofemoralis nov. spec. and Rugiluclivina leonina nov. spec. from Laos (Coleoptera, Carabidae, Scaritinae). Linzer biologische Beitrage, 31: 337 - 344."]}