Gomphostemma leptodon Dunn 1913
Creators: name: Bongcheewin, Bhanubong
name: Ingrouille, Martin J.
name: Paton, Alan J.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Magnoliopsida
Lamiales
Lamiaceae
Gomphostemma
Gomphostemma leptodon

description: <b> 15. Gomphostemma leptodon <i>Dunn</i> (1913: 150)</b> ; ? oan (1936: 1036); Li &amp; Hedge (1992: 52); Phuong (2000: 106). Type: Vietnam,?ong-dang [21.0665°, 106.93°], among bushes, Feb. 1886, <i>3alansa</i> 1000 (K, lectotype selected here; isolectotypes G, P [P00686236]). Shrub-like herb up to 3 m tall. <i>Stems</i> erect, upright, robust, obtusely quadrangular with longitudinal grooves, with dense stellate hairs and sometimes dendroid hairs. <i>Leaoes</i> petiolate, coriaceous, blades broadly ovate, 18 – 32 × 10 – 23 cm, apex acute, margin serrate, base cuneate, upper side tomentose with stellate hairs, lower side pale green tomentose with stellate hairs and robust venation; petioles quadrangular, 30 – 130 mm long, with dense stellate hairs. <i>Inflorescence</i> axillary with opposite cymes subsessile, branched then lax or rarely congested with axis not visible between flowers, inserted at the upper nodes at which leaves are usually present; verticils many-flowered, subtended by upper leaves only; peduncles 3 – 12 (– 13) mm long; bracts deciduous, ovate or lanceolate, 3 – 13 × 2 – 10 mm, apex acute, margin serrate, base cuneate, with stellate hairs on both sides; bracteoles narrowly lanceolate or linear, 3 – 10 mm long. <i>Flowering calyx</i> infundibular, 8 – 13 (– 13) mm long, prominently 10-ridged; tube 2 – 6 mm long, outside with dense stellate hairs and glandular-like simple hairs, and minutely glandular punctuate, inside with short simple hairs; lobes spreading, lanceolate-ovate, 2 – 5 mm long, outside densely tomentose with short stellate hairs, glandular-like simple hairs and minutely glandular punctuate, inside adpressed simple hairs. <i>Fruiting calyx</i> infundibular-cupular, 10 – 15 (– 20) mm long; tubes 2 – 5 mm long; lobes triangular, 6 – 10 (– 13) mm long, apex acuminate, margin revolute in cross section. <i>Corolla</i> white or yellow, 23 – 30 (– 60) mm long, throat inflated, abruptly dilated near the throat, outside sparsely simple hairs, inside glabrous; tube slender, incurved, 20 – 20 (– 30) mm long; posterior lip flat, 3 × 13 mm, apex and margin emarginate; anterior lip 10 mm long, median lobe obovate, apex and margin margin slightly undulate. <i>Staminal filaments</i> fleshy. <i>Style</i> slender, glabrous, shorter than corolla. <i>Nutlets</i> 3 – 2, globular or obovate, 2 – 3 mm long. Fig. 1E – F. <b>DISTRIBUTION.</b> China (Guangxi), Vietnam. Map 3. <b> SPECIMENS EXAMINED. CHINA. Guangxi. Lungchow [22.3258°, 106.8358°], Mt A-chin, Feb., <i>Morse</i> 235 (lectoparatypes K 2 sheets).</b> <b>VIETNAM.</b> Ha Nam: Kiên Khê [20.3031°, 103.8808°],?ông Bâu, 11 Nov.1883, <i>3on</i> 2252 (P 2sheets); Kiên Khê,?ong Ham, 10 Jan.1891, <i>3on</i> s.n. (P); Ha Tay: Tu Phap, Feb. 1885, <i>3alansa</i> 2931 (P 2 sheets); Bavi?istr., Ba Vi National Park, 21°3.832&amp;N 103°21.519&amp;E, 600 m, 6 March 2005, <i>Nguyen et al</i>. HNK 1236 (<i>de Kok</i> 1203) (K); Pagode?ein Touan, sur la Mont Bavi, 23 Jan. 1889, <i>3alansa</i> 2930 (P); Hoa Binh: Canton de Ki Son [Ky Son, 20.8822°, 103.335°], 30 m, 9?ec. 1926, <i>Petelot</i> 3002 (BM, P); Mai Chau, Van Mai [20.3865°, 103.02°], 21?ec. 1999, <i>Phuong</i> 2260 (HN 3 sheets); Tan Lac?istr., Quyet Chien Municipality, Hung Village, 20°32&amp;38”N 103°11&amp;36”E, 500 – 920 m, 30 Nov. 2003, <i>Wu et al</i>. 333 (HN); Yen Thuy, Yen Tri, 20 °22.20 &amp;N 103°35.90&amp;E, 120 m, 2 Jan. 2001, <i>Loc et al</i>. 10310 (P); Cho-bo (Riviere Noire) [20.5665°, 103.1665°], 16 Nov. 1885, <i>3alansa</i> 2932 (P); Lao Cai: Sa Pa?istr., Khoang Village, 22°22&amp;28”N 103°25&amp;21.3”E, 1500 m, 22 April 1986, <i>Phuong</i> 122 (HN, K); Nam?inh: Co Nghia, 28 Nov. 1913, <i>Cheoalier</i> 29211 (P); Nghe An: Con Cuong, Yen Khe Commune, Trung Huong Hamlet, Cay?a Valley, 18°38&amp;10.2”N 102°28&amp;8.8”E, 13 Oct. 2008, <i>Du et al</i>. HNK 2925 (K); Quy Hop?istr. [19.333°, 103.165°], Chau Thai Commune, Ban Nha Village, 15 March 2005, <i>Nguyen et al</i>. HNK 1652 (K 2 sheets); Pu Mat National Park, Khe Kem Waterfall, 18°38&amp;10.2”N 102°28&amp;8.8”E, 13 Oct. 2008, <i>Du et al</i>. HNK 2821 (K); Ninh Binh. Cuc Phuong National Park, 20°21&amp;N 103°33.55&amp;E, 330 m, 22 Nov. 1998, <i>Soejarto et al</i>. 10360 (L, P); Phu Yen. Cho Ganh [13.2833°, 109.23°], Feb. 1923, <i>Petelot</i> 868 (P 2 sheets); Quang Binh.?ong Hoi?istr., Phong Nha-Ke Bang National Park [15.3352°, 106.1313°], Son Trach commune, 230 m, 22 March 2005, <i>Nguyen et al</i>. HNK 1813 (K 2 sheets); Quang Ninh.?ong-dang, Feb. 1886, <i>3alansa</i> 1000 (lectotype K; isolectotypes G, P); Thanh Hoa. Ba Thuoc, Pu Loung Nature Reserve [20.2665°, 103.1525°], 20 Nov. 2002, <i>Phuong</i> 3593 (HN); Thanh Hoa, Ben En National Park [19.6165°, 103.323°], 13 Jan. 2006, <i>Sam</i> HVS 063 (L). <b>HABITAT.</b> In primary lowland, evergreen forest, limestone mountains, 30 – 1500 m. <b>CONSERVATION STATUS.</b> Widespread; Least Concern (LC). <b>PHENOLOGY</b>. Flowering and fruiting October – April. <b>NOTES.</b> <i>Gomphostemma leptodon</i> is recognised by a large corolla, almost 23 – 30 (– 60) mm long and robust venation, especially on the lower side. It is morphologically similar to <i>G. grandiflorum</i> by having fruiting calyx lobes revolute in cross section but <i>G. leptodon</i> has the fruiting calyx lobes longer than the tube at maturity.
description: Published as part of <i>Bongcheewin, Bhanubong, Ingrouille, Martin J. &amp; Paton, Alan J., 2022, A revision of Gomphostemma (Lamiaceae), pp. 27-92 in Kew Bulletin 77 (1)</i> on pages 64-65, DOI: 10.1007/s12225-021-09991-y, http://zenodo.org/record/7615581
description: {"references": ["? oan, T. (1936). Labiatae. In: H. Lecomte (ed.), Flore Generale de L' Indo-Chine 2: 913 - 1026. Masson, Paris.", "Li, X. W. &amp; Hedge, I. C. (1992). Lamiaceae. In: Z. Y. Wu &amp; P. H. Raven (eds), Flora of China 15: 30 - 299. Science Press, Beicing.", "Phuong, V. X. (2000). Lamiaceae (Labiatae). In: N. T. Ban, T.?. Ly, T. C. Khanh, P. K. Loc, N. N. Thin, N. V. Tien &amp; N. K. Khoi (eds), Flora of Vietnam 2: 1 - 258. Science &amp; Technics Publishing House, Hanoi."]}