Hoffeinsodes adamowiczi Skibińska & Szadziewski & Kania & Krzemiński 2020, sp. nov.
Creators: name: Skibińska, Kornelia
name: Szadziewski, Ryszard
name: Kania, Iwona
name: Krzemiński, Wiesław
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Psychodidae
Hoffeinsodes
Hoffeinsodes adamowiczi

description: Hoffeinsodes adamowiczi sp. nov. (Figs. 1–2)
urn:lsid:zoobank.org:act: 91E406F1-1AE9-483D-B66D-67D93417EC6A
Etymology. The new species is dedicated to the late Paweł Adamowicz, ambassador of Baltic amber and promoter of related research, who served many years as the Mayor of the city Gdańsk until his assassination on 13 January 2019.
Diagnosis. The species is characterized by having cross-vein r-m close to the fork M 1+2 (about 5/6 length of M 1+2); M 1 about 2 times longer than M 1+2; epandrium with elongated epandrial lobes greatly extended at its ends; gonocoxite with two lobes. Female unknown.
Description. Male. Head broad with oval eyes (Fig. 1B); palpus four segmented (length in mm: 1/0.15; 2/0.23; 3/0.30; 4/0.63), last segment almost equal in length to the total length of the previous two (Figs. 1B, 2B); antenna with scape (0.04 mm long), pedicel (0.10 mm long) and 14 flagellomeres; flagellomeres long, tubular, decreasing in length (length in mm: 1/0.39; 14/0.12) (Figs. 1B, 2A). Wing length 4.3 mm; width 1.22 mm; Sc ending before half of a wing length and opposite to vein r-m; sc-r about its length before end of Sc; R 1 long, ends opposite about 3/4 length of R 2; fork of Rb into R 1 and Rs before fork of Mb into M 1+2 and M 3+4; R 2+3 more than 2 times longer than Rs; R 2 almost 3 times longer than Rs; r-m located between R 5 and M 1+2; M 1+2 two times longer than Mb; M 1 about 2 times longer than M 1+2; M 3 almost 11 times longer than M 3+4; M 4 about 9 times longer than M 3+4; opening of cells m 1, m 2 and m 3 almost equal in width; Cu ends opposite fork of Rs; m-cu located in fork of M 3+4; A 1 quite long ends clearly after m-cu (Figs. 1C, 2C). Male genitalia. Gonocoxites fused with hypandrium, each with two lobes tapering to a point; compared to each other, lower gonocoxal lobe broad, slightly arched, upper lobe narrow; aedeagus not discernable; gonostyli simple, about twice as long as wide, sub-rectangular in lateral view, slightly narrowing to blunt apices, covered with setae; epandrium with symmetrical, elongated epandrial lobes clearly broadening at apices; proctiger tubular, elongate, extending clearly beyond epandrium; cerci ovate in lateral view (Figs. 1D, 2D, E, F).
Female. Unknown.
Two genera of Bruchomyiinae are known from Baltic amber: Palaeosycorax Meunier, 1905 and Hoffeinsodes. The latter genus, now including seven species, is readily characterized by the fusion of gonocoxites and hypandrium but can also be recognized by an exceptionally large epandrium with elongated lobes. Regarding the wing venation, H. adamowiczi sp. nov. can be easily distinguished by location of cross-vein r-m (Fig. 2C) and proportions of radial and medial veins. Also, external morphology of the male genitalia is unique among described species, especially the epandrium with elongated and apically expanded epandrial lobes. Male genitalia of the new species bear some similarity to those of Hoffeinsodes longicauda from Baltic amber. In the latter species, however, gonocoxites and gonostyli are lacking divisions and gonocoxites have no lobes (Wagner 2017).
We are much indebted to amber dealer Jonas Damzen for donation of the holotype to the Museum of Amber Inclusions at University of Gdańsk. Gregory Curler is greatly appreciated for suggestions to improve the original version of this paper and providing encouragement. Thanks as well to two reviewers for their helpful suggestions. Funding for this research was partially provided by the grant from the National Science Centre of Poland Miniatura 2 no. DEC-2018 /02/ X/ NZ 8/01001.
description: Published as part of Skibińska, Kornelia, Szadziewski, Ryszard, Kania, Iwona & Krzemiński, Wiesław, 2020, A new species of Bruchomyiinae (Diptera: Psychodidae) from Baltic amber, pp. 147-150 in Zootaxa 4838 (1) on pages 147-150, DOI: 10.11646/zootaxa.4838.1.10, http://zenodo.org/record/4403518