Manota arenalensis Jaschhof & Hippa 2005, sp. n.
Creators: name: Jaschhof, Mathias
name: Hippa, Heikki
Publisher: name: Zenodo
Year: 2005
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Mycetophilidae
Manota
Manota arenalensis

description: Manota arenalensis sp. n. (Fig. 12)   
 Description. Large species.   
 Male. Head. Antenna with fourth flagellomere 1.9 times as long as wide. Maxillary palpus with third palpomere bearing 3 curved sensilla; fourth palpomere with a parasegment apically.   
 Thorax. Anterior basalare and laterotergite each bare. Episternum 3 with ventral portion setose. Wing. Length 1.7–2.2 mm. Membrane with a few setae along posterior margin. Sc bare. Junction of CuA2 into CuA1 incomplete or complete.   
 Terminalia (Fig. 12). Sternite 9 almost completely merged with gonocoxites, with only small portions of posterior margin recognisable; margin situated beyond mid length of gonocoxites. Ventro­lateral edges of gonocoxites lobe­like and with some 5 stronger setae; interiorly with 1 curved seta on a separate rim. Disto­lateral lobes of gonocoxites large; with large, straight setae pointing posteriorly and smaller, curly setae pointing inwards. Gonocoxites in position I with 1 medium­sized, blunt megaseta; in position II with 3 short, strong, stiff setae on lobe; in position III with 2 flat megasetae, one megaseta simple and pointed, the other two­branched with one branch broadly rounded and the other branch whip­like; in position IV with 1 pointed megaseta directed inwards. Gonostyli small, slender, bilobed; ventral lobe each with 1 large and 1 smaller setae apically, dorsal lobe with 2+1 setae with one apical seta strongly re­curved. Parastylar lobes large, very weak in some specimens, but seemingly sclerotised in others. Tegmen long and narrow, almost evenly tapering to tip. Hypoproct broad apically; with long setae laterally.   
 Diagnostic characters. Among the species with both anterior basalare and laterotergites bare, it has the terminalia characterised by the combination of very small, slender gonostyli; gonocoxites in position IV with one large, pointed megaseta pointing strictly inwards (see also next species, M. eximia); and sternite 9 almost indistinguishable from the gonocoxites.   
 Etymology. Derived from the type locality, Arenal volcano.   
 Types. Holotype. Male. Costa Rica, Alajuela province, San Carlos, Volcán Arenal National Park, Sendero Pilón, 650 m, 6–7 Nov. 2000, by Malaise trap, G. Carballo (INBio sample #60084). Paratypes. 3 males, same data as the holotype; 3 males, Cartago province, Tapantí­Macizo de la Muerte National Park, Sector la Represa, 1550 m, 26 March 2000, by Malaise trap, D. Rubí & C. Hansson (INBio sample #67894); 2 males, same National Park, but Sendero Arboles Caídos, 1500–1600 m, 9 April 2002, by Malaise trap, D. Rubí & C. Hansson (INBio sample #67891).   
 Other material studied. 1 male, Puntarenas province, La Amistad International Park, Buenos Aires, Sendero a Casa Coca, 13 Oct.–13 Nov. 2001, D. Rubí (INBio sample #74889); 3 males, Limón province, Hitoy Cerere Biological Reserve, Sendero Espavel, 18 Sep.–7 Oct. 2003, E. Rojas, B. Gamboa, W. Arana, M. & C. Jaschhof; 1 male, Limón province, Pococí, Braulio Carrillo National Park, Estación Quebrada González, 21 Aug. 2002, P. Hanson & C. Godoy (INBio sample #70777).
description: Published as part of Jaschhof, Mathias & Hippa, Heikki, 2005, The genus Manota in Costa Rica (Diptera: Mycetophilidae), pp. 1-54 in Zootaxa 1011 (1) on pages 25-27, DOI: 10.11646/zootaxa.1011.1.1, http://zenodo.org/record/5048990