Pusia paraleucozona Harzhauser & Landau 2021, nov. comb.
Creators: name: Harzhauser, Mathias
name: Landau, Bernard
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Neogastropoda
Costellariidae
Pusia
Pusia paraleucozona

description: Pusia paraleucozona (Boettger, 1906) nov. comb.
Figs 14A 1 –A 2, B 1 –B 2
* Mitra (Uromitra) ebenus Lmk. var. paraleucozona n. var. — Boettger 1906: 8.
Vexillum (Uromitra) ebenus paraleucozona (Boettger) — Zilch 1934: 259, pl. 17, fig. 18.
Vexillum (Vexillum) paraleucozona (Boettger, 1906) — Bałuk 1997: 35, pl. 9, fig. 6.
Type material. Lectotype: SMF 360345 (= SMF 12.2208 a), Senckenberg Museum, Frankfurt/Main, Germany, SL: 11.5 mm, MD: 5.0 mm, Coşteiu de Sus (Romania), illustrated in Zilch (1934, pl. 17, fig. 18), figs 14B 1 –B 2. Paralectotypes: SMF 360.2208 b–c, Senckenberg Museum, Frankfurt / Main, Germany, 17+ 7 specimens, Coşteiu de Sus (Romania).
Additional material. NHMW 1865/0036/0010, SL: 9.0 mm, MD: 3.9 mm, Drnovice u Vyškova (Czech Republic), figs 14A 1 –A 2.
Revised description. Shell moderately large, broad fusiform, with distinctly incised suture, and broad conical to weakly cyrtoconoid spire. Protoconch unknown. Teleoconch of seven whorls. Spire whorls weakly convex, with periphery at abapical suture. Axial sculpture of orthocline, convex ribs, separated by interspaces of about same width (about 20–25 axial ribs on last whorl). Adapical tips of axial ribs may be slightly swollen resulting in slightly gradate spire. Last whorl strongly convex, rapidly contracting. Aperture wide, ovoid with indistinct anal sinus. Columellar callus indistinct. Columella with four prominent folds, weakening abapically. Outer lip thin, abapically contracting, about 10–12 lirae within. No spiral sculpture, except four to six spiral cords on weak fasciole, partly extending from columellar folds. Siphonal canal straight, narrow, moderately short, with shallow siphonal notch.
Shell measurements and ratios. SL: 9.0–12.0 mm, MD: 3.9–5.5 mm; AA: 45–55°, SL/MD: 2.3–2.4, AL/AW: 3.7, AH/S: 2.8–3.0.
Discussion. Boettger (1906) described this species as variety of “ Mitra ebenus ” [= Ebenomitra leucozona (Andrzejowski 1830)], which is clearly incorrect. Pusia paraleucozona has four prominent columellar folds and numerous delicate axial ribs, whereas E. leucozona has three folds and few blunt axial swellings. At first glance it is similar to Pusia moravica (Hoernes & Auinger, 1880) in being broadly fusiform with narrow ribs persisting on all whorls, but that species is larger [reconstructed height about 13–16 mm vs 9.0–12.0 mm in Pusia paraleucozona (Boettger, 1906)], at the same size the axial ribs are wider spaced in P. moravica, and the spiral sculpture in the subsutural area is weak, but present, whereas in P. paraleucozona spiral sculpture is restricted to the siphonal fasciole.
Ebenomitra renauleauensis Landau, Ceulemans & Van Dingenen, 2019, from the Tortonian of France, is reminiscent of P. paraleucozona but differs in its smaller size and the coarser axial ribs.
Palaeoenvironment. Unknown; probably offshore settings.
Distribution in Central Paratethys. Badenian (middle Miocene): North Alpine-Carpathian Foredeep: Drnovice u Vyškova (Czech Republic); Korytnica Basin: Korytnica (Poland) (Bałuk 1997); Făget Basin: Coşteiu de Sus (Romania) (Boettger 1906).
description: Published as part of Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, pp. 1-70 in Zootaxa 4982 (1) on pages 41-42, DOI: 10.11646/zootaxa.4982.1.1, http://zenodo.org/record/4921842