cometadata/datacite-rtg-text-other-reclassification-records huggingface dataset:
Diamesa tonsa Haliday in Walker 1856
Creators: name: Montagna, Matteo
name: Urbanelli, Sandra
name: Rossaro, Bruno
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Chironomidae
Diamesa
Diamesa tonsa

description: Diamesa tonsa (Haliday in Walker, 1856)
The illustrations of male genitalia of this species are by Pagast (1947) (as D. thienemanni) and Serra-Tosio (1971b). The species is the most common and widespread Diamesa species in the Alps, and is also common in the Apennines (Rossaro & Lencioni 2015a, b).
description: Published as part of Montagna, Matteo, Urbanelli, Sandra & Rossaro, Bruno, 2016, The species of the genus Diamesa (Diptera, Chironomidae) known to occur in Italian Alps and Apennines, pp. 317-331 in Zootaxa 4193 (2) on page 326, DOI: 10.11646/zootaxa.4193.2.7, http://zenodo.org/record/166939
Diamesa cinerella Meigen in Gisti 1835
Creators: name: Montagna, Matteo
name: Urbanelli, Sandra
name: Rossaro, Bruno
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Chironomidae
Diamesa
Diamesa cinerella

description: Diamesa cinerella (Meigen in Gisti, 1835)
The species is redescribed by Pagast (1947) and Serra-Tosio (1971b). Species delimitation analysis performed with Cytochrome c oxidase subunit I (Montagna et al. 2016) suggested that the species is close to D. tonsa, but the adult males are morphologically well separable to each other. The species is common in glacial streams but in the winter it is also present at low altitudes, as in Ticino River (Rossaro & Lencioni 2015a, b).
description: Published as part of Montagna, Matteo, Urbanelli, Sandra & Rossaro, Bruno, 2016, The species of the genus Diamesa (Diptera, Chironomidae) known to occur in Italian Alps and Apennines, pp. 317-331 in Zootaxa 4193 (2) on page 326, DOI: 10.11646/zootaxa.4193.2.7, http://zenodo.org/record/166939
Phalangopsidae Blanchard 1845
Creators: name: Desutter-Grandcolas, Laure
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Orthoptera
Phalangopsidae

description: Phalangopsidae Blanchard, 1845
Remark. I consider here the family Phalangopsidae as the monophyletic group attested by molecular phylogeny (Chintauan-Marquier et al. 2013, 2015) and redefined in Desutter-Grandcolas et al. (submit.). This family corresponds to the " Phalangopsinae subfamily group" used in the OSF, except that it includes most of the taxa of the Gryllomorphinae subfamily usually classified within the " Gryllinae subfamily group". Its subfamilial and tribal subdivisions are today not fully settled, as most subfamilies and tribes, except the subfamily Phaloriinae Gorochov, 1985, were found para- or polyphyletic with molecular evidence. This subfamily is redefined here, to complete previous definitions which were very short and did not describe most morphological characters.
description: Published as part of Desutter-Grandcolas, Laure, 2015, Phalangopsidae crickets from Tropical Africa (Orthoptera, Grylloidea), with descriptions of new taxa and an identification key for African genera, pp. 451-496 in Zootaxa 3948 (3) on page 454, DOI: 10.11646/zootaxa.3948.3.5, http://zenodo.org/record/241857
Afrophaloria Desutter-Grandcolas
Creators: name: Desutter-Grandcolas, Laure
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Orthoptera
Phalangopsidae
Afrophaloria

description: Genus Afrophaloria Desutter-Grandcolas, n. gen.
(Figs 1–4)
Type species. Afrophaloria amani Desutter-Grandcolas, n. sp.
Other species included. Afrophaloria apiariensis Desutter-Grandcolas, n. sp., Afrophaloria hempae Desutter- Grandcolas, n. sp., Afrophaloria n. sp. 1, Afrophaloria sp.
Etymology. Named after the clade to which these species belong, i.e. the Phaloriinae (see Chintauan-Marquier et al. 2013, 2015) and their place of diversification, i.e. Africa.
Distribution. Eastern and Western Central Africa (Tanzania, Gabon, CAR, Congo).
Diagnosis. Afrophaloria Desutter-Grandcolas, n. gen. can be recognized by the following combination of morphological characters: Males and females apterous (Fig. 1 A, F). Head very high; fastigium wide, rounded, not separated from the vertex (Fig. 1 C), slightly furrowed longitudinally (Fig. 1 B), wider than the scapes (Fig. 1 B), with three large ocelli (Fig. 1 B, C). TI without a tympanum on inner and outer sides. TIII with four pairs of subapical spurs, alternate, subapical spurs 2 and 3 the longest on inner side, subapical spur 1 the shortest on outer side, the others subequal. TIII not serrulate between subapical spurs, with few spines above. Basitarsomeres III with only one row of spines. Second tarsomeres not flattened. Male. Subgenital plate short, with straight apical margin. Supra anal plate without dorsal denticles (Fig. 1 D). Genitalia (Fig. 3): pseudepiphallus partly membranous, with a pair of disto-lateral hooks; pseudepiphallic parameres well-developed, dejected laterally and distally; ectophallic fold very wide and long, much longer than pseudepiphallus, asymmetrical, and with a bifid apex and sclerotized ventral margins; endophallic apodeme lacking; ectophallic dorsal valves lacking. Female. Subgenital plate more or less indented distally (Fig. 2). Ovipositor flattened laterally, apex small and smooth (Fig. 1 G). Genitalia: Copulatory papilla small, flat and very little sclerotized, the apex more of less deeply bisinuated (Figs 2 N–5, 4F–I). Spermathecal duct very short, thick. Spermatheca small, shaped as a thin and elongated kidney (Fig. 2 N, R).
description: Published as part of Desutter-Grandcolas, Laure, 2015, Phalangopsidae crickets from Tropical Africa (Orthoptera, Grylloidea), with descriptions of new taxa and an identification key for African genera, pp. 451-496 in Zootaxa 3948 (3) on page 456, DOI: 10.11646/zootaxa.3948.3.5, http://zenodo.org/record/241857
Afrophaloria Desutter-Grandcolas, 2015, n. sp.
Creators: name: Desutter-Grandcolas, Laure
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Orthoptera
Phalangopsidae
Afrophaloria

description: Afrophaloria n. sp. 1
(Fig. 4 A, Table 2)
I separate here a female that may belong to a new species. It originates from the same locality as Afrophaloria apiariensis Desutter-Grandcolas, n. sp. and is characterized by a very short ovipositor (see measurements below), a subgenital plate only slightly sinuate distally (Fig. 4 A) and a very dark coloration (tergites ochre brown, their distal margin largely bordered with black brown, the last tergites and supra anal plate entirely dark). The only specimen at hand could not be dissected.
Material examined. Gabon, Makandé, affluent Offoué, forêt des abeilles, 11°55'E 0°41'S, 220m, 1 female, 10.vii.1994, jour, fn31, litière (L. Desutter-Grandcolas), MNHN-EO-ENSIF3715.
description: Published as part of Desutter-Grandcolas, Laure, 2015, Phalangopsidae crickets from Tropical Africa (Orthoptera, Grylloidea), with descriptions of new taxa and an identification key for African genera, pp. 451-496 in Zootaxa 3948 (3) on page 466, DOI: 10.11646/zootaxa.3948.3.5, http://zenodo.org/record/241857
Afrophaloria
Creators: name: Desutter-Grandcolas, Laure
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Orthoptera
Phalangopsidae
Afrophaloria

description: Afrophaloria sp.
(Fig. 4 B–I, Table 2)
I separate here female specimens originating from the Central African Republic, Gabon and the Congo, which all present a very long ovipositor, longer than TIII and than FIII (except in the female from Odzala, see measurements below), a subgenital plate (Fig. 4 A–E) and a copulatory papilla more deeply emarginate than in the other species of Afrophaloria Desutter-Grandcolas, n. gen. here described (Fig. 4 F–I). They are globally lightly coloured (head, pronotum, legs); their tergites are brown, more or less marked with dark (last tergites darker, first tergites lighter with their posterior margins bordered by dark brown, planely or as a line of spots, this coloration being highly variable). No corresponding males have been found.
Material examined. Central African Republic, Salo, 40km S Nola, W Sangha, forêt sempervirente, 23.iii.1995, nuit, 1 female, fn49; 30.iii.1995, jour, 2 females, fn6, 60, litière; 30.iii.1995, nuit, 1 female, fn113, litière; 31.iii.1995, jour, 1 female, fn1, litière; 31.iii.1995, nuit, 1 female, fn39, litière; 4.iv.1995, jour, 1 female, fn2, litière, L. Desutter-Grandcolas, MNHN.
Central African Republic, 75km WNW Mbaiki, N'Gotto, W Lobaye, forêt galerie, 17.iii.1995, jour, 1 female, fn25, litière; 18.iii.1995, nuit, 1 female, fn20, sur amadou, L. Desutter-Grandcolas, MNHN.
Central African Republic, 75km WNW Mbaiki, N'Gotto, W Lobaye, forêt galerie inondable, 21.iii.1995, jour, 2 females, fn5, 6, litière; 25.iii.1995, jour, 4 females, fn2, 5–7, litière; 25.iii.1995, nuit, 1 female, fn16, sur plante, L. Desutter-Grandcolas, MNHN.
Central African Republic, 110km W Mbaiki, Bambio, W Lobaye, forêt semi-décidue, 21.iii.1995, nuit, 1 female, fn36, L. Desutter-Grandcolas, MNHN.
Congo, Odzala, Mbamo, 21.xi.1996, 1 female, P. Deleporte, MNHN.
Gabon, Ivindo, 8 km SW Makokou, Plateau d'Ipassa, forêt, 7.vii.1989, jour, 1 female, fn1, litière, P. Grandcolas, MNHN.
Variation. The female from Ipassa has a very small copulatory papilla, compared to other females (Fig. 4 G).
description: Published as part of Desutter-Grandcolas, Laure, 2015, Phalangopsidae crickets from Tropical Africa (Orthoptera, Grylloidea), with descriptions of new taxa and an identification key for African genera, pp. 451-496 in Zootaxa 3948 (3) on page 467, DOI: 10.11646/zootaxa.3948.3.5, http://zenodo.org/record/241857
Kameruloria Gorochov 2003
Creators: name: Desutter-Grandcolas, Laure
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Orthoptera
Haglotettigoniidae
Kameruloria

description: Genus Kameruloria Gorochov, 2003
(Figs 5–7)
Type species. Kameruloria primitiva Gorochov, 2003.
Distribution. Western Central Africa (Cameroon, Congo, Gabon).
Emended diagnosis. Among the Subtiloriini, Kameruloria has been diagnosed (see Gorochov 2003) by its tympana (inner tympanum open, oval, medium-sized; outer tympanum strongly reduced, as a small longitudinal concavity only), its second tarsomeres (weakly widened), the male tegmina with normal stridulatory apparatus (italics mine), developped tegmina in females, the simple supra anal plate of the males (without hook-like spines), its comparatively narrow subgenital plate in males, slightly notched apically, and its male genitalia ("hind epiphallic processes" present but short, "ectoparameres" short, and the "simple and widely-rounded apical part of the guiding rod" (here ectophallic fold) "as well as wide ribbon between left and right endoparameres", see Gorochov 2003, p. 84).
In addition to the general characters of the Phaloriinae for the head, pronotum, tibial apical spurs, TIII and hindtarsomeres III serrulation given above, and those common to African genera (male FW venation: mirror transverse veins parallel over their whole length, harp veins distributed over the whole file width, see Fig. 5 D, E), the following characters could be listed to emend the diagnosis of Kameruloria: TI not widely inflated at the level of the tympana (contra Heterotrypus, Schizotrypus and Subtiloria), TIII inner subapical spurs not very long and almost equal in length, the longest much shorter than the inner dorsal apical spur (contra Heterotrypus, Schizotrypus and Subtiloria); male genitalia (Fig. 6) characterized by a well-developed, wide but short ectophallic fold, shortly going beyond other phallic structures, and short pseudepiphallic structures (distal processes and parameres); ectophallic dorsal valves variably developed (well developped in K. trimaculata Desutter-Grandcolas, n.sp. (Fig. 6 G–I), but very reduced in K. gabonensis Desutter-Grandcolas, n. sp., K. nigricornis Desutter- Grandcolas, n. sp. (Fig. 6 A–C, D–F) and perhaps also in K. primitiva (according to Gorochov 2003, Fig. 15)).
description: Published as part of Desutter-Grandcolas, Laure, 2015, Phalangopsidae crickets from Tropical Africa (Orthoptera, Grylloidea), with descriptions of new taxa and an identification key for African genera, pp. 451-496 in Zootaxa 3948 (3) on pages 467-468, DOI: 10.11646/zootaxa.3948.3.5, http://zenodo.org/record/241857
Upupagryllus alatus Desutter-Grandcolas, n. sp.
Creators: name: Desutter-Grandcolas, Laure
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Orthoptera
Phalangopsidae
Upupagryllus
Upupagryllus alatus

description: Upupagryllus alatus Desutter-Grandcolas, n. sp.
(Figs 13 J–P, 14D–F)
Type locality. Tanzania, Coast. Reg. Kisarame distr., kazimzumbui forets reserve, 39° 03'E 6°57'S.
Type material. Holotype. Tanzania, Coast. Reg. Kisarame distr., kazimzumbui forest reserve,, 39° 03'E 6°57'S, 1 male, Jan–Feb 1991, Frontier Tanzania leg., ZMC. Allotype. Same data as the holotype, 1 female, ZMC. Paratype, 1 male. Same data as the holotype, 1 male, MNHN-EO-ENSIF3742.
Etymology. Species named after male FW condition.
Diagnosis. Within the genus, small, flat and wide species with short legs; light yellowish brown with dark brown abdomen. Male with wide FWs, covering more than half abdomen; stridulatory apparatus (Fig. 13 K): harp with five oblique, parallel veins, mirror low and very wide crossed by one vein only; cell below the mirror longer than mirror width; apical field very reduced, without venation. Lateral field (Fig. 13 L) with high media veins; R and its bifurcations faint. Male genitalia very similar to that of U. subapterus Desutter-Grandcolas, n. sp., the distal sclerites not acute, and less separate from each other; rami shorter distally (Fig. 14). Female apterous; ovipositor longer than TIII. Female genitalia: copulatory papilla very distinctive, with a wide dorsal swelling and a bifid membrane above the apex (Fig. 13 N–P);
description: Published as part of Desutter-Grandcolas, Laure, 2015, Phalangopsidae crickets from Tropical Africa (Orthoptera, Grylloidea), with descriptions of new taxa and an identification key for African genera, pp. 451-496 in Zootaxa 3948 (3) on page 487, DOI: 10.11646/zootaxa.3948.3.5, http://zenodo.org/record/241857
Paragryllinae Desutter 1987
Creators: name: Desutter-Grandcolas, Laure
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Orthoptera
Phalangopsidae

description: Subfamily Paragryllinae Desutter, 1987
Type genus. Paragryllus Guérin-Méneville, 1844.
Remark. Paragryllodes has long been classified in the tribe Endacustini Gorochov, 1986 within the " Phalangopsinae subfamily group", a tribe clearly not monophyletic as then defined (Desutter-Grandcolas 2002). With molecular evidence, Paragryllodes comes within the subfamily Paragryllinae, which monophyly is supported provided a larger definition than currently accepted is considered (see Chintauan-Marquier et al. 2013, 2015; Desutter-Grandcolas et al. submit.).
description: Published as part of Desutter-Grandcolas, Laure, 2015, Phalangopsidae crickets from Tropical Africa (Orthoptera, Grylloidea), with descriptions of new taxa and an identification key for African genera, pp. 451-496 in Zootaxa 3948 (3) on page 488, DOI: 10.11646/zootaxa.3948.3.5, http://zenodo.org/record/241857
Paragryllodes Karny 1909
Creators: name: Desutter-Grandcolas, Laure
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Orthoptera
Haglotettigoniidae
Paragryllodes

description: Genus Paragryllodes Karny, 1909
(Figs 15–17)
Type species. Paragryllodes borgerti Karny, 1909
Other species included. See Kaltenbach (1982) and Desutter-Grandcolas (1999) for argumented species lists. Two additional species close to Paragryllodes amani Desutter-Grandcolas, n. sp. are available from Tanzania, Kimbosa and Tanghanika (MNHN, ZMC).
Distribution. Tropical Western and Eastern Africa, Madagascar, the Andaman island, Western India, Sri Lanka.
Diagnosis. See Kaltenbach (1982) and Desutter-Grandcolas (1999). This genus is characterized by its general shape and short FWs, dark uniform coloration and more or less shiny apparence (Fig. 15). TIII with four outer and three / four inner subapical spurs. Male. FWs thick and more or less corneous; stridulatory apparatus complete, but hardly visible and partly reticulate (Fig. 16 B). Male genitalia characterized by very long, thin and acute ectophallic dorsal valves, extending well beyond the phallic complexe; pseudepiphallus often densely covered with strong setae (Fig. 17). Female. FWs present, short, partly overlapping or not; venation weak, with longitudinal parallel veins (Fig. 15 F). Subgenital plate transverse, the apex bisinuate. Ovipositor with smooth apex. Female genitalia: copulatory papilla triangular, and more or less elongate (Fig. 16 O–N).
Phylogenetic relationships. In Chintauan-Marquier et al. 's (2013, 2015) molecular phylogeny, Paragryllodes is the sister group of a wide clade including present-day Paragryllinae, (Caltathra Otte, 1987 — Protathra Desutter- Grandcolas, 1997), Brevizacla Gorochov, 2003 and Mikluchomaklaia Gorochov, 1986; the whole clade is the sister group of the neotropical Aclodae group (Aclodes Hebard, 1928, Paraclodes Desutter-Grandcolas, 1992 and Uvaroviella Chopard, 1923, see Desutter-Grandcolas 1992). While this topology has to be confirmed with additional data, the position of Paragryllodes within this clade seems well-supported.
description: Published as part of Desutter-Grandcolas, Laure, 2015, Phalangopsidae crickets from Tropical Africa (Orthoptera, Grylloidea), with descriptions of new taxa and an identification key for African genera, pp. 451-496 in Zootaxa 3948 (3) on page 489, DOI: 10.11646/zootaxa.3948.3.5, http://zenodo.org/record/241857
Cryptophialida Kolbasov, Newman & Hoeg 2009
Creators: name: Botha, Thomas P. A.
name: Griffiths, Charles L.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Cryptophialida

description: Order CRYPTOPHIALIDA Kolbasov, Newman & Høeg (Cited in Kolbasov, 2009)
Cryptophialida Kolbasov, Newman & Høeg in Kolbasov, 2009: 241.— Chan, Hsieh & Kolbasov 2014: 2.
Diagnosis. Mainly characterised by females having long, narrow-necked operculum, with small circular aperture and developed lateral and reinforcing bars, tongue-shaped labrum, no caudal appendages and special gastric mill at end of stomach. Rudimentary mouth cirri, thorax with one or two whip-like dorsal processes (Chan, Hsieh & Kolbasov, 2014). Posterior end of dwarf males with circular, cuticular ribs and frequently with mantle teeth; simple attachment antennules without stalk. Cypris larvae posses rudimentary thorax and thoracopods, carapace covers body incompletely. Burrow opening rounded and small.
Remarks. Cryptophialida consists of the monotypic family Cryptophialidae.
description: Published as part of Botha, Thomas P. A. & Griffiths, Charles L., 2021, South African Acrothoracica (Crustacea: Cirripedia), pp. 45-78 in Zootaxa 4949 (1) on page 48, DOI: 10.11646/zootaxa.4949.1.3, http://zenodo.org/record/4635765
Cryptophialidae Gerstaecker 1866
Creators: name: Botha, Thomas P. A.
name: Griffiths, Charles L.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Pygophora
Cryptophialidae

description: Family CRYPTOPHIALIDAE Gerstaecker, 1866
Cryptophialidae Gerstaecker, 1866: 534.
Diagnosis. As for Cryptophialida.
Remarks. Contains the genera Australophialus Tomlinson, 1969 and Cryptophialus Darwin, 1854 with five and 16 species, respectively. In South Africa, only Australophialus observed.
description: Published as part of Botha, Thomas P. A. & Griffiths, Charles L., 2021, South African Acrothoracica (Crustacea: Cirripedia), pp. 45-78 in Zootaxa 4949 (1) on page 48, DOI: 10.11646/zootaxa.4949.1.3, http://zenodo.org/record/4635765
Australophialus , Tomlinson 1969
Creators: name: Botha, Thomas P. A.
name: Griffiths, Charles L.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Pygophora
Cryptophialidae
Australophialus

description: Genus Australophialus, Tomlinson, 1969
Australophialus Tomlinson, 1969: 117.
Diagnosis. Characterised by having four pairs of terminal cirri and one whip-like dorsal body process (Chan, Hsieh & Kolbasov, 2014).
Remarks. Globally, there are currently five species in this genus, two of which are found in South Africa (both endemic).
description: Published as part of Botha, Thomas P. A. & Griffiths, Charles L., 2021, South African Acrothoracica (Crustacea: Cirripedia), pp. 45-78 in Zootaxa 4949 (1) on page 49, DOI: 10.11646/zootaxa.4949.1.3, http://zenodo.org/record/4635765
Lithoglyptida Kolbasov, Newman & Hoeg 2009
Creators: name: Botha, Thomas P. A.
name: Griffiths, Charles L.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Lithoglyptida

description: Order LITHOGLYPTIDA Kolbasov, Newman & Høeg (Cited in Kolbasov, 2009)
Lithoglyptida Kolbasov, Newman & Høeg in Kolbasov, 2009: 239.— Chan, Hsieh & Kolbasov 2014: 12.
Diagnosis. Females with wide aperture, sac-like mantle, operculum without a neck, with opercular bar less than aperture length, well-developed mouth cirri, large, saddle-like labrum and lack special gastric mill at end of stomach (Kolbasov, 2009; Chan, Kolbasov & Cheang, 2012; Chan, Hsieh & Kolbasov, 2014). Thorax without long dorsal processes. Burrow shape elongated. Males have surfaces with several cuticular denticles and lack mantle teeth. Cypris larvae with developed thorax and thoracopods with an unperforated carapace with fronto-lateral pores.
Remarks. Consists of two families, the Lithoglyptidae and Trypetesidae. No members of the Trypetesidae known from South Africa (although reported from south-west of Madagascar).
description: Published as part of Botha, Thomas P. A. & Griffiths, Charles L., 2021, South African Acrothoracica (Crustacea: Cirripedia), pp. 45-78 in Zootaxa 4949 (1) on page 55, DOI: 10.11646/zootaxa.4949.1.3, http://zenodo.org/record/4635765
Lithoglyptidae Aurivillius 1892
Creators: name: Botha, Thomas P. A.
name: Griffiths, Charles L.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Pygophora
Lithoglyptidae

description: Family LITHOGLYPTIDAE Aurivillius, 1892
Lithoglyptidae Aurivillius, 1892: 133.— Kolbasov 2009: 239.— Chan, Hsieh & Kolbasov 2014: 13.
Diagnosis. Females with well-developed opercular bars, comb collar, thoracic lappets and mouth appendages; twosegmented protopod mouth cirri, biramous multi-annulated terminal cirri, caudal appendages present in some species, intestine with anus. Dwarf males pear-shaped, with pair of lobes at base of attachment antennules (Chan, Hsieh & Kolbasov, 2014).
Remarks: Consist of three subfamilies: Berndtiinae, Lithoglyptinae and Kochlorininae, all present in South Africa.
description: Published as part of Botha, Thomas P. A. & Griffiths, Charles L., 2021, South African Acrothoracica (Crustacea: Cirripedia), pp. 45-78 in Zootaxa 4949 (1) on page 55, DOI: 10.11646/zootaxa.4949.1.3, http://zenodo.org/record/4635765
Berndtiinae Utinomii 1950
Creators: name: Botha, Thomas P. A.
name: Griffiths, Charles L.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Pygophora
Lithoglyptidae

description: Subfamily BERNDTIINAE Utinomii, 1950
Berndtiidae Utinomi, 1950: 457.
Weltneriinae Kolbasov & Newman, 2005: 40.
Diagnosis. Females with five pairs of terminal cirri. Caudal appendages present in Weltneria, but not in Berndtia.
Remarks. Currently there are two genera described from this monophyletic subfamily, Berndtia Utinomi, 1950 and Weltneria. No species of Berndtia are reported from Africa (Kolbasov, 2009), however further investigation of coral species could potentially reveal their presence, as corals are their preferred substrate (Chan, Hsieh & Kolbasov, 2014; Chan et al., 2014).
description: Published as part of Botha, Thomas P. A. & Griffiths, Charles L., 2021, South African Acrothoracica (Crustacea: Cirripedia), pp. 45-78 in Zootaxa 4949 (1) on page 55, DOI: 10.11646/zootaxa.4949.1.3, http://zenodo.org/record/4635765
Weltneria Berndt 1907
Creators: name: Botha, Thomas P. A.
name: Griffiths, Charles L.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Pygophora
Lithoglyptidae
Weltneria

description: Genus Weltneria Berndt, 1907
Weltneria Berndt, 1907: 289.
Diagnosis. Females with five pairs of biramous terminal cirri, two-segmented caudal appendages. This is considered plesiomorphic, as this represents the ground pattern for all thecostracans (Kolbasov & Newman, 2005). Lateral bars weak or absent, orificial knob absent. Dwarf males pear-shaped, tapering towards the top, no lateral projections, attachment antennules with stalk absent (Chan, Hsieh & Kolbasov, 2014).
Remarks. At present, there are 12 Weltneria species globally (Kolbasov, Chan & Cheng, 2017). Two of these occur in South Africa, of which W. spinosa is endemic.
description: Published as part of Botha, Thomas P. A. & Griffiths, Charles L., 2021, South African Acrothoracica (Crustacea: Cirripedia), pp. 45-78 in Zootaxa 4949 (1) on page 55, DOI: 10.11646/zootaxa.4949.1.3, http://zenodo.org/record/4635765
Lithoglyptinae Aurivillius 1892
Creators: name: Botha, Thomas P. A.
name: Griffiths, Charles L.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Pygophora
Lithoglyptidae

description: Subfamily Lithoglyptinae Aurivillius, 1892
Lithoglyptinae Aurivillius, 1892: 133.— Kolbasov & Newman 2005: 41.— Kolbasov 2009: 240.
Diagnosis. Lithoglyptids with 4 pairs of terminal cirri and with caudal appendages. Dwarf males can be bottleshaped, pear-shaped or with wing-like lateral projections, with simple attachment antennules, or with long attachment stalk.
Remarks. There are currently 3 genera in this subfamily, Auritoglyptes, Balanodytes and Lithoglyptes. Only Balanodytes is found in South Africa.
description: Published as part of Botha, Thomas P. A. & Griffiths, Charles L., 2021, South African Acrothoracica (Crustacea: Cirripedia), pp. 45-78 in Zootaxa 4949 (1) on page 63, DOI: 10.11646/zootaxa.4949.1.3, http://zenodo.org/record/4635765
Balanodytes Utinomii 1950
Creators: name: Botha, Thomas P. A.
name: Griffiths, Charles L.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Pygophora
Lithoglyptidae
Balanodytes

description: Genus Balanodytes Utinomii, 1950
Balanodytes Utinomi, 1950: 453, 458.— Chan et al. 2013: 226.
Armatoglyptes Kolbasov & Newman, 2005: 47–51.
Diagnosis. Females, terminal cirri with four pairs, two-segmented caudal appendages without pedestals, opercular area with no posterior setose processes or lobes, posterior processes of opercular bars well-developed, orificial knob and lateral bars both present or absent. Dwarf males with no attachment stalk.
Remarks. Utinomi (1950) erected this genus and the species was described without caudal appendages. Initially the genus only included B. taiwanus and subsequently no additional species were added. However, in 2013 Chan et al. discovered that a B. taiwanus specimen from Utinomi in fact had caudal appendages. The diagnosis was thus corrected and then became the same as that of Armatoglyptes Kolbasov & Newman, 2005, which consisted of nine species, but without a type species designated. Thus, all species of Armatoglyptes were moved to Balanodytes and Armatoglyptes was deemed to be a junior synonym. Hence that name is no longer in use (Chan et al., 2013; Lin et al., 2016) and 11 species were included in the genus. Two species are recorded below for the first time in South Africa, one of which is described as new, thus bringing the total in the genus to 12.
description: Published as part of Botha, Thomas P. A. & Griffiths, Charles L., 2021, South African Acrothoracica (Crustacea: Cirripedia), pp. 45-78 in Zootaxa 4949 (1) on page 63, DOI: 10.11646/zootaxa.4949.1.3, http://zenodo.org/record/4635765
Balanodytes flexuosus
Creators: name: Botha, Thomas P. A.
name: Griffiths, Charles L.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Pygophora
Lithoglyptidae
Balanodytes
Balanodytes flexuosus

description: Balanodytes flexuosus (Chan, Kolbasov & Cheang, 2012)
Figs 13, 14
Material examined. CLP14-4, Port Shepstone, Eastern Cape, August 2017, 1 specimen in Charonia lampas shell (specimen used for dissection).
Diagnosis. Balanodytes with opercular bars with bifid teeth, posterior processes L-shaped, strongly bent/recurved at tip.
Description. Female, mantle sac oval-shaped, narrowing below to opercular bars (figs 13, 14A). Length 1.65 mm, width 1.05 mm, opercular bars 500 µm in length, with row of bifid teeth and setae (fig. 14B). Posterior processes hook-like (anteriorly bent/recurved), L-shaped, with bifid teeth and setae along length (Figs 13, 14B). Comb collar with long, feather-like projections, fused at bases. Orificial knob distinct, with setae and simple or bifid teeth (fig. 14B). Lateral area below opercular bar covered with sparse, simple setae and rows of large, multifid scales. Feeble lateral bars. Burrow opening narrow, oval-shaped.
Four pairs of terminal cirri and pair of two-segmented caudal appendages (fig. 14C, D). Mouth cirri with both rami three-segmented, with anterior ramus shorter than posterior ramus, both rami with plumose setae (fig. 14H). Labrum saddle-like, with small, blunt teeth on anterior margin (fig. 14E). Mandible with three large teeth, two smaller teeth close to lower margin, large notch between first and second teeth. Inferior angle with two sharp denticles and setae (fig. 14F). Maxillule with two long, sharp upper and single, smaller lower cuspidate setae. Notch with short, sharp seta, lower 2/3 of cutting edge with numerous short, sharp setae (fig. 14G). Maxilla triangular with setae on both exterior and interior margins. Mandibular palp trapezoidal, with simple setae along exterior margin.
Hosts. Previously found in the coral Pavona sp. from Mozambique Channel (Chan, Kolbasov & Cheang, 2012). Collected in South Africa from Charonia lampas shell.
Distribution. Originally described from Mozambique Channel (2611’00”S, 3501’00”E). Collected in South Africa only in Port Shepstone (3044’38.8”S, 3027’28.7”E). This is the first record of this species from South Africa.
Remarks. Only one individual collected, found in association with Balanodytes sp. A
description: Published as part of Botha, Thomas P. A. & Griffiths, Charles L., 2021, South African Acrothoracica (Crustacea: Cirripedia), pp. 45-78 in Zootaxa 4949 (1) on pages 63-64, DOI: 10.11646/zootaxa.4949.1.3, http://zenodo.org/record/4635765
Kochlorininae Gruvel 1905
Creators: name: Botha, Thomas P. A.
name: Griffiths, Charles L.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Pygophora
Lithoglyptidae

description: Subfamily KOCHLORININAE Gruvel, 1905
Gruvel, 1905: 335.
Diagnosis. Females with three pairs of biramous terminal cirri, two-segmented caudal appendages present.
Remarks. Consists of two genera; Kochlorine Noll, 1872 and Kochlorinopsis Stubbings, 1967, with only the former present in South Africa. Globally these genera contain 7 and 1 species, respectively.
description: Published as part of Botha, Thomas P. A. & Griffiths, Charles L., 2021, South African Acrothoracica (Crustacea: Cirripedia), pp. 45-78 in Zootaxa 4949 (1) on page 69, DOI: 10.11646/zootaxa.4949.1.3, http://zenodo.org/record/4635765
Kochlorine Noll 1872
Creators: name: Botha, Thomas P. A.
name: Griffiths, Charles L.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Pygophora
Lithoglyptidae
Kochlorine

description: Genus Kochlorine Noll, 1872
Kochlorine.— Tomlinson, 1969: 65.
Diagnosis. Females with three pairs of terminal cirri, caudal appendages present (two-segmented), developed lateral bars and orificial knob. Dwarf males with elongated attachment stalk and lateral projections (Chan, Hsieh & Kolbasov, 2014).
Remarks. There are currently seven described species in this genus, of which K. bihamata, was previously thought to occur in South Africa. Below we suggest that report be discounted. However, two other species belonging to this genus are reported for the first time from the region, one of which is considered new to science.
description: Published as part of Botha, Thomas P. A. & Griffiths, Charles L., 2021, South African Acrothoracica (Crustacea: Cirripedia), pp. 45-78 in Zootaxa 4949 (1) on page 69, DOI: 10.11646/zootaxa.4949.1.3, http://zenodo.org/record/4635765
Kochlorine bihamata Noll 1883
Creators: name: Botha, Thomas P. A.
name: Griffiths, Charles L.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Pygophora
Lithoglyptidae
Kochlorine
Kochlorine bihamata

description: Kochlorine bihamata Noll, 1883 (deletion from fauna list)
Kochlorine bihamata.— Barnard 1924: 99.— Tomlinson, 1969: 72.— Kolbasov, 2009: 320.
Diagnosis. Females with no anterior conical processes, mantle aperture with two posterior hooks and spiny teeth on rim; size 5 mm (Tomlinson, 1969).
Hosts. Collected as dried specimens observed in an Haliotis shell. Although not explicitly stated which Haliotis species is referred to, it can be assumed this is the abalone, Haliotis midae, which is known to host other acrothoracicans.
Remarks. This species has not been observed since its description from Cape Town by Noll (1883) and there remains grave doubt regarding the validity of the species. Noll (1883) mentioned that: “Should new specimens of Kochlorine arise from Cape Town, one can well label it Kochlorine bihamata ”. Therefore, it is safe to assume that the genus Kochlorine occurs from Cadiz, Spain to Cape Town, South Africa and burrows into shells of Haliotis spp.. Although Tomlinson (1969) calls into question the validity of the species, he later states that K. bihamata should be treated as a valid species. However, following our survey no trace of this species was found, despite the type host being collected at the type locality and several adjacent localities. Thus, we consider that it is safe to assume that this is a nomen nudum.
description: Published as part of Botha, Thomas P. A. & Griffiths, Charles L., 2021, South African Acrothoracica (Crustacea: Cirripedia), pp. 45-78 in Zootaxa 4949 (1) on page 69, DOI: 10.11646/zootaxa.4949.1.3, http://zenodo.org/record/4635765
Cladophyllus oberthuri Houlbert 1912
Creators: name: Huang, Hao
name: Chen, Chang-Chin
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Lucanidae
Cladophyllus
Cladophyllus oberthuri

description: Cladophyllus oberthuri Houlbert, 1912
(Figs. 1–4, 13–14)
Cladophyllus oberthuri Houlbert, 1912: 139 [type locality: Tsekou (in Lancang valleys, northwestern Yunnan)], figures for habitus, mandible, antenna, canthus, legs, scutellum and mesosternum; Mizunuma & Nagai 1994: plate 148, Fig. 111, “ holotype ”; Boucher 1996: 73, lectotype designation; Fig. 3, male head; Fig. 6, male mentum; Fig. 9, male pronotum; and Fig. 15, male genitalia.
Cladophyllus flavus Didier, 1930: 126 [type locality: “Yunnan, Sseu-tch’ouan, Thibet” (no exact locality)]; Mizunuma & Nagai 1994: plate 148, Fig. 112, “ holotype ”; Boucher 1996: 73, lectotype designation, synonymy.
Material examined. CHINA: Yunnan: 9 3, 4 ƤƤ, Diqing, Weixi County, Pantiange, ca. 2000m, 8.VII. 2010, Xiao-Dong Yang and Wen-Xuan Bi leg.; 2 3 (CHH, CYXW), Diqing, Weixi County, above 2000m, VII. 2006, An-Ming Chen leg.
Identification. Boucher (1996) designated the lectotypes of both Cladophyllus oberthuri and C. flavus and considered them as synonyms; he described and illustrated the male genitalia for the first time. According to the original description and Boucher’s paper, C. oberthuri can be distinguished from C. bousqueti Boucher, 1996 by the more deeply bifurcated antennomere 10, the less transverse mentum, the blunter lateral angles of the pronotum and the markedly longer flagellum of the median lobe with apex not enlarged. The male specimens examined by us match all these characters for C. oberthuri.
The female of Cladophyllus oberthuri are here illustrated and described for the first time; they have been found inside decayed logs together with the male specimens.
Taxonomic notes. All three known species are allopatric in distribution: C. oberthuri is restricted to the Lancang valleys in northwestern Yunnan; C. bousqueti is distributed to the west of the Nujiang River and into northeastern Myanmar; C. nobuhikoi is restricted to the mountain ranges around China-Vietnam border. They are considered as separate because of the marked differences in male and female genitalia.
Distribution. northwestern Yunnan (Lancang valleys).
description: Published as part of Huang, Hao & Chen, Chang-Chin, 2012, A review of the genera Prismognathus Motschulsky and Cladophyllus Houlbert (Coleoptera: Scarabaeoidea: Lucanidae) from China, with the description of two new species, pp. 1-36 in Zootaxa 3255 on pages 3-4, DOI: 10.5281/zenodo.280626
Cladophyllus bousqueti Boucher 1996
Creators: name: Huang, Hao
name: Chen, Chang-Chin
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Lucanidae
Cladophyllus
Cladophyllus bousqueti

description: Cladophyllus bousqueti Boucher, 1996
(Figs. 5–8, 15–16)
Cladophyllus bousqueti Boucher, 1996: 77 [type locality: Chinese Gaoligong, China-Myanmar border (near Pianma, western Yunnan)], Figs. for habitus, head, antenna, mentum, pronotum, mesotibia of male, and male genitalia. Prismognathus bousqueti: Fujita 2010: 169, new combination, plate 101, Figs. 510 (1–2) for males from “Chudu Razi, NE. Kachin, Myanmar ”.
Material examined. CHINA: Yunnan: 3 3, 1 Ƥ, Nujiang, Lushui County, Pianma, Yaojiaping, ca. 2400m, 23.VI. 2010, Xiao-Dong Yang leg.
Identification. The three male specimens examined by us were collected from the same area as the type locality and they match all the characters of the holotype of Cladophyllus bousqueti described by Boucher (1996) including the male genitalic characters.
A female of Cladophyllus bousqueti is illustrated and described here for the first time; it was found inside decayed logs together with the male specimens.
Distribution. Western Yunnan (central Gaoligong Mts.), northeastern Myanmar (Chudu Razi, Kachin).
description: Published as part of Huang, Hao & Chen, Chang-Chin, 2012, A review of the genera Prismognathus Motschulsky and Cladophyllus Houlbert (Coleoptera: Scarabaeoidea: Lucanidae) from China, with the description of two new species, pp. 1-36 in Zootaxa 3255 on page 4, DOI: 10.5281/zenodo.280626
Cladophyllus nobuhikoi Ikeda 2000
Creators: name: Huang, Hao
name: Chen, Chang-Chin
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Lucanidae
Cladophyllus
Cladophyllus nobuhikoi

description: Cladophyllus nobuhikoi Ikeda, 2000
(Figs. 9–12)
Cladophyllus bousqueti nobuhikoi Ikeda, 2000: 42 [type locality: Sapa, northern Vietnam], Fig. (not giving a number), male habitus.
Prismognathus nobuhikoi: Fujita 2010: 169, taxonomic changes, plate 101, Figs. 511-1, male; 511-2, female, from Sapa, Vietnam.
Material examined. CHINA: Yunnan: 1 3, Jinping County, Fenshuiling Nature Reserve, ca. 1800m, 18.IX.2010, Xiao-Dong Yang leg., captured from a decay log.
Identification. This species was described as a subspecies of Cladophyllus bousqueti, only differing for the blackish brown body and the strongly curved mandible with 5–6 inner teeth stated as the diagnostic characters. It was raised to species status by Fujita (2010), who listed more differences. A male specimen from southern Yunnan examined by us matches all the characters of Cladophyllus nobuhikoi. The examination of the male genitalia proved this species to be different from C. bousqueti. It should be noted that Jinping in southern Yunnan is very close to Sapa in northern Vietnam and shares the same lucanid fauna.
A female of Cladophyllus nobuhikoi was illustrated by Fujita (2010) and differs from the female of C. bousqueti by the characters of antenna. The female genitalia of Cladophyllus nobuhikoi were not previously described.
Distribution. Southern Yunnan (Fenshuiling Reserve), northern Vietnam (Sapa).
description: Published as part of Huang, Hao & Chen, Chang-Chin, 2012, A review of the genera Prismognathus Motschulsky and Cladophyllus Houlbert (Coleoptera: Scarabaeoidea: Lucanidae) from China, with the description of two new species, pp. 1-36 in Zootaxa 3255 on pages 4-5, DOI: 10.5281/zenodo.280626
Prismognathus dauricus Motschulsky 1860
Creators: name: Huang, Hao
name: Chen, Chang-Chin
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Lucanidae
Prismognathus
Prismognathus dauricus

description: Prismognathus dauricus (Motschulsky, 1860)
(Figs. 21, 98, 125, 137, 179, 207, 234)
Metopodontus dauricus Motschulsky, 1860: 137 [type locality: “la notre se rencontre au sud de la Daourie, et sur les bord du fl. Amour, aux environs du fort Mariinsk”], plate 9, Fig. 11, female.
Prismognathus subaeneus Motschulsky, 1860: 138 [type locality: “bords du fl. Amour”], plate 9, Fig. 12, male; Didier & Séguy, 1953: 127. Synonym.
Cyclorasis jekelii Parry, 1864: 41 –42 [type locality: “Chowsan, Corea ”], plate 9, Fig. 4, male head. Synonymized by Benesh (1960).
Prismognathus dauricus: Benesh 1960: 56, synonymic list; Mizunuma & Nagai, 1994: 238, plate 51, Fig. 235, males and female; Wan et al. 2007: Figs. 16–19, males and female; Figs. 23–24, male genitalia; Fig. 25, female genitalia; Kim & Kim 2010: 72 -76, locality details in Korea; Fig. 12 p, male and female; Fujita 2010: 164, plate 99, Fig. 488, males and female from Tsushima Island and northern Korea.
Prismognathus davidis: Didier & Séguy 1952: partim [plate 91, Figs. 1–2]. Misidentification.
Material examined. CHINA: Liaoning: 12 3, 8 ƤƤ, Dan-dong City, Laogugou, 26.VII.-11.VIII.2008, H.-C. Shan leg.; 2 3, 2 ƤƤ (CHH, CYXW), Shenyang, no detailed data. Jilin: 2 ƤƤ (IEAS), Changbaishan, Erdaokouhe, 14.VIII.1984, Z.-Y. Liu leg. KOREA: 6 3, 4 ƤƤ, Mt. Dulunsan, VIII.2009.
Identification. This species is distributed in the Russian part of Amur region and in northeastern China; there are no other species of Prismognathus living in the same area.
Diagnosis. The large and medium-size males of P. dauricus are characterized by the absence of the subbasal teeth along the dorsal ridge of the mandible. The small males of P. dauricus are not easy to separate from those of P. klapperichi and P. davidis (apart for the different geographic distribution) unless the genitalia are examined. The females of P. dauricus are indistinguishable in external characters from those of the similar species, especially P. davidis. To identify the female of P. dauricus, a dissection is always needed. For more details see the keys below.
Distribution. Liaoning; Russia (eastern Siberia), Korea, Mongolia, Japan (Tsushima Island only).
description: Published as part of Huang, Hao & Chen, Chang-Chin, 2012, A review of the genera Prismognathus Motschulsky and Cladophyllus Houlbert (Coleoptera: Scarabaeoidea: Lucanidae) from China, with the description of two new species, pp. 1-36 in Zootaxa 3255 on page 5, DOI: 10.5281/zenodo.280626
Prismognathus prossi Bartolozzi & Wan 2006
Creators: name: Huang, Hao
name: Chen, Chang-Chin
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Lucanidae
Prismognathus
Prismognathus prossi

description: Prismognathus prossi Bartolozzi & Wan, 2006
(Figs. 27, 68, 72, 94, 99–100, 114 –116, 162–163, 182 –184, 210 –211, 222, 235)
Prismognathus prossi Bartolozzi & Wan 2006: 61 [type locality: “Xizang, Fa-Mu-Dui, Linzhi” (most probably incorrect locality provided by the specimen dealer)], Figs. 1–7, male, female, male genitalia and female genitalia; Huang & Chen 2010: 115, remarks on the type locality; Fujita 2010: 165, plate 99, partim [Figs. 492-1–2 for males].
Material examined. CHINA: Sichuan: 19 3, 13 ƤƤ, Ya-an City, Yingjing, County, Siping, VII.2007 -2009, X.- D. Yang & W.-X. Lin leg.; 11 3, 4 ƤƤ, Tianquan County, Erlangshan, H.-C. Shan leg., VIII.2007; 2 3, Hanyuan County, VII.2005, C.-J. Dai leg., ex. coll. H. Huang; 1 3, Beichuan County, Xiao-zhai-zi-gou, VII. 2007. Guizhou Province: 2 3, 2 ƤƤ, Tongren City, Jiangkou County, Fanjingshan, VII.2002, W.-I. Chou leg.; 2 3, 15 ƤƤ, Fanjingshan, 10.VII.2009, X.-D. Yang leg. Guangxi Province: 2 3, 1 Ƥ (IEAS), Xing-an County, Mao-er-shan Mts., 900-1500m, 20-21.VIII.1992, X.-W. Liu & H.-S. Yin leg.
Identification. The type locality, as discussed by Huang & Chen (2010), was actually in the mountains around Hanyuan, western Sichuan. This species is very peculiar for the angled preocular margin of the male head. Male specimens from Sichuan examined by us match with the original description of P. prossi. Male specimens from Guizhou and Guangxi have been identified with the examination of genitalia.
The female paratype in the original description was confirmed by us to be P. prossi, with a relatively longer canthus than in other species. Specimens in both sexes have been captured from three widely-separated localities in Sichuan, Guizhou and Guangxi.
Taxonomic notes. This species is separated from all other species not only by the peculiar preocular margin of male, but also by the combination of a very long flagellum and a larger apex of the flagellum itself in male genitalia.
Distribution. Western and northern Sichuan (Hanyuan, Yingjing, Tianquan, Beichuan), Guizhou (Fanjingshan), Guangxi (Mao-er-shan).
description: Published as part of Huang, Hao & Chen, Chang-Chin, 2012, A review of the genera Prismognathus Motschulsky and Cladophyllus Houlbert (Coleoptera: Scarabaeoidea: Lucanidae) from China, with the description of two new species, pp. 1-36 in Zootaxa 3255 on page 11, DOI: 10.5281/zenodo.280626