Polynema (Polynema) grafinya Triapitsyn 2021, sp. n.
Creators: name: Triapitsyn, Serguei V.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Mymaridae
Polynema
Polynema grafinya

description: Polynema (Polynema) grafinya Triapitsyn, sp. n.
(Figs 73–82)
Type material. Holotype female [NZAC] on slide (Fig. 74) labeled: 1. “ NEW ZEALAND: MB, Red Hills, Wairau, 22 Mar. 1972, E. W. Valentine, E1121/4 ENTOMOLOGY DIVISION D.Z.I. R. NZ”; 2. “ ♀ Polynema 28 NOV 1980 ”. The type locality is on South Island.
Paratypes: NEW ZEALAND. South Island: Coronet Peak (OL), 1640 m, i.1981, J.S. Noyes, E.W. Valentine (swept, Tussock/Alpine shrubs, Hebe, mat plants) [1♀ on slide, NZAC]. Wairau (MB), Red Hills, 22.iii.1972, E.W. Valentine (E1121) [11♀ on 3 cards (6, 3, and 2 respectively), 1Ƌ on slide, NZAC].
Description. FEMALE (holotype). Body and appendages dark brown except petiole and legs (the latter partially) brown (as in Fig. 79). Head (Fig. 76) wider than high; face with fine, inconspicuous subtorular lines. Antenna (Fig. 73) with scape, including short radicle, 3.3× as long as wide, smooth; pedicel smooth, longer than F1 and 2.0× as long as wide; F1 the shortest and F2 the longest funiculars, F3 as long as F6 and longer than F4 or F5, length to width ratios of funiculars: F1—2.3, F2—6.1; F3—4.2; F4—2.4; F5—2.2; F6—2.6, F6 with 1 mps; clava 2.6× as long as wide, about as long as combined length of 3 preceding flagellomeres, with 7 mps. Mesosoma about 1.7× as long as wide, mostly smooth except mesoscutum with fine, inconspicuous mesh-like sculpture; pronotum entire; mesoscutum wider than long, with notaular grooves; scutellum shorter than mesoscutum, with campaniform sensilla closer to anterior than to posterior margin and widely separated from each other but at a distance from lateral margins (as in Fig. 81), and with frenal row of small foveae; propodeum (Fig. 75) with a median carina extending from posterior margin for about half length. Fore wing (Fig. 78) 5.0× as long as wide; marginal vein with 1 dorsal macrochaeta; disc slightly infumate, densely setose beyond venation; the discal setae originating behind apex of submarginal vein; longest marginal seta 1.0× greatest width of wing. Hind wing about 33× as long as wide; disc slightly infumate, with an incomplete median row of setae; longest marginal seta 5.3× greatest width of wing. Metacoxa almost smooth. Petiole with anterolateral “wings”. Ovipositor occupying almost entire length of gaster (Fig. 77), exserted beyond gastral apex by 0.14× own total length, and 1.7× length of metatibia.
Measurements of the holotype (µm). Mesosoma: 320; gaster: 449; ovipositor: 486. Scape: 121; pedicel: 60; F1: 30; F2: 85; F3: 55; F4: 38; F5: 41; F6: 55; clava: 136. Fore wing: 898:179; longest marginal seta: 182. Hind wing: 763:23; longest marginal seta: 121.
Variation (paratypes). Body length (dry-mounted specimens, as in Fig. 79) 825–960 µm. In the slide-mounted specimen, scape, including radicle, 3.5× as long as wide, clava 2.3× as long as wide, fore wing 5.5× as long as wide, longest marginal seta 1.3× greatest width of fore wing, and ovipositor exserted beyond gastral apex by about 0.1× own total length, and 1.6× length of metatibia.
MALE (paratype). Similar to female in color except petiole (Fig. 81) brown. Antenna (Fig. 80) with scape, excluding radicle, 2.2× long as wide and shorter than any funicular. Fore wing (Fig. 82) 5.4× as long as wide; longest marginal seta 1.1× greatest width of wing. Genitalia typical for species of P. (Polynema) (Triapitsyn & Fidalgo 2006), 230 µm long; digitus with 2 small denticles apically.
Diagnosis. Polynema grafinya females can be separated from females of other fully winged species of the draperi group by its relatively long ovipositor, at least 1.6× the length of the metatibia, that occupies almost the entire length of the gaster and is exserted beyond the gastral apex by at least 0.1× its own total length (Fig. 77).
Etymology. The species name is a noun in apposition; it means countess in Russian.
Remarks. Three females [NZAC] from South Island, New Zealand, labeled: “ NEW ZEALAND, OL, Kinloch SF, Dart R, Jan. 1981, Nothofagus For Broadleaf, grass P. totara Swept, J. S. Noyes, E. W. Valentine ” may also belong to this species; however, they have a relatively narrower fore wing (5.7–6.0× as long as wide) and a shorter ovipositor (about 1.5× as long as metatibia).
description: Published as part of Triapitsyn, Serguei V., 2021, Taxonomic review of Polynema (Hymenoptera: Mymaridae) in Australia and New Zealand, with descriptions of eight new species, pp. 151-200 in Zootaxa 4915 (2) on pages 178-180, DOI: 10.11646/zootaxa.4915.2.1, http://zenodo.org/record/4454213