Agyrtodes eucalypti Zwick 1979
Creators: name: Seago, Ainsley E.
Publisher: name: Zenodo
Year: 2009
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Leiodidae
Agyrtodes
Agyrtodes eucalypti

description: Agyrtodes eucalypti Zwick   
(Figs. 9, 46, 66, 86, 101)  
 Agyrtodes eucalypti Zwick 1979: 9. Holotype: male, labeled ‘‘ Tolmie north of Whitfield, Victoria, dry sclerophyll forest, under rotting bark of peppermint eucalypt, 1.ix.1972, P. Zwick’ ’ (NMV). Paratype: one male with identical data. (NMV)   
 Distribution. Australian Central Territory, southeastern New South Wales and northern Victoria (Fig. 101). Material examined listed in Appendix 1.   
 Diagnosis. This species is superficially similar in size and shape to A. crassus and A. globosus, new species (described here); in addition to the genitalic characters illustrated, A. eucalypti can be recognized by its lack of conspicuous pronotal punctation, presence of longitudinal strial impressions on the elytra, reduced mesoternal carina and non-expanded apical palpomeres.   
 Redescription. TBL 5 2.4 mm, EW 5 1.6 mm, PNW 5 1.4 mm, HW 5 0.6 mm. (TBL variation: 2.3–2.4 mm)   
Body (Fig. 9) large, ovoid, extremely convex, dorsum covered with fine golden pubescence. Coloration dark brown dorsally, often with pronotum darker brown, elytra reddish-brown with faint, indistinct transverse band of dark brown across disc; some teneral specimens fade to yellow when dried. Elytra with transverse strigae indistinct but marked by irregular puncture rows bearing short, stiff setae.  
Head narrow, shining, finely and shallowly punctate. Eyes small. Epistomal suture present, with extremely short stem (sometimes difficult to discern in darker-colored specimens). Clypeus small, transverse, subovate; labrum short, transverse, lightly cleft distally. Mandibles with dens blunt, mola slightly reduced in size. Maxillae with galea simple, slender, with apical cluster of weakly sclerotized hairs; lacinia with narrow spore-brush of minute teeth. Labial palpi with apical segment twice as long as subapical segment and bearing a basal cluster of minute, stout digitiform sensillae. Distal margin of ligula with four small digitiform sensillae. Mentum transverse, broadly trapezoidal, subrectangular.  
 Antennae (Fig. 46) robust, first two segments light brown, last eight segments darker reddish-brown, third segment intermediate in color. Segments 1 and 2 expanded, broader than segments 3–6; segment 1 longer than all subsequent segments. Segments 3–4 approximately equal in size, 5–6 shorter and slightly wider than 3–4. Segment 7 elongate, nearly triangular, 1.5 3 length of 6. Segment 8 globular, 0.5 3 length of 7; segments 9 and 10 wider than 7, segment 10 subquadrate, segment 11 scarcely longer than 9 and bluntly ovate.   
Pronotum strongly convex, shining, smooth, widest at posterior margin and abruptly narrowing and faintly trilobed anteriorly. Pronotal punctation extremely shallow, with minute punctures only at hair insertions. Hind angles blunt, square, very slightly inflexed, not wrapping tightly against elytral bases.  
 Elytra broad anteriorly, bluntly tapered at apices, with punctate, lightly impressed longitudinal striae easily viewed under oblique lighting. Sutural striae deeply punctate, slightly effaced in basal third. Transverse strigae less distinct than in other Agyrtodes, marked by puncture rows, confused at discal region adjacent to suture. Elytral vestiture of short, stiff setae, no discernable color variation.   
Prosternum short, procoxal separation normal, procoxal cavities ovoid, trochantins exposed. Hypomeron smooth, strongly concave. Mesosternum with median carina low, effaced anteriorly, visible only at mesocoxal separation and anterior (cervical) margin of mesoventrite anterior to transverse mesosternal suture. Mesepimera square laterally, obliquely rectangular, broadly rounded at coxal margins. Metasternum normal, bulging, deeply punctate; metacoxal cavities broadly triangular.  
Abdominal sterna normal, male sternum VII shallowly emarginate, male sternum VIII very deeply cleft. Sternum VII longer than sternites III–VI in both sexes.  
Legs robust, of similar brown to reddish-brown color as rest of venter, femora with sparse vestiture of fine, medium-length hairs. Procoxae ovoid, strongly projecting; mesocoxae ovate, not projecting; metacoxae triangular, with cavities for reception of femora broadly excavate. Tibial armature normal. Profemora and tibiae subequal in length; mesotibiae shorter than mesofemora and very faintly arched; metatibiae barely longer than metafemora. Male protarsi with basal three segments broadly expanded and bearing thick, dense ventral pads of tenent setae; first segment extremely broad, cordate, subequal in width to tibial apex. Male mesotarsi with first segment not laterally expanded but longer than subsequent three segments combined, with a few tenent setae ventrally. Female tarsi simple.  
Male genital segment (Fig. 66) elongate and narrow, parallel-sided, anterior apophysis produced and apically expanded. Aedeagus (Fig. 86) with median lobe extremely long and slender, compressed laterally; median lobe abruptly flared and bent dorsad at apex. Parameres extending only to apex of median lobe, broad, flattened, apically expanded and scoop-shaped, distally pointed, with clusters of long, gently curving setae on internal apical faces. Endophallus with relatively simple dentition, with cluster of well-sclerotized scale-shaped teeth distally and more slender, weakly sclerotized teeth proximally. Basal piece long but rarely as inflated as shown in Zwick (1979) Fig. 10. Female terminalia normal, usually concealed by elytra.  
 Natural History. Collected in window, flight intercept and malaise traps and by Berlese processing of rotten wood and leaf litter in Nothofagus and wet sclerophyll forests.
description: Published as part of Seago, Ainsley E., 2009, Revision Of Agyrtodes Portevin (Coleoptera: Leiodidae), pp. 1-73 in The Coleopterists Bulletin (mo 7) (mo 7) 63 on pages 35-37, DOI: 10.1649/0010-065x-63.sp7.1, http://zenodo.org/record/4912048