Amiota incurva Jones & Grimaldi 2022, sp. nov.
Creators: name: Jones, Lance E.
name: Grimaldi, David A.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Drosophilidae
Amiota
Amiota incurva

description: Amiota incurva, sp. nov.
Figures 17A–B, 19D, 23, 84A
DIAGNOSIS: Medium-sized fly (ThL 1.28– 1.44 mm), black; arista long, plumose; outer paraphysis large, laterally flattened, with apical hook curving inward, proximal appendage long, heavily sclerotized, perpendicular to paraphysis, forming a “0” in dorsal/ventral views; inner paraphyses symmetrical, each composed of a lobe with 3 spines, all curving inward medially, one spine overlaying the others and pointing anteriorly.
DESCRIPTION: Medium-sized fly (ThL 1.28– 1.44 mm), black; legs light yellow. Frons black dorsally, fading to brown ventrally. Facial marking large, depth 0.5× width, semicircular. Cheek whitish gray posteriorly. Palp brown. Arista: Long, plumose; longest branch D1; A.R. 0.53; 4 long dorsal, 2 long ventral branches, none pointed mediad/laterad; arista trunk with short microtrichia along entire length. Male genitalia: Epandrium complete at dorsal midline, this margin graded into membrane below. Cercus large, slightly crescentic, dorsally grading into membrane and epandrium; occupying most of space surrounded by epandrium. Surstylus oval; 10 prensisetae, apices blunt, closely arranged, comblike; with large, lateral fingerlike lobe adjacent to, and extending beyond, prensisetae; scattered setulae along distal margin and process. Subepandrial sclerite a simple plate, without a subepandrial appendage. Paraphyses symmetrical or nearly so, heavily sclerotized. Outer paraphysis large, laterally flattened, with apical hook curving inward; proximal appendage long, perpendicular to paraphysis, forming a “0” in dorsal/ventral views. Inner paraphysis symmetrical, each composed of a lobe with 3 spines, all curving inward medially; one spine overlying the others and pointing anteriorly. Aedeagal apodeme large, midsection and base wide and quadrangular (not constricted), length 1.5× width, slightly curved in lateral view, distal end slightly flared beyond midsection with shallow emargination. Hypandrium evenly narrow, U-shaped, small bulging gonopod in posterior view. Ejaculatory apodeme not studied. Head and thorax measurements: (n = 5; Am 377, 440, 713, 716, 1425) FL/FW 0.84 (0.75–0.96), EL/EW 1.32 (1.26–1.40), EL/ CW 21.81 (12.80–38), FML/FMW 0.38 (0.36– 0.42), PR /RR 0.53 (0.41–0.60), ThL 1.38 (1.28– 1.44 mm).
TYPE MATERIAL: Holotype: male: Sunnyside Canyon, Huachuca Mts., Ariz., [31.339651, -110.491677], VII-9-40, D.E. Hardy, Am 440, [glued to paper point, dissected]. Deposited in the American Museum of Natural History (AMNH).
OTHER MATERIAL EXAMINED: Mexico: Baja California: Sierra La Laguna, 1770–1850m, 1977- 08-29, leg. E. Fisher and R. Westcott, collected from flight trap near stream, 1♂ (Am 377*, CAS). Chiapas: 20–25 mi NE Huixtla, 3000′, 1969-06-01, leg. H.J. Teskey, 1♂ (Am 1425*, CNC); 20–25 mi N Huixtla, 3000′, 1969-06-01, leg. B. V. Peterson, 1♂ (Am 1266*, CNC). Hidalgo: Jacala 7 mi NE, 4600′, 1954-07-27, leg. J.G. Chillcott, “ Holotype ” [not published], 1♂ (Am 1374*, CNC). USA: Arizona: Cochise Co., campground nr. SWRS, 31.872506, -109.233923, 2019-07-19, leg. L.E. Jones and J.L. Hughes, swept around head, 1♂ (Am 713*, AMNH); Cochise Co., Vicinity of SWRS, 31.882018, -109.206636, 2019-07-18, leg. L.E. Jones and J.L. Hughes, swept around head, 1♂ (Am 716*, AMNH); Madera Cyn, Santa Rita Mts., 4–6000′, 1962-07-12, leg. H.E. Milliron, 1♂ (Am 1213*, CNC).
ETYMOLOGY: Formed from in and curva, Latin in feminine form for “curving inward.” In reference to the spines of the paraphyses which all curve medially.
DISTRIBUTION: Amiota incurva has a large distribution, from southeastern Arizona in the United States, throughout Mexico, from Baja California to Chiapas.
COMMENTS: This species exhibits the characteristic behavior of attraction to the eyes and face common to many Amiota.
description: Published as part of Jones, Lance E. & Grimaldi, David A., 2022, Revision Of The Nearctic Species Of The Genus Amiota Loew (Diptera: Drosophilidae), pp. 1-181 in Bulletin of the American Museum of Natural History 2022 (458) on pages 44-45, DOI: 10.1206/0003-0090.458.1.1, http://zenodo.org/record/7400026