cometadata/datacite-rtg-text-other-reclassification-records huggingface dataset:
Penthetria longa Handlirsch 1907
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Penthetria
Penthetria longa

description: ? Penthetria longa (Heer, 1849)
(Figs. 4, 24)
Protomyia longa Heer, 1849: 233.
Plecia longa (Heer) Loew, 1868: 184.
Penthetria longa (Heer) Handlirsch, 1907: 960.
Material: One male in NHMW, not the specimen depicted by Heer, probably non-type. The type described by Heer is a female. Heer mentioned one female in Graz and a detached wing in Vienna.
The specimen at hand is probably not part of the type series, and since it is a male while the type was a female, its identity is quite uncertain and the following redescription is very preliminary. The specimen is also too poorly preserved for a detailed description of its morphology.The genus placement of this species is also highly tentative, the wing venation might suggest genus Plecia. It is tentatively placed in Penthetria here due to its gracile habitus.
Male: Total length 13.1 mm
Head: Length 1.3 mm. Poorly preserved, dark brown.
Thorax: Length 3.3 mm. Dark brown.
Wing (Fig. 24): Length 11.6 mm, width 4.1 mm. Brownish fumose, veins rather weak.
Legs: Long and slender, brown. Fore femur length 3.6 mm, width 0.5 mm, fore tibia length 4.0 mm, width 0.33 mm, hind femur length 4.7 mm, width 0.4 mm, hind tibia length 4.0 mm.
Abdomen: Length 8.8 mm, width 2.1 mm. Brown.
Terminalia: Hypopygium width 1.3 mm. Gonostylus straight.
Female unknown.
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on pages 356-357, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Penthetria murchisonii
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Penthetria
Penthetria murchisonii

description: Penthetria murchisonii (Unger, 1842)
(Figs. 5, 25)
Bibio Murchisonii Unger, 1842: 426.
Bibiopsis Murchisonis (Unger, 1842) Heer, 1849: 230.
Penthetria Murchisoei (Unger, 1842) Loew, 1868: 186.
Penthetria Murchisoni (Unger, 1842) Handlirsch, 1907: 959.
Bibio murchisonis Unger, 1842 Evenhuis, 1994: 125.
Type material: Holotype (female, SLJG 77466), probable paratype (female) GBA 2009/016/0214.
Additional material: 1 female in NHMW (1852 XXIX 45), 1 unnumbered female (feeble impression). Heer (1849) mentioned 9 specimens, 8 in Vienna and one in Graz. Additionally, there is one female (1852. I.1011) in NHMW labeled as Bibiopsis murchisoni, this is Bibio sp.
Male unknown.
Female: Total length 9.9 mm
Head: Width 1.06 mm. Dark brown, oval, a little wider than long. Complex eye not prominent.
Thorax: Length 2.5 mm, width 1.9 mm. Brownish-black, rather short and wide, sulci fine, sharp and distinctive. Pile not apparent. Halter brown.
Wing (Fig. 25): Length 9.8 mm, width 3.2 mm. Brown fumose, rather narrow and pointed. Veins dark brown, distinctive, pterostigma indistinctive. Subcosta distinctive, long and straight. R 2+3 long, slightly oblique. Costa extends a little beyond apex of R 4+5. Vein measurements: Subcosta 5.8 mm, basal R 2.6 mm, distal R 1 3.6 mm, basal R s 1.8 mm, distal R s 0.7 mm, R 2+3 2.2 mm, R 4+5 4.4 mm, R-M 0.4 mm, basal M 3.0 mm, distal M 0.5 mm, M 1 3.0 mm, M 2 2.2 mm, M-Cu 0.25 mm, CuA 2.1 mm, CuA 1 3.9 mm, CuA 2 3.0 mm.
Legs: Black.
Abdomen: Length 6.7 mm, width 2.4 mm. Brownish-black, slightly conical.
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on pages 357-358, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Plecia bucklandi Loew 1868
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Plecia
Plecia bucklandi

description: Plecia bucklandi (Heer, 1849)
Protomyia buklandi (sic!) Heer, 1849: 238.
Plecia bucklandi (Heer) Loew, 1868: 184.
Penthetria bucklandi (Heer) Handlirsch, 1907: 954.
Syn. Protomyia elegans Heer, 1856: 36.
Syn. Protomyia gracilis Heer, 1856: 36.
Syn. Penthetria edwardsii (Oustalet, 1870)
Bibio edwardsii Oustalet, 1870: 130
Plecia edwardsii (Oustalet, 1870); Brongniart, 1878: XLVIII.
Syn. Plecia dilatativentris Théobald, 1937: 338 – 339.
The species was described from both Aix-en-Provence and Radoboj. We have not found any specimens labeled as types, nor have we seen any specimens from Radoboj identified as this species, apart from a specimen in NHMW, which is a hymenopteron, thus obviously mislabeled. The presence of this species from Radoboj seems dubious since the latest revision of the Radoboj locality’s age leaves a> 10 MYr time gap between the two localities. The species was redescribed by Skartveit & Nel (2017) based on a specimen from Aix-en-Provence labeled as Protomyia bucklandi in Oswald Heer’s handwriting. No holotype for the species has been explicitly selected – it would be reasonable to regard the specimen illustrated by Buckland (1837, Pl. 46, Fig. 11), with reference to which the species was formally described, as the holotype. This specimen may possibly be the specimen NHM In 2664, though the illustration in Buckland (1837) is reversed compared to In 2664. Possible reasons for this may be that Buckland’s illustration was based on the counterpart of In 2664, or that the drawing has been accidentally reversed during the engraving process. Though the species was described in a publication (Heer, 1849) dealing principally with the fossils of Öhningen and Radoboj, specimens from Aix-en-Provence were explicitly mentioned in the original description, and since no holotype was designated these should be eligible to be selected as lectotype. The species is named after the rev. William Buckland and should thus be bucklandi. The awkward spelling buklandi was used in the heading of the original description (Heer 1849: 238), however this must be a typographical error. The name is spelled bucklandi in the legends to the illustration (Heer 1849 Pl. XVI, Fig. 22) and subsequently (Heer 1856: 35).
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on page 358, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Plecia lugubris Heer 1849
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Plecia
Plecia lugubris

description: Plecia lugubris Heer, 1849
(Figs. 6, 26)
Plecia lugubris Heer, 1849: 209.
Penthetria lugubris (Heer) Handlirsch, 1907: 960.
Material: Possible type (female, NHMW) in Wien, fits one of the illustrations in Heer except it is reversed, maybe the counterpart of the specimen described by Heer.
Additional material: four females (CNHM 1824, 1825, 1828, 1831)
Female: Total length 7.9–10.3 mm (N=4).
Head: Length 0.84 mm (N=1), width 0.86–1.00 mm (N=2).
Thorax: Length 2.4–3.1 mm (N=3), width 1.9–2.4 mm (N=2). Brown, poorly preserved. Halter brown.
Wing: Length 8.4–9.7 mm (N=4), width 3.1–3.6 mm (N=3). Brownish fumose (colour preservation questionable), rather narrow. Veins fine and distinctive, pterostigma not apparent. Subcosta distinctive, long and straight. R 2+3 straight and oblique. Costa extends to apex of R 4+5. CuA and its continuation CuA 2 very strong and prominent, CuA 2 apically curved. CuP straight, basally strong and prominent. Vein measurements: basal R 2.1–2.7 mm (N=3), distal R 1 2.7–3.5 mm (N=2), basal R s 1.5 –2.2 mm (N=3), distal R s 1.9 –2.0 mm (N=2), R 2+3 0.8–1.3 mm (N=2), R 4+5 2.9–3.1 mm (N=2), R-M 0.41–0.44 mm (N=3), basal M 2.9–4.2 mm (N=3), distal M 0.5–0.7 mm (N=3), M 1 3.2–4.2 mm (N=2), M 2 2.5–3.2 mm (N=2), CuA 1.9–2.3 mm (N=2),CuA 1 3.5–3.7 mm (N=2), CuA 2 2.6–2.7 mm (N=2).
Legs: Brown, slender.
Abdomen: Length 5.4–7.1 mm (N=3), width 1.9–2.3 mm (N=2). Brown, cylindrical, relatively slender.
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on pages 358-359, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Plecia lygaeoides Loew 1868
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Plecia
Plecia lygaeoides

description: Plecia lygaeoides (Heer, 1849)
(Figs. 7, 27)
Bibio lignarius Germar, 1837 sensu Unger, 1841: 427 (nec Germar), pro parte
Protomyia lygaeoides Heer, 1849: 232.
Plecia lygaeoides (Heer) Loew, 1868: 185.
Penthetria lygaeoides (Heer) Evenhuis, 1994: 129.
Protomyia latipennis Heer, 1849: 237 (syn.n.)
Plecia latipennis (Heer) Loew, 1868: 184.
Penthetria latipennis (Heer) Handlirsch, 1907: 960.
Type material: Holotype (female) SLJG 77465. Type and depicted specimen, P. latipennis (female), SLJG (no number). The latter specimen is also labeled as Rhipidia extincta Unger, but this must be a mistake since the specimen does not at all fit the original description of Rhipidia extincta.
Additional material: Female (CNHM 1833).
The type specimen was redescribed by Skartveit & Nel (2017: 51).
The species is habitually similar to Penthetria cimicoides, but easily distinguished from this species by the short vein R 2+3.
Female: Total length 10.1–10.2 mm (N=2)
Head: Length 1.2 mm (N=1), width 1.1 mm (N=2). Width Black, poorly preserved.
Thorax: Length 2.4–2.6 mm (N=3), width 2.1–2.4 mm (N=2). Black.
Wing (Fig. 27): Length 9.3–10.3 mm, width 3.4–3.8 mm, length/width = 2.7–2.9 (N=3). Dark brown fumose, relatively wide. Costa distinctively curved, costal cell wide. Subcosta relatively weak. R 2+3 oblique, with a slight bend in the middle. Pterostigma oval, not very distinctive. Costa extends a little beyond apex of R 4+5. Wing vein measurements: basal R 2.7–3.0 mm (N=3), distal R 1 3.1–3.8 mm (N=3), basal R s 1.7 –1.8 mm (N=3), distal R s 1.5 –1.8 mm (N=3), R 2+3 1.0– 1.2 mm (N=3), R 4+5 2.7–3,0 mm (N=3), R–M 0.5 mm (N=2), basal M 3.0 mm (N=1), distal M 0.9–1.1 mm (N=2), M 1 3.7 mm (N=1), M 2 3.2 mm (N=2), CuA 2.2–2.5 mm (N=2), CuA 1 4.5 mm (N=1), CuA 2 3.2 mm (N=1).
Legs: Black, slender. Fore tibia length 2.4 mm (N=1). Hind femur length 2.8 mm (N=1), slightly thickened, hind tibia 2.8–3.2 mm (N=3), tibia and tarsus quite slender.
Abdomen: Length 6.8–6.9 mm (N=2). Black, oval, poorly preserved in P. latipennis type, covered by wings in P. lygaeoides type.
The species has been recorded from both Radoboj and French Oligocene sites (Skartveit & Nel, 2017). Given that these sites are now thought to be more than 10 MYr older than Radoboj, it seems questionable if the two species are the same. Should they be found to be different, the Radoboj species will keep the name Plecia lygaeoides whereas the French Oligocene species should be called Plecia gracilenta (Heyden & Heyden, 1865).
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on pages 359-361, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Plecia morio Skartveit & Nel 2017
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Plecia
Plecia morio

description: Plecia morio (Heer, 1849)
Bibio morio Heer, 1849: 222.
Plecia morio (Heer) Skartveit & Nel, 2017
Type material: We have not found any type material of this species from Radoboj; specimens identified as this species have turned out to be various other species, none of which fit Heer’s description of the species well. Pongracz (1928: 183) attempted to redescribe the species based on a specimen from «wiener Geologischen Bundesanstalt» (currently, GBA). Pongracz’s figure 38 does not resemble any known genus and obviously is based on misinterpretation of the wing venation. Though the species was described in a publication dealing principally with Öhningen and Radoboj, specimens from Aix-en-Provence were mentioned in the original description (Heer 1849: 222). The species was redescribed by Skartveit & Nel (2017) based on a specimen from Aix-en-Provence, we have not succeeded in locating any specimens from Radoboj, though Heer’s illustration (Heer 1849, pl. XVI, Fig. 13) suggests they would at least be habitually similar to the Aix specimens. Since there is now thought to be a more than 10 MYr difference in age between the localities of Aix-en-Provence and Radoboj, the presence of the same species from both localities seems dubious.
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on page 361, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Bibio crassinervis Skartveit & Pika 2014
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Bibio
Bibio crassinervis

description: Bibio crassinervis Skartveit & Pika, 2014
(Fig. 8)
This species was formally described from Öhningen by Skartveit & Pika (2014), though the name is apparently a cabinet name used by Oswald Heer. The available specimen was identified as Bibio lividus Heer, but does not belong to the same species as the holotype of Bibio lividus.
Material: 1 female (CNHM 1837)
Description of the Radoboj specimen
Female (N=1) Total length 11.6 mm.
Head: Length 1.5 mm. Complex eye of medium size, not prominent. Palp black, rather short and stout. Antenna not preserved.
Thorax: Length 3.2 mm. Dorsum entirely brownish-black. The microsculpture on mesonotum is fine and meshy, laterally with slight, transverse wrinkles. Halter pale brownish.
Wing: Length 9.5 mm, width 3.5 mm, length/width = 2.8. Hyaline with brown, distinctive R-, M- and CuAveins. Pterostigma rather small, black, distinctive. Humeral crossvein apparently absent, Subcosta unpigmented but distinctive, straight. Costa extends to half-way between apices of R 4+5 and M 1. Radial sector about three times as long as crossvein R-M. Crossvein M-Cu joins M 2 some distance beyond furcation of M, M 2 bent at this point. CuA 1 and CuA 2 apically straight. CuP fine but distinctive, straight. Vein measurements: Basal R 3.7 mm, distal R 1 2.0 mm, R s 1.1 mm, R 4+5 4.2 mm, R-M 0.37 mm, basal M 3.7 mm, distal M 1.3 mm, M 1 3.3 mm, M 2 3.0 mm, M-Cu 0.55 mm, CuA 4.5 mm, CuA 1 3.5 mm, CuA 2 2.3 mm. Pterostigma 1.35 mm long, 0.50 mm wide.
Legs: Yellowish-brown.
Abdomen: Blackish-brown. Length 8.5 mm, width 3.0 mm. Tergites with fine, meshy microsculpture.
Terminalia: Last tergite apparently with V-shaped, mesal indentation.
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on pages 361-362, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Bibio enterodelus Unger 1841
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Bibio
Bibio enterodelus

description: Bibio enterodelus Unger, 1841
(Figs. 9–10, 28–30)
Bibio enterodelus Unger, 1841: 428.
Bibio lividus Heer, 1849: 223. Syn.n.
Type material: Holotype Bibio enterodelus (female) (SLJG 77425). Heer mentioned just a single specimen of Bibio enterodelus. Holotype Bibio lividus (female) (GBA 2009/016/0691). Heer (1849: 224) mentioned one specimen from “Cabinet zu Wien” which is currently in GBA.
Additional material: One female (CNHM 1838).
Description
Male unknown.
Female: Total length 15.0– 15.2 mm (N=2).
Head (Fig. 29): Length 1.7 mm (N=2). Brown. Antenna rather slender. Ocellar tubercle prominent.
Thorax: Length 5.1 mm (N=1). Brown, probably faded colour. Halter light brown.
Wing (Fig. 28): Length 12.3–12.7 mm (N=3), width 4.0– 4.4 mm (N=3). Light brownish, veins apparently concolourous with membrane. Relatively slender. Veins rather strong and prominent. Basal R s 1.6 –2.0* as long as R-M. Fork of M relatively short, M-veins apically straight. Crossvein Bm-Cu meets M 2 slightly beyond fork of M. CuA 2 apically strongly curved. Vein measurements (N=3): Subcosta 6.3–6.6 mm, basal R 3.8–4.9 mm, distal R 1 2.4–2.6 mm, basal R s 1.1 –1.5 mm, distal R s (=R 2–5) 5.4–6.0 mm, R-M 0.4–0.8 mm, basal M 3.7–4.6 mm, distal M 0.8–1.0 mm, M 1 4.5–4.9 mm, M 2 3.6–3.8 mm, M-Cu 0.6–1.2 mm, CuA 2.4–2.8 mm, CuA 1 4.5–5.7 mm, CuA 2 3.3–4.5 mm.
Legs: Fore femur (Fig. 29) length 3.1 mm, width 0.8 mm (N=1). Hind femur (Fig. 30) length 4.0– 4.7 mm, width 0.8–0.9 mm (N=2), hind tibia length 4.5 mm, width 0.5 mm (N=1).
Abdomen: Length 9.7–12.0 mm (N=2), width 3.6 mm (N=1). Pale brown, cylindrical.
Discussion: The holotype specimen of Bibio lividus is rather uniformly brown except that the tibiae are darker, not coloured like the recent species Bibio hortulanus as claimed by Heer (1849: 223). The colour is unlikely to be original.
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on page 362, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Bibio firmus Heer 1849
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Bibio
Bibio firmus

description: Bibio firmus Heer, 1849
(Figs. 11, 31)
Bibio firmus Heer, 1849: 226.
Type material: Holotype (female) SLJG 77471.
Description:
Male unknown.
Female: Total length 10.6 mm. The specimen is entirely grayish brown, details difficult to make out as it is a very vague impression.
Head: Length 1.4 mm. Antenna quite thin, flagellum possibly 7-segmented.
Thorax: Length 3.0 mm, width 2.5 mm. No details possible to see in type specimen.
Wing (Fig. 31): Length 7.6 mm, width 2.4 mm, length/width = 3.1. Crossvein R-M nearly equal to basal R s in length.
Legs: Hind legs quite slender, otherwise no details possible to see in type specimen.
Abdomen: Length 6.3 mm, width 3.5 mm. No details possible to see in type specimen.
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on page 364, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Bibio batheri Pongracz 1928
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Bibio
Bibio batheri

description: Bibio batheri Pongracz, 1928: 180, fig. 44.
Described from two specimens in the Natural History Museum, London. The senior author did not find these specimens during a visit to the museum. A medium-sized Bibio with body length 9 mm, wing length 6 mm. Sex of type specimens not stated. Figure (fig. 44) given by Pongracz does not fit any known genus, interpretation of venation probably wrong.
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on page 369, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Bibio incrassatus Heer 1849
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Bibio
Bibio incrassatus

description: Bibio incrassatus Heer, 1849: 221.
Described from a single specimen from “k.k. montan. Cabinet” Vienna. The specimen was depicted by Heer (Pl.
XVI Fig. 12) but it is difficult to tell what it is. Heer’s figure may suggest the presence of a vein R 2+3 and a vertical crossvein R-M, in which case the species should be placed in the genus Plecia, possibly as a synonym of Plecia
morio. On the other hand, if this interpretation of the wing venation is incorrect, it may habitually resemble Bibio
giganteus. Pongracz (1928: 183) considered Bibio morio and Bibio incrassatus as probable synonyms.
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on page 369, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Bibio linearis Heer 1849
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Bibio
Bibio linearis

description: Bibio linearis Heer, 1849: 214.
Heer (1849: 215) mentioned one specimen from “k.k. montan. Cabinet”. We could not find the type in NHMW nor GBA. It is a large female, total length 16 mm, wing length 11.7 mm. One female specimen in CNHM (CNHM 1836) is labeled as this species but does not fit Heer’s description.
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on page 369, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Bibio marmoratus Pongracz 1928
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Bibio
Bibio marmoratus

description: Bibio marmoratus Pongracz, 1928: 186.
Described from a single specimen (female) in Hungarian National Museum. The Radoboj material in the Hungarian National Museum was destroyed by a fire in 1956 (Alfréd Dulai, in litt.). The specimen was illustrated by Pongracz (1928: Fig. 49) and had a total length of 14 mm and wing length 10 mm. Judging from this illustration the species is probably not a bibioninid, the habitus (slender antennae, slender legs with rather long spurs, elongate coxae, wing with dark patterning, pointed abdomen) suggests Mycetophilidae.
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on page 370, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Bibio ungeri Heer 1849
Creators: name: Skartveit, John
name: Krizmanić, Katarina
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Bibionidae
Bibio
Bibio ungeri

description: Bibio ungeri Heer, 1849: 218. (Figs. 18, 38)
Bibio lignarius Germar, 1837 sensu Unger, 1842, pro parte
This species is not a bibionid, the type (SLJG, no catalogue number) is a brachyceran, possibly Xylophagidae. This was commented on already by Heer (1849: 218) who nevertheless placed the species in genus Bibio based on the wing venation. This is not at all clear in the holotype but there appears to be closed discal and anal cells in the wing. From the specimen label it is clear that Unger did not intend to describe the specimen as a new species, but (mis)identified it as Bibio lignarius Germar. Both head and legs are missing in the type specimen, and the wing venation is very difficult to interpret, so this species is likely to remain a nomen dubium.
description: Published as part of Skartveit, John & Krizmanić, Katarina, 2020, Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia, pp. 351-378 in Zootaxa 4759 (3) on page 370, DOI: 10.11646/zootaxa.4759.3.3, http://zenodo.org/record/3741178
Anoplodelphys Lafargue and Laubier 1978
Creators: name: Boxshall, Geoff A.
name: Marchenkov, Andrey
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Cyclopoida
Notodelphyidae
Anoplodelphys

description: Genus Anoplodelphys Lafargue and Laubier, 1978
The two new species described below were all collected from didemnid ascidian hosts from around the coast of the African continent. Both are strongly modified, exhibiting numerous limb losses. Placing such secondarily simplified parasites in a genus can sometimes be problematic since independent reductions and losses in different lineages can result in convergence on highly reduced morphologies. The new species are placed in the genus Anoplodelphys, which was erected by Lafargue and Laubier (1978a) to accommodate three species of highly transformed notodelphyid copepods found in colonial ascidian hosts belonging to the genus Didemnum. This placement is supported by the phylogenetic analysis given below (following the descriptions).
description: Published as part of Boxshall, Geoff A. & Marchenkov, Andrey, 2007, A revision of the Brementia - group of genera (Copepoda: Notodelphyidae), with descriptions of a new genus and four new species, pp. 37-68 in Zootaxa 1459 on page 40, DOI: 10.5281/zenodo.176361
Anoplodelphys corneci Lafargue and Laubier 1978
Creators: name: Boxshall, Geoff A.
name: Marchenkov, Andrey
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Cyclopoida
Notodelphyidae
Anoplodelphys
Anoplodelphys corneci

description: Anoplodelphys corneci Lafargue and Laubier, 1978
Material examined: Holotype female, reg. no. ZMA CO.102.603. Two females in Monniot collection, from Didemnum maculosum (Milne Edwards, 1841) collected at Dinard, France; reg. no. MNHN-Cp2319.
Differential diagnosis: Body vermiform with indistinct segmentation. Cephalosome with narrow frontal margin bearing rostrum ventrally. Rostrum inflated, forming simple lobe, without accessory median lobe. Post-rostral median lobe absent. Labrum inflated, forming elongate tapering lobe. Lateral margin of cephalosome expanded ventrally to form ridge-like swelling produced into process at posterior end. Adjacent antennomedial lobe drawn out into long posterior process. Metasome elongate with about 20% of length extending posterior to origin of leg 4: metasomal somites bearing legs ventrally near lateral margins. No mid-ventral processes present between legs. Urosome vestigial; located terminally, with median anal incision; bearing partly incorporated caudal rami armed with setae distally.
Antennule unsegmented with setose apex. Antenna reduced, 2-segmented, with triangular basal segment and distal subchela incorporating sclerotised distal claw, armed with 2 vestigial setal elements. Mandible to maxilliped lacking. Legs 1–4 biramous; each comprising short basal part (derived from protopod), largely incorporated into somite, carrying unsegmented, lobate rami. Leg 5 not observed.
Body length of female 0.85–3.90 mm. Male unknown.
Remarks: Examination of the holotype of A. corneci (ZMA CO.102.603) confirmed the original description in every respect. The new material comes from the same host species, Didemnum maculosum, collected in the same region. This species is readily characterised by the small size and simple structure of the rostrum (about the same width as the base of the labrum) and by the distribution of legs 1–4 along the metasome.
description: Published as part of Boxshall, Geoff A. & Marchenkov, Andrey, 2007, A revision of the Brementia - group of genera (Copepoda: Notodelphyidae), with descriptions of a new genus and four new species, pp. 37-68 in Zootaxa 1459 on page 46, DOI: 10.5281/zenodo.176361
Anoplodelphys galli Lafargue and Laubier 1978
Creators: name: Boxshall, Geoff A.
name: Marchenkov, Andrey
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Cyclopoida
Notodelphyidae
Anoplodelphys
Anoplodelphys galli

description: Anoplodelphys galli Lafargue and Laubier, 1978
Material examined: none.
Differential Diagnosis: Body with indistinct segmentation. Cephalosome with narrow frontal margin bearing rostrum. Rostrum forming simple, inflated lobe, without accessory median lobe. Post-rostral median lobe absent. Labrum inflated, forming elongate tapering lobe. Lateral margin of cephalosome expanded ventrally to form ridge-like swelling produced into process at posterior end. Antennomedial lobe lacking. Metasome elongate with about 20% of length extending posterior to level of origin of leg 4: metasomal somites bearing legs ventrally near lateral margins. No mid-ventral processes present between legs. Urosome small, unsegmented, located terminally, with median anal incision; bearing partly incorporated caudal rami armed with setae distally. Surface of body, rostrum, labrum, cephalosomic processes and legs densely ornamented with surface setules arranged in triads (with long central setule and short setules either side).
Antennule unsegmented. Antenna 2-segmented, with triangular basal segment and distal subchela incorporating sclerotised distal claw, armed with 2 vestigial setal elements. Mandible to maxilliped lacking. Legs 1–4 biramous; each comprising short basal part (derived from protopod), largely incorporated into somite, with unsegmented, lobate rami. Ratios of gaps between legs along metasome as in Table 2. Leg 5 not observed.
Body length of female 0.70–2.85 mm. Male unknown.
Remarks: Lafargue and Laubier (1978a) concluded that A. galli and A. incerta (both forms) could be separated only by the details of their cuticular ornamentation. They showed that a triad-setule arrangement (a large central setule with 1 shorter setule on each side) was characteristic for A. galli. Their decision to base species-level discrimination on cuticular ornamentation patterns has not been tested subsequently, although their initial conclusions were based on the study of over 40 specimens (30 specimens of A. galli and 13 specimens of A. incerta (both forms) from across the four Didemnum host species).
We regard the spacing of the legs along the metasome as an important character for species discrimination. Anoplodelphys galli and Anoplodelphys cf. galli from Didemnum peyrefittense, D. maculosum, D. fulgens and D. protectum all share a similar pattern of leg distribution, with legs 1 and 2 separated by only 9– 10% of the overall metasomal length, as compared to 18% in A. incerta (Table 2). This character is used here to restrict the definition of A. incerta and to distinguish it from A. galli.
description: Published as part of Boxshall, Geoff A. & Marchenkov, Andrey, 2007, A revision of the Brementia - group of genera (Copepoda: Notodelphyidae), with descriptions of a new genus and four new species, pp. 37-68 in Zootaxa 1459 on pages 47-48, DOI: 10.5281/zenodo.176361
Anoplodelphys galli
Creators: name: Boxshall, Geoff A.
name: Marchenkov, Andrey
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Cyclopoida
Notodelphyidae
Anoplodelphys
Anoplodelphys galli

description: Anoplodelphys cf. galli
Material examined: none.
Remarks: We are uncertain of the status of the specimens collected from D. fulgens, D. protectum, D. peyrefittense and D. maculosum originally described as the “atypical” form of A. incerta by Lafargue and Laubier (1978a: Figs 3 H–J, 4A–E, 5A–B). These specimens share with A. galli characters such as body form, and the size and disposition of legs 1–4 along the body (Table 2). However, they differ in the fine-scale arrangement of setules comprising the cuticular ornamentation on the body surface: the setules are isolated and irregularly arranged, or grouped in twos and threes, but have never been observed in the particular triad arrangement found in A. galli. In addition, the sclerotised apical claw of the antenna has coarser marginal teeth compared to the fine teeth present near the tip only in A. galli.
description: Published as part of Boxshall, Geoff A. & Marchenkov, Andrey, 2007, A revision of the Brementia - group of genera (Copepoda: Notodelphyidae), with descriptions of a new genus and four new species, pp. 37-68 in Zootaxa 1459 on page 48, DOI: 10.5281/zenodo.176361
Paranoplodelphys Boxshall & Marchenkov, 2007, n. gen.
Creators: name: Boxshall, Geoff A.
name: Marchenkov, Andrey
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Cyclopoida
Notodelphyidae
Paranoplodelphys

description: Paranoplodelphys n. gen.
Diagnosis: Body highly transformed; lacking external expression of body segmentation. Cephalosome with broad, strongly-convex frontal margin bearing paired antennule vestiges ventrolaterally. Rostrum lacking. Labrum forming elongate, posteriorly-directed lobe. Metasome extending posteriorly beyond end of dorsally located urosome. Urosome incorporating caudal rami, presented by caudal setae. Surface of body, labrum and legs densely ornamented with surface setules.
Antennules present as pair of wrinkled lobes originating close to labrum on ventral side of cephalosome. Antenna absent. Oral region lacking any vestiges of mouthparts. Leg 1 biramous; rami represented by unsegmented, rounded lobes; exopodal lobe reduced, laterally-directed; endopodal lobe large. Leg 2 uniramous, a simple lobe. Legs 3 and 4 absent. Leg 5 lobe bearing 2 setae.
Type species: Paranoplodelphys simplex n. gen. et n. sp. (by original designation)
Etymology: The genus name refers to the similarity between the new genus and Anoplodelphys.
Remarks: The phylogenetic analysis presented below (Fig. 14) places this new species as the sister-taxon to a lineage comprising Achelidelphys, Cephalodelphys and Syndelphys. It does not cluster with Anoplodelphys species but it lacks the stellate body form (defined by the apomorphic states of characters 1 and 2) of the Achelidelphys – Cephalodelphys – Syndelphys lineage. A new genus is established to accommodate this new species which represents the most extreme reduction known within the lineage. It retains only the antennules on the cephalosome and all other cephalosomic limbs are lost. The legs are also reduced: leg 1 is the only biramous leg, leg 2 is uniramous and legs 3 and 4 are lost. Combined with all these reductions is the retention of a well-defined leg 5, which is apparently lost in many of the less reduced relatives.
description: Published as part of Boxshall, Geoff A. & Marchenkov, Andrey, 2007, A revision of the Brementia - group of genera (Copepoda: Notodelphyidae), with descriptions of a new genus and four new species, pp. 37-68 in Zootaxa 1459 on pages 48-49, DOI: 10.5281/zenodo.176361
Brementia balneolensis Chatton and Brement 1915
Creators: name: Boxshall, Geoff A.
name: Marchenkov, Andrey
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Cyclopoida
Notodelphyidae
Brementia
Brementia balneolensis

description: Brementia balneolensis Chatton and Brément, 1915
Material examined: none.
Differential Diagnosis: Body eruciform, with indistinct segmentation. Cephalosome with narrow frontal margin bearing anteroventrally-directed rostrum. Rostrum forming simple lobe, without accessory median lobe. Post-rostral median lobe absent. Labrum a hemispherical lobe. Lateral margin of cephalosome expanded ventrally to produce ridge-like swelling. Metasome with about 30% of length extending posterior to origin of leg 4. Metasomal somites bearing ventrally-directed legs near lateral margins, lacking mid-ventral processes between legs. Urosome small, narrow and carrying partly-incorporated caudal rami. Caudal rami lobate, armed with setal remnants distally. Surface of body, rostrum, labrum, cephalosomic processes and legs densely ornamented with surface setules.
Antennule tapering with traces of segmentation, with apical segment setose. Antenna indistinctly 3-segmented, with distal segment incorporating reduced distal claw, and bearing 2 vestigial setal elements. Mandible forming large cylindrical lobe with vestigial gnathobase proximally bearing reduced setal elements; mandibular palp with bilobed apex. Maxillule represented by cylindrical lobe, bifid at apex. Maxilla indistinctly segmented, syncoxal part forming triangular lobe bearing vestiges of enditic armature along medial margin; basis bearing reduced claw and vestige of endopod. Maxilliped reduced to tiny conical lobe.
Legs 1 to 3 biramous, ventrally-directed; each comprising long basal part (derived from protopod), carrying short, unsegmented rami distally. Leg 4 reduced, largely incorporated into body somite, weakly bilobed distally, with lobes representing rami. Leg 5 not observed.
Body length of female approximately 2.35 mm. Male unknown.
Remarks: This is a strongly transformed copepod from the colonial ascidian Didemnum commune on the Mediterranean coast of France. It was redescribed to modern standards by Laubier and Lafargue (1974), based on new material collected from Polysyncraton canetense (Brément, 1913) in the same area. Despite the transformed body and swimming legs, this copepod retains a full set of mouthparts, although all are reduced in comparison with basal notodelphyids (Boxshall & Halsey 2004).
description: Published as part of Boxshall, Geoff A. & Marchenkov, Andrey, 2007, A revision of the Brementia - group of genera (Copepoda: Notodelphyidae), with descriptions of a new genus and four new species, pp. 37-68 in Zootaxa 1459 on page 51, DOI: 10.5281/zenodo.176361
Brementia illgi Laubier and Lafargue 1974
Creators: name: Boxshall, Geoff A.
name: Marchenkov, Andrey
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Cyclopoida
Notodelphyidae
Brementia
Brementia illgi

description: Brementia illgi Laubier and Lafargue, 1974
Material examined: none.
Differential Diagnosis: Body with indistinct segmentation. Cephalosome with narrow frontal margin bearing ventrally-directed rostrum. Rostrum forming simple lobe, without accessory median lobe. Post-rostral median lobe absent. Labrum a small lobe, partly concealed beneath rostrum. Lateral margin of cephalosome expanded ventrally to produce ridge-like swelling. Metasome with about 30% of length extending posterior to origin of leg 4: metasomal somites bearing legs near lateral margins. Mid-ventral processes present between legs 2, 3 and 4. Urosome vestigial, largely incorporated into metasomal trunk. Caudal rami lobate, fused to urosome at base, armed with setal remnants distally. Surface of body, rostrum, labrum, cephalosomic processes, and legs densely ornamented with surface setules.
Antennule unsegmented, lobate with broad base, tapering to setose apex. Antenna modified, with traces of segmentation; distal segment reflexed, bearing vestiges of sclerotised distal claw and other setal elements. Mandible represented by tapering lobate palp, lacking gnathobase. Tiny paired lobes located posterior to mandibles possibly representing maxillules. Maxillae and maxillipeds lacking. Legs 1–3 biramous, ventrolaterally-directed; each comprising elongate basal part (derived from protopod), carrying elongate, unsegmented but superficially annulate rami distally. Legs 1–3 each bearing small outgrowths ventrally on protopodal part. Leg 4 reduced, largely incorporated into body somite, weakly bilobed distally, with reduced lobes representing rami. Leg 5 not observed.
Body length of female approximately 2.60 mm. Male unknown.
Remarks: This species was described from Polysyncraton canetense by Laubier and Lafargue (1974), who commented on the numerous differences between this species and B. balneolensis, the type species of the genus. They refrained from establishing a new genus of accommodate this species because of the poor state of knowledge of the phylogeny of the “ophioseidimorph” Notodelphyidae. The phylogenetic analysis performed here (Fig. 14) supports a monophyletic Brementia comprising both B. balneolensis and B. illgi.
description: Published as part of Boxshall, Geoff A. & Marchenkov, Andrey, 2007, A revision of the Brementia - group of genera (Copepoda: Notodelphyidae), with descriptions of a new genus and four new species, pp. 37-68 in Zootaxa 1459 on pages 51-52, DOI: 10.5281/zenodo.176361
Achelidelphys drachi Lafargue and Laubier 1978
Creators: name: Boxshall, Geoff A.
name: Marchenkov, Andrey
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Cyclopoida
Notodelphyidae
Achelidelphys
Achelidelphys drachi

description: Achelidelphys drachi Lafargue and Laubier, 1978
Material examined: Holotype female, reg. no. ZMA CO. 106.637.
Differential Diagnosis: Body highly transformed, stellate (Fig. 8 A–B); segmentation indistinct with segmental boundaries marked by superficial folds. Cephalosome with frontal margin merging into elongate, tapering, anterolaterally-directed antennulary lobes. Rostrum elongate, anteroventrally-directed lobe (Fig. 8 B), without accessory median lobe. Post-rostral median lobe absent. Labrum inflated, forming rounded, slightly-elongate lobe. Lateral margin of cephalosome not produced into ridge-like swellings. Paired antennomedial processes present on ventral surface of cephalosome, anterior and lateral to labrum. Metasome truncated, with urosome hardly extending posterior to origin of leg 4. Legs 1–4 transformed, originating laterally, each occupying entire margin of somite and produced laterally, giving body stellate appearance. Mid-ventral metasomal process present between second legs. Urosome vestigial, located terminally; bearing partly incorporated caudal rami (Fig. 8 C). Surface of body, rostrum, labrum, cephalosomic processes and legs densely ornamented with surface setules.
Antennules forming elongate, tapering lobe directed antero-laterally on either side of frontal margin of cephalosome. Antenna to maxilliped lacking. Legs 1–3 biramous; rami represented by unsegmented, elongate lobes: exopodal lobes laterally-directed, with broad base, carrying elongate endopod ventrally. Leg 4 uniramous, comprising tapering, posterolaterally-directed lobe representing exopod. Exopodal lobes of legs 2–4 each housing internal expansion of uterus, containing eggs visible through body wall. Leg 5 absent.
Body length of female 2.50 mm. Male unknown.
Remarks: This species carries an additional tiny unpaired process on the ventral midline about at the level of the first pair of legs. The homology of this process is uncertain (i.e. whether it is a serial homologue of the processes present between legs 2–4 in A. steinitzi, for example). This species can be distinguished from its congeners by the extreme length of the tapering lobes derived from the antennules, rostrum and leg rami. The paired antenno-medial processes are shared only with A. ampla among the members of Achelidelphys.
description: Published as part of Boxshall, Geoff A. & Marchenkov, Andrey, 2007, A revision of the Brementia - group of genera (Copepoda: Notodelphyidae), with descriptions of a new genus and four new species, pp. 37-68 in Zootaxa 1459 on page 55, DOI: 10.5281/zenodo.176361
Achelidelphys ampla Lafargue and Laubier 1977
Creators: name: Boxshall, Geoff A.
name: Marchenkov, Andrey
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Maxillopoda
Cyclopoida
Notodelphyidae
Achelidelphys
Achelidelphys ampla

description: Achelidelphys ampla Lafargue and Laubier, 1977
Material examined: Holotype female, reg. no. ZMA CO. 102.629.
Differential Diagnosis: Body highly transformed, stellate; segmentation indistinct with segmental boundaries marked by superficial folds. Cephalosome with frontal margin merging into tapering laterally-directed antennulary lobes. Rostrum anteroventrally-directed, simple short lobe, without accessory median lobe. Postrostral median lobe absent. Labrum inflated, forming elongate, posteriorly-directed lobe. Lateral margin of cephalosome not produced into ridge-like swellings. Paired antennomedial processes present on ventral surface of cephalosome, anterior and lateral to labrum. Metasome truncated, with urosome hardly extending posterior to origin of leg 4. Legs 1–4 transformed, originating laterally, each occupying entire margin of somite; produced laterally giving body a stellate appearance. Mid-ventral metasomal processes between legs absent. Urosome vestigial, located terminally; bearing partly incorporated caudal rami armed with vestigial setae (Fig. 9 A). Surface of body, rostrum, labrum, cephalosomic processes, and legs densely ornamented with surface setules.
Antennules forming short, tapering lobe on either side of frontal margin of cephalosome; surface ornamented with short spinules and vestiges of apical setae present (Fig. 9 B). Antenna to maxilliped lacking. Legs 1–3 biramous; rami represented by unsegmented, tapering lobes: exopodal lobe laterally-directed, with broad base and unarmed tip (Fig. 9 C), carrying smaller endopodal lobe ventrally. Leg 4 uniramous, comprising tapering, posterolaterally-directed lobe representing exopod. Uterus not expanded into exopods of legs. Leg 5 absent.
Body length of female 1.55–2.60 mm. Male unknown.
Remarks: Achelidelphys ampla has a compact uterus which does not extend into the exopodal lobes of the legs when full of stored eggs. This character, in combination with the lack of any median ventral processes on the metasome, serves to distinguish A. ampla from all other species. In A. ampla there is a pair of ventral process on the cephalosome originating anterior and lateral to the labrum. The origin of these antenno-medial processes is uncertain but we tentatively interpret them as homologous with the antenno-medial processes present in most Anoplodelphys species. A similar pair of antenno-medial processes is present in A. drachi but is absent in all other Achelidelphys species.
description: Published as part of Boxshall, Geoff A. & Marchenkov, Andrey, 2007, A revision of the Brementia - group of genera (Copepoda: Notodelphyidae), with descriptions of a new genus and four new species, pp. 37-68 in Zootaxa 1459 on pages 55-57, DOI: 10.5281/zenodo.176361