Madates heissi RIDER 2006, nov.sp.
Creators: name: RIDER, D. A.
Publisher: name: Zenodo
Year: 2006
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Pentatomidae
Madates
Madates heissi

description: Madates heissi nov.sp. (Figs 3, 10-13)
Description. Ovoid, black with yellow markings (Fig. 3).
Head subtriangular, lateral margins nearly straight, slightly reflexed, tapering anteriorly to rounded apex; juga apically curving over tylus, and almost contiguous. Eyes stylate, basal area of each eye stalk shallowly impressed. Dorsal surface of head nearly impunctate, with at most a few sparse punctures on vertex, and a few oblique, rugulose lines on juga; coloration yellow with medial, longitudinal band consisting of tylus and very narrow mesial margins of juga, a small triangular spot in front of each compound eye, and base of head including stylate eyes black; apex of each lateral triangular spot does not extend cephalad of adjacent antennifer, or it does so only submarginally. Antennae entirely black, segment I reaching slightly beyond apex of head.
Anterior margin of pronotum nearly straight to slightly concave behind head, obtusely carinate, forming distinct collar which is separated from disk of pronotum by row of deep punctures. Anterolateral pronotal margins nearly straight, distinctly reflexed, anteriorly curving mesad and over lateral margins of pronotal collar. Anterior disk of pronotum slightly tumescent medially, remainder uneven, nearly impunctate; posterior disk provided with a few shallow, coarse punctures. Dorsal surface black with all margins narrowly margined in yellow; sometimes with transverse, thin, yellow band behind pronotal calli (three of four specimens lack this transverse band), thus forming either two or four large black spots (Fig. 3).
Scutellum triangular, apex narrowly rounded, base tumescent, surface black with lateral margins pale yellow (except extreme edges black near base), apex pale yellow, and a narrow, longitudinal line pale yellow (Fig. 3). Basal area of scutellum impunctate, disc and apex shallowly and coarsely punctate. Hemelytra more densely and uniformly punctate, except small area near apex of r + m vein impunctate. Coria black with basal embolar area yellow, this band leaving margin and continuing along lateral margin of r + m vein, then curving mesad and continuing to inner basal angle; apical margin also pale yellow; sometimes obscure pale yellow streak vaguely apparent along claval suture, and another more laterad paralleling claval suture (Fig. 3). Hemelytral membrane fumose, distally hyaline. Connexiva entirely yellow, completely covered by hemelytra when at rest.
Ventral surface of head yellow, except areas around compound eyes and antennifers black, and a black band running along base of bucculae, often with a thin black streak near apex of juga. Rostrum reaching between, and nearly to posterior margins of hind coxae; segment I reaching distinctly beyond apices of bucculae; all segments fuscous to black, except segment I more pale, or mottled with yellow. Prosternum slightly depressed medially with low, vague, obtuse carina on each side, forming v-shape, anterior ends continuing as distinct, obtuse carina along each anterior propleural margin; mesosternum medially carinate, somewhat tumescent on each side of carina, glabrous; metasternum flat medially, medial areas of prosternum, mesosternum, and metasternum covered with short appressed hairs. Thoracic pleura yellow with small black spots near bases of coxae, and larger black spots more laterad. Ostiolar opening small, unattended by ostiolar ruga; evaporative area reduced to small finger-like area caudad of opening, and a thin band along anterior margin of each metapleuron. Coxae brownish, rest of leg segments fuscous to black, perhaps a short obscure yellow streak visible on each femur. Abdominal venter unarmed, impunctate, with two longitudinal rows of large black spots on each side, medial pair on segment VII have fused to form single medial spot. Posterolateral abdominal angles rounded, not spinose.
Posteroventral surface of pygophore divided into dorsal and basal halves by transverse, shallow depression, this depression is interrupted laterally on each side by thin, transverse, ridge running from dorsomesad to lateroventrad (Fig. 10, tr); dorsal half yellow, ventral half may be black, but often mostly yellow with some black lateral markings. Posterior margin of pygophore obtusely rounded laterally, with medial U-shaped depressed area, this depressed area with U-shaped medial emargination (Fig. 10). Posterior lobe of each paramere greatly elongate, nearly parallel-sided, slightly sinuous, with three longitudinal ridges, apex narrowly rounded to spinose (Figs 11-13); anterior parameral arm also somewhat elongate, rim with many small black teeth, apical tooth slightly larger. Proctigeral surface distinctly concave, anterior margin not or only feebly emarginate medially; anterior and posterior halves of proctiger nearly subequal in width. Superior ridge narrowly but distinctly produced over proctiger. Female unknown.
Measurements (mm). Total length 11.90-13.53 (12.64); width across humeri 6.90-7.65 (7.46); width across abdomen 7.42-8.07 (7.52); medial length of pronotum 2.68-2.79 (2.77). Medial length of scutellum 5.25-5.46 (5.25); basal width 4.04-4.27 (4.22); width at distal end of frena 1.23-1.36 (1.36). Length of head 2.25-2.38 (2.31); width 3.12-3.28 (3.28); interocular distance 2.15-2.26 (2.26); interocellar distance 0.94- 1.00 (0.96); ocellar diameter 0.10-0.11 (0.10); distance from ocellus to adjacent eye 0.59-0.64 (0.64). Length of segments I-V of antennae 1.05-1.14 (1.14), 1.59-1.78 (1.59), 2.19-2.48 (2.33), 3.09-3.16 (na), and 2.95 (na) respectively. Length of segments I-IV of rostrum 1.24-1.38 (1.26), 1.83-2.01 (1.98), 1.03-1.11 (1.03), and 1.32-1.33 (1.33) respectively.
Distribution: China (Guangdong, Yunnan), Vietnam.
Holotype: ♂, labeled a) “ Tonkin. Hagiang. May 1914 R. V. de Salvaza.“ (BMNH). Paratypes (3♂♂): China, Guangdong Prov., Lian-xian Co., 24.7 N, 112.3 E (1♂, NKUM); China, Yunnan Prov., Long-hu (1♂, NKUM). Vietnam, 40 km N. Tai-Nguyen (1♂, ZMAS).
Etymology. This species is named in honor of Ernst Heiss, in recognition of the contributions he has made to the systematics of the Aradidae, and also on the Heteroptera fauna of Europe.
Comments. This species is most closely related to M. limbata as they both have rather elongate posterior lobes of the parameres which are either spinose or obtusely spinose apically. The posterior parameral lobe is more elongate in this species, with the lateral margins nearly parallel. The transverse depressed area on the posteroventral surface of the pygophore is interrupted laterally on each side by a narrow, oblique ridge.
description: Published as part of RIDER, D. A., 2006, Review of the genus Madates STRAND with the description of three new species (Heteroptera, Pentatomidae), pp. 599-610 in Denisia 19 on pages 605-606