Sogalabana Stroiński & Świerczewski 2014, gen. nov.
Creators: name: Stroiński, Adam
name: Świerczewski, Dariusz
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Flatidae
Sogalabana

description: Genus Sogalabana gen. nov.   
(Figures 1–7)   Type species   
 Sogalabana ochracea sp. nov., here designated.    Etymology   
 The generic name is a combination of the names Soga (after Pierre Soga, collector of the specimens representing a new genus) and Analabana (locus typicus of newly described species). Gender: feminine (Genetivus singularis: Sogalabanae)    Diagnosis   
 The genus Sogalabana is related to Phaedolus Karsch, 1890 (type species Phaedolus quadripunctatus Karsch, 1890) but differs by the following characters: frons with distinct protrusion in upper part (Phaedolus – frons convex); disc of frons without carinae (Phaedolus – disc of frons with three carinae); veins ScR + M arise as short common stem from basal cell (Phaedolus – veins ScR + M arise at the same point from basal cell).    Description   
 Head. Head with compound eyes in dorsal view narrower than thorax, strongly prolonged (Figure 1B,D).   
Vertex much wider than long at midline, medially partly covered by pronotum (Figures 1D; 2A,B). Anterior margin deeply concave, in form of elevated suture, laterally obsolete; posterior and lateral margins carinate. Disc of vertex without carinae, sensory structures and secretory pores.  
Frons (Figures 1D; 2A,B) in upper part with distinct protrusion, in dorsal and lateral view triangular shape with bluntly rounded apex; upper surface flattened, placed at about same level as thorax. Lateral margins of frons carinate, in frontal view widest about level of antennae. Disc of frons (upper and frontal surface) without carinae, but with small sensory pits.  
Compound eyes rounded, with small callus placed at lower-posterior margin. Ocelli present (Figure 2E). Antennal pedicel elongate, widest medially, with setae and plate organs placed apically and on upper surface (Figure 2F). Clypeus narrower than frons, without carinae (Figure 2C,D). Rostrum with apical segment distinctly shorter than subapical segment, apex reaching hind coxae.  
 Thorax. Pronotum distinctly longer than vertex at midline (Figures 1D; 2A,B); anterior and posterior margins arcuate. Pronotum posteriorly with extremely short median, incomplete carina (clearly visible in female) and median groove (clearly visible in male); disc of pronotum with lateral impressions and triangular postocular eminences.   
Mesonotum triangular (Figures 1D; 2A,B); disc posteriorly with short median, groove; lateral carinae connected basally, running parallel reaching posterior margin; surface between lateral carinae almost flat, with two sensory pits; lateral parts of mesonotum with gibbosities.  
 Tegmen (Figures 1A; 3B–F) coriaceous and weakly convex, with clearly visible venation and bulla, without apical, subapical and nodal lines, with net of irregular transverse veinlets. Costal margin – basal half arcuate, apical part straight, costal angle widely rounded, posterior margin straight, sutural angle acute with rounded apex, postclaval sutural margin straight. Costal area narrower than costal cell, with single, transverse veinlets, terminating a bit before end of clavus. Costal cell wider than costal area with net of transverse veinlets. Basal cell very long and narrow. Longitudinal veins ScR + M arise as short common stem from basal cell; ScR forking just before bulla into ScRA and RP; ScRA strongly elevated, forking into Sc and RA near posterior margin (Figure 3E); M fork before half of tegmen; Cu bifurcated before end of clavus. Claval veins PCu and A 1 fused before end of clavus, vein A 1 elevated; transverse veinlets between claval suture and Pcu. Tubercles on whole tegmen with concentration on costal area, costal cell, between basal Sc + R and M veins and on clavus between PCu-A 1.   
Femora shorter than tibiae; hind tibia arcuate and partly flattened laterally with two lateral spines placed after midlength, apically with row of well-developed teeth (Figure 2G); basitarsomere as cumulative length of second and third tarsomeres with row of apical spines, second tarsomere with two lateral spines (Figures 2H; 3A).  
 Male genitalia. Anal tube (in lateral view, Figures 3G; 5A) massive, longer than wide; basal part distinctly wider than apical; anus placed about midlength. Anal tube (in dorsal view, Figures 3H; 5C) calyx-shaped, basal part narrower than apical part; anus placed about midlength.   
Pygofer (in lateral view; Figure 3G) higher than wide, almost rectangular, dorsoposterior angle bluntly rounded and “elevated”.  
Genital styles (in lateral view; Figures 3G; 5B) longer than wide and bearing distinct, triangular capitulum; dorsal margin without concavity near base of capitulum; ventroposterior angle weakly projected.  
Phallic complex. Periandrium – closed basally to about one-third length, remaining part open dorsally (Figure 5D,E). Dorsal part of periandrium apically with wellsclerotized, vertically oriented lobes and two large processes (dorsal and ventral) oriented basad; dorsal part, near base, with bulba with a few teeth (Figure 4A). Ventral part apically with three lobes; ventral surface with row of denticles. Aedeagus divided into dorsal and ventral parts (Figure 5F,G). Dorsal part with lateral lobes in median portion, apical part bilobate with median incision, subapically with vertical appendage. Ventral part apically widened with lateral lobes.  
 Female genitalia. Pregenital sternite massive, lateral lobes weakly separated (Figures 4E,F; 6A); anterior margin weakly concave, medially with sclerotized lobe (Figure 6A); posterior margin medially with two bluntly triangular lobes separated by shallow concavity.   
Anal tube (in lateral view; Figure 4B,D) flattened, elongate and narrow, reaching end of gonoplac; anus placed about midlength; ventral surface with long setae. Anal tube (in dorsal view; Figures 4C; 6B) pear-shaped; anus placed about midlength.  
Gonoplac unilobate, laterally flattened, elongate (Figures 4B,D; 6C); posterior margin rounded with single row of well-developed teeth (Figure 6D); narrow, membranous part placed alongside ventral margin, extending half of gonoplac; dorsal and ventral part externally with long setae, median portion of internal surface with short and thick setae.  
Gonapophysis VIII sabre-shaped and laterally flattened, tapering apicad (Figures 4F; 6E); apical part of ventral margin folded externally, apical part of dorsal margin with two teeth. Endogonocoxal process a bit shorter than gonapophysis VIII, sabreshaped with spiniferous microsculpture.  
Gonapophyses IX and gonospiculum bridge as in Figure 6F,G.  
 Bursa copulatrix of single, elongately oval, huge pouch; cells clearly visible, without ornamentation (Figure 7A). Spermatheca well developed; ductus receptaculi longer than diverticulum ductus (Figure 7B).    Distribution   
Madagascar: Mahajanga Province.
description: Published as part of Stroiński, Adam & Świerczewski, Dariusz, 2014, Sogalabana ochracea gen. et sp. nov. from Tsaratanana massif in northern Madagascar (Hemiptera: Fulgoromorpha: Flatidae), pp. 1853-1865 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 48 (29 - 30) on pages 1854-1863, DOI: 10.1080/00222933.2014.902141, http://zenodo.org/record/4607600