Chaerilus terueli Kovarik 2012, sp. n.
Creators: name: Kovařík, František
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Scorpiones
Chaerilidae
Chaerilus
Chaerilus terueli

description: Chaerilus terueli Kovařík, sp. n.   
(Figs. 48–61)  
TYPE LOCALITY AND TYPE REPOSITORY. Vietnam, Côn Son Island (Poulo Condore); FKCP.  
TYPE MATERIAL. Vietnam, Côn Son Island (Poulo Condore), III.2012, 1♂ (holotype) 2♂ (paratypes), leg. V. Fura, FKCP.  
ETYMOLOGY. Named after my colleague and friend Rolando Teruel Ochoa, who has contributed to our knowledge of scorpions in many ways.  
DIAGNOSIS. Total length 31–37 mm. Two pairs of lateral eyes and one pair of median eyes. Movable finger of pedipalp with 7–8 cutting edges composed of granules. Chela of pedipalp wide and ampullar, adult male has fingers strongly flexed. Manus of pedipalp in male robust. Chela of pedipalp with 4 complete carinae, patella with 4 or 5 carinae, femur with 4 carinae. Pectinal teeth 5–6 in males. Entire carapace granulated. Larger granules form two symmetrical, longitudinal carinae. Mesosomal tergites granulated. Several large, symmetrical granules on posterior margins of tergites II– VI may be interpreted as incomplete carinae. Sternites smooth, without carinae. First metasomal segments lack ventral carinae; ventral side of first metasomal segment smooth, but may bear several solitary granules. First through third metasomal segments with 6–8 carinae; dorsally with a pair of rows of granules, which may form incomplete carinae. Fourth metasomal segment with 8 carinae and irregular granules on lateral surface.  
 DESCRIPTION. Total males length 31–37 mm. Two pairs of lateral eyes and one pair of median eyes. Pectinal teeth number 5–6 (3x5, 3x6) in males. For the position and distribution of trichobothria see Fig. 50–56. Trichobothrium d2 of the pedipalp patella is located on the edge between dorsal and internal surfaces. Trichobothrium d3 is located on the internal surface. (Fig. 53). The chelicerae are granulated.   
COLORATION. The color is reddish to black, spotted. Younger specimens are lighter-colored. Legs, pedipalps (mainly chela) and telson are red to yellow, lighter than the mesosoma.  
MESOSOMA AND CARAPACE. The entire carapace is covered by large granules, the larger of which form two longitudinal, symmetrical carinae. The anterior margin of the carapace is straight to weakly concave. The mesosomal tergites are granulated. Several large, symmetrical granules on the posterior margins of tergites II–VI may be interpreted as incomplete carinae. The sternites are smooth, without carinae and without glossy zones in the middle of the posterior margin.  
METASOMA AND TELSON. The first through third metasomal segments bear six to eight carinae (two ventral carinae and main lateral carinae may be poorly developed or absent); the fourth metasomal segment bears eight carinae and irregular granules on lateral surface. the fifth segment bears five to seven carinae of which one ventral carina posteriorly branches to form the letter “Y“. All carinae are composed of posteriorly inclined, denticulate granules. The spaces between carinae are irregularly granulated on lateral, ventral and dorsal surfaces; the dorsal surface bears a double row of granules, which may form incomplete carinae. All segments are sparsely hirsute. The telson is elongate, finely granulate and sparsely hirsute.  
PEDIPALPS. The chela of pedipalp is wide and ampullar, the adult male has fingers strongly flexed (Fig. 51). The male manus of pedipalp is robust. The movable finger of pedipalp has seven to eight cutting edges and the fixed finger has six to seven cutting edges composed of granules. The chela of pedipalp has four complete carinae, the patella has four or five carinae and the femur has four carinae. All carinae consist of widely spaced, rounded granules. The spaces between carinae are covered by unevenly spaced small granules, which on the chela form a reticulate pattern.  
LEGS. The legs are hirsute, without bristlecombs and carinae but with macrosetae.  
 MEASUREMENTS IN MM. Total length of male holotype 33.7; carapace length 4.5, width 4.6; metasoma and telson length 17.7; first metasomal segment length 1.6, width 2.5; second metasomal segment length 2.0, width 2.3; third metasomal segment length 2.1, width 2.2; fourth metasomal segment length 2.4, width 1.9; fifth metasomal segment length 4.5, width 1.9; telson length 5.1; pedipalp femur length 3.6, width 1.4; pedipalp patella length 3.6, width 1.6; chela length 7.9; manus width 4.3; movable finger length 4.1.   
 AFFINITIES. The described features distinguish C. terueli sp. n. from all other species of the genus. They are recounted in the key. The male of C. terueli sp. n. has the chela robust (Fig. 50) like the male of C. solegladi sp. n., C. robinsoni and C. variegatus. Of these species only C. solegladi sp. n. has the movable finger with seven or eight cutting edges composed of granules (Fig. 36), like C. terueli sp. n.. C. robinsoni has the movable finger with 11 cutting edges and C. variegatus with 13– 15 cutting edges. However, C. solegladi sp. n. and C. terueli sp. n. cannot be confused, because C. solegladi sp. n. has the fingers straight in both sexes (Fig. 39), whereas the adult male of C. terueli sp. n. has the fingers strongly flexed (Fig. 51).
description: Published as part of Kovařík, František, 2012, Five new species of Chaerilus Simon, 1877 from China, Indonesia, Malaysia, Philippines, Thailand, and Vietnam (Scorpiones: Chaerilidae), pp. 1-14 in Euscorpius 149 on pages 9-11