Atopida proba Sharp 1878
Creators: name: Kiałka, Agata
name: Ruta, Rafał
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scirtidae
Atopida
Atopida proba

description: Atopida proba Sharp, 1878
(Figs 2C–D, 5O–P, 8H–I, 9A, 10G, 15A, 17C)
Atopida proba Sharp, 1878: 47.
Atopida proba: Broun 1880: 313 (reprinted excerpt of Sharp 1878); Pic 1914: 16 (catalogue); Macfarlane et al. 2011: 420 (catalogue); Kiałka & Ruta 2017: 26 (catalogue).
Atopida basalis Broun, 1912: 431 syn. nov.
Atopida basalis: Pic 1914: 16 (catalogue); Macfarlane et al. 2011: 420 (catalogue); Kiałka & Ruta 2017: 21 (catalogue).
Atopida sinuata Broun, 1893: 1455 syn. nov.
Atopida sinuata: Pic 1914: 16 (catalogue); Macfarlane et al. 2011: 420 (catalogue); Kiałka & Ruta 2017: 26 (catalogue).
Type material. Syntype (NHM): card-mounted, “ Atopida proba \ Type D.S. \ New Zealand Broun” [written at base of card in Sharp’s hand], “Type \ H. T.” [round label with red border], “ New Zealand. [red underline]”, “Sharp Coll. \ 1905-313.” Syntype (NHM): card-mounted, “ Atopida proba \ Ind. typ. D.S. \ New Zealand Broun” [written at base of card in Sharp’s hand], “ New Zealand. [red underline]”, “Sharp Coll. \ 1905-313.” Syntype (NHM): cardmounted, “ Atopida proba \ Ind. Type D.S. \ New Zealand Broun” [written at base of card in Sharp’s hand], “ New Zealand. [red underline]”, “Sharp Coll. \ 1905-313.” Holotype of Atopida sinuata (NHM): card-mounted, “2524.” [in Broun’s hand], “Howick”, “ New Zealand. [red underline] \ Broun Coll. \ Brit. Mus. \ 1922-482.”, “ Atopida \ sinuata” [in Broun’s hand]. Syntypes of Atopida basalis (NHM): 2 exx., card-mounted, “3244” [in Broun’s hand], “ New Zealand. [red underline] \ Broun Coll. \ Brit. Mus. \ 1922-482.”, “Kaitoke. \ Wellington ” [in Broun’s hand], “ Atopida \ basalis” [in Broun’s hand].
Other material examined. 1 ex. (NZAC): “ NEW ZEALAND CL \ Thames, Kirikiri \ Saddle, 400m \ 12 Feb 1979 \ J.S. Dugdale ”, “to light”; 2 exx. (NZAC): “ NEW ZEALAND CL \ Thames, 400m \ Kirikiri Sdle \ 12 Feb 1979 \ J.S. Dugdale ”, “ex sprayed logs \ with polypores”; 3 males (NZAC): “ NEW ZEALAND CL \ Thames, Kirikiri \ Saddle, 450m \ 12 Feb 1979 \ J.S. Dugdale”, “to light”; 1 ex. (NZAC): “ Mt. Hobson \ 304 m. \ Gt. Barrier \ I. 27 Nov 57.”, “J.C. Watt”, “Lepto- \ spermum \ scoparium \ ericoides”; 1 ex. (NZAC): “Ohakune Harris”; 1 ex. (NZAC): “ NEW ZEALAND WO \ Mt Pirongia 500m \ 16 Jan 1977 \ A.K. Walker and \ W.H. Benn”, “Litter \ 77/3”; 1 ex. (NMPC): “ NEW ZEALAND: Waikato (WO) \ Pirongia Forest Park \ Mangakara Nature Walk; 177 m \ 37.9667° S, 175/ 149°E \ 18.xi.-21.xi.2019 \ M. Fikáček & M. Seidel lgt. MMO3”, “Lowland broadleaf forest with \ podocarps, nikau palms and \ Ripogonum lianas; sparse \ understory: baited pitfall traps \ (rotten squid)”; 1 ex. (NZAC): “coll. \ A.C. O’Connor \ Feb. 1919 ”, “Karori \ Wellington ”, “3244”, “ Atopida \ basalis \ Broun”, “ A.E. Brookes \ Collection”; 1 ex. (LUNZ): “ NEW ZEALAND, BP \ Te Koau 243 m \ NZMS 260 Z14 778862 \ 24.x.1992 - 31.i.1993 \ R.M. Emberson ”, “ pitfall traps, mixed \ broadleaf/podocarp forest”; 4 exx. (NZAC): “ NEW ZEALAND, BP \ Te Koau \ 1 Dec 1992 –\ 31 Jan 1993 \ J.S. Dugdale ”, “ Malaise trap ”; 4 males (NZAC): “ NEW ZEALAND, BP \ Orete Forest \ Te Puia Hut \ 19 Oct –\ 23 Nov 1992 ”, “ G. Hall \ Malaise trap ”; 1 ex. (LUNZ): “ NEW ZEALAND, TK \ Mount Egmont NP \ Stoney River \ 350m, 25.xii.1985 \ under stones”, “ R.M. Emberson \ P. Syrett ”; 6 exx. (CMNC): “ N. Zealand, TK \ 21 km N. Ohura \ Dec. 14, 1983 \ L. Masner ”, “Undergrowth in \ Nothofagus forest”; 2 exx. (NZAC): “New Zealand WN \ Otaki Forks \ 30-Dec-80”, “Ex John Nunn \ Collection \ Donated 2015”; 2 exx. (NZAC): “ NEW ZEALAND, WN \ Orongorongo V, W. N. \ 15 Dec 1994 ”, “Hard beech \ forest litter \ Tullgren #57”; 2 exx. (NZAC): “New Zealand WN \ Pakuratahi Forks \ Kaitoke \ 06-Jan-93”, “Ex John Nunn \ Collection \ Donated 2015”; 1 ex. (NHM): “N. Auck. \ 1 27”, “ NEW ZEALAND \ C.E. Clarke \ Collection \ B. M. 1957-24.”; 1 male (AMNZ): “Whangarei \ Falls \ 5.I.24 [on the underside of the cardboard with the beetle]”, “ C.E. Clarke \ Collection”; 1 ex. (NHM): “ Whangarei \ 6 1 1927 \ C.E. Clarke”; 1 ex. (NZAC): “Kaimai \ 1500’ \ 3-I-48”, “ A. E. Brookes \ Collection”; 1 ex. (NZAC): “ Parna Bay \ Whangarei \ Fairhum. Dec. 27”, “ A.E. Brookes \ Collection”; 1 ex. (NZAC): “ Balance Reserve \ Manawatu Gorge \ 11/1/67”, “ Atopida \ suffusa”, “R.M. Bull \ Collection”; 1 female (NZAC): “745”, “ NEW ZEALAND. \ Wainuiomata \ 745. \ G.V. Hudson”, “Pres. by \ Comm Inst Ent \ B.M. 1981-315”; 3 males (AMNZ): “ Silverstream \ 22.XII.1923 ”, “ Hutt Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM”, “C.E. Clarke \ Collection”; 1 male (AMNZ): “Silverstream \ Wellington \ 15.12.23 \ Coll. T. [illegible] [on the underside of the cardboard with the beetle]”, “ C.E. Clarke \ Collection”, “ Atopida \ proba \ Sharp \ Id. by J. Armstrong ”; 2 males (NZAC): “Okiwi \ Great Barrier \ Island”, “24 Nov 57 \ J.C. Watt ”, “beating \ flowering \ Leptospermum ”; 1 male (NZAC): “Unuwhao 800’ \ Spirits Bay ”, “Forest \ Litter”; 1 male (NZAC): “ NEW ZEALAND RI \ Ruehine nr \ Rangiwahia \ 4 Jan 1975 \ J.C. Watt ”; 1 male (NZAC): “ NEW ZEALAND BP \ Kaimai Ra \ Lindeman lookout \ and of Lindeman Rd \ 4 Nov 2005 ”, “R Leschen S Nomura \ beating RL992 \ 37°54’ S \ 175°87’ E”; 1 male (NZAC): “Wharite \ 2-2-58 \ J.I. Townsend ”; 1 male (AMNZ): “ NEW ZEALAND AK \ Mataitai Forest S.A. \ pitfall B 1 \ Jan 2002 \ A. Warren”; 2 males (NZAC): “ NEW ZEALAND CL \ Little Barrier I \ Awaroa Stm \ 15 Feb 1977 \ J.A. Macdonald ”, “ MV light trap \ at base of \ waterfall”; 3 males (AMNZ): “ NEW ZEALAND, CL \ Great Barrier I, Little \ Windy Hill, 220m, \ 11.xii.2001 - 18.i.2002, \ P. Sutton.”, “Forest edge, Malaise \ trap. L21044”; 6 males (AMNZ): “ NEW ZEALAND, CL \ Great Barrier I, Little \ Windy Hill, 220m, \ 7.xi.-11.xii.2001, \ P. Sutton & J. Gilbert.”, “Forest edge, Malaise \ trap. L11043.”; 3 males (AMNZ): “ NEW ZEALAND, CL \ Great Barrier I \ Mt Hobson, 500 m \ 17 Dec 2003 \ J.W. Early”, “Agathis / podocarp \ forest \ Swept. L12446”; 1 male (AMNZ): “Waikaraka \ Valley \ Parva Bay \ 14.i.24 [on the underside of the cardboard with the beetle]”, “ C. E. Clarke \ Collection”, “ Atopida \ proba \ Sharp \ Id. by J. Armstrong ”; 1 male (AMNZ): “swept understorey \ Waikaraka Strm.Vy. \ S. of Puketi Forest \ 14.XI.1985 \ R.F. Gilbert”, “ Bay of Island Co \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM”; 1 male (AMNZ): “Wayby Gorge \ N. Auck. \ 28.12.26 [on the underside of the cardboard with the beetle]”, “ Rodney Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM”, “ C.E. Clarke \ Collection”; 2 males (AMNZ): “Clevedon \ 27.2.34 [on the underside of the cardboard with the beetle]”, “ C.E. Clarke \ Collection”.
Diagnosis. Body of moderate size to large (TL 3.9–5.0 mm), uniformly brown. Setation of head and pronotum adherent, covered with somewhat flattened setae, with no suberect setae, what makes this species easy to distinguish from other Atopida.
Redescription. Male. Body elongated, moderate size to large (TL 3.9–5.0 mm), moderately convex. Body uniformly brown, antennae tone lighter at base. Punctation on head and pronotum granulate, strong and dense, punctures touching each other, punctation on elytra simple, punctures separated by 0.5–1.0× diameter of a puncture, interspaces microreticulate, dull. Body covered with moderately long, adherent, yellowish setae, arranged into kind of pattern along the middle of pronotum and sometimes head. Supraantennal ridges well-marked, elevated. Pronotum widest in anterior 1/3, sides curved, anterolateral angles not or only subtly produced, posterolateral angles obtuse. Elytra without carinae, humeri well-marked. Ventrite 5 rounded. Penis long (L 0.63 mm, W 0.24 mm), trigonium narrow, as long as or slightly shorter than parameroids, slightly bent upwards, apex truncate; parameroids elongated, apices broadening and obliquely truncate, slightly serrated, apical part covered with few punctures; pala triangular, much longer than parameroids. Tegmen (L 0.52 mm, W 0.34 mm) with moderately wide subrectangular parameres, apices triangularly narrowing, with scarce punctures and delicate setae in apical part.
Female. Body moderately big to large (TL 4.3–5.0 mm), macropterous. Supraantennal ridges well-marked, elevated. Pronotum strongly transverse, much stronger than in males, punctures along basal margin larger and sparser than on remaining portions of elytra. Setation on elytra very short, setae nor reaching the next puncture, elytral interspaces shining, punctures separated by ca. 0.5× diameter of a puncture. Ventrite 5 rounded, apex rounded, tergite VII with rectangular sclerotization in apical portion. Lateral portions of ventrites 3 and 4 with small, oval setal patches. Bursal sclerite consisting of pair of small (L 0.15 mm) rod-like sclerites hooked in posterior portion.
Measurements and ratios. Males (n = 10) TL 3.90–4.95 mm (4.50 mm), PL 0.70–0.85 mm (0.77 mm), PW 1.00– 1.40 mm (1.22 mm), EL 3.15–4.10 mm (3.73 mm), EW 1.70–2.20 mm (2.04 mm), TL/EW 2.08–2.35 (2.21), PW/PL 1.33–2.00 (1.58), EL/EW 1.76–1.95 (1.83), EL/PL 4.50–5.79 (4.84), EW/PW 1.48–2.15 (1.67). Females (n = 10) TL 4.25–5.00 mm (4.63 mm), PL 0.75–0.90 mm (0.80 mm), PW 1.30–1.55 mm (1.45 mm), EL 3.45–4.15 mm (3.79 mm), EW 1.85–2.15 mm (2.01 mm), TL/EW 2.21–2.38 (2.30), PW/PL 1.67–2.07 (1.82), EL/EW 1.76–1.97 (1.89), EL/PL 4.11–5.47 (4.77), EW/PW 1.34–1.46 (1.39).
Distribution. Common in North Island, known from scattered localities in Northland, Auckland, Coromandel, Waikato, Bay of Plenty, Gisborne, Taranaki, Rangitikei and Wellington areas (Fig. 18H).
Remarks. Broun (1912) correctly noted the similarities of A. sinuata and A. basalis, although he did not notice that these are males and females of the same species.
description: Published as part of Kiałka, Agata & Ruta, Rafał, 2022, Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae), pp. 401-443 in Zootaxa 5174 (4) on pages 426-428, DOI: 10.11646/zootaxa.5174.4.6, http://zenodo.org/record/6986469