Toechorychus heredius Tedesco & Aguiar 2013, sp. nov.
Creators: name: Tedesco, Anazélia M.
name: Aguiar, Alexandre P.
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Toechorychus
Toechorychus heredius

description: Toechorychus heredius Tedesco, sp. nov.   
(Figs 32, 75, 122, 160)  
Description. Holotype MALE. Fore wing length 5.56 mm.  
 Head. Mandible 1.56 as long as basal width, moderately pilose; ventral margin projected as crest; dorsal tooth distinctly longer than ventral tooth. Clypeus 1.62 as wide as high, subquadrate, entirely finely strigulate; apex 1.62 as long as base, truncate; apical margin sharp, medially straight. Supraclypeal area medially strigate, laterally finely punctate, moderately pilose, medially slightly prominent; between antennal foramens with V-shaped carina, medially interrupted; radicle foveolate. Flagellum slender. Supra-antennal area with very faint but distinct median longitudinal carina, medially with slightly longitudinal elevation, near antennal sockets striate; dorsal half medially rugulose. Paraocular area smooth. Vertex rugulose around ocelli, without sulcus; gena and vertex behind ocelli smooth, sparsely pilose; gena in lateral view at level of dorsal portion of occipital carina narrow, ventrally wide; occipital carina stout, dorsally absent, ventrally markedly projected as crest, reaching hypostomal carina far from mandible base; hypostomal carina projected as crest; malar space 1.06 as long as basal width of mandible.   
 Mesosoma. Mesosoma in dorsal view 2.04 as long as wide; in lateral view, mesosoma middle width 0.19 mm. Pronotum centrally smooth, latero-ventrally smooth, margin near mesopleuron entirely corrugated; pronotal swelling dorsally smooth, ventrally strigulate; collar dorso-laterally rounded, distinctly swollen, anteriorly smooth; epomia absent. Mesoscutum sparsely pilose, central lobe dorsally punctate, impression of notaulus markedly corrugated, laterals lobes dorsally striate, central lobe without longitudinal sulcus or carina; notauli deeply impressed, posteriorly abruptly curved, U-shaped. Scutoscutellar groove smooth, polished; scutellum 0.92 as long as wide, smooth; scutellar carina incomplete, restricted approximately to anterior 0.6; postscutellum 0.29 as long as wide; hind margin of metanotum with two short lateral carinae extending towards transverse furrow at base of propodeum. Subalar ridge somewhat elongate; mesopleuron dorsally medially strigate, ventrally strigulate; mesopleural groove corrugated; epicnemial carina stout, dorsally straight; dorsal end of epicnemial carina reaching 0.2 of posterior margin of pronotum; sternaulus complete, reaching base of mid coxa, markedly sinuate, moderately impressed; faintly corrugated; scrobe very shallow, forming sulcus. Mesothoracic venter strigulate; median portion of posterior transverse carina of mesothoracic venter short and straight, regularly shaped, without projections. Transverse furrow at base of propodeum laterally 0.17 as long as anterior portion of propodeum deep and narrow, uniformly corrugated; propodeum 1.18 as long as wide medially; anterior margin of propodeum without teeth; anterior portion of propodeum finely strigulate or with sparse coarse punctures, medially with two parallel longitudinal carinae; lateral longitudinal carina of propodeum represented by short curved carina; posterior portion of propodeum entirely markedly strigate, medially longitudinally swollen; behind posterior transverse carina with two divergent longitudinal carinae; anterior transverse carina of propodeum medially interrupted, faint, straight; posterior transverse carina absent; propodeal spiracle 1.70 as long as wide; pleural carina of propodeum complete, moderately stout; lower division of metapleuron strigulate, sparsely pilose, juxtacoxal carina long and stout.   
 Wings. Fore wing: vein 2+3Rs almost straight; ramellus absent; vein 1M+Rs posteriorly faintly sinuous; vein 1cu-a posteriorly slightly curved, arising far from vein 1M+Rs base, angle with vein M+Cu distinctly obtuse; vein 2Cu 0.50 as long as vein 2cu-a; slightly sinuous, almost straight; cell 1+2Rs 1.07 as high as pterostigma, rectangular; vein 2M distinctly longer than vein 3M; vein 3M distinct, 0.64 as long as vein 2M. Hind wing: vein 1Cu 2.63 as long as vein cu-a; vein 2-1A reaching 0.90 of distance to posterior margin.   
 Legs. Tibia with very sparse short bristles; hind coxa elongate.   
 Metasoma. T1 0.48 as long as hind femur, with faint basolateral tooth; minutely coriarious; spiracle at anterior 0.54, prominent; postpetiole dorso-anteriorly convex; T2–8 minutely coriarious; almost glabrous; T2 1.30 as long as wide at apex; apex of T2 2.00 as wide as base.   
 Color. Head, mesosoma and metasoma black and yellow. Head: pale yellow (229,213,152); mandible base, mandible apex, clypeal apex centrally, supraclypeal area just around clypeus, supra-antennal area medially, interocellar area, vertex and gena, except complete large orbital band, and occiput, black; scape ventrally pale yellow; scape dorsally, pedicel and flagellum, brown. Mesosoma: propleuron pale yellow, dorsal half black; pronotum black, except lateral marks at collar and pronotal swelling, pale yellow; mesoscutum black, except longitudinal rectangular mark at posterior 0.7 of central lobe, centrally interrupted; mesopleuron pale yellow with four black marks: anteriorly around hypoepimeron, at mesopleural groove, rounded spot just behind dorsal end of epicnemial carina, and along posterior 0.6 of sternaulus; remainder of mesosoma pale yellow, except for the following black marks: both axillary trough of mesonotum and metanotum, carinal triangle, anterior portion of propodeum, and three longitudinal marks at propodeum: medially, and laterally along pleural carinae, all marks posteriorly narrow. Legs: mostly pale yellow; fore and mid coxae dorsally with small apical black spot; hind coxa with two basally not connected black marks: ventral mark reaching basal 0.6, dorsal mark reaching apex; all trochanters pale yellow, dorsally with brown basal spot; all femora and tibiae bright yellow; fore and mid t1–5 brownish. Metasoma: black and bright yellow (216,176,075); T1 black, lateral and posterior 0.2 pale yellow; T2–8 black with posterior 0.4 bright yellow; S1 pale yellow; S2–8 yellowish.   
 Comments. Dorsal color pattern of propodeum is unique and characteristic for the species (Fig. 160). The only known T. heredius sp. nov. specimen was identified by I. Gauld as T. abactus in 2001 (label data), but it differs from the original description as follows. Tubercles at posterior portion of propodeum absent (vs. with transverse tubercles in T. abactus); mesopleuron with rounded black spot just behind dorsal portion of epicnemial carina (Fig. 32) (vs. mesopleuron without black marks except around hypoepimeron); posterior half of sternaulus black (vs. sternaulus white); fore and mid tarsi dorsally brown (vs. legs entirely pale yellow, except for all t5 black); T1 dorsally anteriorly black (vs. anterior portion of T1 white).   
 Female. Unknown.   
 Distribution. Known only from Costa Rica (Fig. 229).   
 Biology. Parasitoid of Mischocyttarus collarellus Richards (Vespidae, Polistinae), new record, in La Selva Biological Station, Costa Rica. Collected and reared by E. F. Smith.   
 Material examined. Holotype ♂, COSTA RICA: Heredia, La Selva Biological Station, 11.VII.1999, E.F.Smith, Toechorychus abactus det. I.Gauld, 2001, ex. Mischocyttarus collarellus nest (BMNH). Antennae apical half and right hind leg beyond tibia, missing; mesoscutum somewhat damaged by pin; otherwise in good shape.
description: Published as part of Tedesco, Anazélia M. & Aguiar, Alexandre P., 2013,  Phylogeny and Revision of  Toechorychus  Townes (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of thirty-five new species , pp. 1-138 in Zootaxa 3633 (1) on pages 38-40, DOI: 10.11646/zootaxa.3633.1.1, http://zenodo.org/record/5261931