Cissidium Motschulsky 1855
Creators: name: Darby, Michael
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Ptiliidae
Cissidium

description: Key to species in species group 2
1. Species in which the mesocoxal borders are clearly separated ......................................................... 2
– Species with united mesocoxal borders (join sometimes obscured by longer keels) ........................ 8
2. Keel spatulate (Fig. 37C) .................................................................................... C. noumeae sp. nov.
– Keel narrower, without rounded sides ............................................................................................... 3
3. Keel tapering...................................................................................................................................... 4
– Keel parallel-sided either wholly or in part ....................................................................................... 7
4. Keel smooth, evenly tapering to rounded tip..................................................................................... 5
– Keel rugose either partly or wholly ................................................................................................... 6
5. Anterior angles of mid-keel rounded not reaching collar...................................... C. pinense sp. nov.
– Anterior angles of mid-keel broad reaching collar ........................................... C. amieuense sp. nov.
6. Keel setose throughout ....................................................................................... C. mussardi sp. nov.
– Keel setose and parallel-sided at base then smooth to tip ................................... C. riparium sp. nov.
7. Keel broad with rectangular tip .......................................................... C. subfoveolatum Darby, 2019
– Keel narrower with slightly rounded tip................................................. C. matthewsi Johnson, 2007
8. Lateral margins of mesoventrite serrate............................................................................................. 9
– Lateral margins of mesoventrite without serrations ........................................................................ 20
9. Elytra without foveolae........................................................................................... C. advena sp. nov.
– Elytra with foveolae......................................................................................................................... 10
10. Keel broadening and rounded posteriorly............................................................ C. logunovi sp. nov.
– Keel pointed or parallel-sided or tapering posteriorly......................................................................11
11. Pronotal lateral margins with sharp angles medially....................................................................... 12
– Pronotal lateral margins without sharp medial angles..................................................................... 13
12. One pair of pronotal foveae elongate............................................................... C. banari Darby, 2013
– All the pronotal foveae more or less round ............................................................ C. clareae sp. nov.
13. Pronotal sides distinctly concave before hind angles, sides more sharply rounded (Fig. 28C) .......... .................................................................................................................... C. inexspectatum sp. nov.
– Pronotal sides shallowly concave, almost straight, before hind angles........................................... 14
14. Micropterous species ................................................................................ C. crowsoni Johnson, 1982
– Fully winged or apterous species..................................................................................................... 15
15. Apterous species .............................................................................................................................. 16
– Fully winged species........................................................................................................................ 17
16. Median process of collar long, keel narrow and pointed................................. C. montanum sp. nov.
– Median process of collar very short, keel wider................................. C. okuense Grebennikov, 2009
17. Head with a distinct fovea between the eyes ......................................... C. foveolatum Johnson 1982
– Head without a fovea ....................................................................................................................... 18
18. Elytra appearing almost parallel-sided ................................................ C. ishigakiense Sawada, 2008
– Elytral sides more rounded .............................................................................................................. 19
19. Pronotal foveae small, rounded ....................................................................... C. impressum sp. nov.
– Pronotal foveae larger two medial ones elongate ...................................... C. aristophanousi sp. nov.
20. Species with only one or two pronotal foveae................................................... C. delicatum sp. nov.
– Species with at least four pronotal foveae ....................................................................................... 21
21. Species with a short border along part of the posterior edge of the pronotum................................ 22
– Species without a pronotal posterior border .................................................................................... 23
22. Pronotum with two large basal foveae.......................................................... C. bifoveolatum sp. nov.
– Pronotum with four smaller basal foveae ............................................................... C. yoruba sp. nov. 23. Species with straight/almost straight sided mid keel....................................................................... 24
– Species with a clearly concave sided mid-keel................................................................................ 29
24. Posterior angles of mid-keel with carinae to mesocoxal borders ..................... C ceylonicum sp. nov.
– Posterior angles of mid-keel without carinae .................................................................................. 25
25. Sides of pronotum straight, parallel-sided before hind angles ............................ C. houailou sp. nov.
– Sides of pronotum before hind angles concave ............................................................................... 26
26. Medial process of collar onto mid keel almost reaching to basal tip.................. C. misellum sp. nov.
– Medial process of collar onto mid-keel shorter ............................................................................... 27
27. Elytra distinctly foveolate.................................................................................. C. lawrencei sp. nov.
– Elytra shallowly foveolate .............................................................................................................. 28
28. Pronotum with eight foveae across the basal margin .............................................. C. rogeri sp. nov.
– Pronotum with four foveae across the basal margin................................................ C. deanei sp. nov.
29. Sides of pronotum sharply angled ................................................................................................... 30
– Sides of pronotum rounded or with faint angle ............................................................................... 31
30. Mid-keel broad, medial extension of collar reaching to raised posterior margin of mid-keel ............ ............................................................................................................................ C. robustum sp. nov.
– Medial extension of collar very short/effaced not reaching posterior margin of mid-keel ................. ......................................................................................................................... C. greensladei sp. nov.
31. Keel long, parallel-sided, without pubescence ....................................... C. scutellaris (Deane, 1931)
– Keel pubescent................................................................................................................................. 32
32. Front angles of mid-keel not reaching collar, keel sharply pointed.................. C. davaoense sp. nov.
– Keel narrow, parallel-sided, foveolate ............................................................... C. lamington sp. nov.
description: Published as part of Darby, Michael, 2020, A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species, pp. 1-188 in European Journal of Taxonomy 622 on pages 19-21, DOI: 10.5852/ejt.2020.622, http://zenodo.org/record/3777282