Calomera mamasa Cassola & Brzoska 2008
Creators: name: Moravec, Jiří
name: Dheurle, Charles
name: Schüle, Peter
name: Wiesner, Jürgen
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Calomera
Calomera mamasa

description: Calomera mamasa Cassola & Brzoska, 2008
(Figs 167–199)
Calomera mamasa Cassola & Brzoska, 2008: 33.
Type locality. Indonesia, Sulawesi: Barat, Toraja road 13 km northeast of Mamasa, 02°55.5´S, 119°27.1´E, altitude 1425 m.
Type material. Holotype (examined from photographs Figs 167–171), ♂ in MSNG, labelled: “ Indonesia, Sulawesi / 1425 m, S. Sulawesi – Toraja / Rd., 13 km NE– Mamasa / 02°55.5´S; 119°27.1´E / D. Brzoska 27-IV-2005 ” [printed] // “ Holotypus / Calomera / mamasa n. sp. / F. Cassola & D. Brzoska / dederunt, 2006” [red, printed] // “Dono Avv. / F. Cassola, / 9.II.2008 ” [printed] // “Museo Civico / di Genova” [printed]. Paratypes. Of the 41 paratypes listed by Cassola & Brzoska (2008), following ones were examined: 1 ♂ in CCJM: “ Indonesia, Sulawesi / 895 m, S. Sulawesi-Mamasa / Rd., 44.5km N-Polewali / 03°12.4´S / – 119°19.2´E / D. Brzoska 26-IV-2005 ” [printed]. 1 ♂, 1 ♀ in FCCR-MCZR, 1 ♂ in PSCH: “ Sulawesi Barat 44 km N / Polewali on Mamasa rd: / subvertical roadcut at 03°12.4S- / – 119°19.2E (895m) / 26.IV.05. F. Cassola leg,” [printed] // “ Calomera / cf. cicatroscelis / (Acciavatti i.l.) / det, F, Cassola, 2005” [printed] // Calomera / mamasa / n. sp. / Det. F. Cassola, 2005” [printed]. 1 ♂ in FCCR-MCZR: “ Sulawesi Barat / 11.1. km W Malabo on Mamasa – / Mamuju rd. (1100 m ca) / 0.2°59.8–119°15.2E / 18.III.2007 F. Cassola leg. [printed]. All paratypes labelled: / “ Paratypus / Calomera / mamasa n. sp. / F. Cassola & D. Brzoska / dederunt, 2006” [red, printed].
Other material examined. 1 ♂ in JWCM: “5.- 7.7.2001, S. Sulawesi Prov., 400 m, 20 km NE Sabbang, 2.28.56S 120.12.00E, Bolm lgt.”. 1 ♂ in PSCH: “ Indonesia, Sulawesi Barat / 4.IV.2012, 14km NE Mamasa / P. Schüle & N. Rusdi leg. / 02°55.5S 119°27.5E ”. 2 ♂♂ in CKGC: “ Palolo env. - Palu distr. / Tengah Province / C.- Sulawesi – Indonesia / local collector, 5/2017”. 1 ♂ in AOCW: “ Indonesia, Zentral- / Sulawesi – Tengah / Gunung Sampuraga / 16.-17.I.1995, 1540m / leg. J. Haft ”. 1 ♀ in CDCL: “ Sulawesi / 5/92 / Gueraud leg. / Col. C. Dheurle ”. 1 ♀ in CDCL: “ Sulawesi / Palolo Palu / VI.2012 / Col. Dheurle ”. 1 ♀ in NHMW: “Indonesia-C. Sulawesi / Sampuraga Mts. 1400m / 11-12.2.95 / leg. Tarasov & Sinyaev ” // “ Lophyridia (s. str.) / decemguttata / (Fabricius, 1801) / det. J. Probst ”. 1 ♀ in SDEI: “Magondow / N. Celebes ” [mislabelled?] // “ Staudinger ” // Calomera cf. / mamasa / Cassola & Brzoska, 2008 / det JiřÍ Moravec 2024”. 1 ♂ in NMPC: “ Indonesia Central Sulawesi / Bonebone 110m / 25- 26.V.1996 ” // “ex coll. VÍt Kabourek / National Museum / Prague, Czech Republic ” 1 ♀ in NMPC: “ Indonesia C. Sulawesi, W / Poso, 11-16.iv.1999, 1–400 / m, 5-10km SW Tambarana / Bečvář & Zábranský leg.” // “ex col. Petr Votruba / National Museum / Prague, Czech Republic ” // “ Calomera / mamasa / Cassola & Brzoska, 2008 / det. Votruba 2010” // “Locality label / evidently confused / with labels of other species / stated by JiřÍ Moravec 2024”.
Differential diagnosis. Calomera mamasa belongs to the largest species of the complex. Body 13.5–15.5 (HT 14.0) mm long, 4.95–5.45 (HT 5.05) mm wide. It can be easily distinguished from other species of the species-complex by its remarkable dorsal coloration: head and pronotum deep coppery with reddish reflections, notably prevailingly reddish-cupreous on the pronotum, while elytra almost velvety black, usually with faintly diffusing greenish lustre on anteapical-lateral area and indistinct cupreous lustre on basal area. Setosity: genae glabrous (Fig. 168), proepisterna with setae at their ventral margin, sometimes up to proepisternal half (as in HT, Fig. 168); rather copious or very sparse setae along dorsolateral pronotal margins (Figs 182–186). Elytral maculation yellowish or pale orange-yellow, conspicuously wide, laterally adjacent to outer elytral margin (Figs 167, 172, 179–180, 187–191). Labrum (Figs 174–178) primarily 6–setose as in other species, yet examined specimens possess also 7–8 labral setae (in male holotype only five setae developed). Aedeagus similar to other species in its lateral view (Figs 170, 192–193, 195–196, 198), lacking lateral indentations in its dorsal (or ventral) view (Figs 171, 194, 197, 199).
For the description see Cassola & Brzoska (2008).
Distribution. Calomera mamasa is evidently endemic to the main Sulawesi territory. Most specimens come from the area of type locality along the Toraja road near the village of Mamasa in the Polewali Mamasa Regency, province of West Sulawesi (Sulawesi Barat). It spreads inwards the province of Central Sulawesi (Sulawesi Tengah), known there from Palolo Palu, up to Gunung Sampuraga, a mountainous area situated 180 km northeast from the type locality and in a different valley. This species inhabits altitudes at 770–1540 m, thus considerably higher than those known for C. paradecemguttata sp. nov. which also occurs in Sulawesi. As mentioned by Cassola & Brzoska (2008), it inhabits open and sunny, vertical to subvertical clay roadcuts or loess quarries and has not been collected on riversides.
The female (SDEI) from Northern Sulawesi, labelled “Magondow”, was obviously mislabelled; it is 15.1 mm long, and also its other characters correspond to those in C. mamasa. Also the above-listed female specimen from NMPC was evidently mislabelled. Furthermost, the altitude 110 m on the label of the above-cited male (NMPC) does not agree with the occurrence of C. mamasa; the label was evidently confused with labels of C. paradecemguttata sp. nov. collected at the same area and date (see the same labels cited under the new species above).
description: Published as part of Moravec, Jiří, Dheurle, Charles, Schüle, Peter & Wiesner, Jürgen, 2025, Reassessment of the concept of Calomera decemguttata (Fabricius) with a description of Calomera paradecemguttata sp. nov. (Coleoptera: Cicindelidae), pp. 1-56 in Zootaxa 5570 (1) on pages 37-43, DOI: 10.11646/zootaxa.5570.1.1, http://zenodo.org/record/14734482