Andecladodes cosangensis Bocakova & Campello-Gonçalves & Silveira 2022, SP. NOV.
Creators: name: Bocakova, Milada
name: Campello-Gonçalves, Lucas
name: Silveira, Luiz Felipe Lima Da
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Lampyridae
Andecladodes
Andecladodes cosangensis

description: ANDECLADODES COSANGENSIS SP. NOV.
(FIGS 3F, 4F, 5F, 6B, 7C, 8E, F, 9C, 10E1, E2)
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Holotype: Male, ‘ ECUADOR, Napo Prov., Cosanga, 1900–2100 m, 0 o 36′S 77 o 53′W, Bolm lgt. 12–16 December 2010 (UPOL).
Diagnosis: Body black, only elytral apex brightly orange (Fig. 3F), elytra arcuately acuminate distally, phallus apically bulbous.
Description: Whole body black, only apex of elytra, and a broad medial stain in basal half of abdomen orange. Margin between black and orange elytral coloration arcuate. Head with small eyes, interocular distance 1.5× longer than eye diameter. Antennae short, reaching humeral area, antennomere I stout, with a short horn-like extension as long as half of antennomere width, antennomeres II and III short, length of antennomeres III–X gradually increasing. Branches of antennomeres III–X slender, lamella of antennomere III the longest, length of lamellae decreasing towards apex. Mandibles slightly falcate (Fig. 6B). Labrum connate to frontoclypeus, tongueshaped, forming small, translucent, median extension. Clypeolabral suture indistinct, absent. Maxillary palps slender, terminal palpomere elongate, with apex compressed. Labial palps three-segmented, minute, terminal palpomere diminishing apically, both maxillary and labial palps setose. Pronotum trapezoidal, anterior and lateral margins moderately arcuate, anterior corners widely obtuse (Fig. 4F). Discal area slightly elevated, almost glabrous, covered with fine primary pubescence, anterior and lateral edges of pronotum strongly punctured. Scutellum triangular with apex widely obtuse. Elytra about 2× longer than humeral width, widest in elytral quarter, moderately acuminate distally. Longitudinal costae largely obsolete, costa 1 present only in basal half, costa 2 and marginal costa fully developed, costae 3 and 4 unclear, sutural margins dehiscent in apical one-fifth. Sternum VIII with median point short, slender (Fig. 8E). Pygidium with lateral margins parallel-sided in distal half, median extension large, moderately longer than the lateral (Fig. 8F), syntergite asymmetrical, tergum IX provided with a single, unpaired, spine laterally, suture between tergum IX–X visible, apex of sternum IX moderately emarginated (Fig. 9C–D). Male genitalia with phallus long, broad in basal half, apex bulbous (Figs 10E 1–E 2). Body length: 12 mm, humeral width: 4.6 mm, width in elytral quarter: 5.2 mm. Female unknown.
Etymology: Named for Cosanga, the type locality.
Distribution: Ecuador.
Comments: Andecladodes cosangensis resembles Dodacles elegans by the distally acuminate elytra, but the structure of antennae (antennomere I shortly horned, length of antennomeres increasing distally and length of branches decreasing distally) and the shape of male genitalia demonstrated that A. cosangensis is closely related to the following new species with broadly widened, oval elytra. Therefore, the acuminate shape of elytra of A. cosangensis and Dodacles elegans is assumed to have evolved as a result of convergent evolution.
description: Published as part of Bocakova, Milada, Campello-Gonçalves, Lucas & Silveira, Luiz Felipe Lima Da, 2022, Phylogeny of the new subfamily Cladodinae: neotenic fireflies from the Neotropics (Coleoptera: Lampyridae), pp. 1181-1199 in Zoological Journal of the Linnean Society 195 on page 1196, DOI: 10.1093/zoolinnean/zlab091, http://zenodo.org/record/6993662