Pseudopoda mamillaris Zhang, Jager & Liu 2023
Creators: name: Zhang, Jianshuang
name: Pan, Tianqin
name: Zhang, He
name: Xing, Yuanqian
name: Yu, Hao
name: Zhong, Yang
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Sparassidae
Pseudopoda
Pseudopoda mamillaris

description: Pseudopoda mamillaris Zhang, Jäger & Liu, 2023
Figs 1, 8, 9, 10, 11, 28 B
Pseudopoda mamillaris Zhang, Jäger & Liu, in Zhang et al. 2023 a: 176, figs 159 A, C, 160 A, B (♀).
Holotype.
♀ (CBEE, LJ 202002615), China: • Yunnan Pro.: Honghe Hani and Yi Autonomous Prefecture, Luechun Co., Martyr Cemetery, 22.99 ° N, 102.45 ° E, c. 1940 m, by hand, 11 VII 2020, R. Zhong et al. leg. Examined.
Material examined.
• 2 ♂, 2 ♀ (YNZY 009, YNZY 010, YNZY 022, YNZY 023), same locality as holotype, by hand, 15 IV 2024, Y. Zhong & S. Yang leg.
Diagnosis.
Male of P. mamillaris is similar to that of P. platembola Jäger, 2001, in having a similar sickle-shaped embolus (E) and petal-shaped conductor (C). However, it can be distinguished from the latter by the following characteristics: (1) in ventral view, embolus (E) is apically rugged and as wide as its middle section (vs. apically not rugged, pointed) (cf. Figs 8 A, 10 B, and Jäger 2001: figs 35 b, g, i); (2) dorsal branch of RTA (dRTA) tip not curved (vs. slightly or distinctly curved) (cf. Figs 8 A, B, 9 B, 28 B, and Jäger 2001: figs 35 a – c, f – h); (3) RTA inserted closer to cymbium (Cy), both dRTA and vRTA separated from cymbium (Cy) by approximately half of the tibial (Ti) diameter (vs. inserted farther, both dRTA and vRTA separated from cymbium (Cy) by more than one tibial (Ti) diameter) (cf. Figs 8 A, B and Jäger 2001: figs 35 b, g). For the female diagnosis, see Zhang et al. (2023 a).
Description.
Male (YNZY 009). Total length 7.3. Carapace 3.6 long, 3.5 wide, anterior width 1.8. Opisthosoma 3.8 long, 2.6 wide. Eye sizes and interdistances: AME 0.19, ALE 0.30, PME 0.23, PLE 0.31, AME – AME 0.14, AME – ALE 0.06, PME – PME 0.23, PME – PLE 0.26, AME – PME 0.28, ALE – PLE 0.23, CH AME 0.32, CH ALE 0.27. Spination: palp: 131, 101, 2111; Fe: I – III 323, IV 321; Pa: I – III 101, IV 100; Ti: I – II 2226, III – IV 2126; Mt: I – II 2024, III – IV 3036. Measurements of palp and legs: palp 5.2 (1.6, 0.9, 1.0, 1.7), I 15.3 (4.4, 1.5, 3.8, 4.2, 1.4), II 15.9 (4.5, 1.9, 4.2, 3.8, 1.5), III 13.1 (3.8, 1.4, 3.4, 3.3, 1.2), IV 14.7 (4.0, 1.7, 4.0, 3.6, 1.4). Cheliceral furrow with ~ 26 denticles.
Colouration in ethanol (Fig. 10 D, E). DS yellowish brown, with numerous dark spots; lateral bands and margin slightly darker, clothed with thin hairs; median band bright yellowish-brown, without distinct pattern, not distinctly delimited to lateral bands; fovea and radial grooves distinctly marked. Cheliceral base yellowish white, with red fangs. Sternum uniformly yellowish. Endites and labium reddish orange, darker distally. Legs coloured as DS, with numerous brown spots, and covered by short spines. OS elongate-oval; dorsum with indistinct median band starting from behind pedicel, extending to 4 / 5 of abdomen length, almost reaching the indistinct transverse yellow line; median band anteriorly represented by a longitudinal brown line, with two light brown and reniform patches on each side, the latter half distinctly widened and not clearly delimited, with three pairs of circular spots located at lateral part; ventral OS basically yellowish, centrally marked with three pairs of purplish dots.
Palp (Figs 8, 9, 10 A – C, 28 B). Femur and patella unmodified. Tibia (Ti) moderately long, ca. 3 / 5 cymbium length, with retrolateral apophysis (RTA) arising mesially; RTA bifurcated, ɔ-shaped in retrolateral view, both ventral and dorsal branches distinctly protruding: dorsal branch (dRTA) finger-like, slightly curved and tapering, ca. 3 / 5 of tibia length, extending to cymbial base; ventral branch (vRTA) thumb-like, relatively short, ca. 1 / 2 length of dRTA, apex round. Cymbium (Cy) ca. 1.9 × longer than wide, cymbial bulge (CB) indistinct. Tegulum (T) oval, ca. 1.2 × longer than wide, posteriorly slightly bulged, slightly excavated on prolatero-apical side to accommodate embolus (E) and conductor (C); spermophor (Sp) sinuate, forming a loop along tegular margin. Embolus (E) wide and flattened, nearly Ƨ-shaped in ventral view, ca. 3 / 4 of tegulum length and 1 / 6–1 / 5 of tegulum width; the embolic base (EB) situated prolateral on the tegulum (ca. 9–10 o’clock on tegulum); the free part of the embolus (E) sickle-shaped; the embolic tip (ET) apically rugged, terminated at ca. 12 o’clock position. Conductor (C) membranous, ca. 1 / 3 of embolus length, thick, more or less petal-shaped, inserted apically (approximately 12 o’ lock relative to tegulum), covering embolic tip (ET) in prolateral and ventral views.
Female (YNZY 010). Total length 9.2. Carapace 3.8 long, 3.4 wide, anterior width 2.0. Opisthosoma 5.4 long, 4.0 wide. Eye sizes and interdistances: AME 0.20, ALE 0.31, PME 0.25, PLE 0.33, AME – AME 0.16, AME – ALE 0.06, PME – PME 0.21, PME – PLE 0.29, AME – PME 0.31, ALE – PLE 0.31, CH AME 0.32, CH ALE 0.23. Spination: palp: 131, 101, 2121, 1014; Fe: I – III 323, IV 321; Pa: I – IV 101; Ti: I – III 2026, IV 2126; Mt: I – II 2024, III 2026, IV 3036. Measurements of palp and legs: palp: 4.3 (1.3, 0.7, 0.9, 1.4), I 11.0 (3.4, 1.2, 2.6, 2.8, 1.0), II 12.1 (3.7, 1.6, 3.1, 2.6, 1.1), III 9.6 (2.9, 1.1, 2.4, 2.3, 0.9), IV 10.6 (3.1, 1.3, 2.7, 2.5, 1.0). Cheliceral furrow with ~ 26 denticles. Colouration in ethanol as in males, but body slightly darker (Fig. 11 D, E; see Zhang et al. (2023 a) for others described).
Epigyne (Fig. 11 A – C). Epigynal field ca. 1.27 × wider than long; anterior margin (aEF) distinctly delimited, trilobate, anterolaterally with two large v-shaped incisions that are well separated by at ca. 2 × widths; anterior bands (AB) indistinct, situated at the two incisions. Median field (MF) anterior margin invisible, large, ca. 1 / 2 epigyne length and as wide as epigyne. Lateral lobes (LL) nearly as wide as long; anterior margins (amLL) distinctly recurved; median margins (mmLL) entirely touching each other along the middle line; posterior margins (pmLL) distinctly procurved; posterior incision (PI) absent. Copulatory openings (CO) indistinct, located at basolateral borders of median field (MF). First windings (FW) hyaline, shaped like inverted triangles, starting from copulatory openings (CO), descending longitudinally, tapering posteriorly; two first windings (FW) separated by ca. one diameter. Spermatheca (S) nearly ‘ ∞ ’ - shaped, consisting of inner part and lateral part; inner part spherical, touching median margin of lateral lobe (mmLL), entirely covered by membranous sac (MS) in dorsal view; lateral part shaped like water droplet, anterolaterally with globular appendage (SA), entirely covered by first winding (FW) in dorsal view. Membranous sac (MS) nearly trapeziform, located at posterior portion of vulva; anterior margin almost straight, ca. 1 / 3 epigyne width, separated from epigastric fold by ca. 2 / 5 epigyne length; posterior margin also nearly straight, ca. 2 / 3 epigyne width, reaching posterior margins of lateral lobes (pmLL). Fertilization ducts (FD) acicular, membranous, ca. 1 / 2 length of first windings.
Distribution.
Known only from the type locality (Fig. 1).
description: Published as part of Zhang, Jianshuang, Pan, Tianqin, Zhang, He, Xing, Yuanqian, Yu, Hao & Zhong, Yang, 2025, Species delimitation of newly collected spiders of the genus Pseudopoda (Araneae, Sparassidae) from Honghe Hani and Yi Autonomous Prefecture, Yunnan, China: An integrated morphological and molecular approach, pp. 141-171 in Zoosystematics and Evolution 101 (1) on pages 141-171, DOI: 10.3897/zse.101.136177