Clivina geiseri Balkenohl, 2025, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina geiseri

description: Clivina geiseri sp. nov.
(Figs 19, 54, 88, 114)
Type material. Holotype: ♂, with labels and data: white, black printed “ SARAWAK: [underlined in yellow] foot of Mt.Dulit, junction of rivers Tinjar & Lejok. 29.viii.1932.” / white, black printed “Recent Clearing” / white, black printed “Old secondary forest” / white, black printed “ Oxford Univ. Exp. B.M.Hobby & A.W.Moore. B.M.1933- 254” white, black printed and handwritten in grey ink “ Clivina sp. H. E.Andrewes det.” / white, black printed and handwritten in black ink “ C. subdepressa Kt. det. K. Kult 1950” / red, black framed and black printed “ PARATYPE ” / circular, yellow framed, black printed “Para-type” (NHMUK). Paratypes: 1 ♀, with labels and data: white, black printed SARAWAK (Borneo), ca 25 km E KAPIT III.1994, Kodada leg. (NHMW); 2 ♂♂, same data as HT (NHMW); 2 ♀♀, same data but ca 40 km E KAPIT (NHMW).
Remark. The holotype represents the paratype designated by Kult (1951) as C. subdepressa which is not conspecific with the holotype of C. subdepressa (see remarks under C. subdepressa).
Description. Measurements. Body length 4.49–4.87 mm (ẍ = 4.72 mm *), width 1.31–1.41 mm (ẍ = 1.36 mm *), ratio L/W of pronotum 0.93–0.97 (ẍ = 0.95*), ratio L/W of elytra 1.82–1.88 (ẍ = 1.85*); (*n = 6). Colour glossy, fulvous. Supraantennal plate and clypeal-wing translucent. Antennae, mouthparts, intermediate and hind legs yellowish.
Head a quarter narrower than pronotum. Clypeus with central part straight, with distinctly projecting teeth laterally, separated from clypeal-wing by distinct notch; wing distinct, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin laterally obtuse-angular, clypeus, wing, and supraantennal plate slightly reflexed margined. Supraantennal plate extended posteriorly into a somewhat widened keel, separated from wing by furrow. Clypeus with transverse slightly convex field, clypeal-furrow flattened. Frons moderately convex, with indistinct pore at middle, with scattered small punctures, separated from supraantennal plate and supraorbital carina by wide furrow, with clypeal seta in front; supraorbital carinae narrow, with two supraorbital setigerous punctures at middle of eye and distinctly posterior end of the gena. Neck constriction closed, formed by a step from frons, neck covered with punctures. Eye regularly sized, gena small but somewhat bulgy. Antenna elongate, reaching over base of elytron. Labrum straight anteriorly, laterally rounded, irregularly reticulated, six-setose. Mentum isodiametrically reticulated, lobe laterally convex, tip convex, medially straight, flattened, median tooth wide, slightly higher than lobes.
Pronotum (Fig. 54) with disk distinctly flattened in frontal and lateral view. Distinctly cordate, wider than long. Reflexed lateral margin slightly crenulate, convex, widest at middle; anterior angle widely rounded off, posterior angle distinctly marked by tooth, with short concavity anterior to posterior angle; lateral channel moderately wide, somewhat flattened, irregularly to isodiametrically reticulated, continuing up to base as narrow, indistinctly convex margin. Median line engraved, complete, joining and surpassing anterior transverse line. Anterior transverse line distinct, complete. Disk bilaterally with cloud of medium sized punctures in basal half, with numerous transverse wrinkles, with isodiametric reticulation laterally at base, with impression of posterior setigerous puncture with large diameter, conspicuously distant from margin. Lateral margin from posterior angle to flange of base slanted with angle of around 45°. Basal channel on flange conspicuously widened, rugose, with narrow, obtuse carina.
Elytron (Fig. 19) with disk in frontal and lateral view distinctly flattened. Shape of elytra elongate, about two-fifth longer than wide, with lateral margin nearly straight at middle but slightly diverging, maximum width posterior middle. Humerus rounded off. Reflexed lateral margin slightly crenulate in anterior half. Row of setigerous punctures in the lateral channel regular in size and distance. Scutellar striole distinct; setigerous tubercle at base of first stria distinct, with small tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, indistinctly punctate-striate, one to four free at base, five and six joining at humerus, one and two ending free at apex, three and four, and five and six joining at apex. Intervals slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven and eight not carinate, seventh running to apex as obtuse convexity, convexity of eighth ending before apex. Surface of interval one to seven glossy, interval eight with distinct isodiametric reticulation.
Hind wing fully developed.
Lower surface. Proepisternum isodiametrically reticulated, with indistinct transverse wrinkles. Sternite of abdomen with isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with two slight embayments at margin.
Legs. Protibia slender, with longitudinal reticulation, with sulcus dorsally, with three spines; basal tarsomere distinctly flattened at base, carinate, with acute tubercle and seta at middle of tarsomere. Intermediate tibia with eight tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front and intermediate leg widened.
Male genitalia (Fig. 88). Median lobe moderately and regularly arcuate at middle, apically duckbill-shaped, somewhat flattened. Parameres moderately slender, with small apophysis, slightly distorted, hyaline apically, asetose.
Female gonocoxites and epipleurite (Fig. 114). Basal gonocoxite with two long setae, apical gonocoxite elongated, slightly convex, with short and long seta basally, with two very long setae apically. Epipleurite with two setae.
Variation. In the holotype, the clypeal-furrow is indistinct at middle. In the small series of six specimens, the wing of the clypeus is more or less toothed anteriorly or rounded.
Distribution. Known from Sarawak on the island of Borneo.
Diagnosis. A small sized fulvous species with distinctly cordate pronotum and interval eight of the elytron reticulated. Three of the species with distinctly cordate pronotum, C. geiseri sp. nov., C. subdepressa Kult, and C. depressiforma sp. nov. show in addition a conspicuously elongated flange of the pronotum with flat basal carina. Clivina depressiforma sp. nov. differs mainly by the pronotum with the laterally and acutely projecting teeth of the posterior angles and the coreaceate-like opaque surface, and the conspicuously rounded off anterior angles. Clivina subdepressa Kult differs mainly by the five-setose labrum, the nearly smooth surface of the pronotum, and the smooth lateral margin of the pronotum. Moreover, the male aedeagi of the three species are very different.
Etymology. The name is a patronym in honour of Dr. Michael Geiser (NHMUK) who supported me admirably with full access to the Scaritinae Collection during my stay at the NHMUK, and who helped me to localize a lot of material of this contribution.
description: Published as part of Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), pp. 1-70 in Zootaxa 5592 (1) on pages 33-34, DOI: 10.11646/zootaxa.5592.1.1, http://zenodo.org/record/14953674