Dichotomius (Cephagonus) clypeatus
Creators: name: Nunes, Rafael V.
name: Vaz-de-Mello, Fernando Z.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Dichotomius
Dichotomius clypeatus

description: 3.5.: Dichotomius (Cephagonus) clypeatus (Luederwaldt 1929) (Figure 32 (a-e))
Pinotus (Cephagonus) bicuspis var. clypeatus Luederwaldt 1929 ı p. 129 (description) Pinotus (Cephagonus) clypeatus ı Luederwaldt 1936 ı p. 214
Diagnosis. this species is separated from other in the group by the following combined characters: males pronotal disc with a central pair of tubercles on anterior portion; elytral striae and punctures strongly impressed (as in D. ingens and D. arnaudi); males cephalic horn or tubercle rounded apically (as D. bicuspis); genae borders divergentı posterior angle rounded; paramera lacking basal nor longitudinal excavations and sides of elytra subparallel in dorsal view (as in D. arnaudi).
● HOLOTYPE (male) (Figure 32 (e)): Oxapampa Peru {whiteı printed}/ Pinotus quadraticeps Felsche {whiteı handwritten}/Muséum Paris 1936 Coll. A. Boucomont {white with black marginsı printed}/ Pinotus bicuspis Har. var. clypeatus. Clypeatus nov. var. Luederwaldt 1931 {whiteı handwritten} [MNHN].
Non-type material. ARGENTINA: Salta: Sierra Tartagal. 35 km N Tartagal. 22³19 ʹ 36 ʹ ’ Sı 63³51 ʹ 00 ʹ ’ Wı 874 m. 14.xi.2006. Berger-Dostal [1 Ƌ at CEMT]; BOLIVIA: Santa Cruz: Florida.
18³08 ’ Sı 63³48 ʹ Wı 1730m. 19-xi-2006. WD Edmonds & P Reyes [1 Ƌı 1 ♀ at CEMT]; Tarija. Entre Riosı Quebrada del Abra. 25-ii-1998. Ayzama & F Guerra [1 Ƌı 1 ♀ at CEMT].
Redescription. colour: blackı shiny. Head: fronto-clypealı genal and inter-ocular surfaces smooth. Gena borders not parallel. Cephalic carina conical with rounded apex. Pronotum: borders smoothı not serrated. Pronotal disc shinyı lacking punctures and delimited anteriorly by two central weak tubercles. Metasternum: sides and margins of anterior lobe covered by long hair. Anterior portion of anterior lobe glabrous. Setigerous punctures evenly spaced at sides. Elytra: sub-parallel. Striae deeply impressed having fine weakly impressed punctures spaced by one to three times their diameter. Interstriae convexı lacking microsculpture with very fine puncture (20 x). Abdomen: sides of each ventrite excavated. 1st ventrite with a single row of hair along anterior margin. Each side of ventrites 2 to 5 with groups of 1 – 5 ocelatte setigerous punctures. Puncture of each ventrite interrupted medially as well as groove along the posterior margin of sixth ventrite. Aedeagus (Figure 32 (b-d)). Dorsallyı paramere apexes roundedı convergent; longitudinal and basal excavations lacking; membranous portion taking more than 2/3 of dorsal surface. Laterallyı subtle declivity from basis towards apex. Ventrallyı subgenital plate widely emarginate at apex and rounded at basis.
Morphological variation. Male BL: 13 – 15 mmı PW: 7 – 9 mm. Female BL: 16 mmı PW: 6 mm and distinguished from males as follows: Head: dorsal surface striated. Clypeo-genal junction very obtuseı rounded. Cephalic carina absent forming at most a central weak knob. Pronotum: simply convex. Elytra: interstriae microsculpture chagrinated. Abdomen: sixth ventrite three times larger than fifthı pygidium swollenı lacking modifications at medial portion.
Distribution. Argentina (Salta)ı Bolivia (Santa Cruzı Tarija) and Peru (Oxapampaı type locality)
description: Published as part of Nunes, Rafael V. & Vaz-de-Mello, Fernando Z., 2019, Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini), pp. 2231-2351 in Journal of Natural History 53 (37) on pages 2310-2311, DOI: 10.1080/00222933.2019.1692088, http://zenodo.org/record/3665519