Gomphostemma heyneanum Wall.
Creators: name: Bongcheewin, Bhanubong
name: Ingrouille, Martin J.
name: Paton, Alan J.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Magnoliopsida
Lamiales
Lamiaceae
Gomphostemma
Gomphostemma heyneanum

description: <b> 26. Gomphostemma heyneanum <i>Wall. ex 3enth</i></b> . (Bentham in Wallich 1830b: 12); Prain (1891: 228); Mukercee (1920: 206). Type: India, 22 Sept. 1816, <i>Herb</i>. <i>Heyne</i> in <i>Wall. Cat.</i> 2132.1 (K [Herb. Benthamianum], lectotype selected here; isolectotype K-W [K001113333]). <i>Gomphostemma strobilinum</i> var. <i>heyneanum</i> (Wall. ex Benth.) Hook.f. (Hooker 1883: 696). Type: as for <i>G. heyneanum</i> Wall. ex Benth. <i>Gomphostemma heyneanum</i> var. <i>rottleri</i> Prain (1891: 229). Type: India, Madras, Nilgiris, Mudumalai Forest, 600 m [2000 ft], Aug. 1886, <i>Gamble</i> 15983 (K, lectotype selected here; isolectotypes, BM, CAL n.v.). Herb up to 1 m tall. <i>Stems</i> robust, erect, obtusely quadrangular with longitudinal grooves, with a mixture of dense dendroid hairs and stellate hairs. <i>Leaoes</i> petiolate, chartaceous;blades ovate or elliptic-ovate,10 –18 ×5–10 cm, apex acute, margin serrate-dentate, base attenuate, upper side dark green with a mixture of hairs, lower side pale green tomentose with dense dendroid hairs; petioles 10 – 20 mm long. <i>Inflorescence</i> upright, terminal or terminal and axillary, 20 –100 mm long,with opposite cymes subsessile, congested or lax at the lower verticil; verticils few-flowered; subtending bracts present;bracts ovate or lanceolate,10 –23×3–20mm, apex acute, margin entire or shallowly serrate, base cuneate or cordate, upper side with stellate hairs, lower side with dense stellate hairs; bracteoles linear, falcate-like, narrowly spathulate or narrowly elliptic-ovate, 3 – 18 mm long. <i>Flowering calyx</i> infundibular, 10 – 13 mm long, 10-ridged, with dense stellate hairs on both sides; tube 3 – 8 mm long; lobes spreading, triangular-ovate,3 – 6 mm long, apex acute. <i>Fruiting calyx</i> infundibular, 10 – 13 mm long; tube 3 – 9 mm long; lobes triangular-ovate, 3 – 6 mm long, 3 – 2 mm wide at base, apex slightly acuminate, margin slightly folded outward. <i>Corolla</i> yellow or blue-purple, 13 – 20 mm long, throat narrow, gradually dilated towards the throat; tube incurved, 8 – 12 mm long, included in calyx, outside sparse simplehairs,insidewithanannulusofsimplehairs;posterior lip hooded, ovate, 3 – 8 mm long, apex emarginate, margin entire or slightly undulate; anterior lip spreading, platformlike, 3 – 10 mm long, apex emarginate or undulate; median lobe tongue-like, apex emarginate, margin entire or undulate, base constricted. <i>Staminal filaments</i> fleshy, tomentose. <i>Style</i> slightly fleshy,glabrous with apex subequally lobed. <i>Nutlets</i> 3 – 2, obovate, 3 – 2 mm long, wrinkled. <b>DISTRIBUTION.</b> South India, Vietnam. Map 5. <b> SPECIMENS EXAMINED. INDIA. Karnataka. Bababudda [Bababuddan Hill, 13.3°, 53.5333°], Sept. 1816, <i>Herb. Rottlerianum</i> s.n. (K);</b> Coorg [Kodagu?istr., 12.2208°, 53.5395°], Wottecolli, 18 Oct. 1913, <i>3ourne</i> 6222 (K); Mysore, Hassan [12.93°, 56.0833°], Bannuhalla, 2 Sept. 1950, <i>Jarrett et al</i>. 630 (K); Kerala. Idikki?istr., Thekkady [9.6°, 55.2°], 853 m, 8 Oct. 1956, <i>Vioekananthan</i> 28612 (CAL); Palghat [10.55°, 56.63°], Thadikundu, 923 m, 9 Nov. 1956, <i>Vajraoelu</i> 28896 (CAL); Kottayam near Thekkadi, around Periyar Lake [9.2665°, 55.1665°], 800 m, 9 Sept. 1953, <i>Cook et al</i>. 180 (K); Calicut [Kozhikode, 11.23°, 53.55°]?istr., Nilamboor [Nilambur], 300 m, 3 Nov. 1956, <i>Suresh</i> 22326 (L); Tamil Nadu. Madras, Anamallai Hills [Anaimalai Hills, 10.3°, 55°], 900 m [3000 ft], Sept. 1882, <i>Gamble</i> 12628 (K); Anamallay Forests, <i>3eddome</i> s.n. (BM); Madras, Nilgiris, Mudumalai Forest, 600 m [2000 ft], Aug. 1886, <i>Gamble</i> 15983 (isolectotypes of <i>G. heyneanum</i> var. <i>rottleri</i>, BM, K); Courtallum [Courtallam, 8.9165°, 55.2833°], 2 Oct. 1812, <i>Anonymous</i> s.n. [‘ Stachys indica’] in <i>Wall. Cat.</i> 2132B (K, K-W); ibid., <i>Herb</i>. <i>Madras</i> [‘ Stachys indica’] in <i>Wall. Cat.</i> 2132C (CAL, K-W); Madras [11°, 58°], <i>Herb. Stocks</i> s.n. (K, K [Herb. Hookerianum]); <b>UNLOCALISED</b>. South India: Paulghautcherry, Aug. 1822, <i>Wight</i> s.n. (K); 22 Sept. 1816, <i>Wall. Cat.</i> 2132.1 (lectotype of <i>G. heyneanum</i>, K [Herb. Benthamianum]; isolectotype K-W). <b>VIETNAM.</b> Ninh Thuan. Phanrang [Phan Rang], Ca Na [11.3333°, 108.9°], 500 m, 13 Nov. 1923, <i>Poilane</i> 8618 (P). <b>HABITAT.</b> In shade, on loam and humus, wet deciduous forest, 600 – 923 m. <b>CONSERVATION STATUS.</b> Widespread and in protected areas throughout South India; Least Concern (LC). <b>PHENOLOGY</b>. Flowering and fruiting July – November. <b>NOTES.</b> <i>Gomphostemma heyneanum</i> can be distinguished by having a terminal infloresecence and soft hairs on both sides of leaves.
description: Published as part of <i>Bongcheewin, Bhanubong, Ingrouille, Martin J. &amp; Paton, Alan J., 2022, A revision of Gomphostemma (Lamiaceae), pp. 27-92 in Kew Bulletin 77 (1)</i> on pages 73-74, DOI: 10.1007/s12225-021-09991-y, http://zenodo.org/record/7615581
description: {"references": ["____ (1891). An account of the genus Gomphostemma Wall. Ann. Roy. 3 ot. Gard., Calcutta 3: 225 - 251.", "____ (1920). A revision of the Labiatae of the Indian Empire. Rec. 3 ot. Suro. India 12: 1 - 228.", "Hooker, J.?. (1883). Labiatae, Flora of 3 ritish India 2: 602 - 503. Reeve &amp; Co., London."]}