Puncturella Lowe 1827
Creators: name: Kiel, Steffen
name: Campbell, Kathleen A.
name: Gaillard, Christian
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Lepetellida
Fissurellidae
Puncturella

description: <i>Puncturella</i> (sensu lato) <i>mcleani</i> sp. nov. (Fig 2) <b>Diagnosis:</b> Tall <i>Puncturella</i> with foramen on lower half of anterior slope; apex bent posteriorly and situated directly above posterior margin; apparently no external sculpture. <b>Holotype:</b> UCMP 55740, L = 7 mm, H = 6 mm, W = 4 mm. <b>Paratypes:</b> UCMP 557041, L = 8 mm, H = 5 mm, W = 5.5 mm, from NW Berryessa limpet [UCMP loc. A-4500, loc. 10 in Kiel <i>et al.</i> (2008)]; UCMP 557042, L = 6 mm, H = 4.5 mm, W = 3 mm, from the type locality; LACMIP 13558, L = 6 mm, H = 5 mm, and LACMIP 13559: L = 5 mm, H = 3.5 mm, both from Paskenta [loc. 2 in Kiel <i>et al.</i> (2008)]. <b>Further material examined:</b> Two further specimens from Wilbur Springs and Rocky Creek [locs. 5 and 7 in Kiel <i>et al.</i> (2008)] are deposited in the CAS. <b>Type locality:</b> The Bear Creek seep site, California, USA, 39°03'14"N, 122°24'40" W; Crack Canyon formation of the Great Valley Group; Early Cretaceous, Valanginian (Kiel <i>et al.</i> 2008). <b>Description:</b> Tall limpet, aperture elongate-oval in outline; apex bent posteriorly, situated vertically above posterior margin of aperture; posterior slope slightly concave, anterior slope gently convex; outer surface apparently smooth; foramen on lower half of anterior slope, about four-and-half times longer than wide. <b>Remarks:</b> The absence of sculpture is unusual for <i>Puncturella</i> and fissurellids in general; however, only a single specimen with fragments of shell material was available. If the absence of sculpture is confirmed this species might be better placed in a new genus. The record from the Tithonian (Late Jurassic) site at Paskenta makes this the oldest slit limpet at fossil cold seeps. Species of <i>Puncturella</i> found at modern vents and seeps are quite different from <i>Puncturella</i> (s.l.) <i>mcleani</i>: they are not as tall, have a more centrally located apex and radial sculpture (Okutani <i>et al.</i> 1993; Beck 1996; Warén and Bouchet 2009). Vent-inhabiting fissurellids without radial sculpture belong to the genus <i>Cornisepta</i> McLean &amp; Geiger, 1998; however, they are not smooth but rather have tuberculate sculpture. They are quite different from <i>Puncturella</i> (s.l.) <i>mcleani</i> in being tall, and having a centrally located apex and foramen (McLean and Geiger 1998). Two further fissurellids are known from the Cretaceous of California, which differ from <i>Puncturella</i> (s.l.) <i>mcleani</i> by being lower and having radial sculpture. They include <i>Fissurella</i> (sensu lato) <i>bipunctata</i> Stanton, 1895 from “near Stephenson’s, on Cold Fork of Cottonwood Creek, Tehama county, California ”; this is a seep deposit of Albian age [the Cold Fork of Cottonwood Creek site of Kiel <i>et al.</i> (2008)]. The holotype is figured here (USNM 23065; Fig. 2G); it is worth noting that this species has a seemingly rectangular apical opening and that the two central radial ribs on the anterior slope are stronger than the other radials; Stanton’s (1895) drawing does not show this. These two strong radials, however, do not mark the edges of a slit. The other species is <i>Fissurella gabbi</i> Stewart, 1927, from the Chico Formation (Coniacian-Campanian) and has, according to Stewart (1927), 20 radial ribs while <i>Fissurella</i> (sensu lato) <i>bipunctata</i> has 27. We have one specimen of a tall limpet resembling <i>Puncturella</i> (s.l.) <i>mcleani</i> in general shape from the Bear Creek seep deposit. But in contrast to <i>Puncturella</i> (s.l.) <i>mcleani</i>, this specimen has radial ribs and shows no sign of a slit or foramen. <b>Distribution:</b> Late Jurassic (Tithonian) to Early Cretaceous (Valanginian), Great Valley Group, California, USA. <b>Etymology:</b> For James H. McLean, Los Angeles.
description: Published as part of <i>Kiel, Steffen, Campbell, Kathleen A. &amp; Gaillard, Christian, 2010, New and little known mollusks from ancient chemosynthetic environments, pp. 26-48 in Zootaxa 2390 (1)</i> on pages 29-31, DOI: 10.11646/zootaxa.2390.1.2, http://zenodo.org/record/5305157
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