Bucculentum horstkuscheli Krzemińska & Starzyk & Fraaije & Schweigert & Lukeneder 2021, n. sp.
Creators: name: Krzemińska, Ewa
name: Starzyk, Natalia
name: Fraaije, René H. B.
name: Schweigert, Günter
name: Lukeneder, Alexander
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Bucculentidae
Bucculentum
Bucculentum horstkuscheli

description: Bucculentum horstkuscheli,  n. sp.   
Fig. 1E, F, 6  
 Diagnosis. Hepatic regions large, much wider than metabranchial regions; metabranchial region not expanded laterally; augenrest directed laterad at c. 60 o to body long axis and aligned with proximal portions of hepatic region; suborbital spine wide, directed outwards and downwards, and ending in two short, triangular teeth; cardiac region with one distalmost dominant tubercle accompanied anteriorly by row of small tubercles.   
 Etymology. Trivial name is dedicated to Horst Kuschel (Faurndau), who found and donated the holotype of this species.   
 Material examined. Holotype: # SMNS 70607 (leg. H. Kuschel), Germany: Bad Überkingen-Oberböhringen, near Geislingen a. d. Steige. Age: Late Kimmeridgian: Acanthicum Zone (Fraaije et al. 2017).   
 Description. The holotype represents an internal mould, with only the traces of the white cuticle in the grooves. It is excellently preserved and symmetrical, lacking only the rostrum. The hepatic regions are very wide, c. 1.4x wider than the mesobranchial region and 1.2x wider than the metabranchial region. The frontal portions containing the augenrests and anterior portions of hepatic regions are aligned, obliquely set to the long axis at an angle of 57 o, symmetrical on both sides. Augenrests are almost perpendicular to the plane of carapace, only very slightly shifted dorsally (Fig. 6B), at an angle of c. 80 o to the upper surface of the carapace; they are laterally flanked by single, broad, triangular tubercles. The suborbital spine is very broad, supporting most of the lower margin of the augenrest, directed down- and outwards, and terminating in two short, triangular teeth (Fig. 6B, C, arrows). The epibranchial region is narrow; the cervical groove is almost straight. The cardiac region is equipped at its distal end with a large tubercle (which is the highest point of the carapace) and a median row of small tubercles towards the mesobranchial region. The metabranchial regions are narrow, almost rectangular; the distal end is broad, stretched over the entire distal portion of the metabranchial region and upturned.   
 Remarks. The new species resembles B. bachmayeri in having a large hepatic region, a divided suborbital spine, and a cardiac region with one large tubercle, but differs in having the augenrests directed outwards and positioned obliquely to body long axis. The suborbital spine is divided into two short, triangular lobes, unlike the long, thin prongs in B. bachmayeri.
description: Published as part of Krzemińska, Ewa, Starzyk, Natalia, Fraaije, René H. B., Schweigert, Günter & Lukeneder, Alexander, 2021, Jurassic brachyurans of the genus Bucculentum, pp. 395-410 in Zootaxa 5032 (3) on page 402, DOI: 10.11646/zootaxa.5032.3.5, http://zenodo.org/record/5497036