Furagrion jutlandicus
Creators: name: Archibald, S. Bruce
name: Ware, Jessica L.
name: Rasmussen, Jan A.
name: Sylvestersen, René L.
name: Olsen, Kent
name: Simonsen, Thomas J.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Odonata
Megapodagrionidae
Furagrion
Furagrion jutlandicus

description: Furagrion jutlandicus (Henriksen, 1922)
Figs. 1–4, 6–8, 11–18.
Furagrion morsi Zessin, 2011, syn. nov.
Emended diagnosis. The wings of Furagrion jutlandicus may be distinguished from those of Furagrion ansorgei by shape: both fore- and hind wings have smaller length/width ratios in all measurements except RP2 to apex / width (Tables 2 and 3, and see PCA, above); and by the IR1-RP2 space becoming more than one cell wide at about level of basal end of pterostigma or slightly basal to that [F. ansorgei: distad distal end of pterostigma (tentative, see below)].
Emended description. As for genus description and diagnosis, species diagnosis, and see forewing, hind wing measurements and ratios of these for the proposed neotype and additional specimens in Tables 1 and 2.
Type material. Proposed neotype MM-10752 (previously MHM-17-42) (Figs. 3–4): head mostly indistinct, with faintly preserved left compound eye, complete thorax and abdomen, all four wings, in soft diatomite, coll. Henrik Madsen at Skarrehage mo-clay pit, Mors, Knudeklint Member (informally the Skarrehage bed, 0.6–1.0 m below ash layer -13), Fur Formation in December 2009, in the Fossil and Mo-clay Museum, Museum Mors, Nykøbing Mors, Denmark.
Other material. MM-10750 (previously MHM-17-40) (Figs. 7–8): head missing the compound eyes, complete thorax and almost complete abdomen, most of all four wings, parts of five legs, in soft diatomite, coll. Henrik Madsen at Skarrehage mo-clay pit, Mors, Knudeklint Member (informally Skarrehage bed), Fur Formation, December 2009, in the Fossil and Mo-clay Museum.
MM-4785 (previously MHM-5242A) (Figs. 11–12): head missing, almost complete thorax and abdomen, one forewing with damaged posterior margin and two hind wings in varying states of incomplete preservation, in soft diatomite, coll. by unknown museum guest at the Ejerslev mo-clay pit, Mors, Knudeklint Member, Fur Formation, June 21, 2013, in the Fossil and Mo-clay Museum.
FUM-N 13856 (Fig. 2): an almost complete forewing missing a portion of the posterior at about a third length, in a concretion, coll. Erwin Rettig at Klitgaard Cliff, Mors, Fur Formation, date unknown, in the Fur Museum.
FUM-N 11146 (previously ERK GU E7) (Fig. 13): an almost complete forewing missing parts of the wing basad the nodus, coll. Erwin Rettig at Gullerup; Mors, Fur Formation, date unknown, in the Fur Museum.
FUM-N 16192 (Fig. 14): an almost complete hind wing missing parts basad about Ax2, in a concretion, coll. Jan and Elly Verkleij at the Ejerslev mo-clay pit, Mors, Knudeklint Member, Fur Formation, 2008, in the Fur Museum.
14M-2765 (Fig. 15): a partial thorax and basal fragment of abdomen with a [fore- or hind?] wing attached, the distal posterior portion of which is folded over the rest of the wing, coll. Henrik Madsen at the Ejerslev mo-clay pit, Mors, Knudeklint Member, Fur Formation, November 14, 1991, in the negative series, in the Fossil and Mo-clay Museum, Museum Mors.
FUM-N 11616 (Fig. 16): two very complete hind wings joined by a small fragment of thorax, in a concretion, coll. Jan and Elly Verkleij at Manhøj, new pit, Fur Formation, 2008, in the Fur Museum.
FUM-N 14704 (previously FM 1106) (Fig. 17): an almost complete forewing missing a small portion of the apex, coll. Erik Fjeldsø at Stolleklint Cliff, Silstrup Member, between ash layers +25 and +30, Fur Formation, date unknown, in the Fur Museum.
MM-2085 (previously MHM-i1583AB): (Fig. 18): an almost complete forewing in a concretion, coll. Henrik Madsen at Ejerslev mo-clay pit, Mors, Knudeklint Member, between ash layers +25 and +30, Fur Formation, 2005, in the Fossil and Mo-clay Museum, Museum Mors.
MGUH 1819 (Fig. 1): the distal portions of three wings and much of the abdomen, coll. and date unknown, Struer, loose slab from an unknown level within the Fur Formation, in the Geological Museum, University of Copenhagen, Denmark.
14M-A2163 (Fig. 6) (MM 14M-A2163 of Rust 1999, see his plate 1, fig. b): with a well-preserved head, including compound eyes, thorax, and most of the abdomen, the anterior of a forewing and much of a hind wing, parts of two legs, coll. Henrik Madsen at Ejerslev mo-clay pit, Mors, Knudeklint Member between ash layers -11 and -13, Fur Formation, August 21, 1993, in the Fossil and Mo-clay Museum. Although the distal IR1-RP2 region is missing and the maximum wing width is not known, the preserved portion of the hind wing is shaped like those of F. jutlandicus, clearly not as slender as the F. ansorgei wing.
14M-3801 (Fig. 19): missing the head, thorax, and abdomen mostly complete, one forewing and one hind partly preserved, parts of three legs, in a concretion, coll. Henrik Madsen at the Ejerslev mo-clay pit, Mors, Knudeklint Member between ash layers -13 and -14, Fur Formation, October 2, 1992, in the Fossil and Mo-clay Museum, Museum Mors. Although the distal IR1-RP2 region is missing, its wing shape is not as slender as the F. ansorgei wing.
MOA 769/1, 2 (part, counterpart), “ Furagrion morsi ” is known by the basal portion of a wing (Fig. 20). We could not locate the fossil and based our discussion on the description, photograph and drawing of Zessin (2011). We find its putative differences with F. jutlandicus and F. ansorgei to be within their intraspecific variations (Fig. 20B–D). The shape of its preserved portion is like F. jutlandicus not as slender as the F. ansorgei wing, and so we treat “ Furagrion morsi ” as a junior synonym of F. jutlandicus.
MM-11044 (Fig. 21): basal portion of a wing in a concretion, coll. Henrik Madsen, 2018, Ejerslev mo-clay pit, Mors, Knudeklint Member, between ash layers +25 - +30, Fur Formation, in the Fossil and Mo-clay Museum, Museum Mors. Incomplete and lacking the IR1-RP2 space. Although the distal portion of the wing is missing, we tentatively assign this fossil to F. jutlandicus, as its preserved portion appears not as slender as an equivalent portion of the F. ansorgi wing.
Remarks. The proposed neotype MM-10752 was figured by Pedersen et al. (2012, fig. 46D) who characterised it as a damselfly (translated from Danish: vandnymfe) from the level just below ash layer - 13 in the Skarrehage mo-clay pit, Mors. The specimen was not named or described in further detail.
Range and age. Earliest Ypresian Fur Formation, Jutland, Denmark. The proposed neotype is from the Skarrehage mo-clay pit (N 56°56.65 –, E 8°52.41 –) between 61 and 100 cm below ash layer -13, informally named the Skarrehage bed of the Knudeklint Member, Fur Formation. All specimens: as for genus, above.
description: Published as part of Archibald, S. Bruce, Ware, Jessica L., Rasmussen, Jan A., Sylvestersen, René L., Olsen, Kent & Simonsen, Thomas J., 2023, The damselfly genus Furagrion Petrulevičius et al. (Odonata, Zygoptera) from the early Eocene Fur Formation of Denmark and the dysagrionoid grade, pp. 289-317 in Zootaxa 5278 (2) on pages 304-310, DOI: 10.11646/zootaxa.5278.2.4, http://zenodo.org/record/7906172