Hercostomus golanensis Grichanov, 2015, sp. nov.
Creators: name: Grichanov, Igor Ya.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Dolichopodidae
Hercostomus
Hercostomus golanensis

description: Hercostomus golanensis sp. nov.
(Figs 1–8)
Material. HOLOTYPE ♂, Israel: Golan (Heights), Qatzrin, 30.05.1981, F. Kaplan (TAU). Paratype: 1 ♂, Israel: (Golan Heights), Tel Dan, 10.vii.1975, A. Freidberg (TAU).
Diagnosis. Face under antenna inconspicuously wider than postpedicel height; postpedicel subtriangular, about as long as high; arista-like stylus long-pubescent, 2.2X longer than antennomeres combined; wing distinctly fumose; wing veins R4+5 and M1+2 parallel along most of their length, slightly convergent near wing apex; hypopygium with triangular cercus without scale-like setae; hypandrium smoothly curved, nearly reaching apex of apicoventral epandrial lobe; apicoventral epandrial lobe bearing 2 setae on its knob and 1 very long seta at base of stem.
Etymology. The species is named after the Golan Heights, where the types were collected.
Description. Male (Fig. 1): Head: antenna mostly darkish yellow, distal half of postpedicel brown (Fig. 2); postpedicel subtriangular, hardly longer than height; arista-like stylus mid-dorsal, 2.2X longer than antennomeres combined; distal segment of arista with all hairs at least 2X longer than basal diameter of stylus, sparse at apex; length ratio of scape to pedicel (inner view) to postpedicel to stylus (segments 1 and 2), 11/12/22 /23/49; frons black with greenish reflection, grey pollinose, nearly parallel-sided, slightly wider than height, with distinct dot-like depression in middle of lower part; eyes finely haired; face black, whitish-grey pollinose; ratio of height to width under antennae to width at clypeus, 0.40/0.20/0.15; clypeus small, bulging; palpus small, yellow, with few black hairs and 1 black seta; proboscis short and broad, yellow brown; lower postocular setae yellow, upper postoculars black. Thorax: all setae black; pronotum densely pubescent, with black hairs; mesonotum black with bluish reflection, weakly grey pollinose; 6 dorsocentral setae, 12–13 acrostichals, short, biserial, reaching scutellum; pleura entirely dark, with grey pruinosity, in front of posterior spiracle bare; metapleuron with vertical anteroventral row of about 6 short setae; scutellum mainly brownish-yellow, with black triangular spot on dorsum, with 2 strong medial setae and 2 fine lateral setae. Legs: almost entirely yellow, mid coxa with brown spot, 2–3 distal tarsomeres brown, hairs and setae black; all tarsi simple; foreleg with coxa bearing numerous black hairs anteriorly and 3 strong setae apically, fore femur with preapical posterior seta, fore tibia with 2 anterodorsal, 2 posterodorsal, 2–3 apical setae, with anterodorsal comb-like row of setae distally, no long apicoventral seta; midleg with coxa bearing some black hairs laterally and 2–3 strong setae apically, mid femur with 1 anterior and 1 posteroventral preapical strong seta, mid tibia with 3 anterodorsal, 2 posterodorsal, 1 anteroventral, 1 posteroventral and 5 apical bristles; hindleg with coxa bearing 1 strong seta at middle and few adjacent hairs, hind femur with 1 anterior preapical bristle at apical quarter, hind tibia with 3 anterodorsal, 3 posterodorsal, 3 apical bristles and 1 short ventral seta at middle; hind basitarsus with 2–3 ventral setae, distinctly shorter than next tarsomere. Femur, tibia and tarsomere (from first to fifth) length ratio (in mm): fore leg: 1.03/1.07/0.53/0.19/0.13/ 0.11/0.15, mid leg: 1.37/1.46/0.76/0.40/0.33/0.20/0.17, hind leg: 1.52/1.66/0.52/0.64/0.39/0.30/0.18. Wing (Fig. 3): nearly 3 times longer than wide; membrane regularly fumous, veins brown; costa simple, with short black setae dorsally; R2+3, R4+5 and M1+2 almost straight; R4+5 and M1+2 subparallel; M joining costa at wing apex; ratio of costal section between R2+3 and R4+5 to that between R4+5 and M1+2: 0.38/0.23, distal part of CuA1 1.5X longer than dm-cu; anal vein distinct; anal angle obtuse; lower calypter yellow with black cilia; halter yellow. Abdomen: bluishgreenish-black and shiny, weakly pollinose, with black hairs and long marginal setae; 8th segment pear-like, black, with black cilia dorsally. Male genitalia (Figs 4–8): epandrium black, rhomboid, as long as terga 3–5 combined; hypandrium mainly yellow, mostly free, fused to epandrium at extreme base, smoothly curved, nearly reaching apex of apicoventral epandrial lobe; phallus thin and long; basiventral epandrial lobes symmetrical, each lobe reduced to short seta at base of hypandrium; apicoventral epandrial lobe yellow, arising rather distally, far from base of hypandrium, prominent, with distinct apical expansion, bearing 2 setae on knob and 1 very long seta at base of stem; anteroventral part of postgonite not developed; posterodorsal portion poorly developed, horn-like, slightly curved ventrally, swollen at base, with reduced lateral lobes; surstylus bilobed, ventral lobe elongate-ovate, without dorsal hump, with short apical setae, dorsal lobe thin, expanded at base, slightly curved dorsally, with short apical setae and 1 strong and very long basodorsal seta; cercus yellow-brown in basal half, blackish distally, triangular, nearly 2/3rds epandrium length, nearly 2X as long as wide, with row of long black marginal setae and adjacent inner row of shorter light setae, without modified setae. Measurements: Body length 4.2 mm, antenna length 1.0 mm, wing length 3.8 mm, wing width 1.25 mm. Female: Unknown.
description: Published as part of Grichanov, Igor Ya., 2015, Palaearctic species of the Hercostomus plagiatus group (Diptera: Dolichopodidae) with description of a new species from the Middle East, pp. 424-432 in Zootaxa 3918 (3) on pages 427-429, DOI: 10.11646/zootaxa.3918.3.6, http://zenodo.org/record/232295