Achilia longispina Franz 1996
Creators: name: Kurbatov, Sergey A.
name: Cuccodoro, Giulio
name: Sabella, Giorgio
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Achilia
Achilia longispina

description: Achilia longispina Franz, 1996
Figs 29, 34, 43, 52, 78, 80-82, 84
Achilia longispina Franz, 1996: 116 fig. 65 (aedeagus).
Type material (1 ex.): CENTRAL CHILE: Región Valparaíso: Valparaíso prov.: NHMW (coll. Franz); 1 ♂ (holotype); Mina de la Disputada (now Mina los Bronces), Anden bei Santiago de Chile; 3000-3400 m; 19.XI.1968; H. Franz.
Additional material (12 ex.): See Appendix 1.
Description: Body 1.6-1.7 mm long, entirely reddish or reddish brown, with palpi yellowish. Pubescence decumbent with long setae uniform on entire body. Head wider than long; lacking vertexal sulcus; eyes protruding and slightly pointed, longer than convex temples. Pronotum slightly wider than long, narrower than head, with maximal width at midlength; lateral outlines on anterior portion convergent, on posterior portion subparallel and sinuate; disc slightly convex, smooth and shiny with some punctures; basal margin bordered with row of contiguous shallow impressions; median antebasal fovea smaller than lateral ones. Elytra together wider than long with slightly protruding humeri; disc smooth, shiny, with two big basal foveae; sutural stria entire; discal stria extending to about elytral midlength. Legs elongate. Abdomen smooth, with some minute punctures; tergite I with basal striae slightly divergent, extending to about one-quarter of paratergal length, separated at base by more than one-third of tergal width, with some short and sparse setal brushes between striae.
Male: Head as in Figs 78 & 80-81, with deep lateral pits on each side of eyes, the latter with small median spine on upper margin (Fig. 82); frontal protuberance very large, raised, and flattened, densely covered with large punctures; clypeal area deeply hollowed laterally. Antennae (Fig. 34) with scape and pedicel distinctly longer than wide; antennomeres III-IV as long as wide; antennomeres V-VI longer than wide; antennomere VII as long as wide; antennomere VIII slightly wider than long; antennomere IX wider that long, with protruding mesal margin pointed in middle; antennomere X wider than long, wider and longer than IX, with protruding mesal margin indented and bearing numerous setae; antennomere XI longer than wide, narrower than X and longer than VIII-X combined, its surface bearing small tubercles. Metasternum with deep and broad ovoidal median impression, posterior edge of impression pubescent. Legs with ventral margin of mesotrochanters (Fig. 43) bearing short seta; profemora and mesofemora slightly thickened, finely punctured; protibiae slightly thickened; mesotibiae (Fig. 52) in middle slightly bulging and bearing tuft of short setae; metatibiae slightly thickened and sinuate on basal third. Abdominal tergites and ventrites unmodified. Aedeagus (Fig. 29) 0.26- 0.27 mm long; dorsal plate ovoid with dorsal longitudinal struts divergent; copulatory pieces consisting of pair of long medial sclerites basally recurved and sclerotised and apically pointed, associated on each side with small subapical sclerite. Parameres very wide with very long seta on well-developed outer lobe; tips strongly recurved posteriorly and bearing wide and long subapical seta.
Female: Similar to male except: head, metasternum, and legs unmodified; antennomeres X-XI shorter and thinner than male.
Distribution: Achilia longispina is known only from its type locality: Mina de la Disputada (now Mina los Bronces) in Central Chile, Valparaíso Province (Fig. 84: yellow stars).
Comments: The external morphology and aedeagal features of A. longispina resemble more closely A. cribratifrons Jeannel, 1962 and A. angulifrons Jeannel, 1963 than A. frontalis, as stated by Franz (1996: 116). Therefore we here exclude A. longispina from the A. frontalis group, but at this stage of the revision of the genus we prefer not to assign it to another group.
description: Published as part of Kurbatov, Sergey A., Cuccodoro, Giulio & Sabella, Giorgio, 2018, A revision of the Chilean Brachyglutini – Part 3. Revision of Achilia Reitter, 1890: A. frontalis species group (Coleoptera: Staphylinidae: Pselaphinae), pp. 165-188 in Revue suisse de Zoologie 125 (1) on pages 183-184, DOI: 10.5281/zenodo.1196048