Allometopon
Creators: name: Owen Lonsdale
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Clusiidae
Allometopon

description: Key to species of Allometopon
1. Second dorsocentral inserted closer to transverse suture than to posterior dorsocentral. Anterior dorsocentral approximately 3/ 5 length of posterior dorsocentral; third smaller dorsocentral often present. Posteromedial region between dorsocentrals usually glossy. Two fronto-orbitals present, uncommonly three (Indian A. bharetum).................................. 2
- Second dorsocentral inserted closer to posterior dorsocentral than to transverse suture. In all cases where a third weak dorsocentral is present, all dorsocentrals are obviously displaced posteriorly. Anterior dorsocentral usually 1/2 length of posterior dorsocentral. Posteromedial region between dorsocentrals at least partly setose and microsetulose. Three or four fronto-orbitals, rarely two (A. atromaculatum, A. antennalis)............................................................... 7
2. Three fronto-orbital setae. Scutum setulose between dorsocentrals (Fig. 24). Surstylus extremely long,narrow, apically truncated (Fig. 26). India ................................................................ A. bharetum spec. nov.
- Two fronto-orbital setae. Scutum glossy and bare between dorsocentral setae, sometimes with setulae only slightly reduced (A. hauseri). Surstylus usually pointed, sometimes shorter and apically truncated (Fig. 287), never as elongate not as above.... 3
3. Frons mostly or entirely dark brown. Back of head brown above foramen. Ocellar seta as long as posterior fronto-orbital. Two dorsocentral setae. Two lateral scutellar setae. Surstylus produced posteriorly as a ventrally hooked lobe (as in Fig. 21). Basiphallus/epiphallus developed into compact segmented structure............................................ 4
- Frons yellow. Back of head entirely or almost entirely yellow. Ocellar seta minute. Long, well-developed setula in front of anterior dosocentral. One lateral scutellar seta. Male terminalia not as above...................................... 5
4. Frons entirely dark brown. Scutum dark brown with yellowish spots on postpronotum and notopleuron (Fig. 19). Scutellum dark brown. Male anepisternum entirely yellow (Fig. 20); female antepisternum entirely dark brown. Metanotum entirely dark brown. Tergites 1 and 2 brown. Surstylus strongly arched posteriorly (Fig. 21). Distiphallus elbowed at midpoint (Fig. 23). Paraphallus very long and narrow with apex expanded and serrated. Malaysia (Pahang)........... A. atribatum spec. nov.
- Frons yellow anteromedially. Scutum yellow with brown medial stripe (Fig. 95). Scutellum yellow apically. Male anepisternum at least with faint brownish marking; female unknown. Metanotum yellow with katatergite dark brown and region immediately under scutellum brownish (Fig. 96). Tergites 1 and 2 yellow. Surstylus relatively straight (Fig. 97). Distiphallus straight with base curved (Fig. 99). Paraphallus very small, lobate. Malaysia (Sabah)..................... A. hauseri spec. nov.
5. Notum with strong, dark stripe transverse stripe along anterior margin and with strong medial stripe reaching posterior margin (narrowed on posterior 1/3) (Fig. 69). Posterolateral scutal stripes separate from medial stripe. Surstylus large, especially broad and rounded posterobasally, slightly curved in cross-section and with apex slightly truncated (Figs 71, 72). Malaysia (Sabah)................................................................................... A. gaimarii spec. nov.
- Anterior margin of scutum yellow, but sometimes medial stripe attains anteromedial margin. Medial scutal stripe widening to base, usually confluent with posterolateral stripes when present. Surstylus various, but not as above................... 6
6. Scutellum yellow along entire apical and lateral margins. Lateral scutal stripes sometimes absent, but if present, separate from or faintly confluent with medial stripe (Figs 6–10). Surstylus broad and subtriangular with wide clear distomedial region that makes surstylus appear bifid (Fig. 12). Indonesia (Sulawesi), Malaysia (Perak), Philippines ....... A. antennalis Sasakawa
- Scutellum yellow apically. Lateral scutal stripes broadly fused to longer medial stripe (Fig. 275). Surstylus narrow with apex slightly expanded (Fig. 278). Indonesia (East Kalimantan), Malaysia (Sabah), Philippines ......... A. thrinacos Sasakawa
7. Two very strong lateral scutellar setae that are as long or longer than scutellum; arranged in a transverse row that is straight, or nearly so (Fig. 1). Acrostichal seta large and stout, similar in size to dorsocentral setae. Surstylus small, subtriangular and with slight posterior curve (as in Fig. 3) (in species with known males)................................................ 8
- One or two lateral scutellar setae, shorter than scutellum, not strong or arranged as above. Acrostichal seta small to absent. Surstylus various....................................................................................... 15
8. First flagellomere and scutum entirely yellow.............................................................. 9
- First flagellomere black. Scutum often with with faint to distinct stripe(s)........................................ 10
9. Setae yellowish-brown. No enlarged setula in front of anterior dorsocentral. Fore tarsi yellow or with orange tint (Figs 247, 248). Tergite 2 brown posteromedially. Philippines, Malaysia (Sabah)................................ A. palpale Frey
- Setae brown. Additional enlarged setula in front of anterior dorsocentral, appearing as smaller third dorsocentral. Fore tarsi dark brown (Fig. 246). Tergite 2 yellow (Fig. 245). Malaysia (North Borneo).................. A. sensitivum Sasakawa
10. Three fronto-orbital setae. Medial stripe on scutum very narrowly or distinctly divided medially (ie. bifid) along most of length by yellow line (Figs 168, 169). Wing clouded on distal 1/3. Surstylus with broad, shallow, posteromedial carina along margin (Fig. 171). Malaysia (Sabah, Selangor)................................................... A. magnum spec. nov.
- Four fronto-orbital setae. Medial scutal stripe undivided or absent. Wing clear or lightly infuscated. Surstylus not shallowly produced posteromedially as above..................................................................... 11
11. Scutellum brown (Fig. 229). Notopleuron brown. Wing lightly infuscated, becoming clear posteriorly. Epandrium without spine (Fig. 231). Indonesia (W. Kalimantan), Malaysia (Malaya, Borneo)........................ A. pulcher Sasakawa
- Scutellum yellow. Notopleuron yellow. Wing clear. Epandrium with long, narrow anteroventral spine (males unknown for A. monstrum)......................................................................................... 12
12. Setae dark brown. Frons dark brown medially with yellow spot surrounding tubercle (Fig. 309). Scutum with broad, ill-defined medial stripe attaining anterior margin. Indonesia (West Java)............................... A. monstrum spec. nov.
- Setae yellow. Frons yellow. Scutum entirely yellow or with narrow, faint stripes.................................. 13
13. Scutum with one narrow, faint to indistinct, pointed postsutural stripe and one pair of similarly faint, floating mediolateral stripes that are broken at suture (Figs 1, 2). Male palpus dark brown on distal 1/3. Body length 4.6mm (only one male known). Surstylus narrow and pointed (Fig. 3). Sri Lanka .............................................. A. acum spec. nov.
- Scutum entirely yellow. Male palpus yellow to brownish apically. Body length 3.5–3.6mm (♂), 3.8–4.4mm (♀). Surstylus subrectangular with incurved posterodistal spine (Figs 122, 123) (state unknown for A. flavum)........................ 14
14. Male palpus yellow. Epandrium yellow (Figs 298–300). Seychelles ................................ A. flavum Lamb
- Male palpus sometimes brownish apically. Epandrium pale brown. Laos, Malaysia (Sabah), Philippines. A. hybos Sasakawa
15. Presutural supra-alar seta present. Head msotly dark brown with antenna and face yellow to white; thorax mostly dark brown with yellow stripe from shoulders onto proepisternum and fore coxae (Figs 194–196). Surstylus small and triangular with very long, serrated posterobasal process (Figs 197, 198). Phallus elbowed medially between elongate basiphallus and distiphallus; basiphallus spinulose dorsally and distiphallus with one pair of minute basal processes (Fig. 199). Burma ..... A. nobile Frey
- Presutural supra-alar seta absent. Colour various, but never dark as above. Surstylus and distiphallus never as above..... 16
16. Halter knob brown, at least in part...................................................................... 17
- Halter entirely yellow to white......................................................................... 23
17. Halter knob with brown apical spot. Prosternum brown. Female with tergites 4 and 5 yellow with lateral and sometimes medial stripe brown........................................................................................ 18
- Halter knob entirely or basally brown. Prosternum yellow. Female abdominal tergites 4 and 5 dark brown............. 19
18. Palpus brown apically, yellow basally (Fig. 314). Scutum and scutellum with medial yellow stripe (Figs 311–313). Tergites 2–3 yellow medially. New Caledonia ................................................... A. philomela spec. nov.
- Palpus entirely dark brown (Fig. 316). Scutum and scutellum (except apex) entirely brown (Fig. 315). Tergites 2–3 brown. New Caledonia ....................................................................... A. procne spec. nov.
19. Frons brown with pale anteromedial region. Palpus and clypeus whitish. Scutum brown, sometimes with one pair of narrow yellow lines. Halter knob entirely brown. Surstylus with small rounded basal lobe................................ 20
- Frons yellow or with orange tint. Palpus brown in part. Clypeus brown dorsally. Scutum yellow to brownish with brown vittae. Halter knob brown, sometimes with with apex white. Males unknown.......................................... 21
20. Body length 2.3mm (♂), 2.9–3.1(♀). Scutum entirely brown (Figs 57–59). One lateral scutellar seta. Surstylus narrow, subtriangular (Fig. 61). Paraphallus short, smooth (Fig. 63). Postgonite with thickened setae. Distiphallus slightly more than 1/3 length of phallapodeme. Papua New Guinea ........................................ A. fumihalteratum spec. nov.
- Body length 3.5mm (♂♀). Scutum brown with one pair of narrow, yellow supra-alar stripes (Figs 184, 185). Two lateral scutellar setae. Surstylus subcircular (Fig. 186). Paraphallus long, spinulose (Fig. 187). Postgonite with normal setae. Distiphallus almost ½ length of phallapodeme. Papua New Guinea (New Britain, Manus)................... A. marginale Sasakawa
21. Three dorsocentrals. Halter knob entirely dark brown. Base colour of notum brownish-orange (Figs 329, 330). Medial scutal stripe reaching transverse suture; no sublateral stripes. Madagascar ................................... A. “female 5”
-. Two dorsocentrals. Halter knob brown with apex white. Base colour of notum yellow. Scutum with three complete medial stripes joined apically. Medial scutal stripe reaching anterior margin........................................... 22
22. Three fronto-orbital setae. Two lateral scutellar setae. Palpus brownish (Fig. 308). Lateral brown stripe on scutum separated from brownish lateral margin presuturally by white space (Fig. 307). Scutellum entirely brown. Pleuron with brown spots and femora with basal brownish markings. Papua New Guinea ................................. A. kokodensis spec. nov.
- Four fronto-orbital setae. One lateral scutellar seta. Palpus brownish apically (Fig. 318). Lateral brown stripe on scutum reaching lateral margin, which is slightly yellowish (Fig. 317). Scutellum yellow with brown dorsal spot. Pleuron and femora entirely yellowish. Australia (northern QLD).............................................. A. tetrathrix spec. nov.
23. Two lateral scutellar setae. Anterolateral margin of scutum brown; brown medial stripe on scutum usually short and apically bifid (Fig. 259), but sometimes absent or attaining anterior margin (Fig. 258); scutum uncommonly entirely brown. First flagellomere brown dorsally in both sexes, at least basally. Surstylus subrectangular, angled posteriorly, with apical margin relatively straight; posterodistal corner appearing as subtriangular protrusion near midpoint (Figs 261, 262); distiphallus approximately 3/5 length of phallapodeme (Fig. 263); paraphallus small, smooth, fused to distiphallus. Australia (QLD)............................................................................................. A. striatum McAlpine
- One or two lateral scutellar setae. Scutum various. First flagellomere various. Surstylus various, but never as above; distiphallus and paraphallus never as above...................................................................... 24
24. Scutum entirely brown (with light brown/reddish tint in A. glochis), sometimes excluding venter of postpronotum or one pair of thin, floating yellowish posterolateral stripes. Scutellum entirely brown...................................... 25
- Scutum yellow, usually with brown pattern that sometimes predominates. Scutellum brown or entirely to partially yellow. 33
25. Postsutural scutum with one pair of narrow, floating yellow to yellowish stripes. Pleuron entirely yellow. Epandrium sometimes yellow........................................................................................ 26
- Postsutural scutum entirely brown. Anepisternum usually brown in part (entirely yellow in male A. macalpinei, marking sometimes faint in A. tenebrae). Epandrium brown (males unknown for “female 1” and “female 4”)...................... 28
26. Back of head brown above foramen (Fig. 219). Epandrium brown with distal margin yellow (Fig. 220). Surstylus and epandrium very large, as long as high (Figs 221,222). Postgonite and paraphallus bifid apically (Fig. 223). Papua New Guinea ................................................................................... A. platystylum spec. nov.
- Back of head entirely yellow or with one pair of stripes that may be broadly fused above foramen. Epandrium mostly to entirely yellow. Surstylus and epandrium smaller. Postgonite and paraphallus entire............................... 27
27. One pair of vibrissae. Frons with one pair of distinct brown posterolateral spots that sometimes extend along orbital plate. Back of head with one pair of dark stripes (Fig. 137). Epandrium entirely yellow (Fig. 138). Surstylus appearing folded, narrowest apically (Figs 139–141). Papua New Guinea ........................................ A. lunatum spec. nov.
- Two pairs of vibrissae. Frons yellow, sometimes with one pair of brownish posterolateral spots. Back of head yellow to orange dorsally (Figs 212, 213), sometimes with one pair of brownish spots dorsolaterally. Epandrium with small to elongate brown dorsal spot (Fig. 214). Surstylus narrowest basally, with small outer-basal conical projection (Fig. 215); inner surface with whorl of short, stout setae (Fig. 217). Papua New Guinea ........................... A. phenomena spec. nov. (in part)
28. First flagellomere entirely brown (Figs 327, 328). Frons brown with margin along eye and stripe encompassing base of vertical setae yellow. Coxae, femora and tibiae almost entirely brown. Papua New Guinea ....................... A. “female 4”
- First flagellomere only brown on dorsal half at most. Frons variable, but usually brown along posterolateral margin with brown region enclosing or touching base of vertical setae. Legs yellow, sometimes with femora and base of coxae brown....... 29
29. Frons yellow with brown stripe along posterolateral margin of eye reaching base of posterior fronto-orbital. Back of head yellow centrally and with one pair of marginal dorsolateral stripes (Fig. 321). Anterior fronto-orbital half length of posterior fronto-orbitals. Papua New Guinea ............................................................. A. “female 1”
- Frons yellow to orange, sometimes extensively brown posterolaterally and medially, but not as above. Back of head with one pair of stripes radiating from foramen or entirely brown above foramen. Anterior fronto-orbital 2/3–3/4 length of posterior fronto-orbitals...................................................................................... 30
30. Pleuron with narrow, complete subnotal stripe. Surstylus pointed apically. Ventral lobe of hypandrium strongly produced ventrally. Postgonite small, lobate......................................................................... 31
- Pleuron entirely yellow or with spot on anepisternum; sometimes entirely brown. Surstylus broadly rounded. Ventral region of hypandrium not produced. Postgonite elongate, narrow...................................................... 32
31. Frons orange with lateral margin of orbital plate and posterolateral corner past vertical setae yellow. Male clypeus brownish. Pigment on notum and pleural stripe brown, dark (Figs 64, 65). Surstylus narrow, trifid (Figs 66, 67). Madagascar ............................................................................................ A. fuscinum spec. nov.
- Frons yellow. Male clypeus yellow. Pigment on notum and pleural stripe faint, slightly reddish/brownish (Figs 85, 86). Surstylus with broad posterobasal lobe (Figs 87, 88). Madagascar ................................... A. glochis spec. nov.
32. Postvertical seta as long as ocellar tubercle. Male anepisternum usually entirely yellow (Fig. 149), uncommonly brown dorsally. Surstylus large, flat and broadly rounded (Figs 154,155). Phallapodeme rod-like with small ventral shield. Postgonite long and thin, narrowest near base (Fig. 156). Epiphallus very long. Australia (NSW, southeastern QLD)................................................................................................. A. macalpinei spec. nov.
- Postvertical seta absent. Male anepisternum with posterodorsal infuscation (Fig. 270). Surstylus convoluted, with small posterodorsal bulge and short hooked apex (Figs 271, 272). Phallapodeme dome-like with distal section of shaft atrophied. Postgonite relatively short, narrowest apically (Fig. 273). Epiphallus minute. Papua New Guinea ........ A. tenebrae spec. nov.
33. Scutellum entirely yellow, without any vestige of brown pigment. Not postsuturally bivittate (A. falcistylum with presutural stripes)............................................................................................ 34
- Scutellum brown, at least in part; those species with a very faint lateral brown spot are bivittate postsuturally........... 42
34. Presutural scutum with lateral margin and one pair of dorsocentral stripes brown. Surstylus narrowed and strongly incurved on medial third, and with apex expanded and flat; inner surface with small tubercle-like setae on posterior margin of basal third and apex (Fig. 295). Japan .......................................................... A. falcistylum Sasakawa
- Scutum without paired stripes. Surstylus various but not as above............................................. 35
35. Anteromedial margin of scutum brown................................................................... 36
- Anteromedial margin of scutum yellow.................................................................. 38
36. One lateral scutellar seta. Anepisternum with very small brown spot (Fig. 235). Surstylus very large and subtriangular with posterobasal process (Figs 236, 237). Malaysia (Sarawak)............................... A. punctulatum Sasakawa
- Two lateral scutellar setae. Anepisternum entirely yellow or with large brown spot. Surstylus narrow with pointed apex angled anteriorly.......................................................................................... 37
37. Brown anteromedial margin on scutum wide, extending around margin of postpronotum (Figs 74, 75). Anepisternum with large brown spot. Wing clear. Surstylus with relatively long, narrow, strongly angled apex; without small posterobasal point (Figs 76, 77). Pregonite small, lobate (Fig. 78). Thailand ............................... A. geniculatum (Sasakawa)
- Brown anteromedial margin on scutum narrow, barely reaching postpronotum (Figs 143, 144). Anepisternum entirely yellow. Wing with anterodistal infuscation. Surstylus with short, slightly angled apex; with shallow posterobasal point (Figs 145, 146). Pregonite longer, membranous and partially fused to hypandrium (Fig. 147). Philippines ................ A. lux spec. nov.
38. Female postsutural scutum with large posteromedial spot; male scutum entirely yellow (Figs 253, 254). Surstylus much larger than epandrium and with pronounced posterobasal lobe (Figs 255, 256). Region between hypandrium and pregonite lightly sclerotized, setose and ridged (Fig. 257). Thailand ............................................ A. solare spec. nov.
- Not as above, although male scutum entirely pale or with almost indiscernable posteromedial line.................... 39
39. Notum with slight orange tint, and with narrow, faint, almost indiscernable stripe along margin of notopleuron, postpronotum and anterior margin of scutum, and with short, narrow and very faint posteromedial stripe on scutum (Figs 162, 163). Pleuron with very narrow, faint subdorsal stripe. Surstylus large and subtriangular with anterior margin straight and posterior margin curved (Figs 164, 165). Distiphallus as long as phallapodeme (Fig. 166). Philippines .......... A. macrophallus Sasakawa
- Notum entirely yellow. Pleuron yellow with dark brown spot on anepisternum. Surstylus not as above. Distiphallus much shorter than phallapodeme............................................................................ 40
40. Surstylus narrow and pointed with similar basal process directed distally (Figs 49, 50). Thailand .... A. dicrostylis Sasakawa
- Surstylus thick and hook-shaped with apex relatively broad.................................................. 41
41. Back of head entirely yellow (Fig. 101). Male first flagellomere with vestigial dorsal spot (Fig. 102). Pregonite relatively broad and ovate (Fig. 106). Indonesia (North Kalimantan), Malaysia (Johor, Pahang)......... A. hesperotoxon spec. nov. (males)
- Back of head with faint brownish marking dorsolaterally and below ocelli. First flagellomere with pale, but distinct dorsal stripe (Figs 157, 158). Pregonite narrow, subtriangular (Fig. 161). Ivory Coast ............... A. machaeroges spec. nov.
42. Scutellum brown with yellow arch (Fig. 239). Male scutum with medial stripe enclosing large yellow subrectangular spot between dorsocentral rows; female scutum brown excluding yellow subrectangular spot. Surstylus broad, flat along distal and anterior edges, curved inwards and with shallow, high posterodorsal lobe (Figs 241–243). Papua New Guinea ................................................................................................. A. selenis spec. nov.
- Scutellum, scutum and surstylus various, but never as above................................................. 43
43. Scutum with brown posteromedial stripe that sometimes extends presuturally.................................... 44
- Scutum either yellow with lateral and sometimes anterior margins brown, or bivittate, with vittae sometimes very narrowly separated or united basally along posterior margin.......................................................... 58
44. Anepisternum with brown spot. Surstylus hooked.......................................................... 45
- Anepisternum entirely yellow to white. Surstylus various.................................................... 48
45. Two fronto-orbital setae. Postsutural scutum with broad subquadrate spot. Scutellum only brown dorsally. Taiwan ....................................................................................... A. atromaculatum (Hennig)
- Three fronto-orbital setae. Postsutural scutum with narrow medial stripe. Scutellum brown......................... 46
46. Frons with medial dark yellow to brownish stripe. Palpus brown apically. Back of head with brown spot below tubercle. Medial scutal stripe pointed, not fused basally to one pair of shorter posterolateral stripes; lateral margin of scutum brown to postpronotum (Figs 291, 292). Tergite 2 brown................................ A. hesperotoxon spec. nov. (females)
- Frons yellow medially. Palpus yellow. Back of head vittate. Medial scutal stripe rounded, fused to one pair of small posterolateral stripes basally; lateral margin of scutum yellow. Tergite 2 yellow or light brown.............................. 47
47. Posterolateral corner of frons entirely yellow. Posteromedial stripe basally fused to narrow, fainter posterolateral stripes (Figs 90, 91). Metanotum brown and yellow. Tergite 1 yellow; 2 yellow with posteromedial and posterolateral spots. Laos, Thailand ................................................................................ A. grypostylis Sasakawa
- Posterolateral corner of frons with brown stripe reaching base of outer vertical seta. Posteromedial stripe very broadly fused to dark posterolateral stripes (Figs 319, 320). Metanotum entirely brown. Tergites 1 and 2 light brown. Laos, Thailand ...................................................................................... A. trimaculosum Sasakawa
48. Medial scutal stripe narrow, bounded laterally by dorsocentral rows. Orbital plate yellow........................... 49
- Medial scutal stripe wide, extending beyond dorsocentral rows. Orbital plate sometimes brown in part................ 53
49. Notopleuron yellow.................................................................................. 50
- Notopleuron brown.................................................................................. 51
50. Scutal pattern sometimes reduced to mottled posteromedial stripe; lateromedial stripes absent (Figs 29, 30). Scutellum entirely brown. Ocellar spot limited to tubercle. Surstylus subrectangular with large posteromedial process (Figs 31, 32). Phallapodeme broad, dome-like (Figs 33, 34). Distiphallus shorter than paraphallus. Papua New Guinea ....... A. cavernosium spec. nov.
- Scutal stripe well-developed, sometimes reaching anterior margin; scutum with one pair of narrow lateromedial stripes (Figs 52, 53). Scutellum yellow with dorsal surface brown. Ocellar spot triangular, slightly larger than tubercle. Surstylus hookshaped (Figs 54, 55). Phallapodeme rod-like with flat ventral shield (Fig. 56). Distiphallus longer than paraphallus. Papua New Guinea ............................................................................. A. eotoxon spec. nov.
51. Anteromedial margin of scutum yellow; with one pair of narrow floating posterolateral stripes that are sometimes connected to brown lateral margin by paler brown region (Figs 323, 324). One lateral scutellar seta. Papua New Guinea .... A. “female 2”
- Anteromedial margin of scutum brown; floating posteromedial stripes absent. Two lateral scutellar setae.............. 52
52. First flagellomere entirely white. Medial scutal stripe well-defined, dark, sometimes attaining anterior margin (Figs 35, 36). Metanotum yellow with mediotergite and posterodorsal corner of anatergite brownish. Tergite 1 and epandrium yellow. Sursty-
lus broad, rounded, with long posterobasal process, and outer surface with row of long, thick setae (Figs 37, 38). Papua New Guinea .............................................................................. A. cheiris spec. nov. - First flagellomere with brown dorsal spot. Medial scutal stripe short, pale, mottled (Figs 125, 126). Metanotum brown with mediotergite yellow. Tergite 1 and epandrium brown. Surstylus narrower, slightly pointed apically, and outer surface with small setae on apical and posterior regions (Figs 127, 128). Australia (QLD)..................... A. infernum spec. nov.
53. Two to four pairs of vibrissae; sometimes only one (possibly A. tropalis paratype). Orbital plate yellow, sometimes with faint brownish posteromedial spot. Usually two lateral scutellar setae. Surstylus with circular whorl of setae on inner surface; shape subrectangular with base narrowed...................................................................... 54
- One pair of vibrissae. Orbital plate with long brownish to brown spot or stripe. One lateral scutellar seta. Surstylus without circular whorl of setae on inner surface; shape various but not as above........................................... 55
54. Two pairs of vibrissae with posterior pair joined apically. Medial scutal stripe with complete transverse yellow line at suture (Fig. 212). Postgonite apex short, subtriangular (Fig. 218). Papua New Guinea .......... A. phenomena spec. nov. (in part)
- Four pairs of vibrissae, arranged along a horizontal plane (Figs 284, 285); sometimes one pair (possibly paratype). Medial scutal stripe emargined laterally at suture (Figs 286, 287). Postgonite apex narrow, elongate, lobate (Fig. 290). Papua New Guinea (Bismarck Islands)............................................................. A. tropalis Sasakawa
55. Orbital plate with narrow, well-defined stripe from base of vertical setae to base of posterior fronto-orbital. Posteromedial margin of scutal stripe and scutellum both with yellow notch (Fig. 40). Postvertical seta absent. Surstylus large and subtriangular with apex slightly curved and inner-posterior margin with clusters of tubercle-like setae on distal half and on posterior corner (Figs 44, 45). Australia ............................................................. A. conopeum spec. nov.
- Orbital plate with longer stripe or frons with diffuse posterolateral spot. Scutal stripe entirely brown along posterior margin; scutellum brown, sometimes with yellowish spot at base of setae. Postvertical seta small and thin. Surstylus not as above. 56
56. First flagellomere with brownish basal marking. Orbital plate brown, darkest medially, with anterior edge yellow. Scutellum entirely brown (Figs 107, 108). Epandrium brown with distal margin yellow. Surstylus with long setae in a band across the outer surface and a dense brush along a posteromedial shelf (Figs 109–111). Papua New Guinea ..... A. hirsutum spec. nov.
- First flagellomere entirely yellowish-white. Frons with diffuse brownish spot from orbital plate to ocellar tubercle. Scutellum brown with yellowish spot at base of apical setae. Epandrium entirely yellow. Surstylus not as above................. 57
57. Back of head with one pair of strong, confluent stripes. Scutal stripe shallowly emargined at suture; slightly darker along dorsocentral row (Figs 113, 114). Metanotum yellow with dorsum brownish. Surstylus higher than long, curved posteriorly, with dense brush of tubercle-like setae on raised process (Figs 115–117). Phallapodeme flat and dome-like (Fis 118, 119). Postgonite bifid. Papua New Guinea ....................................................... A. horridum spec. nov.
- Back of head with one pair of small, brownish medial spots. Scutal stripe not emargined at suture, not darker along dorsocentral row (Figs 279, 280). Metanotum yellow with mediotergite brown. Surstylus small, trilobed, without raised processes (Figs 281, 282). Phallapodeme rod-like (Fig. 283). Postgonite entire. Papua New Guinea ............. A. trilobellum spec. nov.
58. Scutum brown with central yellow spot that may reach posteromedial margin. Enlarged setula sometimes present in front of anterior dorsocentral. Male first flagellomere brown dorsally................................................. 59
- Scutum variably vittate, never as above. No enlarged setula in front of anterior dorsocentral. Male first flagellomere usually entirely yellow to white, but sometimes as above........................................................... 60
59. Posterolateral corner of frons dark brown with stripe extending to base of posterior fronto-orbital (Fig. 302). Sucutm with narrow medial yellow stripe, not reaching dorsocentral rows. Male unknown. Papua New Guinea ....... A. fumipenne Kertész
- Posterolateral corner of frons entirely yellow (Fig. 205), female sometimes with brownish spot around base of vertical setae. Scutum with central yellow region very large, extending outside dorsocentral rows laterally (Fig. 208). Surstylus nearly as high as epandrium, stalk-like along most of length (seen laterally), and with distal margin flat and angled dorsally and inwards (Figs 209, 210). Cercus long, pointed. Paraphallus broad and truncated apically (Fig. 211). Postgonite and epiphallus long, projecting. Australia (NSW, QLD)............................................................ A. perkinsi McAlpine
60. Two lateral scutellar setae. Scutum yellow with one pair of very large, dark stripes (Figs 293, 294) that are sometimes confluent on basal half. Papua New Guinea ................................................... A. bivittatum spec. nov.
- One lateral scutellar seta. Scutum not as above............................................................ 61
61. Notopleuron entirely yellow. Postsutural stripes not exceeding transverse suture.................................. 62
- Notopleuron laterally or entirely brown. Postsutural stripe usually exceeding suture, narrowly separated from presutural portion of stripe (more widely spaced in A. conopeum and some male A. malayensis), or at least lateral 1/3 of scutum brown. 66
62. First flagellomere usually brown dorsobasally. Back of head, anterior margin of scutum and mediotergite yellow. Scutal stripes very narrow. Male (only known for A. juxtum; Figs 133–136): epandrium entirely yellow; surstylus longer than high with circular whorl of setae on inner surface and dense comb of setae on posterior process; cercus narrow, finger-like; phallapodeme dome-like subapically; postgonite large, with anterodistal lobe and anteromedial cluster of setae..................... 63
- First flagellomere entirely yellow to white. Back of head with one pair of faint to distinct stripes. Anterior margin of scutum brown. Mediotergite brown. Epandrium brown, sometimes with distal margin yellow. Surstylus higher than wide, without posterior comb of setae. Phallapodeme rod-like with flat ventral shield. Postgonite small, lobate........................ 64
63. Scutum with one pair of very narrow supra-alar stripes (Fig. 305). Scutellum yellow with small brown spot in lateral corner. Anepisternum with brown spot (Fig. 306). Malaysia (Pahang).................................. A. gracile spec. nov.
- Scutum with one pair of pointed stripes inside dorsocentral rows (Figs 131). Scutum yellow with dorsum brown. Pleuron entirely yellow (Fig. 132). Papua New Guinea .............................................. A. juxtum spec. nov.
64. Back of head with one pair of faded brown spots. Scutal stripes narrow, faded to base and widely separated, sometimes also faded anteriorly (Fig. 264). Scutellum yellow with small brown spot in lateral corner. Anepisternum light yellow. Surstylus nearly as high as epandrium, curved, slightly sinuate (Figs 266, 267). Indonesia (E Kalimantan, Sumatra).................
................................................................................. A. suspirium spec. nov. - Back of head with one pair of strong, dark stripes. Scutal stripes large, wider and dark towards base. Scutellum brown. Anepisternum with brown spot. Surstylus narrow, straight, triangular................................................ 65
65. Scutellum brown with margins lightly mottled. Scutal stripes small, dark, well-defined, joined medially by irregular brown patch (Figs 80, 81). Surstylus long, needle-like and curved anteriorly (Figs 82, 83). Vietnam ........... A. giallo spec. nov.
- Scutellum evenly brown. Scutal stripes wide, ill-defined and narrowly separated (Figs 200, 201). Surstylus half height of epandrium, straight (Figs 202, 203). Sri Lanka .................................................... A. nyx spec. nov.
66. Male with brown spot on anepisternum; scutum brown with yellow posteromedial stripe (Figs 14, 15). Female scutum brown with complete central yellow stripe; anepisternum with brown spot; palpus yellow. Surstylus large, broadly rounded apically, narrowest basally (Figs 16, 17). Hypandrium with large, apically truncated ventral lobe (Fig. 18). Distiphallus simple, flat. Sri Lanka .............................................................................. A. asylum spec. nov.
- Male with entirely yellow to white pleuron. Scutum of both sexes yellow with brown vittae; if scutum brown with central yellow stripe (some A. malayensis females), then distal 1/3 of palpus, anepisternum and anepimeron brown. Surstylus and hypandrium not as above. Distiphallus complex, folded (males unknown for “female 3”)................................ 67
67. Ocellar spot larger than tubercle, subtriangular and sometimes extending nearly to anterior margin of frons (Fig. 178). At least basal third of arista white (less in holotype). Female with distal third of palpus, anepisternum and anepimeron brown. Surstylus shorter than epandrium, narrow and pointed apically (Figs 179, 180). Distiphallus dark, clavate (Fig. 181). Indonesia (E Kalimantan, Sumatra), Malaysia (Pahang)................................................. A. malayensis Sasakawa
- Ocellar spot restricted to tubercle, rarely slightly larger. Only base of arista white. Female (only known for “female 3”) palpus and pleuron entirely yellow to white. Male: surstylus as high or higher than epandrium, subtriangular with broad apex, and posterobasal corner slightly exceeding epandrium. Distiphallus made of thinner folds.............................. 68
68. Scutal stripes entire (Figs 325, 326). Anteromedial margin of scutum margined with narrow brown line. Indonesia (Sulawesi), Papua New Guinea ......................................................................... A. “female 3”
- Scutal stripes broken at suture. Anteromedial margin of scutum sometimes with thick brown line.................... 69
69. Postvertical seta absent. Scutal stripes subrectangular, narrowly separated and sometimes partially fused (Figs 41, 42). Scutellum brown with apex yellow, sometimes only around base of apical setae. Surstylus narrow apically, slightly curved anteriorly (Figs 44, 45). Paraphallus sinuate with dorsal subbasal spinules (Fig. 46). Papua New Guinea ..... A. conopeum spec. nov.
- Postvertical seta as long as ocellar tubercle. Scutal stripes narrowing anteriorly, triangular, widely separated. Scutellum brown with complete central yellow stripe, sometimes entirely brown if scutal stripes fused basally. Surstylus thicker, straight apically. Paraphallus arched, dorsally spinulose past base....................................................... 70
70. First flagellomere entirely yellow (Fig. 189). Stripes behind head fused above foramen (Fig. 188). Tergite 1 yellow or light brown. Surstylus slightly shorter than epandrium, apex broadly rounded (Figs 191, 192). Distiphallus folded, but with separate ovate sclerites in membranous lateral membrane (Fig. 193). Philippines ...................... A. megastylis Sasakawa
- First flagellomere with light brown dorsobasal spot. Stripes behind head separate. Presutural scutum sometimes with one pair of narrow stripes divided from posterior stripes at suture (Figs 224, 225). Tergite 1 brown with lateral margin yellow. Surstylus higher than epandrium, slightly pointed at apex (Figs 226, 227). Distiphallus a single folded sclerite (Fig. 228). Philippines .................................................................................... A. plicatum spec. nov.
description: Published as part of Owen Lonsdale, 2016, Revision of the genus Allometopon Kertész (Diptera: Clusiidae), pp. 1-127 in Zootaxa 4106 (1) on pages 7-12, DOI: 10.11646/zootaxa.4106.1.1, http://zenodo.org/record/258766