Euryomma guane Grisales & Wolff & de Carvalho 2012, sp. nov.
Creators: name: Grisales, Diana
name: Wolff, Marta
name: de Carvalho, Claudio J. B.
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Plantae
Rhodophyta
Florideophyceae
Gigartinales
Solieriaceae
Euryomma
Euryomma guane

description: Euryomma guane sp. nov.   
(Figures 2D, 3D, 4D, 5D)  
 Diagnosis   
Frontal vitta dark brown with golden yellow pollinosity, particularly on anterior region; length of postpedicel 3.5 times the length of pedicel; pubescence of arista almost as long as width of base of arista; scutum with five brown vittae, vitta on intraalar region expanded over supra-alar area, median vitta expanded from base to apex of scutellum; wing light brown; calypter whitish with yellow margins; haltere yellowish with base darkened; sternite 5 wide, with two elongate, well sclerotized, rod-shaped apical processes bearing strongly developed setae on inner lateral.  
 Description   
 Male holotype. Body length 3.6 mm; wing length 2.8 mm.   
 Head. Eye with sparse setulae. Frontal vitta dark brown, golden yellow pollinose, particularly on anterior half. Interocular space 0.4 mm. Four pairs of fr, one of which is more developed, with intercalated setulae between them. Orb two pairs, anterior pair lateroclinate, posterior pair reclinate. Ocellar tubercle grey with yellow pollinosity. One pair of oc present. Poc parallel. Fronto-orbital plate, parafacial, gena and lunula golden yellow pollinose. Parafacial with short setae on dorsal half. Scape brown, golden yellow pollinose and with one seta reaching the pedicel. Pedicel brown, yellow around suture. Postpedicel dark brown, golden yellow pollinose, 3.5 times the length of pedicel. Arista brown, bearing long pubescence, rays almost as long as width of base of arista. Palpus dark brown, flat and claviform.   
 Thorax. Greyish with golden yellow pollinosity, scutum grey with faint golden yellow pollinosity, five brown vittae, along acr, dc and intra-alar setae, one vitta expanding over intra-alar area up to supra-alar area, median vitta expanded from base to apex of scutellum. Acr 3:3–4, not organized in rows. Dc 2:3, first presutural seta with half the length of second presutural seta. Postpronotum with two well-developed setae and a weaker one. Pra two, both well developed. Proepimeral setae two, seta closest to spiracle longer, the other with hooked apex, proepimeral area without setulae. Katepisternal setae1:1, with one weakly developed seta between them, katepisternum with short setulae.   
 Wing. Faintly brownish, with apical third, between C vein and apical half of vein R 2+3 faintly darkened. Calypter whitish with margin yellowish. Haltere yellowish with base darkened.   
 Legs. Coxae and femora dark brown, a surface of trochanters, apex of femora and tibiae yellow. Hind tibia with a brown vitta on basal third; tarsi dark brown with yellowish pulvilli. Fore femur with a row of long setae on pd surface, setae becoming longer from base to pre-apex; p surface on basal third with setae shorter than pd setae; pv surface with a row of long setae on apical half; v surface bare. Fore tibia with a pre-apical seta on d surface; ad surface with an apical short seta and a pre-apical seta; av surface without apical seta; pv surface with an apical seta as long as v seta; v surface with an apical seta; pd surface with a developed pre-apical seta. Fore first tarsomere with a long and strong basal seta on v surface. Mid femur on av surface with a row of sparse setae, setae closer together and shorter on apical portion, apex hooked; ad surface with a row of differentiated setae on apical third, with three preapical setae; pv surface with a differentiated basal seta, from basal third to apex with three or four irregular rows of setae with hooked apices forming a strong ctenidium, setae shorter on ventral rows; p and pd surfaces with three differentiated pre-apical setae. Mid tibia on v surface with two developed apical setae, one short and one long; a surface with a short basal seta and longer sub-median seta; ad surface with a developed pre-apical seta; av surface with a developed apical seta; v surface setulose; pd surface with a short pre-apical seta; p surface with a median seta and a short pre-apical seta; pv surface with a short apical seta; d surface without differentiated setae. Hind coxa setulose on posterior margin. Hind femur on median third of ad surface with two developed setae and a less differentiated seta on apical third; av surface with a row of short setae, with two differentiated and well developed setae on apical third; v surface bare; d surface with one differentiated seta on apical third; pd surface on apical third with a differentiated seta shorter than d seta. Hind tibia on d surface with a welldeveloped median and a shorter pre-apical seta; a surface with a short pre-apical seta; ad surface with a median seta at the same level as median d seta and a shorter preapical seta; av surface with a median seta and an apical seta more developed than median seta; pv surface with a weakly developed apical seta.   
Abdomen. Grey, tergites with brown vitta on more than half of tergites 3 and 4, on tergite 5 it is extending from the base to the apex; one pair of lateral marginal setae on tergites 3 and 4. Sternite 1 bare. Sternite 5 (Figure 2D): wide, with two long apical narrow extensions, and with long, well-developed setae, particularly on inner lateral. Terminalia (Figures 3D, 4D): epandrium longer than wide and with short setae, particularly on basal half; surstylus triangular, short and articulated with epandrium, internal surface and apex with short and sparse setae; cercal plate fused on basal third, rounded apically. Hypandrium and associated structures as in Figure 5D.  
 Female. Unknown.   
 Biological notes   
 The biology and habits of species of forest-dwelling Fanniidae are not well-known. Euryomma guane sp. nov. occurs in high Andean Forests from 2290 m to 2390 m. According to collecting data, it is associated with decomposing organic matter such as fish.   
 Comments   
 Euryomma guane sp. nov. and Euryomma uwa sp. nov. were collected in the same locality and substrate. The external morphology of the two species does not differ and does not help in their separation. However, the male terminalia in these two species is different. Euryomma guane has epandrium with short setae, particularly on basal half, surstylus short and triangular with short and sparse setae on inner surface and apex, and cercal plate rounded apically (Figures 3D, 4D). Meanwhile E. uwa has epandrium with long and sparse setae, surstylus narrow apically, with sparse and short setae on apical half and cercal plate elongated and setulose (Figures 3G, 4G). In the holotype the fore right leg and most of setae on the head and thorax are missing.   
 Etymology   
 The specific epithet, guane, refers to the indigenous people known as “Guanes”, who inhabited the department of Santander, type locality of the species, in the Colombian Andes. According to researchers, the Guanes are descendent from the Muiscas, one of the best-developed indigenous cultures in South America until the pre-Hispanic era. Noun in apposition.   
 Type material   
 Male holotype (CEUA). Colombia. Santander. Piedecuesta \ Vereda Cristales. Finca El Rasgón \ Bosque. 2290–2390 m. Trampa VSR [Van Someren-Rydon trap]\ pescado. Agosto 8 / 2001 \ Duque & Vélez. CEUA.   
 Distribution   
Colombia: department of Santander (Sistema Andino, Cordillera Oriental).
description: Published as part of Grisales, Diana, Wolff, Marta & de Carvalho, Claudio J. B., 2012, Neotropical Fanniidae (Insecta: Diptera): new species of Euryomma Stein from Colombia, pp. 803-829 in Journal of Natural History 46 (13 - 14) on pages 819-821, DOI: 10.1080/00222933.2011.651644, http://zenodo.org/record/5199672