Phraepsyche pectinata Oláh & Johanson 2010, new species
Creators: name: Oláh, János
name: Johanson, Kjell Arne
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Odontoceridae
Phraepsyche
Phraepsyche pectinata

description: Phraepsyche pectinata, new species   
Figs 256–263  
 This new species is distinguished from the other 2 species in the genus in the genitalia, by the intersegmental depression between segment IX and segment X having 1 deep step in lateral view, not with 2 steps as in P. danaos or without steps as in P. yitungshana. The phallic apparatus has a subapical bulbous broadening that is absent in P. yitungshana, and the apicoventral setose lobes on segment X are not widening before apex in dorsal view, as they do in P. danaos. The pronotum has 2 separated pairs of setal warts, while in the other species the lateral pair has fused with the mesal pair. There are also significant differences in the groove and setal wart patterns on the head.   
 Male (in alcohol). Body medium sized; light brown; forewings light brown. Head rectangular, almost 2 times broader than long. Ocelli absent. Tentorium without dorsal arm; posterior arms short, robust in dorsal view, ending in pair of large posterior tentorial pits, with strong tentorial bridge armed with small, 2-rayed anteromesal protuberance; anterior tentorial arms posteriorly slender, anteriorly broad, with additional median lamellate process. Facial groove pattern complex. Frontogenal vertical grooves long. Clypeogenal vertical grooves located ventrally of anterior tentorial pits; short, running obliquely laterad; subantennal grooves running oblique mesad, confluent to vertical frontogenal grooves; additional lateral grooves present, between mid-way of ocular groove and clypeogenal groove. Vertexal groove system with thick rim of compact warts; stem of epicranial groove intermittent in middle, anteriorly long, posteriorly short; most pronounced pair of grooves on vertex are lateral, sigmoid vertexal grooves in middle section; occipito-postgenal grooves partly merging with skeletal ring of occipital setal warts, running along ocular grooves. Labrum semi-circular, weakly pigmented anterior part narrow, movable, freely hanging. Broad posterior, basal sclerite of labrum bearing transversely elongate, compact setal wart. Two pairs frontal, lateral, compact setal warts present on frons, anterior warts small, vertically elongate posterior pair large; frontal interantennal warts absent, or represented by anterior warts. Pair of rounded frontogenal, posterior, compact setal warts present on dorsum of pregenae, tangential with subantennal grooves. Vertexal, lateroantennal compact wart small, rounded, located mesally of lateral vertexal groove. Pair of vertexal, medioantennal, diffuse setose warts divided by coronal groove. Large pair of occipital compact warts spreading over posterior half of head dorsum, almost integrating smaller vertexal ocellar setal warts. Vertically elongate, postgenal compact warts curving along posterior section of ocular grooves. Maxillary palps digitiform; maxillary palp formula (I,II)-III-V-IV; each first segment more robust and broader than other segments, without subapicomesal nodule, each 4th segment significantly longer than each 5th segment. Each scape almost as long as head. Each pedicel slightly longer than first segment of each flagellum; basilateral part of first 13 segments of flagellum bearing long branch-like processes, giving pectinated antennae. Pronotum with 2 pairs setal warts. Mesal pronotal compact warts narrow, elongate, transverse, occupying almost entire dorsum; lateral warts small, transverse, oval. Mesoscutal pair of compact warts small, longitudinally elliptical. Pair of mesoscutellar compact warts elongated. Each proepisternum with large, ovoid setose wart. Precoxal warts small. Small, elevated protuberance-like, compact setal wart present on each cervical sclerites. Claws symmetrical. Spur formula 1, 4, 4; midlegs with anterior spur half as long as posterior spurs; hind leg spurs almost equally long. Forewings: length 6.0 mm, broad; membrane light brown, without visible pattern; Sc and R1 meeting in callosity as both veins are thick, fusing, forming broad diffusion area at pterostigma; apical forks III and IV absent; fork I and fork II tangential at short, pentagonal discoidal cell; crossvein m short; M reduced; fork V wide; Cu2 vestigial in holotype, absent in paratype; postanal vein present. Hind wings: Sc and R1 confluent apically of middle of wing; RS reduced, with stem running into M. Abdominal sternum VI with projection (hammer) on posterior margin; forming elongate, robust rod reaching length of segment margin. Sternal hammer reinforced by ventrad curving internal ridge of longitudinal suture.   
Male genitalia. Abdominal segment IX fused annularly, short; tergum slightly shorter than sternum; anterior margins of segment IX triangular with sharply angled process located sub-centrally; posterior margin concave. Antecostae well developed only on sharply angled process; external groove of antecostal sutures well developed; tergum produced into central, triangular lobe in dorsal view; spine rows on posterior margins of segment IX reduced, with 2 to 3 setae near gonocoxites. Intersegmental depression between segment IX and segment X well developed, stepwise. Segment X long in lateral and dorsal views, nearly parallel-sided; slightly triangular in lateral view; in dorsal view slightly broadening from midway. Apicoventral setose lobes with 2 to 3 stout apical spines. Apicodorsal setose lobes located subapically, fused into medial hump. Dorsal interlobular gap between apicoventral setose lobes deep, narrow. Superior appendages 2 times broader in dorsal view than in lateral view, short, not reaching medial hump of apicodorsal setose lobes. Each coxopodite longer than apex of segment X, forming long, stout, leaf-like process with broad basal half; slightly ventrad curving in lateral view; curving mesad in ventral view. Harpagones slender, claw-like, arising before apex of each coxopodite. Phallic apparatus as long as coxopodites, basal section well-sclerotized, short, ventrad curving, with short flange articulating with base of gonocoxites. Tubular phallotheca straight, continuous with membranous phallicata; less pigmented continuation of phallotheca obscure. Endotheca indistinct, forming straight tube, bulbous subapically, with ventrad curving apex. Gonopore indistinct. Ejaculatory duct ending at circular, well-pigmented anterior part of phallotremal sclerite; dorsum of apex excavated. Posterior, narrow elongation of circular phallotremal sclerites exposed in apicodorsal cavity.  
 Holotype male: VIETNAM: Lamdong: Baoloc, Duchma Stream, 23.x.1988, light [J. Oláh] ─ (OPC).   
 Paratypes: VIETNAM: Lamdong: Dalat, Atangla Waterfall, 15.x.1988, light [J. Oláh] ─ 3 males (OPC).   
 Distribution: Vietnam.   
 Etymology: Pectinata, name referring to the pectinated antennae.
description: Published as part of Oláh, János & Johanson, Kjell Arne, 2010, Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457, pp. 1-128 in Zootaxa 2457 (1) on pages 114-117, DOI: 10.11646/zootaxa.2457.1.1, http://zenodo.org/record/5313709