Canthon (Goniocanthon) smaragdulus subsp. subviridis Schmidt 1922, stat. rev.
Creators: name: Nunes, Luis Gabriel de O. A.
name: Nunes, Rafael V.
name: Vaz-De, Fernando Z.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Canthon
Canthon smaragdulus
Canthon (goniocanthon) smaragdulus subviridis schmidt, 1922

description: Canthon (Goniocanthon) smaragdulus subviridis Schmidt, 1922 stat. rev.
Figs 2 B–D, 5, 8
Canthon speculifer var. subviridis Schmidt, 1922: 69.
Canthon speculifer var. subviridis – Blackwelder 1944: 201 (catalogue).
Canthon (Goniocanthon) smaragdulum v. subviride – Vaz-de-Mello 2000: 191 (species list).
Diagnosis
In the subgenus, C. (G.) smaragdulus subviridis stat. rev. is separated by the following combination of characters: ventral surface of meso- and metafemora smooth (Fig. 1A); dorsal surface of pronotum and elytral disc with similar metallic green colouration (Fig. 2B); metasternum anterior lobe strongly convex (Fig. 1B); profemur with a tooth on its anterior border. Paramera: on lateral view (Fig. 1G): flattened; apex truncated; upper angle rounded; lower angle acute, forming a spine or tooth. Internal sac (Fig. 5): FLP involving A forming a central duct with spiniform apex. SA lateral to FLP + A complex present. MP distal to A, not divided, semicircular, widely emarginated; FLP + A complex involved by MP.
Material examined
Lectotype (for Canthon speculifer var. subviridis Schmidt 1922; here designated)
BRAZIL: ♁, Espir. Santo (white label) / 3271 E92+ (blue) / LECTOTYPE, Canthon speculifer var subviridis A. Schmidt, des F.Z. Vaz-de-Mello, 2014 (red with black margins) (NMRS 3271 E92+).
Paralectotypes
BRAZIL: 1 ♁, Espir. Santo (white label) / 3271 E92+ (blue) / PARALECTOTYPE, Canthon speculifer var subviridis A. Schmidt, des F.Z. Vaz-de-Mello, 2014 (yellow with black margins) (NMRS 3271 E92+); 1 ♁, same label data as for preceding, but 3268 E92+ (NMRS 3268 E92+); 1 ♁, same label data as for preceding, but 3269 E92+ (NMRS 3269 E92+); 1 ♁, same label data as for preceding, but 3270 E92+ (NMRS 3270 E92+); 1 ♁, same label data as for preceding, but 3285 E92+ (NMRS 3285 E92+); 1 ♀, same label data as for preceding, but 3277 E92+ (NMRS 3277 E92+); 1 ♀, same label data as for preceding, but 3287 E92+ (NMRS 3287 E92+); 1 ♀, same label data as for preceding, but 3286 E92+ (NMRS 3286 E92+); 1 ♀, same label data as for preceding, but 3288 E92+ (NMRS 3288 E92+).
Non-type material (68 specimens)
BRAZIL – Ceará: 1 ♀, C Pioson, coll. Hangay, Walter Heinz ex coll. and leg. (CEMT). – Alagoas: 1 ♁, Ibateguara, 12 Oct. 2007, B.K.C. Filgueiras leg. (CEMT); 1 ♁, Ibateguara, Serra Grande, 8°59′44″ S, 35º53′15″ W, Mata Atlântica borda, pitfall, 1 Nov. 2011, B.K.C. Filgueiras leg. (CEMT). – Bahia: 3 ♁♁, 4 ♀♀, Itabuna, CEPLAC, Cabruca, 10 Jan. 2003, M. Santos leg. (CEMT). – Minas Gerais: 2 ♁♁, Mariléia, Parque Estadual do Rio Doce, 19º45′19″ S, 42°37′43″ W, 13 Oct. 1992, J.N.C. Louzada leg. (CEMT); 1 ♁, Mariléia, Pq. Est. Rio Doce, Feb. 2000, L. Scoss leg. (CEMT); 1 ♁, same data as for preceding, but Nov. 1992, J.N.C. Louzada leg. (CEMT); 1 ♀, Ipatinga, Aug. 1994, G. Turrer leg. (CEMT). – Espírito Santo: 1 ♁, Soorotema, Reserva Natural Vale, Estrada Cinco Folhas, 19º08′34″ S, 40º03′56″ W, 59 m a.s.l., human faeces, pitfall, 2 Feb. 2015, T. Vargas and L. Lopes leg. (CEMT); 1 ♁, 1 ♀, Soorotema, Reserva Natural Vale, Estrada Cinco Folhas, 19º08′56″ S, 40º04′08″ W, 71 m a.s.l., pitfall, 3 Feb. 2015, T. Vargas and L. Lopes leg. (CEMT); 2 ♀♀, Soorotema, Reserva Natural Vale, Estrada Cinco Folhas, 19º08′57″ S, 40º04′07″ W, 65 m a.s.l., human faeces, pitfall, 3 Feb. 2015, T. Vargas and L. Lopes leg. (CEMT); 1 ♁, 1 ♀, same data as for preceding, but 19°08′58″ S, 40°04′07″ W, 70 m a.s.l., (CEMT); 1 ♀, Soorotema, Reserva Natural Vale, Estrada Cinco Folhas, 19º08′32″ S, 40º03′55″ W, 59 m a.s.l., human faeces, pitfall, 2 Feb. 2015, T. Vargas and L. Lopes leg. (CEMT); 1 ♁, 2 ♀♀, same data as for preceding, but 19º09′09″ S, 40º02′19″ W (CEMT); 1 ♁, Soorotema, Reserva Natural Vale, Estrada Flamengo, 19º09′07″ S, 40º03′00″ W, 11 m a.s.l., human faeces, pitfall, 2 Feb. 2015, T. Vargas and L. Lopes leg. (CEMT); 1 ♀, same data as for preceding, but 17 m a.s.l. (CEMT). – Rio de Janeiro: 1 ♁, Xerém, 17 Aug. 1993, Áureo leg. (CEMT); 6 ♁♁, 2 ♀♀, Guapimirim, PARNA Serra dos Órgãos, -22.530383, -43.001075, 100 m a.s.l., pitfall, 16–18 Dec. 2014, C. Araújo and R. Andrade leg. (CEMT); 1 ♁, 1 ♀, same data as for preceding, but -22.516573, -43.007579, 259 m a.s.l. (CEMT); 2 ♁♁, same data as for preceding, but -22.494686, -43.002185, 400 m a.s.l. (CEMT); 1 ♁, same data as for preceding, but -22.426, -42.992918, 600 m a.s.l. (CEMT); 2 ♀♀, same data as for preceding, but -22.524244, -43.000526, 150 m a.s.l. (CEMT); 1 ♁, same data as for preceding, but -22.468017, -42.998243, 800 m a.s.l. (CEMT); 1 ♀, same data as for preceding, but 22º28′34″ S, 42º59′30″ W, Dec. 2014 (CEMT); 1 ♀, Guapimirim, PARNASO, 22º29′41″ S, 43º0′0″ W, 350 m a.s.l., pitfall, 16–18 Dec. 2014, C. Araújo and R. Andrade leg. (CEMT); 1 ♀, same data as for preceding, but 22º31′05″ S, 43º0′17″ W, 200 m a.s.l. (CEMT). – São Paulo: 3 ♁♁, Ubatuba, Oct. 1998 (CEMT); 1 ♁, Ubatuba, Banana podre, 25 Mar. 2003 (CEMT); 8 ♁♁, 2 ♀♀, Campinas, Bosque São José, 21 Sep. 1997, M. R. Mattos leg. (CEMT); 1 ♁, 1 ♀, same data as for preceding, but 19 Aug. 1997, M. Castro leg. (CEMT).
Distribution
Brazilian Atlantic Forest in the states of Alagoas, Bahia, Ceará, Espírito Santo, Minas Gerais and Rio de Janeiro (Fig. 8).
Taxonomic remarks
Canthon smaragdulus subviridis stat. rev. was described as a variety of C. speculifer (Schmidt 1922). It was also listed as a variety of Goniocanthon smaragdulus in a catalogue, but Vulcano & Pereira (1964) did not explicitly treat this as a taxonomic act, nor did they consult the type material. Finally, this taxon is also treated as a variety of C. smaragdulus in a catalogue by Vaz-de-Mello (2000).
description: Published as part of Nunes, Luis Gabriel de O. A., Nunes, Rafael V. & Vaz-De, Fernando Z., 2018, Taxonomic revision of the South American subgenus Canthon (Goniocanthon) Pereira & Martínez, 1956 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), pp. 1-31 in European Journal of Taxonomy 437 on pages 10-11, DOI: 10.5852/ejt.2018.437, http://zenodo.org/record/3787887