Phlaeothripinae Uzel 1895
Creators: name: Dang, Li-Hong
name: Mound, Laurence A.
name: Qiao, Ge-Xia
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Thysanoptera
Phlaeothripidae

description: Key to Phlaeothripinae genera from China and SE Asia
[excluding Austrothrips and Eothrips q.v.;*indicates no species studied]
1. Compound eyes holoptic, surrounding the ocelli (Fig. 72); mouth cone extending across prosternum; body black with yellow, red and white markings when alive....................................................... Macrophthalmothrips
-. Compound eyes smaller, never fully surrounding ocellar region (Figs 71, 73, 74); mouth cone various; body colour various, but never with yellow, red and white markings.................................................................. 2
2. Tube parallel-sided, long and slender with elongate anal setae, usually 2–4 times as long as tube (Fig. 25); abdominal segment IX usually much longer than wide (Fig. 25) [body with reticulate and tubercles on dorsal surface (Figs 28, 29, 33); basantra absent; usually aptera].................................................................................. 3
-. Tube various, but anal setae usually as long as tube, if long, basantra well-developed; body surface various; abdominal segment IX wider than length (Fig. 45)....................................................................... 8
3. Antenna 5- or 6-segmented, morphological segments III–V united, VII–VIII sometimes fused without sutures (Fig. 90) [head with 1–3 pairs of setae on anterior margin]....................................................... Stephanothrips
-. Antenna 7- or 8-segmented, segments III–V completely separated, VII–VIII sometimes fused without sutures............ 4
4. Anterior margin of head with 1–3 pairs of prominent setae (Figs 28, 29).......................................... 5
-. Anterior margin of head with no prominent setae (Fig. 33)..................................................... 6
5. Anterior margin of head with one pair of prominent setae (Fig. 29)..................................... Bradythrips
-. Anterior margin of head with three pairs of prominent setae (Fig. 28).................................... Baenothrips
6. Head projecting in front of eyes (Fig. 43).......................................................... Habrothrips
-. Head not projecting in front of eyes, but sometimes produced as a hump.......................................... 7
7. Postocular setae well-developed with apex expanded, close together; anal setae 4 times as long as tube or more.. Alerothrips *
-. Head without distinct long postocular setae (Fig. 33); anal setae about twice as long as tube.................... Urothrips
8. Antennal segment III much smaller than IV (Figs 31, 65, 91), these segments either broadly joined or fused.............. 9
-. Antennal segments III and IV normal, distinctly separated from each other....................................... 11
9. Pronotal setae blunt to expanded at apex (Fig. 91); antenna 8-segmented, segment III about 0.5 as long as IV, and widely joined to IV......................................................................................... Preeriella
-. At least pronotal posteroangular setae sickle-shaped at apex (Figs 31, 65); antenna usually 7-segmented, morphological segments III–IV fused with or without sutures................................................................ 10
10. Pronotal notopleural sutures absent; pm setae similar in size to pa setae................................. Crinitothrips
-. Pronotal notopleural sutures present, but incomplete; pm setae much smaller than pa setae................... Hyidiothrips
11. Antennal segment II with campaniform sensorium on median or basal half of segment (Fig. 96)...................... 12
-. Antennal segment II with campaniform sensorium on apical half of segment (Figs 12, 13, 16)........................ 17
12. Body with long barbed setae on elongate dorsal tubercles; tarsal tooth and hamus equally developed in all legs [tergite VIII with 2 pairs of wing-retaining setae]............................................................. Andrethrips *
-. Body with major setae simple; tarsal tooth and hamus not equally developed in any legs............................ 13
13. Antennal segment IV broad apically, with more than 4 prominent sensoria (Fig. 96); female abdominal sternite VIII with row of stout posteromarginal setae................................................................... Streptothrips
-. Antennal segment IV narrowed at apex, with 3 (or 2) prominent sensoria; female abdominal sternite VIII without stout setae.. 14
14. Sternite VIII posterior margin with series of long teeth........................................... Chirothripoides *
-. Sternite VIII simple, without posteromarginal teeth.......................................................... 15
15. Head produced in front of eyes; prosternal ferna large, fused medially................................. Mastigothrips *
-. Head not produced in front of eyes (Fig. 74); prosternal ferna of two distinct plates............................... 16
16. Antennal segment IV with 2 sensoria; fore femora inner margin with tooth............................... Menothrips *
-. Antennal segment IV with 3 sensoria; fore femora inner margin without tooth............................ Plectrothrips
17. Maxillary stylets short and V-shaped, usually scarcely retracted into head capsule (Figs 8, 84, 95)..................... 18
-. Maxillary stylets various, at least clearly visible in head anterior to postoccipital ridge.............................. 21
18. Antennal segment III with 3 sensoria, IV with 4 sensoria, segments VII–VIII broadly joined or fused (Fig. 11); prosternal basantra present (Fig. 8); fore wing constricted medially........................................... Akarethrips gen n.
-. Antennal segment III with 1–2 sensoria, IV with 2–3 sensoria, segments VII–VIII distinct from each other; prosternal basantra absent or weakly developed; fore wing without median constriction............................................. 19
19. Fore femora expanded, with at least one large tooth on inner margin (Figs 92, 93)....................... Veerabahuthrips
-. Fore femora without tooth on inner margin................................................................ 20
20. Metathoracic sternopleural sutures present; abdominal tergites III–VI each with one pair of wing-retaining setae in macropterae (Fig. 94); living on dead branches..................................................... Sophiothrips
-. Metathoracic sternopleural sutures absent; abdominal tergites III–VI each with 2 pairs of wing-retaining setae in macropterae; living on Poaceae ........................................................................... Bamboosiella
21. Abdominal tergite VIII with 1 or 2 pairs of curved wing-retaining setae (Figs 68, 86)............................... 22
-. Abdominal tergite VIII with no wing-retaining setae......................................................... 25
22. Pronotal aa absent; antennal segments III-IV each with 2 sensoria.................................... Solomonthrips
-. Pronotal aa developed; antennal segment III with 2 or 3 sensoria, IV with 3 or 4................................... 23
23. Eyes prolonged on ventral surface of head; tergite VIII with only one pair of curved wing-retaining setae...... Lizalothrips *
-. Eyes not longer ventrally than dorsally; tergite VIII with 2 pairs of curved wing-retaining setae....................... 24
24. Pronotal am setae minute (Fig. 131); antennal segment III with 3 sensoria, IV with 4; fore wing parallel-sided... Phylladothrips
-. Pronotal am well developed, as long as aa (Fig. 84); antennal segment III with 2 sensoria, IV with 3; fore wing slightly contricted medially........................................................................ Propesolomonthrips
25. Antenna 5-segmented, morphological segments III–VI joined (Fig. 77) [body surface bearing numerous tubercles, most setae broadly expanded; meso- and metanotum fused; abdominal tergite I transverse, broadly fused to tergite II; tube encircled by a transverse furrow medially]................................................................... Murphythrips
-. Antenna 7- or 8-segmented............................................................................. 26
26. Fore wing with 3 brown bands, without duplicated cilia, sub-basal setae weak (Fig. 21); abdominal tergites III–VI each with one pair of wing-retaining setae............................................................... Aleurodothrips
-. Fore wing rarely banded, duplicated cilia and sub-basal setae various; abdominal tergites III–VI usually with 2 pairs of wingretaining setae....................................................................................... 27
27. Antennae 7-segmented [body surface strongly reticulated].......................................... Strepterothrips
-. Antennae 8-segmented, but sometimes VII–VIII closely joined, with complete or incomplete suture................... 28
28. Antennal segment III usually with more than 6 stout sensoria (Fig. 12), if with 4, fore femur with a median tubercle in both sexes (Fig. 130)........................................................................... Ecacanthothrips
-. Antennal segment III with no more than 4 sensoria.......................................................... 29
29. Tube elongate, usually more than 10 times as long as basal width, with numerous lateral setae (Fig. 70); abdominal tergites II–V with 2 pairs of wing-retaining setae, usually fan-shaped; antennal segment IV with two sensoria........... Leeuwenia -. Tube less than 5 times as long as basal width; if tube elongate, then abdominal tergites II–V with numerous wing-retaining setae, or antennal segment IV with three sensoria........................................................... 30
30. Head with 2 pairs of prominent postocular setae (Fig. 105); antennal segment IV with 2 sensoria; head of female with ocellar triangle prolonged conically over base of antennae (Fig. 105)....................................... Coryphothrips
-. Head usually with one pair of postocular setae; if with 2 pairs then antennal segment IV with 3 sensoria; head not prolonged into cone over base of antennae......................................................................... 31
31. Antennal segment VIII elongate (Fig. 17); head constricted basally; maxillary stylets far apart from each other; pronotal am (and usually aa) setae minute, notopleural sutures incomplete; fore tarsal tooth absent in both sexes; fore wing parallel-sided...... 32
-. Not above combination of characters..................................................................... 34
32. Maxillary stylets parallel medially in head, about one third of head width apart; mouth cone pointed.......... Athlibothrips
-. Maxillary stylets wide apart and V-shaped in head; mouth cone broadly rounded................................... 33
33. Head about as long as wide (Fig. 71); fore wing sub-basal setae reduced; leaf-feeding species............... Litotetothrips
-. Head longer than wide; fore wing sub-basal setae well-developed; gall-inducing species................... Thlibothrips *
34. Antennal segment II asymmetric, inner margin sharply angulate (Fig. 104); head projecting over bases of antennae, first ocellus directed forwards (Fig. 104); mesopresternum completely absent [antennal segment III with one sensorium, IV with two sensoria; pronotal epimera laterally strongly eroded]................................................ Chelaeothrips
-. Antennal segment II not sharply asymmetric; anterior margin of head different; mesopresternum various, transverse, or reduced to two lateral triangles, or rarely fused to anterior margin of mesoeusternum............................... 35
35. Antennal segment III with one sensorium, IV with 3 (Fig. 89) [basantra usually absent]............................. 36
-. Sensorium number on antennal segments III and IV different from above........................................ 62
36. Basantra present; postocular setae minute......................................................... Jennythrips *
-. Basantra absent; postocular setae usually developed......................................................... 37
37. Eyes with posterior ommatidia enlarged (Figs 81, 82, 119, 120)................................................ 38
-. Eyes without enlarged posterior ommatidia................................................................ 41
38. Mesonotum, metanotum and tergites II–VI yellow, head and abdominal segments VII–X dark brown [maxillary stylets retracted almost to compound eyes, close together medially in head].................................. Praeciputhrips
-. Body uniformly brown................................................................................ 39
39. Posteromarginal setae on abdominal segment IX 0.6 as long as tube..................................... Manothrips *
-. Posteromarginal setae on abdominal segment IX as long as tube, or a little shorter................................. 40
40. Pronotum with notopleural sutures incomplete (Fig. 119)............................................ Gemmathrips
-. Pronotum with notopleural sutures complete (Fig. 120).............................................. Rosingothrips
41. Body strongly reticulate dorsally, usually with fine internal wrinkles; tube usually elongate, at least 5 times as long as basal width, and with fine lateral setae; head about twice as long as width across eyes, cheeks parallel (Fig. 3); antennal segments III–V usually slender; pelta triangular.................................................................... 42
-. Not above combination of characters..................................................................... 44
42. Maxillary stylets elongate, retracted to just behind eyes, close together medially (Fig. 3).................. Agynaikothrips
-. Maxillary stylets short and wide apart, retracted to basal third of head length...................................... 43
43. Abdominal tergites II–V with more than 4 pairs of sigmoid or straight wing-retaining setae (Figs 39, 41, 42)... Gigantothrips
-. Abdominal tergites II–V with 2 pairs of sigmoid wing-retaining setae, usually also with several pairs of accessory smaller wing-retaining setae........................................................................ Gynaikothrips
44. Postocellar setae elongate, much longer than diameter of posterior ocelli.............................. Chaetokarnyia *
-. Postocellar setae not elongate........................................................................... 45
45. Maxillary stylets retracted to compound eyes and close together medially in head (Figs 66, 124)...................... 46
-. Maxillary stylets not reaching compound eyes, about one third of head width apart medially......................... 47
46. Metanotum strongly reticulate; pronotum clearly narrower than prothorax width; female with a large fore tarsal tooth................................................................................................. Horistothrips
-. Metanotum weakly reticulate; pronotum occupying full width of prothorax; female without a fore tarsal tooth..................................................................................................... Psephenothrips
47. Female with sharply recurved, pointed tooth on inner apex of fore tibia (Fig. 142), and fore femora swollen but without tooth on inner margin.............................................................................. Myopothrips
-. Fore tibia of both sexes without a large recurved tooth at inner apex............................................. 48
48. Head strongly constricted basally, forming a neck with one or more stout setae just anterior to the constriction (Fig. 83); fore tibia of male with small triangular tooth at inner apex (Fig. 83)....................................... Sphingothrips
-. Head not constricted to a basal neck with stout setae; fore tibiae of male with no tooth or tubercle at inner apex.......... 49
49. Head with two pairs of equally developed postocular setae (Fig. 118)................................... Adelphothrips
-. Head with no more than one pair of well-developed postocular setae............................................ 50
50. Pronotum with no long major setae, epim no longer than width of fore tibia (Fig. 129); pronotum anterior angles with numerous stout setae (Fig. 129); head much longer than wide, cheeks with numerous stout setae, postocular setae minute (Fig. 129) [both sexes with large fore tarsal tooth].......................................................... Phasmothrips
-. Pronotum anterior angles without numerous stout setae, major setae usually elongate, head usually with postocular setae and cheeks without numerous stout setae..................................................................... 51
51. Postocular setae behind inner margin of compound eyes, and cheeks with numerous short stout setae (Fig. 121) [antennal seg- ments III–IV sensoria 0.75 as long as their segments]................................................ Zelotothrips
-. Postocular setae arising further apart; cheeks without numerous setae........................................... 52
52. Head with one pair of stout setae on basal third of cheeks (Fig. 80).................................. Ponticulothrips
-. Head without such stout cheek setae...................................................................... 53
53. Antennal sensoria on III and IV long and slender, at least 2/3 as long as segment.................................. 54
- Antennal sensoria scarcely half as long as segment.......................................................... 55
54. Male with fore tarsal tooth; head scarcely long than wide (Fig. 126).................................... Eugynothrips
-. Male without fore tarsal tooth; head much longer than width (Fig. 127)................................. Phenicothrips
55. Postocular setae minute, not extending to posterior margin of eye (Fig. 109)...................................... 56
-. Postocular setae well-developed........................................................................ 57
56. Body surface strongly reticulate; fore tarsal tooth present in female; tergite IX setae much shorter than tube... Medogothrips
-. Body surface not reticulate, pronotum with transverse sculpture lines; fore tarsal tooth absent in both sexes; tergite IX setae longer than tube (Fig. 109).................................................................... Pnigmothrips
57. Fore tarsal tooth present in both sexes.................................................................... 58
-. Fore tarsal tooth absent in females, rarely present in males.................................................... 59
58. Pronotal epimeral setae well developed, usually much longer than other major setae (Fig. 32); metathoracic sternopleural sutures well developed....................................................................... Crotonothrips
-. Pronotal 5 pairs of major setae well developed, epimerals slightly longer (Fig. 122); metathoracic sternopleural sutures present, but short................................................................................. Liophloeothrips
59. Male with fore femora enlarged and large fore tarsal tooth; fore tarsus of female without tooth............. Propealiothrips
-. Fore femora of male not enlarged; fore tarsal tooth absent in both sexes.......................................... 60
60. Antennal segment VIII long and slender, constricted basally, as long as VII, or a little longer (Fig. 13)..... Eurhynchothrips
-. Antennal segment VIII short and slightly constricted basally, distinctly shorter than VII............................. 61
61. Male tergite IX with setae S2 as long as setae S1.................................................. Isotrichothrips
-. Male tergite IX with setae S2 much shorter than setae S1............................................... Liothrips
62. Body surface strongly reticulate, many short sculpture lines inside each reticle (Fig. 87); maxillary stylets long, retracted to eyes, close together medially (Fig. 87); pronotal major setae and postocular setae short, broadly expanded at apex (Fig. 87); postocular setae close together (Fig. 87); metathoracic sternopleural sutures present; fore wing, if developed, weakly constricted medially, with duplicated cilia............................................................ Azaleothrips
-. Not above combination of characters..................................................................... 63
63. Mesopresternum fused to mesoeusternum medially (Figs 24, 116).............................................. 64
-. Mesopresternum transverse, or reduced to two lateral triangles, never fused to mesoeusternum (Figs 50, 52, 54).......... 65
64. Antennal segment VIII clearly separate from VII, III with 3 sensoria; pronotal am, aa and ml setae not arising close together (Fig. 19); fore wing constricted medially........................................................... Asianthrips
-. Antennal segment VIII fused to VII, III with 2 sensoria (Fig. 114); pronotal am, aa and ml setae arising close together (Fig. 113); fore wing parallel sided................................................................. Trichinothrips
65. Head strongly reticulate and with cheeks distinctly in-cut just behind eyes (Fig. 78)................................ 66
-. Head not strongly reticulate and with cheeks not in-cut behind eyes............................................. 68
66. Pronotum with 5 pairs of major setae with broadly expanded apices (Fig. 78); postocular setae long and capitate. Mystrothrips
-. Pronotum with only epimeral setae developed; postocular setae minute.......................................... 67
67. Head with cheeks almost parallel-sided; antennal segment III with 3 sensoria; pronotal epimeral setae less than 20 microns long..................................................................................... Sagenothrips *
-. Head with cheeks swollen behind eyes (Fig. 46); antennal segment III with 2 sensoria; pronotal epimeral setae more than 30 microns long (Fig. 46)...................................................................... Heliothripoides
68. Basantra present, usually well developed.................................................................. 69
-. Basantra absent or weak............................................................................... 86
69. Anterior margin of head produced into a prominent ensiform process overhanging base of antennae (Fig. 57)..... Sinuothrips
-. Anterior margin of the head without such a process.......................................................... 70
70. Postocular setae minute; only epimerals developed and expanded on pronotum; forewing without duplicated cilia [antennal segments III–IV with 2 and 4 sensoria respectively]........................................... Ananthakrishnana *
-. Postocular setae usually well developed, at least reaching posterior margin of eyes; all five pairs of pronotal setae developed, sometimes am, ml reduced; forewing with or without duplicated cilia........................................... 71
71. Metathoracic sternopleural sutures present................................................................. 72
-. Metathoracic sternopleural sutures absent................................................................. 75
72. Basantra well developed, longer than wide (Fig. 59)......................................................... 73
-. Basantra usually broader............................................................................... 74
73. Pronotal notopleural sutures complete (Fig. 58); anal setae usually about twice as long as tube (Fig. 141); fore wing constricted medially..................................................................................... Podothrips
-. Pronotal notopleural sutures incomplete (Fig. 133); anal setae slightly longer than tube; fore wing parallel-sided Okajimathrips
74. Anal setae long, about 1.5 times as long as tube (Fig. 140); pronotal am setae minute, ml setae as long as aa setae (Fig. 125).......................................................................................... Praepodothrips
-. Anal setae as long as tube; pronotal am and ml setae minute (Fig. 26)................................... Bagnalliella 75. Pronotal notopleural sutures incomplete (Fig. 60); antennal segment III with one sensorium, IV with 4........... Dyothrips
-. Pronotal notopleural sutures complete; antennal segments III–IV with different sensoria formula than above............ 76
76. Fore femur inner margin with small blunt tubercle either at base or medially (Figs 5, 110)........................... 77
-. Fore femur without tubercle on inner margin............................................................... 78
77. Fore femur inner margin with small blunt tubercle at base (Fig. 5)...................................... Androthrips
-. Fore femur inner margin with small blunt tubercle medially (Fig. 110)................................. Sumatrothrips
78. Head sharply constricted basally, with several stout setae on cheeks (Fig. 56); maxillary stylets short and V-shaped, not reaching postocular setae; stout fore tarsal tooth present in both sexes; antennal segment III with 3 sensoria, IV with 4.. Mesothrips
-. Not this combination of characters....................................................................... 79
79. Mouth cone straight, pointed, usually extending to mesothorax (Fig. 35)................................ Dolichothrips
-. Mouth cone usually short and rounded.................................................................... 80
80. Wings and tergal wing-retaining setae usually absent; antennal III with 1 sensorium, segment IV with 2 or 3; pronotum with 5 well developed setae........................................................................ Apterygothrips
-. Not above combination of characters..................................................................... 81
81. Abdominal segments II–VI with 1 pair of wing-retaining setae (Fig. 69).................................. Plicothrips
-. Abdominal segments II–VI with 2 pairs of wing-retaining setae................................................ 82
82. Anal setae elongate, usually twice as long as tube or more, if shorter then pronotum notopleural sutures incomplete; basantra sometimes as long as wide; pronotal am setae usually minute (Fig. 55)................................. Karnyothrips
-. Anal setae shorter, no more than 1.5 times as long as tube; pronotal am setae usually developed...................... 83
83. Fore tibia with small tooth at inner apex in both sexes (Fig. 112)....................................... Glenothrips
-. Fore tibia with no small tooth at inner apex................................................................ 84
84. Antennal segment III with 1 or 2 prominent sensoria................................................. Haplothrips
-. Antennal segment III with 3 sensoria..................................................................... 85
85. Pronotum with 3 pairs of major setae, am and aa minute (Fig. 108).................................... Logadothrips
-. Pronotum usually with 5 pairs of major setae (am rarely small)....................................... Xylaplothrips
86. Maxillary stylets long, retracted to eyes, close together medially for full length of head (Figs 61, 66)................... 87
-. Maxillary stylets never close together medially for full length of head........................................... 90
87. Antennal segments III and IV each with 3 sensoria (Fig. 88)......................................... Pygmaeothrips
-. Antennal segment IV with 4 sensoria..................................................................... 88
88. Antennal segment III with 4 sensoria............................................................ Oidanothrips
-. Antennal segment III with 2 or 3 sensoria................................................................. 89
89. Fore tarsal tooth present in both sexes............................................................. Holothrips
-. Fore tarsal tooth absent in both sexes (Fig. 1)...................................................... Ablemothrips
90. Female sternite VIII with pair of stout or leaf-like posteromarginal setae (Fig. 102); major setae strongly expanded at apex (Figs 97, 98)............................................................................... Terthrothrips
-. Female sternite VIII without pair of stout posteromarginal setae............................................... 91
91. Wingless; pronotal am, aa and ml setae no larger than discal setae, pa setae often also small or scarcely 0.5 as long as epimeral setae; postocular setae not reaching posterior margin of eyes (Fig. 123)................................ Cephalothrips
-. Usually macropterous; at least pronotal ml, pa and epimeral setae long, also postocular setae extending to posterior margin of eyes............................................................................................... 92
92. Apex of fore femora at inner margin with tooth in both sexes (Fig. 2); head without pair of stout setae on cheeks (Fig. 2).............................................................................................. Acanthothrips
-. Fore femora of female without tooth, male sometimes with tooth; head with pair of stout setae on basal third of cheeks.... 93
93. Maxillary stylets short, wide apart, usually V- or U-shaped.................................................... 94
-. Maxillary stylets long, close together medially............................................................. 97
94. Pronotum with 3 pairs of major setae, am and aa minute (Fig. 73)..................................... Psalidothrips
-. Pronotum with 4 or 5 pairs of major setae, sometimes am minute............................................... 95
95. Antennal segment III with 2 sensoria........................................................... Prosantothrips
-. Antennal segment III with 3 sensoria..................................................................... 96
96. Abdominal tergites with one pair of wing-retaining setae; pronotum with 5 pairs of major setae; antennal segment IV with 3 sensoria................................................................................ Margaritothrips *
-. Abdominal tergites with two pairs of wing-retaining setae; pronotum usually with 4 pairs of major setae, am usually minute; antennal segment IV usually with 4 sensoria...................................................... Deplorothrips
97. Maxillary stylets relatively broad, usually 4–6 microns wide (Fig. 20); pronotal aa and ml setae relatively close together (Fig. 20)...................................................................................... Apelaunothrips
-. Maxillary stylets slender, usually 2–3 microns in diameter; pronotal aa and ml setae well separated.................... 98
98. Postocular setae behind inner margin of eyes (Fig. 4); metathoracic sternopleural sutures absent............ Adraneothrips
-. Postocular setae wider apart; metathoracic sternopleural sutures usually present................................... 99
99. Major setae usually expanded at apex (Figs 63, 64); fore wings usually weakly constricted medially....... Hoplandrothrips
-. Major setae usually acute at apex (Fig. 62); fore wings, if developed, parallel-sided........................ Hoplothrips
description: Published as part of Dang, Li-Hong, Mound, Laurence A. & Qiao, Ge-Xia, 2014, Conspectus of the Phlaeothripinae genera from China and Southeast Asia (Thysanoptera, Phlaeothripidae), pp. 1-82 in Zootaxa 3807 (1) on pages 10-14, DOI: 10.11646/zootaxa.3807.1.1, http://zenodo.org/record/4918122