Nyctiophylax (Paranyctiophylax) kupirah Oláh & Johanson 2010, new species
Creators: name: Oláh, János
name: Johanson, Kjell Arne
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Polycentropodidae
Nyctiophylax
Nyctiophylax kupirah

description: Nyctiophylax (Paranyctiophylax) kupirah, new species   
Figs 23–26  
 This species belongs in the Nyctiophylax (Paranyctiophylax) flavus species group and is most similar to N. padangensis from North Sumatra, but differs by its triangular, not spatulate cerci; ventral paraproctal processes each with shorter ventrad-curving apical part; gonopods each with well-developed ventrobasal “elbow;” the shorter and basally curved ventroapical median lobe of sternite IX; and the presence of a pair of strong, anterad-directed spines and an ovoid phallotremal sclerite in the phallic apparatus.   
 Male. Body uniformly pale brown. Maxillary palp formula IV-II-I-III-V, segment III originating from mesal face basal of apex of segment II. Forewings 4.7 mm long; brown, with hypertrophied longitudinal vein R and pronounced hyaline patch pattern: small, rounded spot below discoidal cell around crossvein r-m in subradial cell; and large, triangular spot along crossvein m-cu and point of basal bifurcation of M; discoidal cell and median cell present; apical forks 2, 3, 4 and 5 present; A1, A2, and A3 looping. Hind wing discoidal cell closed, median cell open; apical forks 2 and 5 present.   
Spur formula 3,4,4.  
Male genitalia. Sternite IX robust, triangular in lateral view with well-developed apicoventral mesal lobe having regular rectangular apex in ventral view and sinuous basal half in lateral view, shorter than ventrobasal “elbows” of gonopods. Tergite IX small, sclerotized; elongated obliquely vertical, discernible dorsal of sternite and anterior of paraproct. Segment X membranous, nearly invisible; with triangular mesal lobe in dorsal view. Cerci setose, triangular in lateral view; clavate in dorsal view; shorter than gonopods and ventral paraproctal processes. Paraproctal complex well-developed, highly sclerotized; consisting of pair of large ventral paraproctal processes curving ventrad, each bearing 4-6 setae in row on outer surface of ventradcurving apical half. Dorsal paraproctal processes forming pair of hyaline digitate processes. Gonopods elongate, slender, uniformly curving in ventral view; each with basal half broadening, forming ventrobasal “elbow;” distal half slender; apex slightly excised, located laterally. Phallic apparatus located dorsally in genitalia, fixed, guided by paraproct and segment X; middle section of phallotheca forming short tube. Phallobase modified into long, narrow, arching dorsal apodeme and short ventral apodeme in lateral view; dorsal apodeme distinctly clear-cut, arching and narrowing into spine-like process in lateral view; weakly narrowing in dorsal view. Phallicata membranous, with pair of anterad-directed, stout spines and ovoid, horizontally elongate phallotremal sclerites embedded in phallic membrane.  
 Holotype male: INDONESIA (Sumatra): Bukit Barisan Selatan National Park, Barisan Mts., Way Pitias, 800 m, 17–19.vi.2008, light trap [J. Oláh, Jr.]— 1 male (OPC, in alcohol).   
 Paratypes: same data as holotype— 10 males (OPC, in alcohol).   
 Distribution: Indonesia (Sumatra).   
 Etymology: Kupirah, from “kupirah,” elbow in Sanskrit, referring to the large, ventrobasal “elbows” of the gonopods.
description: Published as part of Oláh, János & Johanson, Kjell Arne, 2010, Fifteen new Trichoptera (Insecta) species from Sumatra, Indonesia, pp. 1-35 in Zootaxa 2618 (1) on pages 15-16, DOI: 10.11646/zootaxa.2618.1.1, http://zenodo.org/record/5304087