Olmiana Guglielmino & Bückle & Emeljanov 2010, gen. nov.
Creators: name: Guglielmino, A.
name: Bückle, C.
name: Emeljanov, A. F.
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Achilidae
Olmiana

description: Genus Olmiana gen. nov.   
 Type species: Olmiana argentina sp. nov..   
 Description: Head smaller than pronotum, ratio width head: width pronotum 0.58–0.63 (Fig. 1A–D). Vertex subpentagonal, distinctly bordered by sharp carinae; ratio width on posterior margin to length 2.38–2.6 (Figs 1A, 1B); in the middle of posterior margin a triangular bulge extending to about 2/3 of length of vertex with narrow indistinct carina on its sagittal line; between median bulge and lateral margins on both sides with circular grained matt areas. Eye in dorsal view narrower than vertex. Ocellus nearer to eye and antenna than to lateral margin of frons. In lateral view, vertex and frons meet at an angle. Frons elongate, widest near clypeus, slightly and equally narrowed towards vertex (Figs 1C, 1D). Lateral margins acutely carinate; median line with well developed longitudinal carina continuing along postclypeus. Surface between medial carina and lateral margins with two pairs of indistinct, slightly convex areas: one rounded pair near upper margin of frons and one elliptic pair level with antennae. Frontoclypeal suture arcuate, slightly pronounced, evanescent near medial carina. Postclypeus with well developed acute lateral carinae; medial carina distinct, rather acute near frontoclypeal suture, somewhat smoothed in other regions. Anteclypeus tectiform; lateral carinae indistinct, running parallel to indistinct very smoothed medial carina and vanishing at about 1/3 of anteclypeus length. Rostrum long, extending posteriorly beyond metacoxae. Pronotum short and wide, ratio width to length 4.54– 4.90, anteriorly convex, posteriorly with broad obtusely angulate excavation (Fig. 1A). Lateral margin with distinct carina posteriorly, ending anteriorly in slightly protruding gibbosity (Fig. 8D). Disc delimited by elevated carina; its lateral branches converging anteriorly and fused in the middle in rounded or subacute arch immediately behind anterior margin of pronotum, posteriorly not reaching hind margin, but bent sideways towards lateral carina. Median carina distinct along entire length. Most of pronotum, particularly disc, shining, distinctly and deeply wrinkled. Anterolateral sector on each side grained and matt, without wrinkles. Mesonotum (Fig. 1A) in anterior 3/4 with median carina quite elevated in anterior half, interrupted in the middle, vanishing caudally. Lateral carinae diverging and reaching posterior margin of mesonotum. Mesonotum with alternating wrinkled shiny and grained matt areas, the latter consisting in wide transversal band along anterior margin except for region immediately near carinae and rounded area outside lateral carinae near posterior margin. Tegulae without carina, longitudinally wrinkled. Fore wings slender, apical margin broadly rounded-truncate (Fig. 2A). Basal cell elongate. Common stem Sc+R+M short. Sc+R and CuA forking at about the same level. M ramifying near nodal line; anterior branch ramifying again, posterior branch distally fusing with anterior branch of CuA. Hind wings (Figs 2B, 2C) fully developed, with twobranched RP, bifurcate MA, MP and CuA connected by very short transverse vein, and three-branched CuA. Anterior branch of A 1 diverging from 1 st anal wing fold far distally of fork of posterior branch, the latter uniting with A 2. Vein A 2 with 2–4 blind branches (to some degree variable among different specimens) reaching neither wing margin nor 2 nd anal fold. Hind tibiae with tooth just after the middle of outside margin and about a dozen apical teeth arranged on two rows. First hind tarsomere with apical row of teeth, without platellae; second hind tarsomere with apical row of teeth, each tooth with platella except for marginal ones (Figs 1E, 1F).   
Aedeagus with one dorsal lobe, two lateral lobes and two ventral lobes.  
 Diagnosis: On the basis of the peculiar anastomosis between first and second anal veins of hind wings and length of the rostrum we attribute the new genus to the tribe Achilini (see Emeljanov 1991). However, the hind wing of Olmiana has a three-branched CuA, a character separating it from all other genera of the tribe, which display four-branched CuA.
description: Published as part of Guglielmino, A., Bückle, C. & Emeljanov, A. F., 2010, Olmiana argentina, a new genus and species of Achilidae (Hemiptera, Fulgoromorpha) from Argentina., pp. 47-58 in Zootaxa 2661 (1) on pages 48-49, DOI: 10.11646/zootaxa.2661.1.3, http://zenodo.org/record/5301791