Phylloicus ephippium Prather 2003, new species
Creators: name: PRATHER, AYSHA L.
Publisher: name: Zenodo
Year: 2003
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Calamoceratidae
Phylloicus
Phylloicus ephippium

description: Phylloicus ephippium, new species
Figs. 47, 48
Phylloicus ephippium is distinguished by the wing pattern, which in dorsal view appears to have a saddle of dark brown across the middle of the wing, and segments IV and V of the male, which are highly modified (Fig. 47F), with a long, projecting lateral sclerite of tergum IV. Only mesal coremata are present; these are bifurcate and setose.
Adult. Forewing length 12.1 mm, n = 6.
Head golden brown, with dorsomesal crest of black setae. Maxillary palps dark brown. Antenna twice forewing length; dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax dark brown; ventrolateral thorax pale tan. Legs pale tan. Metathoracic leg of male with posterior fringe of long setae, setae pale. Tibial spur formula 2,4,4. Forewing flat; chestnut brown; with two transverse bands; proximal band golden, reaching posterior wing margin, at least 1/2 width of wing, an inverted V­shape; distal band tan, not reaching either wing margin, 1/2 width of wing or less; with two basal stripes, golden; "V" of proximal band encloses “S,addle” of dark brown setae on posterior margin; proximal half of cell Cu 2 clear. Hind wing basal brush present in male, dark brown.
Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum IV with paired lateral sclerites, mesal coremata; lateral sclerite contorted, setose apically; mesal coremata bilobed, mesal lobe setose, capable of inflation to 1/2 length segment V, lateral lobe very long, fully extended reaching to end of abdomen, covered with setae. Tergum V with pair of semi­elliptical ridges on posteromesal corners (Fig. 47F). Sternum VII without anteromesal process. Sternum VIII enclosing base of elongate sternum IX; posteromesal process truncate (Fig. 47A, C). Tergum IX without mesal ridge; posterior margin with round narrow mesal projection; with anterior ridge broken mesally; posterior margin with smooth rounded mesal projection (Fig. 47B); lateral ridge present; dorsal pleural setae approximately 8, ventral pleural setae approximately 5 (Fig. 47A). Preanal appendage approximately length of tergum X, of uniform diameter throughout length, setae long, but not filamentous or longer than appendage (Fig. 47A, B). Tergum X without basal lobes; basodorsal process long and digitate; basolateral processes absent; apex, in lateral view, rounded, in dorsal view, notched, notch triangular (Fig. 47A, B). Harpago rounded; peglike setae few, apical (Fig. 47A, C). Phallic endotheca with paired basolateral lobes, basolateral lobes large and round; phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped (Fig. 47D, E).
Female. Preterminal abdominal terga with anteromesal notch. Sternum VII without anteromesal process. Tergum VIII without posterolateral brush; sternum VIII with shallow posteromesal notch; sternum VIII (Fig. 48C). Tergum IX with very short mesal ridge (Fig. 48B). Sternum IX anterior lobes darkly sclerotized and striate, posterior lobes smooth, without distinct area of thin cuticle or invagination (Fig. 48A). Tergum X appendage shorter than mesal lobe, base marked by faint suture line, apex oblique; mesal lobe lightly sclerotized; digitate lateral processes length approximately equal diameter (Fig. 48B). Sternum X with patches of short fine setae posterolaterally to anal opening (Fig. 48A). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite truncate anteriorly, posterolateral projections acute; posterior sclerite triangular; posterior end of spermatheca a sclerotized expanded tube (Fig. 48A).
Holotype male: ECUADOR: Tungurahua: 13 km E Baños, 1550 m, 15.ix.1990, Flint (NMNH).
Paratypes: ECUADOR: Tungurahua: 13 km E Baños, 1550 m, 15.ix.1990, Flint — 1 female, 3 males (NMNH); 1 male (UMSP).
Distribution. Ecuador.
Etymology. Ephippium, from the Latin, meaning “Saddle,” referring to saddle pattern of the forewings.
description: Published as part of PRATHER, AYSHA L., 2003, Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae), pp. 1-214 in Zootaxa 275 (1) on pages 50-51, DOI: 10.11646/zootaxa.275.1.1, http://zenodo.org/record/5019235