Frutillaria chepuensis Richards
Creators: name: Kits, Joel H.
name: Marshall, Stephen A.
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Sphaeroceridae
Frutillaria
Frutillaria chepuensis

description: Frutillaria chepuensis Richards
(Figs. 16–19, 72, 73, 76, 109)
Frutillaria chepuensis Richards, 1961: 66 –67 Description. Length 1.9–2.6 mm (head + thorax 0.8–1.1 mm)
Occiput brown, face and gena orange, frons intermediately coloured. Frons with shiny spots lateral to ocellar triangle. Scape and first flagellomere orange, pedicel brown. Clypeus orange, prementum brown.
Thorax black, postalar callus and scutellum reddish, wing rudiment pale brown. Mesoscutum and scutellum covered with microtomentum. Anepisternum mostly shiny, posterior margin and part of dorsal margin with microtomentum, sometimes anterior margin with some microtomentum below spiracle. Katepisternum with a large shiny patch behind fore coxa extending posteriorly as a triangle to near bristle, narrowly separated from posterior shiny patch by a band of microtomentum. Meron and metapleuron shiny with the exception of a stripe of microtomentum between posterior thoracic spiracle and base of hind coxa. Laterotergite mostly shiny.
Legs black, fore coxa dark reddish brown, trochanters, joints, and tarsi orange-yellow. Mid femur with 2 anterior bristles.
Male abdomen (Figs. 16–19): Tergites 1–5 with microtomentum. Sternite 2 shiny, sternites 3–4 mostly shiny with some microtomentum medially, sternite 5 mostly covered with microtomentum. Surstylus with broad basal part, clubbed inner arm, outer corner appearing broadly rounded to somewhat pointed, depending on angle of view. Postgonite bilobed, anterior arm short and itself bilobed. Epiphallus short, broadened laterally at tip; preepiphallus long, nearly parallel-sided. Distiphallus with lateral arms extending narrowly from ventral sclerite, projecting past central projecting sclerite and fan-shaped at tip; central projecting sclerite with apical portion strongly ridged; apicoventral discs present, elongate; ventral sacs clear, projecting perpendicularly.
Female abdomen (Figs. 72, 73, 76): Tergites 1–5 covered with microtomentum, with posterior margins shiny, tergite 6 shiny on basal half, covered with microtomentum on apical half. Sternite 2 shiny, sternites 3–5 mostly shiny with microtomentum medially, sternite 6 shiny on basal half, covered with microtomentum on apical half. Tergites 7 and 8 mostly shiny with some microtomentum along posterior margins, tergite 8 divided medially. Epiproct triangular with fairly long anterior arms. Cerci without microtrichia dorsally, but some apically. Sternite 7 mostly shiny, with some microtomentum along posterior margin. Sclerites of sternite 8 round to comma-shaped. Hypoproct with sclerotized lateral spots connected by a weakly sclerotized bar. Spermathecae with long, mushroom-shaped apical invagination, short basal invagination, both annulated.
Type material. Holotype 3: CHILE: [Los Lagos:] Chiloé I., Chepu, 42ºS, 3 Oct 1958, [G. Kuschel] (MEUC, apparently lost). Paratypes: same locality as holotype but dates from 2–17 Oct 1958 (28 3, BMNH, not examined).
Other material examined. CHILE: Los Lagos: Chiloé I., Terao, nr. Chonchi, 50 m, 2nd growth forest, Chile Exp., 19–21 Feb 1988, L. Masner (3 3, 1 Ƥ, CNC); Chiloé I., Terao, 10 km S Chonchi, 42°43'01"S 73°38'47"W, 45 m, high meadow, yellow pans, 13–15 Feb 2005 (13 3, 20 Ƥ, DEBU); Chiloé I., 11 km NW Castro, 200 m, ravine, second growth forest, FMHD #85-948, carrion trap, 27 Dec 1984 – 1 Feb 1985, S. & J. Peck (2 3, 2 Ƥ, FMNH); Chiloé I., Lago Tepuhueico, SE edge, 42°48'06"S 73°55'24"W, 50 m, Valdivian rainforest, FMHD #97-22, berlese, leaf & log litter, 15 Jan 1997, A.F. Newton & M.K. Thayer (2 3, FMNH); Chiloé I., Colonia Yungay, road to (3.6 km W Hwy 5), 42°59'00"S 73°41'00"W, 90 m, grazed secondary Valdivian rainforest remnants, FMHD #97-24, berlese, leaf & log litter, 17 Jan 1997, A.F. Newton & M.K. Thayer (4 3, 9 Ƥ, FMNH).
Comments. This species is only known from Chiloé Island. Although we did not examine any paratypes, its identity is unambiguous based on Richards’ illustration and description. Most of the paratypes are apparently lost, with only 13 specimens currently present in the BMNH (Pitkin 1989; N. Wyatt, pers. comm.)
description: Published as part of Kits, Joel H. & Marshall, Stephen A., 2011, A revision of Frutillaria Richards and Penola Richards (Diptera: Sphaeroceridae: Archiborborinae), pp. 1-34 in Zootaxa 2863 on pages 11-12, DOI: 10.5281/zenodo.203208