Canthophorus impressus subsp. impressus
Creators: name: Gapon, Dmitry A.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Cydnidae
Canthophorus
Canthophorus impressus
Canthophorus impressus impressus (horváth, 1880)

description: Canthophorus impressus impressus (Horváth, 1880) (Figures 1 E, 3 F, 6 N, 6 O, 7 G, 8 J, 8 K, 10 A, 10 B, 12 C)
Material examined
Without locality: “k. V. Yakovleva [V. Jakovlev coll.]”, [date unknown], 1♀ (ZISP); “ Sehirus dubius Scop. ”, [date and collector unknown], 1♀ (ZISP). Germany: Bavaria, Bamberg, 31. V.1953 (Eckerlein), 1J (MNHN). Italy: Lombardia, Bormio, 1933 (Passauro), 1♀ (ZISP). Bulgaria: Sofia Prov., Vitosha Mt., Cherni Vrah, 4.VI.1954 [collector unknown], 2♀ (ZMUM). Ukraine: Odessa Prov.: [?], forest, bottom of gully, 23.VII. [?] (D.V. Znojko), 1♀ (ZISP); Odessa, Khadzhibeyskiy Liman, 29.VII.1920 (Kiritshenko), 1♀ (ZISP); Luzanovka [Odessa], 22.IX.1920 (Kiritshenko), 1♀ (ZISP); Chernovo [Andreevo-Ivanovka Settlm.], Tiligul Riv., 17.I.1921 (Znojko), 2♀ (ZISP); Mykolaiv Prov.: “pered Rakovskim poselkom” [Rakovo Settlm.], 21.VIII. [?] (D.V. Znojko), 1J (ZISP); Trikraty Settlm., Arbuzin[ka] Riv., 1.IX.1923 (A-y Kirichenko), 1♀ (ZISP); Poltava Prov.: Mirgorod, [date unknown] (Taranov), 1♀ (ZISP); Poltava, 1926 (F. Lukjanovitsh), 1♀ (ZISP); Donetsk Prov.: Velikiy Anadol forestry, sweeping over grass, 3.IV., 15. VI.1905 [collector unknown], 2J (ZISP); Sviatogorsk, forest Zaseka, 15.VI.1930 (Arnoldi), 1♀ (ZISP); Lugansk Prov.: Lisichansk, floodplain of Borovaya Riv., 19.VI.1937 (Arnoldi), 1♀ (ZISP); Autonomous Republic of Crimea: Ay-Petri Mt., 6. VI.1911 (I. Levandovskiy), 1♀ (ZISP), 19.X.1966 (K. Arnoldi), 1J (ZMUM); Kerch’, 28.VI.1916 (A. Kiritshenko), 1♀ (ZISP); “ yayla Besh-tekme ”, 15.VII.1927 (Kiritshenko), 1J, 1♀ (ZISP); Krymskiy Nat. Reserve, 18,20,23. VIII.1936 (V. Geptner), 1J, 14♀ (ZMUM); Roman-Kosh Mt., Babugan-Yayla Mts, 19. VI.1947 (K. Arnoldi), 3♀ (ZISP); lower plateau of Chatyrdag, 28.VI.1984 (A.K. Zaguljaev), 1♀ (ZISP); Demir-Kapu Mt., 10. VII.1948 (K. Arnoldi), 1J, 1♀ (ZISP). Russia: Kursk Prov., Tsentral’ no-Chernozemnyy Nat. Reserve, Streletskaya step’, V.1964 (K. Arnoldi), 1♀ (ZMUM); Voronezh Prov., Kamennaya step’, 11 km S Talovaya, 26.VI.1949, 27.VII.1935, (V. Stark), 2♀ (ZISP) [paratypes of C. mixtus]; Rostov Prov., khutor Abramovka, 8.VII.1949 (K. Arnoldi), 1J, 1♀ (ZMUM); Krasnodar Terr.: “ Kubansk. obl.”, [date unknown] (V. Vorob’ ev), 1♀ (ZISP); Krasnodar, 12.IV.1911 (BogdanovKat’ kov), 1♀ (ZISP); Dzhuga Mt., 15.VI.1911 (Volnukhin), 1♀ (ZISP); Kubanskaya Settlm., 14.III.1934 (G. Zimin), 1J (ZISP); Adyghe Republic, 15 km SW Guzeripl’, 25,26. VI.1976 (E. Sugonjaev), 1♀ (ZISP); Stavropol’ Terr.: Kislovodsk, 7.VI.1908 (coll. Kiritshenko), 1♀ (ZISP); Bashanta, 21.VIII.1914 (Uvarov), 1♀ (ZISP); Essentuki, Belyy ugol’, at hills, 21.VIII.1960 (E.S. Sugonjaev), 1♀ (ZISP); Strizhament Mt., 30 km S Stavropol’, 6. VII.1994 (V.A. Krivokhatskiy), 1J (ZISP); Beshtau Mt., Beshtaugorskiy Nat. Reserve, 15.VI.2010 (D. Gapon), 1♀ (ZISP); Karachay-Cherkess Republic, glacier in vicinity of Teberda, valley of Muhu Riv., 5.VIII.2013 (V. Golub), 1J (ZISP); Kabardino-Balkar Republic, “Gundelen” [Kendelen Settlm.], 24.VII.1926 (Berg), 1♀ (ZISP); Republic of North Ossetia-Alania: Il Mt. [Sapitskaya budka] SE Vladikavkaz, 26. IV.1886 (Ananov), 1♀ (ZISP); Republic of Ingushetia, Salgi, “5462′”, 1.VII.1927 (Kiritshenko), 2J (ZISP); Chechen Republic, st. Grebenskaya, 27.V.1960 (Tanasiychuk), 1♀ (ZISP); Republic of Dagestan, “Darg.[inskiy] okr.[ug] ”, Khodzhal-Makhi [Khadzhalmakhi Settlm.], 25.VI.1926 (Rjabov), 1♀ (ZISP); Orenburg Prov., “Orb” [Orenburg] (Eversmann coll.), 2♀ (ZISP); Chelyabinsk Prov. [?], [Chelyabinsk?], Kashtak, 18.V.1936 (S. Rodionoff), 1♀ (ZISP); Krasnoyarsk Terr., “Tomsk. gub, Mar. u.”, Elizavetinskiy vinokurennyy zavod [distilling factory]”, Bogotol’ skiy Distr., Krasnyy zavod Settlm., 15–30.VI.1903 (coll. Kiritshenko), 1♀ (ZISP); Altai Republic, Ongudayskiy Distr., 10 km SE Seminsky pass, nr Tuekta Riv., c. 1300 m, 16. VI.1999 (A.G. Kirejtshuk), 1J (ZISP); Republic of Tyva, Sosnovka, “kkh. [kolkhoz] Kr[asnyy]. part[izan]”., O Tannu-Ola Mts, 2,3.VII,1948 (Dyatlova), 2♀ (ZISP); Irkutsk Prov., Padun Settlm. [Bratsk] “na Verkhney Tunguske” [on Angara Riv.], [18]67 (Chekanovskiy), 1♀ (ZISP). Georgia: “ Swanetia ” (A. Kuschakewitsch coll.), 1♀ (ZISP). Kazakhstan: [Kazakhstan?], “ r.[Riv.] Sarysu, 3.200”, 10.VIII.1961 (Gauzer), 2♀ (ZMUM); Karaganda Prov., Kent Mts SE Karkaralinsk, 21.VII.1962 (L. Arnoldi), 1♀ (ZISP); East Kazakhstan Prov., Tarbagatay, urochishche StaroPyatigorskoe, mountain meadow, 1.VII.1962 (I.M. Kerzhner), 1♀ (ZISP); Almaty Prov., Syugatinskie Mts, urochishche Kokpek [nr Nura Settlm.], 3.VIII.1948 (Bey-Bienko), 1♀ (ZISP). Kyrgyzstan: Issyk-Kul Prov.: “Przheval’ sk” [Karakol] env., “ Semir.[echensk] Prov. ”, [before1922] (A.F. Andreeva), 1♀ (ZISP); Dzhety-oguz, E shore of Issyk-Kul, 7.VII.1930 (M. Berg), 1♀ (ZISP); Dzhergalan Riv., 5.VI.1942 (A. Lubischew), 1♀ (ZISP). Mongolia: Khentiy Prov.,“Syutszukte, yu.-v. Kentey s.-z. Urga ” [Sutszukte, NW of Ulaanbaatar], 1–24.V.1824, 25.IV– 11.V., 11–31.V.1825 (Kozlov), 1J, 2♀ (ZISP).
Diagnosis
Membrane whitish, subtransparent (Figure 1 E). Sensory process of paramere sharply rounded at apex (Figure 3 F). Spicule at dorsolateral lobes of conjunctiva rather sharply curved in middle (Figure 7 G). Posterior pouches of gynatrial sac comparatively long, appreciably longer than wide, with narrowed base (Figure 12 C) or of the same width all over; their bases lying somewhat anterior to posterior margin of gynatrial sac; if bases of pouches lying close to posterior margin of gynatrial sac then inner margins of pouches not looking as prolongations of posterior margin and the latter distinguishable under pouches (Figure 10 A, 10 B).
Variability
Proximal part of spermathecal duct most often as long as or longer then shortened median part; in some specimens, proximal part shorter than median part. In females from Mongolia, posterior pouches slightly shortened.
Remarks
One female corresponding to the diagnosis of the nominotypical subspecies by the whitish membrane, shape, length and position of the posterior pouches was collected south of the general southern boundary of the subspecies natural range, in Transcaucasia (Svanetia).
Measurements
Body 5.55–8.10 in length, 3.45–5.20 in width; pronotum 2.90–6.10 in width; head 1.33–1.71 in width; ocular index 3.52–3.67; length of antennal segments I–V: 0.29–0.38: 0.39–0.64: 0.50–0.71: 0.64–0.98: 0.69–0.99.
Distribution
Central (except the Netherlands), southern (except Portugal and Greece) and eastern (except Hungary and Belarus; in Russia, centre and south of European part) Europe, Great Britain, Sweden, Russian Western and Eastern Siberia, Kazakhstan, and Mongolia (Lis 2006).
description: Published as part of Gapon, Dmitry A., 2018, Morphology of male and female terminalia and taxonomic revision of the burrower bugs genus Canthophorus (Heteroptera: Cydnidae), pp. 183-228 in Annales de la Société entomologique de France (N. S.) (N. S.) 54 (3) on pages 30-31, DOI: 10.1080/00379271.2018.1467788, http://zenodo.org/record/14927017