Fannia quillacingas Grisales & De Carvalho 2019, sp. nov.
Creators: name: Grisales, Diana
name: De Carvalho, Claudio J. B.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Fanniidae
Fannia
Fannia quillacingas

description: Fannia quillacingas sp. nov.
(Figs. 1J, 4 A–C, 5J, 7B, 8J)
Diagnosis. Fore tarsomeres 2–5 flattened and tarsomeres 2–3 laterally expanded; fore tarsomere 1 yellowish, 2–3 partially yellowish; fronto–orbital plate, frontal vitta and facial ridge strongly protruding; gena with a shallow groove under the eye.
Description. Holotype male (Fig. 1J)
Head: eye densely setulose. Frontal vitta velvety black, pollinose. 14–15 fr. 1 pair of oc proclinate and developed, 5 less developed setulae on ocellar area. Fronto–orbital plate silver and pollinose. Parafacial bare, golden pollinose and wide, almost the same width along. Fronto–orbital plate, frontal vitta and facial ridge strongly protruding (Fig. 4A). Fronto–genal suture and gena golden pollinose, gena with a shallow groove under the eye (Fig. 4B). Scape black, 3–4 setae reaching pedicel. Pedicel black pollinose with a dorsal seta developed and surrounded minor setae. Postpedicel black, golden pilose, 2 times the length of pedicel. Arista black, with short pubescence, rays shorter than width of base of arista. Palpus black and slightly spatulate. Prementum long and sclerotized.
Thorax: black. Scutum without vittae. Acr 3:3 not arranged in rows. Dc 2:3, long. Pprn 4, developed. Pra 2. Pro–epimeron setulose.
Wing: brownish, with upper third between C vein and half of R 2+3 vein slightly darker. Calypters brownish with border darker. Haltere brown with stem and base lighter in colour.
Legs: black, pulvilli yellowish and tarsi black, except for first fore tarsomere completely yellowish and tarsomeres 2 and 3 partially yellowish (Fig. 1J). Fore femur setulose on posterior surface with 1 row of long pv and d. Fore tibia with 2 sub–median av and 1 stronger sub–apical; 1 apical p; 1 pd. Fore tarsomere 1 short with a little tuft of short basal v; tarsomeres 2–5 flattened and tarsomeres 2–3 expanded laterally. Mid femur with 2 rows of long pv, strong setae on apical third and 1 row of av. Mid tibia on ventral surface with pronounced constriction on basal half area, densely setulose on apical half, 2 apical v; 3 median a; 1 ad; 4 median p and 1 apical; 1 apical pv. Hind coxa bare setae on posterior margin. Hind femur on ventral surface with a strong almost median protuberance, with tuft of setae and some of them stronger and hook–like, apical half setulose; av, a and ad surfaces with strong sparse setae (Fig. 4C); pv surface setulose. Hind tibia with 1 median d; 2 median ad; 1 a; 2 median av and 1 apical (Fig. 4C); p surface slightly setulose on apical half.
Abdomen: elongate, black, weakly greyish pollinose with developed setae. Sternite 1 densely setulose. Sternite 5 (Fig. 5J). Terminalia (Fig. 7B): epandrium longer as wide with strong and stout setae; cercal plate square apically, fused on the middle portion; surstylus strong and articulated with epandrium, wide apically; bacilliform process spiral.
Female: unknown.
Biology. Label information indicates that this species occurs in southern Ecuador, in the Andes above 2200 m. Nothing else is known about its habits or biology.
Comments. Fannia quillacingas sp. nov. resembles the anthracina species group of Fannia, mainly in the characters of the fore tarsomeres (expanded, flattened and yellowish). This new species differs in the extent of the yellowish color in these tarsomeres and in the lateral expansion.
Etymology. Name in apposition. The species epithet refers to the Quillacingas indigenous people, who once inhabited the higher elevations between Colombia and Ecuador.
Type material. Holotype male (MZSP). ECUADOR: Girón, Portete de/ Tarqui, 2200 –2800 m./ Ecuador, XII. 1970 / L. E. Peña col./ MZSP.
The abdomen is absent in the holotype because it was taken to dissect the terminalia and was fixed under the holotype.
Distribution. Ecuador: Azuay province (Andean region) (Fig. 8J).
description: Published as part of Grisales, Diana & De Carvalho, Claudio J. B., 2019, Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America, pp. 330-360 in Zootaxa 4551 (3) on pages 346-348, DOI: 10.11646/zootaxa.4551.3.4, http://zenodo.org/record/2622947