Ricanula Melichar 1898
Creators: name: Zhang, Huan
name: Wang, Wen-Qian
name: Stroiński, Adam
name: Qin, Dao-Zheng
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Ricaniidae
Ricanula

description: Ricanula Melichar, 1898   
 Ricania (Ricanula) Melichar, 1898: 218.   
 Ricanula Schmidt, 1912: 75.   
 Type species. Ricania noualhieri Melichar, 1898, designated by Schmidt (1912).   
 Remarks. Ricanula can be distinguished from other genera in Ricaniidae by the combination of following characters: costal area of tegmen with sparse and curved transverse veinlets; postero-apical part of tegmen with eyes-spot black cells; tegmen with one line of transverse veinlets; median and posterior parts of tegmen with numerous irregular transverse veinlets.   
 Diagnostic characters. Head. Head including eyes narrower than mesonotum/thorax. Vertex transverse, distinctly wider than long at midline, with all margins well carinate; disc of vertex with or without median carina.   
Frons with all margins well carinated, at upper margin longer than high at midline, widest at the level of lower margin of compound eyes; lateral margins covering base of pedicel, not incised near the level of ocelli. Frontal disc tricarinate, carinae distinctly separated basally, median carina straight; lateral carinae arcuate, almost parallel to lateral margins. Fronto-clypeal suture arched/arcuate.  
Clypeus distinctly narrower than frons, without median carina.  
Compound eyes, with small callus in lower part of posterior margin. Ocelli present. Rostrum-apical segment shorter than subapical one.  
Antenna pedicel elongate, cylindrical, with functional area at the top.  
 Thorax. Pronotum distinctly longer than vertex at midline; disc of pronotum with median carina and two lateral impressions.   
Mesonotum elongate, diamond shape, longer in midlength than wide at lateral angles and longer in midlength longer than combined length of vertex and pronotum at midlength; lateral angles placed before midlength; median carina, lateral and antero-lateral carinae present; median carina and lateral carinae connected basally; median carina reaching scutellum, lateral carinae reaching posterior margin, anterolateral carinae not connected with lateral and not surpassing level of lateral angles of mesonotum.  
 Tegmina membranous, elongately-triangular, flattened, with distinct venation and transverse veinlets. Costal margin weakly arcuate, apical angle broadly rounded, placed distad to claval angle, posterior margin weakly sinuate, postclaval margin (tornus) absent. Costal area with transverse veinlets and oblique shallowly incisions; postero-apical part with eyes-spot black cells. Longitudinal veins ScP+RA, MP and CuA leaving basal cell separated. CuA vein first fork placed before the connection of claval veins Pcu+A 1. Tegmen with single apical line of transverse veinlets, median and posterior part of tegmen with numerous irregular transverse veinlets.   
Hind wing with precostal cell present; ScRA and MP forking distinctly after midlength of wing, CuA forking distinctly before half of wing.  
Hing legs. Metatibia with 2 lateral spines; apically with 6 well developed spines, external lateral spines bigger than internal lateral spines; basitarsomere of metatarsus a little longer than cumulative length of second and apical tarsomeres.  
 Male terminalia. Anal tube with ventral margin strongly concaved in lateral view. Pygofer higher than wide; dorsal part narrower than ventral one. Genital styles broadly triangular (dorsal and ventral margins nearly parallel in R. fujianensis and R. cacaonis), with sharp spine-like process at the end of dorsal margin.   
Periandrium with or without processes, with long lateral split surpassing the half of its length; dorsal periandrium a bit shorter than ventral one. Aedeagus long and narrow, apically with 1–2 pairs of symmetrical, well sclerotized, spinose processes.  
 Female terminalia. Pregenital sternite with well-developed lateral lobes, median portion of the posterior margin with or without processes. Anal tube not surpassing half of upper margin of the gonoplac.   
Gonoplac well developed, laterally flattened; posterior margin of the gonoplac with 2–3 rows of small teeth; membranous parts of gonoplac well developed, placed medially on ventral margin.  
Gonapophysis VIII sabre-like, v-shape in cross section, with teeth at dorsal margin; endogonocoxal process with spiniferous microsculptures, well sclerotized medially; lateral parts membranous, reaching apex of gonapophysis VIII.  
Gonaphophyses IX and gonospiculum bridge well developed.  
Bursa copulatrix with two isometric pouches: first pouch with well visible cell and sclerotized ornamentation; second pouch with numerous pores.
description: Published as part of Zhang, Huan, Wang, Wen-Qian, Stroiński, Adam & Qin, Dao-Zheng, 2021, Two new species and a new combination in the genus Ricanula Melichar, 1898 for Ricaniidae from China (Hemiptera: Fulgoromorpha), pp. 353-369 in Zootaxa 5047 (3) on pages 354-355, DOI: 10.11646/zootaxa.5047.3.7, http://zenodo.org/record/5540953