Odontocheila iodopleura Bates 1872
Creators: name: Moravec, Jiří
name: Brzoska, David
name: Huber, Ronald
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Odontocheila
Odontocheila iodopleura

description: Odontocheila iodopleura Bates, 1872
(Figs 72–121, 208).
Odontocheila iodopleura Bates, 1872: 285.
Type locality. Nicaragua: Chontales.
Odontochila iodopleura: Fleutiaux 1892: 125; Rivalier 1969: 208, fig. 7i, 209. Odontocheila iodopleura: Wiesner 1992: 79.
Type material. Lectotype (designated here) ♂ in BMNH labelled: “Chontales / Nicaragua / T. Belt” [printed] // “B.C.A. Coll., I(1). / Odontocheila / iodopleura, Bates ” [printed/handwritten] // “Lectotype / Cicindela / iodopleura Bates, 1872 / design. Jiří Moravec 2012” [red, printed]. Paralectotypes. 1 ♂, 1 ♀ in BMNH: with same two first labels. 1 ♀ in BMNH: “Type” [circular, red frame, printed] // “ Odontochila / iodopleura / Bates” [handwritten] // “B.C.A. Coll., I(1).” [printed] // “Lectotype [sic!] / Odontocheila / iodopleura Bts / by Ervin ‘76” [printed/ handwritten invalid label]. 2 ♀♀ in MNHN: “Chontales / Nicaragua” [handwritten] // “H.W.Bates / Biol Cent.Amer” [with thin black frame, printed] // “Muséum Paris, 1952 / Coll. R. Oberthür” [printed] // “Paratype” [sic!, red, printed] // “ iodopleura / Bates” [in only one of the females, handwritten].1 ♀ in MNHN: “Chontales” [handwritten] // “Muséum Paris, 1952 / Coll. R. Oberthür” [printed] // “Paratype” [sic!, red, printed]. 1 ♀ in SDEI: “Chontales / Nicaragua / T. Belt” [printed] // “Type! / coll. W. Horn” [printed] // “British Mus. / Godm. Salv. C.”[printed/handwritten] // “Syntypus” [red, printed] // “Coll. W. Horn / DEI Eberswalde” [printed] // “ iodopleura Bat. ” [greenish with black frame] // “ Odontocheila iodopleura / Bates type (DEI- Eberswalde) / borrowed by D. L. Pearson / 23.Oct.1996 (drawer # 59)” [printed]. All paralectotypes labelled: “Revision Jiří Moravec 2011 [2012 respectively]: / Paralectotype / Odontocheila / iodopleura Bates, 1872 ” [red, printed].
Other material examined. Historical data. 1 ♀ in MNHN: “ Muséum Paris / Coll. Chaudoir 1874”. Other data. 1 ♀ in BMNH: “ Nicaragua ”. 1 ♀ in BMNH: “ Rio Grande ” [illegible]. 1 ♀ in BMNH: “ Costa Rica ”. 1 ♂, 3 ♀♀ in BMNH: “ Collection / Schild-Bougdorf / Costa Rica / San Carlos ”. 1 ♂ in SDEI, 1 ♂, 1 ♀ in NMPC: “ Turrialba, Costa Rica / 900 m, IV.”. 1 ♂ in SDEI: “ Cascajal / Costa Rica ”. 1 ♂ in SDEI: “ Cartago Panel”. 1 ♂ in MFNB, 3 ♂♂, 3 ♀♀ in SDEI: “ Costa Rica / San Carlos ”. Recent data. 1 ♂, 2 ♀♀ in RLHC: “ Nicaragua: Rio San Juan / Dept. 8km SE El Castillo / Refugio Bartola, 30m, rn for / N 10°56.6’, W 84°20.4’ / 25–31.V.2002 S. Peck #09”. 2 ♂♂, 1 ♀ in CDCL: “ Nicaragua Rio san Juan / Bartola Lodge / 6–13.VI.2005 / Alaine et Sylvaine Audreau leg.”. 3 ♂♂, 1 ♀: in DBCN: “Nicaragua-R. S. Juan Prov. / Bartola Res. San Juan R./ 10°58.4´N; 84°20.3´W / D. Brzoska 25–30- V-2002 ”. 1 ♂ in CDCL, 1 ♂ in CCJM: “ Nicaragua / Neuva Segovia / Cerro Jesus / 1300 m / 4.V.2013 / C. Dheurle leg.”. 1 ♂ in CMNH: “ Nicaragua: Matagalpa / Dpt. Selva Negra / 13-VI-2002 / F.W. Scillman Jr. ”. 7 ♂♂, 4 ♂♂ in DBCN: ibid., except for: “Selva Negra Bio. Res. / km 140— Carretera Jinnot ega / 12°60.0´N; 85°54.5´W / D. Brzoska 18-22-V-2002 ”. 21 ♂♂, 7 ♂♂ in CJVB, 5 ♂♂ in CCJM: “ Nicaragua / Monte Selva de Negra / Trail Bavaria 1300m / 15.VI.2013 leg. Jan Vybíral. 9 ♂♂, 3 ♀♀ in DBCN, 1 ♂ in INBIO: “ Costa Rica ” Cartago / HY 32— 9 km NE Zurqui Tunnel / D. Brzoska 17-V-1993 ”. 1 ♂ in CCJM: “ Costa Rica: Cartago Prov. / Rancho Naturalista (SE La Suiza) / 0.9°49.9´N, 83°32.8´W / 21.V.2013 leg. M. Mýlek ”. 1 ♀: in INBIO: “ Grano de Oro, Chirripó, Turrialba Prov. / Cartago, Costa Rica, 1120m, VI.1993 / P. Campos, LN 200250 -595900 # 2253”. 2 ♂♂ in CCJM: “ Costa Rica, Cartago Prov. / Las Brisas Res. n. Turrialba / 10°04´N;83°37.8´W, 915m / 23.V.2013, leg. Jiří Moravec“. 3 ♂♂ in CCJM: ibid., except for: “leg. M. Mýlek”. 1 ♂ in CJVB: “ Costa Rica / Turrialba / 20.VI.2008 leg. Ondřej Vybíral. l ♂ in INBIO: “ Río San Lorenzo, 1550 m / Tierra S Morenas, ZP. / Tenorio, Prov. Guanacaste / Costa Rica, M Segura / 28.Mar. a 21 Abr. 1992 L-N-287800, 427600”. l ♂ in INBIO: “ Río San Lorenzo, 1050 m, Tierras / Morenas, ZP. / Tenorio, Prov. / Guanacaste, Costa Rica, Abr. 1991, C. / Alvarado, L-N- 287800”. l ♂ in INBIO: “ R. San Lorenzo, 1500 m, / R. F. Cord. Guanacaste / (Tenorio), Prov. Guanacaste / Costa Rica C. / Alvarado, Jun 1991 / L-N-287800, 427600”. 1 ♂ in CCJM: “ Costa Rica, PN Guanacaste, / NW Hang Cerro Cacao / 900–1000m, submontaner bergwald / Weg. m. krautiger Veg. / 01.IX.1993, M. Franzen leg.”. 2 ♂♂ in CADW: “CR3- Costa Rica, Guanacaste / Est. Biol. Santa Rosa, 310m / Santa Rosa NP, 21.–29.VI.2004 / 10°50´22.3´´N; WO 85°37´04.8´´ / leg. Dostal & Uhler ”. 2 ♂♂, 2 ♀♀ in CCJM: “ Costa Rica—Guanacaste / P.N.G. Cacao Biol. Station / D. Brzoska 25-V- 19963”. l ♂ in INBIO: “ Costa Rica, Prov. Alajuela, San / Carlos, Boca Tapada, Bosque / Ancianos, 50–100 m, 23 May–25 / Jun 2004, B. Hernádez, Tp. Malaise / L-N-298721-516041 #77474. l ♂ in INBIO: “ Costa Rica, Prov. Alajuela, Upala / Alb. Heliconias, Send Heliconias / 700 m, 17–21 Jun 2000, D. Briceffo / Manual (red libre) / L-N-299100-422600 #56784”. 1 ♀ in DBCB: “ Costa Rica—Alajuela / R.F. San Ramon, 900 m., Rio San Lorencito / D. Brzoska 22-V-1996 ”. 1 ♀ in RLHC: “ Costa Rica: Alajuela / Reserva Forestal San Ramón / Rio San Lorencito and tribs / 10.216N, 84.607W / 13–16.vi.1988 el. 980m / C.M. & O.S. Flint, R. Holzenthal ”. l ♂ in INBIO: “ Monumento Nacional Guyabo, A.C. / Amistad, Prov. Cartago, Costa Rica, 1100 m / Jun 1994, G. Forseca / L-N-217400-570000 #2989”. l ♂ in INBIO: “ Costa Rica, Prov. Heredia, Est. Biol. / La Selva 50–150 m, 18 May 1999 / D. Brenes Manual / L-N-268152-535584 #63363”. l ♂ in FCCR: “ Costa Rica: Heredia Prov. / Finca La Selva / 1. 2 mi S Puerto Viejo ” // “ D.R. & M.L. Paulson / 4.IX.1967 ”. l ♂ in CDCL: “ Costa Rica: Heredia Prov. / La Selva Biol. Sta. / 3 km S Pto. Viejo / 10°26´N, 84°01´W / 8.IV.1989 H. A. Hespenheide ”. 1 ♀ in DBCN: “ Costa Rica—Heredia / 3 km N. Puerto Viejo / 10°28.4´N, 84°00.9´W / David Brzoska 22-V-1999 ”. 2 ♂♂, 1 ♀ in CMNH: “ Costa Rica: Heredia Prov. / Pto. Viejo / Finca La Selva / 18–16-VII- 1973 / D.C. Rentz, KR. Brodey ” // “ Howard Boyd / Collection / Acc. 35,563”. 1 ♀ in CMNH: “ Costa Rica / Heredia Prov. / Rio Frio / Standard Fruit Co. / 10°2´N, 83°53´W / 10–17.V.1970 / Michael J. Corn”. 1 ♂ in RLHC: “ Costa Rica, Heredia Pr. / Puerto Viejo, Sarapiqui / VIII-1-65 / E. Ordway coll.”. 1 ♂ in RLHC: “ Costa Rica, Heredia / Chilamaté, 75m / IV-VI-1990 / Hanson & Godoy ”. 2 ♀♀ in USNM (ex INBIO): “ Costa Rica: Prov. Heredia: / 11km SE La Virgen 450 – / 550m, 10°20´N, 84°04´W / 20 Abri. 2003 / INBio-OET_ ALAS transect”. 1 ♂, 1 ♀: in USNM (ex INBIO): ibid., except for 8 Abri 2003”. 1 ♂ in USNM (ex INBIO): ibid., except for: “ 23 Marzo 2003 ”. 1 ♂ in USNM (ex INBIO): ibid., except for: “17 Abri 2003”. 20 spms (♂ ♀) in USNM (ex INBIO): ibid., except for: “ 11 km ESE La Virgen 250 – / 350m, 10°21´N, 84°03´W / and dated: “06, 18, or 21 Abri 2004”. 1 ♂, 2 ♀♀ in USNM (ex INBIO): “ Costa Rica: Prov. Heredia: Est. Biol. La Selva, 50 – / 150 m, 10°26´N, 84°01´W ”. / 21 Abri 2003 / INBio-OET_ ALAS transect”. 1 ♂ in USNM (ex INBIO): ibid., except for: “11 Abri 2003”. 1 ♂ in USNM (ex INBIO): ibid., except for: “ 09 Mar.2004 ”. 3 ♂♂, 1 ♀: in CCJM: “ Costa Rica, Limon Prov. / Veragua Rain Forest Res. / 9°55.5´N;83°11.5´W, 440m / 24.V.2013, leg. Jiří Moravec“. 1 ♀ in RLHC: “ Costa Rica, Limon / 16km W Guápiles / 400m, V-VI-1990 / col. Paul Hanson ”. 1 ♂, 1 ♀ in CCJM: “ Costa Rica / Cariari vic. / Finca La Suerte / VI.2001 leg. Van den Berghe “. 21 ♂♂, 7 ♀♀ in CJVB, 6 ♂♂ in CCJM: “ Costa Rica / Guácimo Earth [University area] / 16–19.VIII.2008 / Ondřej Vybíral lgt.”. 3 ♀♀ in RLHC: “ Costa Rica, San Jose / P.N. Braulio Carrillo / 9.5 km E tunnel, 1000m / VI-1989, col. Hanson ”. 1 ♀ in RLHC: “ Honduras, Sta. Barbara / 7 mi S El Mochito / VII.”. 1 ♀ in RLHC: “ Honduras / Prov. Atlantida / Lancetilla Bot. Gard. / 6 km SW Tela / 30.V.1987 P. Johnson ”.
Differential diagnosis. Because of very similar aedeagi, O. iodopleura is undoubtedly related to O. mexicana and O. potosiana sp. nov. However, it is probably most closely related to O. tawahka Johnson, 1996 known only from Honduras, which, apart from the lookalike aedeagus shape, also has very similar white elytral maculation and pattern of dense elytral punctation of irregularly anastomosing punctures. Particularly some predominantly greener or blue adults of O. iodopleura (see the “Variability” above) may resemble O. tawahka.
Most of the adults of O. iodopleura have either dark copper elytral disc with reddish sublateral area and iridescent-green-blue longitudinal lateral area (apart from the always present violaceous juxtaepipleural stripe), or are conspicuously multicoloured, or with prevailing iridescent green coloration. Only very rarely do they possess almost uniformly green or violaceous-blue elytra (Fig. 99) resembling those in O. tawahka, but the labrum, mandibles, palpi and legs are in O. tawahka generally paler.
O. ignita and O. exilis differ in having moderately, but notably elongate pronotum, almost uniformly reddishcupreous elytra with indistinct greenish lateral stripe, even denser elytral punctation, and very different shape of their aedeagi.
O. potosiana sp. nov., known only from Mexico, has its pronotal disc markedly narrower, with much finer surface sculpture and, moreover, with bright, iridescent reddish-cupreous large median area, elytra bright reddishcupreous on the elytral disc (similar to that in O. mexicana), larger whitish elytral maculae and mostly isolated punctures which are anastomosing only rarely and only on elytral disc.
O. mexicana can be immediately distinguished by its almost smooth and polished surface of the almost globose pronotal disc.
Redescription. Body (Figs 72–76) notably variably coloured and of very variable size, 8.20–10.6 (LT 9.30) mm long, 2.70–3.20 (LT 2.90) mm wide, females mostly distinctly larger than males.
Head (Figs 77–78) with notably pronounced eyes, very large, as wide as body or slightly narrower, rarely wider than body, width 2.70–3.25 mm; all head portions glabrous.
Frons and vertex sculptured and coloured as in O. mexicana, but the rugae on vertex much finer and denser, and coloration much more variable, from reddish-cuprous with green, blue or cyaneous lustre also notably violaceous.
Clypeus and genae as in O. mexicana.
Labrum 4-setose; distinctly bicoloured; male labrum (Figs 79–83) with shape of lateral and anterolateral teeth basically as in O. mexicana, but notably longer and anterior margin with always present median tooth, length 0.75– 0.90 mm, width 1.10–1.30 mm, female labrum (Figs 84–85) 1.10–1.30 mm long, 1.25–1.35 mm wide, with prominent, acutely tridentate median lobe with protruding median tooth.
Mandibles (Figs 77–78) black-brown in male, almost black in female, with distinct, ivory-yellow lateral stripe (more extended in male), normally shaped as in O. mexicana, but sometimes the apex of the right terminal tooth in male is very indistinctly bent inward (Fig. 78).
Palpi (Figs 77–78) with normal (elongate) shape of terminal palpomeres, ivory-yellow to ochre-testaceous, terminal palpomeres in both labial and maxillary palpi metallic black; penultimate palpomeres of maxillary palpi variably yellow-ochre to brownish-darkened, or black; penultimate (longest) palpomeres of labial palpi in females sometimes blackened on their smooth area.
Antennae (Figs 72–78) rather long, in male slightly surpassing elytral half, in female shorter, reaching elytral third; scape with only apical seta, metallic black with strong, blue or green lustre; pedicel mostly concolorous with scape; antennomeres 3–4 (Figs 77–78) almost uniformly metallic-black, usually with violaceous reflections, but often also with purple apical areas, with indistinct, sparse microsetae; antennomeres 5–11 smoky black with usual micropubescence.
Thorax. Pronotum (Figs 86–89) glabrous, variably coloured, the coloration mostly confined to that on elytra: either reddish-cupreous on sublateral areas with usually rather matt metallic-green median area of disc and iridescent green-blue lateral areas, or prevailingly iridescent-green with blue or violaceous lateral areas, as long as wide or slightly wider length 1.65–2.10 mm, width 1.70–2.10 mm; sulci well pronounced; anterior lobe only slightly wider than posterior lobe, shiny reddish-cupreous or metallic green or blue with reddish reflections; surface of anterior lobe rather coarsely, mostly transversely wavy to vermicular but very irregularly rugulose; disc subglobose with distinctly convex lateral margins of dorsally visible proepisterna, while thin, dorsally visible notopleural sutures are mutually less convex; medial line clearly marked; whole discal surface up to the notopleural sutures covered with rather fine rugae which are usually coarser, more stria-like and subparallel along the median line, rugae on lateral areas towards notopleural sutures are coarser and mostly transverse; posterior lobe with distinct basal rim and distinctly raised dorsolateral bulges, variably coloured, its surface coarsely irregularly rugulose; all ventral and lateral sterna glabrous, their surface and coloration as in O. mexicana and other species of this complex.
Elytra (Figs 90–104) elongate, length 5.20–6.70 mm, with rounded or subquadrate humeri, usually more distinctly subquadrate in female, lateral margins as in O. mexicana; dorsal elytral surface moderately convex, particularly on posterior half of elytral disc, discal impression shallow or more distinct and together with rather distinct humeral impression clearly delimiting moderate basodiscal convexity; anteapical-apical impression moderate; whole elytral surface distinctly and notably densely and irregularly punctate: punctures larger and isolated only on antero lateral area with several larger, irregularly placed punctures within the humeral impressions; punctures commonly anastomosing on elytral disc, their thin intervals forming on the large elytral discal area almost cristulate sculpture (apart from different appearance of the sculpture depending on angle of illumination); elytral surface glabrous except for few, but usually long, several white hairlike sensory setae scattered mostly on basal area, and a few others adjacent to epipleura; elytral coloration extremely variable, either dark copper on elytral disc, with narrow, iridescent reddish-cupreous or gold-bronze sublateral area and narrow blue-green lateral stripe, and always present deep violaceous or purple-violet juxtaepipleural area, or the elytra are indistinctly or conspicuously multicoloured, with dull or bright green elytral disc and partly reddish juxtasutural area and with the iridescent reddish-cupreous or gold-bronze sublateral area and blue-green lateral area, or with prevailingly green coloration, rarely black-violet on elytral disc and blue-green on remaining elytral area (except for the always present violaceous juxtaepipleural stripe); whitish elytral maculation in both sexes consisting of three maculae: humeral macula in male distinct, and at least partly visible from above, in female reduced and barely visible from above or only partly as a thin falciform stripe; lateromedian macula mostly smaller than in preceding species, irregularly rounded or triangular; anteapical macula comparatively small somewhat elongate along the margin.
Abdomen basically as in O. mexicana.
Legs as in O. mexicana, but trochanters in female, particularly metatrochanters, sometimes darkened or blackened, faded in old specimens.
Aedeagus (Figs 105–120) 3.30–3.45 mm long, 0.90–1.00 mm wide, shape basically as in O. mexicana, its apex only slightly variable in shape; internal sac (Fig. 107) as in O. mexicana and other species of this species-complex (and generally as in all species of the genus).
Variability. Apart from the usual purple-violet juxtaepipleural stripe which is common for all species of this Odontocheila species-complex, the lectotype of O. iodopleura (Figs 72, 90) and other syntypes from Nicaragua and adults of some other populations both from Nicaragua and Costa Rica, possess elytra with sublateral iridescentgreen-blue longitudinal stripe (passing to gold-bronze) while the lateral areas of the mostly black-copper elytral disc are almost uniformly reddish-cupreous. In contrast, some other populations, mostly from Costa Rica, Zurqui Tunnel (Figs 76, 98, 104) but also some adults from Costa Rica, Hang Cerro Cacao (Figs 75, 93) and in Nicaragua, Cerro Jesus (Fig. 96), as well as the only two females from Honduras, are notably multicoloured and much greener, having wide, bright or cyaneous-green or blue-green elytral disc, often delineated by only narrow longitudinal juxtasutural cupreous area and laterally by sublateral gold-bronze to bronze-cupreous stripe passing to blue-green lateral stripe. Nevertheless, while such notably greener adults, particularly the prevailingly greenish population from Costa Rican Zurqui Tunnel, may indicate a different species, some adults from the Costa Rican province of Guanacaste and from Montes Selva de Negra in Nicaragua possess similar elytra with transiting coloration. Moreover, some syntopic adults of the population from La Virgen of the Costa Rican province of Heredia (mostly deposited in INBIO and USNM) possess a tremendous variability in their elytral coloration (Figs 91–92, 101–102): some of them possess the predominantly reddish-cupreous coloration as in the type specimens, while others are variably multicoloured. In some populations with more uniformly reddish elytral disc, some intermediate elytral colorations also occur, although the discal area is mostly dull-coloured instead of bright green. The populations from the area of La Suiza and Las Brisas of the Costa Rican province of Cartago possesses multicoloured and prevailingly green elytra as in those from the Zurqui Tunnel, but some of them have intermediate coloration (Fig. 95). Rarely (some adults from the Costa Rican province of Alajuela) have their elytra almost uniformly violaceousblue (Fig. 99).
Biology and distribution (Map Fig 208). O. iodopleura occurs in Nicaragua and in the large part of Costa Rica, in the provinces of Guanacaste, Alajuela, Heredia, Limón, Cartago and San José. Within this revision, no specimen was found from the southern province of Puntarenas where it is replaced by O. exilis (see under that species below), and in collections both species were rarely confused.
Only two examined specimens (females) come from Honduras. The Lancetilla Botanical Garden near the town of Tela (province of Atlantida) lies in northwestern Caribbean coast, while El Mochito (Santa Barbara province) is situated in somewhat inner area of western Honduras. Both localities are a long way from the Nicaraguan localities of O. iodopleura, and also from the type locality of O. tawahka in north-eastern corner of Honduras.
As with the majority of species of this genus, adults are found along partial shaded forest paths and open areas.
Remarks. The original description of O. iodopleura by Bates (1872) was clearly based on more specimens of both sexes, and several syntypes are deposited in BMNH, MNHN and SDEI. As none of them was explicitly designated as “Type” in the original description or by subsequent authors including Rivalier (1969), the only male found in BMNH is here designated as the lectotype to assure the stability of the taxon, also because all other syntypes (paralectotypes) in the above mentioned collections are females. One male syntype (MNHN) labelled as “Type”, which was examined years ago by the third author and obviously returned to MNHN (David L. Pearson pers. com.), is no longer deposited in the “Collection Générale” of MNHN, only a label that the type was borrowed by D. L. Pearson in November 1993 is there in the box, and the recent placement of this male syntype in the MNHN collection is unknown. The male lectotype comes from the same series as the other syntypes, also bearing the labels in the accordance with the original description by Bates (1872). The label “Lectotype” attached previously by Erwin to the female syntype in BMNH (see Type Material above) is invalid, as the type designation never was published (Terry Erwin pers. com.); moreover, this female syntype has no original det label—its label “ Odontochila / iodopleura / Bates” was evidently attached additionally (as Bates spelled the genus-group name as Odontocheila).
Odontochila amoena (originally as “ amaena ”), listed without a description by Chaudoir (1865) is an unavailable name (nomen nudum). It was considered to be close to O. iodopleura by Horn (1915) and listed as a synonym of O. iodopleura by Wiesner (1992). There is one female in MNHN, arranged by Rivalier under the name “ Odontochila amaena ” and labelled “ Type ”, and the examination by the first author of this paper has confirmed that the female is conspecific with O. iodopleura. However, the “ type ” locality of this female is Colombia, and the locality label was probably attached to this historical specimen erroneously, because the occurrence of O. iodopleura in Colombia has not been confirmed within this revision, and is almost certainly impossible (see the map of distribution Fig. 208).
Such enormous variability in coloration (stressed in the Variability” above) is in tiger beetles very common, but only rarely occurs in other presently revised species of Odontocheila.
Nevertheless, O. iodopleura is considered here a variably coloured species, and as no other diagnostic differences have been found during this revision, it prevents us from a description of the multicoloured to prevailingly green populations as a new species.
description: Published as part of Moravec, Jiří, Brzoska, David & Huber, Ronald, 2017, Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense — 18. Six Mexican and Central American species related to Odontocheila mexicana Laporte de Castelnau and O. ignita Chaudoir, with a description of O. potosiana sp. nov., pp. 451-499 in Zootaxa 4231 (4) on pages 471-480, DOI: 10.11646/zootaxa.4231.4.1, http://zenodo.org/record/292697