Phaloria tukul Tan & Rahmadi & Robillard 2021, sp. nov.
Creators: name: Tan, Ming Kai
name: Rahmadi, Cahyo
name: Robillard, Tony
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Orthoptera
Haglotettigoniidae
Phaloria
Phaloria tukul

description: Phaloria tukul Tan & Robillard, sp. nov.
(Figs. 1C, 2C, 3C, 4C, 5E, 5F, 6D)
Material examined. Holotype (male, LEN2014 -TR144), New Guinea, Indonesia, West Papua, Lobo, S3.706417, E134.072139 (LOBO4), 189 m. a.s.l., forêt proche PK8/ route Lobo-Kaimana camp à 200 m, forêt primaire [forest near PK8 / Lobo-Kaimana camp road 200 m away, primary forest], 21–30.x.2014, nuit [night], sur plante basse [on low plant], molecular sample P3, T. Robillard (MZB.Orth.21840).
Type locality. New Guinea, Indonesia, West Papua, Kaimana Regency, Lobo
Etymology. The species name refers to the hammer-shaped pseudepiphallic parameres, unique among congeners; tukul = hammer in Bahasa Malay.
Diagnosis. This new species is unique among congeners by male genitalia: pseudepiphallic parameres inverted hammer-shaped, with a rectangular lobe before middle along inner margin (resembling the head of a hammer), with apical half slender and produced with small apical lobule along inner margin (resembling the handle of the hammer).
This new species is most similar to P. beybienkoi from Supiori Island (north of West Papua) in habitus and genitalia (particularly pseudepiphallic parameres), but differs by pseudepiphallic lobes not enlarged at the apex and pseudepiphallic parameres with outer apical process not acute apically. This species is also similar to P. similis and P. parasimilis from New Guinea by pseudepiphallic lobes and pseudepiphallic parameres with rectangular lobe, but differs by long apical process of pseudepiphallic parameres.
Subgeneric status. This species should belong to the subgenus Papuloria based on the pseudepiphallus with a pair of lateral lophi and endophallic sclerite without large unpaired apodeme directed anteriorly.
Description. Very similar to P. dischidia sp. nov. from Lobo in habitus and size (Fig. 1C). Head dorsum yellow brown with 5 very faint brown longitudinal bands at posterior end; vertex with a T-shaped brown pattern (Fig. 2C). Fastigium darker brown. Scapes yellow brown. Antennae yellow brown. Fastigium verticis and frons dark brown; clypeus and mouthparts yellow brown (Fig. 3C). Maxillary palpi pale coloured, apices of segments slightly brownish. Lateral part of head yellow brown without darker patterns. Pronotal disk unicolourous brown with distinctly darkened posterior and anterior margins; posterior margin straight (Fig. 2C). Lateral lobes also generally brown with dark margins. Legs generally unicolourous yellow brown. Inner and outer tympana similar in size and shape: relatively small and oval. FIIs with a faint incomplete ring near knees. TIs with very faint brown rings. FIIIs yellow brown without bands or spots, slightly darker in colouration near knees, knees brown. TIIIs brown with dark spots near bases of spurs. Tergites brown.
Male. FWs 1.7 times longer than wide, surpassing abdominal apex, mostly yellow brown and some parts hyalinous (Fig. 4C). FW venation typical of genus, 8 fairly straight veins in harp; mirror 0.8 times as long as wide, very large and transverse, separated by two dividing veins: basal one mostly straight, distal one obliquely sinuous. Apical field short, 0.9 times shorter than length of mirror. Lateral field with R and M diverging posteriorly, with 6 faint cross-veins. Hind wings reaching apex of FW, not exceeding FWs.
Male genitalia (Figs. 5E, 5F). Pseudepiphallus with lateral margins slightly converging posteriorly; with posterior margin roundly, deeply and broadly incised in the middle. Pseudepiphallic lobe elongated, tapering into a slender process that bends inwards slightly, apex obtuse. Rami short, only slightly longer than half of pseudepiphallus length, not bent apically. Pseudepiphallic parameres sclerotized, inverted hammer-shaped, basal margin truncated (resembling the mallet of a hammer), external margin concave, inner margin with a rectangular lobe before middle (resembling the head of the hammer); apical half slender (resembling the handle of the hammer), with a small apical lobule along inner margin, apical end tapering into an obtuse apex. Ectophallic apodemes fork-shaped; with apical inner and outer arms long and somewhat pointing posteriorly and obliquely. Ectophallic fold with a pair of obtuse lobes at posterior margin.
Female. Unknown.
Measurements (in mm). Male holotype PronL = 2.2, PronW = 3.8, FWL = 9.7, FWW = 5.7, FIIIL = 9.3, FIIIW = 2.6, TIIIL = 9.2.
description: Published as part of Tan, Ming Kai, Rahmadi, Cahyo & Robillard, Tony, 2021, New species of Phaloria (Orthoptera: Phalangopsidae: Phaloriinae) from West Papua (Indonesia), pp. 513-530 in Zootaxa 4985 (4) on pages 517-518, DOI: 10.11646/zootaxa.4985.4.5, http://zenodo.org/record/4964249