Goeldia nigra
Creators: name: Almeida-Silva, Lina M.
name: Brescovit, Antonio D.
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Titanoecidae
Goeldia
Goeldia nigra

description: Goeldia nigra (Mello-Leitão, 1917)
(Figs 1B–D, 16, 26)
Aebutina nigra Mello-Leitão, 1917: 77, figs 22–23 (holotype female from Pinheiro (currently: Piraí, Pinheiral) (22°30’46”S 44°00’03”W), Rio de Janeiro, Brazil, C. Devoto coll., deposited at MNRJ 1358, examined before the fire 2018).
Goeldia nigra: Lehtinen 1967: 209, 236; Silva-Moreira et al. 2010: 31; World Spider Catalog 2020.
Additional Material Examined. BRAZIL. São Paulo: São Paulo (23°32’51”S 46°38’10”W), 1 ♂, VI.1980 (IBSP 2168); 2♂, 1♀, no further data (IBSP 5935, 10001); Instituto de Biociências, Campus da USP, 1♀ 31.VIII.1999, M.A. Vilalva coll. (IBSP 24050); 1♂ 6.XII.2002, A.B. Bonaldo coll. (IBSP 37321); 1♂, VI.2010, F. Yamamoto coll. (IBSP 171107, Fig. 1D); Campus of Instituto Butantan (23°33’50”S 46°43’16”W), 1♂, 12.VI.2007, E.S.S. Álvares coll. (IBSP 81158); Parque Ecológico da Guarapiranga, Ilha dos Eucaliptos (23°44’1”S 46°44’0”W), 1♀, 9–15.IX.1999. R.P. Indicatti coll. (IBSP 131821); Itapevi (23°32’ 57”S, 46°56’14”W), 1♂, 4.IV.1995, C.L.A. Santos coll. (IBSP 6200); Mairiporã (23°19’08”S 46°35’13”W), 1♂, 14/IV/1996, C. Berger coll. (IBSP 11866); same data, 1♀, 26.VII.1996, C. Berger coll. (IBSP 11844); Louveira (23°05’11”S 46°57’02”W), 1♂, 16.I.1996, A.E.C. Farinha coll. (IBSP 12066); Mogi das Cruzes, Parque Natural Municipal Serra do Itapety (23°29’S 46°12’W), 4♂, 2003, Equipe Biota coll. (IBSP 54702, 54591, 54599); Socorro, Pousada da Gruta (22°35’45”S 46°31’01”W), 2♀, 19–27.XII.2010, R. Indicatti coll. (IBSP 171108, Fig. 1B–C); Águas da Prata (21°56’12”S 46°43’00”W), 1♂, 6.IX.1999, V.C. Onófrio coll. (IBSP 24164); Biritiba Mirim (23°34’20”S 46°02’15”W), Barragem do Rio Biritiba, 1♂, 2003, Equipe Biota coll. (IBSP 119423); Jandira (23°31’40”S 46°54’10”W), 1♂, 28.II.2005, Zoonoses Jandira coll. (IBSP 50948). Paraná: Guaraqueçaba (25°18’25”S 48°19’44”W), 1♀, 27.IV.1995, M. Fischer coll. (MCN 27775); Tabagi, Parque Estadual do Guartelá (24°34’S 50°14’W), 4♀, 20–27.I.2008, I. Cizauskas coll. (IBSP 92153); Antonina (25°25’44”S, 48°42’43”W), 1♀, 20–23.IX.2006, E.S.S. Álvares coll. (IBSP 68541); Cruzeiro do Iguaçu, Foz do Chopim (25°33’44”S 53°6’19”W), 1♂, 8–15.X.1998, Equipe Biota coll. (IBSP 21320). Santa Catarina: Piratuba (27°25’12”S 51°46’19”W), 1♀, 1.I.1990, G. A. Martinazzo coll. (MCN 19592). Rio Grande do Sul: São Borja, Batalhão Mecanizado (28°20’00”– 29°04’34”S 55°18’25”– 56°19’38”W), 1♀, 7♂, X–XI/2006 & IX/2007, R. Camargo coll. (IBSP 79481–79485). ARGENTINA. Corrientes: San Cayetano, Estación Biológica de Corrientes (27°32’44.8”S 58°40’52.8”W), elev. 50 m, 2♂, 7♀ 4 juv, 5–10.XI.2007, C Grismado & I Martines coll. (MACN 13331, 13457, 13483, MACN 13538). Jujuy: Yuto (23°38’00”S 64°28’10”W), 1♀, XI.1966, M.E. Galiano coll. (MACN 16212); Fraile Pintado (23°56’00”S 64°47’00”W), 1♀, 1 juv., X.1967, E. Maury coll. (MACN 16092). Salta: no further data, 1♂ (MACN 2320); Santa Maria (22°08’00”S 62°49’00”W), 1♀, VII.1947 (MACN 2265); Aguas Blancas (22°44’10”S 64°21’14”W), 2♀, XI.1967, M.E. Galiano coll. (MACN 16109); Urundel (23°33’00”S 64°25’00”W), 1♀, VIII.1947, Orifila coll. (MACN 2284). Formosa: Laguna Blanca, Parque Nacional Pilcomayo (25°03’52”S 58°08’12”W), 1♀, XI.1990, M. Ramírez coll. (MACN 16023). Misiones: Misiones, Parque Nacional Iguazu (25°41’01”S 54°27’14”W), 1♀, 1 juv., VIII.1985, M. Ramírez coll. (MACN 16053); Barrio 17 de Octubre, 1♀; 14.X.1953, R. Schiapelli & J.A. de Carlo coll. (MACN 16051); Parque Nacional Iguazu (25°41’01”S 54°27’14”W), Ruta 101, 1♂, 1 juv., 22–30.VIII.1986, M. Ramírez coll.(MACN 16045); Puerto Libertad (25°55’00”S 54°36’00”W), 2♀, X.1954, R. Schiapelli & J.A. de Carlo coll. (MACN 5455; MACN 16019); Puerto Libertad (25°55’00”S 54°36’00”W), 16♀, 1 juv., X.1953, J.A. de Carlo, J.M. Viana & M.E. Galiano coll. (MACN 3852); Wanda (25°58’19”S 54°33’58”W), 1♂, X.1985, P.A. Goloboff coll. (MACN 16044). La Pampa: Parque Nacional Lihuel Calel (37°57’S 65°39’W), 1♀, I.1988, P.A. Goloboff & Viñas coll. (MACN 16022).
Diagnosis. Males of Goeldia nigra differ from others species by the slim tegular process, which as an acute and bent tip (Fig. 17A); and by the patellar apophysis short, moderately curved and with an acute tip (Fig. 17B‒C). Females of Goeldia nigra differ from other species by its epigynum with sinuous posterior border (Fig. 17E); enlarged base of the copulatory ducts and long and distally bent vertical spermathecae (Fig. 17F).
Description. Male (IBSP 54702). Coloration: carapace reddish brown with cephalic area, endites and labium brown; sternum caramel brown; legs caramel brown except for femora I dark reddish brown (Fig. 1D). Opisthosoma black. Total length 6.50. Carapace: length 2.81; width 2.63; height 1.31. Clypeus: height 0.18. Opisthosoma: length 3.44; width 2.00. Eye diameters and interdistances: AME 0.07; ALE 0.1; PME 0.11; PLE 0.11; AME–AME 0.11; AME–ALE 0.14; PME–PME 0.175; PME–PLE 0.23; AME–PME 0.17; ALE–PLE 0.08. Chelicerae paturon: length 1.52. Leg measurements: I: femur 2.48/ patella 1.04/ tibia 2.24/ metatarsus 2.00/ tarsus 1.00/ total 8.76; II: 2.10/ 0.94/ 1.68/ 1.88/ 0.88/ 7.47; III: 1.08/ 0.84/ 1.50/ 1.43/ 0.66/ 5.52; IV: 2.24/ 2.20/ 1.00/ 1.68/ 0.72/ 7.84. Leg formula 1423. Leg spination: femur I–II p1ap, tibia I v1r-0-1r-0-1r- 1m, p1sap, r1sap; III v2 sap, r1ap, p1ap; IV v0-0-1p-2, p1sap; metatarsus I v2-1r- 1m; II v2-2- 1m, p1ap, r1ap; III v2-2- 1m, p1ap, r1ap; IV v2-2-2-2- 1m, p1ap, r1ap; tarsus IV v0-2-0. Distally curved setae present only on tibia, metatarsus and tarsus II, and on tibia III and IV. MLT apex partially sclerotized (Fig. 17C). Pars pendula thickening far from embolus tip (Fig. 17D).
Female (MNRJ 1358). Coloration: carapace pale brown with cephalic area and endites brown; chelicerae, labium, and legs pale brown; opisthosoma pale brown. Total length 4.1. Carapace: length 2.05, width, 1.15, height 1.0. Opistosoma: length 1.9, width 1.5. Clypeus: height 0.12. Eye diameters and interdistances: AME 0.06; ALE 0.11; PME 0.07; PLE 0.6; AME–AME 0.08; AME–ALE 0.1; PME–PME 0.14; PME–PLE 0.18; AME–PME 0.1; ALE–PLE 0.08. Paturon 0.91 long. Leg measurements: I: femur 1.16/ patella 0.64/ tibia 1.12/ metatarsus 0.92/ tarsus 0.60/ total 4.44; II: 1.2/ 0.64/ 0.96/ 0.76/ 0.6/ 4.16; III: 1.2/ 0.5/ 0.75/ 0.8/ 0.45/ 3.7; IV: 1.38/ 0.64/ 1.16/ 1.10/ 0.44/ 4.65. Leg formula: 1423. Leg spination: femur I p1ap; metatarsus I r1ap, v1ap; II r1ap; III p1ap, v1ap; IV v1- 1- 1m, p1ap, r1ap. Copulatory duct wall visible at the copulatory opening area (Fig. 17E). Fertilization ducts close to the epigynal rim (Fig. 17E). Vertical spermathecae as high as the copulatory ducts (Fig. 17F), slightly bent outward (Fig. 17F).
Variation. Length (10 ♂): total 4.75–6.50; carapace 2.25–3.25; femur I 1.88–2.50; (3 ♀): total 5.45–5.7; carapace 1.9–2.4; femur I 1.4–2.25.
Distribution. Brazil and Argentina (Fig. 26).
description: Published as part of Almeida-Silva, Lina M. & Brescovit, Antonio D., 2024, Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae), pp. 151-193 in Zootaxa 5428 (2) on pages 177-180, DOI: 10.11646/zootaxa.5428.2.1, http://zenodo.org/record/10847364