Tylodinus mutabilis Luna-Cozar & Anderson & Jones & León-Cortés 2014, sp. nov.
Creators: name: Luna-Cozar, Jesús
name: Anderson, Robert S.
name: Jones, Robert W.
name: León-Cortés, Jorge L.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Curculionidae
Tylodinus
Tylodinus mutabilis

description: 20. Tylodinus mutabilis Luna-Cozar, sp. nov.
(Figures 64–65, 113, 132)
Diagnosis. Length male, 2.3–2.8 mm, female, 2.4–2.7 mm. Width male, 1.1–1.4 mm, female 1.2–1.3 mm. Body approximately 2.3x longer than wide. Head with light brown to yellow scales; vertex with patch of overlapping whitish scales on middle of vertex. Pronotum in dorsal view feebly convex, outline with anterior and posterior constriction; disc somewhat hollowed. Elytra with basal margin sinuate; base of declivity with tubercle on I2 similar in size to tubercle on I4. Metasternum concave. Male abdomen with ventrite 1 concave, ventrite 2 with transverse line of scales along median line. Premucro absent. Male genitalia (Figures 113) with median lobe in lateral view weakly curved, moderately stout; apical process in lateral view moderately large (0.26–0.34 as long as median lobe), slightly curved, abruptly curved downward; in ventral view not constricted beyond the median orifice. Female genitalia with tergite eight trapezoidal, slightly curved at the apex, sternite eight trapezoidal; length of membranous median area 1/3 or ½ as long as basal plate; spermatheca very shallowly curved, ramus and nodulus indistinct.
Comments. This species is variable in the curvature of the front tibiae, sometimes being slightly sinuate; also, the external tooth on the middle tibia is weak or absent.
Geographic distribution. México (Chiapas).
Habitat and elevation. Specimens were collected from leaf litter in various types of forest between 1100–1900 m.
Derivation of specific name. Name derived from Latin word mutabilis = changeable; inconstant.
Material examined. Total 23 males, 19 females. Holotype male (CMNC): México: Chiapas, Villa Corzo, Ejido Sierra Morena, Reserva de la Biósfera La Sepultura, (16° 09' 10.6"N, 93° 35'25.1"W), elev. 1600 m, 19–20.vii.2005. R. Anderson, oak pine ridge forest litter 2005-004A. Paratypes: México: Chiapas, same locality, elev. 1800 m, 18/ 19.vii.2007, R. Anderson — 1 male, 1 female ((CMNC); La Concordia, Reserva El Triunfo, Campamento el Quetzal, elev. 1850 m (15° 43'17"N, 92° 56'21.8"W), 17.v.2008, coll. R. Anderson - 1 female (CMNC); same locality, 19.vii.2007, coll. R. Anderson — 1 female (CMNC); same locality, elev. 1860m, 17.v.2008, coll. R. S. Anderson — 1 male (CMNC); same locality (15.72173N 92.94059W), elev. 1850m, 20.v.2008, coll. R. S. Anderson — 1 female (CMNC); 2 km SE Custepec, elev. 1520 m (15° 43'13.8"N, 92° 57'1.1"W), 17.v.2008, coll. LLAMA (Wa-A-02-1-13) - 1 female (ECOS); same locality (15.72078°N 92.95047°W), 17.v.2008, coll. LLAMA (Wa-A-02-1-18) — 1 male (ECOS); same locality (15.72098°N 92.95054°W), 17.v.2008, coll. LLAMA (Wa-A-02- 1-22) — 1 male (CMNC); same locality (15.72053°N 92.95100°W), 17.v.2008, coll. LLAMA (Wa-A-02-1-41) — 1 female (ECOS); same locality (15.72051°N 92.95104°W), 17.v.2008, coll. LLAMA (Wa-A-02-1-42) — 1 male (CMNC); same locality, elev. 1700 m (15.71522°N 92.93794°W), 17.v.2008, coll. LLAMA (Wa-A-02-2-39) — 1 female (CMNC); same locality, elev. 1700 m (15.71530°N 92.93820°W), 17.v.2008, coll. LLAMA (Wa-A-02-2- 40) — 1 male (CMNC); Villa Flores, Reserva La Sepultura, Ejido Sierra Morena, Parcela Escolar, elev. 1350 (16° 9'10.4"N, 93° 35'25.1"W), 23.vi.2006, coll. J. L. Cozar — 1 male (CMNC); Villa Flores, Ejido Sierra Morena, R. Biosfera La Sepultura, elev. 1400 m (16°9'10.4''N, 93°35'25.1''W), 17-18.vii.2003, coll. R. S. Anderson — 1 female, 1 male (CMNC); same locality (16°09.351’N 93°36.045’W), elev. 1450 m — 1 female (CMNC); same locality (16°09.856’N 93°36.291’W), elev. 1497 m — 3 males, 1 female (CMNC); same locality (16.16121°N 93.60024°W), elev. 1137 m — 2 males, 1 female (CMNC, ECOS); same locality, elev. 1235 m, 19-20.vii.2005, coll. R. S. Anderson — 1 male, 1 female (CMNC); same locality, elev. 1350 m — 1 male, 2 females (CMNC); same locality, elev. 1450 m — 2 males, 1 female (CMNC); same locality, elev. 1600 m — 1 male, 1 female (BMNH, CMNC); same locality (16.15976°N 93.60505°W), 12.v.2008, coll. LLAMA (Wa-A-01-1-26) — 1 male (ECOS); same locality (16.16044°N 93.60516°W), 12.v.2008, coll. LLAMA (Wa-A-01-1-44) — 1 male (CMNC); same locality (16.15458°N 93.60080°W), 12.v.2008, coll. LLAMA (Wa-A-01-2-05) — 1 male (ECOS); same locality (16.15276°N 93.60079°W), 12.v.2008, coll. LLAMA (Wa-A-01-2-36) — 1 male (CMNC); same locality (16.15287°N 93.60059°W), 12.v.2008, coll. LLAMA (Wa-A-01-2-41) — 1 female (ECOS); same locality (16.15292°N 93.60051°W), 12.v.2008, coll. LLAMA (Wa-A-01-2-43) — 1 female (CMNC); Sierra Morena, elev. 1550 m (16.1529°0'N, 93.5903°0'0''W), 17-18.vii.2003, coll. R. S. Anderson — 1 female (CMNC).
description: Published as part of Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W. & León-Cortés, Jorge L., 2014, A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico, pp. 1-63 in Zootaxa 3788 (1) on page 38, DOI: 10.11646/zootaxa.3788.1, http://zenodo.org/record/4913761