Campsurus Molineri & Salles 2017
Creators: name: Molineri, Carlos
name: Salles, Frederico F.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Ephemeroptera
Polymitarcyidae
Campsurus

description: Key to male imagos of Campsurus
1 Pedestals touching each other basally or separated by only a short distance, less than 0.7× the width of a pedestal (e.g. Figs. 35, 75, 85)..............................................................................................2
- Pedestals distinctly separated at the middle, by a distance equal to or greater than the width of a pedestal (e.g. Figs. 1, 25, 100, 117, 138, 151).......................................................................................12
2(1) Pedestal with inner distal corner always more developed than outer corner; outer corner sometimes absent (e.g. Fig. 85).....3
- Pedestal with outer distal corner more developed than inner corner, generally forming a long parastylus (e.g. Figs. 53–59)..6
3(2) FW <7 mm; penes blade-like, not curved (Fig. 85)............................................ .. C. povilla sp. nov.
- FW> 9 mm; penes variable, never blade-like, generally cylindrical or conical and always curved ventrally or medially.... 4
4(3) Penes with only one finger-like lobe, ventrally curved.......................................................................................... violaceus group (see Molineri et al. (2015a) for identification of the six valid species)
- Penes variable, but not finger-like, formed by two lobes (the ventral one may be reduced but always is visible)...........5
5(4) Penes flat, not curved ventrally, and located in the same plane as forceps......................................................................... albifilum group (see Molineri & Salles (2013) for identification of the five valid species)
- Penes curved ventrally, each lobe twisted outwards........................................................................................ major group (see Molineri & Emmerich (2010) for identification of the three valid species)
6(2) Pedestal very long, subrectangular; parastylus relatively short and rounded (Figs. 75–76)............... C. janae sp. nov.
- Pedestal large, but not as above; parastylus longer than above (e.g. Figs. 51–52, 72).................................7
7(6) Penes large, with ear-like base and outer margin granulated (Figs. 71, 73).......................... C. pereirae sp. nov.
- Penes variable, but never with granulated outer margin (e.g., Figs. 53–59, 68–70).................................. 8
8(7) Forceps very short, surpassing the apex of parastylus at a distance less than half of forceps length (Figs. 51–53)............................................................................................... C. cuyuniensis Traver
- Forceps longer, surpassing the apex of parastylus by a distance at least 0.6× the forceps length (Figs. 54, 56)............. 9
9(8) Penes with spatulate apex (Fig. 55); parastylus apically rounded (Figs. 54–55)..................... C. nessimiani sp. nov.
- Main lobe of penes apically pointed (Figs. 36, 39, 70); parastylus apically pointed (in lateral view, may be rounded in ventral view) (Figs. 57, 69)...................................................................................10
10(9) Main lobe of penes relatively long and slender (Figs. 35–36, 39–47).............................. C. essequibo Traver
- Main lobe of penes stouter, main lobe with wider base (Figs. 56–59, 68–70)...................................... 11
11(10) Penes relatively slender at base; parastylus becoming abruptly thinner on apical third in lateral view (Figs. 56–59)............................................................................................... C. dasilvai sp. nov.
- Penes relatively wider at base; parastylus becoming gradually thinner toward the apex in lateral view (Figs. 68–70).............................................................................................. C. zunigae sp. nov.
12(1) Sternum IX strongly projected distally (surpassing pedestals and covering the base of penes) (Figs. 151, 159).................................................................................................. C. litaninensis Spieth
- Sternum IX not projected as above, never surpassing the pedestals (Figs. 117, 123, 100, 103, 107).....................13
13(12) Paraprocts large, clearly protruding as in Figs. 108, 122, 123..................................................14
- Paraprocts of usual size and not protruding................................................................19
14(13) Sternum IX with broad posteromedian emargination (Figs. 117, 123), penes with long and relatively thin lobes (Figs. 120, 126)................................................................................................15
- Sternum IX with straight hind margin (Figs. 100, 103, 107) or with thin median notch (as in C. pedicellarius); penes with short, subtriangullar main lobe and small, rounded secondary lobe (Figs. 101, 104, 109)................................. 16
15(14) Parastylus well-developed (Fig. 124)....................................................... C. indivisus Ulmer
- Parastylus absent (Fig. 119)............................................................. C. inusitatus sp. nov.
16(14) Sternum IX medially notched (see Spieth 1943: Fig. 15).................................... C. pedicellarius Spieth
- Sternum IX without such notch (Figs. 100, 103, 107)......................................................... 17
17(16) Penes dorsally with microspines on outer margin (Figs. 108, 111)................................ C. demeni sp. nov.
- Penes smooth, without microspines as above (Figs. 101, 104)..................................................18
18(17) Parastylus long and slender, at least 2× length of pedestal (Fig. 100)..................... C. lucidus Needham & Murphy
- Parastylus short, less than 1.5× the length of pedestal (Figs. 105–106)............................. C. sinamari sp. nov.
19(13) Distal margin of sternum IX with a subtriangular distomedial projection, sometimes also with a small median triangular point (Figs. 1, 9, 25, 29)....................................................................................20
- Distal margin of sternum IX straight, slightly concave, or with a small truncated median projection (Figs. 138, 140)...... 2 4
20(19) Main lobe of penes with a similar width along its length (Emmerich & Molineri 2011: Figs. 11–13); secondary lobe variable................................................................................................... .. 21
- Main lobe of penes subtriangular, wider at base; secondary lobe reduced in size (Figs. 3, 30).........................22
21(20) Penes with secondary lobe cylindrical and relatively long (Emmerich & Molineri 2011: Figs. 11–13); male fore wings 15.9– 20.5 mm ................................................................ C. vulturorum Emmerich & Molineri
- Secondary lobe of penes apparently absent (Eaton 1871: Plate 4, Fig. 12), size unknown............. C. cuspidatus Eaton
22(20) Penes with dorsal margin smoothly curved in lateral view (Fig. 26)................................ C. salobra sp. nov.
- Penes with indented or sinuous dorsal margin in lateral view (Figs. 4–5)......................................... 23
23(22) Sternum IX projected medially but without an acute point (Figs. 1, 9)........................... C. latipennis (Walker)
- Sternum IX projected medially and with an acute triangular point (Emmerich & Molineri 2011: Figs. 33, 34)........................................................................................ C. evanidus Needham & Murphy
24(19) Main lobe of penes slightly curved, conical; secondary lobe short, less than half the length of main lobe (Emmerich & Molineri 2011: Fig. 50)............................................................ C. jorgenseni Esben-Petersen 1912
- Main lobe of penes strongly curved (Figs. 139, 142, 143) and cylindrical; secondary lobe long (more than 0.6× the length of main lobe)..........................................................................................25
25(24) Main and secondary lobes of penes arising from a large membranous base (Figs. 142, 143)............. C. cotaxe sp. nov.
- Membranous base of penes smaller; secondary lobe arising basally from main lobe (Fig. 139)....................... 26
26(25) Smaller size (male FW max. length 9.8 mm), abdomen darker (brownish to reddish, Figs. 146, 147)............................................................................................... C. segnis Needham & Murphy
- Larger size (male FW length 11 mm), paler abdomen (whitish to yellowish)........................... C. ulmeri Traver
description: Published as part of Molineri, Carlos & Salles, Frederico F., 2017, Review of selected species of Campsurus Eaton 1868 (Ephemeroptera: Polymitarcyidae), with description of eleven new species and a key to male imagos of the genus, pp. 301-354 in Zootaxa 4300 (3) on pages 350-351, DOI: 10.11646/zootaxa.4300.3.1, http://zenodo.org/record/838438