Gomphostemma hirsutum Walsingham
Creators: name: Bongcheewin, Bhanubong
name: Ingrouille, Martin J.
name: Paton, Alan J.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Magnoliopsida
Lamiales
Lamiaceae
Gomphostemma
Gomphostemma hirsutum

description: <b> 13. Gomphostemma hirsutum <i>Walsingham</i></b> (in Walsingham &amp; Bramley 2010: 280); Walsingham (2019: 199). Type: Malaysia, Sabah, Ranau?istr., Kampung Menggis, 29 Aug. 1993, <i>Rumtom</i> [<i>Rumutom</i>] 122 (holotype K [K000259561]; isotype KEP n.v.). Herb up to 1 m tall. <i>Stems</i> dark brown or black, erect, obtusely quadrangular with longitudinal grooves, densely tomentose with a mixture of dendroid hairs with multicelled stalk, stellate hairs with a 1-celled or multi-celled stalk and multi-celled simple hairs. <i>Leaoes</i> petiolate, chartaceous; blades elliptic-obovate, narrowly elliptic or narrowly ellipticoblanceolate, 12 – 26 × 3 – 8 cm, apex acute, margin shallowly dentate, base attenuate; upper side with simple hairs; lower side pale green tomentose with a mixture of branched hairs confined to midrib; petioles 10 – 30 mm long. <i>Inflorescence</i> axillary with opposite cymes sessile, unbranched then congested with axis not visible between flowers, inserted at the upper nodes at which leaves are usually present; verticils 1 – 3-flowered; bracts narrowly ovate or straight falcate, 6 – 10 × 1 – 2 mm, upper side adpressed hirsute or sometimes with bifurcate hairs, lower side with stellate hairs and/or simple hairs; bracteoles linear, 3 – 6 mm long. <i>Flowering calyx</i> infundibular, 10 – 13 mm long, 10-ridged, outside with long simple hairs and long branched hairs, inside adpressed simple hairs; tube 3 – 5 mm long; lobes triangular, 3 – 6 mm long, apex acuminate. <i>Fruiting calyx</i> not seen. <i>Corolla</i> white or creamywhite, 23 – 23 mm long, throat inflated, abruptly dilated near the throat, outside tomentose with stellate hairs, inside glabrous; tube slender, incurved, 20 – 33 mm long; posterior lip ovate, 3 – 8 mm long, apex and margin slightly undulate; anterior lip 3-lobed, longer than posterior lip; median lobe 3 – 9 mm long, apex and margin slightly undulate; lateral lobes shorter than median lobe. <i>Staminal filaments</i> fleshy, sparse tomentose. <i>Style</i> slender, slightly shorter than corolla with apex subequally lobed. <i>Nutlets</i> not seen [round white berries: <i>Darnton</i> 183 (BM)]. <b>DISTRIBUTION.</b> Malaysian Borneo (Sabah). Map 3. <b> SPECIMENS EXAMINED. MALAYSIA. SABAH. Ranau?istr.: 310 m [1500 ft], 16 Feb. 1932, <i>Darnton</i> 183 (BM);</b> Bukit Kulung near Bukit Hampuan at SE base of Mt Kinabalu, 3°39&amp;N 116°21&amp;E, 530 m, 8?ec. 1983, <i>3eaman</i> 5558 (US); near Poring Hot Springs, Kipungit-Langanan Falls trail, 6°3&amp;N 116°22&amp;E, 630 – 530 m, 6 Aug. 1990, <i>3eaman</i> 10893 (K); Kampung Takutan [6.1165°, 116.5165°], 2 Sept. 1993, <i>Tadong</i> 322 (K); Kampung Menggis, 29 Aug. 1993, <i>Rumutom</i> 122 (holotype K); Lahad?atu?istr.: Borneo Rainforest Lodge, Hornbill trail, Borneo Rainforest Lodge, 2°39&amp;20”N 115°30&amp;11”E, 6 July 2006, <i>de Kok et al</i>. SAN 125232 (K);?anum Valley Road, near km 23 [3.2152°, 118.3925°], 10 June 1989, <i>Campbell</i> EC9 (E). <b>HABITAT.</b> In shade and hilly areas of secondary forest, disturbed primary rainforest, lowland forest, dipterocarp forest; alt. 310 – 530 m. <b>CONSERVATION STATUS.</b> Endangered (EN B1ab(iii)) (Walsingham &amp; Bramley 2010). <b>PHENOLOGY.</b> Flowering July – September. Fruiting August – February. <b>NOTES.</b> <i>Gomphostemma hirsutum</i> is a distinctive species recognised by having long white simple hairs, stellate hairs, and dendroid hairs on the flowering calyx, narrow bracts, 1 – 2 mm wide, subtending cymes. It is endemic to Borneo (Sabah). However, the Chinese species, <i>G. stellatohirsutum</i> C.Y.Wu is similar to the Borneo species by having dark or dark brown stems with long dendroid hairs but lacks long simple hairs on the flowering calyx.
description: Published as part of <i>Bongcheewin, Bhanubong, Ingrouille, Martin J. &amp; Paton, Alan J., 2022, A revision of Gomphostemma (Lamiaceae), pp. 27-92 in Kew Bulletin 77 (1)</i> on page 55, DOI: 10.1007/s12225-021-09991-y, http://zenodo.org/record/7615581
description: {"references": ["Walsingham, L. (2019). Gomphostemma. In: G. L. C. Bramley (ed.), Flora Malesiana ser. 1, 23: 1 - 222. Foundation Flora Malesiana, The Netherlands."]}