Koopmanycteris palaeomormoops Morgan & Czaplewski & Simmons 2019
Creators: name: Morgan, Gary S.
name: Czaplewski, Nicholas J.
name: Simmons, Nancy B.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Mammalia
Chiroptera
Mormoopidae
Koopmanycteris
Koopmanycteris palaeomormoops

description: Koopmanycteris palaeomormoops,
gen. et sp. nov.
Figures 2–13
HOLOTYPE: UF 157765, right dentary with p4–m3 and alveoli for i1–p3, from Brooksville 2 LF, early Arikareean, Hernando County, Florida.
PARATYPES: All specimens are from the Brooksville 2 LF. SCSM 2004.35.1, partial right dentary with m1–m3 and complete ascending ramus; UF 209954, right dentary fragment with p3 and alveoli for i1–p1; UF 182782, leπ P4; UF 179949, complete leπ humerus; UF 179922, leπ proximal radius; UF 179948, leπ proximal femur; UF 179947, right distal femur.
REFERRED SPECIMENS: Brooksville 2 Local Fauna: UF 179985, partial right dentary with p4– m2; UF 179951, partial right dentary with p4, m2 and alveoli for c–p3, m1; UF 157767, partial leπ dentary with m1–m2; SCSM 2004.35.2, leπ dentary fragment with m1–m2; UF 157766, partial right dentary with m2–m3, including most of the ascending ramus except for the tip of the coronoid process; SCSM 2004.35.3, partial leπ dentary with m3 and complete ascending ramus; SCSM 2004.35.4, partial right dentary with p3; UF 179957, edentulous leπ dentary with partial ascending ramus; UF 179990, UF 182816, right m1 (2); UF 182815, leπ m1; UF 182810, right m2; UF 179901, right petrosal; UF 179937, right proximal humerus; SCSM 2004.35.5, leπ proximal humerus; UF 179907, 179908, 179938; right distal humerus (3); UF 179905, 179906, leπ distal humerus (2); UF 179917–179919, 1179941, right proximal radius (4); UF 179916, 179920, 179921, 179923–179925, 179943, 179970, 179973, 209955, left proximal radius (10); SCSM 2004.35.6, leπ proximal femur.
I-75 Local Fauna: UF 121718, partial leπ dentary with p4, trigonid of m1, and alveolus for p3; UF 121719, partial leπ dentary with p4 and alveoli for i2–p3; UF 121720, partial right dentary with m1 and alveoli for p3–p4; UF 121733, edentulous partial leπ dentary with alveoli for m2– m3 and anterior edge of ascending ramus; UF 121721, leπ proximal femur; UF 121722, right proximal femur.
TYPE LOCALITY: Brooksville 2 Local Fauna, Hernando County, Florida, late early Arikareean (Ar2), late Oligocene.
ETYMOLOGY: The generic epithet derives from Koopman, in honor of the late Karl F. Koopman, formerly of the American Museum of Natural History, in recognition of his monumental contributions to our knowledge of the systematics, evolution, and biogeography of the Chiroptera, and nycteris, Greek meaning “bat.” The specific epithet is composed of palaeos, Greek for “ancient” or “old,” mormo, Greek for a kind of nighttime monster or ghost, and ops, Greek for “face.”The species name “ palaeomormoops ” or “ancient ghost face” alludes to this species being the oldest known member of the ghost-faced bat family Mormoopidae, and also indicates its phylogenetic relationship with the extant genus Mormoops.
DIAGNOSIS: Lower p3 large, double rooted, outline in occlusal view diamond shaped, with anterior portion of tooth rounded and posterior portion pointed. Lower p4 squarish to slightly rectangular in occlusal view, somewhat narrower anteriorly than posteriorly, with well-developed cingulum on anterior, labial, and posterior margins, but with cingular development weak to absent on the lingual margin. Lower m1 and m2 nyctalodont. Paraconid on m1 highly reduced, positioned labially about halfway between protoconid and metaconid. Metaconid on m1 and m2 taller than entoconid. Dentary with ascending ramus strongly upturned dorsally compared to horizontal ramus; angular process and articular condyle both located well dorsal to alveolar margin of toothrow; coronoid process with its anterior edge essentially vertical; articular condyle located well dorsal to tip of coronoid process; small secondary process present on ascending ramus immediately ventral to angular process. Upper P4 with large, rounded anterior basin, broad talon basin, and tall, sharp central cusp (protocone). Proximal humerus with head elliptical in shape, transversely flattened and directed proximally, and angled toward lesser trochanter. Distal humerus with articular surface moderately offset laterally from humeral shaπ; capitulum not differentiated into medial and lateral ridges; well-developed spinous process separated from medial edge of trochlea by deep notch; epitrochlea a well-developed rounded process with small concavity at proximomedial corner. Proximal end of radius gently rounded, not triangular. Proximal articular surface of radius with two articular facets. Femur with greater trochanter and lesser trochanter highly reduced; greater trochanter larger than lesser trochanter and triangular in shape. Distinct tubercle or short ridge present on posterodistal femoral shaft just proximal to distal articular surface. Femur comparatively long, more than three times longer than alveolar length of toothrow.
description: Published as part of Morgan, Gary S., Czaplewski, Nicholas J. & Simmons, Nancy B., 2019, A New Mormoopid Bat From The Oligocene (Whitneyan And Early Arikareean) Of Florida, And Phylogenetic Relationships Of The Major Clades Of Mormoopidae (Mammalia: Chiroptera), pp. 1-141 in Bulletin of the American Museum of Natural History 2019 (434) on pages 12-13, DOI: 10.1206/0003-0090.434.1.1, http://zenodo.org/record/4613816