Cimbicidae W. Kirby 1837
Creators: name: Vilhelmsen, Lars
name: Smith, David R.
name: Malagón-Aldana, Leonardo A.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Cimbicidae

description: Key to the species of South American Cimbicidae
1. Antennae with at least five antennomeres, one or more apical antennomeres expanded, forming distinct club (Fig. 1 A–B); mesopseudosternal sulcus lateroventrally on mesopleuron absent; fore wing vein 2 r always present, vein Rs always absent between cells 1R and Rs (Fig. 2 A–B); lateral part of abdominal terga separated from median by fold or unsclerotized line, spiracles below line on lateral part (Fig. 3A) …………………………………………………2. Cimbicidae
– Antennae with various numbers of antennomeres, sometimes less than five; apical antennomeres usually not expanded; mesopseudosternal sulcus lateroventrally on mesopleuron present or absent; fore wing 2 r absent or present, vein Rs absent or present (Fig. 2C) between cells 1R and Rs; if antennal club present, then fore wing vein 2 r absent (Fig. 2C); lateral part of abdominal terga rarely separated from median part (Fig. 3B) ………………………………………not Cimbicidae
2. Five antennomeres present (Fig. 1A); median mesoscutal sulcus and notauli always well developed (Fig. 4A); fore wing with vein M always joins Sc+R close to Rs+M junction (Fig. 2A). Exclusively South American ……………………………………………………3. South American Cimbicidae
– Five or more antennomeres present (Fig. 1B); median mesoscutal sulcus and notauli present or absent (Fig. 4B); if present, then fore wing vein M joins Sc+R well separate from Rs+M junction (Fig. 2B). Holarctic or Oriental, never South American …………………………other Cimbicidae
3. Females ……………………………………………………………………………………………4 – Males ………………………………………………………………………………………………10
4. Thorax extensively reddish brown, sometimes with mesoscutum blue-black metallic (Figs 8, 10–11, 14A) ………………………………………………………………………………………………5
– Thorax predominantly dark brown to black (Figs 5A, 6A, 13A), pronotal corners may be creamy white; never extensively reddish brown ……………………………………………………………8
5. Hind basitarsomere shorter than tarsomeres 2–4 combined (Fig. 14A); fore wing vein 1m-cu inserts on vein Rs+M more proximally, distance between 2r-m and 1m-cu insertions on Rs+M (red on Fig. 14D) at least ¼ of distance between M and 1m-cu insertions on Rs+M (green on Fig. 14D); hind wing cell Rs much smaller than cell M, cell M extending to vein Rs proximally (Fig. 14D) …………………………………………… Pseudopachylosticta subflavata Kirby, 1882
– Hind basitarsomere longer than tarsomeres 2–4 combined (cf. Fig. 9A); fore wing vein 1mcu inserts on vein Rs+M distally, distance between 2r-m and 1m-cu insertions on Rs+M (red on Fig. 8A) at most ¼ of distance between M and 1m-cu insertions on Rs+M (green on Fig. 8A); hind wing cells Rs and M of approx. equal size, cell M does not reach vein Rs (Fig. 10A; see also Fig. 9B) ……………………………………………………………6. Pachylosticta (females)
6. Abdomen colored blue-black metallic throughout (Fig. 8A) … Pachylosticta albiventris Klug, 1824
– Abdominal segments 1–3/5 reddish brown, remaining segments blue-black metallic (Figs 10A, 11A) ………………………………………………………………………………………………7
7. Apical antennomere with elongate, lighter colored area on ventral side (Fig. 10 B–C); mesoscutum and mesoscutellum reddish brown (Fig. 10A); apex of fore wing darkly infuscate, proximal part much lighter (Fig. 10A) (male unknown) ……… Pachylosticta apicalis (Westwood, 1835)
– Apical antennomere without elongate, lighter colored area on ventral side (Fig. 11B); mesoscutum mostly blue-black metallic, mesoscutellum reddish brown (Fig. 11A) (male unknown) ………………………………………………………… Pachylosticta plaumanni Malaise, 1939
8. Clypeus predominantly creamy white, contrasting with blackish-brown color of the rest of the head capsule (Fig. 5B); 1 st abdominal tergum subdivided medially by narrow longitudinal line (Fig. 5C) (male unknown) ………………………………… Brasilabia clypealba (Conde, 1932)
– Clypeus blackish brown like the rest of the head capsule (Figs 6B, 13B); 1 st abdominal tergum continuous medially (Figs 6C, 13A) ………………………………………………………………9
9. Dorsomedian margin of torulus pale, overhanging antennal foramen (Fig. 6B); dorsolateral corners of pronotum dark brown to black, as rest of pronotum (Fig. 6C); hind basitarsomere longer than tarsomeres 2–4 combined; hind wing vein M complete, cells Rs and M separated (Fig. 6A; see also Fig. 9B); body length less than 1 cm …… Lopesiana thomasi (Conde, 1937)
– Dorsomedian part of torulus dark, not overhanging antennal foramen (Fig. 13B); dorsolateral corners of pronotum creamy white, contrasting with darker color of the rest of pronotum (Fig. 13C); hind basitarsomere shorter than tarsomeres 2–4 combined; hind wing vein M discontinuous, cells Rs and M partly confluent (Fig. 13A [insert]); body length at least 1.5 cm (male unknown) …….... ............................................................................................……… Pseudabia fusca Schrottky, 1910
10. Antennae inserted very low on face, distance between toruli and median ocellus almost twice the distance between toruli and ventral margin of clypeus (Fig. 7C); body predominantly dark brown to black, no metallic areas (Fig. 7A); body length well below 1 cm … Lopesiana thomasi (Conde, 1937)
– Antennae inserted slightly below middle of face, distance between toruli and median ocellus at most equal to distance between toruli and ventral margin of clypeus (Figs 8B, 10B, 11B, 14B); body with extensive blue-black metallic coloration (Figs 9A, 12); body length approx. 1.5 cm ……………11
11. Hind basitarsomere shorter than tarsomeres 2–4 combined (cf. Fig. 14A); fore wing pterostigma not thickened and protruding from rest of anterior wing margin (Fig. 15); hind wing cell Rs much smaller than cell M, cell M extending to vein Rs proximally (Fig. 15) ……………………………… ……………………………………………………… Pseudopachylosticta subflavata (Kirby, 1882)
– Hind basitarsomere longer than tarsomeres 2–4 combined (Figs 9A, 12); fore wing pterostigma thickened and protruding from rest of anterior wing margin (Figs 9B, 12); hind wing cells Rs and M of approx. equal size, cell M does not reach vein Rs (Fig. 9B) …………12. Pachylosticta (males)
12. Abdominal segments 1–3 and 7–9 in dorsal view blue-black metallic, segments 4–6 reddish brown (Fig.12C) …………………………………………………………………… Pachylosticta sp. (male) – Abdomen blue-black metallic throughout in dorsal view (Fig. 12 A–B) …………………………13
13. Abdomen with sternum and laterotergum 2–4/5 creamy white to orange, contrasting with blueblack metallic color of rest of abdomen (Fig. 9A) …………… Pachylosticta albiventris Klug, 1824 – Abdomen blue-black metallic throughout (Fig. 12) ……………………………………………14
14. Posterodorsal parts of mesopleuron yellowish white, remainder of mesopleuron blue-black metallic; all legs with tibia and most of tarsus yellowish white (Fig. 12A) (female unknown) …………………………………………………………………… Pachylosticta tibialis Klug, 1824
– Mesopleuron blue-black metallic throughout; legs blue-black metallic to dark brown throughout (Fig. 12C) (female unknown) ………………………………… Pachylosticta violacea Klug, 1824
description: Published as part of Vilhelmsen, Lars, Smith, David R. & Malagón-Aldana, Leonardo A., 2018, A review of the South American genera of Cimbicidae (Insecta, Hymenoptera), pp. 1-36 in European Journal of Taxonomy 482 on pages 4-11, DOI: 10.5852/ejt.2018.482, http://zenodo.org/record/3825574