Blepharoneura bipunctata Norrbom & Condon 2010, new species
Creators: name: Norrbom, Allen L.
name: Condon, Marty
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Tephritidae
Blepharoneura
Blepharoneura bipunctata

description: Blepharoneura bipunctata Norrbom & Condon, new species   
Figs. 12, 131  
 Diagnosis. This species resembles B. amplihyalina, apaapa, hyalinella, multipunctata and nigriapex in having a dark brown spot posterodorsally on the anepisternum, a single medial brown spot on the scutellum, and 3 hyaline marginal spots in cell m. It differs from all of them except B. multipunctata in having 2 narrow marginal hyaline marks basally in cell r1 rather than a single broad one, and from B. multipunctata in having a small anepimeral brown spot and the aculeus tip with a convex medial apical lobe.   
 Description. Head: Dark brown area on ocellar tubercle extended more than half distance to postocellar seta or reaching it. Medial occipital sclerite with pair of dark brown submedial vittae on ventral half. Occipital suture narrowly dark brown.   
Thorax: Mostly yellow. Postpronotal lobe with small brown spot at junction with anepisternum and sometimes with small diffuse brown spot on posterolateral margin. Scutum entirely microtrichose, with 2 pairs of dark brown vittae or rows of spots; submedial vitta interrupted slightly posterior to transverse suture and not connected to marks on posterior margin; sublateral vitta interrupted at transverse suture, postsutural part reduced to well separated anterior and posterior spots, latter separated from mark on posterior margin; posterior margin with 2 broad quadrate dark brown marks well separated. Notopleuron with lateral margin dark brown. Small brown spot anterior to postsutural supra-alar seta, large dark brown spot lateral to postsutural seta, small dark brown spot anterior to postalar seta, and large dark brown spot lateral to dorsolateral corner of scutellum present. Scutellum with single, small to moderately large, dark brown medial spot. Subscutellum and mediotergite with pair of dark brown vittae. Anepisternum with small dark brown spot dorsal to anterior seta. Anepimeron with small medial brown spot (faint on 1 side on 1 specimen). Katatergite pale brown dorsally and ventrally. Anatergite with large dark brown area medially and ventrally. Basalare with brown spot or entirely brown. Dorsocentral seta aligned slightly anterior to postalar seta.  
Legs: Mostly yellow. Mid and hind femora with red brown spot on margins of anteroventral and posteroventral apical ridges, extending more faintly slightly more basally very narrowly along ventral margins. Hind tibia dark brown narrowly bordering and on anteroventral and posteroventral basal ridges.  
Wing (Fig. 12): Length 7.55–7.72 mm, width 3.20–3.27 mm, ratio 2.36. Crossvein r-m at 0.55–0.60 distance from bm-cu to dm-cu. Cell c with 2 inverted trapezoidal hyaline spots, both reaching costa and subcosta but fading to pale brown posteriorly; medial brown area almost as dark as area of cell r1 posterior to pterostigma, broader than basal hyaline spot but approximately as broad as distal spot. Pterostigma with large subapical hyaline spot [#3] reaching R1. Cells r1 and r2+3 basally (proximal to apex of R1) with 3 and 2–3 pale brown to hyaline spots, respectively. Radial cells medially with 2 narrow basal marginal hyaline marks in cell r1, both reaching R2+3, sometimes with tiny posterior hyaline spot between them, proximal mark [#5] ovoid; cell r2+3 with 1 very broad hyaline mark posterior to r1 marks (1 wing of 1 specimen) or usually with 1 small hyaline spot [#8] aligned with or slightly distal to basal mark in r1 and 1 broad spot [#9] aligned with second mark in r1; cell r4+5 with small hyaline spot [#14] in anterior half or medially and aligned with basal r1 mark, with small hyaline spot [#15] aligned with or slightly proximal or distal to dm-cu, and also with additional tiny hyaline to pale brown spots, 1–3 near anterior margin and 0–4 on or near posterior margin. Distally cell r1 with 1 [#6] or usually 2 small marginal hyaline spots, on 1 wing of 1 specimen also with 2 minute posterior spots. Cell r2+3 with 2 marginal hyaline marks [#10, 11], proximal mark extending to vein R4+5. Cell r4+5 with small hyaline spot [#16] anteriorly, aligned between apical marks in cell r2+3 or with distal mark; sometimes with small posterior hyaline spot aligned with medial spot in cell m; and with 1–2 small ovoid marginal or submarginal hyaline spots [#18, #18A], more anterior spot sometimes absent or connected away from margin with posterior spot forming somewhat C-shaped mark. Cell m with small subbasal hyaline spot [#49] near midlength of dm-cu and tiny spot near anterobasal corner; with 3 ovoid marginal hyaline spots [#27, #28, #29], medial spot [fusion of at least #28, #26, #26A] largest, irregular, extending to or almost to vein M. Cell br with subbasal pale brown spot [#12]. Cell bm with circular subbasal and subapical hyaline spots [#19, #20]. Cell bcu with or without small hyaline spot in lobe. Posteromedial part of wing with very large hyaline area; cell br subapically with hyaline spot [#13] and small to minute, much more proximal, pale brown spot; cell dm with 1 small anterior ovoid or narrow elongate medial hyaline spot very close to base and with large ovoid medial posterior hyaline mark [fusion of #22?, #23, #24] (ca. 0.25 times as long as cell along vein Cu1) much shorter than hyaline mark in cell cu1 and aligned with its distal part; cell cu1 with very large hyaline area covering medial half or more [broad fusion of at least #31?, #32, #33, #34, #35?, #36, #36A], very broad on posterior wing margin, lacking anterior brown spots and including 1 submarginal pale brown spot subbasally; subapical marginal hyaline spot [#37] relatively small, not reaching vein Cu1. Cell dm with subapical hyaline spot [#25] moderate sized, and with smaller more distal anterior spot and posterior spot [#53] aligned or nearly aligned with subapical mark in cell cu1.  
Abdomen: Mostly yellow, all tergites with 4 rows of evenly spaced dark brown spots, on tergites 3–5 more lateral spot large and connected to or narrowly separated from lateral and posterior markings; tergites 3–5 also with anterolateral spot touching or almost touching lateral margin; all tergites also with pair of spots or band on posterior margin and posterolateral corner, separated medially, on tergites 3–5 extending anteriorly on lateral margin and connected to anterolateral spot, but on tergites 3 and 4 margin with small yellow spot.  
Female terminalia: Oviscape entirely dark brown; length 1.09 mm. Aculeus (Fig. 131) 0.87 mm long, 1.98 times as long as wide, with acute scales dorsally and ventrally on membrane medially; tip angular basolaterally, short triangular (lobed part 0.34 times as long as wide), with small, convex medial lobe and 3 pairs of step-like lobes separated by deep gaps; sublateral lobe larger than submedial lobe; lateral gap 0.90 times as long as wide.  
 Spermathecae subspherical, with straight to slightly convoluted, slender sclerotized neck and large cylindrical basal apodeme (similar to B. femoralis).   
 Distribution. Ecuador. The type specimens were collected at 2000 m elevation.   
 Type data. Holotype ♀ (CMP USNMENT00213904), ECUADOR: Cotopaxi: San Francisco de las Pampas, Otonga, 2000 m, 20 Oct 1996, G. Onore. Paratype: Same data as holotype, 1♀ (USNM USNMENT00213903).   
 Etymology. The name of this species is an adjective referring to the two basal marginal hyaline marks in cell r1.
description: Published as part of Norrbom, Allen L. & Condon, Marty, 2010, Revision of the femoralis group of Blepharoneura Loew (Diptera: Tephritidae) 2374, pp. 1-139 in Zootaxa 2374 (1) on pages 36-39, DOI: 10.11646/zootaxa.2374.1.1, http://zenodo.org/record/5315903