Trypophloeus Fairmaire 1864
Creators: name: Johnson, Andrew J.
name: Hulcr, Jiri
name: Knížek, Miloš
name: Atkinson, Thomas H.
name: Mandelshtam, Michail Yu.
name: Smith, Sarah M.
name: Cognato, Anthony I.
name: Park, Sangwook
name: Li, You
name: Jordal, Bjarte H.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Curculionidae
Trypophloeus

description: Trypophloeus Fairmaire, 1864: 105 (Fig. 21)
Synonymy
= Glyptoderes Eichhoff, 1878b: 34.
Type of genus
Bostrichus binodulus Ratzeburg, 1837.
Diagnosis
This genus can be diagnosed by the emarginated eye, by the coneshaped antennal club with slightly recurved sutures, and by the rounded lateral margins of the pronotum.
Female
Eye emarginated. Frons simple with no extensive sculpturing or of vestiture. Cuticle behind eyes reticulated or aciculated. Antennae with five funicle segments. Antennal club conical, sutures recurved or straight. Pronotum armed with four to six marginal asperities, with the median of the margin slightly projected. Pronotal declivity with more than 30 asperities leading to a slightly elevated summit. Lateral margins of the pronotum rounded, hypomeron with mostly hair-like setae, sometimes with a small number of bifurcating setae. Elytra broadly convex, sometimes with interstriae 4 slightly raised. Vestiture comprised of hair-like or scale-like interstrial bristles and ground vestiture.
Male
Similar to female except anterior margin of pronotum slightly more projected. Elytral interstriae 4 sometimes with small tubercles or spines near declivity. Penis apodemes of a similar length to penis body, separated at apex. Tegmen open dorsally. Median ventral face of tegmen without apodeme or with only a slight projection. Spiculum gastrale with a fork, and of similar thickness to the penis apodemes. Basal sclerites visible.
Distribution
East Asia, Europe, North Africa, North America.
Remarks
Fifteen species known. Found primarily on Salix and Populus (Salicaceae), and Alnus (Betulaceae). Larvae develop communally for the first instar, then independently to complete development (Furniss 2004).
Type material examined
Holotype of Trypophloeus populi Hopkins, 1915 (USNM); holotype of Trypophloeus salicis Hopkins, 1915 (USNM); holotype of Trypophloeus concentralis Hopkins, 1915 (USNM).
Included species
Trypophloeus alni Lindemann, 1875a: 136.
= Trypophloeus holdhausi Wichmann, 1912: 186 (syn: Balachowsky, 1949).
Trypophloeus binodulus (Ratzeburg, 1837: 163) (Bostrichus).
= Cryphalus grothii Hagedorn, 1904: 232 (syn: Reitter, 1913).
= Trypophloeus spiculatus Eggers, 1927d: 122 (syn: Sokanovskiy, 1954).
= Trypophloeus populi Kurentsov, 1941: 164 (syn: Krivolutskaya, 1996).
= Trypophloeus berezinae Stark, 1952: 285 (syn: Sokanovskiy, 1954).
= Trypophloeus kurenzovi Nunberg, 1956b: 208 (syn: Krivolutskaya, 1996).
= Trypophloeus kurenzowi Schedl, 1959d: 42 (syn: Krivolutskaya, 1996).
Trypophloeus bispinulus Eggers, 1927d: 121.
Trypophloeus dejevi Stark, 1936: 152.
= Trypophloeus dejevi Eggers, 1942: 31 (syn: Pfeffer, 1944).
Trypophloeus discedens Palm, 1950: 142.
= Trypophloeus palmi Hansen, 1956: 183 (syn: Süda, 1996).
Trypophloeus grandis Schedl, 1964a: 99.
Trypophloeus granulatus (Ratzeburg, 1837: 164) (Bostrichus).
= Cryphalus granulatus var. tredli Hagedorn, 1904: 232 (unavailable name).
Remarks: Cryphalus tredli Reitter, 1908 was erroneously listed as a synonym of Trypophloeus granulatus, without the original genus in Wood and Bright (1992) (p. 845), in addition to its correct entry as a valid species in Hypothenemus. Cryphalus granulatus var. tredli Hagedorn, 1904 was described without locality data, so cannot be considered even a subspecies.
Trypophloeus klimeschi Eggers, 1915: 188.
Trypophloeus niger Stark, 1936: 152.
Trypophloeus populi Hopkins, 1915: 37.
Trypophloeus rybinskii Reitter, 1895: 72.
⊇ Trypophloeus rybinskii corsicus Eggers, 1912: 113 (syn: Balachowsky, 1949).
= Trypophloeus rybinskii salicis Stark, 1952: 283 (Permanently invalid name due to primary homonymy).
Trypophloeus salicis Hopkins, 1915: 36.
= Trypophloeus concentralis Hopkins, 1915: 36 (syn: Wood, 1954).
Trypophloeus striatulus (Mannerheim, 1853: 235) (Cryphalus).
= Cryphalus nitidus Swaine, 1912: 349 (syn: Wood, 1969).
= Cryphalus punctipennis Hopkins, 1915: 37 (syn: Wood, 1954). Trypophloeus thatcheri (Wood, 1954: 994) (Cryphalus).
Trypophloeus tremulae Stark, 1952: 287.
description: Published as part of Johnson, Andrew J., Hulcr, Jiri, Knížek, Miloš, Atkinson, Thomas H., Mandelshtam, Michail Yu., Smith, Sarah M., Cognato, Anthony I., Park, Sangwook, Li, You & Jordal, Bjarte H., 2020, Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae), pp. 1-81 in Insect Systematics and Diversity 4 (3) on pages 42-44, DOI: 10.1093/isd/ixaa002, http://zenodo.org/record/3826789