Archoleptoneta Gertsch 1974
Creators: name: Ledford, Joel M.
name: Griswold, Charles E.
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Leptonetidae
Archoleptoneta

description: Genus Archoleptoneta Gertsch, 1974
Type species. Archoleptoneta schusteri, Gertsch 1974.
Archoleptoneta Gertsch 1974: 198 (type species Archoleptoneta schusteri Gertsch 1974: 199); Platnick 2008. Archoleptoneta (part): Brignoli 1974: 210 –212; Brignoli 1977: 215 –217; Ledford 2004: 122; Platnick 1994.
Diagnosis. Small (1.16–1.82 mm), three-clawed, cribellate, six-eyed spiders separated from other leptonetids except Darkoneta gen. nov. by having the eyes contiguous (Figs. 6–8, 23, 25); male palpal tarsus cylindrical, lacking a dorsal groove (Fig. 78); bulb suboval with three accessory sclerites (Figs. 76–77, 98, 101, 104, 107); females with simple, paired receptacula (Figs. 109). Separated from Darkoneta gen. nov. by having a cribellum, males with a long, tapering embolus that extends to the tip of the tarsus (Fig. 76–78, 98, 99, 104, 105, 108), and a straight, unmodified prolateral sclerite (PRS, Figs. 98–99, 101–102, 104–105, 107–108). Females with spermathecae that are longer than wide (Figs. 109–110).
Synapomorphies. Archoleptoneta species are united by features of the male palpal bulb, including the presence of a straight prolateral sclerite (PRS, Figs. 98, 101–102, 104–105, 107) and a straight median sclerite (MS, Figs. 98, 101, 104, 107).
Description. Total length 1.16–1.82. Carapace pale to light-brown, round to oval in dorsal view (Figs. 6–8, 23), length 1.14–1.30 x width; smooth; thoracic fovea inconspicuous; rounded along posterior margin in both sexes; six eyes, AME absent; ALE and PME with dark to dusky markings, u-shaped at posterior margin (Figs. 7, 25); OA wider than long, with a single seta anteriad of ALE, two setae posteriad of PLE; chelicerae unmodified, fang promargin with a single large proximal tooth and 6–8 denticles. Sternum smooth, oval, tapering posteriorly, as long as wide. Abdomen (Figs. 6, 8) oval, pale to light-brown, covered with fine setae; spinning organs (Figs. 43–44, 47, 49–56); cribellum oval, divided, with uniformly distributed strobilate spigots, reduced in males (Fig. 53); ALS cylindrical, with 3–4 tartipores, with at least two major ampullate gland spigots and 3–5 piriform gland spigots; PMS tetrahedral, females with 2 anterior cylindrical gland spigots, 2 aciniform gland spigots posteriorly (Fig. 51), males retain 2 aciniform gland spigots; female PLS with 2 posterior cylindrical gland spigots and a single spigot on an elongate pointed base, which is replaced by a nubbin in males. Leg formula IV, I, II, III or I, IV, II, III; legs elongate and thin, sparsely covered with fine setae and lacking spines; male femur I 1.50–1.90 x carapace width, females 1.50–1.89 x carapace width; middorsal integumentary glands elongate, with single, small pores (Fig. 31). Male palpal femur length 0.57– 0.77 x carapace width; palpal tarsus cylindrical in dorsal view (Fig. 78). Basal haematodocha expandable, bulb suboval, bearing three ventral accessory sclerites at base (Figs. 76, 98, 101, 104, 107) embolus straight to sinuous, extending to the tip of the tarsus (Fig. 76, 98, 104–105, 108); tarsal organ trilobed (Figs. 37, 39). Female genitalia (Figs. 109–110) haplogyne, consisting of simple, paired spermathecae.
Composition. Two species, A. schusteri Gertsch, 1974 and A. gertschi sp. nov.
Distribution. Restricted to California (Fig. 115).
description: Published as part of Ledford, Joel M. & Griswold, Charles E., 2010, A study of the subfamily Archoleptonetinae (Araneae, Leptonetidae) with a review of the morphology and relationships for the Leptonetidae, pp. 1-32 in Zootaxa 2391 on pages 10-11, DOI: 10.5281/zenodo.193866