Incertosulcus shiraz Barbosa & Azevedo 2020, sp. nov.
Creators: name: Barbosa, Diego N.
name: Azevedo, Celso O.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Bethylidae
Incertosulcus
Incertosulcus shiraz

description: Incertosulcus shiraz Barbosa & Azevedo sp. nov.   
(Figures 3 (a), 7(c,d), 8(c))  
 Material examined   
 Holotype. ♀ Kenya, Rift Valley Prov [ince], Olorgesailie Nat [ional] Monument, 1.57930°S, 36.44566°E, 982 m, Malaise Trap, Acacia-Commiphora Savan [nah], 30 Oct [ober] – 13 Nov [ember] 2011, R. Copeland (NMKE).   
 Paratype. ♀ South Africa, N[orth] Cape, adjacent to Nieuwoudtvile Flower Reserve, Dump Site (2.78 Km 67° ENE Nieuwoudtvile) 31°21.617’S 19°08.062 ʹ E, Oats field on dolerite, 12–15 October 2000, S. van Noort, Malaise trap, NW00-DA1-M28 (SAM-HYM P0024572).   
 Description   
 Female. Body length 2.8 mm. Forewing 1.7 mm. Head: head longer than wide. Malar space as long as VOL, convergent anteriorly. Mandible with three sharpened apical teeth. Median clypeal lobe quadrate; lateral lobe short; median clypeal carina arched, complete, straight in dorsal view. Antenna short [not reaching the mesoscutum]; scape slightly curved; pedicel caliciform; flagellomere I shorter than pedicel; flagellomeres short. Eye small [0.5 x LH], circular. Frons strongly foveolate. Frontal angle of ocellar triangle right; ocelli very small, DAO 2.0 × POL; ocellar fovea absent. Dorsal half of occipital carina high, with side strigate; ventral half of occipital carina complete, high, with side strigate. Mesosoma: dorsal pronotal area shorter than wide, weakly foveolate, anterior margin outcurved, anterior corner angled, trapezoidal, lateral margin straight, posterior margin sinuous; longitudinal pronotal sulcus complete. Anteromesoscutum coriaceous, longer than mesoscutellum; longitudinal antero-mesoscutal sulcus absent; notaulus deep, narrow, polished; parapsidal signum shallow. Axilla strigate, shallow. Mesoscutummesoscutellar sulcus arched, not laterally dilated; mesoscutellum short, coriaceous. Metapectal-propodeal complex 0.5 × as long as wide; metapostnotum with longitudinal ridge between metaposnotal median carina and metaposnotal-propodeal suture; metapostnotal surface strigate; metapostnotal median carina complete; posterior projection of metapectal-propodeal complex short, thick. Propodeal declivity strigate, with median and lateral carinae. Lateral surface of metapectal-propodeal complex strigate, without subdorsal carina, with vertical carina. Mesopleuron foveolate; posterior oblique sulcus foveolate; mesopleural suture with inner margin strigate. Wings: macropterous; transverse median vein arched; stigma sub-developed, distal margin truncate. Metasoma: tergum I with ventral surface coriaceous, with lateral pilosity; tergum II with dorsal surface polished, with lateral pilosity, ventral surface polished. Genitalia: T9 slender, not overlapping the 2vf. 2vf with anterior area (aa) narrow, posterior projection narrow, dorsal projection (dp) short; anterior section of dorsal flange of second valvifer (asf) transversally oriented. Shaft (1vv+2vv) as long as sting body (2vf+3vv), thick, 2vv wider than 1vv. 3vv longer than wide.   
 Colouration. Wings: bi-banded; light castaneous: scape, pedicel, mandible, palpi, wing venation, tibiae, and tarsi; orange: pronotum and mesonotum; castaneous: head, tegulae, coxae, trochanters, femora; dark castaneous: flagellum and metasoma.   
 Remarks   
 This species is similar to I. consimilis (Móczár, 1970) by having eye small, longitudinal sulcus of anteromesoscutum absent, and posterior projection of metapectal-propodeal complex short. However, Incertosulcus shiraz sp. nov. has body dark castaneous, head longer than wide, median clypeal lobe quadrate, ocelli very small, anteromesoscutum coriaceous, the notaulus narrow, the posterior carina of metapectal-propodeal complex incomplete medially, lateral surface of metapectal-propodeal complex without metapectal carina and mesopleural suture with inner margin strigate.   
 The metapectal-propodeal complex posterior carina incomplete medially is an interesting characteristic observed for this species and genus, because this characteristic was recorded only for Heterocoelia species, within Mesitiinae.
description: Published as part of Barbosa, Diego N. & Azevedo, Celso O., 2020, Synopsis of Incertosulcus Móczár, 1970 (Hymenoptera: Bethylidae), pp. 1363-1393 in Journal of Natural History 54 (21 - 22) on pages 1376-1378, DOI: 10.1080/00222933.2020.1794073, http://zenodo.org/record/5030378