Empis (Enoplempis) subcanaster Sinclair, Brooks & Cumming, 2025, sp. nov.
Creators: name: Sinclair, Bradley J.
name: Brooks, Scott E.
name: Cumming, Jeffrey M.
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Empididae
Empis
Empis subcanaster

description: Empis (Enoplempis) subcanaster sp. nov.
urn:lsid:zoobank.org:act: 687C73F1-77E4-482A-8A82-E7AF62C2C72C
(Figs 121–123, 128, 130)
Type material. HOLOTYPE ♂, labelled: “USA: ID: Shoshone Co. / Hobo Cedar Grove Bot. Area / St. Joe NF, 21.vi.2014, 1360 m / N 47°05.150′ W 116°06.776′/ ex. streams/for. B.J. Sinclair ”; “ CNC/ 1078368 ”; “ HOLOTYPE / Empis (Enoplempis) / subcanaster Sinclair,/ Brooks & Cumming [red label]” (CNC). PARATYPES: USA. Idaho: Same data as holotype (4♂, 4♀, CNC).
Diagnosis. This species is distinguished from other species of the E. (En.) canaster species group by the following male characters: a posterior row of preapical setae on the hind femur, one paint brush-like preapical ventral lobe on the hind femur, swollen foreleg basitarsus, pair of digitiform basal processes on the hind tibia, and the length of the dorsal setae on the hind tibia less than twice width of tibia (Figs 121–123).
Description. Wing length 6.6–7.4 mm. Male. Similar to E. (En.) canaster, except as follows: head with ocellar setae short, slightly longer than ocellar triangle, posterior pair less than half-length of anterior setae, similar to short setulae. Postocular setae with upper 2–3 setae subequal to ocellar setae. Antenna dark; scape slightly more than 2× longer than pedicel, with short, black setae; postpedicel long and tapered, 5× longer than basal width; stylus more than 2× longer than basal width of postpedicel.
Thorax with scutum with all setae very short, except long, strong posterior npl; apex of postpronotal lobe with small shiny brown area; postalar ridge brown. Postpronotum with 1 short seta and several short setulae. Scutum with short, fine uniserial acr; dc uniserial, slightly longer, stronger than acr, increasing in length posteriorly; 1 posterior npl, with 1–3 short anterior npl; 0 presut spal, with several setulae; 0 psut spal, with several setulae; 1 short pal. Scutellum with 1 widely separated pair of short sctl, with 1 outer, shorter pair of marginal setulae. Laterotergite with narrow patch of 3–4 strong, dark setae.
Legs slender, yellowish brown, apex of fore tibia and tarsus dark brown; coxae yellowish brown, extreme base with greyish pruinescence; mid and hindlegs with apical tarsomeres darker (Fig. 121). Coxae with stiff, black setae. Hind trochanter with short setulae. Fore femur clothed in short, black setae, with longer preapical posteroventral seta. Fore tibia with 3–4 long anterodorsal setae on apical half, longer than width of tibia; apical half with long posterior setae; ring of outstanding preapical setae; anterior face clothed in dense short setulae. Fore tarsomere 1 distinctly swollen, broader than basal width of tibia, clothed with dense setulae, long strong ventral setae, longer than width of tarsomere; tarsomeres 2–5 with biserial row of spine-like ventral setae; tarsomeres 1–4 with apicolateral pair of spine-like setae, longer than width of segment. Mid femur clothed in short, black setae, with longer preapical anteroventral and posteroventral setae. Mid tibia with 3–4 posterior and posteroventral setae, shorter than width of femur; several preapical setae surrounding apex; 2–3 anterodorsal and posterodorsal setae, subequal to width of femur. Mid tarsus slender, tarsomeres 2–5 with biserial row of spine-like ventral setae; tarsomeres 1–4 with apicolateral pair of spine-like setae, longer than width of segment. Hind femur with 1 preapical, paint brush-like ventral process and forked, outer posteroventral digitiform process with apical setulae on outer fork; without process or ridge posterior to paint brush-like process; apex of femur with preapical row of 6–8, thin posterior setae, shorter than width of femur; preapical anterior seta strong, shorter than width of femur (Figs 122, 123). Hind tibia with forked posteroventral digitiform process on basal quarter, outer fork with crown of long, black setae, inner fork with setulae; rounded, knob-like anteroventral lobe distal to posteroventral process; seta present in posteroapical comb; row of anterodorsal and posterodorsal setae, increasing in length apically, slightly longer than width of tibia (Figs 121–123). Hind tarsomere 1 with several anterodorsal and posterodorsal erect setae, similar to tibia; tarsomeres 2–5 similar to mid tarsus.
Wing with R 5 and M 1 divergent near wing margin; R 5 ending slightly anterior to or at wing tip.
Abdomen dark, with dense greyish pruinescence, concolourous with scutum; long, pale lateral setae on pregenital segments. Pregenital segments unmodified, except sclerites of segment 8 narrowly separated, weakly narrowly fused anteriorly. Terminalia (Fig. 128): phallus orange brown, epandrium subshiny; apex of cercus darkly pigmented. Cercus short, rectangular; dorsal margin slightly expanded basally; apex truncated, with fewer setae than basally; setae longer than width of cercus; weakly fused anteriorly to epandrium. Subepandrium without process; with row of medium-length setae. Epandrial lamella elongate, higher than long, with truncate posterior margin, slightly projected ventrally; longest setae on posterior half; setae shorter than length of epandrium. Hypandrium short, apical margin rounded, slightly pointed medially; numerous setae of variable lengths present. Phallus very narrow and tubular basally; strongly expanded on basal half, tapered, mostly straight apically; apex emerging beyond cercus; apex not expanded; ejaculatory apodeme longer than length of epandrium, T-shaped, with ventrally positioned lateral apodemes.
Female. Similar to male, except as follows: frons broader, nearly as wide as ocellar triangle; hindleg without modified setae and processes. Abdominal pleural membrane not expanded; apex of tergite 6 and tergites 7 and 8 shiny.
Geographical distribution and seasonal occurrence (Fig. 130). This species is known only from a single collecting event in June at a Cedar reserve in Shoshone County, Idaho.
Etymology. The species name is in reference to the similarity of this species with E. (En.) canaster.
Nuptial gift presentation. Form unknown.
description: Published as part of Sinclair, Bradley J., Brooks, Scott E. & Cumming, Jeffrey M., 2025, Revision of the western Nearctic species of Empis subgenus Enoplempis (Diptera: Empididae), pp. 1-200 in Zootaxa 5615 (1) on pages 60-62, DOI: 10.11646/zootaxa.5615.1.1, http://zenodo.org/record/15217957