Phanagenia cornuta He & Ma 2024, sp. nov.
Creators: name: He, Yi
name: Li, Qiang
name: Ma, Li
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Pompilidae
Phanagenia
Phanagenia cornuta

description: Phanagenia cornuta He & Ma, sp. nov.
urn:lsid:zoobank.org:act: CF0215E1-6CF5-46BA-811F-42CEB282E3CB
Fig. 2
Type material. Holotype, ♀, China, Inner-Mongolia, Alax Left Banner, Helan Mountain, Terrazzo Trench, 38°57′- 38°56′36′′N, 105°57′- 105°57′25′′E, 1931–2067 m, 1.VIII.2010, coll. Dingjie ZHANG. Paratypes: 1♀, same location as holotype, 28.VII.2010, coll. Fangzhou MA; 1♀, China, Inner-Mongolia, Alax Left Banner, Helan Mountain, 38°38′27′′N, 105°47′17′′E, 1954 m, 10.VIII.2010, coll. Fangzhou MA; 1♀, same location and collector as holotype, 26.VII.2010; 1♀, China, Inner-Mongolia, Alax Left Banner, Helan Mountain, Xiangchizi, 25.VIII.2013, coll. Li JIANG; 1♀, China, Hebei, Wei County, Wangxidong, 39°37′N, 114°47′46′′E, 1495 m, 21. VI.2009, coll. Zhenhua ZU.
Diagnosis. The new species Phanagenia cornuta He & Ma, sp. nov. clearly differs from other congeners by the combination of characters: head in frontal view 0.95× as broad as high; pronotum posterior margin angulate medially; propodeum with strong, transverse rugae apically; fore wing without fasciae.
Description. Female (Measurements of the holotype are given in parentheses). Body length 7.5–9.5 (7.5) mm. Fore wing length 6.5–8.5 (6.7) mm (Fig. 2A). Colouration. Integument black except: mandible except basally and apically reddish; labial palpus, maxillary palpus basally and apically (brown in middle), legs and metasomal segments apically fulvous; fore femur apically, all tibiae, all tarsi and all claws apically brown. Fore wing hyaline, without fasciae. Pterostigma brown. Hind wing hyaline, not infuscate (Fig. 2I). Integument. Body with silvery white pubescence covering. Clypeus apically, fore coxa, T1 basally and S2–5 with sparse, short and erect setae. T6 and S6 with sparse, long and erect setae. Head. Head in frontal view 0.95× as broad as high (Fig. 2B). Clypeus convex except apical margin, 2.0× as broad as long, apical margin subarcuately convex and very slightly angulate at middle, its polished apical rim impunctate and with longitudinal striae, broadest at middle. Labrum unexposed. Mandible bidentate and two tooth blunt apically. Last three segments of maxillary palpus much slender than third segment, third segment apically swollen, and fourth segment equal to fifth. Fourth segment of maxillary palpus longest of all palpus, relative length of fourth to sixth palpal segment = 8: 4: 3.5. Fourth segment of labial palpus slender, relative apical width of second to fourth segment = 1.5: 1: 0.75. Prementum with many long, stout and curved bristles. Ratio of malar space and antennal pedicel = 0.5: 2. Frons with median sulcus reaching anterior ocellus. Inner orbits convergent above and below, divergent at middle (Fig. 2B). UID: MID: LID = 1.2: 1.3: 1.2 (Fig. 2B). Ocelli in acute triangle, anterior ocellus larger than posterior ones, ocellar area slightly elevated (Fig. 2C). Vertex slightly convex above level of eye tops (Fig. 2B). Gena in dorsal view, strongly receding posteriorly (Fig. 2C), in profile, relative length of eye and gena medially = 1: 1, lower half of gena wider than its upper part (Fig. 2D). Occipital suture present (Fig. 2D). POD: OOD: Od: OCD = 1.2: 1.7: 0.6: 2.0 (Fig. 2C). F1 longest and 6.3× as long as its apical width. Ratio of scape, pedicel F1 and F2 = 5.5: 2.5: 11: 9.25 (Fig. 2E). Mesosoma. Pronotum shorter than mesoscutum, ratio of length to width medially = 4.5: 12.5, with dorsum flattened medially, gradually sloping anteriorly, its posterior margin angulate medially. Pronotal collar depressed, and much lower plane than thoracic dorsum in lateral view. Pronotum 2.0× as long as mesoscutum at middle (Fig. 2F). Mesoscutum with parapsidal sulcus reaching nearly 2/3 of mesoscutum. Disc of scutellum almost equal to height of mesoscutum in lateral view (Fig. 2G). Metanotum 1.0× as long as metapostnotum at midline, laterally with sparse, oblique striations. Metapostnotum with dense, transverse striations and longitudinal depression medially, its anterior and posterior margins nearly parallel in dorsal view. Propodeum with conspicuous transverse rugae and apical part stronger than basal part, without median groove, sides of propodeum slightly convex in dorsal view, its posterior surface with gradually sloping, not distinct edge, ratio of length to width medially = 12.5: 13.75. Relative length of mesoscutum, metanotum, metapostnotum and propodeum medially = 5: 2: 2: 12.5 (Fig. 2G). Mesopleuron with broad mesopleuraulus (Fig. 2H). Legs. Relative length of fore, mid and hind femur/tibia = 2.6/1.8, 2.4/2.3, 3.0/3.4. All tibiae with fine short spines dorsally. All tarsomeres 1–5 beneath with two rows of spines (Fig. 2J). Longer spur of hind tibia 0.5× hind tarsomere I. All tarsal claws dentate and hind claw sharp apically (Fig. 2J). Wings. Fore wing marginal cell obviously shorter than its outside to wing tip, relative length in middle = 18: 24. Relative length of SMC2: SMC3 = 7.5: 9 on vein M, 5: 5 on vein Rs. SMC2 1.6× as high as long, receiving cross vein 1m-cu at its apex. SMC3 1.7× as high as long, receiving cross vein 2m-cu at its base, distant from outer wing margin by 1.35× its own length. Cross vein 2rs-m almost straight, nearly vertical to vein M. Cross vein 2m-cu slightly curved outward. Cross vein cu-a originating distal to point of separation of vein M+CuA. Hind wing cross vein rs-m nearly vertical to vein M. Cross vein cu-a joins vein M+CuA much before juncture of M with CuA (Fig. 2I). Metasoma. T1 petiolate, with lateral crest on each side. S2 with obvious transverse groove. S6 not compressed laterally. T6 without pygidial area (Fig. 2K).
Male. Unknown.
Distribution. China (Hebei, Inner-Mongolia) (Fig. 3).
Etymology. The specific epithet cornuta is derived from the Latin cornuta (= cornual), referring to the clypeus apical margin angulate at middle, which is one of the main recognition characters of this species.
Remarks. In the similar species Ph. takasago Tsuneki: the head in frontal view 1.1× as broad as high; the pronotum posterior margin gently arcuate medially; the propodeum with weak, transverse rugae apically; the fore wing with two fasciae.
In addition, this new species also has some distinguishing characters from Ph. comes Banks and other congeners by the combination of characters: the body length 7.5–9.5mm; the fore wing length 6.5–8.5 mm; POD: OOD = 1.2: 1.7; the pronotum posterior margin angulate medially; the propodeum with conspicuous transverse rugae; longer spur of hind tibia 0.5× hind tarsomere 1; SMC2 1.6× as high as long; cross vein 2rs-m almost straight. In the species P. comes: the body length 13.5 mm; the fore wing length 12 mm; POD: OOD = 1: 2; the pronotum posterior margin arcuate medially; the propodeum with weak transverse rugae; longer spur of the hind tibia 0.4× hind tarsomere 1; SMC2 2.0× as high as long; cross vein 2rs-m curved.
description: Published as part of He, Yi, Li, Qiang & Ma, Li, 2024, The genus Phanagenia Banks, 1933 (Hymenoptera: Pompilidae) from China with descriptions of two new species and a key to the world species, pp. 542-552 in Zootaxa 5446 (4) on pages 545-547, DOI: 10.11646/zootaxa.5446.4.6, http://zenodo.org/record/11102272