Josa samaria Martínez, Kochalka, Cabra-Garcia & Ramírez, 2025, sp. nov.
Creators: name: Martínez, Leonel
name: Kochalka, John A.
name: Cabra-Garcia, Jimmy
name: Ramírez, Martin
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Anyphaenidae
Josa
Josa samaria

description: Josa samaria sp. nov.
Figures 9–10
Tomopisthes chazaliae Simon, 1897: 11 (only one female paralectotype MNHP; vchLAM-244, misidentification).
Type Material. Holotype: COLOMBIA. Magdalena, from Sierra Nevada de Santa Marta, De Dalmas, 1 ♀, (MNHP; vchLAM-244; paralectotype of Josa chazaliae Simon 1897).
Etymology. The specific epithet refers to the demonym for the Santa Marta people “Samario/a”. It is a noun in apposition.
Diagnosis. The female is similar to that of Josa nadineae sp. nov. by the posterior quadrangular notch of the epigyne (Fig. 13E, compare with Figs 13E, 14D) but can be distinguished by the posterior lateral margins on a straight transversal line (vs. oblique in Josa nadineae sp. nov.), and by the copulatory ducts describing two wide posterior coils, and the small spermathecae, anteromedially placed with long fertilization ducts (Fig. 10A–E).
Description. Female (holotype, MNHP; vchLAM-244) (Fig. 9): Total length 7.33. Carapace length 3.41, width 2.36, height 1.35. Clypeus length 0.19. Sternum length 1.79, width 1.31. Legs. Leg I: ti length 1.84/width 0.42, metatarsus (mt) 1.45. II: ti 1.74/0.40, mt 1.39. III: ti 1.59/0.38, mt 1.55. IV: ti 2.17/0.44, mt 2.41. Spines: legs I–II, femur d 1-1-1, p 2ap; tibia v 2-2-2; metatarsus v 2bas. III, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v 2r-1-2, p 1-d1-1-0 r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-0-2, p d1-1-0-1, r d1-1-0-1. IV=III but metatarsus v 2-2-2. Abdomen: length 4.03, width 2.33, spiracle–epigastrium 0.80. Color: carapace brown, with two wide, longitudinal darker band on sides, narrow in cephalic region. Ocular area on black patch with eyes bordered by black rings. Carapace with lateral region covered by fringe of white setae. Clypeus brown, low with abundant white setae. Chelicerae brown with black patches and fine setae on frontal edge. Endites, labium, and sternum dark brown, labium and endites light in apical region. Sternum with a very thin medial pale stripe. Legs coxae-tarsi dark brown, femora and patellae with longitudinal dark bands. All articles covered by white setae, abundant between coxae. Abdomen with dark brown pattern of spots on a gray background, dorsally with white medial band followed by 4-5 faint light chevrons; lateral regions covered by abundant small dots. Abdomen laterally surrounded by abundant white setae, with a thick bunch of setae on anterior region. Venter uniformly covered by dark and light patches. Epigyne: anterior plate sclerotized with deep and rounded anterior excavations, median notch quadrangular, posteriorly positioned (Fig. 10A, D). Median field transverse, large, oval in posterior view (Fig. 10C). Lateral lobes with sclerotized extensions wide, forming, posterior extensions rounded at base (Fig. 10A, C–D). Copulatory ducts long, forming two basal coils, anteriorly medially expanded, joining with the accessory bulb ducts at medial level. Spermathecae small, rounded, medially positioned in the epigyne, oriented towards longitudinal plane. Ducts of accessory bulbs short, wide, convergent (Fig. 10A–E).
Male. Unknown.
Distribution. Only known for Sierra Nevada de Santa Marta, Colombia (Fig. 31).
description: Published as part of Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy & Ramírez, Martin, 2025, Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America, pp. 201-242 in Zootaxa 5566 (2) on pages 213-216, DOI: 10.11646/zootaxa.5566.2.1, http://zenodo.org/record/14702734