Pantoclis globosa Brazidec & Vilhelmsen 2022, sp. nov.
Creators: name: Brazidec, Manuel
name: Vilhelmsen, Lars
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Diapriidae
Pantoclis
Pantoclis globosa

description: Pantoclis globosa sp. nov.
urn:lsid:zoobank.org:act: 4BF21BF6-7925-4F2A-8A84-5D8C49BBD8D7
Figs 1D, 3D–H, Table 1
Diagnosis
Eye glabrous; female antenna with: scape 5.5 times as long as wide, pedicel shorter than F1, F1 longest, F2–F13 as long as wide (Fig. 3D–E); male antenna with: scape 5.5 times as long as wide, pedicel rounded, F1 longest, emarginate on anterior third, F2–F12 subequal in length (Fig. 3G–H); mesosoma sparsely pubescent, shorter than metasoma; pronotum not elongate with epomia extending from pronotal shoulders to front coxa; notauli on mesoscutum complete and convergent posteriorly; propodeum with plicae; marginal vein shorter than radial cell and as long as three quarters of its distance to basal vein; radial cell closed; Rs inconspicuous proximally to r-rs (Fig. 1D); petiole as long as wide; gaster globous (Fig. 3F–G).
Etymology
The species name refers to the globular gaster of the species. The epithet is to be treated as an adjective.
Type material
Holotype NHMD-608414, a complete female but partially hidden by a milky coat; paratype NHMD- 608394, a complete male.
Locality and horizon
Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma.
Description
Female
BODY. BL = 2.83 mm. Head oval, subtriangular in frontal view (HeL = 0.48 mm), bearing several short hairs, stouter on frons; eye oval, glabrous; antenna inserted on frontal shelf, at level with lower margin of eyes; toruli well separated; antenna bearing numerous short sensilla; scape shorter than head length, 5.5 times as long as wide; pedicel shorter than F1, longer than wide; 13 flagellomeres; F1 longest, two times as long as wide; F2–F13 as long as wide; F13 tapering at apex (antennomeres length of female holotype, in mm: Sc-0.44; P-0.14; F1-0.16; F2-0.10; F3-0.10; F4-0.09; F5-0.09; F6-0.09; F7-0.09; F8- 0.08; F9-0.09; F10-0.09; F11- 0.09; F12-0.09; F13-0.10); mandibles of ordinary form; occipital carina foveate.
MESOSOMA. Smooth and shiny, shorter than metasoma (MsL = 0.97 mm), sparsely pubescent; pronotum not elongate with distinct and uninterrupted epomia; mesoscutum distinctly convex, notauli complete and convergent; anterior scutellar pit suboval, slightly narrower than apical distance between notauli; scutellum without posterior scutellar pit; propodeum with two prominent plicae; median propodeal keel simple. Fore wing hyaline, homogeneously micropubescent, extending beyond metasoma (FwL = 2.71 mm); C, Sc+R, M+Cu, basal vein, marginal vein, postmarginal vein and r-rs pigmented; Cu nebulous; marginal vein length around three quarters of its distance from basal vein; marginal vein shorter than radial cell; radial cell closed by Rs; Rs pigmented distally to r-rs, inconspicuous proximally. Legs slender, with short hairs; tibial spur formula 1-2-2; tarsal claws simple.
METASOMA. Petiole twice as long as wide (PtL = 0.27 mm; PtW = 0.12 mm), longitudinally striated, bearing long hairs dorsally; gaster globular, slightly longer than high (GL = 1.05 mm; GH = 0.67 mm), with sparse pubescence; S2 and T2 longest; four ring-like segments discernible beyond large tergite.
Male
BL = 3.50 mm; similar to female but antennae sexually dimorphic: scape shorter than head length (HeL = 0.53 mm), 5.5 times as long as wide, pedicel rounded, as long as wide, 12 flagellomeres elongate, cylindrical, longer than wide, F1 longest, with marked emargination on anterior third of segment, F2–F12 subequal in length, F12 tapering at apex (antennomeres length of male paratype, in mm: Sc- 0.38 mm; P-0.11; F1-0.25; F2-0.19; F3-0.19; F4-0.19; F5-0.18; F6-0.17; F7-0.16; F8-0.16; F9-0.16; F10-0.16; F11- 0.15; F12; 0.21).
Remaining measurements: HeL = 0.53 mm; MsL = 1.24 mm; FwL = 2.84 mm; PtL = 0.32 mm; PtW = 0.15 mm; GL = 1.41 mm; GH = 0.82 mm.
Comments
Using Nixon’s (1957) key, Pantoclis globosa sp. nov. keys out to Pantoclis because of the following characters: antennae inserted on a frontal prominence, mandibles not forming a beak or sickle-shaped, notauli present, scutellum without foveae along posterior margin, marginal vein nearly as long as its distance from basal vein and presence of ring-like segments beyond the large tergite, marginal vein shorter than radial cell, epomia distinct, median propodeal keel simple, petiole at most one and a third times as long as its apical width. Pantoclis globosa also fits in Pantoclis according to the diagnosis of Hou et al. (2016). It differs from most extant representatives of Pantoclis by having the marginal vein much longer than the r-rs (Nixon 1957; Sharma 1980; Buhl 1998). Pantoclis longiscapa Chambers, 1974 has a similar fore wing configuration but differs by having F1 very long and F2 distinctly longer than the following flagellomeres (Chambers 1974). Pantoclis deperdita Brues, 1906, from the Eocene of Florissant has a shorter radial cell and marginal vein (Brues 1906) and is therefore considered as a different species from P. globosa.
description: Published as part of Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, pp. 57-86 in European Journal of Taxonomy 813 on pages 68-69, DOI: 10.5852/ejt.2022.813.1733, http://zenodo.org/record/6468167