Tryphosella toowoomba Lowry &amp; Stoddart 2011, sp. nov.
Creators: name: Lowry, J. K.
name: Stoddart, H. E.
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Amphipoda
Tryphosidae
Tryphosella
Tryphosella toowoomba

description: <i>Tryphosella toowoomba</i> sp. nov. (Figs 44–46) <b>Type material.</b> HOLOTYPE, female, ovigerous (2 eggs), 4.4 mm, QM W29089, off Cape Moreton, Moreton Bay, Queensland, Australia, approx. 27°01'S 153°28'E, September 1995. PARATYPES: 1 male, 2.0 mm, QM W29090; 9 females, 5 males, QM W20919; same data as Holotype. <b>Type locality.</b> Off Cape Moreton, Moreton Bay, Queensland, Australia, approx. 27°01'S 153°28'E. <b>Etymology.</b> The species is named for the city of Toowoomba in southern Queensland. <b>Diagnostic description. <i>Head</i> lateral cephalic lobe rounded</b> , apically rounded; eyes apparently absent. <i>Antenna 1</i> peduncular article 1 without anterodistal lobe; article 2 without anterodistal lobe; accessory flagellum present, not forming cap, 3-articulate, terminal article not offset; flagellum with strong 2-field callynophore, robust setae absent from proximal articles; calceoli absent. <i>Antenna 2</i> peduncular article 3 long; articles 4 and 5 not enlarged; flagellum short, calceoli absent. <i>Epistome/Upper lip</i> separate; epistome produced beyond upper lip, broadly rounded; upper lip slightly produced, straight. <i>Mandible</i> molar with asymmetrically reduced column, proximally setose, distally triturating; palp attached about midway, article 3 with proximal A3-seta. <i>Maxilla 1</i> outer plate setal-tooth 7 present, setal-tooth 7, left and right symmetrical, cuspidate along most of sinusoidal inner margin; palp distal margin with apical robust setae. <i>Maxilliped</i> basis without recurved hook; outer plate with 1 short apical robust setae. <i>Gnathopod 1</i> subchelate; <b>coxa small (length 1.4 x breadth), subtriangular, tapering distally</b>; basis sparsely setose along anterior margin; ischium short; carpus long, subequal in length to propodus, without posterior lobe; propodus small, margins subparallel, sparsely setose along posterior margin, palm moderately acute, entire, straight. <i>Gnathopod 2</i> palm moderately obtuse. <i>Pereopod 4</i> coxa without distinct lateral ridge, with a well-developed posteroventral lobe. <i>Pereopod 5</i> coxa without distinct lateral ridge, without umbo; basis about as long as broad, without photophore, not posteroproximally excavate, posterior margin not serrate. <i>Pereopod 7</i> basis posterodistally produced less than halfway along merus, not posterodistally excavate. <i>Pleonites 1–3</i> without mid-dorsal carina, not produced dorsodistally. <i>Epimeron 3</i> posterior margin smooth, <b>posteroventral corner acutely produced</b>. <i>Urosomite 1</i> not projecting over urosomite 2, <b>with posteriorlydirected elongate rounded boss</b>. <i>Uropods 1 and 2</i> rami without stout striated robust setae. <i>Uropod 2</i> inner ramus without constriction. <i>Uropod 3</i> peduncle without dorsolateral flange, with 2 short apicomedial robust setae; inner and outer rami well developed, outer ramus article 2 short, with plumose setae on both rami. <i>Telson</i> deeply cleft, with dorsal robust setae, with 2 apical robust setae on each lobe. <b>Male</b> (sexually dimorphic characters). <i>Antenna 1</i> calceoli present, small. <i>Antenna 2</i> flagellum long, calceoli present. <i>Epistome/Upper lip</i> epistome slightly produced beyond upper lip, straight. <b> <i>Gnathopod 1</i> simple</b> ; margins tapering distally, palm absent. <b>Remarks.</b> <i>Tryphosella toowoomba</i> belongs to a group of four Australian species (<i>T. betka, T. martrudan, T. toowoomba</i> and <i>T. wangaratta</i>) in which the male gnathopod 1 is simple. Within this group <i>T. betka</i>, <i>T. martrudan</i> and <i>T. toowoomba</i> have a significantly reduced subtriangular coxa 1. <i>Tryphosella toowoomba</i> and <i>T. martrudan</i> both have an unusually shaped urosomite 1 forming a posteriorly-directed elongate rounded boss. <i>Tryphosella toowoomba</i> has many slight differences from <i>T. martrudan</i> (the slightly longer peduncular article 3 of antenna 2, the symmetrical maxilla 1 setal-tooth 7, the sparsely setose anterior margin of gnathopod 1 basis and the slightly more acute palm of gnathopod 1). The two species differ strongly in the presence of plumose setae on both rami of uropod 3 in the female <i>T. toowoomba</i> (none in <i>T. martrudan</i>) and the long antenna 2 of adult male <i>T. toowoomba</i> (short in <i>T. martrudan</i>). <b>Distribution.</b> South-eastern Queensland, Australia.
description: Published as part of <i>Lowry, J. K. &amp; Stoddart, H. E., 2011, The tryphosine genera Photosella gen. nov. and Tryphosella Bonnier, 1893 (Crustacea: Amphipoda: Lysianassoidea: Lysianassidae: Tryphosinae) in Australian waters 2956, pp. 1-76 in Zootaxa 2956 (1)</i> on pages 63-66, DOI: 10.11646/zootaxa.2956.1.1, http://zenodo.org/record/5281859