Thomomys (Megascapheus) umbrinus
Creators: name: Wilson, Don E.
name: Reeder, DeeAnn
Publisher: name: The Johns Hopkins University Press
Year: 2005
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Mammalia
Rodentia
Geomyidae
Thomomys
Thomomys umbrinus

description: Thomomys (Megascapheus) umbrinus (Richardson 1829)
[Geomys] umbrinus Richardson 1829, Quadrupeds, in: Fauna Boreali-Americana, Vol. 1: 202.
Type Locality: Originally given as Cadadaguios, SW Louisiana, but restricted to Mexico, Veracruz (probably Puebla), vicinity of Boca del Monte, by Bailey (1906).
Vernacular Names: Southern Pocket Gopher.
Subspecies::
Subspecies Thomomys (Megascapheus) umbrinus subsp. umbrinus (Richardson 1829)
Subspecies Thomomys (Megascapheus) umbrinus subsp. arriagensis Dalquest 1951
Subspecies Thomomys (Megascapheus) umbrinus subsp. atrodorsalis Nelson and Goldman 1934
Subspecies Thomomys (Megascapheus) umbrinus subsp. atrovarius (J. A. Allen 1898)
Subspecies Thomomys (Megascapheus) umbrinus subsp. camargensis Anderson 1972
Subspecies Thomomys (Megascapheus) umbrinus subsp. chihuahuae Nelson and Goldman 1934
Subspecies Thomomys (Megascapheus) umbrinus subsp. crassidens Nelson and Goldman 1934
Subspecies Thomomys (Megascapheus) umbrinus subsp. durangi Nelson and Goldman 1934
Subspecies Thomomys (Megascapheus) umbrinus subsp. emotus Goldman 1933
Subspecies Thomomys (Megascapheus) umbrinus subsp. enixus Nelson and Goldman 1934
Subspecies Thomomys (Megascapheus) umbrinus subsp. eximius Nelson and Goldman 1934
Subspecies Thomomys (Megascapheus) umbrinus subsp. extimus Nelson and Goldman 1934
Subspecies Thomomys (Megascapheus) umbrinus subsp. goldmani (Merriam 1901)
Subspecies Thomomys (Megascapheus) umbrinus subsp. intermedius Mearns 1897
Subspecies Thomomys (Megascapheus) umbrinus subsp. juntae Anderson 1972
Subspecies Thomomys (Megascapheus) umbrinus subsp. madrensis Nelson and Goldman 1934
Subspecies Thomomys (Megascapheus) umbrinus subsp. musculus Nelson and Goldman 1934
Subspecies Thomomys (Megascapheus) umbrinus subsp. nelsoni (Merriam 1901)
Subspecies Thomomys (Megascapheus) umbrinus subsp. newmani Dalquest 1951
Subspecies Thomomys (Megascapheus) umbrinus subsp. parviceps Nelson and Goldman 1934
Subspecies Thomomys (Megascapheus) umbrinus subsp. potosinus Nelson and Goldman 1934
Subspecies Thomomys (Megascapheus) umbrinus subsp. pullus Hall and Villa 1948
Subspecies Thomomys (Megascapheus) umbrinus subsp. sonoriensis Nelson and Goldman 1934
Subspecies Thomomys (Megascapheus) umbrinus subsp. supernus Nelson and Goldman 1934
Subspecies Thomomys (Megascapheus) umbrinus subsp. zacatecae Nelson and Goldman 1934
Distribution: SC Arizona and SW New Mexico (USA) south to Puebla and Veracruz (Mexico).
Conservation: IUCN – Lowered Risk (nt) as T. u. emotus, otherwise Lower Risk (lc).
Discussion: Subgenus Megascapheus. Hybridization with bottae reported in Patagonia Mtns of S Arizona by Patton and Dingman (1968), Patton (1973), and Hoffmeister (1969, 1986). Considered a species separate from bottae by these authors and by Anderson (1966, 1972). Hall (1981:469) included baileyi, bottae, and townsendii in this species based on his view of the hybridization data in Hoffmeister (1969), Patton (1973), and Thaeler (1968 b); he reported intergradation between umbrinus and bottae in Nuevo Leon (Mexico), which has not been verified by genetic analyses. Hafner et al. (1987) examined phylogenetic relationships among geographic units of umbrinus. Revised, in part, by Nelson and Goldman (1934 b), Anderson (1972; for Chihuahua), and Castro-Campillo and Ramírez-Pulido (2000; for C Mexico).
description: Published as part of Wilson, Don E. & Reeder, DeeAnn, 2005, Order Rodentia - Family Geomyidae, pp. 859-870 in Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore :The Johns Hopkins University Press on page 870, DOI: 10.5281/zenodo.7316535