Ganomymar De Santis 1972
Creators: name: Triapitsyn, Serguei V.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Mymaridae
Ganomymar

description: Genus Ganomymar De Santis, 1972
Figs 1–15
Ganomymar De Santis, 1972: 1–2
(type species: Ganomymar dessarti De Santis, 1972, by original designation).
Diagnosis
Both sexes
Vertex with a small pit-like depression next to each ocellus (Fig. 2C). Face without a pit next to each torulus. Mandible 3-dentate. Radicle very short, fused with the rest of scape. Pronotum entire; prosternum separated from head by propleura abutting each other anteriorly, and incompletely divided by a weak, short carina posteriorly; mesoscutum wider than long, with fairly wide notaular grooves ending anteriorly in small pits; scutellum separated posteriorly from frenum by transverse row of small foveae, and with campaniform sensilla closer to posterior margin than to anterior margin and very close to each other; frenum short, 0.2–0.33× length of scutellum; propodeum at posterior margin with a pair of small white puffs (spherical structures of unknown nature that are very translucent and thus not visible in slide-mounted specimens) on both sides of petiole attachment (best seen in dry-mounted specimens, Figs 7A, 10B). Tarsi 4-segmented. Petiole attached posteriorly to gastral sternum.
Female Antenna (Figs 2D, 6E, 8D, 14B) 9-segmented (funicle 6-segmented), with scape notably compressed laterally and smooth; F1 the shortest and F2 the longest funiculars and F3 the second longest, all 6 funiculars without mps; clava large, entire, with 6 mps arranged as follows: 2 about in the middle, and
2 subapical pairs. Macropterous or brachypterous. Ovipositor not or at most barely exserted beyond apex of gaster.
Male
Antenna (Figs 5C, 11A) much longer than body, 13-segmented (flagellum 11-segmented), with all flagellomeres much longer than wide and each with several mps. Macropterous. Digiti of genitalia (Figs 5B, 12B) without denticles apically.
The caslot species group
Both sexes macropterous (Figs 1, 3A, 4, 5D, 13, 14E); mesosoma (Figs 2E, 3B, 14D) smooth, with pronotum not enlarged and propodeum either without carinae or with a short median carina extending from posterior margin for at most half length.
The dessarti species group
Females brachypterous (Figs 6A–B, 7C–D, 9A–B) and males macropterous (Figs 10A, 12A); propodeum in both sexes with prominent submedian carinae (Figs 6D, 7A, 8C, 11C); female mesosoma at least partially reticulate (Figs 6D, 8B–C, E), with pronotum notably enlarged; male mesosoma (Fig. 11B–C) mostly smooth, with pronotum not enlarged.
Remarks
Ganomymar belongs to the Polynema Haliday, 1833 genus group, as defined by Lin et al. (2007). Its relationships to other members of the group are unclear, and without having any genetic evidence, it would be premature to make an educated guess. The genus has a unique feature, the propodeum with a pair of small white puffs at posterior margin at both sides of the petiole attachment in both sexes. This feature, and the peculiar fore wings, particularly in females, separates Ganomymar from all other members of the Polynema genus group. The four known species are placed in two distinct, informal species groups (two species in each), as defined above. The males (known only for one species in each group) are quite similar morphologically except for the markedly different configuration of the propodeal carina(e) in both sexes. That is quite unusual for the Polynema genus group in which presence or absence and the configuration of propodeal carina(e) when present, are often important diagnostic features used to distinguish the genera. This variability in propodeal carinae also occurs in species of Cremnomymar Ogloblin, 1952 (Mymaridae) from Juan Fernández Islands, Chile (Huber 2013).
Hosts and biology
Unknown.
Distribution
Afrotropical region: Madagascar.
Key to species of Ganomymar (females)
1 Mesosoma at least partially reticulate (Figs 6D, 8B–C, E), with pronotum notably enlarged and propodeum with prominent submedian carinae (Figs 7A, 8C); brachypterous (Figs 7C–D, 9A–B) (dessarti species group) .................................................................................................................... 2
– Mesosoma smooth (Figs 2E, 14D), with pronotum not enlarged and propodeum either without carinae or with a short median carina extending from posterior margin for at most half length (Fig. 2E); macropterous (Figs 3A, 14E) (caslot species group) ....................................................... 3
2 Head dark brown to black (Fig. 6A–B), vertex smooth (Fig. 6C); rest of body mostly light brown (Fig. 6A–B); fore wing narrow, without ‘cells’ beyond venation (Fig. 7C) ....................................... ................................................................................................................. G. dessarti De Santis, 1972
– Head mostly pale yellow (Fig. 9C), vertex with reticulate sculpture (Fig. 8B); rest of body mostly yellow (Fig. 9C); fore wing wide, with numerous round ‘cells’ beyond venation (Fig. 9A) ......................................................................................................... G. libertatium sp. nov.
3 Clava white, contrastingly lighter than funicle (Figs 1A, 2D), F2 about 6 × as long as wide; fore wing (Fig. 3A) with longest marginal seta at most 0.85 × greatest width of wing ...................................... .................................................................................................................................. G. caslot sp. nov.
– Clava brown, contrastingly darker than funicle (Figs 13B, 14B), F2 about 11 × as long as wide; fore wing (Fig. 14E) with longest marginal seta about 1.1 × greatest width of wing ................................ ............................................................................................................................. G. zuparkoi sp. nov.
description: Published as part of Triapitsyn, Serguei V., 2021, Revision of Ganomymar De Santis, 1972, a remarkable genus of Mymaridae (Hymenoptera, Chalcidoidea) from Madagascar, pp. 127-151 in European Journal of Taxonomy 757 on pages 128-130, DOI: 10.5852/ejt.2021.757.1417, http://zenodo.org/record/5075718