Goja Navás
Creators: name: Carrejo, Nancy
name: Obando, Ranulfo González
name: Aldrete, Alfonso Neri García
name: Mendivil, Julián
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Psocodea
Epipsocidae
Goja

description: Key to males of Goja
(Modified from Casasola González & García Aldrete, 2002)
G. bimaculata, G. pallida and G. trinidadensis, all by Casasola González & García Aldrete, 2002, are not included because only females are known. G. camachensis n. sp., is not included because it lacks the distal part of the hindwing. G. poblana Casasola González & García Aldrete is not included because the male genitalia is missing.
1. Hindwing with caeciliusid (forewing Rs forked, M three branched; hindwing M simple) venation (Figs 35, 74, 80, 113)... 2
- Hindwing Rs with two or more branches; M branched (Figs 2, 13)............................................. 11
2. Forewing R4+5 two-three branched (Fig. 73)............................................................... 3
- Forewing R4+5 unbranched (Figs 34, 79).................................................................. 5
3. Forewing R4+5 distally forked; phallosome concave anteriorly, anterior endophallic sclerites U shaped, with inner arms straight, slender and acuminate; outer arms broad, ribbon-like, distally acuminate (Fig. 131)....................................................................................................... Goja moyai Casasola G. & García A.
- Forewing R4+5 with three-four veins (Figs 73, 112); phallosome convex anteriorly, anterior endophallic sclerites not U shaped (Figs 78, 117)........................................................................................ 4
4. Lateral endophallic sclerites with two posterior processes; outer processes acuminate, curved inward; inner processes distally rounded; aedeagal arch with apical process acuminate (Fig. 78)............................ Goja pillcopatensis n. sp.
- Antero-lateral endophallic sclerites with two acuminate processes directed outward, mesal pair elongate and bent directed anteriorly, apices acuminate (Fig. 117).................................................. Goja yarumosensis n. sp.
5. Endophallic sclerites spiniform (Fig. 38).................................................................. 6
- Endophallic sclerites U-shaped (Fig. 120)................................................................. 9
6. Head vertex well below the upper border of the compound eyes, these projected anteriorly (Fig. 81); endophallic sclerites and aedeagal arch as in Fig. 84 ........................................................... Goja svanhildae n. sp.
- Head vertex straight or slightly convex; compound eyes not projected as above (Fig. 36); endophallic sclerites and aedeagal arch as in Fig. 38..................................................................................... 7
7. Apex of aedeagal arch pointed, with a distal process (Fig. 128)................. Goja mockfordi Casasola G. & García A.
- Apex of aedeagal arch concave, with two distal process (Fig. 38)............................................... 8
8. Posterior endophallic sclerites stout, wide based, distally with three small lobes (Fig. 138)............................................................................................. Goja spinosissima Casasola G. & García A.
- Posterior endophallic sclerites stout throughout, distally acuminate (Fig. 38)............... Goja cuasispinosissima n. sp.
9. Genae unpigmented; apex of aedeagal arch projected (Fig. 132)............... Goja obrienorum Casasola G. & García A.
- Genae with a pigmented area that reaches the antennal fossae; apex of aedeagal arch rounded (Fig. 120)............... 10
10. Anterior endophallic sclerites U-shaped, inner arms close together, about as long as the outer arms; epiproct with three mesal macrosetae....................................................... Goja brailovskyana Casasola G. & García A.
- Anterior endophallic sclerites U-shaped, inner arms separated, shorter than the outer arms, these distally forked (Fig. 120); epiproct with two mesal macrosetae.................................... Goja curvata Casasola G. & García Aldrete
11. Hindwing R4+5 unbranched, M two branched (Figs 51, 57, 68)............................................... 12
- Hindwing R4+5 branched or not, M three-six branched (Figs 13, 45)........................................... 24
12. Phallosome with side struts strongly curved; aedeagal arch extended posteriorly a distance, aedeagal apex broad and rounded; mesal sclerites elongate and curved, each one ended in an acute apex, joined to endophallus by long, twisted sclerotic bands......................................................................... Goja cuspidata Casasola González
- Phallosome with side struts straight or not strongly curved as above; aedeagal arch not extended a distance posteriorly or apex not widened as above, mesal sclerites not as above (Figs 105, 111)............................................. 13
13. Head without a pigmented spot posterior to antennal fossae.................................................. 14
- Head with a pigmented spot posterior or above the antennal fossae (Fig. 108).................................... 19
14. Head with two ochre transverse bands, one between the compound eyes, through the ocellar triangle, and one between the genae, through the postclypeus (Fig. 102); forewings R2+3 and R4+5 branched as illustrated (Fig. 100)............... 15
- Head without transverse bands as above (Figs 52, 69); forewing R2+3 forked or not, R4+5 forked or dichotomously branched (Figs 50, 67)........................................................................................ 16
15. Forewings with R2+3 and Rs, after the R-M crossvein subequal; posterior endophallic sclerites robust, each with two small apical hooks; mesal endophallic sclerites each with a broad, longitudinal area, with a row of spines of unequal length on outer edge, directed outward, and a mesal spine on inner edge, directed anteriorly (Fig. 141)..................................................................................................... Goja vittata Casasola G. & García A.
- Forewings with R2+3 almost twice as long as Rs after the R-M crossvein (Fig. 94); posterior endophallic sclerites not as robust as above, apices rounded, without hooks; mesal endophallic sclerites each a broad, semioval area, without a row of spines as above (Fig. 99)..................................................................... Goja tacanaensis n. sp.
16. Forewings with R2+3 as in Figs 50 and 67; phallosome as in Figs 55 and 72..................................... 17
- Forewings with R2+3 as in Fig. 56; phallosome variable (Fig. 61)............................................. 18
17. Mesal endophallic sclerites of three to four short acuminate teeth directed outward; lateral endophallic sclerites asymmetric, wide proximally, with distally acuminate elongate process curved outward (Fig. 55); aedeagal arch V-shaped, apically acuminate............................................................................ Goja horquetensis n. sp.
- Anterior pair of endophallic sclerites basally broad, irregular, each with a slender, long process on the inner side, directed posteriorly; posterior pair of endophallic sclerites well sclerotized, proximally wide, anterior border concave, with truncate process directed outward, distally with acuminate process curved outward (Fig. 72); aedeagal arch rounded laterally, median process truncate........................................................................ Goja munchiquensis n. sp.
18. Aedeagal arch robust, distally with a large, acuminate process; lateral endophallic sclerites broad, bent inward (Fig. 124)....................................................................... Goja chilensis Casasola G. & García A.
- Aedeagal arch with posterior triangular process; lateral pair of endophallic sclerites kidney-shaped, with truncate posterior border; mesal pair with two elongate, acuminate processes directed outward (Fig. 61)............ Goja korytkowskii n. sp.
19. Forewing R2+3 forked; R4+5 simple, a proximal crossvein between them (Fig. 106); anterior pair of endophallic sclerites wide basally, semicircular, with posterior acuminate processes long and widely separated; posterior pair of endophallic sclerites oval (Fig. 111)......................................................................... Goja toleditensis n. sp.
- Forewing R2+3 simple; phallosome variable.............................................................. 20
20. Forewing Rs about half as long as R2+3; lateral endophallic sclerites long, distally acuminate, each with two mesal spines directed outward, associated anteriorly to a transverse sclerite, each with an acuminate spine directed anteriorly (Fig. 125)................................................................... Goja longispinosa Casasola G. & García A.
- Forewing Rs as long or longer than R2+3; phallosome with endophallic sclerites variable.......................... 21
21. Pigmented genal spots joined to the spots posterior to the antennal fossae; areola postica and M3 joined by a crossvein... 22
- Pigmented genal spots separated from the spots posterior to the antennal fossae; areola postica and M3 not joined by a crossvein.............................................................................................. 23
22. Apex of aedeagal arch with two acuminate distal processes; anterior pair of endophallic sclerites as vertical, membranous bands (Fig. 118)......................................................... Goja acuphallica Casasola G. & García A.
- Apex of aedeagal arch with one rectangular distal process (Fig. 133), anterior pair of endophallic sclerites with anterior processes stout, distally acuminate, posterior processes forked, robust, acuminate distally................................................................................................. Goja ochoterenae Casasola G. & García A.
23. Apex of aedeagal arch wide, truncate, projected posteriorly (Fig. 121), posterior endophallic sclerites wide at base, acuminate distally, each apex with two small hooks.................................. Goja eertmoedi Casasola G. & García A.
- Apex of aedeagal arch rounded, not projected posteriorly (Fig. 136), posterior endophallic sclerites slightly dilated anteriorly, slender, apices without small hooks.................................... Goja omiltemensis Casasola G. & García A.
24. Forewings strongly pigmented, with large transverse hyaline areas; hindwing M three-five branched (Figs 12, 44); two pairs of endophallic sclerites, mesal pair broadly inverted U-shaped (Figs 17, 49)........................................ 25
- Forewings unpigmented (Fig. 23), if pigmented, then without hyaline areas as large as above; hindwing M three-four branched (Fig. 24); phallosome variable.......................................................................... 26
25. Hindwing M five branched (Fig. 13); anterior endophallic sclerites small, slender, distally acuminate, curved outward; mesal endophallic sclerites inverted U-shaped, with outer arms acuminate (Fig. 17)..................... Goja caucana n. sp.
- Hindwing M three-four branched (Fig. 45); mesal endophallic sclerites broadly inverted U-shaped, outer arms short, with a row of transverse lines along outer border; inner arms long, acuminate, slightly curved outward (Fig. 49)........................................................................................................ Goja galarzai n. sp.
26. Forewing areola postica and M3 joined by one or several crossveins (Fig. 23); hindwing R2+3 or R4+5 branched (Fig. 24).. .................................................................................................. 27
- Forewing areola postica and M3 not joined by crossveins; hindwing R2+3 unbranched (Figs 2, 40)................... 30
27. Forewing R2+3 without vein stubs; anterior endophallic sclerites broad, long; inner edge of each with a mesal, robust projection acuminate distally (Fig. 139)............................................... Goja ticana Casasola G. & García A.
- Forewing R2+3 with vein stubs (Fig. 23); anterior endophallic sclerites not as above (Fig. 27)....................... 28
28. Forewing R2 with one vein stub (Fig. 23); aedeagal arch with short, narrow apical process (Fig. 27); mesal endophallic sclerites with four pairs of spines, slender and distally acuminate, two anterior straight and directed outward, two short posterior pairs curved, acuminate............................................................... Goja chiquihuitensis n. sp.
- Forewing Rs and R2+3 with several vein stubs; aedeagal arch apex wide, truncate, or blunt, spines of endophallic sclerites not as above........................................................................................... 29
29. Aedeagal arch apex wide, truncate, with two small distal processes; mesal endophallic sclerites robust, distally with one denticle (Fig. 137), anterior pair slender, sinuous, wide basally, associated to a broad radula.................................................................................................. Goja reticulata Casasola G. & García A.
- Aedeagal arch apex blunt, without terminal processes; mesal endophallic sclerites distally with two denticles, anterior pair broad, irregular, with acicular thickenings anteriorly...................... Goja reticulatoides Casasola G. & García A.
30. Hindwing R4+5 unbranched (Figs 2, 40)................................................................. 31
- Hindwing R4+5 branched (Fig. 29)...................................................................... 35
31. Hindwing R4+5 almost twice as long as Rs after the R-M crossvein................ Goja garcialdretei Casasola González
- Hindwing R4+5 subequal in length to Rs after the R-M crossvein (Figs 2, 40).................................... 32
32. Aedeagal arch apex rounded, without processes (Fig. 41), one pair of endophallic sclerites, anteriorly wide, elongate, acuminate apically and slightly curved inward...................................................... Goja farallones n. sp.
- Aedeagal arch with apical processes pointed (Fig. 6); endophallic sclerites variable................................ 33
33. Phallosome longer than wide, with four pairs of acuminate endophallic sclerites, anterior pair directed caudally and curved outward, the other three pairs directed outward; side struts straight (Fig. 6)..................... Goja caldasensis n. sp.
- Phallosome almost as long as wide, with posterior endophallic sclerites caudally projected, side struts curved (Figs 123, 126) .................................................................................................. 34
34. Apex of posterior endophallic sclerites acuminate; anterior endophallic sclerites horizontal, anteriorly wide, acuminate, directed outward (Fig. 123)............................................... Goja herrerae Casasola G. & García A.
- Apex of posterior endophallic sclerites with two denticles (Fig. 126), anterior endophallic sclerites broad, transverse, pointed at both ends, associated to two mesal ones, inverted Y-shaped.................. Goja martinezi Casasola G. & García A.
35. Pterostigma without pigmented bands; anterior endophallic sclerites directed posteriorly, acuminate, each arm with six pointed processes directed outward (Fig. 134)..................................... Goja ocotensis Casasola G. & García A.
- Pterostigma pigmented or with one-two pigmented bands (Fig. 94); endophallic sclerites variable.................... 36
36. Pterostigma fully pigmented (Fig. 100); anterior endophallic sclerites small, wide at base, acuminate distally, slender and curved outward (Fig. 105).............................................................. Goja tamensis n. sp.
- Pterostigma with pigmented bands, endophallic sclerites variable (Fig. 28)...................................... 37
37. Pterostigma with one pigmented band................................................................... 38
- Pterostigma with two pigmented bands (Fig. 28)........................................................... 39
38. Aedeagal arch apex blunt, without distal processes, posterior endophallic sclerites absent, anterior pair membranous (Fig. 119) ….............................................................. Goja circumvittata Casasola G. & García A.
- Aedeagal arch apex with an acuminate posterior process, and a blunt anterior process; anterior endophallic sclerites membranous, with two posterior, slender projections, on each side of mesal line (Fig. 140).................................................................................................. Goja venezuelensis Casasola G. & García A.
39. Forewing Rs, after R-M crossvein, one fourth or less the length of R2+3 (Fig. 28)................................. 40
- Forewing Rs, after R-M crossvein, about a third the length of R2+3............................................ 42
40. Aedeagal arch without apical process..................................... Goja medellini Casasola G. & García A.
- Aedeagal arch with short apical process (Fig. 33).......................................................... 41
41. Phallosome with mesal endophallic sclerites acuminate distally, curved back over slightly sclerotic area (Fig. 122)........................................................................ Goja guatemalensis Casasola G. & García A.
- Phallosome with mesal endophallic sclerites directed forward, distally acuminate, not curved and directed backward as above (Fig. 33)................................................................... Goja cuasiguatemalensis n. sp.
42. Forewing R4+5 two branched............................................................. Goja ditata Navás
- Forewing R4+5 more than two branched................................................................. 43
43. Phallosome with posterior pair of endophallic sclerites more or less parallel, each with three digitiform small processes distally (Fig. 129); hindwing M eight branched..................................... Goja montieli Casasola G. & García A.
- Phallosome with posterior pair of endophallic sclerites diverging and acuminate distally (Fig. 130); hindwing M four- five branched.......................................................................................... 44
44. Phallosome with posterior endophallic sclerites elongate and curved, each one ended in two conical acute apices, without spine basally (Fig. 130)......................................................... Goja bicuspidata Casasola González
- Phallosome with posterior endophallic sclerites not as above, each with a stout anterior spine, postero-laterally directed, curved inward (Fig. 135)................................................... Goja oculatissima Casasola G. & García A.
description: Published as part of Carrejo, Nancy, Obando, Ranulfo González, Aldrete, Alfonso Neri García & Mendivil, Julián, 2021, New species of Goja Navás (Psocodea: ' Psocoptera': Epipsocidae) from Colombia Mexico and Peru, pp. 501-541 in Zootaxa 4903 (4) on pages 502-505, DOI: 10.11646/zootaxa.4903.4.3, http://zenodo.org/record/4431309