Phylloicus munozi Prather 2003, new species
Creators: name: PRATHER, AYSHA L.
Publisher: name: Zenodo
Year: 2003
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Calamoceratidae
Phylloicus
Phylloicus munozi

description: Phylloicus munozi, new species
Figs. 74, 75
Phylloicus “species 2" Flint, 1991:99.
Phylloicus munozi is similar to P. blahniki, but is distinguished by the unmodified tergum III, the absence of posterior processes of tergum IV and a complex lateral sclerite (Fig. 74F), and the long digitate basodorsal process of tergum X (Fig. 74A, B).
Adult. Forewing length 9.7­10.5 mm, n = 17.
Head golden brown. Maxillary palps dark brown. Antenna twice forewing length; dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax golden brown; ventrolateral thorax pale tan. Legs pale tan. Metathoracic leg of male with posterior fringe of long setae, setae dark. Tibial spur formula 2,4,4. Forewing flat; dark brown; with two transverse bands; proximal band pale orange­yellow, reaching posterior wing margin, at least 1/2 width of wing, narrow, less than 10 setae wide; distal band pale orange­yellow, not reaching either wing margin, 1/2 width of wing or less, narrow, less than 10 setae wide; with two basal stripes, ivory; cell Cu and base of cell Cu 2 clear. Hind wing basal brush present in male.
Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum IV with paired lateral sclerites, mesal coremata and lateral coremata; lateral sclerite spatulate, directed laterally and with acute basomesal process; lateral coremata with basal globose lobes and long tubular posterior lobe; mesal coremata single­lobed, setose, capable of inflation beyond terminalia. Tergum V without sclerotized modifications (Fig. 74F). Sternum VII with short, acute anteromesal process. Sternum VIII similar to anterior sterna, sternum IX not elongate. Tergum IX without mesal ridge; posterior margin continuous with basodorsal process of tergum X (Fig. 74B); lateral ridge present; dorsal pleural setae approximately 2, ventral pleural setae absent (Fig. 74A); sternum IX without mesolateral ridges; sternum IX (Fig. 74C). Preanal appendage approximately length of tergum X, of uniform diameter throughout length, setae long, but not filamentous or longer than appendage (Fig. 74A, B). Tergum X without basal lobes; basodorsal process long and digitate; basolateral processes absent; apex, in lateral view, truncate, in dorsal view, entire (Fig. 74A, B). Harpago slightly tapered; peglike setae many, apical (Fig. 74A, C). Phallic endotheca with paired apicolateral lobes and paired basolateral lobes, basolateral lobes tapered apically, apicolateral lobes large and rounded; phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped (Fig. 74D, E).
Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII without posterolateral brush; sternum VIII posterior margin entire; sternum VIII (Fig. 75C). Tergum IX without mesal ridge (Fig. 75B). Sternum IX anterior and posterior lobes darkly sclerotized and striate, with patch of lightly sclerotized cuticle lateral to vaginal opening (Fig. 75A). Tergum X appendage shorter than mesal lobe, base marked by faint suture line, apex oblique; mesal lobe lightly sclerotized; digitate lateral processes absent (Fig. 75B). Sternum X with patches of short fine setae posterolaterally to anal opening (Fig. 75A). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite rounded anteriorly, posterolateral projections absent; posterior sclerite triangular (Fig. 75A).
Holotype male: COSTA RICA: Cartago: Reserva Tapantí, Quebrada Palmitos & falls, ca. 9 km (road) NW tunnel, 09°43'12”N, 83°46'48"W, 1400 m, 1­2.viii.1990, Holzenthal, Blahnik, & Muñoz (UMSP).
Paratypes: COLOMBIA: Antioquia: Mun. Envigado, Quebrada La Ayura, 1750 m, 2.xii.1983, Matthias — 1 female (NMNH); site B, 1750 m, 9.xii.1983, Matthias — 2 females (NMNH); trap B, 1750 m, 20.iv.1983, Matthias — 1 male (NMNH); 23.viii.1983, Matthias — 2 males (NMNH); COSTA RICA: Límon: Limon, 16 km W Guapiles, 400 m, Ii.­ Mar 1989, Hanson — 1 male (UMSP); Puntarenas: Río Bellavista trib., Las Alturas, road to quarry, 08°57'07”N, 82°50'53"W, 1480 m, 19.iii.1991, Holzenthal, Muñoz, & Huisman — 1 male (INBIO); — 1 male (UMSP); Río Jaba, rock quarry, 1.4 km (air) W Las Cruces, 08°47'24”N, 82°58'12"W, 1150 m, 9.viii.1990, Holzenthal, Blahnik, & Muñoz — 1 male (UMSP); 15.iii.1991, Holzenthal, Muñoz, & Huisman — 2 males (UMSP); Jardín Botanico R & C Wilson, unnamed trib., Sendro del Agua, 08°48'00”N, 82°57'36"W, 1180 m, 8.viii.1990, Holzenthal, Blahnik, & Muñoz — 2 males (UMSP); San José: Reserva Biológica Carara, Río del Sur, 1.5 km (rd) S Carara, 09°46'08”N, 84°31'52"W, 160 m, 13.iii.1991, Holzenthal, Muñoz, & Huisman — 1 female (UMSP); PANAMA: Chiriqui: Fortuna Dam Site nr. Hornitos, 08°55'00”N, 82°16'00"W, 1050 m, 4.i.­ 7.iii.1978, Wolda — 1 male (NMNH).
Distribution. Colombia, Costa Rica, Panama.
Etymology. This species is named for Fernando Muñoz­Quesada, who collected the type specimen.
description: Published as part of PRATHER, AYSHA L., 2003, Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae), pp. 1-214 in Zootaxa 275 (1) on pages 75-77, DOI: 10.11646/zootaxa.275.1.1, http://zenodo.org/record/5019235