Clusiodes Coquillett
Creators: name: Owen Lonsdale
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Clusiidae
Clusiodes

description: Genus CLUSIODES Coquillett
Heteroneura Fallén 1823: 2 [preoccupied by Heteroneura Fallén 1810) (Platypezidae)]. Type species, Heteroneura albimana Meigen, 1830, subsequent designation by ICZN 2010: 344 (Opinion #2262). Gender feminine. Meigen 1830: 126 [redescription, listed in error as independent proposal in Neave (1939b: 640)]; Curtis 1837: column 274 [checklist, Britain], 1838: plate 721 [albimana Meigen 1830 designated as type species, redescription, diagnosis, Britain]; Haliday 1838: 188 [Britain]; Zetterstedt 1838: column 784 [redescription, designation of Heteroneura muscaria Fallén 1823 (= Cnemacantha muscaria (Fallén), Lauxaniidae) as type species], 1848: 2787 [Heteroneura geomyzinus Fallén 1823 designated as type species, redescription], 1852: 4816 [Scandinavia]; Westwood 1840: 152 [Heteroneura albimana Meigen 1830 listed as type species, diagnosis, Britain]; Agassiz 1846: 519 [nomenclator, attributed to Meigen (1830)]; Rossi 1849: 78 [checklist, Austria]; Wilson 1855: 256 [description]; Loew 1862: 42 [catalogue, North America]; Osten Sacken 1878: 197 [catalogue, North America]; Schiner 1862: 37 [redescription, key to species, Austria], 1864: 72 [catalogue, Europe]; Siebke 1877: 181 [catalogue, Norway]; Osten Sacken 1878: 197 [catalogue, North America]; Williston 1896: 386 [key, St. Vincent], 1908: 319 [key, illustrations of head and wing]; Strobl 1897: 27 [catalogue, Transylvania]; Becker 1902: 313 [reexamination of Meigen types], 1905: 38 [catalogue, Palaearctic]; Czerny 1903: 73 [family and genus revision]; Aldrich 1905: 571 [catalogue, North America]; Kertész 1906: 320 [key]; Hendel 1910: 309 [nomenclature]; Schulze et al. 1930: 1540 [nomenclator]; Neave 1939b: 640 [nomenclator]; Lonsdale 2009a: 144 [proposal to ICZN]; ICZN 2010: 344 (Opinion #2262) [designation of H. albimana Meigen, 1830 as type species]).
Heteronevra. Macquart 1835: 587 [diagnosis, key]. Unjustified emendation of Heteroneura Fallén 1823. Blanchard 1840: 628 [Heteroneura Nubila Meigen (= albimana Meigen) designated as type species]; Scudder 1882: 161 [nomenclator], 1884: 148 [nomenclator]; Schulze et al. 1930: 1540 [nomenclator]; Evenhuis et al. 2016: 70 [nomenclatural status, as nomen oblitum, junior synonym of Clusiodes].
Clusiodes Coquillett 1904: 93 [replacement name for Heteroneura Fallén 1823]. Type species, Heteroneura albimanus Meigen 1830, automatic designation. Gender masculine. Coquillett 1910: 525 [type species catalogue, North America]; Johnson 1913b: 97 [family and genus revision, eastern United States], 1925a: 245 (as genus and subgenus) [catalogue, New England]; Wahlgren 1917: 186 [review, Sweden]; Malloch 1918: 5 [key to species], 1922: 47 (as genus and subgenus) [key to subgenera and species]; Melander & Argo 1924: 13 [family and genus revision, key to subgenera and species]; Czerny 1928: 6 (as genus and subgenus) [family and genus revision, Palaearctic]; Schulze et al. 1927b: 730 [nomenclator]; Stackelberg 1933: 381 [key, European USSR], 1988: 505 [key, European USSR]; Tuomikoski 1933: 15 [Finland]; Séguy 1934: 351 [France]; Neave 1939a: 770 [nomenclator]; Kloet & Hincks 1945: 394 [checklist, Britain]; Hennig 1952: 227 [immature stages]; Frey 1960: 18 [family and genus revision, key, Indoaustralia, catalogue, world]; McAlpine 1960: 81 [family and genus revision, Australia—all species presently recognized as Hendelia]; Trojan 1962: 22 (as genus and subgenus) [key to subgenera and species, Poland]; Soós 1964a: 7 [key], 1984: 11 (as genus and subgenus) [catalogue, Palaearctic], 1987: 857 [key, Nearctic]; Steyskal 1965: 807 (as genus and subgenus) [catalogue, America north of Mexico]; Cole & Schlinger 1969: 426 [illustration of head, key to genera of Clusiidae, western North America]; Sasakawa 1977: 236 (as genus and subgenus) [catalogue, Orient], 1987: 100 (as genus and subgenus) [revision, Orient/Pacific], 1998b: 223 [key to subgenera] 2011b: 20 [Oriental fauna]; Stubbs 1982: 89 (as genus and subgenus) [identification guide, biology, Britain]; Ferrar 1987: 121 [notes on biology]; Balthazar 1990: 416 [catalogue, Philippines, only included species Hendelia aberrans (Frey)]; Woznica 1991: 208 [checklist, Poland]; DeBruyn 1991: 146 [catalogue, Belgium]; Caloren & Marshall 1998: 263 [redescription, biology, New world revision, phylogeny, synonymy of Labomyia Frey (now synonymized with Hendelia), Clusiaria Malloch and Columbiella Malloch, illustration of male and female genitalia, photos]; Marshall 2000: 4 [synonymy of Parahendelia McAlpine]; Beuk 2002: 271 (as genus and subgenus) [checklist, Netherlands]; Greve 2005: 3 [Norway]; Sueyoshi 2006: 6 [Japan], 2014: 618 [catalogue, Japan]; Papp et al. 2006: 190 [identified and unidentified species, Thailand]; Lonsdale & Marshall 2007b: 123 [redefinition, maps, review, phylogeny, biology, taxonomic history, zoogeography, data on Nearctic species, discussion of C. melanostomus complex, illustration of immature stages, Old world, incorrect treatment of Gioenia Lioy as synonym], 2010: 1044 [Manual of Central American Diptera]; Lonsdale et al. 2010: 267 [family revision, key]; ICZN 2010: 344 [conservation of Clusiodes in Clusiidae by setting aside all previous type fixations and by designating Heteroneura albimana Meigen 1830 as type species for Heteroneura Fallén 1823]; Kahanpää 2014: 288 [checklist, Finland]; Evenhuis et al. 2016: 70 [nomenclatural status, nomen protectum].
Clusiaria Malloch 1922: 49 (as subgenus) [key to subgenera and species, notes]. Type species, Clusiodes duplicata Malloch 1922, by original designation (= melanostomus Loew). Gender feminine. Johnson 1925a: 245 [catalogue, New England]; Czerny 1928: 6 [key, Palaearctic]; Neave 1939a: 770 [nomenclator]; Kloet & Hincks 1945: 394 [checklist, Britain]; Frey 1960: 16 [key to subgenera and species, Indoaustralia, catalogue, world]; Trojan 1962: 22 [key, Palaearctic]; Stubbs 1982: 89 [key, Britain]; Soós 1984: 12 [catalogue, Palaearctic]; Sasakawa 1987: 101 [key, Orient/Pacific]; Schumann et al. 1999: 144 [checklist, Germany]; Caloren & Marshall 1998: 263 [as synonym of Clusiodes]; Beuk 2002: 271 [checklist, Netherlands].
Columbiella Malloch 1922: 49 [as subgenus]. Type species, Clusiodes apiculata Malloch 1922, by original designation. Gender feminine. Johnson 1925a: 246 [catalogue, New England]; Czerny 1928: 9 [Palaearctic]; Neave 1939a: 806 [nomenclator]; Kloet & Hincks 1945: 394 [checklist, Britain]; Frey 1960: 26 [catalogue, world]; Trojan 1962: 26 [key, Poland]; Steyskal 1965: 807 [catalogue, America north of Mexico]; Stubbs 1982: 89 [key, Britain]; Soós 1984: 12 [catalogue, Palaearctic]; Sasakawa 1987: 101 [key, Orient/Pacific]; Caloren & Marshall 1998: 263 [as synonym of Clusiodes]; Beuk 2002: 271 [checklist, Netherlands].
Clusioides. Misspelling of Clusiodes. Curran 1934b: 363 [key]; Soós 1981: 97 [Hungary, key]. Clusinodes. Misspelling of Clusiodes. Craighead 1950: 541 [as subgenus within Sobarocephala].
albimanus (Meigen). PA: Europe, Russia.
Heteroneura geomyzina var. b Fallén 1823: 3. Sweden. ST ♂♀ [lost]. Zetterstedt 1838: 790 [redescription]; Zetterstedt 1848: 2787 [as synonym of albimana Meigen].
Heteroneura albimana Meigen 1830: 128. Switzerland. Geneva Canton: Bernex, 415m. NT ♂ MHNG [designated by Lonsdale & Marshall 2007b: 129]. Curtis 1837: column 274 [checklist, Britain], 1838: plate 721 [designated as genus type, synonymy of scotica Curtis, illustrations of head, habitus, Britain], 1855: 6403 [bred from rotten willows]; Boie 1847: 331 [discussion of immature stages]; Zetterstedt 1848: 2790 [synonymy of geomyzina var. b Fallén and geomyzina var. c Zetterstedt, Sweden], 1855: 4817 [Sweden]; Curtis 1855: 4603 [notes on insect economy]; Rossi 1849: 78 [checklist, Austria]; Wulp & Snellen van Vollenhoven 1856: 114 [checklist, Netherlands]; Loew 1857: 51 [Germany], 1864b: 339 [synonymy of albimana var b Zetterstedt, redescription, discussion, Germany]; Schiner 1862: 38 [redescription, key, biology, Austria], 1864: 72 [catalogue, Europe]; Brauer 1883: 92 [larva in beetle burrows in wood, records of Vienna Museum]; Girschner 1888: 97 [comparison to alpina Loew (= geomyzinus Fallén)]; Becker 1902: 313 [examination of Meigen types], 1905: 38 [catalogue, Palaearctic]; Czerny 1903: 81 [redescription and biology]; Czizek 1910: 112 [catalogue, Germany]; Haliday 1933: 150 [England]; Lonsdale 2009a: 144 [proposed as type species]; Lonsdale et al. 2010: 240 [genes sequenced for phylogenetic analysis]; Lonsdale & Marshall 2010: 1042 [reproduction of figure from Soós (1987)]; ICZN 2010: 344 [conservation of Clusiodes in Clusiidae by setting aside all previous type fixations and by designating Heteroneura albimana Meigen, 1830 as type species of Heteroneura Fallén 1823].
Heteroneura nubila Meigen 1830: 127. Europe [not given]. ST ♂ MNHNP [lost, probably destroyed]. Rossi 1849: 78 [checklist, Austria]; Wilson 1855: 257 [description, records from Austria and Europe]; Loew 1857: 53 [Germany], 1864b: 343 [as junior synonym of Heteroneura ruficollis Meigen 1830]; Becker 1902: 313 [examination of Meigen material], 1905: 39 [catalogue, Palaearctic]; Czerny 1903: 76 [redescription].
Heteronevra nubile. Macquart 1835: 587 [illustration of head, France].
Heteronevra albimana. Macquart 1835: 588 [France].
Heteroneura scotica Curtis 1837: column 274 [nomen nudum, Britain]; Curtis 1838: pl. 101 [as synonym of albimana Meigen, Britain].
Agromyza geomyzina. Misidentification. Zetterstedt 1838: 790 [Sweden].
Agromyza geomyzina var c Zetterstedt 1838: 790. Sweden. ST ♀ [lost - not found in ZIN]. Zetterstedt 1848: 2787 [as synonym of albimana Meigen].
Heteronevra Nubila. Blanchard 1840: 628 [designated as type of Heteroneura Fallén 1823, France].
Heteroneura albimana var b Zetterstedt 1852: 4817. Sweden. Lund. ST ♀ [lost]. Loew 1864b: 339 [as synonym of albimana Meigen].
Heteroneura albimana var. pallidior Loew 1857: 52 [as “ albimana var. α”]. Germany. Type information unknown. Loew 1864b: 340 [includes albimana sensu Meigen (1830), Zetterstedt (1848) and Schiner (1862), synonymy of albimana var. α Loew]; Czerny 1903: 81 [suggests synonymy of varieties]; Lonsdale & Marshall 2007b: 129 [as synonym of albimana Meigen].
Heteroneura albimana var. obscurior Loew 1857: 52 [as “ albimana var. β”]. Germany. Type information unknown. Loew 1864b: 339 [includes albimana var. b Fallén and pictipes Zetterstedt sensu Schiner (1862), synonymy of albimana var. β Loew]; Czerny 1903: 81 [suggests synonymy of varieties]; Lonsdale & Marshall 2007b: 129 [as synonym of albimana Meigen].
Heteroneura pictipes. Misidentification. Schiner 1862: 37 [Moravia].
Heteroneura albimanus. Aldrich 1905: 570 [gender disagreement, catalogue, North America].
Clusiodes (Clusiodes) albimana. Johnson 1913b: 98 [generic combination, Nearctic records - misidentification of johnsoni Malloch]; Melander & Argo 1924: 15 [includes forms pallidior Loew and obscurior Loew, photo, notes type likely destroyed]; Frey 1928c: 109 [catalogue, Finland], 1960: 26 (in part) [catalogue, world]; Kloet & Hincks 1945: 394 [checklist, Britain]; Trojan 1962: 25 [key, illustrations of male genitalia, Poland]; Chandler 1978: 264 [Ireland]; Hackman 1980: 148 [checklist, Finland]; Stubbs 1982: 92 [Britain]; Greve & Midtgaard 1986: 89 [Norway].
Clusiodes albimana. Wahlgren 1917: 187 [diagnosis, key, photo]; Czerny 1928: 8 [redescription, illustrations of thorax, wing, head and male genitalia]; Stackelberg 1933: 381 [key, European USSR], 1988: 505 [key, European USSR]; Séguy 1934: 352 [illustrations of head and male genitalia, photo of wing, France]; Richards 1934: 2 [puparia in rotting birch trunk]; Tuomikoski 1936: 184 [Finland]; Hennig 1952: 227 [immature stages]; Chandler 1973: 340 [associated with elm, England]; Withers 1985: 63 [bred from Fraxinus excelsior, associated with Fagus sylvaticus, England]; Greve & Midtgaard 1986: 89 [Norway]; Ferrar 1987: 122 [immature stages, illustration of larva and puparium]; Zuska 1987: 235 [Czech Republic]; Smith 1989: 103 [reared from old logs and mossy tree stumps, illustrations of puparium, Britain]; Dunk 1996: 422 [catalogue, Bavaria]; Yagi & Sawada 2000: 41 [Mamaev collection holdings].
Clusiodes (Clusiodes) albimana form pallidior. Melander & Argo 1924: 15 [in part, misidentification in part of johnsoni Malloch].
Clusiodes (Clusiodes) albimana form obscurior. Melander & Argo 1924: 15 [in part].
Clusiodes (Clusiaria) nubila. Melander & Argo 1924: 21 [synonymy, generic combination, distribution, comparison to ruficollis Meigen]; Czerny 1928: 7 [redescription, central Europe]; Frey 1960: 26 [catalogue, world]; Trojan 1962: 22 [key, Poland]; Soós 1984: 12 [catalogue, Palaearctic]; Schumann et al. 1999: 144 [checklist, Germany].
Clusiodes nubila. Séguy 1934: 352 [illustration of wing, France]; Ceianu 1989: 29 [expected in Romania].
Clusiodes (Columbiella) albimanus. Steyskal 1965: 807 [catalogue, America north of Mexico].
Clusioides (Clusioides) albimana. Soós 1981: 99 [Hungary, key, illustration of habitus, male terminalia].
Clusioides (Columbiella) nubila. Soós 1981: 102 [possible occurrence in Hungary, key].
Clusiodes (Clusiodes) albimanus. Soós 1984: 12 [catalogue, Palaearctic (Europe, including Ukraine)]; Roháček 1995: 126 [biology and habitat, male behaviour, Czech and Slovak Republics]; Schumann et al. 1999: 144 [checklist, Germany]; Beuk 2002: 271 [checklist, Netherlands]; Greve 2005: 3 [Norway].
Clusiodes (Clusiodes) albima. Misspelling of albimana. Greve & Midtgaard 1986: 89.
Clusiodes albimanus. Krištofík & Zuska 1986: 165 [Slovakia]; Soós 1987: 855 [illustration of head]; Ceianu 1989: 29 [adults in boggy forests, catalogue, Romania]; Woznica 1991: 208 [checklist, Poland]; DeBruyn 1991: 146 [catalogue, Belgium]; Poole 1996: 126 [checklist, Nearctic]; Barrientos et al. 1996: 118 [biology, Spain]; Merz 1998: 276 [checklist, Switzerland]; Roháček & Barták 2001: 338 [checklist, Czech Republic]; Carles-Tolrá 2002: 155 [catalogue, Spain]; Roháček et al. 2005: 279 [catalogue, habitat, Czech Republic]; Barták & Kubík 2005: 279 [Czech Republic]; Lonsdale & Marshall 2007b: 129 (synonymy of nubila Meigen and varieties pallidior and obscurior, maps, key, notes, neotype, phylogeny, illustrations of genitalia, photo, new records (France, Greece, Netherlands, Russia, Switzerland)]; Roháček 2008: 154 [Italy]; 2009: 218 [Slovakia], 2012: 99 [central Slovakia], 2013: 130 [central Slovakia]; Nilsson 2009: 5 [Sweden]; Rotheray & Horsfield 2013: 403 [illustrations of immatures, photo of puparium; rearing data— Alnus, Betula, Fagus, Fraxinus, Populus, Salix; England, Finland, Scotland]; Kahanpää 2014: 288 [checklist, Finland]; Roháček et al. 2016: 158 [Portugal].
Clusiodes (Clusiodes) albimanus. Roháček 1997: 75 [checklist, Czech and Slovak republics]; Schacht 2004: 276 [Germany].
americanus Malloch. NE: Northeast North America (Ontario to Quebec and Illinois to South Carolina).
Clusiodes americana Malloch 1922: 48. USA. Maryland: Prince Georges Co., Beltsville. HT ♂ unknown [female allotype with “ Type ” label in USNM].
Clusiodes (Clusiodes) americana. Melander & Argo 1924: 16 [redescription, new records (New York, Pennsylvania, Virginia)]; Johnson 1925a: 245 [catalogue, New England (Maine, New Hampshire, Vermont)]; Frey 1960: 26 [catalogue, world].
Clusiodes (Columbiella) americanus. Steyskal 1965: 807 [catalogue, America north of Mexico].
Clusiodes americanus. Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 271 [redescription, illustrations of wing and male and female genitalia, photo, new records (northeastern North America)]; Marshall, 2006: 510 [photo]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis].
apicalis (Zetterstedt). PA: Europe, western Russia.
Heteroneura geomyzina var. apicalis Zetterstedt 1848: 2789. Denmark. “ Dania ”. Type information unknown [MZLU].
Heteroneura apicalis. Czerny 1903: 79 [new status, redescription, illustrations of head and wing]; Becker 1905: 39 [catalogue, Palaearctic].
Clusiodes apicalis. Wahlgren 1917: 187 [diagnosis, key]; Malloch 1918: 6 [Nearctic record - likely ater Melander & Argo]; Stackelberg 1933: 381 [key, European USSR], 1988: 505 [key, European USSR]; Tuomikoski 1933: 18 [Finland], 1936: 183 [Finland]; Séguy, 1934: 352 [France]; Greve et al. 1987: 78 [Norway]; Ceianu 1989: 29 [catalogue, Romania]; Poole 1996: 126 [checklist, Nearctic]; Dunk 1996: 422 [catalogue, Bavaria]; Yagi & Sawada 2000: 41 [Mamaev collection holdings]; Roháček 2005: 326 [listed as endangered in Czech Republic]; Lonsdale & Marshall 2007b: 135 [key, map, synonymy, notes, phylogeny, illustrations of genitalia, new records (Finland, Norway, Russia)]; Rotheray & Horsfield 2013: 404 [illustrations of immatures; rearing data— Alnus, Betula, Populus; Finland, Russia, Scotland]; Kahanpää 2014: 288 [checklist, Finland].
Clusiodes (Clusiaria) apicalis. Melander & Argo 1924: 17 [misidentification—likely ater Melander & Argo, redescription, Idaho]; Czerny 1928: 6 [redescription, Europe]; Frey 1928c: 109 [Finland, catalogue], 1960: 26 [catalogue, world]; Soós 1943: 72 [Hungary], 1984: 12 [catalogue, Palaearctic (Denmark, Finland, Britain, northern European USSR)]; Collin 1953b: 139 [England]; Trojan 1962: 24 [key, illustrations of head and wing, Poland]; Hackman 1980: 148 [checklist, Finland]; Stubbs 1982: 92 [Britain]; Greve & Midtgaard 1986: 89 [biology, Betula verrucosa and Salix spp. as likely hosts, Norway]; Roháček 1995: 128 [Czeck Republic]; Schumann et al. 1999: 144 [checklist, Germany]; Greve 2005: 5 [Norway].
Clusiodes (Clusiodes) apicalis. Steyskal 1965: 807 [catalogue, America north of Mexico].
Clusioides (Columbiella) apicalis. Soós 1981: 102 [possible occurrence in Hungary, key, illustration of wing].
Clusiodes (Clusiaria) pictipes. Misidentification. Máca 1982: 71 [misidentification det. Roháček (1995), thorax with Trombidium cf. mediterraneum (Berlese) mites attached].
apiculatus Malloch. NE: Northeastern North America (Illinois to Nova Scotia and Tennessee to South Carolina).
Clusiodes (Columbiella) apiculata Malloch 1922: 49. USA. New Hampshire: Hillsborough Co., Milford. HT ♂ MCZ [Type No. MCZT 27014]. Melander & Argo 1924: 23 [redescription, North Carolina]; Johnson 1925a: 246 [catalogue, Vermont].
Clusiodes (Columbiella) nigripalpis. Misidentification in part. Malloch 1922: 49 [misidentification of paratypes, det. Caloren & Marshall (1998)].
Clusiodes apiculata. Brimley 1938: 378 [North Carolina];
Clusiodes (Clusiodes) apiculata. Frey 1960: 26 [catalogue, world].
Clusiodes (Columbiella) apiculatus. Steyskal 1965: 807 [catalogue, America north of Mexico].
Clusiodes apiculatus. Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 269 [redescription, photo, illustrations of male and female genitalia, new records (northeastern North America)]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis].
ater Melander & Argo. NE: Widespread in North America north of Mexico (outside of the southeastern USA).
Clusiodes (Clusiaria) atra Melander & Argo 1924: 18. USA. Massachusetts: Franklin Co., Greenfield. HT ♂ USNM. Johnson 1925a: 245 [catalogue, New England]; Frey 1960: 26 [catalogue, world].
Clusiodes (Clusiaria) nitida var. scutellata Melander & Argo 1924: 20. USA. Colorado: Boulder Co., Boulder. HT ♀ USNM [Cat. No. 26264]. Caloren & Marshall 1998: 291 [as “misidentification”, synonymy with atra Melander & Argo].
Clusiodes atra. Johannsen 1928: 844 [catalogue, New York].
Clusiodes (Clusiodes) ater. Steyskal 1965: 807 [catalogue, America north of Mexico].
Clusiodes (Clusiodes) nitidus var. scutellatus. Steyskal 1965: 807 [catalogue, America north of Mexico].
Clusiodes scutellatus. Poole 1996: 126 [checklist, Nearctic, as synonym of nitidus Melander & Argo].
Clusiodes ater. Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 291 [redescription, phylogeny, photo, illustrations of wing and male and female genitalia, new records (North America)]; Marshall, 2006: 510 [photo]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis, photo]; Lonsdale et al. 2010: 240 [genes sequenced for phylogenetic analysis].
caestus Caloren & Marshall. NE: Western North America (Alberta, New Mexico, Utah), Nova Scotia and Ontario.
Clusiodes caestus Caloren & Marshall 1998: 289. Canada. Ontario: Fathom Five National Park, Cove Island. HT ♂ DEBU. Caloren & Marshall 1998: 289 [illustrations of wing and male and female genitalia]; Lonsdale & Marshall 2007b: 146 [map, discussion of C. melanostomus complex, phylogenetic analysis].
caledonicus (Collin). PA: Europe, western Russia.
Heteroneura caledonica Collin 1912: 106 [illustration of male external genitalia]. United Kingdom. Scotland: Inverness; England: Golspie. ST 9♂ 22♀ BMNH, CUMZ, OUMNH.
Clusiodes (Clusiodes) caledonica. Melander & Argo 1924: 17 [generic combination, summary of original description]; Czerny 1928: 9 [redescription, reillustration from original description]; Kloet & Hincks 1945: 394 [checklist, Britain]; Frey 1960: 26 [catalogue, world]; Trojan 1962: 26 [key, illustration of male genitalia, Poland]; Hackman 1980: 148 [checklist, Finland]; Stubbs 1982: 92 [Britain]; Soós 1984: 12 [catalogue, Palaearctic (Britain, Germany, Austria)]; Greve & Midtgaard 1986: 90 [Norway]; Greve & Midtgaard 1986: 90 [Norway]; Roháček 1995: 128 [biology, Czech and Slovak Republics]; Schumann et al. 1999: 144 [checklist, Germany]; Beuk 2002: 271 [checklist, Netherlands]; Greve 2005: 4 [Norway].
Clusiodes caledonica. Tuomikoski 1936: 184 [reared from rotten aspen, adults on birch logs, illustrations of male genitalia, Finland]; Speight & Cogan 1979: 402 [Ireland]; Stackelberg 1988: 505 [key, European USSR].
Clusioides (Clusioides) caledonica. Soós 1981: 100 [possible occurrence in Hungary, key, illustrations of wing, male terminalia].
Clusiodes caledonicus. Ceianu 1989: 29 [expected in Romania]; Merz 1998: 276 [checklist, Switzerland]; Roháček & Barták 2001: 338 [checklist, Czech Republic]; Roháček et al. 2005: 280 [catalogue (considered endangered in Czech Republic), habitat, Czech Republic]; Roháček 2005: 326 [listed as endangered in Czech Republic], 2009: 218 [Slovakia], 2012: 98 [central Slovakia], 2013: 128 [central Slovakia]; Barták & Kubík 2005: 280 [Czech Republic]; Lonsdale & Marshall 2007b: 131 [synonymy, map, key, phylogeny, notes, illustrations of genitalia, new records (Finland, Greece, Italy, Switzerland, Sweden, Spain)]; Nilsson 2009: 5 [Sweden]; Rotheray & Horsfield 2013: 404 [illustrations of immatures, photo of head skeleton in puparium; rearing data (Pinus), Scotland]; Kahanpää 2014: 288 [checklist, Finland]; Chandler 2015: 212 [England]; Roháček et al. 2016: 158 [Portugal].
Clusiodes calcedonicus. Misspelling of caledonicus. DeBruyn 1991: 146 [Belgium].
Clusiodes (Clusiodes) caledonicus. Roháček 1997: 75 [checklist, Czech and Slovak republics].
clandestinus Caloren & Marshall. NE: Northeast North America (Ontario to Nova Scotia and Illinois to North Carolina).
Clusiodes clandestinus Caloren & Marshall 1998: 277. Canada. Ontario: Nipissing District, Algonquin Provincial Park, Swan Lake Research Station. HT ♂ DEBU. Caloren & Marshall 1998: 277 [illustrations of wing and male and female genitalia, on fresh maple log, dogwood leaves and hemlock log]; Marshall, 2006: 510 [photo, female ovipositing in crack in dead wood]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis]; Lonsdale et al. 2010: 240 [genes sequenced for phylogenetic analysis].
coconino Caloren & Marshall. NE: USA (Arizona).
Clusiodes coconino Caloren & Marshall 1998: 280. USA. Arizona: Coconino Co., 10 mi NW Flagstaff, San Francisco Mountains. HT ♂ DEBU. Caloren & Marshall 1998: 280 [illustrations of male genitalia]; Lonsdale & Marshall 2007b: 146 [map, discussion of C. melanostomus complex, phylogenetic analysis].
discostylus Sueyoshi. PA: Japan.
Hendelia beckeri. Misidentification in part. Sueyoshi et al. 2003: 182 [Japan].
Clusiodes discostylus Sueyoshi 2006: 7. Japan. Hokkaido: Shikaribetsu, Shikaoi Town. HT ♂ BLKU. Sueyoshi 2006: 7 [illustrations of head and male and female genitalia, photo of wing], 2011: 8 [Japan, pictoral key], 2014: 618 [catalogue, Japan]; Lonsdale & Marshall 2007b: 132 [key, map, synonymy, phylogeny, illustrations of genitalia].
eremnos Caloren & Marshall. NE: Western North America (Alberta, British Columbia, Oregon, Colorado, Idaho, Montana, Washington).
Clusiodes eremnos Caloren & Marshall 1998: 282. USA. Oregon: Baker Co., 9mi W of Unity, 4500′. HT ♂ CNC. Caloren & Marshall 1998: 282 [illustrations of male genitalia]; Lonsdale & Marshall 2007b: 146 [map, discussion of C. melanostomus complex, phylogenetic analysis].
flaveolus Mamaev. PA: Russia.
Clusiodes flaveola Mamaev 1974: 115. Russia. Far East: Primorskiy Krai. Ussury Res. HT ♂ ZMM [Registration No. Di0160]. Mamaev 1974: 115 [illustration of surstylus]; Yagi & Sawada 2000: 41 [Mamaev collection holdings]. Lonsdale & Marshall 2007b: 129 [lapsus]; Ozerov 2010: 145 [types in ZMM].
Clusiodes flaveolus. Lonsdale & Marshall 2007b: 137 [key, map, synonymy, notes, phylogeny, Russia].
freyi Tuomikoski. PA: Finland, Russia.
Clusiodes (Clusiaria) Freyi Tuomikoski 1933: 16. Finland. Padasjoki, Lohja. HT ♂ UZMH [Type No. 4846].
Clusiodes freyi. Tuomikoski 1936: 183 [Finland]; Stackelberg 1988: 505 [key, European USSR]; Yagi & Sawada 2000: 41 [Mamaev collection holdings, questionable identification]; Lonsdale & Marshall 2007b: 139 [key, map, synonymy, notes, phylogeny, illustrations of genitalia, Finland, Russia]; Rotheray & Horsfield 2013: 404 [illustrations of immatures; rearing data— Betula, Picea; Finland, Russia]; Kahanpää 2014: 288 [checklist, Finland].
Clusiodes (Clusiaria) freyi. Frey 1960: 26 [catalogue, world]; Hackman 1980: 148 [checklist, Finland]; Soós 1984: 12 [catalogue, Palaearctic (Finland, European Russia)].
gentilis (Collin). PA: Europe (not Finland), western and central Russia.
Heteroneura gentilis Collin. 1912: 107. United Kingdom. England: Hampshire, New Forest, Lyndhurst Road. LT ♂ BMNH (designated by Lonsdale & Marshall 2007b: 132)]. Collin 1912: 107 [illustration of male external genitalia].
Clusiodes (Clusiodes) gentilis. Czerny 1928: 9 [redescription, reillustration from original description]; Melander & Argo 1924: 17 [summary of original description]; Frey 1928c: 109 [catalogue, Finland], 1960: 26 [catalogue, world]; Kloet & Hincks 1945: 394 [checklist, Britain]; Trojan 1962: 26 [key, illustrations of male genitalia and larval cephalopharyngeal skeleton, Poland]; Chandler 1978: 264 [Ireland]; Stubbs 1982: 92 [Britain]; Soós 1984: 12 [catalogue, Palaearctic (Britain, Poland, Finland, northern European USSR)]; Beuk 2002: 271 [checklist, Netherlands].
Clusiodes gentilis. Stackelberg 1933: 381 [key, European USSR], 1988: 505 [key, European USSR]; Richards, 1934: 2 [puparia found in rotten birch]; Séguy 1934: 352 [illustrations of puparium and larval cephalopharyngeal skeleton, France]; Hennig 1952: 227 [immature stages]; Withers 1985: 63 [England, reared from Salix sp. in bog and rotting Betula sp., associated with Fagus sylvaticus]; Ferrar 1987: 122 [immature stages, illustration of larva]; Smith 1989: 103 [bred from log, illustration of larval mouthparts, Britain]; Ceianu 1989: 29 [expected in Romania]; Woznica 1991: 208 [checklist, Poland]; DeBruyn 1991: 146 [catalogue, Belgium]; Yagi & Sawada 2000: 41 [Mamaev collection holdings]; Carles-Tolrá 2002: 155 [catalogue, Spain]; Roháček 2004: 250 [Cyprus], 2013: 130 [central Slovakia]; Lonsdale & Marshall 2007b: 132 [synonymy, key, map, phylogeny, lectotype designation, illustrations of genitalia, new records (England, Netherlands, Russia, Switzerland)]; Rotheray & Horsfield 2013: 404 [illustrations of immatures; rearing data— Betula, Fagus, Pinus, Salix, Ulmus; England, Scotland]; Kahanpää 2014: 288 [reiterates Tuomikoski (1936) that previous Finnish records of gentilis were misidentifications of caledonicus]; Roháček et al. 2016: 160 [colour variation, Portugal].
Clusioides (Clusioides) gentilis. Soós 1981: 100 [possible occurrence in Hungary, key, illustrations of wing, male terminalia].
geomyzinus (Fallén). PA: Europe, Russia.
Heteroneura geomyzina Fallén 1823: 2. Sweden. “Smolandia”. ST ♂ ♀ UZMC. Meigen 1830: 129 [Europe]; Zetterstedt 1848: 2787 [larvae and puparia subcortical in conifers, Norway, Sweden, Finland], 1852: 4816 [Sweden]; Loew 1857: 54 [Germany], 1864b: 345 [redescription, Germany]; Schiner 1862: 38 [redescription], 1864: 72 [catalogue, Europe]; Siebke 1877: 181 [catalogue, Norway]; Brauer 1883: 92 [records of the Vienna Museum]; Becker 1902: 313 [discussion of type], 1905: 39 [catalogue, Palaearctic]; Czerny 1903: 77 [redescription, synonymy of alpina Loew, Sweden, Norway, Denmark, Germany, Austria, Switzerland, Italy, illustrations of head and wing], 1928: 7 [redescription]; Czizek 1910: 112 [catalogue, Germany];
Agromyza geomyzina. Zetterstedt 1838: 790 [generic combination].
Heteroneura alpina Loew 1857: 53. Germany [not given]. HT ♀, depository unknown. Loew, 1864b: 344 [discussion, Germany]; Girschner 1888: 97 [comparison to albimana Meigen]; Czerny 1903: 77 [as synonym of geomyzina Fallén].
Clusiodes geomyzina. Johnson 1913b: 97 [generic combination, Nearctic record likely orbitalis Malloch]; Wahlgren 1917: 186 [diagnosis, key]; Malloch 1918: 6 [synonymy, key]; Stackelberg 1933: 381 [key, European USSR], 1988: 505 [key, European USSR]; Séguy 1934: 352 [France]; Tuomikoski 1936: 183 [adults on decayed deciduous and coniferous logs (particularly birch and aspen), Finland]; Hennig 1952: 227 [immature stages, illustration of puparium]; Máca 1985: 397 [Czech Republic]; Ferrar 1987: 122 [illustration of puparium]; Zuska 1987: 235 [possibly present in Czech Republic]; Dunk 1996: 422 [catalogue, Bavaria]; Yagi & Sawada 2000: 41 [Mamaev collection holdings].
Clusiodes (Clusiaria) geomyzinus. Melander & Argo 1924: 18 (in part) [misidentification—possibly orbitalis Malloch or nitidus Melander & Argo, redescription]; Frey 1928c: 108 [catalogue, Finland]; Tuomikoski 1933: 15 [Finland]; Soós 1984: 12 [catalogue, Palaearctic (Europe, Ukraine, northern European Russia)]; Roháček 1995: 128 [biology, habitat, male agonistic behaviour, Czech and Slovak Republics], 1997: 75 [checklist, Czech and Slovak republics], 1999: 291 [Czech Republic]; Poole 1996: 126 [checklist, Nearctic, [misidentification—possibly orbitalis Malloch or nitidus Melander & Argo]; Schumann et al. 1999: 144 [checklist, Germany]; Beuk 2002: 271 [checklist, Netherlands]; Greve 2005: 6 [Norway].
Clusiodes (Clusiaria) geomyzina. Johnson 1925a: 245 [Nearctic records likely misidentifications of orbitalis Malloch]; Czerny, 1928: 7 [redescription, Europe]; Soós 1943: 72 [Hungary]; Kloet & Hincks 1945: 394 [checklist, Britain]; Frey 1960: 26 [catalogue, world]; Trojan 1962: 22 [key, illustration of wing and puparium, Poland]; Hackman 1980: 148 [checklist, Finland]; Stubbs 1982: 92 [Britain]; Greve & Midtgaard 1986: 90 [habitat, Norway].
Clusiodes (Clusiodes) geomyzinus. Steyskal 1965: 807 [catalogue, America north of Mexico - likely Clusiodes orbitalis].
Clusioides (Columbiella) geomyzina. Soós 1981: 102 [possible occurrence in Hungary, key, illustration of wing].
Clusiodes geomyzinus. Ceianu 1989: 29 [catalogue, Romania]; Woznica 1991: 208 [checklist, Poland]; Merz 1998: 276 [checklist, Switzerland]; Roháček & Barták 2001: 338 [checklist, Czech Republic]; Carles-Tolrá & Ventura 2001: 12 [Iberia]; Carles-Tolrá 2002: 155 [catalogue, Spain]; Roháček et al. 2005: 280 [catalogue, habitat, Czech Republic]; Barták & Kubík 2005: 280 [Czech Republic]; Lonsdale & Marshall 2007b: 140 [key, biology, map, synonymy, notes, phylogeny, illustrations of genitalia, new records (Finland, Russia, Spain, Sweden, Switzerland)]; Roháček 2008: 154 [Italy], 2009: 218 [Slovakia], 2012: 99 [central Slovakia]; Nilsson 2009: 5 [Sweden]; Rotheray & Horsfield 2013: 404 [illustration of immature; photo of puparium; rearing data— Betula (?), Picea, Pinus; Finland, Russia]; Rotheray & Wilkinson 2013: 1356 [Scotland]; Kahanpää 2014: 288 [checklist, Finland].
gracilolobus Caloren & Marshall. NE: Alabama, Florida, Georgia, Kentucky, Maryland, Virginia.
Clusiodes gracilolobus Caloren & Marshall 1998: 282. USA. Maryland: Montgomery Co., Rockville. HT ♂ USNM. Caloren & Marshall 1998: 282 [photo and illustrations of male genitalia]; Lonsdale & Marshall 2007b: 146 [notes, map, discussion of C. melanostomus complex, phylogenetic analysis, photo, Alabama, Kentucky].
johnsoni Malloch. NE: Widespread in North America south of Canadian Territories.
Heteroneura albimana. Misidentification. Osten Sacken 1878: 197 [catalogue, North America].
Clusiodes (Clusiodes) albimana. Misidentification. Johnson 1913b: 98 [eastern United States], 1925a: 245 [catalogue, Vermont]; Frey 1960: 26 (in part) [catalogue, world].
Clusiodes albimana. Misidentification. Malloch 1918: 6 [Ontario]; Johannsen 1928: 844 [catalogue, New York].
Clusiodes (Columbiella) johnsoni Malloch 1922: 49. USA. Maine: Kennebec Co., Mt. Desert, Echo Lake. HT ♂ MCZ [Type No. MCZT 27015]. Melander & Argo 1924: 23 [redescription, new records (Ontario, Maryland, New York, Virginia)]; Johnson 1925a: 246 [catalogue, New England (Maine, Massachutsetts, Vermont)]; Steyskal 1965: 807 [catalogue, America north of Mexico].
Clusiodes (Columbiella) nigripalpis Malloch 1922: 49. USA. Maryland: Glen Echo. HT ♂ USNM. Johnson 1925a: 246 [catalogue, New England (Maine, Massachusetts, New Hampshire, Vermont), misidentification in part of apiculatus Malloch].
Clusiodes albimanus var. pallidior. Misidentification in part. Melander & Argo 1924: 16 [photo, new records (Idaho, Washington)].
Clusiodes (Columbiella) johnsoni form nigripalpis. Melander & Argo 1924: 23 [new status]. Caloren & Marshall 1998: 274 [as synonym of johnsoni Malloch].
Clusiodes johnsoni. Johannsen 1928: 844 [catalogue, New York]; Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 274 [redescription, illustrations of wing and male and female genitalia, new records (North America)]; Lonsdale & Marshall 2007b: 146 [phylogenetic analysis, illustrations of puparium and head, photo], 2010: 1045 [illustration of male genitalia]; Lonsdale et al. 2010: 240 [genes sequenced for phylogenetic analysis].
Clusiodes (Clusiodes) johnsoni. Frey 1960: 26 [catalogue, world].
Clusiodes nigripalpis. Poole 1996: 126 [as synonym of johnsoni, checklist, Nearctic].
latus Sasakawa. OR: Laos.
Hendelia latus Sasakawa 2011b: 21. Laos. Vientian: Ban Van Eue, 750m. HT ♂, BPBM [Type No. BPBM 17354]. Sasakawa 2011b: 21 [illustrations of male genitalia].
leptapodemus Caloren & Marshall. NE: New Mexico, Arizona, Nevada.
Clusiodes leptapodemus Caloren & Marshall 1998: 280. USA. New Mexico: Socorro Co., South Baldy Peak, 20mi W of Socorro, alpine meadow, 10,400′. HT ♂ DEBU. Caloren & Marshall 1998: 280 [illustrations of male and female genitalia]; Lonsdale & Marshall 2007b: 146 [map, discussion of C. melanostomus complex, phylogenetic analysis].
leucopeza Frey. OR: Burma.
Clusiodes (Clusiodes) leucopeza Frey 1960: 18. Burma. Kambaiti, 2000 m. ST 1♂ 2♀ UZMH. Frey 1960: 18 [catalogue, world]; Sasakawa 1977: 236 [catalogue, Orient], 1987: 101 [key].
Clusiodes leucopeza. Lonsdale & Marshall 2007b: 132 [key, map, synonymy, phylogeny, notes].
melanostomus (Loew). NE: Northern North America (uncommon in southern United States).
Heteroneura melanostoma Loew 1864a: 260. USA. New Hampshire. LT 1♂ MCZ [accidental designation of lectotype by assumption of holotype (IZCN (1999) Art. 74.6) by Caloren & Marshall (1998); 1 ST originally described]. Osten Sacken 1878: 198 [catalogue, North America]; Czerny 1903: 77 [quotes description]; Aldrich 1905: 570 [catalogue, North America].
Heteroneura flavifascies Slosson 1897: 239. USA. New Hampshire: Coos Co., Mt. Washington. Slosson 1897: 239 [nomen nudum, attributed to Coquillett]; Poole 1996: 126 [checklist, Nearctic, as syn. of melanostoma Loew attributed to Coquillett].
Clusiodes melanostoma. Johnson 1913b: 98 [generic combination, new records (Quebec, Maine, Massachusetts, New Hampshire, Vermont)], 1925b: 95 [Harris collection]; Malloch 1918: 5 [Illinois]; Griffiths 1972: 214 [male dissected for discussion of phylogenetic relationships]; Poole 1996: 126 [checklist, Nearctic].
Clusiodes (Clusiaria) duplicata Malloch 1922: 49. USA. Maryland: Glen Echo. HT ♂ USNM. Melander & Argo 1924: 19 [as synonym of melanostoma Loew].
Clusiodes (Clusiaria) melanostoma. Melander & Argo 1924: 19 [redescription, synonymy of duplicata Malloch, photo, new records (Illinois, Maryland, New Hampshire, New York, Virginia, Washington)]; Johnson 1925a: 245 [catalogue, New England (Maine, Massachusetts, New Hampshire, Vermont)]; Frey 1960: 26 [catalogue, world].
Clusiodes (Clusiaria) ruficollis. Misidentification. Melander & Argo 1924: 22 [redescription, possibly melanostomus Loew].
Clusiodes melanostomus. Johannsen 1928: 844 [catalogue, New York]; Caloren & Marshall 1998: 279 [redescription, illustrations of male and female genitalia, new records (northern North America), designation of LT]; Lonsdale & Marshall 2007b: 146 [notes, map, illustration of puparium, discussion of C. melanostomus complex, phylogenetic analysis]; Lonsdale et al. 2010: 240 [member of “ Clusiodes melanostomus complex” genetically sequenced for phylogenetic analysis].
Clusiodes (Clusiodes) melanostoma. Steyskal 1965: 807 [catalogue, America north of Mexico].
Clusiodes duplicata. Poole 1996: 126 [as synonym of melanostomus, checklist, Nearctic].
microcerus Stackelberg. PA: Finland, Russia.
Clusiodes (Clusiodes) microcera Stackelberg, 1955: 329. Russia. Leningrad Region: Luzhnitsky Rayon, Tolmachevo. HT ♂ ZIN. Frey 1960: 26 [catalogue, world]; Stackelberg 1988: 505 [key, European USSR].
Clusiodes microcera. Mamaev 1987: 114 [Russian Far East].
Clusiodes microcerca. Misspelling. Yagi & Sawada 2000: 40 [Mamaev collection holdings].
Clusiodes microcerus. Lonsdale & Marshall 2007b: 134 [key, map, synonymy, notes, phylogeny, illustrations of genitalia, Russia]; Rotheray & Horsfield 2013: 405 [illustrations of immatures; rearing data— Betula; Finland]; Kahanpää 2014: 288 [checklist, Finland].
niger Melander & Argo. NE: Georgia, New Mexico.
Clusiodes (Clusiodes) nigra. Melander & Argo 1924: 17. USA. New Mexico: Lincoln Co., Rio Ruidoso, White Mts., ~6500′. HT ♂ USNM [Cat. No. 26262]. Frey, 1960: 26 [catalogue, world].
Clusiodes (Columbiella) niger. Steyskal 1965: 807 [catalogue, America north of Mexico].
Clusiodes niger. Poole 1996: 126 [checklist, Nearctic]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis].
Clusiodes nigrus. Caloren & Marshall 1998: 273 [redescription, illustrations of wing and male genitalia].
nitidus Melander & Argo. NE: New Mexico, Arizona, Mexico.
Clusiodes (Clusiaria) nitida Melander & Argo 1924: 20 [with photo]. USA. New Mexico: Rio Arriba Co., Hill above Beulah. HT ♀ USNM [Cat. No. 26263]. Frey 1960: 26 [catalogue, world].
Clusiodes (Clusiodes) nitidus. Steyskal 1965: 807 [catalogue, America north of Mexico].
Clusiodes nitidus. Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 284 [redescription, illustrations of wing and male and female genitalia, new records (Arizona, Mexico)]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis].
orbitalis Malloch. NE: North America (uncommon in southern USA).
Clusiodes (Columbiella) orbitalis Malloch 1922: 50. USA. Maine: Hancock Co., Bar Harbor. HT ♀ MCZ [Type No. MCZT 27016].
Clusiodes (Clusiaria) orbitalis. Melander & Argo 1924: 21 [summary of original description]; Johnson 1925a: 245 [catalogue, Maine]; Frey 1960: 26 [catalogue, world].
Clusiodes (Clusiodes) orbitalis. Steyskal 1965: 807 [catalogue, America north of Mexico].
Clusiodes orbitalis. Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 287 [redescription, illustrations of wing and male and female genitalia, new records (North America)]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis]; Lonsdale et al. 2010: 240 [genes sequenced for phylogenetic analysis].
pictipes (Zetterstedt). PA: Europe, Russia.
Heteroneura pictipes Zetterstedt 1852: 4816. Sweden. “Scania”. ST ♂♀ MZLU. Schiner 1864: 72 [catalogue, Europe]; Slosson 1897: 239 [checklist, New Hampshire - likely orbitalis Malloch]; Czerny 1903: 80 [redescription]; Aldrich 1905: 571 [catalogue, North America - likely orbitalis Malloch]; Becker 1905: 39 [catalogue, Palaearctic]; Czizek 1910: 112 [catalogue, Germany].
Clusiodes pictipes. Johnson 1913b: 98 [generic combination, Nearctic record - likely orbitalis Malloch]; Wahlgren 1917: 187 [diagnosis, key]; Malloch 1918: 7 [doubtful identification—likely orbitalis Malloch]; Tuomikoski 1933: 18 [Finland], 1936: 183 [Finland, adults on rotting birch and aspen trunks]; Stackelberg 1988: 505 [key, European USSR]; DeBruyn 1991: 146 [catalogue, Belgium]; Poole 1996: 126 [checklist, Nearctic, likely misidentification from Malloch (1918)]; Roháček & Barták 2001: 338 [checklist, Czech Republic]; Roháček et al. 2005: 280 [catalogue, habitat, Czech Republic]; Barták & Kubík 2005: 280 [Czech Republic]; Lonsdale & Marshall 2007b: 140 [key, map, synonymy, phylogeny, illustrations of genitalia, illustration of egg, new records (Hungary, Slovakia, Russia, Ukraine)]; Roháček 2008: 154 [Italy—acceptance of record in Palm (1869)], 2012: 99 [central Slovakia], 2013: 128 [central Slovakia]; Kahanpää 2014: 288 [checklist, Finland].
Clusiodes (Clusiaria) pictipes. Czerny 1928: 7 [redescription]; Melander & Argo, 1924: 21 [redescription, USA specimens of uncertain identity]; Johnson 1925a: 245 [Nearctic record—likely misidentification of orbitalis Malloch]; Czerny 1928: 7 [redescription]; Frey 1928c: 109 [catalogue, Finland], 1960: 26 [catalogue, world]; Hackman 1980: 148 [checklist, Finland]; Soós 1984: 12 [catalogue, Palaearctic (northern Europe and northern European Russia)]; Roháček 1995: 130 [biology, Czech and Slovak Republics], 1997: 75 [checklist, Czech and Slovak republics], 1999: 291 [Czech Republic], 2009: 218 [Slovakia]; Schumann et al. 1999: 144 [checklist, Germany]; Greve 2005: 7 [considered rare, Norway].
Clusiodes (Clusiodes) pictipes. Steyskal 1965: 807 [catalogue, America north of Mexico].
ruficollis (Meigen). PA: Europe, Russia.
Heteroneura ruficollis Meigen 1830: 128. Europe. ST? NMW [Winthem collection]. Zetterstedt 1848: 2789 [redescription, Sweden]; Loew 1857: 53 [Germany], 1864b: 343 [synonymy of nubila Meigen and laterella Zetterstedt, Germany]; Schiner 1862: 38 [redescription, key, Austria], 1864: 72 [catalogue, Europe]; Becker 1902: 313 [examination of Meigen types], 1905: 39 [catalogue, Palaearctic]; Czerny, 1903: 75 [redescription, Germany, Sweden].
Heteroneura laterella Zetterstedt 1848: 2791. Sweden. “Smolandia”. ST ♂, depository unknown. Loew 1857: 52 [Germany]; Loew 1864b: 343 [as synonym of ruficollis Meigen].
Clusiodes ruficollis. Wahlgren 1917: 186 [diagnosis, key]; Stackelberg 1933: 381 [key, European USSR], 1988: 505 [key, European USSR]; Séguy 1934: 353 [France, illustrations of puparium and male genitalia (genitalia in illustration those of Clusia flava)]; Tuomikoski 1936: 183 [puparia in rotting alder and birch, Finland]; Hennig 1938: 125 [illustration of male genitalia], 1952: 227 [immature stages]; Vaňhara 1981: 30 [Czech Republic]; Krištofík & Zuska 1986: 165 [Slovakia]; Zuska 1987: 235 [Czech Republic]; Ceianu 1989: 29 [expected in Romania]; Woznica 1991: 208 [checklist, Poland]; DeBruyn 1991: 146 [catalogue, Belgium]; Poole 1996: 126 [checklist, Nearctic, likely representing misidentified melanostoma record in Melander & Argo (1924)]; Merz 1998: 276 [checklist, Switzerland]; Yagi & Sawada 2000: 41 [Mamaev collection holdings]; Roháček & Barták 2001: 338 [checklist, Czech Republic]; Roháček et al. 2005: 280 [catalogue, habitat, Czech Republic]; Barták & Kubík 2005: 280 [Czech Republic]; Lonsdale & Marshall 2007b: 141 [key, map, synonymy, notes, phylogeny, illustrations of genitalia, new records (Estonia, Finland, Netherlands, Russia, Switzerland, Ukraine)]; Nilsson 2009: 5 [Sweden]; Roháček 2009: 218 [Slovakia], 2012: 99 [central Slovakia], 2013: 128 [central Slovakia]; Rotheray & Horsfield 2013: 405 [illustrations of immatures; rearing data— Betula, Fagus, Fraxinus; England, Scotland]; Kahanpää 2014: 288 [checklist, Finland].
Clusiodes (Clusiaria) ruficollis. Czerny 1928: 7 [redescription]; Malloch 1922: 47 [key to subgenera and species, eastern USA—likely melanostomus Loew]; Melander & Argo 1924: 22 [redescription - likey melanostomus Loew)]; Frey 1928c: 108 [catalogue, Finland], 1960: 26 [catalogue, world]; Tuomikoski 1933: 15 [key]; Soós 1943: 72 [Hungary], 1984: 12 [catalogue, Palaearctic (central and northern Europe, northern European Russia)]; Trojan 1962: 23 [key, illustrations of head and leg, Poland]; Hackman 1980: 148 [checklist, Finland]; Withers 1985: 63 [reared from larvae and puparia found beneath bark of felled hornbeam (Carpinus betulus), synonymy of facialis Collin, illustrations of face, England]; Vaňhara 1986: 123 [Czech Republic]; Greve & Midtgaard 1986: 90 [bred from Quercus sp. and Populus tremula, Norway]; Roháček 1995: 131 [biology, habitat, adults on stumps and fallen trunks of various deciduous trees, reared from puparia in decayed birch logs, infestation with mites, mating behaviour, Czech and Slovak Republics], 1997: 75 [checklist, Czech and Slovak republics], 1999: 291 [Czech Republic]; Schumann et al. 1999: 144 [checklist, Germany]; Beuk 2002: 271 [checklist, Netherlands]; Schacht 2004: 276 [Germany]; Greve 2005: 8 [Norway].
Clusiodes (Clusiaria) facialis. Collin 1953a: 139. United Kingdom. England: Cambridgeshire, Chippenham Fen; Sutherland: Golspie; Cheshire: Wilmslow, Rostherne, Cotterhill Clough. Various locations. ST ♂♀, depositories unknown. Collin 1953a: 139 [on decayed fallen trunk]. Frey 1960: 26 [catalogue, world]; Stubbs 1982: 92 [Britain]; Soós 1984: 12 [catalogue, Palaearctic]. Withers 1985: 63 [as synonym of ruficollis Meigen].
Clusiodes (Clusiodes) ruficollis. Steyskal 1965: 807 [catalogue, America north of Mexico].
Clusioides (Columbiella) ruficollis. Soós 1981: 101 [Hungary, key].
spinos Caloren & Marshall. NE: Tennessee, New Hampshire, Vermont.
Clusiodes spinos Caloren & Marshall, 1998: 284. USA. Tennessee: Sevier Co., Great Smoky Mountains National Park, Arch Rock. HT ♂ USNM. Caloren & Marshall 1998: 284 [illustrations of male genitalia]; Lonsdale & Marshall 2007b: 146 [map, discussion of C. melanostomus complex, phylogenetic analysis].
terminalis Melander & Argo. NE: Ontario, New Hampshire.
Clusiodes (Clusiaria) terminalis Melander & Argo 1924: 22. USA. New Hampshire: White Mountains, Morrison. HT ♂ USNM [Cat. No. 26265]. Melander & Argo 1924: 22 [with photo]; Johnson 1925a: 246 [catalogue, New England]; Frey 1960: 26 [catalogue, world].
Clusiodes (Clusiodes) terminalis. Steyskal 1965: 807 [catalogue, America north of Mexico].
Clusiodes terminalis. Poole 1996: 126 [checklist, Nearctic]; Caloren & Marshall 1998: 293 [redescription, illustrations of wing and male genitalia, new record (Ontario)]; Lonsdale & Marshall 2007b: 146 [map, phylogenetic analysis].
tobi Sueyoshi. PA: Japan.
Clusiodes tobi Sueyoshi 2006: 10. Japan. Hokkaido: Shikaribetsu, Shikaoi Town. HT ♂ BLKU. Sueyoshi 2006: 10 [illustrations of head and male and female genitalia, photo of wing], 2011: 8 [Japan, pictoral key], 2012: 26 [Japan], 2014: 618 [catalogue, Japan]; Lonsdale & Marshall 2007b: 142 [key, map, synonymy, notes, phylogeny, illustrations of genitalia].
tuomikoskii Mamaev. PA: Russia (Far East).
Clusiodes tuomikoskii Mamaev 1974: 114. Russia. Far East: Primorsky Krai, Khasansky District, Kedrovaja Pad Nature Reserve. HT ♂ [with puparium] MNHA [not in ZIN, as stated in original publication, examined by author]. Mamaev 1974: 114 [illustration of surstylus]; Yagi & Sawada 2000: 41 [Mamaev collection holdings]. Lonsdale & Marshall 2007b: 142 [key, map, synonymy, phylogeny, illustrations of genitalia].
verticalis (Collin). PA: Europe, Israel, western Russia.
Heteroneura verticalis Collin 1912: 107. United Kingdom. England: Various localities. ST 3♂ [♀?] BMNH [additional depository(ies) unknown]. Collin 1912: 107 [illustration of male external genitalia].
Clusiodes (Columbiella) verticalis. Melander & Argo 1924: 24 [synonymy, generic combination, comparison to albimanus Meigen]; Czerny 1928: 9 [redescription, reillustration]; Frey 1928c: 109 [Finland, catalogue], 1960: 26 [catalogue, world]; Kloet & Hincks 1945: 394 [checklist, Britain]; Trojan 1962: 26 [key, illustration of male genitalia, Poland]; Chandler 1978: 264 [Ireland, reared from rotten wood]; Stubbs 1982: 92 [Britain]; Greve 1983: 55 [habitat, Norway], 2005: 8 [Norway]; Soós 1984: 12 [catalogue, Palaearctic]; Withers 1985: 63 [reared from rotten birch, illustration of posterior end of puparium, Britain]; Greve & Midtgaard 1986: 91 [Norway]; Stackelberg 1988: 505 [key, European USSR]; Smith 1989: 103 [reared from old grass heap, England]; Roháček 1995: 133 [biology, Czech and Slovak Republics], 1997: 75 [checklist, Czech and Slovak republics]; Beuk 2002: 271 [checklist, Netherlands]; Schacht 2004: 276 [Germany].
Clusiodes verticalis. Andersson 1971: 234 [Sweden]; Máca 1985: 398 [adult at fermenting beer in sawdust dump, Czech Republic]; Ceianu 1989: 29 [expected in Romania]; DeBruyn 1991: 146 [catalogue, Belgium]; Barrientos et al. 1996: 120 [biology, Spain]; Merz 1998: 276 [checklist, Switzerland]; Carles-Tolrá 2002: 155 [catalogue, Spain]; Lonsdale & Marshall 2007b: 134 [key, map, synonymy, notes, phylogeny, illustrations of genitalia, new records (Finland, France, Israel, Italy, Switzerland)]; Roháček 2008: 154 [Italy]; 2009: 218 [Slovakia], 2013: 128 [central Slovakia]; Lonsdale et al. 2010: 240 [genes sequenced for phylogenetic analysis]; Rotheray & Horsfield 2013: 405 [illustrations of immatures; rearing data— Alnus, Betula, Fagus, Fraxinus, Pinus, Salix; England, Scotland]; Kahanpää 2014: 288 [checklist, Finland]; Carles-Tolrá & Andrade 2011: 489 [Portugal]; Roháček et al. 2016: 160 [Portugal].
Clusioides (Clusioides) verticalis. Soós 1981: 98 [Hungary, key, associated with fungus Lactarius piperatus, illustrations of male and female terminalia, wing].
description: Published as part of Owen Lonsdale, 2017, World Catalogue of the Druid Flies (Diptera: Schizophora: Clusiidae), pp. 1-85 in Zootaxa 4333 (1) on pages 21-29, DOI: 10.11646/zootaxa.4333.1.1, http://zenodo.org/record/1010066