Manggabora wapitja Dean & Suter, 2004, sp. nov.
Creators: name: Dean, J. C.
name: Suter, P. J.
Publisher: name: Zenodo
Year: 2004
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Ephemeroptera
Leptophlebiidae
Manggabora
Manggabora wapitja

description: Manggabora wapitja sp. nov.   
Figures 14–34  
 Type material. Holotype: male imago (reared from nymph), Coobanrbora Spring, Kakadu National Park, Northern Territory, 12˚24'S 132˚40'E, 21 Aug 1999, J. Dean (NMV T-18491).   
Paratype: male imago (reared from nymph; wings, legs and nymphal exuvia mounted on slides), collected with holotype (NMV T-18492).  
 Other material examined. Northern Territory. 10N, Coobanrbora Spring, Kakadu National Park, Northern Territory, 12˚24'S 132˚40'E, 21 Aug 1999, J. Dean; 1MI, 17N, Koolpin Gorge, Kakadu National Park, 13˚30'S 132˚35'E, 18 Aug 1999, J. Dean; 1MI, 1FSI, 11N, Walker Creek, Litchfield National Park, 13˚05'S 130˚42'E, 31 Aug 1999, J. Dean; 14N, Florence Falls, Litchfield National Park, 13˚06'S 130˚47'E, 31 Aug 1999, J. Dean; 1FSI, 21N, Tolmer Falls. Litchfield National Park, 13˚12 ’ S 130˚43 ’ E, 31 Aug 1999, J. Dean; 2MI, 20N, Mann River, Arnhem Land, 12˚22'S 134˚08'E, 26 Aug 1999, J. Dean; 6N, Liverpool River, Arnhem Land, 12˚21' ’ S 134˚07'E, 26 Aug 1999, J. Dean; 1FI, 22N, Manggabor Creek, Arnhem Land, 12˚17'S 134˚05'E, 26 Aug 1999, J. Dean; 1FI, 56N, Kambolgie Creek, Kakadu National Park, 13˚31'S 132˚23'E, 16–19 Aug 1999, J. Dean; 3N, Kambolgie Creek, Kakadu National Park, 13˚31'S 132˚23'E, 1 May 1990, D. Cartwright; 13N, Barramundie Gorge, Kakadu National Park, 13˚19'S 132˚26'E, 22 Aug 1999, J. Dean; 1FI, 3N, Gerowie Creek, 7 km N of Bukbukluk Lookout, Kakadu Highway, 13˚26'S 132˚16'E, 19 Aug 1999, J. Dean; 32N, creek 2 km N of Bukbukluk Lookout, Kakadu Highway, 13˚29'S 132˚15'E, 19 Aug 1999, J. Dean; 1FI, 1N, South Alligator River, Koolpin Crossing, 13˚32'S 132˚33'E, 18 Aug 1999, J. Dean; 13N, South Alligator River, Gunlom Road Crossing, 13˚30'S 132˚29'E, 19 Aug 1999, J. Dean; 2N, South Alligator River, d/s Gimbat, 13˚35'S 132˚36'E, 1 May 1990, D. Cartwright; 1MI, 1MSI, 1FSI, Jim Jim Creek, 3 km d/s Falls, 1 Sep 1979, J. Blyth; 2N, Magela Creek, u/s Bowerbird Billabong, 3 May 1990, D. Cartwright; 16N, Radon Springs, Kakadu National Park, 12˚45'S 132˚55'E, 23 Apr. 1990, D. Cartwright; 4N, Gulungul Creek, Radon Springs, Kakadu National Park, 12˚45'S 132˚55'E, 13 Aug 1999, J. Dean; 25N, Baroalba Creek, Kubarra Pools, 12˚49'S 132˚52'E, 28 Apr 1990, D. Cartwright; 11N, Baroalba Creek, Gubarra Pools, 12˚49'S 132˚52'E, 14 Aug 1999, J. Dean; 6N, Harris Creek, u/s South Alligator River, 1 May 1990, D. Cartwright. Western Australia. 27N, King Edward River, Mitchell River Road, 25 Sep 1995, L. Metzeling.   
 Description. Imago. Length of male: body 4.7–5.0 mm, forewing 5.1–5.2 mm; Length of female: body 4.1–5.9 mm, forewing 4.9–6.3 mm. Head predominantly medium brown; antennae pale yellow; ocelli white, black at base; male eyes with upper lobes pale orange-brown, in contact dorsally, lower lobes grey. Thorax with meso- and metascutum golden-yellow; lateral surfaces golden with some patches of medium brown. Forelegs with femora reddish-brown, tibiae pale yellow with apex medium brown, tarsi pale yellow; middle and hind legs uniformly pale yellow; forelegs of male with ratios of segment lengths 0.69–0.70; 1.00 (1.74–1.75 mm); 0.05; 0.27–0.31; 0.28; 0.18–0.21; 0.10–0.11; tarsal claws dissimilar, one claw with apical hook and ventral flange and the other expanded and pad-like, without terminal hook (Fig. 16). Forewings hyaline except for pterostigma, which is white and opaque; veins predominantly unpigmented (Fig. 14). Male abdomen reddish, dorsally pale tending to hyaline; each segment with narrow band of medium brown along posterior margin and pair of weakly developed brown lateral markings (Figs 17, 18); abdominal sterna pale, tending to hyaline. Penes lobes (Figs 19–22) narrow, fused almost to apex, each lobe with a stout, retractable apical spine; subapically with a robust, medial projection on the ventral surface. Female abdominal colour pattern similar to male, although generally a little darker, reddish-brown; abdominal sternum 9 strongly projecting, without apical excision (Fig. 23). Subimago. Wings pale greyish-yellow, abdominal colour pattern similar to imago. Mature nymph. General colour yellow. Mouthparts as in Figs 25–29. Legs yellow, all segments banded; foretarsus with 10–15 ventral spine-like setae, relatively uniform in length; tarsal claws with ventral teeth (Figs 30, 31). Abdomen yellow, each segment with pair of brown lateral markings (Fig. 24); gills linear, lateral tracheae very weakly developed or absent (Fig. 32).   
 Etymology. The name wapitja is derived from the word for ‘digging stick’ in the D _ ät_iwuy language of eastern Arnhem land (Ganambarr, 1999), and refers to the appearance of the male genitalia in lateral view.   
 Remarks. The genus is monotypic. Suter (1992) included this species in his keys as “ Leptophlebiidae Genus A sp.1”, and Dean (1999) designated the nymph “Genus V sp.AV1”.
description: Published as part of Dean, J. C. & Suter, P. J., 2004, Descriptions of new species and a new genus of leptophlebiid mayflies (Insecta: Ephemeroptera) from the Northern Territory, Australia, pp. 111-118 in Memoirs of Museum Victoria 61 (1) on pages 114-116, DOI: 10.24199/j.mmv.2004.61.8, http://zenodo.org/record/12209400