Heterogomphus Burmeister 1847
Creators: name: Neita Moreno, Jhon C.
name: Clavijo-Bustos, Julián
name: Ratcliffe, Brett C.
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Heterogomphus

description: Key to the adult males of Heterogomphus species from Colombia
Modified from Ratcliffe et al. (2020)
The absence of developed cephalic or pronotal structures is not a constant character that allows the differentiation of females and males because minor males may appear similar to females. However, males have the last abdominal ventrite medially emarginate (Fig. 10A), while females have the last abdominal ventrite completely arcuate, not emarginate (Fig. 10B).
1. Elytra completely, densely rugopunctate or with punctate rows................................................. 2
1´. Elytra smooth........................................................................................ 7
2. Elytra with long, moderately dense to dense setae............................................................ 3
2´. Elytra glabrous or only with short, sparse setae on apex....................................................... 4
3. Clypeal anterior margin truncate (Fig. 7B); pronotal punctures small (Fig. 7F).................................................................................... H. kogui Neita Moreno, Clavijo-Bustos, & Ratcliffe, new species
3´. Clypeal anterior margin broad, deeply and triangularly emarginate (Fig. 7A); pronotal punctures large (Fig. 7E)........................................................................................... H. carayoni Dechambre
4. Clypeal apex truncate................................................................... H. curvicornis Prell
4´. Clypeal apex broad, deeply and triangularly emarginate....................................................... 5
5. Color dark reddish brown; elytra finely punctate............................................... H. rugicollis Prell
5´. Color black; elytra densely punctate or rugopunctate......................................................... 6
6. Mesotibial apex with 2–3 large teeth; parameres as in Fig. 3L–M......................... H. schoenherri Burmeister
6´. Mesotibial apex with 1 large tooth; parameres as in Fig. 3H–I.............................. H. mniszechi (Thomson)
7. Clypeal apex truncate.................................................................................. 8
7´. Clypeal apex broadly emarginate......................................................................... 9
8. Mesotibia and metatibia with truncate apex (Fig. 10C).................................... H. dilaticollis Burmeister
8´. Mesotibia and metatibia at apex with two large teeth (Fig. 10D)............................ H. telamon (Burmeister)
9. Apex of the medial trough in the elevated section of the pronotum broad, wider than width of head, broadly and deeply emarginate at middle (Fig. 9B); parameres as in Fig. 3P–Q................................... H. ulysses Burmeister
9´. Apex of the medial trough in the elevated section of the pronotum narrower, subequal in width to head in major males or narrower in minor males, weakly emarginate at middle (Fig. 9A); parameres as in Fig. 3A–B..... H. chevrolatii Burmeister
description: Published as part of Neita Moreno, Jhon C., Clavijo-Bustos, Julián & Ratcliffe, Brett C., 2025, A review of the genus Heterogomphus Burmeister, 1847 (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) from Colombia, pp. 345-364 in Zootaxa 5590 (3) on page 352, DOI: 10.11646/zootaxa.5590.3.3, http://zenodo.org/record/14953221