Hyposoter flavicoxa Galsworthy & Shaw & Haraldseide 2023, sp. nov.
Creators: name: Galsworthy, Anthony
name: Shaw, Mark R.
name: Haraldseide, Håkon
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Hyposoter
Hyposoter flavicoxa

description: Hyposoter flavicoxa Galsworthy & Shaw sp. nov.
(Figs. 7, 8)
Diagnosis. Most similar to pale specimens of H. caedator, but easily distinguished by the sculpture of the propodeum, which is granulate throughout, with a broad, ill-defined area superomedia, and by the yellow underside to the scape. The reduced sculpture of the propodeum also distinguishes this species from H. alpicola.
Description. Wing length: 4.5 mm.
Head: face longer than broad, face and clypeus rather coarsely granulate, making punctures almost invisible; clypeus relatively narrow and strongly convex, with narrow and smooth apical margin; malar space shorter than basal width of mandibles; mandibles strongly narrowed towards apex; frons sculptured much as face, but vertex and temples much more finely granulate; occiput descending immediately behind ocelli, and temples very strongly narrowed behind eyes; antenna somewhat broadened in distal half, but distally tapering: 32 flagellomeres.
Mesosoma: pronotum without epomia; mesoscutum finely and evenly granulate throughout, position of notauli marked only by bands of slightly outstanding short ridges; mesopleuron, including speculum, finely granulate virtually throughout without visible punctures, apart from some short longitudinal striae in front of mesopleural fovea and a few longer striae beneath tegular ridge; metapleuron slightly more coarsely rugose, especially in a band along dorsal edge; scutellum finely granulate, keeled only at extreme base; propodeum without clear keels, edges of area superomedia and area petiolaris outlined by irregular lines of rugosity; first lateral areas and front part of area superomedia finely granulate, rest of propodeum strongly transversely rugose; costulae absent, but their position indicated by abrupt change in surface sculpture.
Wings narrow; fore wing 1 cu-a postfurcal; areolet sessile; nervellus vertical, slightly curved but not intercepted.
Metasoma: first metasomal tergite with almost parallel sides to petiole, widening distally to a rectangular postpetiole, all very finely granulate almost to base of petiole; glymma large and obvious; 2nd tergite 1.2x as long as wide, thyridia small and close to base; surface of 2nd tergite onwards smooth but not polished, apart from a slight rugosity in basal half of 2nd tergite; 6th and 7th tergites entire; ovipositor shorter than apical depth of metasoma, very slightly upcurved, with a strong subapical notch.
Colour: black; mandibles, underside of scape, fore and mid coxae, trochanters and trochantelli yellow, sometimes shaded with orange brown on mid coxa; first tergite black with reddish tinge at apex of postpetiole, 2nd tergite orange, darkened at least in basal half, often up to five-sixths, with a narrow black line on apex of tergite; tergites 3–5 orange, 6–7 dark brown to black; all femora, tibiae and tarsi orange, hind tibia usually only faintly darkened at extreme base and apex, sometimes more extensively at apex; visible sternites orange.
Male (based on a single specimen): very similar to female, but with fore and mid legs paler, almost clear yellow, and more extensive darkening on the hind part of the metasoma.
Remarks: biology unknown. We have seen specimens only from the UK and France.
Holotype: ♀, [UK] Tarrington, Herefordshire, SO6140, Malaise trap, 28.vii–6.viii.18, A.C.Galsworthy (NHMUK). Paratypes: 1♀, Chippenham Fen, Cambs, TL 650293, Malaise trap, carr at reedbed edge B, 30.7–11.8.85, J. Field, RMSNH 1986.021 ; 1♀, Abbots Moss, Cheshire, SJ 5868, Malaise Trap 2, Quercus / Betula / Pinus, 5–25.viii.86; R.R.Askew, NMSZ 1988.002 ; 1♀, Lot-et-Garonne, Bernac, Mal. TR. 26.6–3.7.91, R.R.Askew (all NMS). 2♀, Tarrington, Herefordshire, SO6140, light trap, 18–23.viii.16 and 15–22.viii. 2017 ; 1 ♁, 5♀, same location, Malaise trap, 28.vii–6.viii.18, A.C.Galsworthy (all NHMUK).
description: Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on pages 43-45, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256