Plateosaurus trossingensis Fraas 1913
Creators: name: Fernández, Omar Rafael Regalado
name: Stöhr, Henrik
name: Kästle, Benedikt
name: Werneburg, Ingmar
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Reptilia
Saurischia
Plateosauridae
Plateosaurus
Plateosaurus trossingensis

description: Plateosaurus trossingensis Fraas, 1913
Figs 6, 7, 24, 30
Plateosaurus quenstedti von Huene, 1905: 348.
Type material
Holotype
GERMANY • almost complete skeleton; Obere Mühle, Trossingen Formation; SMNS 13200.
Material examined
We treat the following specimens as comparable to Plateosaurus trossingensis but with distinctive morphotypes:
“ GPIT I” (Fig. 24)
GERMANY – Obere Mühle, Trossingen Formation • mounted skeleton in the collection, comprises most of the skeleton, and several cast elements, including a cast of the skull of SMNS 13200; GPIT-PV-30784 • original skull of the skeleton embedded partially in matrix; GPIT-PV- 111840 • left maxillary; GPIT-PV- 111842 • left squamosal; GPIT-PV- 111848 • left postorbital; GPIT-PV- 111849 • left jugal; GPIT-PV- 111845 • left quadratojugal; GPIT-PV- 111846 • left quadrate; GPIT-PV- 111847 • right quadrate; GPIT-PV- 111852 • right squamosal; GPIT-PV- 111853 • right postorbital; GPIT-PV- 111854 • right jugal; GPIT-PV- 111850 • right quadratojugal; GPIT-PV- 111851 • hyoid and tongue bones still in the matrix; GPIT-PV- 60260 • capitulum and tuberculum of a thoracic rib; GPIT-PV- 60257 • fragment of the neural arch of a caudal vertebra; GPIT-PV- 60256 • right distal carpal; GPIT-PV- 60338 • right manual phalanx; GPIT-PV- 60274 • right manual phalanx; GPIT-PV- 60275 • right manual phalanx; GPIT-PV- 60276 • right manual phalanx; GPIT-PV- 60277 • right manual phalanx; GPIT-PV- 60278 • right manual claw; GPIT-PV-60339 • left distal phalanx of manual digit V; GPIT-PV- 60337 • left manual phalanx; GPIT-PV- 60253 • right distal tarsal; GPIT-PV- 60254 • six distal caudal vertebrae with three chevrons; GPIT-PV- 60223.
Note: Plateosaurus trossingensis, gracile and medium-sized morphotype. This specimen was referred to as Pl. ‘ quenstedti ’ (von Huene 1932).
“ GPIT VI ” (Fig. 30)
GERMANY – Obere Mühle, Trossingen Formation • a right femur, a right tibia, a right fibula, a right astragalus, a complete set of the right metatarsals and pedal phalanges, still embedded in the matrix; GPIT-PV- 30789.
Note: Plateosaurus cf. trossingensis, gracile and medium-sized morphotype. Von Huene also referred to this specimen as Pl. ‘ quenstedti ’.
“ GPIT A” (Figs 6–7)
GERMANY – Pfrondorf, (?) Trossingen Formation • atlas; GPIT-PV- 60168 • axis; GPIT-PV- 60162 • anterior cervical 5; GPIT-PV- 60164 • posterior cervical 8; GPIT-PV- 60155 • posterior cervical 9; GPIT-PV-60156 • anterior dorsal 4(?); GPIT-PV- 60154 • fragment of the neural arch of posterior dorsal 14(?); GPIT-PV- 60159 • left scapula; GPIT-PV- 60152 • right ulna; GPIT-PV- 60172 • left ulna; GPIT-PV- 60163 • right pubis; GPIT-PV- 60153 • left femur with a reconstructed shaft; GPIT-PV- 60151 • proximal portion of right femur; GPIT-PV- 60150 • left fibula; GPIT-PV- 60170 • proximal and distal ends of metatarsal II; GPIT-PV- 60165 • proximal and distal ends of metatarsal III; GPIT-PV- 60171 • pedal ungual III with the distal end of phalanx III.3; GPIT-PV- 60161 • possibly pedal phalanx IV.2; GPIT-PV- 60157 • possibly pedal phalanx IV.3; GPIT-PV- 60158 • casts of axis; NHMUK PV R 3062 • cervical vertebra 5; NHMUK PV R 3063 • cervical vertebra 7; NHMUK PV R 3064 • cervical vertebra 8; NHMUK PV R 3065 • left pubis; NHMUK PV R3067 • right pubis NHMUK PV R 5845 • both ischia; NHMUK PV R 3066 • left ulna; NHMUK PV R 3071 • left radius; NHMUK PV R 3074 • left femur; NHMUK PV R 3068 • proximal part of the left tibia; NHMUK PV R 5849 • left fibula; NHMUK PV R 3070 • two phalanges; NHMUK PV R 3075 • left metatarsal; NHMUK PV R 3073 • left metatarsal; NHMUK PV R 3073 • claw; NHMUK PV R 3076.
Note: Plateosaurus cf. trossingensis, a gracile and medium-sized morphotype, holotype of Pl. ‘ quenstedti ’. The GPIT Collection still houses some of the holotype material. However, four cervical vertebrae, a left scapula, two humeri, a radius, the posterior half of the dorsal series, the sacrum, the ilia, the ischia, the right pubis, the left tibia, and the metatarsals are currently lost – the NHMUK in London houses casts of some of these elements.
Remarks
“GPIT I” is referred to as Pl. trossingensis (species 8 in Fig. 3); it is worth noting that a considerable amount of intraspecific variability between the specimens that make Pl. trossingensis needs to be studied in detail. The rest of the material from the Obere Mühle is considered as Pl. cf. trossingensis based on the stratigraphic framework that has been currently proposed for the Trossingen outcrop. Von Huene (1929, 1932) formulated the taxonomy of sauropodomorphs from the Late Triassic more akin to the polytypic approach, where morphotypes are used as criteria to produce what in modern usage would be equivalent to a subspecies. The specimen “GPIT A” from Pfrondorf (label c in Fig. 1), the holotype of Plateosaurus ‘ quenstedti ’ (species 3 in Fig. 3), shares several similarities in the ulna, the ilium and the femoral head morphology with GPIT-PV-30784. The holotype of Pl. ‘ quenstedti ’, however, is now incomplete, with several elements lost, although casts of most of these elements are now available in the Natural History Museum. Given these circumstances, it is reasonable to consider Pl. ‘ quenstedti ’ as a senior subjective synonym of Pl. trossingensis. Furthermore, if future studies conclude that the disparity between the holotype of Pl. trossingensis and GPIT-PV-30784 is substantial enough to warrant the recognition of a new species, then GPIT-PV-30784 can bear the name Pl. ‘ quenstedti ’ of which GPIT-A would be the holotype.
description: Published as part of Fernández, Omar Rafael Regalado, Stöhr, Henrik, Kästle, Benedikt & Werneburg, Ingmar, 2023, Diversity and taxonomy of the Late Triassic sauropodomorphs (Saurischia, Sauropodomorpha) stored in the Palaeontological Collection of Tübingen, Germany, historically referred to Plateosaurus, pp. 1-88 in European Journal of Taxonomy 913 (1) on pages 62-64, DOI: 10.5852/ejt.2023.913.2375, http://zenodo.org/record/10405782