Phanaeus halffterorum Edmonds 1979
Creators: name: Moctezuma, Victor
name: Halffter, Gonzalo
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Phanaeus
Phanaeus halffterorum

description: Phanaeus halffterorum Edmonds, 1979
Figs 1F, 15, 18G, 19G
Phanaeus halffterorum Edmonds, 1978: 321 (nomen nudum).
Phanaeus halffterorum Edmonds, 1979: 99, 102–105, figs 1–3, 6–8 (in part).
Phanaeus halffterorum – Halffter & Edmonds 1982: 88–89 (in part). — Anduaga & Halffter 1991: 157 (in part). — Delgado-Castillo et al. 1993: 125 (in part). — Deloya et al. 1993: 21, 39 (in part); 2014: 77 (in part). — Anduaga 2000: 125, 130 (in part). — López-Guerrero & Halffter 2000: 241 (in part). — Arnaud 2002b: 96 (in part). — Price 2005: 197 (in part); 2007: 17, figs 52–54. — Edmonds 2006: 31–32, 36, fig. 7 (in part). — Ceballos et al. 2009: 397. — Edmonds & Zídek 2012: 5 (in part). — Krajcik 2006: 150. — Moctezuma & Halffter 2017: 52, 54–55, fig. 23 (in part). — Moctezuma et al. 2017: 113–115, 118–119, 122, 130–132, figs 1–5; 2019: 253, fig. 5. — Lizardo et al. 2017: 273, 275, 292, fig.13 (in part). — Kohlmann et al. 2018: 69, 81, 88–89. — Gillett & Toussaint 2020: 2.
Phanaeus (Notiophanaeus) halffterorum – Edmonds 1994: 2, 8–9, 19, 39, 41, 43–44, 101, figs 211, 213, 217–218, 221 (in part); 2003: 61, 65 (in part). — Arnaud 2002b: 95 (in part). — Edmonds & Zídek 2012: 3, 12, figs 134–135, 137, 143–147 (in part). — Lizardo et al. 2017: 272 (in part). — Kohlmann et al. 2018: 80. — Zaragoza-Caballero et al. 2019: 43.
Type material
Holotype (not studied)
MEXICO • ♂, Edmonds 1979: 99; State of Mexico, Temascaltepec; CAS.
Paratypes revised (6 ♂♂, 2 ♀♀)
MEXICO – State of Mexico • 1 ♂; “ 5 km E Temascaltepec, Real de Arriba (2200 m), 10–VII–1976, fungus, oak-pine forest, W. D. Edmonds, P. Reyes, B. Kohlmann cols.”; IEXA • 1 ♀; same collection data as for preceding; TAMU • 1 ♂, 1 ♀; same collection data as for preceding; VMC • 2 ♂♂; “ 8 km W Temascaltepec, 2360 m, 11–VII–76, fungus in pine-oak forest, W. D. Edmonds, P. Reyes, B. Kohlmann cols.”; TAMU • 1 ♂; same collection data as for preceding; VMC • 1 ♂; “ Real de Arriba, VII–1932, 6300 ft, México D. F., Hinton coll., BM 1939–583 ”; TAMU.
Type locality
Mexico, State of Mexico, Temascaltepec.
Distribution
Central Trans-Mexican Volcanic Belt, State of Mexico and Morelos (Fig. 15).
Remarks
Mean length 17.4 mm (13.4–19.9 mm). The specimens from Morelos (Deloya et al. 1993) were not studied by us. This species was erroneously reported from Mexico City (Arnaud 2002b). This mistake is attributed to Hinton (1935), who recorded it from Real de Arriba, Mexico D.F. Real de Arriba is actually located in the State of Mexico. Despite the fact that Moctezuma et al. (2017) split P. halffterorum and P. bravoensis, the colouration pattern of P. halffterorum remains as indicated by Edmonds (1979), with bright metallic green or dark metallic blue specimens. In a review of the immature dung beetles of Scarabaeinae (Edmonds & Halffter 1978), the name P. halffterorum Edmonds, 1978 was published. Nevertheless, this may be considered as a nomen nudum under Article 13 of the Code (ICZN 1999). Consequently, the same name was available later for the same or a different concept under Arts 21, 50; while P. halffterorum Edmonds 1979 must be considered the available authorship and date.
To the original description of the male we add that the right lobe of the endophallite copulatrix is more developed than the left lobe; the right lobe ois btusely triangular; the left lobe is obtusely rectangular; the central ridge lis ess developed than the central column (Fig. 1F). For the female can be added that the trituberculate cephalic carina has conical, nearly aligned tubercles; the middle tubercle is more raised than the lateral tubercles; the pronotal process is trituberculate, with a posterior concavity; all the tubercles are rounded; the middle tubercle is slightly more developed and projected frontally than the lateral tubercles; the pronotal midline is distinctly impressed, with superficially impressed punctures; the pronotal surface is smooth, with almost effaced punctures.
description: Published as part of Moctezuma, Victor & Halffter, Gonzalo, 2021, Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species, pp. 1-71 in European Journal of Taxonomy 747 on pages 21-22, DOI: 10.5852/ejt.2021.747.1333, http://zenodo.org/record/4836744