Tomarus colombianus Lopez-Garcia & Gasca-Alvarez 2014
Creators: name: López-García, Margarita M.
name: Deloya, Cuauhtémoc
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Dynastidae
Tomarus
Tomarus colombianus

description: Tomarus colombianus López-García & Gasca-Álvarez, 2014
(Figs. 9G, 11I, 16D, 19D, 27D; 44)
Tomarus colombianus López-García & Gasca-Álvarez, 2014 in López-García et al. 2014: 579. Original combination. Male holotype (CIUQ) “Colombia—Valle / La Bocana-B/ventura / Altura: 5 m. s.n.m / Fecha: 16/04/04 / Método: Directo / Col: Serna F. // Tomarus colombianus / López-García & / Gasca-Álvarez / HOLOTIPO // CIUQ 1505”. Female paratype (CIUQ) “ COLOMBIA: Dep. Valle del Cauca / Buenaventura / La Bocana / 15.iv.2004 / Light trap / 2 m altitud / Rivas L. Coll // Tomarus colombianus / López-García & / Gasca-Álvarez / PARATIPO // CIUQ 1504”. Female paratype (CIUQ) “Colombia- Valle Cauca / B/ventura-Bocana / Altura: 2 m. s.n.m / Fecha: Mayo/04 / Método: Directo-Bosque / Col: Mendoza L. // Tomarus colombianus / López-García & / Gasca-Álvarez / PARATIPO // CIUQ 1507”. Male parat–pe (ICN) “Colombia—Valle / La Bocana-B/ventura / Altura: 5 m. s.n.m / Fecha: 17/04/04 / Método: Directo / Col: Serna F.” Female paratype (ICN) “ Huila / Neiva / 5.iii.1974 / Bagos A. I. // Tomarus colombianus / López-García & / Gasca-Álvarez / PARATIPO. Type locality: La Bocana, Valle del Cauca, Colombia.
Description. Habitus as in Fig. 27D. Length 18.0–21.0 mm; humeral width 8.5–10.5 mm. Color dark reddish brown. Head: Frons and clypeus coarsely and densely rugose. Frontoclypeal region with 2 conical tubercles separated by about 6 tubercle diameters. Clypeus narrowed towards apex, base 3 times wider than apex. Clypeal teeth triangular, separated by less than a tooth diameter. Mandible with 2 apical teeth and a lateral, narrowly rounded tooth. Mentum abruptly constricted at apical 3rd. Galea of maxilla with teeth 5 and 6 well developed. Interocular distance 3.2 times an eye width. Pronotum: Surface with minute punctures; denser and deeper on anterior and lateral angles, disc impunctate. Apical tubercle small, rounded, not visible in lateral view. Subapical fovea shallow, elongate, narrow (1/3 the interocular distance); equal in both sexes, surface rugopunctate. Scutellum: Surface with deep, large punctures forming 2 lines parallel to margins. Elytra: Punctures on first interval as large as those on other intervals. Inner surface of apex with rounded, large tubercles forming 14–16 parallel lines. Abdomen: Apex of tergite IV with a triangular wide area with small, irregular tubercles forming 9–10 diagonal nearly parallel lines. Pygidium with dense rugosity on basal 3rd (Fig. 9G), apex rounded. Venter: Apex of prosternal process flat, transversely oval to rounded. Metasternum with minute setae on anterior angles. Legs: Protibia tridentate, with a basal, small denticle (Fig. 11I). Apex of metatibia crenulate, with 9–11 spinules. Male genitalia: Spiculum gastrale without basal plates. Phallobase 1.1 times longer than parameres. Parameres with 2 dorsal teeth on each side; basal tooth acute, longer than apical (Figs. 16D, 19D). Internal sac with copulatory lamella and short lamellar spiny belt; with a complex of 7 spine-like accessory lamellae, with dense granules at base.
Diagnosis. Tomarus colombianus can be differentiated by the following character combination: frontoclypeal tubercles conical; pronotal surface nearly smooth (Fig. 27D); pronotal tubercle not visible in lateral view; pronotal fovea shallow and narrow (1/3 as wide as interocular distance); protibia tridentate, with an additional basal denticle (Fig. 11I); apex of metatibia with 9–11 spinules; parameres with 2 dorsal teeth on each side, main teeth long, widely separate from secondary teeth (Figs. 16D, 19D).
Distribution. Colombia (López-García et al. 2014).
Locality records (Fig. 44). Five specimens examined from ICN and CIUQ. COLOMBIA (5). Huila (1): Neiva. Valle del Cauca (4): Buenaventura.
Natural history. The few known specimens were found in tropical humid forest near a mangrove swamp and Guandal forest. They were collected by hand and at light traps.
description: Published as part of López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, pp. 1-119 in Zootaxa 5211 (1) on page 59, DOI: 10.11646/zootaxa.5211.1.1, http://zenodo.org/record/7345007