Acanthagrion Selys 1876
Creators: name: Mauffray, William F.
name: Tennessen, Kenneth J.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Odonata
Coenagrionidae
Acanthagrion

description: Acanthagrion Selys, 1876
A large genus of slender, usually small damselflies, mostly black but usually with blue, green or orange stripes on the thorax and tip of the abdomen (Fig. 12). The appendages of males slant posteroventrally and the apical rim of abdominal segment 10 is often elevated; females usually have paired mesepisternal fossae. Habitat includes vegetated ponds, lake margins, and slow areas of streams. For decades, species identification relied on Leonard (1977), but numerous errors and synonymies have been found in that publication. For recent taxonomic changes and additions, see Lozano (2007, 2013), von Ellenrieder & Lozano (2008) and Machado (2012a). The genus is in need of revision.
Acanthagrion ascendens Calvert, 1909. Publ. Recs. ECU: Paulson (1983 “?”), Tsuda (1986, 1991), (H08), Lozano (2007), (H14 — as A. cf. ascendens), (P16); GY: * Campos (1922 — as A. gracile var ascendens, redetermined as A. floridense by KJT), Leonard (1977); OR: Francisco de Orellana: Tennessen (2004 — “tentatively identified”). PA: Gloger (1967). Also BRA, COL, GUF, GUY, SUR, TTO, VEN: (L06), (P11); plus BOL, PER (H08), (P16); and PRY, (H08).
Addl. Data. OR, Francisco de Orellana, 1 ♂, pond near Río Savaleto, km marker 31, P.N.Y., 17 vii 1996, leg. & det. KJT [FSCA]; adult, T.B.S., Río Tiputini, 55 km SE of Pompeya, 94 km ESE of Coca (-0.6370, -76.1500, 230m), no date provided, leg. & det. WAH & DLW. Note: The ♂ examined has color variation—S9 is orange, as discussed by Tennessen (2004).
Note: A ♂ and ♀ from F. Campos taken at Tena were identified as A. ascendens. We checked the ♂ which is in FSCA and it is A. floridense. The ♀ was sent back to ANSP and may also be A. floridense.
Acanthagrion apicale Selys, 1976. Publ. Recs. ECU: (L06), (Lozano 2007), (H08), (P11), (H14). NA: Machado (2012a — as A. chicomendesi paratypes); OR: * Tennessen (1996b), Muzón & Lozano (2005), von Ellenrieder & Garrison, (2011). Also BOL, BRA, COL, GUF, PER, VEN: (L06), (H08), (P11); plus GUY: (H08); and SUR: (P16).
Addl. Data. MS, Gualaquiza: 3 ♂♂, 3 ♀♀, Gualaquiza (-3.40, -78.58, 800m), vi–v 1935, leg. F. Campos, det. MJW [FSCA]; OR, Francisco de Orellana: 4 ♂♂, 4 ♀♀, [La Nueva Esperanza] [across Río Napo from] near Primavera “Limoncocha” Río Napo & Lake Taracoa (-0.45, -76.77, 300m), 27 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 3 ♂♂, same data but 4 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, same data but swamp N of Lake, 26 viii 1980, leg. & det. SWD [SWD]; 1 ♂, [Limoncocha] [about 40 km east] or 100 km E of Coca [on Río Napo] Lago Garzacocha, swamp, La Selva Jungle Lodge (-0.4819, -76.3733, 235m), 12 x 1988, leg. & det. SWD [SWD]; 1 ♂, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 8, swamp & stream (-0.4694, -76.5869, 253m), 12 vi 1995, leg. & det. WFM [QZAC]; 7 ♂♂, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (-0.60, -76.47, 229m), 17 vii 1996, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 1 ♂, 1 ♀, same data but leg. & det. KJT [KJT]; 1 ♂, swamp pond, km marker 38 on Yasuni Road 12 vi 1995, leg. & det. KJT [KJT]; 5 ♂♂, open & forest swamps, main road, P.N.Y., 15 xi 1997, leg. & det. KJT [KJT]; 1 ♂, just S of Bogi 2 Este River, S of km 42 on Yasuni Road, swamp (-0.38, -76.30), 11 vi 1995, leg. & det. SWD [SWD]; 3 ♂♂, P.N.Y., 2.1 km E of Yasuni Biological Research sta., pond and swamp (-0.6741, -76.3685, 244m), 15 xi 1997, leg. & det. WFM [FSCA, QZAC, USNM]; 1 ♂, stream, road to Waorani Reserve, 5.6 km S of Yasuni Sci. Res. Sta., 10 vi 1995, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [FSCA]; 1 ♀, P.N.Y., 6.8 km E of Yasuni Biological Research Sta., small forested stream (-0.6545, -76.3417, 238m), 15 xi 1997, leg. & det. WFM [QZAC]; 1 ♂, P.N. Y., stream (-0.6542, -76.3542, 200m), 15 xi 1997, leg. & det. TWD [SWD]; 1 ♂, P.N. Y., swamp (-0.6508, -76.3558, 200m), 16 xi 1997, leg. & det. TWD [SWD]; 1 ♂, P.N. Y., stream (-0.7100, -76.4750, 200m), 16 xi 1997, leg. & det. TWD [SWD]; 10 ♂♂, 1 ♀, T. B.S., USFQ, black water stream crossing, Guacamayo trail at marker 2050 (-0.6356, -76.1492, 220m), 15–21 i 2009, leg. & det. RWG & NVE [MLPA, NVE, RWG, USFQ]; 1 ♀, T. B.S., USFQ, swamp at marker 650 of Parahuaco trail (-0.6397, -76.1453, 229m), 21 i 2009, leg. & det. RWG & NVE [NVE]; 1 ♂, T. B.S., USFQ, stream crossing Matapalo trail at marker 450, Ti 15 (-0.6369, -76.1458, 245m), 15 i 2009, leg. & det. RWG & NVE [RWG]; SU, Shushufindi: 1 ♂, Limoncocha, 25 viii 1980, leg. & det. SWD [SWD]; 1 ♀, Limoncocha, Río Napo (-0.40, 76.62, 300m), 21 ix 1974, leg. B. Drummond III, det. SWD [FSCA]; 1 ♀, Limoncocha, swamp, 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♂, trails, small streams 3 km W of Shushufindi, 2+ km N off Hwy. 26 at km marker 17 (-0.1664, -76.6917, 290m), 13 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, same data but leg. & det. KJT [KJT].
Note: Machado (2012a) split A. apicale into several species. After discussion with and evidence provided by Rosser Garrison (pers. comm., 2017), we recognize A. apicale as the species in Ecuador and A. chicomendesi as a possible junior synonym of A. apicale.
Acanthagrion cuyabae Calvert, 1909. Publ. Recs. ECU: (L06 — also as A. lancea), Lozano (2007 — also as A. lancea), (H08 — also as A. lancea), Rojas & Sanchez (2009 — as A. viridescens), (P11 — also as A. lancea) (P11 — as A. viridescens), (H14 — as both A. lancea and A. viridescens). OR: * Tennessen (1996b — as A. lancea), von Ellenrie- der & Garrison (2011 — as A. lancea). Also BOL, BRA: (L06), (H08), (P11); plus COL — as A. viridescens, PER (P16).
Addl. Data. NA, El Chaco: 1 ♂, Chaco, 53 km NE on road to Lago Agrio, ponds and stream, 18 viii 1980, leg. & det. SWD as A. lancea [SWD]; OR, Francisco de Orellana: 1 ♂, [La Nueva Esperanza] [across Río Napo from] near Primavera, Laguna Taracoa (-0.45, -76.77, 300m), 27 vii 1977, leg. DRP & S. Hills, det. DRP as A. viridescens [DRP]; 2 ♂♂, same data but 4 xi 1980, leg. MJW & D. Robinson, det. MJW as A lancea [FSCA]; 4 ♂, same data but 26 viii 1980, leg. & det. SWD as A. lancea [SWD]; 3 ♂♂, P.N.Y., 6.3 km S of Yasuni Road on Waorani Reserve Road, marshy pond (-0.7120, -76.4745, 260m), 10 vi 1995, leg. & det. WFM as A. lancea [FSCA, LSAM, QZAC, SWD]; 4 ♂♂, same data but leg. & det. KJT as A. viridescens [KJT]; 2 ♂♂, swamp pond at km 8, S of Río Napo on Yasuni Road (-0.6333, -76.5000), 12 vi 1995 leg. & det. SWD as A. lancea [RWG]; 3 ♂♂, same data but leg. & det. KJT as A. viridescens [KJT]; 1 cop. pair, P.N.Y., 29 km N of Waorani Road on Yasuni Road roadside ditch and pond (-0.4917, -76.5584, 183m), 11 vi 1995, leg. & det. KJT as A. viridescens [KJT]; 1 ♂, 1 ♀, same data but leg. & det. WFM as A. lancea [FSCA]; 2 ♂♂, [Pompeya Sur], S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (-0.60, -76.47, 229m), 12 vi 1995, leg. & det. WFM as A. lancea [FSCA]; 1 cop. pair, same data but 17 vii 1996, leg. & det. KJT as A. viridescens [KJT]; 3 ♂♂, same data but leg. & det. SWD as A. lancea [SWD]; 2 cop. pairs, 2 ♂♂, same data but leg. & det. WFM as A. lancea [FSCA, QZAC, USNM]; 1 ♂, same data but 17 xi 1997 [FSCA]; 1 ♂, 1 ♀, same data but leg. & det. JJD as A. viridescens [DRP, JJD]; 3 ♂♂, 1 ♀, same data but leg. & det. TWD as A. viridescens [SWD]; 2 cop. pairs, 2 ♂♂, Puerto Francisco de Orellana, 12.3 km W on road to Loreto from Coca-Sachas Road, pond (-0.4750, -77.0747, 290m), 13 vi 1995, leg. & det. WFM as A. lancea [FSCA, USNM]; 1 ♂, same data but leg. & det. SWD as A. lancea [UTIC]; 2 ♂♂, 1 ♀, Francisco de Orellana, 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools & flowing shallow marsh (-0.4367, -77.0017, 258m), 13 vi 1995, leg. & det. WFM as A. lancea [FSCA, QZAC, USNM]; 1 ♂, same data but 18 xi 1997 [FSCA]; 4 ♂♂, Puerto Francisco de Orellana 0.6 km W on road to Loreto from Río Payamino, marsh (-0.4347, -77.0011, 200m), 20 vii 1996, leg. & det. WFM as A. lancea [FSCA, QZAC, USNM]; 1 ♂, same data but 18 xi 1997 [FSCA]; 2 ♂♂, same data but leg. & det. TWD as A. viridescens [FSCA, SWD]; SU, Shushufindi: 6 ♂♂, Limoncocha (-0.40, -76.62, 300m), 27 xii 1971, leg. D. Pearson, det. DRP as A. viridescens [DRP]; 1 ♀, same data but 6 i 1972 [DRP]; 2 ♂♂, 11–16 viii 1972 [DRP]; 6 ♂♂, same data but 3–16 vi 1977, leg. P. Spangler & D. Givens, det.? as A. lancea [USNM]; 3 ♂♂, same data but 23 viii 1980, leg. & det. SWD as A. lancea [SWD]; 2 ♂♂, same data but 25 viii 1980 [SWD]; 1 ♂, same data but 14 v 1975, leg. T. Emmel, det. MJW as A. lancea [FSCA]; 1 ♀, same data but 23 viii 1980, leg. SWD, det. KJT as A. viridescns [KJT]; 16 cop. pairs, 62 ♂♂, 4 ♀♀, same data but 1–27 xi 1980, leg. MJW & M. Robertson, det. MJW as A. lancea [FSCA]; 2 cop. pair, 52 ♂♂, 10 ♀♀, same data but Limoncocha, lakeshore, 23–26 vii 1977, leg. & det. DRP as A. viridescens [DRP]; 3 ♂♂, [Limoncocha] [about 40 km E] 100 km E of Coca [on Río Napo], Lago Garzacocha, La Selva Jungle Lodge (-0.4819, -76.3733, 235m), 12 x 1988, leg. & det. SWD as A. lancea [FSCA, SWD]; 1 ♂, same data but 23 xi 1997, leg. & det. TWD as A. viridescens [SWD]; 1 ♂, La Selva, Heron Lake, 4 v 1990, leg. E Smith, det. WFM [SWD].
Note: Leonard (1977) considered A. cuyabae and its two subspecies as synonyms of A. lancea Selys, 1876. Federico Lozano (pers. comm., 2019) found that A. cuyabae is a valid species and that A. viridescens is a synonym of A. cuyabae. Most published records and many specimens in various collections from Ecuador are misidentified as either A. viridescens or A. lancea; probably all such records should be referred to A. cuyabae.
Acanthagrion floridense Fraser, 1946. Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (L06), Lozano (2007 — also as A. gracile), (H08), Rojas & Sanchez (2009), (P11), (H14). NA: Gloger (1967 — as A. gracile); Leonard (1977 — as A. peruvianum); OR: Tennessen (1996b), von Ellenrieder & Garrison (2011 — as A. peruvianum); PA: * Leonard (1977); ZC: Tennessen (2000), Tennessen & Johnson (2008). Also ARG, PER: (L06), (H08), (P11); plus COL: (P11, P&P); and BRA: (H08); and URY: (L06), (P11).
Addl. Data. LJ, Loja: [UTIC]; MS, Santiago: [FSCA, TWD, USNM]; NA, Carlos Julio Arosemena Tola: [USNM; El Chaco: [SWD]; Tena: [DRP, FSCA, UTIC, JTJ, KJT, RWG, SWD, TWD, USNM]; OR, Francisco de Orellana: [DRP, FSCA, UTIC, KJT, RWG, SWD, TWD, USNM]; Loreto: [FSCA, KJT, SWD, SWD]; PA, Mera: [USNM]; Pastaza: [USNM]; Santa Clara: [USNM]; SU, Cascales: [SWD, USNM]; Lago Agrio: [FSCA, KJT, SWD, USNM]; Shushufindi: [FAL, FSCA, KJT, SWD, USNM]; TU, Baños: [DRP, RWG]; ZC, El Pangui: [FSCA]; Yantzaza: [FSCA, USNM]; Zamora: [FSCA, UTIC, JTJ, KJT, USNM].
Note: All the published records and a majority of the specimens in various museums under “Addl. Data” above are labeled A. peruvianum which has only recently been determined as a junior synonym of A. floridense (Lozano et al. 2017). A ♂ and ♀ from F. Campos collected at Tena were labeled as A. ascendens; we checked the ♂ which is in FSCA and it is A. floridense. The ♀ was sent back to ANSP and may be labeled A. ascendens but it is probably A. floridense also.
Acanthagrion longispinosum Leonard, 1977. Publ Recs. BOL, BRA (H08).
Addl. Data. OR, Francisco de Orellana: 1 ♀, T.B.S. Río Tiputini, 55 km southeast of Pompeya, 94 km ESE of Coca (-0.6370, -76.1500), 21 i 2012, leg. DLW, John Ascher, WAH, Fred Morrison, det. RWG, data provided by RWG.
Acanthagrion obsoletum (Förster, 1914). Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), Lozano (2007 — also as A. hartei), (P11 — also as A. hartei), (H14 — also as A. hartei); MS: * Muzón & Lozano (2005 — as A. hartei, Holotype & Allotype: Bomboiza, 17 km S of Gualaquiza (-3.4333, -78.5167, 880m), 10 ix 1990, leg. OSF [USNM]); NA: * Förster (1914 — type locality: “ Napo ”), Bridges (1993), Gloger (1967 — as A. leonora); Leonard (1977), Muzón & Lozano (2005), Machado (2012a); OR: Tennessen (1996b), von Ellenrieder & Garrison (2011); PA: * Förster (1914 — ALLOTYPE), Leonard (1977); SU: * Muzón & Lozano (2005 — as A. hartei); ZC: Tennessen (2000), Tennessen & Johnson (2008 — as A. hartei). Also BOL, COL, PER: (H08), (P11).
Addl. Data. MS, Huamboya: [FSCA, KJT, USNM]; Morona: [JJD]; Santiago: [FSCA, SWD]; NA, Carlos Julio Arsemena Tola: [FSCA, USNM]; El Chaco: [SWD]; Tena: [DRP, FSCA, UTIC, KJT, RWG, SWD, TWD, USNM]; OR, Francisco de Orellana: [DRP, FAL, FSCA, KJT, RWG, SWD, TWD, USNM]; SU, Lago Agrio: [FSCA, KJT, RWG, SWD]; Shushufindi: [DRP, FSCA, KJT, SWD]; PA, Pastaza: [FSCA, JTJ, KJT]; Santa Clara: [FSCA, UTIC, RWG, USNM]; ZC, El Pangui: [FSCA]; Yantzaza: [FSCA, USNM]; Zamora: [FSCA, UTIC, USNM].
Note: According to Garrison (pers. comm. 2017), A. hartei Muzón & Lozano is a junior synonym of A. obsoletum.
Acanthagrion phallicorne Leonard, 1977. Publ. Recs. ECU: *(L06), Lozano (2007), (P11), (H14). Also BRA, PER: (L06), (H08), (P11).
Addl. Data. NA, Tena: 1 ♂, Río Chinchipino, oxbow pond, 27 km E of Puerto Napo (-1.0728, -77.5864, 385m), 13 xi 1997, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 1 ♂, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto [road to Yasuni Research station] (-0.60, -76.47, 230m), 12 vi 1995, leg. & det. WFM [FSCA].
Acanthagrion trilobatum Lenoard, 1977. Publ. Recs. ECU: Lozano (2007), (P11), (H14). SD: * Daigle et al. (1997); ES: * Daigle et al. (1997); IM: * Daigle et al. (1997); LJ: Tennessen & Johnson (2008). Also CEN, COL, VEN: (H08), (P11).
Addl. Data. BO, Caluma: [FSCA, KJT]; CT, Pangua: [FSCA, JTJ, KJT]; EO, Arenillas: [FSCA, KJT]; Santa Rosa: [FSCA, JTJ, KJT]; ES, San Lorenzo: [FAL, FSCA, KJT, USNM]; GY, Balao: [FSCA, JTJ, KJT]; Naranjal: [FSCA]; LJ, Loja: [FSCA, KJT]; LR, Buena Fé: [DRP, FAL, SWD, USNM]; MN, El Carmen: [FSCA, JTJ, KJT]; Flavio Alfaro: [JTJ]; PC, Quito: [FSCA, KJT, USFQ]; San Miguel Bancos: [FSCA, KJT, RWG, USFQ]; SD: [FSCA, JJD, JTJ, KJT, SWD, USNM].
Acanthagrion yungarum Ris, 1918. Publ. Recs. ECU: Lozano (2007), (P11), (H14); NA: Tennessen (1996b), *Daigle (1996); ZC: Tennessen & Johnson (2008). Also PER: (H08), (P11); plus COL: (P16).
Addl. Data. MS, Gualaquiza: 1 ♂, 1 ♀, Gualaquiza (-3.40, 78.58, 800m), iv–v 1935, leg. F. Campos, det. KJT 2011—labeled A. gracile by P. Calvert, [ANSP, FSCA]; Morona: 1 cop. pair, 1 ♂, 1 ♀, Río Copueno, E side of Macas (-2.3019, -78.1133), 17 ix 2005, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. JJD [JJD]; NA, Archidona: 1 ♂, [Mondayacu], 28 km N of Tena on road between Tena & Baeza (-0.80, -77.77, 884m), 27 x 1980, leg. & det. MJW [FSCA]; 2 cop. pairs, 9 ♂♂, 1 ♀, [Osayacu] 33 km N of Tena on road between Tena & Baeza (0.78, -77.78, 975m), 27 x 1980, leg. & det. MJW [FSCA]; 1 ♂, 1 cop. pair, [Narupa] E on road to Loreto, 43.5 km W of Loreto, seep near large rocky river [Río Hollin] (-0.7022, -77.7189, 1036m), 14 vi 1995, leg. & det. WFM [FSCA, USNM]; 2 ♂♂, same data but 15 vii 1996 [QZAC, USNM]; 2 ♂♂, same data but leg. & det. SWD [SWD]; 2 ♂♂, [Narupa] E on road to Loreto, 43.5 km W of Loreto, seep near large rocky river [Río Hollin] (-0.7022, - 77.7189, 1036m), 14 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, pool, Loreto Road, 44 km E of Baeza / Tena Road, 15 vii 1996, leg. & det. KJT [KJT]; 1 ♂, km 45 on Hollen-Loreto Road (-0.7228, -77.5653, 1130m), 22 xii 1987, leg. M. & J. Wasbauer, det. RWG [RWG]; 2 ♂♂, [Narupa] E off road to Loreto, 1 km NW Bridge, Río Hollin (-0.6875, -77.7333, 914m), 4 xi 2001, leg. & det. F. Sibley [FSCA]; El Chaco: 12 ♂♂, 1 ♀, El Chaco 28.3 km N on Hwy. 45 near Salado, adjacent to Río Quijos, marsh and borrow pit (-0.1831, -77.6789, 1237m), 7 vi 1995, leg. & det. WFM [FSCA, QZAC, USNM]; 3 ♂♂, same data but leg. & det. SWD [RWG, SWD]; 5 ♂♂, 1 ♀, same data but leg. & det. KJT [KJT]; OR, Loreto: 1 ♂, Loreto, 30.7 km W on road to Narupa, trickle (-0.7381, -77.5119, 920m), 14 vi 1995, leg. & det. WFM [FSCA]; 2 ♂♂, pond & small stream, 34 km W of Loreto (835m), 14 vi 1995, leg. & det. SWD [FAL]; 1 cop. pair, 5 ♂♂, same data but leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 2 ♂♂, same data but leg. & det. KJT [KJT]; 1 cop. pair, 2 ♂♂, Loreto, 38.9 km W on road to Narupa, pond (-0.7411, -77.5569, 988m), 14 vi 1995, leg. & det. WFM [FSCA, USNM]; 1 ♂, same data but leg. & det. KJT [KJT]; 3 ♂♂, same data but leg. & det. SWD [SWD]; 1 ♂, Loreto, 35.2 W on road to Narupa, pond (-0.7414, -77.5481, 927m), 19 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. KJT [KJT] 1 ♂, same data but leg. & det. TWD [FSCA]; 6 ♂♂, 2 ♀♀, pond along Loreto-Archidona road (-0.7417, -77.5517, 900m), 19 xi 1997, leg. & det. TWD [SWD]; PA, Mera: 1 ♂, Shell (-1.52, -78.07, 914m), 7 xi 2001, leg. & det. F. Sibley [FSCA]; Pastaza: 1 ♂, 1 ♀, pond near Comunidad Chuwitayo, S of Puyo, 9 xi 1997, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, pond 3 km N of Comunidad Chu- witayo, S of Puyo, 9 xi 1997, leg. & det. KJT [KJT]; 1 ♂, pond, 8.7 km S of Veracruz, Hwy. 45, 11 xi 1997, leg. & det. KJT [KJT]; 1 ♂, pond, about 4 km SE off Hwy. 45, N of Puyo (-1.3422, -77.8681, 945m), 8 x 2009, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. JTJ [JTJ]; 2 ♂♂, Puyo, 16 km N on Tena Road (Hwy. 45), near Teniente Hugo Ortiz, small stream [listed at 2000 feet], (-1.3475, -77.9183, 1036m), 26 x 1980, leg. & det. MJW [FSCA]; 2 ♂♂, Cuasha, about 1.5 km N on Hwy. 45 [91.5 km N of Sucua], hard bottomed pond (-1.85, -77.82, 884m), 9 xi 1997, leg. & det. WFM [FSCA, QZAC]; 2 tan. pairs, 3 ♂♂, Cuasha approx. 4 km N on Hwy. 45 [94 km N of Sucua], cut grass pond (-1.83, -77.82, 884m), 9 xi 1997, leg. & det. WFM [FSCA, QZAC, USNM]; 2 ♂♂, pond on road between Macas and Puyo (-1.8417, -77.8283, 900m), 9 xi 1997, leg. & det. TWD [SWD]; 1 ♂, 1 ♀, 12 km E of Puyo, 2 ii 1976, leg. P. Spangler et al., det.? [USNM]; Santa Clara: 1 ♂, 1 ♀, 39 km NE of Puyo, 5 xi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; SU, Gonzalo Pizarro: 1 ♂, 65 km W of Lago Agrio, roadside pond, 18 v 1975, leg. A Gurney, det.? [USNM]; ZC, Zamora: 1 ♂, roadside ditch, 2.1 km ENE of Zamora (915m), 5 xi 1997, leg. & det. KJT [KJT]; 1 ♂, pool near Río Sabanillo, 10 km W of Zamora (1220m), 5 xi 1997, leg. & det. KJT [KJT]; 1 ♂, tilapia ponds and small streams, Hosteria El Arenal, about 2 km S of Cumbaratza (-4.0017, -78.8753, 950m), 2 iv 2008, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. JTJ [JTJ].
description: Published as part of Mauffray, William F. & Tennessen, Kenneth J., 2019, A Catalogue and Historical Study of the Odonata of Ecuador, pp. 1-265 in Zootaxa 4628 (1) on pages 29-33, DOI: 10.11646/zootaxa.4628.1.1, http://zenodo.org/record/3335707