Abida secale subsp. ionicae Kokshoorn & Gittenberger 2010, subspec. nov.
Creators: name: Kokshoorn, Bas
name: Gittenberger, Edmund
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Chondrinidae
Abida
Abida secale
Abida secale ionicae

description: Abida secale ionicae subspec. nov. (pl. 7 figs D–K)   
Type series.— Andorra. Holotype (RMNH 109059): Arinsal (= 8 km NNW of Andorra la Vella), Alt de la Capa mtn (= 2.8 km WSW of Arinsal), 2,500 m alt., 42.56203°N / 1.45567°E, B. Kokshoorn leg., 16-ix-2007. Paratypes: type locality (RMNH 109060 /20); Alt de la Capa mtn, 2,350 m alt., 42.56180°N / 1.45981°E, B. Kokshoorn leg., 16-ix-2007 (RMNH 109061 /16); Alt de la Capa mtn, 2,100 m alt. 42.56355°N / 1.46659°E, B. Kokshoorn leg., 16-ix-2007 (RMNH 109062 /31); Pal (= 5.6 km NW of Andorra la Vella), N-side above town, 42.54618°N / 1.47136°E, B. Kokshoorn leg., 15-ix-2007 (RMNH 109063 /30); Pas de la Casa, 2,100 m alt., “ Wageningse Studenten leg.”, vi-2002 (RMNH 109064 /3); Pico Mainera mtn, M. Vilella leg. (RMNH 111881 / 1 paratype).  
Distribution.—See fig. 2.  
Description (n=58).—Shell slender cylindrical to slightly fusiform, with 7–8 3/4 weakly inflated whorls, sculptured with regularly placed, moderate to strong axial ribs. The sculpture is often partially obsolete. Body whorl obliquely flattened and more or less narrowed, with a prominent keel. A slight external indentation accompanies the palatalis superior; at the position of the infraparietalis the apertural lip is mostly slightly indented. The palatal side of the aperture appears vertical (in lateral view). Umbilicus open. Shell height, 5.1– 7.1 mm, width, 2.0– 2.7 mm.  
Apertural lip thickened, continuous at parietal side or connected by a more or less prominent callus. Occasionally the aperture slightly protrudes. Angularis connected with spiralis. Subangularis lamelliform and more or less strongly developed. Infraparietalis prominent. Columellaris reaching only slightly beyond the columella, at its front end clearly thicker than the other lamellae; ending abruptly and sometimes connected with an indistinct fold on the apertural lip. Infracolumellaris longer, often reaching the apertural edge. Infrapalatalis, palatalis inferior and palatalis superior clearly indented and reaching the edge of the aperture. At the weak anterodorsal palatal center a suprapalatalis and often a suturalis are present. At the apertural edge several more or less indistinct folds may be present.  
 Differentiation.— Abida s. ionicae differs clearly from A. s. andorrensis. Although these subspecies are connected by a series of intermediates, the extremes are easily distinguishable. Shells of Abida s. ionicae are smaller than A. s. andorrensis (5.1–7.1 mm versus 9.6–11.8 mm in height) and much more slender. The former subspecies is found at c. 1,600 m altitude and higher, whereas A. s. andorrensis is known from c. 900– 1,500 m altitude. Abida s. ionicae differs from the other high-altitude forms by its slightly more cylindrical shape and the very prominent axial ribbing. A few shells are known from the province of Lérida, Spain: Pico Mainera mtn, NW of Sort (pl. 7 fig. K).   
Ecology.—The subspecies is known from altitudes between 1,600 and 2,700 m, where it occurs in Andorra on calcareous schist with limonitized pyrite crystals.  
Derivatio nominis.—This subspecies is named after Ms. Ionica Smeets, a good friend of the first author, to celebrate 15 years of friendschip.  
Genetic barcode.—GenBank accession number EU395371.
description: Published as part of Kokshoorn, Bas & Gittenberger, Edmund, 2010, Chondrinidae taxonomy revisited: New synonymies, new taxa, and a checklist of species and subspecies (Mollusca: Gastropoda: Pulmonata) 2539, pp. 1-62 in Zootaxa 2539 (1) on page 22, DOI: 10.11646/zootaxa.2539.1.1, http://zenodo.org/record/10094211