Triepeolus kathrynae , Rozen 1989
Creators: name: Onuferko, Thomas M.
name: Rightmyer, Molly G.
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Apidae
Triepeolus
Triepeolus kathrynae

description: Triepeolus kathrynae Rozen, 1989
Figs 11E, 12E, 19, 20A
Triepeolus kathrynae Rozen, 1989: 10 (♀, ♂), figs 7–8, 18–21, 26–32.
Triepeolus species B – Rozen 1966: 14, figs 15–18 (description, illustrations of postdefecating larva).
Triepeolus kathrynae – Rozen 2001: figs 7–13 (illustrations of mature larva).
Proposed common name
Kathryn’s triepeolus.
Diagnosis
The following morphological features in combination tell T. kathrynae apart from all other species of Triepeolus: the axillae have pale pubescence all along their margins and are large, each with the tip extending well beyond the midlength of the mesoscutellum and the free portion distinctly pointed, hooked (i.e., concave along the medial margin), and mesally unattached to the mesoscutellum for approximately half the medial length of the axilla (Figs 19B, 20A). In T. bimorulus Rightmyer, 2008 and T. simulatus Rightmyer, 2008, which are not in the simplex species group, the axillae also have pale pubescence all along their margins but extend little if at all beyond the midlength of the mesoscutellum. Males of T. kathrynae are very similar in overall appearance to those of T. grandis (Friese, 1917) (outside of the simplex species group), but in both sexes of T. grandis the axillae do not have pale pubescence all along their margins.
Etymology
The specific epithet (declined in the genitive case) honors Kathryn Gail Rozen-Gagnon, granddaughter of the taxonomic authority.
Material examined
Primary type material
USA • ♀, holotype; New Mexico, 11 mi N of Rodeo, Hidalgo County; 19 Aug. 1968; J.G. Rozen and M. Favreau leg.; AMNH AMNH_IZC 00324363.
Secondary type material
MEXICO – Chihuahua • 1 ♀, paratype; 18 mi W of Jiménez; 10 Aug. 1951; H.E. Evans leg.; KUNHM M.G.R. Database No. 3838. – Durango • 1 ♂, paratype; 8 mi S of Canutillo; 9 Aug. 1951; H.E. Evans leg.; KUNHM SEMC26/M.G.R. Database No. 5579 • 1 ♀, paratype; San Juan del Río; 7 Aug. 1951; H.E. Evans leg.; KUNHM M.G.R. Database No. 3836 • 1 ♂, paratype; same data as for preceding; 7 Aug. 1951; H.E. Evans leg.; KUNHM M.G.R. Database No. 3839. – Guerrero • 1 ♀, paratype; Mezcala; 29 Jun. 1951; H.E. Evans leg.; KUNHM M.G.R. Database No. 3837. – Morelos • 1 ♀, paratype; 4.3 mi W of Yautepec; 17 Aug. 1962; Ordway and Marston leg.; KUNHM SEMC25 /M.G.R. Database No. 5578.
USA – Arizona • 1 ♀, paratype; 1 mi E of Douglas, Cochise County; 16 Aug. 1974; Rozens and Favreau leg.; AMNH M.G.R. Database No. 1711 • 1 ♀, paratype; same data as for preceding; 20 Aug. 1975; J.G. Rozen leg.; AMNH M.G.R. Database No. 1710 • 1 ♂, paratype; same data as for preceding; 20 Aug. 1975; J.G. Rozen leg.; AMNH M.G.R. Database No. 1727 • 1 ♂, paratype; 2 mi NE of Portal, Cochise County; 1 Aug. 1959; M. Statham leg.; AMNH M.G.R. Database No. 1726 • 1 ♂, paratype; 7 mi E of Elfrida, Cochise County; 3 Aug. 1961; E.G. Linsley leg.; AMNH M.G.R. Database No. 1725 • 3 ♂♂, paratypes; Madera Canyon (Santa Rita Mountains), Pima County; 13 Aug. 1952; M. Cazier, R. Schrammel, and C. and P. Vaurie leg.; AMNH M.G.R. Database Nos 1480, 1728, 1729 • 1 ♂, paratype; Southwestern Research Station (5 mi W of Portal), Cochise County; 24 Jul. 1956; C. and M. Cazier leg.; AMNH M.G.R. Database No. 1717 • 1 ♂, paratype; same data as for preceding; 24 Jul. 1956; E. Ordway leg.; AMNH M.G.R. Database No. 1722 • 1 ♂, paratype; same data as for preceding; 25 Jul. 1956; C. and M. Cazier leg.; AMNH M.G.R. Database No. 1720 • 1 ♂, paratype; same data as for preceding; 3 Aug. 1956; E. Ordway leg.; AMNH M.G.R. Database No. 1715 • 1 ♂, paratype; same data as for preceding; 5 Aug. 1956; E. Ordway leg.; AMNH M.G.R. Database No. 1713 • 1 ♂, paratype; same data as for preceding; 6 Aug. 1956; E. Ordway leg.; AMNH M.G.R. Database No. 1723 • 2 ♂♂, paratypes; same data as for preceding; 10 Aug. 1956; C. and M. Cazier leg.; AMNH M.G.R. Database Nos 1718, 1721 • 1 ♂, paratype; same data as for preceding; 10 Aug. 1956; E. Ordway leg.; AMNH M.G.R. Database No. 1716 • 1 ♂, allotype; same data as for preceding; 15 Aug. 1956; E. Ordway leg.; AMNH AMNH_IZC 00324364 • 1 ♂, paratype; same data as for preceding; 18 Aug. 1956; C. and M. Cazier leg.; AMNH M.G.R. Database No. 1714 • 1 ♀, paratype; same data as for preceding; 21 Aug. 1956; C. and M. Cazier leg.; AMNH M.G.R. Database No. 1712 • 1 ♂, paratype; same data as for preceding; 24 Aug. 1956; E. Ordway leg.; AMNH M.G.R. Database No. 1719 • 1 ♀, paratype; Tumacacori, Santa Cruz County; 25 Aug. 1981; E.G. and J.M. Linsley leg.; USNM 0930-0959 /M.G.R. Database No. 5715 • 1 ♀, paratype; same data as for preceding; 29 Aug. 1981; E.G. and J.M. Linsley leg.; USNM 0900-0929 /M.G.R. Database No. 5714. – New Mexico • 1 ♀, paratype; 1 mi N of Rodeo, Hidalgo County; 23 Aug. 1963; M.A. Cazier and M. Mortenson leg.; USNM 0930-0959 /M.G.R. Database No. 5716 • 1 ♀, paratype; 5 mi N of Carlsbad, Eddy County; 21 Sep. 1956; J.W. MacSwain leg.; USNM M.G.R. Database No. 5713. – Texas • 1 ♂, paratype; 17 mi S of Kent, Culberson County; 3 Jul. 1963; J.G. and B.L. Rozen leg.; AMNH M.G.R. Database No. 1724 • 1 ♀, paratype; Carrizo Springs, Dimmit County; 10 Jun. 1948; M. Cazier leg.; AMNH M.G.R. Database No. 1709.
DNA barcoded material
Available. BIN: BOLD:AAW8068. Specimens examined and sequenced:
MEXICO • 1 ♀; Sonora, 30 km E of Agua Prieta; 31.3267° N, 109.2520° W; 26 Jul. 2000; K. Toal leg.; BOLD sample ID: CCDB-34570 D05; RLM.
USA – Arizona • 1 ♀; 9 km W of Bisbee on Hwy 92, Cochise County; 31.3788° N, 109.9635° W; 30 Aug. 2013; L. Graham leg.; BOLD sample ID: CMNTO_005; PCYU • 1 ♀; Bisbee (Hwy 92), Cochise County; 31 Aug. 2013; BOLD sample ID: CMNTO_007; PCYU • 1 ♀; Portal Road (7 mi marker), Cochise County; 28 Aug. 2004; BOLD sample ID: 04-AZ-0942; PCYU.
Non-barcoded material
MEXICO – Baja California Sur • 2 ♂♂; 3.7 mi N of La Burrera; 7–8 Oct. 1975; R.R. Snelling leg.; LACM M.G.R. Database Nos 5426, 5428 • 1 ♀; 4 km N of Los Barriles; 4 Sep. 1977; R.R. Snelling leg.; LACM M.G.R. Database No. 3381 • 1 ♀; Estación de Microondas Ligüí vicinity (48 km S of Loreto); 7 Sep. 1977; R.R. Snelling leg.; LACM M.G.R. Database No. 3382. – Chihuahua • 1 ♂; 17 km S of Samalayuca; 31 Aug. 1992; B.N. Danforth leg.; USNM M.G.R. Database No. 6038 • 1 ♂; 18 mi W of Jiménez; 10 Aug. 1951; H.E. Evans leg.; KUNHM M.G.R. Database No. 4578. – Jalisco • 2 ♂♂; 3 mi SE of Plan de Barrancas; 8 Jul. 1963; F.D. Parker and L.A. Stange leg.; UCBME M.G.R. Database Nos 5429, 5431. – Nuevo León • 1 ♂; Vallecillo; 2–5 Jun. 1951; H.E. Evans leg.; KUNHM M.G.R. Database No. 4577. – Sonora • 1 ♀; 106 km E of Hermosillo; 16 Aug. 1991; J.G. Rozen and N. Pember leg.; KUNHM SM0326509 /M.G.R. Database No. 7129 • 1 ♀; 50 km NE of Hermosillo; 14 Aug. 1991; R. Ayala leg.; KUNHM SM0326511 /M.G.R. Database No. 7131 • 1 ♀; 55 km NE of Hermosillo; 13 Aug. 1991; J.L. Neff leg.; KUNHM SM0326512 /M.G.R. Database No. 7132 • 1 ♀; Mazocahui; 13 Aug. 1991; L. Godinez leg.; KUNHM SM0326513 /M.G.R. Database No. 7130.
USA – Arizona • 1 ♀; 1 mi E of Douglas, Cochise County; 13 Aug. 1969; M.A. Cazier leg.; UCBME 1645-1700 /M.G.R. Database No. 11617 • 1 ♀; same data as for preceding; 15 Aug. 1994; J.G. Rozen, S.A. Budick, and G. LeBuhn leg.; AMNH M.G.R. Database No. 1739 • 1 ♂; 1 mi N of Douglas, Cochise County; 16 Aug. 1967; E.G. Linsley leg.; UCBME M.G.R. Database No. 5427 • 1 ♀; same data as for preceding; 16 Aug. 1969; E.G. Linsley leg.; UCBME M.G.R. Database No. 11619 • 2 ♀♀; 12 mi NW of Douglas, Cochise County; 30 Aug. 1989; J.G. and B.L. Rozen and R.L. Foster leg.; AMNH M.G.R. Database Nos 1736, 1737 • 1 ♀; 13 mi S of San Simon, Cochise County; 32.0719° N, 109.1797° W; 1 Sep. 2004; PCYU • 1 ♂; 14 km NNE of Portal, Cochise County; 31 Aug. 1989; KUNHM M.G.R. Database No. 5423 • 1 ♂; 15 mi SW of Apache, Cochise County; 18 Aug. 1997; J.G. Rozen and B. Adams leg.; AMNH M.G.R. Database No. 3290 • 1 ♂; 16 mi NE of Douglas, Cochise County; 18 Aug. 1964; C.D. Michener leg.; KUNHM M.G.R. Database No. 5417 • 1 ♀; 18 mi SW of Apache, Cochise County; 18 Aug. 1994; J.G. Rozen and J.S. Ascher leg.; AMNH M.G.R. Database No. 3711 • 5 ♀♀; 18 mi SW of Apache, Cochise County; 18 Aug. 1994; J.G. Rozen and J.S. Ascher leg.; AMNH M.G.R. Database Nos 1730 to 1734 • 6 ♀♀; 18 mi SW of Apache, Cochise County; 15 Aug. 1994; J.G. Rozen and J.S. Ascher leg.; AMNH M.G.R. Database Nos 1738, 1740 to 1744 • 1 ♂; 4 mi E of Willcox, Cochise County; 32.2386° N, 109.7708° W; 18–25 Aug. 2002; RSKM RSKM _ ENT_E-168251 • 1 ♀; 4 mi S of Apache, Cochise County; 18 Aug. 1982; R.M. Bohart leg.; UCBME M.G.R. Database No. 11615 • 1 ♂; 4 mi SW of Apache, Cochise County; 12 Aug. 2003; J.G. and B.L. Rozen leg.; AMNH M.G.R. Database No. 3289 • 1 ♀; 5 mi E of Sahuarita, Pima County; 15–16 Aug. 1989; J.G. Rozen and R.L. Foster leg.; AMNH M.G.R. Database No. 1735 • 1 ♂; 5 mi N of Douglas, Cochise County; 17 Aug. 1982; R.M. Bohart leg.; UCBME M.G.R. Database No. 5425 • 1 ♀; 8 mi NE of Portal, Cochise County; 23–25 Aug. 1989; J.G. Rozen and R.L. Foster leg.; AMNH M.G.R. Database No. 3712 • 1 ♂; Cochise County; 8 Aug. 1980; J.H. Cane leg.; KUNHM M.G.R. Database No. 5418 • 2 ♀♀; Continental; 2–4 Aug. 1959; K.V. Krombein leg.; USNM M.G.R. Database Nos 6035, 6036 • 1 ♀; Hereford, Cochise County; W.M. Mann leg.; USNM M.G.R. Database No. 6037 • 1 ♂; Hwy 80 & State Line Rd, Cochise County; 31.8005° N, 109.0527° W; 18–25 Aug. 2002; RSKM RSKM _ENT_E-168244 • 1 ♂; Madera Canyon (Santa Rita Mountains); 9 Aug. 1947; L. Martin leg.; LACM M.G.R. Database No. 3385 • 1 ♂; Paradise, Cochise County; 31.9333° N, 109.2167° W; 11–16 Aug. 2008; W. Steiner and J.M. Swearingen leg.; BIML USGS-DRO 147147 • 1 ♂; Portal (Road site 2), Cochise County; 26 Aug. 2007; J. Gibbs leg.; WRME JBWM0380539 • 11 ♀♀; Silver Bell Bajada (International Biological Program Desert Scrub Site), Pima County; J.L. Neff leg.; LACM 97646 /M.G.R. Database No. 3370, 97638 /M.G.R. Database No. 3371, 96503 /M.G.R. Database No. 3372, 96507 /M.G.R. Database No. 3373, 96506 /M.G.R. Database No. 3374, 97644 /M.G.R. Database No. 3375, 96505 /M.G.R. Database No. 3376, 97645 /M.G.R. Database No. 3377, 96491 /M.G.R. Database No. 3378, 97639 /M.G.R. Database No. 3379, 96504 /M.G.R. Database No. 3380 • 1 ♀; Silver Bell Bajada (International Biological Program Desert Scrub Site), Pima County; 31 Jul. 1974; J.L. Neff leg.; LACM 69441 /M.G.R. Database No. 3369 • 1 ♀; Southwestern Research Station (5 mi W of Portal), Cochise County; 4 Aug. 1956; E. Ordway leg.; AMNH M.G.R. Database No. 2451 • 1 ♀; same data as for preceding; 8 Aug. 1956; C. and M. Cazier leg.; AMNH M.G.R. Database No. 2537 • 1 ♂; same data as for preceding; 3 Aug. 1966; D.R. Miller leg.; UCBME M.G.R. Database No. 5430 • 1 ♂; same data as for preceding; 22 Aug. 2007; H.T. Ngo and J. Gibbs leg.; PCYU • 1 ♀; Southwestern Research Station (5 mi W of Portal), Cochise County; 31.8830° N, 109.2070° W; 23 Aug. 2019; J. Gibbs leg.; WRME WRME0500912 • 1 ♂; Tucson, Pima County; 28 Aug. 1951; C.D. MacNeill leg.; UCBME M.G.R. Database No. 5424 • 2 ♂♂; W Tucson-Ajo Hwy (22 mi W of Three Points), Pima County; 6 Aug. 1992; L. Packer leg.; PCYU. – New Mexico • 1 ♂; 1 mi N of Rodeo, Hidalgo County; 13 Aug. 1974; J.G. and K.C. Rozen leg.; AMNH M.G.R. Database No. 3291 • 1 ♀; 3 mi NE of Lordsburg, Hidalgo County; 21 Aug. 1971; E.E. Grissell and R.F. Denno leg.; UCBME M.G.R. Database No. 11616 • 1 ♀; 6 mi N of Rodeo, Hidalgo County; 26 Aug. 2011; WRME JBWM0380529 • 2 ♂♂; 7.5 mi W of Hachita, Grant County; 17 Aug. 1967; R.R. Snelling leg.; LACM M.G.R. Database Nos 3384, 3386 • 1 ♂; 20 mi SW of Alamogordo, Otero County; 14 Aug. 1971; M. and T.M. Favreau leg.; AMNH M.G.R. Database No. 3727 • 1 ♀; Animas, Hidalgo County; 26 Aug. 1975; R.J. McGinley leg.; USNM M.G.R. Database No. 6039 • 1 ♀; ca 4 mi N of Rodeo, Hidalgo County; 31.8803° N, 109.0356° W; 15 Aug. 2000; R. Brooks leg.; KUNHM SM0196592 / ARIZ 1B00001/M.G.R. Database No. 1323 • 1 ♀; Community Road (NE of Portal), Hidalgo County; 23 Aug. 2006; A.I. Gravel leg.; PCYU • 1 ♀; Granite Pass, Hidalgo County; 25 Aug. 1958; P.D. Hurd leg.; UCBME M.G.R. Database No. 11618 • 1 ♂; Hwy 80, Hidalgo County; 32.0935° N, 108.9684° W; 15 Aug. 2007; J. Gibbs and C.S. Sheffield leg.; PCYU PYU-3537 • 1 ♀; S of Animas, Hidalgo County; 2003; L. Packer leg.; PCYU. – Texas • 1 ♂; La Puerta Tract, Lower Rio Grande Valley National Wildlife Refuge, Starr County; 26.3450° N, 98.7317° W; 4 May 2017; M. Buck leg.; RAM pmae00141635.
Non-preserved material
MEXICO • 1 ♀; Nuevo León, Lampazos de Naranjo; 4 Oct. 2021; R. Castillo obs.; iNaturalist record #97206163.
USA – Arizona • 1 ♀; Cochise County; Aug. 2023; “ samwilhelm ” obs.; iNaturalist record #181775336 • 1 ♂; same data as for preceding; 20 Aug. 2023; K. Parys obs.; iNaturalist record #179819569 • 1 ♂; Southwestern Research Station (5 mi W of Portal), Cochise County; 31.8836° N, 109.2068° W; 27 Aug. 2019; J. Gibbs obs.; iNaturalist record #31641363. – New Mexico • 1 ♀; S end of Sonoma Ranch Boulevard, Las Cruces, Doña Ana County; 17 Aug. 2020; P. De Ley obs.; BugGuide record # 1885693 / 1885695 / 1885697 / 1885698.
Redescription
MEASUREMENTS OF HOLOTYPE. Body length 12.2 mm; ITW 2.4 mm; head length 3.0 mm; head width 4.0 mm; fore wing length 10.1 mm.
Both sexes
INTEGUMENT COLORATION. Dark brown to black except as follows. Mandible with apical half golden yellow and middle quarter in basal half reddish orange. F1 extensively, coxae to some extent, trochanters to tarsi (excluding brown meso- and metatibial spurs) partially to entirely, metasomal terga laterally, pygidial plate to some extent, and metasomal sterna to some extent orange. F2 with orange spot basally. Tegula reddish brown (dark brown in some specimens). Fore wing membrane dusky subhyaline throughout. Hind wing membrane dusky subhyaline to hyaline.
PUBESCENCE. Face with tomentum densest around antennal socket. Tomentum slightly sparser on clypeus; upper paraocular and frontal areas and vertexal area mostly exposed. Pronotal collar with tomentum uniformly pale yellow. Mesoscutum with well-defined paramedian band of pale-yellow tomentum (tapering slightly toward but not attaining anterior margin in holotype; attaining anterior margin in allotype); pale tomentum otherwise mostly restricted to lateral and posterior margins. Axilla with off-white to pale-yellow, appressed, branched setae all along its margins. Mesopleuron with off-white to pale-yellow, appressed, branched setae in upper half and patches of off-white and brown, appressed, branched setae ventrally; upper half densely setose, except behind pronotal lobe, with setae slightly sparser on hypoepimeral area; ventrolateral half nearly bare, except along margins. Metanotum with tomentum rubbed off medially in holotype, but uninterrupted and uniformly off-white in allotype and other specimens. Propodeal triangle mostly glabrous, with (pale) setae restricted to small lateral patches. T1 with basal and apical transverse bands of pale-yellow tomentum interrupted medially and subparallel, discal patch trapezoidal (crescent-shaped in some specimens). T2–T4 with complete apical transverse bands of pale-yellow tomentum, those of T2–T3 medially somewhat removed from apical margins of terga, that of T2 with pair of basomedially convergent anterolateral extensions. S2–S3 with apical transverse bands of white tomentum.
SURFACE SCULPTURE. Labrum coarsely and densely (most i <1d) rugose-punctate. Clypeus densely punctate (most i <1d) but interspaces well defined, shining. Vertexal area somewhat sparsely punctate (some i>2d), especially around ocelli. Mesoscutum and axilla with punctures more or less equally dense (most i ≤ 1d); interspaces well defined, shining. Mesoscutellum with punctures nearly contiguous (most i <1d). Mesopleuron with punctures in upper half nearly contiguous and denser (i <1d) than in ventrolateral half (i≤ 2d); interspaces shining where punctures not contiguous; punctures similar in size throughout. Discs of metasomal terga with punctures very fine, dense (i≈ 1d), and evenly distributed; interspaces somewhat dull due to tessellate surface microsculpture.
STRUCTURE. Labrum with pair of small subapical denticles, each preceded by discrete longitudinal ridge. Pronotal collar somewhat elongate (medial length ~ ¾ MOD). Mesoscutellum moderately bigibbous. Axilla extending well beyond midlength of mesoscutellum but not as far back as its posterior margin; tip distinctly pointed and hooked (i.e., concave along medial margin), mesally unattached to mesoscutellum for approximately half medial length of axilla; lateral margin somewhat arcuate. Fore wing with three submarginal cells.
Female Scape and pedicel extensively orange. T5 with concave apical margin and large patch of off-white to pale-yellow tomentum on each side lateral to pseudopygidial area. Pseudopygidial area trapezoidal, with posteromedially directed coppery brown setae on each side of glabrous midline and apical row of posteriorly directed, dense, suberect golden to silvery setae. Pygidial plate apically truncate. S4 with apical transverse band of white tomentum. S5 straight in lateral view, covered in mostly brown tomentum and with apical fimbria of coppery bristle-like setae.
Male
Scape and pedicel to some extent orange. T5 with complete apical transverse band of pale-yellow tomentum. T6 with complete apical transverse band or small posteromedial patch of off-white to pale-yellow tomentum. Pygidial plate apically rounded and slightly downturned, with lateral margin somewhat sinuate. S4–S5 each with apical/subapical fringe of dense, long (> 1 MOD), curved, coppery to silvery setae, not contrasting strongly with bands of preceding sterna.
Distribution
Arizona to Texas in the United States to south-central Mexico (Fig. 12E).
Ecology
Host records
Rozen (1966) described an unknown Triepeolus postdefecating larva taken from the nest of Protoxaea gloriosa (Fox, 1893). In the original description of T. kathrynae, Rozen (1989) identified the larva as this species based on the collection of an adult female (paratype, USNM 0930-0959/M.G.R. Database No. 5716) from the same locality and year investigating the entrance of a Protoxaea burrow.
Floral records
Rozen (1989) and labels of examined voucher specimens indicate that this species has been collected from the following flowering plant species: in Apocynaceae Juss., Asclepias subverticillata (A.Gray) Vail; in Asteraceae, Baccharis sp., Engelmannia peristenia (Raf.) Goodman & C.A.Lawson, Gaillardia pulchella Foug., Picradeniopsis absinthifolia, and Verbesina encelioides (Cav.) A.Gray; in Fabaceae, Eysenhardtia sp., Melilotus albus, and Psorothamnus scoparius Rydb.; in Malvaceae Juss., Sphaeralcea sp. or spp.; in Polygonaceae Juss., Eriogonum sp. and Eriogonum deflexum Torr.; and in Zygophyllaceae R.Br., Kallstroemia sp. or spp., K. grandiflora Torr. ex A.Gray, and Larrea sp.
Remarks
Based on known records, adults of T. kathrynae are active from May to October.
description: Published as part of Onuferko, Thomas M. & Rightmyer, Molly G., 2024, A revision of the simplex species group of the cleptoparasitic bee genus Triepeolus Robertson, 1901 (Hymenoptera: Apidae), pp. 1-106 in European Journal of Taxonomy 950 (1) on pages 34-40, DOI: 10.5852/ejt.2024.950.2643, http://zenodo.org/record/13691805