Sohmaea teres H. Ohashi & K. Ohashi 2018
Creators: name: Saisorn, Witsanu
name: Chantaranothai, Pranom
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Magnoliopsida
Fabales
Fabaceae
Sohmaea
Sohmaea teres

description: 4. Sohmaea teres (Wallich 1831: 196 ex Bentham 1852: 225) H.Ohashi & K.Ohashi (2018b: 163). Fig. 2C–D.   
 ≡ Desmodium teres Wallich ex Bentham Type:— MYANMAR. Taong Dong, Wallich 5694 (lectotype K-W! K001121790, designated by Rafidah et al. (2020), isolectotypes BM! BM000946774, CAL, n.v., G!, K! K000858870, K-W! K001121791).   
 ≡ Meibomia teres (Wall. ex Benth.) Kuntze (1891: 198).   
 = Desmodium zonatum Meeuwen (1961: 97), non Miq. (1855: 250). Specimen cited:— Dorgeh 1755 (L, n.v.), fide Ohashi (2004).   
 Herb, up to 2 m high; stems and twigs terete, sparsely to densely uncinate hairy. Leaves unifoliolate, alternately arranged; stipules narrowly triangular, 5–12 × 2–3 mm, apex long acuminate to caudate, surface glabrescent; petioles 0.3–0.8 cm long, ascending pubescent and minutely uncinate. Leaflets: stipels narrowly triangular, 2–5 × 0.5–1 mm, apex long acuminate, surface glabrescent, margin not fimbriate; petiolules 1.5–3 mm, ascending pubescent and uncinate hairy; leaf blade lanceolate to narrowly lanceolate or oblong, (3–)9–20 × (0.5–) 2.5–7 cm, apex attenuate, base obtuse, margin entire, upper surface nearly glabrous or sparsely uncinate hairy, with straight hairs on mid- and lateral veins, lower surface spreading uncinate hairy, with straight hairs on mid- and lateral veins; lateral veins 9–14 per side. Inflorescences 10–40 cm long, terminal or axillary; rachis densely uncinate hairy. Primary bract narrowly ovate, 3.5–4 × 1–1.5 mm, apex long acuminate to attenuate, surface pubescent, conspicuously veined, enclosing 5-immature flowers and secondary bract. Secondary bract narrowly ovate to narrowly triangular, 1.5–2 × 0.5 mm, pubescent. Flowers 6–7 mm long, 5-flowered fascicles; bracteoles absent; pedicels 4–5 mm long, densely uncinate and straight hairy (on lower half part). Calyx yellowish green, ca. 4 mm long, base cuneate; outside ascending pubescent and minutely uncinate, inside glabrescent, tube ca. 1.5 mm long; teeth 2–3 mm long, acuminate to needle-like at the apex, prominently longer than tube length, two lateral teeth shorter than others, lower tooth longest, upper one entire. Corolla standard white, wings and keels violet to blue; standard broadly obovate to oblanceolate, 4.5–8 × 3.5–5.2 mm, apex obtuse to rounded, base attenuate, not auriculate, claw 0.3–1 mm long; wings slightly oblong, 3.5–6 × 1–2 mm, apex obtuse, base auriculate, ca. 0.3 × 0.5 mm, claw 0.5–1 mm long; keels slightly curved, 4–7 × 1–1.5 mm, prominently longer than wings, apex acute to slightly obtuse, base shortly auriculate, claw ca. 2.5 mm long. Stamens 4–7.5 mm long; anthers ellipsoid, ca. 0.3 × 0.2 mm. Gynoecium 4.5–9 mm long; ovary oblong, laterally compressed, pubescent, stipitate, 0.5–1 mm long, 10–13-ovulate; style 1.5–3 mm long, pubescent; stigma minutely capitate. Pods light brown, indehiscent, 8–11-articulate, subterete, 4–6.5 cm long, 1–1.5 mm wide, surface densely uncinate, not reticulate-patterned (but longitudinal striate), both sutures less constricted between articles and thickened; isthmus ca. 4/5 as broad as the pod; articles narrowly oblong to linear, 5–6 mm long; pod stipe 1.5–2 mm long, glabrescent; fruiting pedicels 5–7 mm long, ascending pubescent and uncinate hairy. Seeds brown, transversely oblong, 4.5–5 × 1 mm, ca. 0.7 mm thick.   
 Distribution:— Myanmar, Thailand, Malaysia, and Indonesia.   
 Ecology:—In sunny places of limestone mountain and evergreen, dry evergreen, and mixed deciduous forests; 50–700 m elev.   
 Phenology:—Flowering in September–November. Fruiting in October–January.   
 Notes:— Sohmaea teres is a new record species for Cambodia.   
 Specimens examined:— THAILAND. Chiang Mai: Mae Rim, QBG, BGO. Staff 4828 (QBG-2 sheets), Doi Chiang Dao, 480 m, 21 Oct. 1955, Garrett 1460 (K, L, P-2 sheets), Doi Chiang Dao, 600–1300 m, 25 Sept. 1971, Murata et al. T-14911 (KYO), Doi Chiang Dao, 600–1300 m, 25 Sept. 1971, Murata et al. T-14936 (KYO), Chiang Dao, Ban Bing Kong, 500 m, 29 Oct. 1987, Maxwell 87-1304 (CMU, L), Doi Suthep, 630 m, 17 Oct. 1999, Srapratet 24 (CMUB), Fang, Doi Pha Hom Pok National Park, 21 Dec. 2014, Saisorn 352 (KKU), Mae Rim, QSBG, 860 m, 13 Nov. 2014, Tanming 732 (QBG) & Chiang Dao, Tham Chiang Dao, 350 m, 15 Feb. 1958, SØrensen et al. 1137 (C); Chiang Rai: Doi Pha Hom Pok, 700 m, 15 Sept. 1967, Iwatsuki et al. T-9724 (KYO) & Pan, Doi Luang National Park, 610 m, 3 Nov. 2015, Muangyen 214 (QBG); Phayao: Mae Chai, Doi Luang National Park, 800 m, 4 Nov. 2015, Muangyen 318 (QBG), Doi Luang National Park, 600 m, 4 Nov. 2015, Muangyen 327 (QBG) & Mueang Phayao, Doi Luang National Park, 575–625 m, 25 Sept. 1997, Petrmitr 123 (BKF, CMUB); Nan: Tham Pha Tok, 250 m, 16 Nov. 1993, Larsen et al. 44405 (AAU); Lampang: Chae Hom, 325 m, 26 Oct. 2012, Maxwell 12-279 (CMUB, QBG), en route from Pang La to Huai Tak, 350 m, 24 Sept. 1967, Shimizu et al. T-10752 (BKF, KYO) & without locality, 250 m, 22 Nov. 1922, Winit 811 (ABD, BK, K); Sukhothai: Si Satchanalai, Ban Mae San, 17 Nov. 2015, Maknoi 8391 (QBG) & Ban Mae San, 28 Jan. 2016, Maknoi 8514 (QBG); Nakhon Sawan: Ban Na Kluay, 50 m, 7 Nov. 1924, Kerr 4537 (ABD, BK, BM, E, K); Phetchabun: Nam Nao National Park, 3 Nov. 2013, Mattapha 448 (KKU) & Nam Nao National Park, 18 Oct. 2004, Mattapha 696 (KKU-2 sheets); Loei: Pha Nok Khao, 250–350 m, 6 Dec. 1970, Charoenphol et al. 4564 (AAU, BKF, C) & Wat Tham Pha Sawan, 381 m, 14 Nov. 2011, Clark et al. 240 (K, QBG); Khon Kaen: Phu Pha Man National Park, 25 Oct. 2007, Norsaengsri & Lakoet 2807 (QBG), Phu Pha Man National Park, 410.6 m, 22 Nov. 2007, Norsaengsri & Lakoet 3069 (QBG), Phu Pha Man National Park, 467 m, 24 Dec. 2011, Norsaengsri & Tathana 8403 (QBG), Wiang Kao, Phu Wiang National Park, 11 Nov. 2014, Saisorn et al. 294 (KKU), Wiang Kao, Phu Wiang National Park, 15 Feb. 2015, Saisorn 376 (KKU), Chum Phae, Dong Lan, 300 m, 22 Nov. 1958, Smitinand 4919 (BKF, L) & Chum Phae, Dong Lan, 400 m, 22 Dec. 1958, SØrensen et al. 6122 (C); Nakhon Ratchasima: Pak Chong, 600–650 m, 20 Nov. 1982, Koyama et al. T-30147 (KYO), Pak Chong, 700–730 m, Koyama et al. T-30094 (KYO) & without locality, 18 Dec. 1962, Phengklai 584 (C, K, L); Kanchanaburi: Wang Po, 26 Oct. 1969, Kasem 648 (BK, L, SING), Srisawat, 580–600 m, 26 Nov. 1982, Koyama et al. T-30304 (KYO), Sai Yok, Mahidol University Kanchanaburi Campus, 225 m, 20 Dec. 2005, Maxwell 05-739 (CMUB), Si Sawat, Ban Khao Lek, 500–600 m, 21 Oct. 1994, Nisit 94102118 (BK), Si Sawat, Ban Khao Kaeng Khaep, 500 m, 2 Nov. 1979, Shimizu et al. T-21399 (BKF, KYO, L), Si Sawat, Erawan National Park, 100–300 m, 3 Nov. 1979, Shimizu et al. T-21540 (BKF, KYO), Si Sawat, Erawan National Park, 100–300 m, 3 Nov. 1979, Shimizu et al. T-21567 (KYO), Sai Yok, 160 m, 6 Nov. 1979, Shimizu et al. T-21760 (BKF, KYO), Sai Yok, Near Sai Yok Arboretum, 160 m, 6 Nov. 1979, Shimizu et al. T-21829 (KYO) & Si Sawat, 150 m, 5 Nov. 1971, van Beusekom et al. 3413 (BKF, C, K, KYO, L-2sheets, P); Prachuap Khiri Khan: Kui Buri, Sam Roi Yot, 23 Oct. 1973, Sutheesorn 2703 (BK); Lop Buri: Chai Badan, 280 m, 4 Dec. 1982, Koyama & Terao T-30595 (BKF, KYO) & Subchampa Hill, 170–190 m, 19 Nov. 1984, Murata et al. T-50933 (BKF, KYO); Saraburi: Muek Lek, 200 m, 10 Nov. 1924, Marcan 1867 (ABD, BK, BM, KKU); Chon Buri: Si Racha, Khao Khiao, 300 m, 18 Dec. 1974, Maxwell 74-1077 (BK) & Si Racha, Khao Khiao, 100 m, 20 Oct. 1975, Maxwell 75-1052 (AAU, BK, L); Chanthaburi: Pong Nam Ron, 700 m, 11 Oct. 1955, Sangkhachand 528 (BKF, L), Pong Nam Ron, Ban Thalee, 200 m, 24 Oct. 1956, Smitinand 3616 (BKF) & without locality, 200 m, 28 Nov. 1956, Smitinand 3648 (BKF, L); LAOS. Luang Prabang: Massie s.n (Dy Phone et al., 1994); CAMBODIA . Mondulkiri: Sen Monorom, 527 m, 5 Nov. 2016, Yahara et al. 7382 (FU); VIETNAM. Annam: Ca-na, Phanrang, 500 m, 22 Nov. 1923, Poilane 8652 (P-2 sheets).
description: Published as part of Saisorn, Witsanu & Chantaranothai, Pranom, 2022, A taxonomic revision of two genera, Pleurolobus and Sohmaea (Leguminosae) in Thailand and Indo-China, pp. 231-246 in Phytotaxa 573 (2) on pages 241-242, DOI: 10.11646/phytotaxa.573.2.4, http://zenodo.org/record/7349951