Anomala divisa Filippini, Galante, and Mico, new species
Creators: name: Filippini, V.
name: Galante, E.
name: Micó, E.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Rutelidae
Anomala
Anomala divisa

description: Anomala divisa Filippini, Galante, and Micó, new species   
(Figs. 3, 11, 19, 27, 36)  
 Type Material. Holotype: ♂ " Alajuela. Costa Rica Cinco Esquinas de Carrizal, 1800m 16 abril de 1988 Col: G. Barrantes / INBIOCRI002517128" deposited at MNCR. Paratypes (6): 2♂ " Zarcero, Alfaro Ruiz, 1700m Alajuela Pr. COSTA RICA. Apr - May 1989. A. Del Valle & A. Solis, L- N 240500, 493500" / INBIOCRI000022463 and CEUA00106199; 3♂ 1♀ " Est. Biol. Las Alturas 1500 m, Coto Brus, Prov. Puntarenas, Costa Rica, F. Araya, 23 mar a 2 may 1992, L- S 322500,591300" / CEUA00106198, INBIOCRI000909434, INBIO CRI000916198 and INBIOCRI000909406. Four paratypes are deposited at MNCR, and two paratypes are deposited at Universidad de Alicante.   
 Holotype. Male (Fig. 3). Body oval. Length 12.87 mm, width 6.47 mm. Head reddish brown with ochre apical fringe and clypeus. Pronotum ochre with 2 large, trapezoidal, reddish brown maculae with bronze luster touching anterior and posterior margins. Scutellum ochre with thin, reddish sides. Elytra ochre with reddish suture. Pygidium ochre. Legs ochre with light to reddish brown apical part of tibia and tarsus. Venter reddish to dark brown, sternum with sides ochre. Head: Clypeus trapezoidal, anterior angles widely curved, surface densely punctate-reticulate, convex; anterior margin straight, thin in frontal view, ratio width/length 1.99. Frons densely punctate, flat. Ocular canthus long, thin, with acute apex. Ratio of interocular width/width of eye 3.52. Ratio antennal funiculus/club 0.79. Pronotum: Subtrapezoidal. Ratio width/length 1.81. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles quadrate, posterior angles quadrate. Basal margin sinuate, subapical bead complete. Surface with deep, dense punctures. Scutellum: Subtriangular, with rounded sides and acute apex. Ratio width/length 1.56. Surface sparsely punctate. Elytra: Costae defined by regular rows of punctures. Subsutural interstice with 2–4 rows of sparse punctures, 2 nd –4 th interstices with 1–2 irregular, secondary stria(e). Surface covered with secondary punctures. Pygidium: Finely granulated with long setae on posterior margin. Ratio width/length 1.57. Venter: Space between mesocoxae narrow, slightly convex. Mesometasternal suture well-defined at base of mesocoxae. Abdominal sternites with 1 row of setae (2 rows on second sternite) and 5–7 rows of punctures per sternite. Last sternite strigate, with subapical bead thin and sinuate, apical membrane wide. Legs: Protibia (Fig. 11) bidentate, apical tooth long and curved, basal tooth below internal apex of protibia triangular, acute. Metatibia slender, fusiform, length/width ratio 3.39. First external carina with a row of sparse setae, surface punctate. Internal protarsal claw bifurcate, with upper branch of same length and half width of the lower one; inferior margin with a sharp angle at half length. Male genitalia: Parameres with blunt apex in lateral view (Fig. 19), ventral angle defined. Endophallus (Fig. 27) with 1 ventral and 1 dorsal inflation. On dorsal inflation, a long curved, claw and a patch of sclerotized, thick setae; ejaculatory duct opening located on ventral inflation. Median lobes very thin and long, a dorsally located sclerotized plate between their apices.   
 Female. Similar to male except protibia (Fig. 11) with wider teeth; internal protarsal claw narrower with curved inferior margin; last sternite with curved apex.   
 Variation. Six paratypes. Maculae on pronotum can fuse into one large macula with a bronze or green luster. Venter of female specimen is ochre. Claw on endophallus can have a bifurcated tip. Body length 12.04–13.56 mm, body width 6.47–7.18 mm. Clypeal width/length ratio 1.85–2.01. Ratio of interocular width/width of eye 3.42–3.93. Ratio of antennal funiculus/club 0.70–0.79. Pronotal width/length ratio 1.79–1.87. Scutellar width/length ratio 1.22–1.56. Pygidial width/length ratio 1.57–1.66. Metatibial length/ width ratio 3.16–3.61.   
 Etymology. From the past participle of the Latin verb divido, to divide. The ochre median line on the pronotum and dark suture on the elytra visually divide in half the dorsal aspect of this species.   
 Distribution. Anomala divisa occurs in the Central and Talamanca mountain ranges of Costa Rica, from 1,500 to 1,800 m elevation (Fig. 36).   
 Diagnosis. This species is recognized by the presence of two large, trapezoidal, dark maculae on the pronotum and the parameres with blunt apex and defined ventral angle. Similar species, also present in Costa Rica, are Anomala jansoni Ohaus, 1897 and Anomala quiche Ohaus, 1897. The former species is slightly larger, with smaller maculae on the pronotum and long, slender, curved parameres in lateral view. The latter species has smaller and more separated maculae on the pronotum, maculae on the elytral calli, and parameres with a wider and blunter apex (aedeagi illustrated in Filippini et al. 2015a).
description: Published as part of Filippini, V., Galante, E. & Micó, E., 2015, Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica, pp. 463-476 in The Coleopterists Bulletin 69 (3) on pages 470-471, DOI: 10.1649/0010-065x-69.3.463, http://zenodo.org/record/10106050