Chandleromyia anomala Grimaldi 2018, new species
Creators: name: Grimaldi, David A.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Platypezidae
Chandleromyia
Chandleromyia anomala

description: Chandleromyia anomala, new species
Figures 8, 33
DIAGNOSIS: As for genus, by monotypy.
DESCRIPTION: (male only). Head: Large, broader than thorax. Eyes bare, large, holoptic for ~30 facets (virtually entire length of frons except short segment above antennae). Dorsal facets (~ 70% of eye) large, 2× diameter of ventral facets. Frontal margins of eyes emarginate around bases of antennae, margins slightly divergent ventral to this. Median furrow of frons, interfrontal, fronto-orbital setae (presence/ absence) not observable. Face entirely bare. Ocelli on tubercle, with pair of short posterior setae (length shorter than diameter of ocellus). Vertical setae absent; postoculars minute, except for short row of 4–5 short, thick seate on dorsal portion. Occiput with fine, short, scattered, light setae (lengths ca. equal to diameter of large eye facets); occipital setae do not extend to gena. Vibrissa, genal setae absent. Antenna: Scape small, barely exposed, apparently bare; pedicel small, conical, with apical circlet of setulae; arista terminal; basal flagellomere small, tapered, with incised portion mesally; two small basal aristomeres (lengths ~3× the diameter); apical aristomere long, fine, bare of any micropubescence.
Mouthparts: Oral margin squared; clypeus a broad triangle; labrum broad, apex pointed and sclerotized; labellar lobes deeply separated anteriorly, ~0.65× lobe length; palp very small.
Thorax: Scutum convexed but not arched, flatter anteriorly. Acrostichals completely absent, even minute ones. Only 1 pair of dorsocentrals present, short, prescutellar, no smaller, graded dorsocentrals present. Supraalar setae absent; 4–5 stout notopleural setae present (4 on one side of specimen, 5 on other side); anepisternum and katepisternum bare. One pair of short, stiff postalar setae present. Prescutellum absent. Scutellum with one pair of apical setae only; setae short, stout.
Wing: Length/width 2.4; hyaline, except for faint tint between Sc and R 1. Membrane microtrichia well developed over entire surface of wing, even in cell Sc. Two prehumeral costal setae at base of C; C with spinules, end near apex of R 2+3. Sc relatively short, apex slightly beyond level of crossvein r-m; R 1 straight, bare, length 0.72× the wing length; bases of Sc and R 1 close, divergent for apical half. Pterostigma absent. Vein R 4+5 ends at wing tip; cells rm and dm closed; crossvein br-m slightly basal to level of Sc apex; crossvein dm-cu close to wing margin, separated from margin 0.75× its length. M not forked (M 2 lost); M 1 bent in middle, deflected anteriad. Tip of CuA 1 incomplete; cell cup longer than vein at apex, cell not extremely acute or narrow; CuP slightly curved, not straight; CuP+A 1 tip incomplete. Legs: All femora without macrosetae, spines or tubercles. Dark bifid scales on all tibiae and tarsi: scattered on partial, lateral surface of tibiae, encircling ventral portion of tibial apex; in dense lateral row on tarsi. Metatarsomeres 1–4 flattened laterally, expanded; tarsomere 2 with blunt spine on lateroapical corner; tarsomere 3 with small, light, clavate seta preapically near row of bifid scales. Mesotibia with 2 ventroapical spurs, large one mesally (its length ca. diameter of tibia). Pulvilli well developed; empodium not observable (probably minute, setiform).
Abdomen: Tergites without macrosetae or setulae; 8 tergites visible; anterior 6 broad (1–4 broadest); tergite 7 narrow, longer than wide, tubular, ventroflexed (presumably circumverted); epandrium small, capsular. Male genitalia not observable (preservation insufficient). Sternites well developed, but lateral margins do not meet tergites (pleural membrane exposed).
SPECIMENS EXAMINED: Holotype, male, AMNH Bu-KL30-27, in Burmese amber. The fly is extremely well preserved, complete, though some details are obscured (e.g., small structures of the genitalia).
ETYMOLOGY: In reference to the anomalously early appearance of a platypezine in the mid-Cretaceous.
description: Published as part of Grimaldi, David A., 2018, Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi, pp. 1-97 in Bulletin of the American Museum of Natural History 2018 (423) on pages 1-97, DOI: 10.1206/0003-0090-423.1.1, http://zenodo.org/record/4613008