Leucogeorgia lobata Antić & Reip 2020, sp. nov.
Creators: name: Antić, Dragan Ž.
name: Reip, Hans S.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Julida
Julidae
Leucogeorgia
Leucogeorgia lobata

description: Leucogeorgia lobata sp. nov.
urn:lsid:zoobank.org:act: A52F58C6-7035-48FA-924D-6F237089CE7C
Figs 1D, 2A, 25–26, 27 A–C, 28, 56, 58
Diagnosis
This species belongs to the group of Leucogeorgia spp. with neither modified mouthparts nor teeth on the mesomeral claw (vs teeth present in L. abchasica, L. borealis sp. nov., L. gioi sp. nov., L. oculata sp. nov. and L. satunini). Leucogeorgia lobata sp. nov. differs from L. golovatchi sp. nov. and L. prometheus sp. nov. by having a strongly developed lobe on the mesomeral lamella, with a deep rift between mesomeral claw and the lobe (vs absence of such a lobe or rift in L. golovatchi sp. nov. or sometimes a poorly developed lobe without a deep rift between the mesomeral claw and lobe in L. prometheus sp. nov.).
Etymology
From the Latin ʻ lobata ʼ (= ʻhaving lobesʼ, ʻlobateʼ), reflecting the presence of a well-developed lobe on the mesomeral lamella. Adjective in feminine gender.
Material examined
Holotype CENTRAL-WEST GEORGIA – Samegrelo-Zemo Svaneti • ♂; Chkhorotsku District, near Chkhorotsku, Odishi Plateau, Kalitshona (= Konglomeratnaya) Cave; 42.53° N, 42.16° E; 14 Aug. 2016; D.M. Palatov leg.; ZMUM.
Paratypes
CENTRAL-WEST GEORGIA – Samegrelo-Zemo Svaneti • 1 ♀, 1 juv.; same collection data as for holotype; ZMUM • 3 ♂♂, 2 ♀♀; same collection data as for holotype but 29 Oct. 1991; V. Kiselev leg.; ZMUM • 2 ♂♂, 4 ♀♀, 5 juvs; same collection data as for holotype but 3 Feb. 2017; D.M. Palatov leg.; ZMUM • 1 ♂; same collection data as for holotype; IZB • 1 ♂; same collection data as for holotype; SMNG; • 1 ♂; village Garakha, Odishi Plateau, Garakha Cave; 42.53° N, 42.17° E; 30 Apr. 2017; G. Nebieridze and S. Barjadze leg.; IZISU • 1 ♂; same collection data as for preceding but 2 Feb. 2017; D.M. Palatov leg.; ZMUM.
Other material
CENTRAL-WEST GEORGIA – Samegrelo-Zemo Svaneti • 1 juv.; Martvili District, Askhi karst Massif, near Balda village, Motena Cave; 42.47° N, 42.39° E; 19 Oct. 1978; R.A. Djanashvili leg.; ZMUM • 1 ♀; same collection data as for preceding except 12 Aug. 2016; ZMUM • 4 ♂♂, 5 ♀♀; same collection data as for preceding except 30 Jan. 2017; D.M. Palatov leg.; ZMUM • 3 ♂♂; same collection data as for preceding except 13 Jun. 2019; H. Reip leg.; SMNG • 2 ♀♀; Askhi karst Massif, Martvili village, Jortsku Cave; 42.51° N, 42.42° E; 2 Feb. 2017; D.M. Palatov and A.M. Sokolova leg.; ZMUM.
Description
SIZE AND NUMBER OF BODY RINGS. Holotype male 20 mm long, vertical diameter of largest body ring 1.5 mm, body with 37 podous rings + 1 apodous ring + telson. Paratype males 14.5–22 mm long, vertical diameter of largest body ring 1.1–1.6 mm, body with 32–41 podous rings + 1–2 apodous rings + telson. Paratype females 20–22 mm long, vertical diameter of largest body ring 1.5–1.7 mm, body with 37–40 podous rings + 0–2 apodous rings + telson. Additional males 19–33 mm, vertical diameter of largest body ring 1.4–2 mm, body with 40–46 podous rings + 1–2 apodous rings + telson. Additional females 21–34 mm long, vertical diameter of largest body ring 1.6–2.3 mm, body with 39–46 podous rings + 1–2 apodous rings + telson.
COLOUR (Figs 1D, 25). Yellowish white.
HEAD (Figs 25B, 26C). Without ommatidia. Frontal setae absent. Labrum with three labral teeth, four supralabral setae and 12–14 labral setae. Gnathochilarium with rhomboid promentum; lamellae linguales with 4+4–5+5 setae in one row, stipites each with 3 distolateral setae and 6–7 medial setae. Antennae 2.3 mm long in holotype male, their length ca 150% of vertical diameter of largest body ring. Lengths of antennomeres I–VIII (in mm): 0.12 (I), 0.45 (II), 0.46 (III), 0.4 (IV), 0.46 (V), 0.28 (VI), 0.11 (VII) and 0.02 (VIII). Length/width ratio of antennomeres I–VII: 0.75 (I), 3.7 (II), 3.5 (III), 3 (IV), 2.7 (V), 1.8 (VI) and 1 (VII). Antennomeres V and VI each with a terminal corolla of large sensilla basiconica bacilliformia; antennomere VII with a terminal corolla of small sensilla basiconica bacilliformia.
BODY RINGS (Fig. 25D). Entire metazonal area with longitudinal striations. Length of midbody setae ca 7% of vertical diameter of rings.
TELSON (Fig. 25C). Epiproct with a short and blunt preanal process, covered with dorsal and lateral setae. Paraprocts rounded, with numerous setae. Hypoproct without any modifications.
LEGS IN MALES. First pair of legs modified, hook-shaped (Figs 26 A–B, 28E–F), with three podomeres; coxae each with one seta; prefemora each with 6–9 setae; femora, postfemora and tibiotarsi coalesced, with 4+4 setae (3+3 on remnants of femora and 1+1 on remnants of postfemora). Tibiotarsal part with a small distal lobe. Tip slightly tuberculated. Postfemoral and tibial ventral pads well-developed on anterior legs, then gradually disappearing towards posterior legs.
PENES (Fig. 28D). In the form of a short trapezoid, apically with two small subtriangular lobes.
VENTRAL MARGIN OF MALE BODY RING 7 (Fig. 25E). Strongly developed, more or less rounded in lateral view.
GONOPODS (Figs 2A, 26D, 27 A–C, 28A–C). Promere (p) long and slender, with a flagellum (f); apical part spatulate, with denticulated margin; basal half with two developed ridges. Mesomere (m) with a slender mesomeral claw (mc) lacking teeth, more or less curved anteriad; mesomeral lamella (ml) with a strongly developed lobe (l) and a deep rift between mesomeral claw and lobe, distal margin smooth, posterior part finely fimbriate. Opisthomere (o) bipartite. Anterior branch of o with a solenomere (s) with a short tip, and a well-developed and fimbriate velum (v). Posterior branch of o in form of a shield-like protective lamella (pl). Mesomere and opisthomere connected basally with an accessory membrane (am).
Distribution
Known only from four caves in the Samegrelo-Zemo Svaneti region in Central-West Georgia (Fig. 58, violet triangles).
Remarks
In the Motena and Jortsku caves,this species has been found together with another taxon of Leucogeorgiini, Martvilia parva gen. et sp. nov.
See also under Leucogeorgia aff. lobata sp. nov.
description: Published as part of Antić, Dragan Ž. & Reip, Hans S., 2020, The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae), pp. 1-106 in European Journal of Taxonomy 713 on pages 41-47, DOI: 10.5852/ejt.2020.713, http://zenodo.org/record/4020752