Toktokkus Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith 2021
Creators: name: Kamiński, Marcin J.
name: Gearner, Olivia M.
name: Kanda, Kojun
name: Swichtenberg, Kali
name: Purchart, Luboš
name: Smith, Aaron D.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Tenebrionidae
Toktokkus

description: KEY TO THE SPECIES OF THE GENUS TOKTOKKUS 1. Apex of elytra not sloped (dorsal view) and depressed with clear margin (Fig. 3B)............... T. congolensis - Apex of elytra sloped (dorsal view) and flat to slightly convex (e.g. Fig. 3A)...............................................2 2. Margin of prosternal collar expanded and folded out into a large lip (Fig. 3C); elytra round, almost heart- shaped (Fig. 3C); declivous portion of elytra with little to no tubercles..................................... T. tschinkeli - Margin of prosternal collar not expanded, only occasionally folded out; elytra round or elongate; declivous portion of elytra with tuberculate rows.........................................................................................................3 3. Disc of pronotum with prominent punctures (Fig. 3D); gold setae present on elytra (Fig. 3D)..... T. herero - Disc of pronotum only with micropunctures; elytra not covered with setae................................................44. Tubercles on elytral sides round/globular, vertically direct (Fig. 2B, C)......................................................5- Tubercles on elytral sides pointed, directed posteriorly (e.g. Fig. 3F)..........................................................6 5. Elytral tubercles dense (1.0–1.5 diameters apart), laterally reaching humerus; elytra round (Fig. 2B)........................................................................................................................................................... T. mulleri - Elytral tubercles sparse (2–3 diameters apart), laterally terminating prior to humerus; elytra elongate to round (Fig. 2C)................................................................................................................................ T. schultzei 6. Elytral tubercles sparse (4–6 lateral rows), deeply angled...........................................................................7- Elytral tubercles dense (more than 6 lateral rows), slightly angled (Fig. 2B–F).........................................8 7. Elytral tubercles distinct, not confluent, nearly reaching humeri (Fig. 3A)................................ T. barclayi - Elytral tubercles small and short, confluent into rows, terminating well before humeri (Fig. 2A)...................................................................................................................................................... T. mashunus 8. Microtubercles present between tuberculate rows (e.g. Fig. 3E), tuberculate rows rarely elevated on ridges...............................................................................................................................................................9- Elytral tubercles all relatively the same size, no microtubercles present between rows, tuberculate rows often elevated on ridges................................................................................................................................10 9. Body size fairly small (18.0–23.0 mm); elytral tubercles relatively dense, almost confluent; tubercles relatively short.................................................................................................................... T. tuberculipennis - Body size medium to large (28.0–32.0 mm); elytral tubercles less dense, only occasionally confluent; tubercles taller................................................................................................................................ T. makuya 10. Elytral tubercles large, often confluent (Fig. 3F); tuberculate rows extend over humeri to scutellum; humeri prominent; disk of elytra where smooth often flat (Fig. 3F)......................................... T. waclawae - Tubercles medium sized (Fig. 2E, F), not confluent; tuberculate rows end at humeri; humeri not prominent; disc of elytra where smooth generally convex.................................................................................... T. vialis
description: Published as part of Kamiński, Marcin J., Gearner, Olivia M., Kanda, Kojun, Swichtenberg, Kali, Purchart, Luboš & Smith, Aaron D., 2021, First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group, pp. 883-901 in Zoological Journal of the Linnean Society 191 on page 894