Holepyris raonis Singh & Colombo & Shreevihar & Pandey & Tribull 2021, comb. nov.
Creators: name: Singh, Sudhir
name: Colombo, Wesley D.
name: Shreevihar, Santhosh
name: Pandey, Vijay P.
name: Tribull, Carly M.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Bethylidae
Holepyris
Holepyris raonis

description: 35. Holepyris raonis (Kurian, 1955) comb. nov.    
Fig. 35, A–J  
 Apenesia raonis Kurian, 1955, 4: 100, 101, Figs: 118–123.   
 Sclerodermus raonis (Kurian, 1955), Azevedo et al. (2018): 250.   
Type locality: St. John’s College, Agra, India (♀ holotype, NFIC-FRI).  
 Diagnosis. Female. Length about 4.3 mm. General colour dark reddish-brown to black. Head as long as wide; frons coriaceus; ocelli distinct, posterior to supra-ocular line; eye glabrous; median clypeal lobe conspicuous, anterior margin rounded, as long as lateral ones; antenna with sparse pubescence, scape 2.4 × longer than pedicel, pedicel longer than first flagellomere. Pronotum coriaceous, posterior pronotal sulcus foveolate; mesonotum shorter than pronotal dorsal area; notauli absent; mesoscuto-mesoscutellar suture sulcate, trabeculate, deep. Metapectalpropodeal disc longer than wide; metapostnotal median carina present; first abdominal spiracle laterally placed. Tarsal claw bifid. Wings nearly hyaline; forewing with pterostigma light brown; hind wing with three hamuli. Metasoma polished. Male. Unknown.   
 Material examined. Holotype: [INDIA, Uttar Pradesh,] Agra, St John’s College, 23-III-1950, S.N. Rao coll.   
 Type conditions. Holotype partly dissected on slide Nº. 23/B and remaining rested on the pin. Pinned: body complete with left antenna and left forewing attached. On slides: S1 C1—one forewing and one hind wing. S1 C2—one antenna. S2 C1—one pro and one metaleg, both without coxae. S2 C2—two meso and one metaleg all with separated coxae.   
 Remarks. This species was originally allocated in Apenesia Westwood, 1874. Azevedo et al. (2018, p. 250) transferred this species to Sclerodermus. Kurian (1955) provided a detailed description of the holotype, with drawings. He illustrated the forewings with anterior margin angular, which is common for sclerodermines, and probably for this reason, Azevedo et al. (2018) removed this species from Pristocerinae, since all the females of Pristocerinae are apterous. We detected the clypeus trilobate, eyes mid-sized and glabrous, the dorsal pronotal with posterior pronotal sulcus foveolate, and with humeral angle not projected, the mesoscuto-scutellar suture sulcate, the metanotum undeveloped medially, the wings are fully developed, the forewings with three closed cells (C 2 c, R 2 c, 1Cu 2 c), and the metapostnotal median carina is long. These observations support the placement of this species in Holepyris.   
 Hosts. Unknown.   
  Genus Laelius Ashmead, 1893    
 Laelius Ashmead, 1893, 45: 50.   
 Type species: Laelius trogodermatis Ashmead, 1893 by original designation.
description: Published as part of Singh, Sudhir, Colombo, Wesley D., Shreevihar, Santhosh, Pandey, Vijay P. & Tribull, Carly M., 2021, Rediscovery of Kurian's types of Hymenoptera at Forest Research Institute, Dehradun, India with notes on Indian bethylid fauna, pp. 1-90 in Zootaxa 5019 (1) on page 60, DOI: 10.11646/zootaxa.5019.1.1, http://zenodo.org/record/5222728