Upembarus Koch 1956
Creators: name: Kamiński, Marcin Jan
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Tenebrionidae
Upembarus

description: SUBGENUS UPEMBARUS KOCH, 1956   
 Type species: Upembarus saegeri Koch, 1956; by original designation.   
 Diagnostic characters: See diagnostic characters for subgenus Pseudoselinus.   
 Distribution: Democratic Republic of the Congo and Zambia.   
 Species composition (four species): Upembarus saegeri Koch, 1956, Upembarus sympatrius Koch, 1956, Upembarus upembaensis Koch, 1956, Upembarus wittei Koch, 1956.   
ACKNOWLEDGEMENTS  
The author is grateful to Dariusz Iwan (MIIZ) for the supervision during the PhD project. Additionally, for the valuable comments on the manuscript, the author thanks Joanna Ma˛ kol (Wrocław University of Environmental and Life Sciences), Grzegorz Pas´nik (Institue of Systematics and Evolution of Animals, PAS), Rolf Aalbu (California Academy of Sciences), and the anonymous reviewers assigned by the Zoological Journal of the Linnean Society. The author would also like to thank the curators of the institutions cited above for the loan of specimens in their care. This research received support from the SYNTHESYS Project (ES-TAF- 3891; http://www.synthesys.info/), which is financed by European Community Research Infrastructure Action under the FP7 ‘Capacities’ Programme at the Museo Museo Nacional de Ciencias Naturales (CSIC).  
KEY TO THE AFROTROPICAL PLATYNOTOID PLATYNOTINA GENERA AND SUBGENERA  
1. Pronotal disc without basal indentations. Ratio of the pronotal disc’s total width to its height <5.0. Ratio of pronotal disc height to prothorax height> 0.4................................................................................................. (2)  
 Pronotal disc with a pair of basal indentations. Ratio of the pronotal disc’s total width to its height> 6.0. Ratio of pronotal disc height to prothorax height <0.3.................................................. Ectateus generic group (6)   
 2. Anaepisternum and metaventrite long (Fig. 25). Wings present (reduced in Z. exalatus and Z. royi)....... Zidalus   
 Anaepisternum and metaventrite reduced. Wings absent................................... Upembarus generic group (3)   
 3. Ventral margin of eyes with a groove below (Fig. 11). Distribution: Madagascar................................. Lechius   
Ventral margin of eyes without a groove below. Distribution: continental Africa........................................(4)  
 4. Middle of mentum not projecting anteriad. Anterior tentorial pits invisible. Intercoxal process of prosternum narrow............................................................................................................................. Paraselinus   
 Middle part of mentum projecting anteriad (Fig. 7). Anterior tentorial pits longitudinal (Fig. 9). Intercoxal process of prosternum wide (Fig. 17)............................................................................................ Upembarus (5)   
 5. The upper edge of the elytral base disappearing before humerus (cf. Kamin´ ski, 2013c). Fifth abdominal ventrite without bordering. Bursa copulatrix without sclerites..................................................subgenus Upembarus   
 The upper edge of the elytral base reaching the humerus. Fifth abdominal ventrite bordered. Bursa copulatrix with longitudinal sclerites...................................................................................subgenus Pseudoselinus   
 6. Anaepisternum and metaventrite long. Wings reduced (but present) (Fig. 30)..............  Pteroselinus gen. nov.    
Anaepisternum and metaventrite short. Wings absent..........................................................................(7)  
7. Anterior tentorial pits deep, clearly visible as circular depressions (Fig. 10)............................................. (8)  
Anterior tentorial pits invisible or visible as longitudinal depressions..................................................... (9)  
 8. Pronotum widest at the base, sides evenly narrowing towards apex. Fifth abdominal ventrite bordered. Penis narrow; clavae developing from two-thirds of the aedeagal tegmen length (cf. Kamin´ ski, 2014)............. Selinus   
 Pronotum widest in the middle, sides rounded. Fifth abdominal without bordering. Penis wide; clavae developing from half of the aedeagal tegmen length (cf. Kamin´ ski, 2014)...................................................... Monodius   
 9. Dorsal side of frons with conspicuous depressions (Fig. 6). Bursa copulatrix with several small sclerites (Fig. 27)........................................................................................................................... Nesopatrum   
Dorsal side of frons without depressions. Bursa copulatrix with a single sclerite or without sclerites.......... (10)  
 10. Submentum pentagonal (Fig. 5). Last segment of maxillary palpi strongly widened (Fig. 4)... Anchophthalmus....................................................................................................................................................(11)   
Submentum triangular. Last segment of maxillary palpi narrow...........................................................(12)  
 11. Lateral wings of mentum developing at the level of two-thirds of mentum length (Fig. 5). Base of the fourth elytral interval protruding towards the front of the body (cf. Kamin´ ski & Iwan, 2013a)............. subgenus Kochogaster   
 Lateral wings of mentum visible from the base. Base of the fourth elytral interval not protruding towards the front of the body............................................................................................subgenus Anchophthalmus   
 12. Pronotal disc and elytra densely covered with setae. Elytral humeri strongly protruding sideways (cf. Iwan, 2001a). Clavae developing from half of the aedeagal tegmen length (Fig. 29)....................................... Phallocentrion   
Pronotal disc and elytra densely glabrous or covered with fine and sparsely distributed setae. Elytral humeri not protruding outwards. Clavae developing from two-thirds of the aedeagal tegmen length.................................................................................................................................................................................(13)  
 13. Apical part of mentum with a median indentation (Fig. 8)......................................................... Angolositus   
Apical part of mentum straight.......................................................................................................(14)  
 14. Third antennomere short (ratio of its length to the length of second antennomere ≥ 2.0). Elytral humeri strongly protruding towards body apex (Fig. 20)................................................................................. Glyptopteryx   
Third antennomere long (ratio of its length to the length of second antennomere <3.0). Elytral humeri not protruding towards body apex (Fig. 19).................................................................................................(10)  
 15. Bases of the first four elytral rows connected in the following way: 1–2, 3–4 (Fig. 19). Scutellum dented. Elytral epipleuron flat over its whole surface. Apex of the elytral epipleuron emarginated............................. Ectateus   
Bases of the first four elytral rows not connected. Scutellum situated at the level of elytra. Elytral epipleuron curved at the level of the base of fifth ventrite. Apex of the elytral epipleuron not emarginated.................(16)  
 16. Antennomeres from seventh to 11th elongated (Fig. 12B). Parallel basal halves of the lateral elytral sides. Elytral intervals covered with coarse tubercles (Fig. 22)................................................................... Phymatoplata   
Antennomeres from seventh to 11th transverse (Fig. 12A). Rounded basal halves of the lateral elytral sides. Elytral intervals without tubercles.............................................................................................................(17)  
 17. Antennomeres from seventh to 11th triangular. Scutellum small (ratio of the width of a single elytra to the base of the scutellum> 15.0; Fig. 18). Base of parameres strongly widened (Fig. 28)..................... Anchophthalmops   
 Antennomeres from seventh to 11th rounded. Scutellum large (ratio of the width of a single elytra to the base of the scutellum <12.0). Base of parameres slightly widened (cf. Kamin´ ski, 2014)............................. Eleoselinus
description: Published as part of Kamiński, Marcin Jan, 2015, Phylogenetic reassessment and biogeography of the Ectateus generic group (Coleoptera: Tenebrionidae: Platynotina), pp. 73-106 in Zoological Journal of the Linnean Society 175 (1) on pages 93-94, DOI: 10.1111/zoj.12263, http://zenodo.org/record/10831727