Lispe caespitosa Pont 2019, sp. nov.
Creators: name: Pont, Adrian C.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Anthomyiidae
Lispe
Lispe caespitosa

description: Lispe caespitosa sp. nov.
(Figs 284–294)
Diagnosis. In the group of species with a seta on posterior apical margin of hind coxa, the male of L. caespitosa can be recognised by its dark palpus and the conformation of hind tibia and tarsomeres (Fig. 287).
Etymology. This name is from the Latin “caespes” (“turf”) and refers to the tufted armature at the tip of the male hind tibia and tarsomere 1 (Fig. 287).
Material examined. Holotype ♂. AUSTRALIA: South Australia: Arno Bay, mangrove, 33.92S 136.57E, 8– 9.ii.2013, N. Vikhrev (ANIC). Paratypes 5♂ 5♀. AUSTRALIA: data as for the holotype, 3♂ 5♀ (ZMUM). Victoria: Lake Tyrrell, 21.xi.1975, K.R. Norris, 1♂ (ANIC); Pink Lakes, 73 km W of Ouyen, 13.x.1975, Z. Liepa, 1♂ (ANIC).
Dr N.E. Vikhrev has kindly presented the holotype of this Australian species to the ANIC.
Additional material (N.E. Vikhrev, pers. comm.): AUSTRALIA: South Australia, Arno Bay, mangrove, 33.92S 136.57E, 8–9.ii.2013, N. Vikhrev, 13♂ 14♀ (ZMUM).
Description ♂ ♀. Head (Fig. 285). Ground-colour black. Frons at broadest point 0.34 (♂) or 0.30 (♀), at lunula 0.27, of maximum head-width (frontal view). Fronto-orbital plate uniformly pruinose, together with parafacial, face, gena and most of occiput silvery-white pruinose, more yellow in ♀ except for occiput, face and facial ridges tinged with yellow, and a distinct brown spot at lunula; frontal triangle yellowish- to brownish-grey, very broad, much of frontal vitta suppressed. Lower orbital seta short. Parafacial narrow, half as broad as postpedicel; entirely setulose. Antenna black; postpedicel short, 2.0 times (♂) or 2.7 times (♀) as long as broad, falling short of mouthmargin by its length (♂) or slightly less than this (♀). Arista plumose, the hairing at widest point twice width of postpedicel. Vibrissa short. Gena broad, 0.18 of vertical eye-length; genal setae strong and dense. Palpus brown, stalk yellow, strongly swollen at tip (Fig. 286).
Thorax. Ground-colour black. Scutum rather densely grey dusted; postpronotal lobe, notopleuron and pleura very light grey; scutellum entirely brown dusted. Viewed from behind, scutum thinly and irregularly grey dusted, with a broad dark brown patch occupying the whole area between dc, from neck to 1st post dc, this dark area with a pair of paler dusted narrow paramedian vittae (♂) or with a single broad median pale vitta (♀). Anterior spiracle yellowish. Acr in 4–5 irregular rows at suture. Dc 2+3. 2 pprn. 1+1 ia. 1+1 sa, both strong. 1 strong and 1 weaker proepisternal setae. 1 strong proepimeral, slightly shorter than the strong proepisternal, with 4 adjacent setulae. Anepisternum with a very short seta in upper anterior corner; posterior row with 3 strong and 3 weak setae. 3 strong kepst, lower one closer to posterior than to anterior. Meron bare below spiracle and above hind coxa. Scutellum with disc densely setulose, 2 irregular rows descending on to sides, otherwise sides and ventral surface bare.
Legs. Black, except for reddish knees. Fore coxa bare behind. Fore femur with a row of short spinules on av surface; pv row complete, the setae long and fine, longer than femoral depth, interspersed with multiserial v to pv spinules; pd row complete. Fore tibia without submedian seta; with d, pd and pv apical setae, and also with a distinct ad apical. Fore tarsomeres rather short, without modifications. Mid femur with only short setulae on av surface; on pv surface with fine setae in basal half and thereafter only with short setulae; basal half also with a row of short strong a setae, culminating in a strong seta at middle, these all much weaker in ♂ than in ♀; 2 p preapicals. Mid tibia with 1 strong p seta; with strong ad, av, v, pv and p apicals, the p short, the ad apical a little before apex. Mid tarsomeres without modifications. Hind coxa with a seta on posterior apical margin. Hind femur setulose on av surface, and 1 fine seta at middle in ♂ and another before apex; pv surface with 4–5 long fine setae in basal half, otherwise only with setulae; ad row complete; 1 d preapical. Hind tibia of ♂ with 1 ad, with the ground-setulae on this surface long and erect; 1–2 av seta; tibia at and just before apex with numerous long fine setae and setulae on all surfaces, and thus with no distinct apical setae (Fig. 287); in ♀ with 1 ad and with d and ad preapicals, all three very strong, 1 av, and av and pv apical setae. Hind tarsus of ♂ with basal tarsomere slightly swollen at tip of pv surface and there covered with a short brush of setulae (Fig. 287); without modifications in ♀.
Wing. Clear. Tegula black, basicosta yellow. Crossvein r-m placed below the point where vein R1 enters costa; dm-cu weakly sinuous, longer than apical section of vein CuA1. Vein M running straight to wing-margin. Calypters white, margins yellow. Knob of haltere black.
Abdomen. Ground-colour black. Tergites light grey dusted, with a pair of small poorly defined spots on tergite 3 in ♂, these spots larger and more semi-circular in ♀, and a broad black central spot on tergite 4 that runs down on to sides of tergite and encroaches broadly on tergite 5 (Figs 288–289). ♂ epandrium grey dusted. ♀ ovipositorwith the exposed tergites grey dusted. Sternites 1–5 grey dusted, sternite 5 of ♂ shining. Setae only present on tergite 5, with 2 pairs of median and 2 of lateral discals (♂) or 2–3 pairs of discals (♀), and 4–5 (♂) or 2 (♀) pairs of erect marginals. Sternites 2–4 with sparse setulae (Fig. 290).
Ƌ terminalia. Epandrium separated from tergite 5 by syntergosternite 8, which has 2 spiracles; without large setae; weakly produced lobe-like ventrally (Fig. 291). Tergite 6 absent. Sternite 6 present as a small plate, withdrawn beneath sternite 5 and connected to it by a pair of broad outgrowths from the sternite. Surstylus absent, i.e. fused to epandrium without trace (Fig. 291). Cercal plate divided dorsally and ventrally (Fig. 292). Hypandrium attached at two points: to a rod originating from wall of epandrium close to upper outer edge of cercal plate; and via a tiny apparent praegonite to lower anterior corner of epandrium. Phallic complex (Figs 293–294): hypandrium with a pair of processes at posterior end that meet and unite with base of epiphallus; praegonite and postgonite present, the postgonite curved outwards (Fig. 294); phallapodeme large, at mid-length joined to hypandrium by a small bridge; phallus long, tube-like, simple, juxta large, without spinules, without paired processes at base.
♀ ovipositor. Not examined.
Measurements. Wing-length, 4.0– 4.5 mm (♂), 4.0 mm (♀). Body-length, 6.0– 6.5 mm, 5.5 mm (♀).
Biology. Adults have been collected among mangroves.
Distribution. Known only from Australia (V, SA).
description: Published as part of Pont, Adrian C., 2019, Studies on the Australian Muscidae (Diptera). VIII. The genus Lispe Latreille, 1797, pp. 1-232 in Zootaxa 4557 (1) on pages 135-138, DOI: 10.11646/zootaxa.4557.1.1, http://zenodo.org/record/2584520