Torodora hana Park & Koo & Minet & Razafindranaivo 2023, sp. nov.
Creators: name: Park, Kyu-Tek
name: Koo, Jun-Mo
name: Minet, Joël
name: Razafindranaivo, Victor
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Lecithoceridae
Torodora
Torodora hana

description: 1. Torodora hana Park &Koo, sp. nov.
(Figs. 2, 3)
LSID: urn:lsid:zoobank.org:act: 161117FD-7E30-49EA-A923-9E375D8B2D87
Type specimen. Holotype: Male, [East] Madagascar, Alaotra-Mangoro, Mantadia National Park, 18°52′24.55“S, 48°36′19.11“E, alt. 721 m, 26 xi 2019, leg. Jun-Mo Koo; gen. slide no. CIS-7422 (in NIBR). Paratypes: 17♁, 13♀, same data as holotype; gen. slide no. CIS-7165 (♁), COI barcode CBNU125; gen. slide no. CIS-7419 (♁), COI barcode CBNU146; gen. slide no. CIS-7420 (♁), COI barcode CBNU144; gen. slide no. CIS-7321 (♀), COI barcode CBNU145; gen. slide nos. CIS-7423 (♀), -7424 (♀); COI barcode CBNU143; wings slide no. CIS-7421 (in NIBR); 1♁, [East] Madagascar, near 3rd Integral Natural Reserve [Zahamena National Park], road to Manakambahiny E, near Nosivola, alt. 1,000 m, 18 ii [19]54, leg. P. Viette; gen. slide no. CIS-7396 (in MNHN). Additional material: 1♁, [East] Madagascar, Andasibe, 845 m, 200 km from the coast, no coll. date, a photo provided by Maik Bippus (Réunion).
Diagnosis. The new species can be distinguished from its congeners by the bifurcated uncus of the male genitalia with slender latero-caudal processes, and the narrowly elongated cucullus.
Description. Male and female (Figs. 2A–D). Wingspan, 14.0– 16.5 mm. Head: yellowish brown (grayish brown centrally on vertex), with dense yellowish-white erect scales laterally. Antenna longer than forewing, about 1.2 times the length of forewing; basal segment elongated, golden yellow dorsally, yellowish brown on both sides; flagellum golden yellow, without distinct annulations. 2 nd segment of labial palpus thickened, yellowish brown on outer surface; 3 rd segment slender, longer than 2 nd segment, upturned. Thorax: tegula and thorax yellowish brown dorsally. Hind tibia with yellowish-brown rough scales dorsally. Forewing broadened distally; ground color yellowish-brown, with scattered dark-brown scales; a pair of (often indistinct) dark spots at middle and at end of discal cell; costa slightly arched in basal 1/3, then oblique beyond 2/3; apex angulate; termen slightly concave medially; fringe concolorous with ground color, with yellowish-white basal line; venation (Fig. 2E) with R 1 arising from near middle of cell, R 2 arising from near upper corner of cell; distance between R 1 and R 2 about twice that separating R 2 from stem R 3+4+5; R 3, stalked with R 4+ 5 in basal 1/3; R 4 and R 5 stalked in proximal 2/3; R 5 to termen; M 1 remote from R 3+4+5, nearly parallel to M 2; base of M 2 close to M 3; M 3 free, arising from lower corner of cell; CuA 1 and CuA 2 stalked for proximal 3/5; 1A+2A narrowly forked basally; cell closed. Hind wing ground color as in the forewing but slightly lighter; apex angulate; base of vein M 2 close to M 3 +CuA 1; M 3 and CuA 1 stalked for about basal half; cell weakly closed before middle of wing. Abdomen (Fig. 3C): yellowish-brown, speckled with dark-brown scales dorsally, yellowish white on segments V-VIII laterally; narrow spinose zones on dorsal surface along posterior edges; sternum VIII trapezoidal, lacking specially modified sclerites, but with small triangular lobes on anterior margin and a narrow, crescent-shaped, linear sclerite anteriorly; posterior margin more or less convex medially.
Male genitalia (Figs. 3A, B): Uncus bifurcate, Y-shaped, with narrow latero-caudal processes, apices rounded. Basal plate of gnathos fan-shaped, with round caudal margin; median process small, about as long as process of uncus. Tegumen broad, with inverted V-shaped anterior margin. Valva broad at base; costa abruptly oblique from near base; ventral margin nearly straight, slightly emarginated at lower corner of cucullus; sacculus weakly developed; cucullus elongated (about 1.5 times the length of basal part of valva), gradually broadened distally, densely setose, with strong setae along margins; apex rounded. Juxta quadrate, with short, digitate latero-caudal processes; caudal margin slightly convex medially. Aedeagus stout, broadened basally; ventral margin strongly produced basally; cornuti consisting of several rows with minute spinules.
Female genitalia (Fig. 3D): Caudal margin of abdominal sternum VIII deeply emarginated (with U-shaped notch), several close-set setae along caudal margin. Antrum funnel-shaped, membranous. Ductus bursae narrowed in posterior 1/5, broadened beyond, once coiled in anterior 1/5; ductus seminalis arising from near posterior 1/3. Corpus bursae large, ovate, slightly shorter than ductus bursae; signa consisting of a crescent-shaped, heavily sclerotized sclerite (near at posterior entrance extended to an inverted hat-shaped plate at left side), and an elliptical spinose plate at right side beyond half of corpus bursae.
Distribution. Madagascar (eastern region).
Etymology. The species name is derived from the Korean term hana, meaning the first or a single.
description: Published as part of Park, Kyu-Tek, Koo, Jun-Mo, Minet, Joël & Razafindranaivo, Victor, 2023, Lecithoceridae (Lepidoptera) of Madagascar: I. Descriptions of eighteen new species of Torodora Meyrick, 1894, pp. 151-184 in Zootaxa 5264 (2) on pages 153-156, DOI: 10.11646/zootaxa.5264.2.1, http://zenodo.org/record/7836316