Tingidae , Drake and Ruhoff 1965
Creators: name: Shofner, Ryan
name: Cassis, Gerasimos
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Tingidae

description: Tingidae genera associated with the Proteaceae
Tingid clades are often associated with a narrow range of plant genera (Cassis and Symonds 2011, Cassis et al. 2017), which are often closely related. Based on recent surveys (Cassis et al. 2007) numerous tingid taxa are associated with the plant family Proteaceae. Host records from the PBI database were harvested for a list of tingid taxa associated with Proteaceae (Supporting Information, Table S4). Three genus-group taxa were recovered from our molecular analysis that are specialized on the Proteaceae. In addition, there are several genera that have one to several species associated with the Proteaceae but are not exclusively associated with the family. The following key accounts for all genera that have at least one species associated with the Proteaceae.
Key to Tingidae associated with Proteaceae
1 Paranota highly modified, extending dorsally onto pronotal disc and exceeding lateral carinae, inflated to produce paired bulbous structures (Fig. 4A, B, para) .........................................................................................................................................................2
2 Paranota not as above ....................................................................................................................................................................................3
3 Frons porrect, extending beyond antenniferous tubercles (Fig. 4A, B, frn) ................................................... Cysteochila Stål 1873
4 Frons flattened to slightly protruding, but never exceeding antenniferous tubercles (Fig. 4G, frn) .......... Oncophysa Stål 1873
5 Pair of spines present on collum, their bases proximal to medial carina (Fig. 4C, pdsp) ....................... Engynoma Drake 1942b
6 Collum without spines ..................................................................................................................................................................................4
7 Frons porrect, extending beyond antenniferous tubercles.....................................................................................................................5
8 Frons flattened to slightly protruding, but never exceeding antenniferous tubercles.......................................................................6
9 Subcostal area four areolae wide; dorsal surface of pronotum and hemelytra glabrous ......................... Chorotingis Drake 1961
10 Subcostal area three or four areolae wide, if four then dorsal surface of pronotum and hemelytra with short hairs................... ....................................................................................................................................................................................... Euaulana Drake 1945
11 Trochanters fused to femurs (Fig. 4D).............................................................................................................. Epimixia Kirkaldy 1908
12 Trochanters differentiated from femurs (Fig. 4E)....................................................................................................................................7
13 Paranota absent or reduced to carina-like structure on pronotal disc; if present anteriorly, upturned and adpressed to collum (Fig. 4F, G, para).............................................................................................................................................................................................8
14 Paranota present, areolate over entire length; if reduced posteriorly on pronotal disc, paranota never adpressed to collum (Fig. 5A, B, E, F, para)................................................................................................................................................................................. 10
15 Peritreme of metathoracic gland present (Fig. 4G); medial spine present (Fig. 4F, G); costal area areolate................................ ................................................................................................................................................................... Eritingis Drake and Ruhoff 1962
16 Peritreme of metathoracic gland absent (Fig. 5d); medial spine absent or greatly reduced in size (Fig. 5c, d); if medial spine present, costal area reduced to carina-like structure................................................................................................................................9 17 Costal area areolate for at least portion of length; mesosternal and metasternal carinae straight, forming narrow rostral channel; dorsal surface of pronotum and hemelytra glabrous .......................................................................... Malandiola Horváth 1925
18 Costal area reduced to carina-like structure, lacking areolae; mesosternal and metasternal carinae curved, forming broad rostral channel; dorsal surface of pronotum and hemelytra usually with setae or hairs ......................... Nethersia Horváth 1925
19 Collum raised anteriorly, forming hood (Fig. 5B, hd); paranota never more than two areolae wide anteriorly, one areola wide at humeral angles (Fig. 5a); costal area one areolae wide over entire length.................................................. Proteatingis gen. nov.
20 Collum flattened, or if raised, only posteriorly, never forming hood; paranota two or more areolae wide, one to three areolae wide at humeral angles or carina-like; costal area one or two areolae wide, at least partially ...................................................... 11
21 Costal area one areolae wide over entire length; dorsal surface of pronotum glabrous .......... Ulonemia Drake and Poor 1937a
22 Costal area two areolae wide anteriorly; dorsal surface of pronotum with scale-like setae........................... Cercotingis gen. nov.
description: Published as part of Shofner, Ryan & Cassis, Gerasimos, 2024, Phylogenetics and taxonomy of the Indo-Australian genus Ulonemia sensu Drake (Hemiptera: Tingidae), with the recognition of new genera and species collected from Proteaceae in Australia, pp. 360-425 in Zoological Journal of the Linnean Society 200 (2) on pages 375-376, DOI: 10.1093/zoolinnean/zlad080, http://zenodo.org/record/11240237