Draculoides nosferatu Abrams and Harvey 2020, n. sp.
Creators: name: Abrams, Kym M.
name: Huey, Joel A.
name: Hillyer, Mia J.
name: Didham, Raphael K.
name: Harvey, Mark S.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Schizomida
Hubbardiidae
Draculoides
Draculoides nosferatu

description: Draculoides nosferatu Abrams and Harvey, n. sp.
(Figs. 1–8, 36–37)
Zoobank Code: http://zoobank.org/NomenclaturalActs/ 51471C86-8899-4E6A-B0DB-8F9F369C05F6
Draculoides SCH 061: Abrams et al. 2019 MPE 106532: 8, fig. 2.
Material examined. Holotype female. AUSTRALIA: Western Australia: 1 ♀, Robe Valley, ca. 27 km ESE. of Pan- nawonica, 21°44’27.72”S, 116°33’24.81”E, 6/06/2015, 8/08/2015, troglofauna trap, J. Alexander (WAM T138516) (DNA: COI, 18S, 28S, ITS2).
Other material. AUSTRALIA: Western Australia: 1 juvenile, Robe Valley, ca. 27 km ESE. of Pannawonica, 21°44’31.19”S, 116°33’41.85”E, 6 June–8 August 2015, troglofauna trap, J. Alexander (WAM T138518) (DNA: COI, 18S, ITS2); 1 juvenile, Robe Valley, ca. 27 km ESE. of Pannawonica, 21°44’37.56”S, 116°33’49.11”E, 6 June–8 August 2015, troglofauna trap, J. Alexander (WAM T138519) (DNA: COI, 18S, ITS2); 1 juvenile, Robe Valley ca. 27 km ESE. of Pannawonica, 21°44’31.19”S, 116°33’41.85”E, 6 June–8 August 2015, troglofauna trap, J. Alexander (WAM T139921).
Diagnosis. Males are unknown and the flagellum is missing from the only available female. Draculoides nosferatu is the only Draculoides species known that does not have a guard tooth on the movable finger of the chelicera. Draculoides nosferatu can be diagnosed from all other Draculoides species that were sequenced at COI by the 50bp mini-barcode shown in Figure 3. Draculoides nosferatu can be diagnosed from all other Draculoides species that were sequenced at ITS2 (Fig. 7) except for D. claudiae, D. carmillae and D. mesozeirus, which are not distinguishable using the ITS2 mini-barcode.
Description (adult female). Colour. Yellow-brown; propeltidium and pedipalps darker.
Cephalothorax. Propeltidium with 2+1 apical setae in a triangular formation on anterior process and 2 + 2 + 2 setae; eye spots absent. Mesopeltidia separated. Metapeltidium divided. Anterior sternum with 14 setae (including 2 sternapophysial setae); posterior sternum triangular with 6 setae.
Chelicera. Fixed finger with 2 large teeth plus 5 smaller teeth between these, proximal and distal teeth without small lateral teeth; membranous area between fixed and movable fingers with 3 large, lanceolate, terminally pilose setae (G1); G2 composed of 6 setae; G3 composed of 4 setae; internal face of chelicera with 5 short whip-like setae (G4); brush at base of fixed finger composed of 7 setae (G5A), each densely pilose in distal half and G5B composed of 10 setae; G6 with one seta; G7 composed of 2 setae. Movable finger serrula composed of ca. 18 long lamellae, guard tooth absent; 3 accessory teeth present, most distal large, most proximal tiny.
Pedipalp. Without apophyses; trochanter with sharply produced ventro-distal extension, ventral margin with ca. 9 stout setae, without mesal spur; tarsus and tibia without spines; tarsal spur present; claw 0.36 × length of tarsus.
Legs. Tarsus I with 6 segments; baso-dorsal margin of femur IV produced at about a 90° angle.
Abdomen. Chaetotaxy of tergite I: 2 macrosetae + 4 microsetae (microsetae diagonal), tergite II:? (damaged) macrosetae + 6 microsetae (microsetae in column), tergites III–IX: 2: 2: 2: 2: 2: 2: 2.
Female genitalia. Two pairs of spermathecae with short, equal-sized lobes, each pair connected basally before connection with bursa (Fig. 37A), distally round and smooth; sparsely covered with small pores; gonopod short, rectangular, distally slightly bifurcate.
Flagellum. Collected without flagellum.
Dimensions (mm). Holotype female (WAM T138516): Body length 3.56. Propeltidium 1.21/0.62. Chelicera 0.69. Flagellum broken. Pedipalp: trochanter 0.46, femur 0.50, patella 0.54, tibia 0.44, tarsus 0.27, claw 0.10, total excluding claw 2.21.
Remarks. Draculoides nosferatu is known from three locations within the most southern mesa of the Middle Robe mesas (Fig. 1D). The phylogeny suggests that D. nosferatu ’ s closest relatives are the other species from the Middle Robe area including D. minae, D. claudiae, D. carmillae and D. mesozeirus (Fig. 2 Clade H). The juvenile specimens listed above are associated with this species by locality and, in many cases, by sequence data.
Other names. WAM SCH061 (Abrams et al., 2019).
Etymology. The specific epithet refers to the seminal 1922 German horror film “Nosferatu: A Symphony of Horror,” and relates to the species living in the dark.
description: Published as part of Abrams, Kym M., Huey, Joel A., Hillyer, Mia J., Didham, Raphael K. & Harvey, Mark S., 2020, A systematic revision of Draculoides (Schizomida: Hubbardiidae) of the Pilbara, Western Australia, Part I: the Western Pilbara, pp. 1-75 in Zootaxa 4864 (1) on pages 63-65, DOI: 10.11646/zootaxa.4864.1.1, http://zenodo.org/record/4416849