Aquattuor Frederiksen 2013
Creators: name: Enghoff, Henrik
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Spirostreptida
Odontopygidae
Aquattuor

description: Genus Aquattuor Frederiksen, 2013
Diagnosis (modified after Enghoff & Frederiksen 2015)
Differs from all other Odontopygidae by the structure of the limbus, which consists of large, rectangular, easily detachable flaps. Gonopods of the Prionopetalini type, i.e., basomere of telopodite with a torsate flexible zone followed by a posttorsal narrowing; solenomere elongated. Gonopodal proplica apically expanded into a distal palette; metaplica with an oblique flange; solenomere simple, thin and whip-like; telomere terminating in a long, curved, gutter-like to tubular part.
Descriptive notes
For characters not mentioned here, see Enghoff & Frederiksen (2015).
Mandibular stipes (Fig. 2): in males, but not in females with a disto-ventral expansion.
First pair of male legs: in the genus Aquattuor the first pair of legs is remarkably diverse. Figs 3–7 illustrate the first pair of legs in all known species of the genus, with a focus on the prefemur. In all of them, the prefemur carries a more or less finger-shaped process (pfp) which projects orad from the anterior surface of the prefemur (Figs 3–7). The distal (ventral) surface of the process is smooth, the basal (dorsal) surface is more or less scaly-rugose. The shape of the process varies from slender fingershaped to relatively broad/triangular, and in one species (A. nguruensis sp. nov.) there is a clearly delimited depression (pfd) on the distal surface of the process (Fig. 4 A–C). In A. major Enghoff, 2015, A. longipala Enghoff, 2015 and A. mollilobus sp. nov. there is in addition a second, more distal, more irregularly shaped prefemoral lobe (dpl, Figs 5–6).
Another type of variation is seen in the setation of the telopodites of the first pair of legs (Fig. 8). Here, A. mollilobus sp. nov. (Fig. 8C) stands out by having very short setae on all podomeres. Aquattuor major and A. longipala have long setae on the tarsus, but very short ones on the femur (Fig. 8B). The remaining species have long setae (same length as on the following leg-pairs) on all podomeres (Fig. 8A). These differences in setation do not seem to reflect the setation of normal walking legs.
In one species (A. udzungwensis), the podomeres distal to the prefemur of the first leg pair seem to be invariably missing (several specimens examined) (Fig. 7 D–E).
Adhesive pads on walking legs are absent.
Included species (alphabetically)
Aquattuor claudiahempae Enghoff & Frederiksen, 2015
A. denticulatus Frederiksen, 2013 (type species)
A. fasciatus (Attems, 1896)
A. longipala Enghoff, 2015
A. major Enghoff, 2015
A. mollilobus sp. nov.
A. nguruensis sp. nov.
A. stereosathe Enghoff, 2015
A. submajor Enghoff, 2015
A. udzungwensis Enghoff, 2015
description: Published as part of Enghoff, Henrik, 2020, A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae), pp. 1-32 in European Journal of Taxonomy 626 on pages 4-5, DOI: 10.5852/ejt.2020.626, http://zenodo.org/record/3754989