Drapetis incompletoides Zouhair & Grootaert & Gençer & Kettani 2025, sp. nov.
Creators: name: Zouhair, Laila
name: Grootaert, Patrick
name: Gençer, Lütfiye
name: Kettani, Kawtar
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Hybotidae
Drapetis
Drapetis incompletoides

description: Drapetis incompletoides Zouhair & Grootaert sp. nov.
(Figs 11, 12)
Type material. HOLOTYPE. Sivas prov. ♂, Yýldýzeli, 39.788 N, 36.21 E, 1185 m, grassland, 07.vi.2009, sweep net, leg. L. Gençer. PARATYPES: Sivas prov. 10♂, same data as holotype; 1 ♂, Taþlýdere, 39.564 N, 37.041 E, 1338 m, grassland, 24.v.2009, sweep net, leg. L. Gençer.
Diagnosis. A dark-legged species of the exilis-group with upper crossvein distinctly beyond middle of second basal cell (bm). Antennae blackish brown, pedicel with a circlet of short brownish setae. Mid femur simple, with row of thick brown setae in basal half, becoming shorter towards apex and 2 long black subapical setae. Hind femur weakly curved in basal third. Wing with longitudinal veins thickened, vein R 4+5 slightly undulating, pale and faded shortly before wing margin, vein M 1+2 straight.
Description. Male. Length: body 2.2 mm, wing 2.2 mm. Head black in ground colour. Occiput densely greyish pollinose, covered with short black setulae, becoming longer below. A pair of long convergent black verticals, with pair of long black postocular setae at sides, slightly shorter than verticals. Frons densely greyish pollinose, broadened towards ocellar tubercle, narrowing downwards to antennae. Face grey dusted, linear, very narrower than pedicel, widening immediately above clypeus. Clypeus densely grey dusted. Ocellar tubercle densely greyish pollinose, with two pairs of black long ocellars (anterior pair longer, crossing; posterior pair slightly shorter than anterior and divergent). Antenna blackish brown, pedicel with a circlet of short brownish setae, postpedicel nearly quadrangular, about as long as wide, with long apical stylus, about 2.5X as long as all antennal segments combined. Palpus blackish, slightly elongate, rather ovate, covered with pale setae and longer subapical pale seta present. Proboscis blackish.
Thorax mostly shiny black, except on prothorax, proepimeron, scutellum, upper half of anepisternum, anepimeron and margins of katepisternum, meron densely greyish dusted. No upturned seta on proepisternum. Mesonotum densely set with brownish setulae. Acrostichals not distinct from surrounding setulae. Prominent setae: pair of long black prescutellar dorsocentrals, notopleurals (2 long black thick setae, 1 shorter and faint seta), 1 pair of long black scutellars, 1 pair of long black postalars. Legs extensively blackish brown, but fore legs somewhat slightly light brown. Leg chaetotaxy mainly consisting of short pale setulae; coxae and trochanters with unmodified setation. Fore femur somewhat thickened, slightly thicker than mid femur, with 1 thick brown posteroventral subapical seta and ring of apical brown setae, ventral setae lacking. Fore tibia slender with 1 thick brown posterovental subapical seta and 1 posterodorsal subapical seta. Mid femur simple, with row of somewhat thick brown setae in basal half, becoming shorter towards apex and 2 long black subapical setae. Mid tibia simple, with 1 thick brown subapical seta. Hind femur weakly curved on basal third, with ventral pale setae and some short dorsal preapicals. Hind tibia slightly widened toward apex, making a short posteroventral apical tooth-like projection. Hind tarsomere 1 about as long as combined length of hind tarsomeres 2–3. Wing faintly brownish tinged, covered with uniform brown microtrichia, with brown veins, but costa and vein R 1 darker and longitudinal veins thickened. Vein R 4+5 slightly undulating, pale and faded shortly before wing margin, vein M 1+2 straight. Vein CuA+CuP present, distinctly indicated by pale vein, not reaching wing margin. Squama blackish. Haltere brownish.
Abdomen grey dusted, covered with pale setulae (tergite 9 with long black marginal setae, sclerites blackish brown with basal sclerites somewhat yellowish. Tergite I not sclerotized, sternite I very short. Terminalia (Fig. 12) with right cercus much smaller and thinner than left cercus (Fig. 12B). Left cercus large with truncate tip, broadest at middle. Left epandrial lamella with 6 strong setae near apical border. Right epandrial lamella densely set with long fine setae (Fig. 12A).
Female. Unknown.
Etymology. This species is named after its similarity to D. incompleta.
Remarks. In the key to males of species of Drapetis of Northwest Europe provided by Michelsen & Grootaert (2019), the new species will run to the exilis -group as the upper cross vein ends beyond the middle of the basal medial cell (bm). Since the new species has mid femur and tibia simple, and the wing with longitudinal veins thickened and vein R 4+5 pale and faded shortly before wing margin, it will run to D. incompleta. Both species are very close morphologically, but genitalia are distinctly different (compare Fig. 12 with figs 611–613 in Chvála 1975).
description: Published as part of Zouhair, Laila, Grootaert, Patrick, Gençer, Lütfiye & Kettani, Kawtar, 2025, Turkish Hybotidae (Diptera: Empidoidea): description of six new species with additional data, pp. 503-524 in Zootaxa 5609 (4) on pages 515-516, DOI: 10.11646/zootaxa.5609.4.3, http://zenodo.org/record/15215427