Asiobaccha tripartita, comb. nov.
Creators: name: Mengual, Ximo
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Syrphidae
Asiobaccha
Asiobaccha tripartita

description: <i>Asiobaccha tripartita</i> (Walker, 1861) comb. nov. (Figures 11c, d, f, 14d) <i>Baccha tripartita</i> Walker, 1861: 285. Holotype: &amp;male;, BMNH. Type locality: Indonesia: Misool Island. Schiner 1868: 344; Bigot 1892: 166; Wulp 1896: 121; Kertész 1910: 157. <i>Baccha flavipes</i> Doesburg, 1959: 233. Syn. nov. <i>Baccha sulica</i> Austen, 1893: 144 (in part). Syn. nov. Wulp 1896: 121; Kertész 1910: 157. <i>Baccha</i> (<i>Allobaccha</i>) <i>sulica</i> of Knutson et al. 1975: 323 (in part). <i>Baccha</i> (<i>Allobaccha</i>) <i>tripartita</i> of Knutson et al. 1975: 323. <i>Baccha papuana</i> Hull, unpublished of Greve and Ismay 1983: 74. <i>Episyrphus</i> (<i>Asiobaccha</i>) <i>sulica</i> of Thompson and Vockeroth 1989: 443 (in part). <i>Episyrphus</i> (<i>Asiobaccha</i>) <i>tripartita</i> of Thompson and Vockeroth 1989: 443. <i>Allobaccha flavipes</i> of Thompson and Vockeroth 1989: 441. <i>Allobaccha papuana</i> (Hull, unpublished name) of Rojo et al. 2003: 20. <i>Differential diagnosis</i> This species has a yellow face (Figure 11f), the alula is bare, narrower than costal cell, the wing is partly bare basally (Figure 14d), and it does not have a mesonotal fringe. The abdominal pattern is identical to that of two other species, <i>A. bicolor</i> and <i>A. loriae</i>, with tergum 4 entirely black (Figure 11c, d), but differs by the wing microtrichia as stated in the key. <i>Asiobaccha bicolor</i> and <i>A. tripartita</i> differ from <i>A. loriae</i> by having the wing bare basally, easy to distinguish because both always have the cell CuP and anal lobe partly bare basally (Figure 14c, d); sometimes <i>A. loriae</i> specimens may have very small bare areas in the cell BM but the cell CuP and the anal lobe are always microtrichose. On the other hand, <i>A. tripartita</i> has a more extensive microtrichia and bare basal areas of the wing are on basal 1/3 <i>–</i> 1/2 cell BM only (Figure 14d), while <i>A. bicolor</i> has the cell R bare on basal 1/3 <i>–</i> 1/2 and the cell BM bare on basal 1/2 or more (Figure 14c). <i>Variation.</i> Wing microtrichia is a bit less variable in this species: the bare area of cell BM varies between basal 1/3 to basal 1/2. Furthermore, a few specimens have costal cell bare very basally. The specimen from New Ireland has the scutum entirely yelloworange. Wing colouration varies from very dark to almost hyaline. <i>Length (N = 4).</i> Body, 11.0 <i>–</i> 13.0 (12.3) mm; wing, 10.0 <i>–</i> 12.0 (11.1) mm. <i>Biology</i> Greve and Ismay (1983) mentioned one species of <i>Baccha</i> with the following information: <i>‘</i> Ex <i>Tiracola</i> (Lep., Noctuidae) larva, Brevaturu, N.P., August, 1971 <i>’</i>. The species mentioned is <i>Baccha</i> ? [sic] <i>papuana</i> Hull. This record is compiled by Rojo et al. (2003) but with a different name, <i>Allobaccha papuana</i> (Hull, unpublished name). In the opinion of this author, this species might refer to the unpublished name of <i>Baccha papuana</i> Hull that appears in two specimens from Mafulu kept at the BMNH and CNC, although there is no information in the original labels about <i>Tiracola</i>. After studying the work of Greve and Ismay (1983), this citation may refer to a larva of a syrphid species parasiting a larva of <i>Tiracola</i> Moore, 1881 (Lepidoptera: Noctuidae) from Barevaturu N.P. in Papua New Guinea. However, there is no species of <i>Asiobaccha</i> or <i>Baccha</i> that parasitises butterfly larvae. Consequently, this record by Greve and Ismay (1983) may not refer to a species of <i>Baccha, Allobaccha</i> or <i>Asiobaccha</i>, or, on the other hand, it might indicate that the larva of <i>Baccha</i> was preying on the <i>Tiracola</i> larva and the adult emerged from a puparium. <i>Geographical distribution</i> Kai Islands, Misool Island, New Guinea Island, New Ireland Island and Woodlark Island. <i>Type locality</i> Indonesia: West Papua Province, Raja Ampat Regency, Misool Island (Mysol), between 01°40 <i>’</i> and 02°04 <i>’</i> S, and between 129°43 <i>’</i> and 130°26 <i>’</i> E. <i>Material examined</i> <i>Type material.</i> <i>Holotype</i>, male, deposited in The Natural History Museum, formerly British Museum (Natural History) (London, United Kingdom) and labelled: <i>‘W’</i> [round, handwritten] <i>‘Mysol</i>. // <i>Wallace</i>. <i>’ ‘ Baccha</i> // <i>tripartita</i> // <i>Wlk</i>. <i>’</i> [on the reverse of previous label, handwritten] <i>‘tripartita’</i> [handwritten] <i>‘</i> HOLOTYPE // <i>Asiobaccha</i> // <i>tripartita</i> // (Walker) // det. X. Mengual 20 <i>14’</i> [red]. <b> <i>Type material of</i> Baccha sulica <i>Austen, 1893</i> <i>.</i> <i>Syntype</i>, female, deposited in The Natural History Museum (London, UK) and labelled: <i>‘</i> SYN- // TYPE <i>’</i> [round, blue margin] <i>‘</i> Mysol <i>’</i> [round, light blue] <i>‘</i> hl // 124 <i>’</i> [on the reverse of previous label, handwritten] <i>‘ Mysol</i>. // (Wallace.) // 61.124. <i>’ ‘ Baccha</i> // <i>sulica</i> // <i>Aust</i>. <i>’</i> [on the reverse of previous label, handwritten].</b> <b> <i>Type material of</i> Baccha flavipes <i>Doesburg, 1959</i> <i>.</i> <i>Holotype</i>, female, deposited in the Naturalis Biodiversity Center (Leiden, The Netherlands) and labelled: <i>‘</i> HOLOTYPE <i>’</i> [yellow] <i>‘ G.den Hoed // Ifar 12-57’</i> [handwritten] <i>‘</i> Museum Leiden // Collectie // Van Doesburg // rec. 1973 <i>’ ‘ Baccha</i> &amp;female; // <i>flavipes Dobg</i>. // det. v. Doesburg <i>’</i>.</b> <i>Nontype material.</i> INDONESIA: Maluku Prov., Great Kai Island, Mountain Daab, April 1922, T. Mortensen [2&amp;female;, ZMUC]; Papua Prov., 11 km SE of Oerberfaren, Bodem, 100 m, 7 <i>–</i> 17 July 1959, T.C. Maa [&amp;female;, BPBM];..., Wataikwa River, August 1910, A.F.R. Wollaston 1911 <i>–</i> 229 [1&amp;female;, BMNH];..., Araucaria River, SW von Bernhard Camp, 800 m, 6 January 1939, L.J. Toxopeus [1&amp;female;, RMNH]. PAPUA NEW GUINEA: Central Prov., Owen Stanley Range, Goilala, Loloipa, 1 <i>–</i> 15 February 1958, W.W. Brandt [1&amp;male; 1&amp;female;, BPBM; 1&amp;male;, ZFMK, ZFMKDIP 00011948];..., Goilala, Tapini, 975 m, 16 <i>–</i> 25 November 1957, W.W. Brandt [1&amp;male;, BPBM]; Central Prov., Mafulu, 4000 ft., January 1934, L.E. Cheesman [1&amp;female;, BMNH; 1&amp;female;, CNC]; West Sepik Prov., Telefomin District, Eliptamin Valley, 1200 <i>–</i> 1350 m, 19 <i>–</i> 30 June 1959, W.W. Brandt [1&amp;female;, BPBM]; Madang Prov., Finisterre Range, Saidor, Sibog village, 27 May <i>–</i> 5 June 1958, W.W. Brandt [1&amp;female;, BPBM]; Jiwaka Prov., Upper Jimi valley, Tsenga, 1200 m, 15 July 1955, J.L. Gressitt, light trap [1&amp;male;, BPBM]; Morobe Prov., Bulolo Gorge, 800 m, 17 January 1962, G. Monteith [1&amp;male;, BPBM];..., 6 miles NW Lae, Rain forest, 15 m, 9 July 1957, D.E. Hardy [1&amp;male;, ZFMK, ZFMKDIP 00011952];..., Bulolo, Manki Logging area, 2 June 1983, H. Roberts [1&amp;male;, BMNH] (specimen photographed);..., Wau, 6 <i>–</i> 14 March 1974, H. Hippa, P.T.Lehtinen [1&amp;female;, MZH];..., Wau, April 1968, M. Sedlacek [1sp., BPBM];..., Wau, Nakata Range, 4700 ft., 9 June 1968, J.W. Boyes [1&amp;male;, CNC]; Milne Bay Prov., Woodlark Island, Kulumadau Hill, 4 <i>–</i> 9 March 1957, W.W. Brandt [1&amp;male; 1&amp;female;, BPBM; 1&amp;female;, ZFMK, ZFMKDIP 00011950];..., 16 March 1957,... [1&amp;female;, ZFMK, ZFMKDIP 00011953]; New Ireland Prov., Danu, Kalili Bay, 3 April 1962, Noona Dan Exp. 61 <i>–</i> 62 [1&amp;male;, ZMUC];..., 30 April 1962,... [1&amp;female;, ZMUC]. <i>Remarks</i> Austen (1893) described <i>Baccha sulica</i> based on two female specimens, but only the female from Sula Island had Walker <i>’</i> s handwriting. Austen (1893) stated that he found the type of this species, but he could not find if it was already described. In my opinion, he decided to describe a new species based on Walker <i>’</i> s handwriting. The most interesting fact is that Austen (1893) used the same argument to separate <i>sulica</i> from <i>Baccha moluccana</i> as to separate <i>bicolor</i> from <i>moluccana</i>, but he did not compare his two new species. In the BMNH there are the two syntypes of <i>Baccha sulica</i>, but one belongs to <i>A. bicolor</i> (specimen from Sula Island and Walker <i>’</i> s handwriting) and the other syntype belongs to <i>A. tripartita</i> (specimen from Misool). Specimens from the New Ireland at ZMUC have a manuscript name given by P.H. van Doesburg, <i>Baccha pleuralis</i>, never published. Specimens from Mafulu have a manuscript name given by F. M. Hull, <i>Baccha papuana</i>, again never published. None of these three names is available. As already mentioned, <i>A. tripartita</i> might belong to a widespread taxon and might include also <i>A. loriae</i> and <i>A. bicolor</i> (see Remarks under <i>A. bicolor</i> and <i>A. loriae</i>). There are specimens of <i>A. tripartita</i> and <i>A. bicolor</i> from the same localities, but, more interestingly, specimens of <i>bicolor</i> and <i>tripartita</i> from New Ireland Island have an orange scutum. Both females from Great Kai Island also have the scutum orange.
description: Published as part of <i>Mengual, Ximo, 2016, A taxonomic revision of the genus Asiobaccha Violovitsh (Diptera: Syrphidae), pp. 2585-2645 in Journal of Natural History 50</i> on pages 2632-2634, DOI: 10.1080/00222933.2016.1206634, http://zenodo.org/record/3994636
description: {"references": ["Walker F. 1861. Catalogue of the dipterous insects collected in Batchian, Kaisaa and Makian, and at Tidon in Celebes, by Mr. A. R. Wallace, with descriptions of new species. J Proc Linn Soc Zool. 5: 270 - 303.", "Schiner IR. 1868. Diptera. In: von Wullerstorf-Urbair B, editor. Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wullerstorf-Urbair. Zoologischer Theil, 2 Band. Vienna: Carl Gerold ' s Sohn.", "Bigot JMF. 1892. Catalogue of the Diptera of the Oriental Region. Part II. J Asiat Soc Bengal. 61: 133 - 178.", "van der Wulp FM. 1896. Catalogue of the described Diptera from south Asia. The Hague: M. Nijhoff.", "Kertesz K. 1910. Catalogus dipterorum hucusque descriptorum, vol. VII. Budapestini [= Budapest]: G. Engelmann.", "Doesburg PJ van. 1959. Passalidae (Col.) en Syrphidae (Dipt.) van Ned. Nieuw-Guinea. Entomol Ber. 19: 231 - 235.", "Austen EE. 1893. Descriptions of new species of dipterous insects of the family Syrphidae in the collection of the British Museum, with notes on species described by the late Francis Walker. - Part I. Bacchini and Brachyopini. Proc Zool Soc Lond. 1893: 132 - 164.", "Knutson LV, Thompson FC, Vockeroth JR. 1975. Family Syrphidae. In: Delfinado MD, Hardy DE, editors. A catalog of the Diptera of the Oriental Region, Volume II, Suborder Brachycera through Division Aschiza, Suborder Cyclorrhapha. Honolulu: The University Press of Hawaii; p. 306 - 373.", "Greve JE van S, Ismay JW. 1983. Crop insect survey of Papua New Guinea from July 1 st 1969 to December 31 st 1978. P N G Agric J. 32: 1 - 120.", "Thompson FC, Vockeroth JR. 1989. Family Syrphidae. In: Evenhuis NL, editor. Catalog of the Diptera of the Australasian and Oceanian Regions. Hawaii: Bishop Museum Special Publication 86; p. 437 - 458.", "Rojo S, Gilbert F, Marcos-Garcia MA, Nieto JM, Mier MP. 2003. A world review of predatory hoverflies (Diptera, Syrphidae: Syrphinae) and their prey. Alicante: CIBIO Ediciones."]}