Petrarctus veliger Holthuis 2002, n. sp.
Creators: name: Holthuis, Lipke B.
Publisher: name: Zenodo
Year: 2002
Subjects: Biodiversity
Taxonomy
Animalia
Petrarctus
Petrarctus veliger

description: <i>Petrarctus veliger</i> n. sp. (Figs 15-17) TYPE MATERIAL. — Holotype: 19 mm, RV <i>Anton Bruun</i>, cruise 1, stn 38 (USNM); paratypes: 1 18 mm, 1 ov. 17 mm (ZRC 2001.0326). TYPE LOCALITY. — Andaman Sea, off South Burma, 14°07'N, 97°05'E, 69- 73 m. ETYMOLOGY. — The specific name <i>veliger</i> is Latin, meaning sail-bearer, in reference to the very high cardiac and gastric teeth towering above the body. MATERIAL EXAMINED. — <b>Andaman Sea.</b> Off South Burma, IIOE, RV <i>Anton Bruun</i>, cruise 1, stn 38, 14°07'N, 97°05'E, trawl, 69-73 m, 30.III.1963, 1 holotype, 19 mm (USNM). <b>Philippines.</b> Bohol, Balicasaq Id, off Panglao Id, <i>c.</i> 200-300 m, tangle nets, XII.2000, from local fishermen, 1 18 mm, 1 ov. 17 mm (ZRC 2001.0326). DISTRIBUTION AND HABITAT. — There are only two depth records for this species: 69-73 m (Andaman Sea) and <i>c.</i> 200-300 m (Philippines). So far the species is only known from the types. DESCRIPTION The body is very robust, heavy and rough. The tubercles are quite high and the grooves and depressions deep. The rostrum is bluntly rounded and bears a short and blunt tubercle, which is about as large as the tubercle found on the extreme inner part of the ophthalmic somite. There is no pregastric tooth, but the gastric tooth is very high and triangular ending in a single sharp point. The posterior surface of the gastric tooth is flattened and delimited by two curved rows of three or four tubercles, between which one or two smaller tubercles may be found. The posterior part of the gastric tooth is high and separated from the anterior part of the cardiac tooth by the cervical groove, which here forms a very narrow slit and curves up over the postrostral ridge. The cardiac tooth is even much higher and larger than the gastric, but of about the same shape. On its posterior margin it bears two rows of four or five distinct tubercles; the lateral surface of the cardiac tooth bears a few tubercles. On either side of the posterostral carina the cervical groove, which is very narrow on the ridge, strongly widens laterally forming a large smooth area between the postrostral and branchial ridges. The anterior submedian ridge lies close to the anterior postrostral ridge and ends in a distinct tubercle, which reaches about as far forward as the gastric tooth; it is followed by some smaller tubercles, some of which are interconnected. The intermediate row consists of four tubercles, the anterior of which jots out into the cervical groove; one or two tubercles are present between the anterior part of the intermediate row and the posterior branchial carina. The space between the anterior postrostral and the anterior branchial ridges shows very few tubercles apart from a transverse row of very small tubercles behind the anterior margin. The branchial carina is widely interrupted by the cervical groove, the gap showing a distinct tubercle. The anterior branchial carina ends in the usual two teeth on the inner orbital margin; both teeth are sharply angled. Behind the posterior tooth the carina shows some three low tubercles. The orbital margin is finely tuberculate and between it and the branchial carina another tuberculated ridge is present. There are two postorbital tubercles. The intercervical ridge is covered by about 10 small, rather high tubercles. The posterior branchial carina ends anteriorly in a strong blunt tooth, behind which the carina bears two slightly diverging rows of tubercles; the inner row has three tubercles, the last of which being elongate and ridge-like; the outer row consists of five or six distinct tubercles. The anterior tooth of the lateral margin of the carapace is strong and followed by one or two more or less distinct anterolateral teeth. The anterior of the mediolateral teeth is triangular, the two or three following are rounded. The posterolateral margin bears five or six strong teeth which show some grooves; it ends in a group of tubercles. The marginal groove FIG. 16.— <i>Petrarctus veliger</i> n. gen.,n.sp.,Andaman Sea off Burma, holotype carapace length 19 mm (USNM); <b>A</b>, dorsal view; <b>B</b>, lateral view. along the posterior margin of the carapace is quite distinct; before and behind it is a single row of tubercles. The posterior margin of the carapace has a blunt triangular median incision. The first abdominal somite is entirely smooth and has no transverse groove, except for an indication of one near the base of the pleura. Also the longitudinal grooves are absent or only very vaguely indicated. The posterior margin of the somite shows a distinct blunt triangular median incision. Abdominal somites II to V have a distinct median longitudinal carina. The carina of somite II is lowest and shows a central groove in all but the anteriormost part; its lateral margin is lobulated. The median carina of somite III is highest, it is bluntly triangular and somewhat produced posteriorly, it slightly overhangs the posterior margin; it has an indistinct and interrupted median groove dorsally, and tubercles and short grooves on its lateral surface. The carinae of somites IV and V are lower, they are also slightly produced posteriorly. The tergites of somites II to V show a rather wide transverse median groove with a row of large tubercles before it, and behind it a broad transverse ridge, which in somites II and III has faint tubercles and is smooth in somites IV and V. The pleura of somite I have tubercles on the outer surface and three incisions in the lateral margin. FIG. 17. — <i>Petrarctus veliger</i> n. gen., n. sp., Andaman Sea off Burma, holotype carapace length 19 mm (USNM); <b>A -E</b>, pereiopods 1 to 5. The pleura of the other abdominal somites end in a blunt top. Those of somite II show two wide transverse grooves, separated by rows of flat tubercles. In somites III to V there is only one transverse groove and two ridges; before the anterior ridge the surface of the pleuron is smooth. The sixth somite has a continuous transverse groove with a row of tubercles before it and three tubercles on the posterior margin. The hard part of the telson shows four low and rounded tubercles: two anterior in the median area, and two more behind and more lateral. The teeth at the end of the hard part are placed in one line, the outer being widest. The ophthalmic somite shows four distinct blunt tubercles, one on either side of the rostrum and one laterally. The anterior margin of the antennular somite shows a strong triangular tooth in each half. The sixth (last) segment of the antenna ends in six teeth. The outer is wide and blunt. The following four are narrow and taper gradually towards the top. The inner tooth is short and triangular and has a small incision at its base. The fifth segment is short and ends in two small teeth, the upper of which carries a carina. The fourth segment has a distinct oblique dorsal carina, which is somewhat curved at its base and carries a tubercle there. The carina traverses the segment in its middle and the outer and inner halves of the dorsal surface are of about equal width, they are deeply concave; there is no additional carina or row of tubercles in the outer half, sometimes there are a few scattered small tubercles. The outer margin of the fourth antennal segment bears two or three distinct teeth (the apical tooth of the segment excepted), sometimes an additional minute tooth is present distally. The anterior margin of the fourth segment bears a strong teeth mediad of the articulation with the fifth segment. The margin between this articulation and the apex of the segment bears one large tooth near the articulation and four smaller acute teeth on the rest of the margin. The anterior margin of the epistome is triangularly incised in the middle, the two halves are convex. The first pereiopod is robust. The dactylus is almost as long as the propodus. The surface of propodus and carpus is smooth with at most a few minor depressions. The outer surface of the merus has a wide ventral groove proximally and some depressions in the rest of the surface which thereby becomes rather rough. The second leg has the dactylus somewhat longer than in the other legs; dactylus, propodus and carpus are naked; the merus has a short dorsal hairy fringe and is rather roughly pitted. In P.3 the dactylus has a short dorsal hairy fringe, while the propodus has distinct dorsal and ventral fringes. The carpus has a dorsal area of hairs and does not carry a spur. P.4 and P.5 have hairs on the dorsal surface of the dactylus and the distal part of the propodus. The merus of P.2 to P.5 has dorsal hairs which are placed more densely in the posterior than in the anterior legs. A groove in the dorsal part of the outer surface of the merus, if present, is very inconspicuous. The anterior end of the sternum is U-shapedly concave with a median incision flanked on each side by an inconspicuous tubercle. The anterior margin is slightly swollen. On either of the anterolateral teeth there is a V-shaped carina, which narrows anteriorly and ends abruptly some distance behind the top of the anterolateral tooth. The central part of the sternum is deeply concave and somewhat pitted; at the borders of the sternites it is somewhat sunken. In the male specimen a small median tubercle is present on somites II to V; in the female only the one on the fifth somite is discernable. The elevated sides of the sternum along the bases of the legs show an intricate pattern of rounded knobs and grooves. The posterior margin of the sternum is straight. <i>Size</i> The carapace length of the examined specimens lies between 17 and 19 mm. The ovigerous female has cl. 17 mm. <i>Colour</i> In the preserved specimens almost all traces of the coloration have disappeared, except for the specimens from the Philippines, which show a very indistinct darkish rounded median spot on the smooth anterior part of the first abdominal somite. REMARKS The species is only known from the three specimens cited above. No details are known about colour, habitat, variation, etc. <i>P. veliger</i> n. sp. is most closely related to <i>P. brevicornis</i> n. comb., but can be distinguished by the very high cardiac and gastric teeth, the presence of four tubercles on the ophthalmic somite, the small number of lateral teeth on the fourth antennal segment, etc. <i>Petrarctus demani</i> (Holthuis, 1946) n. comb. (Figs 18; 19) <i>Scyllarus demani</i> Holthuis, 1946: 91, pl. 7 fig. d, pl. 8 fig. b, pl. 9 fig. a. — Burukovsky 1974: 106; 1983: 149. — Ritz 1977: 229, 231, figs 1-6. — Phillips <i>et al.</i> 1980: 69; 1981: 418, 422. — Barnett <i>et al.</i> 1986: 599, fig. 2. — Sekiguchi 1986a: 1290; 1987a: 331; 1987b: 415, 417; 1989b: 457; 1990a: 113; 1992: 212. — Barnett 1989: 124, fig. 1. — Phillips &amp; McWilliam 1989: 357, 358. — Ito &amp; Lucas 1990: 144, figs 1- 10. — Jones 1990: 181. — Sheehy, 1990: 618. — McWilliam <i>et al.</i> 1995: 564. — Burton 1996: 499, 503, figs 14-16. <i>Scyllarus tuberculatus –</i> Nobili 1903: 12. — De Man 1924: 53, fig. 18 [non <i>Arctus tuberculatus</i> Bate, 1888 = <i>Petrarctus rugosus</i> (H. Milne Edwards, 1837) n. comb.]. TYPE MATERIAL. — Holotype:, H. Van der Horst leg., 1912 (ZMA). TYPE LOCALITY. — Indonesia. Banka Id, East of Sumatra. MATERIAL EXAMINED. — <b>Singapore.</b> Received 31.VII.1902, E. Deschamps leg., 1 13 mm, 1 14 mm, 4 juv. 7-8 mm (USNM). <b>Sarawak.</b> Borneo, W of Tandjong Sirik, RV <i>Lanessan</i>, 2°40'N, 110°39'E- 2°38'N, 110°29'E, A. Krempf No. 35, 5.IX.1926, 1 24 mm (MNHN-Pa 795). <b>Indonesia.</b> Banka Id, East of Sumatra, 1912, H. Van der Horst leg., 1 holotype (ZMA). — Jedan, Aru Ids, dredged, 10.II.1907, P. N. Van Kampen leg., 1 19 mm (ZMA). — Off west coast of Wasir Id, Wokam, Aru Ids, Mariel King Memorial Expedition, stn AWI/HS, 5°30'S, 139°12'E, 55-59 m, mud, 15.VI.1970, 1 juv. 6 mm (WAM 251-70). <b>Australia.</b> Western Australia, Shark Bay, 1964, E. Barker leg., 2 19 and 22 mm, 1 non-ov. 21 mm, 4 ov. 22-27 mm (WAM 93-66); 15.X.1964, 1 ov. 26 mm (WAM 95-66); 20.X.1964, 1 ov. 24 mm (WAM 96-66). — RV <i>Lancelin</i>, in prawn trawl, 17.IX.1957, 2 21 and 24 mm, 1 24 mm (WAM 65-58). — FV <i>Bluefin</i>, VII.1963, W. Poole, 1 25 mm, 2 19 and 23 mm (WAM 94-66). — FV <i>Bluefin</i>, trawled, IX.1963, W. &amp; W. Poole, 1 25 mm (WAM 325- 64). — IV.1957, G. K. Bowen leg., 1 20 mm, 1 18 mm (WAM 327-64). — In prawn trawl, 7.XI.1970, D. A. Ritz, 1 24 mm, 2 ov. 25 and 26 mm (WAM 1-71). — FRV <i>Péron</i>, haul 1, 1.III.1962, 2 ov. 25 and 27 mm (WAM 319-64). — FRV <i>Péron</i>, haul 5, 2.III.1962, 1 20 mm (WAM 321-66). — FRV <i>Péron</i>, haul 105, 14.VI.1962, 1 24 mm (WAM 326-64); VI-VII.1961, Poole brothers, 2 19 and 23 mm (WAM 330-64). — RV <i>Lancelin</i>, trawled, R. McKay, 1 19 mm (WAM 320-64). — CSIRO, Danish seine, 1 25 mm, 1 juv. 13 mm (AM-P12482). — Hopeless Reach, FRV <i>Péron</i>, 25°40'S, 113°45'E, 1.X.1965, 3 ov. 21-25 mm (WAM 239-700). — Denham Sound, Shark Bay, <i>c.</i> 25°35'S, 113°15'E, trawl, 2-11 m, VII. 1955, 1 20 mm, 1 non-ov. 26 mm, 2 ov. 24 and 27 mm (WAM 333-64 and 166/7-57). — Near Homestead, Shark Bay, FV <i>Davena</i>, Honolulu dredge, 11 m, VI. 1960, 1 22 mm (WAM 332-64). — Carnarvon, 16.V.1979, J. C. Miquel leg., 4 18- 22 mm, 1 23 mm (RMNH D 32258). — 40 miles SW of Carnarvon, trawled, VI.1960, A. Snell, 1 22 mm (WAM 328-64). — Exmouth Gulf or Shark Bay, FRV <i>Péron</i>, 1960, R. McKay, 1 17 mm (WAM 329-64). — Between Shark Bay and Onslow, trawled, 1966, W. &amp; W. Poole, 1 juv. 5 mm (WAM 236-70). East end of Mary Ann Passage, off Onslow, FV <i>Davena</i>, dredged, 13 m, 15.V.1960, B. R. Wilson, 1 20 mm (WAM 324-64). — SE of Rosemary Id, Dampier Archipelago, dredged, 5-9 m, sand and shells, 26.VIII.1961, G. W. Kendrick &amp; B. R. Wilson, 1 16 mm (WAM 331-64). — NE of Mallus Id, FV <i>Davena</i>, Honolulu dredge, 18 m, 31.V.1960, R. D. Royce, 1 12 mm (WAM 394-64). — Off Gantheaume Point near Broome, dredged, 7 m, VIII.1929, 1 ov. 21 mm (AM). — Near entrance of Roebuck Bay, dredged, 9-15 m, <i>Lithothamnion</i> reef bottom, 26.IX.1929, 1 23 mm (AM). NW Coast. FV <i>Dorothea</i>, <i>c.</i> 21.5 miles ENE of Troughton Id, 13°45'S, 126°09'E, 59 m, mud, 23.X.1962, R. W. George, 1 17 mm (WAM 323- 64). Northern Territory. Darwin, 20 miles 320°, 4.IX.1965, E. Barker, 1 24 mm (WAM 101-66). — Darwin, trawled, IX.1975, E. Barker, 1 ov. 25 mm (WAM 231-70). — Chambers Bay, Darwin, FV <i>Paxia</i>, 13 m, 7.XI.1959, V. Wells, 1 ov. 25 mm (WAM 335-64). Queensland. SE corner of Gulf of Carpentaria, FV <i>Rama</i>, stn 1527, up to 22 m, XI.1964, R. W. George, 1 21 mm (WAM 98-66); stn 1540, up to 22 m, 3 ov. 23-26 mm (WAM 97-66); stn 1541, up to 22 m, 1 22 mm (WAM 99-66); stn 1542, 1 ov. 24 mm (WAM 100-66). — Trawled, 21 m, sandy mud bottom, 20.II.1983, C. Jones, 1 24 mm (RMNH D 39342). — Trawled, 15 m, soft muddy bottom, 16.III.1983, C. Jones, 1 26 mm (RMNH D 49591). — Wellesley Ids, 5 m, 13.II.1983, C. Jones, 1 14 mm (RMNH D 39341). — Off Karumba, trawled, less than 26 m, XII.1963, J. C. Yaldwyn, 2 19 and 20 mm, 1 non-ov. 23 mm, 1 ov. 21 mm (RMNH D 49594). NE Gulf of Carpentaria. Trawled, 14 m, soft muddy bottom, 15.III.1983, C. Jones, 1 ov. 25 mm (RMNH D 49592). East coast of Queensland. Off Cairns, about 16°55'S, 145°46'E, trawl, 24 m, soft muddy bottom, 16.V.1984, C. Jones, 1 24 mm, 1 22 mm (RMNH D 49593). — Off Cairns, trawled, 26 m, soft muddy bottom, 18.V.1984, C. Jones, 1 25 mm C B (RMNH D 49594). — Between Cape Gloucester (about 20°04'S, 148°27'E) and Cape Cleveland (19°11'S, 147°01'E), RV <i>Challenge</i>, CSIRO, trawled, 18-46 m, K. de Witte, 1 24 mm (AM P.12992). — 7 miles NNE of Bowen, RV <i>Endeavour</i>, stn E 3098, 20°01'S, 148°14'E, 29 m, 2 19 and 24 mm (AM P.3518). — Bowen Harbour, Port Denison, about 20°02'S, 148°15'E, F. H. Rainford, 1 juv. 11 mm (AM P.6017). — Between Hayman Id (20°03'S, 148°53'E) and Eshelby Id (20°02'S, 148°37'E), Cumberland Group, RV <i>Challenge</i>, trawl, 37-46 m, K. de Witte, 1 ov. 18 mm (AM P.12988). — SE of Mackay, 21°09'S, 149°12'E, dredged, 24.II.1964, W. Goode, 1 ov. 24 mm (WAM 334-64). — Great Sandy Id (= Fraser Id), Platypus Bay, RV <i>Endeavour</i>, stn E 3111, 25°15'S, 153°10'E, 9-16 m, 1 23 mm, 1 24 mm (AM). — Platypus Bay, RV <i>Endeavour</i>, 28.VII.1910, 2 23 and 25 mm (AM P.3525). — Off Tin Can Bay, 25°49'S, 153°01'E, 46 m, P. T. Garrard, 1 24 mm (AM). — Tin Can Bay, S end of Great Sandy Str., prawn trawl, P. T. Garrard, 1 27 mm (AM P.12990). — Moreton Bay, II.1967, A. J. Bruce, 2 22 and 23 mm, smallest ov. (RMNH D 23379). DISTRIBUTION. — The species has been found in several localities in the Malay Archipelago (Sarawak, Singapore, Banka and the Aru Ids), and is taken in considerable numbers along the coasts of Western Australia (from Shark Bay north), Northern Territory, and Queensland (from Moreton Bay north). Nobili (1903: 12) reported it from Singapore, incorrectly assigning it to <i>Scyllarus tuberculatus</i> (Bate, 1888). De Man (1924) brought a specimen from Banka to the same species; Holthuis (1946) based his new species on De Man's (1924) material, so that Banka, E of Sumatra, is the type locality. Ritz (1977) described phyllosoma stages 1 to 6 from Shark Bay, from the Gulf of Carpentaria and Moreton Bay; he had also described adult material from Shark Bay. Jones (1990) listed many specimens from Shark Bay (Homestead, Carnarvon and SW of it, Denham Sound, Hopeless Reach, between Shark Bay and Onslow, and material originating either from Shark Bay or Exmouth Gulf), all of this material has been examined for the present paper. Barnett (1989) mentioned the species from N Queensland, and Burton (1996) studied the sperm of material from the east coast of Queensland. McWilliam <i>et al.</i> (1995) gave the Australian range of the species as Western Australia to S Queensland, including Torres Strait, from where the species had so far not been reported. Barnett <i>et al.</i> (1986) and Ito &amp; Lucas (1990) reported larvae from off Townsville, Queensland at about 19°S, 147°E. Phillips &amp; McWilliam (1989) reported a final (gilled) phyllosoma stage of this species from 20 km off the SW coast of Oahu, Hawaiian Ids. HABITAT. — The species has been reported from depths between 5 and 59 m; more than two thirds of the records are from 10 to 25 m. The species was found on bottoms described as: soft mud, sandy mud, mud, sand and shells, lithothamnion reef (in this order of frequency). DESCRIPTION The rostrum is not very narrow, the anterior margin is convex or bilobed, it is constricted behind the top. The dorsal surface bears a distinct rostral tooth. Apart from the rostral tooth there are two more teeth in the median line of the carapace. The cardiac tooth is very high, single or twotopped and overhangs the cervical groove; behind it there is a row of four to seven double tubercles. The gastric tooth ends in a single blunt point, it is likewise strong and forwards directed; behind it is a double row of three or four tubercles, the posterior pair being separated from the anterior by a distinct incision so that it seems as if two teeth are placed here, one right behind the other. A few tubercles are seen behind the rostral tooth. The branchial carina is deeply and rather narrowly interrupted by the cervical groove; there is no tubercle in the gap, but two tubercles are placed in the groove to the inner side of the gap. Anteriorly the branchial carina ends in two blunt teeth that are placed, one behind the other, on the inner orbital margin.
description: Published as part of <i>Holthuis, Lipke B., 2002, The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae), pp. 499-683 in Zoosystema 24 (3)</i> on pages 541-551, DOI: 10.5281/zenodo.4689240
description: {"references": ["HOLTHUIS L. B. 1946. - The Stenopodidae, Nephropsidae, Scyllaridae and Palinuridae. The Decapoda Macrura of the Snellius Expedition. I. Biological results of the Snellius Expedition. XIV. Temminckia 7: 1 - 178, text-figs 1, 2, pls 1 - 11.", "BURUKOVSKY R. N. 1974. - Keys for the Identification of Shrimps, Spiny Lobsters and Lobsters. Pishchevaya Promyshlennost Publishers, Moscow, 126 p., 189 figs (in Russian).", "BURUKOVSKY R. N. 1983. - Key to shrimps and lobsters. Russian Translation Series 5, A. A. Balkema, Rotterdam: i-xi, 1 - 174, figs 1 - 189 (English translation of Burukovsky 1974).", "RITZ D. A. 1977. - The larval stages of Scyllarus demani Holthuis, with notes on the larvae of S. sordidus (Stimpson) and S. timidus Holthuis (Decapoda, Palinuridea). Crustaceana 32: 229 - 240, figs 1 - 8.", "BARNETT B. M., HARTWICK R. F. &amp; MILWARD N. E. 1986. - Descriptions of the nisto stage of Scyllarus demani Holthuis, two unidentified Scyllarus species, and the juvenile of Scyllarus martensii Pfeffer (Crustacea: Decapoda: Scyllaridae), reared in the laboratory; and behavioural observations of the nistos of S. demani, S. martensii and Thenus orientalis (Lund). Australian Journal of Marine and Freshwater Research 37 (5): 595 - 608, figs 1 - 5.", "SEKIGUCHI H. 1986 a. - Identification of late-stage phyllosoma larvae of the scyllarid and palinurid lobsters in the Japanese waters. Bulletin of the Japanese Society of Scientific Fisheries 52 (8): 1289 - 1294.", "SEKIGUCHI H. 1987 a. - Life histories of the scyllarid and palinurid lobsters - 11. Aquabiology, Tokyo 9 (5): 330 - 335, figs 46, 47.", "SEKIGUCHI H. 1987 b. - Life histories of the scyllarid and palinurid lobsters - 12. Aquabiology, Tokyo 9 (6): 415 - 419, figs 48 - 50.", "BARNETT B. M. 1989. - Final-stage phyllosoma larvae of Scyllarus species (Crustacea: Decapoda: Scyllaridae) from shelf waters of the Great Barrier Reef. Invertebrate Taxonomy 3 (2): 123 - 134, figs 1 - 9.", "PHILLIPS B. F. &amp; MCWILLIAM P. S. 1989. - Phyllosoma larvae and the ocean currents off the Hawaiian Islands. Pacific Science 43 (4): 352 - 361, figs 1, 2.", "ITO M. &amp; LUCAS J. S. 1990. - The complete larval development of the scyllarid lobster, Scyllarus demani Holthuis, 1946 (Decapoda, Scyllaridae), in the laboratory. Crustaceana 58 (2): 144 - 167, figs 1 - 10.", "JONES D. S. 1990. - Annotated checklist of marine Decapod Crustacea from Shark Bay, Western Australia, in BERRY P. F., BRADSHAW S. D. &amp; WILSON B. R. (eds), Research in Shark Bay, Report of the France-Australe Bicentenary Expedition Committee. Western Australian Museum, Perth: 169 - 208.", "SHEEHY M. R. J. 1990. - Widespread occurrence of fluorescent morphological lipofucsin in the Crustacean brain. Journal of Crustacean Biology 10 (4): 613 - 622, fig. 1.", "MCWILLIAM P. S., PHILLIPS B. F. &amp; KELLY S. 1995. - Phyllosoma larvae of Scyllarus species (Decapoda, Scyllaridae) from the shelf waters of Australia. Crustaceana 68 (5): 537 - 566, figs 1 - 11.", "BURTON T. E. 1996. - Comparative spermiomorphism and taxonomic implications within the genus Thenus (Decapoda: Scyllaridae). Journal of Submicroscopic Cytology and Pathology 28 (4): 499 - 506, figs 1 - 19.", "NOBILI G. 1903. - Crostacei di Singapore. Bolletino dei Musei di Zoologia ed Anatomia comparata delle R. Universita di Torino 18 (455): 1 - 39, 1 pl.", "DE MAN J. G. 1924. - On a collection of macrurous Decapod Crustacea, chiefly Penaeidae and Alpheidae from the Indian Archipelago. Archiv fur Naturgeschichte 90 (1): 1 - 60, figs 1 - 20.", "BATE C. S. 1888. - Report on the Crustacea Macrura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger, during the Years 1873 - 76 (Zool.). Her Majesty's Stationery Office, London, 24, xc + 942 p., 76 text-figs, 150 pls.", "MILNE EDWARDS H. 1837. - Histoire naturelle des Crustaces, comprenant l'Anatomie, la Physiologie et la Classification de ces Animaux 2. Librairie encyclopedique de Roret, Paris, 532 p.", "PHILLIPS B. F., BROWN P. A., RIMMER D. W. &amp; BRAINE S. J. 1981. - Description, distribution and abundance of late larval stages of the Scyllaridae (Slipper Lobsters) in the south-eastern Indian Ocean. Australian Journal of Marine and Freshwater Research 32: 417 - 437, figs 1 - 14."]}