Anisoscelini Latreille 1829
Creators: name: Leavengood, John M.
name: Brailovksy, Harry
name: Barrera, Ernesto
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Coreidae

description: Key to the genera of Anisoscelini (incl. Leptoscelini)
1. Hind tibia expanded, even if only weakly dilated with a flattened ridge limited to the outer surface [“ Anisoscelini group”] (Figs. 13, 15–18)........................................................................................... 2
1. Hind tibia simple or curved, perhaps quadrate in cross-section, but never dilated with a flattened lanceolate or foliate ridge [“Leptoscelini group”] (Figs. 14, 26–31).................................................................. 12
2. Hind tibia only weakly dilated with a flattened ridge limited to the outer surface (Fig. 46)............................ 3
2’. Hind tibia conspicuously dilated, lanceolate or phylliform, at each side of the leg................................... 5
3. Corium hyaline, except the veins (Figs. 16–17); posterior margin of pronotal disk with a rounded triangular projection on either side of scutellar base (Fig. 47)................................................. Holhymenia Lepeletier & Serville
3’. Corium leathery, never hyaline; posterior margin of pronotal disk with..or without tiny triangular projections at either side of scutellar base........................................................................................ 4
4. Antennal segment I robust, shorter than or subequal to head, non-granulate; scutellum flat; femora ventrally armed with one or two acute subapical spines (Fig. 30).............................................................. Tarpeius Stål
4’. Antennal segment I elongate, about 1.5x longer than head, cylindrical, irregularly granulate; scutellum with a depression near the apex; femora ventrally without acute subapical spine, usually with robust setae resembling a spine (Fig. 13).. Baldus Stål
5. Anterior angles of pronotal disk projecting forward, lateral margins dentate; tylus spine-like (Figs. 32, 48).............................................................................................. Bellamynacoris Brailovsky
5’. Anterior angles of pronotum without such projections; tylus usually obtuse (except Leptoglossus clypealis Heidemann).... 6
6. Body color dull-light-brown to dark-brown or black; never metallic, often less ostentatious species; light color limited to pale spots or stripe on the anterior pronotal disk, and/or a pale mid-corial fascia (zig-zag or straight), sometimes absent; antennal segments without flattened expansions.....................................................................7
6’. Body coloration highly variable, often with at least head or legs reddish to light-yellowish-orange (perhaps as stripes); usually metallic, colorfully ostentatious species; with or without pronotal marking and lacking a mid-corial fascia; antennal segments with or without flattened expansions...................................................................... 8
7. Antennal segment I shorter than preocular length, measurement from anterior border of eye to apex of tylus; rostral segment IV the longest (Fig. 33)........................................................................... Narnia Stål
7’. Antennal segment I longer than preocular length, measurement from anterior border of eye to apex of tylus; rostral segment I the longest (Fig. 18)........................................................... Leptoglossus Guérin-Méneville
8. Area between calli with two large tubercles; outer expansion of hind tibia lanceolate (Fig. 34)........ Onoremia Brailovsky
8’. Area between calli smooth, without tubercle; outer expansion of hind tibia phyliform................................9
9. Outer expansion of hind tibia as wide as the internal expansion; head, pronotum and scutellum metallic green with two yellowish longitudinal vittae extending from the apical jugum to nearly apex of the scutellum (Fig. 15)............... Diactor Perty
9’. Outer expansion of hind tibia always wider than inner expansion; lacking the combination of color above.............. 10
10. Outer expansion of hind tibia extending over most of tibial length and tapering apically; surface partially with green or bluish metallic iridescence (Figs. 1–6)........................................................... Anisoscelis Latreille
10’. Outer expansion of hind tibia short, not reaching the apex of tibia; surface without metallic iridescence (except Chondrocera mirabilis Brailovsky & Barrera)......................................................................... 11
11. Antennal segment I twice as long as the head; hind femur with one row of acute spines (Figs. 7-12)….......... Bitta Osuna
11’. Antennal segment I less than twice as long as the head; hind femur with only two subapical spines (Fig. 35)................................................................................................. Chondrocera Laporte
12. Anterior pronotal calli with two conspicuous spines-like projections, each lateral to midline (Fig. 49); metallic-blue body with a bright-orange mid-corial fascia, alternating orange and blue connexival markings, and a mostly orange central pronotal disc (Fig. 36); known only from Cuba .................................................... Rhytidophthia Brailovsky
12’. Anterior pronotal calli without two conspicuous spines-like projections; body coloration variable; broad geographic range... .................................................................................................. 13
13. Anterior angles of pronotal disk with triangular projections or spines directed anteriorly (or anterolaterally if part of a series of dentate lateral margin) (Figs. 50–52)......................................................................14
13’. Anterior angles of pronotal disk rounded or straight, not projecting anteriorly or spinose, at most with a small tubercle (Dalmatomammurius Brailovsky)....................................................................... 18
14. Abdomen conspicuously expanded; anterolateral margins of pronotal disk moderately emarginated....................15
14’. Abdomen elongate, not expanded, or only weakly rounded laterally; anterolateral margins of pronotal disk not emarginated.....................................................................................................16
15. Tylus acutely projecting beyond and above the juga (Fig. 50); postocular tubercle well developed; antennal segment I elongate, robust, longer, length 1.90–2.21 mm; distance between antenniferous tubercle and anterior border of eye shorter (0.55–0.58 mm) (Fig. 31)................................................................................ Ugnius Stål
15’. Tylus not acutely projecting beyond and above the juga; postocular tubercle not visible; antennal segment I broad, shorter, length 1.04–1.09 mm; distance between antenniferous tubercle and anterior border of eye longer (0.79–0.81 mm) (Fig. 37)........................................................................................... Plunentis Stål
16. Mandibular plate well developed, close to the antenniferous tubercle; pronotal collar laterally with a short, acute, tooth-like tubercle on each side (when viewed dorsally); body elongate (Fig. 38)....................... Kalinckascelis Brailovsky
16’. Mandibular plate flat, not raised; pronotal collar laterally without a short acute spine on each side; body robust.......... 17
17. Pronotal disk piceous with reddish lateral margins, densely pubescent; antennal segments II and IV subequal; buccula slightly raised, evanescent; humeral angles obtuse, truncate, not laterally protruding (Fig. 51); posterolateral margins (behind humeri) of pronotal disk sinuate, not serrate or dentate (Fig. 27).................................... Phthiacnemia Brailovsky
17’. Pronotal disk brown, largely glabrous; antennal segment II longer than IV; buccula quadrate, widely raised; humeral angles laterally protruding, apically with large to medium pointed projection (Fig. 39); posterolateral margins of pronotal disk serrate, even if weakly so (Fig. 52)................................................................... Mamurius Stål
18. Humeral angles of pronotal disk expanded or produced into a long acute spine or conspicuously foliate (Fig. 53–54)...... 19
18’. Humeral angles of pronotal disk usually obtuse, rounded, and sometimes with a small subacute expansion (Leptoscelis Laporte) (Figs. 57, 59, 61)..................................................................................... 23 19. Body robust, larger species (27.80–32.00 mm)............................................................. 20
19’. Body elongate or slightly broad, smaller species (10.50–20.80 mm)............................................ 21
20. Humeral angles well-developed, wing expanded, dorsoventrally flattened and curving upward, coming to an acute spine with serrations anteriorly and posteriorly; anterior pronotal calli with two black tubercles at each side of midline; pronotum with a moderately produced but distinct transverse carina, between and behind the humeral angles; femora ventrally armed; yellowish-orange to dull-brown body, ventrally paler with black discoidal spot at mesopleuron, metapleuron and below each abdominal spiracle (Fig. 22–25, 53–54)................................................................... Malvana Stål
20’. Humeral angles not broadly expanded, but with a laterally projecting spine; pronotal calli without two black tubercles; anterior angles of pronotal disk densely granulate; femora unarmed; shiny orange and black body (Fig. 40)... Sephinioides Brailovsky
21. Humeral angles moderate to foliate, not produced into an acute spine; body always with green, bluish or purple metallic reflections; antennal segment IV conspicuously much longer than II (maximum length of II 5.00 mm, maximum length of IV 7.32 mm) (Fig. 41).................................................................. Malvanoides Brailovsky
21’. Humeral angles produced into a spine; body never with metallic reflections; antennal segment IV not much longer than II... .................................................................................................. 22
22. Pronotal humeral angles with acute, anteriorly projecting spines; body more elongate, parallel-sided; anterolateral margins of pronotal disk almost smooth; head clearly longer than wide; outer margin of antenniferous tubercle smooth (Fig. 42)........................................................................................ Leptopelios Brailovsky
22’. Pronotal humeral angles coming to a broad but sharp point; body robust; anterolateral margins of pronotal disk serrate; head as wide or wider than long; outer margin of antenniferous tubercles serrate (Fig. 43)..................... Coribergia Casini
23. Body always with green, bluish or purple metallic reflections................................................. 24
23’. Body without green, bluish or purple reflections............................................................ 26
24. Metathoracic peritreme undivided, not bilobate (Fig. 58); clavus and corium entirely with green metallic reflections (Figs. 28, 57)................................................................................ Phthiadema Brailovsky
24’. Metathoracic peritreme bilobate (Figs. 60, 62); clavus and corium rather dull or scattered with metallic reflections or entirely with greenish-blue metallic reflections................................................................... 25
25. Clavus and corium rather dull, except for a yellow transverse fascia; dorsal surface of head dull; anterolateral borders of pronotum smooth; abdominal sterna III–VI lateral to midline with rectangular yellowish to reddish-orange maculae (Figs. 26, 55, 59)...................................................................................... Phthia Stål
25’. Clavus and corium entirely with green to bluish reflections, unless with a yellow-orange transverse fascia sometimes expanded to occupy most of the clavus and corium; dorsal surface of head with green to bluish reflections; anterolateral borders of pronotum dentate; abdominal sterna III–VI lateral to midline without yellowish to reddish-orange maculae (Figs. 29, 56, 61)................................................................................... Phthiarella Brailovsky
26. Antennal segment I robust, shorter than total length of head................................................... 27
26’. Antennal segment I elongate, slender, longer than total length of head........................................... 28
27. Antenniferous tubercle with a blunt external spine (Fig. 63); anterior angles of pronotal disk acute, sometimes forming a tubercle; pronotal disk finely punctate, punctures not deeply excavated; tylus clearly raised above juga (Fig. 14)..................................................................................... Dalmatomammurius Brailovsky
27’. Antenniferous tubercle unarmed (Fig. 64); anterior angles of pronotal disk rounded; pronotal disk heavily punctate, punctures deeply excavated; tylus not raised above juga (Fig. 44)........................................... Amblyomia Stål
28. Body elongate, slender; dorsal body coloration dull-brown; antennal segment IV pale yellowish-orange; posterior half of hind femur armed dorsally with two rows of spines (Fig. 45).................................... Leptostellana Brailovsky
28’. Body not slender, relatively broad; dorsal body coloration brown to black, often with contrasting markings of bright colors on the corium; antennal segment IV mostly red to yellow, if darker, then all veins of corium apically orange; posterior half of hind femur unarmed dorsally (Figs. 19–21)..................................................... Leptoscelis Laporte
description: Published as part of Leavengood, John M., Brailovksy, Harry & Barrera, Ernesto, 2024, Studies in the New World tribe Anisoscelini Laporte, 1832 (Heteroptera: Coreidae: Coreinae): updated distributions, nomenclatural changes, and keys to the genera of the tribe and the species of Anisoscelis Latreille, 1829 and Bitta Osuna, 1984, pp. 241-268 in Zootaxa 5415 (2) on pages 262-264, DOI: 10.11646/zootaxa.5415.2.2, http://zenodo.org/record/10692976