Cossula duplexata Davis & Gentili-Poole & Mitter 2008, SP. NOV.
Creators: name: Davis, Steve R.
name: Gentili-Poole, Patricia
name: Mitter, Charles
Publisher: name: Zenodo
Year: 2008
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Cossula
Cossula duplexata

description: COSSULA DUPLEXATA SP. NOV.  (FIGS 43, 81, 82, 103, 143)  
 Male (Fig. 43): Forewing length: 22 mm.   
 Head: Antenna light brown; labial palpus rustybrown dorsally and ventrally; frons rusty-brown; vertex dark rusty-brown to fuscous.   
 Thorax: Patagia same colour as vertex; tegula and dorsum mixed with white, brown and fuscous scales; pleura mostly brown, with some white along base of forewing; venter mostly brown, mixed with creamy white and light brown; a dark brown to fuscous patch below labial palps, ventrally. Legs mostly rustybrown, lighter brown mesally. Forewing a mix of different shades of brown dorsally, mixed with creamy white; several short fuscous striae traversing wing, with few striae along costal and inner margins; an elongate fuscous spot beginning at base of discal cell and extending towards inner angle; terminal patch with outer margin greyish brown and inner margin dark brown to fuscous; a brown to fuscous spot separate from terminal patch between CuA 1 and CuA 2; fringe different shades of brown; forewing mostly brown ventrally, with a mix of cream along wing; costal margin cream and brown banded. Hindwing brown dorsally and ventrally; fringe different shades of brown.   
 Abdomen: Brown to dark brown dorsally and laterally; mostly light brown and cream ventrally; A7–8 light brown and cream.   
 Male genitalia (Figs 81, 82, 103): Valva oblong and rounded apically; length of valva approximately 0.7¥ length of genital capsule; gnathos–uncus expanse approximately 0.03¥ length of genital capsule; sacculus with a lenticular ridge running lengthwise and developing terminally into a blunt spinose process; juxta process elongate and wide; gnathos arms short; gnathos bridge elongate and narrow, broadly downcurved as viewed laterally, subacute terminally and apressed to uncus; uncus elongate and narrow, downcurved with gnathos bridge, with apex truncate and shallowly bifurcate. Aedoeagus becoming wider towards rostellum, as viewed laterally; rostellum short, knob-like and blunt apically; vesica with a dense covering of small spines at distal end where ductus ejaculatorius emerges.   
 Female: Unknown.   
 Distribution: Known only from the type locality, Costa Rica.   
 Holotype: ♂; COSTA RICA: CARTAGO: Moravia de Chirripo, 1100 m: 16.iv.1983, A. M. Chavarria, slide USNM 96036 (INBio).   
 Host: Unknown.   
 Flight period: April (unique record).   
 Etymology: Derived from the Latin suffix ‘-atus’ meaning ‘imitate’ or ‘copy’. Cossula duplexata refers to its similarity to C. duplex.   
 Discussion: This species most closely resembles C. duplex in wing pattern and colour, and an undescribed species from Brazil in male genital morphology. Cossula duplexata differs from duplex in possessing a smaller, fuscous scale patch at the base of the forewing discal cell (Fig. 43). The male genitalia of duplexata differs from the Brazilian species in possessing a more curved, caudal apex of the gnathos and in the distinctly shorter, broader saccular process of the valvae (Fig. 81). In the Brazilian species the gnathos is not curved and the saccular process is long and slender.
description: Published as part of Davis, Steve R., Gentili-Poole, Patricia & Mitter, Charles, 2008, A revision of the Cossulinae of Costa Rica and cladistic analysis of the world species (Lepidoptera: Cossidae), pp. 222-277 in Zoological Journal of the Linnean Society 154 (2) on pages 269-270, DOI: 10.1111/j.1096-3642.2008.00406.x, http://zenodo.org/record/5446371