Toechorychus darienus Tedesco & Aguiar 2013, sp. nov.
Creators: name: Tedesco, Anazélia M.
name: Aguiar, Alexandre P.
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Toechorychus
Toechorychus darienus

description: Toechorychus darienus Tedesco, sp. nov.   
(Figs 30, 78, 92, 162, 167)  
Description. Holotype FEMALE. Fore wing length 6.06 mm.  
 Head. Mandible 1.38 as long as basal width, densely pilose; ventral margin projected as crest; dorsal tooth distinctly longer than ventral tooth. Clypeus 1.51 as wide as high, subquadrate, entirely finely strigulate; apex 1.45 as long as base, truncate; apical margin sharp, medially straight. Supraclypeal area entirely strigulate, also densely punctate, moderately pilose, medially slightly prominent; between antennal foramens with V-shaped carina, medially interrupted; radicle foveolate. Antenna with 25 flagellomeres; white band starting at flagellomere 4, reaching flagellomere 11; flagellum slender; subapical flagellomeres slightly flattened. Supra-antennal area with very faint but distinct median longitudinal carina, medially without longitudinal elevation, near antennal sockets striate; dorsal half medially rugulose. Paraocular area smooth. Vertex rugulose between ocelli, at anterior third with short, longitudinal sulcus, posterior two-thirds smooth; gena and vertex behind ocelli minutely strigulate, sparsely pilose; gena in lateral view at level of dorsal portion of occipital carina narrow, ventrally wide; occipital carina faint, dorsally absent, ventrally slightly projected as crest, reaching hypostomal carina at mandible base or nearly so; hypostomal carina slightly projected as crest; malar space 1.06 as long as basal width of mandible.   
 Mesosoma. Mesosoma in dorsal view 1.80 as long as wide; in lateral view, mesosoma middle width 0.20 mm. Pronotum centrally smooth, latero-ventrally smooth, margin near mesopleuron ventrally corrugated; pronotal swelling finely rugulose; collar dorso-laterally rounded, distinctly swollen, anteriorly smooth; epomia absent. Mesoscutum sparsely pilose, lobes dorsally punctulate, laterally markedly corrugated, central lobe with dense coarse punctures, central lobe without longitudinal sulcus or carina; notauli deeply impressed, posteriorly abruptly curved, U-shaped. Scutoscutellar groove smooth, polished; scutellum 0.98 as long as wide, scarcely punctate; scutellar carina incomplete, restricted approximately to anterior 0.6; postscutellum 0.30 as long as wide; hind margin of metanotum with two lateral teeth. Subalar ridge somewhat elongate; mesopleuron entirely markedly strigate, ventrally strigulate; mesopleural groove corrugated; epicnemial carina stout, dorsally curving forwards; dorsal end of epicnemial carina reaching 0.2 of posterior margin of pronotum, behind epicnemial carina with rounded confused-rugulose area; sternaulus complete, reaching base of mid coxa, markedly sinuate, deeply impressed; markedly corrugated; juxtacoxal carina of mesopleuron stout and long; scrobe deeply impressed, forming pit. Mesothoracic venter strigulate; median portion of posterior transverse carina of mesothoracic venter V-shaped, regularly shaped, without projections. Transverse furrow at base of propodeum laterally 0.09 as long as anterior portion of propodeum, moderately deep and narrow, medially wide, very faintly corrugated, almost smooth; propodeum 1.16 as long as wide medially; anterior margin of propodeum with two faint lateral teeth; anterior portion of propodeum punctulate or with sparse coarse punctures, medially with two posteriorly convergent faint longitudinal carinae; lateral longitudinal carina of propodeum represented by short curved carina; posterior portion of propodeum entirely markedly strigate, medially without distinctly stout longitudinal carina; behind posterior transverse carina with two divergent longitudinal carinae; anterior transverse carina of propodeum complete, stout, medially slightly arched forwards; posterior transverse carina absent, only with lateral slight swellings; propodeal spiracle 1.45 as long as wide; pleural carina of propodeum irregular and faint, obsolescent posteriorly, fused with sculpture of lower division of metapleuron and propodeum; lower division of metapleuron strigulate or sparsely punctate, sparsely pilose, juxtacoxal carina long and stout.   
 Wings. Fore wing: vein 2+3Rs slightly concave; ramellus absent; vein 1M+Rs posteriorly faintly sinuous; vein 1cu-a straight, arising far from vein 1M+Rs base; vein 2Cu 0.46 as long as vein 2cu-a; vein 4Rs 0.93 as long as vein 4M, uniformly slightly convex; cell 1+2Rs 0.72 as high as pterostigma, rectangular; vein 2M approximately as long as vein 3M; vein 3M distinct, 0.64 as long as vein 2M. Hind wing: vein 1Cu 1.33 as long as vein cu-a; vein 2- 1A reaching 0.83 of distance to posterior margin.   
 Legs. Tibia with sparse short bristles; hind coxa globose.   
 Metasoma. T1 0.52 as long as hind femur, with basolateral tooth; anteriorly smooth, posteriorly coriarious; spiracle at anterior 0.54, slightly prominent; postpetiole dorso-anteriorly convex; T2–8 minutely coriarious; almost glabrous; T2 1.09 as long as wide at apex; apex of T2 1.93 as wide as base. Dorsal valve of ovipositor with nodus indistinct; apex of lower valve with series of teeth; notch present.   
 Color. Head and mesosoma yellow and black, metasoma mostly orange. Head: pale yellow (208,133,077); mandible, ventral half of malar space, apex of clypeus, M-shaped mark at ventral half of supraclypeal area, supraantennal area medially, vertex, and occiput, black; scape ventrally pale yellow; pedicel, and flagellum brown; flagellum dorsally entirely white at f5–10 entirely, partially at f4 and f11. Mesosoma: black, except for the following pale yellow marks: propleuron centrally, collar laterally, pronotal swelling, longitudinal narrow mark at posterior 0.9 of central lobe of mesoscutum, tegula, axillary carina, scutellum, postscutellum, large mark extending from subalar ridge to posterior ventral margin of mesopleuron and over epicnecium, dorsal 0.8 of hypoepimeron, mesothoracic venter, upper division of metapleuron, lower division of metapleuron except ventral rounded spot, hind margin of metanotum, and two lateral longitudinal marks at posterior portion of propodeum. Legs: mostly orange (212,147,043); fore coxa pale yellow, posteriorly fulvous; mid coxa pale yellow, laterally with black strip reaching first trochanter; hind coxa pale yellow with lateral black stripes at entire length. Metasoma: mostly orange; T1 dorsally black, laterally and at posterior 0.2, pale yellow, brownish around spiracle; T2–8 yellowish towards apex; S2–3 laterally dark brown (108,062,031).   
 Variation. Mandible basally white. Occipital carina dorsally widely interrupted, completely absent. Mid coxa dorsally black only at apical 0.4. Hind coxa laterally with small black spot. T1 black around spiracle.   
 Comments. Very similar to T. amapaeus sp. nov., from which it can be isolated by having supraclypeal area ventrally with M-shaped mark (vs. with two lateral longitudinal black stripes in T. amapaeus sp. nov.; Fig. 66); lateral black stripes at supraclypeal area reaching antennal foramens, even if medially interrupted (vs. not reaching antennal foramens); malar space ventrally black (vs. completely yellow); occipital carina near mandible black (vs. bright yellow); juxtacoxal carina of mesopleuron stout and long (vs. faint and short); scrobe deeply impressed, forming a pit (vs. very shallow, forming a sulcus); lower division of metapleuron with ventral rounded black spot (vs. entirely yellow; Fig. 13).   
 Male. Unknown.   
 Distribution. Costa Rica and Panama. Record from Heredia and Darién. The type locality is Darién National Park, Darién (7°44’10”N 77°32’50”W) (Fig. 235). These records comprise a range of 3°42’ in latitude.   
 Biology. Collected near Mischocyttarus collarellus Richards (Vespidae, Polistinae) nest, in La Selva Biological Station, Costa Rica.   
 Material examined. Holotype ♀, PANAMA: Darién, Darién National Park, Pirre, Est. Rancho Frio, 17.I.2000, 80 m, R.Cambra & A.Santos leg., Toechorychus, Townes det. (AEIC). Complete, in good shape, except for mesoscutum somewhat damaged by pin. Paratype ♀ from COSTA RICA: Heredia, La Selva Biological Station, 8.VI.1999, E.F.Smith Toechorychus abactus, det. I.Gauld 2001, flying near Mischocyttarus collarellus nest, Nest P. 03B (BMNH).
description: Published as part of Tedesco, Anazélia M. & Aguiar, Alexandre P., 2013,  Phylogeny and Revision of  Toechorychus  Townes (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of thirty-five new species , pp. 1-138 in Zootaxa 3633 (1) on pages 33-35, DOI: 10.11646/zootaxa.3633.1.1, http://zenodo.org/record/5261931