Gomphostemma japanicum
Creators: name: Bongcheewin, Bhanubong
name: Ingrouille, Martin J.
name: Paton, Alan J.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Magnoliopsida
Lamiales
Lamiaceae
Gomphostemma
Gomphostemma japanicum

description: <b>16. Gomphostemma japanicum</b> (<i>3lume</i>) <i>3enth.</i> (Bentham 1833: 630); Prain (1891: 261); Keng (1969: 86, 1958: 322); Murata (1956: 188); Phuong (2000: 111); Walsingham &amp; Bramley (2010: 283); Walsingham (2019: 199). Type: as for <i>Prasium jaoanicum.</i> <i>Prasium jaoanicum</i> Blume (1826: 820). Type: Indonesia, Java, Mont. Gede [-6.5992°, 106.9689°], <i>3lume</i> s.n. (lectotype L (L0003500), selected by Keng (1969); isolectotypes 2 sheets [L0003696, L0003695, L0003698, L0003699]). <i>Prasium phlomoides</i> Reinw. ex Blume (1826: 820). Type: Indonesia, Java, <i>Reinwardt</i> s.n. (lectotype L [L 0003692], selected by Keng (1969); isolectotypes 2 sheets L [L0003693, L0003693]). <i>Gomphostemma lucidum</i> Wall. ex Benth. (Bentham in Wallich 1830b: 12). Type: India, Pundua [Pandua, 23.1165°, 91.5333°], May 1822, <i>de Siloa</i> s.n. in <i>Wall. Cat.</i> 2136.2 (lectotype K [Herb. Benthamianum], selected here; isolectotype K-W [K001113320]). <i>Gomphostemma oblongum</i> Wall. ex Benth. (Bentham in Wallich 1830b: 12). Type: Burma, Tavoy, 12 Aug. 1825, <i>Gomez</i> s.n. in <i>Wall. Cat.</i> 2132 (lectotype K [Herb. Benthamianum, K000301086]; isolectotype K-W [K001113335]). <i>Gomphostemma phlomoides</i> (Reinw. ex Blume) Benth. (Bentham 1833: 629). Type: as for <i>Prasium phlomoides</i>. <i>Gomphostemma philippinarum</i> Benth. (Bentham in A.P. de Candolle 1828: 331). Type: Philippines, Island of Bohol [9.8333°, 122.1665°], 1821, <i>Cuming</i> 1820 (K [Herb. Benthamianum, K000836198] lectotype selected here; isolectotypes C, E [E00301969], G [G00189005], K [Herb. Benthamianum, K000836200]). <i>Gomphostemma oblongum</i> var. <i>helferi</i> Hook.f. (Hooker 1883: 695). Type: Burma, Tenasserim, <i>Helfer</i> 2023 (holotype K [Herb. Hookerianum, K000826319]). <i>Gomphostemma scortechinii</i> Prain (1890: 313). Type: Malaysia, Perak, Gunong Tucok, Sept. 1882, <i>Scortechini</i> 1223 [CAL [CAL0000005131], lectotype selected here; isolectotypes 3 sheets CAL [CAL0000005130, CAL 0000005132, CAL0000005133], K [K00030182], SING). <i>Gomphostemma scortechinii</i> var. <i>helferi</i> Prain (1891: 260). Type: as for <i>G. oblongum</i> var. <i>helferi</i> Hook.f. <i>Gomphostemma rugosum</i> Prain (1891: 262). Type: Burma, Tenasserim, Chapadong [Chappedong] Hill, 23 Feb. 1825, <i>Wall. Cat.</i> 2136.1 (K-W [K001113339], lectotype selected here; isolectotype CAL n.v.). <i>Gomphostemma lucidum</i> Wall. ex Benth. var. <i>canescens</i> Prain (1891: 263). Type: India, Lakhimpur, <i>Griffith</i> 2038 (K [Herb. Hookerianum], lectotype selected here; isolectotypes C, CAL n.v., P [P02061923]). <i>Gomphostemma intermedium</i> Craib (1910: 23). Type: Thailand, Chiang Mai,?oi Sootep, in evergreen cungle, 530 m, 15 July 1909, <i>Kerr</i> 533 (K, lectotype selected here; isolectotypes AB?, BM). <i>Gomphostemma cinereum</i> Elmer (1919: 3086). Type: Philippines, Luzon, Province of Laguna, Los Banos (Mt Maquiling), along stream beds of dense forests half way up the mountain, June – July 1915, <i>Elmer</i> 15836 (K [K000836203], lectotype selected here; isolectotypes 2 sheets BM [BM 000930332, BM 000930333], L [U0226339], NY [00000332], P, US [00119222]). <i>Gomphostemma oblongum</i> var. <i>setosum</i> Ridl. (Ridley 1923a: 633). Type: Malaysia, Selangor, Semangkok Pass [3.6833°, 101.53°], 22 Jan. 1921, <i>Ridley</i> s.n. (holotype K [K000301132]). <i>Gomphostemma lacteum</i> Ridl. (Ridley 1922: 300). Type: Malaysia, Pahang, Pine Tree Hill [3.5165°, 101.6833°], near Fraser Hill, 1220 m [2800 ft], 25 Aug. 1923, <i>3urkill</i> 11053 (holotype K [K000301133]; isotype SING). <i>Gomphostemma lucidum</i> var. <i>intermedium</i> (Craib) C.Y. Wu (1963: 123). Type: as for <i>G. intermedium</i> Craib. <i>Gomphostemma arbusculum</i> C.Y. Wu (1963: 126). Type: China, Yunnan, Lan-tsang, Lan-tsang Hsien [22.3333°, 99.9333°], 1800 m, May 1933, <i>Wang</i> 56253 (holotype PE; isotype A n.v.). <i>Gomphostemma latifolium</i> C.Y. Wu (1963: 125). Type: China, Yunnan, Jinghong, Che-li Hsien, 930 m, Aug. 1936, <i>Wang</i> 53330 (holotype PE; isotype A n.v.). <i>Gomphostemma jaoanicum</i> f. <i>phlomoides</i> (Reinw. ex Blume) Murata (1956: 189). Type: as for <i>Prasium phlomoides</i> Reinw. ex Blume. <i>Gomphostemma jaoanicum</i> var. <i>helferi</i> (Hook.f.) Murata (1956: 189). Type: as for <i>G. oblongum</i> var <i>helferi</i> Hook.f. <i>Gomphostemma jaoanicum</i> subsp. <i>lucidum</i> (Wall. ex Benth.) Murata (1956: 189). Type: as for <i>G. lucidum</i> Wall. ex Benth. <i>Gomphostemma jaoanicum</i> var. <i>rugosum</i> (Prain) Murata (1956: 190). Type: <i>Gomphostemma rugosum</i> Prain. <i>Gomphostemma jaoanicum</i> var. <i>breoidens</i> Murata (1989: 101). Type: Thailand, Chiang Mai,?oi Inthanon, in evergreen forest, 1800 – 2300 m, 3 Oct. 1951, <i>Murata et al.</i> T- 13988 (holotype KYO [KYO00020216]; isotypes BKF [SN029508], KYO [KYO00020215]). Perennial herb 0.3 – 2 m tall. <i>Stems</i> erect, upright, woody at base, obtusely quadrangular with longitudinal grooves, with dense stellate hairs with a 1-celled or multi-celled stalk and sometimes dendroid hairs. <i>Leaoes</i> petiolate, chartaceous or coriaceous; blades oblong-ovate, elliptic, elliptic-oblong, oblong, elliptic-ovate, elliptic-obovate or obovate, 5 – 30 × 3 – 13 cm, apex acute or broadly acuminate, margin entire, subentire, serrate, obtusely serrate or crenate, base cuneate, attenuate or obtuse, upper side tomentose with a mixture of soft hairs, lower side pale green tomentose with a mixture of dense branched hairs, sometimes with robust venation, rugose-like; petioles 10 – 30 mm long, with stellate hairs and/or dendroid hairs. <i>Inflorescence</i> axillary with opposite cymes sessile or sometimes pedunculate, branched then lax or unbranched then congested with axis not visible between flowers, exceeding 20 mm wide when pressed, excluding corolla, inserted at the upper nodes at which leaves are usually present; verticils few – manyflowered, subtended by upper leaves only; pedicels 0 – 10 (– 13) mm long; bracts deciduous, slightly falcate, ovate, narrowly obovate, obovate or spathulate, 3 – 13 × 3 – 5 mm, upper side with shortly simple hairs, lower side with stellate hairs; bracteoles linear, 2 – 10 mm long. <i>Flowering calyx</i> infundibular, 5 – 13 mm long, prominently 10-ridged, outside dense tomentose with stellate hairs with a 1-celled stalk or sessile stellate hairs or short simple hairs, inside with short simple hairs; tube 3 – 10 mm long; lobes triangular, 2 – 3 mm long, apex acute or acuminate. <i>Fruiting calyx</i> broadly infundibular or cupular, 10 – 20 mm long; tube 6 – 13 mm long; lobes 2 – 8 mm long, longer than the tube at maturity or not longer than the tube; apex acute or acuminate, margin flat in cross section. <i>Corolla</i> white, pale yellow, pale pink or scarlet, 23 – 20 (– 30) mm long, throat inflated, abruptly dilated near the throat, outside tomentose with stellate hairs or simple hairs, inside glabrous; tube incurved, 13 – 33 mm long; posterior lip ovate, 3 – 3 mm long, apex emarginate or undulate, margin undulate or entire; anterior lip 3-lobed, spreading beyond posterior lip, 5 – 13 mm long; median lobe tongue-like, spreading, broadly ovate, apex and margin undulate, base constricted. <i>Staminal filaments</i> fleshy, tomentose. <i>Style</i> fleshy, glabrous, shorter than corolla with apex subequally lobed. <i>Nutlets</i> 1 – 2, elliptic or oblong, 2 – 6 × 3 – 2 mm, sparse tomentose or glabrescent at apex, wrinkled. Fig. 1? &amp; 3. <b>DISTRIBUTION.</b> India (Arunachal Pradesh, Assam, Meghalaya, Andaman and Nicobar Islands), Bangladesh, China (Yunnan, Hainan, Guangdong), Burma, Thailand, Vietnam, Laos, Cambodia, Peninsular Malaysia, Malaysian Borneo (Sabah, Sarawak), Brunei, Indonesia (Sumatra, Java, Sulawesi), Philippines (Luzon, Mindanao, Visayas, Sulu Archipelago). Map 2. <b> SPECIMENS EXAMINED. INDIA. Arunachal Pradesh. Nizamghat [28.2665°, 93.5°], <i>Ward</i> 8532 [in parte] (K);</b> Lohit Valley, 28°N 96°33&amp;E, 600 – 900 m [2000 – 3000 ft], 3?ec. 1926, <i>Ward</i> 5689 (K); Lakhimpur, <i>Griffith</i> 2038 (isolectotypes of <i>G. lucidum</i> var. <i>canescens</i>, C, K [Herb. Hookerianum], P); Assam. <i>Griffith</i> 290 (BM), 1012 (BM) &amp; 1830 (K [Herb. Hookerianum]); ibid., <i>Masters</i> s.n. (lectoparatype <i>G. lucidum</i> var. <i>canescens</i>, K [Herb. Hookerianum]); Seelsagar, Borpatta [Barpeta, 26.3333°, 91°], 90 m [300 ft], 13 Oct. 1883, <i>Clarke</i> 20563A (BM); Meghalaya. Jaintea, 1200 m [2000 ft], 19 Oct. 1865, <i>Clarke</i> 3930 (K); Khasia, 1200 – 1300 m [2000 – 3000 ft], <i>Hooker &amp; Thomson</i> s.n. (G, K 2 sheets [Herb. Hookerianum], P); Khasia Hills, <i>Griffith</i> s.n. (L, P); Khasia, Churra, 1200 – 1300 m [2000 – 3000 ft], 13 Aug. 1830; <i>Hooker &amp; Thomson</i> 2058 (K [Herb. Hookerianum]); Khasia, Mammai, 1030 m [3300 ft], 9 May 1886, <i>Clarke</i> 23530 (G); Khasia, Maodadoom, 900 m [3000 ft], 3 Feb. 1886, <i>Clarke</i> 22936 (BM); Cherrapuncee, 1200 m [2000 ft], 12 May 1932, <i>Koelz</i> 29833 (E); <b>ANDAMAN AND NICOBAR ISLANDS</b>. South Andaman, Ranga Chang [11.3833°, 92.53°], 1 April 1891, <i>Prain</i> s.n. (G); South Andaman, Bacacagdah Hill, 18 Aug. 1892, <i>King's Collector</i> s.n. (P); Middle Andamans, camp no. 13, 22 km from Rangat [12.30992°, 92.912°], 23 m, 3 Nov. 1955, <i>3hargaoa &amp; Nooteboom</i> 6311 (CAL, L); Mt Harriet, Port Blair [11.6665°, 92.53°], 20 Sept. 1882, <i>Kunstler</i> 232 (BM, K). <b>BANGLADESH.</b> Sylhet. Pundua, May 1822, <i>de Siloa</i> s.n. in <i>Wall. Cat.</i> 2136.2 (lectotype of <i>G. lucidum</i>, K [Herb. Benthamianum]; isolectotype K-W); <b>UNLOCALISED</b>. <i>Griffith</i> 2038 (C, K [Herb. Hookerianum], P). <b>CHINA.</b> Yunnan. Foo-ning [23.6225°, 103.6252°], Jar-gui, 500 m, 31 May 1920, <i>Wang</i> 89228 (KUN); Hekou, Nanxi [22.6203°, 103.9328°], Machang to Liangzi, 500 m, 6 May 2002, <i>Shui &amp; Chen</i> 20883 (KUN); Kuen-ger, Che-li Hsien [21.9833°, 100.8165°], 1200 m, Oct. 1936, <i>Wang</i> 59289 (KUN); Lan-tsang Hsien [22.3333°, 99.9333°], 1800 m, May 1936, <i>Wang</i> 56253 (holotype of <i>G. arbusculum</i>, PE); Mengla [21.2665°, 101.3833°], 1300 m [3000 ft], <i>Henry</i> 11026 (K); Meng-soong,?ah-meng-lung, Che-li Hsien, 1300 m, Sept. 1936, <i>Wang</i> 55982 (KUN); Ping-bien Hsien [22.9°, 103.6665°], Hwang-tian-bien, 1300 m, 5 Oct. 1939, <i>Wang</i> 82329 (KUN); Szemao [22.5665°, 101.0833°], 1330 m [2300 ft], <i>Henry</i> 12609 (K); Jinghong, Che-li Hsien, 930 m, Aug. 1936, <i>Wang</i> 53330 (holotype of <i>G. latifolium</i>, PE); Hainan. Lam Ko, Cheung Wo and vicinity, 8 Aug. 1925, <i>Tsang</i> 200 (GH); ad pagum Fan lun, territ, indigenarum Lai diatorum, 13 Nov. 1882, <i>Henry</i> 133 (BM); Hong Fa, July 1893, <i>Chinese Collector</i> 262 (K); Lai area, Hung Mo Shan and vicinity, 2 June 1929, <i>Tsang &amp; Fung</i> 225 (GH, K); Ling Shui [18.3036°, 110.0322°] (Ling-tui), Chim Shan, Fan Maan Ts&amp;uen and vicinity, 3 – 20 May 1932, <i>Fung</i> 20110 (GH, K, P); Lingshuixian, Nanqiaoqu Shuangximu zi?acong, 630 m, <i>Anonymous</i> s.n. (AAU); Loktung, 2 July 1936, <i>Lau</i> 25392 (GH, KUN); Po-ting [18.6203°, 109.6992°], 180 m [600 ft], 12 April 1933, <i>How</i> 51890 (GH, P); Tai Pin, foot of Loi Mother Mt, 290 m, 20 July 1933, <i>Gressitt</i> 1062 (BM, E, G, GH); Tamm-chau [19.5113°, 109.3125°], Nodoa and vicinity, 20 July 1925, <i>Tsang</i> 109 (G, K); Tingon [19.6995°, 110.3183°], 31 Oct. 1936, <i>Lau</i> 28123 (KUN 2 sheets); Yaichow, 16 July 1933, <i>Liang</i> 62113 (E, K); Ban-chiou-chian, Che-li Hsien [21.9833°, 100.8165°], 820 m, Oct. 1936, <i>Wang</i> 59523 (KUN); Ching-tung [22.2665°, 100.9°], 1500 m, 30 Oct. 1939, <i>Li</i> 930 (KUN); Guangdong. 28 July 1935, <i>Liang</i> 69858 (IBSC); Fang Ch&amp;eng, T&amp;aan Faan, Kung P&amp;ing Shan and vicinity, 10 – 18 Sept. 1936, <i>Tsang</i> 26823 (C, E, GH, IBSC, K, P). <b>MYANMAR.</b> Tanintharyi. Eastern Tenasserim [11.8°, 99.23°], 30 m, 31 May 1932, <i>Kerr</i> 21623 (BK, BM, K, L); Tenasserim [12.0833°, 99.0165°], <i>Griffith</i> 2021 (K [Herb. Hookerianum]); ibid., <i>Helfer</i> 2021 (K [Herb. Hookerianum], P); ibid., <i>Helfer</i> 2023 (holotype of <i>G. oblongum</i> var. <i>helferi</i>, K [Herb. Hookerianum]); Tavoy, 12 Aug. 1825, <i>Wall. Cat.</i> 2132 (holotype of <i>G. oblongum</i>, K [Herb. Benthamianum]; K-W); Tenasserim, Tavoy, Hill W of Paungdaw Power Station, 12°N 98°30&amp;E, 630 – 690 m [2100 – 2300 ft], Oct. 1961, <i>Keenan et al</i>. 1603 (E); Mon. Chapadong Hill, 23 Feb. 1825, <i>Wall. Cat.</i> 2136.1 (isolectotype of <i>G. rugosum</i>, K-W); Mandalay. Maymyo [22.0333°, 96.2665°], 1110 m [3500 ft], 31 July 1923, <i>Parkinson</i> 1393 (K); Shan. S Shan States, Keng Tung [Keng Tang, 21.3°, 99.63°], 1300 m [3000 ft], June 1909, <i>MacGregor</i> 392 (E). <b>VIETNAM.</b> ?a Nang. Sur de Tourane, June 1909, <i>D'alleizette</i> s.n. (L); Ba Na pres Tourane, 1300 m, 8 July 1923, <i>Poilane</i> 6931 (P); Ravine du Massif Mautaguong de la pres qu&amp; la de Tourane, May – June, <i>Couderc</i> s.n. (P 2 sheets);?ak Lak. Krong Pak, Hoa lo, 23?ec. 1959, <i>Phuong</i> 699 (HN 2 sheets);?ong Nai. ad Gia lau me in prov Bienhoa [10.93°, 106.8165°], Sept. 1869, <i>Pierre</i> s.n. (K, P); Ha Tay. Mont Bavi, 300 m, May 1885, <i>3alansa</i> 2932 (K, P); Mont Bavi, pres de la pagode de?ein Touan, June 1885, <i>3alansa</i> 2933 (K, P 3 sheets); Hoa Binh. Mai chau, Pa co, Ban Phuong, Khoi, Binh, 12 June 1999, <i>Phuong</i> 1561 (HN 2 sheets); Hang Tram [20.215°, 103.5°], 3 May 1950, <i>Lien</i> s.n. (HN); Quang Ninh. Ha-coi [21.2201°, 105.5619°], Chuk-phai, Taai Wong Mo Shan and Vicinity, 16 – 22 Oct. 1936, <i>Tsang</i> 25020 (C, E, K, P); Ha-coi, Shui Mei Village, NE of Chuk-phai, Taai Wong Mo Shan and vicinity, 23 June – 31 Aug. 1939, <i>Tsang</i> 29398 (BKF, C, E, G, K, L, P, SING); Lam?ong.?i Linh,?inh Trang Thuong [11.5628°, 105.9303°], 16 May 1980, <i>Nguyen</i> 28 (HN);?uc Trong?istr., Xa Hiep An (Elephant Mountain), 11°30&amp;33”N 108°23&amp;23”E, 1300 – 1600 m, 31 Aug. 2001, <i>Gardner et al</i>. 80 (E); Lao Cai. Kilo 16, route de Ta Phinh Chapa [Sa Pa], 1230 m, 3 Aug. 1926, <i>Poilane</i> 12802 (P 3 sheets); Chapa [Sa Pa, 22.33°, 103.8665°], 1200 – 1300 m, 6?ec. 1913, <i>Cheoalier</i> 29210 (P 2 sheets); Nghe An. Co Ba (Ke Nhe), <i>Cheoalier</i> 32220 (P 2 sheets); Thua Thien-Hue. Hue and vicinity, May – July 1925, <i>Clemens &amp; S. Clemens</i> 3693 (G 2 sheets, K, P); Son La. Ngoc Chien, Ban Pieng, 21°33&amp;23.11”N 102°13&amp;32.2”E, 3 Oct. 2008, <i>Du et al</i>. HNK 2302 (K). <b>LAOS.</b> Vientiane. Pu Bia [19°, 103.1665°], 1300 m, 13 April 1932, <i>Kerr</i> 21058 (K 2 sheets, P); Champasak. Khong [12.1165°, 103.83°], Khon Island, Southern tip, Hill at Hang Khong Island, Mekong R., 123 m, 12 Oct. 1995, <i>Maxwell</i> 95-1165 (CMUB); Pak Song [13.1833°, 106.2333°], July 1929, <i>Vidal</i> 1022B (P); Salavan. Ban Munh Pak?i [13.2333°, 106.2°] &amp; Ban Honh, 22 Sept. 1928, <i>Poilane</i> 13518 (P); <b>UNLOCALISED</b>. Cahn Trap, <i>Spire</i> 1333 (P 2 sheets). <b>CAMBODIA.</b> Angkor [13.2165°, 103.8665°], <i>Godefroy-Lebeuf</i> s.n. (K); montibus Tamir in Prov [Province] Tpong, 800 m, May 1850, <i>Pierre</i> 1181 (P). <b>THAILAND.</b> Chiang Mai. Chom Thong,?oi Song Mea, 1300 m, 21 Aug. 1999, <i>Watthana et al.</i> 399 (QBG); Chom Thong, Mae Soi Subdistr., Mae Soi Ridge, near Ban Bah Gluay (Meo Village), 1253 m, 12 Aug. 1991, <i>Maxwell</i> 91-535 (E, L, P);?oi Angka [?oi Inthanon, 18.3833°, 98.2833°], 1200 m, 18 July 1922, <i>Kerr</i> 6325 (BK, BM, K); ibid., 1800 – 2300 m, 3 Oct. 1951, <i>Murata et al.</i> T-13988 (holotype of <i>G. jaoanicum</i> var. <i>breoidens</i>, KYO; isotype BKF); ibid., 2330 m, 12 July 1990, <i>Phengklai et al.</i> 5303 (BKF 2 sheets);?oi Angka [?oi Inthanon], Mae Wang drainage [18.6123°, 98.5530°], 1900 m, 29 July 1925, <i>Garrett</i> 209 (BM 2 sheets, K, SING);?oi Inthanon, along the road to Mae Chaem, km 1 – 2, 18°31&amp;N 98°29&amp;E, 2000 m, 20 Sept. 1993, <i>Larsen et al.</i> 26529 (AAU, BKF);?oi Inthanon, en route to Mae Jaem [Mae Chaem] km 38, 1800 m, 6 Jan. 1998, <i>Srisanga et al.</i> 33 (QBG);?oi Inthanon, S side of Wachirathan Waterfalls [18.3218°, 98.3992°], 500 – 860 m, 1 Aug. 1988, <i>Tsugaru</i> T-61811 (BKF, GH, L);?oi Sootep, 520 m [2200 ft], 15 July 1909, <i>Kerr</i> 533 (holotype of <i>G. intermedium</i>, K; isotypes AB?, BM, K 2 sheets);?oi Suthep, 18°30&amp;N 98°33&amp;E, 1300 m, 15 Aug. 2002, <i>Nielsen et al.</i> 1512 (BKF);?oi Suthep, E side, Pah Laht [18.8012°, 98.9238°], 600 m, 23 July 1995, <i>Maxwell</i> 85-526 (BKF, L);?oi Suthep, E side, Mohntatahn Waterfalls [18.8165°, 98.9°], 600 m, 19 Nov. 1985, <i>Maxwell</i> 85-1263 (L); Huai Chang Kien [18.8238°, 98.8886°], 900 m [3000 ft], 23 July 1921, <i>Kerr</i> s.n. (AB? 2 sheets); Fang, 19°35&amp;N 99°11&amp;E, 600 m, 22 July 1968, <i>Larsen et al.</i> 2636 (AAU, L, P); Chiang?ao [19.3665°, 98.9665°], 1020 m, 23 June 1936, <i>3unchai</i> 203 (AB?, BKF); Mae On,?oi Lahn, W side, above Mae Lio Village, Mae Lio Valley; Huay Gayo Subdistr., 1023 m, 12 July 2003, <i>Maxwell</i> 03-226 (BKF); Queen Sirikit Botanic Gardens, Huai Puu [18.8991°, 98.8603°], 900 m, 25 Aug. 1992, <i>3GO staff</i> 1262 (QBG); Me Sa [18.9032°, 98.9293°], July 1913, <i>Kerr</i> s.n. (BM); Ban Mae Saa Mai, 1000 m, 12 Aug. 1998, <i>Pongamornkul</i> 265 (QBG); Chiang Rai. Chiang Rai, Khun Korn Waterfalls [19.8821°, 99.6169°], 653 m, 11 Oct. 1995, <i>KK</i> 200 (BCU); Mae Suai?istr., Ban San Sa-ard, 16 July 1965, <i>Sangkhachand</i> 911 (BK); Wat Phra Thard?oi Tung [20.3221°, 99.8328°], 1313 m, 22 June 2002, <i>Chamchumroon</i> 1393 (BKF); Wiang Pa Pao?istr., Mae Chadee [19.1833°, 99.3165°], 1200 m, 31 Sept. 1993, <i>Vial-Debas</i> 36 (CMUB);?oi Luang National Park, [19.2°, 99.5165°], 1153 m, 8 Sept. 1995, <i>Maxwell</i> 95- 982 (BKF, CMUB); Phayao.?oi Luang National Park, E side, 630 m, 22?ec. 1995, <i>Petrmitr</i> 182 (CMUB);?oi Luang National Park, E side, Jahm Bah Tawng Falls, 353 m, 23 July 1995, <i>Petrmitr</i> 22 (CMUB, L); Nan. Bo Kleu,?oi Phu Kha National Park, Sapan Waterfalls, 19°12&amp;N 101°12&amp;E, 830 m, 6 July 2001, <i>Srisanga</i> 1933 (QBG);?oi Phu Kha National Park, 19°13&amp;N 101°06&amp;E, 1630 m, 1 July 1999, <i>Srisanga et al.</i> 539 (QBG); Nam Hang [18.3333°, 100.6165°], 600 m, 11 July 1926, <i>Winit</i> 1523 (BK, BKF, BM); Lamphun. Khun Tan [18.2508°, 99.2825°], 1000 m, 31 Oct. 1955, <i>Chermsirioathana</i> s.n. (BK); Lampang. Me Ngow, 220 – 260 m, 26 Aug. 1922, <i>Winit</i> 522 (BK, BKF, BM); Muang Pan, Chae Sorn National Park, along Mae Mawn Stream, near Jae Sawn Waterfalls, 330 m, 22 July 1996, <i>Panatkool</i> 132 (CMUB, L); Wang Nua [19.13°, 99.6165°], 630 m, 26 July 1996, <i>Maxwell</i> 96-1022 (CMUB); Tak.?oi Mussor, 900 m, 5?ec. 1960, <i>Smitinand</i> 5038 (BKF); Yanhee?am [15.23°, 98.9665°], <i>Youngboonkird&lt;</i>
description: Published as part of <i>Bongcheewin, Bhanubong, Ingrouille, Martin J. &amp; Paton, Alan J., 2022, A revision of Gomphostemma (Lamiaceae), pp. 27-92 in Kew Bulletin 77 (1)</i> on pages 55-64, DOI: 10.1007/s12225-021-09991-y, http://zenodo.org/record/7615581
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