Cleome Roalson, 2021, s.s.
Creators: name: Roalson, Eric H.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Magnoliopsida
Brassicales
Cleomaceae
Cleome

description: Key to Cleome s.s.   
1. Leaves compound, 3-foliolate...........................................................................................................................................................2.  
1. Leaves simple..................................................................................................................................................................................16.  
2. Fruits> 4 mm wide; seeds villous......................................................................................................................................................3.  
2. Fruits <4 mm wide; seeds villous or papillate...................................................................................................................................5.  
3. Fruits pendulous................................................................................................................................................................................4.  
 3. Fruits erect to erect-spreading........................................................................................................ C. socotrana Balfour (1882: 501)   
 4. Petals 3–4 mm long; fruits oblong............................................................................................ C. amblyocarpa Barratte & Murbeck   
 4. Petals 4–8 mm long; fruits linear................................................................................. C. ramosissima Parlatore ex Webb (1854: 22)   
5. Seeds villous (sometimes pubescence very short).............................................................................................................................6.  
5. Seeds papillate...................................................................................................................................................................................8.  
 6. Fruits pendulous........................................................................................................................... C. ramosissima Parlatore ex Webb   
6. Fruits erect to erect-spreading...........................................................................................................................................................7.  
 7. Leaves glaucous (usually also with noticeable glands, at least along margin); pedicels glabrous; petals 3–4 mm long; seeds with long hairs.............................................................................................. C. omanensis (Chamberlain & Lamond) Thulin (2003: 217)   
 7. Leaves green (not glaucous); pedicels glandular; petals ca. 5 mm long; seeds with short hairs, often with glabrous patches.............................................................................................................................................................. C. hadramautica Thulin (2003: 216)   
 8. Fruits linear, very long, reaching 11 cm; upper petals white (or slightly yellowish) on upper half, clearly differentiated by coloration from lateral petals; veins clearly evident on petals............................................................................................ C. violacea Linnaeus   
8. Fruits linear, but no more than 4 cm long; petals white or yellow to pink or purple, but all petals of similar coloration; veins evident or not on petals..................................................................................................................................................................................9.  
9. Petal blades obovate to oblong with a distinct narrowly linear claw..............................................................................................10.  
9. Petal blades spatulate to linear, but without a distinct narrowly linear claw...................................................................................12.  
 10. Fruits spreading; gynophore 2.5–10 mm long; sepals triangular-lanceolate; glabrous or with few glands................................................................................................................................................................................................................. C. stevensiana Schultes   
10. Fruits pendulous; gynophore 0.5–2(– 3 mm) long; sepals lanceolate or oblong; variable in the presence of glands......................11.  
 11. Sepals lanceolate, acuminate; ± densely clothed with glands.............................................................. C. ornithopodioides Linnaeus   
 11. Sepals oblong, obtuse, or acuminate; glabrous or with few glands...................................................................... C. iberica Candolle   
 12. Leaflets linear to narrow linear-elliptic.................................................................................................. C. ariana Hedge & Lamond   
12. Leaflets obovate to broadly elliptic to +/- circular..........................................................................................................................13.  
 13. Style up to 0.5 mm long; sepals c. 1 mm long........................................................... C. khorassanica Bunge & Bienert ex Boissier   
13. Style 1.5–5 mm long; sepals 3–4 mm long.....................................................................................................................................14.  
 14. Fruits 10–17 mm long........................................................................................................................................ C. foliolosa Candolle   
14. Fruits (20–) 28–40 mm long.............................................................................................................................................................15.  
 15. Annual; upper leaves 3-foliolate; style in fruit (1.5–) 3–5 mm long................................ C. heratensis Bunge & Bienert ex Boissier   
 15. Perennial, small shrub; upper leaves often unifoliate; style in fruit <1 mm long or absent............................................................................................................................................................................................................ C. pakistanica (Jafri) Khatoon & Perveen   
16. Leaves tomentose, particularly on young material..........................................................................................................................17.  
16. Leaves not tomentose......................................................................................................................................................................18.  
 17. Flowers in rather dense racemes; filaments much elongated, 2–3 times as long as petals; gynophores 1.5–1.8 cm long................................................................................................................................................................................................ C. gordjagini Popov   
 17. Flowers in very loose racemes, occupying a large part of the stem; filaments elongated, slightly longer than petals; gynophores ca. 0.7–1 cm long...................................................................................................................................................... C. turkmena Bobrov   
 18. Fruits 10–35 mm wide, for the most part inflated, indehiscent....................................................................... C. coluteoides Boissier (Note: If young and without fruits, the flowers are typically subtended by large, often spathulate bracts not found in other species.)   
18. Fruits 3–10 mm wide, not inflated, dehiscent.................................................................................................................................19.  
 19. Fruits erect........................................................................................................................................................ C. albescens Franchet   
19. Fruits pendant..................................................................................................................................................................................20.  
20. Fruits glabrous; plants glaucous (young upper stems and pedicels often glandular)......................................................................21.  
20. Fruits with dense sessile or stalked glands, plants rarely glaucous, typically glandular throughout..............................................22.  
 21. Fruits 3–5 mm wide; sepals 1–2.5 mm long; petals acute to acuminate........................................................... C. oxypetala Boissier   
 21. Fruits 5–10 mm wide; sepals 3–5 mm long; petals acute............................................................................. C. glaucescens Candolle   
 22. Plant covered in dense with long stalked glandular hairs; fruits 35–60 × 4–5(–8) mm...................................... C. arabica Linnaeus   
 22. Plant with sparse sessile or short stalked glands; fruits 25–50(–60) × (3–) 4–10 mm............................................ C. rupicola Vicary
description: Published as part of Roalson, Eric H., 2021, A revised synonymy, typification, and key to species of Cleome sensu stricto (Cleomaceae), pp. 54-68 in Phytotaxa 496 (1) on pages 56-57, DOI: 10.11646/phytotaxa.496.1.2, http://zenodo.org/record/5423496