Hydropsyche igunapali Olah & Schefter 2008, new species
Creators: name: Oláh, J.
name: Johanson, K. A.
Publisher: name: Zenodo
Year: 2008
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Hydropsychidae
Hydropsyche
Hydropsyche igunapali

description: Hydropsyche igunapali Oláh & Schefter, new species   
Fig. 163–167  
 This species is morphologically close to H. nepalarawa, new species from Nepal. Hydropsyche igunapali is characteristic in having a broad, bilobed median keel not tapering and angular in dorsal view; partly covering the intersegmental depression, not sloping distally in lateral view. The harpagones of H. igunapali are almost parallel-sided and clearly capitate, not narrowing in ventral view. The sclerous ventral process of the phallotheca is longer than the phallotremal sclerites, while it is shorter in H. nepalarawa.   
Male. Large, body and wings dark brown. Forewing membrane dark brown with pale maculation forming scattered pattern. Maxillary palp formula I-II-(III, IV)-V, segment V as long as segments I–IV together. Head dorsum dark brown, with 9 paler setal warts. Swollen setal wart present on proepisternum. Pretarsal claws asymmetrical, laterally flanked by setal bundle. Spur formula 244. Both protibiae with posteroapical spur equal to anteroapical spur.  
Wings. Venation typical for the genus. Forewing crossveins m-cu and cu located distantly. Sc and R, and Cu2 and A running separate before costa. Hind wing Sc and R meeting before costa about at crossvein r. Stem of M and Cu1 running closely. Median cell closed. Fork 1 present. Forewing length 11.2 mm.  
Male genitalia. Abdominal segment IX fused annularly, moderately long. Median keel short, parallelsided, deeply bilobed apically in dorsal view (Fig. 164); dorsal and lateral surfaces granulose; anterior margin arciform, with short ventrum and dorsum (Fig. 163). Antecosta well developed, at each side forming double line narrowing dorsally before margin, with well visible external groove of antecostal suture. Apical lobe on posterolateral margins triangular, directed slightly ventrad (Fig. 163). Posterior spine row almost completely continuous, with short area without spines dorsally at apical lobes and dorsum of median keel. Depression between segments IX and X deep, wide; U-shaped; apical end of median keel overhanging and partly covering anterior part of intersegmental depression (Fig. 163, 164). Dorsal margin of segment X short, ventral margin long, convex (Fig. 163); slightly curving dorsad in lateral view; rounded quadrangular in dorsal view (Fig. 164); slightly constricting basally. Lateral setose areas (preanal appendages) forming compact, pale pigmented, well delineated, elliptical surface centrally on both sides of segment, and distal, loosely packed group of separate setae below apicoventral setose lobes. Apicoventral setose lobes modified into pair of short, digitiform processes; very short, slightly directed ventrad. Dorsal interlobular gap deep, widely U-shaped. Apicodorsal setose lobe forming fused, setose, double ridge or crest, dominating dorsum in lateral view (Fig. 163); located below median keel on segment IX. Glabrous, smooth, shallow cavity present on segment X. No darkly pigmented transverse suture running from bottom of intersegmental depression present. Sinuous, longitudinal suture runs below glabrous, shallow cavity, above Lateral setose areas on both lateral surfaces, followed by second sinuous suture running towards ventrum. Coxopodites slightly exceeding apex of segment X; both with slightly sinuous dorsal margin at basal half; ventral margins straight along its length, both dilating distally from mid-length; nearly straight in ventral view (Fig. 165). Harpagones straight, basal half broad, narrowing distad; parallel sided along its length in ventral view (Fig. 165). Phallic apparatus simply bent (Fig. 166); flat, short, ventrad curving basal section; broadening along short ventrad curving base. Phallotheca with slightly concave dorsum near apex, apex dilating. Endophallus distally attached to dorsal wall of phallotheca; reaching over phallic apparatus bend. Phallotremal sclerites falciform; broad in lateral view; ending before apex of sclerous ventral phallothecal lobe (Fig. 166, 167). Pair of sclerous, basally broad, distally narrowing endothecal processes arise from side of phallotremal sclerites; with slightly granulose apex. Sclerous ventral process of phallotheca longer than phallotremal sclerites, single lobe with rounded apex in ventral view (Fig. 167).  
 Holotype male: INDIA: Assam, Kameng, Nyukmadong, 22.iv.1961 [F. Schmid] (ROM, pinned with cleared abdomen in plastic genital vial).   
 Distribution. India (Assam).   
 Etymology. igunapali, “iguna”, meaning twofold; and “pali”, meaning lobe in Sanskrit, named after its peculiar median keel having bilobed ending in dorsal view.
description: Published as part of Oláh, J. & Johanson, K. A., 2008, Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae), pp. 1-248 in Zootaxa 1802 on pages 106-108