Landouria nodifera Nurinsiyah & Neiber & Hausdorf 2019, sp. nov.
Creators: name: Nurinsiyah, Ayu Savitri
name: Neiber, Marco T.
name: Hausdorf, Bernhard
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Camaenidae
Landouria
Landouria nodifera

description: Landouria nodifera sp. nov.
urn:lsid:zoobank.org:act: 36465425-2C50-4721-9B73-3719E2D4A78A
Figs 40, 54, 61, 64; Tables 3–4
Landouria rotatoria – Nurinsiyah 2015: 32 (in part). (not Pfeiffer, 1842)
Diagnosis
Landouria nodifera sp. nov. is characterized by a strongly depressed conical, brownish-corneous, sharply keeled shell and a penis that becomes broader towards is proximal end, the presence of a coecum at the proximal end of the penis and a node at the base of the flagellum.
Etymology
Landouria nodifera sp. nov. is named after the node at its flagellum (Latin: ʻnodusʼ = ʻnodeʼ, ʻferreʼ = ʻto carryʼ; used as an adjective).
Material examined
Holotype
INDONESIA • holotype (det. anat.); Central Java, Pati, Larangan Village, Lanang Spring; 6°54′19″ S, 111°03′22″ E; 100 m a.s.l.; 15 Aug. 2014; A.S. Nurinsiyah, F.L.H. Irsyad, F.J. Sari and E. Nurlaela leg.; measurements: D = 15.0 mm, H = 7.0 mm; MZB 19227.
Paratypes
INDONESIA – Central Java • 1 spec.; same data as for holotype; MZB 19233 • 6 spec.; same data as for holotype; MZB 19228 • 5 spec.; same data as for holotype; ZMH 133428 • 9 spec. (det. anat.); Pati, Larangan Village, Lowo Cave; 6°54′19″ S, 111°03′24″ E; 112 m a.s.l.; MZB 19229 • 1 spec.; same data as for preceding; MZB 21266.
Description
SHELL (Fig. 61; Tables 3–4). Depressed conical, with 5–5.75 hardly convex whorls; protoconch almost smooth; teleoconch with irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes sparse, mainly at periphery; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture rhombic; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 26–38% of shell diameter, hardly obscured by columellar edge.
GENITALIA (Figs 40, 64; Table 4). Atrium short; penis long, consisting of a cylindrical distal part covered by an additional sheath, connected by muscular tissue to proximal part of epiphallus, and a dilated proximal part containing a short penial papilla; inside proximal part of penis with longitudinal pilasters, which pass into a more complex sculpture in distal part of penis; at proximal end of penis with a blunt penis coecum; penial retractor runs from diaphragm to distal part of epiphallus, which tapers proximally; flagellum moderately long, broadest in its middle part, where it is equipped with a node, tapering towards its proximal end; vas deferens narrow and long; vagina long; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.
Remarks
Landouria nodifera sp. nov., L. tonywhitteni sp. nov., L. pacitanensis sp. nov., L. sukoliloensis sp. nov., L. sewuensis sp. nov. and L. intumescens share strongly depressed, brownish-corneous, sharply keeled shells. These species cannot be reliably distinguished by shell characters. Landouria nodifera sp. nov. shares having a coecum at the proximal end of the penis with L. pacitanensis sp. nov., from which it differs in the lack of the narrow constriction subdividing the penis of L. pacitanensis sp. nov. and the presence of a node at the base of the flagellum. Landouria nodifera sp. nov. shares having a node at the base of the flagellum with L. tonywhitteni sp. nov., from which it differs in the presence of the coecum at the proximal end of the penis and the proximally broader penis.
Distribution
Landouria nodifera sp. nov. is endemic to Sukolilo Karst (Fig. 54).
description: Published as part of Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, pp. 1-73 in European Journal of Taxonomy 526 on pages 47-48, DOI: 10.5852/ejt.2019.526, http://zenodo.org/record/3152217