Clivina dilutureticulata Balkenohl, 2025, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina dilutureticulata

description: Clivina dilutureticulata sp. nov.
(Figs 15, 50, 84)
Type material. Holotype: ♂, with labels and data: white, black printed “ PHILIPPINEN - Mindoro 28km S Calapan Balete 100-700m leg. Jäch 27.- 29.11. 1992 ” (NHMW). Paratypes: 1 ♂, same data as holotype (CMBB); 1 ♂, “ PHILIPPINEN: Mindoro or. Baco, Hidden Paradise 19.- 20.11.1993 leg. H. Zettel (27)” / “ Clivina VII. subdepressa Kult det. Alex. Dostal 1994” (CADW).
Description. Measurements. Body length 5.72–6.04 mm (ẍ = 5.91 mm *), width 1.67–1.74 mm (ẍ = 1.71 mm *), ratio L/W of pronotum 0.97–0.99 (ẍ = 0.98*), ratio L/W of elytra 1.88–1.93 (ẍ = 1.90*); (*n = 3). Colour glossy, fuscous. Supraantennal plate and clypeal-wing at the margin slightly translucent. Intermediate and hind legs somewhat paler, mouthparts and first four antennomeres yellowish.
Head about a quarter narrower than pronotum. Clypeus with central part indistinctly excised, with distinctly projecting somewhat acute teeth laterally, separated from clypeal-wing by distinct obtuse-angles notch; wing wider than teeth of clypeus, rounded, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin laterally convex, clypeus, clypeal-wing, and anterior part of supraantennal plate distinctly reflexed margined. Supraantennal plate extended posteriorly into blunt keel, separated from wing by narrow furrow. Clypeus transverse, slightly convex, smooth, clypeal-furrow flattened. Frons moderately convex, with flat pore-like depression at middle, with scattered fine punctures, separated from supraantennal plate by moderately wide furrow, with clypeal seta in front; supraorbital carinae blunt, separating frons from supraorbital furrow, with two supraorbital setigerous punctures at middle of eye level and posterior to gena-level. Neck constriction closed, with distinct sharp step, with second narrow carina. Eye relatively small, somewhat flattened, gena distinct, enclosing eye posteriorly by nearly one quarter. Antenna elongate, reaching over base of elytron. Labrum straight anteriorly, laterally rounded, finely isodiametrically reticulated, six-setose. Mentum with lobes oval, angled at tip, obtusely margined, with isodiametric reticulation, median tooth wide, higher than lobe, hollowed out at tip.
Pronotum (Fig. 50) with disk distinctly flattened in lateral view, flattened at middle and slightly convex laterally in frontal view. Shape sub-cordate, nearly as long as wide. Reflexed lateral margin sub-crenulate (80-fold), moderately convex at middle, widest at middle; anterior angle distinct, posterior angle indicated by distinct anteriorly projecting tooth; lateral channel moderately narrow, reticulated, continuing up to base as sharp, distinctly convex margin. Median line complete, engraved. Anterior transverse line distinct, joining median line. Disk with finely scattered punctures, baso-medially with group of few median sized punctures, with few fine transverse wrinkles laterally, with isodiametric reticulation at base and laterally, with posterior setigerous puncture somewhat removed from margin. Lateral margin at flange slanted with angle of around 30°. Basal channel on flange of regular width, rugose, with sharp carina.
Elytron (Fig. 15) with disk flattened in anterior half in lateral view, flattened at middle and moderately convex in frontal view. Shape of elytra elongate, nearly two times longer than wide, with lateral margin straight anterior middle but slightly diverging, maximum width posterior middle. Humerus distinct, rounded. Reflexed lateral margin sub-crenulate (80-fold) posterior humerus. Lateral channel slightly narrower at middle, row of setigerous punctures more distant at middle, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with small tubercle at base of third interval, basal declivity falling perpendicularly to pedunculus, with isodiametric reticulation. Striae moderately deep, impunctate, one to four free at base, five and six joining at humerus, one and two ending free at apex, three and four, and five and six joining apically. Intervals slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven convex at apex, eight carinate at apex. Surface of intervals two to eight with isodiametric reticulation, reticulation tapering in intensity towards middle, first interval nearly smooth.
Hind wing fully developed.
Lower surface. Proepisternum isodiametrically reticulated, with fine flat transverse wrinkles laterally. Sternite of abdomen with isodiametric reticulation laterally, transverse at middle. Last visible sternite with the two marginal setigerous punctures widely separated, with moderately wide projection posteriorly at middle where it is flattened, with narrow marginal furrow at middle.
Legs slender, protibia with fine longitudinal reticulation, with indistinct sulcus dorsally, with three spines, spatulate-like, with obtuse tooth, movable spur moderately arcuate, basal tarsomere slender at base, carinate, with acute tubercle with two setae and a small tubercle with one seta at middle of tarsomere. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg somewhat widened.
Male genitalia (Fig. 84). Median lobe slender, distinctly curved in basal half in dorsal view, nearly straight in apical half, slightly cracked at beginning of apical spatula, apex acute, in lateral view apically regularly acuminate, somewhat flattened. Parameres robust, with distinct apophyses, distorted, with small seta at apex, each.
Female gonocoxites and epipleurite unknown.
Variation. In one of the paratypes, the neck constriction is not doubled.
Distribution. Known from the island of Mindoro, Philippines.
Diagnosis. A medium sized slender fuscous species with sub-cordate pronotum and intervals two to eight with isodiametric reticulation. A species sharing most of the basic characters is C. bauri sp. nov. However, and beside other characters, in C. bauri sp. nov. the frons of the head is completely flat with the supraorbital carinae nearly continuing to the flat frons, the neck constriction is open at middle and the frons continues to the neck, and the antennae are much shorter, i.e. reaching up to base of the pronotum.
Etymology. The name refers to the elytra with its reticulation on the intervals which is more and more fading to the middle (Latin: dilutus = diluted; reticulum = small sized net).
description: Published as part of Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), pp. 1-70 in Zootaxa 5592 (1) on pages 27-28, DOI: 10.11646/zootaxa.5592.1.1, http://zenodo.org/record/14953674