Daedalma drusilla subsp. krugeriana Pyrcz 2011, n. ssp.
Creators: name: Pyrcz, Tomasz W.
name: Greeney, Harold F.
name: Willmott, Keith R.
name: Wojtusiak, Janusz
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Nymphalidae
Daedalma
Daedalma drusilla
Daedalma drusilla krugeriana pyrcz

description: Daedalma drusilla krugeriana Pyrcz, n. ssp.   
(Figs. 7E, 7F, 16B)  
 [Daedalma drusilla Hewitson; Pyrcz, 1999: 228]   
 Material examined: COLOMBIA: HOLOTYPE female: Tolima, Albania, Central Cordillera, East slopes, 14.VIII.1916, 2800 m, E. Krüger leg., red, rectangular label saying: Holotype, MIIZ; ECUADOR: PARATYPE male: Carchi, Tulcán-Maldonado km 40–50, 2800–3200 m, 07.V.1999, P. Boyer leg., PBF.   
 Description: MALE (Fig. 7E): Head, thorax and abdomen: not differing from nominate subspecies. Wings (FW length: 26 mm): FWD blackish brown, a shade lighter in distal one-third, as compared to uniform blackish brown in nominate. FWD uniform blackish brown, lustrous, same as in nominate. FWV ground colour blackish brown; patches in distal part of discal cell and submarginal area barely lighter than ground colour, same as in some specimens of nominate, somewhat variable in this respect. HWV with more prominent chestnut and chocolate brown pattern than in other subspecies, especially in subapical and marginal area; submarginal milky white patches slightly larger than in nominate, similar as in tinta; three subapical-apical ocelli in Sc-Rs, Rs-M1 and M1-M2 form an arch pointing distally at costa, similarly to tinta and contrary to the nominate whose ocelli form a straight line.   
 Genitalia: Not examined.   
 FEMALE (Fig. 7F): Head, thorax and abdomen: not differing from nominate subspecies. Wings: FW length: 27.5 mm; FWD dark brown in basal half, light brown along outer margin; an irregular yolk-yellow patch in distal 2/3 of discal cell connected through base of cell M3-Cu1 to a large lighter yellow patch covering most of wing surface from postmedian to submarginal area except for an incision of brown along vein M3, gradually turning into light brown in subapical area, enclosing a faint brownish spot in M1-M2; fringes alternately brown and yellow, dark brown at vein ends. HWD medium brown, slightly darker in basal one-third; traces of rich yellow forming faint submarginal streaks in cells M1-M2 and M2-M3; fringes yellow in apical area, turning orange towards apex, vein ends brown. FWV ground colour slightly lighter brown and duller than on upperside; yellow markings as on upperside except that discal cell patch is disconnected from yellow postmedian area; subapical and apical area dusted with milky white scales, and different shades of brown; two dark brown subapical ocelli in R5-M1 and M1- M2; a zigzagging subapical line gradually turning into a wider area of brown in M2-M3. HWV ground colour light brown with a nondescript pattern of brown, chestnut and milky white lines and bands, similar to other taxa, with a somewhat better marked dark brown median diffused band and a milky white roughly triangular submarginal patch contiguous to vein M3. Genitalia (Figs. 16B): This subspecies differs from nominate by significantly longer ductus bursae and narrowly bent central part of batten on lamella postvaginalis. Sinus vaginalis with narrow entrance, flattened anteriorly, with wrinkles on lateral and ventral walls. Antrum opens to sinus vaginalis posteriorly, not at its base. Ductus bursae connects with bursa copulatrix centrally. Central part of lamella postvaginalis widely concave. Coliculum well developed at posterior part of ductus bursae. Narrow central part of a batten on lamella postvaginalis well pronounced. Outer walls of papillae anales at base of reduced apophyses posteriores set off as well marked bumps. Cuticule around anus with pronounced wrinkles. Bursa copulatrix with two ribbon-like, parallel signa consisting of minute teeth.   
 Remarks: This subspecies of D. drusilla is described based on a female, which bears some resemblance to the "dora type " form of the nominate subspecies, with its wide yellow markings on the FWD. However, contrary to that form and other females of D. drusilla, there is no trace of yellow on the HWD. The only known individual was collected by Edwin Krüger at Albania in Tolima on the eastern slopes of the Colombian Central Cordillera (Pyrcz, 1999). Available data indicate that Central and Eastern Cordillera populations of most species of Pronophilina, especially inhabiting high elevations, belong to separate subspecies (Adams, 1986), and Pyrcz & Rodríguez (2007) confirm that the valley of Magdalena is an important zoogeographic barrier. Given the distinctive wing pattern of the only known female and biogeographic data from other pronophilines, we feel justified in describing it here as a new subspecies. Curiously, Krüger (1924) lists a male of " D. dora " (= D. drusilla), collected in the area of Bogotá, and states that he did not collect any individual of D. drusilla, yet mentions no female of either. Daedalma drusilla is apparently a rare species in the central departments of the Central Cordillera, as it was not collected by Adams (1986) who sampled extensively in Tolima, and it was not listed as occurring in Tolima or Caldas by Vargas & Salazar (2004).
description: Published as part of Pyrcz, Tomasz W., Greeney, Harold F., Willmott, Keith R. & Wojtusiak, Janusz, 2011, 2898, pp. 1-68 in Zootaxa 2898 on pages 30-31