Coccidae Fallen 1814
Creators: name: Choi, Jinyeong
name: Soysouvanh, Pheophanh
name: Lee, Seunghwan
name: Hong, Ki-Jeong
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Coccidae

description: Key to adult females of species of Coccidae in Laos
1. Stigmatic clefts each with a single stigmatic spine............................................................ 2
- Stigmatic clefts each with at least 3 stigmatic spines.......................................................... 3
2. Antenna with 4th and 5th segments each longer than wide; preopercular pores eXtending anteriorly onto thoraX; multilocular disc-pores absent from metathoraX............................................. Drepanococcus chiton (Figs 4, 5)
- Antenna with 4th and 5th segments each shorter than wide, or of similar length to width; preopercular pores restricted to abdomen; multilocular disc-pores present on metathoraX........................................... D. cajani (Figs 2, 3)
3. Marginal setae each fan-shaped.......................................................................... 4
- Marginal setae not fan-shaped............................................................................ 5
4. Preopercular pores present anterolateral to anal plates, in 3 groups on each side; legs entirely absent.............................................................................................. Paralecanium quadratum (Fig. 34)
- Preopercular pores present anterolateral to anal plates, in 2 groups on each side; legs represented by leg stubs............................................................................................. P. expansum (Figs 32, 33)
5. Marginal setae truncate, of 2 types (slender and long, or thick and same length as slender ones or shorter); anal plates each with apical setae truncate..................................................... Megapulvinaria maxima (Figs 35, 36)
- Marginal setae of a single type, tips pointed or frayed; anal plates each with apical setae sharply spinose................. 6
6. Body entirely covered with thick waX test; without white ovisac; anal plates situated on sclerotized caudal process at maturity; stigmatic spines conical or bullet-shaped................................................................... 7
- Body without obvious waXy test; sometimes with white ovisac; anal plates not situated on sclerotized caudal process; stigmatic spines normally spinose............................................................................... 11
7. WaX test star-shaped, with 7 rays; interantennal setae numbering about 7-10 pairs......... Ceroplastes stellifer (Figs 14, 15)
- WaX test more or less round to oval; interantennal setae numbering about 1–3 pairs................................. 8
8. Stigmatic clefts each with a large medial seta; legs poorly developed, each with tibia and tarsus fused...................................................................................................... C. rubens (Figs 12, 13)
- Stigmatic clefts each with medial seta, if present, never more than twice as large as other stigmatic setae; legs normally devel- oped, each with tibia and tarsus distinct.................................................................... 9
9. Ventral tubular ducts each with a swollen inner ductule, present in lateral submarginal areas of body; with about 8–10 slender marginal setae present between anterior and posterior stigmatic clefts on each side............. C. floridensis (Figs 11, 12)
- Ventral tubular ducts each with a narrow or filamentous inner ductule, present in submarginal area of posterior abdomen and head only; with about 2–4 slender marginal setae present between anterior and posterior stigmatic clefts on each side..... 10
10. WaX test forming a distinct horn at maturity; stigmatic spines arranged in about 6 rows by each cleft; antenna 6 segmented; legs without tibio-tarsal articulatory scleroses.................................................. C. ceriferus (Figs 6, 7)
- WaX test without a horn at maturity; stigmatic spines arranged in 2 or 3 rows by each cleft; antenna 7 segmented; legs with tibio-tarsal articulatory scleroses.................................................... C. cirripediformis (Figs 8, 9)
11. Anal plates together kite-shaped, each plate with anterolateral margin over 1.7 times as long as posterolateral margin............................................................................. Milviscutulus mangiferae (Figs 37, 38)
- Anal plates together approXimately quadrate, each plate with anterolateral margin similar in length to posterolateral margin 1 2
12. Ventral tubular ducts, each with a broad inner ductule, present in medial areas of head, thoraX and abdomen; in life, mature adult female with white ovisac.......................................................................... 13
- Ventral tubular ducts each with a broad inner ductule absent; or if present, either (i) present in medial area of thoraX only (e.g. Coccus celatus and C. viridis), or (ii) mainly present in submarginal areas of body, as well as few ducts present around meso- and procoXa (e.g. Saissetia coffeae); in life, mature adult female without white ovisac.............................. 15
13. Dorsum with polygonal reticulations; with 4 or 5 stigmatic spines present in each stigmatic cleft................................................................................................ Pulvinaria polygonata (Figs 41, 42)
- Dorsum without polygonal reticulations; with only 3 stigmatic spines present in each stigmatic cleft................... 14
14. Multilocular disc-pores mostly each with 7 loculi; with ventral tubular ducts of 3 types (type I: each with a broad inner ductule; type II: each with a narrow inner ductule; type III: each with a filamentous inner ductule)........ P. floccifera (Figs 39, 40)
- Multilocular disc-pores mostly each with 10 loculi; with ventral tubular ducts of only 2 types (type III with filamentous inner ductules absent)....................................................................... P. psidii (Figs 43, 44)
15. Submarginal band of ventral tubular ducts present........................................................... 16
- Submarginal band of ventral tubular ducts absent........................................................... 19
16. Dorsum with polygonal reticulations; anal plates each without a discal seta; in life, without H-shaped ridge on dorsum................................................................................ Parasaissetia nigra (Figs 45, 46)
- Dorsum without polygonal reticulations; anal plates each with a discal seta; in life, young specimens with H-shaped ridge on dorsum............................................................................................. 17
17. With ventral tubular ducts of 3 types (type I: each with a broad inner ductule; type II: each with a narrow inner ductule; type III: each with a filamentous inner ductule), present in submarginal and medial areas of thoraX and abdomen............................................................................................ Saissetia coffeae (Figs 47, 48)
- With ventral tubular ducts of only 1 type (type I: each with a narrow inner ductule), present in submarginal areas only..... 18
18. Marginal setae between anterior and posterior stigmatic clefts on each side numbering 17–23...... S. miranda (Figs 49, 50)
- Marginal setae between anterior and posterior stigmatic clefts on each side numbering 5–13.......... S. oleae (Figs 51, 52)
19. Dorsum of mature adult female formed of polygonal sclerotized plates; ventral tubular ducts absent........................................................................................... Eucalymnatus tessellatus (Figs 30, 31)
- Dorsum of mature adult female without polygonal plates; ventral tubular ducts present, of 1 or 2 types................. 20
20. Pregenital setae numbering 1 or 2 pairs; ventral tubular ducts absent from medial area of thoraX, present only in submarginal area of posterior abdomen...................................................... Coccus capparidis (Figs 16, 17)
- Pregenital setae numbering 3 pairs; ventral tubular ducts present in medial area of thoraX............................ 21
21. Dorsal submarginal tubercles present..................................................................... 22
- Dorsal submarginal tubercles absent...................................................................... 25
22. Ventral tubular ducts in medial area of thoraX each with a broad or filamentous inner ductule......................... 23
- Ventral tubular ducts in medial area of thoraX each with a narrow inner ductule.................................... 24
23. Multilocular disc-pores mostly each with 10–12 loculi; ventral tubular ducts of 2 types (type I: each with a broad inner ductule; type II: each with a filamentous inner ductule); antenna 8 segmented........................... C. celatus (Figs 18, 19)
- Multilocular disc-pores mostly each with 7 loculi; ventral tubular ducts of 1 type (type I: each with a broad inner ductule) only; antenna 7 segmented.................................................................. C. viridis (Figs 28, 29)
24. Antennae each 8 segmented....................................................... C. gymnospori (Figs 22, 23)
- Antennae each 7 segmented....................................................... C. hesperidum (Figs 24, 25)
25. Dorsal setae tapering, sharply setose; marginal setae with pointed apices; ventral tubular ducts abundant, a continuous transverse band of ducts present on mesothoraX; legs without tibio-tarsal articulatory scleroses........ C. formicarii (Figs 20, 21)
- Dorsal setae cylindrical, bluntly spinose; marginal setae with bifid or fimbriate apices; ventral tubular ducts scarce, a continu- ous transverse band of ducts absent from mesothoraX, a few present near each mesocoXa; legs with tibio-tarsal articulatory scleroses................................................................... C. latioperculatum (Figs 26, 27)
description: Published as part of Choi, Jinyeong, Soysouvanh, Pheophanh, Lee, Seunghwan & Hong, Ki-Jeong, 2018, Review of the family Coccidae (Hemiptera: Coccomorpha) in Laos, pp. 1-62 in Zootaxa 4460 (1) on pages 6-7, DOI: 10.11646/zootaXa.4460.1.1, http://zenodo.org/record/1459506