Cheumatopsyche chitawana Olah & Johanson 2008, new species
Creators: name: Oláh, J.
name: Johanson, K. A.
Publisher: name: Zenodo
Year: 2008
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Hydropsychidae
Cheumatopsyche
Cheumatopsyche chitawana

description: Cheumatopsyche chitawana Oláh & Johanson, new species   
Fig. 341–344  
 Cheumatopsyche chitawana belongs to the Cheumatopsyche ningmapa, new species cluster. Here we separate this new species cluster as characterized by slender and highly sinuate harpagones, reduced mesoapical smooth lobe and shortened apicoventral setose lobes with knob-like apex. The species cluster contains totally 4 species: C. chitawana, C. ningmapa Schmid from Bhutan, C. naumanni Malicky from Afghanistan, Pakistan and Nepal, and C. charites Malicky & Chantaramongkol from Thailand, Laos, Vietnam and Malaysia (Oláh et al. 2008). Cheumatopsyche chitawana is easily recognized by having knob-like apex of the apicoventral setose lobes not produced. Instead they originate posteriorly on segment X. As a result, the dorsal interlobular gap is shallower. Furthermore, segment X broadening before apex due to the longitudinal and lateral rim of the apicoventral lobes, easily seen in dorsal view.   
Male. Body and wings pale brown with lighter pubescence; legs and antennae stramineous. Maxillary palp formula I-(III, IV)-II-V, segment V as long as sum of segments I–IV. Head dorsum pale brown, with 9 pale brown setal warts. Swollen setal wart absent on each proepisternum. Setal wart present on each precoxale. Pretarsal claws asymmetrical, laterally flanked by setal bundle. Mid- and hind-leg claws asymmetrical.  
Wings. Forewing evenly pale, with slightly darker veins. Forewing crossveins m-cu and cu almost meeting in oblique line. Cu2 and A1 running separately before costa. Hind wing Sc and R meeting slightly posteriorly of crossvein r. Crossveins r and s tangential. Fork 1 absent. Forewing length 7.0 mm.  
Male genitalia. Abdominal segment IX fused annularly, tergum as long as sternum (Fig. 341). Anterior margins of segment IX regularly bow-shaped, rather flat dorsally in lateral view (Fig. 341). Apical lobe on posterolateral margins blunt right-angled, located above less sclerotized articulation cavity of gonocoxites. Spine row on posterior margin of segment IX intermittent, heterogeneous. Spines on dorsolateral spiny lobes 2 times longer than spines on apical lobes; in dorsal view dorsolateral spiny lobes protruding and surrounding deep, wide, concave indentation or diverticulum (Fig. 342). Inter-segmental step between segments IX and X pronounced, deep, sharply angled. Segment X long, nearly triangular in lateral view (Fig. 341); regularly quadrangular in dorsal view (Fig. 342); broadening apical half produced by ventral rim of apicoventral setose lobes; bilobed in dorsal view (Fig. 342). Dorsal interlobular gap wide and shallow. Sutures well developed on both sides, transverse, crossing segment X obliquely; longitudinal sutures continuing from apicoventral setose lobes; meeting transverse suture into Y. Smooth dorsomesal plate reduced, ending in concavity. Mesal part of apicoventral setose lobes fusing with segment X; apex free, blunt, knob-like, curving slightly mesad and pressed into body of segment X (Fig. 342); setae restricted to apical tip. Lateral setose areas (superior or preanal appendages) forming small, circular, elevated warts; shifted distad, basally surrounded by rim of apicoventral setose lobes. Coxopodites slightly exceeding apex of segment X; forming slightly sinuous rod with dilating apex in lateral view (Fig. 341); weakly curving mesad apically in ventral view (Fig. 343). Harpagones sinuate in lateral view (Fig. 341); basal half broad, S-shaped in ventral view (Fig. 343). Phallotheca with straight dorsum (Fig. 344). Endophallus long, running through entire phallotheca; ending in narrow tube at gonopore. Chitinized endothecal process elongating ovoid; curving ventrad. Phallotremal sclerites vertically broad in lateral view. Vestigial, membranous, ventral endothecal lobes nearly invisible.  
 Holotype male: NEPAL: Chitawan District, Royal Chitawan National Park, 1.xi.1985 [J. Louton] - (NMNH).   
Paratypes: same data as holotype - 9 males, 15 females (NMNH); 2 males, 1 female (OPC).  
 Distribution. Nepal.   
 Etymology. chitawana, named after the type locality, Chitawan National Park.
description: Published as part of Oláh, J. & Johanson, K. A., 2008, Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae), pp. 1-248 in Zootaxa 1802 on page 205