Lamprocystis (Lamprocystis) guttula
Creators: name: Hyman, Isabel T.
name: Ponder, Winston F.
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Euconulidae
Lamprocystis
Lamprocystis guttula

description: Lamprocystis (Lamprocystis) guttula (Pfeiffer, 1853)   
Figures 11L, 40J–L, 41A  
 Material examined. Vanuatu: FMNH 174940 (two specimens dissected, one radula examined), behind Ranmawat, SW Pentecost, dense forest, alt. 120 m, 10 Nov. 1972, L. Price.   
 Description. External morphology: Shell 4.5 whorls, light golden brown, glossy, spire and apex slightly raised. Protoconch smooth; teleoconch sculptured with faint microscopic spiral grooves. Whorl profile rounded above and below evenly convex periphery. Internal walls of early whorls complete; umbilicus closed. Animal primarily pale grey with white sole and darker grey on top of head. Right mantle lap moderately long, left mantle lap moderately short, both mantle laps narrow, tapering to a point, not fused, uniform in colour. Mantle lobes very small. Sole of foot and caudal apparatus as for family; caudal horn large; caudal foss horizontal slit in tail.   
 Mantle cavity: As for family. Pigmentation absent, mantle gland absent.   
 Digestive system: Oesophageal crop absent.   
 Genital system (Figure 41A): As for family. Carrefour and talon both not embedded in albumen gland. Oviduct non-glandular, containing three embryos. Post-uterine oviduct very long, internal longitudinal pilasters absent. Bursa copulatrix inserted on penis, very short, shorter than the free oviduct; duct of bursa copulatrix wide, not distinguishable from bursa copulatrix, internally smooth. Penis short; epiphallus enters penis through two fleshy lips; penis internally smooth, at least two longitudinal penis pilasters present, penial diverticulum absent. Penial sheath present; penis retractor muscle attached to epiphallus near penis junction. Epiphallus longer than penis, simple internally. Epiphallic retractor caecum absent; Flagellum and lime-sac absent. Spermatophore not present in dissected specimens.   
 Radula (Figure 40J–L): Relatively long and narrow. Central tooth with moderately small ectocones; mesocone lanceolate, longer than tooth base. Lateral and marginal tooth fields distinguishable. Lateral teeth with endocone smaller than central tooth ectocones; ectocone equal in size to those on central tooth; mesocone longer than tooth base. Marginal teeth with endocones absent; ectocones approximately same length and breadth as mesocone, subdivided into three to four extra teeth. Radular formula (26.7.1.7.26) × 92 rows.   
 Remarks. The type species of Lamprocystis sensu stricto was dissected by Baker (1940) but was not figured due to its poor preservation condition. However, Baker (1940) stated that it was very similar to L. ensifera (Mousson, 1869), which was figured. The species examined here shows anatomical similarities to L. ensifera and can be assigned with confidence to this subgenus.   
 Lamprocystis is divided by Schileyko (2002a) into nine subgenera. Lamprocystis sensu stricto contains around 40 species and is distributed in the Philippines and many islands of the Pacific.   
 In the phylogenetic analysis in the current study (Figures 1–3) and in the molecular phylogenetic analysis of Hyman et al. (2007) (Figure 44A), Lamprocystis guttula always grouped with the other members of Microcystinae.
description: Published as part of Hyman, Isabel T. & Ponder, Winston F., 2010, A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462, pp. 1-148 in Zootaxa 2462 (1) on pages 114-116, DOI: 10.11646/zootaxa.2462.1.1, http://zenodo.org/record/5312710