Helodon (Helodon) Enderlein
Creators: name: Adler, Peter H.
name: Crosskey, Roger W.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Simuliidae
Helodon

description: Subgenus HELODON Enderlein s. str.
Helodon alpestris (Dorogostaisky, Rubtsov & Vlasenko)
Chubareva (1977 a, 1977d, 1978b, idiogram, notes on general polytene features; as Prosimulium alpestre and P. altaicum): Russia
Chubareva & Petrova (1979, 2003, notes on general polytene features, polytene chromosome lengths; as Ahaimophaga alpestris and A. altaica): Russia
Rothfels (1979, cytodendrogram, whole-arm interchange [IIL + IIIS, IIS + IIIL]): North America
Chubareva & Petrova (1999, comparison with H. ferrugineus [as H. rufum], notes on general polytene features, photos of metaphase and polytene complements, polytene chromosome lengths; A. altaica): Russia
Chubareva & Petrova (2008a, description, photos of metaphase and polytene complements, polytene chromosome lengths): Russia
Helodon beardi Adler, Currie & Wood
Newman (1983, characterization, hybrid with H. newmani [as P. onychodactylum ‘sibling species 5’], inversions mapped, photo of partial IIL, resolution relative to H. onychodactylus standard; as P. onychodactylum ‘sibling species 3’): USA
Henderson (1985, 1986a, characterization, cytodendrogram, inversions mapped; as P. onychodactylum ‘form 3’): USA
Adler, Currie & Wood (2004, chromosomal characters in larval key, formal name for Newman’s 1983 P. onychodactylum ‘sibling species 3’, notes on geographic variation): USA
Helodon chaos Adler, Currie & Wood
Newman (1983, characterization, hybrid with H. mccreadiei [as P. onychodactylum ‘sibling species 7a/7b’], inversions mapped, photo of partial IIL, resolution relative to H. onychodactylus standard; as P. onychodactylum ‘sibling species 1a and 1b’): USA
Henderson (1985, 1986a, characterization, cytodendrogram, inversions mapped; as P. onychodactylum ‘forms 1a and 1b’): [USA]
Adler, Currie & Wood (2004, chromosomal characters in larval key, formal name for Newman’s 1983 P. onychodactylum ‘sibling species 1a and 1b’, notes on geographic variation): USA
Helodon clavatus (Peterson)
Rothfels (1979, cytodendrogram, whole-arm interchange [IIL + IIIS, IIS + IIIL]): North America
Newman (1983, photo of hybrid complement with H. mccreadiei [as P. onychodactylum]; as P. albertense): USA
Helodon diadelphus Adler, Currie & Wood
Newman (1983, characterization, hybrid with H. mccreadiei [as P. onychodactylum ‘sibling species 7a/7b’], inversions mapped, photo of partial IIL, resolution relative to H. onychodactylus standard; as P. onychodactylum ‘sibling species 2b’): USA
Henderson (1985, characterization, cytodendrogram, demonstration of homosequential sibling species, inversions mapped, polymorphisms; as P. onychodactylum ‘form 2b’): Canada, USA
Henderson (1986a, characterization, cytodendrogram, inversions mapped, polymorphisms; as P. onychodactylum ‘form 2b’): Canada, USA
Henderson (1986b, characterization, demonstration of homosequential sibling species, polymorphisms; as P. onychodactylum ‘form 2b’): Canada, USA
Adler, Currie & Wood (2004, chromosomal characters in larval key, formal name for Newman’s 1983 P. onychodactylum ‘sibling species 2b’): North America
Helodon ferrugineus (Wahlberg)
Chubareva & Petrova (1969, comparisons with P. hirti pes and partially with H. rubicundus, description, idiogram, photos of metaphase and polytene complements): Russia
Chubareva (1977a, idiogram, low-resolution photo of polytene complement): Russia
Chubareva (1977b, 1977d, 1978b, idiogram, notes on general polytene features): Russia
Chubareva & Petrova (1979, notes on general polytene features, polytene chromosome lengths): Russia
Rothfels (1979, note on comparison with Helodon standard, whole-arm interchange [IIL + IIIS, IIS + IIIL]): Norway
Chubareva & Petrova (1999, notes on general polytene features, photos of metaphase and polytene complements, polytene chromosome lengths; as H. rufum): Russia
Chubareva & Petrova (2000b, note on general polytene features; as H. rufus): Russia
Chubareva & Petrova (2003, notes on general polytene features, polytene chromosome lengths; as H. rufus): Russia
Chubareva & Petrova (2004, notes on general polytene features, polytene chromosome lengths): Russia
Chubareva & Petrova (2008a, description, photos of metaphase and polytene complements, polytene chromosome lengths): Russia
Helodon irkutensis (Rubtsov)
Rothfels (1979, cytodendrogram, whole-arm interchange [IIL + IIIS, IIS + IIIL]; as H. martini and H. perspicuum): North America
Helodon kamtshaticus (Rubtsov)
Chubareva (1977 a, 1978b, notes on general polytene features): Russia
Chubareva (1977d, idiogram, notes on general polytene features): Russia
Chubareva & Petrova (1979, 2003, notes on general polytene features): Russia
Helodon mccreadiei Adler, Currie & Wood
Newman (1983, characterization, hybrids with H. chaos, H. diadelphus, H. newmani, and H. protus [as P. onychodactylum ‘sibling species 1a/1b’ ‘2b’, ‘5’, and ‘4a/4b’, respectively], inversions mapped, photo of IIIS and hybrid complement with H. clavatus [as P. albertense], resolution relative to H. onychodactylus standard; as P. onychodactylum ‘sibling species 7a and 7b’): USA
Henderson (1985, 1986a, characterization, cytodendrogram, inversions mapped; as P. onychodactylum ‘forms 7a and 7b’): [USA]
Adler, Currie & Wood (2004, chromosomal characters in larval key, formal name for Newman’s 1983 P. onychodactylum ‘sibling species 7a and 7b’, notes on geographic variation): USA
Helodon multicaulis (Popov)
Chubareva (1977a, notes on general polytene features, photos of metaphase and low-resolution polytene complements): Russia
Chubareva (1977d, 1978b, idiogram, notes on general polytene features): Russia
Chubareva & Petrova (1979, 2003, notes on general polytene features, polytene chromosome lengths): Russia
Chubareva & Petrova (1999, comparison with H. ferrugineus [as H. rufum], notes on general polytene features, photos of metaphase and polytene complements, polytene chromosome lengths): Russia
Chubareva & Petrova (2008a, description, photos of metaphase and polytene complements, polytene chromosome lengths): Russia
Helodon newmani Adler, Currie & Wood
Newman (1983, characterization, hybrids with H. beardi, H. mccreadiei, and H. onychodactylus [as P. onychodactylum ‘sibling species 3’ ‘7a/7b’, and ‘6’, respectively], inversions mapped, photo of IIIS, resolution relative to H. onychodactylus standard; as P. onychodactylum ‘sibling species 5’): USA
Henderson (1985, 1986a, characterization, cytodendrogram, inversions mapped; as P. onychodactylum ‘form 5’): [USA]
Adler, Currie & Wood (2004, chromosomal characters in larval key, formal name for Newman’s 1983 P. onychodactylum ‘sibling species 5’, notes on geographic variation and possible sibling species): USA
Helodon onychodactylus (Dyar & Shannon) s. str.
Newman (1970, initial evidence for sibling species, notes on sex chromosomes): USA
Rothfels (1979, cytodendrogram, whole-arm interchange [IIL + IIIS, IIS + IIIL]; as H. onychodactylum ‘B’): North America
Newman (1983, characterization, hybrids with H. newmani and H. susanae [as P. onychodactylum ‘sibling species 5’ and ‘2a’, respectively], inversions mapped, photos of mitotic and meiotic metaphase I and all polytene arms; as P. onychodactylum ‘sibling species 6’): USA
Henderson (1985, 1986a, characterization; cytodendrogram; inversions mapped; photos of all arms; polymorphisms; as P. onychodactylum ‘forms 6’ and ‘10’): Canada, USA
Adler, Currie & Wood (2004, chromosomal characters in larval key, recognition of Newman’s 1983 P. onychodactylum ‘sibling species 6’ as H. onychodactylus): North America
Helodon protus Adler, Currie & Wood
Newman (1983, characterization, hybrid with H. mccreadiei [as P. onychodactylum ‘sibling species 7a/7b’], inversions mapped, photo of IIIS, resolution relative to H. onychodactylus standard; as P. onychodactylum ‘sibling species 4a and 4b’): USA
Henderson (1985, 1986a, characterization, cytodendrogram, inversions mapped, polymorphisms, recognition as distinct species; as P. onychodactylum ‘forms 4a and 4b’): USA
Adler, Currie & Wood (2004, chromosomal characters in larval key, formal name for Newman’s 1983 P. onychodactylum ‘sibling species 4a and 4b’): USA
Helodon rubicundus Rubtsov
Chubareva & Petrova (1969, notes on general polytene features, partial comparison with H. ferrugineus, photo of IIIS polymorphism): Russia
Chubareva & Petrova (1979, 2003, notes on general polytene features): Russia
Helodon susanae (Peterson)
Rothfels (1979, cytodendrogram, whole-arm interchange [IIL + IIIS, IIS + IIIL]; as ‘ H. onychodactylum ‘A’): North America
Newman (1983, characterization, hybrid with H. onychodactylus [as P. onychodactylum ‘sibling species 6’], inversions mapped, resolution relative to H. onychodactylus standard; as P. onychodactylum ‘sibling species 2a’): USA
Henderson (1985, characterization, cytodendrogram, demonstration of homosequential sibling species, inversions mapped, polymorphisms; as P. onychodactylum ‘form 2a’): Canada, USA
Henderson (1986a, characterization, cytodendrogram, inversions mapped, polymorphisms; as P. onychodactylum ‘form 2a’): Canada, USA
Henderson (1986b, characterization, demonstration of homosequential sibling species, polymorphisms; as P. onychodactylum ‘form 2a’): Canada, USA
Adler, Currie & Wood (2004, chromosomal characters in larval key, notes on geographic variation, recognition of Newman’s 1983 P. onychodactylum ‘sibling species 2a’ as H. susanae): North America
Helodon trochus Adler, Currie & Wood
Henderson (1985, 1986a, characterization, cytodendrogram, inversions mapped, low-magnification photo of polytene complement, recognition as distinct species; as P. onychodactylum ‘form 8’): Canada
Adler, Currie & Wood (2004, chromosomal characters in larval key, formal name for Henderson’s 1986a P. onychodactylum ‘form 8’, notes on geographic variation): North America
description: Published as part of Adler, Peter H. & Crosskey, Roger W., 2015, Cytotaxonomy of the Simuliidae (Diptera): a systematic and bibliographic conspectus, pp. 1-139 in Zootaxa 3975 (1) on pages 11-14, DOI: 10.11646/zootaxa.3975.1.1, http://zenodo.org/record/192364