Quichuana mariliae Ricarte & Marcos-García & Hancock & Rotheray 2012, SP. NOV.
Creators: name: Ricarte, Antonio
name: Marcos-García, M. Ángeles
name: Hancock, E. G.
name: Rotheray, Graham E.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Syrphidae
Quichuana
Quichuana mariliae

description: QUICHUANA MARILIAE RICARTE & HANCOCK SP. NOV.   
FIGURES 47–50  
 Description   
Male  
 Head: Upper quarter of eyes with dense, brownish white hairs; occasional white hairs at the middle of the eyes; lower margin of eyes with sparse, straight, short black hairs dorsally; metatrochanter with a conspicuous process bearing apically a row of black setae (Fig. 50); metatibia inflated towards the apex, constricted ventrally and with an apical, short process bearing black, adpressed hairs (Fig. 49); metafemur with long yellow hairs, longer on anterior surface; a few short, spiny, black setae on ventral third of femora; metatarsus yellow, with black setae dorsally; wings extensively microtrichose, with small, bare areas in cells bc, C, and CuP; wings with light-brown pigmentation in cells SC and R 1, and in cells R 2+3 and R anteriorly; ventral calypteres white centrally, but black with short, black hairs in the margin; halteres whitish yellow apically, brown on the basal third.   
white hairs; vertical triangle with long, black hairs; a row of long, golden yellow hairs just posterior to the ocellar triangle; frontal triangle black, with long, anteriorly directed, golden yellow hairs laterally and scattered, long, black hairs centrally; antenna elongate, basoflagellomere 1.3 ¥ longer than pedicel and pedicel 2.7 ¥ longer than scape; ventral surface of frontal prominence yellowish laterally; face black, with lateral, white pollinosity delimiting a central, rhomboidal, shiny area; long, bright-yellowish white hairs on both the pollinose and shiny surfaces of face, except for a median, narrow, bare stripe; occiput with long, golden yellow hairs, shorter and paler when further down the occiput (upper section of occiput with a row of black setae intermixed with yellow hairs); from the midpoint of the occiput to the lower end, hairs progressively longer and sparser.  
 Thorax: Scutum with two grey pollinose stripes, nearly reaching the posterior margin of scutum; scutum with golden yellow hairs as long as those on scutellum; NP, PAPT posterodorsally, and PC with a tuft of golden yellow hairs; scutellum brownish black, with long, golden yellow hairs; proleg red, except for the black coxa and trochanter; profemur narrowly yellow apically; meso- and metaleg red, except for the black coxa, trochanter, and basal two-thirds of the femur; pro- and mesofemur and tibia with long, yellow hairs posteriorly; tibiae with black hairs dorsoapically (in mesoleg also ventro-apically); tarsi with Abdomen: All terga dull, bronzy black, except for shiny surface of terga I and II anteriorly; tergum I with a moustache arrangement of long, golden yellow hairs; tergum II only with yellow hairs, very long on the anterior section of lateral margins; tergum III only with golden yellow hairs; tergum IV with golden yellow hairs and scattered black hairs posteriorly; tergum V with both whitish yellow hairs and scattered black hairs; sternum I with long white hairs; sterna II and III with long yellowish white hairs; sternum IV with very few lateral hairs, both pale and black intermixed.   
 Genitalia: Superior lobes short and more or less conical; secondary processes of hypandrium developed with respect to those in the genitalia of species such as Q. angustiventris; surstyli slender, narrowed at the base, but widening towards the apex; base of the surstyli with a tooth in the interior side.   
Female  
 Same as male except upper fourth of eyes with sparse hairs; basoflagellomere 1.6 ¥ longer than pedicel, and pedicel 2.1 ¥ longer than scape (N = 4) (Fig. 47); ocellar triangle with black hairs; frons shiny black; anterior part of the frons with long, anteriorly directed, golden yellow hairs, and posterior part with long, posteriorly directed, golden yellow hairs; frons with an anteriorly directed chevron of pollinosity; scutum with two strong grey pollinose stripes reaching the posterior margin (Fig. 48); scutum with hairs shorter than those on scutellum; metatrochanter and metatibia without processes or constrictions; dorsoapical third of protibia darkened; basal half of mesofemur sometimes black; metafemur with long, white to yellow hairs, longer ventrally; metatibia with short, black hairs intermixed with yellow hairs; wing membrane weakly pigmented in cells bc and C, sometimes nearly hyaline; tergum II usually with scattered black, adpressed hairs posteriorly; posterior quarter of tergum III usually with black, adpressed hairs; tergum V with only whitish yellow hairs; shiny surface of tergum II smaller than that in male; sterna I–III with long white hairs centrally; sternum IV with yellowish-white hairs all over.   
 Etymology   
 The epithet ‘ mariliae ’ refers to ‘Marili’, the nickname for Dr M. Ángeles Marcos-García. This species is named for her because she collected some material and, in addition, encouraged me to undertake this revision (A Ricarte).   
 Material examined   
 Holotype: 1m, Costa Rica, Prov. Guanacaste, Rincón de la Vieja, Upala, Dos Ríos, San Cristóbal, 600– 620 m, 17.v.1998, leg. F.A. Quesada, en flor, L_N_ 318056383200#63528/INB0003341160 (INBio).   
 Paratypes: COSTA RICA: 1m with puparium, Las Flores, Alajuela, Upala, 2.ii.2008 (puparium 21.ii.2008, adult 2.iv.2008), ex liquid contained in a live bromeliad, leg. MAM (CR170); 1m with puparium, El Pilón, 700 m, ex bromeliad, larva 21.vi.09, puparium 17.vii.09, adult 25.vii.2009, leg. MAM (ref. 718), CEUA00089988; 2m with puparia, Guatuso, Finca Blanco, ex bromeliad, larva 17.vi.09, puparium 2.vii.2009 (1m) & 7.vii.2009 (1m), adult 2.vii.2009, leg. MAM (ref. 717 and 719), CEUA00089987 and 89989; 1m, Albergue Heliconias, Volcán Tenorio, 2.ii.2008, leg. MAM, collected with a light trap, CR181, CEUA00089984 (CEUA); 4f with puparia, Las Flores, Alajuela, Upala, ex live bromeliads, larva 2.ii.2008, puparium 13.ii.2008, adult 27.ii.2008 (CR172); puparium 14.ii.2008, adult 1.iv.2008 (CR168); puparium date not recorded, adult 14.iv.2008 (CR174) (for CR172 breeding site unknown), leg. MAM; 1f, Las Flores, Alajuela, Upala, ex bromeliad, larva 2.ii.2008, adult 1.iii.2008, leg. MAM (CR162) (CEUA); 1f, Prov. Alajuela, P.N. Volcán Tenorio, Sector El Pilón, 700–800 m, 29–31.vii.2003, L N 298212 427913#74561, leg. J. Azofeifa with light trap (INB0003743854) (INBio); 1f, Est. Pitilla, 700 m, 9 km S, Sta. Cecilia, P.N. Guanacaste, Prov. Guanacaste, 1 to 11.ix.1992, L-N 330200, leg. C. Moraga (INBio CRI 000849090) (INBio).   
 Range   
Costa Rica.  
 Taxonomic notes   
 Medium size species (10.2 mm, holotype) with slender abdomen; males of this species can be readily separated from males of any other species by the presence of a conspicuous process on the metatrochanter (Fig. 50), as well as by having the apical section of the metatibia inflated and then constricted ventrally, to end with a short but conspicuous process (Fig. 49); the long pedicel and basoflagellomere (bf = 2.3, holotype; for female, see Fig. 47) separate this species from similar species such as Q. angustiventris, Q. picadoi, or Q. quixotea, but females of Q. mariliae sp. nov. are readily distinguishable amongst females of species with a slender abdomen by the presence of two strong pollinose stripes reaching the posterior margin of the scutum (Fig. 48).
description: Published as part of Ricarte, Antonio, Marcos-García, M. Ángeles, Hancock, E. G. & Rotheray, Graham E., 2012, Revision of the New World genus Quichuana Knab, 1913 (Diptera: Syrphidae), including descriptions of 24 new species, pp. 72-131 in Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) (Zool. J. Linn. Soc.) 166 (1) on pages 99-101, DOI: 10.1111/j.1096-3642.2012.00842.x, http://zenodo.org/record/5408533