Gedanochila museisucini Brysz & Bourgoin & Szwedo 2022, sp. nov.
Creators: name: Brysz, Alicja Magdalena
name: Bourgoin, Thierry
name: Szwedo, Jacek
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Achilidae
Gedanochila
Gedanochila museisucini

description: Gedanochila museisucini sp. nov.
(Figs 1–5)
Etymology. Specific epithet derived from Greek museion —museum, and Latin sucinus —amber, and refers to the Museum of Amber in Gdańsk.
Type material. Holotype. Specimen No. MHMB-827, deposited in Museum of Amber—Department of Museum of Gdańsk, Gdańsk, Poland. Piece of Baltic amber, with inclusion partly covered by milky veil and other impurities, which cover part of the head, most of the abdomen and parts of legs. Paratype. Specimen No. GPIH-SBS-0424, deposited in Centrum für Naturkunde (CeNak), Geologisch-Paläontologisches Institut der Hamburg Universität. Piece of amber, with inclusion partly covered with milky veil ventrally, right tegmen on layer with some mineral impurities (markasite?), one proleg detached; syninclusions: Microlepidoptera; isolated leg of Diptera (?).
Diagnosis. As for genus and in addition: pronotum 3.5× as wide as long, tegmen with cell C5 slightly longer than cell C1; metatibio-tarsal formula LT2-10/Mt1-9(7)/Mt2-7(5); body dark, tegmina light, with regular colour pattern, with dark border at the tip and pterostigmal area, and two series of dots: three parallel to margin on costal cell, in the basal part of the wing and two on radial and median cells respectively, at basal ⅓ of tegmen.
Description. Coloration. Body dark; tegmina light, with regular colour pattern (Figs. 3A, 4 E-H, 5A), with dark border at the tip and pterostigmal area, and two series of dots: three parallel to margin on costal cell, in the basal part of the wing and two on radial and median cells respectively, at basal ⅓ of tegmen.
Head. Head with compound eyes narrower than pronotum, about 0.6 as wide as pronotum. Vertex 2.5× as wide as long in mid line. Anterior margin of vertex widely arcuately angulate, smoothly transiting to lateral margins, lateral margins diverging posteriad; posterior margin arcuately incised, reaching level of 1/3 of compound eye length; all margins carinate; disc of vertex depressed, flat, with median carina widened at base, not reaching half of vertex length (Figs 1A, 3A, 4 E-H, 4A, B). Frons in mid line 1.2× as long as clypeus, 1.2× as long as wide, subrectangular, with lateral margins slightly arcuate, slightly diverging ventrad; widest at level of antennal bases, laterally carinate; median carina distinct, reaching frontoclypeal suture. Frontoclypeal suture widely angulate. Postclypeus with distinct median carina, lateral carinae as prolongation of lateral margins of frons, converging ventrad, but not fused, anteclypeus about 2× as long as wide at base (Figs. 1B, C, 4A, 5D). Rostrum reaching slightly behind base of metacoxae, apical segment longer than subapical one. Suture between gena and loral plate oblique. Compound eye with ventral incision, without subocular callosity. Lateral ocellus placed at level of lower margin of compound eye. Base of antenna below compound eye, scapus short, pedicel truncately subglobose, base of flagellum slightly inflated, flagellum about 4× as long as pedicel (Figs. 1B, C, 2B, 3A, 5C, D).
Thorax. Pronotum 3.5× as wide as long; disc in shape of depressed arch, elevated, not concave; lateral carinae not reaching posterior margin; posterior margin elevated, incised with an angle ~120 degrees. Mesonotum 1.2× as wide as long, slightly humpy anteriorly, then sunken after half of its length, scutellum depressed; median carina not entering scutellum, lateral carinae full and straight (Fig. 1A, 3A, 4 E-H, 5A, B)).
Metatibia with single lateral spine at ⅔ of its length and with 10 apical teeth (6 in arcuate row and 4 based slightly above them). Basimetatarsomere with row of 9 apical teeth, basimetatarsomere distinctly (ca. 1.5×) longer that combined length of mid- and apical metatarsomeres; midmetarsomere with row of 7 apical teeth; basi- and midmetatarsomeres bearing subapical setae, except lateral teeth. Metatibiotarsal formula LT2-10/Mt1-9(7)/Mt2- 7(5) (Fig. 1D, 4 B-D, 5E).
Tegmina and wings. Tegmina 3× as long as wide; costal margin (fused veins Pc+CP+CA) curved at base than straight, strengthened at base, basal portion of Pc+CP carinate; anteroapical angle acutely rounded, posterior margin obliquely convex, posteroapical angle widely angulate, tornus merely convex; angle between A 2 and tornus ca. 160°. Basal cell narrow, about 4× as long as wide, tapered apicad. Longitudinal veins elevated, stem ScP+R+MP leaving basal cell with a short common stalk, about ½ length of basal cell, stem ScP+R forked at about 1/3 of tegmen length, slightly basad to CuA fork, basad of claval veins junction; single prenodal branching (forked on left tegmen); branch ScP+RA with 5-6 terminals; branch RP with 3 terminals; branch MP with 7 terminals. Claval veins connecting at the level of ScP+R fork, slightly basad to CuA fork. Veinlet mp 4 -cua vein basad to rp-mp 1 r one. Basal cell 4× as long as wide; cell C1 more or less even in its width, curved; cell C5 lanceolate, with widened apical part. Cells C1/C3 length ratio 1.58; cells C1/C5 length ratio 0.98; apical line parallel to wing margin in apical part, then convex; apical cells narrow. Membrane covering about 40% of tegmen’s length (Fig. 2A, B; 3A, B, 4 E-H, 5A, C).
Venation differs in both tegmina—left tegmen: branch RA with 7 terminals (6 terminals on right tegmen); RP with 4 terminals (3 terminals on right tegmen), branch MP 1+2 forked slightly apicad of branch MP 3+4 forking (branch MP 1+2 forked much more basad of branch MP 3+4 forking on right tegmen); branch MP 4 forked twice, resulting in 8 terminals of MP, with branching MP 4b intercepted by terminal CuA 1 in apical section (7 terminals of MP on right tegmen); cell C3 closed with double crossvein near its base, at level of nodal line, cell C3a with crossvein in the middle, cell C4’ tapering and divided by a crossvein in apical part, cell C5 with one complete crossvein on the narrowing and one incomplete in basal part; right tegmen: RA with 5 terminals, additional recurrent vein between MP and CuA stems in cell C4, cell C1 with oblique additional veinlet before half of its length (Fig. 2A, B; 3A).
Hind wing. Anteroapical angle widely angulate. ScP+RA single reaching margin distinctly basad of apex, branch RP vein with 2 terminals (3 terminals on the left hindwing), reaching margin basad of apex; stem MP with 3 terminals (MP 3+4 not branched), terminal MP 2 forked before margin on left hind wing; branch CuA with 4 terminals, CuA 1 forked, CuA 2 single, CuP not branched. Veinlet rp-mp long, reaching stem MP slightly basad of MP forking, veinlet mp-cua 1 merely basad of rp-mp veinlet, in line with stalk of CuA 1b. Anal region not visible (Fig. 2C, D; 3A, B. 5A, C).
Age and occurrence. Lutetian, Eocene, Baltic amber, Gulf of Gdańsk area, secondary deposit. For the discussions on age of the Baltic amber and its deposits see e.g., Szwedo & Drohojowska (2016), Wolfe et al. (2016), Grimaldi and Ross (2017), Bukejs et al. (2019), Sadowski et al. (2017, 2020).
description: Published as part of Brysz, Alicja Magdalena, Bourgoin, Thierry & Szwedo, Jacek, 2022, Spotted beauty-Gedanochila museisucini gen. et sp. nov. - a new Achilidae from Eocene Baltic amber (Hemiptera: Fulgoromorpha) and its relation to Achilini, pp. 295-312 in Zootaxa 5125 (3) on pages 298-300, DOI: 10.11646/zootaxa.5125.3.3, http://zenodo.org/record/6443793