Stenonartonia mimica Kohl 1907, comb. nov.
Creators: name: Garcete-Barrett, Bolívar R.
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Eumenidae
Stenonartonia
Stenonartonia mimica

description: Stenonartonia mimica (Kohl 1907), comb. nov.
(Figs 4, 16, 17, 40, 41, 71, 84, 91, 102, 116, 130, 143, 154, 179)
Nortonia mimica Kohl 1907: 248, 247 [holotype, Ƥ, Brazil: Rio Grande do Sul (NMW), examined]. Paranortonia mimica: Giordani Soika 1941: 159.
Diagnosis and comments. This species is outstanding in the group by being the only one with a small and exposed anterior axillary fossula as well as: parategula narrow, sharp above; propodeum with sides regularly convex in dorsal view, with a moderately well differentiated medial channel devoid of reflective pilosity. The propodeal valvula is separated from the apical propodeal lamella by a blunt crease and is hardly dislocated from it along the margin. They are small wasps with brown and partly amberish wings and a color pattern reminiscent of either typical Agelaia multipicta (Haliday) [pale form] or A. vicina (de Saussure) [melanic form].
Description. FEMALE. Color pattern (pale form). Black integument with diffuse dark reddish brown areas as follows: mandibular margins; propodeal sides, disc and sides of the declivity of T1; base of T2; sides of S2; thin basal suffusion on yellow bands of metasomal segments 3–5; metasomal segment 6. Yellow markings as follows: disc of mandible; scape ventrally; broad lateral line on clypeus; inner orbit from clypeal border to the level of the hind ocelli; frontal line from between the antennal sockets to the mid ocellus; whole genae from almost the mid vertex down to mandible and extending to the hypostomal region; thin anterior band and posterior margin of pronotum including lateral lobe; thick humeral line; anteriorly outcurving submedial mesonotal line; tegula except for a light brown medial spot; large lateral scutellar blotch; thick metanotal band; mesepimeron and posterior two thirds of mesepisternum (except for the sulci); most of metapleura; large submedial propodeal spot below; mid propodeal carina partly; submarginal carina at sides; whole anterior leg and mid and hind tarsi; mid and hind coxa largely; anterior face of mid and hind femora (the latter with a large darkening at middle); whole mid and hind tibia except for a small inner subapical darkening; hind and lateral margin of T1; whole S1; apical band on metasomal segments 2–5 (thinner on terga); medial spot on metasomal segment 6. Ventral face of flagellum brown, gradually lighter toward apex. Fore wing membrane light amberish brown, more intense along the costal region. Fore wing venation light amberish brown, turning brown toward the apex.
Measurements and proportions. Body length 9.1 mm. Wing length 9.4 mm. Clypeus 1.1 × wider than high and with its apical margin 0.9 × as wide as the interantennal distance. T1 as wide as long and 0.7 × as wide as T2. S1 equilateroid, 2.25 × wider than its medial length and 1.55 × wider than its total length.
Structure. Clypeus cordiform, moderately convex, with sides of its free part almost straight, with a slight apical depression and with a small but definite apical emargination between two rather blunt, hardly carenate, down pointing teeth. Humeral angle weak, obtuse in frontal view. Pronotal carina almost straight below the humeral angle. Parapsidal line very weak. Parategula sickle-shaped, careniform above. Scutellum slightly convex. Anterior axillary fossula small, shallow and exposed behind the parategular emargination. Lateral axillary panel poorly produced mesoposteriorly. Metanotum almost right angled in side view and with a well marked though low toothed ridge. Propodeum semi oval, regularly convex and almost not angled at sides as seen from above (there is no crease between the lateral and sublateral surfaces). Mid propodeal channel rather shallow but distinguishable and strongly narrowed above. Mid propodeal carina complete. Apical propodeal lamella projecting backward and separated from the valvula by a blunt longitudinal crease, both hardly dislocated along the margin. T1 conical, smoothly convex above, with a well marked transverse carina projecting backward at the middle and curving backward at the level of the spiracle.
Tegumental sculpture. Clypeus subshiny, covered with thin reticulation, very sparse and shallow micropunctures and moderately spaced macropunctation. Head and mesosomal sculpture differing from S. apicipennis in: overall surface much less opaque due to a sparser and thinner micropunctation; anterior metanotal band better defined, very densely and confluently macropunctate; tegula subshiny, covered with moderately spaced micropunctation all over; metapleura with just a few thin macropunctures below; lateral propodeal surface more sparsely macropunctate; sublateral propodeal surface transversely shagreened and covered with dense and moderately thin macropunctation which is gradually sparser toward the spiracle; mid propodeal channel sub opaque, densely micropunctate. Metasoma subshiny, covered with moderately dense micropunctation which is sparser at sterna. T1 with thin macropunctures scattered over its anterior declivity and moderately sparse on its disc. T2 covered with thin sparse macropunctures becoming coarser but not much denser mesoposteriorly. S1 coarsely and densely macopunctate. S2 covered with scattered macropunctures which become coarser and denser mesoposteriorly. Metasomal segment 3–5 covered with moderately dense macropunctures. T6 with a few scattered thin macropunctures. S6 without macropunctures.
Pilosity. Body covered with a short, sparse and semidecumbent to semi erect brownish pubescence which becomes lighter and appressed on the thoracic sides and the propodeum and is appressed and thinner on metasoma. Posterior propodeal declivity covered with non-reflective upward and outward pointing pilosity.
MALE. Unknown.
Variation. The holotype is the only know individual with the pale pattern. The remaining of the examined material show a melanic pattern differing as follows: most of the yellow markings are rather orange tinged (on mandible, clypeus and legs) or ferrugineous (on antennae, genae and mesosoma except for the pronotum). Many markings show reduction as follows: clypeal line broken at middle or present just below; orbital line reaching only up to the ocular sinus, an isolated upper spot can also exist; frontal line reduced to an interantennal spot and sometimes also a mid frontal spot; anterior pronotal band absent; posterior pronotal margin interrupted at middle; humeral line diffuse, almost inexistent; mesoscutal line very thin and vanishing far from the pronotal margin; scutellar blotch small and dark or absent; T1 with diffuse lateral markings and very thin posterior band; remaining of metasoma with very thin and medially interrupted bands, those on T2 and beyond S3 rather ferrugineous. The melanic form is reported from places above 300 meters above the sea level. The pale form probably comes from lesser altitudes, as also seems to happen with S. flavotestacea and the social wasp Polybia fastidiosuscula de Saussure.
Material examined. BRAZIL: Paraná: Londrina, 4.x.1985 (Exc. Dep. Zool.) [1 Ƥ: DZUP]; Rio Grande do Sul: without precise locality (Stieglmayr) [1 Ƥ Holotype: NMW]; Santa Catarina: Nova Teutônia, 300–500 m, 27°11'S, 52°23'W, ii.1965 (F. Plaumann) [1 Ƥ: UCD]; São Paulo: Cajuru, Fazenda Rio Grande, 21°12'S, 47°09'W, 2–18.xii.1999 (G.A.R. Melo & Nascimento) [2 Ƥ: DZUP].
Distribution. Southern and South Eastern Brazil. Probably mostly in highlands.
description: Published as part of Garcete-Barrett, Bolívar R., 2011, A revision of the genus Stenonartonia Giordani Soika 1973 (Hymenoptera: Vespidae: Eumeninae), pp. 1-50 in Zootaxa 2868 on pages 16-18, DOI: 10.5281/zenodo.277490