Clivina spatulifigura Balkenohl, 2025, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina spatulifigura

description: Clivina spatulifigura sp. nov.
(Figs 23, 58, 91, 117)
Type material. Holotype: ♂, with labels and data: yellow, black printed “ BORNEO: SABAH Bingkor N Keningau 400-500m, 19.-20.XI.1996 leg. W. SCHAWALLER“ / “ Clivina group helferi nr. wallacei PUTZ. det. Balkenohl, 1998” (SMNS). Paratypes: 1 ♀, 1 ♂, same data as holotype; one of the paratypes with additional information “at light” (SMNS, CMBB); 1 ♂, “ BORNEO, SABAH, C., Telupid Miruru, 25 km 8.v.1999. Z.Smrź leg.” / “helferi-Gr.” (CADW); 1 ♂, “BRUNNEI-Temburong Belalong Field Res. C. Mix.Dipterocarp. For. ~ 100m, 2-8.5.95 Heiss ” (CADW).
Remark. In the female paratype, the right front and intermediate leg is missing.
Description. Measurements. Body length 4.91–5.05 mm (ẍ = 4.99 mm *), width 1.38–1.39 mm (ẍ = 1.39 mm *), ratio L/W of pronotum 0.98–0.99 (ẍ = 0.98), ratio L/W of elytra 1.91–1.95 (ẍ = 1.94*); (*n = 5). Colour glossy, fuscous. Supraantennal plate and clypeal-wing at margin slightly translucent. Antennae, mouthparts, intermediate and hind legs yellowish.
Head about a fifth narrower than pronotum. Clypeus with central part straight, projection of lateral teeth distinct, separated from clypeal-wing by distinct notch; wing rounded, distinctly less high as central part of clypeus, separated from supraantennal plate by obtuse-angled notch, supraantennal plate slightly elongated, with margin convex at middle, clypeus, wing and supraantennal plate reflexed margined. Supraantennal plate extended posteriorly into a blunt keel, separated from wing by small furrow. Clypeus transverse, convex, smooth, clypeal-furrow wide, flat, with small punctures, frons moderately convex, with pore at middle, separated from supraantennal plate and supraorbital carina by flat furrow, with clypeal seta in front; supraorbital carinae moderately wide, blunt, with two supraorbital setigerous punctures at middle of eye and at middle of gena-level. Neck constriction closed, developed as step from frons, with indistinct punctures posteriorly. Eye prominent, gena small. Antenna elongate, reaching just over base of elytron. Labrum excised anteriorly, irregularly reticulated, seven-setose. Lobe of mentum slightly convex laterally, rectangular acuminated at tip, medially straight, with fine isodiametric reticulated, median tooth as high as lobe, wide, hollowed out anteriorly.
Pronotum (Fig. 58) with disk indistinctly convex in lateral view, in frontal view moderately convex, flattened at median line. Shape square to sub-cordate, as long as wide. Reflexed lateral margin smooth, widest at beginning of posterior third, nearly straight in anterior half but slightly converging; anterior and posterior angles distinct, posterior one with distinct projecting somewhat rounded tooth; lateral channel narrow, reticulated, continuing up to base as sharp, convex margin. Median line complete, engraved at middle, joining and surpassing anterior transverse line. Anterior transverse line distinct, complete. Disk with few scattered punctures, with transverse wrinkles, with isodiametric reticulation at base, with indistinct impression in basal third, with posterior setigerous puncture situated beside tooth of posterior angle. Lateral margin from posterior angle to flange of base slanted with angle of around 40°. Basal channel on flange regularly developed, rugose, with sharp carina.
Elytron (Fig. 23) with disk flattened in anterior half in lateral view, convex in frontal view. Shape of elytra elongate, around two times longer than wide, with lateral margin nearly straight at middle, slightly diverging posteriorly, maximum width posterior middle. Humerus distinct, rounded. Reflexed lateral margin slightly crenulate behind humerus (50-fold). Row of setigerous punctures in lateral channel regular in size, slightly more distant at middle, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, nearly impunctate, one to four free at base, five and six joining humerus, one and two ending free at apex, three and four and five and six joining apically. Intervals moderately convex, convexity more distinct laterally, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven with step-like carina at apex, eight carinate at apex. Surface of all intervals with distinct isodiametric reticulation already well visible at 40-fold.
Hind wing fully developed.
Lower surface. Proepisternum with fine isodiametric reticulation, with fine transverse wrinkles. Sternite of abdomen with regular isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with slight flattened projection posteriorly at middle, with narrow marginal furrow at middle (in both sexes).
Legs. Protibia moderately developed, with longitudinal reticulation, with wide sulcus dorsally, with three apically rounded spines and indistinct tubercle, movable spur moderately arcuate, basal tarsomere slender at base, carinate, with acute tubercle with two setae. Intermediate tibia with eight tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front legs slightly widened.
Male genitalia (Fig. 91). Median lobe regularly arcuate in basal half, nearly straight in apical half, slightly bent to apex. Apex formed as elongate spatula. Parameres moderately slender, with small apophyses, slightly distorted, with flat hyaline elongation at apex, with two setae apically.
Female gonocoxites and epipleurite (Fig. 117). Basal gonocoxite with apex curved laterally into a tubercle with long seta, medially with two long and two medium sized setae, apical gonocoxite elongated, curved in basal third, oval in cross section, with one small seta in basal third and one long seta at end of apical third, with two long setae apically. Epipleurite trapezoid, with one seta.
Variation. On the head and pronotum the density of punctures varies slightly. In one of the paratypes, the labrum is five-setose due to a gap of punctures on the left side. In the specimen from Temburong the reticulation at middle of the elytron is slightly less expressed.
Distribution. Known from Sabah and Brunei on the island of Borneo.
Diagnosis. A small to medium sized fuscous species with sub-cordate pronotum, all intervals of the elytron reticulated, and with seven-setose labrum. The most similar species is C. hartmanni sp. nov. from Irian Jaya also with isodiametric reticulation on all intervals of the elytron. In C. spatulifigura sp. nov. however, the main differences are as follows: the pronotum is as long as wide, the clypeus with its teeth, wings and notches are distinctly developed, there is a distinct clypeal-furrow between the clypeus and the frons, the neck constriction is closed, and the eyes are more prominent. In addition, the male and female genitalia are different.
Etymology. The name refers to the form (Latin: forma = form) of the male aedeagus with its distinctly elongated spatulate apex (Latin: spatula = spatula, scoop).
description: Published as part of Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), pp. 1-70 in Zootaxa 5592 (1) on pages 38-39, DOI: 10.11646/zootaxa.5592.1.1, http://zenodo.org/record/14953674