Campsicnemus spectabulus Evenhuis, sp. nov.
Creators: name: Evenhuis, Neal L.
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Dolichopodidae
Campsicnemus
Campsicnemus spectabulus

description: Campsicnemus spectabulus Evenhuis, sp. nov.
(Figs. 5, 10, 11, 15, 19)
Campsicnemus fumipennis: Williams, 1938a: 120, 1940: 295; Smith, 1952: 430; Wirth & Stone, 1956: 450; Hardy & Kohn, 1964: 92; Tenorio, 1969: 3; Evenhuis, 1996: 55; Anonymous, 2009: 80. Misidentifications.
Diagnosis. Similar to C. fumipennis in wing and abdominal patterning but can be easily distinguished by the characteristic long hairs of the male mid tibia (Fig. 19) (much shorter in C. fumipennis) and the predominantly yellowish brown laterotergite (predominantly dark brown in C. fumipennis).
Description. Male: Body length: 3.5–5.0 mm. Wing length: 4.0–5.0 mm. Head: Black, face, front and clypeus yellow, face and clypeus silvery golden tomentose; oc and vt black, about two-thirds length of antennal arista; occiput, and vertex black with blue-green to brassy highlights; postgena with long black hairs; face constricted at middle, holoptic for a length of 6 ommatidia; palpus small, brown; proboscis brown, extending below eye in lateral view; antenna (Fig. 5) with scape and pedicel yellow, postpedicel brown to gray-brown; scape flared apically, length ca. 2X greatest width; pedicel subspherical, with ring of short spiky black setae subapically; postpedicel long, lanceolate, length about 2.2–2.5X width, acute apically; arista slightly longer than head height. Thorax: Mesonotum yellowish brown, scutellum brown; pleura yellow except anepimeron dark brown, laterotergite with spot of brown laterally; dorsum of mesonotum with pair of brown admedian vittae (cf. Fig. 10), each vitta narrow anteriorly, flaring posteriorly to tear-drop shape near prescutellar area; thoracic setae very long, black: 1 + 4 dc; 2 np; 1 + 2 ph; 1 pa; 1 + 1 + 1 sc; 5–6 ac, ending at middle of mesonotum; halter stem and knob yellowish white. Legs: CI yellowish, spot of brown at extreme base; coxae with normal setation, CII brown, CIII yellow with some brown on posterior margin; fore and hind legs unmodified and without MSSC; FI and FII yellowish brown, FIII yellowish brown with darker brown at apex, without MSSC; TiII and TiIII yellow, remainder of legs brown; FII with 2 rows of 15–18 black setae along ventral surface in middle (MSSC), setae absent at apex and toward base; TiII (Fig. 19) very slightly sinuous, with long, slightly curved, black hairs along two-thirds of lateral surface (MSSC), mesal surface with short, stiff black spiky setae on basal half, longer curved hairs on apical half, becoming appressed on apical fourth (MSSC), apex with pair of strong long black setae on mesal surface. IIt1 slightly longer than IIt2, without MSSC. Remaining leg segments unmodified and without MSSC. Wing (Fig. 15): Subhyaline, with brownish infumation surrounding apical one-half to one-third of R2+3 and R4+5 and coalesced between those two veins to wing margin; infuscation faint along distal section of M1, cloud of infuscation on M1 at posterior crossvein; infuscation also along CuA1 just before, at, and after junction with posterior crossvein and posterior crossvein clouded with brown; posterior crossvein longer than distal section of CuA1. Abdomen (cf. Fig. 10): Dark brown dorsally with yellow laterally and posterolaterally, yellow color most extensive on basal tergites, becoming less extensive on succeeding tergites; sternites with brown medially and basally, otherwise yellow; tergal vestiture black, strongest along posterior margin of tergite I. Hypopygium grayish brown with pale brown cerci, not dissected. Female: As in male except for lack of MSSC and the following: eyes holoptic, separated at medial constriction by width of one ommatidium; antennal postpedicel subtriangular, length subequal to width, rounded apically; all brown areas of thorax darker and broader or more extensive than in males; TiII brown; wing with darker infuscation than male; abdominal coloration (Fig. 10) more contrasting than in male.
Types. HOLOTYPE 3 (BPBM 16,663) and 13 paratype from HAWAIIAN ISLANDS: O‘ahu: Lulumahu Canyon, 4 Apr 1937, on banana stem, F.X. Williams (BPBM). Other paratypes: O‘ahu: 131Ƥ, same pin, Lulumahu Str, Konahanui, 1800 ft [550 m], 20 Sep 1936, banana grove, F.X. Williams (BPBM); 13, same data (HDOA); 13, same data except 29 Sep 1836 (HDOA); 13, 1Ƥ, same pin, Lulumahu Str, 1800 ft [550 m], 27 Sep 1936, F.X. Williams (HDOA); 1Ƥ, same data (HDOA); 13, Lulumahu, 1750 ft [530 m], 21 Feb 1937, F.X. Williams (HDOA); 13, Lulumahu Val, 1900 ft [580 m], 11 Oct 1936, on banana stem, F.W. Williams (HDOA); 13, same data except, in banana grove (HDOA); 23, same data except 4 Oct 1936, on banana stem (HDOA); 23, on same pin [one only thorax and abdomen remaining], same data except 27 Sep 1936 (HDOA); 33, Lulumahu Val, 28 Mar 1937, on banana stem, F.X. Williams (HDOA); 13, Lulumahu Val, 1850 ft [564 m], 8 Nov 1936, on banana stem, F.X. Williams (HDOA); 1Ƥ, Lulumahu Val, 21 Feb 1937, on banana stem, F.X. Williams (HDOA); 13, Kaluanui Val, 2000 ft [610 m], 18 Oct 1936, on banana stem, F.W. Williams (HDOA); 13, 1Ƥ, Castle Trail, 11 Apr 1970, reared ex rotten bark of Charpenteria, S.L. Montgomery (UHM); 1Ƥ, same data except reared ex rotten bark of Tetraplasandra (UHM); 23, Castle Trail, 15 Mar 1970, reared ex Clermontia bark, S.L. Montgomery (HUM); 2 3, same data except reared ex rotten fruit of Pritchardia (UHM); 1Ƥ, Makaleha, 23 Jan 1970, reared ex Pisonia leaves, fed on Dettopsomyia [Drosophilidae], S.L. Montgomery (UHM); 1Ƥ, S. Kahana Val, 31 May 1970, reared ex Tetraplasandra bark, S.L. Montgomery (UHM); 13, Hidden Val, S. Kahana, 31 May 1970, reared ex Tetraplasandra bark, S.L. Montgomery (UHM); Kaluaa Pu‘u Hapapa, 15 Feb 1970, reared ex fern frond, S.L. Montgomery [with associated pupal case] (UHM); 13, Pu‘u Kaua, 13 Aug 1972, reared ex Pisonia stem, S.L. Montgomery (UHM); 1Ƥ, Pia, Niu V[alley], 1400 ft [425 m], 23 Jan 1971, reared ex Touchardia bark w/ Drosophila, S.L. Montgomery (UHM); 1Ƥ, Palehua, 2200 ft [670 m], 18 Apr 1970, reared ex koa sap flux, S.L. Montgomery (UHM); Mt Tantalus, 9 Oct 1963, M.R. Wheeler (UHM).
Etymology. The specific epithet derives from a combination of the Latin, spectabilis = “showy, notable” + fabulous, from the Latin fabulor = “story, tale”, in reference to the fabulous and showy wing and abdominal pattern and strikingly hirsute male mid tibia.
Remarks. The biology of this species [as “ C. fumipennis ”] and its immature stages were described in detail by Williams (1938) based on specimens observed and collected at Lulumahu Stream. Rearing records of immatures are shown in Table 1. One prey record is found on one label from a reared immature feeding on a species of the drosophilid Dettopsomyia at Makaleha, O‘ahu.
description: Published as part of Evenhuis, Neal L., 2012, Review of the Campsicnemus fumipennis group (Diptera: Dolichopodidae) in the Hawaiian Islands, with descriptions of new species and corrections of misidentifications, pp. 1-16 in Zootaxa 3497 on pages 11-15, DOI: 10.5281/zenodo.208996