cometadata/datacite-rtg-text-other-reclassification-records huggingface dataset:
Dendrohyrax interfluvialis Oates & Woodman & Gaubert & Sargis & Wiafe & Lecompte & Dowsett-Lemaire & Dowsett & Gonedelé Bi & Ikemeh & Djagoun & Tomsett & Bearder 2022
Creators: name: Oates, John F
name: Woodman, Neal
name: Gaubert, Philippe
name: Sargis, Eric J
name: Wiafe, Edward D
name: Lecompte, Emilie
name: Dowsett-Lemaire, Françoise
name: Dowsett, Robert J
name: Gonedelé Bi, Sery
name: Ikemeh, Rachel A
name: Djagoun, Chabi A M S
name: Tomsett, Louise
name: Bearder, Simon K
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Mammalia
Hyracoidea
Procaviidae
Dendrohyrax
Dendrohyrax interfluvialis

description: Dendrohyrax interfluvialis
Ghana. Kalakpa Resource Reserve: 6°23′N, 0°19′E to 6°28′N, 0°25′E (F.D. -L., R.D., 2008; S.B., J.O., 2016) (N = 23). Nyagbo Anyigbe: 6°47′N, 0°22′E (S.B., J.O., 2016) (N = 13). Biakpa: 6°50′N, 0°24′E (S.B., J.O., 2016) (N = 45).
Benin. Lama Forest Reserve: 6°59′N, 2°05′E (F.D. -L. & R.D., 2009) (N = 1). Lokoli: swamp forest 07°03′N, 2°15′E (F.D. -L. & R.D., 2009) (N = 1). Niaouli: 6°44′N, 2°08′E (F.D. -L. & R.D., 2009) (N = 1). Pahou Forest Reserve: 6°23′N, 2°09′E (F.D. -L., 2015) (N = 1).
Nigeria. Omo Forest Reserve: 6°55′N, 4°19′E (J.O., 1992) (N = 1). Okomu National Park: 6°19′N, 5°21′E (S.B., J.O., 2009) (N = 41).
APPENDIX 2
description: Published as part of Oates, John F, Woodman, Neal, Gaubert, Philippe, Sargis, Eric J, Wiafe, Edward D, Lecompte, Emilie, Dowsett-Lemaire, Françoise, Dowsett, Robert J, Gonedelé Bi, Sery, Ikemeh, Rachel A, Djagoun, Chabi A M S, Tomsett, Louise & Bearder, Simon K, 2022, A new species of tree hyrax (Procaviidae: Dendrohyrax) from West Africa and the significance of the Niger-Volta interfluvium in mammalian biogeography, pp. 527-552 in Zoological Journal of the Linnean Society 194 (2) on page 551, DOI: 10.1093/zoolinnean/zlab029, http://zenodo.org/record/6352101
Dendrohyrax J. E. Gray 1868
Creators: name: Oates, John F
name: Woodman, Neal
name: Gaubert, Philippe
name: Sargis, Eric J
name: Wiafe, Edward D
name: Lecompte, Emilie
name: Dowsett-Lemaire, Françoise
name: Dowsett, Robert J
name: Gonedelé Bi, Sery
name: Ikemeh, Rachel A
name: Djagoun, Chabi A M S
name: Tomsett, Louise
name: Bearder, Simon K
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Mammalia
Hyracoidea
Procaviidae
Dendrohyrax

description: SKULLS OF DENDROHYRAX EXAMINED FOR THIS STUDY
For institutional abbreviations, see Material and Methods.
description: Published as part of Oates, John F, Woodman, Neal, Gaubert, Philippe, Sargis, Eric J, Wiafe, Edward D, Lecompte, Emilie, Dowsett-Lemaire, Françoise, Dowsett, Robert J, Gonedelé Bi, Sery, Ikemeh, Rachel A, Djagoun, Chabi A M S, Tomsett, Louise & Bearder, Simon K, 2022, A new species of tree hyrax (Procaviidae: Dendrohyrax) from West Africa and the significance of the Niger-Volta interfluvium in mammalian biogeography, pp. 527-552 in Zoological Journal of the Linnean Society 194 (2) on page 551, DOI: 10.1093/zoolinnean/zlab029, http://zenodo.org/record/6352101
Dendrohyrax dorsalis subsp. sylvestris
Creators: name: Oates, John F
name: Woodman, Neal
name: Gaubert, Philippe
name: Sargis, Eric J
name: Wiafe, Edward D
name: Lecompte, Emilie
name: Dowsett-Lemaire, Françoise
name: Dowsett, Robert J
name: Gonedelé Bi, Sery
name: Ikemeh, Rachel A
name: Djagoun, Chabi A M S
name: Tomsett, Louise
name: Bearder, Simon K
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Mammalia
Hyracoidea
Procaviidae
Dendrohyrax
Dendrohyrax dorsalis
Dendrohyrax dorsalis sylvestris (temminck,

description: Dendrohyrax dorsalis sylvestris (Temminck,
1855) (N = 12)
Western population (N = 2). Liberia. Nimba County: Mt. Nimba: 7°04′N, 8°25′W (NHMUK 70.76). Western Rivercess County: 5°53′N, 9°14′W (RMCA 35862).
Eastern population (N = 10). Côte d’Ivoire. Adiopodoumé: 5°20′N, 4°08′W (AMNH M-239584). Ghana. Western Province: Papasi: 6°35′N, 2°55′W (NHMUK 76.1782). Ashanti Province: Mampong: 7°03′N, 1°25′W (NHMUK 35.10.22.152, 35.10.22.153). Eastern Province: Oda: 5°56′N, 1°01′W (NHMUK 46.481; MCZ 42709; FMNH 54448, 54679, 54681, 62807).
description: Published as part of Oates, John F, Woodman, Neal, Gaubert, Philippe, Sargis, Eric J, Wiafe, Edward D, Lecompte, Emilie, Dowsett-Lemaire, Françoise, Dowsett, Robert J, Gonedelé Bi, Sery, Ikemeh, Rachel A, Djagoun, Chabi A M S, Tomsett, Louise & Bearder, Simon K, 2022, A new species of tree hyrax (Procaviidae: Dendrohyrax) from West Africa and the significance of the Niger-Volta interfluvium in mammalian biogeography, pp. 527-552 in Zoological Journal of the Linnean Society 194 (2) on page 551, DOI: 10.1093/zoolinnean/zlab029, http://zenodo.org/record/6352101
Dendrohyrax dorsalis subsp. nigricans
Creators: name: Oates, John F
name: Woodman, Neal
name: Gaubert, Philippe
name: Sargis, Eric J
name: Wiafe, Edward D
name: Lecompte, Emilie
name: Dowsett-Lemaire, Françoise
name: Dowsett, Robert J
name: Gonedelé Bi, Sery
name: Ikemeh, Rachel A
name: Djagoun, Chabi A M S
name: Tomsett, Louise
name: Bearder, Simon K
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Mammalia
Hyracoidea
Procaviidae
Dendrohyrax
Dendrohyrax dorsalis
Dendrohyrax dorsalis nigricans (peters,

description: Dendrohyrax dorsalis nigricans (Peters,
1879) (N = 11)
Nigeria. Rivers State: Odieke: 5°01′N, 6°26′E (NHMUK 1998.298, 1998.299). Cross River State, Oban: 5°13′N, 8°33 ′E (NHMUK 12.10.28.53). Cameroon. Edéa: 3°48′N, 10°07′E (AMNH M-69542); Mbonge: 4°33′N, 9°03′ E (NHMUK 60.56). Equatorial Guinea. Rio Muni: uncertain locality (USNM 598583); Benito (Mbini) River: 1°51′N, 10°13′E (NHMUK 99.4.6.40). Gabon. Estuaire: Kango: 0°10′N, 10°07′E (AMNH M-119958, M-119961, M-119962). Republic of the Congo. Komono: 3°19′S, 13°15′E (AMNH M-119959).
description: Published as part of Oates, John F, Woodman, Neal, Gaubert, Philippe, Sargis, Eric J, Wiafe, Edward D, Lecompte, Emilie, Dowsett-Lemaire, Françoise, Dowsett, Robert J, Gonedelé Bi, Sery, Ikemeh, Rachel A, Djagoun, Chabi A M S, Tomsett, Louise & Bearder, Simon K, 2022, A new species of tree hyrax (Procaviidae: Dendrohyrax) from West Africa and the significance of the Niger-Volta interfluvium in mammalian biogeography, pp. 527-552 in Zoological Journal of the Linnean Society 194 (2) on page 551, DOI: 10.1093/zoolinnean/zlab029, http://zenodo.org/record/6352101
Dendrohyrax dorsalis dorsalis (Fraser
Creators: name: Oates, John F
name: Woodman, Neal
name: Gaubert, Philippe
name: Sargis, Eric J
name: Wiafe, Edward D
name: Lecompte, Emilie
name: Dowsett-Lemaire, Françoise
name: Dowsett, Robert J
name: Gonedelé Bi, Sery
name: Ikemeh, Rachel A
name: Djagoun, Chabi A M S
name: Tomsett, Louise
name: Bearder, Simon K
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Mammalia
Hyracoidea
Procaviidae
Dendrohyrax
Dendrohyrax dorsalis

description: Dendrohyrax dorsalis dorsalis (Fraser,
1854) (N = 7)
Equatorial Guinea. Bioko: 3°30′N, 8°41′E (NHMUK 3.1.6.3, 4.7.1.119, 4.7.1.121, 4.7.1.123, 4.7.1.124, 55.12.26.183, 58.5.3.1).
description: Published as part of Oates, John F, Woodman, Neal, Gaubert, Philippe, Sargis, Eric J, Wiafe, Edward D, Lecompte, Emilie, Dowsett-Lemaire, Françoise, Dowsett, Robert J, Gonedelé Bi, Sery, Ikemeh, Rachel A, Djagoun, Chabi A M S, Tomsett, Louise & Bearder, Simon K, 2022, A new species of tree hyrax (Procaviidae: Dendrohyrax) from West Africa and the significance of the Niger-Volta interfluvium in mammalian biogeography, pp. 527-552 in Zoological Journal of the Linnean Society 194 (2) on page 552, DOI: 10.1093/zoolinnean/zlab029, http://zenodo.org/record/6352101
Dendrohyrax dorsalis subsp. emini Thomas 1887
Creators: name: Oates, John F
name: Woodman, Neal
name: Gaubert, Philippe
name: Sargis, Eric J
name: Wiafe, Edward D
name: Lecompte, Emilie
name: Dowsett-Lemaire, Françoise
name: Dowsett, Robert J
name: Gonedelé Bi, Sery
name: Ikemeh, Rachel A
name: Djagoun, Chabi A M S
name: Tomsett, Louise
name: Bearder, Simon K
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Mammalia
Hyracoidea
Procaviidae
Dendrohyrax
Dendrohyrax dorsalis
Dendrohyrax dorsalis emini

description: Dendrohyrax dorsalis emini Thomas (1887)
(N = 24)
Democratic Republic of Congo. Gemena Zone, Tandala: 2°59′N, 19°21′E (USNM 537895). Buta: 2°48′N, 24°45′E (RMCA 9382). Yaenero: 0°12′N, 24°46′E (RMCA 93091 M38). Niapu: 2°25′N, 26°28′E (AMNH M-53823, M-53824, M-53826, M-53828, M-53829, M-53830, M-53831, M-53832, M-53835, M-53836, M-53837, M-53838, M-53840, M-53844). Gamangui: 2°10′N, 27°17′E (AMNH M-53809, M-53810). Medje: 2°23′N, 27°18′E (AMNH M-52120, M-53814). Ngayu: 1°45′N, 27°33′E (AMNH M-53818). Upper Ituri: 1°24′N, 29°45′E (NHMUK 7.4.23.1, 12.7.4.4).
description: Published as part of Oates, John F, Woodman, Neal, Gaubert, Philippe, Sargis, Eric J, Wiafe, Edward D, Lecompte, Emilie, Dowsett-Lemaire, Françoise, Dowsett, Robert J, Gonedelé Bi, Sery, Ikemeh, Rachel A, Djagoun, Chabi A M S, Tomsett, Louise & Bearder, Simon K, 2022, A new species of tree hyrax (Procaviidae: Dendrohyrax) from West Africa and the significance of the Niger-Volta interfluvium in mammalian biogeography, pp. 527-552 in Zoological Journal of the Linnean Society 194 (2) on page 552, DOI: 10.1093/zoolinnean/zlab029, http://zenodo.org/record/6352101
Dendrohyrax dorsalis subsp. latrator
Creators: name: Oates, John F
name: Woodman, Neal
name: Gaubert, Philippe
name: Sargis, Eric J
name: Wiafe, Edward D
name: Lecompte, Emilie
name: Dowsett-Lemaire, Françoise
name: Dowsett, Robert J
name: Gonedelé Bi, Sery
name: Ikemeh, Rachel A
name: Djagoun, Chabi A M S
name: Tomsett, Louise
name: Bearder, Simon K
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Mammalia
Hyracoidea
Procaviidae
Dendrohyrax
Dendrohyrax dorsalis
Dendrohyrax dorsalis latrator (thomas,

description: Dendrohyrax dorsalis latrator (Thomas,
1910) (N = 11)
Democratic Republic of Congo. Bikoro, Lac Tumba: 0°44′S, 18°08′E (RMCA 22114). North of Boende, Tshuapa: 0°10′S, 20°53′E (FMNH 219683). Bumba Zone, Yalosemba: 2°35′N, 21°47′E (USNM 537894). Amadjabe: 0°04′S, 25°17′E (RMCA 89023 M24, 89023M 32, 89023M34, 89023M37, 89023M 38, 89023M47, 89023M48). Bomboma, Bombolo: 2°05′S, 20°55′E (NHMUK 12.7.26.12).
description: Published as part of Oates, John F, Woodman, Neal, Gaubert, Philippe, Sargis, Eric J, Wiafe, Edward D, Lecompte, Emilie, Dowsett-Lemaire, Françoise, Dowsett, Robert J, Gonedelé Bi, Sery, Ikemeh, Rachel A, Djagoun, Chabi A M S, Tomsett, Louise & Bearder, Simon K, 2022, A new species of tree hyrax (Procaviidae: Dendrohyrax) from West Africa and the significance of the Niger-Volta interfluvium in mammalian biogeography, pp. 527-552 in Zoological Journal of the Linnean Society 194 (2) on page 552, DOI: 10.1093/zoolinnean/zlab029, http://zenodo.org/record/6352101
Dendrohyrax interfluvialis Oates & Woodman & Gaubert & Sargis & Wiafe & Lecompte & Dowsett-Lemaire & Dowsett & Gonedelé Bi & Ikemeh & Djagoun & Tomsett & Bearder 2022
Creators: name: Oates, John F
name: Woodman, Neal
name: Gaubert, Philippe
name: Sargis, Eric J
name: Wiafe, Edward D
name: Lecompte, Emilie
name: Dowsett-Lemaire, Françoise
name: Dowsett, Robert J
name: Gonedelé Bi, Sery
name: Ikemeh, Rachel A
name: Djagoun, Chabi A M S
name: Tomsett, Louise
name: Bearder, Simon K
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Mammalia
Hyracoidea
Procaviidae
Dendrohyrax
Dendrohyrax interfluvialis

description: Dendrohyrax interfluvialis (N = 4)
Nigeria. Ogun State: Ilaro Forest Reserve: 6°47 ′ N, 3°04 ′ E (NHMUK 46.360, holotype). Oyo State: Ibadan: 7°30 ′ N, 3°53 ′ E (NHMUK 61.461). Bayelsa State: Uzere: 5°20 ′ N, 6°13 ′ E (NHMUK 1998.300). Edo State: Sapoba Forest Reserve: 6°05 ′ N, 5°49 ′ E (USNM 377548).
description: Published as part of Oates, John F, Woodman, Neal, Gaubert, Philippe, Sargis, Eric J, Wiafe, Edward D, Lecompte, Emilie, Dowsett-Lemaire, Françoise, Dowsett, Robert J, Gonedelé Bi, Sery, Ikemeh, Rachel A, Djagoun, Chabi A M S, Tomsett, Louise & Bearder, Simon K, 2022, A new species of tree hyrax (Procaviidae: Dendrohyrax) from West Africa and the significance of the Niger-Volta interfluvium in mammalian biogeography, pp. 527-552 in Zoological Journal of the Linnean Society 194 (2) on page 552, DOI: 10.1093/zoolinnean/zlab029, http://zenodo.org/record/6352101
Ampittia matudai Murayama 1943
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Ampittia
Ampittia matudai

description: Ampittia matudai Murayama, 1943
Zephyrus 9(3): 170.
Labels: “ Taiwan Musha (=Wushe, in Chinese Character)/ 22-VI-1941 /S MURAYAMA,” “ Ampittia /matudai/ HOLOTYPE.”
A male specimen was designated as the holotype by Murayama (1943), and the specimen was retrieved in LBM (Figs. 1–3; Dried Insect Database Reg. no. 1500022019).
Additional specimens of the species were not reported until Murayama (1955) found a pair in Sugitani’s collection deposited at Kyushu University. HC tried to locate those specimens, but they were removed from the collection and no record was kept as to who loaned or took them. Subsequent records of matudai (Lin 1994, Lin & Su 2013) turned out to be either A. inachus (Ménétriés, 1858) (Hsu et al. 2019) or A. bandaishanus Murayama & Shimonoya, 1968 (Hsu 2013, Sugisaka 2018). YFH undertook investigations at the type locality of the taxon every year since 1982 searching for the species but in vain. The vicinity of the area was covered with decent vegetation before the end of World War II but was later severely deforested and disturbed due to horticultural practice.
We regard matudai as a valid specific name in the genus Aeromachus de Nicéville, 1890 following the treatment in Chiba et al. (1992) and Hsu et al. (2019). Huang (2009) considered that the taxon should be treated as a subspecies of A. jesta (Evans, 1939). This point of view awaits further evaluation.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on page 281, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Erynnis montanus subsp. neomontanus Murayama & Yoshisaka 1959
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Erynnis
Erynnis montanus
Erynnis montanus neomontanus murayama & yoshisaka, 1959

description: Erynnis montanus neomontanus Murayama & Yoshisaka, 1959
Tyô to Ga 10(1): 14.
Labels: “ Taiwan Hori (=Puli, in Chinese character)/1955/S. MURAYAMA,” “ E. montanus /neomontanus/ Mur&Yoshisa/ka HOLOTYPE.”
A male specimen was designated as the holotype by Murayama & Yoshisaka (1959), and the specimen was retrieved in LBM (Figs. 4–6; Dried Insect Database Reg. no. 1500022024).
This taxon was described based on 2 males sent from Mr. Masato Watanabe to Mr. Michio Yoshisaka. They were among worn out butterfly specimens from Taiwan, and the authors claimed that it was improbable that specimens from Honshu, Japan were mixed in. The collecting locality was not certain and Puli, central Taiwan, was assigned with a question mark. In those days, Puli, a small town located in central Taiwan, was the place to accumulate specimens, and butterflies and some moths came from all over Taiwan. Butterflies also arrived from foreign countries in order to utilize their wings for art display. The authors of the taxon claimed that the type specimens were extremely similar but different from the Japanese and Korean samples in having the yellow dots on hindwings unclearly edged. Such variation, however, is also found in Japanese specimens, and there are a few examples in which Taiwanese and Japanese (except for the southern islands of Okinawa) specimens are so similar as in this case except for some wide-spread species like Eurema hecabe (Linnaeus, 1758) or wanderers such as Lampides boeticus (Linnaeus, 1767). Erynnis montanus (Bremer, 1861) obviously does not belong to such cases for this species is known as a montane dweller.As we have already regarded this taxon as a junior subjective synonym of E. montanus in previous publications (Tsukiyama et al. 1997, Hsu et al. 2019), we concluded that both specimens in the type series were mislabeled.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on page 281, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Ochlodes siva subsp. yuchingkina Murayama & Shimonoya 1963
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Ochlodes
Ochlodes siva
Ochlodes siva yuchingkina murayama & shimonoya, 1963

description: Ochlodes siva yuchingkina Murayama & Shimonoya, 1963
Tyô to G a 13(3): 53.
Labels: “ Taiwan, Tsugitakayama (= Xueshan, in Chinese character)/ June 1961,” “ Ochlodes /siva/yuchingkina/ HOLOTYPE.”
A male specimen was designated as the holotype by Murayama & Shimonoya (1962), and the specimen was retrieved in the Shimonoya collection (Figs. 7–9).
Having examined the male genitalia, Chiba & Hsu (1989) indicated that yuchingkina is not a subspecies of siva Moore, 1878, but of bouddha Mabille, 1876. Subsequent publications follow this view (Chiba et al. 1992, Chiba & Tsukiyama 1996, Hsu 2013, Hsu et al. 2019). We regard yuchingkina as a subspecies of O. bouddha.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on pages 281-282, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Aeromachus bandaishanus Murayama & Shimonoya 1968
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Aeromachus
Aeromachus bandaishanus

description: Aeromachus bandaishanus Murayama & Shimonoya, 1968
New Entomologist 17(1): 10.
Labels: “Bandaisha/ Formosa /1, VII, 1965,” “Gen. No./ 66079,” “ Aeromachus / bandaishanus/ Mur et Shim./ HOLOTYPE.”
A male specimen was designated as the holotype by Murayama & Shimonoya (1968), and the specimen was retrieved in the Shimonoya collection (Figs. 10–12).
Though this species might be related to a continental species such as A. nanus (Leech, 1890), we regard bandaishanus as a valid species name.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on page 282, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Pseudocoladenia pinsbukana Shimonoya & Murayama 1976
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Pseudocoladenia
Pseudocoladenia pinsbukana

description: Pseudocoladenia pinsbukana Shimonoya & Murayama, 1976
Tyô to Ga 27(2): 44.
Labels: “Shinseimura Kyusenkei (= Xinshengcun Jiuxianxi, in Chinese character)/ Formosa/APR.15.1970,” “ Pseudocoladenia pinsbukana,/SHIMONOYA et/ MURAYAMA 1976,” “ Holotype.” However, the type locality given in the original description was different, reading “Amuh-chi”, a locality not recognized on any map of Taiwan, presumably a local name of Jiuxianxi.
A male specimen was designated as the holotype by Shimonoya & Murayama (1976), and the specimen was retrieved in the Shimonoya collection (Figs. 13–15).
Shimonoya & Murayama (1976) considered that pinsbukana was close to Coladenia sheila Evans, 1939 from China. In describing the new species, however, they somehow compared it with Pseudocoladenia dan (Fabricius, 1787) from India. The wing markings, especially the 2 costal hyaline spots over the cell on the forewing recognized in the holotype, are a diagnostic character of Coladenia hoenei Evans, 1939. The discal dots on the hindwing are reduced in pinsbukana, whereas those of hoenei are more prominent. A slight difference is also seen in the male genitalia, if their drawing is accurate. The dorsal process of valva is narrower than the ventral process in hoenei, whereas those of pinsbukana are nearly of the same width (Hsu et al. 2019).
We regard pinsbukana as a valid species name in the genus Coladeni a Moore, 1881 for the present.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on page 282, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Apostictopterus fulginosus subsp. jiangxiensis Murayama 1983
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Apostictopterus
Apostictopterus fulginosus
Apostictopterus fulginosus jiangxiensis murayama, 1983

description: Apostictopterus fulginosus jiangxiensis Murayama, 1983
Entomotaxonomia 5(4): 284.
Labels: “Shusui (= Xiushui, in Chinese character)/ Jiangxi Province (in Chinese character)/ VI-1982 /MURAYA- MA,” “ A. fuliginosus / jiangxiensis/ HOLOTYPE.”
A male specimen was designated as the holotype by Murayama (1983), and the specimen was retrieved in LBM (Figs. 19–21; Dried Insect Database Reg. no. 1500022015).
Murayama (1984) corrected his identification of the type and considered the name a synonym of Astictopterus jama chinensis (Leech, 1890). We follow his treatment.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on page 283, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Aeromachus inachus subsp. jiujiangana Murayama 1983
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Aeromachus
Aeromachus inachus
Aeromachus inachus jiujiangana murayama, 1983

description: Aeromachus inachus jiujiangana Murayama, 1983
Entomotaxonomia 5(4): 284.
Labels: “Kyukou (= Jiujiang, in Chinese character)/ Jiangxi Province (in Chinese character)/ IX-1982 /MU- RAYAMA,” “ A. inachus / jiujiängana/ HOLOTYPE.”
A male specimen was designated as the holotype by Murayama (1983), and the specimen was retrieved in LBM (Figs. 22–24; Dried Insect Database Reg. no. 1500022013).
As Bridges (1994) noted, Evans (1949) overlooked the Chinese taxon, nanka Evans, 1943 that he described, and treated inachus Ménétriés, 1858 from China as formosanus Matsumura, 1931. As a consequence, Murayama (1983) was not aware of nanka and described jiujiangana. Tsukiyama et al. (1997) overlooked jiujiangana, and the name was not listed. We find no difference among south eastern Chinese specimens of inachus (e. g. specimens shown in Tsukiyama et al. 1997). Therefore, we consider jiujiangana as a junior subjective synonym of nanka.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on page 283, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Pedesta masuriensis subsp. cuneomaculata Murayama 1995
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Pedesta
Pedesta masuriensis
Pedesta masuriensis cuneomaculata murayama, 1995

description: Pedesta masuriensis cuneomaculata Murayama, 1995
Nature & Insects 30(14): 33.
Labels: “ Yunnan Province (in Chinese character)/Dokanmura (= Tuguancun, in Chinese character)/ 24-V-1995 / S MURAYAMA,” “ Pedesta masuri-/ensis/cuneoma/culata/ HOLOTYPE.”
A male specimen was designated as the holotype by Murayama (1995), and the specimen was retrieved in LBM (Figs. 25–27; Dried Insect Database Reg. no. 1500022032).
Huang (2011) treated cuneomaculata as a junior subjective synonym of Thoressa masuriensis tali (Swinhoe, 1912). We agree that cuneomaculata is a synonym of tali. The nominate subspecies masuriensis Moore, 1878 and subspecies tali are significantly different in appearance (Figs. 46–49), but Evans (1949) considered them as conspecific. However, masuriensis (Figs. 46–47, 50) and tali (Figs. 48–49, 51) are two different species, supported by distinctions in genitalia (Figs. 50–51) and the genetic divergence of COI barcode between them (12–13%, Table 1). They may be separated by the following features: 1) uncus bent with blunt tips in masuriensis (Fig. 50) versus straight with narrowed tips in tali (Fig. 51); 2) left harpe with inner one of terminal processes as small lobe in masuriensis (Fig. 50) versus produced serrate teeth in tali (Fig. 51); 3) phallus short and stout in masuriensis (Fig. 50) versus elongate and slender in tali (Fig. 51). Those masuriensis molecular samples in China (Li et al. 2019) all belong to tali.
Huang (2011) was correct in merging the Chinese taxa of Thoressa Swinhoe, 1913 and Pedesta Hemming, 1934. His taxonomic treatment, however, was not complete because he did not include the type species of Thoressa. The genus Thoressa sensu Evans (1949) was polyphyletic and is now divided into three genera based on molecular evidences (Huang et al. 2019). Those species assigned to Thoressa by Huang (2011) indeed should be included in Pedesta. Therefore, Pedesta tali comb. nov. is the valid name for this taxon.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on page 283, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Thoressa dianchiana Murayama 1995
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Thoressa
Thoressa dianchiana

description: Thoressa dianchiana Murayama, 1995
Nature & Insects 30(14): 33.
Labels: “ Yunnan Province /Tuguan Village (in Chinese)/ 12-V-1995 /S MURAYAMA,” “ T. dian/chiana/HOLO- TYPE.”
A male specimen was designated as the holotype by Murayama (1995), and the specimen was retrieved in LBM (Figs. 28–30; Dried Insect Database Reg. no. 1500022025).
Huang & Zhan (2004) treated dianchiana as a synonym of Thoressa baileyi (South, 1913), and we follow their treatment. As mentioned above, we consider baileyi a species of Pedesta.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on page 284, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Thoressa nanshaona Murayama 1995
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Thoressa
Thoressa nanshaona

description: Thoressa nanshaona Murayama, 1995
Nature & Insects 30(14): 33.
Labels: Yunnan Province (in Chinese character)/ Dokanmura (= Tengchongcun, in Chinese character)/ 24-V- 1995 / S MURAYAMA,” “ T. nan-/ choana [sic!]/ HOLOTYPE.”
A male specimen was designated as the holotype by Murayama (1995), and the specimen was retrieved in LBM (Figs. 31–33; Dried Insect Database Reg. no. 1500022026).
Huang & Zhan (2004) examined the type and Huang (2011) treated nanshaona as a junior synonym of Thoressa pandita (de Nicéville, 1885). We follow his treatment. Again, we retain the conventional view and consider panditia a species of Pedesta.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on page 284, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Lobocla bifasciata subsp. disparalis Murayama 1995
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Lobocla
Lobocla bifasciata
Lobocla bifasciata disparalis murayama, 1995

description: Lobocla bifasciata disparalis Murayama, 1995
Nature & Insects 30(14): 34.
Labels: “ Yunnan Province Dairi (= Tali, in Chinese character)/ 28-V-1988 / SMURAYAMA,” “Labocla [sic!]/ bifasciata/ disparalis/ HOLOTYPE.”
A male specimen was designated as the holotype by Murayama (1995), and the specimen was retrieved in LBM (Figs. 34–36; Dried Insect Database Reg. no. 1500022020).
Huang (2003) synoymized disparalis Murayama, 1995 with bifasciatus Bremer & Grey, 1853, and we follow his treatment.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on page 284, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Carterocephalus dieckmanni subsp. soyosanus Murayama 1983
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Carterocephalus
Carterocephalus dieckmanni
Carterocephalus dieckmanni soyosanus murayama, 1983

description: Carterocephalus dieckmanni soyosanus Murayama, 1983
Entomotaxonomia 5(4): 284.
Labels: “Soyosan (in Chinese character)/ S. Korea (in Chinese character)/ 24-4-1944 / MURAYAMA,” “dieckmanni/ soyosanus/ HOLOTYPE (in red).”
A male specimen was designated as the holotype by Murayama (1983), and the specimen was retrieved in LBM (Figs. 37–39; Dried Insect Database Reg. no. 1500022017).
The subspecific name soyosanus has not been popularized in Korea. No subspecific name is assigned to dieckmanni Graeser, 1888 in two recently published butterfly books in Korea (Joo et al. 1997, Kim 2002).
Evans (1949) recognized 4 subspecies of dieckmanni based on the difference in the central band on the ventral side of the hindwing. Even a brief examination of specimens tells us that some of the subspecific names should be synonymized due to clinal variation.After examining 57 specimens (1 from Russia, 3 from Korea, 2 from Northeast China, 13 from North China, 37 from West China, and 1 from Myanmar) in the Tsukiyama Collection, we found that the markings used by Evans (1949) as diagnostic characters to distinguish subspecies actually represent a clinal variation. The difference between ssp. dieckmanni and ssp. gemmatus Leech, 1891 is not clearly defined, and so is the difference between ssp. minor Evans, 1932 and ssp. watsoni Evans, 1932. Consequently, no subspecific name is necessary for Korean populations of dieckmanni.
We consider soyosanus a junior subjective synonym of nominotypical dieckmanni Graeser, 1888, syn. nov.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on pages 284-287, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Sebastonyma suthepiana Murayama & Kimura. Valid 1990
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Sebastonyma
Sebastonyma suthepiana

description: Sebastonyma suthepiana Murayama & Kimura, 1990
Nature & Insects 25(2): 24.
Labels: “Doi Suthep near/ Chieng Mai, Thail./ 12 th April 1976 / Y. Kimura leg.,” “ Sebastonyma suthepiana / Murayama & Kimura, 1990,” “ Holotype / specimen.,” “9600050[with genitalia vial].” (Figs. 40–42)
Additional specimens were collected in Laos (Onodera, personal communication). We regard suthepiana as a valid species name. The specific name of this taxon was spelled on p. 23 of the original description as “ sutlepiana ”, an incorrect original spelling.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on page 287, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Halpe nuydai Murayama & Okamura 1973
Creators: name: Chiba, Hideyuki
name: Tsukiyama, Hiroshi
name: Liang, Jia-Yuan
name: Wang, Shou-Ming
name: Shen, Zong-Yu
name: Hsu, Yu-Feng
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Halpe
Halpe nuydai

description: Halpe nuydai Murayama & Okamura, 1973
Tyô to Ga 24(1): 24.
Labels: “ St Thomas /Luzon/ 2-IV-1972 /S. MURAYAMA,” “ Halpe /nuydai/ HOLOTYPE.”
A male specimen was designated as the holotype by Murayama & Okamura (1973), and the specimen was retrieved in LBM (Figs. 43–45; Dried Insect Database Reg. no. 1500022031).
Though Murayama & Okamura (1973) claimed that the genitalia of nuydai resembled that of Thoressa horishana (Matsumura, 1910) from Taiwan, they somehow classified nuydai in the genus Halpe Moore, 1878. The taxon was synonymized by Kawazoé (1973), and we follow his treatment. Halpe nuydai Murayama & Okamura, 1973 is a junior subjective synonym of Thoressa justini Inoué & Kawazoé, 1969.
description: Published as part of Chiba, Hideyuki, Tsukiyama, Hiroshi, Liang, Jia-Yuan, Wang, Shou-Ming, Shen, Zong-Yu & Hsu, Yu-Feng, 2020, The types of skippers described by Shu-Iti Murayama (Lepidoptera: Hesperiidae), pp. 280-290 in Zootaxa 4801 (2) on pages 287-288, DOI: 10.11646/zootaxa.4801.2.4, http://zenodo.org/record/3990957
Haderonia sukharevae Varga 1974
Creators: name: Varga, Zoltán
name: Gyulai, Péter
name: Ronkay, Gábor
name: Ronkay, László
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Noctuidae
Haderonia
Haderonia sukharevae

description: The sukharevae -group
Diagnosis – Relatively large moths with robust body, very long, in male strongly (sukharevae, oxyptera, juliannae) or shortly (longicornis, lukhtanovi, stenocera) bipectinate antennae and elongate triangular forewings with colourful appearance. The noctuid forewing pattern is complete and regular, in some species sharply expressed (sukharevae, oxyptera, juliannae), in others rather faint (lukhtanovi, stenocera). Male brush organ reduced.
Male genitalia – uncus mostly short and acute triangular (exceptionally obtuse in lukhtanovi); valvae with normally developed rounded cucullus and corona without incision, short and broad ampulla (exception: oxyptera). Saccular processes symmetrical, short and obtuse. Species of this group can be mostly differentiated by the length of pectination of male antennae, by the shape and length of uncus and ampulla but also by the length of the field of fasciculate cornuti on the distal part of the tubular vesica.
Checklist
Ctenoceratoda longicornis (Graeser, 1892, Mamestra), Berliner Entomologische Zeitschrift 37: 306.Type-locality: [Kirghisia] Kysyl-Yart; [China, Xinjiang] Kashgar;
Ctenoceratoda sukharevae (Varga, 1974, Haderonia), Annales historico-naturales Musei nationalis hungarici 66: 301, pl. 7, fig. 7. Type-locality: Mongolia, Bayankhongor aimak, Zhinst Mts;
Haderonia sukharevae excellens (Varga, 1974, Haderonia), Annales historico-naturales Musei nationalis hungarici 66: 302, pl. 1, figs 7–8.Type-locality: Mongolia, Khövsgöl aimak, Delger Mörön River;
Ctenoceratoda oxyptera Varga, 1992, Acta Zoologica Academiae Scientiarum Hungaricae 38 (1–2): 99, pl. 3, figs 17–18, gen. figs 37–43.Type-locality: Mongolia, Govi Altai aimak, Govi Altai Mts;
Ctenoceratoda juliannae Varga, 1992, Acta Zoologica Academiae Scientiarum Hungaricae 38 (1– 2): 99, pl. 3, fig. 19, gen. figs 33–36. Type-locality: Mongolia, Khovd aimak, Dzhungar Govi, Bulgan sum;
Ctenoceratoda lukhtanovi Varga et Gyulai, 1999, Acta Zoologica Academiae Scientiarum Hungaricae 45 (2): 172, figs 1–2, 23, 41, 50–52, 82. Type-locality: Tadjikistan, Pamir Mts, Muzkol Mts, Ak-Baital Pass;
Ctenoceratoda stenocera Varga et Gyulai, 2002, Esperiana 9: 230, pl. 20, fig. 16. Type-locality: China, Kunlun Mts, 60 km NW of Xaidullah village.
Bionomics and distribution – A mostly Inner Asiatic species group, typical for extreme cold-continental semi-desert or rupicolous habitats in medium or high altitudes.
description: Published as part of Varga, Zoltán, Gyulai, Péter, Ronkay, Gábor & Ronkay, László, 2018, Review Of The Species Groups Of The Genus Ctenoceratoda Varga, 1992 With Description Of Four New Species And A New Subspecies (Lepidoptera, Noctuidae), pp. 51-74 in Acta Zoologica Academiae Scientiarum Hungaricae 64 (1) on page 53, DOI: 10.17109/AZH.64.1.51.2018, http://zenodo.org/record/5734636
Ctenoceratoda khorgossi
Creators: name: Varga, Zoltán
name: Gyulai, Péter
name: Ronkay, Gábor
name: Ronkay, László
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Noctuidae
Ctenoceratoda
Ctenoceratoda khorgossi

description: The khorgossi -group
Diagnosis – This species-group is most similar in its external and genital characters to the sukharevae -group. They are medium-sized moths with robust body, very long and strongly bipectinated male antennae and relatively narrow triangular, apically acute forewings with brownish-bluish-ochreous or silvery grey variegated colouration. Orbicular and reniform stigmata conspicuous, reniform stigma often with acute “pipe-shaped” extension in basal direction, orbicular stigma narrow and oblique. Male brush organ reduced.
In the male genitalia, the uncus is relatively short, triangular, pointed apically, the cucullus is rounded triangular or slightly inflated with narrow neck and regular corona; the ampulla is mostly elongate, acute or obtuse. In male vesica the stripe of fasciculate cornuti and in females the sausage-shaped appendix bursae are relatively short.
Checklist
Ctenoceratoda khorgossi (Alphéraky, 1882, Mamestra), Horae Societatis Entomologicae Rossicae 17: 65, pl. 2, fig. 49. Type-locality: [China, Xinjiang] Kuldja district; Khorgoss;
Ctenoceratoda argyrea Varga, 1992, Acta Zoologica Academiae Scientiarum Hungaricae 38 (1-2): 98, pl. 2, figs 9–10, gen. figs 5–10, 16–17.Type-locality: Mongolia, Govi Altai aimak, Govi Altai Mts;
Ctenoceratoda persephone sp. n.
Ctenoceratoda scotosparsa sp. n.
Bionomics and distribution – One species of the group is widely distributed in Central Asia, the range of three others seems to be confined to western Mongolia with some phenological and vertical differentiation.
description: Published as part of Varga, Zoltán, Gyulai, Péter, Ronkay, Gábor & Ronkay, László, 2018, Review Of The Species Groups Of The Genus Ctenoceratoda Varga, 1992 With Description Of Four New Species And A New Subspecies (Lepidoptera, Noctuidae), pp. 51-74 in Acta Zoologica Academiae Scientiarum Hungaricae 64 (1) on page 54, DOI: 10.17109/AZH.64.1.51.2018, http://zenodo.org/record/5734636
Haderonia tancrei
Creators: name: Varga, Zoltán
name: Gyulai, Péter
name: Ronkay, Gábor
name: Ronkay, László
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Noctuidae
Haderonia
Haderonia tancrei

description: The tancrei -group
Diagnosis – Medium-sized or large moths with robust body, very long and strongly bipectinated male antennae and elongated-triangular forewings. Colouration of forewings mostly greyish-ochreous-brownish, rarely rufous with regular but often faint maculation. Reniform stigma often shows an acute whitish stripe basally. Male brush organ regularly present.
In the male genitalia, the uncus is relatively long, the cucullus is rounded triangular, often slightly inflated or with small incision; the ampulla is elongate, more or less curved. Vesica tubular, extremely long with long stripe of fasciculate cornuti.
The species of this group can be recognised externally by the greyish or rufous-brownish colouration, the presence or absence of whitish marking at the basal part of the reniform stigma. The group features of the male genitalia are the characteristic shape of the “head” of cucullus, and configuration of the ampulla and the length of the fascia of cornuti in the vesica.
Checklist
Ctenoceratoda tancrei (Graeser, 1892, Mamestra), Berliner Entomologische Zeitschrift 37: 305. Type-locality: [Kirghisia] Alexander Mts;
Ctenoceratoda graeseri (Püngeler, 1898, Haderonia, tancrei var.), Societas Entomologica 13: 58. Type-locality: [Kazakhstan or China] Ili region;
Ctenoceratoda thermolimna (Boursin, 1964, Haderonia), Zeitschrift der Wiener Entomologischen Gesellschaft 49: 174, pl. 22, figs 1, 4. Type-locality: Kirghisia, Issyk-Kul;
Ctenoceratoda anthracina Varga et Gyulai, 1999, Acta Zoologica Academiae Scientiarum Hungaricae 45 (2): 180, figs 12–13. Type-locality: Kirghisia, Naryn region, Maly Naryn;
Ctenoceratoda naryna Varga et Gyulai, 1999, Acta Zoologica Academiae Scientiarum Hungaricae 45 (2): 189, figs 15–16, 33, 47, 66–67, 87. Type-locality: Kirghisia, Naryn region, Maly Naryn;
Ctenoceratoda transalaica Varga et Gyulai, 1999, Acta Zoologica Academiae Scientiarum Hungaricae 45 (2): 189, figs 17, 34, 48, 68–69, 88. Type-locality: Kirghisia, Transalai, Aram-Kungei.
Ctenoceratoda cyanochrea sp. n.
Bionomics and distribution – Almost all species are confined to the Tien-Shan mountain system where they are mostly typical for the semi-desert like habitats of low or moderate altitudes (contrasting to the next group!). Two species are relatively widely distributed in Central Asia (tancrei, graeseri), the other species seem to be rather localised and infrequent.
description: Published as part of Varga, Zoltán, Gyulai, Péter, Ronkay, Gábor & Ronkay, László, 2018, Review Of The Species Groups Of The Genus Ctenoceratoda Varga, 1992 With Description Of Four New Species And A New Subspecies (Lepidoptera, Noctuidae), pp. 51-74 in Acta Zoologica Academiae Scientiarum Hungaricae 64 (1) on pages 54-55, DOI: 10.17109/AZH.64.1.51.2018, http://zenodo.org/record/5734636
Ctenoceratoda turpis Staudinger 1999
Creators: name: Varga, Zoltán
name: Gyulai, Péter
name: Ronkay, Gábor
name: Ronkay, László
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Noctuidae
Ctenoceratoda
Ctenoceratoda turpis

description: The turpis -group
Diagnosis – Large or medium-sized moths with not extremely long and moderately or strongly bipectinated male antennae, moderately elongate, apically acute forewings. Colouration simple, nearly concolorous grey or brownish, markings regular, often faint. Male brush organ present.
In the male genitalia, uncus long and narrow; cucullus rounded triangular with moderately elongate “neck”; ampulla elongate, often slightly falcate. Since the genital capsule of the different members of the group is very similar, the pectination of antennae appears as important specific character.
Checklist
Ctenoceratoda turpis (Staudinger, 1900, Phoebophilus), Deutsche Entomologische Zeitschrift. Gesellschaft Iris zu Dresden 12: 341, pl. 7, fig. 8. Type-locality: [China, Xinjiang] Korla;
Ctenoceratoda brassicina (Draudt, 1934, Scotogramma), in A. Seitz, Die Gross-Schmetterlinge der Erde 3: 98, pl. 14, row c. Type-locality: [Russia or Kazakhstan] “Altai occ.”;
Ctenoceratoda gandhara (Hacker et Varga, 1990, Haderonia), Esperiana 1: 340, pl. E, fig. 16. Type-locality: Pakistan, Karakoram Mts, Kunjerab pass;
Ctenoceratoda weigerti (Hacker et Varga, 1990, Haderonia), Esperiana 1: 340, pl. E, fig. 17. Type-locality: Pakistan, Karakoram Mts, Kunjerab pass;
Ctenoceratoda peregovitsi Varga et Gyulai, 1999, Acta Zoologica Academiae Scientiarum Hungaricae 45 (2): 174, figs 4–5, 25–26. Type-locality: Mongolia, Ömnögovi aimak, Tost Mts;
Ctenoceratoda psychrogena Varga et Gyulai, 1999, Acta Zoologica Academiae Scientiarum Hungaricae 45 (2): 175, figs 6–8, 27–29, 42–44, 58–62, 84. Type-locality: Tadjikistan, Pamir Mts, Vakhanskiy Mts;
Ctenoceratoda aksakal Varga et Gyulai, 1999, Acta Zoologica Academiae Scientiarum Hungaricae 45 (2): 179, figs 11, 41, 57, 85. Type-locality: Tadjikistan, E Pamir Mts, Sarykolskiy Mts.
Bionomics and distribution – All species are confined to the high mountains of Central and Inner Asia, they regularly occur in high altitudes.
description: Published as part of Varga, Zoltán, Gyulai, Péter, Ronkay, Gábor & Ronkay, László, 2018, Review Of The Species Groups Of The Genus Ctenoceratoda Varga, 1992 With Description Of Four New Species And A New Subspecies (Lepidoptera, Noctuidae), pp. 51-74 in Acta Zoologica Academiae Scientiarum Hungaricae 64 (1) on pages 55-56, DOI: 10.17109/AZH.64.1.51.2018, http://zenodo.org/record/5734636