Pseudodryinus parisiensis Peinado & Nel & Waller 2006, sp. n.
Creators: name: Peinado, Julie
name: Nel, Andre
name: Waller, Alain
Publisher: name: Zenodo
Year: 2006
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Dryinidae
Pseudodryinus
Pseudodryinus parisiensis

description: Pseudodryinus parisiensis sp. n.
(figs 1–7)
Diagnosis: Mandible with four teeth; palpal formula 5/2; maxillary palpomeres long; rudimentary tarsal claw of front leg present; enlarged tarsal claw of front leg with a subapical tooth, as long as fifth tarsal segment, both bearing lamellae; front tarsal segments 2 and 3 with a strong hook; tibial spur formula (1/1/2); ocelli present; occipital carina complete; mesosomal segments not fused; notauli clearly visible on mesoscutum; epicnemium visible; fore wing cells C, R, 1Cu and 2R1 closed, cells 1R1, 1M and 2Cu distally closed by nebulous veins; cells 2Rs, 2M, and 3Cu present but distally opened. The combination of tibial spur (1/1/2) and palpal (5/2) formulae is diagnostic for the new species.
Description. Body and legs brown yellow; body 4 mm long; head hypognathous, longer than wide; eyes oval, large, occupying nearly the whole length of the head; three ocelli between eyes, making an obtuse equilateral triangle, the median one distinctly larger; occiput short, vertical, and dorsally bordered by a carina; frons occupied by a depression that ends in clypeus; OL = 12; POL = 9; OOL = 30; OPL = 10; TL = 15; antenna with 10 bare segments; scape robust; pedicel and flagellum cylindrical, progressively shortening, with segments in the following proportions 28:10:40:30:22:19:15:14:14:11; antenna much shorter than body, approximately 3.3 times as long as head (length of head measured dorsally from occipital carina behind the ocelli to distal apex of the mandibles): 203:61; maxillary palp with four visible segments but with a distinct angle between second and third visible palpomeres, implying the presence of a not visible basal palpomere, visible palpomeres long and thin, in the following proportions 15:20:20:15; labial palp two–segmented; head broader than pronotum; neck robust; mandibles with three strong teeth and a smaller basal one, third tooth smaller than second and fourth.
Mesosomal segments delimited by sutures, not fused; mesosoma partly hidden by legs; pronotum apparently bare; notauli complete; propodeum strongly depressed near petiole; epicnemium visible (Olmi 1992, 1993).
Fore wing 1.8 mm long; pterostigma narrow; only three cells are closed by tubular veins, i.e. cells C, R, 1Cu and 2R1; cell 1R1 closed but bordered anteriorly and basally by tubular veins, posteriorly and distally by nebulous veins; cells 1M and 2Cu both bordered basally by a tubular vein and by nebulous veins on their three other sides; cells 2Rs, 2M, and 3Cu open with their anterior and posterior sides bordered by nebulous veins; anal margin of wing not visible; wing covered with short and dense bristles on whole surface.
Hind wing narrow not fully visible; four hamuli on anterior edge at ¾ of distance from base to wing apex, with only vein Sc+R present; wing covered with short and dense bristles on whole surface.
Legs. Front leg very long, with a terminal chela; the two arms of chela of equal length; first arm (enlarged tarsal claw) with a small subapical tooth and at least one row of lamellae, and distinctly shorter than front tibia; apex of enlarged tarsal claw pointed and curved, sheltered in the terminal “cup” of the other branch of chela (fifth tarsomere); this “cup” is nearly circular with a rather large diameter, it is covered with lamellae as is the inner face of the fifth tarsomere; fifth tarsomere with a complete series of long spines on its outer face; arolium present between the two arms of the chela, rather long; second tarsal claw atrophied, very small, half length of arolium; front tarsal segments 2 and 3 both with a strong hook turned upside–down, both hooks bearing five long bristles at their extremity; fore leg segments in the following proportions: 5:3:12:12:5:1:1.5:6:9 for coxa:trochanter:femur:tibia:tarsomeres 1–5 respectively; mid leg segments in the following proportions: 12:13:56:60:46:13:13:5:16; hind leg segments in the following proportions: 40:15:75:97:40:20:15:5:12; tibial spur formula 1/1/2; median and posterior legs with an apical pair of claws separated by an arolium.
Abdomen with a clearly visible ovipositor; two antepenultimate segments ventrally with four very long bristles (sensillae?).
Material. Female holotype specimen PA 2378 1/2, in the same piece of amber with a specimen belonging to the Mantodea, mounted in Canada balsam, in collection De Ploëg and Indivision Langlois –Meurine, deposited in Muséum National d'Histoire Naturelle, Paris. Specimens collected in Le Quesnoy all bear the letter PA for Paris (meaning Paris Basin).
Locality deposit. Le Quesnoy, Chevrière, region of Creil, Oise department, France.
Geological age. Earliest Eocene, Sparnacian, level MP7 of the mammal fauna of Dormaal. We have demonstrated that the amber is autochthonous and very different from the Baltic amber in age, chemical composition and origin (Feugueur 1963; De Ploëg et al. 1998; Nel et al. 1999).
Remarks. The recent Dryinidae are parasitoids of Hemiptera: Auchenorrhyncha. Fossil Cicadellidae and Fulgoroidea are rather diverse and frequent in the ëOiseí amber.
Etymology. The species is named after the city of Paris.
description: Published as part of Peinado, Julie, Nel, Andre & Waller, Alain, 2006, A dryinid wasp in Early Eocene amber from the Paris basin (Hymenoptera: Dryinidae), pp. 31-41 in Zootaxa 1168 (1) on pages 32-36, DOI: 10.11646/zootaxa.1168.1.3, http://zenodo.org/record/5058582