Fimbriatella fimbriata
Creators: name: Harzhauser, Mathias
name: Landau, Bernard
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Heterostropha
Mathildidae
Fimbriatella
Fimbriatella fimbriata

description: Fimbriatella fimbriata (Michelotti, 1847)
Figs 27A–F
* Cerithium fimbriatum mihi — Michelotti 1847: 193, pl. 16, figs 23– 231.
Mathilda fimbriata Michelotti —Semper 1865: 337.
Mathilda fimbriata Michelotti —de Boury 1883: 116.
[Mathilda] Fimbriatella fimbriata (Micht.) —Sacco 1895: 37, pl. 3, figs 35, 36.
[Mathilda] Fimbriatella fimbriata (Micht.) —Sacco 1904: 126, pl. 25, figs 43, 44.
Fimbriatella fimbriata (Michelotti) —Montanaro Gallitelli & Tacoli 1951: 175, pl. 2, fig. 1.
Fimbriatella fimbriata (Michtti) var. elegans n. var. – Montanaro Gallitelli & Tacoli 1951: 175, pl. 2, fig. 2. Mathilda (F [imbriatella].) fimbriata (Micht.) —Sieber 1958: 139.
Mathilda (Fimbriatella) fimbriata (Micht.) —Sieber 1960: 266, pl. 1, figs 20, 21.
Mathilda (Fimbriatella) fimbriata Micht. — Csepreghy-Meznerics 1969a: 20, pl. 2, figs 14–15.
Fimbriatella fimbriata Micht. var. lapusiyensis var. sp.— Moisescu 1955: 277. pl. 2, fig. 3.
Mathilda (Fimbriatella) fimbriata Micht. — Csepreghy-Meznerics 1969b: 69, pl. 1, fig. 7.
Mathilda fimbriata (Michelotti, 1847) —Tabanelli et al. 2021: 7, figs 7a–f, 8a–d.
Type material. Holotype: SL: 7 mm, illustrated in Michelotti (1847: pl. 16, fig.23), Sacco (1895: pl. 3. Fig. 35) and Sacco (1904: pl. 25, fig. 43), Stazzano (Italy), Tortonian (Late Miocene). The specimen was probably stored in the Museo Regionale di Scienze Naturali Torino (Italy), because Sacco (1895, 1904) had the holotype at hand. Ferrero Mortara et al. (1984: 239), however, did not list this specimen in their catalogue. The type specimen of Fimbriatella fimbriata elegans Montanaro Gallitelli & Tacoli, 1951 might have been stored at the University of Modena (Italy) but cannot be traced.
Illustrated material. NHMW 2013 /0078/0125a, SL: 8.8 mm, MD: 4.1 mm, Baden (Austria), Figs 27A 1 –A 2. NHMW 2013 /0078/0125b, SL: 6.6 mm, MD: 3.8 mm, Baden (Austria), Figs 27B 1 –B 2. NHMW 1854 /0035/0307a, SL: 14.1 mm, MD: 6.8 mm, Lăpugiu de Sus (Romania), Fig. 27C. NHMW 1864 /0001/0376b, SL: 10.5 mm, MD: 5.0 mm, Jaroměřice (Czech Republic), Figs 27D 1 –D 2. NHMW 1864 /0001/0376a, SL: 11.0 mm, MD: 5.9 mm, Jaroměřice (Czech Republic), Figs 27E 1 –E 2. NHMW 1854 /0035/0307b, SL: 16.0 mm, MD: 6.9 mm, Lăpugiu de Sus (Romania), Fig. 27F.
Revised description. Large, solid, broad turritelliform; apical angle about 28–36°. Protoconch multispiral, of about 2.5 smooth whorls, heterostrophic, nucleus exposed, tilted at angle of about 110°. Teleoconch of ten angled whorls separated by impressed suture. Whorl profile slightly concave, with broad sutural ramp. Sculpture of first teleoconch whorl of three primary spiral cords: S1 moderately developed, placed abutting upper suture; S2 placed mid-whorl weakest; S3 strongest, placed close above abapical suture. Later whorls with carinate S3, coinciding with periphery. Secondary spiral cord appears between S2 and S3 on third teleoconch whorl, forming a pair of same strength with S2. Additional secondary spiral cord appears between suture and S1 on fifth teleoconch whorl, often forming prominent, bifid cord. Tertiary spiral threads may occur below S1 and S3. Axial sculpture of about 40–60 close-set, narrow, arcuate axial lamellae that overrun cords, forming small tubercles at intersections. Abapically further secondary cords intercalated between primaries forming finely reticulated sculpture. Last whorl with double peribasal cord. Base rapidly contracting, nearly flat, bearing about ten weak cords. Aperture subquadrate, slightly angled at S3. Columella slightly twisted. Columellar callus indistinct. No parietal callus. Siphonal canal short, wide. Outer lip not preserved.
Discussion. Fimbriatella fimbriata is a rather poorly known species. Sacco (1895: pl. 35, fig. 36) and Montanaro Gallitelli & Tacoli (1951: pl. 2, fig. 2) illustrated two specimens from the Tortonian of S. Agata and Montegibbio (Italy) close to the type locality. These show a comparatively stout specimen with delicate and rather uniform sculpture, a simple subsutural cord and a weak keel (Montegibbio), and a relatively slenderer specimen with strongly differentiated spiral sculpture, bifid subsutural cord and prominent keel (S. Agata). The Paratethyan specimens are characterized by a bifid subsutural cord, a feature, which is also depicted in the detailed illustration of the holotype by Michelotti (1847). In addition, Tabanelli et al. (2021) illustrated Pliocene specimens, which are stout, have a strongly differentiated spiral sculpture and a prominent subsutural cord adjoined by a much weaker spiral thread. The nucleus of the protoconch is covered in these Pliocene species but fully exposed in Paratethyan specimens. For the Italian Miocene specimens this information is not available. Based on these somewhat contradicting data we prefer to treat all these specimens as single variable species until a revision of the Mediterranean occurrences is available. Consequently, Fimbriatella fimbriata elegans Montanaro Gallitelli & Tacoli, 1951, from Žabčice (Czech Republic), is treated herein as subjective junior synonym of F. fimbriata, although we note that it differs from all other Paratethyan specimens by the more delicate cancellate sculpture and the weaker S3, and hence weaker keel.
Paleoecology. Middle to outer neritic environments (Tabanelli et al. 2021).
Distribution. Middle Miocene of the Central Paratethys Sea and Late Miocene to Late Pliocene in the (Proto-)Mediterranean Sea.
Central Paratethys Sea. Middle Miocene (Badenian): Alpine-Carpathian Foredeep: Žabčice, Jaroměřice (Czech Republic) (Montanaro Gallitelli & Tacoli 1951; hoc opus); Vienna Basin: Baden (Austria); Eisenstadt-Sopron Basin: Walbersdorf, Forchtenau (Austria) (Sieber 1960); Pannonian Basin: Szob, Sámsonháza (Hungary) (Csepreghy-Meznerics 1969a); B̧kk Mountains: Borsodbóta, B̧kkmogyorósd (Hungary) (Csepreghy-Meznerics 1969b); Făget Basin: Lăpugiu de Sus (Semper 1865).
Proto-Mediterraean Sea. Tortonian (Late Miocene): Po Basin: S. Agata, Stazzano, Montegibbio, Tortona (Italy) (Sacco 1895). Early Pliocene: central Mediterranean, Italy (Tabanelli et al. 2021). Late Pliocene: western Mediterranean, Estepona Basin (Spain).
description: Published as part of Harzhauser, Mathias & Landau, Bernard, 2023, The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition, pp. 1-74 in Zootaxa 5370 (1) on pages 54-56, DOI: 10.11646/zootaxa.5370.1.1, http://zenodo.org/record/10147814