Pupinidius pupinella subsp. altispirus Moellendorff 1901
Creators: name: Wu, Min
name: Zheng, Wei
Publisher: name: Zenodo
Year: 2009
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Enidae
Pupinidius
Pupinidius pupinella
Pupinidius pupinella altispirus (moellendorff, 1901)

description: 14. Pupinidius pupinella altispirus (Moellendorff, 1901)
Figs 1, 6 D–H, 8G, 15
Buliminus (Pupinidius) pupinella altispirus Moellendorff, 1901: 344, pl. 14, figs 4, 5; Kobelt, 1902: 848. Pupinidius pupinella altispirus Yen, 1939: 84, pl. 7, fig. 50.
Material examined. HBUMM-05723 (20 shell measured, 3 with broken apex; specimen-1 for dissection; 1 young): 1661 m a.s.l., 34°0΄127.6΄ N, 104°25΄16.4΄ E, Dangchang, Gansu, loessial area, 2006-X-4, coll. J.Liu & W.Zheng; ZI RAS-N602: type material, det. Moellendorff, 1885-VI-19, coll.? Potanin. SMF-41990: Lectotype. Tan-tshang, Kansu, China. Slg. O.v.Moellendorff. SMF-41991: Paratypes. 3 fmss. Tan-tshang (Dangchang of new spelling), Kansu, China. Slg. O.v.Moellendorff (measurement of types: ewh 1.500–1.656–1.750, wh 6.375–6.500–6.625, h 15.4–16.8–17.9, diam. maj. 11.4–11.8–12.0, aph 8.7–9.3–9.7, apw 7.2–7.8–8.6, h/diam. maj. 1.35–1.40–1.49, h/aph 1.68–1.77–1.87).
Distribution. Gansu (type locality, between Li-dshia-pu and His-gu-tsheng, between Li-dshia-pu and Guan-ting).
Type locality. Dangchang.
Diagnosis. Shell bandless; not or faintly striate. Aperture more or less distinctly with angular tubercle. Most swollen part at penultimate and body whorls. With 6.375–7.000 whorls; height 15.4–18.7, diam. maj. 11.4–14.0. Penis without penial caecum; internally with more than 2 fine longitudinal pilasters. Papilla developed by fused proximal pilasters forming V-shaped structure absent. Epiphallus forming a few loops. Epiphallic caecum absent.
Re-description of shell. Ovate; apex protruded. Most swollen part at penultimate and body whorls. Shell dextral; thin; solid; opaque; glossy; not speckled; with spiral grooves (only at umbilical region); with 6.375–6.588–7.000 whorls. Whorls not shouldered; flattened. Growth lines usually not very clear. Embryonic shell smooth; polished; with 1.375–1.529–1.750 whorls. Postnuclear whorls smooth. Suture normal, without narrow defined zone on beneath whorl. Last whorl gradually ascending towards aperture; rounded at periphery; without smoothed spiral peripheral depression nor rugate region. Shell 16.4–17.5–18.7 high; 12.0–12.7–14.0 in diam. maj. Height/width ratio 1.30–1.38–1.48. Aperture in a plane; completely attached to body whorl; subcircular or truncate-ovate; not adnate; oblique; not armed; more or less distictly with angular tubercle; 9.0–10.0–10.9 high; 7.2–8.4–9.3 wide. Ratio of shell height to aperture height 1.60–1.75–1.92. Peristome reflexed, without distinct cuff; expanded; sharp. Parietal callus indistinct. Columellar margin reflexed. Columella vertical or slightly arched. Outer edge of columellar lip vertical. Umbilicus very narrowly open. Apical whorls normally coloured to differently tinted. Shell multicoloured; and umbilical region. in light greyish brown. From about the third or the fourth whorl on, with a broad whitish spiral band which occupies nearly the whole whorl. Aperture white.
Genitalia. Vas deferens long; of even thickness; entering epiphallus apically with distinct demarcation. Epiphallus long; cylindrical; of uniform thickness; forming a few loops; externally smooth; without caecum. Flagellum short; tubular; proximally normal; with tip blunt. Penis with terminal entrance of epiphallus; clavate; evenly thick; without penial caecum; thin-walled; internally with more than 2 fine longitudinal pilasters; with 1 V-shaped pilaster. V-shaped pilaster with proximal free end approaching at penial retractor insertion; distally not developing into a papilla. Penial process nipple-shaped; situated at boundary between epiphallus and penis. Penial verge absent. Penial appendix long; branched off from penis at some distance from atrium; divided into sections sections including A-1+A-2, A-3 and A-4+A-5. A-1 and A-2 fused; interiorly with more than 2 fine longitudinal pilasters. A-2 near A-3 internally without a ring of papilla. A-2 and A-3 not fused. A-3 opening into A-2 by no papilla. Boundary between A-4 and A-5 indistinct. A-5 long; convoluted. Penial retractor biramous; attaching to penis distally and to A-1+ A-2; with penial retractor arms arising from diaphragm closed to each other. Muscular band connecting vagina and epiphallus absent.. Additional retractor other than penial or appendicular absent. Atrium short; with weak atrial retractor. Free oviduct short; much shorter than vagina. Vagina long; not swollen; straight; not lined with loose, spongy tissue; unpigmented. Bursa copulatrix duct rather long; proximally twisted. Bursa copulatrix with stalk; without apical ligament; normal in size; with long neck; well defined. Diverticle normally present; longer than reservoir; expanded. Bursa copulatrix and diverticle distinguishable; forked more distally from their base.
Measurement of genitalia. P: 4.4; Ep: 19.0; Fl: 0.7; VD: 13.0; Va: 3.9; FO: 2.2; BCD: 26.1; BC: 1.6; D: 4.5; A-1+A-2: 6.0; A3: 1.6. A-4+A-5: 20.2.
description: Published as part of Wu, Min & Zheng, Wei, 2009, A review of Chinese Pupinidius Moellendorff (Gastropoda, Stylommatophora: Enidae), with the description of a new species, pp. 1-31 in Zootaxa 2053 on pages 19-20, DOI: 10.5281/zenodo.186646