Tanyproctus (Tanyproctus) arher Bezdek, Sehnal & Kral, new species
Creators: name: Bezděk, Aleš
name: Sehnal, Richard
name: Král, David
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Melolonthidae
Tanyproctus
Tanyproctus arher

description: Tanyproctus (Tanyproctus) arher Bezděk, Sehnal & Král, new species
(Figs 1, 2, 5, 6, 9)
Type locality. Yemen, Socotra Island, Hala [= Haalla] area, Arher, 12°33.0′N, 54°27.6′E, 5 m a.s.l., freshwater spring in sand dune.
Type material. Yemen (Socotra Isl.): Holotype ♂ (NMPC): “ Yemen, SOCOTRA Island / Hala area, ARHER / freshwater spring in sand dune / 9–10.+ 15-vi-2012 / 12°33.0′N, 54°27.6′E, 5 m // SOCOTRA expedition 2012 / J. Bezděk, J. Hájek, V. Hula, / P. Kment, I. Malenovský, / J. Niedobová & L. Purchart leg. [printed label]”. Paratype ♂ [pinned specimen, left middle leg preserved in pure alcohol] (ZFMK): same data as holotype.
Description of holotype (♂). Body length 8.5 mm. Body elongate; surface dark brown to black coloured, appendages and sutural elytral interval somewhat lighter; shiny; macrosetation pale (Fig. 1).
Head. Labrum small, bilobed; lobes round, coarsely, irregularly punctate. Outline of clypeus almost trapezoidal, with distinctly upturned margin; clypeus remarkably depressed along margin and excavated centrally; anterior margin shallowly emarginate, anterior angles round, sides broadly arcuate (Fig. 2). Genae narrow, round. Frontoclypeal suture feebly arcuate, considerably impressed. Eyes relatively large, distinctly exceeding genae externally in dorsal aspect (Fig. 2); distance between eyes in ventral aspect extended remarkably diameter of eye. Punctation of clypeus coarse and dense, almost evenly distributed but impunctate centrally; punctures separated by approximately their diameter, each puncture bearing short, semierect macroseta. Vertex rather rugopunctate; punctures separated by less than their diameter to confluent, each puncture bearing very short, erect macroseta; macrosetation becoming shorter posteriad. Antennae with 10 antennomeres; antennomere 2 short, approximately as long as wide; antennomeres 3–5 elongate. Club pentamerous, straight, shorter than antennal shaft (antennomeres 1–5 combined). Antennomeres 1–5 with sparse, long macrosetae, club with sparse, short macrosetae. Terminal maxillar palpomeres elongate, rounded apically, absent from depression, approximately of same length as palpomeres 2 and 3 combined.
Pronotum moderately convex, transversal, broadest just anterior to middle except broad basal interruption all around border; anterior bead flat, narrow, distinctly widened medially. Punctures irregular; each puncture bearing long, erect macroseta; macrosetae becoming regularly shorter mediad. Lateral margin considerably coarsely crenate, with row of long macrosetae; basal margin with row of finely and irregularly distributed punctures bearing long, recumbent macrosetae in posterior angles and short macrosetae medially. Anterior angles prominent, projecting over anterior margin, acute-angled, with round apex; sides in approximately anterior half almost straight, divergent posteriad to very broadly obtuse posterior angles; posterior margin broadly round. Surface glabrous and finely, coarsely, slightly irregularly punctate, punctures separated by 2–4 their diameters, area along medial longitudinal line smooth.
Scutellum approximately as wide as long, triangulate, sides broadly arcuate, apex acute, surface with several punctures basally and laterally.
Elytra convex, slightly dilated posteriad, sutural angle rounded; striae excepting sutural stria missing or only very feebly indicated; distinctly microsculptured with disc feebly transversally wrinkled; punctation coarse, dense, almost regular; punctures separated by 1–2 their diameters. Sutural interval slightly convex; sutural stria with row of irregularly distributed punctures; lateral margin distinctly bordered with row of long, erect setae.
Macropterous.
Legs. Femora moderately shiny; metafemora with entire surface sparsely, almost regularly punctate; punctures bearing long, recumbent macrosetae. Protibia tridentate; basal tooth considerably weak; terminal calcar long, sharp, slightly curved externally, acute apically, inserted against emargination between basal and medial teeth. Mesotibia and metatibia slightly expanded apicad, with two setiferous transversal carinae. Mesotibial terminal calcars equal in length, flattened, acute apically. Metatibial terminal calcars equal in length, considerably flattened, acute apically. Protarsomeres 2–4 considerably dilated, protarsomere 2 with length-width ratio: 1:2. Mesotarsomeres 2–4 remarkably more slightly dilated that those of protibiae; protarsomeres and mesotarsomeres 1-4 with shortly and densely macrosetaceous pads ventrally; metatarsomeres covered with long sparse macrosetae ventrally. Claws bifid.
Ventral surface of thorax with dense, long, recumbent macrosetation.
Propygidium microsculptured, coarsely, sparsely, irregularly punctate; pygidium finely microsculptured; border complete; surface coarsely and irregularly punctate.
Ventrit es almost bare, remarkably coarsely and irregularly punctate.
Male genitalia (Figs 5–6). Aedeagus symmetrical; parameres slender, relatively long, only slightly shorter than phallobase, distal part not dilated in dorsal aspect.
Female. Unknown.
Varability. The only known paratype slightly differs in size (total body length: 8.4 mm) and with lighter coloured elytra.
Differential diagnosis. Tanyproctus (T.) arher is morphologically very similar to the other endemic Socotran Tanyproctus species (Král et al. 2012). In the key to males of Socotran Tanyproctini (Král et al. 2012: 180), T. (T.) arher will key to the couplet with T. (T.) wraniki. For differentiation from this species see the complex of diagnostic characters in Table 1.
Sehnal & Bezděk, 2012
Etymology. Derived from area of origin of the new species, the Arher freshwater spring, Socotra (Yemen); noun in appositon.
Collecting circumstances. Both specimens were captured at light at about 6 p. m. (J. Hájek, personal communication).
Geographical distribution. Endemic species to the Socotra Island; both so far known specimens originates from the coastal area of Haalla (E Socotra), for details see Bezděk et al. (2012).
description: Published as part of Bezděk, Aleš, Sehnal, Richard & Král, David, 2013, Tanyproctus (Tanyproctus) arher (Coleoptera: Scarabaeidae: Melolonthinae: Tanyproctini), a new species from the Socotra Island, Yemen, pp. 191-196 in Zootaxa 3737 (2) on pages 192-196, DOI: 10.11646/zootaxa.3737.2.7, http://zenodo.org/record/224230