Naarda hallasana Tóth & Ronkay, 2015, sp. n.
Creators: name: Tóth, Balázs
name: Ronkay, László
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Noctuidae
Naarda
Naarda hallasana

description: Naarda hallasana sp. n.
male: fig. 30, female: fig. 31
Type material. Holotype. ♂: South Korea, No. 1631–1634; Prov. Cheju, Cheju-do Island, Mts. Halla-san, at the eastern edge of the Halla-san National Park; Songpanak, 750 m, 23.VIII.1992; at light, leg. L. Ronkay L. & A. Vojnits; slide No. TB 811m (coll. HNHM).
Paratypes. ♂, with data as holotype, slide No. TB 838m (coll. HNHM), ♀: Korea, Prov. Cheju, Mt. Hallasan, 5 km SW of Cheju city, 500 m, 24.VII.1992, No. 1639, leg. L. Ronkay & A. Vojnits; slide No. RL7660f (coll. HNHM).
Diagnosis. Overall external appearance of N. hallasana is similar to N. maculifera (Staudinger, 1892) due to the size and shape of forewing and labial palps. The new species is, however, has more brownish ground colour than N. maculifera, moreover the reniform stigma contains two black dots and the base of male forewing has a light yellow brush while the stigma of N. maculifera is monochromatic yellow (sometimes with single dark scales) and the males do not have any brush. The male genitalia show clear relationship with N. pectinata Sugi, 1982; the uncus is even broader, the structure originating from editum is very strong, broad and rigid while in N. pectinata this structure is longer, weaker and its sclerotisation is not stronger than that of the valval plate. Additional common feature of the two species is the fusion of saccular and costal processes beyond the base of cucullus. This fused structure forms a huge, intensively dentate process reaching the mid-point of cucullus in N. hallasana, while in N. pectinata this part is plate-like, rounded and hardly extends beyond the basal third of cucullus.
Description. Wingspan 16 mm, length of forewing 8 mm. Antennae filiform, in male cilia 1.5 times longer than diameter of flagellum and arranged into three bands per segment; male setae absent; female antennae lost from the only known specimen. Labial palps 4.5 times longer than diameter of eye, with elongate and apically light 3rd segment in both sexes, dorsal edge of palp domed, obtuse-angled in male, in female apparently straight due to loss of dorsal scales. Scale-hood of vertex short in male, very short in female. Foretibiae slightly hairy in male. Characteristic wing pattern features: weak fasciae and relatively small stigmata on greyish-brown forewing; subterminal line lighter than ground colour, fragmented to distinct spots arranged to a straight line; postmedial line relatively broad, minutely sinuous (nearly straight); medial and antemedial lines not visible; reniform stigma oval, with bigger black dot in lower third and tiny dot in top third, filling of reniform stigma deep ochreous in male, light ochreous (faded?) in female; orbicular stigma small but visible, its ground colour like that of reniform; hindwing ground colour slightly lighter, medial line slightly more conspicuous than those of forewing. Forewing costa of male not concave. Forewing underside with basal brush composed by pale yellow, elongate scales being slightly longer than the quarter of forewing length. Female specimen in a worn condition. Male genitalia (fig. 3). Uncus very broad, straight, with average length, truncate, junction of ventral and distal margins with tiny hook. Scaphium straight, as long as uncus. Tegumen longer than vinculum; saccus broad-based, long, gradually tapering to a rounded tip. Juxta large, plain, triangular, dorsal edge straight, ventral edge right-angled (damaged in the figure). Dominant character of the genitalia being a large, very strong, broad-based, slightly curved process with pointed tip and granulose (dentate at edges) surface. This process having complex system of connections: ventral corner of its based fused with the dorso-medial corner of sacculus but connected also to juxta and, in certain extent, to the ventral end of tegumen. Originating from editum and transtilla. Valva relatively narrow-based, comparatively long, abruptly tapering. Cucullus narrow, free in the lateral half section of valva, its tip rounded. Sacculus short, narrowbased, middle section very broad, extending towards dorsal edge of valva, then abruptly tapering and ending at the base of free section of cucullus. Saccular extension and costa fused beyond the base of cucullus, raising as huge, very strong process; this process being long, broad, minutely curved, covered by spines of various size in irregular arrangement. Aedeagus (damaged) stout, straight, vesica (partially everted) smooth, with two small scobinate areas oppositely. Female genitalia (fig. 17). Papillae anales angular. Apophyses posteriors ca 1.5 times longer than apophyses anteriores. Sternum A7 narrow, bilobed, anterior margin scobinate, lobes triangular; sinus very broad and very abruptly tapering anteriorly; lamella antevaginalis present, U-shaped, thick, wrinkled with creasy appearance; antrum broad; ductus bursae broad and short; corpus bursae ovoid, relatively large, surface of its posterior 2/3 part densely scobinate by tiny grains, remaining section of bursa with signum field composed by radially oriented, small, sparse spines (overwhelmed by the spermatophore in the figure).
Distribution. Korea: Jeju Island.
Etymology. The species was hitherto found only in the Mt. Halla-san.
Taxonomic remarks. The female genitalia of this species were illustrated by Kononenko & Han (2007) as those of Gynaephila maculifera Staudinger, 1892. The lack of setae on male antennae is a very rare character state in the genus. For systematic consideration see the Diagnosis.
description: Published as part of Tóth, Balázs & Ronkay, László, 2015, Revision of the Palaearctic and Oriental species of the genus Naarda Walker, 1866 (Lepidoptera: Erebidae, Hypeninae). Part 5. Description of 13 new species from Asia, pp. 179-201 in Zootaxa 3925 (2) on pages 181-182, DOI: 10.11646/zootaxa.3925.2.2, http://zenodo.org/record/232614