Lesbosoxylon kemaliyensis Akkemik & Mantzouka 2020, sp. nov.
Creators: name: Akkemik, Ünal
name: Mantzouka, Dimitra
name: Yildirim, Demet Kiran
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Pinopsida
Pinales
Protopinaceae
Lesbosoxylon
Lesbosoxylon kemaliyensis

description: Lesbosoxylon kemaliyensis Akkemik & Mantzouka, sp. nov. (Figs 3-6)
Plant Fossil Names Registry Number: PFN001780
HOLOTYPE. — Specimen KEM01; repository: Istanbul University-Cerrahpasa, Faculty of Forestry, Department of Forest Botany (designated here).
ETYMOLOGY. — The epithet “ kemaliyensis ” comes from the name of the main city “Kemaliye” nearby the fossil site.
TYPE HORIZON. — Divriği Formation.
AGE. — Early-middle Miocene.
TYPE LOCALITY. — Kemaliye (Eğin), Erzincan, Turkey.
DIAGNOSIS. — Coniferous wood with resin canals. Growth ring boundaries distinct, transition from earlywood to latewood mostly gradual, axial and radial resin canals with thin-walled epithelial cells, latewood tracheids thin to thick walled, bordered pits on radial walls of tracheids 1-2(-3) seriate, crassulae common, rays heterocellular, uniseriate, partly biseriate, uniseriate rays up to 27 cells high, fusiform rays up to 30 seriate high. Axial parenchyma occasionally present. Ray tracheids 2-3 rows, cell walls of ray tracheids smooth. Cross-field pitting pinoid, 1-2(-6) pits per cross-field, horizontal and end walls of ray parenchyma cells smooth and slightly nodular, horizontal walls occasionally pitted, indentures not observed.
DESCRIPTION
The petrified wood sample has grey color; dimensions are 7 × 8 × 19 cm (Fig.3). In cross section,growth ring boundaries are distinct, transition from earlywood to latewood is mostly gradual (Fig.4A, B). Measured diameters of tracheids and pits are given inTable 1. Axial resin canals with thin walled epithelial cells present mostly in latewood zone (Fig. 4A, B). Traumatic resin canals present (Fig. 4C). In tangential section, rays uniseriate (Fig.4D), and fusiform rays present (Fig. 4E). Rays occasionally biseri-ate or partly biseriate (Fig. 5A). Big rays with idioblast-like cells present (Fig 5 B-D). Ray cells uniform in some rays, and heterocellular, wider or narrower (Fig. 5 D-E). Height of uniseriate rays up to 27 cells (Fig. 4C), height of fusiform rays up to 30 cells and wider (Fig. 4D and Fig. 5D). Average height of rays 4-15 cells. Radial resin canals with thin–walled epithelial cells. Axial parenchyma cells occasionally present, and generally smooth transverse end walls (Fig. 5 F-G and I), and slightly nodular (Fig. 5H). Axial resin canal visible in radial section (Fig. 6A). Tracheid pitting in radial walls of earlywood 1-2 (-3) seriate opposite and crassulae formation common (Fig. 5 B-D). Cross-field pitting pinoid in 1(-2) rows of usually 1-2(-6) pits per cross-field (Fig. 6E, F). End walls of ray parenchyma cells smooth, no indentures observed. Rays heterocellular. Ray tracheids present, 1-3 seriate, smooth walled (Fig. 6G). Ray cells smooth walled (Fig. 6 E-H) and occasionally pitted. These descriptive features are also provided in Appendix A based on “Criteria used according the IAWA for fossil conifers” by Wheeler et al. (2004).
description: Published as part of Akkemik, Ünal, Mantzouka, Dimitra & Yildirim, Demet Kiran, 2020, The first report of Lesbosoxylon Süss & Velitzelos from the early-middle Miocene of eastern Anatolia, pp. 427-441 in Geodiversitas 42 (23) on page 432, DOI: 10.5252/geodiversitas2020v42a23, http://zenodo.org/record/4447671