Macroscytus brunneus
Creators: name: Lis, J. A.
Publisher: name: Zenodo
Year: 2000
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Cydnidae
Macroscytus
Macroscytus brunneus

description: Macroscytus brunneus (FABRICIUS)
(Figs 94­98)
Cydnus brunneus FABRICIUS, 1803: 185.
Macroscytus brunneus: FIEBER 1861: 362.
Cydnus proximus RAMBUR, 1839: 112 (syn. by AMYOT 1845: 427).
Aethus opacus STÃL, 1854: 214 (syn. by SIGNORET 1881: 644).
Macroscytus scutellaris HORVÁTH, 1919: 238 (syn. by LINNAVUORI 1993: 25).
Macroscytus exiguus HORVÁTH, 1919: 238 (syn. by LINNAVUORI 1993: 25).
Macroscytus subaeneus, not of DALLAS: AHMAD & MOIZUDDIN 1977: 76; AHMAD et al. 1979: 15 & 18; MOIZUDDIN & AHMAD 1985: 163, 1990: 325 & 326.
DESCRIPTION
Body 6.00­ 9.13 mm in length, 3.25­4.86 mm in width, from pale brown to blackish brown, corium usually paler than pronotum and scutellum. In general appearance (habitus and body puncturation) very similar to M. badius.
Head dorsally impunctate, sometimes with distinct wrinkles and several almost indistinct tiny punctures on paraclypei; clypeus without subapical setigerous punctures (in several aberrant specimens one or two setigerous punctures on clypeus were found); each paraclypeus with a row of 4­8 submarginal hair­like setae (Fig. 95); ocular index 2.20­3.35; length of antennal segments (in mm): 0.30­0.43: 0.38­0.61: 0.28­0.51: 0.48­0.71: 0.54­0.77.
Pronotum weakly punctate, sometimes almost impunctate, transverse discal impression behind calli absent or very shallow; lateral margins with 8­13 submarginal setigerous punctures (Fig. 94). Anterior convexity of propleuron impunctate or almost impunctate, posterior convexity with a few punctures, propleural depression striated, sometimes bearing a row of more or less distinct punctures.
Scutellum more or less punctate, puncturation usually denser than that of pronotal disc.
Corium distinctly punctate; clavus with one complete and two partial rows of punctures; mesocorial disc almost evenly punctate with small punctures; exocorium almost evenly punctate, punctures larger than those on mesocorium; costa narrow, almost entirely separated from exocorium, bearing 4­7 setigerous punctures (Fig. 94).
Hind femora with clearly visible subapical tooth on dorsal margin, that may vary in size; hind tibiae simple (Fig. 96).
Abdominal sterna laterally slightly transversely wrinkled, impunctate, except for a few small punctures posterior to spiracles.
Male genitalia as in Figs 97­98.
TYPE DATA
Lectotype male of Cydnus brunneus FABRICIUS (designated by LIS, 1994a: 239): Morocco, Tanger (ZMUC). Lectotype male of Macroscytus scutellaris HORVÁTH (designated by LINNAVUORI, 1977: 41): Mozambique, East Africa, Zambesi (HNHM). Syntype (s) of Cydnus proximus RAMBUR: Spain, env. of Cadiz (BMNH). Syntype (s) of Aethus opacus STÃL: South Africa, Caffraria (NNML). Lectotype male of Macroscytus exiguus HORVÁTH (designated by LINNAVUORI, 1977: 41): South Africa, Natal (HNHM). Lectotype male of Macroscytus exiguus HORVÁTH (designated by LINNAVUORI, 1977: 41): South Africa, Natal (HNHM).
MATERIAL EXAMINED
Lectotype of Cydnus brunneus, Lectotype of Macroscytus scutellaris, Lectotype of Macroscytus exiguus, and 379 other specimens: Afghanistan: KizylKha, 2 exx. (UO). Algeria: Laghouat, 1 ex. (MNHN); Randon, 1 ex. (MNHN). Botswana: Gaborone, 1 ex. (MZLU). Canary Islands: Gran Canaria, Artedara b. Fataja, 2 exx. (TLM); Gran Canaria, Ginamar, 3 exx. (MNHN). Cyprus: Paphos, 11 exx. (CMNH, UO). Egypt: Cairo, 2 exx. (NNML). Greece: Corfu, 3 exx. (NNML). Iran: 25 km S Minab, 1 ex. (NHMW); Bandar Langeh, 12 exx. (MNHN, UO). Iraq: Bagdad, 3 exx. (HNHM, UO). Israel: Askhelon, 6 exx. (CMNH); Hullah Valley, 2 exx. (ZSMC); EnGedi, 1 ex. (ZSMC). Italy: Sardinia, 1 ex. (HNHM). Jordan: Wadi Sir, n. Amman, 1 ex. (HNHM); Arda Road, 1 ex. (HNHM). Morocco: O El Moddah, 1 ex. (UO); Safi­Agadir road, 1 ex. (UO); Oudjda, 3 exx. (MNHN, UO). Mozambique: Lourenco Marques, Matola, Vila Salazar, 1 ex. (JEC). Namibia: Bushmanland, Klein Dobe, 11 exx. (MNHU, UO); Fish­River­Canyon, 6 exx. (MNHU, UO); Etoscha N.P., Namutoni, 6 exx. (MNHU, UO); Kavango, Popa Falls, 3 exx. (MNHU); East Caprivi, Mudumu N.P., Makatwa, 3 exx. (MNHU, UO); East Caprivi, Katima Mulilo, 2 exx. (MNHU); Ongongo Fall, Warmbad, Kaokoveld, 3 exx. (MNHU, UO); Kunene, 9km W Ruacana, 2 exx. (MNHU). Oman: Ruwi, 2 exx. (MZLU). Pakistan: Karachi, 4 exx. (NHMUK); Baluchistan, Quetta, 1 ex. (BMNH). Portugal: Algarve, Sta Barbara de N., 17 exx. (MNHN, UO). Saudi Arabia: Najd Region, 50 mi SE Riyadh, 6 exx. (JEC). Senegal: Dakar Bel­Air, 1 ex. (MNHN). South Africa: Natal, Empangeni area, 4 exx (NMP, UO); Natal, Mntunzini Garland Fm, 1 ex. (NMP); Natal, Mkuzi Game Reserve, 1 ex. (NMP); Mkuze Game Res., 1 ex. (UO); Natal, Itala Game Reserve, Louwsburg, 2 exx. (MNHU); Cape Prov., Oranje River, Vioolsdrif, 1 ex. (MNHU); Transvaal, Zoutpansberg [= Soutpansberg], 1 ex. (ZMPA). Tunisia: Boughrara, 4 exx. (HNHM); Maknassy, 8 exx. (MNHN, UO). Turkey: Prov. Urfa, Halfeti, valley of Euphrat, 2 exx. (HNHM). Turkmenistan: Kopet­Dagh Mts, 6 km S of Ipay­Kala, 1 ex. (HNHM); Kopet­Dagh Mts, 5 km S of Chull, 1 ex. (HNHM). Zambia: South Luangwa N.P., Mfuwe, Crocodile Farm, 1 ex. (MNHU); Muchinga Escarpment, 1 ex. (MNHU). Zimbabwe: St. Benedict, 30km NE Macheke, 2 exx. (TLM); Chinhoyi Caves, 1 ex. (NMP); Victoria Falls, Zambesi Nat. Park, 1 ex. (MNHU); Matabeleland, Lake Mutirikwi bei Masvingo, Kyle N.P., 11 exx. (MNHU, UO); Matabeleland, Matopos Nat. Park, 25 exx. (MNHU, UO); S. Rhodesia, Kariba, 182 exx. (JEC, UO).
DISTRIBUTION
Afghanistan, Albania, Algeria, Azerbaidjan, Botswana, Cameroon, Canary Islands, Cape Verde Islands, Central African Republic,?China (North), Congo, Croatia, Cyprus, Egypt, Ethiopia, France, Gambia, Ghana, Greece, Guinea, Iran, Iraq, Israel, Italy, Ivory Coast, Jordan (first record), Kazakhstan, Kenya, Liban, Libya, Macedonia, Mali, Malta, Morocco, Mozambique, Namibia (first record), Nigeria, Oman (first record), Pakistan, Portugal, Saudi Arabia, Senegal, Socotra, Somalia, South Africa, Spain, Sudan, Switzerland, Syria, Tadzhikistan, Tanzania (incl. Zanzibar), Turkey, Turkmenistan, Tunisia, Uganda, Uzbekistan, Yemen, Yugoslavia, Zaire, Zambia (first record), Zimbabwe (first record).
description: Published as part of Lis, J. A., 2000, A revision of the burrower-bug genus Macroscytus Fieber, 1860 (Hemiptera: Heteroptera: Cydnidae), pp. 359-509 in Genus (Wrocław) 11 (3) on pages 401-403