Therevidae Irwin & Winterton, 2024, gen. nov.
Creators: name: Irwin, Michael E.
name: Winterton, Shaun L.
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Therevidae

description: Key to genera of Malagasy Therevidae and species of Rinhatiana gen. nov. and Tianarinha gen. nov.
1. Femora with regular filiform setae of varying lengths, lacking adpressed scale-like setae; costal vein ending between veins R 4 and CuA+CuP, never circumambient around wing; female terminalia with, at most, a single set of acanthophorite spines ..............................................................2 (Therevinae)
– Femora with regular filiform setae of varying lengths, admixed with adpressed scale-like setae, especially on mid and hind legs; costal vein circumambient around wing; female terminalia with two distinct sets of acanthophorite spines on each side (A1 and A2) ...................................................... 3
2. Dorsal surface of wing vein R 1 with setulae; palpus with apical pit; female sternite 8 without strong macrosetae; genitalia with three spermathecae and no spermathecal sac ........................................... ........................................................................................ Rueppellia Wiedemann, 1830 (Phycusinae)
– Wing vein R 1 without setulae; palpus without an apical pit; female sternite 8 with strong macrosetae; genitalia with two spermathecae and spermathecal sac is present ...................................................... ................................................................................. Microgephyra Lyneborg, 1972 (Xestomyzinae)
3. Wing vein R 1 without setulae ............................................................................................................ 4
– Dorsal surface of wing vein R 1 with setulae ..................................................................................... 5
4. Sternum with setae present medially in furrow; mid coxae with setae on posterior surface .............. ................................................................................................................... Irwiniella Lyneborg, 1976
– Sternum with setae present laterally only, rarely in medial furrow; mid coxae without setae on posterior surface .................................................................................................. Stenopomyia Lyneborg, 1976
5. Prosternum with setae medially; one pair of scutellar macrosetae; frons with dark setae, pubescence grayish-brown with a dark band from base of antenna to eye margin (Fig. 14); gena with dark setae ventrally; abdomen predominantly brown .................................................. Tianarinha gen. nov. ...6
– Prosternum without setae; two pairs of scutellar macrosetae; frons usually lacking setae, pubescence silver or metallic tan around antennal bases, matte black above this and frequently grayish-silver dorsally to ocellar tubercle (e.g., Fig. 9); gena with pale setae; abdomen predominantly bright yellow, sometimes brown laterally along tergites ................................................... Rinhatiana gen. nov. ...7
6. Extensive dark setae on the lower part of occiput, gena (Figs 10; 14A–B) and coxae; flagellum style relatively elongate; few dark setae on frons just above antennae; setae on posterior surface of hind coxae (Fig. 11B) ..................................................................................... T. goodmani gen. et sp. nov.
– Setae on the lower head and coxae fewer in number and shorter (Fig. 12A, 13A); flagellum style relatively short; frons only with setae dorsally, no setae just above antennae; setae absent on posterior surface of hind coxae (Fig. 13A) ................................................................... T. micet gen. et sp. nov.
7. Two pairs of supra alar macrosetae; dorsocentral macrosetae present (Figs 5–6); Male frons width at narrowest point slightly wider than anterior ocellus (Fig. 9E); short, dark setae present dorsally on frons .................................................................................. R. distincta (Lyneborg, 1976) comb. nov.
– One pair of supra alar macrosetae; dorsocentral macrosetae absent; male frons width at narrowest point narrower than anterior ocellus, eyes almost contiguous (Fig. 9A, C, G); frons lacking setae .. ........................................................................................................................................................... 8
8. Frons with metallic-tan pubescence around base of antennae (Fig. 9C, D); abdomen dark brown (yellow dorsally in female), cream-white posteriorly on each segment, silver pubescence absent (Figs 3–4) ................................................................................................ R. cracentis gen. et sp. nov. – Frons with silver pubescence around base of antennae and matte black area medially (Fig. 9A, G); abdomen predominantly yellow, sometimes brown laterally along tergites, triangular patches of silver pubescence laterally on at least tergites 2 and 3 ............................................................................... 9
9. Male frons with matte-black pubescence relatively narrow, brown pubescence dorsally (Fig. 9A); female frons with dorsomedial stripe of matte-black pubescence relatively narrow, suffused with brown pubescence laterally (Fig. 9B); male terminalia black (Figs 1–2) ...... R. arctifestuca gen. et sp. nov.
– Male frons with matte-black pubescence broader, lacking brown pubescence dorsally (Fig. 9G); female with dorsomedial stripe of matte-black pubescence broader, mostly lacking brown pubescence in upper portion (Fig. 9H); male terminalia yellow (Figs 7–8) ............ R. latifestuca gen. et sp. nov.
description: Published as part of Irwin, Michael E. & Winterton, Shaun L., 2024, New genera of stiletto flies endemic to Madagascar (Therevidae: Therevinae), pp. 1-33 in European Journal of Taxonomy 932 (1) on pages 30-31, DOI: 10.5852/ejt.2024.932.2507, http://zenodo.org/record/11001504