Acantholycosa sayanensis Marusik, Azarkina &amp; Koponen 2004
Creators: name: Fomichev, Alexander A.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Lycosidae
Acantholycosa
Acantholycosa sayanensis

description: <i>Acantholycosa sayanensis</i> Marusik, Azarkina &amp; Koponen, 2004 Figs 1–2, 14–15, 20–23, 32–33, 54–56, 72, 74–76 <i>A. sayanensis</i> Marusik, Azarkina &amp; Koponen, 2004: 107, f. 10–14 (&amp;male;). <i>A. sayanensis</i> — Marusik &amp; Kovblyuk 2011: 184, f. 22.12, 15, 21 (&amp;male;). <b>Material examined.</b> RUSSIA: <b>Krasnoyarsk Krai</b>: holotype &amp;male; (ISEA, 000.622), Yermakovski District, Western Sayan Mt. Range, Oiski Pass, 35–40 km SW of Oiskoe Lake, 11.07.1990, N.A. Gladkevich, S.E. Chernyshov; 11&amp;male; 3&amp;female; (ISEA, 001.8651), 1&amp;female; (ISEA, 001.8721), Borus Mt. Range, Malyi Borus and Bol’shoi Borus (= Poilova) Mountains, 52°47–48’N, 91°31–33’E, scree, 1500–2300 m, 16.07.2020, A.A. Fomichev; 5&amp;male; 12&amp;female; (ZMMU), Borus Mt. Range, Pelekhova Mt. Pass, 52°49’N, 91°31’E, scree, 1300 m, 13, 15.07.2020, A.A. Fomichev, V.V. Sergeev; <b>Tuva Republic</b>: 3&amp;male; 3&amp;female; (ISEA, 001.8652), Western Sayan Mt. Range, Sayanskyi Mt. Pass, 5 km SSW from Sabalkias Mt., 51°43’N, 89°54’E, scree and alpine meadow, 2400–2600 m, 07.06.2017, A.A. Fomichev. <b>Comparative material:</b> <i>Acantholycosa azyuzini</i> Marusik, Hippa &amp; Koponen, 1996 (Figs 57–59): RUSSIA: <b>Altai Republic</b>: 1&amp;female; (ISEA, 001.8663) Katunskyi Mt. Range, watershed of Mul’ta River and Kuyguk River, 49°59.019’N, 85°51.096’E, scree, 2400 m, 26.07.2019, A.A. Fomichev, E.A. Nepaeva, Yu.V. Dyachkov. <b>FIGURES 20–31.</b> Bulb (20, 24, 28) and embolic division (21–23, 25–27, 29–31) of <i>Acantholycosa sayanensis</i> (20–23), <i>A. sidorovi</i> <b>sp. n.</b> (24–27) and <i>A. levinae</i> (28–31). 20, 23–24, 27–28, 31—retrolateral; 21, 25, 29—ventral, 22, 26, 30—anterior. Scale bars= 0.2 mm. Abbreviations: <i>Em</i> embolus, <i>Pl</i> palea, <i>Po</i> paleal outgrowth, <i>Sp</i> spine, <i>Ta</i> terminal apophysis, <i>Ts</i> triangleshaped projection of the palea. <b>Diagnosis.</b> <i>Acantholycosa sayanensis</i> is similar to <i>A. altaiensis</i> and <i>A. azyuzini</i>. The female of <i>A. sayanensis</i> is similar to these species in having diamond-shaped epigynal fovea (<i>Fo</i>). The female of <i>A. sayanensis</i> differs from those of <i>A. altaiensis</i> by having septal base (<i>Sb</i>) almost as large as the epigynal fovea (vs. much smaller) and diverging receptacles (<i>Re</i>) (cf. Figs 54–56 and figs 8, 9 in Marusik <i>et al.</i> 2004). It can be distinguished from the female of <i>A. azyuzini</i> by the rugose (vs. smooth) surface of the septal base (cf. Figs 54, 57). The male of <i>A. sayanensis</i> is similar to those of <i>A. altaiensis</i> and <i>A. azyuzini</i> in having broad and straight embolus (<i>Em</i>), claw-like paleal outgrowth (<i>Po</i>) and spine-like terminal apophysis (<i>Ta</i>). The males of <i>A. sayanensis</i> differ from those of <i>A. altaiensis</i> in having rounded and solid anterior part of the tegular apophysis (<i>Tg</i>) (vs. triangular and laminar anterior part), a small, almost indistinct triangular spine on the base of the embolus (vs. large outgrowth) and terminal apophysis bent outward (vs. directed inward) (cf. Figs 21–22, 32 and figs 2–3, 6 in Marusik et al. (2004)). Finally, the male of <i>A. sayanensis</i> can be distinguished from <i>A. azyuzini</i> by the embolus with a widened distal part (vs. pointed) and by the presence of a small spine on the base of embolus (vs. absence, cf. Figs 21–22 and fig. 15 in Marusik <i>et al.</i> 2004). <b>Description.</b> Female. Total length 9.6. Carapace: 4.4 long, 3.6 wide. General appearance as in Fig. 2. Coloration. Carapace, chelicerae and sternum dark brown. Eye field black. Endites and labium yellow-brown. Coxae and palps brown with yellow spots. Legs dark brown. Femora with yellow-brown annulation. Abdomen dark gray, almost black dorsally, with dark brown cardiac mark. Venter of abdomen yellow-gray. Spinnerets dark gray, almost black. Spination of leg I: Fe d1-1-1 p0-0-2 r1-0-1; Pa r1; Ti p1-0-0 r1-0-0 v2-2-2-2-2-2; Mt p1-1-0 r1-1-0 v2-2-0. Leg measurements: I 4.35, 1.95, 4.05, 3.8, 1.8 (15.95); II 4.4, 1.95, 3.75, 3.95, 1.75 (15.8); III 4.3, 1.8, 3.5, 4.8, 1.9 (16.3); IV 5.25, 1.95, 4.6, 7.25, 2.55 (21.6). Epigyne and internal genitalia as in Figs 54–56. Fovea as wide as long. Septal base fills almost the entire fovea. Septal stem (<i>Ss</i>) as long as septal base. Lips of the epigyne (<i>Li</i>) not touching each other. Apical pocket (<i>Ap</i>) two times narrower than septal stem length, with one hood. Receptacles not clearly delineated from copulatory ducts (<i>Cd</i>). Copulatory ducts as long as the receptacles. Male. See Marusik <i>et al.</i> (2004) for description. General appearance as in Fig. 1. Palp as in Figs 14–15, 20– 23, 32–33. Tegular apophysis hammer-shaped, without apical arm and with a short basal arm (<i>Ba</i>). Palea (<i>Pl</i>) rounded. Paleal outgrowth large, hooked. Terminal apophysis spine-like in the apical part, almost touching the paleal outgrowth. Embolus expanded distally, with a hook-like tip in anterior view. <b>Distribution.</b> The species is known from three localities in the Western Sayan Mountains, which are located more than 100 km apart (Figs 74–76). This species appears to have one of the broadest ranges among its endemic to Siberia congeners.
description: Published as part of <i>Fomichev, Alexander A., 2021, New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia, pp. 567-585 in Zootaxa 5026 (4)</i> on pages 568-572, DOI: 10.11646/zootaxa.5026.4.7, http://zenodo.org/record/5300792
description: {"references": ["Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148.", "Marusik, Y. M. &amp; Kovblyuk, M. M. (2011) Spiders (Arachnida, Aranei) of Siberia and Russian Far East. KMK Scientific Press, Moscow, 344 pp.", "Marusik, Y. M., Hippa, H. &amp; Koponen, S. (1996) Spiders (Araneae) from the Altai area, Southern Siberia. Acta Zoologica Fennica, 201, 11 - 45."]}