Clivina depressiforma Balkenohl, 2025, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina depressiforma

description: Clivina depressiforma sp. nov.
(Figs 18, 53, 87, 113)
Type material. Holotype: ♂, with labels and data: yellow, black printed “ BORNEO: SABAH Bingkor N Keningau 400-500 m, 19.-20.XI.1996 leg. W. SCHAWALLER” / white, black printed “ Clivina nr. vigil DARL. det. Balkenohl, 1998” / “ Coll. M.Balkenohl ” (SMNS). Paratypes: 8 specs, same data as holotype (SMNS, CMBB); 2 specs, with labels and data: yellow, black printed “ BORNEO: SABAH, Bingkor N Keningau 400-500 m, at light; 20.XI.1996, leg. D.Grimm ” / “ Clivina nr. vigil DARL. det. Balkenohl, 1998” / “comp. ≠ vigil aus AUS/Baehr” (SMNS); 2 specs, with labels and data: white, black printed “ BORNEO, Sabah, Danum Valley, Rainforest Lodge, 19.-20.X.2009, leg. U.+ H.J. BREMER” (SMNS); 1 ♂, 1 ♀, with labels and data: white, black printed “SARAWAK (Borneo) ca 40 km SE KAPIT 03.1994, J. Kodada leg.” (NHMW); 2 ♀♀, with labels and data: white, black printed “Malaysia, Sabah, Crocker Range National park, Long-kogungan env., ca 750-850 m a.s.l., 19.-21.VI.1996, 7 c (NHMW); 1 ♀, with labels and data: white, black printed “BRITISH N. BORNEO Tawau Residency Kalabakan R., Tawau 48km W, 9-18.XI.’58” / “Primary Forest” / “ T. C. Maa Collector BISHOP” (NHMUK); 2 ♀♀, with labels and data: white, black printed “ MALAYSIA – Sabah prov. Banjaran Crocker Mts. GUNUNG ALAB peak 30.IV.-27.V.1996. 1650- 1800m M. Strba & R. Hergovits leg.” / “VII. helferi Gr. det. Dr. A. DOSTAL 2000” / red, handwritten “sp. nov. ad. depressa” (CADW).
Description. Measurements. Body length 5.23–6.02 mm (ẍ = 5.47 mm *), width 1.55–1.77 mm (ẍ = 1.66 mm *), ratio L/W of pronotum 0.89–0.93 (ẍ = 0.90), ratio L/W of elytra 1.74–1.81 (ẍ = 1.77*); (*n = 10). Colour glossy, fuscous to ferruginous. Supraantennal plate at margin and clypeal-wing translucent. Antennae, mouthparts, intermediate and hind legs yellowish. Antennomeres darker from joint four onwards.
Head a quarter narrower than pronotum. Clypeus with central part straight, with slightly projecting sharp teeth laterally, separated from clypeal-wing by distinct notch; wing distinct, much less projecting as clypeal-teeth, rounded, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin laterally obtuse-angularly rounded, clypeus, wing, and supraantennal plate slightly reflexed margined. Supraantennal plate extended posteriorly into a somewhat widened keel, separated from wing by furrow. Clypeus and frons developed as continuous moderately convex field, with scattered punctures, separated from supraantennal plate and supraorbital carina by wide roughly reticulated furrow, with clypeal seta in front; supraorbital carinae narrow, with two supraorbital setigerous punctures at middle of eye and distinctly posterior end of the gena. Neck constriction closed, formed by a step from frons, neck covered with irregular partly connected punctures. Eye prominent, regularly sized, gena distinct. Antenna elongate, reaching distinctly over base of elytron. Labrum nearly sub-circular, irregularly reticulated, seven-setose. Mentum isodiametrically reticulated, lobe with askew wrinkles, convex anteriorly, median tooth nearly rectangular at tip, not as high as lobes.
Pronotum (Fig. 53). Conspicuously flattened but still slightly convex in frontal and lateral view. Distinctly cordate, wider than long. Reflexed lateral margin slightly crenulate, convex, widest at middle and at teeth of posterior angles; anterior angle conspicuously and widely rounded off, bent ventrally, posterior angle marked by distinct laterally projecting tooth, with short concavity anterior to posterior angle; lateral channel moderately wide, somewhat flattened, irregularly to isodiametrically reticulated, continuing up to base as narrow, straight margin. Median line engraved, complete, joining anterior transverse line. Anterior transverse line distinct, complete. Disk covered with partly connected punctures, with numerous transverse wrinkles, with isodiametric rough reticulation basally and laterally, with impression of posterior setigerous puncture distinct, conspicuously distant from margin. Lateral margin from posterior angle to flange of base slanted with angle of around 45°. Basal channel on flange conspicuously widened, coreaceate, with narrow, obtuse carina.
Elytron (Fig. 18) with disk in frontal view moderately convex, in lateral view flattened in anterior half. Shape of elytra elongate, about two-fifth longer than wide, with lateral margin nearly straight at middle but slightly diverging, maximum width slightly posterior middle. Humerus rounded off. Reflexed lateral margin slightly crenulate in anterior half. Row of setigerous punctures in the lateral channel wider separated at middle. Scutellar striole distinct, impressed anteriorly; setigerous tubercle at base of first stria distinct, with tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, indistinctly punctate-striate, one to four free at base, five and six joining at humerus, one and two ending free at apex, three and four, and five and six joining at apex. Intervals somewhat flattened but still slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven and eight not carinate, seventh running to apex as obtuse convexity, convexity of eighth ending before apex. Surface of interval one to seven glossy but covered with indistinctly and evanescent reticulation, interval eight with distinct and clear isodiametric reticulation.
Hind wing fully developed.
Lower surface. Proepisternum isodiametrically reticulated, with indistinct transverse wrinkles laterally. Sternite of abdomen with isodiametric reticulation, weaker medially. Last visible sternite with the two marginal setigerous punctures widely separated, with two small but distinct lappets at margin (in both sexes).
Legs. Protibia slender, with longitudinal reticulation, with moderately deep sulcus dorsally, with three slender spines, basal tarsomere fractuate and flattened at base, carinate laterally, with acute tubercle and two setae at middle of tarsomere. Intermediate tibia with eight tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg widened.
Male genitalia (Fig. 87). Median lobe relatively short, moderately arcuate at middle, apically regularly rounded, slightly flattened. In lateral view moderately acuminated. Parameres moderately slender, with small apophysis, slightly distorted, slightly more arcuate and hyaline apically, asetose.
Female gonocoxites and epipleurite (Fig. 113). Basal gonocoxite crescent-shaped, with two long setae, apical gonocoxite elongated, slightly convex, with long seta at middle and shorter one sub-medially, with two very long setae apically. Epipleurite lappet-like, with two setae close together and one isolated towards middle.
Variation. The clypeus with central part is straight in the holotype but in some of the paratypes slightly undulate. On the elytron, the reticulation on intervals one to seven varies in distinctness from clearer to seemingly washed out.
Distribution. Known from Sabah and Sarawak on the island of Borneo
Diagnosis. A medium sized fuscous to ferruginous species with distinctly cordate pronotum and intervals of the elytron more or less distinctly reticulated. Three of the species with distinctly cordate pronotum, C. geiseri sp. nov., C. subdepressa Kult, and C. depressiforma sp. nov. show in addition a conspicuously elongated flange of the pronotum with flat basal carina. C. depressiforma sp. nov. differs distinctly as only species by the pronotum with the laterally and acutely projecting teeth of the posterior angles, the coreaceate-like surface, and the conspicuously rounded off anterior angles. In addition, in C. depressiforma interval eight of the elytron is clearly and distinctly reticulated isodiametrically and all of the other intervals are indistinctly and evanescent reticulated. Moreover, the male aedeagi of the three species are very different.
Etymology. The name refers to the pronotum with its extraordinary depressed dorsal surface. (Latin: depressus = depressed; forma = form).
description: Published as part of Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), pp. 1-70 in Zootaxa 5592 (1) on pages 31-32, DOI: 10.11646/zootaxa.5592.1.1, http://zenodo.org/record/14953674