Hyposoter naso Galsworthy & Shaw & Haraldseide 2023, sp. nov.
Creators: name: Galsworthy, Anthony
name: Shaw, Mark R.
name: Haraldseide, Håkon
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Hyposoter
Hyposoter naso

description: Hyposoter naso Galsworthy & Shaw sp. nov.
(Figs. 23–25)
Diagnosis. Superficially similar to some paler specimens of H. caedator, though distinguished from them by the dark trochanters and trochantelli on all legs. The unusual shape of the clypeus, which is unique in Hyposoter, is the most distinctive feature. From some angles this can appear to be similar to the clypeal tooth of Campoletis, but seen in profile it is clearly a nose-like bulge, not a tooth.
Description. Female. Fore wing: 5 mm.
Head: face broader than long, clypeus narrow, the central part strongly and narrowly prominent subdistally, surface of both rugosely reticulate; malar space slightly broader than width of mandibles at base, the latter strongly narrowed at about two thirds; frons and vertex strongly granulate, temples lightly so, the latter strongly narrowed behind the eyes, making the temples very short in dorsal view; antenna slender, slightly tapered in apical half. 31–32 flagellomeres.
Mesosoma: mesoscutum closely and rugosely punctate throughout; speculum smooth and shining; area above and in front of speculum with strong transverse striation; front and ventral area of mesopleuron, and metapleuron, with close strong punctation; all areas of propodeum rugose, petiolar area with strong transverse rugosity; propodeal carinae complete except for hind carina of area superomedia; basal area narrowly trapezoid; area superomedia about as broad as long.
Wings narrow; 1 cu-a very slightly postfurcal; areolet small and squarish, long-stalked; nervellus slightly curved.
Metasoma: basal part of first tergite parallel-sided, smooth and shining, sharply narrowing before postpetiole; latter a broad oval, with a linear depression in the basal half, lightly granulate; glymma a very shallow linear impression at base of postpetiole. 2nd tergite longer than broad, widening to apex; third tergite quadrate, the rest transverse; 2nd tergite with light granulation, the rest almost smooth, but dull with very fine microsculpture; sixth tergite entire, seventh with a semicircular emargination at apex; ovipositor shorter than height of metasoma.
Colour: black; tegulae yellow; fore and mid legs orange except for black trochanters and trochantelli; hind femur orange; hind tibia with a pale yellow spot dorsally at the base, rather uniform brownish dorsally, orange ventrally, with faint darkening subproximally and distally; hind tarsi brown dorsally, orange ventrally; tergite 1 black; tergite 2 black in basal half, orange distally, apart from a narrow black band at extreme apex, which extends to base of tergite 3; rest of tergite 3 orange, as are tergites 4–6; tergite 7 brown; sternites 2–3 yellow with broad pale brown bands.
Male. Very similar to female, but sides of postpetiole orange.
Holotype: ♀, Spain: Madrid, 600m, Ciempozuelos, G.E.King, ex Perigune convergata (de Villers) (Lepidoptera, Geometridae), Thymus, 16.4.2005, coc 23.4.2005, em 2.5.2005 (NMS). Paratypes: 1♀, Spain, Madrid, 600m, Ciempozuelos, G.E.King, ex Perigune narbonea (Linnaeus) (Lepidoptera, Geometridae), Teucrium, 28.11.2004, coc 3.12.04, em 31.12.04; 1♀, Ciempozuelos, ex Petrophora convergata, (de Villers) (Lepidoptera, Geometridae), Thymus, 8.4.2004, em by 20.5.2004, G.E.King ; 1♀, Spain, Madrid, 600m, Ciempozuelos, G.E.King, Perigune convergata (de Villers), Thymus, 8.4.2004, coc 27.4.04, em 16/ 20.5.04 ; 1♀, Spain, Madrid, 600m, Ciempozuelos, “ Scopula ” Glossotrophia asellaria Herrich-Schäffer (Lepidoptera, Geometridae), Antirrhinum, 13.5.05 [sic: presumably 13.3.05], pupa 3.4.05, em 1.5.05 ; 1♀, Spain: Madrid, 600m, Ciempozuelos,? Chemerina caliginearia (Rambur) (Lepidoptera, Geometridae), 4.5.08, em 21.5.08, G.E.King ; 1♁, Spain, Madrid, 600m, Ciempozuelos, G.E.King, ex Antilurga alhambrata (Staudinger) on Helianthemum hirtum, 6.3.2005 em. 29.3.2005 ; 1♁, Spain, Madrid, 600m, Ciempozuelos, G.E.King, uncertain host, coc. 12.4.08, em 18.4.2008 ; 1♁, Spain, Madrid, 600m, Ciempozuelos, G.E.King, ex Petrophora convergata (de Villers) (all in NMS).
All the above are accompanied by cocoons and host remains, except for the single female reared from the sterrhine G. asellaria which is mounted with an intact lepidopteran pupa. This raises a doubt as to the identity of the actual host in that case. The rearing data make clear that H. naso is a plurivoltine species.
Remarks. First identified by Horstmann as an undescribed species.
description: Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on pages 54-56, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256