Arpactophilus haveke Breitkreuz, Ohl & Engel, 2016, sp. nov.
Creators: name: Breitkreuz, Laura C. V.
name: Ohl, Michael
name: Engel, Michael S.
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Crabronidae
Arpactophilus
Arpactophilus haveke

description: Arpactophilus haveke sp. nov.
(Figs. 59–60)
Diagnosis. Arpactophilus haveke is the only New Caledonian species in the genus that has a stalked triangular submarginal cell II in the forewing, and particularly when taken in combination with the light reddish-brown metasoma further distinguishes itself from its congeners. Another distinctive feature is the apical margin of thee clypeus which is projecting medially and medioapically forming a point, without a medial incision (Fig. 59).
Description. FEMALE: Total length 3.8 mm; forewing length 2.3 mm.
Body black, with areas of light yellow and light reddish-brown. Light yellow: apical margin of clypeus (Fig. 59); mandible; palpi; scape in front; pronotal lobe; tegula; coxae apically; trochanters; femora partially; tibiae, except apical area on metatibia; tarsi. Light reddish-brown: posterior of scape; pedicel; flagellum; most of coxae; most of femora; apical area on metatibia; metasoma (Fig. 60). Wings hyaline; pterostigma light brown. Labrum and anterior pronotal margin not visible.
Head about as long as wide in frontal view. Apical margin of clypeus projecting medially, medioapically forming a point (not incised or with a straight margin as in some other species) (Fig. 59). Labrum not visible. Ventral mandibular tooth about ¼–1/5 of total mandibular length, not reaching opposite mandibular base. Palpal formula 4:3. Frons imbricate with sparse punctation and short setae. Frontal carina present from median ocellus to apical margin of clypeus. Scape 3.7 × as long as wide. Ocellar triangle anterior of tangent between upper posterior orbits of compound eyes, lateral ocelli anterior of tangent by approximately their diameter. Occipital carina interrupted dorsally (Fig. 6). Gena imbricate with sparse punctation and sparse setae, without tubercles, spines, or dorsoventral carina. Hypostomal midventral line faint, not angulate.
Mesosoma about 1.7 × as long as wide in dorsal view. Propodeum about 0.8 × as long as wide in dorsal view. Mesosoma imbricate except on propodeum (Fig. 60); lateral and dorsal surface of propodeum transversely carinulate; dorsal surface of propodeum also finely reticulate with transverse carinulae apically (similar to pattern in Fig. 20). No groove present posterior to mesoscuto-mesoscutellar sulcus. Hypersternal sulcus absent. Metafemur 2.6 × as long as wide. Metatibia with a light brown area apically. Forewing with two submarginal cells; submarginal cell II triangular, not reaching media but connected to it by a single crossvein (i.e., cell is stalked). Hind wing with four distal hamuli.
Metasoma polished, punctation and associated setae sparse (Fig. 60). Metasomal sternum II without bulge (as in Fig. 1). Pygidium with narrow row of silk setae.
MALE: Difference from female aside from genitalic structures: yellow marking on clypeus larger; yellow markings present on lower frons and malar space; flagellum yellow; metasoma brown. Metasomal sternum VIII neither broadened nor narrowed.
Remarks. Arpactophilus haveke can easily be distinguished from the other species with a reddish metasoma by its stalked second submarginal cell. The only species that has a similarly triangular (but not stalked) second submarginal cell is A. tiri, which has a much more slender body form (Fig. 102).
Material examined. HOLOTYPE ♀: “Nouvelle Calédonie, Pinda 30m, 18.IV.1995, Réc. Chazeau & Jourdan [MNHN].
PARATYPE 1♂: “Nouvelle Calédonie, Pinda 30m, 18.IV.1995, Réc. Chazeau & Jourdan [MNHN].
Etymology. The specific epithet is taken from the name Haveke, one of the New Caledonian native languages. It is treated as a noun in apposition.
description: Published as part of Breitkreuz, Laura C. V., Ohl, Michael & Engel, Michael S., 2016, A review of the New Caledonian Arpactophilus (Hymenoptera: Crabronidae), pp. 1-66 in Zootaxa 4063 (1) on page 27, DOI: 10.11646/zootaxa.4063.1.1, http://zenodo.org/record/270412