Tomarus maimon Erichson 1847
Creators: name: Neita-Moreno, Jhon César
name: Alexander, Recursos Biológicos
name: Agustín, San
name: de, Villa
name: Leyva
name: Boyacá
name: Ratcliffe, Brett C.
name: Collections, Systematics Research
name: of, University
name: Museum, Nebraska State
name: Hall, Nebraska
name: of, University
name: Nebraska
name: Lincoln
name: Ne
name: Usa
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Dynastidae
Tomarus
Tomarus maimon

description: Tomarus maimon Erichson, 1847   
(Fig. 3h, 4d, 5d, 6g –h, 7d, 8d, 9b, 10g –h, 11d, 12d, 13d, 14d, 15d, 16c)  
 Tomarus maimon Erichson 1847: 96 (original combination).   
 Heteronychus fossator Burmeister 1847:101 (synonym).   
Redescription. Male. Length 22.0–26.0 mm; width 11.0–12.0 mm. Color dark reddish brown to black. Head: Frons and clypeus coarsely rugose. Frontoclypeal region with 2 distinctly transverse tubercles; tubercles broadly separated, usually about 4 tubercle diameters apart (Fig. 4d). Clypeus strongly rugopunctate to transversely rugose, tapering to broad apex with 2 nearly contiguous, reflexed teeth. Mandibles with 2 teeth, apical tooth subtruncate or rounded, subapical notch relatively narrow (Fig. 6g –h); labium with lateral edge densely setose, base setose, paraglossa developed, apex slightly convex at center (Fig. 7d); maxilla with galea developed and with 6 teeth (Fig. 5d). Pronotum: Surface with disc moderately punctate, sides and anterior angles densely punctate to rugopunctate; punctures moderate in size on disc, becoming large laterally; fovea rugose within. Anterior margin with small, median tubercle and with small, narrow fovea (narrower than distance between eyes) (Fig. 8d). Elytra: Surface densely punctate; punctures mostly moderate in size with sparser, minute punctures between larger punctures; larger punctures usually ocellate, and with 3 pairs of double rows. Pygidium: Surface densely punctate, punctures becoming denser in basal and lateral angles; punctures small to moderate in size, weakly ocellate. In lateral view, male with convex surface, females with weakly convex surface. Legs: Protibia tridentate, teeth subequally spaced from each other (Fig. 9b). Protarsus in male simple, not enlarged. Mesotibia on transverse carinae with long, slender spinules (Fig. 10g –h). Metatibia with 1 basal, transverse carina interrupted at middle (carina with spine-like setae) and 1 medial, transverse carina; apex truncate with about 21 spinules (Fig. 10h). Venter: Prosternal process long, thick, apex longitudinally oval. Mesosternal surface glabrous, slightly concave at middle. Metasternal surface with sparse, ocellate punctures (Fig. 2f). Male Genitalia: Parameres (Fig. 11d, 12d, 13d) subtriangular, but apical half of shaft parallel, apices blunt. Internal sac with 3 copulatory lamellae (Fig. 15d).  
 Female. Length 23.1–28.2 mm; width: 10.8–13.6 mm. Similar to male, but pronotal surface with punctures slightly less dense than in males. Gonocoxite larger than gonocoxal sternite, with a semicircular furrow (Fig. 14d).
description: Published as part of Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, pp. 1-36 in Insecta Mundi 2017 (547) on pages 7-8, DOI: 10.5281/zenodo.5172580