Acrographinotus Holmgren 1917
Creators: name: Acosta, Luis E.
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Opiliones
Gonyleptidae
Acrographinotus

description: Identification key for males of Acrographinotus *
* Acrographinotus opacus not included (male unknown)
1. DS set with large, pustular tubercles that fill areas I–IV, in marked contrast with the whitish background in those areas. Coxa IV with a large, sigmoid apophysis, diagonally oriented; trochanter IV articulates fairly straight. FeIV and TiIV heavily armed, in the former the armature includes a large retrolateral apophysis on the middle (cf. Roewer 1929: fig. 28)........ A. ornatus
- DS of uniform color, areas I–IV completely smooth or with scattered granules to small round tubercles. Coxa IV with a short prolateral apophysis, often truncate and quite transverse, if diagonal it is not sigmoid; trochanter IV articulates obliquely outwards (less accentuated in A. mitmaj). FeIV without a large retrolateral apophysis on the middle; armature of TiIV moderate..... 2
2. VAP unarmed. Leg IV covered with acute tubercles or apophyses, what gives this appendage a spiny look (cf. Roewer 1929: fig. 27; Acosta 2002a).................................................................................. 3
- VAP armed (a pair of horn-like apophyses and/or a shelf-like projection; Figs. 3B, D; 5B). Leg IV never spiny, it is covered with rows of rounded tubercles or truncate apophyses (see e.g., Figs. 3A–C, 5A–C)..................................... 5
3. Prolateral apophysis of coxa IV diagonal, it emerges close to the articulation with trochanter, so it hides its prolateral corner if seen from above. Femur IV covered with uniformly-sized acute grains and a single retro-subapical apophysis—no subapical hiatus exists. FT-III with a modest apophysis; armature on trochanter IV little developed. Penis: VPS with a membranous ‘crest’ (cf. Acosta 2002a: figs. 1–12)..................................................................... A. mitmaj
- Prolateral apophysis of coxa IV transverse, truncate, or slightly diagonal, very short, it emerges separated from the articulation with trochanter, leaving a gap and the prolateral corner uncovered (cf. Roewer 1929: fig. 27). FeIV with large distal apophyses that leave a subapical hiatus (cf. Roewer 1957: Pl. 3, fig. 10). Single apophysis of FT-III robust; strong armature on trochanter IV. Penis: VPS without ‘crest’............................................................................ 4
4. Apophysis on the ocular mound high, it is always a single one. FeIV with moderate subapical hiatus, the large apical apophysis is conic, similar to the adjacent ones; strong proventral subapical apophysis, with foliaceous and truncate look (cf. Roewer 1929: fig. 27).............................................................................. A. curvispina
- Apophysis on the ocular mound low or absent, its place may be taken by a notch that gives the ocular mound a paired appearance. FeIV with wide subapical hiatus, apical apophysis hooked and larger than the rest; no proventral subapical apophysis (cf. Roewer 1957: Pl. 3, figs. 9–10)..................................................................... A. ortizi
5. VAP armature: a pair of independent, horn-like acute apophyses (cf. Acosta 2001: figs. 2, 3). DS with scattered granules, they may be almost small round tubercles toward the median line................................................... 6
- VAP armature: a platform-like projection (‘shelf’) extends beneath the anus, normally flanked by the apophyses (cf. Acosta 2001: fig. 25; this paper: Fig. 3B), these are rarely lacking (Fig. 5B). DS sparsely covered with tiny granules to almost smooth... 7
6. Apophyses of VAP strong and diverging, pointing sideways. Apophysis on FT-III very large, it expands the tergite itself into a conical shape and has a row of short apophyses each side. FeIV arched in lateral view. Retroapical apophysis of trochanter IV rudimentary. Ocular mound acute (cf. Soares & Bauab 1972: figs. 10–15).............................. A. ceratopygus
- Apophyses of VAP subparallel, directed backwards. Apophysis on FT-III large but not expanding the tergite itself, it has a row of round granules (not apophyses) each side. FeIV straight in lateral view. Retroapical apophysis of trochanter IV large. Ocular mound blunt (cf. Acosta 2001: figs. 1–10)....................................................... A. erectispina
7. Robust. FeIV thick, with a retrodorsal row of large truncate tubercles and a dorsal row of blunt apophyses; TiIV thick and heavily armed with retroventral row of apophyses. Trochanter IV with bilobated (anvil-shaped) prodorsal apophysis, and an acute, large retroapical one (cf. Acosta 2001: figs. 23–32)............................................. A. niawpaq
- Slender. FeIV and TiIV slim; FeIV with similar dorsal and retrodorsal rows, but with tubercles or apophyses much smaller. Prodorsal and retroapical apophyses of trochanter IV less developed............................................. 8
8. Medium-sized (DS length 5.9–7.2 mm; mean 6.4; Fig. 2), long-legged and with fairly robust habitus. FeIV substraight and long (FeIV/DS RatiO in α males: 0.94–1.15; Fig. 2), with a defined subapical naRROwing (Figs. 3A–C). ApOphYsis On FT-III sigmOid in lateRal view (Fig. 3D), RaRelY hORizOntal OR cuRved (Figs. 3H, J). ApOphYses On VAP laRge, in α-males theY fuse tO bOth sides of the shelf-like process, most frequently exceeding it in size (Figs. 3B, D). Penis: curvature of beak-like process of VPS marked (Fig. 4C)............................................................................. A. tariquiae sp. nov.
- Small-sized (DS length 5.7–6.2 mm; Fig. 2), with shorter legs and habitus comparatively weaker. FeIV slightly arched (Figs. 5A–B), shorter than DS (FeIV/DS ratio: 0.84–0.89; Fig. 2), with no discernible subapical narrowing. Apophysis on FT-III curved or horizontal in lateral view, not clearly sigmoid (Fig. 5D). VAP with a small shelf-like projection, apophyses not recognizable except for a pair of small subapical tubercles (Figs. 5B, D). Penis: curvature of beak-like process of VPS mild (Fig. 6C)................................................................................. A. calilegua sp. nov.
description: Published as part of Acosta, Luis E., 2024, The two southernmost species of the Andean genus Acrographinotus Holmgren (Arachnida, Opiliones, Gonyleptidae) described from Bolivia and Argentina, pp. 547-562 in Zootaxa 5481 (5) on page 551, DOI: 10.11646/zootaxa.5481.5.4, http://zenodo.org/record/12783058