Desmoxytes pilosa Attems 1937
Creators: name: Likhitrakarn, Natdanai
name: Golovatch, Sergei I.
name: Panha, Somsak
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Polydesmida
Paradoxosomatidae
Desmoxytes
Desmoxytes pilosa

description: Desmoxytes pilosa (Attems, 1937)
Figs 9–10.
Centrodesmus pilosus Attems, 1937: 124 (D).
Centrodesmus pilosus — Attems, 1938: 238 (D).
Pratinus pilosus — Jeekel, 1964: 62 (D, M); 1968: 61 (M).
Hylomus pilosus — Jeekel, 1980: 657 (M, R); Golovatch, 1983: 181 (M).
Desmoxytes pilosa — Golovatch & Enghoff, 1994: 48, 59 (D, M); Enghoff et al., 2004: 38 (R); Nguyen et al., 2006: 257 (D); Nguyen & Sierwald, 2013: 1242 (R).
Holotype male (NHMW-7989), Vietnam, Ninhthuan Province, Phan Rang City, 1000 m a.s.l., 31.07.1930, leg. C. Dawydoff.
Redescription. Length 19 mm (male), width of midbody pro- and metazonae 2.2 and 4.7 mm (male), respectively (versus width of midbody pro- and metazonae 1.65 and 4.3 mm (male), respectively, as given in the original description (Attems, 1937, 1938)).
Coloration of alcohol material upon long-term preservation red-brown, venter and legs pale yellowish (Fig. 9) (versus dark chestnut brown, as given in the original descriptions (Attems, 1937, 1938)).
Clypeolabral region and vertex densely setose, epicranial suture distinct. Antennae very long, reaching segment 7 when stretched dorsally. In width, head <collum <segment 2 <3 <4 <5 <6–15; thereafter body gently and gradually tapering. Collum with three transverse rows of prominent setigerous spines: 3+3 anterior, 1+1 intermediate, and 3+3 posterior; paraterga antler-shaped, directed dorsolaterad, with three strong spines anteriorly, median one being largest (Fig. 9 A & B). Tegument invariably dull, prozonae coarsely shagreened, metazonae coarsely granulate, surface below paraterga coarsely microgranulate, sterna delicately microgranulate (Fig. 9 A–G). Metaterga with two transverse rows of spines, 3+3 small anterior and 3+3 thorn-shaped posterior spines, lateral spines of posterior rows being much longer than others, metaterga 2–5 each with 2+2 anterior and 3+3 posterior spines; following segments with 2+2 anterior and 4+4 posterior spines, except for metaterga 6 and 11 with 2+2 anterior and 3+4 posterior spines (Fig. 9 A, B, C, E & F). Tergal setae mostly abraded. Axial line invisible. Paraterga antler-shaped, upturned, directed dorsolaterad, tip bent posteriad; segments 2–9 each with a prominent anterior spine at about 3/4–2/3 extent of paraterga and another evident spine near tip; following segments with a prominent anterior spine near base and two evident spines at 2/3 extent of paraterga (Fig. 9 A–G). Ozopores (Op) lateral, located distal to notch, but lower than base of large spine (Fig. 9 D). Transverse sulcus traceable on segments 2–18, very shallow, line-shaped, rather wide, not reaching bases of paraterga. Stricture between pro- and metazonae narrow, shallow, ribbed at bottom down to base of paraterga. Pleurosternal carinae complete crests only in segment 2, roughly granulate crests with a caudal tooth in segments 3 and 4, remaining only a front tooth in segments 5–8, absent thereafter. Epiproct unusual (Fig. 9 E–G), flattened dorsoventrally, with 4+4 setigerous spines near a subtruncate tip. Hypoproct subtriangular, evident setiferous knobs at a slightly convex caudal edge rather large and well-separated.
Sterna sparsely setose, without modifications; cross-impressions shallow; two evident, rounded, fully separated, but nearly contiguous, setose cones between male coxae 4, carrying a rounded tubercle at each corner (Fig. 9 H & I). Legs very long and slender, midbody ones ca 1.9–2.0 (male) as long as body height. Prefemora without modifications. Male tarsal brushes absent.
Gonopods (Fig. 10) suberect. Coxite rather large. Prefemur densely setose, as long as femorite. Femorite stout, slightly curved, strongly enlarged distad, seminal groove running entirely on mesal face. Solenophore (= lamina medialis) well-developed, with two evident processes; Solenomere rather short, flagelliform, well separated at base from solenophore.
description: Published as part of Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak, 2015, Two new species of dragon millipedes, genus Desmoxytes Chamnerlin, 1923, from Laos (Diplopoda: Polydesmida: Paradoxosomatidae), with redescriptions of all four species of Attems from Vietnam, pp. 483-504 in Zootaxa 3931 (4) on pages 495-497, DOI: 10.11646/zootaxa.3931.4.2, http://zenodo.org/record/234300