Anemesia incana Zonstein 2001
Creators: name: Zonstein, Sergei
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Cyrtaucheniidae
Anemesia
Anemesia incana

description: Anemesia incana Zonstein, 2001
Figs 12, 24, 32–33, 35, 43–44, 59, 73, 84–85, 101, 113, 129, 138–142, 147–150, 156–157, 171–172, 184, 215–216, 251–254, 285–286, 302–305, 318–321, 338–344, 363, 373
Anemesia incana Zonstein, 2001: 11, figs 1–8 (Ƌ ♀)
Anemesia incana – Zonstein 2007: 211, fig. 9 (♀). — Marusik et al. 2014: 3, fig. 9 (Ƌ).
Diagnosis
Anemesia incana differs from the closely related and similar-looking A. infumata sp. nov. and A. infuscata sp. nov. in a considerably wider clypeus and noticeably smaller eyes both in males and females (Figs 59, 73; cf. Figs 60–62, 74–77). Additionally, males of A. incana are well distinguishable due to their wider palpal organ (Figs 215–216; cf. Figs 217–223), and the conspecific females due to their oblate receptacles (Figs 251–254; cf. Figs 255–265).
Material examined
Holotype
TAJIKISTAN: Ƌ, foothills of Aruktau Mts, 1 km NE of Ganjina Village, 750 m, 37°58′ N, 68°32′ E, 13 Apr. 1987, S. Zonstein leg. (AMNH).
Paratypes (15 ƋƋ, 15 ♀♀)
TAJIKISTAN: 12 ƋƋ, 9 ♀♀, Aruktau Mts, surroundings of Ganjina, 700–1000 m, 9–13 Apr. 1987, S. Zonstein leg. (SMNH: 9 ƋƋ, 6 ♀♀; MNHL: 1 Ƌ; ZISP 1 ♀; ZMMU: 2 ƋƋ, 2 ♀♀); 1 Ƌ, 4 ♀♀, same collection data as for preceding but 11 Apr. 1988 (SMNH); 1 Ƌ, 1 ♀, same collecting data as for preceding but 22 Apr. 1992 (SMNH).
Additional material (3 ƋƋ, 7 ♀♀)
TAJIKISTAN: 1 ♀, Aruktau Mts, surroundings of Ganjina, 800–950 m, 19 Apr. 1989, S. Zonstein leg. (SMNH); 1 Ƌ, same collection data as for preceding but 18 Apr. 1992 (SMNH); 1 Ƌ, 2 ♀♀, same collection data as for preceding but 22 Apr. 2015 (SMNH); 3 ♀♀, same collection data as for preceding but 27 Apr. 2002, S. Zonstein and A. Feodorov leg. (SMNH); 1 Ƌ, Panj Karatau Mts, vicinity of Mt Astana, 1600 m, 37°23′ N, 69°15′ E, 23 Apr. 1991, S. Zonstein leg. (SMNH); 1 ♀, Vahsh Karatau Mts, summit area 4 km NNE of Mt Hojamaston, 1670 m, 38°00.9′ N, 68°58.7′ E, 24 Apr. 2015, S. Zonstein leg. (SMNH).
Redescription
Male (holotype)
HABITUS. See Fig. 12.
MEASUREMENTS. TBL 14.50, CL 6.87, CW 5.63, LL 0.63, LW 1.05, SL 4.25, SW 3.30.
COLOUR. Carapace, chelicerae, sternum, labium, most part of palps and legs intensely dark reddish brown; maxillae, coxae, metatarsi, tarsi and sternal sigilla dark brownish orange; eye tubercle blackened; abdomen dorsally dark grey with brown pattern consisting of wide median longitudinal spot and few paired transverse and slightly inclined short stripes, ventral part of abdomen dark yellowish grey, spinnerets pale yellow.
PROSOMA. Clypeus wide. Eye tubercle as in Fig 59. Eye diameters and interdistances: AME 0.19(0.27), ALE 0.28, PLE 0.15, PME 0.14, AME–AME 0.24(0.16), ALE–AME 0.22(0.18), ALE–PLE 0.22, PLE– PME 0.08, PME–PME 0.57. Weak cheliceral rastellum consists of about 20 spikes grouped in front of cheliceral fang and on low mound. Each cheliceral furrow with 7 promarginal teeth and 3–4 smaller teeth confined to basal retromargin. Sternum, labium and maxillae as shown in Fig. 101. Sternal sigilla oval; posterior pair relatively large, long oval, and located distantly from sternal margin. Maxillae with 4–5 cuspules each.
LEGS. Tibia and metatarsus I as shown in Fig. 129. Scopula distal on metatarsi I and II, very narrowly divided on tarsi I and II, widely divided on tarsus III, mixed, rudimentary, and widely divided on tarsus IV. Trichobothria: 2 rows of 9–10 each on tibiae, 14–17 on metatarsi, 14–17 on tarsi, 8 on cymbium. PTC I–IV with 6–7 teeth on each margin.
SPINATION. Palp: femur d3, p2; patella p2; tibia p3, r2, v3; tarsus d5–6. Leg I: femur d5, pd2; tibia p3, v6–7+m; metatarsus p2, v7. Leg II: femur d5, pd1; patella p3–4; tibia p3, v8; metatarsus p3, v6. Leg III: femur d4–5, pd3, r2; patella p4–5; tibia d1, p3, r3, v10–11; metatarsus d1, p3, r3, v10; tarsus v9–10. Leg IV: femur d5, p2, r3; tibia r3, v9–11; metatarsus p4, r2, v7–8; tarsus v7–10. Patellae I and IV, and tarsi I–II aspinose.
PALP. Tibia, cymbium and palpal organ as shown in Figs 184, 215–216. Palpal tibia long, cylindrical and spinose (Fig. 184). Palpal organ with wide asymmetrical bulb and tapering and slightly curved embolus (Figs 215–216).
SPINNERETS. See Fig. 285. PMS: length 0.63, diameter 0.30. PLS: maximal diameter 0.53; length of basal, medial and apical segments 1.45, 1.07, 0.95; total length 3.47; apical segment shortly digitiform.
LEG MEASUREMENTS. Ƌ(♀)
Female (paratype)
HABITUS. See Fig. 24.
MEASUREMENTS. TBL 17.50, CL 8.15, CW 7.07, LL 0.85, LW 1.63, SL 4.75, SW 3.97.
COLOUR. Similar to that of male, except even darker chelicerae.
PROSOMA. Clypeus and eye tubercle as in Fig 73. Eye diameters and interdistances: AME 0.23(0.30), ALE 0.28, PLE 0.20, PME 0.18, AME–AME 0.20(0.13), ALE–AME 0.20(0.17), ALE–PLE 0.22, PLE–PME 0.05, PME–PME 0.53. Weak cheliceral rastellum as in male. Each cheliceral furrow with 7–8 promarginal teeth and 5–6 smaller retromarginal teeth. Sternum, labium and maxillae as shown in Fig. 113. Sternal sigilla as in male. Maxillae with 7–10 cuspules each.
SPINATION. Palp: tibia p5, v18–20; tarsus v1. Leg I: tibia v8–10; metatarsus p1, v10. Leg II: tibia p3, v8–9; metatarsus p3, v8–10. Leg III: patella r2; tibia p2–3, r2, v9–10; metatarsus p3–4, r3, v12–14; tarsus v6–8. Leg IV: tibia r2, v10–12; metatarsus r2, v11–12; tarsus ventrally ca 20. All femora and most part of patellae with long and relatively thin bristles instead of spines; tarsi I and II unarmed.
LEGS. Scopula distal on metatarsi I and II, entire on palpal tarsus, very narrowly divided on tarsus I, narrowly divided on tarsus II, elsewhere absent. Trichobothria: 8–10 in each row on tibiae, 16–20 on metatarsi, 15–18 on tarsi. Palpal claw with 4 promarginal teeth. PTC I–II with 6–7 teeth on each margin. PTC III with 6 teeth on outer margin, 3 teeth on inner margin; PTC IV with 3 teeth and one tooth, respectively.
SPERMATHECAE. Entire with short stout stalks and considerably oblate receptacles (Fig. 251).
SPINNERETS. See Fig. 286. PMS: length 0.67, diameter 0.50. PLS: maximal diameter 0.93; length of basal, medial and apical segments 1.55, 1.13, 0.95; total length 3.63; apical segment shortly digitiform.
Variation
The length of the carapace varies from 6.30 to 7.92 in males and from 5.96 to 8.43 in females. The colour of the carapace and legs varies from very dark reddish brown to deep blackish-brown. Number of maxillary cuspules varies from 2 to 11.
Habitat
Foothills and low mountain ridges at an altitude of 600–1700 m, covered with ephemerous grasslands and subshrubs (mostly species of Artemisia), often with tall shrubs or low open forest (Pistacia vera L., different species of Prunus, Crataegus and Acer). The typical biotopes preferred by these spiders are shown in Figs 318–321.
Distribution
South-western Tajikistan (Fig. 273).
Notes
Some illustrations of Anemesia incana used in this study are partially based on the additional (non-type) material. Live spiders are presented in Figs 32–33, 35, 43–44. The dentition of the cheliceral furrow in female and male paratypes is shown in Figs 84 and 85, respectively. The separated female palps and legs I–IV are shown in Figs 138–142. The SEM micrographs include those of the palpal organ (147, 149), the tip of the embolus (Figs 148, 150), the tarsal claws (Figs 156–157), the trichobothrial base and the tarsal organ of leg I (Figs 171 and 172, respectively), and spigots of PLE (Fig. 302) and PME (Figs 303–305).
description: Published as part of Zonstein, Sergei, 2018, A revision of the spider genus Anemesia (Araneae, Cyrtaucheniidae), pp. 1-100 in European Journal of Taxonomy 485 on pages 39-42, DOI: 10.5852/ejt.2018.485, http://zenodo.org/record/3829992