Colobostema infumatum Jean-Paul Haenni 2013, comb. n.
Creators: name: Jean-Paul Haenni
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Scatopsidae
Colobostema
Colobostema infumatum

description: Colobostema infumatum (Haliday, 1833) comb. n.
(Figs 6 A-G)
Sccitopse infurnata Haliday, 1833: Entomol. Mag. 1: 157.
Scatopse nigripennis Meigen: Walker, 1856: Insecta britannica III: 143; Edwards 1925: Ann. appl. Biol. 12: 274: Duda 1928: Scatopsidae, Fliegen pal. Reg. 2(1)5: 31.
Colobostema nigripenne (Meigen): Cook 1974: J. nat. Hist. 8:64; Freeman 1985: Hdbk Ident. Br. Ins. 9(7):27.
Colobostema infumatum (Haliday, 1833) Haenni & Greve 1995: 78; Laurence & James 1996: Entomol. Mon. Mag. 132: 73, Figs 2-3; Krivosheina 2000: Entomol. Obozr. 79(2): 491, Figs 3 (7-12); Krivosheina 2001: Int..1. Dipteral. Res. 12(2): 75, Figs 6-8, 10.
tree: Reichertella infurnata (Haliday) sensu Enderlein 1912: Zool. Anz. 40: 272,
Type locality. Northern IRELAND, Down: Holywood.
Material examined. Lectotype ♂, labelled: «fuscinervis» [handwritten, green paper] / « Ireland » [green paper] / «Type» / «Haliday 20.2.82» / « Colobostema nigripenne det. P.J. Chandler».
This specimen is designated here as lectotype and l have labelled it accordingly: « Scatopse infurnata Hal., lectotype, Haenni des. 1990» / « Colobostema infumatum (Hal.) det. Haenni 1990». The lectotype in fair condition is glued on a pinned card and is preserved in the NMID, Dublin.
Other material. CZECH REPUBLIC. Moravia sept.: Beskydy-Mufnkovy, 49°31 ’ N, 18°39’ E, 950 m, 13.VI.1987, 1 ♂, M. Bartâk; Sumava-Jezerny slat, 49°02’ N. 13°34’ E, 980 m, 2-18.VI.1995, 4 ♂♂, M. Bartâk, CMB; same, 20-21.VIE 1995, 1 ♂, M. Bartâk; MHNN; Sumava, Zhufské slât, 1130-1140 m, dates ranging from 18.V to 21.' VIII. 1999, 8 ♂♂, M. Bartâk & Kubik, CMB and MHNN; Sumava Mts, Nova Hurka, 850-870 m, dates ranging from 18.V to 22.VII.1999, 6 ♂♂, M. Bartâk & Kubik, CMB and MHNN; Sumava, Horskâ Kvilda, 1 100 m, 20.V-17.VI. 1999, 1 ♂, M. Bartâk & Kubik, CMB; Sumava Mts, Rokytecka slât, 1100 m, dates ranging from 18.V to 22.VII.1999, 3 ♂♂, M. Bartâk & Kubik, CMB and MHNN.— " GREAT BRITAIN. Wales: Cader Idriss, 1-3.VIII.1993, 4 ♂ ♂, B.R. Laurence, BMNH (Laurence & James, 1996). The following series of 24 ♂ ♂ and 5 ♀ ♀ are all labelled Colobostema cornuta Freeman, and include the « ♂ holotype » and 3 ♂ ♂ and 1 ♀ « paratypes » of this unpublished species (see under ‘Taxonomic remarks’ below); Dyfed: Llyn Eiddwen, W.T.
SN607674, 30. VII 1987, 2 ♂♂, « holotype » and « paratype »; Cors Caron, SN 685623, 27, VII.1987, 1 ♂, « paratype »; Puncheston Common, SN 0 0 5302, 20.VII.1987, 1 ♀, « paratype »; Cors Grairanos, 28.VI1.1988, 1 ♂; Cwm Ystwyth SN856757, 7.X.1987, 1 ♂ « paratype »; Denbigh: Blaen y Wergloedd, SH 913633­ 4, 23.VI. 1988, 1 ♂, 1 ♀; same, 4.VIII.1988, 2 ♂♂, 1 ♀; Gors Naen Muyrd, SH 981587, 4.VIII.1988, 3 ♂♂, 1 ♀; Montgomery: Llanbrynmair, SH 940049, 13 ♂♂, 1 ♀; Gwent: Cleddon Bog, SO 509041, 21.VII. 1988, 1 ♂; all labelled W.T., in the BMNH. — NORWAY. The author has studied 14 specimens from provinces AK, HOT, RY, NTI and F0 (details in Haenni & Grevel995, 2000).— SWITZERLAND. BE: Beatenberg, 2000, 2 ♂♂, P.Duel!i, CGB;GL: Klöntal, 850 m, I 1-14.IX.1974, 1 ♂, G. Bächli, MFINN; Schwändital, 2000, 3 ♂♂, P. Duelli, CGB / MHNN; Le Cachot, 1050 m, 13 ♂♂, dates ranging from 1.VI to 4.VIII.1973, W. Matthey, MHNN; same, 1- 11.VIII. 1983, 1 ♂, Y.Basset, MHNN; same, 22.VII.1982, 1 ♂, IP Haenni, MHNN; Le Lode, Pouillerel (Le Saignolet), 1 1.VIII.1987, 1 ♂, JP.Haenni, MHNN; Vers Maix Rochat, 5 ♂♂, 1 ♀, dates ranging from 6.V11 to 17.VIII, 1992, C. Vaucher-von Ballmoos; same, 5-12.VIT 1993, T ♂; same but 13E, 25.V-1.VI.1992, 1 ♂; Le Cachot, dates ranging from 20.VII to I 7.VIII.1992, 1 ♂, 2 ♀♀, C. Vaucher-von Ballmoos; same, dates ranging from 31.V-7.VI to 14-21.VI.1993, 3 ♂♂, 1 ♀; same, 27.VI1-3.VIM. 1992, 1 ♀, C. Vaucher-von Ballmoos, all CCV / MHNN; SZ: Alptal, 2000, 1 ♂, P. Duelli, CGB.
Diagnosis. This species has large individuals, with typical large brownish wings, and belongs to the nigripenne- group, being recognized especially in males by the shape of sternite 7 (Fig. 6 B), which has strongly developed postero-lateral projections, with prominent spiracles, and medially largely truncate posterior margin; the pointed gonocoxites (Fig. 6 C) and the strongly asymmetrical posterior projection of the epandrium (Fig. 6 D) are also characteristic of this species. The female has unique transversally wrinkled valvifers on sternite 8 (Fig. 6 G).
Description. Male. 1.8 mm long; dull black in general colour, with pilosity lighter; a pair of light spots on notum posteriorly to wing bases; wings blackishbrown, with lighter zones between anterior stem vein on M, and apex of R4+5, at the base of wing between basal part of M and CuA2 and along vena spuria; anterior veins blackish brown, well marked, posterior veins brownish, base of wing more yellowish; halters dark, with contrasting fulvous stem; legs concolourous with body, except basal fourth or third of tibiae and tarsi lighter.
Head. Eyes separated on frons above antennae by width of anterior ocellus; antennae shorter than head and thorax together, widening apically, flagellomeres well separated, easy to count, wider than long from 4lh flagellomere on, last flagellomere about twice preceding one.
Thorax quadrate, slightly wider than long; wing (Fig. 6 E), 2.5 mm; no visible anteriorly directed stem of vein on M,, which has a strongly marked bend at basal iourth; M1+2 about as long as medial fork.
Abdomen. Pregenital segment with a complete basal sclerotized ring; tergite 7 (Fig. 6 A) with slightly asymmetrical digitiform posterior projection; sternite 7 (Fig. 6 B) with strongly developed lateral projections encompassing tergite, bearing prominent spiracles; genital capsule (Fig. 6 C) with pointed gonocoxites, elongate parameres, posterior process of epandrium strongly asymmetrical (Fig. 6 D); aedeagus long, often strongly projecting.
Female. As male in general colour and appearance; tergite 7 with posterior margin deeply notched medially (Fig. 6 F); stemite 8 bearing a pair of transversally wrinkled elongate valvifers (Fig. 6 G).
Taxonomic remarks. Former authors have all accepted the synonymy of C. infurnata with C. nigripenne, first established by Walker in 1856 (sec. Cook 1974). Although it is not possible to ensure with certainty the identity of the specimen in the Haliday collection considered here as the primary type (see O’Connor & Nash 1982), there are good reasons for its designation as lectotype of Scatopse infurnata Haliday. This specimen fits well with the very short original description and is preserved as such in Haliday’s collection in the National Museum of Ireland in Dublin. It belongs not to C. nigripenne (Meigen), but to a distinct species with very typical segment 7 and genitalia.
The specimens of C. infurnata from Wales have been studied by Paul Freeman in 1989, who correctly recognized that they were not conspecific with C. nigripenne and prepared their description under the name Colobostema cornuta n.sp. (Freeman in litt.). However, he has retired his paper before publication in order to avoid the creation of a supplementary synonym (Freeman in litt.) and so the latter name will remain only a collection name.
Distribution. Known presently from the British Isles (Ireland, Wales) (Laurence & James 1996), Norway (Haenni & Greve 1995, 2000), Switzerland (Vaucher-von Ballmoos 1997, Haenni 1998), Finland (Krivosheina 2001), Northern Russia (Krivosheina 2000, 2001), Czech Republic (Haenni & Bartâk 2006). This species is probably present in the whole of Central Europe and widespread in Northern Europe.
Ecology. Estival species with a flight period extending from beginning of June to end of August. It appears to be restricted to peat-bogs in Western and Central Europe (Wales, Switzerland and Czech Republic), where these biotopes are relic.
description: Published as part of Jean-Paul Haenni, 2013, A revision of the West Palaearctic species of Colobostema Enderlein, 1926 (Diptera, Scatopsidae). Part I. European subregion, pp. 199-242 in Mitteilungen Der Schweizerischen Entomologischen Gesellschaft 86 on pages 217-220, DOI: 10.5169/seals-403072, http://zenodo.org/record/270241