Anemesia pallida Zonstein 2018, sp. nov.
Creators: name: Zonstein, Sergei
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Cyrtaucheniidae
Anemesia
Anemesia pallida

description: Anemesia pallida sp. nov.
urn:lsid:zoobank.org:act: 8CF0BF65-A760-49B5-9B6F-1DB683314DD4
Figs 10, 23, 57, 72, 99, 112, 127, 182, 211–212, 250, 282–283, 313, 318–321, 337, 372
Diagnosis
Anemesia pallida sp. nov. is a minute, pale-coloured species, with a dark dorsal abdominal pattern reduced to a longitudinal median spot. The configuration of the male palpal organ in this new taxon resembles that in males of A. castanea sp. nov. and A. karatauvi, but both the male palpal tibia and the embolus in A. pallida sp. nov. are considerably shorter (Figs 211–212, cf. Figs 206–210). Anemesia pallida sp. nov. differs from A. parvula sp. nov., another member of the same species group known only from males, in the shape of the bulb (which is pegtop-shaped, with a strongly bulging anterior lobe vs comma-shaped bulb with a weakly convex anterior lobe in A. parvula sp. nov.; Figs 211–212, cf. Figs 213–214). Females of A. pallida sp. nov. differ from all other known female congeners by their unique diverging and divided receptacles (Fig. 250).
Etymology
The specific name is derived from the Latin adjective ‘ pallidus ’ meaning ‘pale’, and reflects the general light colouration of the specimens; the gender is feminine.
Material examined
Holotype
TAJIKISTAN: Ƌ, foothills of Aruktau Mts, 1 km NE of Ganjina Village, 750 m, 37°58′ N, 68°32′ E, 14 Apr. 1986, S. Zonstein leg. (SMNH).
Paratypes (11 ƋƋ, 2 ♀♀)
TAJIKISTAN: 4 ƋƋ, 1 ♀, same collection data as for preceding but 13–19 Apr. 1986 (SMNH); 1 Ƌ, same collection data as for preceding but 9 Apr. 1988 (SMNH); 3 ƋƋ, same collection data as for preceding but 15 Apr. 1991 (SMNH, ZMMU); 1 Ƌ, southernmost tip of Aruktau Mts, Garavuti, 3 Apr. 1974, A. Kononenko leg. (ZMMU); 1 Ƌ, Rangontau Mts, Yavan Pass, 1200 m, 38°24′ N, 68°58′ E, 29 Apr. 1986, S. Zonstein leg. (SMNH); 1 ♀, Vahsh River valley, Tigrovaya Balka (Tiger Gorge) Nature Reserve, Palvan-Tugai, 350 m, 37°08.5′ N, 68°17.5′ E, 16 Apr. 1989, S. Zonstein leg. (SMNH); 1 Ƌ, Vahsh Karatau Mts, western slope of Mt Hojamaston, 1040 m, 38°01.2′ N, 68°57.1′ E, 24 Apr. 2015, S. Zonstein leg. (SMNH).
Description
Male (holotype)
HABITUS. See Fig. 10.
MEASUREMENTS. TBL 10.80, CL 4.75, CW 4.03, LL 0.40, LW 0.75, SL 2.15, SW 2.03.
COLOUR. Carapace pale yellowish orange with brownish clypeus and blackish-brown spots surrounding AMEs and lateral eyes; chelicerae, palps and legs dorsally yellowish orange with femora and patellae slightly darker; sternum, labium, maxillae, palps and legs ventrally pale brownish yellow; abdomen dorsally greyish-yellow with simple brown pattern consisting of relatively narrow median lanceolate spot only, ventral part of abdomen pale yellowish grey, spinnerets pale brownish yellow.
PROSOMA. Clypeus and eye tubercle as in Fig. 57. Eye diameters and interdistances: AME 0.14(0.20), ALE 0.27, PLE 0.12, PME 0.07, AME–AME 0.16(0.10), ALE–AME 0.09(0.06), ALE–PLE 0.14, PLE– PME 0.03, PME–PME 0.44. Cheliceral rastellum consists of 20–25 spikes located in front of fang base and on low mound. Each cheliceral furrow with 7 promarginal teeth and 3–4 smaller medial teeth. Sternum, labium and maxillae as in Fig. 99. Sternal sigilla very small but well defined. Maxillae with 22–27 cuspules each.
LEGS. Tibia and metatarsus I as in Fig. 127. Scopula entire and distal on metatarsus I, narrowly divided on tarsi I and II, elsewhere absent. Trichobothria: 2 rows of 7–8 each on tibiae, 11–12 on metatarsi, 11–12 on tarsi, 7–8 on cymbium. PTC I–IV with 5–6 teeth on each margin.
SPINATION. Palp: femur d6–7, pd4, rd2; patella p1; tibia d2–3, p3, r4–5, v9–10; tarsus d13–15. Leg I: femur d6–7, pd3, rd3; tibia p3–5, r3, v7+m; metatarsus p3, v6. Leg II: femur d5, pd3, rd1; tibia p3, v8; metatarsus p3, v7. Leg III: femur d4, pd3, rd3; patella p3, r2; tibia d1, p3, r3, v8; metatarsus pd4, p3, r2, v9; tarsus p5, r3. Leg IV: femur d5, pd3, rd3; tibia d1, p3, r3, v8; metatarsus pd2, p3, r7, v7; tarsus p5, r6, v3. Patellae I–II and IV, and tarsi I–II aspinose.
PALP. Tibia, cymbium and palpal organ as shown in Figs 182, 211–212. Palpal tibia short cylindrical and spinose (Fig. 182). Palpal organ with embolus tapering and slightly curved (Figs 211–212).
SPINNERETS. See Fig. 282. PMS: length 0.37, diameter 0.15. PLS: maximal diameter 0.40; length of basal, medial and apical segments 0.73, 0.75, 0.44; total length 1.92; apical segment triangular.
LEG MEASUREMENTS. Ƌ(♀)
Female (paratype from Ganjina)
HABITUS. See Fig. 23.
MEASUREMENTS. TBL 13.30, CL 4.43, CW 3.65, LL 0.46, LW 0.93, SL 2.41, SW 2.20.
COLOUR. Similar to that of male, but slightly paler.
PROSOMA. Clypeus and eye tubercle as in Fig 72. Eye diameters and interdistances: AME 0.14(0.20), ALE 0.28, PLE 0.14, PME 0.07, AME–AME 0.14(0.06), ALE–AME 0.13(0.10), ALE–PLE 0.12, PLE– PME 0.05, PME–PME 0.39. Cheliceral rastellum strong and consists of about 15 stout thornlike spikes located in front of fang base and on low mound. Each cheliceral furrow with 6 promarginal teeth and 0–1 smaller teeth alongside. Sternum, labium and maxillae as in Fig. 112. Sternal sigilla very small, submarginal, and lacking certain edges. Maxillae with 30–32 cuspules each.
LEGS. Scopula: distal on metatarsi I and II, entire on palpal tarsus, narrowly divided on tarsi I and II, elsewhere absent. Trichobothria: 2 rows of 6–8 each on tibiae, 14–15 on metatarsi I and II, 12 on metatarsus III, 13 on metatarsus IV, 13 on tarsi I and II, 10 on tarsus III and on palpal tarsus, 12 on tarsus IV. Palpal claw with 4 promarginal teeth. PTC I–II with 3–4 teeth on each margin. PTC III with 3–4 teeth on outer, 3 teeth on inner margins; PTC IV with 3 teeth and 1 tooth, respectively.
SPINATION. Palp: femur pd1–2; patella p1; tibia p3–4, v9–10; tarsus v2–3. Leg I: femur pd1; tibia p1, v5; metatarsus v6. Leg II: femur pd1; patella p1; tibia p2–3, v4–5; metatarsus p1, v7. Leg III: patella d1, p3; tibia d1, p2, r2, v6–7; metatarsus pd2, p2–3, rd4, v7–8; tarsus p1. Leg IV: tibia r2, v6; metatarsus pd 2, rd1–2, v11–12; tarsus p+r+v: ca 20. All femora with dorsal median row of 5–8 long slender bristles; femora III and IV also with pro- and retrodorsal rows of 2–4 spine-like bristles in each row; patellae I and IV, and tarsi I and II aspinose.
SPERMATHECAE. With diverging and branched stalks (see Fig. 250).
SPINNERETS. See Fig. 283. PMS: length 0.48, diameter 0.17. PLS: maximal diameter 0.50; length of basal, medial and apical segments 0.84, 0.60, 0.39; total length 1.83; apical segment triangular.
Variation
The length of the carapace varies from 4.10 to 4.55 in paratype males, the dorsal abdominal pattern may be developed somewhat better and represent the more contrasted variant than that shown in Fig. 10. The number of tarsal spines varies as follows: leg II: 0–3; leg III: 6–12; leg IV: 9–20; the number of the maxillary cuspules ranges as 21–27 in males and 28–32 in females.
Habitat
See the corresponding data for A. incana and A. karatauvi.
Distribution
Southwestern Tajikistan. See Fig. 372.
description: Published as part of Zonstein, Sergei, 2018, A revision of the spider genus Anemesia (Araneae, Cyrtaucheniidae), pp. 1-100 in European Journal of Taxonomy 485 on pages 35-37, DOI: 10.5852/ejt.2018.485, http://zenodo.org/record/3829992