Anemesia Pocock 1895
Creators: name: Zonstein, Sergei
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Cyrtaucheniidae
Anemesia

description: Key to species of Anemesia
Males
1. Palpal tibia distinctly swollen and mostly aspinose (Figs 173–176, 178–179) ................................ 2
– Palpal tibia subcylindrical or cylindrical, and generally with numerous spines and thick bristles (Figs 177, 180–187) .......................................................................................................................... 7
2. Embolus shorter and widely tapering (Figs 188–196) ...................................................................... 3 – Embolus longer and narrowly tapering (Figs 199–205) ................................................................... 6
3. Distance AME–AME subequal to diameter of AME (Figs 48–50) .................................................. 4 – Distance AME–AME considerably larger then diameter of AME (Fig. 51) ...................................... .................................................................................................................... A. tubifex (Pocock, 1889)
4. Carapace and legs light to medium brown; abdomen with distinct dorsal pattern (Figs 1, 3–4) ..... 5 – Carapace and legs dark brown; abdominal pattern indistinct (Fig. 2)................................................. ............................................................................ A. koponeni Marusik, Zamani & Mirshamsi, 2014
5. Carapace and legs paler, light brown (Fig. 3); embolus longer (Figs 193–194) .. A. pococki sp. nov. – Carapace and legs darker, medium brown (Fig. 1); embolus shorter (Figs 188–190) ........................ ................................................................................................................... A. birulai (Spassky, 1937)
6. Palpal tibia aspinose (Fig. 179). Embolus gradually tapering and uniformly curved (Figs 203– 205) ..................................................................................................................... A. sogdiana sp. nov.
– Palpal tibia with numerous spines (Fig. 178). Distal part of embolus more sharply tapering and curved than its proximal part (Figs 199–202) ........................................................ A. oxiana sp. nov.
7. Cymbium and/or palpal tibia with spines (Figs 180–187). Embolus long to very long, with slightly curved and gradually tapering distal part (Figs 206–223) ................................................................ 8
– Palpal tibia and cymbium aspinose (Fig. 177). Embolus relatively short and bent medially, with very thin and narrow distal part (Figs 197–198) ....................................................... A. andreevae sp. nov.
8. Cymbium with numerous dorsal and few lateral spines (Figs 180–183). Subapical embolic keel very small and gradually elevated (Figs 206, 208, 210, 213–214). PMS small and slender (Figs 279–280, 282–284) ........................................................................................................................................... 9
– Cymbium without lateral spines (Figs 184–187). Subapical embolic keel larger and noticeably dilated (Figs 147–150, 215, 218, 220). PMS stouter (Figs 285, 289) ............................................ 12
9. Carapace and legs paler, light brownish (Figs 10–11). Palpal tibia shorter: 3.0–3.2 times as long as wide (Figs 182–183). Tarsus IV with numerous (5–20) spines ...................................................... 10
– Carapace and legs darker, chestnut brown (Figs 8–9). Palpal tibia longer: 4.3–4.6 times as long as wide (Figs 180–181). Tarsus IV with few (1–4) spines ...................................................................11
10. Legs longer:tibia I 4.9–5.0 times as long as wide (Fig. 127). Palpal organ pegtop-shaped, with certainly bulging anterior lobe (Figs 211–212). Dorsal abdominal pattern as in Fig. 10 ..... A. pallida sp. nov.
– Legs shorter: tibia I 4.2 times as long as wide (Fig. 128). Palpal organ comma-shaped, with weakly convex anterior lobe (Figs 213–214). Dorsal abdominal pattern as in Fig. 11 ..... A. parvula sp. nov.
11. Legs longer: tibia I 5.7–5.9 times as long as wide (Fig. 125). Palpal organ with strongly bulging anterior lobe (Fig. 208) ....................................................................................... A. castanea sp. nov.
– Legs shorter: tibia I 5.1–5.2 times as long as wide (Fig. 126). Palpal organ with weakly convex anterior lobe (Fig. 210) ...................................................................... A. karatauvi (Andreeva, 1968)
12. Palpal bulb noticeably (1.3–1.4 times) wider than long (Fig. 215). Clypeus wide, about 2 times exceeding diameter of ALE (Fig. 59). Posterior sigilla long, oval, and located closer to sternal margin (Fig. 101) ....................................................................................... A. incana Zonstein, 2001
– Palpal bulb as wider as long, or very moderately (1.1–1.2 times) wider than long (Figs 218, 220). Clypeus considerably narrower, shorter than diameter of ALE, or only slightly exceeding it (Figs 60– 62). Posterior sigilla smaller, rounded or irregular, and located more distantly from sternal margin (Figs 102–103) ................................................................................................................................ 13
13. Legs longer: tibia I 4.8–4.9 times as long as wide (Fig. 130). Embolus more tapering, with base located closer to retrolateral side of palpal organ (Figs 217–219). Apical segment of PLS longer (Fig. 287) ............................................................................................................ A. infumata sp. nov.
– Legs shorter: tibia I 4.0–4.3 times as long as wide (Figs 131–132). Embolus less tapering, with base located closer to prolateral side of palpal organ (Figs 220–223). Apical segment of PLS shorter (Fig. 289) ........................................................................................................... A. infuscata sp. nov.
Females
Females of A. koponeni Marusik, Zamani & Mirshamsi, 2014 and A. parvula sp. nov. are unknown.
1. Spermathecae 2–3 branched (Figs 224–234)..................................................................................... 2 – Spermathecae entire (Figs 235–249, 251–265) or divided (Fig. 250) .............................................. 5
2. Spermathecae strictly 3-branched, with long stalks and well sclerotised receptacles (Figs 233–234). Habitus as in Fig. 18 ......................................................................................... A. andreevae sp. nov.
– Spermathecae mostly 2-branched, with shorter stalks and poorly sclerotised receptacles (Figs 224– 232). Habitus differs (Figs 15–17) .................................................................................................... 3
3. Distance AME–AME considerably larger then diameter of AME (Fig. 65). Spermathecae as in Figs 230–232. Habitus as in Fig. 17 .......................................................... A. tubifex (Pocock, 1889)
– Distance AME–AME subequal to diameter of AME (Figs 63–64). Habitus and spermathecae differ (Figs 15–16, 224–229) ...................................................................................................................... 4
4. Carapace and legs paler, light brown (Fig. 16). Spermathecae as in Figs 228–229 ........................... ............................................................................................................................... A. pococki sp. nov.
– Carapace and legs darker, medium brown (Fig. 15). Spermathecae as in Figs 224–227 ................... ................................................................................................................... A. birulai (Spassky, 1937)
5. Spermathecae divided (Fig. 250). Dorsal abdominal pattern reduced to longitudinal median band. Habitus as in Fig. 23 .............................................................................................. A. pallida sp. nov.
– Spermathecae entire (Figs 235–249, 251–265). Habitus, including more developed dorsal abdominal pattern, differs (Figs 19–22, 24–26) ................................................................................................. 6
6. Carapace light yellowish brown to medium chestnut brown, chelicerae darker coloured (Figs 19– 22). Spermathecae with relatively long and narrow stalks (Figs 235–249) ..................................... 7
– Carapace and chelicerae uniformly dark brown (Figs 24–26). Spermathecae with relatively short and wide stalks (Figs 251–265) ...................................................................................................... 10
7. Carapace and legs paler, light yellowish brown (Fig. 20). Maxillae with 10–20 cuspules each. Spermathecae with moderately long stalks (Figs 235–240) ............................... A. sogdiana sp. nov.
– Carapace and legs darker, foxy to chestnut brown (Figs 19, 21–22). Maxillae with 28–85 cuspules each. Spermathecae with long to very long stalks (Figs 241–249) .................................................. 8
8. Habitus as in Figs 19, 29. Dorsal abdominal pattern with dark longitudinal median spot having two large lateral extensions. Spermathecae with more narrowly spaced bases and conical stalks (Figs 241–244). Apical segment of PLS triangular (Fig. 276) .............................. A. oxiana sp. nov.
– Habitus as in Figs 21–22. Dorsal abdominal pattern with smaller lateral extensions. Spermathecae with more broadly spaced bases and cylindrical stalks (Figs 245–249). Apical segment of PLS shortly digitiform (as in Fig. 281) ..................................................................................................... 9
9. Clypeus broader, AME and ALE relatively smaller (Fig. 70). Maxillae with 65–70 cuspules each. Spermathecae with narrower spaced, stronger inclined and more tapering receptacles (Fig. 245) ..................................................................................................................... A. castanea sp. nov.
– Clypeus narrower, AME and ALE relatively larger (Fig. 71). Maxillae with 40–55 cuspules each. Spermathecae with broader spaced, lesser inclined and less tapering receptacles (Figs 246– 249) .................................................................................................... A. karatauvi (Andreeva, 1968)
10. Clypeus wide, about 2 times exceeding diameter of ALE (Fig. 73). Posterior sigilla long, oval, and located closer to sternal margin (Fig. 113). Spermathecae with widened bases and oblate receptacles (Figs 251–254) ........................................................................................... A. incana Zonstein, 2001
– Clypeus considerably narrower, shorter than diameter of ALE, or only slightly exceeding it (Figs 74– 77). Posterior sigilla smaller, rounded or irregular, and located more distantly from sternal margin (Figs 114–116). Spermathecae differ (Figs 255–265)......................................................................11
11. Tarsus IV with 3–4 spines. Spermathecae shorter, cap-like, with poorly defined receptacles (Figs 255–257) ................................................................................................... A. infumata sp. nov.
– Tarsus IV with 7–10 spines. Spermathecae longer, mushroom-like, with dilated receptacles (Figs 258–265) ................................................................................................... A. infuscata sp. nov.
description: Published as part of Zonstein, Sergei, 2018, A revision of the spider genus Anemesia (Araneae, Cyrtaucheniidae), pp. 1-100 in European Journal of Taxonomy 485 on pages 8-10, DOI: 10.5852/ejt.2018.485, http://zenodo.org/record/3829992