Clivina wallacei subsp. wallacei Putzeys 1867
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina wallacei
Clivina wallacei wallacei putzeys, 1867

description: Clivina wallacei wallacei Putzeys, 1867
(Figs 30, 65, 97, 123)
Clivina wallacei Putzeys, 1867: 127; Andrewes, 1926: 374; 1929b: 355; 1930: 119; Csiki, 1927: 127; Kult, 1951: 27; Darlington, 1962: 365; 1971: 255; Balkenohl, 2001: 17; Lorenz, 2005: 145; 2024: 5 ZB 8W.
Type material. Lectotype by present designation: ♂, with labels and data: yellow, handwritten in black ink “ C. Wallacei Ptz. Mak. (Wall. Schm.) ” / grey, handwritten in black ink and printed “ Clivina Wallacei Ptz. dét. J. Putzeys” / white, black framed, red printed “ Syntype ” / white, dark blue framed and printed “Soc.Ent.Belg. Coll. PUTZEYS” / white, handwritten in black ink and printed “ Clivina Wallacei Putz. cotype H.E. Andrewes 1928 ” (RBINS).
Additional material. 1 ♀, Dorey / Bowring 63.47* / Clivina Wallacei Putz. Compared with type H.E.A. (NHMUK); 1 ♂, 59.58. Dory, New Guinea. / Clivina Wallacei Putz det. K. Kult 47 (NHMUK); 1 ♂, Bantimoeroeng Haros 4. 30 W. Celebes / Clivina wallacei Putz. H.E.Andrewes det. (CADW); 1 ♂, Celebes Dorey. / Compared with TYPE K. Kult 1946 / C. Wallacei Putz. det. K. Kult 47 / COLLECTIO KAREL KULT COLL.A.DOSTAL, 1999 (CADW); 2 ♂♂, 2 ♀♀, N – SULAWESI Dumoga Bone NP / Doluduo, Licht (16) 22.IV.1992 leg. Jäch (NHMW); 2 specs, S-Sulawesi Prov., Wasuponda env., 600 m, 2.33.13S 121.13.05E, 8.-9.Jul2001, Bolm lgt. (SMNS).
Remarks. In the lectotype, the right front leg is missing.
Putzeys (1867, p. 127) writes at the end of his description [translated from French]: “ M. Schaum found two exemplars in the collection Wallace which he acquired. They came from Makassar (Célèbes). He gave me one. M. de Chaudoir also has two individuals which came from Dorey (Célèbes).” The specimen from M. Schaum is available in RBINS and served for the designation of the lectotype. The whereabouts of the second specimen is unclear. Inquiry at the MFNB revealed the second specimen is not in MFNB in the Schaum collection (written confirmation by the curator Bernd Jäger). The two specimens Putzeys quoted to be in the collection Chaudoir could not be located while checking the Chaudoir collection during my personal visit at the MNHN in November 2021.
Andrewes (1926, p. 373 and 1929b, p. 355) obviously misinterpreted ‘ Dorey’ as a location in New Guinea. He quotes two specimens to be located in RBINS, including the ‘type’. The type was located in RBINS and is now designated as lectotype. The second specimen was dirty and after cleaning identified as member of the Clivina lobata -species group. The two other specimens from ‘Dorey’ Andrewes quoted to be deposited in NHMUK were localized there.
Darlington (1962, p. 330, 331, 366) cleared the misinterpretations regarding the geographic position of ‘Dorey’ also spelled as ‘Dory’. He thinks, this location is located in Celebes (today Sulawesi) and not in New Guinea.
Future collections will confirm if the species occurs outside Sulawesi.
The species is described by Putzeys (1867), redescribed by Andrewes (1926), and keyed out with descriptive information by Darlington (1962).
Compendious description. A medium sized fuscous species with the following measurements: Lectotype: Body length 5.39 mm, width 1.52 mm, ratio L/W of pronotum 1.93, ratio L/W of elytra 1.94. All material: Body length 5.39–5.74 mm (ẍ = 5.54 mm *), width 1.52–1.79 mm (ẍ = 1.61 mm *), ratio L/W of pronotum 0.89–0.94 (ẍ = 0.92*), ratio L/W of elytra 1.79–1.96 (ẍ = 1.91*); (*n = 5).
Head with labrum seven-setose, straight anteriorly. Median part of clypeus separated from clypeal-wing by distinct notch, lateral teeth of clypeus moderately projecting, rounded, clypeal-wing not as far projecting as clypeal-teeth, separated from supraantennal plate by distinct notch. Eye prominent, gena small but distinct. Clypeo-frontal furrow flattened but distinct; clypeus transverse; neck constriction closed, developed as step with punctures posteriorly. Frons with flat pore at middle, with scattered small punctures. Antennae subelongate, not reaching base of pronotum.
Pronotum (Fig. 65) square to sub-cordate, wider than long, slightly convex in frontal and moderately convex in lateral view, with lateral margin slightly convex, widest at middle, angles distinct, posterior angle with tooth somewhat rounded and projecting anteriorly, lateral margin sub-crenate, disk with short impression bilaterally in basal half, with isodiametric reticulation laterally and at base; lateral margin from posterior angle to flange of base slanted with angle of around 30°, basal channel and carina of flange of regular size. Proepisternum with isodiametric reticulation, laterally with some transverse wrinkles.
Elytron (Fig. 30) elongate, flattened in basal half in lateral view, lateral margin slightly convex, subcrenulated in anterior quarter, striae one to four ending free at base, interval three with four setigerous punctures; striae nearly impunctate; intervals slightly to moderately convex, interval seven and eight, and half of interval six as well as base and apical third of elytron isodiametrically reticulated, interval eight carinate in apical fifth, interval seven with shortened step-like carina at apex, striole distinct, setigerous tubercle at base distinct, with distinct tubercle at base of third interval.Abdomen with isodiametric reticulation, transverse at middle, two apical setigerous punctures widely separated.
Hind wing fully developed.
Legs. Intermediate tibia with nine tubercles with setae above the spur at the lateral side.
Male genitalia (Fig. 97). Median lobe relatively short, regularly arcuate in basal half, nearly straight in apical half, apex parable-like acuminated. Parameres moderately slender, distorted and hyaline in apical quarter, with medium long seta each.
Female gonocoxites and epipleurite (Fig. 123). Basal gonocoxite with apex acuminated into a tubercle with long seta, with second long seta in apical third, apical gonocoxite elongated, slightly curved, cylindrical in cross section, with two setae basally, with one medium sized seta in basal third and one at middle, with long seta at end of apical third, with two long setae apically. Epipleurite trapezoid, with small seta at each edge.
Variation. In one of the male specimens, the lateral margin of the pronotum is slightly more convex. On the frons of the head the number of scattered punctures vary.
Distribution. The species occurs on the island of Sulawesi. The occurrence in New Guinea needs confirmation (see remarks above).
description: Published as part of Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), pp. 1-70 in Zootaxa 5592 (1) on pages 47-49, DOI: 10.11646/zootaxa.5592.1.1, http://zenodo.org/record/14953674