Landouria zonifera Nurinsiyah & Neiber & Hausdorf 2019, sp. nov.
Creators: name: Nurinsiyah, Ayu Savitri
name: Neiber, Marco T.
name: Hausdorf, Bernhard
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Camaenidae
Landouria
Landouria zonifera

description: Landouria zonifera sp. nov.
urn:lsid:zoobank.org:act: 81A2A657-B12B-43BE-B19D-22AE36E2525C
Figs 44, 54, 68, 82; Tables 3–4
Diagnosis
Landouria zonifera sp. nov. is characterized by a keeled, corneous shell with a brownish zone at the periphery, a wide umbilicus and a flagellum that is more than twice as long as the short epiphallus.
Etymology
Landouria zonifera sp. nov. is named after the brownish zone at its shell periphery (Latin: ʻzonaʼ = ʻzoneʼ, ʻferreʼ = ʻto carryʼ; used as an adjective).
Material examined
Holotype
INDONESIA • holotype (det. anat.); Yogyakarta, Kulonprogo, Sibolong, snakefruit plantation; 7°44′44″ S, 110°08′02″ E; 716 m a.s.l.; 9 Jan. 2013; A.S. Nurinsiyah and T. Setiadi leg.; D = 13.2 mm, H = 7.6 mm; MZB 17727.
Paratypes
INDONESIA • 1 spec.; Yogyakarta, Kulonprogo, Sibolong, Sibolong Hill, agroforestry; 7°45′00″ S, 110°07′00″ E; 721 m a.s.l.; MZB 17762 • 13 spec.; same data as for holotype; MZB 17720 • 1 spec.; same data as for holotype; ZMH 133420 • 4 spec.; Kulonprogo, Jonggrangan; 7°44′20″ S, 110°08′32″ E; 730 m a.s.l.; ZMH 140031.
Description
SHELL (Fig. 82; Tables 3–4). Depressed conical, with 5.5 –5.75 hardly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles and indistinct incised spiral lines; scaly processes sparse, mainly at periphery; tuberculate around umbilicus; corneous with a brownish zone at periphery; body whorl keeled; aperture rounded rhombic; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 40% of shell diameter, hardly obscured by columellar edge.
GENITALIA (Figs 44, 68; Table 4). Atrium short; penis long, cylindrical, slightly dilated at proximal end, without penis coecum; membranaceous sheath around distal part of penis connected with proximal part of epiphallus by tissue; penial retractor runs from diaphragm to proximal part of short, cylindrical epiphallus; flagellum more than twice as long as epiphallus, broadest near its proximal end, then abruptly tapering, slightly crenated, without node; vas deferens narrow and long; vagina very long, cylindrical; oviduct shorter, broad, proximally tapering; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.
Remarks
Landouria zonifera sp. nov. clusters in the molecular phylogeny (Fig. 1) with L. abdidalem sp. nov. (Fig. 56) and L. tholiformis sp. nov. (Fig. 51). Landouria zonifera sp. nov. and L. abdidalem sp. nov. have overlapping ranges. Landouria zonifera sp. nov. differs from L. abdidalem sp. nov. in the smaller (D 12.5–13.2 mm vs 13.1–16.4 mm in L. abdidalem sp. nov.), more depressed (D/H 1.73–1.81 vs 1.93–2.14 in L. abdidalem sp. nov.), corneous shell with a brownish zone at the periphery, the shorter penis and a flagellum that is more than twice as long as the short epiphallus. Furthermore, Landouria zonifera sp. nov. occurs sympatrically with L. menorehensis sp. nov. (Fig. 88) from which it can easily be distinguished by the keeled, depressed conical, corneous shell with a brownish zone at the periphery.
Distribution
Landouria zonifera sp. nov. is endemic to the Menoreh karst area (Fig. 54).
description: Published as part of Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, pp. 1-73 in European Journal of Taxonomy 526 on pages 60-61, DOI: 10.5852/ejt.2019.526, http://zenodo.org/record/3152217