cometadata/datacite-rtg-text-other-reclassification-records huggingface dataset:
Lesteva morvani Jarrige 1971
Creators: name: Shavrin, Alexey V.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Lesteva
Lesteva morvani

description: Lesteva (s.str.) morvani Jarrige, 1971
Fig. 9
Lesteva morvani Jarrige, 1971: 489
Type material examined: Holotype of Lesteva morvani, ♂ [beetle glued on rectangular plate, eight abdominal segment glued under the beetle; aedeagus missing]: ‘massif du KUH-I-KAHAR’, ‘a l'ouest de GASH-I-SAR’, ‘accolé a' ur névé’, ‘AIT. 3000 m. versant NORD’, ‘ 16.7.1968 ’, ‘ ELBURS IRAN’, ‘Morvan Pierre’ , ‘TYPE’ , ‘ Lesteva morvani Jarr [ige].’  (MHNH).
Material examined: IRAN: 1 ♀: ‘ ELBURS IRAN’, ‘massif du KUH-I-KAHAR’, ‘a l'ouest de GASH-I- SAR’, ‘ 16.7.1968 ’, ‘AIT. 3000 vers. NORD’, ‘sous une Pierre très humide’, ‘Morvan Pierre’ (MHNH).
Remarks. This species was described from a single male by Jarrige (1971) from North Iran. Based on the shape of aedeagus, it was compared with L. luctuosa Fauvel, 1871. Unfortunately, the aedeagus of the holotype is missing. It is possible that it had been glued above the beetle as there is a small piece of glue remaining there. In Paris Museum I found a female from the same locality as the holotype, which I identified as L. morvani based on external characters. The species has very fine punctation of the forebody, which was not observed in other species mentioned in this study, and has moderately elongated atennomeres (Fig. 9). Similar species are known from the Caucasus, for example L. zolotarevi Roubal, 1910, and some related, undescribed species with the same character of punctation of the forebody and a wide aedeagus, which was figured by Jarrige (1971; Fig. 6). At the present time, a redescription of the species is not possible and additional comparative material is required.
description: Published as part of Shavrin, Alexey V., 2014, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) of Iran and adjacent territories, pp. 122-140 in Zootaxa 3884 (2) on page 136, DOI: 10.11646/zootaxa.3884.2.2, http://zenodo.org/record/4951550
Tiramnia Grunt 1977
Creators: name: Poletaev, Vladyslav
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Brachiopoda
Rhynchonellata
Spiriferida
Martiniidae
Tiramnia

description: Genus Tiramnia Grunt, 1977
Type species
Martinia uralica Chernyshev, 1902; original designation.
Diagnosis
Shells small to medium-sized, brachythyrid outlines, without plates or septa in either valve; fold and sulcus weakly or moderately developed; furrow in sulcus is very weak or absent. Surface smooth or with the weak, short ribs on anterior edge. Muscle field of ventral valve comparatively large, lanceolate; radial vascular imprints ramiform; axial mantle canal (vascula media) singular.
Included Pennsylvanian species
Arctic Russia, the northern Ural (Chernyshev 1902; Grunt 1977): Tiramnia uralica (Chernyshev, 1902), T. semiglobosa (Chernyshev, 1902) Permian, Cisuralian; T. tschaikensis Grunt, 1977; upper Pennsylvanian, Cape Chaika.
Arctic Canada and Southeastern Alaska (Carter & Poletaev 1998): Tiramnia waltery Poletaev, 1998 and T. grunti Poletaev, 1998; Pennsylvanian.
Greenland (Grunt 1977): Tiramnia greenlandica Grunt, 1977; Permian.
China (Chao 1929): Tiramnia changchiacouensis Chao, 1929; Pennsylvanian.
Ukraine (this paper) Tiramnia davidi sp. nov., Lower Pennsylvanian.
Remarks
Tiramnia Grunt, 1977 differs from Martinia M’Coy, 1844 by the presence of a ramified vascular system; Tiramnia differs from Jilinmartinia Li & Gu, 1980 by having single (connected) vascula media rather than paired vascula developed in the ventral interior of the latter. Tiramnia differs from the somewhat similar Postamartinia Wang & Yang, 1993 by having round, elongated, but not transverse shells, by the absence of low ribs on the flanks, and by a much shorter tongue of sulcus.
description: Published as part of Poletaev, Vladyslav, 2024, New and revised taxa of Carboniferous spiriferides (Brachiopoda, Spiriferida) from the Donets Basin (Ukraine) and South Urals (Russia), pp. 132-155 in European Journal of Taxonomy 968 on page 136, DOI: 10.5852/ejt.2024.968.2723, http://zenodo.org/record/14047314
Brachythyridoidea Frederiks 1924
Creators: name: Poletaev, Vladyslav
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Spiriferida

description: Superfamily Brachythyridoidea Frederiks, 1924
Diagnosis emended
Outline commonly ovate; hinge line shorter than width, weakly denticulated; cardinal extremities rounded; lateral slopes usually with few simple coarse ribs or with numerous flat bifurcated ribs. Microornament absent or weakly developed. Delthyrium partially covered by deltidium or stegidial plates; subdelthyrial plates absent; ventral adminicula absent; vascular imprints pinnate or ramiform; miophragm or septum in ventral valve usualy absent. Dorsal adminicula and crural plates absent.
Remarks
The presence of a denticulated hinge line in the ventral interarea of Spirifera ovalis Phillips, 1836 has not yet been demonstrated, but this character has been found and is supported by some other data. Angiolini et al. (2011: 71) revised the Carboniferous brachiopod fauna from historical sections in England.As a result, they established the genus Latibrachythyris Angiolini et al., 2011. The type species of Latibrachythyris is Spirifer pinguis Sowerby, 1821 and has a denticulated hinge line. Carniti et al. (2022: 865) attributed Spirifer crassus de Koninck, 1883 to Latibrachythyris and, following Angiolini et al. (2011), also Spirifer rotundatus Sowerby, 1825. There is no doubt that the authors’ definitions of these species are correct, but the exact definition of some Brachythyris -like genera is problematic. An examination of the brachiopod collections of de Koninck (1883, 1887) at the Royal Belgian Institute of Natural Sciences in Brussels, Belgium, in 1992 showed that specimens of Spirifer subrotundatus M’Coy, 1844 (sensu de Koninck, 1887) have a primary median costa in the sulcus and a distinct short septum in the ventral valve. Therefore, this species must now be assigned to Phragmobrachythyris Poletaev 1999 from the family Skelidorygmidae Carter, 1994 (Carter & Gourvennec 2007). A correct genus definition of all Brachythyris -like spirifers is only possible by using serial cross sections of the apical part of the shell. The data presented above have shown that Latibrachythyris has a weakly denticulated hinge line and Phragmobrachythyris has a miophragm or septum in the ventral valve. Altogether, these data do not allow to assign the families Brachythyrididae Frederiks, 1924 and Skelidorygmidae to the superfamily Martinioidea Waagen, 1883, as I had done earlier (Poletaev 2022).
Stratigraphic distribution
Late Devonian to Permian.
description: Published as part of Poletaev, Vladyslav, 2024, New and revised taxa of Carboniferous spiriferides (Brachiopoda, Spiriferida) from the Donets Basin (Ukraine) and South Urals (Russia), pp. 132-155 in European Journal of Taxonomy 968 on pages 138-139, DOI: 10.5852/ejt.2024.968.2723, http://zenodo.org/record/14047314
Brachythyrididae Fredericks 1924
Creators: name: Poletaev, Vladyslav
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Brachiopoda
Rhynchonellata
Spiriferida
Brachythyrididae

description: Family Brachythyrididae Fredericks, 1924
Diagnosis
Primary median costa in sulcus absent.
Stratigraphic distribution
Late Devonian (Famennian) to Permian (Lopingian).
description: Published as part of Poletaev, Vladyslav, 2024, New and revised taxa of Carboniferous spiriferides (Brachiopoda, Spiriferida) from the Donets Basin (Ukraine) and South Urals (Russia), pp. 132-155 in European Journal of Taxonomy 968 on page 139, DOI: 10.5852/ejt.2024.968.2723, http://zenodo.org/record/14047314
Meristorygma Carter 1974
Creators: name: Poletaev, Vladyslav
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Brachiopoda
Rhynchonellata
Spiriferida
Brachythyrididae
Meristorygma

description: Genus Meristorygma Carter, 1974
Type species
Meristorygma arctica Carter, 1974; original designation.
Diagnosis
Medium-sized shell with diverse outline; fold and sulcus moderately developed. Well-defined sulcus with two, rarely four primary ribs and median furrow; uniplicate anterior commissure usually with subquadrate tongue; fold with three distinct ribs; the central with weakly furrow. Ribs on sides are few and weak. Hinge line of short, smooth ventral interarea non-denticulated. Ventral adminicula and dorsal crural plates are absent; vascula ramose, radially arranged with a pair of strong vascular media, which correspond to the pair of costae in the sulcus. Microornament is absent.
Species included
Arctic Canada and southeastern Alaska (Carter 1974: 674): Meristorygma arctica Carter, 1974; Pennsylvanian.
Russia, Northern and Polar Ural (Kutorga 1844; Kalashnikov 1980): M. panduriformis (Kutorga, 1844); upper Pennsylvanian–Lower Permian; M. pajkchoica Kalashnikov, 1980; Moscovian.
South Ural (Bashkortostan)and Middle Asia (Yanishevsky 1900; Poletaev 2018): M. gracilis (Yanishevsky, 1900) upper Serpukhovian–lower Bashkirian; (this paper) M. donakovae Poletaev, 2018; uppermost Visean–Serpukhovian, Magnitogorsk district.
Ukraine, Donets Basin (Rotai 1951; Poletaev 2018): M. miussensis (Rotai, 1951) upper Bashkirian; M. postgracilis (Rotai, 1951); Moscovian.
Remarks
The analysis of the ventral valve of the natural mould of Meristorygma arctica (Carter & Poletaev 1998) has shown that the species has a ramose, radially arranged vascular system with a twin of strong vascular media, in contrast to Brachythyris ovalis (Phillips, 1836), which has a weak, simple, pinnate vascular system. The variability of the vascular system of related taxa may mostly be the result of evolution (Lazarev & Poletaev 1982).
description: Published as part of Poletaev, Vladyslav, 2024, New and revised taxa of Carboniferous spiriferides (Brachiopoda, Spiriferida) from the Donets Basin (Ukraine) and South Urals (Russia), pp. 132-155 in European Journal of Taxonomy 968 on page 139, DOI: 10.5852/ejt.2024.968.2723, http://zenodo.org/record/14047314
Meristorygma donakovae Poletaev 2018
Creators: name: Poletaev, Vladyslav
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Brachiopoda
Rhynchonellata
Spiriferida
Brachythyrididae
Meristorygma
Meristorygma donakovae

description: Meristorygma donakovae Poletaev, 2018
Fig. 4A
Meristorygma donakovae Poletaev, 2017: 74 (nom. nud.).
Meristorygma donakovae Poletaev, 2018: 88, pl. 112 fig. 3.
Spirifer sp. – Yanishevsky 1900: 150, pl. 1 fig. 7.
Spirifer aff. gracilis – Yanishevsky 1910: 96, pl. 12 fig. 17.
Brachythyris integricostus – Donakova 1983: pl. 2 figs 6– 7.
Diagnosis
Transversely-ovate shell with one to two pairs of ribs in the sulcus, small rounded tongue and relatively broad, scanty ribs on the lateral slopes.
Material examined
Holotype (Fig. 4) RUSSIA • South Urals, Magnitigorsk region, right bank of Ural River; Kizil Formation, upper Visean and Serpukhovian; Donakova 1983 Coll.; CNIGR Museum 12065/8.
Paratypes RUSSIA • 2 specs; same data as for holotype; CNIGR 12065/9, 12065/10.
Additional material
RUSSIA • 20 specs; same data as for holotype; CNIGR 12065.
Description
Shell small to medium-sized; ventral valve with transversely-ovate outline, approximately equidimensional (holotype with length 26 mm, width 28 mm, thickness 18 mm), thin-walled. Hinge margin shorter than greatest width; interarea low, triangular, anacline to apsacline, smooth. Sulcus moderately developed, well delineated, with median furrow and two, rarely four primary ribs arising from parietal ribs. Anterior margin uniplicate with a small rounded tongue. Ribs on lateral slopes are few (5–7), well expressed. Dorsal valve oval in outline, almost such convex as ventral valve. Fold with three distinct ribs, of which two lateral ribs weakly bifurcate; interspaces between ribs broad. Surface of ribs smooth with concentric traces of growth. Ventral adminicula absent. Features of vascular system not established.
Remarks
This species differs from Meristorygma arctica by wider transversely oval outlines of the shell and a short, rounded tongue with a pair of weak additional ribs on its slopes, whereas M. arctica has a long, flat and subquadrate tongue with a pair of ribs. Meristorygma donakovae differs from the geographically and stratigraphically closest species M. gracilis by wider and less numerous ribs on the lateral slopes of both valves. The specimens examined resemble M. postgracilis in outline and rib pattern, but the latter has a narrower shell, more frequent bifurcation of the ribs in the sulcus, and a subquadrate tongue. Meristorygma parva (Rotai, 1951) differs in small size and low total number of ribs (not more than eight); M. miussensis and M. panduriformis in contrast are much larger and have very rare bifurcation of ribs in the sulcus.
description: Published as part of Poletaev, Vladyslav, 2024, New and revised taxa of Carboniferous spiriferides (Brachiopoda, Spiriferida) from the Donets Basin (Ukraine) and South Urals (Russia), pp. 132-155 in European Journal of Taxonomy 968 on pages 140-141, DOI: 10.5852/ejt.2024.968.2723, http://zenodo.org/record/14047314
Betaneospirifer Gatinaud 1949
Creators: name: Poletaev, Vladyslav
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Brachiopoda
Rhynchonellata
Spiriferida
Spiriferidae
Betaneospirifer

description: Genus Betaneospirifer Gatinaud, 1949
Type species
Spirifera moosakhailensis Davidson, 1862: 28; original designation.
Diagnosis
Shells relatively thick-walled, medium to large in size; with transverse outlines; cardinal extremities from weakly rounded to mucronate or alate; ventral area triangular or trapezoidal with subparallel lower and upper edges, usually apsacline; hinge line denticulate; lateral slopes moderately or sharply folded, covered with numerous fasciculated almost uniform in width ribs, microornament regularly imbricate. Ventral valve has short deltirial plate, composed of short, rapidly diverging adminicula and dental flanges.
Remarks
The remark in the description of Betaneospirifer (Carter et al. 2006: 1790) “The validity of this genus is contingent on confirmation and completion of the diagnosis of Neospirifer ” was related to the error in the choice of the lectotype of Spirifer fasciger Keyserling, 1846 as the type species of Neospirifer Frederiks, 1924. The revision of Keyserling’s (1846) collection published by Poletaev (1997) showed that the specimen SPGGI 129/46, chosen by Archbold & Thomas (1984) as the lectotype of Spirifer fasciger, most probably does not belong to the real syntypes of this species, because its image is missing among the primary good quality drawings of Keyserling’s specimens. In addition, this specimen does not show any characters of the species and has to be rejected as lectotype. Specimen SPGGI 128/46 has been proposed (Poletaev 1997) as neolectotype of Spirifer fasciger (not paralectotype as decided by Carter et al. 2006: fig. 1185, 1b–c). It is the “ragged” ventral valve, which has an ovate overline, a few weak radial plicae on the flanks, and obviously belongs to Neospiriferinae. This ventral valve was first shown in dorsal view by Poletaev (1997) and has a very unusual type of short triangular ventral area and a trace of a delthyrial plate. The difference between the confirmed type species of Neospirifer and Spirifera moosakhailensis, the type species of Betaneospirifer, is obvious. The genus Neospirifer includes only the type species and some closely related species from the Productus Limestone (Late Permian) of Salt Range (Pakistan), described by Reed (1944: 678) and Waterhouse (2004: 1–265). Most of the thick-walled species of Neospiriferinae with long, trapezoidal ventral area belong to Betaneospirifer and other neospiriferid genera.
Distribution
Carboniferous (late Moscovian) to Early Permian of Russia and Pakistan.
description: Published as part of Poletaev, Vladyslav, 2024, New and revised taxa of Carboniferous spiriferides (Brachiopoda, Spiriferida) from the Donets Basin (Ukraine) and South Urals (Russia), pp. 132-155 in European Journal of Taxonomy 968 on page 141, DOI: 10.5852/ejt.2024.968.2723, http://zenodo.org/record/14047314
Paeckelmanelloidea Ivanova 1972
Creators: name: Poletaev, Vladyslav
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia

description: Superfamily Paeckelmanelloidea Ivanova, 1972
Diagnosis (emended)
Moderately to strongly transverse shell with maximum of width at hinge line; flanks usually with simple ribs, rarely smooth; stratigraphically younger forms sometimes with bifurcations; fold and sulcus smooth or with median rib in sulcus and on fold, more rarely multicostate or with pair of opposed plica on both valves; ventral interarea usually moderately to very high, often catacline to procline or weakly apsacline; hinge line denticulate; microornament usually consisting of capillae and anteriorly free growth lamellae; dental adminicula usually present; vascular markings simple, radial; crural plates and dorsal adminicula lacking; ctenophoridium present.
description: Published as part of Poletaev, Vladyslav, 2024, New and revised taxa of Carboniferous spiriferides (Brachiopoda, Spiriferida) from the Donets Basin (Ukraine) and South Urals (Russia), pp. 132-155 in European Journal of Taxonomy 968 on page 143, DOI: 10.5852/ejt.2024.968.2723, http://zenodo.org/record/14047314
Johnsoniana Poletaev, 2024, nom. nov.
Creators: name: Poletaev, Vladyslav
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Spiriferida
Oceaniidae
Johnsoniana

description: Genus Johnsoniana nom. nov.
Type species
Spirifer oceani d’Orbigny, 1849; original designation (by monotypy).
Diagnosis
Shells small to medium-sized, pentagonal or trapezoidal outline with equibiconvex profiles; ventral interarea triangular, medium high, concave only under apex, apsacline to catacline with denticulate hinge line equal to the width of shell; cardinal extremities acute but not mucronate; delthyrium narrowly elongate, triangular, high, partly covered by pseudodeltidium; shallow ventral sulcus and median depression on fold clearly limited by coarse rounded opposite plica, divided each valve into three depressed areas; flank plications absent; anterior commissure subrectilinear. Surface smooth or with concentric wrinkles and obscure irregular radial striae (10–12 per mm). In ventral valve strong deltirial ridges, boarding delthyrium, septum and dental plates absent; in dorsal valve short base of crural plate and flat ctenophoridium. Shell substance impunctate.
Stratigraphic and geographic range
Western Europe (Belgium), Russia, Urals (Bashkortostan); Mississippian, from the Late Tournaisian to Viséan (late Warnantian or late Asbian).
Included species
Type species only.
description: Published as part of Poletaev, Vladyslav, 2024, New and revised taxa of Carboniferous spiriferides (Brachiopoda, Spiriferida) from the Donets Basin (Ukraine) and South Urals (Russia), pp. 132-155 in European Journal of Taxonomy 968 on page 146, DOI: 10.5852/ejt.2024.968.2723, http://zenodo.org/record/14047314
Condyloderes Higgins 1969, sp. nov.
Creators: name: Neuhaus, Birger
name: Zotto, Matteo Dal
name: Yamasaki, Hiroshi
name: Higgins, Robert P.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Cephalorhyncha
Kinorhyncha
Cyclorhagida
Centroderidae
Condyloderes

description: Condyloderes Higgins, 1969
Included species. Condyloderes kurilensis Adrianov & Maiorova, 2016; C. multispinosus (McIntyre, 1962) Higgins, 1969; C. paradoxus Higgins, 1969; C. setoensis Adrianov, Murakami & Shirayama, 2002; C. shirleyi sp. nov. Neuhaus & Higgins; C. storchi Higgins, 2004 in Martorelli & Higgins, 2004.
Emended diagnosis. Mouth cone with 9 individual outer oral styles with broader base and thin and flexible anterior part; 14 trichoscalids each originating from trichoscalid plate and accompanied anteriorly by two pointed filamentous appendages; 16 placids with broader midventral placid, neighboured by two narrower placids and alternatingly a broader and a narrower placid; all placids with knobby projections (= condyles) in one or two rows in narrower placids and two or three rows in broader placids; midventral placid with three apical condyles, two in intermediate row, and 3–6 in basal row; ventromedial to midventral free flap and primary pectinate fringe of segment 1 at least partly reduced; acicular spine middorsally on segments 1–9, midterminally on segment 11 (= midterminal spine), lateroventrally on segments 1–9, laterodorsally on segment 10 (male only in species with irregularly distributed or almost no cuticular hairs), and lateral accessorily on segment 11 (= lateral terminal accessory spine); cuspidate spines present; type-3 sensory spot ventrolaterally next to lateral terminal accessory spine and subdorsally on segment 11; type-6 sensory spot elevated above trunk surface giving the impression to be half-drowned in the trunk cuticle; ventromedial appendage at least on two segments of segments 5–8 in female; area of micropapillae ventromedially on segment 9 in female of species with irregularly distributed or almost no cuticular hairs.
description: Published as part of Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi & Higgins, Robert P., 2019, Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov., pp. 1-91 in Zootaxa 4561 (1) on pages 13-14, DOI: 10.11646/zootaxa.4561.1.1, http://zenodo.org/record/2627892
Condyloderes kurilensis Adrianov & Maiorova 2016
Creators: name: Neuhaus, Birger
name: Zotto, Matteo Dal
name: Yamasaki, Hiroshi
name: Higgins, Robert P.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Cephalorhyncha
Kinorhyncha
Cyclorhagida
Centroderidae
Condyloderes
Condyloderes kurilensis

description: Re-description of Condyloderes kurilensis Adrianov & Maiorova, 2016
(Figs 4–7; Tables 1, 2, 7)
Condyloderes kurilensis Adrianov & Maiorova, 2016: pp. 13–18, figs 1–4, table 1.
Condyloderes kurilensis — Zeppilli et al. 2018: p. 56.
Diagnosis. Neck placids with condyles in two rows in narrower placids and 2–3 rows in broader placids; each narrower placid with one apical and one basal condyle; primary pectinate fringe of all segments reduced, free flap with numerous short spinose or broad extensions on most parts of a segment; acicular spine middorsally on segments 1–9 (and on segment 10 in male only), midterminally on segment 11, laterodorsally on segment 10 (in male only), and lateroventrally on segments 1–9; cuspidate spine ventrolaterally on segment 5 and lateral accessorily on segment 8; type-6 sensory spot ventromedially on segments 1 (slightly more paraventrally), 2, 4, 6, 8 and 9, lateroventrally on segment 10, sublaterally on segments 3, 5–7 and 9, midlaterally on segments 1 and 2, laterodorsally on segments 2, 3 and 7–9, and paradorsally on segments 1 (almost subdorsally), 4–6, 8 and 9; ventromedial appendage on segments 7 and 8 and area of micropapillae ventromedially on segment 9 in female only.
Material examined. Holotypic male (MIMB 32339) and paratypic female (MIMB 32340) mounted for light microscopy (Fig. 1; Table 1).
description: Published as part of Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi & Higgins, Robert P., 2019, Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov., pp. 1-91 in Zootaxa 4561 (1) on page 14, DOI: 10.11646/zootaxa.4561.1.1, http://zenodo.org/record/2627892
Condyloderes multispinosus Higgins 1969
Creators: name: Neuhaus, Birger
name: Zotto, Matteo Dal
name: Yamasaki, Hiroshi
name: Higgins, Robert P.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Cephalorhyncha
Kinorhyncha
Cyclorhagida
Centroderidae
Condyloderes
Condyloderes multispinosus

description: Condyloderes multispinosus (McIntyre, 1962) Higgins, 1969
Centroderes multispinosus McIntyre, 1962: p. 505, fig. 1.
Condyloderes multispinosus comb. nov. — Higgins, 1969: pp. 3–6, figs 1–9, table 1.
Condyloderes multispinosus — Higgins 1971: p. 26.— Moore 1973: p. 341, 350.— Brown 1985: p. 93.— Higgins 1986: p. 115.— Higgins 1988: fig. 28.1d.— Huys & Coomans 1989: pp. 211, 213, fig. 1.— Nebelsick 1993: p. 221, 226.— Adrianov & Malakhov 1994: p. 229, fig. 130b.—— Bamber 1997: pp. 65, 424.—Adrianov & Malakhov 1999: pp. 114, 253, 254, fig. 5.10B, C.— Adrianov et al. 2002: p. 205, 206, 214, 215.— Martorelli & Higgins 2004: pp. 88–90, tables 1, 2.— Sørensen & Pardos 2008: pp. 52, 53.— Sørensen et al. 2010b: pp. 234, 241, [not Centroderes].—Neuhaus 2013: pp. 291, 326, tables 6, 11 [not Centroderes multispinosus].— Neuhaus & Sørensen 2013: p. 73.— Adrianov & Maiorova 2016: pp. 11, 18.— Dal Zotto & Todaro 2016: p. 99.
description: Published as part of Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi & Higgins, Robert P., 2019, Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov., pp. 1-91 in Zootaxa 4561 (1) on page 20, DOI: 10.11646/zootaxa.4561.1.1, http://zenodo.org/record/2627892
Condyloderes multispinosus Neuhaus & Zotto & Yamasaki & Higgins 2019
Creators: name: Neuhaus, Birger
name: Zotto, Matteo Dal
name: Yamasaki, Hiroshi
name: Higgins, Robert P.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy

description: Re-description of Condyloderes multispinosus based on type material
(Figs 8–10; Tables 1–3, 7)
Diagnosis. Neck placids with condyles in one row in narrower placids and three rows in broader placids; each narrower placid with one basal condyle; each broader placid with two condyles both in apical, intermediate, and basal row; midventral placid with three apical and basal condyles and two in intermediate row; acicular spine middorsally on segments 1–10, midterminally on segment 11, lateroventrally on segments 1–9, and laterodorsally on segment 10; cuspidate spine ventrolaterally on segments 2, 5 and 9 and lateral accessorily on segment 8; type-6 sensory spot ventromedially on segments 1 (slightly more paraventrally), 2–4 and 6–9, lateroventrally on segment 10, sublaterally on segments 1–3 and 5–9, midlaterally on segments 1 and 2, laterodorsally on segments 1–9, subdorsally on segment 10, and paradorsally on segments 1 (almost subdorsally), 2 and 49; gland cell outlet ventromedially on segment 10.
Material examined. Holotypic male (ZOO 1962.82, previously identified as adult by McIntyre (1962) and Higgins (1969)) mounted for light microscopy (Fig. 1; Table 1).
description: Published as part of Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi & Higgins, Robert P., 2019, Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov., pp. 1-91 in Zootaxa 4561 (1) on page 21, DOI: 10.11646/zootaxa.4561.1.1, http://zenodo.org/record/2627892
Condyloderes paradoxus Higgins 1969
Creators: name: Neuhaus, Birger
name: Zotto, Matteo Dal
name: Yamasaki, Hiroshi
name: Higgins, Robert P.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Cephalorhyncha
Kinorhyncha
Cyclorhagida
Centroderidae
Condyloderes
Condyloderes paradoxus

description: Condyloderes paradoxus Higgins, 1969
Condyloderes paradoxus Higgins, 1969: pp. 6–9, figs 10–20, table 2.
Condyloderes paradoxus — Moore 1973: p. 350. Higgins & Rao 1979: p. 76.— Higgins 1983: p. 2. Higgins 1986: p. 115.— Adrianov & Malakhov 1994: p. 229, fig. 130c.— Rao 1998: p. 82.—Adrianov & Malakhov 1999: pp. 114, 253, 254, fig. 5.10D, E.— Adrianov et al. 2002: p. 205, 206, 214, 215.— Martorelli & Higgins 2004: pp. 88–90, tables 1, 2.— Sørensen 2007: pp. 165, 166, table 3.— Sørensen & Pardos 2008: p. 53.— Sørensen et al. 2010b: pp. 235, 241.—Neuhaus 2013, pp. 329, 292, 329, figs 5.5.2C, 5.8.2A, table 6.— Neuhaus & Sørensen 2013: p. 73.— Venkataraman et al. 2015: p. 3.— Sørensen et al. 2016: pp. 3, 4, 12, 13, fig. 6, table 1.— Adrianov & Maiorova 2016: pp. 11, 18.— Jeeva & Mohan 2016: p. 251.
Condyloderes cf. paradoxus — Sørensen et al. 2012a: p. 163, table 1.—this paper: Late juvenile stage of Condyloderes paradoxus or C. setoensis [either C. paradoxus or C. setoensis].
description: Published as part of Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi & Higgins, Robert P., 2019, Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov., pp. 1-91 in Zootaxa 4561 (1) on page 31, DOI: 10.11646/zootaxa.4561.1.1, http://zenodo.org/record/2627892
Condyloderes paradoxus Higgins 1969
Creators: name: Neuhaus, Birger
name: Zotto, Matteo Dal
name: Yamasaki, Hiroshi
name: Higgins, Robert P.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy

description: Re-description of Condyloderes paradoxus based on type material
(Figs 13–17; Tables 1, 2, 7)
Diagnosis. Neck placids with condyles in two rows in narrower placids and three rows in broader placids; narrower placids with one apical and one basal condyle; broader placids with two condyles both in apical, intermediate, and basal row; midventral placid with three apical and basal condyles and two in intermediate row; acicular spine middorsally on segments 1–10, midterminally on segment 11, lateroventrally on segments 1–9 (and on segment 10 in female only), and laterodorsally on segment 10 (male only); cuspidate spine ventrolaterally on segments 2, 5 and 9 and lateral accessorily on segments 4 and 8; type-6 sensory spot ventromedially on segments 1 (slightly more paraventrally) and 2–9, lateroventrally on segment 10 (male only), sublaterally on segments 1–3 and 5–9, midlaterally on segments 1 and 2, laterodorsally on segments 1–9, subdorsally on segment 10, and paradorsally on segments 1 (almost subdorsally), 2 and 4–9; ventromedial appendage on segments 7 and 8 and area of micropapillae ventromedially on segment 9 in female only.
Material examined. Holotypic male (USNM 37460), four paratypic females (USNM 37461, 1209809; USNM 1209808 and 1209812, previously identified as males), six paratypic males (USNM 1209804, 1209805, 1209807, 1209810, 1209813, 1209814), and one juvenile moulting (USNM 1209811) (Fig. 1; Table 1).
description: Published as part of Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi & Higgins, Robert P., 2019, Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov., pp. 1-91 in Zootaxa 4561 (1) on page 31, DOI: 10.11646/zootaxa.4561.1.1, http://zenodo.org/record/2627892
Condyloderes setoensis Adrianov, Murakami & Shirayama 2002
Creators: name: Neuhaus, Birger
name: Zotto, Matteo Dal
name: Yamasaki, Hiroshi
name: Higgins, Robert P.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Cephalorhyncha
Kinorhyncha
Cyclorhagida
Centroderidae
Condyloderes
Condyloderes setoensis

description: Condyloderes setoensis Adrianov, Murakami & Shirayama, 2002
Condyloderes setoensis Adrianov, Murakami & Shirayama, 2002: pp. 208–215, figs 2–6.
Condyloderes setoensis — Martorelli & Higgins 2004: pp. 88–90, tables 1, 2.— Sørensen & Pardos 2008: p. 53, fig. 14.— Sørensen et al. 2010b: pp. 235, 241.—Yamasaki et al. 2012: p. 23.—Yamasaki & Kajihara 2012: p. 109.— Adrianov & Maiorova 2016: pp. 11, 18.— Yamasaki 2017: pp. 549, 552, table 21.1.
Condyloderes megastigma Sørensen, Rho & Kim, 2010b: pp. 236–241, figs 2–6, tables 1, 2.—this paper: not Condyloderes megastigma Sørensen, Rho & Kim, 2010b but Condyloderes setoensis Adrianov, Murakami & Shirayama, 2002. Condyloderes megastigma — Sørensen et al. 2010a: p. 53.— Sørensen et al. 2012b: p. 231.—Sørensen & Thormar 2010: p. 273.—Neuhaus 2013: p. 232, 254, 291, table 6.—Neuhaus et al. 2013: p. 129.— Neuhaus et al. 2014: pp. 57, 64.— Neuhaus & Sørensen 2013: p. 73.— Adrianov & Maiorova 2016: pp. 11, 18.— Yamasaki 2016: p. 79.—Neuhaus 2017: p. 147, table 5.
Condyloderes cf. megastigma — Sørensen et al. 2012a: p. 163, table 1.—this paper: not Condyloderes cf. setoensis but Condyloderes sp. 2.
description: Published as part of Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi & Higgins, Robert P., 2019, Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov., pp. 1-91 in Zootaxa 4561 (1) on page 39, DOI: 10.11646/zootaxa.4561.1.1, http://zenodo.org/record/2627892
Condyloderes setoensis Adrianov, Murakami & Shirayama 2002
Creators: name: Neuhaus, Birger
name: Zotto, Matteo Dal
name: Yamasaki, Hiroshi
name: Higgins, Robert P.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy

description: Re-description of Condyloderes setoensis based on type material
(Figs 19–21; Tables 1, 2, 7)
Diagnosis. Neck placids with condyles in one basal row in narrower placids and in three rows in broader placids; each narrower placid with one basal condyle; each broader placid with two condyles each in apical, intermediate, and basal row; midventral placid with three condyles in apical and basal row and two in intermediate row; acicular spine middorsally on segments 1–10, midterminally on segment 11, lateroventrally on segments 1–9, midterminally on segment 11, and laterodorsally on segment 10; cuspidate spine ventrolaterally on segments 2, 5 and 9, and lateral accessorily on segment 8 and sometimes on segment 4; type-5 (?) sensory spot ventromedially on segment 11; type-6 sensory spot ventromedially on segments 1 (slightly more paraventrally) and 2–9, lateroventrally on segment 10, sublaterally on segments 1–3, 5–7 and 9, midlaterally on segments 1 and 2, laterodorsally on segments 2–9, subdorsally on segment 10, and paradorsally on segments 1 (almost subdorsally), 2–9; gland cell outlet ventromedially on segment 10.
Material examined. Three paratypic males (MIMB AVA-CM-JAP-02, AVA-CM-JAP-03, AVA-CM-JAP-04) mounted for light microscopy (Fig. 1; Table 1).
description: Published as part of Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi & Higgins, Robert P., 2019, Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov., pp. 1-91 in Zootaxa 4561 (1) on page 40, DOI: 10.11646/zootaxa.4561.1.1, http://zenodo.org/record/2627892
Condyloderes storchi Higgins, 2004 in Martorelli & Higgins 2004
Creators: name: Neuhaus, Birger
name: Zotto, Matteo Dal
name: Yamasaki, Hiroshi
name: Higgins, Robert P.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Cephalorhyncha
Kinorhyncha
Cyclorhagida
Centroderidae
Condyloderes
Condyloderes storchi

description: Re-description of Condyloderes storchi Higgins, 2004 in Martorelli & Higgins, 2004
(Figs 29–31; Tables 1, 2, 7)
Condyloderes storchi Higgins, 2004 in Martorelli & Higgins, 2004: pp. 86–88, figs 2–5, tables 1, 2.
Condyloderes storchi — Sørensen & Pardos 2008: pp. 53, 54 [species erroneously cited as C. storchi Martorelli & Higgins, 2004].— Shirley 2009: p. 1129 [species erroneously cited as C. storchi Martorelli & Higgins, 2004].— Neuhaus et al. 2010: p. 468.— Sørensen et al. 2010b: pp. 235, 241 [species erroneously cited as C. storchi Martorelli & Higgins, 2004].— Neuhaus 2013, pp. 297, 332, tables 6, 12.— Adrianov & Maiorova 2016: pp. 11, 18 [species erroneously cited as C. storchi Martorelli & Higgins, 2004].
Diagnosis. Neck placids with condyles in one basal row in narrower placids and three rows in broader placids; narrower placids with one basal condyle; broader placids with two condyles each in apical, intermediate, and basal row; midventral placid with three apical and basal condyles and two in intermediate row; acicular spine middorsally on segments 1–10, midterminally on segment 11, lateroventrally on segments 1–9, and laterodorsally on segment 10; cuspidate spine ventrolaterally on segments 5 and 9, lateral accessorily on segments 1, 4 and 8, and middorsally on segments 5 and 7; type-5 (?) sensory spot ventromedially on segment 11; type-6 sensory spot ventromedially on segments 1 (slightly more paraventrally) and 2–9, lateroventrally on segment 10, sublaterally on segments 1–3, 5–7 and 9, midlaterally on segments 1 and 2, laterodorsally on segments 1–9, subdorsally on segment 10, and paradorsally on segments 1 (almost subdorsally), 2, 4–6, 8 and 9.
Material examined. Holotypic male (USNM 1025879) mounted laterally (Fig. 1; Table 1).
description: Published as part of Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi & Higgins, Robert P., 2019, Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov., pp. 1-91 in Zootaxa 4561 (1) on page 56, DOI: 10.11646/zootaxa.4561.1.1, http://zenodo.org/record/2627892
Condyloderes megastigma Sorensen, Rho & Kim 2010
Creators: name: Neuhaus, Birger
name: Zotto, Matteo Dal
name: Yamasaki, Hiroshi
name: Higgins, Robert P.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Cephalorhyncha
Kinorhyncha
Cyclorhagida
Centroderidae
Condyloderes
Condyloderes megastigma

description: Synonymy of C. megastigma with C. setoensis
Condyloderes megastigma differs from C. setoensis in the possession of a lateral accessory cuspidate spine on segment 4 versus the lack of this character in C. setoensis (Tables 2, 7). Female C. megastigma display a paraventral type-1 (?) sensory spot on segments 2–5 (Figs 24A, 25A), but this character is only barely visible in the single female of C. setoensis because of its staining (Fig. 22C). One paratypic male of C. setoensis (AVA-CM- Jap02) exhibits a lateral accessory cuspidate spine on segment 4 on one side only, so the specimen shows a character intermediate between C. megastigma and C. setoensis. This finding means that characters to distinguish C. megastigma from C. setoensis are lacking. Therefore, we herewith conclude that C. megastigma is conspecific with C. setoensis and represents a junior synonym of this species. We also interpret the minor differences in sensory spot and spine occurrence between the populations recorded from relatively close areas in the Korea Strait as variation (comp. also chapters Original material of “ Condyloderes megastigma Sørensen, Rho & Kim, 2010b ” and Additional material of “ Condyloderes megastigma ”; Tables 2, 4, 7).
description: Published as part of Neuhaus, Birger, Zotto, Matteo Dal, Yamasaki, Hiroshi & Higgins, Robert P., 2019, Revision of Condyloderes (Kinorhyncha, Cyclorhagida) including description of Condyloderes shirleyi sp. nov., pp. 1-91 in Zootaxa 4561 (1) on page 82, DOI: 10.11646/zootaxa.4561.1.1, http://zenodo.org/record/2627892
Alloxysta fuscicornis
Creators: name: Ferrer-Suay, Mar
name: Selfa, Jesús
name: Notton, David G.
name: Pujade-Villar, Juli
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Charipidae
Alloxysta
Alloxysta fuscicornis

description: Alloxysta fuscicornis (Hartig, 1841)
Xystus fuscicornis Hartig, 1841: 352.
Allotria ancylocera Cameron, 1886: 85 (synonymized in Evenhuis 1982: 23).
Allotria (Allotria) ancylocera − Dalla Torre & Kieffer 1902: 40.
Charips ancylocera − Hafez 1961: 85.
Alloxysta ancylocera − Andrews 1978: 77.
Type material
Holotype of Allotria ancylocera
♂, with the following labels: “ Holotype ” (round label with red margins), “ancylocera” (handwritten), “Cameron 96 76, Carribber Glen” (handwritten), “ ancylocera Cam. ” (handwritten), “ Holotype ♂ Allotria ancylocera Cam. det. J. Quinlan 1973” (handwritten), “B.M. TYPE HYM. 7.119.”, “ Alloxysta fuscicornis (Hartig, 1841) det. M. Ferrer-Suay 2012”.
Remarks
Allotria ancylocera was synonymized with Alloxysta fuscicornis (Hartig, 1841) by Evenhuis (1982: 23). Its type material has been revised and the validity of this synonymy is confirmed here.
description: Published as part of Ferrer-Suay, Mar, Selfa, Jesús, Notton, David G. & Pujade-Villar, Juli, 2013, Revision of the types of species of Alloxysta described by Cameron and Fergusson (Hymenoptera: Figitidae: Charipinae) and deposited in the Natural History Museum (London), including a key to the fauna of Great Britain, pp. 1-27 in European Journal of Taxonomy 53 on pages 11-12, DOI: 10.5852/ejt.2013.53, http://zenodo.org/record/3822925
Alloxysta macrophadna
Creators: name: Ferrer-Suay, Mar
name: Selfa, Jesús
name: Notton, David G.
name: Pujade-Villar, Juli
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Charipidae
Alloxysta
Alloxysta macrophadna

description: Alloxysta macrophadna (Hartig, 1841)
Xystus macrophadnus Hartig, 1841: 352.
Allotria filicornis Cameron, 1889: 57 (synonymized in Fergusson 1986: 10).
Type material
Lectotype of Allotria filicornis
♂, designated by Quinlan (1978: 125) with the following labels: “ Holotype ” (round label with red margins), “Cladish” (handwritten), “Cameron 96 76 Cladish”, “ filicornis Cam ” (handwritten), “ Holotype of Allotria filicornis Cam. ♂ det. J. Quinlan, 1977”, “B.M. Type Hym. 7.169”, “ Alloxysta macrophadna (Hartig, 1841) ♂ det. Ferrer-Suay, 2012” (white label).
Remarks
Alloxysta filicornis is represented by the holotype male and five paralectotype males at BMNH.According to Quinlan’s notes (1978: 125), there is another conspecific male from England, but it is not considered a paralectotype, because the Cameron’s original description stated that the specimens were collected only in Scotland. This species was synonymized by Fergusson (1986) with A. macrophadna (Hartig, 1841) and the validity of this synonymy is confirmed here.
description: Published as part of Ferrer-Suay, Mar, Selfa, Jesús, Notton, David G. & Pujade-Villar, Juli, 2013, Revision of the types of species of Alloxysta described by Cameron and Fergusson (Hymenoptera: Figitidae: Charipinae) and deposited in the Natural History Museum (London), including a key to the fauna of Great Britain, pp. 1-27 in European Journal of Taxonomy 53 on page 12, DOI: 10.5852/ejt.2013.53, http://zenodo.org/record/3822925
Alloxysta victrix
Creators: name: Ferrer-Suay, Mar
name: Selfa, Jesús
name: Notton, David G.
name: Pujade-Villar, Juli
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Charipidae
Alloxysta
Alloxysta victrix

description: Alloxysta victrix (Westwood, 1833)
Allotria victrix Westwood, 1833: 495.
Allotria curvicornis Cameron, 1883: 366 (synonymized in Fergusson 1986: 11).
Type material
Holotype of Allotria curvicornis
♂, with the following labels: “ Holotype ” (round label with red margins), “ Glen Lyon ” (handwritten), “ Cameron 96-73 Glen Lyon ”, “ curvicornis Cam ” (handwritten), “ Allotria curvicornis Cameron ” (handwritten), “ Holotype Allotria curvicornis C. ♂ det. J. Quinlan, 1977”, “ B.M. Type Hym. 7.168”, “ Alloxysta victrix (Westwood 1833) ♂ det. Ferrer-Suay, 2012” (white label).
Remarks
Alloxysta curvicornis is represented only by the holotype male at BMNH. This species was synonymized with A. victrix (Westwood, 1833) by Fergusson (1986: 11). The type material of this species has been revised and the validity of this synonymy is here confirmed.
description: Published as part of Ferrer-Suay, Mar, Selfa, Jesús, Notton, David G. & Pujade-Villar, Juli, 2013, Revision of the types of species of Alloxysta described by Cameron and Fergusson (Hymenoptera: Figitidae: Charipinae) and deposited in the Natural History Museum (London), including a key to the fauna of Great Britain, pp. 1-27 in European Journal of Taxonomy 53 on pages 20-21, DOI: 10.5852/ejt.2013.53, http://zenodo.org/record/3822925
Gnaphosa wiehlei Schenkel 1963
Creators: name: Marusik, Yuri M.
name: Omelko, Mikhail M.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Gnaphosidae
Gnaphosa
Gnaphosa wiehlei

description: Gnaphosa wiehlei Schenkel, 1963
Figs 10–12, 16–26, Map 1.
Gnaphosa wiehlei Schenkel, 1963: 95, figs 55a–c (♂); Ovtsharenko et al. 1992: 52, figs 171–172, 177–180 (♂♀); Marusik & Logunov 1995: 191, figs 69–70 (♀); Song et al. 1999: 451, figs 262L, Q (♂♀); Song et al. 2004: 126, figs 74A–D (♂♀). Gnaphosa proxima Kulczyński: Marusik & Logunov 1995: 189, figs 47–50 (♂♀).
Material examined. MONGOLIA: Omnogov Aimag 1♂ (IBPN), Mandal-Ovoo Somon, 44o51.56’N 104o05.70’E, 1140 m, 26.05.1997 (Yu.M.Marusik); Bayanhkogor Aimag: 1♂ (IBPN), Bayanlig Somon, Bogd Somon, Ikh-Bogd Mt. Range, Khar-Obot Mt., 44°54’N 100°34’E, 2500 m, 4–6.1997 (Yu.M. Marusik); 1♂ 1♀ (MNHB, Nr. 555 Kaszab), Changaj Gebirge, 120 km W von Somon Zag, 2280 m, 21.06.1966 (Z. Kaszab).
Description. Male described by Schenkel (1963) and female described by Ovtsharenko et al. (1992).
Comment. Male of this species is similar to those of G. koponeni sp. n. and can be recognised by shorter tibial apophysis, smooth retrobasal edge of cymbium, very small or lacking spines on retrolateral side of embolus and strong tooth (spur, Sp) on the prolateral edge of embolic base. Shape of the embolic spur is variable (cf. Figs 16–17, 19, 22–24). Females of this species are distinct, due to the shape of epigyne.
The exact type locality of this species was not accurately recorded and only given as “Tal des Sining ho unterhalb Sining, Kansu” (=Valley of Sining River below Sining, Gansu). Sining refers to the modern city Xining. Distribution. This species is known from Tuva, Mongolia and Gansu (Map 1).
description: Published as part of Marusik, Yuri M. & Omelko, Mikhail M., 2014, A survey of East Palaearctic Gnaphosidae (Araneae). 3. On new and poorly known Gnaphosa Latreille, 1804 in Zootaxa 3894 (1), DOI: 10.11646/zootaxa.3894.1.4, http://zenodo.org/record/251800
Gnaphosa sticta Kulczynski 1908
Creators: name: Marusik, Yuri M.
name: Omelko, Mikhail M.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Gnaphosidae
Gnaphosa
Gnaphosa sticta

description: Gnaphosa sticta Kulczyński, 1908
Figs 13–15, 27–30.
Gnaphosa sticta: Ovtsharenko & Marusik 1988: 208, figs 16–17, 20 (♂♀); Ovtsharenko et al. 1992: 48, figs 163168 (♂♀); Almquist 2006: 404, figs 350a–e (♂♀).
For a complete synonym listing and references see Platnick (2014).
Material examined. FINLAND: 1♂, 2♀ (ZMUT), Utsjoki, Kevo, indoors, 5.09.1973-6.06.1974 (S. Koponen). RUSSIA: Chukotka: 1♀ (ZMMU), 74 km W of Anadyr’ City, Anadyr’ River near Omochi River mouth, 613 m, 64.82N 175.967E, 7-8.2013 (O. Khrulyova).
Description. The species has been described by Almquist (2006).
Comments. Here we provide figures of this species to show the similarity to females of G. koponeni sp. n. Figures of epigynes provided by Ovtsharenko et al. (1992), Almquist (2006) and this study (Figs 13–15, 29–30) differ slightly in shape of lateral pockets, size and direction of glands and shape of receptacles, but all illustrated specimens have the same shaped insemination (copulatory) ducts (Id). Males of G. sticta have a very large median apophysis (Ma).
Distribution. The present records from Chukotka coincide with the previous easternmost known locality of the species, which was in the upper reaches of Bolshaya Osinovaya River. This species is known from Norway to East Chukotka and south to Tuva and Hokkaido.
description: Published as part of Marusik, Yuri M. & Omelko, Mikhail M., 2014, A survey of East Palaearctic Gnaphosidae (Araneae). 3. On new and poorly known Gnaphosa Latreille, 1804 in Zootaxa 3894 (1), DOI: 10.11646/zootaxa.3894.1.4, http://zenodo.org/record/251800