Thiodia ursulana Bidzilya & Blackstein & Budashkin & Šumpich & Tsvetkov 2022, comb. nov.
Creators: name: Bidzilya, Oleksiy
name: Blackstein, Hans
name: Budashkin, Yuriy
name: Šumpich, Jan
name: Tsvetkov, Eugene
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Tortricidae
Thiodia
Thiodia ursulana

description: Thiodia ursulana (Kennel, 1919), comb. nov.
Figs 1–5
Semasia ursulana Kennel, 1919: 85.
Eucosma (Phaneta) ursulana (Kennel, 1919) — Obraztsov 1968: 12.
Eucosma ursulana (Kennel, 1919) — Razowski 1999: 475, 2003: 93.
ursulana [no genus, Eucosmini unplaced]— Gilligan & Wright 2013: 331.
= Thiodia aequilibris Tsvetkov, 2017: 3, syn. nov.
Type material. Lectotype of Semasia ursulana, ♂, here designated, labelled: “ ursulana Kenn. Type”, “Uralsk”, “Lectotypus Semasia ursulana Kennel, 1919, des. O.V. Bidzilya, H. Blackstein, Yu.I. Budashkin, J. Šumpich, E.V. Tsvetkov, 2022” (MfN). Paralectotype, ♂, here designated, labelled: “ursulana n. sp. Kenn. Type. 89.90.”, “Uralsk”, “ Paralectotypus Semasia ursulana Kennel, 1919, des. O.V. Bidzilya, H. Blackstein, Yu.I. Budashkin, J. Šumpich, E.V. Tsvetkov, 2022 (MfN).
Additional material examined. ♂, Indersk, 28.v, ex coll Kröne, gen. slide 18/028, JŠ (HNHM). 1 ♂, [without locality data], “ Semasia ursulana Kenn., N. Filipjev det”, “A. Bang-Haas” (ZIN). 1 ♂, 1 ♀, Russia, Astrakhan Prov., 23 km NE of vill. Khosheutovo, N 47°10’48’’, E 48°01’50’’, 16.v.2018 (Tsvetkov) (RCET); 17 ♂, 6 ♀, Russia, Astrakhan Prov., 18 km of vill. Dosang, N 47°00’28’’, E 48°06’26’’, 12.v.2018 (Tsvetkov) (RCET).
The species was originally described in the genus Semasia Stephens, 1829. Later, Obraztsov (1968:12) transferred it to the subgenus Phaneta Stephens, 1852 of the genus Eucosma Hübner, 1823 based on external morphology of adults. Razowski (1999: 475; 2003: 93) followed Obraztsov (1968), but listed ursulana under „unplaced species“ of Eucosma. He also stated that the species was known only from the original description and that neither the type nor conspecific specimens were found. Gilligan & Wright (2013: 331) placed ursulana in the group of species with unclear generic assignment (“Eucosmini unplaced”), and noticed that the holotype was deposited in MfN (as MNHU).
Semasia ursulana Kennel, 1919 was described from two males collected in the environs of Uralsk (W Kazakhstan) (Kennel 1919: 85, Pl. III, fig. 17). These syntypes, both without abdomens, are kept in the collection of MfN. One of syntypes is in good condition, other with both left fore-and hindwings brocken at base. We here designate the better preserved specimen as lectotype (see above under “Type material” and Fig. 1), and the other (with wings brocken) as paralectotype. Another male from Western Kazakhstan (Indersk) labelled as “ ursulana ” was found by the authors in the collection of HNHM. The specimen (Fig. 2) agrees externally in all details with two type specimens of S. ursulana, so that we have no doubt of their conspecificity.
Thiodia aequilibris was described from the male holotype collected in Atyrau Province and a male paratype from Mangistau Province of W Kazakhstan. The original description is accompanied by the photographs of adults and drawings of the male genitalia (Tsvetkov 2017: 3–4, Pl. 1), which fully agree with S. ursulana. Hence, the following synonymy is proposed Thiodia aequilibris Tsvetkov, 2017, syn. nov. of Thiodia ursulana (Kennel, 1919), comb. nov.
Remarks. The assignment of ursulana to the genus Thiodia and the position of the species within the genus was discussed by Tsvetkov (2017: 4, as T. aequilibris). The species can be clearly separated from its congeners both by the male genitalia (Fig. 4) and by the wing pattern. The female of T. ursulana was illustrated in the original description of T. aequilibris (Tsvetkov 2017: 4, Pl. 1, fig. 2). However, the specimen was later lost, but its genitalia have been described, though not illustrated. Below we provide a drawing and re-description of the female genitalia of T. ursulana based on additional material from Astrakhan Province of Russia.
Female genitalia (Fig. 5). Papillae anales elongate, covered by small bristles. Posterior apophyses nearly equal to anterior apophyses, strongly broadened in posterior 1/2. Lamella postvaginalis short and small, anterior margin with a tiny, sharp central protrusion, posterior margin straight. Lamella antevaginalis small, evenly bent or V-shaped. Antrum small, heart-shaped. Sternum VII trapezoidal with bent sides and rounded angles, two vane-like protrusions of posterior margin vary in length. Ductus bursae membranous, cingulum tapering posteriorly, weakly sclerotised or membranous ventrally, usually obliquely cut anteriorly, covered by sternum VII (in ventral view). Corpus bursa spherical, membranous, with a small horn-like signa.
Distribution. Kazakhstan: Western Kazakhstan, Atyrau and Mangystau Provinces; Russia: Astrakhan Province (new record), Buryatia: Munku-Sardyk Mountain (Filipjev 1929: 6).
description: Published as part of Bidzilya, Oleksiy, Blackstein, Hans, Budashkin, Yuriy, Šumpich, Jan & Tsvetkov, Eugene, 2022, New synonyms and redescription of two little-known Thiodia species (Lepidoptera, Tortricidae): T. ursulana (Kennel, 1919) comb. nov. and T. klapperichi (Razowski, 1967) comb. nov., pp. 458-470 in Zootaxa 5124 (4) on pages 460-462, DOI: 10.11646/zootaxa.5124.4.3, http://zenodo.org/record/6413668