Clivina thenmala Balkenohl 2021, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina thenmala

description: <i>Clivina thenmala</i> sp. nov. urn:lsid:zoobank.org:act: 33C279EC-1465-49E4-9809-2EF2A136A2A0 (Figs 35, 55, 75, 102, 119) ETYMOLOGY. The name is a noun and refersto the type locality, Thenmalain Kerala, Southwest of India. TYPE MATERIAL. Holotype: • &amp;male;, with labels and data: white, black printed “South India: V. 93 Karala State, Quilon distr. Thenmala, T.R.S.N. Coll.” / “Coll. J. Saltin in Coll. CBA 1997 ” (CMBB). Paratypes: • 1 &amp;male;, 2 &amp;female;&amp;female;, 21 specs., same data as holotype (CMBB); • 1 &amp;male;, 1 &amp;female;, INDIA – KERALA THEMALA [sic], 5.5.2005 08°58,0’N, 77°03,5’E leg. M. Halada 100 m (CTAA). DIAGNOSIS. Alarge sized species with smooth lateral margin of the pronotum and elytron, and isodiametric reticulation on intervals five to eight of the elytron. Distinguished from the most similar species <i>C. acutimentum</i> sp. nov. by the bigger size, the lobe of the mentum not sharp angled, and the median tooth of the mentum which is distinctly obtuse angled and not straight. The other species with reticulation on intervals five to eight, <i>C. agumbea</i> sp. nov., shows a pronotum with lateral margin straight and converging in anterior half. In addition, the shape of the lobe of the mentum is very different. DESCRIPTION. <i>Measurements</i>. Holotype: Body length 8.94 mm, width 2.73 mm; ratio length/width of pronotum 0.89; ratio length/width of elytra 1.89. Other material: Body length 8.18-8.94 mm (: = 8.7 mm *), width 2.45-2.73 mm (: = 2.63 mm *), ratio length/width of pronotum 0.88-0.89 (: = 0.89*), ratiolength/widthofelytra 1.86-1.89 (: = 1.88); (*n = 10). <i>Colour</i>: Glossy, laterally and apically semi-opaque. Piceous. Legs dark fuscous, antenna light fuscous. <i>Head</i>: Athird narrower than pronotum. Clypeus slightly excised anteriorly; wing prominent, slightly less protruding as clypeus, separated from clypeus by distinct notch; supraantennal plate moderately convex; clypeus laterally with carina, with double reflexed margin in over two thirds of the specimens, wing amargined, supraantennal plate reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by distinct obtuse notch. Clypeus transverse, slightly convex and glossy, separated from frons by moderately deep furrow. Frons moderately convex, with small pore at middle, with almost parallel lateral carina at posterior eye-level. Clypeus and frons smooth, with minute scattered punctures, separated from wing and supraantennal plate by deep moderately wide furrow continuing posteriorly as wide supraorbital furrow with two supraorbital setigerous punctures at posterior eye level. Neck constriction developed as small step, with row of small punctures at middle, reticulated laterally. Eye hemispherical projecting. Gena small, without projecting carina. Antenna relatively short, reaching over second third of pronotum, antennomeres four to ten of moderate length, shape like a barrel with convex sides, (ratio L/W around 1.74). Labrum nearly straight anteriorly, indistinctly pointed at middle, with isodiametric reticulation, seven setose. Mentum (Fig. 55) with isodiametrically reticulated surface, with lobe slightly hollowed out, shape of lobe like a nearly regular but slightly askew trapezium with distinctly convex anterior margin and obtuseangled latero-anterior angle, lateral and anterior margin carinate, medial angle obtuse but distinct, epilobe of moderate width, developed at anterior margin and medial angles, median tooth wide, nearly as protruding anteriorly as lobe, straight anteriorly, almost with minute knob at middle. <i>Pronotum</i> (Fig. 75): Disk slightly and regularly convex in lateral view, moderately convex in frontal view. Distinctly wider than long. Reflexed lateral margin smooth, regularly curved in anterior half, widest in posterior third; anterior angles not projecting; posterior angle marked by relatively small tooth, tooth much less protruding laterally as convexity of lateral margin. Lateral channel moderately wide, reticulated, distinctly convex to disk. Median line distinct, narrow; anterior transverse line punctured, joining median line, finely joining basal constriction. Surface with few transverse wrinkles, with distinct isodiametric reticulation at base and laterally up to anterior setigerous punctures. <i>Elytron</i>: Disk flattened in anterior half in lateral view; in frontal view regularly convex at middle and distinctly convex laterally. Nearly twice as long as wide, not dilated in apical half with maximum width at middle. Reflexed lateral margin smooth. Scutellar striole distinct, with setigerous tubercle at base of first stria and small tubercle at base of third interval situated down the declivity. Striae moderately deep, punctuate-striate and reticulated, one to three free at base, four to seven joining at humerus, one joining with seven at apex, two to four ending free at apex, five and six joining apically. Intervals moderately convex, eights with short carina at humerus, carina at apex distinct, reaching nearly up to last setigerous puncture. Third interval with four setigerous punctures, first situated at second stria near base, the other three approaching third stria. Intervals on disk with isodiametric reticulation of decreasing intensity towards middle, most distinct on interval eight, very faint on interval five (best visible along striae), intervals four to one glossy. Reticulation distinct in apical third on all intervals. <i>Hind wings</i>: Fully developed. <i>Lower surface</i>: Proepisternum with transverse wrinkles and longitudinal rough wrinkles, in lateral half with distinct isodiametric reticulation. Terminal sternite of abdomen developed apically as flat narrow transverse sulcus. <i>Legs</i>: Legs covered with very fine irregular reticulation. Protibia with three spines of moderate length, with indistinct sulcus anteriorly, movable spur regularly arcuate. <i>Male genitalia</i> (Fig. 102): Median lobe short, distinctly arcuate at base, with nearly straight part before middle, arcuate at middle, apex developed as long straight parallel spatula, ventral rim indistinct, base with slender apophysis, with small group of microtrichia at middle of surface. Endophallus with long row of short trichia. Ventral paramere moderately wide, regularly arcuate, lateral apophysis small, apex with four setae. Dorsal paramere slightly curved in basal part, distinctly curved to apex, at apex with four long setae, one of them situated preapically. <i>Female coxostylus and laterotergite</i> (Fig. 119): Coxostylus slender, moderately curved, dorsally with seven long to medium sized and one distinctly elongated and robust nematiform setae in basal half, ventrally with four long and numerous smaller setae, with two SSOs. Laterotergite with three long and four moderately long setae basally and one isolated seta towards coxostylus. VARIATION. On the clypeus the reflexed lateral margin is doubled in over two thirds of the specimens. The central pore on the frons of the head varies in size. DISTRIBUTION. Known from Thenmala in Kerala, India.
description: Published as part of <i>Balkenohl, Michael, 2021, Belgian Journal of Entomology, pp. 1-83 in Belgian Journal of Entomology 115</i> on pages 58-60, DOI: 10.5281/zenodo.5645953