Derris taiwaniana Z. Q. Song. A. 2022, comb. nov.
Creators: name: Song, Zhuqiu
name: Pan, Bo
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Magnoliopsida
Fabales
Fabaceae
Derris
Derris taiwaniana

description: 2. Derris taiwaniana (Hayata) Z.Q. Song, comb. nov.
≡ Pongamia taiwaniana Hayata (1913: 79) ≡ Millettia taiwaniana (Hayata) Hayata (1920: 22) ≡ Whitfordiodendron taiwanianum (Hayata) Ohwi (1936: 660). Type: — CHINA. Taiwan, Taibei, Sankakuyu, 1902, K. Nagai s.n. (syntype, not seen); CHINA. Taiwan, 13 March, S. Yokoyama 27 (syntype, K001414935!).
= Millettia pachycarpa Bentham (1852: 250), non Derris pachycarpa Merrill (1922: 312). Type: — INDIA. Upper Assam. Jenkins s.n. (Lectotype: K001415586!, designated here; isolectotypes: K001415584!, K001415585!, L0019105!, NY!, P 02141837!, P 02141838!).
= Millettia dunnii Merrill (1918: 139). Type: — CHINA. Guangdong, Boluo, Loh Fau [Luofu] Mountain, in thickets near So Liu Koon, 200 m, 13 August 1917, E. D. Merrill 10861 (holotype: PNH, may be destroyed, photo in A!; isotypes: CAS0007371!, NY00016340!, US 00003978!).
= Millettia fooningensis Hu (1955: 360). Type: — CHINA. Yunnan, Foo-ning [Funing County], 550 m, 11 April 1940, Chi-Wu Wang 88378 (holotype: PE00022404!; isotypes: KUN0754399!, KUN0754400!, KUN0754601!, KUN0754602!, KUN0754603!, KUN0755303!).
Liana large, climbing. Stems cylindric, dark brown, densely scattered with white to brown lenticels, pithy inside; young shoots yellow tomentose, glabrescent when mature. Stipules ovate-triangular, ca. 3.5 × 3.0 mm. Leaves imparipinnate, 13–17-foliolate (usually 13), juvenile at anthesis, reddish when young; rachis 21.8–38.8 cm long, including petiole 8.6– 14.0 cm long; leaflet blades oblanceolate to oblong, 5.8–20.3 × 2.3–6.1 cm (ratio usually 2–4), papery to subleathery, glabrous above, appressed pubescent to densely hairy beneath, acute to acuminate at apex, cuneate to rounded at base; lateral veins 10–16 on each side of midvein, obvious, looped near margin; petiolules ca. 5 mm long; stipels absent. Pseudoracemes 10.9–39.8 cm long, usually inserted in the leafless axil of leaf on the lower part of branchlets of current year; rachis appressed hairy; brachyblasts distinct, wart-like or knob-like, up to 7–11 mm long on the lower part, with 3–5 flowers on the top of each brachyblast; bracts inserted at base of brachyblasts and pedicels; bracteoles 2, inserted at the base of calyx. Flowers pink, pale purplish white to white, ca. 2.1–2.6 cm long; pedicels 0.7–1.2 cm long, appressed tawny hairy; calyx campanulate, dark red, 5-lobed, pubescent outside, with short calyx lobes, the lower lobe longest, ca. 2 mm long, 4 mm wide, the tube ca. 6 mm long; standard blade glabrous, suborbicular, ca. 2.3 × 2.1 cm, reflexed, emarginate at the apex, with a yellow-green patch and a 3.5–4.5 mm long claw; wing blade pubescent on the outer surface at apex, oblong, ca. 1.8 × 0.7 cm, with a 7 mm long claw; keel blade pocketed at side, pubescent on the outer surface at apex, oblong, 1.6 × 0.7 cm, with a 7 mm long claw; stamen 10, monadelphous, 2.2 cm long, with two small openings (fenestrae) at base; floral disk indistinct; ovary appressed hairy, 2.2 cm long, with 6–8 ovules; style inflexed, glabrous; stigma minutely capitate. Pods inflated, slightly to deeply constricted between seeds, 6.0–14.6 × 3.2–4.1 × 3 cm, glabrous when mature, with a distinct beak at apex, densely covered with warts, usually indehiscent; fruiting pedicels 0.9 cm long. Seeds 1–5, large, dark brown, reniform, 2.1–3.3 × 1.8–2.4 cm.
Phenology and habitat: —Flowering from March to June, and fruiting from April to next March. The flowers and leaves develop together. It grows in evergreen forests at elevations of 0–2500 m.
Distribution: — Bhutan, China, India, Laos, Myanmar, Nepal, Thailand and Vietnam (Fig. 5).
Notes: —Transfer of Millettia pachycarpa Bentham (1852) to Derris Lour. would lead to a later homonym because of the existence of Derris pachycarpa Merrill (1922: 312). Pongamia taiwaniana Hayata (1913) is the next earliest legitimate name for this species. Therefore, according to Art. 11.4 of the ICN (Turland et al. 2018), we made the new combination Derris taiwaniana (Hayata) Z.Q. Song based on Pongamia taiwaniana Hayata. This new combination is not threatened by the nomen nudum “ Derris taiwaniana Matsum. ”, which is merely cited as a synonym of Pongamia taiwaniana, and thus, an invalid name (Art. 36.1, Turland et al. 2018).
Additional specimens examined.
BHUTAN. Punakha District: 1350 m, 11 Oct. 1984, I. W. J. Sinclair & D. G. Long 5598 (E, K); Tashigang: 1360 m, 18 Jun. 1979, A. J. C. Grierson & D. G. Long 2039 (E, K), 1050 m, 28 Jun. 1979, A. J. C. Grierson & D. G. Long 2350 (E), 3000 ft, 17 Aug. 1915, R. E. Cooper & A. K. Bulley 4504 (BM, E). CHINA. Chongqing: Banan, 19 Nov. 1940, C. Pei 7722 (NAS); Dazu, 100 m, 12 Jul. 1978, Anonymous 440 (SM); Fengdu, 590 m, 12 May 1964, J. A. Wang & Y. X. Wang 59 (CDBI); Hechuan, 11 May 1959, 1-Section 1586 (CDBI); Nanchuan, 500 m, 30 Oct. 1957, G. F. Li 65000 (IBSC, KUN, NAS), 800 m, 12 Oct. 1995, S. R. Yi 15680 (MO), 750 m, 21 Aug. 1990, Z. Y. Liu 12864 (IMC), 200 m, 20 Aug. 1996, Z. Y. Liu 2029201 (IMC); Rongchang, 27 May 1978, Anonymous 30 (SM); Tongliang, 500 m, 27 Aug. 1978, Anonymous 549 (SM); Wulong, 24 Sep. 1978, Anonymous 1224 (SM); Fujian: Dehua, 1250 m, 5 May 1930, P. C. Tsoong 193 (AU, IBSC, PE); Fuzhou, 2 Jun. 1951, T. J. Liu 112 (FJSI), Huaan, 10 Jun. 1959, S. M. Huang 5118 (IBSC), 710 m, 15 Apr. 1987, W. D. Han 20425 (NF), Huai Pin Hsiang, 16 Apr. 1937, H. Migo s.n. (NAS), 11 Jul. 1937, H. Migo s.n. (NAS); Longyan, 260 m, X. F. Zeng ZXF13580 (CZH); Nanan, 850 m, 31 May 1965, Fujian Exped. 2164 (NAS), Nanjing, 400 m, 26 Feb. 1989, H. B. Chen 2252 (FJSI), 450 m, 10 Apr. 1991, H. B. Chen 2553 (MO), 17 May 1942, J. He 1938 (FJSI, PE); Shanghang, 340 m, 5 Sep. 1983, Meihuashan Exped. 9 (FJSI); Xiamen, 27 Oct. 2019, Z. W. Mao 363 (AU); Yanping, 280 m, 24 May 1980, H. Y. Zou 477 (NF), 280 m, 21 Oct. 1980, H. Y. Zou 858 (NF), 14 May 1905, S. T. Dunn 2564 (IBSC); Yongchun, 702 m, 24 Oct. 2018, X. F. Zeng ZXF41394 (CZH); Yongtai, 203 m, 7 Nov. 2012, Q. Tian et al. TQ02522 (CSH), 230 m, 30 May 1931, Y. Lin 316 (AU, IBSC, PE); Guangdong: Boluo, 125 m, 13 Aug. 1917, C. O. Levine 1371 (A), Feb. 27 1930, H. T. Ho 60155 (IBK, NY, IBSC), 30 Jul. 1930, N. K. Chun 41461 (IBK, IBSC, SYS), 26 Feb. 1930, S. P. Ko 50112 (NY, IBSC); Chaoan, 690 m, 6 Dec. 2009, X. F. Zeng 8719 (CZH), 510 m, 3 May 2009, X. F. Zeng ZXF6573 (CZH); Conghua, 24 Nov. 1958, Sino-Germany Exped. 1158 (IBSC); Dabu, 500 m, 2 Nov. 1996, B. H. Chen et al. 83 (IBSC), 350 m, 11 May 1984, Dabu Exped. 489 (IBSC); Dongguan, 2500 ft, 16–17 Mar. 1932, S. Y. Lau 20102 (A, NY, IBSC, SYS); Gaozhou, 6 May 1929, Y. Tsiang 2163 (IBK, IBSC, SYS); Guangzhou, 27 May 1924, To & Tsang 12170 (A, BM, E, MO, NAS, P, US), 16 May 1935, Y. Tsiang 392 (IBK); Heping, 298 m, 1 Jul. 2002, C. M. Tam Y06622 (JJF, SZG); Heyuan, 14 Apr. 1930, C. L. Tso 21552 (IBSC); Huidong, 24 May 1983, B. Y. Chen et al. 461 (IBSC); Jiaoling, 650 m, 25 May 1957, L. Teng 4898 (IBSC, NAS, PE); Lechang, 450 m, 4 Aug. 1986, B. Y. Chen et al. 2626 (IBK, IBSC), 8 May 1929, C. L. Tso 20268 (NY, IBSC), 8 May 1929, C. L. Tso 20271 (IBSC), 17 May 1934, S. P. Ko 54519 (IBK, IBSC, NAS), 300 m, 24 May 1934, S. P. Kuo 80676 (IBSC), 23 Aug. 1950, T. S. Chu 104 (IBSC); Liannan, 18 May 1951, T. S. Chu 60986 (IBSC); Lianzhou, 27 May 1951, T. S. Chu 730 (IBSC); Longchuan, 24 Jul. 1990, B. H. Chen 992 (IBSC); Longmen, 250 m, 17 May 1986, Nanling Exped. 2155 (IBSC); Maoming, 280 m, 30 Apr. 1957, Zhanjiang Exped. 4087 (IBSC); Nanxiong, 21 Aug. 1985, Z. Y. Li 739 (MO); Qujiang, 14 Jun. 1985, Z. Y. Li 624 (MO); Raoping, 16 Apr. 1931, N. K. Chun 42677 (IBK, IBSC); Ruyuan, 20 Aug. 1935, C. S. Chung 10867 (IBSC), 25 Oct. 1938, S. K. Lau 29096 (IBK, IBSC), 21 Jul. 1941, S. K. Lau 29618 (IBSC); Shenzhen, 9 Dec. 1999, F. W. Xing & Y. X. Zhang 12126 (IBSC), 400 m, 17 Oct. 1992, J. F. Chen 2275 (SZG), 100–200 m, 25 Apr. 2005, S. Z. Zhang et al. 587 (PE, SZG), 100–200 m, 10 May 2005, S. Z. Zhang et al. 1291 (PE, SZG); Shixing, 150 m, 11 Aug. 1958, L. Teng 7166 (IBSC, NAS), 27 Jul. 1985, X. B. Ye 35010 (MO); Wengyuan, 22 Sep. 1933, S. K. Lau 2382 (A, SYS), 23 Nov. 1939, S. K. Lau 25293 (IBK, IBSC); Xinfeng, 1–19 Jun. 1938, Y. W. Taam 859 (P, IBSC); Xinxing, 3 Oct. 1958, Y. S. Lau 2434 (IBSC); Xinyi, 18 Jul. 1931, C. Wang 30979 (IBK, IBSC, SYS), 25 Apr. 1932, C. Wang 32211 (IBK), 29 Nov. 1951, T. S. Chu 1137 (IBSC); Yangchun, 4 Nov. 1935, C. Wang 38641 (IBK, IBSC), 300 m, 6 Sep. 1990, Yunkai Exped. 46 (IBSC); Yangshan, 29 Jul. 1936, L. Teng 206 (IBSC), 400 m, 17 Sep. 1985, Nanling Exped. 1463 (IBSC), Jul. –Sep. 1932, T. M. Tsui 762 (A, MO, NY, IBSC, NAS, PE, SYS); Yingde, 18 Oct. 1931, Anonymous 61400 (IBSC), 11 May 1931, H. Y. Liang 60953 (IBK, IBSC, PE), 14 Jan. 1929, Y. K. Wang 510 (A, IBSC, K, MO, P, PE), 9 Jun. 1985, Z. Y. Li 576 (MO); Yunfou, 10–22 Feb. 1928, Y. Tsiang 1892 (NAS); Yunfu, 26 Mar. 1955, C. Wang & L. Teng 501 (IBSC), 19 Nov. 1934, L. Teng 10116 (IBSC); Zhaoqing, 6 Jun. 1975, G. L. Shi 11457 (IBSC), 9 Aug. 1977, G. L. Shi 13089 (IBSC); Guangxi: Baise, 20 Apr. 1936, Kwangsi Prov. Mus. 11188 (IBSC), 2000 ft, 12 Sep. 1928, R. C. Ching 7359 (A, NY, IBSC, NAS); Bama, 26 Apr. 1957, Y. K. Li P01014 (IBK); Cangwu, 200 m, 21 Jul. 1956, S. H. Chun 9954 (IBSC), 200 m, 21 Jul. 1956, S. H. Chun 9956 (IBK, KUN, LBG); Chongzuo, 251 m, 27 Mar. 2015, Y. T. Peng et al. 451402150327042LY (GXMG); Dahua, 355 m, 23 Apr. 2018, D. X. Nong et al. 451029180423017LY (GXMG); Daxin, 29 Jul. 2016, Y. T. Peng et al. 451424160729013LY (GXMG), 26 Jul. 1958, Z. X. Zhang et al. 3842 (IBK); Debao, 613 m, 26 Jun. 2015, Debao Exped. 451024160626122LY (GXMG, IBK); Donglan, 200 m, 18 Jan. 1958, C. C. Chang 11364 (IBK, IBSC); Du’an, 21 Apr. 1978, Du’an Exped. 4-10-0232 (GXMI), 5 Jul. 1957, Y. K. Li P01613 (IBK); Fengshan, 933 m, 2 Apr. 2013, B. Y. Huang et al. 451223130402046LY (GXMG), 696 m, 25 Aug. 2013, D. X. Nong et al. 451223130825037LY (GXMG), 750 m, 26 Oct. 2012, H. Z. Lv et al. 451223121026048LY (GXMG); Fusui, 24 Apr. 1957, S. H. Chun 12047 (IBK, IBSC, KUN); Guanyang, 209 m, 26 Nov. 2015, Guanyang Exped. 450327151126006LY (GXMG, IBK); Guilin, 28 Oct. 1950, C. S. Chung 808606 (IBK, IBSC, PE); Hechi, 11 Jun. 1928, Anonymous 92059 (IBK); Hengxian, 30 Apr. 1957, Z. Z. Chen 50352 (IBK); Hexian, 200 m, 20 Oct. 1989, Daguishan Exped. 81028 (GXMI); Huanjiang, 500 m, 23 Oct. 1991, Dian-Qian-Gui Exped. 70158 (IBK), 236 m, 20 Jul. 2013, Huangjiang Exped. 451226130720003LY (GXMG, IBK); Jingxi, 26 Jul. 1977, Y. Lin 3-54276 (GXMI); Jinxiu, 24 Feb. 1954, Anonymous 24 (IBK), 18 Apr. 1982, Dayaoshan Exped. 14381 (IBSC), 310 m, 19 Apr. 1982, Dayaoshan Exped. 14411 (MO, IBSC), 500 m, 22 Nov. 1981, Dayaoshan Exped. 811983 (GXMI), 9 May 1929, S. S. Sin 8222 (IBSC), 28 Jun. 1934, S. S. Sin 23356 (IBK, IBSC), 11 Dec. 1958, Y. C. Chen 1163 (IBK); Laibin, 3 Aug. 1977, Y. A. Shi 574-92 (GXMI); Leye, 5 May 1960, Lingle Exped. 33022 (IBK), 1250 m, 14 May 1960, Lingle Exped. 33115 (IBK), 23 Jun. 1960, N. K. Liang 10173 (GXMI), 19 Jun. 1960, N. K. Liang 11115 (GXMI), 882 m, 10 Sep. 2013, X. Y. Huang et al. 451028130910030LY (GXMG); Lingchuan, 615 m, 17 May 2013, Lingchuan Exped. 450323130517016LY (GXMG, IBK); Lingui, 12 Jan. 1953, L. H. Chun 93247 (IBK, MO, IBSC, PE), 189 m, 11 Aug. 2015, Lingui Exped. 450322150811023LY (GXMG), 12 May 1964, X. Z. Zheng 249 (GXMI); Lingyun, 18 Jul. 1933, A. N. Steward & H. C. Cheo 711 (P), 21 Dec. 1958, C. T. Ting & J. Z. Wang 1336 (NAS), 17 Apr. 1957, C. Wang 43053 (IBSC), 8 Dec. 1984, Guangxi Exped. 327 (GXMI), 350 m, 10 Sep. 1989, Huanan Exped. 1180 (IBSC), 450 m, 8 Oct. 1989, Huanan Exped. 2338 (IBSC), 617 m, 21 Mar. 2013, Lingyun Exped. 451027130321036 (GXMG, GXMI), 14 Jul. 1937, S. K. Lau 28639 (IBK, IBSC, KUN, PE); Longan, 24 Apr. 1978, X. X. Chen & Y. P. Huang 2-281 (GXMI); Longlin, 650 m, 24 Jul. 1959, Anonymous 604 (HGAS), 24 Jul. 1959, Anshun Exped. 604 (KUN), 800 m, 5 Nov. 1957, C. C. Chang 10812 (IBK, IBSC, KUN), 1 May 1957, C. F. Liang & T. L. Wu 32033 (IBK, IBSC), 560 m, 14 Oct. 1957, Nanzhidi 4593 (IBK, IBSC), 1000 m, 10 Sep. 1987, S. Q. Tang et al. 130 (IBK); Longsheng, 480 m, 13 May 2014, Longsheng Exped. 450328140513016LY (GXMG), 3 May 1987, X. X. Chen & D. R. Liang 6133 (GXMI); Longzhou, 550 m, 23 Dec. 1957, P. C. Tam 57590 (IBSC); Luocheng, 21 Apr. 1978, Luocheng Exped. 4-1-086 (GXMI), 179 m, 21 Apr. 2013, Luocheng Exped. 451225130421004LY (IBK); Nandan, 2500 m, 22 Jun. 1937, C. Wang 40840 (IBK, IBSC, PE); Nanning, 25 Mar. 1959, D. Fang 1907 (GXMI); Napo, 1400 m, 9 Dec. 1958, C. C. Chang 13506 (IBK, IBSC), 19 Oct. 1962, C. C. Lee 1215 (IBK), 22 Dec. 1958, C. T. Li 602312 (IBK, PE), 1250 m, 4 Jan. 1959, C. T. Li 602533 (IBK), 1200 m, 23 Apr. 1981, D. Fang et al. 22420 (GXMI), 10 Oct. 1935, S. P. Ko 55892 (IBSC); Ningming, 12 Oct. 1958, C. C. Chang 12167 (IBK, IBSC, KUN), 9 Jun. 1959, H. Q. Li 40911 (IBK), 15 Nov. 1959, X. F. Deng 10591 (IBK); Pingguo, 13 Oct. 2015, H. Z. Lv et al. 451023151013061LY (GXMG); Pingle, 22 Dec. 1958, S. Q. Zhong A62834 (KUN, PE); Pingxiang, 377 m, 15 Apr. 2016, Y. Y. Xie et al. 451481160415005LY (GXMG); Qinxian, 330 m, 1 Aug. 1958, C. C. Chen 490 (IBSC); Quanzhou, 344 m, 30 Jan. 2013, Quanzhou Exped. 450324130130027LY (GXMG, IBK); Rongshui, 700 m, 5 Jun. 1959, Liuzhou Exped. 2346 (IBK); Shanglin, 19 Apr. 1978, Shanglin Exped. 2-432 (GXMI), 20 Apr. 1978, Shanglin Exped. 2-439 (GXMI); Shangsi, 13 Jul. 1959, H. Q. Li 40865 (IBK), 30 m, 7 Jan. 1944, S. H. Chun 4288 (IBSC); Tiandeng, 730 m, 29 Apr. 2015, H. Z. Lv et al. 451425150429044LY (GXMG); Tiandong, 17 Sep. 1977, S. G. Wang 3-15226 (GXMI), 146 m, 26 Mar. 2018, Tiandong Exped. 451022180326024LY (GXMG); Tian’e, 23 Aug. 1958, C. T. Li 601326 (IBK, KUN), 17 Jul. 1977, Tiane Exped. 4-6-613 (GXMI); Tianlin, 200 m, 5 Jun. 1958, C. T. Li 600614 (IBK, IBSC, KUN), 16 Jun. 1936, H. Y. Liang 67725 (IBK, IBSC), 600 m, 18 Apr. 1989, Hongshuihe Exped. 89-69 (PE), 320 m, 21 Mar. 2013, Tianlin Exped. 451029130321001 (GXMG), 1072 m, 22 Apr. 2013, Tianlin Exped. 4510291300422040 (GXMG); Tianyang, 21 Nov. 2002, N. Li et al. 11280 (SZG), 676 m, 23 May 2016, Tianyang Exped. 451021160523031LY (GXMG); Tze-an, 1400 ft, 3 Jul. 1928, R. C. Ching 6341 (A, NY, US, IBSC, NAS, PE); Wuming, 21 Apr. 1978, G. G. Xie 2-106 (GXMI); Xingan, 400 m, 16 May 2016, Xingan Exped. 450325160516005LY (GXMG); Yangshuo, 9 May 1954, C. S. Chung 31292 (IBK), 280 m, 1 Nov. 1963, Z. Z. Chen 53271 (IBK); Yao-shan, S. S. Sin 592 (IBSC); Yongfu, 425 m, 25 Mar. 2013, Yongfu Exped. 450326130325030LY (GXMG, IBK); Yulin, 26 May 1959, Y. K. Li 404459 (IBK); Zhaoping, 26 Oct. 1958, Y. K. Li 403556 (IBK, IBSC). Guizhou: Anlong, 1000 m, 12 May 1960, Guizhou Exped. 2393 (NAS, PE), 500 m, 14 May 1960, Guizhou Exped. 2616 (NAS), 1100 m, 9 Jun. 1960, Guizhou Exped. 3185 (HGAS, NAS), 700 m, 16 May 1960, Guizhou Exped. 3449 (HGAS), 1200 m, 15 Jun. 1960, Guizhou Exped. 3534 (HGAS), 1300 m, 20 May 1960, Guizhou Exped. 3646 (HGAS), 900 m, 24 May 1960, Guizhou Exped. 3741 (HGAS), 1100 m, 12 Jun. 1960, Guizhou Exped. 4343 (HGAS), 1700 m, 15 Jun. 1960, Guizhou Exped. 4534 (NAS), 28 Sep. 1985, J. B. Zuo 75-007 (GFS); Bijie, 648 m, 22 Nov. 2016, S. Y. Peng 522401161122009LY (GZTM); Ceheng, 600 m, 3 May 2005, F. C. Wang 637 (HITBC, PE), 4 Nov. 1930, Y. Tsiang 9204 (A, NY, IBSC, NAS), 23 Oct. 1980, Z. L. Ding & H. H. Zhang 25 (GFS); Chishui, 500 m, 10 Apr. 1959, Bijie Exped. 1162 (KUN), 850 m, 25 Sep. 1959, Bijie Exped. 1744 (KUN), 430 m, 27 Jul. 1998, D. X. Wang 2073 (GFS), 28 Mar. 1986, H. H. Zhang 1204 (GFS), 5 Apr. 1986, H. H. Zhang 1322 (GFS), 500 m, 22 Jul. 1995, Z. T. Wang & Z. Y. Cao 57 (PE); Congjiang, 430 m, 7 May 1982, J. M. Yuan 838 (HGAS); Daozhen, 393 m, 13 May 2016, Pucha Sect. 520325160513545LY (GZTM), 650 m, 7 Jun. 2003, Z. Y. Liu et al. 2032994 (IMC); Dejiang, 720 m, Guizhou Uni. Exped. DJ-0135 (GZAC), 493 m, 2 May 2016, M. C. Wang 520381160502097LY (GZTM), 473 m, 21 May 2016, X. G. Lu 522227160521025LY (GZTM); Dushan, 984 m, M. T. An DS-0561 (GZAC), 330 m, 25 Aug. 1930, Y. Tsiang 6679 (A, IBSC, NAS); Guanling, 24 Nov. 1935, S. W. Teng 1675 (IBSC); Guiding, 1908, J. Cavalerie 3340 (K); Huangping, 1299 m, 23 Mar. 2017, M. C. Wang 520402170323407LY (GZTM); Jinping, 9 Jun. 1965, J. M. Wang 1196 (GFS); Kaiyang, 610 m, 23 Sep. 1983, M. Z. Yang 1570 (HGAS), 900 m, 24 Sep. 1983, M. Z. Yang 1664 (HGAS); Libo, 25 Jul. 1959, Libo Exped. 1384 (HGAS, PE), 730 m, 8 May 1981, M. Z. Yang 810274 (HGAS), 350 m, 2 May 1983, X. H. Song 657 (NF), 650 m, 22 May 1983, X. H. Song 857 (NF), 780 m, 1 Oct. 1983, X. H. Song 1111 (NF), 350 m, 2 May 1983, Xiang-Hou Song et al. 657 (K, MO), 650 m, 22 May 1983, Xiang-Hou Song et al. 857 (K), 545 m, 14 May 1982, Y. K. Li 9427 (HGAS, PE); Liping, 350 m, 16 Jun. 1981, D. F. Huang 766 (HGAS, NAS); Luodian, 200 m, 27 Mar. 1960, Guizhou Exped. 44 (HGAS), 180 m, 29 Mar. 1960, Guizhou Exped. 375 (HGAS), 621 m, 2 Nov. 2015, Luodian Exped. 522728151102005LY (GZTM), 1026 m, 23 Oct. 2015, M. C. Wang GYZ20151023017 (GYBG), 600 m, 6 Apr. 1959, S. Guizhou Exped. 257 (HGAS, KUN), 200 m, 8 Apr. 1959, S. Guizhou Exped. 368 (KUN), 10 Apr. 1959, S. Guizhou Exped. 766 (HGAS); Pingtang, 22 Apr. 1965, Anonymous 31 (HGAS), 700 m, 22 Apr. 1965, Anonymous 74 (HGAS); Renhuai, 10 Oct. 1929, P. C. Tsoong 146 (PE); Rongjiang, 480 m, 2 Aug. 1965, Z. P. Jian et al. 51357 (HGAS, KUN); Sandu, 630 m, 29 Aug. 2004, J. Y. Ping 31 (HITBC, PE), 6 Jan. 1960, S. Guizhou Exped. 21 (HGAS), 500– 303 m, 27 Oct. 1980, Y. K. Li et al. 8541 (HGAS, IBSC, KUN); Suiyang, 1123 m, 3 Jun. 2015, M. C. Wang GYZ201506030103 (GYBG); Taijiang, 12 May 2001, M. T. An 210 (GFS); Tongzi, 200 m, 7 Feb. 1931, Y. Tsiang 8054 (A, NY, IBSC, NAS); Wanmo, 28 Jun. 1973, Anonymous 79 (HGAS), 750 m, 21 Jun. 1965, Anonymous 843 (HGAS), 800 m, 11 Jul. 1981, D. J. Liu 129 (HGAS), 400 m, 19 Apr. 1960, Guizhou Exped. 1813 (HGAS), 19 Jun. 1999, M. T. An 991064 (GFS); Weining, 2100 m, 22 Jun. 2014, M. C. Wang YD760622-554 (GYBG); Wudang, 1200 m, 17 Aug. 2014, Wudang Exped. 52011213100403LY (GZTM); Xingren, 14Aug. 1960, Guizhou Exped. 8188 (HGAS, PE); Xingyi, 7 Aug. 1959, Anshun Exped. 876 (KUN), 1141 m, 11 Jul. 2015, C. Y. Deng 522301150711684LY (GZTM), 1050 m, 14 Jul. 1960, Guizhou Exped. 6165 (HGAS), 15 Jul. 1960, Guizhou Exped. 6165 (HGAS), 1300 m, 12 Jul. 1960, Guizhou Exped. 6992 (KUN, NAS), 10 Aug. 1960, Guizhou Exped. 7471 (HGAS), 800–1000 m, 23 Jun. 1917, Handel-Mazzetti 118 (A); Xishui, 850 m, 25 Sep. 1959, Bijie Exped. 1746 (HGAS, KUN); Yanhe, 850 m, 10 Sep. 2003, J. W. Xiao 5834 (HITBC, PE); Zhenan, 1400 m, 15 Oct. 2015, H. Liu 520324151015016LY (GZTM); Zhenning, 1 Mar. 1993, L. Yang 149 (GNUG); Zhijin, 1103 m, 11 Nov. 2015, B. B. Luo 522425151111010LY (GZTM), 14 Aug. 1975, Y. L. Tu 338 (GNUG); Ziyun, 1124 m, 19 Jun. 2017, M. C. Wang GYZ201706190008 (GYBG), 1237 m, 08 Jun. 2017, X. Q. Hou 520425170608351LY (GZTM); Zunyi, 783 m, 17 Mar. 2016, P. F. Zhang 522121160317019LY (GZTM); Hunan: Baojing, 19 Oct. 2012, D. G. Zhang 1019036 (JJU), 3 Jul. 2009, D. G. Zhang 907030535 (JJU), 500 m, 16 Sep. 1958, L. H. Liou 9819 (KUN, IBSC, PE), 320 m, 14 Nov. 2012, X. J. Su & H. B. Li 433125D00181114000 (JJU); Dong’an, 660 m, 7 Oct. 2004, J. K. Liu 767 (PE), 500 m, 15 Jul. 2002, X. L. Yu et al. 2121 (CSFI); Guzhang, 334 m, 7 Oct. 2018, C. B. Ma & L. J. Shi HY20180710-0389 (JJU), 253 m, 13 Apr. 2017, D. G. Zhang & D. F. Zhang LX20170413017-0017 (JJU); Jianghua, 6 Aug. 1958, C. J. Qi 3691 (CSFI); Jiangyong, 800 m, 9 Jul. 1959, P. C. Tam 62239 (IBK), 350 m, 11 Aug. 1997, X. L. Yu 1323 (CSFI), 250 m, 25 Apr. 1984, Y. T. Xiao 40584 (CSFI); Jishou, 18 May 2008, H. S. He 2008051808 (JJU), 193 m, 3 Nov. 2018, S. Xiang JS20180311-0168 (JJU); Longshan, 1 Aug. 2013, Y. Xiao et al. LS-2464 (CSH); Luxi, 15 Aug. 2017, D. G. Zhang ZZ170815852 (JJU); Sangzhi, 200 m, Sep. 1963, C. J. Qi 30324 (CSFI); Shuangpai, 420 m, 30 Aug. 2002, X. L. Yu & Q. H. Xu 1878 (CSFI); Suining, 440 m, 7 Jul. 2013, J. J. Zhou & D. Zhou 13597 (CSFI); Tongdao, 250 m, 13 Sep. 1984, K. W. Liu 32815 (CSFI), 29 May 1988, T. C. Chen 580 (IBSC); Yanling, 550 m, Nov. 1988, K. W. Liu 33867 (CSFI); Yizhang, 14 Mar. 1941, P. H. Liang 83066 (IBK, IBSC), 200 m, 26 Jan. 1942, S. H. Chun 90 (IBSC, PE), 105 m, 12 May 1942, S. H. Chun 1013 (IBK, IBSC), 12 Jul. 1942, S. H. Chun 1871 (IBK, IBSC); Yongshun, 220–270 m, 2 May 1988, Beijing Exped. 778 (PE), 16 Oct. 1990, D. G. Zhang 4443 (JJU), 24 Jul. 1959, S. C. Lee 204581 (IBK, IBSC); Jiangxi: Dayu, 600 m, 10 Jul. 1965, Anonymous 9450 (KUN); Xunwu, 260 m, 3 Oct. 2016, X. F. Zeng ZXF22192 (CZH), 305 m, 7 Jan. 2017, X. F. Zeng ZXF24228 (CZH); Sichuan: Anyue, 490 m, 24 May 1978, Anonymous 1180 (SM); Changning, 820 m, 20 May 1959, Anonymous 560 (PE); Dechang, 1800 m, 15 May 1959, S. G. Wu 870 (CDBI, KUN); Ebian, 28 Oct. 1938, T. N. Liou 12666 (PE); Emeishan, 22 Jun. 1931, C. L. Zhou 6289 (HGAS), 1 Sep. 1939, C. W. Yao 4942 (PE), 1 Sep. 1939, C. W. Yao 4949 (NAS), 30 Sep. 1938, Cheo & Tsu 726 (NAS), 600 m, 31 Aug. 1957, G. H. Yang 57036 (IBSC, KUN, NAS, PE), 600 m, 26 May 1995, H. G. Xu 1995434 (MO), 13 Oct. 1952, J. H. Xiong et al. 33133 (IBK, IBSC, NAS, PE), 18 Oct. 1952, J. H. Xiong et al. 33274 (IBK), 900 m, 9 Oct. 1957, K. C. Liu 3132 (CDBI), 6 Oct. 1939, T. N. Liou & C. Wang 1335 (PE, WUK), 7 Jun. 1941, W. P. Fang 17478 (PE), 480 m, Nov. 1941, W. P. Fang 18038 (IBSC, JXU), 9 Aug. 1935, X. B. Zhang 319 (NAS), 750 m, Nov. 1928, Y. Chen 7570 (NAS, NF), 27 Jul. 1935, Y. Y. Ho 5496 (NAS), 30 Nov. 1946, W. K. Hu 9167 (E, US), 22 Jan. 1929, W. P. Fang 6393 (K, SYS); Gulin, 800 m, 2 Jul. 1976, Anonymous 279 (SM); Huili, 1 Apr. 1914, C. Schneider 627 (K), 1550 m, 1 Apr. 1914, Handel-Mazzetti 1064 (E); Junlian, 500 m, 22 Jul. 1977, Anonymous 1075 (SM); Leibo, 29 May 1959, Anonymous 443 (PE); Leshan, 200 m, 3 Sep. 1959, C. T. Kuan 6123 (PE), 400 m, 19 Nov. 1934, T. T. Yü 4368 (K), 400 m, 19 Nov. 1934, T. T. Yü 4368 (PE), 15 Sep. 1981, Z. X. Li et al. 7 (CDBI); Mabian, 8 May 1931, F. T. Wang 22823 (PE), 700 m, 2 Apr. 2000, H. G. Xu 240043 (MO), 600 m, 25 May 1978, Mabian Sect. 179 (SM); Miyi, 7 Jul. 1958, S. Y. Chen et al. 10649 (NAS, SM); Miyi, 1500 m, 22 Aug. 1958, Z. He et al. 11370 (NAS, SM); Muchuan, 800 m, 15 Aug. 1999, Y. F. Xu 180050 (MO); Nanchong, 500 m, 29 Apr. 1958, Nanchong Exped. 133 (CDBI, KUN, PE); Pingshan, 800 m, 1 Jun. 1959, Anonymous 743 (PE), 770 m, 6 Jun. 1934, T. T. Yü 1919 (K), 600 m, 6 Jun. 1934, T. T. Yü 2939 (PE), 600 m, 6 Jun. 1934, T. T. Yü 2939 (K), 13 May 1930, W. P. Fang 170 (A, E, K, NY, US, IBSC, PE); Weiyuan, 400 m, 18 May 1978, Weiyuan Sect. 77 (SM); Xinjin, 600 m, 22 Jul. 1938, T. P. Wang 8291 (WUK, PE); Xuyong, 475 m, 15 May 1959, Luzhou Exped. 797 (CDBI), 22 Aug. 2013, W. B. Ju & X. N. Deng HGX12364 (CDBI), 470 m, 15 Mar. 2013, X. F. Gao et al. HGX11278 (CDBI); Ya’an, E. H. Wilson 4608 (A, K), 1000 m, 2 Sep. 1999, Y. F. Xu 190020 (MO); Yingjing, 23 Nov. 1951, W. G. Hu & Z. He 12093 (PE); Taiwan: Pingdong, 1 Jan. 1917, Matuda Eizi 1056 (TAI); Taibei, 11 Oct. 1918, E. H. Wilson 10783 (A), 300–500 m, 28 Dec. 1984, S. F. Huang 2303 (TAI); Tainan, 700–800 m, 11 Dec. 1992, H. F. Yen 7197 (MO); Taizhong, 500–600 m, 21 Sep. 1990, T. C. Huang 14784 (TAI); Taoyuan, 520 m, 16 Sep. 1996, C. M. Wang & H. M. Lin 2191 (MO), 500–800 m, 9 Apr. 1990, J. M. Hu & K. C. Yang 748 (A, MO), 2030 m, 28 Mar. 1997, M. F. Loa 193 (PE); Xinzhu, 1000 m, 17 Jan. 1992, C. C. Liao & C. C. Wang 146 (A, MO), 565 m, 13 Jan. 2003, C. M. Wang 6470 (MO, PE), 30 Dec. 1932, S. Sasaki s.n. (MO); 13 Mar., S. Yokoyama 27 (K); Xizang: Chayu, 1500 m, 13 Jul. 1980, Z. C. Ni et al. 602 (XZ); Motuo, 650 m, 14 Sep. 1974, Anonymous 2530 (PE), 1100 m, 12 Apr. 1983, B. S. Li & S. Z. Cheng 4037 (PE), 800 m, 12 Oct. 1992, H. Sun et al. ETM-0389 (KUN), 1400 m, 12 Nov. 1992, H. Sun et al. ETM-1252 (KUN), 1000 m, 25 Feb. 1993, H. Sun et al. ETM-3965 (KUN), 1400 m, 25 Feb. 1993, H. Sun et al. ETM-4019 (KUN), 1400 m, 16 Oct. 2015, J. Hu et al. HJ05282 (NAS), 1000 m, 6 Aug. 1974, Kezi Exped. 1656 (KUN, PE), 800 m, 11 Aug. 1974, Qinghai-Xizang Exped. 74-4197 (PE); Yunnan: Baoshan, Apr. 1914, G. Forrest 12353 (BM, E, K, IBSC); 7000–8000 ft, Apr. 1921, G. Forrest 19371 (A, E, K); Bijiang, 1200 m, 15 Jul. 1978, Bijiang Exped. 1286 (KUN); Cangning, 1330 m, 28 Sep. 1988, Anonymous 1039 (KUN); Cangyuan, 1700 m, 28 Mar. 2005, S. S. Zhou 2527 (HITBC); Daguan, 700 m, 21 Jun. 1973, B. Y. Sun et al. 738 (KUN, PE); Eshan, 1500 m, 7 Jul. 1989, Yuxi Exped. 89-295 (KUN); Fengqing, 1900 m, 4 Oct. 1958, J. Chen 128 (KUN), 1200 m, 21 Nov. 1977, Q. Lin 7730 (KUN), 1800 m, 13 May 1938, T. T. Yü 15859 (KUN); Fugong, 1300 m, 2 Jun. 1978, Bijiang Exped. 342 (KUN); Fugong, 5 Nov. 2011, S. S. Zhou 11037 (HITBC); Funing, 1000 m, 1 Nov. 1964, Q. A. Wu 9696 (KUN), 14 Jun. 1964, S. Z. Wang 971 (KUN); Gongshan, 1600 m, 30 May 1950, Anonymous 10044 (KUN), 1400 m, 11 Jul. 1979, Q. Lin & X. F. Deng 790826 (KUN), 1300 m, 2 Oct. 1965, 65-Wenshan Exped. 16 (KUN); Guangnan, 1100 m, 14 Jun. 1964, S. Z. Wang 1063 (KUN); Hekou, 23 Dec. 1974, Hekou Exped. 93 (KUN, PE); Huaning, 1507 m, 24 Dec. 2012, C. Liu et al. 12CS4767 (KUN); Jingdong, 2300 m, 4 Nov. 1956, P. Y. Chiu 53427 (KUN), 1200 m, 8 Nov. 1963, Z. H. Yang et al. 101483 (KUN), 1000 m, Aug. 1936, C. W. Wang 76363 (A, IBSC, NAS, PE), 900 m, 18 Aug. 1977, G. D. Tao et al. 17659 (KUN, HITBC), 9 Apr. 2011, S. S. Zhou 8684 (HITBC); Lincang, 1300 m, 23 Aug. 1957, S. S. Sin 270 (KUN, IBSC, PE); Lushui, 840 m, 2 Apr. 1957, W. X. Liu 133 (KUN, PE), 150 m, 7 Arp. 1957, W. X. Liu 165 (KUN, PE); Menghai, 1000 m, 4 Apr. 1996, B. G. Li & L. Sun 960268 (HITBC), 1300 m, 9 Mar. 1957, Sino-USSR Exped. 7151 (KUN); Mengla, 300 m, 12 Apr. 1955, K. M. Feng 20248 (KUN, IBSC, PE), 620 m, 19 Sep. 1961, Y. H. Li 3571 (KUN, HITBC), 900–1000 m, 2 Nov. 1974, Z. H. Yang et al. 12131 (HITBC); Mengzi, 6000 ft, A. Henry 10521 (A, K, MO, NY, US); Mojiang, 1717 m, 2 Apr. 2021, Z. Q. Song 2021023 (IBSC), 1697 m, 2 Apr. 2021, Z. Q. Song 2021027 (IBSC); Nanjian, 1390 m, 13 Jun. 2012, Z. F. Xu et al. 3732 (KUN); N’mai kha-Salwin divide, 26°N, 7000–8000 ft, May 1919, G. Forrest 17858 (E, K); Pingbian, 760 m, 21 Apr. 1954, P. I. Mao 3943 (WUK, PE); Pu’er, 4500 ft, 16 Apr., A. Henry 13000 (A, E, K, MO, NY), 4000 ft, A. Henry 13530 (A, K, K, MO, NY, US), 1500 m, 5 Apr. 2000, H. Wang 3890 (HITBC), 1000 m, 2 Oct. 1955, Kunming Station 6384 (KUN); Shizong, 12 Nov. 1984, H. Sun et al. 53 (KUN), 800 m, 15 Nov. 1984, H. Sun et al. 137 (KUN), 970 m, 17 Aug. 1985, S. C. Ho 85326 (IBSC), 800 m, 3 Jul. 1987, Xiangke Exped. 870381 (KUN); Shuangbai, 1750 m, 19 Apr. 1957, W. Q. Yin 704 (KUN, WUK); Shuangjiang, 1600 m, 19 Sep. 1957, S. S. Sin 1045 (KUN, IBSC); Shweli-Salwin divide, 25°20’ N., 7000–8000 ft, Aug. 1912, G. Forrest 9030 (E, PE), 8000–9000 ft, Jun. 1919, G. Forrest 17891 (E); Tengchong, 5000 ft, May 1912, G. Forrest 7576 (E, K, PE), 5000 ft, Jul. 1912, G. Forrest 8570 (BM, E, K, PE); Jun. 1913, G. Forrest 12031 (E), 5000 ft, Jun. 1913, G. Forrest 12081 (BM, K), 1580 m, 5 Oct. 1977, P. H. Yu 20227 (HITBC); Weishan, Weishan Exped. 5329271085 (IMDY); Wenshan, 1350 m, 5 May 1962, K. M. Feng 22450 (KUN, WUK); Yangbi, 29 Mar. 1933, H. D. McLaren C176 (A, BM, E), 7 Nov. 1941, T. N. Liou 22611 (PE), 6 Mar. 1988, Y. H. Li et al. 541 (KUN); Yanjin, 560 m, 1 Sep. 1964, N. E. Yunnan Exped. 968 (KUN); Yiliang, 450 m, 24 Sep. 1972, N. E. Yunnan Exped. 974 (KUN, PE); Yongde, 840 m, 6 Jul. 2005, E. D. Liu et al. 1090 (KUN); Yongping, 1500 m, 20 Oct. 1985, Q. Su 309 (KUN); Yuanjiang, 1380 m, 6 Nov. 1964, Y. H. Li 5894 (KUN, HITBC), 1500 m, 4 Jul. 2012, Yuanjiang Exped. 5304280853 (IMDY); Zhenxiong, 550 m, 8 Sep. 1987, Xiangke Exped. 870596 (KUN); Zhejiang: Taishun, 250 m, 21 Apr. 1990, Li et al. 428 (ZM). INDIA. Assam: Cachar, R. L. Keenan s.n. (K); Grwhatly, 29 Mar. 1886, C. B. Clarke 43242D (BM); Jeypore, 1898, Prain Collector s.n. (A); Meghalaya: Khasia Hills, 2000 ft, 2 Apr. 1886, C. B. Clarke 43280A (BM); Griffith Kew Dist. 1767 (K, P); Nagaland: Naga Hills, 5000 ft, Dec. 1907, A. E. Meebold 7123 (K, K), 7000 ft, 4 Sep. 1950, W. N. Koelz 26069 (L); Kala Naga Hills, 4000 ft, May 1882, G. Watt 7257 (E, K, L); Sikkim: 1–3000 ft, J. D. Hooker s.n. (A, BM, K, L, P); West Bengal: Choowbultee, Darjeeling, 12 Jun. 1870, C. B. Clarke 12274 (BM); Darjeeling, 2500 ft, May 1875, J. S. Gamble 2102A (K); Mangbar, 3000 ft, Apr. 1904, H. H. Haines 2029 (E, K); Selimpahar, 3000 ft, 30 Jun. 1876, J. S. Gamble 765A (K); Swoke, 11 Arp. 1910, Ribu & Rhomoo 4081 (L); Tista, 1000 ft, 10 Apr. 1923, G. H. Cave s.n. (A, E); W. Duars, 11 Feb. 1879, J. S. Gamble 6669B (K). LAOS. Phongsali: Phou Den Dinh Phinh Ha, 27 Apr. 1936, M. Poilane 25904 (P); Xieng Khouang: Phou Khe, 1500 m, 3 Apr. 1952, J. E. Vidal 1479 (P). MYANMAR. Chin State: Natma Taung National Park, 887 m, 12 Mar. 2014, P. Srisanga et al. 97919 (E); Kachin State: Myitkyina District, 3100 ft, 11 Apr. 1927, Maung Mya 5365 (K); Myitkyina District, 4000 ft, 3 Apr. 1919, R. Farrer 844 (E); Nansau, 500 ft, Apr. 1911, M. Kyaw 41 (E); Mandalay Region: Maymyo, 3700 ft, 18 Apr. 1917, C. G. Rogers 881 (E), 3500 ft, 13 Jul. 1908, J. H. Lace 3289 (E, K); J. H. Lace 5821 (E, K), 3500 ft, 22 Apr. 1913, J. H. Lace 6162 (A, E, E, K); Tanintharyi: Tenasserim, 4000 ft, 14 Apr. 1877, G. Gallatly 725 (BM). NEPAL. Morang District: 26.48N, 87.55E, 1000 ft, 20 Apr. 1971, J. D. A. Stainton 6805 (BM). THAILAND. Chiang Mai: Chiang Dao District, 26 Sep. 1971, J. E. Vidal 5189 (P); Doi Phra Kao, 1470 m, 1 Sep. 1931, H. B. G. Garrett 714 (K, L); Mae Rim District, 1300 m, 16 Jun. 2009, J. F. Maxwell 09-163 (L); Mae Wang District, 1275 m, 20 Mar. 2003, J. F. Maxwell 03-46 (A, L); Me Chem, 900 m, 12 May 1921, A. F. G. Kerr 5423 (BM, K, TCD); Me Kong, 1190 m, 2 Apr. 1925, Winit 1318 (K); Muang District, 1610 m, 15 Sep. 1989, J. F. Maxwell 89-1087 (A, E, L, MO), 1600 m, 22 Mar. 1991, J. F. Maxwell 91-289 (A, E, L, P); Mueang Chiang Mai District, 5200 ft, 9 Apr. 1911, A. F. G. Kerr 1770 (BM, K, TCD); Sameung, 1350 m, 20 Jul. 1997, J. F. Maxwell 97-787 (A, L, KUN); Chiang Rai: Mae Sai District, 550 m, 20 Aug. 2004, I. C. Nielsen et al. 1856 (L, P); Phan District, 575 m, 10 Sep. 1997, J. F. Maxwell 97-1007 (A, L); Lampang: Mueang Pan District, 550 m, 4 Dec. 1995, J. F. Maxwell 95-1303 (L); Mae Hong Son: Pai District, 1450 m, 6 Apr. 1990, J. F. Maxwell 90-396 (A, E, L, MO); Phayao: Mueang Phayao District, 650 m, 7 May 1997, J. F. Maxwell 97-483 (A, L); Ranong: Mueang Ranong District, 10 m, 22 Mar. 1993, Sino-Thai Botanical Exped. 1335 (KUN). VIETNAM. Cao Bang: Cho Ra, Phan Ke Loc P7052 (HN); Lai Chau: Phong Tho, 600 m, 6 Apr. 1936, M. Poilane 25533 (P); Lang Son: Huu Lung District, 100 m, D. K. Harder & P. K. Loc 4089 (MO, PE); Ninh Binh: Cuc Phuong, 12 Jul. 1998, Chin Cuong Ha 98798 (CPNP).
description: Published as part of Song, Zhuqiu & Pan, Bo, 2022, Transfer of Millettia pachycarpa and M. entadoides to Derris (Fabaceae), supported by morphological and molecular data, pp. 230-248 in Phytotaxa 531 (3) on pages 238-239, DOI: 10.11646/phytotaxa.531.3.4, http://zenodo.org/record/5886324