Calvariomorphus malayanus Ruta & Yoshitomi 2022, sp. nov.
Creators: name: Ruta, Rafał
name: Yoshitomi, Hiroyuki
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scirtidae
Calvariomorphus
Calvariomorphus malayanus

description: Calvariomorphus malayanus sp. nov.
(Figs 1A–B, 2C–E, 3, 4 A–B, 5A–B, E, 7A, 8, 9)
Type material. Holotype, male (ZMUC): “ SINGAPORE: \ Ubin Island \ 1°24’N 103°58’E \ 3-5.xi.1991. O. Martin leg. \ Zoologisk Museum, København”. Paratypes, 2 males, 1 female (ZMUC): same data as holotype. Paratype, male (MNHN): “COLL. Comtesse de BÉARN \ CROISIÈRE DU “NIRVANA” \ SINGAPORE \ E. CORDIER 7.IV.1908 ”; “MUSEUM PARIS \ Comtesse de BÉARN 1909”; “Dascillide”; “? Cyphon \ sp” [handwritten by Pic]. Paratype, male (BPBM): “MALAYA: Kuala \ Lumpur, 19km S of \ Subang, 23.XII.58”; “ T. C. Maa \ Collection \ BISHOP”. Paratype, 5 males (ZMHB): “Hist.-Coll. (Coleoptera) \ Nr. 49600 \ Cyphon spec. \ Bintang, Roettger \ Zool. Mus. Berlin”. Paratype, male (MHNG): “BORNEO—Sabah \ Sepilok \ IV-V.82, M. Horak ”.
Diagnosis. Small species (TL ~ 1.6–1.9 mm), body widest in the middle of elytra, trigonium of penis slightly longer than parameroids, females with two excitators shaped as subtle depressions in apical region of each elytron.
Description. Body very small, TL 1.60–1.87 (1.73, n = 4) mm, TL/EW ~ 1.5–1.6 (1.5), oval, brown, legs and antennae paler, yellowish, covered with yellowish semierect setae. Head strongly transverse, punctation sparse, punctures separated by ca. 2 diameters. Pronotum transverse, anterolateral and posterolateral angles right angled, lateral carinae almost straight, punctation sparse, similar to that on head. Angle between pronotum and elytra not marked in dorsal view. Scutellar shield subtriangular, impunctate. Elytra elongate, punctation stronger than on pronotum, punctures separated by ca. 1.5 diameter.
Penis (L 0.35 mm, W 0.10 mm) elongate, symmetrical, pala as long as parameroids, parameroids narrowly triangular, trigonium slightly longer than parameroids, narrow; tegmen (L 0.23 mm, W 0.15 mm) u-shaped, apices of parameres pointed; sternite IX (L 0.20 mm, W 0.12 mm) u-shaped, with subtrapezoidal basal portion; tergite VIII (L 0.29 mm, W 0.25 mm) with parameres as long as apical portion, apical portion covered with microtrichia, apical margin with short setae; tergite IX (L 0.23 mm, W 0.21 mm) with lightly sclerotized, membranous apical portion, apical margin covered with indistinct setae.
Female. Size similar to that in males, TL 1.73–1.83 (1.78, n = 2) mm, TL/EW ~ 1.5. Each elytron with two subtle excitators shaped as depressions in apical portion, each depression with micropores (Fig. 4A, B). Ovipositor long (L 1.37 mm), bursal sclerite elongate (L 0.17 mm).
Distribution. A widely distributed species, known from the Malayan Peninsula, Singapore, Riau Archipelago and Borneo.
Etymology. After Malayan Peninsula, the area where the species occurs.
Remarks. Bintang is sometimes called Bintam or Bintan; the area in the Riau Archipelago, close to Singapore (Kuijten 1992).
description: Published as part of Ruta, Rafał & Yoshitomi, Hiroyuki, 2022, Calvariomorphus-a new genus of marsh beetles (Coleoptera: Scirtidae) with remarkable elytral excitators, pp. 65-82 in Zootaxa 5120 (1) on page 73, DOI: 10.11646/zootaxa.5120.1.4, http://zenodo.org/record/6388863