Phylloicus elegans Hogue and Denning
Creators: name: PRATHER, AYSHA L.
Publisher: name: Zenodo
Year: 2003
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Calamoceratidae
Phylloicus
Phylloicus elegans

description: Phylloicus elegans Hogue and Denning
Figs. 43, 44, 109
Phylloicus elegans Hogue and Denning, 1983 in Denning et al. 1983:184 [Type locality: Panama, Canal Zone, Barro Colorado Island; WSU; male]. — Flint 1991:98 [male, distribution].
The wing pattern of P. elegans (Fig. 109) is identical to that of P. lituratus. However, the male abdomen of P. elegans is very distinctive: tergum IV has distinctively shaped posteriorly projecting lateral sclerites and multilobed lateral coremata (Fig. 43F). The females of the two species are indistinguishable, and as several series from different localities contain males of both P. elegans and P. lituratus, all determinations of females are tentative. Hybrids may occur; this is discussed under P. lituratus.
Adult. Forewing length 10.4­12.1 mm, n = 106.
Head chestnut brown. Maxillary palps dark brown. Antenna twice forewing length; chestnut brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax chestnut brown; ventrolateral thorax golden. Femora golden; foretibiae dark brown; mesotibiae dark brown; metatibiae dark brown; foretarsi dark brown; mesotarsi white proximally, dark distally; metatarsi dark brown. Metathoracic leg of male with posterior fringe of long setae, setae dark. Tibial spur formula 2,4,4. Forewing flat; dark brown; with two transverse bands; proximal band ivory, reaching posterior wing margin, at least 1/2 width of wing, an inverted V­shape; distal band ivory, beginning at anterior wing margin, at least 1/2 width of wing; with two basal stripes, ivory (Fig. 109). Hind wing basal brush present in male and female; male brush light brown; female brush dark brown.
Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum IV with paired posterior processes and paired lateral sclerites, lateral coremata; posterior process truncate; lateral sclerite spatulate, directed laterally; lateral coremata with basal globose lobes. Tergum V without sclerotized modifications (Fig. 43F). Sternum VII with short, acute anteromesal process. Sternum VIII enclosing base of elongate sternum IX; posteromesal process notched, notch deep and round (Fig. 43A, C). Tergum IX with mesal ridge extending full length of segment; posterior margin with round narrow mesal projection (Fig. 43B); lateral ridge present; dorsal pleural setae approximately 4, ventral pleural setae approximately 2 (Fig. 43A). Preanal appendage at least 11/2 times length of tergum X, widest apically, setae filamentous, longer than appendage (Fig. 43A, B). Tergum X without basal lobes; basodorsal process short and digitate; basolateral processes short, length less than or equal to diameter; apex, in lateral view, truncate, in dorsal view, entire or notched, notch shallow; with paired dorsolateral ridges (Fig. 43A, B). Harpago rounded; peglike setae many, apical (Fig. 43A, C). Phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped (Fig. 43D, E).
Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII without posterolateral brush; sternum VIII with shallow posteromesal notch, or posterior margin entire; sternum VIII (Fig. 44C). Tergum IX without mesal ridge (Fig. 44B). Sternum IX anterior and posterior lobes darkly sclerotized and striate, with patch of lightly sclerotized cuticle lateral to vaginal opening (Fig. 44A). Tergum X appendage shorter than mesal lobe, base indistinct, apex oblique; mesal lobe lightly sclerotized; digitate lateral processes absent (Fig. 44B). Sternum X with patches of short fine setae posterolaterally to anal opening (Fig. 44A). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite truncate anteriorly, posterolateral projections absent; posterior sclerite triangular; posterior end of spermatheca a sclerotized cone (Fig. 44A).
Material examined. COLOMBIA: Antioquia: Quebrada Honda, 12 km SW Fredonia, 1450 m, 22.ii.1983, Flint — 3 males, 1 female (NMNH); Quebrada La Jimenez, Mun. Sopetran, trap C, 780 m, 22.v.1983, Matthias — 1 male (NMNH); COSTA RICA: Alajuela: 20 km S Upala, 15.vii.1990, Parker — 2 males (EMUS); 6.i.1991, Parker — 1 male (EMUS); 30.v.1991, Parker — 1 male (EMUS); 1­15.vii.1991, Parker — 1 male (EMUS); 1­10.viii.1991, Parker — 1 male (EMUS); 21­31.viii.1991, Parker — 1 male, 1 female (EMUS); unnamed river, Cerro Campana ca. 6 km (air) NW Dos Rios, 10°54'00”N, 85°24'00"W, 640 m, 22­23.vii.1987, Holzenthal, Morse, & Clausen — 1 male (UMSP); Guanacaste: Maritza, Quebrada Marilin, 10°57'04”N, 85°29'24"W, 600 m, 12.i.1992, Sweeney, MacLeod, & Villalobos — 3 males (NMNH); 9.xi.1992, de la Rosa — 1 male (UMSP); Río Los Ahogados, Río Los Ahogados, 11.3 km ENE Quebrada Grande, 10°51'54”N, 85°25'23"W, 470 m, 7.iii.1986, Holzenthal & Fasth — 1 male (UMSP); Parque Nacional Guanacaste, Río Orosí, Estación Pitilla, 10°59'28”N, 85°25'41"W, 700 m, 22­25.v.1990, Holzenthal & Blahnik — 1 male (UMSP); Límon: Río Uatsi, ca. 8 km (air) W Bribri, 09°37'12”N, 82°54'00"W, 60 m, 25.v.1987, Holzenthal, Hamilton, & Heyn — 1 male (UMSP); Puntarenas: Río Singrí, ca 2 km (air) S Finca Helechales, 09°03'25”N, 83°04'55"W, 720 m, 21.ii.1986, Holzenthal, Morse, & Fasth — 1 male (UMSP); San José: Escazu, 11­18.iv.1988, Parker — 2 males (EMUS); 15.iv.1988, Parker — 2 males (EMUS); 15­21.v.1988, Parker — 1 male, 1 female (EMUS); Reserva Biológica Carara, Río Carara, Carara, 09°46'41”N, 84°31'52"W, 200 m, 14.iii.1991, Holzenthal, Muñoz, & Huisman — 1 male (UMSP); Río del Sur, 1.5 km (rd) S Carara, 09°46'08”N, 84°31'52"W, 160 m, 13.iii.1991, Holzenthal, Muñoz, & Huisman — 1 male (UMSP); ECUADOR: Pichincha: Río Palenque Biological Station, Río Palenque, Santo Domingo (47 km), 229 m, 29.vii.1976, Cohen — 1 male (NMNH); NICARAGUA: Chontales: Rt. 7, km 206 E of Villa Somosa, 29.vii.1967, Flint & Ortiz — 1 male (NMNH); Jinotega: Cerro Mazú, 14°33'00”N, 85°07'00"W, 220 m, 7­10.ix.1997, Maes & Hernández — 2 males, 1 female (UMSP); Zelaya: Cerro Saslaya, 13°44'00”N, 85°01'00"W, 700 m, 1.iv.1996, Maes & Hernández — 1 male, 1 female (NMNH); PANAMA: Darien: Río Tacarcuna, 579 m, 11.vii.1963, Fairchild — 1 male (MCZ); Panama: Canal Zone, Barro Colorado Island, 28­ 30.iv.1964, S & W Duckworth — 1 male (NMNH); 10­17.v.1964, S & W Duckworth — 1 male (NMNH); 21.ii.1967, Akre — 12 male, 21 female paratypes (CAS); 22­24.ii.1967, Akre — holotype male (WSU); 12.iii.1967, Akre — allotype female (WSU); 18.iii.1967, Akre — 10 male, 14 female paratypes (CAS); 24­25.ii.1969, Akre — 8 male, 9 female paratypes (CAS); 12.iii.1969, Akre — 1 male, 2 female paratypes (CAS); 8.v.1977, Silberglied & Aiello — 1 male (NMNH); 25.vi.1978, Silberglied & Aiello — 1 male (NMNH); Snyder­Molino trail, marker 3, 28.ix.1988 ­ 3.i.1989, Wolda 7 males (NMNH); 13.xii.1989 ­ 13.ii.1990, Wolda — 6 males (UMSP); 12.xii.1990 ­ 5.ii.1991, Wolda — 5 males (NMNH); Windowpane trap­4A, 31.i.­5.ii.1986, Wolda — 1 male (NMNH).
Distribution. Colombia, Costa Rica, Ecuador, Nicaragua, Panama.
description: Published as part of PRATHER, AYSHA L., 2003, Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae), pp. 1-214 in Zootaxa 275 (1) on pages 46-48, DOI: 10.11646/zootaxa.275.1.1, http://zenodo.org/record/5019235