Eotrogaspidia adhabar Okayasu, Lelej & Williams 2021, sp. nov.
Creators: name: Okayasu, Juriya
name: Lelej, Arkady S.
name: Williams, Kevin A.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Mutillidae
Eotrogaspidia
Eotrogaspidia adhabar

description: Eotrogaspidia adhabar Okayasu, Lelej & Williams, sp. nov.
(Figs 1A, 1E, 5A, 7A, 8 A–B, 10A, 10E)
Diagnosis. MALE. Clypeus medially gibbous in pentagonal shape (Fig. 5A); scape with one strong longitudinal carina ventrally (Fig. 5A). Metacoxa evenly convex with long sparse setae (Fig. 7A). FEMALE. Unknown.
Description. MALE. Body length: 13.0 mm. Forewing length: 10.9 mm.
Color and setae. Black with mandible medially, premetum, stipes, palpi, scape except apex and pedicel dark brown, apex od scape yellow, tegula brownish black, protibial spur pale brown, meso- and metatibial spurs yellowish, T1–4 and S1–4 yellowish red except medial carina on S1 black, T5–7 and S5–8 brownish black. Frons, vertex and gena entirely covered with sparse long erect silvery setae, dense on antennal scrobe, malar space and clypeus laterally; frons with dense long appressed silvery setae; vertex behind ocellar region with dense short appressed silvery setae; anterior margin of clypeus fringed with long erect golden setae; prementum, stipes and palpi with sparse very short erect pale golden setae; scape with sparse long recumbent and sparse long erect silvery setae; pedicel and F1 with dense short sub-appressed silvery setae; F2–11 with dense very short erect pale golden setae. Mesosoma mostly with sparse erect silvery setae; pronotal collar posteriorly glabrous; dorsum of pronotum, mesopleuron and dorsum of propodeum between sublateral carinae with dense long appressed silvery setae; mesoscutum with sparse short recumbent and sparser long erect black setae; tegula with sparse short recumbent black setae; mesoscutellum with sparse long erect black setae on anterior half, with sparse long erect yellowish setae on posterior half; axilla and metanotal trough with dense short recumbent silvery setae; lateral face of pronotum, metapleuron and propodeum except dorsum with sparse short recumbent silvery setae. Wings pale brown, veins brown, with short erect black, regularly distributed setae. Legs entirely with sparse long erect silvery setae; procoxa with dense short appressed silvery setae; meso- and metatrochanters, femora and tibiae with sparse short appressed silvery setae; tarsi with dense long recumbent silvery setae, ventrally more or less yellowish. T1, T2–5 laterally and S1–8 with sparse long erect white setae; T2–5 medially with sparse long erect golden setae; T6–7 with sparse long recumbent black and sparser long erect white setae; T1–4 with posterior fringe of sparse long golden setae; S2–4 with posterior fringe of sparse long white setae; S5–8 with posterior fringe of sparse long pale golden setae; T2 with lateral golden felt line.
Structure. Head 1.79 × wider than long; lateral margins strongly convergent behind eye; vertex rounded posteriorly; POD/OOD = 0.68; OD:MOD:LOD = 1.00:1.38:1.00; ocellar region slightly convex, each ocellus without posterior carina; antennal tubercle and antennal scrobe with weak continuous dorsal carina not reaching inner eye margin; genal carina absent; frons and gena with minute sparse and large dense punctures; ocellar region and vertex laterally to it with large sparse punctures; vertex posteriorly to ocelli with minute dense and large sparse punctures; post genal bridge without medial projection; hypostomal carina weakly developed. Clypeus medially gibbous in pentagonal shape, laterally flattened; anterior margin medially widely protruding anteriorly; medial protruding part depressed ventrally, laterally not defined by carina; surface of clypeus impunctate, polished, except anterior margin and posterior margin of medial convexity with minute dense punctures. Mandible slender, narrowed to apex, apically bidentate, without subbasal denticle; inner denticle smaller than apex; ventral margin excised into large basal tooth; outer face of mandible with minute dense punctures. Scape bending medially, with one strong carina beneath not widened apically; surface of scape with small dense punctures dorsally, impunctate and polished ventrally; surface of pedicel with minute sparse punctures; F1 compressed; ration between length and width of pedicel, F1 and F2 = 1.00:1.18:3.53:1.59:2.29:1.47; F2–11 almost equal in width; F11 as long as F10, conical; surface of flagellum with minute dense punctures. Prementum flattened, micro-reticulate, with posterior impunctate convex line; stipes with minute sparse punctures.
Mesosoma broadest at mesonotum including tegulae; ratio between head width, humeral width, pronotal width, mesonotal width = 88:75:84:100; humeral angle rounded, with weak humeral carina ventrally; propodeum abruptly convergent posteriorly; mesoscutal width 1.17 × its length, 0.70 × mesonotal width; tegula short rounded, posteriorly reaching mesoscuto-scutellar articulation, 1.38 × longer than wide; mesoscutum distinctly convex; notaulus evenly wide, reaching anterior 3/8 of mesoscutum, subparallel; parapsidal line length 0.2 × mesoscutal length; parascutal carina forming wide, apically rounded, weakly elevated process on postero-lateral angle of mesoscutum; mesoscutellum gibbous with medial longitudinal carina posteriorly elevated; mesopleuron laterally with distinct transverse mesopleural suture, ventrally with precoxal swelling; precoxal ridge absent; pronotal collar anteriorly with minute dense punctures, posteriorly with small sparse punctures; dorsum of pronotum with minute dense and large dense punctures; mesoscutum and mesoscutellum with large confluent punctures; tegula impunctate polished except anterior and inner margins with small sparse punctures; axilla with minute sparse punctures; metascutellum medially with minute sparse punctures, laterally with small confluent punctures; metanotal trough striate; posterior propodeal face reticulate, with sublateral carina, antero-medially with long triangular cell posteriorly forming small process; dorsal and lateral propodeal faces not separated by carina; lateral face of pronotum transversely striate; anterior part of lateral face of mesopleuron, dorsal half of metapleuron and ventral half of lateral propodeal face with minute dense punctures; medial part of lateral face of mesopleuron and ventral face of mesopleuron with large confluent punctures; posterior part of lateral face of mesopleuron and ventral half of metapleuron with minute dense and small sparse punctures; dorsal half of lateral propodeal face reticulate.
Wings well developed; distance between origin of RS on vein SC and the base of stigmatic cell equal to stigmatic cell length and first abscissa RS length.
Procoxa with minute dense punctures; meso- and metacoxae with minute sparse and small sparse punctures, ventro-medially largely impunctate polished; femora and tarsi with minute dense and small sparse punctures; tarsi with small sparse punctures; metacoxa with weak inner carina throughout; tibiae without outer spines; claws simple, without inner denticles.
T1 with distinct dorsum and anterior slope; T1 spiracle weakly tuberculate; T2 evenly convex, 1.33 × wider than long; S1 with straight medial longitudinal carina with its outer margin emarginate medially and elevated posteriorly forming sharp tooth; S2 evenly convex without medial swelling or carina; T1, T2 laterally and S1–2 with large dense punctures; T2 disc with small sparse punctures; T3–7 and S3–8 with small dense punctures except T7 medially with narrow longitudinal impunctate carina not forming tubercle posteriorly; S7 with lateral rounded tubercle; S8 with lateral pointed tubercle reaching posterior 1/2 of sternum. Paramere down-curved posteriorly, without inner protuberance; inner margin of paramere dorsally with row of short erect golden setae; ventral lobe posteriorly projecting, pointed; parapenial lobe large, rounded; cuspis long slender, apex pointed, posteriorly up-curved, clearly surpassing apex of penis valve, ventrally with dense long erect golden setae; digitus S-shaped, posteriorly with dense short erect golden setae on outer face; paracuspis tuberculate with dense long erect golden setae; penis valves slender, weakly asymmetrical, right penis valve slightly longer than left, with posterior ventral hook and subapical short erect setae.
FEMALE. Unknown.
Type material. 3♂. Holotype, ♂, Nepal: BARA DISTRICT: Adhabar, 21.X.1975, S. Takagi leg. [HUM]. Paratypes: India: KERALA: 2♂, Walayar Forests, 700 ft., X.1959, P.S. Nathan [CASC].
Distribution. India: Kerala. Nepal: Bara.
Etymology. This new species is named after the type locality in Nepal. Treat as a noun in apposition.
Remarks. This new species is readily recognized in Eotrogaspidia by having the clypeus gibbous and the metacoxa evenly convex with long sparse setae; in these features, it is similar to Trogaspidia. The genitalia, however, are consistent with other Eotrogaspidia males.
description: Published as part of Okayasu, Juriya, Lelej, Arkady S. & Williams, Kevin A., 2021, Review of Eotrogaspidia Lelej (Hymenoptera: Mutillidae: Trogaspidiini), pp. 56-90 in Zootaxa 4920 (1) on pages 59-62, DOI: 10.11646/zootaxa.4920.1.2, http://zenodo.org/record/4471588