Landouria menorehensis Nurinsiyah & Neiber & Hausdorf 2019, sp. nov.
Creators: name: Nurinsiyah, Ayu Savitri
name: Neiber, Marco T.
name: Hausdorf, Bernhard
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Camaenidae
Landouria
Landouria menorehensis

description: Landouria menorehensis sp. nov.
urn:lsid:zoobank.org:act: E09F6255-8F7A-49E7-A3BC-E5B795EE2015
Figs 47, 55, 71, 88; Tables 3–4
Diagnosis
Landouria menorehensis sp. nov. is characterized by a whitish, globose shell with a distinct brown band above the rounded periphery and a moderately narrow umbilicus.
Etymology
Landouria menorehensis sp. nov. is named after its occurrence in the Menoreh karst area.
Material examined
Holotype
INDONESIA • holotype (det. anat.); Yogyakarta, Kulonprogo, Sibolong, agroforestry; 7°44′52″ S, 110°08′12″ E; 687 m a.s.l.; 8 Jan. 2013; A. Nurinsiyah leg.; measurements: D = 13.4 mm, H = 8.6 mm; MZB 19201.
Paratypes
INDONESIA – Central Java, Yogyakarta • 1 spec.; Kulonprogo, Sibolong, agroforestry along the way to Sibolong Hill; 7°45′00″ S, 110°07′00″ E; 721 m a.s.l.; ZMH 133441 • 2 spec.; same data as for preceding; ZMH 133442 • 1 spec.; same data as for preceding; ZMH 133443 • 12 spec. (det. anat.); Kulonprogo, Sibolong, agroforestry; 7°44′44″ S, 110°08′02″ E; 716 m a.s.l.; MZB 17686 • 9 spec.; same data as for preceding; MZB 17702 • 5 spec.; same data as for preceding; ZMH 133311 • 1 spec. (det. anat.); Kulonprogo, Sibolong, snakefruit plantation; 7°44′44″ S, 110°08′02″ E; 716 m a.s.l.; MZB 17724 • 1 spec.; same data as for preceding; MZB 17725 • 2 spec. (det. anat.); Kulonprogo, Sibolong, near Kiskendo Cave, shrubs; 7°44′50″ S, 110°08′09″ E; 721 m a.s.l.; MZB 17644 • 1 spec.; same data as for preceding; ZMH 133444 • 6 spec.; same data as for holotype; MZB 19202 • 3 spec.; same data as for holotype; ZMH 133439 • 1 spec.; same data as for holotype; ZMH 133440 • 2 spec.; Kulonprogo, Sibolong, agroforestry; 7°44′51″ S, 110°08′12″ E; 673 m a.s.l.; MZB 18039 • 1 spec.; same data as for preceding; ZMH 133445 • 2 spec.; Kulonprogo, Jonggrangan; 7°44′20″ S, 110°08′32″ E; 730 m a.s.l.; ZMH 140034.
Description
SHELL (Fig. 88; Tables 3–4). Slightly depressed conical-globose, with 5.25–5.75 convex whorls; protoconch smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; without scaly processes; whitish with a distinct brown band above periphery; body whorl rounded; aperture almost circular; upper insertion of peristome distinctly descending; peristome expanded, reflexed and thickened; umbilicus slightly eccentric, moderately narrow, comprising 16–24% of shell diameter, hardly obscured by columellar edge.
GENITALIA (Figs 47, 71; Table 4). Atrium short; penis short, distal part cylindrical, proximal part slightly dilated, without penis coecum; membranaceous sheath around penis connected with proximal part of epiphallus by tissue; penial retractor runs from diaphragm to distal part of short epiphallus; flagellum long, broadest in its middle part, almost smooth, without node, tapering towards its proximal end; vas deferens narrow and long; vagina long; oviduct short; peduncle of bursa copulatrix subdivided into a short, broad distal part and a narrow and long proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.
Remarks
Landouria menorehensis sp. nov. shares with L. pakidulan sp. nov. and L. ciliocincta the whitish shell with a brown band at the periphery. For differences, see remarks under L. pakidulan sp. nov.
Distribution
Landouria menorehensis sp. nov. is endemic to the Menoreh karst area (Fig. 55).
description: Published as part of Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, pp. 1-73 in European Journal of Taxonomy 526 on pages 66-67, DOI: 10.5852/ejt.2019.526, http://zenodo.org/record/3152217