Liogenys cartwrighti Frey 1969
Creators: name: Cherman, Mariana Alejandra
name: Basílio, Daniel Silva
name: Mise, Kleber Makoto
name: Frisch, Johannes
name: Almeida, Lúcia Massutti De
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Melolonthidae
Liogenys
Liogenys cartwrighti

description: Liogenys cartwrighti Frey, 1969
Figs. 2; 26.
Liogenys cartwrighti Frey, 1969: 57; Evans 2003: 207 (checklist); Evans & Smith 2009: 176 (checklist); Krajčík 2012: 144 (checklist); Cherman et al. 2017: 5, 20 (generic history, natural history).
Type material. Liogenys cartwrighti male holotype (USNM): [white handwritten] “Sgo. d. Estero / Argentinien ”, [white, typeset and handwritten] “ Liogenys / cartwrighti / n.sp. / Type / det. G Frey, 1968”, [red typeset] “TYPUS”, [pink typeset] “ Cartwright / Smithson.”, [red, typeset and handwritten] “ Type No / 74011 / USNM”, [white typeset] “ Loan from / USNMNH / 2065060”. Paratypes (2): one female paratype (NHMB) [white handwritten] “ Salta / Dep. Rivadavia / XII.950 Daguerre ”, [orange empty], [white typeset] “ ♀ ”, [pink typeset] “Property / USNM”, [red typeset] “ PARATYPE ”, [white typeset and handwritten] “ Liogenys / cartwrighti / n.sp. / det. G. Frey, 1968”, [white typeset] “Museum Frey / M̹nchen”. One female paratype (NHMB) [white handwritten] “ Salta / Dep. Rivadavia”, [white typeset] “ ♀ ”, [pink typeset] “Cartwrighti / Smithson.”, [red typeset] “ PARATYPE ”, [white, typeset and handwritten] “ Liogenys / cartwrighti / n.sp. / det. G. Frey, 1968”.
Non-type material (12). ARGENTINA: Jujuy: without locality and date, R. Richter 1 male and one female (MLPA); Los Perales, I.1951, F. Justus Jor., 1 female (DZUP); Parque Nacional Calilegua, 29.XI.1986, J. Muzón, 1 male (IADIZA); near Parque Nacional Calilegua, 10.XII.2008, F.C. Ocampo, G. San Blas, & F. Campón, 1 male (IADIZA); Dique la Ciénaga, III.1953, A. Martínez, 1 male (MZSP); Salta: Orán, 11–12.X.1968, L. Peña, 2 males (NHMB); Dique Itiyuro- 5 km Salvador Mazza [22.0509ºS, 63.7101ºW], 15.XI.1995, S. Flores & S. Roig, 1 male (IADIZA); Formosa: Ruta Provincial 39, 23 km S Ingeniero Juárez, 24º05’27’’S 61º56’49’’W, 13–17.XII.2008, F. Ocampo, G. San Blas, & F. Campón, mercury vapor & ultraviolet light trap, 1 male (IADIZA); Tucumán: Lules, XII.1957, A. Martínez, 1 male (MZSP); Santiago del Estero: Ojo de Agua, III.1969, A. Martínez, 1 male (CMNC).
Diagnosis. Body elongate, brown to dark brown, shiny (Fig. 2A); clypeal emargination deep but not reaching the base of the teeth, rounded and wide; lateral margin strongly convex and produced, forming a tooth-like projection; pronotal posterior corners in obtuse angle (Fig. 2B); basal apophysis of metacoxa produced; metatibia weakly or not carinate along the inner margin; superior tooth of a tarsal claw slightly longer than the inferior, distance between teeth shorter than the inferior tooth; pygidium with sparse bristles throughout (Fig. 2 D–E); ventrites with abundant bristles (Fig. 2C); in males protarsomere II as long as it is wide; parameres lyre shaped; ventral surface visible in dorsal view, concave (Fig. 2F).
Redescription. Length 9.4–12.3 mm; width: 4.4–5.9 mm. Dark brown. Head: distance between eyes twice as wide as one eye; clypeal emargination deep, rounded and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye; lateral margin strongly convex and produced, forming a tooth-like projection; distance between clypeal lateral projection and anterior margin of the eye as long as one eye; distance between clypeal lateral projection and apex of the adjacent anterior tooth longer than the basal width of the anterior tooth; acute angle between outer side of anterior teeth and clypeal lateral projection; distal maxillary palpomere, maximum width twice the apical width; fovea shallow, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and longer than the funicle in males, as long as in females. Thorax: anterior margin of pronotum straight; glabrous, punctures fine and very sparse, denser and coarser near the anterior margin; pronotal posterior corners in obtuse angle; hypomere with long bristles, scales near the inner margin; mesepisternum scaly; sides of metaventrite with brisltes and scales; distance between mesocoxae and metacoxae slightly longer than the length of the metacoxa; scutellum ogival, weakly punctate at the sides. Elytra: shiny, glabrous, brown or dark brown; elytral suture and elytron unicolored, weakly elevated; all elytral ridges weakly defined. Legs: three protibial teeth, the middle and apical equal in size, basal one slightly shorter; distance between basal and middle teeth slightly longer than between middle and apical; mesofemural surface setose; mesotibia subquadrate or quadrate in cross-section, surface coarsely sculptured; two transverse carinae, the apical one incomplete in males and complete in females; metacoxa scaly, scales long; basal apophysis of metacoxa produced; metatibia not carinate along the inner margin, apical inner surface setose; metatibial transverse carinae present posteriorly; metatibial apical spurs of different lengths; the larger longer than the diameter of the tibial apex; in males protarsomeres and mesotarsomeres I to IV strongly enlarged; metatarsomeres slightly enlarged; protarsomere II as long as it is wide; metatarsomere I two times shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw slightly longer than the inferior and equal in width, distance between teeth shorter than the inferior tooth. Abdomen: ventrites with abundant bristles on disc; propygidium slightly visible, with bristles; pygidium more or less convex; subquadrate, as wide as it is long; pygidial width equal in distance or slightly wider than the distance between spiracles of propygidium; pygidial disc with sparse bristles throughout; pygidial apex quadrate. Parameres: basal region narrower than both sections of the parameres at its maximum width, parameral split at 2/3, lyre shaped; ventral surface seen from dorsal view, concave; parameres convex in lateral view (Fig. 2G).
Type locality. ARGENTINA, Santiago del Estero.
Geographical distribution. ARGENTINA (Jujuy, Salta, Formosa, Tucumán, Santiago del Estero).
Remarks. Liogenys cartwrighti is similar to L. denticeps Blanchard, 1851 (Fig. 5) in the body size and color and in the clypeus and pygidium in shape. Liogenys cartwrighti differs from L. denticeps (in parenthesis) by the obtuse (rouded) pronotal posterior corners; pronotum and elytra finely punctate (coarsely); pygidium with bristles throughout (glabrous); males with protarsomeres and mesotarsomeres strongly enlarged (slightly wider than the metatarsomeres); and in the different shape of the parameres (Figs. 2F, G; 5K, L). After examining the specimen at NHMB, from which Frey (1969) had drawn the genitalia of L. denticulata Moser, 1918 (sensu Frey 1969), we realized that it is a misidentified specimen of L. cartwrighti. Frey (1969) mentioned “ Salta, Rivadavia” [Argentina] as the type locality of the complete series, and that the type [holotype] is deposited at the USNM. The paratypes examined at NHMB match with this locality, but the holotype at the USNM bears “ Santiago del Estero ” [Argentina]. As this type specimen bears all the original labels of Frey, we confirm that Santiago del Estero is the type locality of L. cartwrighti, and not Salta.
Besides the Chacoan region, L. cartwrighti can be also found at the Yungas Province, in sympatry with L. denticeps and their similar species L. latitarsis Moser, 1918.
description: Published as part of Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, pp. 1-59 in Zootaxa 4938 (1) on pages 6-8, DOI: 10.11646/zootaxa.4938.1.1, http://zenodo.org/record/4561258