Lethades kanagawensis Watanabe, 2024, sp. nov.
Creators: name: Watanabe, Kyohei
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Lethades
Lethades kanagawensis

description: Lethades kanagawensis sp. nov.   (SJN: Kanagawa-hirata-maru-himebachi)  
(Figs. 34A–E, 39L, 40Y, 41I)  
 Type series. Holotype: JAPAN, KPM-NK 81378, F, Kanagawa Pref., Hadano City, Mt. Koubou-yama, 15. IV. 2013, K. Watanabe leg. Paratypes: JAPAN, KPM-NK 84894, F, Tokyo, Akiruno City, Ninomiya, 3. V. 2010, K. Watanabe leg.; KPM-NK 84895, M, Kanagawa Pref., Hadano City, Mt. Koubou-yama, 18. IV. 2010, K. Watanabe leg.; KPM-NK 84896, F, ditto, 18. IV. 2010; KPM-NK 84897, F, Kanagawa Pref., Hiratsuka City, Mt. Komayama to Shonandaira, 21. IV. 2007, K. Watanabe leg.; KPM-NK 84898, 84899, 2 M, Kanagawa Pref., Aikawa Town, Nakatsu, 11. IV. 2014, K. Watanabe leg.; KPM-NK 84900, F, Kanagawa Pref., Atsugi City, Funako, 14. IV. 2007, K. Watanabe leg.; KPM-NK 84901, M, Kanagawa Pref., Atsugi City, Sanda, 26. IV. 2008, H. Katahira leg.; KPM-NK 84902, M, Kanagawa Pref., Atsugi City, Nakaogino, 20. IV. 2008, H. Katahira leg.; KPM-NK 84903, 84904, 2 F, ditto, 26. IV. 2008, M. Gunji leg.   
 Description. Female (n = 7). Body length 4.75–5.8 (HT: 5.7) mm, matt and covered with silver setae.   
Head 0.6 × as long as wide in dorsal view. Clypeus 3.0 × as broad as high, sparsely punctate, polished, lower margin rounded (Fig. 40Y). Face 1.75–2.0 (HT: 2.0) × as broad as high, slightly convex medially in lateral view. Dorsal profile of gena as Fig. 39L. Occipital carina complete. Length of malar space 0.4–0.45 (HT: 0.45) × as long as basal width of mandible. Base of mandible flat. POL 1.5 × as long as OD. OOL 1.9–2.0 (HT: 2.0) × as long as OD. Antenna with 31–34 (HT: 34) flagellomeres. FL I 3.3× as long as maximum depth, 1.2 × as long as FL II.  
Mesosoma densely punctate. Epomia absent. Mesoscutum with notaulus, its posterior end reaching the middle of mesoscutum. Epicnemial carina present laterally and ventrally, its dorsal end situated slightly behind of anterior margin of mesopleuron. Speculum with smooth area. Mesopleuron largely coriaceous around speculum. Propodeum with all carinae except for anterior transverse carina sometimes partly obscured or absent. Fore wing length 4.2–5.0 (HT: 4.8) mm. Areolet present, shortly petiolated, received vein 2m-cu slightly based of the outer angle (Fig. 34C). Fore wing vein 1cu-a postfurcal to vein M&RS. Nervellus inclivous, intercepted posterior the middle. Tarsal claws simple. Hind femur 4.25–4.4 (HT: 4.25) × as long as maximum depth in lateral view. Ratio of length of hind first to fifth tarsomeres 4.8–5.0 (HT: 4.8): 2.0: 1.6: 0.9–1.0 (HT: 1.0): 1.1–1.2 (HT: 1.1). Hind TS III distinctly longer than hind TS V (Fig. 41I).  
Metasoma. T I 1.2–1.3 (HT: 1.2) × as long as maximum width, densely and finely punctate. T II 0.6–0.65 (HT: 0.63) × as long as maximum width, largely densely and finely punctate. Ovipositor sheath 0.25–0.3 (HT: 0.25) × as long as hind tibia.  
Colouration (Figs. 34A–C). Body (excluding wings and legs) black. Mandible except for teeth, ventral surfaces of scape and pedicel, palpi, postero-dorsal corner of pronotum, and tegula yellow. Lower margin of clypeus and ventral surface of flagellum more or less tinged with reddish brown. T II to T IV and ovipositor sheath red to reddish brown. Wings hyaline. Veins and pterostigma blackish brown except for yellow wing base. Legs reddish brown. Trochanter and trochantellus yellow.Apices of hind tibia and hind femur blackish brown. Hind tarsus largely tinged with blackish brown.  
 Male (n = 5). Similar to female. POL 1.5–1.8 × as long as OD. Length of malar space 0.6 × as long as basal width of mandible. FL I 2.85 × as long as maximum depth, 1.25–1.3 × as long as FL II. Hind femur 4.3–4.6 × as long as maximum depth in lateral view. Ratio of length of hind first and second tarsomeres 4.6: 2.0. T II 0.7–0.8 × as long as maximum width. Face, clypeus, malar space, and subtegular ridge yellow (Figs. 34D, E). Fore and mid legs largely yellowish brown (Fig. 34D). Red area of metasomal tergites sometimes partly reduced (especially T II and T IV).   
 Distribution. Japan (Honshu).   
 Bionomics. Host is unknown. Adults usually collected from grassland.   
 Etymology. The specific name is from Kanagawa Prefecture.   
 Remarks. This species resembles Let. nigricoxis Sheng & Sun, 2013 in the following combination of characters: flagellum with more than 30 segments; hind coxa black; ovipositor sheath shorter than hind TS I, but can be distinguished by the following combination of characters: T IV entirely red (basal 0.3 red in Let. nigricoxis); lateral side of T V red (not red in Let. nigricoxis); hind TS III distinctly longer than hind TS V (same length in Let. nigricoxis); face yellow in male (black in male of Let. nigricoxis).
description: Published as part of Watanabe, Kyohei, 2024, Taxonomic Study of Japanese Ctenopelmatinae (Hymenoptera, Ichneumonidae), with Descriptions of 22 New Species, pp. 39-102 in Bulletin of the Kanagawa Prefectural Museum (Natural Science) 53 on page 91, DOI: 10.5281/zenodo.13237363