Hastula H. Adams &amp; A. Adams 1858
Creators: name: Fedosov, Alexander E
name: Malcolm, Gavin
name: Terryn, Yves
name: Gorson, Juliette
name: Modica, Maria Vittoria
name: Holford, Mandë
name: Puillandre, Nicolas
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Neogastropoda
Terebridae
Hastula

description: Genus <i>Hastula</i> H. Adams &amp; A. Adams, 1858 (Fig. 6 A–E) <i>Type species:</i> <i>Buccinum strigilatum</i> Linnaeus, 1758; SD, Cossmann (1896). <i>Synonyms</i>: <i>Impages</i> E.A. Smith, 1873 (type species <i>Terebra caerulescens</i> Lamarck, 1822 = <i>Buccinum hecticum</i> Linnaeus, 1758; SD, Cossmann, 1896); <i>Acuminia</i> Dall, 1908 (type species <i>Buccinum lanceatum</i> Linnaeus, 1767; OD); <i>Hastulina</i> Oyama, 1961 (type species <i>Terebra casta</i> Hinds, 1844; OD); <i>Egentelaria</i> Rehder, 1980 (type species <i>Terebra stylata</i> Hinds, 1844; OD). <i>Definition:</i> Includes all species in clade D of Modica <i>et al.</i> (2019), and those that show a combination of conchological, anatomical and distribution characteristics closely comparable with <i>Hastula strigilata</i> or any genetically proven member of the clade. <i>Diagnosis</i>: Diagnostic nucleotide combinations provided for entire clade with exception of four specimens of <i>H</i>. <i>albula</i> (see Material and Methods) and, separately, for subclade <i>H</i>. <i>albula</i> <i>–</i> <i>H</i>. <i>natalensis</i> <i>–</i> <i>H</i>. aff. <i>casta</i> of clade D (Table 4). <i>Shell</i>: Small to medium-sized, shiny, 12–90 mm. Protoconch paucispiral or multispiral, about 1–5 whorls. Axial sculpture of crenulations on subsutural portion of whorl, or fine ribs on part or throughout whorl height; spiral sculpture of single subsutural groove, occasionally with additional row of punctations. Siphonal canal stout, very short, with well-developed fasciole. Aperture elongate to rounded. Genus Hastula (except four specimens of H. albula) Species/sequences analysed 18/91 Diagnostic nucleotides: 28: A, 67: T, 95: C, 181: T, 221: C, 222: C, 253: A, 581: T, 598: A Subclade H. albula <i>–</i> H. natalensis <i>–</i> H. aff. casta <i>Anatomy</i>: Rhynchodeal introvert, proboscis, venom gland, salivary glands and radula well developed. Radula of hypodermic marginal teeth attached to thin, reduced membrane; walls of marginal teeth often with multiple perforations (Fig. 6E, F) Species/sequences analysed 3/17 Diagnostic nucleotides 49: G, 347: C, 349: T, 517: T <i>Distribution</i>: Indo-Pacific, West Africa, Caribbean. <i>Remarks</i>: Species of <i>Hastula</i> are rather easy to recognize among Terebridae by their typically shiny shells, with fine close-set axial ribs and usually by the lack of a clearly defined subsutural band. The shell morphology in the <i>Partecosta macleani</i> group closely resembles that of <i>Hastula</i>; however, the latter is usually larger at maturity, while shells of comparable size can be differentiated by texture—a polished surface in <i>Hastula vs</i> dull in <i>Partecosta</i>. Furthermore, species of <i>Hastula</i> possess a venom gland and hypodermic radular teeth, whereas <i>Partecosta</i> species lack a venom gland and their radulae teeth are solid. <i>Included species:</i> <i>Hastula aciculina</i> (Lamarck, 1822) 2; <i>H. acumen</i> (Deshayes, 1859) 1; <i>H. alboflava</i> Bratcher, 19882; <i>H. albula</i> (Menke, 1843) 1; <i>H. androyensis</i> Bozzetti, 20082; <i>H. anomala</i> (Gray, 1834) 2; <i>H. anosyana</i> (Bozzetti, 2016) 2; <i>H. apicitincta</i> (G. B. Sowerby III, 1900) 2; <i>H. bacillus</i> (Deshayes, 1859) 2; <i>H. casta</i> (Hinds, 1844) 1; <i>H. celidonota</i> (Melvill &amp; Sykes, 1898) 2; <i>H. cernohorskyi</i> Burch, 19652; <i>H. cinerea</i> (Born, 1778) 1; <i>H. continua</i> Deshayes, 18592; <i>H. crossii</i> (Deshayes, 1859) 1 ∗; <i>H. cuspidata</i> (Hinds, 1844) 2; <i>H. denizi</i> Rolán &amp; Gubbioli, 20002; <i>H. engi</i> Malcolm &amp; Terryn, 20172; <i>H. escondida</i> (Terryn, 2006) 2; <i>H. exacuminata</i> Sacco, 18912; <i>H. filmerae</i> (G. B. Sowerby III, 1906) 2; <i>H. hamamotoi</i> Tsuchida &amp; Tanaka, 19992; <i>H. hastata</i> (Gmelin, 1791) 1; <i>H. hectica</i> (Linnaeus, 1758) 1; <i>H. imitatrix</i> (Auffenberg &amp; Lee, 1988) 2; <i>H. inconstans</i> (Hinds, 1844) 2; <i>H. knockeri</i> (E. A. Smith, 1872) 2; <i>H. lanceata</i> (Linnaeus, 1767) 1; <i>H. leloeuffi</i> Bouchet, 19832; <i>H. lepida</i> (Hinds, 1844) 2; <i>H. marqueti</i> (Aubry, 1994) 2; <i>H. maryleeae</i> R. D. Burch, 19652; <i>H. matheroniana</i> (Deshayes, 1859) 1; <i>H. nana</i> (Deshayes, 1859) 2; <i>H. natalensis</i> (E. A. Smith, 1903) 1 ∗; <i>H. parva</i> (Baird, 1873) 1; <i>H. penicillata</i> (Hinds, 1844) 1; <i>H. philippiana</i> (Deshayes, 1859) 2; <i>H. puella</i> (Thiele, 1925) 1; <i>H. raphanula</i> (Lamarck, 1822) 1; <i>H. rufopunctata</i> (E. A. Smith, 1877) 2; <i>H. salleana</i> (Deshayes, 1859) 1; <i>H. sandrogorii</i> Ryall, Terryn &amp; Rosado, 20172; <i>H. solida</i> (Deshayes, 1857) 1; <i>H. strigilata</i> (Linnaeus, 1758) 1; <i>H. stylata</i> (Hinds, 1844) 1; <i>H. tenuicolorata</i> Bozzetti, 20081; <i>H. tiedemani</i> Burch, 19652; <i>H. venus</i> Aubry, 20082; <i>H. verreauxi</i> (Deshayes, 1857) 1; <i>H. westralica</i> (Aubry, 1999) 2; <i>H. daniae</i> (Aubry, 2008) 3.
description: Published as part of <i>Fedosov, Alexander E, Malcolm, Gavin, Terryn, Yves, Gorson, Juliette, Modica, Maria Vittoria, Holford, Mandë &amp; Puillandre, Nicolas, 2019, Phylogenetic classification of the family Terebridae (Neogastropoda: Conoidea), pp. 359-388 in Journal of Molluscan Studies The Malacological Society of London 85 (4)</i> on pages 12-14, DOI: 10.1093/mollus/eyz004, http://zenodo.org/record/4469844
description: {"references": ["COSSMANN, M. 1896. Essais de paleoconchologie comparee, Vol. 2. Author and Comptoir Geologique, Paris.", "DALL, W. H. 1908. Subdivisions of the Terebridae. Nautilus, 21: 124 - 125.", "OYAMA, K. 1961. On some new facts of the taxonomy of Terebridae. Venus, 21: 176 - 189.", "MODICA, M. V., GORSON, J., FEDOSOV, A. E., MALCOLM, G., TERRYN, Y., PUILLANDRE, N. &amp; HOLFORD, M. 2019. Macroevolutionary analyses suggest environmental factors, not venom apparatus, play key role in Terebridae marine snail diversification. Systematic Biology. DOI: https: // doi. org / 10.1093 / sysbio / syz 059."]}