Hydropsyche semkala Olah & Johanson 2008, new species
Creators: name: Oláh, J.
name: Johanson, K. A.
Publisher: name: Zenodo
Year: 2008
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Hydropsychidae
Hydropsyche
Hydropsyche semkala

description: Hydropsyche semkala Oláh & Johanson, new species   
Fig. 231–235  
 This species is similar to H. klimai Mey from West Indonesia, representing the only two known Hydropsyche having a sclerous, half ring attached to the middle dorsum of the phallotheca. This structure forms a part of the dorsal membranous phallocrypt, a ventral continuation of segment X. Hydropsyche semkala has a short, deep and oblique intersegmental depression. The apicodorsal setose lobes on segment X are not higher than the median keel on segment IX; the apicoventral setose lobes are slender, not robust in dorsal view; and the trilobed apicoventral lobe of phallotheca has lateral lobes that are longer than broad, and a plate-like median lobe.   
Male. Body and wings brown with pale pubescence. Maxillary palp formula I-(III, IV)-II-V, segment V as long as sum of segments I–IV. Head dorsum light, almost yellow, with 9 warts of the same colour, the anteromesal setal wart difficult to discern. Swollen setal wart present on proepisternum. Pretarsal claws asymmetrical, laterally flanked by setal bundle. Spur formula 244. Protibiae with posteroapical spur smaller than anteroapical spur.  
Wings. Venation typical for the genus. Forewing crossveins m-cu and cu separate. Sc and R, and Cu2 and A separate before costa. Hind wing Sc and R meeting before costa about at crossvein r. Stem of M and Cu1 running closely. Median cell closed. Fork 1 present. Forewing length 7.1 mm, hind wing length 5.8 mm.  
Male genitalia. Abdominal segment IX fused annularly, rather long laterally. Median keel long (Fig. 232); dorsal surface very broad, granulose; lateral surfaces flat. Anterior margin deeply arciform; ventrum short; dorsum 2 times longer than ventrum (Fig. 231). Antecosta well developed, widens before margin. External groove of antecostal suture present. Apical lobe on posterolateral margins robust, broad, nearly triangular (Fig. 231). Posterior spine row intermittent, present on segment X and apical lobes, absent between. Depression between segments IX and X very narrow, deep. Body of segment X broadly rounded (Fig. 231), dorsal margin short, ventral margin long (Fig. 231). Lateral setose areas (preanal appendages) located ventrodistally, diffusing into scattered groups of setae. Digitiform, apicoventral setose lobes small, slender, directed ventrad; shorter than median keel on segment IX. Dorsal interlobular gap long, wide, V-shaped (Fig. 232). Apicodorsal setose lobes shifted posterad to middle of dorsum, forming narrow, separate pair of elongate setose areas. Smooth cavity absent. Longitudinal sutures absent, transverse sutures short, each running ventrad from bottom of intersegmental depression. Coxopodites short, robust, with slightly sinuous dorsal margin (Fig. 231); nearly straight in ventral view (Fig. 233). Harpagones elongate, tapering, straight (Fig. 231, 233). Phallic apparatus sinuate, forming almost equally thick tube (Fig. 234); exposed phallotremal sclerites heavily pigmented, rounded in lateral view (Fig. 234); twin sclerites double-rounded in ventral and dorsal view (Fig. 234, 235); tube narrowing at twin sclerites; strongly constricting in ventral view (Fig. 235); sclerotized, ventral phallothecal lobe hiding eversible membrane with cluster of slender spines. Ventral lobe widening strongly after constriction, ending in 3 lobes (Fig. 235); lateral lobes longer than broad, plate-like, less sclerotized median lobe. Dorsolateral membranous lobe located at mid-length of phallotheca; dorsally with pair of pale windows with single, small, blunt spine. Additional structure attached to posterior part of light windows, with small, thick spine. Medium, sclerotized half ring present dorsally; reaching half-way laterally; possibly forming sclerotized ventral tip of otherwise membranous ventrum of segment X.  
 Holotype male: INDIA: Karnataka, Shimoge District, Jog Falls, 02.iii.1980, collected at MV lamps at the bungalows [Gy. Topal] (OPC, alcohol).   
 Paratype: same data as holotype — 1 male (OPC, alcohol).   
 Distribution. India.   
 Etymology. semkala, from “semkala”, meaning “to add” in Sanskrit, referring to the species additional sclerous structure present on the middle dorsum of the phallotheca.
description: Published as part of Oláh, J. & Johanson, K. A., 2008, Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae), pp. 1-248 in Zootaxa 1802 on pages 140-142