cometadata/datacite-rtg-text-other-reclassification-records huggingface dataset:
Phragmacossia bandeamiri Yakovlev, Pljustch, Skrylnik & Pak, sp. nov.
Creators: name: Yakovlev, Roman V.
name: Pljustch, Igor G.
name: Skrylnik, Yuriy
name: Pak, Oleg
name: Witt, Thomas J.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Phragmacossia
Phragmacossia bandeamiri

description: Phragmacossia bandeamiri Yakovlev, Pljustch, Skrylnik & Pak, sp. nov.
(Figs 69−70, 87, 89, 110)
Material. Holotype: male, Afghanistan, Bamian, Band-e-Amir, Sabzel vill., 2900 m, 19.0 7.2013, leg. I. Pljustch, Yu. Skrylnik & O. Pak (ZISP); paratypes: 4 males, same locality, 19− 20.07.2013 (RYB).
Description. Forewing length 11–13 mm. Thorax and abdomen densely covered with pale-brown hairs. Antenna cup-shaped, bipectinate from base to border of middle and distal third, distal third filamentous. Forewing pale brown, with thin pale edge on outer margin, patternless. Fringe pale brown, monochrome. Hindwing patternless, pale grey, with thin pale edge on outer margin. Fringe pale brown, monochrome.
Male genitalia. Uncus long, thin, with rostral apex; tegumen medium-sized; processes of transtilla short, teniform, free; valva of medium width, symmetrical, with parallel sides and broadly rounded apex; juxta with long dorsally directed teniform lateral processes; saccus broad, semicircular; aedeagus short, thick, curved in medial third, with large cornutus on lateral surface of vesica, wall of aedeagus opposite cornutus folded.
Female unknown.
Diagnosis. The new species differs from other members of the genus by its very small size, completely reduced wing pattern, and evenly parallel costal and abdominal edges of the valva.
Annotated list of species (* new for Afghanistan)
description: Published as part of Yakovlev, Roman V., Pljustch, Igor G., Skrylnik, Yuriy, Pak, Oleg & Witt, Thomas J., 2015, The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park, pp. 41-72 in Zootaxa 3990 (1) on page 44, DOI: 10.11646/zootaxa.3990.1.3, http://zenodo.org/record/289113
Catopta Staudinger 1899
Creators: name: Yakovlev, Roman V.
name: Pljustch, Igor G.
name: Skrylnik, Yuriy
name: Pak, Oleg
name: Witt, Thomas J.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Catopta

description: Genus Catopta Staudinger, 1899
Catopta Staudinger, 1899: 157 −159. Type species: Catopta albimacula Staudinger, 1899. Designation by monotypy.
Catopta cashmirensis (Moore, 1879) (Figs 1, 91)
Cossus cashmirensis Moore, 1879: 86.
Type locality: Tawi, Kashmir [Tawi, Kashmir, NW India]. Type material: holotype (male) in MHUB, examined.
Distribution: N. India, Pakistan, Afghanistan, Nepal, Bhutan, China (Tibet, N. Yunnan). Reported for Pandshir-Tal, Parian (Daniel 1964).
Habitat and biology. Flight period: September. Altitude: 2350−2600 m.
Material examined: 1 male, SO Afghanistan, Safed-Koh, Südseite Kotkal, 2350 m, 21.06.−1.07.1969, leg. Vartian (MNHW).
Catopta eberti Daniel, 1964 (Figs 2, 92)
Daniel, 1964: 4−5.
Type locality: Afghanistan, Hazaradjat, Koh-i-Baba, Pandjao Umg. Type material: holotype (male) in ZSSM, examined.
Distribution: Afghanistan.
Habitat and biology. Flight period: end of June. Altitude: 2500 m. Material examined: 1 male (holotype), Afghanistan, Hazaradjat, Koh-i-Baba, Pandjao Umg., 2500 m, 26.06.−1.07.1961, leg. G. Ebert (ZSSM).
Catopta kendevanensis Daniel, 1937 (Figs 3−5, 93)
Daniel, 1937: 50.
Type locality: Persia s., Elburs mts. C., Kendevan pass. [pass Kendevan, Elburs Mts. Range, Iran]. Type material: holotype (male) in ZSSM, examined.
Distribution: Northern and Central Iran and Afghanistan.
In Afghanistan: subspecies anjumanica Daniel, 1964: 4.
Type locality: Anjuman Pass, Anjuman-Gebirge, Badakschan, NO-Afghanistan. Type material: holotype (male) in ZSSM, examined.
Distribution: NE and Central Afghanistan.
Reported for Band-e-Amir (Daniel 1965).
Habitat and biology. Flight period: July −August. Altitude: 3000−4200 m (Daniel 1964; 1965). Material examined: 8 males, 4 females (holotype, allotype and 10 paratypes), NO Afghanistan, Anjuman Pass, Anjuman-Gebirge, Badakschan, 4000−4200 m, 11−13.08.[19]52, leg. Klapperich (ZSSM); 4 males, 1 female, Afghanistan, Badakhshan, Bala-Kuran loc., 10.0 7.2003, leg. local coll. (MWM); 1 male, Afghanistan, Anjuman, 3000 m, 10.0 8.1963, leg. Omoto (ZSSM).
Catopta rocharva Sheljuzhko, 1943 (Figs 6, 94)
Sheljuzhko, 1943: 83.
Type locality: Rocharv im Pjandzh-Tale (Ruschan) [Rushan, Tadzhikistan]. Type material: holotype (male) in ZMKU, examined.
Distribution: Tadzhikistan (Gissar, W Pamir), NE Afghanistan (Daniel 1964). Reported for Badakshan, Sarekanda (Daniel 1964).
Habitat and biology. Flight period: July. Altitude: 3600 m (Daniel 1964). Material examined: 1 male, Hindukush, Panjir, 2500, 29.0 7.1971, leg. Flauger (MWM).
description: Published as part of Yakovlev, Roman V., Pljustch, Igor G., Skrylnik, Yuriy, Pak, Oleg & Witt, Thomas J., 2015, The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park, pp. 41-72 in Zootaxa 3990 (1) on pages 44-45, DOI: 10.11646/zootaxa.3990.1.3, http://zenodo.org/record/289113
Kotchevnik Yakovlev 2004
Creators: name: Yakovlev, Roman V.
name: Pljustch, Igor G.
name: Skrylnik, Yuriy
name: Pak, Oleg
name: Witt, Thomas J.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Kotchevnik

description: Genus Kotchevnik Yakovlev, 2004
Yakovlev, 2004: 358.
Type species: Cossus modestus Staudinger, 1887, by original designation.
Kotchevnik tapinus (Püngeler, 1898) (Figs 7, 95)
Cossus tapinus Püngeler, 1898: 57.
Type locality: Transcaspien, Merw, Sefir-kuh [Tadzhikistan, Kulyab]. Type material (syntypes) in MHUB, examined.
Distribution: Tadzhikistan, Turkmenistan, Iran, Afghanistan, Pakistan (Yakovlev 2004). Reported for Sarobi (Daniel 1964). Reports of “ Cossus tapinus ” from Nuristan, Barikot (Daniel 1965) and Nimla, 40 km SW Dschelalabad (Daniel 1971) were based on misidentifications of specimens from the genus Dervishiya Yakovlev, 2006: D. cadambae (Moore, 1881) and D. vartianae Yakovlev, 2011 (description was based on the specimens mentioned by Daniel (1971)), respectively. Habitat and biology. Flight period: June −July. Altitude: 1100−1800 m (Daniel 1964; 1965).
Material examined: 1 male, Afghanistan, Kabul Umg., 29.0 9.51, leg. Derele, Heydemann Kiel, GenPr – Het, 9105 (MWM); 2 males, Afghanistan, Iran Grenze, Umg. Kohsan, 1000 m, 13.5.77, de Freina (MWM).
description: Published as part of Yakovlev, Roman V., Pljustch, Igor G., Skrylnik, Yuriy, Pak, Oleg & Witt, Thomas J., 2015, The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park, pp. 41-72 in Zootaxa 3990 (1) on pages 45-46, DOI: 10.11646/zootaxa.3990.1.3, http://zenodo.org/record/289113
Cossus Fabricius 1794
Creators: name: Yakovlev, Roman V.
name: Pljustch, Igor G.
name: Skrylnik, Yuriy
name: Pak, Oleg
name: Witt, Thomas J.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Cossus

description: Genus Cossus Fabricius, 1794
Fabricius, 1794: 3.
Type species: Phalaena cossus Linnaeus, 1758. Designation by monotypy.
Cossus afghanistana Daniel, 1953 (Figs 8−10, 88, 96)
Cossus cossus afghanistana Daniel, 1953: 256.
Type locality: Afghanistan, Wardik [Central Afghanistan, Paghman Mts.].
Type material: holotype (male) in ZSSM, examined.
General distribution: Afghanistan, Pakistan (Yakovlev 2011).
Reported for Kabul, Paghman, Herat, Gulbahar and Polichomri (30 km NW Kabul) (Daniel 1953; 1963; 1964; 1965).
Habitat and biology. Flight period: April −September. Altitude: 700−2100 m (Daniel 1953; 1963; 1964). Systematic note. The taxon аfghanistana Daniel, 1953 was described as a subspecies. We found strong differences in the male genitalia between the taxon in question and C. cossus (L.), namely absence of developed crest on costal edge of valva, short processes of transtilla and slender aedeagus. Cossus afghanistana Daniel, 1953 was raised to species level (Yakovlev 2009b). External differences are less pronounced and include dark brown spots in the postdiscal area of the forewing which are fused and form a broad transverse band.
Material examined: 1 male (holotype), Afghanistan, Wardik, 7.0 7.1950, Volk leg. (ZSSM); 1 male, Kabul (ZMMU); 9 males, Afghanistan, Paghman, 30 km NW v. Kabul, 2100 m, 20- 30.07.1963, leg. Kasy & Vartian (MNHW); 2 males, Hindukush, bei Rukha, 2400 m, 21.0 5.1977, de Freina (MWM); 3 males, 30 km NW Kabul, 2500, 15.5.65 (MWM); 1 male, Paghman, 3000 m, 6.7.73 (MWM); 2 males, Kabul prov., Tangi (MWM); 3 males, Khurd-Kabul (MWM); 2 males, Panjir Valley, Kotul e Shava, Astana, 2500 m, 2005 (MWM).
description: Published as part of Yakovlev, Roman V., Pljustch, Igor G., Skrylnik, Yuriy, Pak, Oleg & Witt, Thomas J., 2015, The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park, pp. 41-72 in Zootaxa 3990 (1) on page 46, DOI: 10.11646/zootaxa.3990.1.3, http://zenodo.org/record/289113
Deserticossus Yakovlev 2006
Creators: name: Yakovlev, Roman V.
name: Pljustch, Igor G.
name: Skrylnik, Yuriy
name: Pak, Oleg
name: Witt, Thomas J.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Deserticossus

description: Genus Deserticossus Yakovlev, 2006
Yakovlev, 2006: 32.
Type species: Cossus arenicola Staudinger, 1897, by original designation.
Deserticossus arenicolus (Staudinger, 1879) (Figs 19, 101)
Cossus Arenicola Staudinger, 1879: 317.
Type locality: [Narun, nordostlich von Astrachan zwischen Wolga- und Ural-Flu., etwa 15 deutsche Meilen ostwarts von der Wolga] (NW Kazakhstan, european part).
Type material: lectotype (male) in MHUB, examined. Designation by Yakovlev (2006a). Distribution: Turkmenistan, Kazakhstan, Uzbekistan, Kyrgyzstan, N. Afghanistan, NW China (Xinjang) (Hua et al. 1990; Yakovlev 2006a).
In central and southern Afghanistan: subspecies iranica (Austaut, 1897)
Deserticossus arenicolus iranica Austaut, 1897
Cossus iranicus Austaut, 1897: 44.
Type locality: Sefis Kuh, Perse [S. Iran, Fars prov.].
Type material: holotype (male) in BMNH, examined.
Distribution: Iran, Afghanistan, Pakistan (Yakovlev 2006a).
Reported for “ 35 km nördlich Kandahar, Arghadab Dam” (Daniel 1964) and “SO v. Shindan” (Daniel 1965). Habitat and biology. Flight period: May −June. Altitude: 1150 m. Host plants: in Tadzhikistan (Tigrovaya balka Reserve) and Uzbekistan: Tamarix ramosissima, T. hispida (Tamaricaceae), Caragana sp. (Fabaceae) (Sinadsky 1960; Stschetkin 1963; Falkovitch 1986).
Material examined: 2 males, 20 km N. Kandagar, Afghan-Dab dam, 1150 m, 23.0 5.1961, Ebert (ZSSM); 1 male, W. Afghanistan, SO v. Shindan, 25.0 6.1963, leg. Kasy & Vartian (MNHW).
description: Published as part of Yakovlev, Roman V., Pljustch, Igor G., Skrylnik, Yuriy, Pak, Oleg & Witt, Thomas J., 2015, The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park, pp. 41-72 in Zootaxa 3990 (1) on pages 47-48, DOI: 10.11646/zootaxa.3990.1.3, http://zenodo.org/record/289113
Cryptoholcocerus Yakovlev 2006
Creators: name: Yakovlev, Roman V.
name: Pljustch, Igor G.
name: Skrylnik, Yuriy
name: Pak, Oleg
name: Witt, Thomas J.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Cryptoholcocerus

description: Genus Cryptoholcocerus Yakovlev, 2006
Yakovlev, 2006: 61, 68.
Type species: Cossus mongolicus Erschoff in Alpheraky, 1882. Designation by monotypy.
Cryptoholcocerus mongolicus (Erschoff in Alpheraky, 1882) (Figs 20, 102) Cossus mongolicus Erschoff in Alpheraky, 1882: 33.
Type locality: Kouldja [NW China, Kuldzha].
Type material probably lost.
Distribution: SE Kazakhstan, Kirgizstan, Uzbekistan, Tadzhikistan, Pakistan, Afghanistan, NW China (Yakovlev 2006a; 2011).
Reported for Badakhshan, Barak and Sarobi (Daniel 1964).
Habitat and biology. Flight period: July −September. Altitude: 1100−1600 m (Daniel 1964). Material examined: male, NO Afghanistan, Badakhshan, Barak, 1600 m, 23.0 7.1957, leg. G. Ebert (ZSSM); male, Kabul, 1800 m, 4.0 7.1963, Omoto (ZSSM); 2 males, Badakhschan, Bela-Kuran, 2900 m, 12.0 7.1963, Omoto leg. (ZSSM); male, Sarobi (O. Afghanistan), 1100 m, 27.0 8.1961, leg. Ebert (ZSSM).
description: Published as part of Yakovlev, Roman V., Pljustch, Igor G., Skrylnik, Yuriy, Pak, Oleg & Witt, Thomas J., 2015, The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park, pp. 41-72 in Zootaxa 3990 (1) on page 48, DOI: 10.11646/zootaxa.3990.1.3, http://zenodo.org/record/289113
Dervishiya Yakovlev 2006
Creators: name: Yakovlev, Roman V.
name: Pljustch, Igor G.
name: Skrylnik, Yuriy
name: Pak, Oleg
name: Witt, Thomas J.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Dervishiya

description: Genus Dervishiya Yakovlev, 2006
Yakovlev, 2006c: 197.
Type species: Cossus cadambae Moore, 1865. Designation by monotypy. * Dervishiya cadambae (Moore, 1865) (Figs 43, 112)
Cossus cadambae Moore, 1865: 822.
Type locality: Calcutta.
Type material: holotype (male) in BMNH, examined. Distribution: India, Pakistan, new for Afghanistan (Yakovlev 2011). Habitat and biology. Flight period: June.
Material examined: 1 male, Afghanistan, Nuristan, Barikot, 1800 m, 12− 17.07.1963, leg. Kasy & Vartian (MNHW).
Dervishiya vartianae Yakovlev, 2011 (Figs 44−45, 113)
Yakovlev, 2011: 30.
Type locality: Afgh[anistan]., Nimla, 40 km SW v. Dschelalabad. Type material: holotype (male) in MWM, examined.
Distribution: Afghanistan (Yakovlev 2011).
Erroneously reported as Catopta pallidalae Hampson, 1892 (Daniel 1965). Сossus pallidalae Hampson, 1892 (Fig. 46) was described on the single specimen from Sikkim and later moved to the genus Sansara Yakovlev, 2004 (subfamily Zeuzerinae).
Habitat and biology. Flight period: June.
Material examined: 13 males (holotype and 12 paratypes), 4 females (paratypes) Afgh[anistan]., Nimla, 40 km SW v. Dschelalabad, 23.6.1965, E. & A. Vartian leg. (MWM; MNHW).
description: Published as part of Yakovlev, Roman V., Pljustch, Igor G., Skrylnik, Yuriy, Pak, Oleg & Witt, Thomas J., 2015, The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park, pp. 41-72 in Zootaxa 3990 (1) on pages 50-51, DOI: 10.11646/zootaxa.3990.1.3, http://zenodo.org/record/289113
Dyspessa Hubner 1820
Creators: name: Yakovlev, Roman V.
name: Pljustch, Igor G.
name: Skrylnik, Yuriy
name: Pak, Oleg
name: Witt, Thomas J.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Dyspessa

description: Genus Dyspessa Hübner, [1820] 1816
Hübner, [1820] 1816: 194.
Type species: Phalena pantherina Hübner, 1790, by subsequent designation by Kirby (1892).
Dyspessa sp. (Figs 56, 84, 89, 110)
Material examined: male, Afghanistan, Bamian, Band-e-Amir, Sabzel vill., 2900 m, 19.0 7.2013, leg. I. Pljustch, Yu. Skrylnik & O. Pak (RYB).
Notes. We were unable to identify the single specimen from the large Palaearctic genus Dyspessa. The specimen resembles the Iranian D. serica Brandt, 1938 (type locality: Fort Sine-Sefid [Iran, Fars, Strasse Chiraz-Kazeroun]). Differences in the genitalia are insufficient, and external characters cannot be used due to the poor condition of the specimen. It will be possible to determine the species when new specimens are obtained. Daniel listed four specimens of the genus for Afghanistan: D. tristis A. Bang-Haas, 1912 (type locality: Karagai-tau [Kazakhstan]) for Herat, D. nigritula (Staudinger, 1887) (type locality: Transalai [S. Kyrgyzstan]) for Sarekanda and Kandaharkuna, Dyspessa sp. for Balkh (Daniel 1964) and D. pallidata Staudinger, [1892] (type locality: Amasia: «Hadjin, Marasch und Kara Hissar» [Turkey]) for Khurd-Kabul (Daniel 1971). Poor quality black-and-white photos pictured in the papers do not allow to identify the species with certainty. The material was not examined. Thus the composition of the Afghan fauna of Dyspessa cannot be ascertained.
description: Published as part of Yakovlev, Roman V., Pljustch, Igor G., Skrylnik, Yuriy, Pak, Oleg & Witt, Thomas J., 2015, The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park, pp. 41-72 in Zootaxa 3990 (1) on page 52, DOI: 10.11646/zootaxa.3990.1.3, http://zenodo.org/record/289113
Dieida Strand
Creators: name: Yakovlev, Roman V.
name: Pljustch, Igor G.
name: Skrylnik, Yuriy
name: Pak, Oleg
name: Witt, Thomas J.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Dieida

description: Genus Dieida Strand in Stichel, 1911
Strand in Stichel, 1911: 162−163.
Type species: Dieida persa Strand, 1911. Designation by monotypy.
Dieida persa Strand in Stichel, 1911 (Figs 57, 90, 110)
Strand in Stichel, 1911: 163.
Type locality: Prov. Arrak [NW Iran, Arrak].
Type material: syntypes (2 females) in MHUB, probably lost. Distribution: NW Iran, Afghanistan.
Habitat and biology. Flight period: May. Altitude: 2900 m. Material examined: 1 female, Afghanistan, Bamian, Band-e-Amir, H- 2900 m, 18.0 5.2012, leg. I.G. Pljustch (RYB).
Dieida afghana Yakovlev & Pljustch, 2014 (Figs 58, 90, 110)
Yakovlev et Pljustch, 2014: 75−76.
Type locality: Afghanistan, Bamian, Band-e-Amir.
Type material: holotype (female) in ZISP, examined.
Distribution: Central Afghanistan.
Habitat and biology. Flight period: June. Altitude: 3100 m. The new species was collected flying in the daytime on bushy north-east slopes with mugworth and prickly shrubs plant formations in the north end of Zulfikar Mts. 5 km NE main office of the Band-e-Amir National Park.
Material examined: 1 female (holotype), Afghanistan, Bamian, Band-e-Amir, H- 3100 m, 1.0 6.2012, leg. I.G. Pljustch (ZISP).
description: Published as part of Yakovlev, Roman V., Pljustch, Igor G., Skrylnik, Yuriy, Pak, Oleg & Witt, Thomas J., 2015, The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park, pp. 41-72 in Zootaxa 3990 (1) on pages 52-53, DOI: 10.11646/zootaxa.3990.1.3, http://zenodo.org/record/289113
Phragmataecia Newman 1850
Creators: name: Yakovlev, Roman V.
name: Pljustch, Igor G.
name: Skrylnik, Yuriy
name: Pak, Oleg
name: Witt, Thomas J.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Phragmataecia

description: Genus Phragmataecia Newman, 1850
Newman, 1850: 2931.
Type species: Noctua arundinis Hübner, [1808]. Designation by monotypy.
Phragmataecia albida Erschoff, 1874 (Figs 71, 123)
Phragmataecia castaneae albida Erschoff, 1874: 34.
Type locality: Kisil-Kum desert [SW Uzbekistan].
Type material: lectotype (male) in ZISP, examined. Designation by Yakovlev (2005). Distribution: Iran, Turkmenistan, Uzbekistan, Kazakhstan, NW China, Afghanistan, Russia (S. Volga reg.) (Yakovlev 2011).
Erroneously reported as Ph. roborowskii Alpheraky, 1897 for Hindukush Doab; Herat, Bala Murghab (Daniel 1964; 1969).
Habitat and biology. Flight period: May −June, September −October. Altitude: 470−1400 m (Daniel 1964; 1969). Material examined: 1 male, N-Afghanistan, Prov. Badakhshan, Khwahan (Darwaz), 1000m, 11.VII.1972, leg. Brade & Naumann (ZFMK); 1 male, O-Afghanistan, Pr. Kunar, Nuristan, ob. Lindai Sin-Tal, vic. Barg e Matal, Flussaue, UV-Licht, 2200m. 9.VII.1970, leg. C. Naumann (ZFMK).
Phragmataecia dushman Yakovlev, 2009 (Figs 72, 124)
Phragmataecia dushman Yakovlev, 2009: 357 −358.
Type locality: O. Afghanistan, prov. Nengrahar, Jalalabad.
Type material: holotype (male) in MWM, examined.
Distribution: E. Afghanistan.
Habitat and biology. Flight period: April −June. Altitude: 450−1100 m. Material examined: male (holotype), O. Afghanistan, prov. Nengrahar, D. Povolny (MWM); 1 female, O- Afghanistan, Sarobi, 1100 m, 28.0 6.1956, Amsel leg. (ZSSM).
Notes. Reported for Sarobi (Daniel 1964) as Phragmataecia furia Groum-Grshimaïlo, 1890 (type locality – “rive du Sourkhan” [banks of the Surkhob, Tadzhikistan]). Externally it differs from the latter by the darker colouration and brown fringe of the hindwing.
Phragmataecia furia Groum-Grshimaïlo, 1890 (Figs 73, 125) Groum-Grshimaïlo, 1890: 542 –543.
Type locality: “rive du Sourkhan” [banks of the Surkhob river, Tadzhikistan]. Type material (cotypes) in BMNH and ZISP.
Distribution: Uzbekistan, Tadzhikistan, Afghanistan (Yakovlev 2011). Material examined: 1 male, N-Afghanistan, Prov. Takhar, Amu Darya-Auen, Darqad Gebiet, 450m, 29.IV.1971, leg. C. Naumann (ZFMK);
description: Published as part of Yakovlev, Roman V., Pljustch, Igor G., Skrylnik, Yuriy, Pak, Oleg & Witt, Thomas J., 2015, The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park, pp. 41-72 in Zootaxa 3990 (1) on pages 54-55, DOI: 10.11646/zootaxa.3990.1.3, http://zenodo.org/record/289113
Zeuzera Latreille 1804
Creators: name: Yakovlev, Roman V.
name: Pljustch, Igor G.
name: Skrylnik, Yuriy
name: Pak, Oleg
name: Witt, Thomas J.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Zeuzera

description: Genus Zeuzera Latreille, 1804
Latreille, 1804: 186.
Type species: Phalaena aesculi Linnaeus, 1767. Designation by monotypy.
Zeuzera nuristanensis Daniel, 1964 (Figs 74, 126)
Zeuzera nuristanensis Daniel, 1964: 6 −7.
Type locality: Afghanistan, Nuristan, Bashgultal.
Type material: holotype (male) in ZSSM, examined.
Distribution: Afghanistan, N. Pakistan (Yakovlev 2011).
Reported for Nuristan, Bashgultal, 25 km N Barikot and Kutiau (Daniel 1964). Habitat and biology. Flight period: April −July. Altitude: 1150−1800 m (Daniel 1964). Material examined: male (holotype), Afghanistan, Nuristan, Bashgultal, 1150 m, 9.0 5.1953, leg. J. Klapperich (ZSSM); 2 males (paratypes), Afghanistan, Nuristan, 25 km N v. Barikot, 1800 m, 12.0 7.1963, leg. Kasy & Vartian (MNHW); 1 male (paratype), Afghanistan, Nuristan, Kutitau, 1450 m, 2.0 5.1953, leg. J. Klapperich (ZSSM).
description: Published as part of Yakovlev, Roman V., Pljustch, Igor G., Skrylnik, Yuriy, Pak, Oleg & Witt, Thomas J., 2015, The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park, pp. 41-72 in Zootaxa 3990 (1) on page 55, DOI: 10.11646/zootaxa.3990.1.3, http://zenodo.org/record/289113
Azygophleps Hampson 1892
Creators: name: Yakovlev, Roman V.
name: Pljustch, Igor G.
name: Skrylnik, Yuriy
name: Pak, Oleg
name: Witt, Thomas J.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Azygophleps

description: Genus Azygophleps Hampson, 1892
Hampson, 1892: 309.
Type species. Hepialis scalaris Fabricius, 1775. By original designation.
Azygophleps afghanistanensis (Daniel, 1964) (Figs 75, 127)
Zeuzera regia afghanistanensis Daniel, 1964: 6.
Type locality: O-Afghanistan, Sarobi, Gulbahar.
Type material: holotype (male) in ZSSM, examined.
Distribution: Afghanistan.
Reported for Gulbahar and Sarobi (Daniel 1964) and 30 km NW Kabul (Daniel 1971). Habitat and biology. Flight period: June −July. Altitude: 1100−2500 m (Daniel 1964; 1971). Material examined: male (holotype), O-Afghanistan, Gulbahar, 1700 m, 7.56, Amsel (ZSSM); male (paratype), O- Afghanistan, Sarobi, 1100 m, 3.0 7.56, Amsel leg. (ZSSM); 1 male, Afghan., Prov. Parwan, 69°30, 35°20, Panjishirtal, 1800 m, Charikar, 16.6.1971, leg. Vartian (MNHW).
description: Published as part of Yakovlev, Roman V., Pljustch, Igor G., Skrylnik, Yuriy, Pak, Oleg & Witt, Thomas J., 2015, The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park, pp. 41-72 in Zootaxa 3990 (1) on page 55, DOI: 10.11646/zootaxa.3990.1.3, http://zenodo.org/record/289113
Meharia Chretien 1915
Creators: name: Yakovlev, Roman V.
name: Pljustch, Igor G.
name: Skrylnik, Yuriy
name: Pak, Oleg
name: Witt, Thomas J.
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Cossidae
Meharia

description: Genus Meharia Chrétien, 1915
Chrétien, 1915: 367.
Type species. Meharia incurvariella Chrétien, 1915. Designation by monotypy.
Meharia incurvariella Chrétien, 1915 (Figs 76−78, 128) Meharia incurvariella Chrétien, 1915: 368.
Type locality: Biskra [Algeria].
Type material: holotype (male) in MNHN, examined.
In Afghanistan subspecies persica (Wiltshire, 1946)
Blalia vittata persica Wiltshire, 1946: 120.
Type locality: Shiraz [Fars, SW Iran].
Type material: holotype (male) in BMNH, examined.
Distribution: Iran, Afghanistan, Pakistan (Yakovlev 2011). Reported for “SO v. Shindan” and “ 80 km NO v. Kandahar” (Daniel 1965). Habitat and biology. Flight period: June.
Material examined: 2 females, SO v. Shindan, 25.0 6.63, leg. Kasy & Vartian (MNHW); 1 female, SO-Afghanistan, 80 km NO v. Kandahar, 27.0 6.1963, leg. Kasy & Vartian (MNHW).
description: Published as part of Yakovlev, Roman V., Pljustch, Igor G., Skrylnik, Yuriy, Pak, Oleg & Witt, Thomas J., 2015, The Cossidae (Lepidoptera) of Afghanistan with description of three new species and special notes on the fauna of Bande-Amir National Park, pp. 41-72 in Zootaxa 3990 (1) on page 56, DOI: 10.11646/zootaxa.3990.1.3, http://zenodo.org/record/289113
Metapheretima Michaelsen 1928
Creators: name: Nguyen, Tung T.
name: Tran, Binh T. T.
name: Nguyen, Anh D.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Clitellata
Opisthopora
Megascolecidae
Metapheretima

description: Genus Metapheretima Michaelsen, 1928
Pheretima (Metapheretima) (partim) Michaelsen, 1928: 8.
Metapheretima — Sims & Easton 1972: 205, 233; Easton 1979: 79.
Type species. Perichaeta neoguinensis Michaelsen, 1892, by original designation.
Diagnosis. " Megascolecidae with an oesophageal gizzard in VIII, intestinal caeca and gizzards absent. Body cylindrical, setae never excessively crowded ventrally, creeping sole absent. Male pores lacking porophores, occasionally on slim penes which may be within copulatory pouches. Crescentic genital markings associated with the male pores on mature individuals of most species. Spermathecal pores small or large, spermathecal diverticula variable in structure and origin" (Easton 1979: 79).
Distribution. North and southeast New Guinea, New Britain, Lombok, Carolines, Solomon Islands and New Hebrides (Easton 1979).
description: Published as part of Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D., 2014, Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta, pp. 105-121 in Zootaxa 3866 (1) on page 106, DOI: 10.11646/zootaxa.3866.1.5, http://zenodo.org/record/227564
Metapheretima vietnamensis Thai 1984
Creators: name: Nguyen, Tung T.
name: Tran, Binh T. T.
name: Nguyen, Anh D.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Clitellata
Opisthopora
Megascolecidae
Metapheretima
Metapheretima vietnamensis

description: Metapheretima vietnamensis Thai, 1984
(Fig. 1)
Metapheretima vietnamensis Thai, 1984: 614, Fig. 1 B; Blakemore 2007: 104.
Type material. 1C (W.270, Zoological Museum of M. V. Lomonosov Moscow State University, Russia) (Thai, 1984)
Records from Vietnam. Gia Lai Province (An Khe town); Da Nang City (Hoa Vang); Quang Nam Province (Hiep Duc; Tra My) (Thai 1984; Pham 1995a).
Distribution. Only known from Vietnam.
Remarks. The species is relatively similar to Metapheretima speiseri (Michaelsen, 1913) and Metapheretima apunae Lee, 1981, but differs from both in having two pairs of spermathecal pores in 5/6 and the spermathecal diverticulum originating from the spermathecal duct (Thai 1984).
description: Published as part of Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D., 2014, Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta, pp. 105-121 in Zootaxa 3866 (1) on page 108, DOI: 10.11646/zootaxa.3866.1.5, http://zenodo.org/record/227564
Pheretima arboricola Rosa 1890
Creators: name: Nguyen, Tung T.
name: Tran, Binh T. T.
name: Nguyen, Anh D.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Clitellata
Opisthopora
Megascolecidae
Pheretima
Pheretima arboricola

description: Planapheretima arboricola (Rosa, 1890)
(Fig. 2)
Perionyx arboricola Rosa, 1890: 119; Beddard 1895: 438; Michaelsen 1900: 209; Michaelsen 1903: 89; Gates 1936: 465; Gates 1972: 141.
Pheretima arboricola — Gates 1936: 399; Gates 1972: 169; Thai 1997: 5, Fig. 3 B. Planapheretima arboricola — Sims & Easton 1972: 233; Easton 1979: 76, Figs 27j, 30a.; Nakamura 1999: 5; Blakemore 2007: 8.
Type locality. Myanmar (Mt Carin).
Type material. Genoa 44017, Museo Civico di Storia Naturale “Giacomo Doria”, Italy. (Rosa 1890)
Material examined. 12A (SORC-V.206.1) Vinh Phuc province, Tam Dao National Park, on the way to the Rung Rinh peak, 20 June 1995.
Records from Vietnam. Vinh Phuc Province (Tam Dao National Park) (Thai 1997).
Distribution. Myanmar; Vietnam.
Remarks. The populations in Vietnam differ from the description of Gates (1936) in having spermathecal pores paired in VI, VII, additional genital markings in VI, and more setae (Thai 1997).
description: Published as part of Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D., 2014, Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta, pp. 105-121 in Zootaxa 3866 (1) on page 109, DOI: 10.11646/zootaxa.3866.1.5, http://zenodo.org/record/227564
lacertina Chen 1946
Creators: name: Nguyen, Tung T.
name: Tran, Binh T. T.
name: Nguyen, Anh D.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Clitellata
Opisthopora
Megascolecidae

description: Planapheretima lacertina (Chen, 1946)
(Fig. 2)
Pheretima lacertina Chen, 1946: 109, Plate IV Fig. A; Thai, 1997: 4, Fig. 2 C. Planapheretima lacertina — Sims & Easton 1972: 233; Easton 1979: 78, Figs 27k, 30d; Nakamura 1999: 5; Blakemore 2007: 57.
Type locality. China (Szechuan).
Material examined. 2C (HNUE) Son La Province,Ta Sua Natural Reserve, forest, 30 December 2012, coll. Tran T.T. Binh.
Records from Vietnam. Lao Cai Province (Sapa)—(Thai, 1997); Son La Province (Ta Sua).
Distribution. China; Vietnam.
Remarks. The population in Vietnam differs from the original description of the species (Chen 1946) in smaller body size, fewer setae and in having spermathecal pores paired in 5/6/7/8 (Thai 1997).
description: Published as part of Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D., 2014, Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta, pp. 105-121 in Zootaxa 3866 (1) on page 109, DOI: 10.11646/zootaxa.3866.1.5, http://zenodo.org/record/227564
Pheretima plumatomusculata
Creators: name: Nguyen, Tung T.
name: Tran, Binh T. T.
name: Nguyen, Anh D.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Clitellata
Opisthopora
Megascolecidae
Pheretima
Pheretima plumatomusculata

description: Planapheretima plumatomusculata (Thai, 1982)
(Fig. 2)
Pheretima plumatomusculata Thai, 1982: 829, Fig. 3 g; Thai 1997: 2, Fig. 2 B. Planapheretima plumatomusculata — Blakemore 2007: 81.
Type locality. Vietnam (Ninh Binh).
Type material. In the collection of Zoological Museum of M. V. Lomonosov Moscow State University, Russia.
Material examined. 8C, 7A (CTU) Vinh Phuc Province, Tam Dao National Park, on the way to the observation tower, 25 July 1998, coll. Tran T.T.Binh.
Records from Vietnam. Nghe An Province (Ky Son; Quy Hop); Ninh Binh Province (Cuc Phuong); Vinh Phuc Province (Tam Dao) (Thai 1997; Do 1994; Le 1995).
Distribution. Only known from Vietnam.
Remarks. The species is fairly similar to Planapheretima lacertina (Chen, 1946), but differs in having spermathecal pores paired in 5/6/7 dorsally and pigments dorsally (Thai 1997).
description: Published as part of Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D., 2014, Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta, pp. 105-121 in Zootaxa 3866 (1) on pages 110-111, DOI: 10.11646/zootaxa.3866.1.5, http://zenodo.org/record/227564
Pheretima rarusa
Creators: name: Nguyen, Tung T.
name: Tran, Binh T. T.
name: Nguyen, Anh D.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Clitellata
Opisthopora
Megascolecidae
Pheretima
Pheretima rarusa

description: Planapheretima rarusa (Thai, 1997)
(Fig. 2)
Pheretima rarusa Thai, 1997: 1, Fig. 2 A.
Planapheretima rarusa — Blakemore 2007: 85.
Type material. Holotype: 1C (SORC-V.202) Nghe An province, Ky Son District, Muong Long commune, Tham Lang village, forest, 21 June 1986, coll. Tran Minh Khoi. Paratypes: 1C, 1A, same data as holotype; 1C, Nghe An province, Ky Son District, Chau Son, primary forest (Thai 1997).
Description. Body length 75mm, diameter 3.5–4mm, segments 92. Body flattened ventrally with creeping sole. Prostomium epilobous. First dorsal pore in 9/10. Clitellum in XIV–XVI. Setae numerous, more distributed ventrad, 65(57) in V, 81(71) in VIII, 27–30 in XVIII, 68 in XXV, yz=7ab, zz=11aa. Porophores in shallow pouches of parietal invagination. Genital markings small, oval papillae, ranged in transversal presetal and postsetal lines, 2 on XVII, 3+2 on XVIII and 2 on XIX. Spermathecal pores three pairs, dorsolateral in 5/6/7/8. Ampulla more or less heart-shaped, diverticula as long as or longer than ampulla, attaching to ducts. Testes sacs ventral, not separated. Seminal vesicles well developed in XI–XVI. Prostate glands well-developed, strongly lobuled. Ovisacs well developed. Last hearts in XIII. Intestine beginning at XV. Intestinal caeca in XXVI–XXVII. Septa 8/9/10 present, other septa thinned. Pharyngeal micronephridia in 6/7. (Data from Thai 1997: 1–2.)
Records from Vietnam. Nghe An Province (Ky Son: Muong Long) (Thai 1997).
Distribution. Only known from Vietnam.
Remarks. The species is very close to Planapheretima plumatomusculata (Thai, 1982) but differs in violet color dorsally, having copulatory pouch, genital markings in male regions, and accessory glands with long duct (Thai 1997). Types were rechecked for this study.
description: Published as part of Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D., 2014, Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta, pp. 105-121 in Zootaxa 3866 (1) on page 111, DOI: 10.11646/zootaxa.3866.1.5, http://zenodo.org/record/227564
Polypheretima tenebrica Chen 1946
Creators: name: Nguyen, Tung T.
name: Tran, Binh T. T.
name: Nguyen, Anh D.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Clitellata
Opisthopora
Megascolecidae
Polypheretima
Polypheretima tenebrica

description: Planapheretima tenebrica (Chen, 1946)
(Fig. 2)
Pheretima tenebrica Chen, 1946: 93, Plate III Fig. A; Thai 1997: 4, Fig. 3 A. Planapheretima tenebrica — Sims & Easton 1972: 233; Easton 1979: 77, Figs 27l, 30e; Nakamura 1999: 5; Blakemore 2007: 97.
Type locality. China (Szechuan).
Records from Vietnam. Vinh Phuc Province (Tam Dao) (Thai 1997).
Distribution. China; Vietnam.
Remarks. The population in Vietnam differs from the original description (Chen 1946) in bigger size, more setae and having simple intestinal caeca (Thai 1997).
description: Published as part of Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D., 2014, Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta, pp. 105-121 in Zootaxa 3866 (1) on page 111, DOI: 10.11646/zootaxa.3866.1.5, http://zenodo.org/record/227564
Polypheretima Michaelsen 1934
Creators: name: Nguyen, Tung T.
name: Tran, Binh T. T.
name: Nguyen, Anh D.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Clitellata
Opisthopora
Megascolecidae
Polypheretima

description: Genus Polypheretima Michaelsen, 1934
Pheretima (Polypheretima) Michaelsen, 1934a: 25. Metapheretima (part)— Sims & Easton 1972: 205, 233. Polypheretima — Easton 1979: 28.
Type species. Perichaeta stelleri Michaelsen, 1892, by original designation.
Diagnosis. " Megascolecidae with an oesophageal gizzard in VIII, intestinal caeca and gizzards absent. Body cylindrical, setae never excessively crowded ventrally, creeping sole absent. Male pores on circular porophores which may be within copulatory pouches. Crescentic genital markings absent. Spermathecal pores small, spermathecal diverticula simple and usually ectal in origin" (Easton 1979: 28).
Distribution. Philippines, Indonesian archipelago, Papua New Guinea, Malaya peninsula and Vietnam (Easton 1979).
description: Published as part of Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D., 2014, Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta, pp. 105-121 in Zootaxa 3866 (1) on pages 111-112, DOI: 10.11646/zootaxa.3866.1.5, http://zenodo.org/record/227564
Polypheretima annamensis Stephenson 1931
Creators: name: Nguyen, Tung T.
name: Tran, Binh T. T.
name: Nguyen, Anh D.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Clitellata
Opisthopora
Megascolecidae
Polypheretima
Polypheretima annamensis

description: Polypheretima annamensis (Stephenson, 1931)
(Fig. 3)
Pheretima annamensis Stephenson, 1931: 60, text-fig. 13.
Metapheretima annamensis — Sims & Easton 1972: 233.
Polypheretima annamensis — Easton 1979: 56; Nakamura 1999: 7; Blakemore 2007: 7.
Type locality. Vietnam (Lam Dong).
Type material. BMNH 1930.2.7.10-11, British Museum of Natural History, London, UK. (Stephenson, 1931) Records from Vietnam. Lam Dong Province (Lang Bian Peaks) (Stephenson 1931). Distribution. Only known from Vietnam.
Remarks. There are no other records from Vietnam since its description.
description: Published as part of Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D., 2014, Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta, pp. 105-121 in Zootaxa 3866 (1) on page 112, DOI: 10.11646/zootaxa.3866.1.5, http://zenodo.org/record/227564
Polypheretima colonensis
Creators: name: Nguyen, Tung T.
name: Tran, Binh T. T.
name: Nguyen, Anh D.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Clitellata
Opisthopora
Megascolecidae
Polypheretima
Polypheretima colonensis

description: Polypheretima colonensis (Thai, 1996)
(Fig. 3)
Pheretima colonensis Thai, 1996: 1, fig. 2A.
Polypheretima colonensis — Nakamura 1999: 7; Blakemore 2007: 24.
Type material. 7A (SORC-HV.222) Ba Ria-Vung Tau province, Conlon Island, Nha Bang slope, 6200m asl, 24–26 June 1989, coll. Thai Tran Bai (Thai 1996).
Description. Body cylindrical, unpigmented. Prostomium prolobous. First dorsal pore in 11/12. Setae all fine and regularly distributed on pre- and postclitellar segments, 97(22) in V, 89 in VIII, 72 in XIII, 50+ 4 in XVIII, 80(16) in XIX, 74 in XXV, aa=2ab, zz=2zy. Male porophore superficial, on an eyelike circular plate. Crescentic genital markings present in association with male porophores. No other genital markings in spermathecal and male regions. Spermathecal pores 2 pairs, ventral in furrows 5/6/7, about 0.25 body circumference apart. Ampulla heartshaped, divercula short, attaching to duct. Metandric. Testes sacs ventral in XI, not separated. Seminal vesicles in XII. Ovaries and ovisacs well developed in XIII. Last hearts in XIII. Intestine beginning at XVII, without caeca. Typhlosole simple, lamelliform, T=0.12. Septa 8/9 and 10/11 absent, 5/6/ 7/8 and 9/10 thickened, 11/12/13 slightly thickened. Pharyngeal micronephridia in 7/8. (Data from Thai 1996: 1–2.)
Records from Vietnam. Ba Ria—Vung Tau Province (Conlon Island).
Distribution. Only known from Vietnam.
Remarks. The species is fairly similar to Polypheretima sepikensis Ude, 1924, but it is metandric, genital markings and accessory glands are absent, and the first dorsal pore is in 11/12. There are crescentic markings in front of and behind the male pores, but they differ from the crescentic genital markings of Metapheretima sensu Easton (1979). Types were rechecked for this study.
description: Published as part of Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D., 2014, Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta, pp. 105-121 in Zootaxa 3866 (1) on page 112, DOI: 10.11646/zootaxa.3866.1.5, http://zenodo.org/record/227564
Polypheretima grandisetosa
Creators: name: Nguyen, Tung T.
name: Tran, Binh T. T.
name: Nguyen, Anh D.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Clitellata
Opisthopora
Megascolecidae
Polypheretima
Polypheretima grandisetosa

description: Polypheretima grandisetosa (Thai, 1996)
(Fig. 3)
Pheretima grandisetosa Thai, 1996: 3, Fig. 2 B.
Polypheretima grandisetosa — Nakamura 1999: 7; Blakemore 2007: 45.
Type material. Holotype: 1A (SORC-HV.223) Ba Ria-Vung Tau province, Conlon Island, slope near Datdoc Bay, 10m asl, 25 June 1989, coll. Thai Tran Bai. Paratype: 1A, same data as holotype (Thai 1996).
Further material. 5A (SORC), Ba Ria-Vung Tau province, Baycanh Island, 210m asl, 20 June 1989, coll. Thai Tran Bai (Thai 1996).
Description. Body cylindrical, small size, unpigmented. Prostomium prolobous. First dorsal pore in 11/12. Setae all fine and regularly distributed on pre- and postclitellar segments, 60(16) in V, 54 in VI, 50 in VIII, 40+ 4 in XVIII, 44 in XXV, aa=1.52ab, zz=zy; segment VI with 4–6 enlarged setae on the dorsal side. Male pores on transversely oval porophores. No genital markings in both spermathecal and male regions. Spermathecal pores 2 pairs, ventral in furrows 5/6/7, about 0.25 body circumference apart. Metandric. Seminal vesicles in XII. Ovaries small in XIII. Last hearts in XIII. Intestine beginning at XVII, without caeca. Typhlosole simple, lamelliform, T=0.15. Septa 8/9/10 absent, 5/6/7/8 thickened. Pharyngeal micronephridia in 7/8. (Data from Thai 1996: 3–4.)
Records from Vietnam. Ba Ria-Vung Tau Province (Conlon Island; Baycanh Island).
Distribution. Only known from Vietnam.
Remarks. The species is relatively close to Polypheretima colonensis Thai, 1996, but differs in smaller size and the presence of big setae in VI. Types were rechecked for this study.
description: Published as part of Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D., 2014, Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta, pp. 105-121 in Zootaxa 3866 (1) on page 115, DOI: 10.11646/zootaxa.3866.1.5, http://zenodo.org/record/227564
Polypheretima kyhaensis
Creators: name: Nguyen, Tung T.
name: Tran, Binh T. T.
name: Nguyen, Anh D.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Clitellata
Opisthopora
Megascolecidae
Polypheretima
Polypheretima kyhaensis

description: Polypheretima kyhaensis (Thai, 1996)
(Fig. 3)
Pheretima kyhaensis Thai, 1996: 4, Fig. 3 B.
Polypheretima kyhaensis — Nakamura 1999: 7; Blakemore 2007: 61.
Type material. Holotype: 1A (SORC-HV.224) QUANG NAM, Nui Thanh District, Ky Ha commune, 24 November 1989, coll. Pham Thi Hong Ha. Paratype: 1A, same data as holotype (Thai 1996).
Description. Body cylindrical, colour pale. Prostomium epilobous. First dorsal pore in 11/12. Setae numerous, regularly distributed on segments, 160(40) in V, 126(50) in VIII, 32+ 16 in XVIII, 60(20) in XIX, 62 in XXV, aa>ab, zz>zy. Male porophores superficial. Spermathecal pores 4 pairs, ventral in 5/6-8/9. Genital markings presetal, paired, one pair each on VIII and IX, two pairs on XVIII and XIX, and one pair on XVII, XX and XXI. Ampulla heart-shaped, divercula small, attaching to duct. Holandric. Testes sacs extending to the lateral line. The first seminal vesicles more developed and larger than the other seminal vesicles. Ovaries large on septum 12/13. Prostate glands in XVII-XIX. Intestine beginning at XV, without caeca. Typhlosole simple, lamelliform, T=0.35. Last hearts in XII. Septa 8/9/10 absent, 4/5–7/8 thickened. Pharyngeal micronephridia in 5/6/7/8. Lymph glands not developed (Data from Thai 1996: 4–5.).
Records from Vietnam. Quang Nam Province (Nui Thanh, Ky Ha).
Distribution. Only known from Vietnam.
Remarks. The species is fairly similar to Polypheretima monticola (Beddard, 1912), but differs in the absence of septa 8/9/10, more complicated arrangement of genital markings in the male pore region. Types were rechecked for this study.
description: Published as part of Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D., 2014, Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta, pp. 105-121 in Zootaxa 3866 (1) on pages 115-116, DOI: 10.11646/zootaxa.3866.1.5, http://zenodo.org/record/227564
Polypheretima parataprobanae
Creators: name: Nguyen, Tung T.
name: Tran, Binh T. T.
name: Nguyen, Anh D.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Clitellata
Opisthopora
Megascolecidae
Polypheretima
Polypheretima parataprobanae

description: Polypheretima parataprobanae (Thai & Nguyen, 1993)
(Fig. 3)
Pheretima parataprobanae Thai & Nguyen, 1993: 7, Fig. 1 C. Polypheretima parataprobanae — Blakemore 2007: 77.
Type material. Holotype: 1C (SORC-HV.225) Thua Thien Hue province, Hue city, Nguyet Bieu, sandy garden, 13 January 1985, coll. Nguyen Van Thuan. Paratypes: 1C, 1A, same data as holotype (Nguyen 1993).
Description. Medium size, body cylindrical, colour pale. Prostomium prolobous. First dorsal pore in 12/13. Setae numerous, regularly distributed on segments, 104–126 in V, 124–134 in VIII, (64–78) + (18–22) in XVIII, 104–120 in XIX, 96–110 in XXV, aa≥ab, zz=1–1.5zy. Porophores raising strongly with small slits around the foot. No genital markings in male region. Spermathecal pores 1 pair ventral in 7/8. Genital markings oval-shaped, one pair, presetal on VIII. Ampulla heart-shaped, ducts short and small, diverticula short, attaching to ducts. Testes sacs ventral, not separated. Protate glands slightly lobuled. Typhlosole lamelliform, T=0.03. Intestine beginning at XVI, without caeca. Septa 5/6/7/8 thickened, 8/9/10 absent. (Data from Nguyen 1993: 7.)
Records from Vietnam. Thua Thien Hue Province (Hue city, Bach Ma National Park); Da Nang city (Hoa Vang) (Nguyen 1993) (Pham 1995a).
Distribution. Only known from Vietnam.
Remarks. The species is fairly similar to Polypheretima taprobanae (Beddard, 1892), but differs in absence of genital markings in the male region, and having a pair of genital markings near the spermathecal pores. Types were rechecked for this study.
description: Published as part of Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D., 2014, Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta, pp. 105-121 in Zootaxa 3866 (1) on page 116, DOI: 10.11646/zootaxa.3866.1.5, http://zenodo.org/record/227564