Paramanota trilobata Hippa &amp; Kaspřák &amp; Kahar &amp; Ševčík 2016, new species
Creators: name: Hippa, Heikki
name: Kaspřák, David
name: Kahar, Siti Rafhiah Haji Abd
name: Ševčík, Jan
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Mycetophilidae
Paramanota
Paramanota trilobata

description: <i>Paramanota trilobata</i>, new species (Figs. 2A–D) <b>Type material.</b> Holotype. Male (T 4371 in QSBG). THAILAND, Petchaburi, Kaeng Krachan NP, Panernthung [km 27 water pump], 12°49.151’N 99°22.483’E, 970 m, Malaise trap, coll. Sirichai &amp; Chusak, 12–19 September 2008. <b>Female.</b> Unknown. <b>Etymology.</b> The name is Latin, <i>trilobata</i>, three-lobed, referring to the three-lobed gonostylus. <b>Distribution.</b> Thailand. <b>Description.</b> Male. <b>Colour</b>. Head yellowish brown, vertex and antennal flagellum darker brown; setae and other vestiture dark. Thorax yellowish, scutum, scutellum and the medial part of mediotergite brown, anterior part of anepisternum infuscated; thoracic setae mostly worn out, the few existing ones dark, the short setae anteriorly on scutum seem pale. Legs yellowish, hind femur with an infuscated subapical patch ventrally, the setae and other vestiture dark which makes the apical part of tibiae and all of tarsi seem dark under low magnification. Wing yellowish brownish; haltere yellowish with dark brown knob. Abdomen brown, tergites and tergites concolorous, setae dark. <b>Head.</b> Similar to fig 7a in Hippa et al. (2005) but the number of facial setae higher. Antennal flagellomere 4, Fig. 2D. Maxillary palp similar to fig 7a in Hippa et al. (2005), ultimate palpomere 1.9 times longer than the penultimate one. The strong postocular setae are not countable in the specimen. <b>Thorax</b> similar to Fig. 8a in Hippa et al. (2005). <b>Legs</b>. Front tibial organ not well visible in the specimen, only one seta observable. <b>Wing</b> similar to fig. 1D in Hippa (2010). Wing length 2.6 (3.0) mm. <b>Hypopygium</b>, Figs 2A–C: Gonocoxae ventrally separated by a membranous area, each side with a very large lobe which is posteriorly extending as far as the gonostylus, with the posterior part simple, not divided into sub-lobes; the ventral surface of the lobe as well as the other ventral parts of gonocoxa evenly covered with strong setae, the dorsal surface of the lobe with a postero-mesial area of numerous megasetae enlarging in size towards the base, a few rather unmodified setae flanking on the lateral side the area with megaseta. Gonocoxa dorsally simple with the setae similar to those of the ventral side. Tergite 9 simple, with strong setae. Tergite 10 fused with tergite 9 and appearing as a small setose lobe at the postero-lateral corner of the latter. Cercus simple. The hypoproct with one seta. Gonostylus with a dorsal lobe, a ventral lobe and a median lobe, each with difficultly observable largely membranous sub-lobes; the dorsal lobe in dorsal view elongated, nearly parallel sided, with membranous apical part showing a striated or lamellar structure; the ventral lobe narrow, oblique, at anterior margin with a long comb-like row of narrow lamellae, the median lobe narrow, oblique, without comb-like structures. Aedeagus elongate, with a broader basal part and narrower almost parallel sided apical part, the details not discernible in the mount. <b>Discussion.</b> In the key to the species of <i>Paramanota</i> (Hippa, 2010), <i>P. trilobata</i> fits couplet 1 including only <i>P. orientalis</i> because the ventral gonocoxal lobe is posteriorly simple, not divided into a more lateral and a more mesial sub-lobes. In this respect <i>P. trilobata</i> is similar to <i>P. rodzayi,</i> the other new species described in the present paper. Both these species are distinguished from <i>P. orientalis</i> by having all the megasetae on the dorsal side of the ventral gonocoxal lobe short, the longest ones at most one fourth the width of the lobe while in <i>P. orientalis</i> they are double of that length. <i>Paramanota trilobata</i> is distinguished from <i>P. rodzayi</i> by having the antennal flagellomeres longer, the flagellomere 4 being as long as broad instead of only half of that length, by having the dorsal aspect of the dorsal gonostylar lobe elongate subquadrangular, not subtriangular, and by lacking a transverse comb-like row of lamellae apically on the median lobe of the gonostylus. Also many other details in the hypopygium distinguish the two species (Figs. 1, 2).
description: Published as part of <i>Hippa, Heikki, Kaspřák, David, Kahar, Siti Rafhiah Haji Abd &amp; Ševčík, Jan, 2016, Two new Oriental species of Paramanota Tuomikoski (Diptera: Mycetophilidae), with DNA sequence data, pp. 360-367 in Raffles Bulletin of Zoology 64</i> on pages 363-366, DOI: 10.5281/zenodo.5355716
description: {"references": ["Hippa H, Jaschhof M &amp; Vilkamaa P (2005) Phylogeny of the Manotinae, with a review of Eumanota Edwards, Paramanota Tuomikoski and Promanota Tuomikoski (Diptera Mycetophilidae. Studia Dipterologica, 11 (2004): 405 - 428."]}