Tomosvaryella planca Foldvari, Skevington & Motamedinia 2023, sp. nov.
Creators: name: Motamedinia, Behnam
name: Földvari, Mihaly
name: Skevington, Jeffrey H.
name: Kelso, Scott
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Pipunculidae
Tomosvaryella
Tomosvaryella planca

description: Tomosvaryella planca Földvari, Skevington & Motamedinia sp. nov.
urn:lsid:zoobank.org:act: 73676891-FBE8-4A5A-9A17-80147F090C16
Figs 78A–E, 115, 146E, 151E
Diagnosis: This species can be recognized by the two patches of spines on the hind trochanter (Fig. 146E); long and condensed bristles on membranous area; surstyli elongated in dorsal view (Fig. 78A); phallic guide with some different-sized spines (Fig. 78B); one of ejaculatory ducts with some saw-like teeth (Fig. 78B); both gonopods with a finger-like projection in ventral view (Fig. 78B).
Description: MALE: Body length: 3.0– 3.1 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black, lower part distinctly silver pollinose; eyes touching for distance equal to 1.5–1.8 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper half. Thorax. Postpronotum pale, with 4–5 distinct pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part silver pollinose, slightly grayish also from the side. Scutellum silver pollinose and with 5–7 weak bristles along the distal edge. Dorsocentral bristles well developed, dark, longer on frontal part and numerous dark bristles around postpronotum. Halter, knob pale, stem black. Legs. Mid coxa with 2–3 spines. Trochanters and femora black, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees yellow, tibiae black, tarsal segments dark brown, only yellowish ventrally; fore metatarsus is transparent in ventral view. Hind trochanter with a few short dark spines (5–8) ventrobasally and (2–3) shorter ones ventroapically (Fig. 146E). Ventroapical row of spines on fore femur missing, 4–6 very short, black spines on mid femur; no spines on hind femur, only 6–8 pale bristles posteriorly on distal half (as long as the width of hind tibia at the distal end); ventrobasal spines (2) present on fore and weaker on mid femur; hind tibia slightly bent at middle in posterior view. Subapical (distal) spines on first four tibiae present, very short. Hind metatarsus extremely flattened, spoon-like (extending posterior edge, widest at basal 1/3) (Fig. 151E), second tarsomere slightly longer than 3–5. Pulvilli as long as last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with one dark and long bristles. Fourth costal section 1.5–2 times as long as third costal section, fourth section appears to be unusually short. Cross-vein r-m slightly distal to middle of discal cell. 4–5 dark setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 silvery grey, sides silvery pollinose on tergites 4 and 5 (the latter with larger spots). Dispersed long dark bristles on tergites present, the longest on ST8 and up to 3.5–4 times as long as width of hind tibia at base. Lateral bristles on first tergite present, 6–8 brown bristles in a patch up to as long as 1–1.2 times the width of hind femur at distal tip in lateral view. Postabdomen in dorsal view: T6, S7 invisible; T5 1–1.5 times as long as syntergosternite 8 (ST8). Genitalia without dissection: ST8 large, black, hairy with longest bristles on posterior part; membranous area round, posteriorly directed, the longest bristles originate from membranous area; tip of epandrium brown; surstyli pale, appear elongated and uniformly broad. Genitalia. Genital capsule in dorsal view: epandrium wider than long (MLE: MWE = 0.8). Surstyli asymmetrical, elongated, left longer than right one (Fig. 78A). Genital capsule in ventral view: subepandrial sclerite covered with some transverse streaks in middle; both gonopods with a finger-like projection extended towards surstyli, left rather longer than right one (Fig. 78B), phallic guide with 5–6 different-sized spines dorsolaterally (longer ones extended towards left surstylus); phallus with three ejaculatory ducts, one bears 10–11 saw-like teeth along its side. Genital capsule in lateral view: left surstylus straight in basal two thirds, bent towards sternite in apical one third; right surstylus curved towards sternite (Fig. 78D–E); ejaculatory apodeme flat, sperm pump elongated, linear (Fig. 78C).
FEMALE: Unknown.
Etymology: The name planca is derived from the Latin plancus, flat-footed, in reference to the flattened hind metatarsus.
Examined material: HOLOTYPE: AUSTRALIA: Queensland: 3.5 km SW by S Mount Baird, 15°10’S, 145°7’E, 3.v.1981, D.H. Colless, Malaise trap, JSS8576 (1♂, ANIC); PARATYPES: AUSTRALIA: Queensland: 20 km E Mareeba, 17°0’S, 145°26’E, 16.iv.1980, G.F. Hevel & J.A. Fortin, JSS10535 (1♂, USNM); 3.5 km SW by S Mount Baird, 15°10’S, 145°7’E, 3.v.1981, D.H. Colless, Malaise trap, JSS8577, JSS8585 (1♂, ANIC; 1♂, QM); Edgbaston National Reserve (EBM1), 22°44’S, 145°26’E, Melaleuca floodplain, 29.X–15. XII.2010, C. Lambkin et al., Malaise trap, 18528, CNC596962 (1♂, CNC); Western Australia: Martin’s Well, West Kimberley, -16.56604, 122.85010, 28.IV.1977, D.H. Colless, Malaise trap, JSS9223 (1♂, ANIC).
Distribution: Australia (Queensland, Western Australia) (Fig. 115).
Notes: This species is mostly restricted to NE Queensland, but a disjunct record is from the West Kimberley region.
description: Published as part of Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H. & Kelso, Scott, 2023, Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species, pp. 1-271 in Zootaxa 5599 (1) on pages 173-175, DOI: 10.11646/zootaxa.5599.1.1, http://zenodo.org/record/14970961