Compsophorus (Xenojoppa) quadrimaculatus Riedel 2023, nov. sp.
Creators: name: Riedel, Matthias
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Compsophorus
Compsophorus quadrimaculatus

description: Compsophorus (Xenojoppa) quadrimaculatus nov. sp.
(Figs 19A–D)
Holotype. (♂) Indonesia: Sulawesi, nr Bantaeng, Borong Rappoa, Gn. Daulu, Mal. trap 16, c 700 m, 4–24.iv.1991, C. van Achterberg, RMNH´91 (Leiden).
Paratype. (♂) similar label (Leiden).
Etymology. The species name refers to the four ivory spots on the mesoscutum.
Description. ♂. Body length 14–15 mm.
Head. Flagellum with 37–39 flagellomeres, bristle-shaped, with distinctly nodose flagellomeres. 1 st flagellomere 2.0 × and 2 nd flagellomere 2.4 × longer than wide. Tyloids not visible. Head large. Temple long, slightly widened behind eye and moderately narrowed apically (fig. 19B). OED 1.2–1.6 × and OOD 0.7–1.0 × ocellar diameter. Vertex and frons with sparse punctures, strongly shining. Face and clypeus densely punctate, face dull. Clypeus flat, c 1.8 × wider than long; apical margin sharp and slightly bilobed (fig. 19A). Mandible large; ventral tooth large and only slightly shorter than dorsal tooth. Malar space 0.8 × as long as width of mandibular base. Gena with rather dense punctures. Genal carina reaching mandibular base.
Mesosoma. Mesoscutum with coarse vermiform rugae. Notaulus indistinctly but widely impressed. Mesopleuron rugose-punctate ventrally, smooth and with sparse punctures dorsally; speculum large, smooth. Metapleuron with superficial punctures, shining; juxtacoxal carina present. Scutellum distinctly elevated, almost square, coarsely punctate or rugose, with complete lateral carina. Propodeum short and strongly sloping downwards to apex, with reticular-rugose surface (fig. 19C), without apophysis; spiracle slit-shaped. Area superomedia small, oval, c 2.0 × longer than wide; costula in its middle. Legs slender. Hind coxa densely punctate. Hind femur 5.0–5.3 × longer than wide. Claws simple. Areolet pointed or shortly stalked frontally. Vein 1 cu-a strongly postfurcal by 3–4 × its width. Nervellus of hknd wing distinctly reclivous, intercepted in apical 0.7.
Metasoma. Slender.Petiole with lateral carina in distal half. Postpetiole distinctly widened, without latero-median carina, finely rugose or aciculate, median field not separated. 2 nd tergite 1.0–1.2 × as long as wide. Gastrocoelus distinctly impressed, with fine ridges. Thyridium large, slightly oblique, 1.7–2.3 × wider than the interval between thyridia (fig. 19D). 2 nd tergite finely and densely rugose-punctate, finely aciculate medially, dull. Following tergites finely punctate, ± shining. Only 2 nd sternite with median fold. Hypopygium unmodified, apical margin straight.
Colour. Black with extended ivory coloration. Flagellum black. Head and mesosoma black with ivory colour pattern. Ivory are palps, mandible except teeth, clypeus, face, gena, frontal orbit, scape ventrally, collar, upper margin and ventral third of pronotum, tegula, subtegular ridge, epicnemium, ventral stripe and dorsal spot of mesopleuron, median stripe of mesosternum, short anterolateral and paramedian spots of mesoscutum, lateral spots of scutellum, postscutellum, upper division and central spot of metapleuron. Propodeum mainly ivory, area superomedia brownish, subbasal transverse band of propodeum and spot on area petiolaris blackish (fig. 19C). In holotype, 1 st to 4 th tergites except black base of petiole red, with yellow apical bands. 5 th to 7 th tergites black, with narrow ivory apical margins. In paratype, 1 st and 2 nd tergites reddish-brown, following tergites blackish, apical bands of 1 st to 4 th tergites yellow, following tergites with narrow yellow apical margins. Legs reddish; fore and mid coxae and trochanters ivory; hind coxa reddish-brown, with ivory dorsal spot; fore tibia with yellow stripe frontally; hind tibia infuscate in apical 0.2; hind tarsus ochreous or black. Wings with weak yellowish suffusion; pterostigma brown.
Remarks. This new species belongs to Xenojoppa Cameron, 1902 which has been synonymized with Compsophorus by Townes & Townes (1973) but separated by Heinrich (1975). Xenojoppa is preliminarily included as a subgenus in the genus Composphorus here.
In his key of the known Xenojoppa species (Heinrich 1975: 487–489) it runs to Xenojoppa parvidens Heinrich, 1975 from Sulawesi but differs by yellowish anterolateral and paramedian spots of mesoscutum, apical yellow bands or margins of all tergites, entirely black flagellum and extended ivory coloration of the propodeum. Since many Xenojoppa species show a considerable colour variability and the ♀ of the new species is still unknown, it is possible that the new species represents a colour variant of Xenojoppa parvidens only.
It seems necessary that the separation of different Xenojoppa species as shortly presented by Heinrich (1975: 487–488) needs further confirmation by additional material and genomic studies.
Distribution. Only known from Sulawesi.
description: Published as part of Riedel, Matthias, 2023, Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia, pp. 1-94 in Zootaxa 5363 (1) on pages 46-48, DOI: 10.11646/zootaxa.5363.1.1, http://zenodo.org/record/10146851