Pygoluciola hamulata
Creators: name: Ballantyne, L. A.
Publisher: name: Zenodo
Year: 2008
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Lampyridae
Pygoluciola
Pygoluciola hamulata

description: Pygoluciola hamulata (Olivier)   
(Figs. 1, 2)  
 Luciola hamulata Olivier, 1885: 367; 1902: 80; 1907: 52; 1913: 59. Luciola (Luciola) hamulata Olivier. McDermott, 1966: 105.   
 Luciola (Pygoluciola) hamulata Olivier. Ballantyne, 1968: 121.   
 Pygoluciola hamulata (Olivier). Ballantyne & Lambkin, 2006: 29.   
  Lectotype. –  Male, Borneo, Sarawak, 1865–1866, coll. G. Doria (MSNG). This is the only remaining specimen of the original two in MSNG on which Olivier based his original description. Labels: 1. black ink on card – “Borneo Sarawak 1865–66 coll. G Doria ”; 2. red printed label on red margined card ‘Typus; 3. black ink male symbol; 4. handwritten in black ink “ hamulata E. Oliv. ” Male specimen is hereby designated as the lectotype of Luciola hamulata Olivier, 1885.   
 Distinctive features. – Superficially most similar to P. stylifer, distinguished by the non curved tibiae of all legs, the entire apex of abdominal tergite 8 (that of stylifer is emarginate); distinguished from P. satoi by features listed under that species.   
  Redescription of lectotype male. –  Glue runs along the dorsal surface between abdomen and elytra and the abdomen is not attached to the thorax but to the elytra; the abdomen was not removed and certain features thus not ascertained on this specimen.   
Elongate, moderately slender, 2.5 times as long as wide; 9.1 mm long.  
Dorsal surface pale yellowish brown; pronotum semitransparent where fat body is retracted, with pale brown markings near posterior margin to either side of the median line; elytral punctures dark at their bases, more noticeably so in an area that extends posteriorly from elytral base to the mid point of the suture and laterally as far as interstitial line 2; head, antennae, palpi reddish brown, apical segment of maxillary palpi pale yellow on inner surface; apical segment of labial palpi pale light brown; venter of thorax dingy yellow except for red-brown metasternum; legs yellowish except for reddish brown apices of femora 1, and all of tibiae and tarsi of legs 1, 2, and brown tarsi only of legs 3; V2 pale brown, V3, 4 light red brown, semitransparent, V5 dark reddish brown; light organ in V6, 7 deep yellow (post mortem), remainder light brown; visible posterior portion of tergite 8 pale yellowish, rest of tergites not examined.  
Pronotum: (Fig. 1); 3.0 mm wide, 1.5 mm long; width/length = 2; 1/6 as long as whole body length; wider across posterior margin than elsewhere; width across posterior margin subequal to width across elytral humeri (elytra closed); median anterior margin rounded, not emarginate, scarcely projecting beyond anterolateral corners, which are narrowly rounded and slightly obtuse, and do not project strongly anteriorly; lateral margins smoothly divergent along their length; posterolateral corners angulate acute, projecting slightly beyond posterior margin, and separated from the slightly emarginated mid posterior margin by shallow emarginations which do not follow the anterior margin of the elytral humerus; dorsal surface mainly smooth, median sulcus not depressed, lateral margins flat and median area largely flat and slightly raised above lateral margins, with irregular small, low, smooth shiny elevations in posteromedian areas; punctures small shallow, some contiguous, some separated by up to their width.  
Elytra: (Fig. 2); 7.6 mm long; elytral lateral margins divergent in anterior 1/4, subparallel in next 1/4, and gently convergent in posterior 1/2; sutural margins divergent slightly in apical 1/4; when viewed from below with specimen horizontal, the elytral epipleuron at base covers the elytral humerus (which is thus not visible from beneath); when viewed from above (specimen horizontal) the most anterior portion of the epipleuron becomes visible at the humeral corner and in front of the posterior margin of the MS.  
 Head: gently depressed between eyes; retracted into the prothoracic cavity but 1/3 of head visible in front of the pronotum; GHW 1.9mm; SIW 0.4 mm; GHW/SIW 5; ASD subequal to ASW; labrum transverse, lateral margins overlie the inner edges of the (closed) mandibles; frons-vertex junction not defined, this area smoothly rounded. Mouthparts well developed; apical segment of labial palpi laterally flattened and bearing on their inner, longest edge, 5–6 projections on both (basal projection 3 × as broad as next; 3rd projection from this end is very narrow and short, with longer wider projections to each side; apical projection is narrowly bifurcate). Antennae longer than 2 × GHW but <3 × GHW.   
Legs: femora and tibiae straight, not swollen or curved.  
Abdomen: MPP of V7 appearing short and broad from beneath, curving strongly dorsally with its posterior curved area (which is not visible from below) about as long as the anterior area (which is visible from beneath); posterior face of MPP with narrow sharp edged margins and median area shallowly depressed, and engulfed at its tip by the strongly downturned, slender, and apically rounded apex of tergite 8. No dissections were made of this fragile specimen.  
Female and larva not associated.
description: Published as part of Ballantyne, L. A., 2008, Pygoluciola Satoi, A New Species Of The Rare Southeast Asian Firefly Genus Pygoluciola Wittmer (Coleoptera: Lampyridae: Luciolinae) From The Philippines, pp. 1-9 in Raffles Bulletin of Zoology 56 (1) on pages 3-4, DOI: 10.5281/zenodo.5354269