Dirhinus quadrhinus Delvare & Copeland 2018
Creators: name: Delvare, Gérard
name: Copeland, Robert S.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Chalcididae
Dirhinus
Dirhinus quadrhinus

description: The quadrhinus species group
Diagnosis, females (character number in parentheses). Body with long setae on mandibles, labrum and procoxa (1) (Fig. 25); clypeus with long setae on both sides, oriented forwards (2) (Figs 119, 148); mandibles about as long as procoxa (3) (Fig. 28); apices of mesal mandibular tooth broadly rounded (4) (Fig. 143); inner tooth of mandible absent (5); outer mandibular tooth hardly projecting, not visible in frontal view (6) (Fig. 143), head with four horns (8) (Fig. 31); inner horns quite broad (9); depression between inner horns very narrow, forming an incision (10); inner horns without apical notch (11); ocellar triangle not raised (12) (Fig. 39); lateral margin of scrobal depression without tooth (13) (Fig. 39; what could be mistaken here for an additional tooth is actually the profile of the outer horn, compare Fig. 42); flagellomeres short, transverse (14) (Fig. 43); clava narrowly rounded apically, with very small area of micropilosity (15); pro- and most of mesonotum with lanceolate, thick and adpressed setae, except on apical half of scutellum (16) (Fig. 49); mesonotum with extensive smooth surfaces, especially on mid lobe of mesoscutum anteriorly, disc of lateral lobes and oblong surface on centre of mesoscutellum (17) (Fig. 49); anteromedian areola of propodeum short, at most as long as wide (19) (Figs 123, 135, 144); lateral areolae on propodeum with setation similar to the rest of surface (20); medioventral tooth of prepectus emarginate apically (21) (Fig. 61); epicnemium virtually flat (22) (Fig. 66); mesodiscrimenal groove wide, appearing as an oblong fovea, truncate ventrally, tapering dorsally and without dorsal fovea (23) (Fig. 66); ventral section of epicnemial carina without median notch (24); subcoxal tooth on posterior margin of metepimeron projecting and narrowly rounded (25); posterior projection of metepisternum vestigial (26); metacoxa short and faintly strigose dorsally (27) (Fig. 76); metafemur with sparse setation (28) (Figs 76, 124); metafemur without engraved network (29) (Fig. 76); metatibia with evident outer groove in addition to tarsal sulcus, the outer edge of which has a row of short and adpressed setae (30) (Fig. 84); fore wing hyaline (31) (Fig. 85); setation on upper side of wing moderately sparse (32); under side of wing with microtrichiae only (33); Rs fold setose (34) (Fig. 87); very few admarginal setae on fore wing (35), setation on underside of marginal cell reduced (36); only two sensilla placodea at apex of submarginal vein (37) (Fig. 86); four aligned sensilla placodea on stigma (38); dorsal surface of petiole sloping backwards (39) (Fig. 139); petiolar surface strongly emarginate on anterior margin (40) (Fig. 118); median sulcus of petiole present, delimited by submedian carina (sulcus secondarily lost in some species) (41) (Fig. 118); dorsal surface of petiole bare (42); petiole with vestigial basoventral tooth (43) (Fig. 139); strigose surface on first gastral tergite well expanded, about as long as wide (44) (Figs 112, 140, 147); lateral ridges of first gastral tergite that delimit strigose surface somewhat curved outwards (45); hypopygium with long and diverging apical submedian setae (46).Dirhinus(Dirhinus)anthraciaWalker GDEL0154 Cameroon Centre Messamendongo, (surrounds of Yaoundé) 03°57'07" N 011°29'27" E Malaise trap 31/03- 14/04/2003 Tussac M.Dirhinus(Dirhinus) aff.hesperidumMorocco Fez - Meknès Moulay Idriss, Volubilis antic city 34°04'34" N 05°04'14" W 17/05/1995 Tussac H.Dirhinus(Hontalia) aff.caeruleusGDEL0261 French Guiana Regina Roura, Patawa camp, Montagne de Kaw 4°33'N 52°10' W (7) Mandible, outer tooth apex rounded (as in Fig. 152) rounded (as in Fig. 23) rounded (Fig. 25) rounded (as in Fig. 25) rounded (Fig. 23) rounded (Fig. 22) (12) Ocellar triangle not raised (as in Fig. 39) not raised (as in Fig. 39) not raised (as in Fig. 39) not raised (as in Fig. 39) not raised (as in Fig. 39) slightly raised (Fig. 40) (13) Margin of scrobal depression, secondary tooth absent but outer horn present (as in Fig. 39) absent (as in Fig. 40) absent (as in Fig. 40) absent (as in Fig. 40) absent (as in Fig. 40) absent (Fig. 40) (14) Flagellomeres, relative length short, transverse (as in Fig. 43) short, strongly transverse (as in Fig. 44) short, strongly transverse (Fig. 44) short, strongly transverse (as in Fig. 44) short, strongly transverse (as in Fig. 44) short, transverse except F1 and F2 (Fig. 45) (15) Clava, apex narrowly rounded (as in Fig. 43) narrowly rounded truncate (Fig. 44) truncate (as in Fig. 44) narrowly rounded (as in Fig. 43) moderately rounded (Fig. 45)
…….continued on the next page ……continued on the next page Dirhinus (Dirhinus) Dirhinus (Dirhinus) Dirhinus (Dirhinus) Dirhinus (Dirhinus) aff. Dirhinus (Dirhinus) Dirhinus (Dirhinus) aff.
Characters quadrhinus himalayanus claviger claviger anthracia hesperidum 16) Pronotum and setae thick, lanceolate and setae thick, lanceolate and setae thick, lanceolate and setae thick, lanceolate and setae thick, lanceolate setae slender, adpressed mesonotum, setation adpressed (as in Fig. 49) adpressed (as in Fig. 49) adpressed (as in Fig. 49) adpressed (as in Fig. 49) adpressed (Fig. 50) (Fig. 51) with smooth surfaces on with smooth surfaces on with smooth surfaces on with smooth surfaces on With small smooth surfaces with reduced smooth 17) Mesonotum, lateral lobes of mesoscutum lateral lobes of mesoscutum lateral lobes of mesoscutum lateral lobes of mesoscutum on lateral lobes of surface on center of pattern of puncturation and mesoscutellum (as in and mesoscutellum (as in and mesoscutellum (as in and mesoscutellum (as in mesoscutum and mesoscutellum (Fig. 51) Fig. 49) Fig. 49) Fig. 49) Fig. 49) mesoscutellum (Fig. 50)
18) Propodeum, present (as in Fig. 123) present present (Fig. 55) present (as in Fig. 55) present (Fig. 56) present (Fig. 57) spiracular tooth 19) Propodeum, as long as wide (as in Fig. wider than long (as in Fig. wider than long (as in Fig. as long as wide (Fig.
as long as wide (Fig. 55) as long as wide (Fig. 56) anteromedian areola 135) 134) 135) 57) 20) Propodeum, setae thin as on rest of setae thin as on rest of setae thin as on rest of setae lanceolate and thick in setae thin as on rest of setae thin as on rest of
setation of lateral propodeal surface (as in propodeal surface (as in propodeal surface (as in contrast with rest of propodeal surface (Fig.
propodeal surface (Fig. 55)
areola Fig. 135) Fig. 55) Fig. 55) propodeal surface (Fig. 56) 57)(21) Prepectus, apical margin of medioventral tooth distinctly emarginate (as in Fig. 61) distinctly emarginate (as in Fig. 62) distinctly emarginate (as in Fig. 62) distinctly emarginate (as in Fig. 62) distinctly emarginate (Fig. 62) distinctly emarginate (Fig. 63) (22) Mesepisternum, mesal elevation of epicnemium absent, epicnemium virtually flat (as in Fig. 66) epicnemium only very slightly raised mesally (as in Fig. 67) epicnemium only very slightly raised mesally (as in Fig. 67) epicnemium only very slightly raised mesally (as in Fig. 67) epicnemium only very slightly raised mesally (Fig. 67) epicnemium only very slightly raised mesally (Fig. 68) (23) Mesodiscrimenal groove wide and superficial (as in Fig. 66) wide and deep (as in Fig. 67) wide and deep (as in Fig. 67) wide and deep (as in Fig. 67) wide and deep (Fig. 67) moderately wide, deep (Fig. 68) (27) Metacoxa, habitus short (as in Fig. 76) short (as in Fig. 76) short (as in Fig. 76) short (as in Fig. 76) moderately long (Fig. 77) moderately long (Fig. 78) (28) Metafemur, setation sparse (Fig. 112) very sparse sparse (as in Fig. 76) very sparse dense (Fig. 77) dense (Fig. 78)
piliferous points only, piliferous points only, piliferous points only, piliferous points only, piliferous points only, piliferous points only, 29) Metafemur, without engraved network without engraved network without engraved network without engraved network without engraved network without engraved sculpture (as in Fig. 76) (as in Fig. 76) (as in Fig. 76) (as in Fig. 76) (Fig. 77) network (Fig. 78) present but setose, visible 30) Metatibia, outer present and evident (as in present and evident (as in present and evident (as in present and evident (as in through outer ridge only absent groove Fig. 84) Fig. 126, 127) Fig. 126, 127) Fig. 126, 127) (Fig. 83) ……continued on the next page Dirhinus (Pareniaca) aff. Dirhinus (Pareniaca) aff. Dirhinus (Hontalia) aff.
Characters Dirhinus (Pareniaca) inflexus Dirhinus (Pareniaca) 'Patawa' ehrhorni schwarzi caeruleus narrow, on top of mesal quite narrow, on top of mesal quite narrow, on top of mesal quite narrow, on top of mesal quite narrow, on top of mesal(20) Propodeum, setation of lateral areola setae thin as on rest of propodeal surface (as in Fig. 57) setae thin as on rest of propodeal surface (as in Fig. 57) setae lanceolate, thick (Fig. 58) setae lanceolate, thick (Fig. 59) setae thin, of medium length (Fig. 60) (21) Prepectus, apical margin of medioventral tooth convex (Fig. 64) strongly convex (as in Fig. 65) strongly convex (as in Fig. 65) strongly convex (Fig. 65) strongly convex (as in Fig. 65) (22) Mesepisternum, mesal elevation of epicnemium epicnemium moderately raised mesally (Fig. 69) epicnemium strongly raised (Fig. 70) epicnemium strongly raised (as in Fig. 70) epicnemium strongly raised (Fig. 71) epicnemium strongly raised (Fig. 72)
) Mesodiscrimenal groove elevation of epicnemium (Fig. elevation of epicnemium (Fig. elevation of epicnemium (as in elevation of epicnemium (Fig. elevation of epicnemium (Fig.
69) 70) Fig. 70) 71) 72)
) Metacoxa, habitus moderately long (Fig. 79) moderately long (as in Fig. 79) moderately long (Fig. 80) long (Fig. 81) very long (Fig. 82)
) Metafemur, setation dense (Fig. 79) dense (as in Fig. 79) dense (Fig. 80) dense (Fig. 81) dense (Fig. 82)
with engraved network in with engraved network in with engraved network in with engraved network in without engraved network, only
) Metafemur, sculpture addition to piliferous points addition to piliferous points (as addition to piliferous points addition to piliferous points piliferous points (Fig. 82)
(Fig. 79) in Fig. 79) (Fig. 80) (Fig. 81)
) Metatibia, outer groove absent absent absent absent absent
……continued on the next page ……continued on the next page) Hypopygium, pair of apical absent absent absent (Fig. 108) absent (Fig. 109) absent
description: Published as part of Delvare, Gérard & Copeland, Robert S., 2018, Four-horned wasps, description of some remarkable Dirhinus (Hymenoptera, Chalcididae) from Kenya, with a discussion of their taxonomic placement, pp. 301-349 in Zootaxa 4374 (3) on pages 309-323, DOI: 10.11646/zootaxa.4374.3.1, http://zenodo.org/record/1155252