Typhoctes evansi Quintero & Rodriguez 1990
Creators: name: Fernández, Fernando
name: Brothers, Denis J.
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Mutillidae
Typhoctes
Typhoctes evansi

description: Typhoctes evansi Quintero & Rodriguez, 1990
(Figs 4, 5)
Typhoctes evansi Quintero & Rodríguez, 1990: 30–34; holotype ♂, Caballero [approx. 8°32’N 80°12’W], Antón Distrito, Coclé Province, Rep. Panama (NMNH) (Figs 5A–D, but not examined).
Material examined. PANAMA, Tranquilla Norte [approx. 8°37’N 80°24’W], Antón, Coclé, 10.iii.1991, Ricci Rodriguez (det. R. Cambra 1991) (2♂, DJBC).
Diagnosis. Male. (Figs 4, 5A–D). Frons with fairly sparse white simple decumbent setae forming an indistinct lateral whorl; clypeus fairly densely covered with long slightly thickened semi-decumbent subplumose white setae scarcely concealing the cuticle. Lateral face of pronotum entirely fairly coarsely punctate, punctures merging with irregular longitudinal costulation on posterior half; disk of propodeum almost entirely densely longitudinally costulate but merging with dense coarse punctures on lateral one-sixth, declivity finely transversely costulate; lateral face of fused metepisternum-propodeum entirely densely longitudinally costulate. Mid and hind tibial spurs white with tip blackened. Female (Figs 5E–H). See Quintero & Rodríguez 1990.
Description. Male. Length 5.5 & 5.4 mm. Body shining black; appendages and wing veins very dark reddish brown; mandible and tegula dark reddish brown; mid and hind tibial spurs white with tip blackened; wings infumated throughout but slightly darker on apical third. Vestiture of decumbent and erect, silvery white and fuscous to black hairs, forming the following notable patterns: frons with fairly sparse white simple decumbent setae forming an indistinct whorl on each side, numerous scattered long thin erect dark sparsely subplumose setae and a few erect subplumose white and black setae between antennal bases, clypeus fairly densely covered with long slightly thickened semi-decumbent subplumose white setae scarcely concealing the cuticle, gena with simple decumbent and erect finely subplumose white setae, mandible with erect finely subplumose fuscous setae; mesosoma generally with fairly dense decumbent and erect apparently simple to finely subplumose white setae (some pale fuscous on scutum) and sparse apparently simple to finely subplumose dark fuscous erect setae antero-dorsally; legs with white setae except tarsal setae pale fuscous; T1–T6 fairly sparsely covered with erect finely subplumose white and pale fuscous setae; T2–T6 with larger scattered erect finely subplumose fuscous setae near posterior margin becoming denser posteriorly; T7 (pygidium) with dense short erect subplumose fuscous setae; T1 and T2 with posterior fringe of semi-erect thickened white setae; T3 with sparse fringe of fine whitish setae; T4–T6 with very inconspicuous sparse fringe of fine fuscous setae; metasomal sterna with sparse erect finely subplumose white setae, a very few scattered long erect finely subplumose fuscous setae on S3–S6; S8 (hypopygium) with scattered erect finely subplumose and denser shorter semi-decumbent fuscous setae.
Head. 1.19 & 1.15 × as wide as high, 1.00 &1.00 × width of mesosoma; narrowest part of frons 0.77 & 0.68 × eye height. Clypeus fairly densely and moderately punctate, almost flat but weakly convex medio-dorsally. Frons slightly convex, with fine well separated punctures; ventral width between eyes 0.83 & 0.75 × dorsal width between eyes. Median ocellus 1.38 & 1.22 × as wide as lateral ocellus, 0.23 & 0.24 × narrowest width of frons; POL 0.68 & 0.67 × OOL, POL 1.89 & 1.60 × LOL. Eye large, shallowly reniform, eye height 0.72 & 0.77 × head height; malar space short, 0.05 & 0.05 × eye height. Antennal sockets large, oval; ratios of lengths of antennal segments 1–4 about 22:7:14:18 and 20:7:13:17, scape 3.14 & 2.86 × as long as wide.
Mesosoma. 1.56 & 1.67 × as long as wide. Pronotum 1.48 & 1.53 × as wide as long, slightly widening posteriad, 2.57 & 2.50 × as wide as mid-length, posterior margin evenly shallowly concave; epaulet an inconspicuous diagonal tuft of short pale setae on weakly convex humeral tubercle; dorsum with well-marked moderate punctures irregularly interspersed with fine punctures, punctation sparse medially but denser elsewhere; lateral face scarcely concave dorsally, entirely fairly coarsely punctate, punctures merging with irregular longitudinal costation on posterior half. Mesoscutum densely moderately punctate anteriorly, punctures becoming larger and sparser posterad and surface smooth postero-medially, without notauli, deep well defined parapsidal groove extending three-quarters scutum length from postero-lateral angle. Scutellum densely punctate-costulate medially, more distinctly longitudinally costate laterally, posterior margin smooth. Metanotum densely punctate-costulate medially becoming more distinctly costate laterally. Disk of propodeum almost entirely densely longitudinally costulate but merging with dense coarse punctures on lateral one-sixth, declivity entirely transversely costulate. Mesopleuron evenly gibbose, coarsely and densely punctate but less so ventrally. Metepimeron weakly convex, clearly distinguishable, densely and finely punctate. Lateral face of fused metepisternum-propodeum entirely densely longitudinally costulate.
Wings. Fore wing 2.12 & 1.97 × as long as mesosoma, 3.21 & 3.34 × as long as broad; marginal cell very slightly wider basally than apically, with weakly rounded apex and sometimes with minute free stub at postero-lateral corner, 3.13 & 3.00 × as long as high, 0.72 & 0.77 × as long as first submarginal cell; second submarginal cell 1.97 & 2.33 × as long as high, 0.90 & 0.84 × as long as marginal cell, 0.65 & 0.64 × as long as first submarginal cell; third submarginal cell 1.05 & 1.31 × as long as high, 0.54 & 0.56 × as long as marginal cell, 0.39 & 0.43 × as long as first submarginal cell. Hind wing with (3&3) & (2&2) basal hamuli, and (7&8) & (6&7) apical hamuli.
Legs. Dorsal tooth on hind coxa a low convex elongate lamina postero-dorsally. Mid and hind tibiae each with 2 longitudinal rows of 2 or 3 black bristles dorsally and 2 or 3 similar bristles apically. Longer spur of hind tibia 0.78 & 0.79 × as long as hind basitarsus. Claws slender, with small tooth near middle of ventral margin.
Metasoma. 3.02 & 2.97 × as long as wide. T1 strongly constricted into anterior narrow neck, bulging dorsally on posterior third, constricted again at junction with T2, 1.36 & 1.34 × as long as wide and 0.68 & 0.69 × as wide as T2; indistinguishably fused with S1 on anterior third; entirely sparsely finely punctate, smooth interspaces at least the width of punctures. T2 similarly punctate as T1; anterolaterally with short indistinct sparse and pale felt line 0.17 & 0.15 × lateral length of T2, its anterior extremity at 0.06 & 0.14 × lateral length of T2. T3–T6 with sparse mixture of small and very fine punctures. Pygidium (T7) almost square, dorsally flattened and posteriorly slightly retuse, with dense small punctures, posterior margin straight medially and abruptly convex laterally. S1 mainly flattened, smooth and shining with very few widely scattered minute punctures at about anterior third. S2–S6 punctation similar to corresponding terga but much sparser. S7 very short, smooth. Hypopygium (S8) almost square, flattened, posterior margin straight to scarcely concave, with fairly dense small punctures.
Remarks. Although the specimens do not agree in some respects with the original description and figures (e.g. pronotal proportions), the anomalies appear to be simple errors, differences in interpretation or simple intra-specific variation. The identification is additionally supported by the fact that these specimens were collected not very far from the type locality by the collector of the original type series, although some time later, and were identified by Roberto Cambra who was also involved with the original collections.
Distribution. Panama.
description: Published as part of Fernández, Fernando & Brothers, Denis J., 2025, New species of Typhoctes (Hymenoptera, Thynnoidea, Chyphotidae) from Colombia and Jamaica, pp. 123-138 in Zootaxa 5633 (1) on pages 130-134, DOI: 10.11646/zootaxa.5633.1.6, http://zenodo.org/record/15372929