Quadrophenia townshendi , Rifkind 2017, new species
Creators: name: Rifkind, Jacques
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Melyridae
Quadrophenia
Quadrophenia townshendi

description: Quadrophenia townshendi Rifkind, new species   
(Fig. 5, 6, 7, 17)  
 Type material. Holotype, male. Panama, Panama Pr., Cerro Campana, 2700′, May 17–19, 1987, E. Giesbert, coll. Holotype deposited in CSCA. Paratypes: 3 (JNRC), 2 (FSCA), same data as holotype; PANAMA: 1 (CSCA), Panama Prov., Cerro Campana, 2100 ft., April 27–May 4, 1992, E. Giesbert, coll.; 1 (LACM), Panama Prov., Cerro Jefe, ± 2800′, May 21, 1987, E. Giesbert, coll.; 3 (WFBM), Pan [amá] Prov., Altos de Cerro Campana, V-17–18-1994, F. T. Hovore; 1 (JNRC), Coclé, Cerro Gaital, VI-10–12- 1985, E. Riley & D. Rider; 1 (RHTC), Chiriqui, 4.7 km N Valle de las Minas, 8 July 1997, R. Turnbow; COSTA RICA: 1 (FMNH), La Caja, S. Jose, V-1932, H. Schmidt, leg., F. Nevermann [collection?]; 2 (MNHN), 1920, Paul Serre, Museum Paris; 1 (JNRC), Est. Cacao, 1000–1400 m, Lado SO Vol. Cacao, P. N. Guan., Prov. Guanacaste, P. Rios, C. Moraga y M. M. Chavarria, 21 a 28 May, 1992, L–N 323300, 375700, INBIO CR1000 /734267; 1 (JNRC), Est. Pitilla, 700 m, 9 km S. Sta. Cecilia, P. N. Guanacaste, Prov. Guan., 19 May–3 Jun 1993, C. Moraga, L–N. 330200, 380200, INBIO CR1001 /315248; 1 (WOPC), Est. Pitilla, 700 m, 9 km S Sta. Cecilia, P. N. Guanacaste, Prov. Guan., R. W. Flowers, 15 May, 1991, L–N–330200, 380200, INBIO CR1000 /516354; 1 (MNCR), Est. Cacao, 1000–1400 m, Lado Suroeste del Volcan Cacao, Prov. Guan., II Curso Parataxon, Jun 1990, L–N–323300, 375700, INBIO CR1000 /250223; 1 (MUCR), Est. Cacao, 1000–1400 m, SW side Volcan Cacao, Guanac. Pr., Mar. 1988, GNP Biod. Sur, 323300, 375700, INBIO CR1000 /028684; 1(CASC), Est. Cacao, S. O. de Volcán Cacao, Prov. Guana. 800 –1600 m, Jul 1993, G. Mora, G. Fonseca, J. Saborio, G. Varela, LS 323300_375700, #2222, INBIO CR1001 /180362; MEXICO: 1 (CIUM), Chiapas, 20 km NNO Ocozocoautla, 25-VII-2000, Cols. V. H. Toledo y A. M. Corona.; 1 (JNRC), Chiapas, Sumidero Cyn., 4000′, June 14, 1987, E. Giesbert, coll.; 1 (JNRC), Chiapas, El Sumidero, Mirador La Coyota, 24 June, 1990, R. Turnbow; 1 (CSCA), Chiapas, Sumidero Cyn., 4000′, June 15, 1987, E. Giesbert, coll.; 1 (MEM), Chiapas, Sumidero Nat. Pk., 19 June 1989, S. Testa, P. K. Lago; 1 (EMEC), Veracruz, Cordoba, VII-6-1966, J. S. Buckett, M. R. & R. C. Gardner, colls.; 1 (WFBM), Veracruz, 250′, Los Tuxtlas Biol. Sta. UNAM, 20 May 1983, C. & L. O’Brien & G. Marshall; NICARAGUA: 1 (WOPC), Matagalpa, Selva Negra, beating, 1280 m, 20- V-2006, Weston Opitz.   
 Diagnosis. The only member of the genus displaying a combination of reddish pronotal ground color, infuscate or piceous antennal club, femora (usually) darkened at distal end, and scutellum (usually) pale.   
 Description. (Holotype). Length: 8.25 mm. Color: reddish-orange; antennal club infuscate; the following black: mandibles, eyes, antennomeres 2–3 (in part) and 4–9 (entirely), legs (except proximal 4/5 of profemora and mesofemora, and proximal 3/4 of metafemora); pronotum with piceous longitudinal macula at middle, extending from anterior margin to beyond posterior limit of transverse impression; elytra with 4 black maculae (Fig. 5) that do not attain epipleura laterally. Abdomen: ventrite 5 with posterior angles rather broadly rounded, hind margin with a rather shallow, broad, V-shaped emargination; ventrite 6 narrower than ventrite 5, with hind angles more acute, and posterior emargination deeper at middle; tergite 6 concave ventrally, with hind margin narrowly rounded, surpassing posterior margin of ventrite 6.   
 Variation. Length of specimens range from 7.6 mm–10.8 mm. The female pygidium has ventrite 5 feebly, broadly emarginate posteriorly, ventrite 6 rounded or subtruncate posteriorly, sometimes with a shallow indentation apically at middle, and tergite 6 with the hind margin broadly rounded or subtruncate, and sometimes slightly inflected at middle. Coloration and markings are highly variable in this species. Fig. 5–7 illustrate the range of variation, which includes individuals with no pronotal or elytral markings, some with expanded markings, and others with the elytra entirely black. Three specimens from Chiapas, Mexico have the scutellum darkened. Three other specimens have the elytral ground color testaceous rather than reddish or black.   
 Etymology. I take great pleasure in naming this species for Pete Townshend, guitarist and primary composer for the British rock group The Who.
description: Published as part of Rifkind, Jacques, 2017, New genera and species of mimetic Cleridae from Mexico and Central America (Coleoptera: Cleroidea), pp. 1-18 in Insecta Mundi 2017 (591) on pages 5-6, DOI: 10.5281/zenodo.5169515