Tomosvaryella patula Foldvari, Skevington & Motamedinia 2023, sp. nov.
Creators: name: Motamedinia, Behnam
name: Földvari, Mihaly
name: Skevington, Jeffrey H.
name: Kelso, Scott
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Pipunculidae
Tomosvaryella
Tomosvaryella patula

description: Tomosvaryella patula Földvari, Skevington & Motamedinia sp. nov.
urn:lsid:zoobank.org:act: 9F645DC0-660C-4972-92BE-FBA870D01E99
Figs 74A–E, 134, 149D, 155D
Diagnosis: This species can be recognized by the hind trochanter having a keel covered by short dark spines on the edge (Fig. 149D); hind metatarsus distinctly flattened; elongated surstyli, left surstylus with a lobe on inner margin in dorsal view (Fig. 74A); both gonopods with a triangular projection on inner margin; phallic guide with 3–5 dorsolateral spines (Fig. 74B, D–E).
Description: MALE: Body length: 2.9–3.1 mm. Head. Flagellum acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1/3. Thorax. Postpronotum pale, with 3–4 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side. Scutellum brownish pollinose and with 6–8 bristles along the distal edge. Dorsocentral bristles short, dark, somewhat longer along frontal edge, numerous pale bristles around postpronotum present. Halter, knob pale, stem brown. Legs. Mid coxa with 2–3 long dark bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1/4 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow, ventrally paler, last segment brown. Hind trochanter with shiny edge (keel) at 45 degrees to the long axis of trochanter, with short dark spines on the edge (Fig. 149D); weak ventrobasal spines (2) present on fore and missing on mid femur. Ventroapical row of spines on fore femur absent; 4–5 small, black spines on mid femur; hind femur without ventroapical spines, except 8–10 equally spaced bristles on distal half posteroventrally on a widened femur in the middle (almost like a small keel), longest up to 1.5 times the width of hind tibia at distal end (sometimes adpressed to femur). Subapical (distal) spines on first four tibiae present (short). Hind metatarsus distinctly flattened, as long as 2–5 combined and with strong bristles dorsally, scrub-like bristles ventrally (tarsomeres 1–2 only). Pulvilli shorter than last tarsal segment. Wing. Length: 3.0 mm. Upper side of basal costal cell with one long dark bristle. Fourth costal section 2–2.5 times as long as third costal section (Fig. 158E). Cross-vein r-m at middle of discal cell (sometimes slightly distal to middle). 2–3 short brown setulae on tegula. Abdomen. Viewed obliquely from front tergites brownish pollinose, tergite 1 silvery grey, silvery pollinose spots on tergites 4 and 5 (spot on t5 larger). Dispersed strong dark bristles on all tergites present, longest on tergite 5 in length up to 1/4 the width of hind tibia at distal end. Lateral bristles on first tergite present, 4–6 short dark bristles up to as long as 2/3 of hind femur’s width at base. Postabdomen in dorsal view: T6, S7 invisible; T5 0.5–0.7 times as long as ST8. Genitalia without dissection: ST8 large, rounded in dorsal view, brown and with a few distinct bristles, otherwise with velvet-like coverage; membranous area with round part extended dorsally and narrow part approaching epandrium, mostly directed posteriorly; epandrium brown, surstyli yellow. Genitalia. Genital capsule in dorsal view: epandrium slightly as long as wide (MLE:MWE = 1.08). Surstyli elongate, left surstylus with a distinct lobe on inner margin in middle and longer than right one, right surstylus with a small denticle on inner margin after basal fourth, both surstyli strongly bent to each other at apex (Fig. 74A). Genital capsule in ventral view: gonopods equal in height, extended towards surstyli, both gonopods with a triangular projection on inner margin; phallic guide with 3–6 dorso-dorsolateral spines, one is longer, distinct subepandrial sclerite (Fig. 74B); Genital capsule in lateral view: both surstyli broadened at base, rather straight, right surstylus moderately narrowed to apex, left surstylus bent towards sternite before apex (Fig. 74D–E); ejaculatory apodeme tube-like, bent, with a bulb in its middle (Fig. 74C).
FEMALE: Body length: 2.4 mm (without head). As male except for the following characters. Frons, eyes: NO HEAD. Pulvilli and claws about 2.5–3 times as long as last tarsal segment, smaller on metatarsi. Female abdomen with silver spots on sides of tergites 4–6. Ovipositor. Straight in ventral view, straight and robust in lateral view (yellow piercer, brown base), reaching middle of 3 rd segment; base silvery pollinose with dispersed bristles. LP:LB = 3.0. LDP:LPP = 5.0. (Fig. 155D).
Etymology: From the Latin patulus, spread out, broad, in reference to the distinctly flattened hind metatarsus.
Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Warrumbungle National Park, 31°14’S, 149°1’E, 19.X–3.XI.1997, J. Skevington, & S. Winterton, Malaise trap, JSS8439 (1♂, QM); PARATYPES: AUSTRALIA: New South Wales: Colo Heights, Putty Road, 33°23’S, 150°45’E, 6.II.1968, D.H. Colless, JSS8667 (1♂, ANIC); Culgoa National Park, 13.5km SW Cawwell Homestead (CGN1 M), 29°7’S, 146°56’E, Callitris, 30.I– 18. V.2010, QM Team, B. Shiebaan & R. Ohlsen, Malaise trap, CNC576617 (1♂, QM); Culgoa National Park, 8km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN2 M), 29°3’S, 146°60’E, Coolibah, 30.I–18. V.2010, C. Lambkin & R. Olsen, B. Shieban, Malaise trap, 19322, CNC575139 (1♂, QM); Bundjalung National Park; near Evans Head, 29°2’S, 153°26’E, 8.I.1997, C.L. Lambkin, Malaise trap, JSS6068 (1♀, QM); Queensland: Carnarvon National Park, Mount Moffatt Section, The Tombs, 25°5’S, 147°52’E, 10.X.2002, J. Skevington, hand collected, JSS12635 (1♂, CNC); Carnarvon National Park, Mount Moffatt Summit, 25°4’S, 148°3’E, 1097m, hilltop, 2. XII.1997, J. Skevington & C. Lambkin, hand collected, JSS1821 (1♂, QM); Carnarvon National Park, Mount Moffatt Summit, 25°4’S, 148°3’E, 1097m, hilltop, 13.X.2002, J. Skevington, JSM 466, CNC484088 (1♀, CNC); Isla Gorge National Park, 25°11’S, 149°58’E, 320m, hilltop near lookout, 13.XI.1999, J.&A. Skevington, hand collected, JSS7256 (1♂, QM); Western Australia: West of Norseman, 32°11’S, 121°38’E, Eucalyptus woodland, 1–17.XI.2003, C. Lambkin & J. Recsei, Malaise trap, JSS16212 (1♂, ANIC).
Distribution: Australia (New South Wales, Queensland, Western Australia) (Fig. 134).
Notes: Intraspecific genetic distance ranges from 0.3% to 1.5%. This species is genetically most similar to T. unda sp. nov. (1.6–8.3% pairwise divergence) (Supplementary file 3). This is a forest species that is apparently widespread but rare. It has been found hilltopping.
description: Published as part of Motamedinia, Behnam, Földvari, Mihaly, Skevington, Jeffrey H. & Kelso, Scott, 2023, Revision of Australian Tomosvaryella Aczél (Diptera: Pipunculidae) with description of 100 new species, pp. 1-271 in Zootaxa 5599 (1) on pages 164-165, DOI: 10.11646/zootaxa.5599.1.1, http://zenodo.org/record/14970961