Clivina bilobata Balkenohl, 2025, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina bilobata

description: Clivina bilobata sp. nov.
(Figs 11, 46, 81)
Type material. Holotype: ♂, with labels and data: white, black printed “ Malaysia, Sabah, Batu Punggul Resort env., 24.VI.-1.VII.1996, 11c. vegetation debris and forest floor litter accumulated around large trees near river” (NHMW).
Description. Measurements. Body length 5.77 mm, width 1.59 mm, ratio L/W of pronotum 0.89, ratio L/W of elytra 1.99. Colour glossy, piceous. Supraantennal plate indistinctly translucent. Antennae, mouthparts, intermediate and hind legs rufous-fuscous.
Head a sixth narrower than pronotum. Clypeus with central part straight, with slightly projecting teeth laterally, separated from clypeal-wing by distinct notch; wing distinct, rounded, wider but not as projecting as clypeal-teeth, separated from supraantennal plate by distinct notch, supraantennal plate with margin laterally obtuse-angular, clypeus, wing, and supraantennal plate reflexed margined. Supraantennal plate extended posteriorly into a convex keel, separated from wing by flat furrow. Clypeus continuing into frons without furrow, slightly convex, with scattered small punctures. Frons with pore at middle, separated from supraantennal plate and supraorbital carina by sharp furrow, with clypeal seta in front; supraorbital carinae moderately long, narrow, widened posteriorly, with two supraorbital setigerous punctures at middle of eye and at end of the gena-level. Neck constriction closed, formed by a step from frons. Eye relatively large, gena small. Antenna elongate, reaching up middle of pedunculus. Labrum excised anteriorly, conspicuously bilobate, isodiametrically reticulated, seven-setose. Mentum isodiametrically reticulated, with oblique wrinkles, lobe with lateral margin latero-basally straight, with obtuse angle, latero-apically straight, with apex rounded, medially straight to cleft, median tooth wide, much lower than lobes, at apex bent dorsally.
Pronotum (Fig. 46) with disk flattened in lateral view, moderately convex in frontal view. Shape cordate, wider than long. Reflexed lateral margin smooth, slightly convex, widest at middle; anterior angle distinctly angled, posterior angle nearly completely rounded off, visible by knob-like tooth; lateral channel moderately narrow, isodiametrically reticulated, continuing up to base as narrow, convex margin. Median line engraved, complete, joining anterior transverse line. Anterior transverse line distinct, complete. Disk nearly completely covered with small sized punctures, with some transverse wrinkles, with isodiametric reticulation at lateral margin and at base, with posterior setigerous puncture removed from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 45°. Basal channel on flange moderately narrow, rugose, with sharp carina.
Elytron (Fig. 11) with disk indistinctly convex in lateral view, distinctly convex in frontal view. Shape of elytra elongate, about two-fifth longer than wide, with lateral margin nearly parallel, maximum width at middle. Humerus distinct, rounded. Reflexed lateral margin slightly crenulate behind humerus. Row of setigerous punctures in the lateral channel regular in size, slightly more distant at middle. Scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, impunctate, one to four free at base, five and six joining at humerus, one and two ending free at apex, three and four and five and six joining at apex. Intervals moderately convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as very short carina, interval seven with rim at apex, eight carinate at apex. Surface of interval one to six glossy, interval seven and eight with distinct isodiametric reticulation, apical quarter of elytron isodiametrically reticulated at all intervals.
Hind wing fully developed.
Lower surface. Proepisternum isodiametrically reticulated, with fine wrinkles laterally. Sternite of abdomen with isodiametric reticulation, more scaly medially. Last visible sternite with the two marginal setigerous punctures widely separated, with slight projection posteriorly at middle where it is flattened, with narrow marginal furrow at middle.
Legs. Protibia slender, with longitudinal to irregular reticulation, with narrow sulcus dorsally, with three spines, movable spur nearly straight, basal tarsomere slender at base, carinate, with acute tubercle and two wide setae at middle of tarsomere. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg widened.
Male genitalia (Fig. 81). Median lobe distinctly arcuate in basal part, straight in apical half, apically regularly tapered, somewhat flattened. Parameres moderately slender, with small apophysis, slightly distorted, hyaline apically, with one minute seta at apex.
Female gonocoxites and epipleurite unknown.
Variation not observed.
Distribution. Known from Sabah on the island of Borneo.
Diagnosis. A medium sized piceous species with peltate pronotum and intervals seven and eight of the elytron reticulated. It is the only species with peltate pronotum and having the labrum seven-setose. In addition, the frons is covered by medium sized punctures. Moreover, it is the only species of the group with a distinctly bilobed labrum.
Etymology. The name refers to the labrum which is distinctly bilobed (Latin: bilobata).
description: Published as part of Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), pp. 1-70 in Zootaxa 5592 (1) on pages 21-22, DOI: 10.11646/zootaxa.5592.1.1, http://zenodo.org/record/14953674