Slaterocoris atritibialis
Creators: name: Schwartz, Michael D.
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Miridae
Slaterocoris
Slaterocoris atritibialis

description: Slaterocoris atritibialis (Knight)   Figures 13, 30; plate 1; map 6  
 Strongylocoris stygicus: Van Duzee: 1921: 135 (disc., in part).   
 Strongylocoris atratus: Van Duzee: 1921: 135 (disc., in part).   
 Strongylocoris atritibialis Knight, 1938: 2, fig. 6 (orig. desc.).   
 Slaterocoris atritibialis: Kelton, 1968: 1122 fig. 3 (disc.); Knight, 1970: 244, fig. 12, (disc.); Reid, 1974: 239 (parasit.); Reid et al., 1976: 561 (biol.); Messina, 1978: 139 (biol.); Henry and Smith, 1979: 215 (list); Wheeler et al., 1983: 142 (list); Henry and Wheeler, 1988: 448 (catalog); Schuh, 1995: 197 (catalog); Cappuccino 1987: 1646 (biol.); Wheeler 2001: 153, 284 (biol.); Henry et al., 2005: 62 (list); Forero, 2008: 152, 155–156 (disc.).   
 DIAGNOSIS: Recognized among breviatus group species by the arcuate costal margin of both sexes (pl. 1), the black antennal segments, hind tibia, and tarsomeres, the female usually having the fore- and middle tibiae dusky yellowish brown, the male genitalia with the tergal process truncate with two lateral points (fig. 30J), apical spines of the right paramere tightly grouped, moderately long with basalmost apical spine stouter than in others (fig. 30C–G), dorsal lobe of endosomal spicule bifurcate, larger (lateral) ramus flattened and marginally serrate, smaller (medial) ramus usually one-fourth length of lateral ramus (fig. 30A), sometimes much smaller (fig. 30B); ventral lobe of endosomal spicule evenly curved at recurved middle region, apex reaching to base of dorsal lobe. Female with the head and vertex wider and tarsomeres usually darker than in the female of S. stygicus and S. breviatus.   
 REDESCRIPTION: Male: Large, ovate, length 4.00–4.88, width 1.85–2.38 (pl. 1). COLORATION: Shining black, including antenna and legs, except diffusely pale on apices of femora (fig. 13); in female fore- and middle tibiae often yellowish brown to reddish black and antennal segment II pale medially. VESTITURE: Subglabrous, sparsely distributed, short, suberect, brown simple setae. STRUCTURE: Frons striate; labium reaching base of middle coxa; costal margin arcuate. GENITALIA: Tergal process: Apical points conspicuous (fig. 30J). Endosomal spicule: Ventral lobe long, marginal serration strong; dorsal lobe with lateral ramus long, flat, relatively wide, apex truncate, lateral margin strongly serrate, medial ramus short, with variable length (fig. 30A), sometimes present as large spine (fig. 30B). Right paramere: Apical spines tightly grouped, moderately long, discrete, splayed, with basalmost spine stouter than and slightly removed from others (fig. 30C–G); basal lobe with serrate region produced, somewhat constricted, strongly serrate on posterior surface (fig. 30C–G). Left paramere: Long; apex somewhat spatulate (fig. 30H, I).   
 Female: Large, ovate; length 4.00–4.95, width 2.03–2.63 (pl. 1). COLORATION: Antennal segment II, fore- and middle tibiae often paler than in male. STRUCTURE: Costal margin strongly arcuate than male, base of embolium wide. GENITALIA: As in species group except: First gonapophyses: Right greater than left at overlap. Second gonapophyses: Anterior medial surface strongly convex. Interramal sclerite: Posteromedial portion with tumid process abutting ovipositor bulb.   
 HOSTS: The predominance of records indicate that Solidago species (predominantly S. altissima and S. canadensis) are breeding hosts. Five other genera of Asteraceae are also noted from far fewer localities: Ambrosia sp., Artemisia (sp. and A. cana), Aster sp., Eurybia macrophylla, and Leucanthemum vulgare. The Artemisia records are from five localities in Colorado, Idaho, North Dakota, and South Dakota. Only the collection event at American Falls, Idaho (00113953) was from a series of specimens (an additional 78, 6♀) and may indicate that in the arid western states this unidentified species of Artemisia is also a breeding host or at least is a significant feeding host. Wheeler (2001) reported personal observations of foliar chlorosis on Artemisia vulgaris as a result of feeding by S. atritibialis. The scattered sampling of nonasteraceous plant occurrences are all considered to be sitting records.   
DISTRIBUTION: From westcentral British Columbia spanning east across the Prairie Provinces (as far north as 56.86 N in Alberta) to the Great Lakes region, southern Quebec, and Maine. Widely distributed across the eastern and central U.S. as far south as northern Georgia and Alabama, then west to Oklahoma and in the upper Great Plains as far south as Nebraska. Widely distributed in the Rocky Mountain cordillera from Cloudcroft, New Mexico, in the south, north through Colorado to Montana, then west across the Snake River plain to eastern Oregon (map 6).  
 COLLECTION SUMMARY: 1760 specimens (54 % with USI numbers) from 767 collection events. Phenologically suitable collection times occur in April and May for southern localities, June to August for the majority of localities, with the latest collection events in September from northern localities.   
 DISCUSSION: In western females, the body width is not as great and the curvature of the costal margin is not as arcuate as in eastern females of S. atritibialis. Such specimens can be confused with female S. stygicus, but ultimately distinguished by the mostly dark fore- and middle tibiae and dark antennal segment II. These appendages are usually pale in the latter species. When antennal segment II in the female of these species is partially pale, then the pale region is somewhat more diffuse in S. atritibialis and more starkly contrasting in S. stygicus. The length of segment II in the female tends to be shorter on average in S. atritibialis (1.09, 0.93–1.25) than in S. stygicus (1.16, 1.05– 1.28) and the vertex of the female is wider in S. atritibialis (0.72, 0.62–0.78) than in S. stygicus (0.63, 0.58–0.66). Western specimens, especially those from Wyoming, tend to have somewhat longer, more densely distributed dorsal vestiture than eastern specimens. Some males from Alberta and British Columbia have tibial coloration more typical of S. stygicus with fore- and middle tibiae pale and hind tibia black; however, the male genitalia, particularly the endosomal spicule, is identical to male S. atritibialis with all black tibiae.   
 Slaterocoris atritibialis was listed as a facultative predator of Uroleucon caligatum (Richards) and U. nigrotuberculatum (Olive) (Sternorrhyncha: Aphididae) colonies feeding on Solidago canadensis var. (as S. altissima) in central New York (Cappuccino, 1987). It was the only predaceous plant bug species in Solidago arthropod community studies. In laboratory colonies, originating from goldenrod, S. atritibialis was also observed to feed on the eggs of Ophraella sexvittata (Leconte) (Coleoptera: Chrysomelidae) (Messina, 1978). The phenology of S. atritibialis in several Solidago studies (Reid, 1974; Reid et al., 1976; Messina, 1978) was earlier than for the sympatric S. breviatus and S. stygicus.   
 HOLOTYPE:  USA: New York: Tompkins Co.:  Ithaca [42.440566 N 76.496946 W], 23 Jun 1920, H.H. Knight, 18 (AMNH _ PBI 00134026) (USNM).   
 OTHER SPECIMENS EXAMINED: CANADA: Alberta: Armena, 53.12296 N 112.951986 W, 750 m, 17 Aug 1920, C.E. Hendrickson, 1♀ (00134315) (USNM). Banff, 51.166666 N 115.566666 W, 1615 m, 28 Sep 1925, O. Bryant, 1♀ (00134592) (USNM). Barrhead, 54.117166 N 114.381816 W, 640 m, 11 Jul 1938, R.W. Salt, Trifolium sp. (Fabaceae), 1♀ (00113862) (CNC). Brooks, 50.566666 N 111.90006 W, 13 Jun 1957, Brooks and McNay, 18 (00113851) (CNC). Conrad, 49.526 N 111.976 W, 21 Jun 1951, L.A. Konotopetz, 18 (00113847) (CNC). Coutts, 49.00456 N 111.964056 W, 1073 m, 15 Jun 1952, A.R. Brooks, 28 (00113846, 00115511), 2♀ (00113863, 00317195) (CNC). Cypress Hills Provincial Park, Elkwater Park Campground at Horse[shoe] Canyon trailhead, 49.659896 N 110.317946 W, 1282 m, 03 Sep 1993, M.D. Schwartz, Solidago spp., 1♀ (00113869) (CNC). Drumheller, 51.466666 N 112.70006 W, 18 Jun 1957, Brooks and McNay, 18 (00242558) (CAS), 18 (00113853) (CNC). Elkwater Lake, Cypress Hills Provincial Park, 49.662936 N 110.301676 W, 1219 m, 19 Jul 1956, O. Peck, 1♀ (00113858) (CNC); 20 Jul 1956, O. Peck, 1♀ (00113857) (CNC). Elkwater Park, Cypress Hills Provincial Park, 49.636 N 110.20006 W, 1427 m, 07 Jul 1952, L.A. Konotopetz, Solidago sp., 28 (00113849, 00317194), 1♀ (00317197) (CNC); 14 Jul 1952, L.A. Konotopetz, 1♀ (00113865) (CNC); 29 Jul 1952, L.A. Konotopetz, 18 (00113845) (CNC). Grande Prairie, 55.166666 N 118.80006 W, 26 Jul 1961, A.R. Brooks, 1♀ (00113859) (CNC). Hines Creek, 56.233336 N 118.60006 W, 16 Jun 1961, A.R. Brooks, 18 (00113855) (CNC). Hobbema, 52.832746 N 113.429726 W, 794 m, 11 Jul 1939, R.W. Salt, Trifolium sp. (Fabaceae), 1♀ (00113861) (CNC). Kananaskis Hwy, 50.915556 N 115.141666 W, 25 Jul 1973, L.A. Kelton, 18 (00113318) (CNC). Lethbridge, 49.70006 N 112.833336 W, 28 Jul 1928, G.F. Mason, 1♀ (00113864) (CNC); 07 Jul 1956, O. Peck, 1♀ (00113868) (CNC). Medicine Hat, 50.033336 N 110.683336 W, 14 Jun 1930, J.H. Pepper, 1♀ (00113866) (CNC); 23 Jul 1930, J.H. Pepper, 1♀ (00113860) (CNC). Milk River, 49.133336 N 112.083336 W, 18 Jul 1929, J.H. Pepper, 1♀ (00113867) (CNC). Nevis, 52.330936 N 113.03696 W, 798 m, 27 Jul 1973, L.A. Kelton, 18 (00318771) (CNC). Nordegg, 52.466666 N 116.083336 W, 29 Jul 1921, J. McDunnough, paratype, 1♀ (00113856) (CNC); 31 Jul 1921, J. McDunnough, paratype, 18 (00113842) (CNC); 06 Aug 1923, J. McDunnough, paratype, 18 (00113843) (CNC). North Star, 56.854116 N 117.630846 W, 487 m, 17 Jul 1940, C.L. Neilson, 18 (00113854) (CNC). Onefour, 49.066666 N 110.456 W, 14 Jun 1952, L.A. Konotopetz, Aster sp., 18 (00113848), 1♀ (00317198) (CNC). Red Deer, 52.270246 N 113.804696 W, 856 m, 25 Jun 1957, Brooks and McNay, 18 (00113850) (CNC). Stettler, 52.333336 N 112.683336 W, 03 Aug 1957, A. and J. Brooks, 18 (00113852) (CNC). Waterton Lakes National Park, Belly River Campground, 49.02496 N 113.686736 W, 1383 m, 07 Jul 2005, D. Langor, 18 (00113750) (CNC). Waterton Lakes National Park, Dardanelles, 49.066726 N 113.850556 W, 1280 m, 31 Jul 1967, David J. and M.A. Larson, 18 (00113751) (WLNP). Waterton Park, 49.056 N 113.916666 W, 04 Jul 1970 – 06 Jul 1970, L.A. Kelton, 18 (00113844) (CNC). British Columbia: Anarchist Mountain, Osoyoos, 49.033336 N 119.466666 W, 13 Jul 1970, L.A. Kelton, 1♀ (00317196) (CNC). Chilcotin, 52.115496 N 122.562386 W, 1020 m, 12 Jul 1930, unknown, 18 (UBC); 15 Jul 1978, L.A. Kelton, 18 (CNC). Eastgate, 49.13756 N 120.61256 W, 15 Jul 1986, unknown, 18 (UBC). Fernie, 49.50006 N 115.066666 W, 16 Aug 1935, J.F. Brimley, Fagus sp. (Fagaceae), 18 (00113840) (CNC); 23 Jul 1959, L.A. Kelton, 18 (00113841) (CNC). Houston, 54.40006 N 126.666696 W, 19 Aug 1967, unknown, 18 (UBC). Lizard Creek, Fernie, 49.466756 N 115.066676 W, 984 m, 07 Jul 1949, Hugh B. Leech, 18 (00121679) (CAS). Midday Val[- ley]., Merritt, 49.983336 N 120.933336 W, 736 m, 24 Jul 1923, R. Hopping, Pinus ponderosa Laws. (Pinaceae), 1♀ (00134295) (USNM). Nicola, 50.176 N 120.676 W, 21 Aug 1932, unknown, 18 (UBC). Osoyoos, Anarchist Mountain, 49.033336 N 119.333336 W, 13 Jul 1970, L.A. Kelton, Solidago sp., 18 (00113581) (CNC). Prince George, 53.916666 N 122.766666 W, 579 m, Aug 1929, unknown, 18 (UBC). Quesnel, 52.983336 N 122.483336 W, 07 Jul 1949, unknown, 18 (UBC). Rolla, 55.891996 N 120.144576 W, 664 m, 11 Jul 1927, P.N. Vroom, 18 (00113839) (CNC). Tyhee Lake [Park], 54.70006 N 127.033296 W, 24 Jul 1983, unknown, 18 (CNC). Williams Lake, 52.116666 N 122.156 W, 09 Jul 1930, unknown, 18 (UBC). Manitoba: 4.7 km E of Tolstoi off Rt 209, 49.07536 N 96.743626 W, 292 m, 07 Jul 1990, M.D. Schwartz and R.G. Foottit, 18 (00329797), 1♀ (00113913) (CNC). 6.4 km W of Russell on Rt 16, 50.771536 N 101.381386 W, 533 m, 18 Jul 1990, M.D. Schwartz and R.G. Foottit, Symphoricarpos occidentalis Hook. (Caprifoliaceae), 18 (00113909) (CNC). 30 mi N of Roblin, 51.667036 N 101.356 W, 13 Jul 1954, Brooks, Wallis, 1♀ (00113912) (CNC); 14 Jul 1954, A.R. Brooks, Wallis, 18 (00113897) (CNC). Boissevain, 49.233336 N 100.056 W, 17 Jul 1953, A.R. Brooks, L.A. Kelton, 18 (00113899) (CNC); 24 Jul 1953, Brooks and Kelton, 18 (00113898) (CNC). Brandon, 49.833336 N 99.956 W, 25 Jun 1970, unknown, 18 (00113910) (CNC). Carberry, 49.866666 N 99.356 W, 23 Jun 1953, Brooks and Kelton, 18 (00113907) (CNC); 30 Jul 1953, Brooks and Kelton, 18 (00113906) (CNC); 09 Aug 1953, Brooks and Kelton, 1♀ (00113916) (CNC). East Braintree, 49.616666 N 95.616666 W, 30 Jun 1972, L.A. Kelton, Solidago sp., 18 (00113911) (CNC). Oak Lake, 49.666666 N 100.756 W, 09 Jul 1953, Brooks and Kelton, 1♀ (00113915) (CNC); 11 Jul 1953, Brooks and Kelton, 1♀ (00113914) (CNC). Pembina, 49.755696 N 97.150896 W, 230 m, unknown, 18 (00164927) (USNM). Russell, 50.783336 N 101.283336 W, 17 Jul 1954, Brooks and Wallis, 18 (00113896) (CNC). Souris, 49.616666 N 100.256 W, 23 Jul 1953, A.R. Brooks, 18 (00113908) (CNC). Turtle Mountain, 496 N 100.333336 W, 17 Jul 1953, A.R. Brooks, L.A. Kelton, 18 (00113900) (CNC). Turtle Mountain Forest Reserve, International Peace Gardens, 496 N 100.333336 W, 17 Jul 1958, R.B. Madge, 1♀ (00114013) (CNC). Virden, 49.856 N 100.933336 W, 08 Jul 1953, Brooks and Kelton, 18 (00113905) (CNC); 09 Jul 1953, Brooks and Kelton, 18 (00113903) (CNC); 10 Jul 1953, Brooks and Kelton, 18 (00113902) (CNC); 12 Jul 1953, Brooks and Kelton, 18 (00113904) (CNC); 14 Jul 1953, Brooks and Kelton, 18 (00113901) (CNC). Ontario: 5 mi S of Crozier, 48.55096 N 93.515796 W, 343 m, 15 Jun 1960, Kelton and Whitney, 1♀ (00114019) (CNC). 6 mi S of Devlin, 48.532356 N 93.673256 W, 339 m, 28 Jun 1960, Kelton, Whitney, 18 (00113973) (CNC). 10 mi N of Emo, 48.761236 N 93.833336 W, 28 Jun 1960, Kelton, Whitney, 18 (00113979) (CNC). Arkell, 43.535216 N 80.156246 W, 355 m, 17 Jun 1991, J. Taylor, 28 (00329831–00329832) (DEBU). Ausable River, 43.07836 N 81.758486 W, 203 m, 10 Jun 1976, J.F. Fortin, 1♀ (00329818) (DEBU). Bergland, 48.951726 N 94.386866 W, 325 m, 22 Jun 1960, Kelton and Whitney, 18 (00113978) (CNC). Bluevale, 43.856136 N 81.250366 W, 327 m, 10 Jun 1962, Kelton and Thorpe, 1♀ (00114024) (CNC). Brockville, 44.60296 N 75.660736 W, 87 m, 18 Jun 1963, J.R. Vockeroth, 1♀ (00114047) (CNC). Bryanston, 43.126 N 81.276 W, 20 Jun 1962, Kelton and Brumpton, 18 (00113998) (CNC). Carp, 45.356 N 76.033336 W, 22 Jun 1981, D.J.E. Brown, (Poaceae), 18 (00113924), 1♀ (00113716) (CNC). Chatham, 42.40006 N 82.183336 W, 15 Jun 1928, G.M. Stirrett, 1♀ (00114046) (CNC). Clinton, 43.60006 N 81.533336 W, 09 Jun 1961, L.A. Kelton, 2♀ (00114006, 00114022) (CNC). Coldstream, 43.019156 N 81.499766 W, 248 m, 01 Jun 1922, A.A. Wood, 18 (00114041) (CNC). Dashwood, 43.356 N 81.636 W, 12 Jul 1962, Kelton and Thorpe, Salix sp. (Salicaceae), 18 (00113995) (CNC). Dundas, 43.266666 N 79.933336 W, 19 Jun 1978, M. Lichtenberg, 28 (00329822– 00329823) (DEBU); 20 Jun 1979, B. Merchant, 1♀ (00329824) (DEBU). Eagle, 42.567376 N 81.558286 W, 200 m, 26 Jun 1985, G.G.E. Scudder, 1♀ (00114015) (CNC). Eden, 42.80006 N 80.756 W, 02 Jul 1962, H. Blanchard, 1♀ (00114012) (CNC). Elora, 43.682426 N 80.433646 W, 384 m, 27 Jun 1975, S. Allan, 18 (00382310) (DEBU). Erin, 43.756 N 80.116666 W, 25 Jun 1979, J. Ernst, 1♀ (00329828) (DEBU). Fisher Glen, 45.353886 N 75.705666 W, 84 m, 16 Jun 1931, G.S. Walley, 1♀ (00113684) (CNC). Fonthill, 43.033336 N 79.283336 W, 26 Jun 1915, H.G. Crawford, 1♀ (00114011) (CNC); 08 Jun 1962, Kelton and Thorpe, 18 (00113971), 1♀ (00114025) (CNC). Fort Frances, 48.616666 N 93.416666 W, 28 Jun 1960 – 29 Jun 1960, Kelton and Whitney, 1♀ (00114023) (CNC). Foxboro, 44.246236 N 77.43396 W, 111 m, 24 Jun 1970, D.G. Reid, 18 (00114038) (CNC); 26 Jun 1971, D.G. Reid, Solidago sp., 18 (00114039) (CNC). Freelton, 43.404226 N 80.040356 W, 280 m, 26 May 1962, L.A. Kelton, 18 (00113988) (CNC). Fuller, 44.40006 N 77.426 W, 14 Jun 1968, C.C. Loan, Solidago sp., 18 (00114031) (CNC); 26 Jun 1968, C.C. Loan, Solidago sp., 18 (00114032) (CNC); 22 Jun 1970, D.G. Reid, 18 (00114036) (CNC); 09 Jun 1971, D.G. Reid, 18 (00114037) (CNC); 14 Jun 1971, D.G. Reid, 18 (00114033) (CNC); 16 Jun 1971, D.G. Reid, 18 (00114034) (CNC); 02 Jul 1971, D.G. Reid, 18 (00114035) (CNC). Galt, 43.362036 N 80.332086 W, 304 m, 19 Jun 1961, Kelton and Brumpton, 18 (00113985) (CNC). Grimsby, 43.192716 N 79.562266 W, 93 m, 19 Jun 1979, K.L. Bailey, 18 (00382311) (DEBU). Guelph, 43.556 N 80.256 W, 323 m, 21 Jun 1976, E.A. Innes, 1♀ (00329810) (DEBU); 30 May 1977, W.A. Attwater, 18 (00329800), 3♀ (00329801–00329803) (DEBU); 07 Jun 1977, K.M. Ralph, 1♀ (00329809) (DEBU); 11 Jun 1977, P.R. Heels, 1♀ (00329808) (DEBU); 01 Jun 1979, G.W. Annette, 18 (00329807) (DEBU); 13 Jun 1979, B. Merchant, 18 (00329806) (DEBU); 13 Jun 1979, D. Lewis, 18 (00329804), 1♀ (00329805) (DEBU). Hamilton-Wentworth Region, Flamborough, 5th Concession, Lawson Farm, 43.316116 N 80.040556 W, 251 m, 01 Jul 2003, M. Buck, 18 (00329798 [cat debu00224719]) (DEBU). Hanlon [Creek] Conservation Area, Guelph, 43.502626 N 80.224516 W, 320 m, 14 Jun 1984 – 21 Jun 1984, K. Gilbert, 1♀ (00329811) (DEBU). Harrow, 42.033336 N 82.916666 W, 09 Jun 1993, B. Larson, 1♀ (00329819) (DEBU). Hespeler, 43.433336 N 80.316666 W, 18 Jun 1961, Kelton and Brumpton, 18 (00113994) (CNC). Hillsdale, 44.58466 N 79.765566 W, 253 m, 29 Jun 1962, G. Thorpe, 1♀ (00114026) (CNC). Hilton Beach, 46.256 N 83.883336 W, 08 Jul 1961, G. Brumpton, 18 (00113986) (CNC). Ingleside, 45.000016 N 75.000056 W, 86 m, 27 Jun 1977 – 29 Jun 1977, L.A. Kelton, 18 (00114040) (CNC). Ipperwash, 43.208056 N 81.976396 W, 185 m, 11 Jul 1962, Kelton and Thorpe, Solidago sp., 1♀ (00114021) (CNC). Kapuskasing, 49.416666 N 82.433336 W, 18 Jul 1961, G. Brumpton, 18 (00113977) (CNC). Kincardine, 44.166666 N 81.633336 W, 13 Jun 1961 – 14 Jun 1961, Kelton and Brumpton, 18 (00114000) (CNC); 28 May 1962, L.A. Kelton, Thorpe, 18 (00113970) (CNC). Kingsville, 42.033336 N 82.756 W, 23 May 1962, L.A. Kelton, Thorpe, 18 (00113974) (CNC); 18 Jun 1962, Kelton and Thorpe, 18 (00113990) (CNC); 19 Jun 1962, Kelton and Thorpe, 18 (00113980) (CNC). Kirkland Lake, 48.156 N 80.056 W, 17 Jul 1961, G. Brumpton, 18 (00113984) (CNC). Leamington, 42.056 N 82.583336 W, 07 Jun 1929, L.J. Milne, 18 (00113922) (CNC); 09 Jun 1929, L.J. Milne, 18 (00113920) (CNC); 10 Jun 1929, L.J. Milne, 18 (00113921) (CNC); 10 Jun 1929, G.S. Walley, 18 (00113918) (CNC); 11 Jun 1929, G.S. Walley, 18 (00113919) (CNC); 05 Jun 1961 – 06 Jun 1961, L.A. Kelton, 18 (00113993) (CNC). London, 42.983336 N 81.256 W, 24 Jun 1985, G.G.E. Scudder, 18 (00114004) (CNC). Marmora, 44.483336 N 77.683336 W, 07 Jul 1952, C. Boyle, 1♀ (00114048) (CNC); 10 Jul 1952, C. Boyle, 18 (00114045) (CNC); 14 Jul 1952, J.R. McGillis, 1♀ (00114010) (CNC); 19 Jul 1952, J.R. McGillis, 18 (00114044) (CNC). Meaford, 44.60006 N 80.583336 W, 13 Jun 1962, Kelton and Thorpe, 18 (00113991) (CNC). Mount Pleasant, 44.233336 N 77.033336 W, 10 Jul 1958, L.A. Kelton, 18 (00113972) (CNC). New Glasgow, 42.516666 N 81.633056 W, 27 Jun 1961, Kelton and Brumpton, 18 (00111412), 28 (00113989, 00113999) (CNC), 18 (00121793) (UCB). North Bay, 46.30006 N 79.456 W, 14 Jul 1961 – 15 Jul 1961, G. Brumpton, 18 (00114017) (CNC). Norway Point, Lake of Bays, 45.256 N 79.056 W, 28 Jun 1922, J. McDunnough, 18 (00113923) (CNC), 18 (00382300), 1♀ (00382313) (DEBU); 12 Jul 1922, J. McDunnough, 18 (00113925) (CNC). Norwich, 42.983336 N 80.60006 W, 19 Jul 1962, Kelton and Thorpe, Solidago sp., 18 (00114020) (CNC). Ohsweken [Six Nations of the Grand River First Nation], 43.064556 N 80.114916 W, 207 m, 08 Jun 1979, D. Morris, 18 (00329829), 1♀ (00329830) (DEBU). Ojibway Prairie [Provincial Nature] Reserve, 42.267226 N 83.074236 W, 179 m, 17 Jun 1980, K.H. Harvey, 18 (00329833) (DEBU); 19 Jun 1980, J.D. Cashaback, 1♀ (00329834) (DEBU). Ottawa, 45.390796 N 75.703246 W, 71 m, 29 Jun 1912, E.P. Van Duzee, 18 (00242559) (CAS); 28 Jun 1951, E.H.N. Smith, (Poaceae), 1♀ (00113712) (CNC). Ottawa, [E of] Ryder St., Green Belt, 45.383056 N 75.634066 W, 83 m, 28 Jun 1999, M.D. Schwartz, Solidago canadensis, 18 (00113917) (CNC). Oxenden, 44.764936 N 81.092156 W, 212 m, 30 Jun 1962, G. Thorpe, 1♀ (00114018) (CNC). Parry Sound, 45.333336 N 80.033336 W, 10 Jul 1915, H.S. Parish, paratypes, 38 (00134198–00134200), 1♀ (00134201) (USNM); 10 Jul 1915, unknown, paratypes, 2♀ (00134202–00134203) (USNM); 15 Jul 1915, unknown, 38 (00134192–00134194), 3♀ (00134195–00134197) (USNM). Pitopiko River Picnic Area on Rt 11 between Longlac and Hearst, 49.770516 N 83.540216 W, 217 m, 21 Jul 1990, M.D. Schwartz, Solidago sp., 18 (00113926) (CNC). Point Pelee, 41.966666 N 82.516666 W, 28 Jun 1961 – 29 Jun 1961, L.A. Kelton, G. Brumpton, 18 (00113987) (CNC). Port Burwell, 42.656 N 80.816666 W, 03 Jul 1962, G. Thorpe, 18 (00113997) (CNC). Prince Edward County, no specific locality, 446 N 77.256 W, 20 Jul 1927, J.F. Brimley, 18 (00114043) (CNC); 24 Jun 1953, J.F. Brimley, 1♀ (00114049) (CNC); 15 Jun 1955, J.F. Brimley, Solidago sp., 1♀ (00114051) (CNC); 04 Jul 1956, J.F. Brimley, 1♀ (00114050) (CNC). Puslinch, 43.432586 N 80.085926 W, 294 m, 10 Jun 1962, L.A. Kelton, Thorpe, 18 (00113992) (CNC); 30 May 1982, G.M. Grant, 2♀ (00329826–00329827) (DEBU); 28 Jun 1982, G.M. Grant, 1♀ (00329825) (DEBU). Queenston, 43.166666 N 79.056 W, 24 Jun 1961, L.A. Kelton, G. Brumpton, 18 (00113983) (CNC). Rainy River, 48.716666 N 94.566666 W, 22 Jun 1960, Kelton and Whitney, Solidago sp., 18 (00113976) (CNC). Sault Ste. Marie, 46.526 N 84.336 W, 05 Jul 1961 – 06 Jul 1961, G. Brumpton, 18 (00113975) (CNC). Simcoe, 42.833336 N 80.30006 W, 23 Jun 1915, H.G. Crawford, 18 (00114001) (CNC); 02 Jul 1915, H.G. Crawford, 18 (00114003) (CNC); 12 Jul 1915, H.G. Crawford, 18 (00113969) (CNC); 13 Jun 1931, G.S. Walley, 18 (00113996) (CNC). Smithville, 43.10006 N 79.533336 W, 19 Jun 1979, D. Lewis, 18 (00329815), 2♀ (00329816–00329817) (DEBU). South Gloucester, Blossom Park, Greenbelt, 45.343796 N 75.646356 W, 105 m, 09 Jun 1991, M.D. Schwartz, Solidago sp., 28 (00318762–00318763) (CNC). St. Lawrence Islands National Park, Grenadier Island Center, 44.389136 N 75.897936 W, 73 m, 10 Jun 1975, E. Wilson, 18 (00114029) (CNC); 18 Jun 1975, E. Wilson, 18 (00114028) (CNC); 16 Jul 1975, E. Wilson, 18 (00114030) (CNC). Strathroy, 42.956 N 81.616666 W, 25 Jul 1917, H.G. Crawford, 18 (00114002) (CNC). Sundridge, 45.766666 N 79.40006 W, 13 Jul 1961, G. Brumpton, 18 (00113981) (CNC). Thamesford, 43.076 N 816 W, 29 Jun 1961, Kelton and Brumpton, 1♀ (00114016) (CNC). Thedford, 43.162166 N 81.860036 W, 207 m, 03 Jun 1957, D.H. Pengelly, 38 (00329812–00329814) (DEBU). Tillsonburg, 42.866666 N 80.733336 W, 11 Jul 1958, L.A. Kelton, 2♀ (00113717, 00114009) (CNC); 20 Jun 1962, Kelton and Thorpe, 1♀ (00114008) (CNC). Vienna, 42.686 N 80.80006 W, 15 Jul 1961, L.A. Kelton, 1♀ (00114007) (CNC). Wallbridge, 44.214156 N 77.495326 W, 116 m, 09 Jun 1957, unknown, 18 (00114042) (CNC). Quebec: Gatineau National Park, 46.50006 N 766 W, 09 Jun 1981, L.A. Kelton, (Poaceae), 18 (00113932) (CNC). Gatineau National Park, Harrington, 46.55716 N 75.926926 W, 172 m, 22 Jun 1954, E.E. Sterns, 1♀ (00113936) (CNC); 22 Jun 1954, H.J. Huckel, 1♀ (00113935) (CNC). Impreuil [coordinates unknown], 19 Jul 1902, unknown, 1♀ (00116333) (CUIC). Kazubazua, 45.956 N 76.026 W, 18 Aug 1927, G.S. Walley, 18 (00113927) (CNC). Knowlton, 45.216666 N 72.516666 W, 30 Jun 1936, G.S. Walley, 1♀ (00113933) (CNC). Lac Mondor, Ste. Flore, 46.616666 N 72.766666 W, 21 Jun 1951, E.G. Munroe, 18 (00113929) (CNC); 24 Jun 1951, E.G. Munroe, 18 (00113930) (CNC). Laniel, 47.033336 N 79.266666 W, 03 Jul 1963 – 04 Jul 1963, L.A. Kelton, 1♀ (00113934) (CNC). Magog, 45.266666 N 72.156 W, 02 Aug 1961, G. Brumpton, 1♀ (00114014) (CNC). Nominingue, 46.401176 N 75.037986 W, 259 m, 12 Jul 1922 – 22 Jul 1922, J. Ouellet, 1♀ (00291136) (TAMU); 12 Jul 1932, J. Ouellet, 1♀ (00093771) (TAMU). Quyon, 45.516666 N 76.233336 W, 22 Jul 1958, L.A. Kelton, Salix sp. (Salicaceae), 18 (00113931) (CNC). Rigaud, 45.478916 N 74.307366 W, 30 m, 26 Jul, unknown, 18 (00116328) (CUIC). Sainte-Agathe, 46.033336 N 74.283336 W, 09 Aug 1961, G. Brumpton, 18 (00113928) (CNC). Sherbrooke, 45.347336 N 71.936986 W, 306 m, 05 Jul 1916, unknown, 18 (00116327) (CUIC). Saskatchewan: Assiniboia, 49.616666 N 105.983336 W, 20 Jun 1955, A.R. Brooks, 18 (00113882) (CNC); 25 Jun 1955, A.R. Brooks, 18 (00113881) (CNC). Big River, 53.833336 N 107.033336 W, 05 Jun 1959, A. and J. Brooks, 18 (00113873) (CNC); 03 Jul 1959, A. and J. Brooks, 18 (00113874) (CNC). Broadview, 50.383336 N 102.566666 W, 10 Jul 1951, L.A. Konotopetz, 18 (00113872) (CNC). Buffalo Pound, 50.58336 N 105.38896 W, 29 Jun 1955, A.R. Brooks, 1♀ (00113885) (CNC). Christopher Lake, 53.566666 N 105.833336 W, 11 Jul 1959, A.R. and J.E. Brooks, 18 (00113883) (CNC); 15 Jul 1959, A.R. and J.E. Brooks, 18 (00113884) (CNC). Elbow, 51.116666 N 106.60006 W, 20 Jun 1960, A.R. Brooks, 18 (00113878) (CNC). Good Spirit Lake, 51.556 N 102.666666 W, 10 Jul 1954, A.R. Brooks, Wallis, 18 (00113876) (CNC). Indian Head, 50.533336 N 103.666666 W, 07 Jul 1957, A.R. Brooks, 1♀ (00113894) (CNC). Kandahar, 51.776 N 104.356 W, 29 Jul 1954, Brooks and Wallis, 18 (00113880) (CNC). McGee, 51.50006 N 108.256 W, 03 Jul 1952, L.A. Konotopetz, 1♀ (00113888) (CNC). Mossbank, 49.933336 N 105.966666 W, 23 Jun 1955, A.R. Brooks, 18 (00242557) (CAS). Saint Victor, 49.433056 N 105.866666 W, 25 Jun 1955, A.R. Brooks, 18 (00113877) (CNC). Saskatoon, 52.133336 N 106.666666 W, 26 Jun 1951, L.A. Konotopetz, 18 (00113870) (CNC); 06 Jul 1957, A.R. Brooks, 18 (00113871) (CNC). Snowden, 53.48346 N 104.683396 W, 447 m, 17 Jul 1950, L.A. Konotopetz, Solidago sp., 1♀ (00113895) (CNC). Torch River, 53.856 N 103.10006 W, 11 Jul 1950, L.A. Konotopetz, Solidago sp., 1♀ (00113886) (CNC); 20 Jul 1950, L.A. Konotopetz, Solidago sp., 1♀ (00113887) (CNC). Willows, 49.60006 N 105.856 W, 19 Jun 1955, A.R. Brooks, 18 (00113875) (CNC).  USA: Alabama: Clay Co.:  Cheaha State Park, 33.453916 N 85.826926 W, 449 m, 28 May 1984, C.B. Barr, Light Trap, 18 (00119542) (UCB). Cullman Co.: 1 mi S of Hwy 278 on I-65, 34.156246 N 86.872416 W, 230 m, 18 May 1983, D.A. Rider, 38 (00242945–00242947), 1♀ (00242948) (DAR).  Colorado: Arapahoe Co.:  Cherry Creek Reservoir, 39.61346 N 104.8226 W, 1707 m, 21 Jun 1980, D.A. Polhemus, 18 (00116888) (JTP). Englewood, 39.647786 N 104.987226 W, 11 Jun 1978, D.A. and J.T. Polhemus, 18 (00116887) (JTP). Boulder Co.: 4.5 mi N of Boulder, 40.081036 N 105.281456 W, 1676 m, 10 Jun 1961, J.R. Stainer, 18 (00113960) (CNC); 13 Jun 1961, J.R. Stainer, 18 (00113961), 1♀ (00113959) (CNC). Costilla Co.: Fort Garland, Ute Creek Ranch, 37.432746 N 105.438466 W, 11 Aug 1925, H.H. Knight, paratypes, 48 (00134183– 00134185, 00134270), 4♀ (00134186–00134189) (USNM). Custer Co.: Ophir Creek Campground, 38.10006 N 105.156 W, 29 Aug 1991, G.F. and J.F. Hevel, 18 (00134127) (USNM). Denver Co.: Denver, 39.739176 N 104.984176 W, 20 Jun 1969, J.T. Polhemus, 18 (00093784) (TAMU). Denver, 39.739176 N 104.984176 W, Jul, N. Banks, 1♀ (00108879) (AMNH). Dolores Co.: Rico, 37.692786 N 108.029726 W, Aug, E.D. Ball, 18 (00134483) (USNM). Douglas Co.: 3 mi E of Larkspur, 39.228616 N 104.830686 W, 03 Jul 1979, D.A. Polhemus, 18 (00116891) (JTP). Roxborough Park Road near Chatfield State Park, 39.473896 N 105.084726 W, 1707 m, 01 Jun 1981, J.T. Polhemus, 18 (00116886) (JTP). Waterton, 39.493616 N 105.088066 W, 15 Jun 1981, D.A. Polhemus, 18 (00116890) (JTP); 22 Jun 1981, D.A. Polhemus, 18 (00116889) (JTP). nr Waterton, Roxborough Road, 39.493616 N 105.088066 W, 1707 m, 04 Jun 1981, D.A. Polhemus, 18 (00116885) (JTP); 22 Jun 1981, D.A. Polhemus, 18 (00116884) (JTP). Jackson Co.: Muddy Pass, Routt National Forest, 40.376676 N 106.579176 W, 2652 m, 21 Aug 1968, L.A. Kelton, Artemisia sp., 1♀ (00113958) (CNC). Jefferson Co.: Indian Hills, 39.616676 N 105.236676 W, 2134 m, 15 Jul 1983, R.T. Schuh and D.A. Polhemus, 1♀ (00170126) (AMNH). Larimer Co.: Fort Collins, 40.585286 N 105.083896 W, 12 Jun, unknown, paratype, 18 (00134170) (USNM); 28 Jun, unknown, paratype, 18 (00134171) (USNM). Fort Collins, Dixon Canyon, 40.55446 N 105.1486 W, 1657 m, 19 Aug 1898, E.D. Ball, paratypes, 48 (00134174– 00134177), 5♀ (00134178–00134182) (USNM). Little Beaver [Creek], 40.623586 N 105.523886 W, 2402 m, 18 Jul 1898, unknown, 1♀ (00134296) (USNM). Montrose Co.: 27 mi E of Nucla, 38.269446 N 108.050056 W, 2804 m, 14 Aug 1987, T.J. Henry, Ligusticum porteri Coult and Rose (Apiaceae), 18 (00069991) (USNM). Routt Co.: Steamboat Springs, 40.4856 N 106.831116 W, 2134 m, 22 Aug 1968, L.A. Kelton, Solidago sp., 28 (00113949–00113950) (CNC). Steamboat Springs, 40.4856 N 106.831116 W, 2103 m, 15 Jul 1964, H.H. Knight, 18 (00093786) (TAMU), Solidago sp., 18 (00134276), 1♀ (00134277) (USNM). Unknown co.: Colo 1690, unknown, 18 (00134293) (USNM).  Connecticut: Fairfield Co.:  Danbury, 41.384236 N 73.453436 W, 134 m, 18 Jun 1909, C.W. Johnson, 18 (00170146) (AMNH). Greenwich, 41.017866 N 73.625856 W, 09 Jul 1907, Cushman, 18 (00170147) (AMNH). Ridgefield, 41.279836 N 73.495576 W, 219 m, 24 Jun 1916, I.N. Gabrielson, 18 (00134303) (USNM). Litchfield Co.: Lakeville, 41.962376 N 73.44546 W, 228 m, 22 Jun 1935, R.F. Hussey, 1♀ (00127505) (UMMC). Middlesex Co.: Killingworth, 41.358156 N 72.56376 W, 117 m, 27 Jun 1920, W.E. Britton, paratype, 1♀ (00134227) (USNM). New Haven Co.: 2 mi E of Seymour, 41.377686 N 73.058516 W, 109 m, 30 May 1965, J. and W. Ivie, 1♀ (00170182) (AMNH). Cheshire, 41.498896 N 72.901116 W, 78 m, 20 Jun 1959, J.A. Slater, 1♀ (00170160) (AMNH). Orange, 41.279756 N 73.029596 W, 55 m, 22 Jun 1920, P. Garman, paratype, 1♀ (00134235) (USNM); 30 May 1977, P.W. Kovarik, 18 (00093787) (TAMU). West Haven, 41.26366 N 72.95826 W, 15 m, 27 Jun 1905, H.L. Viereck, 1♀ (00242508) (CAS). Tolland Co.: Eagleville, 41.786676 N 72.276946 W, 22 Jun 1977, D. Leston, 18 (00170148) (AMNH). Mansfield Center, 41.765286 N 72.198616 W, 84 m, 22 Jun 1954, J.A. Slater, 1♀ (00170159) (AMNH); 21 Jun 1959, J.A. Slater, 18 (00170156) (AMNH). Storrs, 41.808336 N 72.256 W, 02 Jun 1954, J.A. Slater, 1♀ (00170161) (AMNH); 02 Jul 1954, J.A. Slater, 1♀ (00170165) (AMNH); 29 Jul 1954, J.A. Slater, 1♀ (00170166) (AMNH); 06 Jul 1964, unknown, 18 (00170153) (AMNH); 14 Jul 1977, D. Leston, 18 (00170149) (AMNH). Union, vicinity of Yale Forest, 41.992346 N 72.127176 W, 218 m, 01 Jun 1977, R. Veranes, 18 (00170150) (AMNH). Windham Co.: Bibbins Pond, 41.662596 N 72.170896 W, 90 m, 02 Jun 1974, K. Schmidt, 18 (00169992), 1♀ (00170163) (AMNH). Unknown co.: Conn, unknown, 1♀ (00070114) (USNM). District of Columbia: Brightwood, 38.956856 N 77.038896 W, 80 m, 17 May 1905, O. Heidemann, 18 (00116275), 1♀ (00116279) (CUIC); 12 Jun 1906, O. Heidemann, 18 (00242560) (CAS), 18 (00116273) (CUIC). Rock Creek, Washington, 38.908176 N 77.051056 W, 27 May 1891, unknown, 1♀ (00116277) (CUIC). Rock Creek Park, 38.6786 N 77.04686 W, 46 m, 28 May 1905, unknown, 1♀ (00069966) (USNM). Washington DC, 38.891786 N 77.008316 W, 20 May 1888, unknown, 1♀ (00116284) (CUIC); 26 Jun 1897, unknown, 1♀ (00116281) (CUIC); 22 May, unknown, 18 (00116271) (CUIC); 22 Jun, unknown, 1♀ (00116285) (CUIC), 1♀ (00069967) (USNM); 04 Jun 1902, O. Heidemann, 1♀ (00116283) (CUIC); 21 May 1903, unknown, 1♀ (00116280) (CUIC); 06 Jun 1903, O. Heidemann, 1♀ (00069968) (USNM); 15 Jun 1903, O. Heidemann, 1♀ (00242561) (CAS); 05 Jun 1904, O. Heidemann, 18 (00116269) (CUIC); 06 Aug 1904, unknown, 1♀ (00116282) (CUIC); 18 May 1905, D.H. Clemons, 18 (00069927) (USNM); 25 May 1905, D.H. Clemons, 1♀ (00069964) (USNM); 08 Jun 1905, O. Heidemann, 18 (00116270) (CUIC); 24 Jun 1926, H.H. Knight, paratype, 1♀ (00113966) (CNC).  Georgia: Baldwin Co.:  Baldwin County, no specific locality, 33.086 N 83.2326 W, 102 m, May 1979, C.L. Smith, 18 (USNM). Clarke Co.: Clarke County, no specific locality, 33.966 N 83.3786 W, 228 m, May 1979, C.L. Smith, 18 (USNM). Jackson Co.: Jackson County, no specific locality, 34.1166 N 83.5726 W, 228 m, Apr 1979, C.L. Smith, 18 (USNM). Madison Co.: Madison County, no specific locality, 34.1246 N 83.2216 W, 220 m, Jun 1979, C.L. Smith, 18 (USNM). Rabun Co.: Pine Mountain, 34.940376 N 83.18716 W, 427 m, 04 May 1957, W.R.M. Mason, 18 (00113965) (CNC). Union Co.: Union County, no specific locality, 34.8766 N 83.9586 W, 574 m, May 1979, C.L. Smith, Solidago sp., 18 (USNM).  Idaho: Clark Co.:  13 mi E of Kilgore, 44.39956 N 111.816016 W, 1949 m, 15 Jul 1956, W.F. Barr, 18 (00122391) (UID). Idaho Co.: Lowell, Clearwater National Forest, 46.02256 N 115.896 W, 30 Jul 1972, L.A. Kelton, 18 (00113955) (CNC). Latah Co.: Moscow, 46.73256 N 116.999176 W, 10 Jul 1932, T.A. Brindley, 18 (00134285) (USNM); 14 Jun 1936, T.A. Brindley, 18 (00134492) (USNM); 11 Jul 1936, T.A. Brindley, 18 (00134286) (USNM); 26 Jul 1936, L.A. Aitken, 18 (00134284) (USNM). Lemhi Co.: 9.5 mi SE of Leadore, 44.553166 N 113.316386 W, 1953 m, 13 Jul 1956, W.F. Barr, 18 (00122392) (UID). Nez Perce Co.: Lewiston, 46.416676 N 117.016676 W, 168 m, 13 Jul 1936, B.F. Coon, 1♀ (00134292) (USNM). Power Co.: American Falls, 42.786116 N 112.853616 W, 05 Aug 1972, L.A. Kelton, Artemisia sp., 18 (00113953) (CNC). Teton Co.: Pine Creek Pass, 43.571676 N 111.214446 W, 06 Aug 1972, L.A. Kelton, Artemisia sp., 1♀ (00113956) (CNC). Valley Co.: McCall, 44.911116 N 116.098616 W, 1528 m, 25 Jul 1952, S.E. Knapp, 18 (00122395) (UID).  Illinois: Clark Co.:  Dolson, Rocky Branch (Clarkeville), 39.467786 N 87.765286 W, 25 Jun 1932, Frison and Mohr, paratype, 1♀ (00243468) (INHS). Coles Co.: Charleston, 39.46846 N 88.17966 W, 205 m, 08 Jun 1931, Frison, paratype, 18 (00243494) (INHS). Cook Co.: Park Ridge, 42.013446 N 87.861846 W, 190 m, 03 Jun 1942, J.A. Slater, 18 (00170157) (AMNH). Edwards Co.: Browns, 38.38136 N 87.9876 W, 125 m, 25 Jun 1932, Ross, Dozier, and Park, paratype, 18 (00243508) (INHS). Gallatin Co.: Shawneetown, 37.71316 N 88.18676 W, 122 m, 27 May 1928, T.H. Frison, paratype, 1♀ (00243495) (INHS). Grundy Co.: Morris, 41.354856 N 88.42346 W, 149 m, 28 Jun 1968, L.A. Kelton, Solidago sp., 18 (00113941) (CNC). Hancock Co.: Hamilton, 40.396396 N 91.338896 W, 27 Jun 1968, L.A. Kelton, Ambrosia sp., 18 (00113940) (CNC). Hardin Co.: Elizabethtown, 37.445836 N 88.3056 W, 27 May 1932 – 31 May 1932, H.L. Dozier, paratype, 18 (00243467) (INHS). Iroquois Co.: Sheldon, 40.76926 N 87.563926 W, 208 m, 04 Jun 1932, Frison and Mohr, paratypes, 2♀ (00243496) (INHS). Jackson Co.: Fountain Bluff, 37.681396 N 89.507226 W, 15 May 1932, Frison, Ross, Mohr, paratype, 18 (00243497) (INHS). Giant City Park, Makanda, 37.617556 N 89.208976 W, 136 m, 21 May 1932, H.L. Dozier, paratype, 18 (00243486) (INHS). Giant City Park, Makanda, 37.61246 N 89.18926 W, 198 m, May 1932, H.L. Dozier, paratype, 1♀ (00243469) (INHS). Jo Daviess Co.: Elizabeth, 42.31786 N 90.22156 W, 244 m, 06 Jul 1917, unknown, paratype, 18 (00243466) (INHS). Galena, 42.416676 N 90.428896 W, 30 Jun 1932, Dozier and Mohr, paratype, 18 (00243498) (INHS); 01 Jun 1933, Ross and Townsend, paratype, 18 (00243459) (INHS). Johnson Co.: Cypress, 37.365056 N 89.018136 W, 119 m, 17 May 1932, H.L. Dozier, paratype, 18 (00243463) (INHS). Kankakee Co.: sand area east of Saint Anne, 41.025036 N 87.713926 W, 203 m, 04 Jun 1932, Frison and Mord, paratype, 18 (00243499) (INHS). Knox Co.: Galesburg, 40.956116 N 90.349796 W, 234 m, 07 Jul 1892, Stromberg, paratype, 18 (00243500) (INHS). Lake Co.: Antioch, 42.477226 N 88.095566 W, 05 Jul 1932 – 07 Jul 1932, T.H. Frison, paratype, 18 (00243501) (INHS); 05 Jul 1932 – 07 Jul 1932, Frison et al., paratype, 1♀ (00243574) (INHS); 10 Jul 1933, Mohr and Townsend, paratypes, 28 (00243460, 00243573) (INHS). Grays Lake, 42.349116 N 88.040216 W, 237 m, 10 Jun 1936, Ross and Burke, paratype, 18 (00243507) (INHS). Volo Bog, 42.35146 N 88.18876 W, 238 m, 15 Jul 1928, T.H. Frison, paratype, 1♀ (00243470) (INHS); 11 Jun 1936, Ross and Burks, paratype, 1♀ (00243471) (INHS). Macon Co.: Decatur, 12 Jun 1935, Frison and DeLong, 18 (00243472) (INHS). Madison Co.: Glen Carbon, 38.748336 N 89.983066 W, 15 Jun 1946, J.A. Slater, 1♀ (00170171) (AMNH). Godfrey Estes Kennels, 38.960136 N 90.189596 W, 182 m, 09 May 1976, D.C. Iftner, 18 (00243098) (UMRM). Mason Co.: Havana, 40.30006 N 90.060836 W, 31 May 1933, C.A. Mohr, paratype, 1♀ (00243502) (INHS). McHenry Co.: Algonquin, 42.16566 N 88.29426 W, 07 Jul 1909, Nason, paratype, 18 (00243462) (INHS). Mercer Co.: Keithsburg, 41.099446 N 90.94256 W, 08 Jun 1932, Ross and Mohr, 18 (00243474) (INHS). Ogle Co.: Grand Detour, Castle Rock, 41.896676 N 89.411676 W, 02 Jul 1932, Dozier and Mohr, paratype, 1♀ (00243475) (INHS). Piatt Co.: Monticello, 40.027786 N 88.573336 W, 201 m, 28 Jun 1968, L.A. Kelton, 18 (00113942) (CNC). Pope Co.: Herod, 37.580286 N 88.436116 W, 29 May 1936, Ross and Mohr, paratype, 18 (00243503) (INHS). Pulaski Co.: Pulaski, 37.2146 N 89.20586 W, 10 May 1932, H.L. Dozier, paratype, 18 (00243465) (INHS); 25 May 1932, H.L. Dozier, paratype, 18 (00243464) (INHS). Union Co.: Dongola, 37.361166 N 89.165916 W, 120 m, 12 May 1916, C.A. Hart, paratype, 18 (00243504), 1♀ (00243575) (INHS); 11 May 1917, unknown, paratype, 18 (00243461) (INHS). Unknown co.: ‘‘N. Ill.’’, unknown, 1♀ (00164926) (USNM). Vermilion Co.: Muncie, Stoney Creek, 40.115596 N 87.844756 W, 197 m, 24 May 1914, C.A. Hart, paratype, 18 (00243505) (INHS). Washington Co.: Du Bois, 38.22396 N 89.21226 W, 152 m, 22 May 1917, unknown, 18 (00243473) (INHS). Will Co.: Joliet, 41.5256 N 88.081676 W, 09 Jun 1933, Mohr and Townsend, paratype, 18 (00243506) (INHS).  Indiana: Brown Co.:  Yellowwood Forest, nr Belmont, 39.16116 N 86.34256 W, 25 May 1963, F.N. Young, 18 (00243719), 4♀ (00243714–00243717) (FSCA). Clark Co.: Clark County, 38.295676 N 85.731496 W, 136 m, 13 Jun 1932 – 14 Jun 1932, B.E. Montgomery, 1♀ (00127506) (UMMC). Henryville, Clark Co. State Forest, 38.55416 N 85.836 W, 61 m, 13 Jun 1933, B.E. Montgomery, 18 (00127496) (UMMC). Lawrence Co.: Lawrence County, no specific location, 38.87316 N 86.47216 W, 213 m, 17 Jun 1931, Montgomery, 18 (00134075) (USNM).  Iowa: Boone Co.:  Ledges State Park, 41.983336 N 93.886676 W, 09 Jul 1956, J.C. Schaffner, 1♀ (00291099) (TAMU). Pilot Mound, 42.164446 N 94.016396 W, 01 Jun 1957, A.H. Barnum, 1♀ (00291100) (TAMU). Cerro Gordo Co.: Clear Lake, 43.15576 N 93.37966 W, 366 m, 30 Jun 1927, Harris and Johnston, 18 (00113963) (CNC). Clayton Co.: McGregor, 43.039116 N 91.120426 W, 228 m, 01 Jul 1950, Hicks and Slater, 2♀ (00170170, 00170173) (AMNH). Dickinson Co.: Cayler Prairie, 43.396396 N 95.244446 W, 09 Jun 1963, J.C. Schaffner, 18 (00093785) (TAMU). Iowa Lakeside Laboratory, 43.379176 N 95.186116 W, 11 Jun 1963, J.C. Schaffner, 18 (00093783) (TAMU). Henry Co.: 5 mi S of Mount Pleasant, 40.894286 N 91.546956 W, 186 m, 20 Jun 1966, J.C. Schaffner, 18 (00093774) (TAMU). Oakland Mills State Park, 40.936116 N 91.616396 W, 31 May 1963, J.C. Schaffner, 1♀ (00291098) (TAMU). Iowa Co.: Amana, 41.803266 N 91.875386 W, 251 m, 07 Jul 1927, Harris and Johnston, 1♀ (00134350) (USNM). Johnson Co.: Iowa City, 41.661116 N 91.536 W, 203 m, 17 Jun 1915, L. Stoner, 1♀ (00242507) (CAS). Lake MacBride State Park, 41.800736 N 91.571516 W, 244 m, 08 Jul 1950, Laffoon, Slater, Hicks, 18 (00170155) (AMNH). Monona Co.: Lewis and Clark State Park, 42.033336 N 96.166666 W, 319 m, 06 Jun 1960, N.R. Austin, 1♀ (00133820) (USNM). Turin, nr loess bluff, 42.033336 N 95.956 W, 320 m, 06 Jun 1960, J.C. Schaffner, 1♀ (00093796) (TAMU); 06 Jun 1960, J.L. Laffoon, 1♀ (00133815) (USNM). Story Co.: Ames, 42.034726 N 93.619726 W, E.D.B., 18 (00116330) (CUIC); 03 Jun 1886, Osborn, 1♀ (00069963) (USNM); 18 Jun 1925, H.H. Knight, paratype, 18 (00093779), 1♀ (00093781) (TAMU); 04 Jun 1926, H.H. Knight, paratype, 18 (00113962) (CNC), paratype, 18 (00093777), 1♀ (00093780) (TAMU); 01 Jun 1927, H.G. Johnston, 1♀ (00093795) (TAMU); 31 May 1932, Floyd Andre, 18 (00134491) (USNM). Doolittle Prairie [1 mi N of 150th on 560th Ave], 42.151526 N 93.580276 W, 295 m, 01 Jun 1989, M.E. Rice, 1♀ (00093794) (TAMU), 18 (00122393), 1♀ (00122394) (UID). Warren Co.: 3 mi NE of Hartford, 41.49016 N 93.363776 W, 03 Jun 1994, J.C. Schaffner, 18 (00093775) (TAMU). Woodbury Co.: Sioux City, 42.50036 N 96.39396 W, 10 Jun 1949, Slater and Laffoon, 28 (00002609–00002610), 1♀ (00002608) (AMNH). Sioux City, 42.500286 N 96.393896 W, 10 Jun 1949, Slater and Laffoon, 18 (00170151) (AMNH), 18 (00093782) (TAMU); 12 Jun 1949, Slater and Laffoon, 1♀ (00170169) (AMNH).  Kansas: Anderson Co.:  Garnett, 38.280566 N 95.241676 W, 16 Jun 1931, P.W. Oman, 18 (00134306) (USNM); 20 Jun 1931, P.W. Oman, 1♀ (00134318) (USNM). Douglas Co.: 5 mi NE of Lawrence, 39.01146 N 95.184656 W, 254 m, 18 Jun 1971, V.P. Gapud, 1♀ (00117933) (KU). 6 mi NE of Lawrence, Rockefeller Tract, 39.0336 N 95.156046 W, 05 Jun 1976, T.W. Oldham, 18 (00093792) (TAMU). Douglas County, no specific locality, 38.866676 N 95.233336 W, 274 m, 26 Jun 1975, T.W. Oldham, 1♀ (00291137) (TAMU). Neosho Co.: Chanute, 37.684046 N 95.472226 W, 295 m, 29 Aug 1939, J.D. Beamer, 1♀ (00117931) (KU).  Kentucky: Anderson Co.:  1800 ft WNW of Tyrone, 38.04636 N 84.860516 W, 241 m, 24 May 2003, C. Wright, 18 (DBUL). Bullitt Co.: Bernheim Arboretum and Research Forest, site 11A, 37.912586 N 85.662816 W, 164 m, 03 Jun 1999, C.V. Covell and team, 38 (DBUL); 02 Jun 2000, S. Berla, J. Lewis, J. Bennett, 38 (DBUL); 09 Jun 2000, S. Berla and C.V. Covell, 18 (DBUL). Bernheim Arboretum and Research Forest, site 11B, 37.911796 N 85.662086 W, 170 m, 03 Jun 1999, C.V. Covell and team, 18 (DBUL). Christian Co.: PSF 370553 874122, 36.872276 N 87.476756 W, 160 m, 30 May 2003, C. Wright, 28 (DBUL). Fayette Co.: Lexington, 37.988696 N 84.477726 W, 294 m, 12 May 1921, K.S.B., paratype, 1♀ (00134252) (USNM); 13 Jun 1925, T.H. Hubbell, 18 (00127488) (UMMC); 07 Jun 1937, L.H. Townsend, 38 (UK); 01 Jun 1951, H.H.W., Trifolium pratense L. (Fabaceae), 18 (UK). Harlan Co.: Black Mountain, 36.848516 N 83.14636 W, 442 m, 06 Jun 1951, B.L. Monroe, 18 (DBUL). Jefferson Co.: Louisville, 38.215376 N 85.763456 W, 139 m, 10 May 1942, T. Spilman, 18 (DBUL); 23 Apr 1948, R. Hoskinson, 18 (DBUL); 02 Jun 1948, N.L. Beckman, 18 (DBUL); 07 May 1954, L.J. Cooper, 18 (DBUL); 12 Jun 1954, P.J. Christian, 18 (DBUL). Martin Co.: 2.6 mi SW of Davella, 37.776246 N 82.607546 W, 251 m, 02 Jun 2003, C. Wright, 18 (DBUL). Owen Co.: Kleber Wildlife Management Area, 38.357856 N 84.783956 W, 208 m, 26 May 2003, C. Wright, 58 (UK).  Maine: Androscoggin Co.:  Greene, Rt 202, 44.191236 N 70.143876 W, 98 m, 01 Jul 1989, T.J. Henry and A.G. Wheeler, Jr, Solidago sp., 1♀ (00069982) (USNM). Lewiston, 44.107166 N 70.208326 W, 93 m, 01 Jul 1989, A.G. Wheeler, Jr., Solidago sp., 1♀ (00069981) (USNM). Cumberland Co.: Pownal Center, 43.892486 N 70.187046 W, 41 m, 18 Jun 1986, Larochelle and Lariviere, 18 (00113937) (CNC). Waldo Co.: Rt 139, 1.5 mi W of Unity, 44.611326 N 69.336856 W, 62 m, 03 Jul 1989, T.J. Henry and A.G. Wheeler, Jr, Solidago sp., 18 (00069971) (USNM).  Maryland: Allegany Co.:  Green Ridge State Forest, 39.59886 N 78.49986 W, 415 m, 22 Jun 1985, T.J. Henry and A.G. Wheeler, Jr, Solidago sp., 18 (00069992) (USNM). Little Orleans Campground, 39.631816 N 78.392276 W, 177 m, 20 May 1995, L.L. Deitz, 18 (00242830) (NCSU). Anne Arundel Co.: South River, 4 mi S of Annapolis, 38.903896 N 76.56066 W, 35 m, 10 Jun 1934, H.G. Barber, 1♀ (00164924) (USNM). Montgomery Co.: Glen Echo, 38.967486 N 77.141726 W, 32 m, 24 Aug, unknown, 18 (00116274) (CUIC). Silver Spring, 38.990566 N 77.026396 W, 02 Jun 1967, P.W. Oman, 18 (00118158 [cat UCRC ENT 00035344]), 2♀ (00118143 [cat UCRC ENT 00035344], 00118144 [cat UCRC ENT 00035344]) (ORSU). nr Plummers Island, 38.976 N 77.1786 W, 06 Jun 1914, R.C. Shannon, 18 (00134488) (USNM). Prince George’s Co.: Beltsville, 39.034726 N 76.907786 W, 20 May 1941, unknown, 1♀ (00133814) (USNM). Beltsville, BARC [Beltsville Agricultural Research Center], 39.02476 N 76.92656 W, 47 m, 22 May 1982, T.J. Henry and A.G. Wheeler, Jr, 1♀ (00133740) (USNM). Bladensburg, 38.939266 N 76.934166 W, 06 Jul 1906, unknown, 1♀ (00116278) (CUIC).  Massachusetts: Middlesex Co.:  Arlington, 42.415376 N 71.156446 W, 20 m, 26 Jun 1897, C. Bullard., 18 (00170131) (AMNH). Framingham, 42.279176 N 71.416676 W, 01 Jun 1929, unknown, 18 (00244237) (SDNH). Holliston, 42.20006 N 71.4256 W, 09 Jun, N. Banks, 18 (00170141) (AMNH); 20 Jun, N. Banks, 1♀ (00170167) (AMNH); 22 Jun, N. Banks, 18 (00170143) (AMNH); 03 Jul, N. Banks, 1♀ (00170168) (AMNH); 04 Jul, N. Banks, 18 (00170140) (AMNH); 13 Jul, N. Banks, 18 (00170142) (AMNH). Natick, 42.297156 N 71.341996 W, 67 m, 09 Jun 1929, unknown, 18 (00244235), 1♀ (00244236) (SDNH). Sherborn, 42.238886 N 71.369726 W, 61 m, 07 Jul 1928, C.A. Frost, paratype, 1♀ (00134228) (USNM).  Michigan: Arenac Co.:  Arenac County, no specific locality, 44.005836 N 83.857646 W, 181 m, 26 Jun 1940, R.R. Dreisbach, 18 (00127484) (UMMC). Bay Co.: Bay County, no specific locality, 43.59796 N 83.89166 W, 178 m, 13 Jul 1940, R.R. Dreisbach, 1♀ (00133819) (USNM). Benzie Co.: Benzie County, no specific locality, 44.718336 N 86.116486 W, 184 m, 04 Jul 1957, R. and K. Dreisbach, 1♀ (00069986) (USNM). Berrien Co.: E.K. Warren Preserve, Warren Woods, 41.835996 N 86.636056 W, 195 m, 20 Jun 1919, R.F. Hussey, 1♀ (00127508) (UMMC). Calhoun Co.: Wise Lake, 42.234336 N 84.989156 W, 282 m, 06 Jun 1959, G.C. Eickwort, 18 (00134314) (USNM). Charlevoix Co.: Boyne City, 45.224976 N 85.036996 W, 177 m, 08 Jul 1923, T.H. Hubbell, 1♀ (00127511) (UMMC). Cheboygan Co.: Cheboygan County, no specific locality, 45.646946 N 84.474446 W, 18 Jul 1950, George Garoian, 1♀ (00170174) (AMNH). Green Timbers State Park, 45.199776 N 84.517296 W, 288 m, 12 Jun 1988, M.L. Niebylski, 18 (00242950) (DAR). Lancaster Lake, 45.61976 N 84.71126 W, 219 m, 03 Aug 1965, W.R. Anderson, 1♀ (00170179) (AMNH). Clare Co.: Clare County, no specific locality, 43.819446 N 84.768616 W, 01 Jul 1945, R.R. Dreisbach, 18 (00127490) (UMMC). Gladwin Co.: Gladwin County, no specific locality, 43.980836 N 84.486396 W, 14 Jun 1958, R. and K. Dreisbach, 1♀ (00069987) (USNM). Grand Traverse Co.: Grand Traverse County, no specific locality, 44.683346 N 85.466736 W, 285 m, 09 Jul 1960, R. and K. Dreisbach, 18 (00069979) (USNM). Traverse City, 44.746246 N 85.550776 W, 180 m, 14 Jun 1969, L. and C.W. O’Brien, 1♀ (00119541) (UCB). Huron Co.: Pigeon, 43.832216 N 83.270996 W, 189 m, 01 Jul 1922, R.F. Hussey, 18 (00127486) (UMMC); 03 Jul 1922, R.F. Hussey, 18 (00127494) (UMMC). Sand Point, 43.915136 N 83.404266 W, 176 m, 26 Jun 1922, R.F. Hussey, 1♀ (00127510) (UMMC). Ionia Co.: Ionia County, no specific locality, 42.987256 N 85.071126 W, 218 m, 04 Jul 1951, R.R. Dreisbach, 1♀ (00127509) (UMMC). Isabella Co.: Isabella County, no specific locality, 43.643536 N 84.995116 W, 274 m, 05 Jun 1945, R.R. Dreisbach, 18 (00127487) (UMMC); 07 Jun 1958, R. and K. Dreisbach, 18 (00069973) (USNM). Lake Co.: Lake County, no specific locality, 43.890866 N 85.846776 W, 250 m, 12 Jul 1947, R.R. Dreisbach, 1♀ (00134313) (USNM). Lenawee Co.: Lenawee County, no specific locality, 41.877746 N 84.052066 W, 231 m, 08 Jun 1952, R.R. Dreisbach, 1♀ (00127507) (UMMC). Livingston Co.: E.S. George Reserve, 42.460286 N 83.994376 W, 269 m, 29 Jun 1947, K. Bohnsack, 18 (00127501) (UMMC); 30 Jun 1957, T.E. Moore, 18 (00127504) (UMMC); 15 Jun 1961, unknown, 18 (00127500) (UMMC); 19 Jun 1961, unknown, 18 (00127502) (UMMC); 05 Jul 1961, unknown, 18 (00127503) (UMMC); 28 May 1964, F.C. Evans, 18 (00127492) (UMMC). Manistee Co.: 8 mi N of Manistee, 44.345736 N 86.267376 W, 197 m, 15 Jun 1969, L. and C.W. O’Brien, 18 (00119540) (UCB). Manistee County, no specific locality, 44.244476 N 86.324256 W, 202 m, 22 Jun 1957, R. and K. Dreisbach, 18 (00069974) (USNM); 05 Jul 1957, R. and K. Dreisbach, 18 (00069975) (USNM). Mecosta Co.: 2 mi W of Remus [off of] —20, 43.596986 N 85.185186 W, 309 m, 11 Jul 1953, R.F. Hussey, 1♀ (00127512) (UMMC). Mecosta County, no specific locality, 43.615366 N 85.237216 W, 291 m, 17 Jul 1950, R.R. Dreisbach, 18 (00127495) (UMMC); 15 Jun 1957, R. and K. Dreisbach, 1♀ (00069983) (USNM). Menominee Co.: Banat, Upper Michigan, 45.51886 N 87.69636 W, 244 m, 07 Jul 1972, W.F. Chamberlain, 1♀ (00245359) (TAMU). Midland Co.: Midland County, no specific locality, 43.615566 N 84.247226 W, 09 Jun 1939, R.R. Dreisbach, 1♀ (00133818) (USNM); 28 Jun 1958, R.R. Dreisbach, 1♀ (00069984) (USNM); 12 Jul 1960 – 16 Jul 1960, R.R. Dreisbach, 18 (00069976) (USNM). Missaukee Co.: Missaukee County, no specific locality, 44.335296 N 85.215056 W, 381 m, 11 Jul 1959, R. and K. Dreisbach, 18 (00069977) (USNM). T23N R6W Sec36, 44.34486 N 84.98576 W, 348 m, 20 Jun 1957, T.H. Hubbell, 18 (00127499) (UMMC). Muskegon Co.: Muskegon County, no specific locality, 43.295586 N 86.144136 W, 199 m, 30 Jun 1951, R.R. Dreisbach, 18 (00127493) (UMMC). Osceola Co.: Osceola County, no specific locality, 43.983356 N 85.333386 W, 401 m, 29 Jun 1957, R. and K. Dreisbach, 18 (00069978) (USNM). Roscommon Co.: Roscommon County, no specific locality, 44.333356 N 84.600026 W, 353 m, 05 Jun 1955, R.R. Dreisbach, 18 (00069980) (USNM). Tuscola Co.: Tuscola County, no specific locality, 43.326416 N 83.657186 W, 192 m, 22 Jun 1952, R.R. Dreisbach, 18 (00127489) (UMMC). Washtenaw Co.: Steere’s Swamp, 42.389986 N 84.153846 W, 280 m, 28 Jun 1920, R.F. Hussey, 1♀ (00243727) (FSCA). Washtenaw County, no specific locality, 42.250596 N 83.849946 W, 286 m, 28 Jun 1920, R F. Hussey, 1♀ (00127513) (UMMC). Wayne Co.: Detroit, 42.340686 N 82.96696 W, 175 m, 26 May 1892, unknown, 1♀ (00116334) (CUIC). Wexford Co.: Wexford County, no specific locality, 44.333346 N 85.566736 W, 426 m, 30 Jun 1957, R. and K. Dreisbach, 18 (00069972) (USNM).  Minnesota: Beltrami Co.:  2 mi N of Hines, Rt 71, 47.683336 N 94.633336 W, 430 m, 23 Jul 1995, T.J. Henry and A.G. Wheeler, Jr, Solidago sp., 18 (00133737) (USNM). Crow Wing Co.: 5 mi SE of Pequit Lakes, 46.625686 N 94.268416 W, 391 m, 06 Jul 1957, J.L. Laffoon, 18 (00133744) (USNM). Hennepin Co.: Hennepin County, no specific locality, 45.016676 N 93.456 W, 28 Jun 1921, A.T. Hertig, paratype, 18 (00134254), 1♀ (00134262) (USNM). Minneapolis, 44.986 N 93.263616 W, 18 Jun 1922, H.H. Knight, paratype, 18 (00134255) (USNM). Lac qui Parle Co.: Madison, 45.009726 N 96.195566 W, 25 Jun 1921, H.H. Knight, paratype, 18 (00134256) (USNM). Morrison Co.: Morrison County, no specific locality, 45.976356 N 94.36256 W, 339 m, 10 Jul 1920, A.A. Nichol, 18 (00134261), 1♀ (00134264) (USNM). Ramsey Co.: Mid Hills Golf Club, 44.992196 N 93.195226 W, 299 m, 25 Jun 1922, Arthur Hartig, 18 (00134260) (USNM). Saint Anthony Park, 44.978056 N 93.190836 W, 02 Jul 1923, H.H. Knight, paratype, 1♀ (00134257) (USNM). Redwood Co.: Redwood Falls, 44.533336 N 95.116666 W, 313 m, 19 Jun 1995, T.J. Henry and A.G. Wheeler, Jr, 18 (00134067) (USNM). Renville Co.: Olivia, 44.776396 N 94.989446 W, 28 Jun 1921, H.H. Knight, paratype, 18 (00134229), 1♀ (00134236) (USNM). Rice Co.: Faribault, 44.2956 N 93.268616 W, 12 Jun 1922, H.H. Knight, paratype, 18 (00134258) (USNM). Hunt Lake, 9 mi W of Faribault, 44.332466 N 93.452656 W, 332 m, 18 Jun 1995, T.J. Henry and A.G. Wheeler, Jr, Solidago sp., 18 (00069989) (USNM). Roseau Co.: Roseau, 48.846096 N 95.762776 W, 320 m, 27 Jun 1920, J.P. Jensen, 1♀ (00134263) (USNM). St. Louis Co.: 6 mi NNW of Cook on Rt 53, 47.900126 N 92.784536 W, 411 m, 26 Jun 1995, T.J. Henry and A.G. Wheeler, Jr, Eurybia macrophylla, 18 (00069988) (USNM). Steele Co.: Owatonna, 44.09276 N 93.21936 W, 351 m, 21 Jun 1921, A.A. Nichol, paratype, 1♀ (00134226) (USNM); 24 Jun 1921, A.A. Nichol, paratype, 18 (00134259) (USNM).  Missouri: Adair Co.:  Sublette, 40.302036 N 92.573076 W, 293 m, 22 Jun 1940, E. Herbold, 1♀ (00134328) (USNM). Bollinger Co.: Grassy, 37.264646 N 90.124586 W, 216 m, 30 May 1943, E.H. Froeschner, 18 (00134320) (USNM). Boone Co.: 1.3 mi N of Ashland Wildlife Area, 38.780586 N 92.196836 W, 247 m, 21 May 1981, R.L. Blinn, 18 (00243090) (UMRM). 5.5 mi N of Columbia, Wheaton’s Farm, 39.031316 N 92.322366 W, 212 m, 27 May 1981, R.L. Blinn, 18 (00243106) (UMRM). Ashland Wildlife Area, 38.76286 N 92.20046 W, 244 m, 12 Jun 1968, unknown, 1♀ (00243116) (UMRM). Boone County, no specific locality, 39.03556 N 92.33356 W, 195 m, 08 Jun 1988, B.D. Robbins, 1♀ (00127579) (UMRM). Columbia, 38.951676 N 92.333896 W, 02 Jun 1941, R.C. Froeschner, 1♀ (00134327) (USNM); 03 Jun 1972, S.O. Swadener, 18 (00243562) (INHS); 07 May 1981, R.L. Blinn, 18 (00243105) (UMRM); 19 May 1981, R.L. Blinn, Solidago sp., 18 (00243088) (UMRM); 05 Jun 1984, R.L. Blinn, 18 (00243094) (UMRM). Columbia, University of Missouri Campus, 38.942746 N 92.334076 W, 225 m, 12 Jun 1993, Usmani, 1♀ (00243109) (UMRM). Entomology Farm, Columbia, 38.895046 N 92.211456 W, 266 m, 16 May 1968, unknown, 18 (00243093) (UMRM). Mount Hope Baptist Church, 39.022046 N 92.231226 W, 256 m, 10 May 1974, J.W. Smith, 18 (00243092) (UMRM). Rudolph Bennet Wildlife Area, 38.927566 N 92.309816 W, 215 m, 03 Jun 1974, Smith and Thewke, 18 (00243101) (UMRM). Sturgeon, 39.230366 N 92.274626 W, 246 m, 01 Jun 1973, S.E. Thewke, Asclepias sp. (Asclepiadaceae), 18 (00243091) (UMRM). Three Creeks State Forest, 38.83886N 92.30746 W, 213 m, 14 May 1986, R.L. Blinn, 18 (00243102) (UMRM). Butler Co.: 5 mi S of Qulin, 36.594136 N 90.251086 W, 94 m, 06 Jun 1973, Enns, Swadener, Thewke, 1♀ (00243113) (UMRM). Callaway Co.: Little Dixie Lake, 38.908096 N 92.126236 W, 254 m, 20 May 1975, E.G. Riley, 18 (00243100) (UMRM). Reform, 38.758066 N 91.792666 W, 251 m, 02 Jun 1975, D.D. Kopp, 18 (00243099) (UMRM). Tucker Prairie [Natural Area], 38.948266 N 91.984116 W, 264 m, 10 May 1968, unknown, 1♀ (00243120) (UMRM); 14 May 1968, unknown, 18 (00243082) (UMRM); 17 May 1968, unknown, 1♀ (00243121) (UMRM); 01 Jun 1968, unknown, 18 (00243084) (UMRM); 04 Jun 1968, unknown, 18 (00243083) (UMRM); 06 Jun 1968, unknown, 1♀ (00243119) (UMRM); 11 Jun 1968, unknown, 18 (00243086) (UMRM); 03 Jun 1969, unknown, 18 (00243085) (UMRM); 10 Jun 1969, unknown, 18 (00243087) (UMRM); 06 May 1980, E.G. Riley, 1♀ (00243112) (UMRM); 21 May 1986, R.L. Blinn, 1♀ (00243108) (UMRM). Cooper Co.: Bunceton, 38.795426 N 92.802186 W, 233 m, 18 May 1967, W.S. Craig, 18 (00243107) (UMRM). Crawford Co.: Steelville, 37.96816 N 91.354876 W, 13 May 1939, R.C. Froeschner, 18 (00134321) (USNM). Harrison Co.: New Hampton, 40.262516 N 94.197646 W, 279 m, 13 May 1968, R.E. Munson and L.R. Hanning, (Poaceae), 1♀ (00243115) (UMRM). Hickory Co.: Hermitage, 37.933716 N 93.310756 W, 227 m, 04 Jun 1968, R.E. Munson and L.R. Hanning, (Fabaceae), 1♀ (00243114) (UMRM). Jackson Co.: Atherton, 39.186116 N 94.305286 W, 223 m, 31 May 1915, unknown, paratype, 18 (00242640) (CAS); 13 Jun 1915, C.F. Adams, 18 (00242504) (CAS); 27 Jun 1915, C.F. Adams, 18 (00242505) (CAS). Kansas City, 39.099726 N 94.578336 W, 25 May 1975, H. Greenbaum, 18 (00093790) (TAMU). Jasper Co.: Carthage, 37.157116 N 94.320446 W, 317 m, 18 May 1941, R.C. Froeschner, 18 (00134319) (USNM). Macon Co.: La Plata, 40.0236 N 92.49956 W, 274 m, 10 Jun 1968, R.E. Munson, (Fabaceae), 18 (00243104) (UMRM). Monroe Co.: Mark Twain State Park, 39.48786 N 91.79136 W, 198 m, 14 Jun 1984, R.L. Blinn, 18 (00243089) (UMRM). Randolph Co.: Moberly, 39.413626 N 92.414156 W, 259 m, 16 May 1972, T.J. Riley, 1♀ (00243117) (UMRM). Shelby Co.: 0.7 mi W of Lakenan, 39.675386 N 91.962876 W, 227 m, 22 May 1975, S.O. Swadener, 1♀ (00243563) (INHS). St. Louis Co.: Glencoe, 38.543436 N 90.626186 W, 123 m, 16 May 1937, R.C. Froeschner, 18 (00134322) (USNM); 23 May 1937, R.C. Froeschner, 1♀ (00134323) (USNM); 18 May 1947, R.C. Froeschner, 18 (00170154) (AMNH). Overland, 38.693886 N 90.371936 W, 183 m, 14 May 1938, B.H. Piaeul, 1♀ (00134305) (USNM). Ranken, 38.534876 N 90.511096 W, 128 m, 23 May 1954, E.P. Meiners, 1♀ (00243111) (UMRM). Saint Louis, 38.627226 N 90.197786 W, 19 Jun 1937, R.C. Froeschner, 1♀ (00134324) (USNM); 27 May 1939, R.C. Froeschner, 1♀ (00134326) (USNM); 25 Jun 1939, R.C. Froeschner, 1♀ (00134325) (USNM). Vernon Co.: 4 mi E of Milo, 37.756116 N 94.232686 W, 14 Jun 1966, J.C. Schaffner, 1♀ (00093797) (TAMU).  Montana: Fergus Co.:  6 mi S of Grass Range, 46.939366 N 108.803896 W, 06 Jul 1972, G.C. Gaumer, 18 (00118219), 18 (00093773), 1♀ (00291135) (TAMU). Gallatin Co.: Montana [Range] Experimental Station, Bozeman, 45.67976 N 111.03786 W, 1467 m, 30 Jun 1922, unknown, 1♀ (00134173) (USNM). Moose Flat Campground, 26 mi S of Bozeman Hot Springs on Rt 191, 45.355276 N 111.170556 W, 1737 m, 10 Aug 1986, Schuh, Schwartz, and Stonedahl, 18 (00170120) (AMNH). Mineral Co.: 18 mi S of Dry Creek, 47.051656 N 114.759886 W, 914 m, 20 Jun 1919, M.A. Hanna, 18 (00164888) (USNM). Park Co.: Park County, 45.483336 N 110.566676 W, 1829 m, 14 Aug 1926, A.A. Nichol, 18 (00134283) (USNM). Phillips Co.: 1 mi W of Landusky, 47.895176 N 108.638586 W, 1277 m, 06 Jul 1972, G.C. Gaumer, 18 (00093772) (TAMU). Ravalli Co.: Hamilton, 46.247226 N 114.153896 W, 03 Jul 1928, unknown, 18 (00134294) (USNM).  Nebraska: Cherry Co.:  Valentine, 42.872786 N 100.550566 W, 10 Jun 1950, Hicks, Slater, and Laffoon, 1♀ (00170180) (AMNH). Cuming Co.: West Point, 41.8456 N 96.731666 W, 397 m, Jun 1884, unknown, 1♀ (00164889) (USNM).  New Hampshire: Coos Co.:  Mount Washington, 44.292326 N 71.280826 W, 1282 m, 06 Jul 1914, C.A. Frost, 1♀ (00242497) (CAS).  New Jersey: Bergen Co.:  Campgaw, 41.059456 N 74.198666 W, 177 m, 02 Jun 1925, F.M. Schott, paratype, 18 (00134219) (USNM). Mercer Co.: Hopewell, Stony Brook-Millstone Watershed, Adventure Trail, 40.359046 N 74.774896 W, 55 m, 28 Jun 2001, J. Lento, 1♀ (00329799) (DEBU). Morris Co.: Madison, 40.759726 N 74.41756 W, 16 Jun 1898, H.G. Barber, 1♀ (00134490) (USNM). Passaic Co.: Paterson, 40.916676 N 74.172226 W, 31 May, unknown, 1♀ (00170176) (AMNH). Ramapo, 41.042296 N 74.249216 W, 187 m, 03 Jun 1905, unknown, 1♀ (00164925) (USNM); 05 Jun 1905, unknown, 18 (00133810) (USNM). Sussex Co.: Duttonville, 41.351486 N 74.6866 W, 148 m, 08 Jun 1910, unknown, 1♀ (00170175) (AMNH).  New Mexico: Otero Co.:  6 mi NE of Cloudcroft, 32.991546 N 105.672136 W, 2468 m, 04 Jun 1980, D.W. Webb, 38 (00243561, 00243569–00243570), 1♀ (00243571) (INHS). San Miguel Co.: Panchuela Canyon [Campground], Cowles, 35.845916 N 105.653926 W, 2743 m, 01 Jul 1935, H. Ruckes, 1♀ (00134237) (USNM).  New York: Cattaraugus Co.:  Allegany State Park, 07 Jul 1936, unknown, 1♀ (00134484) (USNM); 21 Jul 1936, unknown, 1♀ (00134486) (USNM). Four Mile, 42.020246 N 78.464586 W, 713 m, 04 Jul 1916, H.H. Knight, 1♀ (00134301) (USNM). Rock City, 42.014446 N 78.474726 W, 04 Jul 1915, H.H. Knight, 18 (00134300) (USNM); 05 Jul 1915, H.H. Knight, 1♀ (00134316) (USNM). Erie Co.: Buffalo, 42.886396 N 78.878616 W, Jun 1886, E.P. Van Duzee, 18 (00170129) (AMNH), 18 (00116331) (CUIC); 05 Jun 1891, E.P. Van Duzee, 48 (00242562–00242565) (CAS). Hamburg, 42.715836 N 78.829726 W, 26 Jun 1898, E.P. Van Duzee, 1♀ (00242571) (CAS); 06 Jun 1909, E.P. Van Duzee, 38 (00242566–00242567, 00242569), 2♀ (00242568, 00242570) (CAS). Essex Co.: Lake Placid, 44.279446 N 73.980286 W, 610 m, 04 Jul 1991, M.D. Schwartz, Solidago sp., 1♀ (00111413) (CNC). Underwood, 44.087856 N 73.676276 W, 336 m, 26 Jun 1936, H. Dietrich, 1♀ (00116337) (CUIC). Wilmington, 44.388386 N 73.815426 W, 94 m, 30 Jun 1922, J.M. Aldrich, 1♀ (00134489) (USNM). Genesee Co.: Batavia, 42.998066 N 78.187786 W, 271 m, 12 Jul 1914, H.H. Knight, paratype, 18 (00134239) (USNM); 14 Jun 1915, H.H. Knight, paratype, 1♀ (00134216) (USNM); 25 Jun 1915, H.H. Knight, 18 (00242506) (CAS). Greene Co.: Freehold, 42.35816 N 74.0526 W, 131 m, 09 May 1976, S.E. Thewke, 18 (00243103) (UMRM). Livingston Co.: Portage, 42.576676 N 78.046676 W, 27 Jun 1915, H.H. Knight, paratype, 18 (00134204) (USNM); 27 Jul 1915, H.H. Knight, 28 (00134205–00134206), 1♀ (00134207) (USNM). Monroe Co.: Charlotte, 43.255066 N 77.616956 W, 82 m, 20 Jun 1907, J.L. Zabriskie, 18 (00170127) (AMNH). Honeoye Falls, 42.952226 N 77.590566 W, 14 Jun 1915, M.D. Leonard, 1♀ (00116335) (CUIC). Rochester Junction, 42.996346 N 77.56386 W, 205 m, 27 Jun 1914, M.D. Leonard, 18 (00116319) (CUIC); 06 Jul 1914, M.D. Leonard, 18 (00116320) (CUIC); 07 Jul 1914, M.D. Leonard, 1♀ (00116332) (CUIC); 25 Jul 1914, M.D. Leonard, 18 (00116318) (CUIC). Richmond Co.: Egbertville, Staten Island, 40.583096 N 74.131116 W, 35 m, 07 Jun 1970, V. Sapienza, 1♀ (00170164) (AMNH). Schuyler Co.: Hector Land Use Area, nr Reynoldsville, 42.50216 N 76.74166 W, 395 m, 30 Jun 1982, G. and K. Eickwort, 18 (00116321) (CUIC). Suffolk Co.: Flanders, 40.903436 N 72.617596 W, 2 m, 11 Sep 1946, Roy Latham, 18 (00134485) (USNM). Napeague, Long Island, 41.009556 N 72.068136 W, 2 m, 07 Sep 1946, R. Latham, 18 (00116323) (CUIC). Northwest, 41.01016 N 72.24566 W, 3 m, 12 Jun 1951, Roy Latham, 18 (00134307) (USNM). Riverhead, 40.917046 N 72.662046 W, 3 m, 09 Jun 1946, R. Latham, 1♀ (00116336) (CUIC). Sag Harbor, 41.000636 N 72.285756 W, 2 m, 04 Apr 1946, Roy Latham, 18 (00134311) (USNM). Tompkins Co.: Ithaca, 42.440566 N 76.496946 W, 24 May 1977, F.J. Messina, 18 (00116325) (CUIC); 23 Jun 1977, F.J. Messina, 18 (00116324) (CUIC); 07 Jul 1977, F.J. Messina, 18 (00116326) (CUIC). McLean Bogs Res. [Preserve], 42.541856 N 76.298946 W, 335 m, 03 Jul 1920, H.H. Knight, paratype, 18 (00134238) (USNM); 21 Jul 1923, J.L. Buys, paratype, 1♀ (00134349) (USNM). Tompkins Co. Field 2, 42.359736 N 76.411946 W, 328 m, 18 Jun 1975, unknown, Leucanthemum vulgare, 18 (00116322) (CUIC). Westchester Co.: Armonk, 41.126486 N 73.714026 W, 118 m, 14 Jun 1924, unknown, 1♀ (00134224) (USNM). Armonk, Calder Ecology Study Center, 41.126396 N 73.714446 W, 11 Jun 1979 – 12 Jun 1979, K. Schmidt, 18 (00170138) (AMNH). White Plains, 41.033896 N 73.763336 W, 14 Jun 1919, J.R. de la Torre Bueno, paratype, 1♀ (00134215) (USNM); 07 Jun 1924, J.R. de la Torre-Bueno, 18 (00134214) (USNM). Wyoming Co.: Wyoming, 42.827366 N 78.089926 W, 308 m, 25 Jun 1916, H.H. Knight, 18 (00134299) (USNM).  North Carolina: Avery Co.:  Banner Elk, 36.157336 N 81.872466 W, 1135 m, 16 Jun 1964, J.F. Cornell, 18 (00242825) (NCSU); 21 May 1995, C.R. Bartlett, 18 (00242829) (NCSU). Buncombe Co.: Swannanoa, 35.597786 N 82.40006 W, 914 m, 20 Jun 1916, R.W. Leiby, paratype, 1♀ (00134230) (USNM). Chatham Co.: Hwy 64 at Haw River, 35.731276 N 79.108176 W, 101 m, 07 May 1993, R.L. Blinn, 1♀ (00242839) (NCSU). Columbus Co.: Hwy 701 at SR 1545, 34.438756 N 78.730636 W, 20 m, 04 May 2005, P. Alvarez, 18 (00242833) (NCSU). Harnett Co.: Hwy 401 at Upper Little River [S of Harnett], 35.33556 N 78.78356 W, 39 m, 19 Apr 2000, R.L. Blinn, Solidago sp., 18 (00242831) (NCSU). Haywood Co.: Lake Junaluska, 35.52786 N 82.95976 W, 23 May 1954, H.V. Weems, Jr., 18 (00243718) (FSCA). Henderson Co.: Henderson County, no specific locality, 35.31936 N 82.4756 W, 679 m, 06 May 1955, D.H. Babeck, 1♀ (00242837) (NCSU). Hendersonville, 35.318616 N 82.461116 W, 01 Jun 1907, F. Sherman, paratype, 18 (00134213) (USNM). Macon Co.: Highlands, 35.05256 N 83.196946 W, 1158 m, 03 Jun 1957, J.R. Vockeroth, 18 (00113964) (CNC). Madison Co.: Rt 209 between Trust and Hot Springs, 35.837256 N 82.857396 W, 592 m, 20 May 1979, Wygodzinsky, Schuh, and Schmidt, 18 (00170145) (AMNH). McDowell Co.: Old Fort E of Ashville on I-40, 35.629486 N 82.16896 W, 430 m, 22 May 1984, C.B. Barr, 18 (00242951) (DAR). Montgomery Co.: Dark Mountain, 35.472626 N 79.95186 W, 271 m, 16 May 2000, R.L. Blinn, 18 (00242832) (NCSU). Uwharrie National Forest, SR 1306 at Barnes Creek, 35.479946 N 79.95736 W, 148 m, 23 May 1995, R.L. Blinn, Solidago sp., 18 (00242828), 1♀ (00242868) (NCSU). Moore Co.: SR 1459, 35.50896 N 79.595246 W, 118 m, 30 May 1996, R.L. Blinn, Solidago sp., 18 (00242835) (NCSU). Randolph Co.: SR 2458, 35.80816 N 79.598966 W, 203 m, 30 May 2003, L.L. Deitz, 1♀ (00242840) (NCSU). Robeson Co.: Hwy 74, 3.2 km E of Hwy 71, 34.657046 N 79.208036 W, 46 m, 10 Apr 1997, L.L. Deitz, 18 (00242834) (NCSU). Rockingham Co.: Jct. SR 2571 and SR 2572, 36.353086 N 79.567916 W, 213 m, 16 May 1995, R.L. Blinn, Solidago sp., 18 (00242827) (NCSU). Swain Co.: Cherokee, 35.47296 N 83.31336 W, 610 m, 29 May 1957, W.R.M. Mason, 1♀ (00113968) (CNC). Great Smoky Mountain National Park, Clingmans Dome, 35.55766 N 83.49376 W, 1924 m, 21 Jun 1941, A.L. Melander, 18 (00119974) (UCR). Great Smoky Mountain National Park [rest area Rt 441], 35.59826 N 83.41356 W, 1250 m, 25 Jun 1940, C.T. Brues, 18 (00170144) (AMNH). Transylvania Co.: Brevard, 35.236036 N 82.725936 W, 644 m, 21 May 1965, R.C. Froeschner, 18 (00133742) (USNM). Wilkes Co.: Wilkes County, no specific locality, 36.149376 N 81.137326 W, 296 m, 15 May 1958, D.A. Young, 18 (00242826) (NCSU). Yancey Co.: Yancey County, without specific locality, 35.913416 N 82.298566 W, 872 m, 28 Apr 1990, T. Daggy, 1♀ (00242838) (NCSU).  North Dakota: Billings Co.:  Bad Lands [Badlands Scenic Area], 47.217796 N 103.560466 W, 713 m, 15 Jun 1920, A.A. Nichol, 2♀ (00134217–00134218) (USNM). Bottineau Co.: 3 mi NE of Bottineau, Turtle Mountain foothills, 48.856 N 100.366666 W, 594 m, 23 Jun 2000, T.J. Henry, 18 (00134069) (USNM). Cass Co.: Fargo, 46.877226 N 96.789446 W, 22 Jun 1923, Wm. Baker, 18 (00134302) (USNM). Fargo, N at 19th St. and University, 46.905886 N 96.79686 W, 272 m, 17 Jun 2000, T.J. Henry, Solidago sp., 18 (00134068) (USNM). Dunn Co.: Killdeer [Mountain State Game] Management Area, 10 mi NW of Killdeer, 47.444666 N 102.978696 W, 823 m, 24 Jun 2000, T.J. Henry, Symphoricarpos occidentalis Hook. (Caprifoliaceae), 18 (00069990) (USNM). Kidder Co.: Kidder County, no specific locality, 46.760326 N 99.505146 W, 577 m, 14 Jul 1920, A.A. Nichol, paratypes, 28 (00134190–00134191) (USNM). Ramsey Co.: Devils Lake, 48.112786 N 98.864726 W, 19 Jul 1920, T.H. Hubbell, 18 (00127491) (UMMC). Stark Co.: 127 R Ave. 4 mi N of 48 R St. SW [9 mi S of South Heart], 46.733336 N 103.016666 W, 27 Jun 2000, T.J. Henry, Artemisia cana, 1♀ (00133739) (USNM). Wells Co.: Wells County, no specific locality, 47.645546 N 99.629786 W, 490 m, 03 Jul 1960, D. Kuske, 1♀ (00242952) (DAR).  Ohio: Ashland Co.:  Mohican Memorial State Forest, 40.607326 N 82.313896 W, 377 m, 30 May 1987, P.W. Kovarik, 18 (00093788) (TAMU). Ashtabula Co.: Crooked Creek Farm nr Hartsgrove, 41.6246 N 80.9796 W, 29 May 1963, J.C. and D. Pallister, 18 (00170136) (AMNH); 30 May 1963, J.C. and D. Pallister, 18 (00170137) (AMNH); 01 Jun 1963, J.C. and D. Pallister, 18 (00170132) (AMNH); 13 Jun 1963, J.C. and D. Pallister, 18 (00170133) (AMNH); 14 Jun 1963, J.C. and D. Pallister, 18 (00170134) (AMNH); 16 Jun 1963, J.C. and D. Pallister, 18 (00170135) (AMNH). Athens Co.: Athens County, no specific locality, 39.319576 N 82.096666 W, 191 m, 15 May 1955, unknown, 1♀ (00133813) (USNM). Clinton Co.: Wilmington, 39.442346 N 83.82426 W, 312 m, 17 Jul 1926, S.A. Watson, paratype, 18 (00134241) (USNM). Cuyahoga Co.: Bedford, 41.39316 N 81.53676 W, 10 Jun 1931, J.C. Pallister, 3♀ (00113248 [cat CMNH Acc 1906]–00113250 [cat CMNH Acc 1906]) (CMNH). Cleveland, 41.478496 N 81.690336 W, 207 m, unknown, 18 (00116329) (CUIC). Erie Co.: Resthaven Preserve, 41.39236 N 82.83386 W, 191 m, 06 Jun 2003, T. Pucci, 18 (00113253) (CMNH). Resthaven Wildlife Area, nr Herr Rd, 41.39966 N 82.83456 W, 191 m, 06 Jun 2003, T. Pucci, 1♀ (00113252) (CMNH). Franklin Co.: Columbus, 39.961116 N 82.998896 W, 29 May 1954, H.V. Weems, Jr., 18 (00243720) (FSCA). Portage Co.: Towner’s Woods [Park], 41.1716 N 81.30256 W, 335 m, 07 Jun 2003, T. Pucci, 18 (00113254) (CMNH). Summit Co.: Northfield, 41.32866 N 81.53916 W, 290 m, 30 May 1925, J.C. Pallister, 3♀ (00113244 [cat CMNH 36500]–00113246 [cat CMNH 36502]) (CMNH); 20 Jun 1927, J.C. Pallister, 1♀ (00113247 [cat CMNH 21850]) (CMNH).  Oklahoma: Nowata Co.:  6 mi N of Childers, 36.858086 N 95.539776 W, 212 m, 25 Apr 1972, D.C. Arnold, (Fabaceae), 18 (00242309) (OSEC). Payne Co.: Stillwater, 36.115566 N 97.058066 W, 28 Apr 1954, F.A. Fenton, 1♀ (00134310) (USNM); 28 May 1954, F.A. Fenton, 18 (00242310), 1♀ (00242311) (OSEC); 09 May 1978, D.C. Arnold, 68 (00242295– 00242300), 8♀ (00242301–00242308) (OSEC).  Oregon: Grant Co.:  Izee, Keerin’s Ranch, 44.067226 N 119.3856 W, 25 Jun 1935, Joe Schuh, 18 (00170158) (AMNH). Harney Co.: Frenchglen, 42.826946 N 118.915556 W, 1372 m, 21 Jun 1951, Borys Malkin, 18 (00242542) (CAS). Steens Mts, Loop Road (S leg) at Blitzen River Crossing, 42.639596 N 118.764096 W, 1568 m, 09 Jul 1999, J.D. Pinto, 18 (00119975 [cat UCRC ENT 00035344]) (UCR).  Pennsylvania: Berks Co.:  Reading, 40.352926 N 75.93066 W, 114 m, 11 Jun 1934, R.P. Seibert, 1♀ (00122396) (UID). Bucks Co.: Neshaminy Creek, E of Jamison, 40.269516 N 75.074066 W, 49 m, 03 Jul 1966, J. and W. Ivie, 1♀ (00170181) (AMNH). Centre Co.: Milesburg, Rt 144, 40.929576 N 77.782276 W, 242 m, 06 Jun 1974, T.J. Henry, 18 (00134076) (USNM). Crawford Co.: Hartstown Bog, 41.54716 N 80.362666 W, 309 m, 26 Jun 1919, D.M. DeLong, 1♀ (00134225) (USNM). Erie Co.: Charter Oak [s], 42.093546 N 80.151896 W, 218 m, 17 Jun 1917, J.N. Knull, 18 (00134212) (USNM); 11 Jul 1917, H.B. Kirk, 18 (00134211) (USNM). Franklin Co.: Chambersburg, 39.93756 N 77.661396 W, 14 May 1921, E.M. Craighead, 18 (00134220) (USNM); 08 Jun 1921, J.R. Stear, paratype, 18 (00134208) (USNM); 13 Jun 1921, J.R. Stear, paratype, 1♀ (00134223) (USNM); 07 Jun 1923, J.R. Stear, paratype, 18 (00134209), 1♀ (00134210) (USNM); 08 Jun 1923, J.R. Stear, paratype, 1♀ (00134222) (USNM); 23 Jun 1923, J.R. Stear, paratype, 1♀ (00134221) (USNM). Monroe Co.: Delaware Water Gap, 40.96756 N 75.12256 W, A.T. Slosson, 18 (00170128) (AMNH). Philadelphia Co.: Philadelphia, 40.01496 N 75.1056 W, 24 m, 21 Jun 1926, J.C. Lutz, 18 (00133811) (USNM); 08 Jun 1927, J.C. Lutz, 1♀ (00133821) (USNM); 26 Jun 1927, J.C. Lutz, 1♀ (00134308) (USNM).  South Carolina: Lexington Co.:  Cayce, 33.960366 N 81.077076 W, 70 m, 13 May 1988, A.G. Wheeler, Jr., Solidago sp., 1♀ (00133812) (USNM). Swansea, 33.740976 N 81.105346 W, 94 m, 13 May 1988, A.G. Wheeler, Jr., 18 (00133741) (USNM). Newberry Co.: Newberry, 34.274446 N 81.618896 W, 15 May 1988, A.G. Wheeler, Jr., 1♀ (00133822) (USNM). Pickens Co.: Rocky Bottom, 35.044216 N 82.804696 W, 527 m, 22 May 1934, J.N. Todd, 18 (00134298) (USNM).  South Dakota: Brookings Co.:  Brookings, 44.311396 N 96.798066 W, 493 m, 16 Jun 1891, unknown, 28 (00242502– 00242503) (CAS); 12 Jun 1923, H.H. Knight, paratype, 18 (00134233) (USNM); 12 Jun 1923, H.C. Severin, paratype, 1♀ (00134234) (USNM); 14 Jun 1931, H.C. Severin, 1♀ (00134329) (USNM). White, 44.434726 N 96.649446 W, 26 Jul 1922, H.C. Severin, 1♀ (00134332) (USNM). Butte Co.: Castle Rock, 44.964436 N 103.424356 W, 957 m, 29 Jun 1973, L.A. Kelton, Artemisia sp., 1♀ (00113948) (CNC). Custer Co.: Custer, 43.766676 N 103.598336 W, 19 Aug 1927, H.H. Knight, 3♀ (00134267–00134269) (USNM); 27 Jul 1991, D.C. Arnold, 2♀ (00242312– 00242313) (OSEC). Haakon Co.: Philip, 44.039436 N 101.66516 W, 660 m, 23 Jun 1923, H.C. Severin, paratype, 18 (00134232) (USNM). Harding Co.: Buffalo, 45.584176 N 103.545566 W, 19 Jun 1925, H.C. Severin, 1♀ (00134317) (USNM); 20 Jun 1925, H.C. Severin, 18 (00134312) (USNM). Jones Co.: Capa, 44.108896 N 100.977786 W, 538 m, 01 Jun 1921, H.C. Severin, 48 (00242498– 00242501) (CAS). Lake Co.: Chester, 43.890466 N 96.92546 W, 485 m, 17 Jun 1930, G.I. Gilbertson, 1♀ (00134331) (USNM). Lawrence Co.: Black Hills, 44.416676 N 103.708336 W, 29 Jun 1973, L.A. Kelton, 18 (00113945) (CNC). Deadwood, 44.376386 N 103.729446 W, 1381 m, 29 Jul 1927, H.H. Knight, 18 (00134240), paratype, 1♀ (00134231) (USNM). Englewood, 44.297966 N 103.783576 W, 1704 m, 18 Jun 1926, H.C. Severin, 18 (00134333) (USNM). Englewood, Black Hills, 44.416676 N 103.708336 W, 05 Aug 1971, L.A. Kelton, 18 (00113944) (CNC). Pennington Co.: Pactola Lake, Black Hills, 44.0736 N 103.47346 W, 1341 m, 05 Aug 1971, L.A. Kelton, 18 (00113943) (CNC). Roberts Co.: Browns Valley, 45.593826 N 96.830486 W, 299 m, 23 Jun 1927, H.C. Severin, 1♀ (00134330) (USNM). Shannon Co.: Pine Ridge, 43.019326 N 102.5656 W, 980 m, 20 Jun 1928, H.C. Severin, 5♀ (00134271–00134275) (USNM). Union Co.: 3 mi S of Jefferson, 42.559126 N 96.558896 W, 11 Jun 1949, Slater and Laffoon, 18 (00170152) (AMNH). Unknown co.: Dac., P.R. Uhler, 1♀ (00069969) (USNM).  Tennessee: Giles Co.:  Pulaski, 35.19976 N 87.03086 W, 25 May 1932, H.L. Dozier, paratype, 18 (USNM). rest stop 3.2 mi S of Hwy 129 on I-65, 35.327286 N 86.881696 W, 254 m, 18 May 1983, D.A. Rider, 18 (00242949) (DAR). Knox Co.: Knoxville, 35.960566 N 83.920836 W, 15 May 1917, Geo. G. Ainslie, (Cyperaceae), paratype, 1♀ (00134253) (USNM). Sevier Co.: Gatlinburg, 35.711666 N 83.515136 W, 413 m, 15 May 1988, D.A. Dean, 1♀ (00093793) (TAMU). Great Smoky Mountains National Park, Indian Gap, 35.61116 N 83.43856 W, 1585 m, 18 Jun 1957, W.R.M. Mason, 18 (00113967) (CNC). Warren Co.: Warren County, no specific locality, 35.682176 N 85.782526 W, 268 m, 22 May 1940, unknown, Solidago sp., 18 (00134304) (USNM). Wilson Co.: nr Gladeville, Rt 165, 36.133676 N 86.415046 W, 192 m, 29 May 1985, T.J. Henry and A.G. Wheeler, Jr, 18 (00133738) (USNM).  Utah: Box Elder Co.:  Snowville, 41.965566 N 112.708896 W, 09 Jun 1969, G.F. Knowlton, 18 (00134281) (USNM). Cache Co.: Logan, 41.735566 N 111.833616 W, 06 Jul 1955, S.L. Wood, 18 (00113957) (CNC); 16 Jul 1955, G.F. Knowlton, 18 (00134287) (USNM); 23 Jul 1963, G.F. Knowlton, 18 (00134288) (USNM). Wellsville, 41.632226 N 111.934446 W, 1389 m, 01 Jul 1948, R.S. Bailey, 18 (00134289) (USNM). Duchesne Co.: Currant Creek, 40.196686 N 110.894346 W, 2021 m, 30 Jul 1960, G.F. Knowlton, 18 (00134290) (USNM). Rich Co.: Garden City, 41.889176 N 111.3856 W, 06 Jul 1968, G.F. Knowlton and H.H. Knight, 18 (00134280) (USNM). Laketown, 41.825566 N 111.321676 W, 23 Jul 1963, G.F. Knowlton, 1♀ (00134291) (USNM); 06 Jul 1968, G.F. Knowlton and H.H. Knight, 18 (00134279) (USNM). Utah Co.: Hwy 80, 24 Aug 1966, L. and C.W. O’Brien, 1♀ (00134282) (USNM). Weber Co.: Huntsville, 41.260836 N 111.769176 W, unknown, 18 (00070115) (USNM). Ogden, 41.223066 N 111.973066 W, 24 Jun 1941, W. Don Fronk, 18 (00164894) (USNM).  Vermont: Bennington Co.:  5 mi N of Bennington, 42.938426 N 73.231946 W, 214 m, 25 Jun 1989, T.J. Henry and A.G. Wheeler, Jr, Solidago sp., 18 (00069970) (USNM). Windham Co.: Halifax, 42.768056 N 72.745376 W, 481 m, 03 Jul 1948, L.B. Burbank, 1♀ (00170172) (AMNH).  Virginia: Arlington Co.:  Chain Bridge, 38.929596 N 77.116846 W, 5 m, 05 Jun, N. Banks, 1♀ (00170178) (AMNH); 05 Jun 1906, D.H. Clemons, 18 (00069926) (USNM). Glencarlyn, 38.861676 N 77.129446 W, 30 May, N. Banks, 18 (00170130) (AMNH); 30 May 1906, D.H. Clemons, 1♀ (00069965) (USNM). Fairfax Co.: Annandale, off of Duncan Drive, 2 mi from Rt 236, 38.822926 N 77.230056 W, 81 m, 28 May 2004, K. Menard, 1♀ (00133931) (USNM).  Washington: Columbia Co.:  Dayton, 46.323896 N 117.971396 W, 1447 m, 06 Jul 1937, R.E. Miller, paratypes, 28 (00134166–00134167), 1♀ (00134168) (USNM).  West Virginia: Jefferson Co.:  Halltown, 39.31396 N 77.79776 W, 125 m, 03 Jun 1907, L.J. Condit, 18 (00134487) (USNM). Mineral Co.: Elk Garden, 39.38176 N 79.1556 W, 701 m, 25 Jun 1968, R. and E. Froeschner, 18 (00133743) (USNM). Monongalia Co.: Monongalia County, no specific locality, 39.6296 N 79.9566 W, 275 m, 19 May 1983 – 17 Jul 1983, T.L. Mason, Jr., 18 (USNM). Pendleton Co.: Seneca Creek, 38.715896 N 79.547166 W, 1158 m, 26 Jun 1966, O.R. Taylor, 1♀ (00170162) (AMNH). Pocahontas Co.: Cranberry Glades (Visitor Center), 38.200836 N 80.268336 W, 04 Jun 1955, H.V. Weems, Jr., 1♀ (00243726) (FSCA). Edray, 38.274566 N 80.091456 W, 734 m, 04 Jul 1986, Larochelle and Lariviere, 18 (00113946) (CNC). Preston Co.: Preston County, no specific locality, 39.4716 N 79.6836 W, 552 m, 19 May 1983 – 17 Jul 1983, T.L. Mason, Jr., 18 (USNM). Randolph Co.: Dryfork, 38.968726 N 79.502836 W, 672 m, 02 Jul 1986, Larochelle and Lariviere, 1♀ (00113947) (CNC). Ritchie Co.: Ritchie County, no specific locality, 39.2096 N 81.0516 W, 259 m, 19 May 1983 – 17 Jul 1983, T.L. Mason, Jr., 18 (USNM). Tucker Co.: Tucker County, no specific locality, 39.0966 N 79.6816 W, 505 m, 19 May 1983 – 17 Jul 1983, T.L. Mason, Jr., 18 (USNM).  Wisconsin: Columbia Co.:  Columbia County, no specific locality, 43.549886 N 89.484696 W, 250 m, Jul 1963, T.R. Yonke, 18 (00243095) (UMRM). Dane Co.: Dane County, no specific locality, 43.074316 N 89.484236 W, 305 m, 04 Jul 1962, T.R. Yonke, 18 (00243097) (UMRM); 17 Jun 1970, A.E. Akingbohungbe, 1♀ (00242069) (IRCW). Madison, 43.04546 N 89.427276 W, 279 m, 03 Jun 1948, W.S. Marshall, 18 (00242062) (IRCW). Madison, University of Wisconsin Arboretum, Wingum Marsh - Stevens Pond, 43.04866 N 89.43666 W, 267 m, 26 Jun 1967, C.B., D.W. Schwehr, 18 (00242063) (IRCW). Jefferson Co.: Fort Atkinson, 42.93126 N 88.85876 W, 259 m, 17 Jun 1916, J.G. Sanders, 18 (00242065) (IRCW). Jefferson County, no specific locality, 43.01276 N 88.79256 W, 259 m, 14 Jun 1962, J.T. Medler, 18 (00242064) (IRCW). Kenosha Co.: Benedict Prairie, E of Woodworth, 42.557546 N 87.99316 W, 223 m, 16 Jun 1998, W. Suter, 2♀ (00242054–00242055) (IRCW). Chiwaukee Prairie, T1N R23E Sec32 U:5, S of Phil Sander W of 2nd Ave., 42.50576 N 87.813646 W, 176 m, 11 Jul 1996, unknown, 18 (00242052), 1♀ (00242053) (IRCW). Kenosha, 42.57316 N 87.87166 W, 213 m, 12 Jun 1970, A.E. Akingbohungbe, 1♀ (00242068) (IRCW). Outagamie Co.: Mack State Wildlife Area, 44.46226 N 88.51236 W, 233 m, 15 Jun 1999, K. Kirk, 28 (00242056– 00242057) (IRCW). Rock Co.: Janesville, 42.67246 N 89.00576 W, 244 m, 17 Jun 1916, J.G. Sanders, 1♀ (00242067) (IRCW). Walworth Co.: Young Prairie, T4N R16E Sec5 NW4, 42.847026 N 88.636786 W, 246 m, 14 Jun 1995, unknown, 48 (00242046– 00242049), 2♀ (00242050–00242051) (IRCW). Waukesha Co.: Waukesha County, no specific location, 42.988376 N 88.193926 W, 272 m, 26 Jul 1962, T.R. Yonke, 18 (00243096) (UMRM).  Wyoming: Albany Co.:  Albany County, no specific locality, 41.666396 N 105.700556 W, 2133 m, 12 Aug 1952, R.R. Dreisbach, 18 (00127497), 1♀ (00127498) (UMMC). Big Horn Co.: Big Horn Mountains, 44.754996 N 107.782396 W, 1981 m, 17 Aug 1927, H.H. Knight, 18 (00134266) (USNM). Crook Co.: Sundance, 44.406396 N 104.375286 W, 30 Jul 1927, H.H. Knight, 28 (00134248–00134249), 3♀ (00134250– 00134251, 00134265) (USNM). Fremont Co.: Shoshone National Forest, 43.809136 N 109.563066 W, 14 Aug 1927, H.H. Knight, paratype, 18 (00093776) (TAMU), 18 (00134169), paratypes, 58 (00134242– 00134246), 1♀ (00134247) (USNM). Lincoln Co.: Alpine Junction, 43.172426 N 111.018546 W, 1725 m, 08 Aug 1972, L.A. Kelton, Solidago sp., 18 (00113954) (CNC). Niobrara Co.: Niobrara Co. stop 13, 42.780336 N 104.472466 W, 1568 m, 07 Jul 1943, R.E. Pfadt, 18 (00116600) (ESUW). Niobrara Co. stop 25, 42.780336 N 104.472466 W, 1568 m, 08 Jul 1943, R.E. Pfadt, 1♀ (00116601) (ESUW). Park Co.: Yellowstone National Park, 44.766676 N 110.233336 W, 16 Jul 1920, A.A. Nichol, paratype, 1♀ (00134172) (USNM). Sheridan Co.: Sheridan, 44.797226 N 106.955566 W, Metz, 38 (00121664–00121666), 1♀ (00121667) (CAS); 25 Jun 1947, O.A. Hills, 18 (00134297) (USNM). Teton Co.: Grand Teton National Park, Teton Science School near Kelly, 43.623066 N 110.622786 W, 24 Jul 1982 – 25 Jul 1982, S.C. Williams, 38 (00121670– 00121672), 1♀ (00121673) (CAS). Jackson, 43.50006 N 110.766666 W, 1920 m, 13 Jul 1920 – 17 Jul 1920, unknown, 18 (00170121) (AMNH). Uinta Co.: Fort Bridger, 41.316626 N 110.384326 W, 2034 m, 13 Jul 1965, Harry H. Knight, 1♀ (00134278) (USNM). Unknown co.: Wyoming, 16 Jul 1949, W.F. Chamberlain, 18 (00093789) (TAMU).
description: Published as part of Schwartz, Michael D., 2011, Revision And Phylogenetic Analysis Of The North American Genus Slaterocoris Wagner With New Synonymy, The Description Of Five New Species And A New Genus From Mexico, And A Review Of The Genus Scalponotatus Kelton (Heteroptera: Miridae: Orthotylinae), pp. 1-290 in Bulletin of the American Museum of Natural History 2011 (354) on pages 130-133, DOI: 10.1206/354.1, http://zenodo.org/record/5407316