Nototriadophlebia pritykinae Béthoux & Anderson 2023, n. gen., n. sp.
Creators: name: Béthoux, Olivier
name: Anderson, John M.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Odonata
Permophlebiidae
Nototriadophlebia
Nototriadophlebia pritykinae

description: Nototriadophlebia pritykinae n. gen., n. sp.
(Figs 7-9)
urn:lsid:zoobank.org:act: BB1717F8-0D90-41B5-9D88-68689FB9B410
HOLOTYPE. — Specimen BP /2/20950ab, adpression preserving four wings (Anderson collection, Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg, South Africa; Figs 7, 8).
PARATYPE. — Specimen BP /2/20986ab, adpression preserving a single wing (Anderson collection, Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg, South Africa; Fig. 9).
E ScP
RA RP1 1 -rp 2
CuA
ScP F
RA RP1 1-rp2
DIAGNOSIS. — Area between the anterior wing margin and RA (distal to the nodus) widening gradually to ª/5 of wing length, where it is as wide as the RA-RP1 area (most prominent in forewing); first I- in the RP3+4-MA area (i.e. RP3+4 pseudo-fork; large orange arrows on Fig. 8A, F) arising before RP3+4 mid-length; CuP+AA stem splitting into CuA and CuP+AA (as opposed to Cu and AA; large red arrow on Figs 8F, 9A, C, D); AA reaching the posterior wing margin basal to the end of ScP; between the endings of MA and AA, posterior wing margin comparatively straight.
ETYMOLOGY. — The species epithet honours Lyudmila N. Pritykina, for her contribution to our knowledge of fossil Odonata, and of Triadophlebiomorpha in particular.
TYPE LOCALITY AND STRATIGRAPHY. — Both holotype and paratype were collected at Birds River (locality code ‘Bir 111’; see Anderson & Anderson 1984), South Africa; Molteno Formation; lower Carnian, Upper Triassic (Anderson et al. 1998).
GENERAL DESCRIPTION
Fore- and hindwings subequal, very elongated, broadest at mid-length; wing length about 74 mm (as derived from holotype specimen; possibly down to 70 mm), maximum width about 11.0 mm (possibly down to 10.2 mm), width opposite nodus 7.1-7.6 mm); petiole long (about 12.8 mm) and narrow (width about 2.4 mm); ScP reaching the anterior wing margin basal to the first third of wing length; distal to the nodus, area between anterior wing margin and RA broadening gradually, at its broadest opposite the end of Irp 1+2 -rp 3+4; structure of the nodus and origin of RP+MA, RP and MA unknown; RP1/ RP2 fork in a position more distal than that of the point of origin of the first I- in the RP3+4 area; Irp 1 -rp 2 weakly differentiated, zigzagging along RP1; occurrence of 3 I- (and I+ between them) forming RP2 posterior pseudo-branches in the distal part of the area between RP2 and Irp 1+2 -rp 3+4; RP3+4 with a marked angle opposite the origin of the first I- (between RP3+4 and MA; large orange arrows on Fig. 8A, F); the two first I- in the area between RP3+4 and MA branched; in this area, total of about 8 I- reaching the posterior wing margin; MA simple, rectilinear, very close to RP3+ 4 in its basal third; marked inflexion of the posterior wing margin opposite the ending of MA; MP originating from MP +Cu+AA with a slight obliquity; branches of MP and CuA not clearly distinguishable (probable occurrence of a CuAprlg), CuA with 2 branches at least, and with a I- distally branched between its two (first posterior) branches; MP and CuA covering more than a quarter (less than a third) of the posterior wing margin length; Cu+AA splitting into CuA and CuP+AA, CuA diverging very obliquely; CuP rectilinear, with about 4-5 posterior branches and I+ between them; AA gently curved, reaching the posterior wing margin basal to the end of ScP.
SPECIMEN DESCRIPTIONS
Specimen BP/2/20950 (Figs 7; 8)
Adpression preserving four wings, with side BP/2/20950a preserving the right forewing (isolated) and left wings in ventral aspect, and right hindwing in dorsal aspect (the three latter largely overlapping); and its reverse side BP/2/20950b.
Right forewing (Figs 7; 8A, B). Well preserved, with area basal to the nodus and small portion of apex missing; preserved length 55.6 mm, maximum width 10.7 mm, width opposite nodus about 7.6 mm.
Left forewing (Figs 7; 8 C-E). Only anterior-most area decipherable; preserved length 73.2 mm; estimated total length about 74.2 mm.
Right hindwing (Figs 7; 8C, D, F). Well preserved, with nodal area and part of apex missing, and venation in the petiole difficult to decipher; basal to the nodus, RP and MA not distinguishable from each other (assumed to closely overlap); cubito-anal area preserved, showing a CuP+AA stem.
Left hindwing (Figs 7; 8C, D). Barely decipherable but for the posterior wing margin, with a sharp inflexion opposite the end of MA.
description: Published as part of Béthoux, Olivier & Anderson, John M., 2023, New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata), pp. 479-496 in Geodiversitas 45 (17) on pages 491-494, DOI: 10.5252/geodiversitas2023v45a17, http://zenodo.org/record/8399030