Tylopus flavolineatus Likhitrakarn & Golovatch & Panha 2016, sp. nov.
Creators: name: Likhitrakarn, Natdanai
name: Golovatch, Sergei I.
name: Panha, Somsak
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Polydesmida
Paradoxosomatidae
Tylopus
Tylopus flavolineatus

description: Tylopus flavolineatus sp. nov.
urn:lsid:zoobank.org:act: 4A4DA74B-774B-45F5-A60F-96EB516022D2
Figs 1–3, 26
Diagnosis
Differs in the presence of middle and lateral yellow stripes on the body, by the sterna bearing evident cones caudally near each coxa, and in ♂ femur 5 showing a large distovental adenostyle, coupled with the gonopods being rather slender, with a quite large, slightly curved and pointed process h.
Etymology
To emphasize the middle and lateral yellow stripes on the body; adjective.
Material examined
Holotype
THAILAND: ♂, Wat Tham Santisuk, Nong Muang, Lopburi, 15°12'10" N, 100°39'50" E, 145 m a.s.l., 7 Jun. 2008, leg. C. Sutcharit & N. Likhitrakarn (CUMZ).
Paratypes
THAILAND: 2 ♂♂, 2 ♀♀ (CUMZ), same data as for holotype.
Description
MEASUREMENTS AND COLOUR. Length 13.8–16.4 (♂) or 14.2–16.5 mm (♀), width of midbody pro- and metazonae 0.83–0.92 and 1.21–1.33 mm (♂) or 1.19–1.28 and 1.68–1.73 mm (♀), respectively. Coloration of alcohol material after six years of preservation dark brown; a pair of paramedian longitudinal stripes flanking a narrow, sometimes vague, brown, axial line; paraterga and area of pleurosternal carinae contrasting light yellow to yellowish; legs and venter dark brown to dark yellow (Fig. 1).
HEAD. Clypeolabral region densely, vertex sparsely, setose; epicranial suture distinct. Antennae rather short (Fig. 1B), clavate (antennomere 6 highest), extending behind body segment 3 (♂) or 2 (♀) when stretched dorsally. In width, segment 2 = 3 <head <collum <segment 4 <5–15; thereafter body gently and gradually tapering. Collum with three transverse rows of very small setigerous tubercles: 6+6 anterior, 4+4 intermediate and 5+5 posterior; with a small lateral denticle at about 1/3 collum length in front of caudal corner; the latter very broadly rounded anteriorly, declined ventrad, caudal corner not surpassing rear tergal margin.
BODY. Tegument smooth and shining, prozonae and surface below paraterga largely finely shagreened, metaterga rather smooth and leathery, posterior halves faintly rugulose, surface below paraterga finely microgranulate (Fig. 1 A–G). Postcollum metaterga with four transverse rows of very small setigerous tubercles or setae: an anterior transverse row of 4–6+4–6, intermediate transverse rows (one pre-sulcus, the other post-sulcus) of 3–4+3–4, and a posterior transverse row of 5–7+5–7; tergal setae long, strong, slender, about 2/3 of metatergal length (Fig. 1A, D, F). Axial line visible both on pro- and metazonae. Paraterga well-developed (Fig. 1A, D, F), especially so in ♂, lying rather high (at upper 1/3 of body height), anterior edge rounded and narrowly bordered, fused to callus; caudal corner very narrowly rounded to pointed. Paraterga 2 and 3 each with two evident setigerous incisions at lateral edge (Fig. 1A). Following poreless segments with two evident setigerous incisions; following pore-boring segments often with one incision in front of pore (Fig. 1C – D). Calluses on paraterga delimited by a sulcus both dorsally and ventrally. Ozopores evident, lateral, lying in an ovoid groove at about 1/4 metatergal length in front of posterior edge of metaterga. Transverse sulcus usually distinct (Fig. 1A, D, F), slightly incomplete on segments 4 and 19, complete on segments 5–18, rather wide, line-shaped, rather deep, not reaching bases of paraterga, beaded at bottom. Stricture between pro- and metazonae evident, wide and rather deep, ribbed at bottom down to base of paraterga (Fig. 1A, C–F). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2 and 3, thereafter increasingly reduced and broken, remaining as a front bulge and a caudal tooth until segment 10 (♂) or 6 (♀), on following segments retained as a small caudal tooth until segment 18 (♂) or absent starting with segment 7 (♀). Epiproct (Fig. 1 E–G) conical, flattened dorsoventrally, with two evident, small, rounded apical papillae; tip subtruncate; pre-apical lateral papillae evident, lying close to tip. Hypoproct roundly subtriangular, setigerous knobs at caudal edge small and well-separated (Fig. 1G).
STERNA. Densely setose, with a small, but evident cone caudally near each coxa, rear cones being a little larger than fore ones (Fig. 2E); an entire, large, tongue-shaped sternal lobe between ♂ coxae 4 (Fig. 1H, I). Legs rather long and slender, midbody ones ca 1.2–1.3 (♂) or 0.9–1.0 (♀) as long as body height (Fig. 2G); ♂ femora 5 (Fig. 2F) each with a very strong, distovental, densely pilose adenostyle in distal 1/3, ♂ prefemora 10–12 each with a small parabasal adenostyle (Fig. 2G); ♂ prefemora 2 until segment 12 each with microgranulations; tarsal brushes present until ♂ leg 6.
GONOPOD. Rather simple (Figs 2 A–D, 3); coxa slightly curved caudad, sparsely setose distoventrally. Prefemur as usual, densely setose, about 1/3 as long as femorite + postfemoral part. Femorite slightly curved and rather slender, with an evident mesal groove and a strong distolateral sulcus demarcating a postfemoral part; lobe l rather small, rounded apically, process h large, slightly curved and pointed; solenophore (sph) clearly curved mesad, long, vaguely bifid, lamina medialis supporting a long flagelliform solenomere (sl).
Remark
In the normal condition, the solenomere is sheathed by the solenophore (Fig. 2 A–D), broken off in Fig. 3.
description: Published as part of Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak, 2016, The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina, pp. 1-47 in European Journal of Taxonomy 195 on pages 3-6, DOI: 10.5852/ejt.2016.195, http://zenodo.org/record/3837846