Pseudodrobna natator Audo & Winkler & Charbonnier 2021, n. comb.
Creators: name: Audo, Denis
name: Winkler, Norbert
name: Charbonnier, Sylvain
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Penaeidae
Pseudodrobna
Pseudodrobna natator

description: Pseudodrobna natator (Glaessner, 1945) n. comb. (Figs 1 A-H; 2A)
Penaeus natator Glaessner, 1945: 700-702, fig. 3.
Penaeus libanensis – Roger 1946: 23-26, figs 16-17, pl. 1 fig. 4 not pl. 3 fig. 1 (pro parte).
Drobna sp. I – Roger 1944: 848; 1946: 29-31, fig. 21, pl. 1 fig. 6.
Metapenaeopsis natator – Charbonnier et al. 2017: 69-74, figs 121-141. TYPE MATERIAL. — Holotype NHMUK 59690 (part and counterpart, Fig. 1A); paratypes NHMUK 59511, In 29563, In 29578 (Fig. 1B). As stated byCharbonnier et al. (2017), the original material also included three additional paratypes, now excluded from the type series, and assigned to Palaeobenthonectes arambourgi (Roger, 1946) (NHMUK 59688, 59689) and Carpopenaeus septemspinatus (Dames, 1886) (NHMUK I 148), respectively.
TYPE LOCALITY. — Sahel Alma, Kesrouane district, Lebanon.
TYPE AGE. — Late Cretaceous, late Santonian, Nannofossil biozone UC13 of Burnett (1998) - UC17 pro parte Sissingh (1977).
DISTRIBUTION. — Santonian (Lebanon).
ADDITIONAL MATERIAL. — Same as Charbonnier et al. (2017): five specimens MNHN.F.B18859 (Fig. 2A) (also figured by Roger 1946: pl. 1, fig. 4), MNHN.F.B18886 (figured by Roger 1946: pl. 1, fig. 6),MNHN.F. A30696 (Figs 1 F-H), A30697 (Fig. 1C), A30702 (Arambourg coll., Fig. 1E). All are from Sahel Alma.
DESCRIPTION
See Appendix 1.
DISCUSSION
Pseudodrobna natator n. comb. was assigned to Penaeus Fabricius, 1798 without real justification by Glaessner (1945), as it has often been the case for numerous fossils (Pérez Farfante & Kensley 1997; Charbonnier et al. 2017). Charbonnier et al. (2017) justified the assignment to Penaeidae, mostly by elimination, as, unfortunately, diagnostic characters of Penaeidae are difficult to observe and mostly defined on extant species (see Pérez Farfante & Kensley 1997). Later, Charbonnier et al. (2017), not aware of the work by Winkler (2017), assigned P. natator n. comb. to Metapenaeopsis Bouvier, 1905 based upon the rostrum with only suprarostral teeth (Fig. 1E), the epigastric tooth separated from rostral teeth, the marked antennal and pterygostomial spines and pubescent (i.e. covered by setae) exoskeleton (Fig. 1F). However, in the light of the work by Winkler (2017), it appears that P. natator n. comb. is more closely allied to Pseudodrobna Winkler, 2017 than Metapenaeopsis: indeed, as Pseudodrobna kenngotti Winkler, 2017 (type species and only species assigned to Pseudodrobna by Winkler 2017 – Fig. 1I, J), P. natator n. comb. possesses a rather short cephalothoracic shield, almost as long as high in lateral view (vs distinctly longer in Metapenaeopsis), a rather short, upturned rostrum (vs longer in Metapenaeopsis). For these reasons, we assign Penaeus natator to Pseudodrobna.
description: Published as part of Audo, Denis, Winkler, Norbert & Charbonnier, Sylvain, 2021, Pseudodrobna natator n. comb., a new link between crustacean faunas from the Jurassic of Germany and Cretaceous of Lebanon, pp. 209-218 in Geodiversitas 43 (8) on pages 210-212, DOI: 10.5252/geodiversitas2021v43a8, http://zenodo.org/record/4701081