Hendelia Beckeri Czerny 1903
Creators: name: Owen Lonsdale
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Clusiidae
Hendelia

description: Genus HENDELIA Czerny
Hendelia Czerny 1903: 83. Type species, Hendelia beckeri Czerny 1903, by monotypy. Gender feminine. Becker 1905: 39 [catalogue, Palaearctic]; Kertész 1906: 320 [key]; Williston 1908: 320 [comparison to Heteroneura]; Melander & Argo 1924: 27 [family and genus revision]; Czerny 1928: 4 [family and genus revision, Palaearctic]; Schulze et al. 1930: 1517 [nomenclator]; Neave 1939b: 617 [nomenclator]; Hennig 1952: 227 [discussion of immature stages in Clusiidae]; Frey 1960: 11 (as genus and subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, world]; Trojan 1962: 21 [key, Poland]; Soós 1963a: 177 [redefinition, key, relationship to Prohendelia and Parahendelia], 1964a: 7 [key], 1981: 96 [Hungary, key], 1984: 11 [catalogue, Palaearctic]; Cole & Schlinger 1969: 426 [citation]; Sasakawa 1977: 238 [catalogue, Orient], 2011b: 13 [Oriental fauna]; Ferrar 1987: 122 [immature stages, illustrations of larva and puparium]; Stackelberg 1988: 505 [key, European USSR]; Greve 2005: 3 [Norway]; Sueyoshi 2006: 4 [Japan], 2014: 619 [as genus and subgenus, catalogue, Japan]; Papp et al. 2006: 190 [identified and unidentified species (including possible new subgenus), Thailand]; Lonsdale & Marshall 2007b: 120 [review, biology, redefinition, comparison to Clusiodes, synonymy of Aristohendelia Frey, Burmanochorea Frey, Labomyia Frey, Parahendelia Frey, Prohendelia Frey, Microclusaria Frey, Steleoclusiodes Frey and Xenoclusia Frey, maps for Palaearctic species], 2008b: 5 [Fiji], 2011 [revision, New world], 2010: 1044 [Manual of Central American Diptera]; Lonsdale 2009b: 230 [key to genera in Australia]; Lonsdale et al. 2010: 267 [family revision, key]; Kahanpää 2014: 288 [checklist, Finland].
Labomyia Frey 1928a: 14. Type species, Labomyia mirabilis Frey 1928, by original designation. Gender feminine. Neave 1939b: 842 [nomenclator]; Hennig 1952: 227 [discussion of immature stages in Clusiidae]; Frey 1960: 26 [catalogue, world]; McAlpine 1960: 67 [key]; Soós 1964a: 6 [key], 1968: 4 [catalogue, Americas south of USA]; Caloren & Marshall 1998: 263 [as junior synonym of Clusiodes]; Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].
Aristohendelia Frey 1960: 14 (as subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, World]. Gender feminine. Type species, Hendelia extensicornis Frey 1960, by original designation. Sasakawa 1977: 238 [catalogue, Orient]; Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].
Burmanochorea Frey 1960: 14 (as subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, world]. Type species, Hendelia variabilis Frey 1960, by original designation. Gender feminine. Sasakawa 1977: 238 [catalogue, Orient]; Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].
Prohendelia Frey 1960: 13 (as subgenus) [family and genus revision, key to subgenera and species, family and genus revision, Indoaustralia, catalogue, world]. Type species, Hendelia orientalis Frey 1960, by original designation. Gender feminine. Soós 1963a: 177 [new status as genus, key, relationship to Hendelia and Parahendelia], 1964a: 7 [key]; Sasakawa 1977: 238 [catalogue, Orient]; Balthazar 1990: 417 [catalogue, Philippines]; Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].
Microclusiaria Frey 1960: 19 (as subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, world]. Gender feminine. Type species, Clusiodes obscuripennis Frey 1960, by original designation. Sasakawa 1977: 237 [catalogue, Orient]; Balthazar 1990: 417 [catalogue, Philippines]; Papp et al. 2006: 190 [identified and unidentified species; Thailand]. Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].
Steleoclusiodes Frey 1960: 18 (as subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, world]. Type species, Clusiodes nigrifrons Frey 1960, by original designation. Gender masculine. Sasakawa 1977: 237 [catalogue, Orient], 1987: 101 [key, Orient/Pacific]; Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].
Xenoclusia Frey 1960: 21 [catalogue, world]. Type species: Xenoclusia latifrons Frey 1960 [by original designation]. Gender feminine. Sasakawa 1977: 239 [catalogue, Orient]; Soós 1964a: 7 [key]; Balthazar 1990: 417 [catalogue, Philippines]; Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].
Parahendelia McAlpine 1960: 83. Type species, Parahendelia latifrons McAlpine 1960, by original designation (= Hendelia armiger Lonsdale & Marshall). Gender feminine. Soós 1963a: 179 [key, relationship to Hendelia and Prohendelia], 1964a: 4 [key]; Pitkin & Evenhuis 1989: 536 [catalogue, Australasia/Pacific]. Marshall 2000: 5 [as junior synonym of Clusiodes); Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].
Prophedelia. Misspelling of Prohendelia. Hua 2006: 160 [catalogue—not known from China]. Lonsdale & Marshall 2007b [citation].
aberrans (Frey). OR: Malaysia (Sabah), Philippines.
Clusiodes aberrans Frey 1928b: 106. Philippines. Oriental Mindoro Province: “Calopan” [= Calapan]. HT ♀, depository unknown [originally deposited UZMH]. Delfinado & Hardy 1971: 29 [catalogue Philippine types]. Clusiodes (Microclusiaria) aberrans. Frey 1960: 19 [Philippines, catalogue, world]; Sasakawa 1977: 237 [catalogue, Orient]; Balthazar 1990: 417 [catalogue, Philippines].
Clusiodes (Clusiaria) aberrans. Sasakawa 1987: 103 [key, Orient/Pacific].
Hendelia aberrans. Lonsdale & Marshall 2007b: 147 [generic combination]; Sasakawa 2011b: 14 [Malaysia].
aenigma Frey. OR: Burma.
Hendelia (Burmanochorea) aenigma Frey 1960: 15. Burma. NE Burma: Kambaiti, 2000m. HT ♀ UZMH. Frey 1960: 15 [catalogue, world]; Sasakawa 1977: 238 [catalogue, Orient].
Hendelia aenigma. Lonsdale & Marshall 2007b: 147 [list].
amerinx Lonsdale & Marshall. AU: Fiji.
Hendelia amerinx Lonsdale & Marshall 2008b: 7. Fiji. Viti Levu: Naitasiri Province, 4km WSW Colo-i-Suva Village, Mt. Nakobalevu, 372 m. HT ♂ FNIC. Lonsdale & Marshall 2008b: 7 [illustrations of male genitalia].
angulosa (Sueyoshi). PA: Japan.
Clusiodes angulosus Sueyoshi 2006: 6. Japan. Honshu: Sarugajo, Kitaibaraki City, Ibaraki Prefecture, Honshu. HT ♂ BLKU. Sueyoshi 2006: 6 [map, illustrations of head and male and female genitalia, photo of wing], 2011: 7 [Japan, pictoral key], 2014: 618 [catalogue, Japan]; Lonsdale & Marshall 2007b: 154 [lapsus]. Hendelia angulosa Lonsdale & Marshall 2007b: 147 [generic combination]; Sueyoshi 2012: 27 [Japan].
arguta (McAlpine). AU: Australia (New South Wales, Victoria).
Clusiodes arguta McAlpine 1960: 82. Australia. New South Wales: Blue Mountains, Wentworth Falls. HT ♂ AMS. Clusiodes (Clusiodes) arguta. Sasakawa 1987: 102 [key, Orient/Pacific].
Clusiodes argutus. Pitkin & Evenhuis 1989: 535 [catalogue, Australasia/Pacific].
Hendelia arguta. Lonsdale & Marshall 2007b: 147 [generic combination].
armiger Lonsdale & Marshall. AU: Australia (New South Wales, Queensland).
Parahendelia latifrons McAlpine 1960: 84. Australia. New South Wales: Blue Mountains, Wentworth Falls. HT ♂ AMS. McAlpine 1960: 84 [illustrations of head], 1976: 75 [discussion of vibrissal structure]; Pitkin & Evenhuis 1989: 536 [catalogue, Australasia/Pacific].
Clusiodes latifrons. Marshall 2000: 3 [generic combination, male agonistic behaviour, photos]. Hendelia armiger Lonsdale & Marshall 2007b: 118 [generic combination, replacement for name preoccupied by Frey (1960)].
beckeri Czerny. PA: Austraia and central Europe to Norway, Finland, eastern and northern coasts of the Black Sea, central and eastern Russia, Japan, China (Sichuan, Zeijiang)*, South Korea *.
Hendelia Beckeri Czerny 1903: 84. Austria. Bad Hall. ST ♂♀, depository unknown. Czerny 1903: 84 [illustrations of body and heads of both sexes], 1928: 4 [redescription, illustrations of head and body, redescription of females as nigripalpis Czerny]; Becker 1905: 39 [catalogue, Palaearctic].
Hendelia beckeri. Melander & Argo 1924: 27 [summary of original description]; Tuomikoski 1936: 183 [Finland]; Trojan 1962: 21 [key, redescription, illustrations of male and female heads]; Sasakawa 1963: 18 [Japan]; Hackman 1980: 148 [checklist, Finland]; Soós 1981: 97 [Hungary, illustrations of habitus, head], 1984: 11 [catalogue, Palaearctic]; Greve 1983: 55 [habitat, Norway], 2005: 3 [considered rare, Norway]; Greve & Midtgaard 1986: 91 [Norway]; Zuska 1987: 235 [Czech Republic]; Mamaev 1987: 115 [Russian Far East]; Stackelberg 1988: 505 [key, illustration of female head, European USSR]; Ceianu 1989: 29 [catalogue, Romania]; Roháček 1995: 125 [puparia in decaying alder and birch, adults on aspen trunks, biology and habitat, Czech and Slovak Republics], 1997: 75 [checklist, Czech and Slovak republics], 2009: 219 [photo of male, Slovakia], 2012: 99 [central Slovakia], 2013: 128 [central Slovakia]; Merz 1998: 276 [checklist, Switzerland]; Yagi & Sawada 2000: 40 [Mamaev collection holdings]; Gärdenfors 2000: 222 [Red List of Swedish species]; Roháček et al. 2005: 280 [catalogue, habitat, considered local and rare in Czech Republic, Czech Republic]; Barták & Kubík 2005: 280 [Czech Republic]; Sueyoshi 2006: 14 [Japan], 2011: 6 [Japan, pictoral key], 2012: 28 [Japan], 2014: 619 [catalogue, Japan]; Lonsdale & Marshall 2007b: 147 [biology, illustrations of genitalia and heads, relationships]; Schacht et al. 2007: 66 [Germany]; Nilsson 2009: 5 [Sweden, illustration of female]; Adaschkiewitz 2009: 287 [Germany]; Kahanpää 2014: 288 [checklist, Finland].
Hendelia nigripalpis Czerny 1928: 5 [illustration of head]. Austria. ST ♀ [= ♀ ST of H. Beckeri Czerny], depository unknown. Frey 1928c: 108 [catalogue, Finland]; Tuomikoski 1936: 183 [as synonym of beckeri Czerny]. Hendelia (Hendelia) beckeri. Frey 1960: 13 [redescription, catalogue, World].
bisetosa (Mamaev). PA: Russia (Far East).
Clusiodes bisetosa Mamaev 1974: 115. Russia. Far East: Kedrovaja Pad Nature Reserve. HT ♂ MNHA [not in ZIN, as stated in original publication, examined by author]. Mamaev 1974: 115 [illustration of surstylus]; Yagi & Sawada 2000: 41 [Mamaev collection holdings].
Hendelia bisetosa. Lonsdale & Marshall 2007b: 147 [generic combination, map].
bismarckensis (Sasakawa). AU: Papua New Guinea (Lavongai Island).
Clusiodes bismarckensis Sasakawa 1974: 160. Papua New Guinea. Bismarck Archipelago: Lavongai Island, Banatam. HT ♂ UZMC. Sasakawa 1974: 160 [illustrations of male genitalia], 1987: 101 [key, Orient/Pacific]; Pitkin & Evenhuis 1989: 535 [catalogue, Australasia/Pacific].
Hendelia bismarckensis. Lonsdale & Marshall 2007b: 147 [generic combination].
boliviensis Lonsdale & Marshall. NT: Bolivia.
Hendelia boliviensis Lonsdale & Marshall 2011: 4. Bolivia. La Paz Department: Caranavi, ca. 10km NW road to ENTEL tower, 1400 m, 15°46′35ʺS, 67°35′48ʺW. HT ♀ ANCB. Lonsdale & Marshall 2011: 4 [illustration of female genitalia].
campa Lonsdale & Marshall. NT: Mexico.
Hendelia campa Lonsdale & Marshall 2011: 4. Mexico. Veracruz State: Xalapa. HT ♂ TAUI. Lonsdale & Marshall 2011: 4 [illustrations of male genitalia, photo].
chaetostylotis (Sasakawa). AU: Indonesia (West Papua).
Clusiodes (Clusiodes) chaetostylotis Sasakawa 1987: 103. Indonesia. West Papua: Kebar Valley, Vogelkop, W of Manokwari, 550 m. HT ♂ BPBM [Type No. 13580]. Sasakawa 1987: 103 [illustration of male genitalia]. Hendelia chaetostylotis. Lonsdale & Marshall 2007b: 147 [generic combination].
clara (McAlpine). AU: Australia (Queensland).
Clusiodes clara McAlpine 1960: 83. Australia. Queensland: Silkwood, near Innisfail. HT ♀ AMS. Clusiodes (Clusiodes) clara. Sasakawa 1987: 103 [key, Orient/Pacific].
Clusiodes clarus. Pitkin & Evenhuis 1989: 535 [catalogue, Australasia/Pacific].
Hendelia clara. Lonsdale & Marshall 2007b: 147 [generic combination].
dasyta (Sasakawa). AU: Solomon Islands (New Georgia Island).
Clusiodes (Columbiella) dasytus Sasakawa 1987: 113. Solomon Islands. New Georgia Island: Munda, 1– 30m. HT ♂ BPBM [Type No. 13587]. Sasakawa 1987: 113 [illustration of male genitalia].
Hendelia dasyta. Lonsdale & Marshall 2007b: 147 [generic combination].
extensicornis Frey. OR: Burma, Thailand.
Hendelia (Aristohendelia) extensicornis Frey 1960: 14. Burma. NE Burma, Kambaiti, 2000 m. ST 6♂ 3♀ UZMH. Frey 1960: 14 [illustrations of head, catalogue, world]; Papp et al. 2006: 191 [Thailand].
Hendelia extensicornis. Lonsdale & Marshall 2007b: 147 [relationships].
femorata (Sasakawa). AU: Indonesia (Papua).
Clusiodes (Clusiodes) femoratus Sasakawa 1987: 105. Indonesia. Papua Province: Ifar, east end of Cyclops Mts., 150 m.
HT ♀ BPBM [Type No. 13581].
Hendelia femorata. Lonsdale & Marshall 2007b: 147 [generic combination].
formosana (Hennig). OR: Taiwan.
Clusiodes (Microclusiaria) formosana Hennig 1938: 128. Taiwan. Paiwan District: northern Paroe. HT ♂ DEI. Clusiodes formosana. Hennig 1941: 146 [Taiwan]; Hua 2006: 160 [catalogue—not known from China]. Clusiodes (Microclusiaria) formosana. Frey 1960: 19 [catalogue, world]; Sasakawa 1977: 237 [catalogue, Orient]. Clusiodes (Clusiaria) formosana. Sasakawa 1987: 103 [key, Orient/Pacific].
Hendelia formosana. Lonsdale & Marshall 2007b: 147 [generic combination].
freyi (Soós). OR: Taiwan.
Prohendelia (Prohendelia) freyi Soós 1963a: 179. Taiwan. Kosempo. HT ♂ HNHM. Soós 1963a: 179 [illustrations of head]; Sasakawa 1977: 239 [catalogue, Orient].
Hendelia freyi. Lonsdale & Marshall 2007b: 147 [generic combination].
fulvifasciata Sasakawa. OR: Malaysia (Sabah).
Hendelia fulvifasciata Sasakawa 2011b: 14. Malaysia. Sabah: 19 km N of Kalabakan, forest camp. HT ♂, BPBM [Type No. BPBM 17352]. Sasakawa 2011b: 14 [illustrations of male genitalia].
geniculata Sasakawa. OR: Malaysia (Sabah).
Hendelia geniculata Sasakawa 2011b: 16. Malaysia. Sabah: 9.8 km SW of Tenom, forest camp. HT ♂, BPBM [Type No. BPBM 17355]. Sasakawa 2011b: 16 [illustrations of male genitalia].
gladiator (McAlpine). AU: Australia (New South Wales, Queensland, Tasmania, Victoria). Clusiodes gladiator McAlpine 1960: 81. Australia. New South Wales: National Park, south of Sydney. HT ♂ AMS. McAlpine 1960: 81 [illustrations of head and wing], 1976: 75 [discussion of spiral vibrissae and sexual dimorphism, illustrations of head and male postabdomen]; Pitkin & Evenhuis 1989: 535 [catalogue, Australasia/Pacific]; Marshall 2000: 3 [male agonistic behaviour, photos].
Clusiodes (Clusiodes) gladiator. Sasakawa 1987: 102 [key, Orient/Pacific].
Hendelia gladiator. Lonsdale & Marshall 2007b: 147 [generic combination]; Lonsdale et al. 2010: 241 [genes sequenced for phylogenetic analysis].
heliconiae Lonsdale & Marshall. NT: Bolivia, Costa Rica, Ecuador.
Hendelia heliconiae Lonsdale & Marshall 2011: 6. Ecuador. Pichincha Province: Maquipucuna Biological Research Station, river trail, 1200 m. HT ♂ QZAC. Lonsdale & Marshall 2010: 1046 [illustration of female genitalia as Hendelia sp.], 2011: 6 [illustrations of male and female genitalia, photo].
iotoides (Sasakawa). AU: Malaysia (Sabah).
Clusiodes iotoides Sasakawa in Sasakawa & Pong 1990: 55. Malaysia. Sabah: Sepilok Forest Reserve. HT ♂ FRC. Sasakawa & Pong 1990: 55 [illustrations of male genitalia].
Hendelia iotoides. Lonsdale & Marshall 2007b: 147 [generic combination].
kinetrolicros (Caloren & Marshall). NT: Costa Rica, Mexico (Chiapas, Veracruz).
Clusiodes kinetrolicros Caloren & Marshall 1998: 295. Costa Rica. Guanacaste Province: Guanacaste Conservation Area, Cacao Field Station, 900 m. HT ♂ CNC. Caloren & Marshall 1998: 295 [illustrations of head, wing and male and female genitalia].
Hendelia kinetrolicros. Lonsdale & Marshall 2007b: 147 [generic combination], 2010: 1043 [Manual of Central American Diptera], 2011: 6 [photo, reproduction of genitalic figures].
laosica Sasakawa. OR: Laos.
Hendelia laosica Sasakawa 2011b: 16. Laos. Vientian: Gi Sion Village, de Tha Ngone. HT ♂, BPBM [Type No. BPBM 17353]. Sasakawa 2011b: 16 [illustrations of male genitalia].
lateralis (Sasakawa). OR: Philippines.
Prohendelia lateralis Sasakawa 1974: 153. Philippines. Palawan: Pinigisan, 600 m. HT ♂ UZMC. Sasakawa 1974: 153 [illustrations of head and male genitalia], 1977: 239 [catalogue, Orient].
Hendelia (Prohendelia) lateralis. Balthazar 1990: 417 [catalogue, Philippines].
Hendelia lateralis. Lonsdale & Marshall 2007b: 147 [genus redefinition].
latifrons (Frey). OR: Philippines.
Xenoclusia latifrons Frey 1960: 21. Philippines. Mindanao: Suriago. HT ♀ UZMH. Frey 1960: 21 [catalogue, world]; Delfinado & Hardy 1971: 29 [catalogue Philippine types]; Balthazar 1990: 417 [catalogue, Philippines]. Hendelia latifrons. Lonsdale & Marshall 2007b: 147 [generic combination, illustration of head].
marginalis (Sasakawa). AU: Papua New Guinea (Bougainville Island), Indonesia (Biak Island). Clusiodes (Clusiodes) marginalis Sasakawa 1987: 106. Papua New Guinea. Bougainville Island: Kokure, 690 m. HT ♂ BPBM [Type No. 13582]. Sasakawa 1987: 106 [illustration of male genitalia]. Hendelia marginalis. Lonsdale & Marshall 2007b: 147 [generic combination].
masneri Lonsdale & Marshall. NT: Costa Rica.
Hendelia masneri Lonsdale & Marshall 2011: 8. Costa Rica. Puntarenas Province: Monteverde Research Station, 1550 m. HT ♂ INBio. Lonsdale & Marshall 2011: 8 [illustrations of male genitalia].
megaspilos (McAlpine). AU: Australia (New South Wales, Queensland).
Clusiodes megaspilos McAlpine 1960: 83. Australia. New South Wales: Blue Mountains, Wentworth Falls. HT ♂ AMS.
Sasakawa 1987: 102 [key, Orient/Pacific].
Hendelia megaspilos. Lonsdale & Marshall 2007b: 147 [generic combination].
megastylotis (Sasakawa). AU: Indonesia (West Papua), Papua New Guinea.
Clusiodes (Clusiodes) megastylotis Sasakawa 1987: 107. Indonesia. West Papua: Manokwari, Vogelkop, 75 m. HT ♂ BPBM [Type No. 13583]. Sasakawa 1987: 107 [illustrations of male genitalia]. Hendelia megastylotis. Lonsdale & Marshall 2007b: 147 [generic combination].
melanospilus (Sasakawa). AU: Papua New Guinea (Bismarck Isl., Morobe).
Clusiodes (Clusiodes) melanospilus Sasakawa 1987: 109. Papua New Guinea. NE New Guinea: Morobe District, Wau, 1200 m. HT ♂ BPBM [Type No. 13584]. Pitkin & Evenhuis 1989: 535 [catalogue, Australasia/Pacific]. Hendelia melanospilos. Misspelling of melanospilus. Lonsdale & Marshall 2007b: 147 [generic combination].
mirabilis (Frey). NT: Bolivia, Costa Rica, Ecuador, Peru.
Labomyia mirabilis Frey 1928a: 16. Peru. Eastern Andes, Roque. LT 1♂ UZMH [accidental designation of lectotype by assumption of holotype (IZCN (1999) Art. 74.6) by Caloren & Marshall (1998); 2♂ ST originally described]. Frey 1960: 26 [illustration of head, catalogue, world]
Clusiodes mirabilis. Caloren & Marshall 1998: 297 [generic combination, redescription, new record (Peru), illustrations of head, wing and male and female genitalia].
Hendelia mirabilis. Lonsdale & Marshall 2007b: 147 [generic combination, habitus photos], 2010: 1043 [Manual of Central American Diptera], 2011: 9 [description of variation, photos, illustrations of male genitalia, new records (Bolivia, Costa Rica, Ecuador)].
napo (Caloren & Marshall). NT: Ecuador, Venezuela.
Clusiodes napo Caloren & Marshall 1998: 305. Ecuador. Napo Province: Baeza, 2000 m. HT ♂ DEBU. Caloren & Marshall 1998: 305 [illustrations of wing and male and female genitalia].
Hendelia napo. Lonsdale & Marshall 2007b: 147 [generic combination], 2011: 12 [reproduction of genitalic illustrations, new records (Venezuela)].
nigriceps (McAlpine). AU: Australia (New South Wales).
Parahendelia nigriceps McAlpine 1960: 85. Australia. New South Wales: Blue Mountains, Mt. Wilson. HT ♂ AMS. McAlpine 1960: 85 [illustration of head], 1976: 75 [discussion of spiral vibrissae]; Pitkin & Evenhuis 1989: 536 [catalogue, Australasia/Pacific].
Clusiodes nigriceps. Marshall 2000: 3 [generic combination, male agonistic behaviour, photos]. Hendelia nigriceps. Lonsdale & Marshall 2007b: 147 [generic combination].
nigrifrons (Frey). OR: Burma.
Clusiodes (Steleoclusiodes) nigrifrons Frey 1960: 18. Burma. NE Burma: Kambaiti, 2000 m. ST 7♂ 3♀ UZMH. Frey 1960: 18 [catalogue, world]; Sasakawa 1977: 237 [catalogue, Orient]; Papp et al. 2006: 190 [Thailand]. Hendelia nigrifrons. Lonsdale & Marshall 2007b: 147 [generic combination].
nigripalpus Lonsdale & Marshall. NT: Bolivia.
Hendelia nigripalpus Lonsdale & Marshall 2011: 12. Bolivia. La Paz Department: Caranavi, ca. 10 km NW, road to ENTEL tower, 1400 m, 15°46′35ʺS, 67°35′48ʺW. HT ♂ ANCB. Lonsdale & Marshall 2011: 12 [illustrations of male genitalia].
notata (Sasakawa). OR: Malaysia (Sarawak).
Clusiodes (Clusiodes) notatus Sasakawa 1987: 111. Malaysia. Sarawak: Kampong, Sadong, Tapuh, 300–450 m. HT ♂ [Type No. 13585], depository unknown [originally deposited in BPBM].
Hendelia notata. Lonsdale & Marshall 2007b: 147 [generic combination]; Sasakawa 2011b: 18 [Malaysia, illustration of male genitalia].
obscuripennis (Frey). OR: Burma, Thailand.
Clusiodes (Microclusiaria) obscuripennis Frey 1960: 19. Burma. Tenasserim, Sukli, 75 km E of Moulmein, 600 m. HT ♀, depository unknown [originally deposited at UZMH]. Frey 1960: 19 [catalogue, world]; Sasakawa 1977: 237 [catalogue, Orient]; Papp et al. 2006: 190 [Thailand].
Clusiodes (Clusiaria) obscuripennis. Sasakawa 1987: 103 [key, Orient/Pacific].
Hendelia obscuripennis. Lonsdale & Marshall 2007b: 147 [generic combination].
orientalis (Frey). OR: Burma.
Hendelia (Prohendelia) orientalis Frey 1960: 14. Burma. NE Burma: Kambaiti, 2000 m. ST 1♂ 2♀ UZMH. Frey 1960: 14 [catalogue, world].
Prohendelia orientalis. Soós 1963a: 182 [key, illustrations of head]; Sasakawa 1977: 239 [catalogue, Orient]. Hendelia orientalis. Lonsdale & Marshall 2007b: 147 [generic combination].
orientalis form conjuncta Frey. OR: Burma.
Hendelia (Prohendelia) orientalis form conjuncta Frey 1960: 14. Burma. Kambaiti. ST 1♂ 2♀ UZMH.
orientalis form obscurior Frey. OR: Burma.
Hendelia (Prohendelia) orientalis form obscurior Frey 1960: 14. Burma. Kambaiti. ST 1♀ UZMH.
palliceps Frey. OR: Burma.
Hendelia (Burmanochorea) palliceps Frey 1960: 14. Burma. NE Burma: Kambaiti, 2000 m. ST 5♂ 2♀ UZMH. Frey 1960: 14 [catalogue, world]; Sasakawa 1977: 238 [catalogue, Orient].
Hendelia palliceps. Lonsdale & Marshall 2007b: 147 [list].
personata Frey. OR: Burma.
Hendelia (Burmanochorea) personata Frey 1960: 15. Burma. NE Burma: Kambaiti, 2000 m. HT ♂ UZMH. Frey 1960: 15 [catalogue, world]; Sasakawa 1977: 238 [catalogue, Orient].
Hendelia personata. Lonsdale & Marshall 2007b: 147 [list].
phrenzinus (Caloren & Marshall). NT: Costa Rica, Venezuela.
Clusiodes phrenzinus Caloren & Marshall 1998: 307. Venezuela. Aragua State: Parque Nacional Henry Pittier, Highway Maracay-Choroni, km 19, 1330 m. HT ♂ DEBU. Caloren & Marshall 1998: 307 [illustrations of wing and male and female genitalia].
Hendelia phrenzinae. Lonsdale & Marshall 2007b: 147 [generic combination, unjustified emendation]. Hendelia phrenzinus. Lonsdale & Marshall 2011: 13 [reproduction of genitalic figures, new record (Costa Rica)].
philippinensis Sasakawa. OR: Philippines.
Hendelia philippinensis Sasakawa 2011b: 18. Philippines. Luzon: Nueva Vizcaya, Dalton Pass, 915 m. HT ♂, BPBM [Type No. BPBM 17350]. Sasakawa 2011b: 18 [illustrations of male genitalia].
plumipes (Sasakawa). OR: Japan (Ryukyus), Malaysia (Sabah).
Clusiodes (Clusiodes) plumipes Sasakawa 1987: 112. Malaysia. Sabah: Ranau, 500 m. HT ♀ BPBM [Type No. 13586]. Hendelia plumipes. Lonsdale & Marshall 2007b: 147 [generic combination]; Sasakawa 2011b: 20 [Oriental revision]; Sueyoshi 2014: 619 [catalogue, Japan].
Clusiodes usikumuri Sueyoshi 2006: 11. Japan. Ryuku: Irimote Island, Ushikumori, Taketomi Town. HT ♀ BLKU. Sueyoshi 2006: 11 [illustrations of head and female genitalia, photo of wing]; Sasakawa 2011b: 20 [as synonym of plumipes].
Hendelia usikumuri. Lonsdale & Marshall 2007b: 147 [generic combination], 2011: 7 [Japan, pictoral key].
plumosa (Sasakawa). OR: Japan (Ryukyus). PA: Japan (widespread).
Clusiodes (Clusiodes) plumosus Sasakawa 1964: 14. Japan. Honshu: Ashiu, Kyoto. HT ♀ OMNH. Sasakawa 1964: 14 [lateral illustration], 1990: 540 [description of male]; Soós 1984: 12 [catalogue, Palaearctic]; Matsumoto & Sasakawa 2006: 19 [notes holotype deposition in OKAKA].
Clusiodes plumosus. Sueyoshi 2006: 9 [illustrations of head and male genitalia, photo of wing]. Hendelia plumosa. Lonsdale & Marshall 2007b: 147 [generic combination, map]; Sueyoshi et al. 2009: 101 [log emergence trap, Japan]; Sueyoshi 2011: 8 [Japan, pictoral key], 2012: 28 [Japan], 2014: 619 [catalogue, Japan].
pterygion (Caloren & Marshall). NT: Southern Brazil.
Clusiodes pterygion Caloren & Marshall 1998: 299. Brazil. Rio de Janeiro: Rio de Janeiro. HT ♂ AMNH. Caloren & Marshall 1998: 299 [illustrations of male genitalia and wing].
Hendelia pterygion. Lonsdale & Marshall 2007b: 147 [generic combination], 2011: 13 [reproduction of genitalic figures].
punctifrons (Frey). OR: Burma.
Clusiodes (Clusiodes) punctifrons Frey 1960: 18. Burma. NE Burma: Kambaiti, 2000 m. ST 8♂ 7♀ UZMH. Frey 1960: 18 [catalogue, world]; Sasakawa 1977: 237 [catalogue, Orient], 1987: 102 [key, Orient/Pacific]. Hendelia punctifrons. Lonsdale & Marshall 2007b: 147 [generic combination].
putealis Lonsdale & Marshall. NT: Costa Rica.
Hendelia putealis Lonsdale & Marshall 2011: 15. Costa Rica. Cartago Province: Cerro de la Muerte La Cangreja, 1950 m. HT ♀ INBio. Lonsdale & Marshall 2011: 15 [illustration of female genitalia, photo].
quatuorsetosa (Mamaev). PA: Russia (Far East).
Clusiodes quatuorsetosa Mamaev 1974: 114. Russia. Far East: Ussury Krai. HT ♂ [with puparium] MNHA [not in ZIN, as stated in original publication, examined by author]. Mamaev 1974: 114 [illustration of surstylus]; Yagi & Sawada 2000: 41 [Mamaev collection holdings].
Hendelia quatuorsetosa. Lonsdale & Marshall 2007b: 147 [generic combination, map].
salebrosa Lonsdale & Marshall. NT: Bolivia.
Hendelia salebrosa Lonsdale & Marshall 2011: 15. Bolivia. La Paz Department: Zongo, 16°06′32ʺS, 68°04′30ʺW. HT ♂ ANCB. Lonsdale & Marshall 2011: 15 [illustrations of male and female genitalia].
saopaulo (Caloren & Marshall). NT: Brazil (São Paulo, Rio de Janeiro).
Clusiodes saopaulo Caloren & Marshall 1998: 303. Brazil. São Paulo: São Paulo, Jaragua. HT ♂ MNRJ. Caloren & Marshall 1998: 303 [illustrations of wing and male and female genitalia].
Hendelia saopaulo. Lonsdale & Marshall 2007b: 147 [generic combination], 2011: 16 [reproduction of genitalic figures].
similis Lonsdale & Marshall. AU: Fiji.
Hendelia similis Lonsdale & Marshall 2008b: 8. Fiji. Viti Levu: Nukura Forest, 60– 130m. HT ♂ BPBM [Type No. 16838]. Lonsdale & Marshall 2008b: 8 [illustrations of egg and male and female genitalia, photo].
spilota Sasakawa. OR: Malaysia (Sabah).
Hendelia spilota Sasakawa 2011b: 20. Malaysia. Sabah: 19 km N of Kalabakan. HT ♀, BPBM [Type No. BPBM 17351]. Sasakawa 2011b: 20 [illustrations of male genitalia].
stimulator (Caloren & Marshall). NE: Mexico (Tamaulipas).
Clusiodes stimulator Caloren & Marshall 1998: 301. Mexico. Tamaulipas State: “ Mpla ”, Gomez Farias, Rancho de Cielo. HT ♂ TAMU. Caloren & Marshall 1998: 301 [illustrations of male genitalia and wing].
Hendelia stimulator. Lonsdale & Marshall 2007b: 147 [generic combination], 2011: 16 [reproduction of genitalic figures].
unica Mamaev. PA: Russia (Far East).
Hendelia (Burmanochorea) unica Mamaev 1974: 112. Primorsky Krai: Kedroveja [= Kedrovaya] Pad Nature Reserve. HT ♂ MNHA [not in ZIN, as stated in original publication, examined by author]. Mamaev 1987: 115 [Russian Far East]. Hendelia unica. Lonsdale & Marshall 2007b: 147 [list, map].
variabilis Frey. OR: Burma.
Hendelia (Burmanochorea) variabilis Frey 1960: 15. Burma. NE Burma: Kambaiti, 2000 m. ST 1♂ 3♀ UZMH. Frey 1960: 15 [catalogue, world]; Sasakawa 1977: 238 [catalogue, Orient].
Hendelia variabilis. Lonsdale & Marshall 2007b: 147 [list].
description: Published as part of Owen Lonsdale, 2017, World Catalogue of the Druid Flies (Diptera: Schizophora: Clusiidae), pp. 1-85 in Zootaxa 4333 (1) on pages 35-41, DOI: 10.11646/zootaxa.4333.1.1, http://zenodo.org/record/1010066