Lipseuma josianae Golovatch & Geoffroy & Mauriès 2006, n. sp.
Creators: name: Golovatch, Sergei I.
name: Geoffroy, Jean-Jacques
name: Mauriès, Jean-Paul
Publisher: name: Zenodo
Year: 2006
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Chordeumatida
Kashmireumatidae
Lipseuma
Lipseuma josianae

description: Lipseuma josianae n. sp.   
(Figs 5; 6)  
TYPE MATERIAL. — Holotype: China, Hubei Prov., Banqiao County, Cave Chuan Dong Zi, Nos. 482, 485, 494, 9.VIII.1999, leg. J. & B. Lips, ♂ (MNHN DA155). Paratypes: same locality, 1 ♂ (MNHN DA155); 1 ♂, 2 juvs (MNHN DA155); ♀ (MNHN DA155); 2 juvs (MNHN DA155).  
ETYMOLOGY. — To honour Josiane Lips.  
 DIAGNOSIS. — Differs from the only congener (L. bernardi n. gen., n. sp.) by the slightly larger size, as well as in certain details of leg and gonopod structure.   
DESCRIPTION  
Length 9.3-9.8 (♂) to 10.1 mm (♀); width of midbody segments, 0.75-0.8 (♂) to 0.9 mm (♀), maximum width of head, 0.95 mm (holotype). Vertical diameter of midbody segments, 0.75 (♂) to 0.88 mm (♀). Coloration entirely pallid.  
Head width >> collum> segment 2> 3> 4> 5> 6> 7> 8> segment 9 = 18; thereafter body very gradually tapering toward telson.  
Body with 28 segments (46 leg-pairs). Head sparsely setose, clypeolabral region slightly convex. Eyes completely reduced. Antennae very long and slender (Figs 5A; 6C), reaching beyond body segment 6 (♂) or 5 (♀) dorsally; antennomere 7 with a peculiar flattened seta dorsally in distal third; antennomere 8 with four unusually long and slender sensory cones. Gnathochilarium without promentum.  
 Collum usual, obcordate in shape, with rudimentary paraterga. Tegument smooth, modestly shining, non-areolated, translucid. Metatergal setation 3 + 3, typical, macrochaetae very long, pointed, positioned on minute knobs; exterior macrochaeta longest, c. 0.7 times as short as body width and nearly three times longer than median or interior one; stricture between pro- and metazona very shallow, inconspicuous (Fig. 5B, C).   
Paraterga very small dorsolateral shoulders, rather regularly rounded in dorsal view (Fig. 5C), slightly better developed on anterior body third.  
 CIX (15) ≈ 0.9; MIX (15) = 1.25; MA (15) ≈ 175°; PIX impossible to evaluate as paraterga too poorly developed.   
Axial suture distinct, usual; segment 7 inconspicuous, pleurite without ventral apophyses. Pleural crests evident. Limbus invisible.  
 Legs generally very long and slender, c. 1.8 times as long as midbody height. Legs 1 and 2 slightly reduced as usual, tarsi with usual ventral brushes, but without papillae; male coxa 2 perforated by gonopore orifice. Tarsal papillae absent. Male legs 3-5 unmodified. Male legs 6 (Fig. 5D) exceptionally incrassate, with a distoventral process both on coxa and prefemur; acropodite strongly clasper-like, tarsus with two bulges subventrally and a modified claw. Male legs 7 (Fig. 5E) slightly incrassate, with evidently enlarged coxa and prefemur; coxa with a pointed lobe ventrally. Male legs 10 (Fig. 6A) and 11 normal, both pairs with coxal glands. Claws very long, sabre-shaped, simple.   
Anterior gonopods (leg-pair 8) unusually simple, small; coxites with traces of median division basally on caudal face, completely fused on front face; distally subsecuriform telopodites laterocaudal, with a few setae distally, nearly as high as the unpaired, linguiform, distally pilose syncolpocoxite, but somewhat higher than the voluminous angiocoxal sacs with a ridge on top; sternum plate-like, with each lateral bulge on front face supporting a strong seta (Fig. 5 F-H).  
Posterior gonopods (leg-pair 9) also very simple (Fig. 6B), placed on a plate-like sternum; coxae contiguous medially, each crowned with a simple, long, slender, erect process ventrally; telopodite reduced to a single voluminous segment filled with glandular material.  
Female legs 2 normal. Vulva (Fig. 6D, E) very broad; operculum small, devoid of outgrowths.  
REMARKS  
In one of the male paratypes, leg 6 is normal, not modified, whereas the anterior gonopod syncolpocoxite shows a small median notch apically, but there are no lateral setae on the coxite. Some of these traits are probably anomalous, likely neotenic.
description: Published as part of Golovatch, Sergei I., Geoffroy, Jean-Jacques & Mauriès, Jean-Paul, 2006, Four new Chordeumatida (Diplopoda) from caves in China, pp. 75-92 in Zoosystema 28 (1) on pages 86-90, DOI: 10.5281/zenodo.5393502