Ischnosoma quadriguttatum subsp. quadriguttatum quadriguttatum (Champion 1923
Creators: name: Kocian, Matúš
name: Schülke, Michael
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Ischnosoma
Ischnosoma quadriguttatum
Ischnosoma quadriguttatum quadriguttatum (champion, 1923)

description: Ischnosoma quadriguttatum quadriguttatum (Champion, 1923)
Figs 28, 45.
NEPAL: Baglung: Dhaulagiri, Baglung Lekh, 10–15 km NW Baglung, 2350–2550 m, 28°18'36''N, 83°31'56''E, 10.–12.V.2004, leg. J. Schmidt, 10 exs. (cKle, cSch); Bhojpur: Valley NW Phedi, 1900 m, 25.V.1997, leg. W. Schawaller, 1 ex. (SMNS); Dailekh: Talpokhari S Dailekh, 1800 m, 29.V.1988, leg. W. Schawaller, 1 ex. (SMNS); Dhawalagi: Annapurna South Himal, SE of Narcheng, left riverside of Bele Khola, 2300 m, 28°30'40''N, 83°41'33''E, 25./ 26.V.2001, leg. G. Hirthe, 1 ex. (cHir); Dolakha: Rolwaling Himal, Lapchi Kang, range below Ting Sang La, 2300–2500 m, 1.IX.1999, leg. J. Schmidt, 1 ex. (NME); Dolpa–Mustang–Myagdi: right banks of Lethe Khola near Lethe, 2400 m, 5.–7.V.1995, leg. J. Martens & W. Schawaller, 1 ex. (SMNS); S slope, Dhaulagiri mountains, N Banduk, 2400–2600 m, 28°28'35''N, 83°35'05''E, 8.V.2009, leg. J. Schmidt, 8 exs. (NME); Dhaulagiri mountains, upper Marang Khola valley., 28°29'50''N, 83°27'37''E, 2500–2700 m, 16.V.2009, leg. J. Schmidt, 1 ex. (NME), 1 ex. (cSch); Dhaulagari Himal, S-slope, N Banduk village, 1900–2300 m, 28°27'22''N, 83°35'13''E to 28°28'07''N, 83°35'10''E, 6.V.2009, leg. J. Schmidt, 4 exs. (NME), 1 ex. (cSch); SW Dhaulagiri Himal, Dhara Khola Vali, 28°30'36''N, 83°18'16''E, 1900 m, 21.–22.V.2012, leg. J. Schmidt, 6 exs. (NME); S slope, Dhaulagari mountains, above Pathlekharka, 2500–2700 m, 28°32'19''N, 83°29'25''E, 12.V.2009, leg. J. Schmidt, 2 exs. (NME); SW Dhaulagiri, Maraini, 2400–2800 m, 11.V.2012, 28°31'07''N, 83°15'49''E, leg. J. Schmidt, 2 exs. (NME); Gorkha: Darondi Khola, above Barpak, 3300– 3000 m, 11.VIII.1983, rhododendron forest, leg. J. Martens & W. Schawaller [243], 1 ex. (SMNS); Darondi Khola, below Barpak to Doreni, 1100– 900 m, 12.VIII.1983, forest remnants, leg. J. Martens & W. Schawaller [246], 1 ex. (SMNS); Buri Gandaki, Nyak to lower Chuling Khola valley, 2450–2870 m, 2.VIII.1983, Pinus excelsa, mountain pasture, leg. J. Martens & W. Schawaller [227], 1 ex. (SMNS); same data, but 2270–2450 m, 1 ex. (SMNS); Ilam: road Ilam to Birtamode, Karphok, 1700 m, bushes, agricultural area, 2.IV.1980, leg. J. Martens & A. Ausobsky, 1 ex. (SMNS); Mai Pokhari, 2100–2200 m, 9.– 10.IV.1988, Castanopsis forest remnant, leg. J. Martens & W. Schawaller [319], 1 ex. (SMNS); Kaski: above Dhumpus, Berlese, 2100 m, 8.–10.V.1980, leg. J. Martens & A. Ausobsky, 1 ex. (SMNS); above Dhumpus, Laubwald, 2100 m, 8.–10.V.1980, leg. J. Martens & A. Ausobsky, 1 ex. (SMNS); above Pothana, 2000 m, 27.– 29.IV.1995, leg. J. Martens & W. Schawaller, 1 ex. (SMNS); Pokhara, souther bank of Phewa Lake, creek, sifted, 800–900 m, 8.V.2001, leg. G. Hirthe, 2 exs. (cHir, cSch); Annapurna region, Khare to Pothana, rhododendron forest, 1950–2000 m, 10. V. 2001, leg. G. Hirthe, 1 ex. (cHir); Annapurna mountains, Tadapani to Chuile, 2450– 2550 m, 4.V.1999, leg. C. Krüger & G. Hirthe, 1 ex. (cHir); Lalitpur: Mount Phulchoki, 1800–2000 m, 25.IV.1995, leg. J. Martens & W. Schawaller, 1 ex. (SMNS); above Godawari, Kunako Khola, sifted, 1750 m, 6.V.2001, leg. G. Hirthe, 2 exs. (cHir, cSch); Lamjung: Nawronkot near Larjung, Thakhola area, 2900–3000 m, leg. H. Franz, 1 ex. (NHMW); Manang: Annapurna mountains, Bagarchhap–Temang, 2400–2700 m, 17.IV.1999, leg. C. Krüger & G. Hirthe, 1 ex. (cHir); Panchthar: Paniporua, 2300 m, 16.–20.IV.1988, mixed broadleaved forest, leg. J. Martens & W. Schawaller [328], 1 ex. (SMNS); Parbat: Annapurna mountains, Bhickok to Deurali, 2100 m, 7.V.1999, leg. C. Krüger & G. Hirthe, 3 exs. (cHir, cSch); Annapurna region, West Mardi Himal, above Deurali, 2100–2200 m, 28°18'57''N, 83°49'59''E, 10. V. 2001, leg. G. Hirthe, 4 exs. (cHir, cSch); Rasuwa: Bagmati province, Gul Ghanjyang, 2600 m, 6.IV.1981, leg. I. Löbl & A. Smetana, 1 ex. (MHNG); Sankhuwasabha: above Pahakhola, 2600–2800 m, 31.V.–3.VI.1988, Quercus semecarpifolia, Rhododendron, leg. J. Martens & W. Schawaller [404], 3 ex. (SMNS); Arun Valley, between Mure and Hurure, 2050–2150 m, 9.–17.VI.1988, mixed broadleaved forest, leg. J. Martens & W. Schawaller [412], 2 ex. (SMNS); N Furure, floodplain near creek, 27°30'44''N, 87°16'14''E, 2100 m, 7.XII.1998, sifted, leg. M. Hartmann, 1 ex. (NME); Arun Valley, W above Tashigaon, 2500–2700 m, 26.– 27.V.2014, 27°36'33''N, 87°13'28''E, leg. J. Schmidt, 1 ex. (NME); Arun Valley, near Tashigaon, 2200–2400 m, 12.+ 28.–29.V.2014, 27°37'N, 87°14'E, leg. J. Schmidt, 2 ex. (NME); Sindhupalchok: Barahbise region, Ting – Sang – La, leg. H. Franz, 1 ex. (NHMW); Solukhumbu: below Pangum, 2500 m, 14.–15.V.1997, leg. W. Schawaller, 3 exs. (SMNS); Umg. Sutje near Lughla, Khumbu, leg. H. Franz, 2 exs. (NHMW); Surkhet: Bheri Khola bridge, 500 m, 24.–25.V.1998, leg. W. Schawaller, 1 ex. (SMNS); N Surkhet, 1600–2000 m, 28.V.1998, leg. W. Schawaller, 1 ex. (SMNS); Syangja: Manaslu mountains, Dudh Pokhari Lekh, upper Dardi Khola valley, 2600– 2300 m, 15.–17.IV.2003, leg. J. Schmidt, 2 exs. (NME, cSch); Manaslu mountains, Dudh Pokhari Lekh, upper Phulinagiri Madi, 2500 m, 19.–21.IV.2003, leg. J. Schmidt, 10 exs. (NME, cSch); Manaslu mountains, Dudh Pokhari Lekh, below Helam Pokhari, 2000 m, 22.IV.2003, leg. J. Schmidt, 1 ex. (NME); Manaslu mountains, S of Bara Pokhari, 2300 m, 8.IV.2003, leg. J. Schmidt, 24 exs. (NME, cSch); Manaslu mountains, Bara Pokhari lekh, Chandi Khola valley, 11.–12.IV.2003, 2000–2300 m, leg. J. Schmidt, 1 ex, (NME); Manaslu mountains, S of Bara Pokhari, 2000 m, 7.IV.2003, leg. J. Schmidt, 1 ex. (NME); Bara Pokhari Lekh, 2500 m, 2.IV.1999, leg. G. Hirthe & C. Krüger, 2 exs. (cHir, cSch); Taplejung: confluence of Kabeli and Tada Khola, 1000 m, 23.–25.IV.1988, mixed broadleaved forest, leg. J. Martens & W. Schawaller [344], 5 exs. (SMNS); ascent to pasture Lassetham from Omje Khola, 2400–3150 m, 6.V.1988, mixed Quercus - Tsuga - Rhodonendron forest, leg. J. Martens & W. Schawaller [357], 1 ex. (SMNS).
Remarks. I. quadriguttatum quadriguttatum is widespread in Southeast Asia. In Nepal, it is the most common species and present everywhere (Fig. 45). A second subspecies, I. quadriguttatum japonicum Kocian, 2003 was described from Japan.
description: Published as part of Kocian, Matúš & Schülke, Michael, 2018, New species and records of Ischnosoma Stephens (Coleoptera: Staphylinidae: Tachyporinae) from Nepal, pp. 501-522 in Zootaxa 4442 (4) on pages 516-518, DOI: 10.11646/zootaxa.4442.4.1, http://zenodo.org/record/1304822