Prothyma (Parasymplecthyma) johnsoni Wiesner & Anichtchenko 2023, sp. nov.
Creators: name: Wiesner, Jürgen
name: Anichtchenko, Alexander
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Prothyma
Prothyma johnsoni

description: Prothyma (Parasymplecthyma) johnsoni sp. nov. (Fig. 18)
Type material. Holotype male: “ March 2021 / Mt Talomo / Davao del Sur / Mindanao, Philippines // “HOLOTYPUS Prothyma / (Parasymplecthyma) / johnsoni / design.Wiesner & /Anichtchenko 2022 [printed, red]” (JWc). Paratypes: 2 females: “ June 2019 / Gumitan, Davao / Mindanao / Philippines ” and “ April 2021 / Mt Talomo / Davao del Sur / Mindanao, Philippines ” (WJc); 1 male: “ Filippine Mindanao / Aguson [Agusan] del Norte / V.2012 ” (RSc); 1 female: “ Filippine Mindanao / South Cotobato / Arakan IV. 2014 ” (RSc); 1 female: “ Philippines Mindanao / Davao del Sur / Mt. Apo VI.2014 ” (RSc). All labelled with “PARATYPUS Prothyma / (Parasymplecthyma) / johnsoni design. Wiesner & / Anichtchenko 2022 [printed, red]”.
Etymology. This new species is cordially dedicated to Walter N. Johnson, Minneapolis, Minnesota, a specialist of Nearctic tiger beetles, who made three specimens available to us.
Diagnosis. Prothyma (Parasymplecthyma) johnsoni sp. nov. is unique because: 1, the inner sac (aedeagus) with a long, filiform flagellum forming a helical spiral with six coils; 2, the yellow labrum; 3, yellow legs and 4, basal four antennal segments yellow.
Description. Size: TL (without labrum) 11.2–13.9 mm (mean = 12.8 mm, n = 3).
Head: Color shining black; with greenish-blue or violet reflections; surface glabrous, with two setigerous punctures next to eye, one along frontal margin and one dorsally at center; strong longitudinal striae on orbital plates and vertex; strong transversal striae behind the eyes and on neck; striae becoming obsolete on genae, frons, and clypeus. Head a little smaller than the elytra (mean EW/HW = 1.1). Labrum yellow brownish. Labrum of male (Fig. 18f) wider than long (mean LW/LL = 1.4); with four setae located near the margin, two lateral and two apical; with two blunt anterolateral teeth and two outward directed anterior teeth at the anteromedian lobe, intermediate space slightly bulged twice. Labrum of female (Fig. 18g) as wide as long (mean LW/LL = 1.0), with four setae located near the margin, two lateral and two apical; with two blunt anterolateral teeth and two outward directed thorn-like anterior teeth at the anteromedian lobe, intermediate space with a large triangular tooth. Mandibles brownish, with four teeth, margins darker. Labial and maxillary palpi yellow brownish. Antennae slender, reaching posteriorly to just behind the shoulders; scape and antennomeres two to four yellow brownish; scape with one apical seta, the following three antennomeres almost glabrous; antennomeres five to eleven black, dull, finely, and evenly pubescent.
Thorax: Pronotum slightly longer than wide (mean PW/PL = 0.9); sides not rounded between the transversal constrictions, nearly parallel sided; glabrous, with transverse striae above; shiny black with blue greenish or violet reflections; sternae and episternae glabrous, shiny black with bluish reflections; Mesepisternum of female with a long, shallow depression in the center.
Elytra: (Figs 18a–c) slightly more than twice as long as wide (mean EL/EW = 2.1), Elytra with a basal hump, laterally from the end of the humeral spot to the beginning of the apical lunule gently and arcuately indented, enclosing the latero-marginal spot; color shining black, with violet or greenish-blue reflections; elytral testaceous maculation consists of humeral and lateromedial dots and apical lunule; the humeral spot is broad and extends from the lateral margin to the middle suture in the male; the latero-marginal spot is large and trapezoidal, oriented obliquely downwards; the apical lunule is broad and encompasses the entire elytral apex to the middle suture; surface with dense and evenly spaced punctures, becoming confluent and forming grooves near the suture; these grooves extends from the basal hump along the middle suture to the apex and is bounded toward the lateral margin by the edge of the arcuate indentation; apical margin distinctly rounded and slightly restricted towards middle suture, with a tiny sutural spine. Epipleurae yellow-brownish.
Ventral aspect: Venter glabrous, yellow-brownish; legs yellow-brownish with apex of segments slightly brownish; coxa and trochanter glabrous, femur, tibia, and tarsi sparsely setose, mesotibiae more densely bristled in apical half.
Aedeagus: (Figs 18d–e) in lateral view with a short slender base, from there strongly and cylindrically inflated, tapering in apical quarter to a point at upper lateral margin, taper not rectilinear but bulging. The tip is short, curved upwards, thickened and knob-like. The inner sac is half-filled by a flagellum in six helical coils; the flagellum encloses a sustaining membrane, with an apical chitinous plate, shield and piece are crowded together (total length 3.5 mm).
Distribution. MINDANAO - few localities near Mt. Apo.
description: Published as part of Wiesner, Jürgen & Anichtchenko, Alexander, 2023, The Philippines species of the genus Prothyma Hope, 1838 (Coleoptera: Cicindelidae) Part 1: Subgenera Prothyma Hope, 1838, Genoprothyma Rivalier, 1964 and Parasymplecthyma subgen. nov., pp. 1-36 in Zootaxa 5315 (1) on page 33, DOI: 10.11646/zootaxa.5315.1.1, http://zenodo.org/record/8129911