Clytus nigritulus Kraatz 1879
Creators: name: Danilevsky, M. L.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Cerambycidae
Clytus
Clytus nigritulus

description: Clytus (s. str.) nigritulus Kraatz, 1879   
Figs 7–19  
 Clytus nigritulus Kraatz, 1879: 109 [“Ost-Sibirien”]; Pic, 1900a: 9 [Sibérie]; 1900b: 60 [Sibérie]; Aurivillius, 1912: 371, part. [Amur]; Lee, 1987: 113 [Korea, Siberia, Northern China]; Hua, 2002: 202, part. [China: Heilongjiang, Jilin; Siberia]; Danilevsky & Smetana, 2010: 171 (= fulvohirsutus Pic) [Eastern Siberia, Far East Russia, Korean Peninsula, North-eastern China]; Lim et al., 2012: 195 [Korea]; Hwang, 2015: 219 [Korea]; Jang et al., 2015: 236 [Korea]; Chen et al., 2019: 151 (= fulvohirsutus Pic) [North-eastern China, Korean Peninsula, Russia: Siberia]; Danilevsky, 2020: 234 (= fulvohirsutus Pic) [Eastern Siberia, Far East Russia, Korean Peninsula, North-eastern China]; Anisimov & Bezborodov, 2020: 150 [Amur region].   
 Clytus fulvohirsutus Pic, 1904a: 18 [Sibérie]; 1904b: 15 [sur les bords du fleuve Amour]; Aurivillius, 1912: 370, part. [Sibirien]; Plavilstshikov, 1932: 191 [Eastern Siberia]; 1940: 394, 417, 729 [Eastern Siberia from Raddevka on Amur to Pacific Ocean and probably in neighbor regions of Korea and Manchuria]; Gressitt, 1951: 252, 255 [SE Siberia, NE China]; Heyrovský, 1974: 34 [“Sibirien (Amur- und Ussuri-Gebiet)] Korea ”]; Tsherepanov & Tsherepanova, 1975 [Ussuri Region]; Lobanov et al., 1982: 261 [Far Eastern Russia, Korea, Northern China]; Lee, 1982: 40 [Korea]; Tsherepanov, 1982: 101 [low Amur valley, Ussuri Region]; 1996: 107 [Amur Region, Khabarovsk Region, Primorye Region]; Sundukov, 1998: 40 [Lazo Natural Reserve]; Hua, 2002: 202, part. [China: Heilongjiang, Jilin; Siberia]; Niisato & Koh, 2003: 297 (? = nigritulus Kr.) [SE Siberia, Ussuri-Primorye and Amur, NE China, Korean Peninsula]; Danilevsky, 2021: 333 [from Raddevka on Amur River (now Radde in Jewish Region of Russia, 48°36'06" N, 130°35'42" E) to Pacific Ocean, and probably penetrates to China and Korea].   
 Clytus (Clytus) nigritulus : Smirnov, 2009: 186 (= fulvohirsutus Pic) [Lazo Natural Reserve; Amur Region, Primorye Region, Korea, North-eastern China].   
 Clytus (Paulomaculatus) fulvohirsutus : Özdikmen, 2023: 54, part. [Russia: Siberia].   
 Clytus (Paulomaculatus) nigritulus : Özdikmen, 2023: 54, part. [Russia; Siberia].   
TYPE LOCALITY: Russia, Eastern Siberia (most probably Amur Region).  
 MATERIAL EXAMINED. Author’s collection: 1 ♂, Russia, Primorye Reg., Vladivostok, Akademgorodok, 8.6.1970, Crataegus, Azarova; 1 ♀ (Fig. 18), Russia, Primorye Reg., Kamenushka River, 30.7.1971, A. I. Tsherepanov; 1 ♀, Russia, Jewish Region, Pashkovo, 9.6.1978, S. Murzin. 1 ♀ (Fig. 19), Russia, Primorye Reg., Barabash, 24.3.1982, ex larva, Alnus, S. Murzin; 1 ♂, Russia, Primorye Reg., Lazo Distr., Kievka, 18.6.1991, Filimonov; 1 ♀, Russia, Primorye Reg., Ussuriysk Distr., Gorno-Tayozhnaya Station, 31.5.1991, Filimonov; 1 ♀, Russia, Amur Region, Blagoveshchensk, Chigiri, 12.6.2021, M. Anisimova. Collection of Zoological Museum of Moscow University (all specimens under the bottom label “ Clytus fulvohirsutus ”): 1 ♂, “Ussuri / Ossinovka / 28. V.917 / P. Elsky ”; 1 ♂, “Ussuri / Ossinovka / 16. VI.917 / P. Elsky ”; 1 ♂, Primorye Region, Piankov Zavod, Nikolsk-Ussuriysk, 2.6.1925, T. Samoylov; 1 ♀, “Sibiria or. / Sichote Alin mer / Romanovka / 15. VI.930”; 1 ♀, “Ussuri / Ozernaja / 22. VI.34 / S. Sevastjanov ”; 1 ♂, “Mandzhuria / prov. Girin / Hsialoling / 3.VIII.937 / ex coll. Alin ”; 1 ♂, 3 ♀, “Mantshzhuria / st. Kaolingtzu / prov. Girin / 10.VII.940 V. Alin”; 1 ♀, Sikhote-Alin, Sibichi, 16.6.1951; 1 ♂, Primorye Region, Sibichi, 13.6.1951; 3 ♂, 4 ♀, each with a single date label from 15. V. to 7.VII. Collection of S.M. Murzin (Moscow): 1 ♀, Russia, Jewish Region, Dichun, ex larva, 6.1.1978; 1 ♀, Russia, Primorye, Kamenushka, 4.6.1979, S. Murzin; 1 ♀, Russia, Jewish Region, Malyi Khingan Ridge, Dichun River, 21.6.1979, S. Murzin.   
 DIAGNOSIS. Body small; all femora usually totally black, antennae and tibiae often partly reddish; male antennae extend far beyond long oblique central elytral stripe, female antennae can be about same relative length or rather shorter; prothorax convex, with less rounded sides than in C. arietoides, never bordered with pale pubescence anteriorly, but with several white setae near hind angles; in males about as long as wide or slightly elongated, in females always transverse; regular small cells of honeycomb sculpture are distinct in females only; male pronotum consists of very dense fine grainy sculpture with obliterated or conjugated cells; pronotum covered with very dense and long erect setae, looks furry; scutellum with dense white recumbent pubescence; male elytra about 2.4-2.6 times longer, than humeral width, female elytra – about 2.2–2.5 times; each elytron with 2 dense narrow setae stripes (central and postmedian) and poorly developed disperse apical stripe, humeral stripes always totally absent; long oblique central stripe often extended along suture; elytral setae stripes usually white, rarely – yellowish; hind femora usually reach or even go beyond elytral apices; body length in males: 5–8 mm; in females: 6.8–9 mm.   
DISTRIBUTION. Russian Far East: Primorye Region, Southern Khabarovsk Region and Amur Region, Korea, North-eastern China.  
 TYPE MATERIAL. Clytus nigritulus Kraatz, 1879; ♂ (Fig. 7) lectotype (present designation) with 5 labels (Fig. 8): 1) Amur / Christoph 77, 2) [red] Syntypus, 3) coll. Kraatz, 4) Clytus / nigritulus / 1879 Kraatz, 5) SDEI Coleoptera / # 303529; ♂ (Fig. 9), paralectotype (present designation) with 4 labels (Fig. 10): 1) Amur / Christoph 77, 2) [red] Syntypus, 3) coll. Kraatz, 4) SDEI Coleoptera / # 303530; ♂ (Fig. 11), paralectotype (present designation) with 4 labels (Fig. 12): 1) Sibirien, 2) [red] Syntypus, 3) coll. Kraatz, 4) SDEI Coleoptera / # 303531. All three types are preserved in Senckenberg Deutsches Entomologisches Institut (SDEI), Müncheberg, Germany. Clytus fulvohirsutus Pic, 1904; ♂ (Fig. 13) holotype with 5 labels (Fig. 14): 1) Amour: Radde / (Korb), 2) Museum Paris / coll. M. Pic, 3) type / 4) fulvohirsutus / Pic, 5) [red] holotype.
description: Published as part of Danilevsky, M. L., 2023, SPECIES OF THE ARIETOIDES-GROUP OF THE GENUS CLYTUS LAICHARTING, 1784 (COLEOPTERA: CERAMBYCIDAE) FROM THE RUSSIAN FAR EAST, pp. 17-28 in Far Eastern Entomologist 490 on pages 21-23, DOI: 10.25221/fee.490.3, http://zenodo.org/record/10135152