Paraindopamphantus Malipatil & Scudder 2018, gen. nov.
Creators: name: Malipatil, M. B.
name: Scudder, G. G. E.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy

description: Genus Paraindopamphantus gen. nov.
Type-species: Paraindopamphantus bruneiensis sp. nov.
Description
Body strongly myrmecomorphic (Figs. 1 & 2).
Head: Evenly sloping downward from posterior margin (Fig. 4); bucculae produced near base as flaps (Fig. 5), then gradually diminishing posteriorly to almost to surface of head near base of antenna; head below with gular area broadly conspicuously swollen increasingly from apex to base on either side of narrow median labial groove (Figs. 4–6); eyes not reniform and placed on posteriorly rounded head well away from anterolateral corners of pronotum (e.g., Fig. 3). Labium with 3rd segment shortest. Labrum about as long as 1 st labial segment. Antenna with 1 st segment shortest, 2nd slightly longer than 4th (Fig. 4).
Thorax: Pronotum with broad deep median transverse impression separating anterior and posterior lobes with a long stalk at about half length, lateral margins almost rounded and non-carinate (Fig. 3). Thoracic pleura with sparse punctures (Fig. 7), metathoracic scent gland auricle conspicuously projecting above surface of pleuron with spout turning upwards; evaporative area large, dull (Fig. 7). Scutellum with transverse impression on basal ½, coarse deep punctures on sides and on posterior area, apex up curved and projecting above surface as a pointed cone (e.g., Fig. 7). Hemelytra extending to posterior margin of tergum VII (Figs. 1 & 8); corium with more or less three rows of thick-set brown punctures; clavus with two rows of coarse punctures, one complete row along claval suture, 2nd row complete on inside apical ½, inner margin impunctate (Fig. 8). Hind wing venation reduced. All femora almost similarly uniformly incrassate, unarmed (Fig. 1).
Abdomen: Vasiform (Fig. 10). First visible (i.e., II) and anterior part of second visible (i.e., III) segment strongly constricted to a parallel sided tube, and strengthened with longitudinal ridges on ventral surface (Fig. 9). Spiracles II dorsal on membrane as connexivum almost absent, III dorsal on connexivum (Fig. 11), IV, V, VI and VII ventral. Sutures between all tergal segments distinct, those between terga III-IV only slightly and those between IV-V and V-VI strongly curved caudad from margin to meson and with distinct scent gland scars which are subequal (the latter fractionally wider) in width, scar between III-IV indistinct (e.g., Fig. 10). Inner laterotergites greatly reduced. Sterna II, III and IV with intersegmental sutures faint, sutures reaching abdominal margin (Figs. 12–14). Trichobothria on sternum III in loose triangle and those on IV almost linear to open triangle (Fig. 14), variable in spacing; trichobothria on V, VI and VII typically lygaeoid, posterior to spiracle one above the other, spiracle of V and VI situated almost equidistant between anterior and posterior trichobothria.
In female, seventh sternum cleft with ovipositor.
Female genitalia —Ovipositor with first ramus traversing to almost 4/5 length of first gonapophysis (Fig. 16); second gonapophysis spatulate with sparse minute setae in two rows (Fig. 17).
Notes. Paraindopamphantus is the only second genus described from the Oriental Region, and differs from the first genus Indopamphantus in the characters in the key, as well as by having the pronotum with a long stalk that is about ½ as long as anterior and posterior lobes (Fig. 3) (short stalk, much less than ½ as long as anterior and posterior lobes in Indopamphantus), and the posterior lobe of the pronotum that is almost impunctate.
description: Published as part of Malipatil, M. B. & Scudder, G. G. E., 2018, ParaindOPaMPHantus bruneiensis gen. nov. et sp. nov., as the first representative of the subfamily Pamphantinae from South East Asia (Hemiptera: Heteroptera: Geocoridae: Pamphantinae: Indopamphantini) in Zootaxa 4415 (1), DOI: 10.11646/zootaxa.4415.1.3, http://zenodo.org/record/1241791