Tachyusa balteata Erichson, Austria-Helvetia, Schuppel (ZMHB 1839
Creators: name: Pa ṡnik, Grzegorz
Publisher: name: Zenodo
Year: 2006
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Tachyusa
Tachyusa balteata

description: Tachyusa balteata Erichson, 1839 (Figs. 26–28)   
 Tachyusa balteata Erichson, 1839: 71.   
 Tachyusa balteata elongata Kolenati, 1846: 5 (synonym confirmed).   
 Tachyusa balteata: Redtenbacher, 1849: 654; Redtenbacher, 1856: 123.   
 Tachyusa (Caliusa) balteata: Mulsant & Rey, 1875: 404; Ganglbauer, 1895: 248; Reitter, 1909: 74; Fenyes, 1920: 173; Bernhauer & Scheerpeltz, 1926: 586; Lohse, 1974: 69.   
 Type material. Tachyusa balteata Erichson. Lectotype (here designated): ♂: Hist.­Coll. (Coleoptera) Nr. 5330, Tachyusa balteata Erichson, Austria­Helvetia, Schüppel (ZMHB). Paralectotypes: 3♂♂ and ♀: same data as the lectotype (ZMHB).   
 Tachyusa balteata elongata Kolenati. Holotype: ♀: Georgia, Kolenati (NMW).   
 Additional material. Albania: Elbasan, Mader, 1 ex (NMW). Austria: Ehrenhansen, 2 exx (NMW); Feistritz, 31.VII.1910, 1 ex (MSNM); Goritschach, VI.1976, 3 exx (MHNG); Graz, Netolitzky, 1 ex (MTD); 5 exx (DEI); Graz, Krauss, 1 ex (MSNM); 4 exx (ZFMK); Graz, VI.1895, 2 exx (MSNM); Graz, Mur river, 1 ex (CAS); Graz, Weber, 1 ex (NMPC); Grunburg, Bernhauer, 6 exx (FMNH); Kampthal, 2 exx (NMW); Karawanken, Drautal Ferlach, 21.VII.1939, Linke, 1 ex (MTD); Karnten, Eisenkappel, VII.1922, 1 ex (ZMHB); Karnten, Eisenkappel, 4.VIII.1924, Bernhauer, 1 ex (FMNH); Karnten, Eisenkappel, VII–VIII.1926, O. Scheerpeltz, 13 exx (NMW); Karnten, Eisenkappel, VII– VIII.1931, O. Scheerpeltz, 27 exx (NMW); Karnten, Eisenkappel, Mader, 1 ex (NMW); Klagenfurt, 1 ex (MSNM); Leithagebirge, 2 exx (NMW); Leitha river near Ebenfurth, 31.V.1934, P. Meyer, 1 ex (NMW); Lichtenwald, Wradatsch, 1 ex (MZLU); Sachsenburg, 1 ex (NMPC); Scheiblingberg, Linke, 1 ex, (MTD); Steyr, Petz, 2 exx (NMW); Tirol, Meran, Lokay, 1 ex (MTD); Vorarlberg, Bregenz, A.J. Muller, 10 exx (NMPC); Waidhofen an der Ybbs, Bernhauer, 1 ex (FMNH); Wien, Breit, 1 ex (DEI); 4 exx (ZMHB); 1 ex (NMW); Wolfsberg environs, Lavant river, 1 ex (ZMHB); 4 exx (DEI). Belgium: Comblain­au­Pont (Rivage), 21.VII.1938, Fagel, 1 ex (IRSNB); Gent, Escher, 2 exx (ETH); Vieuxville (Sy), Ourthe river, 25.IV.1952, G. Fagel, 1 ex (IRSNB). Bosnia and Herzegovina: “Central­Bosnien”, Reitter, 2 exx (HNHM); Ilidža, Linke, 15 exx (MTD); Ilidža, Apfelbeck, 3 ex (FMNH); Jablonica, 14.VI.1936, Linke, 6 exx (MTD); Prozor, 1902, O. Leonhard, 1 ex (NMW); Sarajevo, Linke, 1 ex (MTD). Bulgaria: Kazanlik, Netolitzky, 4 exx (NMW); 2 exx (MTD); Pirin, 2 km N Gara, 11–12.VI.1983, Hieke, 12 exx (ZMHB); Rumelia: Sliven, Tundze, 28.VI.1908, Rambousek, 1 ex (DEI); 1 ex (NMPC); Rupite near General Todorow, 4.V.1984, Hieke, 3 exx (ZMHB); Rupite near General Todorow, 23–28.IV.1984, Hieke, 1 ex (ZMHB); Sandanski, 6–11.V.1984, Jaeger, 1 ex (ZMHB); Sejmen Marica, 21.V.1909, Rambousek, 1 ex (NMPC); Simitli, Strum river, 25.VI.1979, M. Uhlig, 15 exx (ZMHB); Sofia, 19.III.1909, Rambousek, 2 exx (NMPC); Strumjani, 30.IV.1985, B. Jaeger, 2 exx (ZMHB); Strumjani, 1985, Wrase, 4 exx (ZMHB). Czech Republic: Brandeis [Brandys], Skalitzky, 1 ex (MNHN); Brandeis [Brandys], Skalitzky, 1 ex (NMW); Choceň, 10.V.1904, 2 exx (NMPC); Chrnolein, 8.V.1905, 1 ex (NMPC); Dubna, 9.VII.1909, Klička, 1 ex (NMPC); Modřany, 1.VIII.1918, 2 exx (NMPC); Pisek, Dr Skalizky, 2 exx (NMPC); Praha, 1 ex (NMW); Praha, 16.I.1920, Klička, 2 exx (NMPC); Praha, 1.VII.1912, 2 exx (NMPC); Praha Bubny, 17.VII.1907, 2 exx (NMPC); Radotin, 10.V.1909, 1 ex (NMPC); Št chovice, Zeman, 4 exx (NMPC); Toušeň, Heyrovsky, 1 ex (NMPC); Vrané, 21.V.1911, Heyrovsky, 2 exx (NMPC); Wilken, Hildesheim, 1 ex (NMPC); Žamberk, 1 ex (MNHN). France: Aigues Mortes, IV.1946, Levasseur, 1 ex (MNHN); Aimargues, E Tisson, 1 ex (NMPC); Albertville, Savoy, 2 exx (BMNH); Allier, 4 exx (MNHN); Argelès, 15 exx (MNHN); Ariège, Moulis, IX.1962, 2 exx (MNHN); Ariège, Massiag, VII.1959, 1 ex (MNHN); Ariège, Nalzen, 1.VIII.1951, Coiffait, 3 exx (MNHN); Arles, Amélie, Tech river, 25.III.1931, Levasseur, 1 ex (MNHN); Audes, 19.VII.1972, 7 exx (MHNG); Aure, 5 exx (MNHN); Avignon, Rhône, 24.IX.1919, 3 exx (MNHN); Bec. d'Allia (Cher), Deville, 2 exx (MNHN); Belmont, Savoie, Poussielgue, 1 ex (NMW); Briançon, 1 ex (MNHN); Briare, Loiret, Fongond, 10 exx (MNHN); Briare Loiret, III.1941, 1 ex (MNHN); Brignais, Rhône, G. Audras, 8 exx (MHNL); Brivé, V.1944, Coiffait, 1 ex (MNHN); Carcassonne, 4 exx (MNHN); Chabanais, VII.1940, 5 exx (MNHN); Choranche, Isère, 2 exx (MHNL); Cogolin, Var, 12.VI.1953, 2 exx (MHNG); Dourbies, Milhau­Aveyr, 1 ex (MHNG); Écully, Rhône, G. Audras, 1 ex (MHNL); Elne, 28.VII.1951, Coiffait, 3 exx (MNHN); Embrun, 2 exx (MNHN); Feurs, Rhône, 1 ex (MNHN); Fréjus, Var, G. Audras, 8 exx (MHNL); Gagnieres / St. Ambroix Cevennen, 27–28.V.1972, Schawaller, 4 exx (SMNS); Georges du Loup (Alpes­Maritimes), 2 exx (MNHN); Gevrieux, G. Audras, 1 ex (MHNL); Hautes Alpes, Buch near Aspres, 22.VI.1976, 3 exx (MHNG); Hautes Prov., Annot 900 m., Vaire, VI.1988, P. Wunderle, 5 exx (VAPC); Isère, Sassenage, 7.VI.1903, Agnus, 8 exx (MNHN); Isère, Grenoble, Poussielgue, 2 exx (NMW); La Mole, Var, Deville, 4 exx (MNHN); Labastille de Seine, 3.VI.1924, 2 exx (MNHN); Lent, IX.1968, G. Audras, 1 ex (MHNL); Les Allies, 20.III.1903, 2 exx (MNHN); Lyon, II.1950, G. Audras, 1 ex (MHNL); Millas, G. C. Champion, 1 ex (BMNH); Moulins, Baudelin, 1 ex (MHNL); Ossas, 8.VIII.1945, Tempere, 1 ex (MHNG); Pau, VI.1962, Coiffait, 6 exx (MNHN); Pau, VI.1859, H. Clark, 6 exx (BMNH); Pau, 5.VI.1942, Coiffait, 4 exx (MNHN); Pont du Gard, 19.IV.1925, 6 exx (MNHN); Pont du Gard, 9.IX.1923, 4 exx (MNHN); Pont du Gard, IX.1923, Levasseur, 2 exx (MNHN); Pont du Gard, IV.1946, Levasseur, 5 exx (MNHN); Pont du Gard, V.1987, Rougemont, 4 exx (GRPC); Pyrénées, Assas, VI.1946, Coiffait, 8 exx (MNHN); Saint Florent, Cher, Deville, 4 exx (MNHN); Saint Germain, 1 ex (MNHN); Saint Loup, 5 exx (MNHN); Saint Martin, G. Audras, 5 exx (MHNL); Saint Rambert, Ain, 7 exx (MNHN); Strasbourg, V.1953, 1 ex (MHNG); Strasbourg, V.1954, 4 exx (MNHN); Tardets, VI.1939, Coiffait, 16 exx (MNHN); Tech near Elne, 5.VII.1976, 1 ex (MHNG); Toulouse, III.1954, Coiffait, 1 ex (MNHN); Ussat­Ariege, Pyrénée, IX.1926, V. Jedlicka, 3 exx (NMPC); Villefranche de Conte, 10.VI.1948, 3 exx (MNHN); Walbourg, Rhône, 10.VI.1952, Coiffait, 8 exx (MNHN); “ Corse ” (localities not specified), 2 exx (MNHN); 2 ex (CAS). Germany: Bayern, Farchant, VII.1925, Langer, 1 ex (ZMHB); Bayern, Loisach river, 30.VI.1932, Ihssen, 23 exx (ZMHB); 4 exx (ISEA); Bayrische Alpen, Friedergries, 24.IV.1987, J. Wolf, 1 ex (ZMHB); Büren, Ratzer, 6 exx (CAS); Darmstadt, 2 exx (DEI); Dresden, Kirsch, 7 exx (MTD); Eschenlohe, Murnau O.B., 3.VIII.1937, 8 exx (ZMHB); Freiburg, V.1939, 1 ex (ETH); Garmisch, 4.VII.1932, 3 exx (ZMHB); 18.VII.1934, 22 exx (ZMHB); 22.VI.1935, 2 exx (ZMHB); 26.V.1942, 3 exx (ZMHB); Garmisch, 1 ex (FMNH); Gross­Gerau, 11.V.1957, H. Vogt, 1 ex (ZMHB); Huckeswagen, Eigen, 3 exx (CAS); Mittenwald, Isartal, VIII.1935, Benick, 4 exx (MHNG); München, 8.IX.1906, 3 exx (ZMHB); 6.VI.1910, 1 ex (ZMHB); 8.VI.1910, 7 exx (ZMHB); 17.VI.1910, 4 exx (ZMHB); 10.VI.1931, 4 exx (ZMHB); 24.VIII.1931, 1 ex (ZMHB); 4.V.1949, H. Freude, 1 ex (DEI); Ost­Alpen, Grunberg, 4 exx (NMW); Schleching, Tirder Ache, 10.V.1987, J. Wolf 1, 1 ex (ZMHB); Ulm, Württemberg, 2 xx (NMPC); 6 exx (MTD); “ Saxonia ” (localities not specified), 1 ex (BMNH); 1 ex (HNHM); 3 exx (ZMUC). Greece: Fthiotis, 470 m, 30 km SW Lamia, bank of Inachos river, 5.IV.2001, V. Assing, 2 exx (VAPC); Kalavryta, Morea, Holtz, 1 ex (NMPC); Makedonien, Nomos Kavala, Pangeo Gipfelregion, 1700 m, Matten, 30.IV.1994, Schawaller, 1 ex (SMNS); Peloponnsos, 8 km NE Kalavrita, Vouraikos river, 700 m, 30.III.1997, V. Assing, 11 exx (VAPC). Holland: Eijgelshoven, VII.1923, Ebner, 1 ex (NMW); Valck Lucassen, Valkenburg, 1.VI.1929, 1 ex (FMNH). Hungary: " Hungary ” (localities not specified), 4 exx (NMW); 1 ex (FMNH). Italy: Abruzzes, L'Aquila, torr. Aterno, 600 m, 4.VI.1952, Fagel, 12 exx (IRSNB); Anghebeni, Leno, 12.V.1992, A. Zanetti, 1 ex (AZPC); Appennino, Vignole Borde, VI.1962, 1 ex (MSNM); Aspromonte (Calabria), S. Eufemia, VI.1958, 12 exx (MSNV); Bergsoe, 9 exx (ZMUC); Brozzo, Valtrompia, Val Mala, 17.VI.1962, Bucciarelli, 1 ex (MCSN); Capriati, F. Volturno, 4.VII.1975, Rossi, 2 exx (AZPC); Carnia, Tolmezzo Casanova, torr. But, 300 m, 30.VI.1950, Fagel, 4 exx (IRSNB); Carnia, Moggio, Fella river, 400 m, 23.VI.1950, Fagel, 5 exx (IRSNB); Carnia, Arta, Cedarchis: torr. Chiarso 400 m, 22.VI.1950, Fagel, 1 ex (IRSNB); Carnia, Villa Santina, torr. Degano, 400 m, 20.VI.1950, Fagel, 12 exx (IRSNB); Cassano Spinola, Piemonte, V.1932, G. B. Moro, 2 exx (MCSN); Civita Cast, 25.V.1975, Rossi, 1 ex (AZPC); Colle del Forno, Montelibretti, 17.VI.1977, Rossi, 1 ex (AZPC); Cusercoli, Fiume Ronco, 8.V.1976, Sama, 1 ex (AZPC); Episcopia, Sinni river, 500 m, 22.VII.1993, A. Zanetti, 1 ex (AZPC); Fiume Sesia, Carpignano, 1.VII.1958, Rosa, 1 ex (NMW); Florency, Linke, 1 ex (MTD); Frisanco, v. Colvera­rive, 1.V.1990, A. Zanetti, 1 ex (AZPC); Fumane, Rive Torrente, 1.V.1974, A. Zanetti, 1 ex (AZPC); Goriziano, 25.V.1909, 2 exx (NMPC); Goriziano, Isola Morosini, 25.IV.1909, 1 ex (MSNM); Guarcino, 28.V.1998, 1 ex (FMNH); Gubbio, 9.VIII.1891, 1 ex (MSNM); Isola Farnese, 2 exx (MNHN); Isonzo, 22.VI.1947, Sagrado, 1 ex (MSNM); Lecce, D. Barros, 3 exx (MNHN); Liguria, S. Martino, 1.V.1967, Binaghi, 5 exx (MCSN); Lombardia, Prov. Varese, Gavirate, V.1933, G. Binaghi, 7 exx (MCSN); Lombardia, Brianza Case, Bracchi, VII.1959, R. Nava, 3 exx (MSNM); Lombardia, Monza­Parco, V.1957, Brivio, 2 exx (MSNM); Lombardia, Massiccio Grigna, 1000 m, 5.VII.1960, 1 ex (MSNM); Lombardia, Correzzana, 5.IV.1959, 1 ex (MSNM); Milano, Parco Monza, IV.1957, 1 ex (MSNM); Montelibretti (Roma), 15.IV.1975, W. Rossi, 3 exx (AZPC); Passo rest, 2.VI.1970, Dioli, 1 ex (AZPC); Piemonte, Valdengo, Torr. Quargnasca, 4.VIII1959, 1 ex (MSNM); Prato de Mollo, 5.VI.1948, 6 exx (MNHN); Roma, 1916, 1 ex (MSNM); Ronchi, 31.V.1990, 1 ex (AZPC); Rovereto, 20.VI.1939, Brundin, 4 exx (MZLU); San Canzian, F. me Isanzo, 21.IV.1973, A. Zanetti, 3 exx (AZPC); Sant Eufemia, 26.VI.1958, M. Magistretti, 1 ex (MSNV); Sicilia, Floresta, VII.1959, M. Nebrodi, 1 ex (MSNV); Stazzano Scrivia, 18.IX.1875, Ferrari, 2 ex (MCSN); Stazzano Scrivia, 12.VI.1869, Ferrari, 2 exx (MCSN); Tarcentoiuss, Musi valley, 13.IX.1953, 5 exx (MSNM); Torino, 8.VI.1909, T. Sangone, 2 exx (MNHN); Torino, T. Sangone, 8.VI.1909, 2 exx (MSNM); Torino, 1871–72, L. Fea, 7 exx (MCSN); Torre Pellice, 25.VII.1932, 1 ex (MSNM); Toscana, Viareggiu, 9.III.1924, 1 ex (MSNM); Toscana, Guazzino, IV.1922, Marchi, 1 ex (MSNM); Tra Toano e Carbineri, Rive f. Secchid, 26.V.1985, A. Zanetti, 1 ex (AZPC); Trentino, Leo, 1.V.1996, A. Zanetti, 5 exx (AZPC); Turania, 15.VI.1977, Rossi, 4 exx (AZPC). Lithuania: Kuzy, VIII.1922, Klička, 1 ex (NMPC). Marocco: Marakesch, 1 ex (NMW). Portugal: Algarve, Caldas de Monchique, V–VI.1960, Fagel, 1 ex (IRSNB); Algarve, Odelouca, 4 km NW Silves, 2.V.1998, Hieke & Wendt, 13 exx (ZMHB); 5 exx (ISEA); Algarve, Caldas de Monchique, V–VI.1960, Fagel, 3 exx (IRSNB); Coimbra, 4.VI.1962, 3 exx (NMW); Estremadura, Serra de Montejunto, 2 km N Alcoentre, 100 m, 17.VII.1966, M. E. Bacchus, 1 ex (BMNH); Guarda, Gouveia, 11.VI.1970, Comellini, 1 ex (MHNG); Portimao, Algarve, 13.IV.1969, Comellini, 9 exx (MHNG); Viseu, 11.VI.1970, A. Comellini, 1 ex (MHNG). Romania: Banat, 1 ex (NMW); Drobeta­Turnu Severin, 10–11.VI.1995, Prudek, 11 exx (MSPC); Herkulesbad, Breit, 1 ex (NMW). Slovenia: Celje, 14.VII.1931, V. Koovic, 1 ex (MTD); Kamnik, 10.VIII.1907, Rambousek, 1 ex (NMPC). Spain: Andalucía, Algeciras, 17.IV.1981, Hozm, 1 ex (VAPC); Andalucía, Algeciras, Sierra de Luna, 350 m, 28.III.1994, V. Assing, 11 exx (VAPC); Andalucía, Benacjan, 5.X.1952, Coiffait, 7 exx (MNHN); Andalucía, Ronda, Arriate rio Guadalcobacin, 8–20.V.1956, G. Fagel, 63 exx (IRSNB); Aragon, Albarracin, 1200 m, VI–VII.1925, 2 exx (ZMHB); Barcelona, Tordera river, 2.IV.1986, 3 exx (VAPC); Buferrada, Valdefrancos, 1.VI.1970, Comellini, 3 exx (MHNG); Cadiz, Los Barrios, 20.V.1978, G. Sama, 1 ex (SMNS); Cadiz, Jimena de la Frontera, 5–6.VI.1991, Wrase, 14 exx (VAPC); Cadiz, Rio Alberite river, 9 km SE Alcala de los Gazules, 7.V.1991, F. Hieke, 4 exx (ZMHB); Cangas, 5 exx (BMNH); Catalogne, Seo de Urgel, Segre, V–VI.1962, Fagel, 4 exx (IRSNB); Cerro de Cuenca, 2 ex (BMNH); Cerro de Cueuca, 16.VI.1970, 3 exx (MHNG); Cordoba, Palma de Rio, 2.VI.1967, A. Comellini, 1 ex (MHNG); El Barco de Valdeorras, G. C. Champion, 2 ex (BMNH); Estremadura, Guadalupe, Guadalupejo river, V.1958, Fagel, 10 exx (IRSNB); Gerona, Ribas de Freser, 21.IX. 1989, 900 m, Schawaller, 2 exx (SMNS); Gerona, Ripoll, V.1965, Comellini, 1 ex (MHNG); Granada, Granada­Mulhacen Rd, 8 km E of Granada, 800–900 m, 21.V.1967, R. Genil, 3 exx (BMNH); “ Grenade ”, 1 ex (MNHN); Huelve, Rio de Cafa, 5 km N Santa Olalla del Cala, 450 m, 9.V.1991, F. Hieke, 1 ex (ZMHB); Jaen, Valdepenas de Jaen, Trib. of R. Guadajoz, 900–1000 m, 25.V.1967, M. E. Bacchus & B. Levey, 4 exx (BMNH); La Coruna, Arzna, 2.VI.1970, A. Comellini, 1 ex (ISEA); Madrid, 1 ex (MNHN); Ponferrada, Paganetti, 3 exx (FMNH); Potes, 2 exx (BMNH); Seo de Urgell, Lerida, 15.VI.1969, Comellini, 6 exx (MHNG); Soria, G. C. Champion, 5 exx (BMNH); Teba / Andalucía, Prov. Malaga, 26.III.1998, Esser, 1 ex (VAPC); Tragacete, G. C. Champion, 2 exx (BMNH); Zamora, Lago di Sanabria, 3.VI.1971, Comellini, 3 exx (MHNG). Switzerland: Aarberg, VIII.1935, 1 ex (ETH); Basel, 1 ex (NHMB); Büren an der Aare, 3 exx (NMW); Jhrich, 11.V.1909, 4 exx (NHMB); Laupen, 16.VII.1972, 4 exx (MHNG). Tunisia: El Kef, 14.V.1988, 1 ex (ZMUC). Turkey: Alanya, 15 km W Turkler, 5.IV.1996, A. Kopetz, 2 exx (NMEG); Zonguldak, Alapli, 15.V.1976, Busuchet & Loebl, 2 exx (MHNG). Ukraine: Kiev, Linke, 2 exx (MTD).   
 Redescription. Body. Length 2.5–3.2 mm; convex, parallel­sided, glossy; head, pronotum and elytra pitchy brown to black, abdomen black with two basal tergites reddish yellow, legs testaceous, antennae brown with antennomeres 1–3 red.   
Head subquadrate in outline; convex, eyes large, moderately protruding from lateral contours of head, length of each seen from above subequal to that of postocular region; surface of head without microsculpture; puncturation fine and moderately dense. Antennae short, clearly increasing in width apically, antennomeres 2 and 3 subequal in length, antennomeres 4–8 longer than wide, antennomeres 9–10 sexually dimorphic, quadrate in male and slightly transverse in female, antennomere 11 nearly conical.  
Pronotum quadrate, lateral sides gradually narrowed in straight line to obtuse hind angles; before base with small and shallow transverse impression; surface without microsculpture; puncturation fine, moderately scattered and weakly asperate; pubescence at midline directed posteriorly in apical half and anteriorly in basal half.  
Elytra subquadrate, at suture as long as pronotal length at midline; surface lacking microsculpture; puncturation fine, moderately dense and slightly asperate.  
Abdomen parallel­sided, bases of tergites 3–5 each with deep transverse impression, impressions with 7–8 longitudinal ridges, tergal puncturation fine and very scattered, punctures very small and scarcely visible, surface without microsculpture.  
Male. Aedeagus as in Figs. 26–27.  
Female. Spermatheca as in Fig. 28.  
 Remarks. Tachyusa balteata is similar to T. ferialis Erichson, 1839, from which it can be easy distinguished by the black pronotum (red in T. ferialis), the larger eyes, the stouter body, the more distinct and denser pronotal puncturation and by the shape of the genitalia.   
 Distribution. T. balteata is widespread in Europe (except north) and North Africa.
description: Published as part of Pa ṡnik, Grzegorz, 2006, A revision of the World species of the genus Tachyusa Erichson, 1837 (Coleoptera, Staphylinidae: Aleocharinae), pp. 1-152 in Zootaxa 1146 on pages 40-45