Glyphiulus intermedius Golovatch & Geoffroy & Mauriès & Spiegel 2007, n. sp.
Creators: name: Golovatch, Sergei I.
name: Geoffroy, Jean-Jacques
name: Mauriès, Jean-Paul
name: Spiegel, Didier Van Den
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Spirostreptida
Cambalopsidae
Glyphiulus
Glyphiulus intermedius

description: Glyphiulus intermedius n. sp.   
(Figs 25; 26)  
 TYPE MATERIAL. — China. Sichuan Prov., Chengdu County, Huan Long Dong Cave (No. 593), 26.VIII.1999, leg. J. & B. Lips, holotype ♂ (MNHN GA 053); paratypes 1 ♂, 2 ♀♀, 1 juv (MNHN GA 053); 1 ♀ (SEM); 1 ♀ (ZMUM).   
 ETYMOLOGY. — To emphasize a somewhat intermediate position between Podoglyphiulus species and the remaining species of the javanicus -group.   
DIAGNOSIS. — Differs from congeners in the particularly long ozoporiferous tubercles, the special carinotaxy formula on the collum, coupled with certain details of gonopod structure (see also key below).  
DESCRIPTION  
Holotype with 43p+4a+ T, about 26 mm long and 1.6 mm wide; adult paratypes with 38p+4a+ T, about 22 mm long and 1.5 mm wide (♂), or with 43p+4a+ T, about 26 mm long and 1.6 mm wide (♀); with 53p+2a+ T, about 37 mm long and 2.0 mm wide (♀); with 56p+1a+ T, about 36 mm long and 2.2 mm wide (♀).  
Coloration from uniformly pallid to marbled light yellow- or grey-brown, ozoporiferous cones often contrastingly dark brown; antennae, legs and venter yellowish to light yellow-brown; in adults, 8-12 blackish ocelli in a subtriangular eye patch on each side of head.  
 Antennae (Fig. 26A), gnathochilarium (Fig. 26B), postcollar constriction (Fig. 25B), dorsal crests and carinotaxy pattern on postcollar segments (Fig. 25), segment’s texture and cross-section (Fig. 25F), paraprocts, ventral flaps of male segment 7, male legs 1-3 (Fig. 26 C-E), penes (Fig. 26D) generally as in G. percostulifer n. sp., but collum (Fig. 25A, B) with a slightly deviating carinotaxy formula (I+2c+III-IV+5c +6a+pc+ma+pc+6a+5c+IV-III+2c+I); all tergal crests (Fig. 25 A-E) on subsequent body segments higher, subdivided (2/2+I/i+3/3+I/i+2/2), ozoporiferous cones higher than wide; anterior half of paramedian dorsal crests particularly high, subtriangular on first 10-11 postcollar segments (Fig. 25A, B), thereafter somewhat lower (Fig. 25 C-F); epiproct with a small but distinct tubercle at midway dorsally (Fig. 25E), very broadly rounded in caudal part in ventral view; hypoproct slightly but evidently emarginate mediocaudally; legs about as long as body diameter (Fig. 25F), always with a short accessory spine at base of claw (Fig. 26 C-F).   
Anterior gonopods (Fig. 26G) with coxosternum not plate-shaped but narrowed/elongate distally; telopodites straight. Posterior gonopods (Fig. 26H, I) also slightly elongate, flagella long and bare.  
REMARKS  
The juvenile paratype is haplopodous, with 32p+2a+T. As usual, the ozoporiferous cones on segments 5 and 6 are considerably larger than the others.  
 In the absence of clear troglomorphic traits, this species seems to be troglophilic at most. In addition, the cave is described as being “very small, full of bats and guano” (J. Lips in litt.).   
 Even though the gonopods of this new species are not very typical of the group concerned, their similarity to, and ready derivation from, the basic patterns is unquestionable. An assignment to Podoglyphiulus can be excluded on both morphological and zoogeographical grounds (see above).
description: Published as part of Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group, pp. 417-456 in Zoosystema 29 (3) on pages 446-449, DOI: 10.5281/zenodo.5393630