Myopsocidae Pearman 1936
Creators: name: González-Obando, Ranulfo
name: Calderón-Martínez, Nadia R.
name: Carrejo-Gironza, Nancy S.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Psocodea
Myopsocidae

description: Key to South American species of Myopsocidae
(M. cinereus Navas, 1932, not included because there are no illustrations or photographs in the original description)
1. Hindwing with Rs and M veins fused for a distance (Fig. 68).................................................... Myopsocus .......................................................................................... 2
- Hindwing with Rs and M veins joined by a transverse vein (Fig. 2)............................................... Lichenomima ....................................................................................... 25
2. Species in which only females are known.................................................................. 3
- Species in which males and females or only males are known.................................................. 7
3. Forewings festooned apically, mainly on the apical margin of cell m2 (Fig. 79).................................... 4
- Forewings not festooned apically (Fig. 67)................................................................. 6
4. Valve V1 basally narrower, with a dorsal tooth-shaped projection............................ M. vespertilio Mockford
- Valve V1 with uniform width, without dorsal projections...................................................... 5
5. Forewing with large areas of depigmentation on M1 and M2 cells............................ M. albomaculatus (New)
- Forewing with or without small depigmented areas M1 and M2 cells; pterostigma darker in distal half..... M. cinctus (New)
6. Apical sclerite process subgenital plate long, narrow and with setae......................... M. albus (Garcia-Aldrete)
- Apical sclerite process subgenital plate very short and without setae........................... M. circularis Mockford
7. Forewings festooned apically, especially on the apical margin of cell M2. (Figs 73, 79, 85)........................... 8
- Forewings without festooned apical margin (Fig. 133)....................................................... 12
8. Hypandrium with latero-posterior processes (Figs 77, 113).................................................... 9
- Hypandrium without latero-posterior-processes (Figs 83, 119)................................................. 10
9. Hypandrium widened apically, with widely separated latero-posterior processes (Fig. 113); phallosome with side struts convergent basally (Fig. 114)........................................................... M. colombianus n. sp.
- Hypandrium narrow apically, with latero-posterior processes close together (Fig. 77); phallosome with side strut well separated basally, external parameres and side struts more widened than in the previous one (Fig. 78).......... M. amazonensis n. sp.
10. Hypandrium V-shaped, apically widened (Fig. 119); phallosome with side struts basally fused, very narrow proximally (Fig. 120), external parameres broadly rounded apically.......................................... M. garcialdretei n. sp.
- Hypandrium V-shaped, apically narrow (Fig. 83); phallosome with side struts basally separated but medially fused, external parameres not widened apically as above................................................................. 11
11. Forewing with M1 and M2 cells with apical hyaline area reaching to the wing margin (Fig. 79); endophallic membrane without spicules (Fig. 84)...................................................................... M. andaquies n. sp.
- Forewing without hyaline areas in cells M1 and M2 as above (Fig. 127); endophallic membrane with abundant spicules (Fig. 132)...................................................................................... M. inga n. sp.
12. Epiproct with conspicuous posterior horn.................................................. M. dispar (Enderlein)
- Epiproct without posterior horn (Fig. 136)................................................................. 13
13. Hypandrium subtrapezoidal, posterior margin with small tubercles on both sides of a narrow medial process (Fig. 137).......................................................................................... M. insularis n. sp.
- Hypandrium with variable shape, without small tubercles and medial process as described above (Figs 95, 107, 168)..... 14
14. Clunium male with a pair of rounded lobes on the dorsum, covered by small spicules and setae (Fig. 178); phallosome oval (Fig. 177).............................................................................. M. minutus (Mockford)
- Clunium male without a pair of rounded lobes on the dorsum; phallosome with variable shape, sometimes oval (Figs 96, 108, 150, 169)........................................................................................... 15
15. Hypandrium with latero-posterior processes (Fig. 168)....................................................... 16
- Hypandrium without latero-posterior processes (Fig. 71)..................................................... 17
16. Hypandrium with widely separated latero-posterior processes, directed outward (Fig. 168), epiproct with strongly convex anterior margin (Fig. 166)............................................................... M. tukunare n. sp.
- Hypandrium with closely spaced latero-posterior processes, directed caudally, these are short, widened, with rounded lateralposterior margin and small tooth-shaped processes (Fig. 107); epiproct with concave anterior margin (Fig. 106).............................................................................................. M. chocoensis n. sp.
17. Phallosome very narrow basally (Figs 72, 96, 173).......................................................... 18
- Phallosome basally widened or greatly widened (Figs 102, 150, 162, 173)....................................... 20
18. Hypandrium with small medial apical bifurcated process......................................... M. pennyi (New)
- Hypandrium without medial apical bifurcated process (Fig. 95)................................................ 19
19. Phallosome with external parameres apically briefly separated by membrane, endophallus elongated, flattened and apically more splayed (Fig. 96)................................................................... M. caliensis n. sp.
- Phallosome with external parameres broadly fused medially and its endophallus is not elongated and apically more broadened (Fig. 72)............................................................................ M. amacayacu n. sp.
20. Phallosome basally broadened, curved inward, apically rounded, with narrow posterior lobes (Fig. 150), endophallus membranous........................................................................................ 21
- Phallosome basally narrower than distally or membrane-bound side struts, posterior lobes broadened (Figs 102, 156, 162)... .................................................................................................. 22
21. Epiproct with convex anterior margin, projecting onto the clunium; posterior process of paraprocts with rounded apex (Fig. 148); phallosome with widened side struts (Fig. 150)............................................. M. katios n. sp.
- Epiproct semioval, with anterior margin straight to concave; paraprocts with posterior process acuminates; phallosome with narrow side struts (Fig. 176)....................................................... M. minor (New & Thornton)
22. Phallosome not piriform, side struts basally separate or widely united external parameres broad distally (Fig. 162)....... 23
- Phallosome piriform, with side struts basally fused or joined by membrane (Figs 102, 156).......................... 24
23. Epiproct hexagonal, distal-lateral sides longer than others and heavily sclerotized; phallosome with side struts basally separated, joined by membrane, external parameres with internal margin gently curved and apically joined by membrane (Fig. 174)....................................................................................... M. parvus Mockford
- Epiproct semioval (Fig. 160); phallosome with side strut basally united by broad sclerotized area, external parameres with internal margin strongly curved, apically converging mesally (Fig. 162)........................ M. putumayensis n. sp.
24. Phallosome with side struts basally fused (Fig. 102); forewing with length/width ratio of approximately 2.8 (Fig. 97).......................................................................................... M. caquetensis n. sp.
- Phallosome with side struts basally not fused, joined by membrane (Fig. 156); forewing with length/width ratio of approximately 2.4 (Fig. 151)........................................................................... M. panchei n. sp.
25. Species in which only females are known................................................................. 26
- Species in which males and females or only males are known................................................. 29
26. Forewing with large depigmented area from the rear angle pterostigma-stigmasac, diagonal to the posterior margin, interrupted only by a point around the vein M before the intersection between the vein M and Cu1a (see Fig. 1 in Williner 1910).......................................................................................... L. argentina Williner
- Forewing without large depigmented area from the rear angle pterostigma-stigmasac, diagonal to the posterior margin.... 27
27. Vertex emarginated; forewing with M1 – M3 cells and apical half of R5 completely pigmented (see Fig. 175 in New & Thornton 1975)....................................................................... L. pulchella (New & Thornton)
- Vertex not emarginated, forewing with speckled pattern of coloration........................................... 28
28. Apical sclerite of subgenital plate with a medial group of setae and truncated apically................ L. timmei Mockford
- Apical sclerite of subgenital plate without setae and rounded apically........................... L. thorntoni Mockford
29. Phallosome with side struts fused at least at base, narrow or greatly enlarged, basally rounded; hypandrium variable in its shape (Figs 11, 29, 41)..................................................................................... 32
- Phallosome with side strut separate, joined only by membrane (Fig. 172); hypandrium triangular, longer than wide....... 30
30. Hypandrium with lateral processes long and slender.............................................. L. varia (Navas)
- Hypandrium with lateral processes short, pointed corners..................................................... 31
31. Hypandrium with lateral processes scarcely protruding beyond edges; forewing without clear band parallel to apical margin...................................................................................... L. onca Mockford
- Hypandrium with lateral processes obviously protruding as pair points; forewing with clear band parallel to apical margin.................................................................................... L. conspersa Enderlein
32. Hypandrium shorter than wide, bilobed apically (Fig. 29); phallosome widely enlarged, with rounded anterolateral angles, with medial sclerotization (Fig. 30), external parameres with elongated tooth-like processes on their inner margin................................................................................................... L. huilensis n. sp.
- Hypandrium as long as wide or longer than wide; phallosome elongated......................................... 33
33. Epiproct vase-shaped (Fig. 179), hypandrium triangular, phallosome with external parameres present as short, slender arms (Fig. 170)...................................................................... L. punctipennis Mockford
- Epiproct variable in shape, not vase-shaped; variable phallosome.............................................. 34
34. Epiproct with superficial dorsal flap (Fig. 40); hypandrium with extensive basally projecting sclerotization (Fig. 41)...... 35
- Epiproct without superficial dorsal flap (Figs 4, 10 and 34); hypandrium is different, not like the above (Figs 11, 41)..... 37
35. Phallosome rounded basally (Fig. 175)......................................................... L. ariasi (New)
- Phallosome concave basally (Fig. 42).................................................................... 36
36. Phallosome broadened basally (Fig. 42), external parameres narrow, curved inward, sharply angled, aedeagus distally widened, endophallic membrane with well sclerotized apical spinous processes.............................. L. lienhardi n. sp.
- Phallosome narrow basally, external parameres distally widened, curved inward but not strongly angled as above, aedeagus apically bilobed, each lobe with elongated processes, endophallic membrane without apical spinous processes.................................................................................................... L. ariasi (New)
37. Phallosome with side strut apically distinct, elongated, slender, or distally curved inward (Fig. 12); hypandrium U-shaped, usually longer than wide (Fig. 11)....................................................... L. gorgonaensis n. sp.
- Phallosome with side strut apically undifferentiated in its apical part of the external parameres, hypandrium with variable shape.............................................................................................. 38
38. Phallosome sledgehammer-shaped, external parameres as two flattened and converging lobes (Fig. 5)................. 39
- Phallosome with variable shape but not sledgehammer-shaped, variable external parameres (Figs 24, 36 and 48)......... 41
39. Phallosome with side strut basally close together (Fig. 5)..................................... L. bonaverensis n. sp.
- Phallosome with side strut separate basally, bound together by a membrane (Fig. 60) or somewhat broadened and rounded basally (Fig. 171).................................................................................... 40
40. Phallosome with external parameres with small apical lobes, endophallus with large sclerotized area and membrane with abundant spicules as illustrated (Fig. 60)..................................................... L. pearmani n. sp.
- Phallosome with external parameres without apical lobes as above, endophallus without sclerotized area and spicules as above (Fig. 171)...................................................................... L. chelata Thornton & Woo
41. Hypandrium with preapical constriction, forming a broad distal lobe (Fig. 35); phallosome with mesal endophallic process, more sclerotized and somewhat acuminate apically (Fig. 36)...................................... L. iscuande n. sp.
- Hypandrium without preapical constriction (Figs 23, 47); phallosome without mesal endophallic process (Fig. 48)....... 42
42. Phallosome with side strut widened, external parameres apically more broadened (Fig. 48), epiproct semioval (Fig. 46)........................................................................................ L. occidentalis n. sp.
- Phallosome with side strut narrow, external parameres evenly broadened (Fig. 24); subtrapezoidal epiproct (Fig. 22)............................................................................................ L. guapiensis n. sp.
description: Published as part of González-Obando, Ranulfo, Calderón-Martínez, Nadia R. & Carrejo-Gironza, Nancy S., 2023, New species of Myopsocidae (Insecta: Psocodea: " Psocoptera ") from Colombia, pp. 1-61 in Zootaxa 5377 (1) on pages 7-9, DOI: 10.11646/zootaxa.5377.1.1, http://zenodo.org/record/10208826