Palaeogronotoma nordlanderi Peñal & Fontal-Ca & Pujade- 2013, n. sp.
Creators: name: Peñal, Enrique
name: Fontal-Ca, Félix M.
name: Pujade-, Juli
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Hexapoda
Hymenoptera
Figitidae
Palaeogronotoma
Palaeogronotoma nordlanderi

description: Palaeogronotoma nordlanderi n. sp. (Figs 1; 2)
HOLOTYPE. — Alate adult MPZ-97/547 (additional number RM-RR-276), in a small slab of oil-shale with dimensions 4 × 3 × 0.3 cm, housed in Museo de Paleontología de la Universidad de Zaragoza (Zaragoza, Spain). Specimen in lateral position, virtually complete and well preserved, having lost only some portions of the wings and the mesosoma (some of the legs and one antenna are overlapped under the body). The same surface with the holotype also contains a Mycetophilidae (Diptera) of the genus Exechia (MPZ-96/19) (currently on a separate portion of the original slab) and shells of ostracods.
DERIVATION OF NAME. — The new species is named after Dr. Göran Nordlander, specialist in Cynipoidea. The epithet nordlanderi is in the genitive case.
TYPE HORIZON. — Specimen discovered in lacustrine oil-shales from the Early Miocene (Early Burdigalian) of Rubielos de Mora Basin (Iberian Chain).
TYPE LOCALITY. — “Río Rubielos 2” outcrop, Rubielos de Mora village, Teruel Province, Spain.
DIAGNOSIS. — As for genus.
DESCRIPTION (EXCLUDING
THE DIAGNOSTIC CHARACTERS)
Body dark and large, 1.5 mm long (Fig. 2A). Female antenna with 6-segmented club; club segments distinguished by presence of rhinaria (Fig. 2C, D). Antenna 1.11 mm long. Measures of the antennal segments (length × width, in mm) are: S1:? × 0.06, S2: 0.03 × 0.04, S3: 0.10 × 0.02, S4: 0.05 × 0.03, S5: 0.06 × 0.03, S6: 0.06 × 0.03, S7: 0.07 × 0.03, S8: 0.07 × 0.05, S9: 0.08 × 0.04, S10: 0.08 × 0.04, S11: 0.07 × 0.05, S12: 0.07 × 0.05, S13: 0.15 × 0.07. Mesosoma 0.59 mm long and 0.57 mm high (Fig. 2A, E). Estimated forewing length 1.38 mm. Radial cell closed on front margin and about two times longer than wide (internal measures) (Fig. 2B). Hind leg 1.32 mm long (femur about 0.34 mm, tibia 0.46 mm, tarsus 0.52 mm). Metasoma, 0.78 mm long and 0.65 mm high, with the segment 1 obscured by segment 2, that occupies the main visible part of metasoma (Fig. 2A). Metasomal segments 3, 4 and 5 visible.
PALAEOBIOLOGY
Unknown, but all recent genera closely related to this new genus are parasitoids of Agromyzidae (Diptera). The family Agromyzidae is commonly referred to as the leaf-miner flies, for the feeding habit of larvae, most of which are leaf miners on various plants. No agromyzid adult specimens or leaves with distinctive leaf-mines by agromyzids have been identified from the Rubielos de Mora fossil record up to now. Diptera, the most abundant and diverse group in Rubielos de Mora site containing 13 families identified, was monographed by Peñalver (2002), but several fossil specimens of small morphotypes remained unstudied and many others were found later, thus possibly the family Agromyzidae is also recorded.
description: Published as part of Peñal, Enrique, Fontal-Ca, Félix M. & Pujade-, Juli, 2013, Palaeogronotoma n. gen. from the Miocene of Spain, the first Tertiary fossil record of the subfamily Eucoilinae (Hymenoptera: Figitidae), pp. 643-653 in Geodiversitas 35 (3) on pages 646-647, DOI: 10.5252/g2013n3a7, http://zenodo.org/record/5380573