Illiesonemoura bituberculata Wang & Du 2023, sp. nov.
Creators: name: Wang, Zhi-Jie
name: Du, Yu-Zhou
name: Ji, Xiao-Yu
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Plecoptera
Nemouridae
Illiesonemoura
Illiesonemoura bituberculata

description: Illiesonemoura bituberculata Wang & Du sp. nov.  
Materials  
 Type status: Holotype. Occurrence: recordedBy: Yu-zhou Du; individualID: Insect collection of Yangzhou University (ICYZU), Jiangsu Province, China; individualCount: 1; sex: male; lifeStage: adult; occurrenceStatus: present; occurrenceID: 82D320BE-8137-54E6-B600-64B17A0A86DE; Taxon: kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Plecoptera; family: Nemouridae; genus: Illiesonemoura; specificEpithet: bituberculata; Location: continent: Asia; country: China; countryCode: CN; stateProvince: Sichuan; locality: 15 km west of Kangding, Zheduo River; minimumElevationInMeters: 2920; verbatimLatitude: 29.994447N; verbatimLongitude: 101.892415; Identification: identifiedBy: Zhi-jie Wang, Yu-zhou Du & Xiao-yu Ji; Event: year: 1996; month: 6; day: 8; Record Level: language: en Type status: Paratype. Occurrence: recordedBy: Yu-zhou Du; individualID: Insect collection of Yangzhou University (ICYZU), Jiangsu Province, China; individualCount: 1; sex: male; lifeStage: adult; occurrenceStatus: present; occurrenceID: 7053928D-FDFD-51B3-AF16-C74A29F497D6; Taxon: kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Plecoptera; family: Nemouridae; genus: Illiesonemoura; specificEpithet: bituberculata; Location: continent: Asia; country: China; countryCode: CN; stateProvince: Sichuan; locality: 15 km west of Kangding, Zheduo River; minimumElevationInMeters: 2920; verbatimLatitude: 29.994447N; verbatimLongitude: 101.892415; Identification: identifiedBy: Zhi-jie Wang, Yu-zhou Du & Xiao-yu Ji; Event: year: 1996; month: 6; day: 8; Record Level: language: en   
Description  
 Male. Head brown, antennae slightly brown (Fig. 2); pronotum paler than head, anterior margin wider than posterior margin. A single unbranched cervical gill as typical of the genus present on each side of mid-line (Fig. 1 A). Wings subhyaline, fumose, veins brown. Forewing length ca. 6 mm, hind wing 5.5 mm. Legs brown (Fig. 2). Tergum 9 slightly sclerotised, extending posteriorly as a broad lobe over tergum 10. Tergum 10 slightly sclerotised anteriorly, but being more sclerotised along the posterior margin where a pair of lateral knobs and a pair of strong median prongs are present, both most noticeable in dorsal view, a field of 5-6 tiny spines present anteromedially (Fig. 1 B). Hypoproct broad basally and tapering to a small, narrow tip, vesicle plump, length 2x width (Fig. 3 B). Cerci mostly membranous, long thin and bent inwards (Fig. 1 B, Fig. 3 A-C). Paraprocts divided into two lobes; inner lobe slightly sclerotised, thin and long; outer lobe broad basally, large triangular, outer part curving dorsally, with outer margin darkly sclerotised and with outward directed sclerotised prong from outer margin near cerci (Fig. 3 B and D). Epiproct short and broad in dorsal aspect; dorsal face with a pair of sclerotised bands each side of mid-line and distal margin of dorsal sclerite darkly sclerotised, extending anterolaterally, enveloping tip of ventral sclerite, which is more distinct in lateral and ventral views; basal sclerite membranous; ventral sclerite strongly sclerotised, broad basally and with lateral knobs at basolateral corners, becoming narrower towards apex, forming a pair of ridges each side of mid-line, each bearing a row of spines on anterior half (Fig. 3 E-G).   
Diagnosis  
 The male of this species is characterised by two pairs of small knobs arising posteromedially from tergum 10. They are similar to I. tuberostyla (Wu 1962) in the lateral view of the epiproct, both looking like tumours, but can be separated from the latter by the knobs protruding from the posterior margin of tergum 10. In I. tuberostyla, there was no mention of knobs on the terga.   
Etymology  
 The name refers to the pair of knobs and strong prongs located posteromedially on tergum 10. The Latin adjective " tuberculata " means having knobs.   
Distribution  
China: Sichuan Province.
description: Published as part of Wang, Zhi-Jie, Du, Yu-Zhou & Ji, Xiao-Yu, 2023, Three new species of Illiesonemoura Baumann, 1975 (Plecoptera, Nemouridae) from China, pp. 112020 in Biodiversity Data Journal 11 on page 112020, DOI: 10.3897/BDJ.11.e112020