Clivina hartmanni Balkenohl, 2025, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina hartmanni

description: Clivina hartmanni sp. nov.
(Figs 22, 57, 90, 116)
Type material. Holotype: ♂, with labels and data: white, black printed “ INDONESIA, Irian Jaya Jayapura district Genyem, 50m NN, IV.-V. 1999,leg. M.Balke ” / “Sammlung NATURKUNDE- MUSEUM ERFURT” (NKME). Paratype: 1 ♀, same data as holotype (CMBB).
Description. Measurements. Body length 5.86 / 5.91 mm, width 1.79 / 1.77 mm, ratio L/W of pronotum 0.88 / 0.88, ratio L/W of elytra 1.77 / 1.78. Colour glossy, fuscous. Supraantennal plate at margin slightly translucent. Antennae, mouthparts, intermediate and hind legs yellowish.
Head about a fifth narrower than pronotum. Clypeus with central part straight, projection of lateral teeth just traceable, separated from clypeal-wing by small notch; wing rounded, nearly as high as central part of clypeus, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin convex at middle, clypeus reflexed margined.Supraantennal plate extended posteriorly into a blunt decreasing convexity, separated from wing by small furrow. Clypeus and frons forming continuous moderate convexity, with scattered small punctures, clypeal-furrow not traceable. Frons with pore at middle, separated from supraantennal plate and supraorbital carina by flat furrow, with clypeal seta in front; supraorbital carinae moderately developed, with two supraorbital setigerous punctures at middle of eye and at middle of gena-level. Neck constriction widely open, indicated laterally. Eye prominent but somewhat flattened, gena enclosing eye posteriorly by nearly a third. Antenna elongate, reaching over base of elytron by one to two antennomeres. Labrum excised anteriorly, finely irregularly reticulated, seven-setose. Lobe of mentum oval acuminated, flat, median tooth flat, distinctly lower than lobe, rectangular at apex.
Pronotum (Fig. 57) with disk flattened in lateral view, in frontal view laterally moderately convex, flattened at middle. Shape square to sub-cordate, wider than long. Reflexed lateral margin smooth, widest at middle; anterior and posterior angles distinct, posterior one with distinct projecting sharp tooth; lateral channel moderately narrow, continuing up to base as sharp, sigmoid margin. Median line complete, engraved at middle, joining anterior transverse line. Anterior transverse line distinct, complete. Disk covered with scattered micro-punctures, with transverse wrinkles, with isodiametric reticulation at base, with indistinct impression in basal third, with posterior setigerous puncture situated directly at tooth of posterior angle. Lateral margin from posterior angle to flange of base slanted with angle of around 30°. Basal channel on flange somewhat widened, rugose, with sharp carina.
Elytron (Fig. 22) with disk flattened in anterior half in lateral view, moderately convex in frontal view. Shape of elytra elongate, slightly less than two times longer than wide, with lateral margin indistinctly convex at middle, diverging posteriorly, maximum width posterior middle. Humerus rounded off, base retracted. Reflexed lateral margin slightly crenulate behind humerus. Row of setigerous punctures in lateral channel regular in size, more distant at middle, with about every fourth seta extraordinarily elongated, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with small but distinct tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, striate-punctate, one to four free at base, five joining humerus, six ending free at humerus, one and two ending free at apex, three and four and five and six joining apically. Intervals moderately convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven with step-like carina at apex, eight carinate at apex. Surface of all intervals with distinct isodiametric reticulation already well visible at 25-fold, slightly less reticulated at disk of first interval.
Hind wing fully developed.
Lower surface. Proepisternum regularly isodiametrically reticulated, with very fine wrinkles laterally. Sternite of abdomen with regular isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with indistinct projection posteriorly at middle, with narrow marginal furrow at middle (in both sexes).
Legs. Protibia moderately slender, with longitudinal reticulation, with sulcus dorsally, with three slender acute spines and indistinct tubercle, movable spur moderately arcuate, basal tarsomere convex, slender at base, carinate, with acute tubercle with three setae. Intermediate tibia with ten tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of intermediate and hind leg laterally widened, looking serrate.
Male genitalia (Fig. 90). Median lobe moderately arcuate at basal half, more straight in apical half, indistinctly bent to apex. Apex like an elongated cone. Parameres slender, with small apophyses, slightly distorted, with flat hyaline elongation at apex.
Female gonocoxites and epipleurite (Fig. 116). Basal gonocoxite with one long, three medium long, and two small setae, apical gonocoxite elongated, slightly curved, oval in cross section, with one long seta at end of basal third, with one small seta basally, with two long setae apically. Epipleurite triangle-like, with two setae arising from flat tubercles.
Variation. With exception of measurements, not observed.
Distribution. Known from Irian Jaya, Indonesia.
Diagnosis. A medium sized fuscous species with sub-cordate to square pronotum, all intervals of the elytron reticulated, and with seven-setose labrum.
The most similar species with completely reticulated elytra are C. spatulifigura sp. nov., C. opacielytra sp. nov., and C. dealata antecessor Darlington. The last mentioned one is a member of the vulgivaga species-group due to its falcate gonocoxites. It also differs mainly by the smaller eyes and enlarged genae, coreaceate episternum and the acute tubercle at the front trochanter. Differences in C. opacielytra are mainly the six-setose and pentagonal labrum, and the distinctly peltate shaped pronotum. C. spatulifigura differs by its distinctly closed neck constriction, the slightly excised clypeus, and the smaller body size. Moreover, all of the genitalia are different.
Etymology. The name is a patronym in honour of Matthias Hartmann (NKME) who provided me with full support during many years and loaned material of this and other species.
description: Published as part of Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), pp. 1-70 in Zootaxa 5592 (1) on pages 37-38, DOI: 10.11646/zootaxa.5592.1.1, http://zenodo.org/record/14953674