Leptocera finalis
Creators: name: Buck, Matthias
name: Marshall, Stephen A.
Publisher: name: Zenodo
Year: 2009
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Sphaeroceridae
Leptocera
Leptocera finalis

description: <i>Leptocera finalis</i> (Collin, 1956) (Figs. 155–156, 200) <i>Paracollinella finalis</i> Collin, 1956: 175 (lectotype &amp;male;, OXUM, not examined). <i>Leptocera finalis</i>.— Rohá&amp;ccaron;ek, 1982: 24 (redescription); Marshall, 1997: 666 (first Nearctic record); Rohá&amp;ccaron;ek <i>et al</i>., 2001: 153 (World catalog). <b>Taxonomy</b>. In the Nearctic the name “ <i>L. finalis</i> ” has been misapplied to the following five species (records from Yukon Territory in Marshall, 1997): <i>L. angulispina</i> sp.n., <i>L. neofinalis</i> sp.n., <i>L. neovomerata</i> sp.n., <i>L. kanata</i> sp.n., and <i>L. parafinalis</i>. <b>Description</b>. A detailed redescription of the species was provided by Rohá&amp;ccaron;ek (1982). Outstanding paramedian acrostichals moderately enlarged, longest one about 2/3 as long as lower orbital bristle; prescutellar acrostichals also somewhat enlarged. Mid tibia with bristle above distal dorsal shorter than anteroapical bristles; dorsal posteroapical bristle shorter than ventral one (rarely almost of same length), extending well beyond socket of ventrobasal metatarsal bristle (as in Fig. 26). Male terminalia (Figs. 155, 156): Sternite 5 with posteromedial desclerotized area somewhat small (see Rohá&amp;ccaron;ek, 1982: Fig. 36). Anterior section of surstylus with anterior process in lateral view more or less rectangular and relatively short; bare lateral ridge at apex of anterior process prominent and darkened (best seen in ventral view); ventral lobe triangular and prominent, with short bristles; no bristles between long, basal, posterior bristle and apex of ventral lobe. Posterior section of surstylus with few bristles on dorsal (posterior) surface and slightly unequal apical bristles. Postgonite as in Fig. 200. Female terminalia (indistinguishable from <i>L. neofinalis</i> sp.n., Figs. 174–176): Hind margin of sternite 7 straight, simple. Sternite 8 with lateral margins angulate, posterolateral lobes well developed, triangular; median process rounded and of moderate width. Spermathecae pear-shaped with basolateral invagination, without or with small and shallow invagination near apex; spicules noticeably crowded towards base. <b>Material examined</b>. (Identification of females through association with males) CANADA. Labrador: 1 &amp;male;, 1 &amp;female;, Cartwright, 29.vi.1955, E.F. Cashman (CNCI). Nova Scotia: 1 &amp;male;, 2 &amp;female;&amp;female;, Kingsport, 26.vi.1980, pan trap, Dondale &amp; Redner (DEBU). Quebec: 1 &amp;male;, Thunder River [= Rivière-au-Tonnerre], 11.vi.1929, W.J. Brown (CNCI). Manitoba: 2 &amp;male;&amp;male;, 2 &amp;female;&amp;female;, Churchill, 21/ 22.vi.1948 G.E. Shewell and 17.viii.1952, J.G. Chillcott (CNCI); 1 &amp;female;, Churchill, 28.v.1952, muskeg, E.E. Wiffen (DEBU); 1 &amp;male;, 4 &amp;female;&amp;female;, Fort Churchill, 17.vi., 29.vi. and 11.viii.1952, J.G. Chillcott (CNCI); 1 &amp;male;, Hudson Bay Railway mi 505, 13.vi.1952, J.G. Chillcott (CNCI); 1 &amp;female;, same as previous but mi 504, 21.vi.1952. Northwest Territories: 1 &amp;male;, Hyndman Lake, 68°15’N 131°3’W, 30.vii.1969, G.E. Shewell (CNCI). Yukon Territory: 2 &amp;male;&amp;male;, 1 &amp;female;, Rampart House, 2.vi.1951, C.C. Loan (CNCI). U.S.A. Alaska: 1 &amp;male;, Healy, 26.vi.1921, J.M. Aldrich (USNM); 1 &amp;male;, Steese Hwy mile post 98.4, 4.ix.1948, G. Jefferson (USNM); 5 &amp;male;&amp;male;, Matanuska, 7–17.v.1944, rotary trap and 5–11.viii.1945, J.C. Chamberlin (USNM) [Note: part of a mixed series of 52 &amp;male;&amp;male;, 104 &amp;female;&amp;female;, with roughly 60% of the specimens pertaining to <i>L. finalis</i>; the remainder are <i>L. neofinalis</i> sp.n.]. We have also examined numerous specimens from the Novosibirsk area in Russia (DEBU). <b>Distribution</b> (Map 6). Transcontinental in boreal and arctic Canada (records from LB, NS, QC, MB, NT, YT), Alaska; Palaearctic region from Great Britain to the Russian Far East (Rohá&amp;ccaron;ek <i>et al</i>., 2001). Most records from Yukon Territory listed in Marshall (1997), except those from Rampart House (see above), actually belong to other species in the <i>L. fontinalis</i> group, especially the Nearctic sister species <i>L. neofinalis</i> sp.n. In the Nearctic, <i>L. finalis</i> extends less far southward and appears to be much rarer than <i>L. neofinalis</i> sp.n. Due to the fact that both species can only be separated by male genitalic characters it is possible that some examined series of <i>L. neofinalis</i> sp.n. (see below) might include further specimens of <i>L. finalis</i>. Even though the range of <i>L. finalis</i> seems to overlap the range of <i>L. neofinalis</i> sp.n. entirely, Rampart House (Yukon Territory) and Matanuska (Alaska) are presently the only localities known where both species occur together. <b>Discussion</b>. <i>Leptocera finalis</i>, <i>L. neofinalis</i> sp.n. and <i>L. parafinalis</i> differ from all other Nearctic members of the <i>L. fontinalis</i> group in possessing a more of less angulate (not rounded) ventral lobe of the anterior section of the surstylus, a simple female sternite 7 (hind margin neither sharp-edged nor medially produced) and pear-shaped spermathecae with spicules mostly restricted to basal half (other species have subcylindrical spermathecae with more or less evenly spiculate surface). For characters separating <i>L. finalis</i> from <i>L. neofinalis</i> sp.n. and <i>L. parafinalis</i> see Discussion under these species.
description: Published as part of <i>Buck, Matthias &amp; Marshall, Stephen A., 2009, Revision of New World Leptocera Olivier (Diptera, Sphaeroceridae), pp. 1-139 in Zootaxa 2039 (1)</i> on pages 77-78, DOI: 10.11646/zootaxa.2039.1.1, http://zenodo.org/record/5311868
description: {"references": ["Collin, J. E. (1956) Some new British Borboridae (Diptera). Journal of the Society for British Entomology, 5 (5), 172 - 178.", "Rohacek, J. (1982) Revision of the subgenus Leptocera (s. str.) of Europe (Diptera, Sphaeroceridae). Entomologische Abhandlungen, Staatliches Museum fur Tierkunde in Dresden, 46, 1 - 44.", "Marshall, S. A. (1997) Sphaerocerid flies (Diptera: Sphaeroceridae) of the Yukon. In: Danks, H. V. &amp; Downes, J. A. (Eds.), Insects of the Yukon. Biological Survey of Canada (Terrestrial Arthropods), Ottawa, pp. 663 - 685.", "Rohacek, J., Marshall, S. A., Norrbom, A. L., Buck, M., Quiros, D. I. &amp; Smith, I. (2001) World Catalog of Sphaeroceridae (Diptera). Slezske Zemske Muzeum, Opava, 414 pp. (PDF version available at: www. uoguelph. ca / debu / catalog. htm)"]}