Mecodema haunoho Seldon & Leschen 2011
Creators: name: Seldon, David S.
name: Buckley, Thomas R.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Mecodema
Mecodema haunoho

description: Mecodema haunoho Seldon & Leschen, 2011.
Figures 4, 15B and 30.
Diagnosis: Distinguished from other North Island Mecodema species by having: 1, the prothoracic carina narrow the entire length, moderately crenulated with 3 setae along each side (curviden s species group); 2, distinctive shape of the penis lobe (Fig. 30 PL).
Description: Length 20.5–24.1 mm, pronotal width 5.1–5.9 mm, elytral width 6.3–6.8 mm. Colour of entire body matte to glossy black.
Head: Narrow and flat (Fig. 14C). Vertex with very fine wrinkles forming an isodiametric pattern (may be indistinct); vertexal groove narrow and well-defined the entire length; small supraorbital puncture bearing 2–3 setae; 1–2 well-defined supraorbital grooves (especially the one that encompasses the supraorbital puncture); frons with a large and shallow depression each side of midline (Fig. 30); frontoclypeal suture well-defined, tentorial pits absent; anterior area of clypeus (Fig. 9) with shallow and narrow grooves, very faint in some specimens; clypeus with 1 setose puncture each side bearing 2 setae. Labrum rectangular, anterior edge outwardly curved with 2 proximate central setae, 2 setae each side evenly spaced. Mentum lobes squared (Fig. 12G), median process short and narrow, slightly angled upward (15°), notched indentation (Fig. 12D); mentum setae variably present or absent. Submentum sclerite constriction narrow with 6–8 setae, medial setae pair distantly spaced, 2 lateral setae pairs anterad the others. Stipes with 2 basal setae. Gula pits (Fig. 8) large, suture poorly defined, gula flat and smooth. Gena with fine wrinkles forming an isodiametric pattern across entire area.
Prothorax: Prothoracic carina narrow the entire length, moderately crenulated with 3–4 setae along each side (Fig. 30), extended beyond anterior angle; posterior lateral sinuation evidently carinate, slightly angled inward; pronotum broad and laterally deflected, overall shape squared; midline weakly impressed, medial impressions absent, disc with very fine wrinkles laterally; pronotal foveae shallow and narrow; anterior edge slightly inwardly curved, posterior edge slightly inwardly curved medially. Prosternum (Fig. 2) flat and smooth; proepisternum with indistinct wrinkles. Procoxal setae absent; protibia distally expanded and shovel-like (Fig. 10B).
Elytra: Narrow and slightly deflected; humeral angle evenly convex (Fig. 6B), basal margin slightly curved and bevelled to base, interval 1 extended to margin; lateral carina narrow the entire length, extended to humeral angle; humerus with 2 setose punctures within angle (Fig. 30); suture impressed but no more so than striae; striae with small, regularly spaced asetose punctures; intervals 1–4 slightly convex, 5–8 moderately convex, interval 9 strongly convex; interval microsculpture absent; 7 th strial setal pattern with 0–1 setose puncture in anterior ½, 4 setose punctures in posterior ½ (Fig. 30), setose punctures small.
Ventral surface: Mesepisternum and metepisternum with corrugose wrinkles; setose punctures of mesocoxae (Fig. 2) and metacoxae absent. All abdominal ventrites finely lineate; ventrites 3–5 setose punctures absent each side of midline; ventrite 6 setae present: ♂ with 2 setose punctures each side of straight portion of apical edge, ♀ with 2 setose punctures each side, distantly spaced at edge of bluntly rounded apical edge; lateral foveae absent. Anterior metaventrite process a short rounded triangle with a very broad (apically) and distinctly defined carina.
Male genitalia: Apical portion of penis lobe symmetrically rounded with a moderate deflection to right of vertical axis (VV) (Fig. 15B), dorsal process slightly more flattened than ventral apical curve (Fig. 30 PL); overall length of penis lobe shaft with curve to right absent (VV); overall length of penis lobe shaft short and broad, moderately curved ventrally (LV) (Fig. 16B). Structures of the endophallus (Fig. 4): lateral form of the apex of the central spicule pointed; dorsal form of the apex of the central spicule short and narrow; setal (scales) coverage of the apical plate moderate (26–75%); size of left setose flange small; size of right setose flange large. Left paramere basal lobe a very slender rectangle (only slightly broader than right paramere), short slope to arm (Fig. 30 LP); arm relatively long and broad, gradually narrowed to terminal lobe; short and narrow terminal lobe with a tuft of medium length setae (Fig. 30 LP); sparse shorter setae extend along apical ¾ of ventral edge; ventral edge straight. Right paramere narrow and triangular with setae extending along apical ¾ of ventral edge (Fig. 30 RP).
Female genitalia: Basal gonocoxite 1 short and broad, apically flattened with a few grooves present on ventral surface, internal dorso-lateral carina with 1 seta. Gonocoxite 2 (Fig. 5) a short, broadly rounded triangle, apical edge strongly reflexed. Ramus long and relatively broad.
Comments: Mecodema haunoho is endemic to Te Hauturu-o-Toi/Little Barrier Island, and its very close sister taxa, M. aoteanoho is endemic to Aotea.
Distribution: New Zealand, North Island, Coromandel, Te Hauturu-o-Toi/Little Barrier Island (only).
Holotype: NZAC, male labelled: NEW ZEALAND CL, Little Barrier Island, Tirikakawa Stream, 2005, D.S. Seldon ♂ / HOLOTYPE Mecodema haunoho n. sp. design. by Seldon & Leschen 2010 [red label].
Paratypes: (see Seldon & Leschen 2011).
Material examined: 1♀, NEW ZEALAND CL, Little Barrier Is., Awaroa Stream PTs 22.11.06 – 22.1.07, D.S. Seldon / NZAC04127983.
description: Published as part of Seldon, David S. & Buckley, Thomas R., 2019, The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand, pp. 1-148 in Zootaxa 4598 (1) on pages 53-55, DOI: 10.11646/zootaxa.4598.1.1, http://zenodo.org/record/2668063