Clivina circumreticulata Balkenohl, 2025, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina circumreticulata

description: Clivina circumreticulata sp. nov.
(Figs 25, 60, 93, 119)
Type material. Holotype: ♂, with labels and data: white, black printed “ BORNEO – SABAH 1995 Ban jaran Mattland Balu Pungul 25.-27.5. Ivo Jenis leg.” / “ Clivina dealata DARL. det. Balkenohl X. 97” (CMBB).
Paratypes: 5 ♂♂, 3 ♀♀, “BRUNEI-Temburong Belalong Field Res. C. Mix. Diperocarp For. ~ 100m, 2-8.5.95 Heiss” (CADW); 1 ♂, 1 ♀, “ BORNEO BRUNEI Temburong Kuala Belalong R. Borcherding leg.” / “VI / VII 1995 KBFSC” / “ Clivina wallacei PUTZ. det. M. Baehr’ 12” / “COLL. WRASE BERLIN” / “COLL. WRASE GUSOW-PLATKOW GERM.” (CDWGP); 1 ♀, “ SARAWAK Kapit dist. Rumah Ugap vill., Sut riv., 3.- 9.3.1994 J. Horák leg.” (CADW); 1 ♂, “ BORNEO: SARAWAK Nanga Pelagus nr. Kapit 180-585m, 7-14.VIII. ‘58” / “ T. C. Maa Collector ” / “ Clivina (s.str.) helferi group P.Bulirsch det.2015” (NHMUK).
Remark. In one of the paratypes (from NHMUK), the right front and hind leg, the tarsomeres of the right intermediate leg, two apical tarsomeres of the left front leg, and two right apical antennomeres are missing. The specimen was pinned and shows considerable damages on the ventral side.
Description. Measurements. Body length 5.17–6.38 mm (ẍ = 5.76 mm *), width 1.5–1.82 mm (ẍ = 1.61 mm *), ratio L/W of pronotum 0.86–0.91 (ẍ = 0.88*), ratio L/W of elytra 1.84–1.95 (ẍ = 1.91*); (*n = 10). Colour glossy, piceous. Supraantennal plate slightly translucent. Antennae, mouthparts, intermediate and hind legs yellowish.
Head a fifth narrower than pronotum. Clypeus with central part slightly concave, with slightly projecting rounded teeth laterally, separated from clypeal-wing by small notch; wing wider than teeth of clypeus, rounded, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin laterally convex, more distinct at middle, clypeus and wing indistinctly reflexed margined. Supraantennal plate extended posteriorly into an obtuse keel, separated from wing by flat furrow. Clypeus moderately convex, with scattered small punctures, clypeal-furrow indistinct, flattened. Frons slightly to moderately convex, with indistinct flat pore at middle, with scattered small punctures, separated from supraantennal plate and supraorbital carina by distinct furrow, with clypeal seta in front; supraorbital carinae moderately developed, with two supraorbital setigerous punctures at middle and end of eye level. Neck constriction nearly closed, formed by a step from frons and punctures posteriorly. Eye well developed, gena small but distinct. Antenna elongate, just reaching up to middle of pedunculus. Labrum excised anteriorly, finely isodiametrically reticulated, seven-setose. Mentum with irregularly askew reticulated surface, lobe slightly hollowed out, laterally reflexed margined, laterally convex, medially straight, apex acute, median tooth acute, hollowed out apically, distinctly lower than lobes, clefts moderately developed.
Pronotum (Fig. 60) with disk slightly convex in lateral view, moderately convex in frontal view. Shape square to cordate, wider than long. Reflexed lateral margin smooth, slightly convex, widest at middle; anterior angle distinctly marked, posterior angle indistinct, marked by tooth; lateral channel narrow, indistinctly reticulated, continuing up to base as narrow, indistinctly convex margin. Median line complete, engraved at middle, narrow at extremities, joining anterior transverse line. Anterior transverse line distinct, complete. Disk completely covered with medium sized punctures, with some transverse wrinkles, with isodiametric reticulation at base, with posterior setigerous puncture removed from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 35°. Basal channel on flange regularly developed, somewhat rugose, with sharp carina.
Elytron (Fig. 25) with disk distinctly flattened in lateral view, moderately convex in frontal view with laterally increasing convexity. Shape of elytra elongate, slender, around two-fifth longer than wide, with lateral margin indistinctly convex at middle but slightly diverging, maximum width posterior middle. Humerus rounded but distinct. Reflexed lateral margin indistinctly crenulate behind humerus (120-fold). Row of setigerous punctures in the lateral channel regular in size and distance, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct small tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, impunctate, one to four free at base, five and six joining at humerus, one to four ending free at apex, five and six joining apically. Intervals slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven with obtuse step at apex, eight carinate at apex. Surface of interval one to four glossy at middle, all other intervals with distinct isodiametric reticulation, apically nearly half of elytron reticulated as well as base.
Hind wing fully developed.
Lower surface. Proepisternum isodiametrically reticulated, with few fine wrinkles laterally. Sternite of abdomen with isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with moderately wide projection posteriorly at middle where it is transversely flattened, with narrow marginal furrow at middle (in both sexes). Legs. Protibia slender, with fine longitudinal reticulation, with slight sulcus dorsally, with three slender acute spines and obtuse tooth, movable spur acute, distinctly curved, basal tarsomere slender at base, carinate, with acute tubercle and three setae at middle of tarsomere. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg widened.
Male genitalia (Fig. 93). Median lobe slender, distinctly arcuate at basal part and at middle, nearly straight to apex, apically flattened-oval in dorsal view, duck-bill like in lateral view. Parameres moderately slender, with moderate apophyses, slightly distorted, hyaline apically, each with one apical seta.
Female gonocoxites and epipleurite (Fig. 119). Basal gonocoxite with two long and three medium sized setae, apical gonocoxite elongated, nearly cylindrical in cross section, with one long and one short seta at base and one long seta at beginning of apical third, with two very long setae apically. Epipleurite sub-trapezoid shaped, with one long seta laterally.
Variation. In one of the paratypes, the lateral margin of the pronotum is more distinctly convex and the tooth of the posterior angle is more distinctly developed.
Distribution. Known from the island of Borneo (Sabah, Sarawak, Brunei).
Diagnosis. A small to medium sized piceous species with square to cordate pronotum and intervals one to four smooth and glossy on the disk of the elytron. The most similar species with smooth and glossy intervals one to four of the elytron and a similar body size is C. circumopaca sp. nov. from Sulawesi. It differs by the straight labrum which is in addition six-setose. Moreover, the pronotum is distinctly trapezoid-shaped and the elytra are more convex.
Etymology. The name refers to the elytra with its reticulation on the intervals basally, apically, and laterally (Latin: circum = surrounded by; reticulum = small sized net).
description: Published as part of Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), pp. 1-70 in Zootaxa 5592 (1) on pages 41-42, DOI: 10.11646/zootaxa.5592.1.1, http://zenodo.org/record/14953674