cometadata/datacite-rtg-text-other-reclassification-records huggingface dataset:
Bathyglycinde Fauchald 1972
Creators: name: Meissner, Karin
name: Schwentner, Martin
name: Göưing, Miriam
name: Fiege, Thomas Knebelsberger and Dieter
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Polychaeta
Phyllodocida
Goniadidae
Bathyglycinde

description: Genus Bathyglycinde Fauchald, 1972
Type species: Bathyglycinde mexicana Fauchald, 1972. Gender: female.
Diagnosis: (See: Böggemann 2022.) Segments uni- or biannulate. Prostomium smooth or indistinctly annulated; tip with biarticulate appendages. Proboscis with several different types of papillae, arranged in distinct longitudinal rows; macrognaths and dorsal micrognaths present; chevrons absent. First segment sometimes with only a pair of small lateral cirri and without parapodia or chaetae. Anterior part of body with uniramous parapodia, following region with biramous parapodia. Notochaetae capillaries; neurochaetae compound spinigers.
description: Published as part of Meissner, Karin, Schwentner, Martin, Göưing, Miriam & Fiege, Thomas Knebelsberger and Dieter, 2023, Polychaetes distributed across oceans-examples of widely recorded species from abyssal depths of the Atlantic and Pacific Oceans, pp. 906-944 in Zoological Journal of the Linnean Society 199 on page 920, DOI: 10.1093/zoolinnean/zlad069, http://zenodo.org/record/10470369
Bathyglycinde sibogana Boggemann 2005
Creators: name: Meissner, Karin
name: Schwentner, Martin
name: Göưing, Miriam
name: Fiege, Thomas Knebelsberger and Dieter
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Polychaeta
Phyllodocida
Goniadidae
Bathyglycinde
Bathyglycinde sibogana

description: Bathyglycinde sibogana (Augener & Peflibone in Peflibone, 1970)
(Fig. 10A, B)
Glycinde sibogana Augener and Peưibone in Peưibone, 1970: 244, figs 40, 41.
Bathyglycinde sibogana Böggemann, 2005: 196–198, figs 115, 116. – Böggemann 2009: 311, figs 43, 44, 47. – Fiege et al. 2010: appendix, tables 3 and 5 (name only). – Moreira and Parapar 2015: 61–62, fig. 25A–M. – Böggemann and Dietz 2016: 1508– 1509, fig. 1.
Material examined: Central Atlantic Ocean, Mid-Atlantic Ridge, East, SO 237 (VEMA-Transit), station 4-8, 5735 m (SMF 30491, SEM 1340, SMF 30492, ZMH-P 29694). – for details and additional specimens see the Supporting Information, Table S2.
Measurements for largest specimen studied (ZMH-P 29694): length 14.5 mm, width 0.5 mm (excl. parapodia), 67 chaetigers, 35 uniramous parapodia.
Description: Prostomium in all three specimens with four irregular indistinct annulations and longitudinal lateral grooves along each side. Posterior chaetigers with two neuropodial prechaetal lobes from chaetiger 36 in specimen ZMH-P 29694 but absent in specimen SMF 30491 with only 37 chaetigers. Proboscidial papillae in area I small conical papillae (Fig. 10A, B; long arrows) and area II with five rows of long, slender partly sclerotized papillae similar in shape to those of B. profunda (Fig. 10A, B; short arrows). Some with protruding bulge in lower part (Fig. 10A, B; asterisk).
Remarks: Bathyglycinde sibogana was originally described from bathyal depths of 1886 m in the Gulf of Boni (Indonesia) (Augener and Peưibone in Peưibone 1970) and has been reported to occur in the Central and SE Pacific (Böggemann 2005) and the Central and SE Atlantic (Böggemann 2009; this study) as well as the Atlantic Sector of the Southern Ocean (Böggemann and Dietz 2016). The three specimens studied agree well with the description published by Böggemann 2005. Molecular data confirm and extend the distribution of Bathyglycinde sibogana to the Atlantic (Cape Basin (s. Böggemann 2009) and the central Atlantic) (Fig. 2).
Distribution: Indo-Pacific, Gulf of Boni (1886 m, Augener and Peưibone in Peưibone 1970), Central Pacific [4978 m, Böggemann 2005], SE Pacific [3739–3824 m, Böggemann 2005], Central Atlantic (Vema Fracture Zone: 5735 m, this study), South Atlantic [Cape and Angola Basin: 5058–5648 m, Böggemann (2009)], and Scotia Sea [3789–4720 m, Böggemann and Dietz (2016)].
description: Published as part of Meissner, Karin, Schwentner, Martin, Göưing, Miriam & Fiege, Thomas Knebelsberger and Dieter, 2023, Polychaetes distributed across oceans-examples of widely recorded species from abyssal depths of the Atlantic and Pacific Oceans, pp. 906-944 in Zoological Journal of the Linnean Society 199 on page 924, DOI: 10.1093/zoolinnean/zlad069, http://zenodo.org/record/10470369
Progoniada Hartman 1965
Creators: name: Meissner, Karin
name: Schwentner, Martin
name: Göưing, Miriam
name: Fiege, Thomas Knebelsberger and Dieter
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Annelida
Polychaeta
Phyllodocida
Goniadidae
Progoniada

description: Genus Progoniada Hartman, 1965
Type species: Progoniada regularis Hartman, 1965. Gender: female.
Diagnosis: (See: Böggemann 2022.) Segments bi- or triannulate. Prostomium annulated, consisting of eight rings; tip with biarticulate appendages. Proboscis with a few different types of papillae; with macrognaths and dorsal and ventral micrognaths; chevrons present. First segment with only a pair of small lateral cirri and without parapodia or chaetae. All parapodia uniramous. Notochaetae absent; neurochaetae compound falcigers and/or spinigers in all parapodia.
description: Published as part of Meissner, Karin, Schwentner, Martin, Göưing, Miriam & Fiege, Thomas Knebelsberger and Dieter, 2023, Polychaetes distributed across oceans-examples of widely recorded species from abyssal depths of the Atlantic and Pacific Oceans, pp. 906-944 in Zoological Journal of the Linnean Society 199 on page 924, DOI: 10.1093/zoolinnean/zlad069, http://zenodo.org/record/10470369
Scirtes japonicus Kiesenwetter 1874
Creators: name: Watts, Chris H. S.
name: Cooper, Steven J. B.
name: Saint, Kathleen M.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scirtidae
Scirtes
Scirtes japonicus

description: Scirtes japonicus species group (Nyholm 2002)
Nyholm (2002) based this group primarily on the form of the penis: “symmetric with parameroids longitudinally deeply cleft and tegmen with a lateral lobe (paramere) on each side”. We would add: first segment of labial palpus shorter than second, mandible incisivus moderately long, caudal margin of metacoxal plate concave and caudal margin of sternite 7 moderately concave; larvae with tip of mandible bifid and maxillary palpus threesegmented (Watts 2004; Yoshitomi 2005). The prehensors and bursal sclerites of all three Australian species (Figs 3–5) are closely similar to those of S. japonicus (Yoshitomi 2005).
The penis of no Australian species has this morphology. The aedeagus of S. emmaae is simple and appears plesiomorphic as it conforms closely to the scirtid ground plan (Zwick 2016). Scirtes kaytae and S. tindaleensis have asymmetric aedeagi resembling those of species in the S. helmsi group.
The aedeagus of S. emmaae, particularly the apical triangular lobes of the parameres, is similar to those of S. micronesianus Yoshitomi, which was placed in a separate group by Yoshitomi (2009) and named the S. micronesianus group. However, other than their aedeagi, S. emmaae and S. kaytae are morphologically and genetically close and S. tindaleensis is also close on both larval and adult morphology yet only S. emmaae could be classified in the S. micronesianus group. We think that on this evidence these three species belong in the same species group and have placed all three Australian species in the S. japonicus group based on sequence data and adult and larval morphology but recognise that future study is needed to confirm this placement, particularly in light of the conflicting morphology of the male genitalia.
Scirtes nehouensis Ruta & Yoshitomi, 2010 from New Caledonia appears to us on both general morphology and male and female genitalia (Ruta & Yoshitomi 2010b) to be conspecific with S. emmaae. We herein place the species in synonymy under Scirtes emmaae Watts, 2004.
description: Published as part of Watts, Chris H. S., Cooper, Steven J. B. & Saint, Kathleen M., 2017, Review of Australian Scirtes Illiger, Ora Clark and Exochomoscirtes Pic (Coleoptera: Scirtidae) including descriptions of new species, new groups and a multi-gene molecular phylogeny of Australian and non-Australian species, pp. 511-532 in Zootaxa 4347 (3) on pages 518-519, DOI: 10.11646/zootaxa.4347.3.5, http://zenodo.org/record/1048629
Scirtes haemisphaericus
Creators: name: Watts, Chris H. S.
name: Cooper, Steven J. B.
name: Saint, Kathleen M.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scirtidae
Scirtes
Scirtes haemisphaericus

description: Scirtes haemisphaericus species group (Yoshitomi 2005)
Yoshitomi (2005) defined this group as having the “body oval, parameres simple, penis symmetrical with a long slender pala (basal piece)”. We add: mandibles blunt, first segment of labial palpus shorter than second segment, caudal margin of metacoxal plates weakly concave, metatrochanters elongate (about 2.5× as long as wide), caudal margin of sternite 7 deeply concave; females with relatively short coxites and broad short styles and a moderately sized, compact, well sclerotized prehensor and bursal sclerite (Fig. 10; Yoshitomi 2005); larvae with a pronounced spindle-shape with multidentate mandibles, maxillary palpus three-segmented, anal palpi>5 (Klausnitzer 2009; Watts 2014).
Both the sequence data and adult and larval morphology support a relationship of the single Australian species, S. auratus, with the European S. haemisphaericus, the type species of the genus Scirtes. The Japanese S. sobrinus Lewis clearly also belongs in this group on larval characters, prehensor and bursal sclerite although less clearly on the male genitalia (Yoshitomi 2005). The sequence data also placed two closely related species from Vietnam, each represented by a single male, in this grouping (Fig. 1). The male genitalia of both are similar and closely resemble the illustration of the male genitalia of S. sobrinus in Yoshitomi (2005).
description: Published as part of Watts, Chris H. S., Cooper, Steven J. B. & Saint, Kathleen M., 2017, Review of Australian Scirtes Illiger, Ora Clark and Exochomoscirtes Pic (Coleoptera: Scirtidae) including descriptions of new species, new groups and a multi-gene molecular phylogeny of Australian and non-Australian species, pp. 511-532 in Zootaxa 4347 (3) on page 519, DOI: 10.11646/zootaxa.4347.3.5, http://zenodo.org/record/1048629
Scirtes elegans
Creators: name: Watts, Chris H. S.
name: Cooper, Steven J. B.
name: Saint, Kathleen M.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scirtidae
Scirtes
Scirtes elegans

description: Scirtes elegans species group (Yoshitomi 2009)
Yoshitomi (2009) defined the S. elegans group as “having the tegmen connected to an asymmetrical penis”. We add: first segment of labial palpus as long as second, mandible incisivus long, metatrochanter elongate, caudal margin of sternite 7 moderately concave. The relationship between tegmen and penis perhaps could be described as having the elements of each complexly intermingled (Fig. 21). Within the group the female of only one species is known—that of the only Australian species, S. albamaculatus, from North Queensland. Assuming that the female of S. albamaculatus is representative of the group we could also add: members are dark coloured with white/cream macula or stripes on the elytra and females with elytral excitators (Fig. 31) and a ring-like bursal sclerite (Fig. 6). No larvae have been described.
The Scirtes elegans group as presently defined has only three species, S. elegans from Malaysia, S. albamaculatus from Australia and S. albotaeniatus Yoshitomi from Guam.
The sequence data place S. albamaculatus as unique within Australian Scirtes. However, it also strongly links S. albamaculatus to S. teruhisai Yoshitomi & Ruta from Borneo (Fig. 1) which is the only representative of the species-rich Southeast Asian S. flavoguttatus group (Yoshitomi & Ruta 2010; Ruta et al. 2014) included in our study. This placement is not unexpected as a number of species currently placed in this group have female excitators, pale colour patterns on the elytra, all have fused tegmen and penis and some, e.g. S. sarawakensis Ruta & Yoshitomi, have an aedeagus approaching the complexity seen in S. albamaculatus. The form of the mandible and labial palpus also point to membership of the S. flavoguttatus group. With the discovery of additional species in the S. flavoguttatus group in Southeast Asia (Ruta et al. 2014) the distinction between these two groups has become rather tenuous and relies on the asymmetric rather than symmetric penis. The strong linkage of S. albamaculatus and S. teruhisai by the sequence data suggests that the two species belong in the same species group. However, with only one species of each group available to us we feel it is premature to formally combine these species groups on the results of this study.
description: Published as part of Watts, Chris H. S., Cooper, Steven J. B. & Saint, Kathleen M., 2017, Review of Australian Scirtes Illiger, Ora Clark and Exochomoscirtes Pic (Coleoptera: Scirtidae) including descriptions of new species, new groups and a multi-gene molecular phylogeny of Australian and non-Australian species, pp. 511-532 in Zootaxa 4347 (3) on page 519, DOI: 10.11646/zootaxa.4347.3.5, http://zenodo.org/record/1048629
Scirtes helmsi Blackburn 1892
Creators: name: Watts, Chris H. S.
name: Cooper, Steven J. B.
name: Saint, Kathleen M.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scirtidae
Scirtes
Scirtes helmsi

description: Scirtes helmsi species group (Watts 2004)
Defined here as: broadly oval; mandible incisivus relatively short, first segment of labial palpus shorter than second, caudal margin of metacoxal plates straight or nearly so, caudal margin of sternite 7 truncated rather than concave, metatrochanters elongate-oval, 2.0× as long as wide, aedeagus asymmetrical, pala well developed, one or two parameroids, typically the right hand one longer and with an apical hook; tegmen variable, usually simple, parameres simple, usually long and thin but sometimes broader and tending to wrap around the penis but are never fused with it (Watts 2004) (Figs 18–20); female coxites and styles elongate, prehensor moderately developed, occasionally absent, basal sclerite typically pelican-bill-like but occasionally ring-shaped with internal prongs or bullet-shaped with internal spines (Figs 7–16). Larva with maxillary palpus four-segmented, five anal papillae and mandible with single-pointed incisivus.
Most of the species of Australian Scirtes studied fell into this group. The sequence data identified two major subgroupings with some internal substructure within each. There is no correlation between the different female prehensors and bursal sclerites and the sequence data. The male terminalia of S. zwicki are quite unlike those of other species in the group (and from any other known Scirtes species), having elongate, symmetrical genitalia and a greatly modified sternite 9. Sternite 8 appears to be missing (Fig. 23). Genetically it is unequivocally within the S. helmsi subgroup (Fig. 1).
There are 27 described species in the Scirtes helmsi group in Australia (Watts 2004 & this paper). Yoshitomi & Ruta (2010) noted the close similarity of 11 New Caledonian species with Australian species. From the illustrations of the male genitalia, S. babeldaobensis Yoshitomi from Micronesia in Yoshitomi (2009) and possibly S. zerchei Klausnitzer from Luzon Island may also belong in the S. helmsi group.
description: Published as part of Watts, Chris H. S., Cooper, Steven J. B. & Saint, Kathleen M., 2017, Review of Australian Scirtes Illiger, Ora Clark and Exochomoscirtes Pic (Coleoptera: Scirtidae) including descriptions of new species, new groups and a multi-gene molecular phylogeny of Australian and non-Australian species, pp. 511-532 in Zootaxa 4347 (3) on page 520, DOI: 10.11646/zootaxa.4347.3.5, http://zenodo.org/record/1048629
Ora improtecta Watts 2004
Creators: name: Watts, Chris H. S.
name: Cooper, Steven J. B.
name: Saint, Kathleen M.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scirtidae
Ora
Ora improtecta

description: Ora improtecta Watts, 2004
Notes. This species remains little known and no specimens were available for our DNA study. Morphologically it is distinct from the two other Australian species in the genus, which appear to be more typical of the genus, and its position in Ora is problematic. In this context it needs to be recognized that Ora is a poorly characterized cosmopolitan genus much in need of a revision. There is a recent specimen in the South Australian Museum’s collection from Halmahera, Indonesia. It is thus probable that the species is widespread in the islands to the north of Australia.
description: Published as part of Watts, Chris H. S., Cooper, Steven J. B. & Saint, Kathleen M., 2017, Review of Australian Scirtes Illiger, Ora Clark and Exochomoscirtes Pic (Coleoptera: Scirtidae) including descriptions of new species, new groups and a multi-gene molecular phylogeny of Australian and non-Australian species, pp. 511-532 in Zootaxa 4347 (3) on page 520, DOI: 10.11646/zootaxa.4347.3.5, http://zenodo.org/record/1048629
Scirtes albamaculatus Watts. However 2004
Creators: name: Watts, Chris H. S.
name: Cooper, Steven J. B.
name: Saint, Kathleen M.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scirtidae
Scirtes
Scirtes albamaculatus

description: Scirtes albamaculatus Watts, 2004
(Figs 6, 21, 25)
Supplementary description. Male. Tergite 8: transverse, apical portion with a few small setae, apodemes strongly sclerotized, about twice as long as apical portion, diverging basally. Tergite 9: apical portion relative large, moderately long seta on apical margin; apodemes strongly sclerotized, about as long as apical portion, diverging basically. Sternite 9 small, broadly bilobed, apex with a few long setae, apodemes short, merging. Aedeagus complex, asymmetric, basal piece greatly reduced, trigonium relatively broad, weakly sinuate, apex pointed, one edge with well-developed setae; parameroids absent (possibly represented by irregular projections on anterior edge of basal piece). Tegmen well developed, parameres well developed, long, thin, robust, and connected at their bases by a narrow stirrup-like bridge which partially surrounds the penis, one side of bridge fused to penis (Fig. 21).
Female. Apices of elytra with a transverse, bare, depression surrounded by slightly longer setae (excitator of Ruta 2008) (Fig. 25). Coxites elongate, styli small, oval, apodemes long, simple, prehensor weakly sclerotised broad with two thin serrated lateral ridges, bursal sclerite moderately sclerotized, signet-ring-shaped with or without an internal prong (Fig. 6).
Specimens examined. 6, ‘Eubenangee Swamp, Queensland, at light, 13/3/13, CHS Watts’. Notes. A relatively rare species so far known only from coastal swamps of North Queensland. Larva not known.
description: Published as part of Watts, Chris H. S., Cooper, Steven J. B. & Saint, Kathleen M., 2017, Review of Australian Scirtes Illiger, Ora Clark and Exochomoscirtes Pic (Coleoptera: Scirtidae) including descriptions of new species, new groups and a multi-gene molecular phylogeny of Australian and non-Australian species, pp. 511-532 in Zootaxa 4347 (3) on pages 520-522, DOI: 10.11646/zootaxa.4347.3.5, http://zenodo.org/record/1048629
Scirtes auratus Watts 2004
Creators: name: Watts, Chris H. S.
name: Cooper, Steven J. B.
name: Saint, Kathleen M.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scirtidae
Scirtes
Scirtes auratus

description: Scirtes auratus Watts, 2004
(Figs 2, 17)
Supplementary description. Male. Tergite 8: small, apical portion very weakly sclerotized, hind edge with dense row of extremely small fine setae or microtrichia. Apodemes relatively short, strongly sclerotized, parallel. Tergite 9: short, broad, apical portion short with apical margin with long setae, apodemes strongly sclerotized, about three times length of apical portion, weakly diverging. Sternite 9: small, weakly sclerotized, broadly bilobed, apical third with numerous pores and thin setae, without obvious apodemes. Aedeagus complex, symmetrical, basal piece large, expanding anteriorly; trigonium long, thin, with complex set of huge curved spines at base, apex with a few small stout spines; parameroids in two parts, inner lobe longer than trigonium, broad in basal half, narrow in apical half, bulbous at apex; outer lobe broad, flat, about half as long as inner lobe numerous pores and spines on outer margin and towards apex. Tegmen consists of two separate narrow rods, about as long as aedeagus, central portions covered with short strong spines, apical portion with outer spur of varying length (Fig. 17).
Female. Coxites elongate, styles relatively short and broad. Tergite 9: long, with spurs; tergite 8 long; tergite 8 shorter, apex with thick row of moderately long setae. Prehensor moderately sclerotized, long, apical end with two triangular plates. Bursal sclerite elongate, strongly sclerotized, triangular apical end with two strong prongs, surface strongly pimpled (Fig. 2).
Notes. The species is found in coastal swamps from the Northern Territory around the northern coast to at least Townsville and has been collected in large numbers at light. The larvae are found in shallow grassy swamps (Watts 2014). It is not known if the pupae of the species use a snorkel as described by Zwick & Zwick (2008) for S. haemisphaericus. In the laboratory without access to plant stems the larvae form pupal cells in damp sand or pupate on the undersides of leaves.
description: Published as part of Watts, Chris H. S., Cooper, Steven J. B. & Saint, Kathleen M., 2017, Review of Australian Scirtes Illiger, Ora Clark and Exochomoscirtes Pic (Coleoptera: Scirtidae) including descriptions of new species, new groups and a multi-gene molecular phylogeny of Australian and non-Australian species, pp. 511-532 in Zootaxa 4347 (3) on page 523, DOI: 10.11646/zootaxa.4347.3.5, http://zenodo.org/record/1048629
Scirtes brisbanensis Pic 1956
Creators: name: Watts, Chris H. S.
name: Cooper, Steven J. B.
name: Saint, Kathleen M.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scirtidae
Scirtes
Scirtes brisbanensis

description: Scirtes brisbanensis Pic, 1956 / Scirtes triangularis Watts, 2004
Notes. In Cooper et al. (2014) it was noted that two closely related taxa, each represented by a single female and both identified as Scirtes brisbanensis, were present at Mt Molloy in North Queensland. With access to additional specimens, including males, the present study shows that one specimen was S. brisbanensis and the other S. triangularis (Fig. 1).
Larva. A single larva of S. triangularis is now known, identified by DNA sequencing. In the key to the larvae of Australian Scirtes in Watts (2014) it runs to the S. helmsi / S. orientalis couplet. However, it is smaller than S. helmsi (head width 0.9 mm vs 1.2 mm) and with fewer setae on tergite 1 (8 vs 30–40). From the similar sized S. orientalis it differs in having a more uneven row of palisade setae on the clypeolabrum (Fig. 28) and eight rather than nine claw teeth but is otherwise very similar.
description: Published as part of Watts, Chris H. S., Cooper, Steven J. B. & Saint, Kathleen M., 2017, Review of Australian Scirtes Illiger, Ora Clark and Exochomoscirtes Pic (Coleoptera: Scirtidae) including descriptions of new species, new groups and a multi-gene molecular phylogeny of Australian and non-Australian species, pp. 511-532 in Zootaxa 4347 (3) on page 526, DOI: 10.11646/zootaxa.4347.3.5, http://zenodo.org/record/1048629
Scirtes pinjarraensis Watts 2004
Creators: name: Watts, Chris H. S.
name: Cooper, Steven J. B.
name: Saint, Kathleen M.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scirtidae
Scirtes
Scirtes pinjarraensis

description: Scirtes pinjarraensis Watts, 2004
= Scirtes pygmaeus Watts, 2004, syn. nov.
Notes. Cooper et al. (2014), using sequence data, suggested that Scirtes pinjarraensis and S. pygmaeus Watts were probably conspecific. Examination of further specimens of both taxa confirms this. Watts (2004) separated the two taxa by the presence of a distinctly bent tip to the trigonium in S. pygmaeus which was absent in S. pinjarraensis. Examination of additional specimens of both taxa shows that this bent tip is actually a distinct spine projecting at right angles to the trigonium and surrounded by a number of much smaller spines (Fig. 22). Dissection of more paratypes of S. pinjarraensis showed that a number had identical structures and that the absence of a spine was caused by it being broken off, probably during dissection. We consider, on both morphological and sequence grounds, that S, pinjarraensis and S. pygmaeus are conspecific. Therefore, S. pygmaeus on page preference becomes a junior synonym of S. pinjarraensis.
This terminal structure on the trigonium is unique within Australian Scirtes and readily identifies the species, which is often the commonest scirtid in swamps in southwest Australia. Other than by the male genitalia, the species is recognized from other species in southwest Australia by its small size, dark testaceous colour with pale antennal base and pale sides to the pronotum.
In size, colour and male genitalia S. pinjarraensis resembles S. lynnae, but differs from this species in the hooked left parameroid, presence of a small right parameroid and spines at the apex of the trigonium.
Sequence data place it close to S. helmsi and S. brisbanensis (Fig. 1).
description: Published as part of Watts, Chris H. S., Cooper, Steven J. B. & Saint, Kathleen M., 2017, Review of Australian Scirtes Illiger, Ora Clark and Exochomoscirtes Pic (Coleoptera: Scirtidae) including descriptions of new species, new groups and a multi-gene molecular phylogeny of Australian and non-Australian species, pp. 511-532 in Zootaxa 4347 (3) on page 526, DOI: 10.11646/zootaxa.4347.3.5, http://zenodo.org/record/1048629
Scirtes rutai Watts, Cooper & Saint, 2017, nom. nov.
Creators: name: Watts, Chris H. S.
name: Cooper, Steven J. B.
name: Saint, Kathleen M.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scirtidae
Scirtes
Scirtes rutai

description: Scirtes rutai nom. nov.
= Scirtes beccus Ruta, Kiałka & Yoshitomi, 2014
Notes. Scirtes beccus Ruta, Kiałka & Yoshitomi, 2014 from Kinabalu Park, Ranu, Sabah is preoccupied by Scirtes beccus Watts, 2004. We propose the replacement name, Scirtes rutai, after Rafał Ruta of Wroclaw in recognition of his extensive and important work on the Scirtes of Southeast Asia.
description: Published as part of Watts, Chris H. S., Cooper, Steven J. B. & Saint, Kathleen M., 2017, Review of Australian Scirtes Illiger, Ora Clark and Exochomoscirtes Pic (Coleoptera: Scirtidae) including descriptions of new species, new groups and a multi-gene molecular phylogeny of Australian and non-Australian species, pp. 511-532 in Zootaxa 4347 (3) on page 527, DOI: 10.11646/zootaxa.4347.3.5, http://zenodo.org/record/1048629
Scirtes exoletus Waterhouse 1880
Creators: name: Watts, Chris H. S.
name: Cooper, Steven J. B.
name: Saint, Kathleen M.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scirtidae
Scirtes
Scirtes exoletus

description: Scirtes exoletus Waterhouse, 1880
Notes. A very widespread, quite common species found in all states and mainland territories of Australia. Specimens from the Northern Territory, New South Wales and Victoria were identical in the genes sequenced; however, the specimen from Wongabel on the Atherton Tableland of North Queensland differed by 7.2% (pdistance) for COI, potentially indicative of a separate species.
description: Published as part of Watts, Chris H. S., Cooper, Steven J. B. & Saint, Kathleen M., 2017, Review of Australian Scirtes Illiger, Ora Clark and Exochomoscirtes Pic (Coleoptera: Scirtidae) including descriptions of new species, new groups and a multi-gene molecular phylogeny of Australian and non-Australian species, pp. 511-532 in Zootaxa 4347 (3) on page 527, DOI: 10.11646/zootaxa.4347.3.5, http://zenodo.org/record/1048629
Zodariellum asiaticum
Creators: name: Zamani, Alireza
name: Marusik, Yuri M.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Zodariidae
Zodariellum
Zodariellum asiaticum

description: Zodariellum asiaticum (Tyshchenko, 1970)
Figs 1A, 2A, 3A, 4A, 5A, 6A, 9A, 12A
Zodarion asiaticum Tyshchenko, in Marikovski & Tyshchenko 1970: 199, fig. 1 (♂ ♀).
Material examined. KAZAKHSTAN: Jambyl Region: 1♂ (ZMMU), Korday Dist., 10 km SE of Korday Vil., Yeradzhy well, [43°02'N 74°42'E], 20.04.1997 (A.V. Gromov).
Description. See Marikovski & Tyshchenko (1970). Habitus of male as in Fig. 1A. Male palp as in Figs 2A, 3A, 4A, 5A, 6A, 9A.
Comments. Previously illustrated only in the original description. The type material was collected from the nest of Polyergus rufescens (Latreille, 1798) (Hymenoptera: Formicidae). Marikovski & Tyshchenko (1970) and Pekár (2009) described its hunting and courtship behaviours.
Distribution. Kazakhstan (type locality; Fig. 12A), Kyrgyzstan (Zonstein et al. 1996) and Tajikistan (Andreeva 1976). The record from Tajikistan is based on a singleton female that is most likely misidentified, and therefore not included in the map.
description: Published as part of Zamani, Alireza & Marusik, Yuri M., 2022, New taxonomic considerations in Zodariellum Andreeva & Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera, pp. 161-177 in Zootaxa 5178 (2) on pages 164-165, DOI: 10.11646/zootaxa.5178.2.3, http://zenodo.org/record/7022237
Zodariellum hunanense Zamani & Marusik 2022, comb. n.
Creators: name: Zamani, Alireza
name: Marusik, Yuri M.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Zodariidae
Zodariellum
Zodariellum hunanense

description: Zodariellum hunanense (Yin, 2012) comb. n.
Fig. 12B
Zodarion hunanensis Yin, in Yin et al. 2012: 1148, figs 611a–e (♀).
Comments. The southernmost species of the genus, currently known only from the original description based on material from southern China (Fig. 12B). Here, it is transferred to Zodariellum due to its screw-like copulatory ducts.
Distribution. Southern China (Hunan) (Fig. 12B).
description: Published as part of Zamani, Alireza & Marusik, Yuri M., 2022, New taxonomic considerations in Zodariellum Andreeva & Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera, pp. 161-177 in Zootaxa 5178 (2) on page 170, DOI: 10.11646/zootaxa.5178.2.3, http://zenodo.org/record/7022237
Zodariellum mongolicum Marusik & Koponen 2001
Creators: name: Zamani, Alireza
name: Marusik, Yuri M.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Zodariidae
Zodariellum
Zodariellum mongolicum

description: Zodariellum mongolicum Marusik & Koponen, 2001
Fig. 12B
Zodariellum mongolicum Marusik & Koponen, 2001: 45, figs 3–5, 8–9, 17–18, 26 (♂).
Zodarion mongolicum: Jocqué & Henrard 2015: 21.
Zodariellum mongolicum: Zamani & Marusik 2021: 185.
Zodarion apertum Zhang & Zhang, 2019: 69, figs 1A–D, 2A–E, 3A–E (♂ ♀). Syn. n.
Comments. Accounting the original descriptions, we hereby synonymize Z. apertum Zhang & Zhang, 2019 syn. n. (described from Xinjiang, northwestern China) with Z. mongolicum (described from Mongolia), as no noticeable differences can be detected between the illustrated male palps (cf. Zhang & Zhang 2019: fig. 2C and Marusik & Koponen 2001: figs 17–18).
Distribution. Mongolia and northwestern China (Xinjiang) (Fig. 12B).
description: Published as part of Zamani, Alireza & Marusik, Yuri M., 2022, New taxonomic considerations in Zodariellum Andreeva & Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera, pp. 161-177 in Zootaxa 5178 (2) on pages 170-171, DOI: 10.11646/zootaxa.5178.2.3, http://zenodo.org/record/7022237
Zodariellum bactrianum
Creators: name: Zamani, Alireza
name: Marusik, Yuri M.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Zodariidae
Zodariellum
Zodariellum bactrianum

description: Zodariellum bactrianum (Kroneberg, 1875)
Figs 1B, 2B, 2F, 3B, 4B, 5B, 6B, 7A–C, 8A–C, 9B–C, 12A
Zodarion bactrianum Kroneberg, 1875: 12, plate 1, fig. 5 (♂ ♀).
Zodarion bactrianum: Andreeva & Tyshchenko 1968: 687, fig. 4 (♂).— Andreeva 1976: 35, fig. 42 (♂; figure from Andreeva & Tyshchenko 1968).
Zodariellum bactrianum: Zamani & Marusik 2021: 185, fig. 28D–F (♂).
Material examined. TAJIKISTAN: Khatlon Region: 1♂ (ZMMU), Shaartuz Dist., Babatag Mt. ridge, 37°04'32.9"N 68°01'10.6"E, 427 m, cay cliffs, 20.04.2015 (Y.M. Marusik); Sughd Region: 1♂ (ZMMU), Buston, ca. 40°31'N 69°20'E, 18.05.2002 (S.L. Zonstein).
Description. See Andreeva & Tyshchenko (1968). Habitus of male as in Fig. 1B. Male palp as in Figs 2B, 2F, 3B, 4B, 5B, 6B, 7A–C, 8A–C, 9B–C.
Distribution. This species was described from eastern Uzbekistan, and later reported from Tajikistan (Spassky & Luppova 1945; Andreeva & Tyshchenko 1968; Andreeva 1976; Zamani & Marusik 2021) (Fig. 12A). According to Mikhailov (2021), this species also occurs in Kazakhstan and Kyrgyzstan.
description: Published as part of Zamani, Alireza & Marusik, Yuri M., 2022, New taxonomic considerations in Zodariellum Andreeva & Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera, pp. 161-177 in Zootaxa 5178 (2) on pages 167-168, DOI: 10.11646/zootaxa.5178.2.3, http://zenodo.org/record/7022237
Zodariellum continentale
Creators: name: Zamani, Alireza
name: Marusik, Yuri M.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Zodariidae
Zodariellum
Zodariellum continentale

description: Zodariellum continentale (Andreeva & Tyshchenko, 1968)
Fig. 12A
Zodarion continentalis Andreeva & Tyshchenko, 1968: 687, fig. 5 (♂).
Zodarion martynovae Andreeva & Tyshchenko, 1968: 684, figs 1–2 (♀). Syn. n.
Zodarion martynovae: Andreeva 1976: 37, figs 45–46 (♀).
Zodarion continentalis: Andreeva 1976: 35, fig. 43 (♂).
Zodarion continentale: Brignoli 1983: 212.— Jocqué & Henrard 2015: 21.
Zodariellum continentale: Marusik & Koponen 2001: 41.
Zodariellum continentalis: Zamani & Marusik 2021: 185.
Comments. The proposed synonymy is due to the fact that the type materials of Z. martynovae (described on the basis of female) and Z. continentale (described on the basis of male) were collected in the same locality. The holotype female of Z. martynovae is in the Institute of Zoology of the Polsih Academy of Sciences (W. Wawer, pers. comm.).
Distribution. Uzbekistan, Tajikistan (Fig. 12A).
description: Published as part of Zamani, Alireza & Marusik, Yuri M., 2022, New taxonomic considerations in Zodariellum Andreeva & Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera, pp. 161-177 in Zootaxa 5178 (2) on page 169, DOI: 10.11646/zootaxa.5178.2.3, http://zenodo.org/record/7022237
Zodariellum nenilini
Creators: name: Zamani, Alireza
name: Marusik, Yuri M.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Zodariidae
Zodariellum
Zodariellum nenilini

description: Zodariellum nenilini (Eskov, 1995)
Figs 1D–E, 2C, 3C, 4C, 5C, 6C, 7D–F, 8D–F, 10A, 12A–B
Zodarion nenilini Eskov, in Eskov & Marusik 1995: 62, figs 27–29 (♂ ♀).
Zodariellum nenilini: Marusik & Koponen 2001: 42, figs 1, 6, 10–13, 19–24 (♂ ♀).— Esyunin et al. 2007: 57, figs 21–23 (♂).— Marusik & Kovblyuk 2011: 277, figs 44.2. 4–5 (♂ ♀).
Material examined. KAZAKHSTAN: East Kazakhstan Region: 2♂ (ISEA), W outreaches of Narym Mt. Range, Kaindy River Valley, 10–15 km SE of Slavyanka, [43°N 78°E], steppes, 6– 7.05.1999 (R. Dudko, I. Lyubechanski); 1♀ (ISEA), northern Cis-Zaisan (=N cosat of Zaisan Lake), N part of Kuludzhun sands, [48°40'N 83°20'E], 8– 10.05.1999 (R. Dudko, I. Lyubechanski).
Description. See Eskov & Marusik (1995) and Marusik & Koponen (2001). Habitus of male and female as in Fig. 1D, E. Male palp as in Figs 2C, 3C, 4C, 5C, 6C, 7D–F, 8D–F. Epigyne as in Fig. 10A: fovea (Fo) wider than long; copulatory ducts (Cd) contiguous and forming angle>90º.
Distribution. Known from South Urals (51°54'N, 55°20'E; Esyunin et al. 2007), east Kazakhstan (type locality) and Gobi-Altai and Bayankhongor Aimags (ca 101ºE) of Mongolia (Marusik & Koponen 2001) (Fig. 12A–B). It is possible that the records from South Ural belong to Z. volgouralense Ponomarev, 2007.
description: Published as part of Zamani, Alireza & Marusik, Yuri M., 2022, New taxonomic considerations in Zodariellum Andreeva & Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera, pp. 161-177 in Zootaxa 5178 (2) on pages 172-173, DOI: 10.11646/zootaxa.5178.2.3, http://zenodo.org/record/7022237
Zodariellum planum Zamani & Marusik 2022, comb. n.
Creators: name: Zamani, Alireza
name: Marusik, Yuri M.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Zodariidae
Zodariellum
Zodariellum planum

description: Zodariellum planum (Zhang & Zhang, 2019) comb. n.
Fig. 12B
Zodarion planum Zhang & Zhang, 2019: 73, figs 4A–E, 5A–C (♂).
Comments. Currently known only from the original description, on the basis of material from central China (Fig. 12B). Here, it is transferred to Zodariellum due to the conformation of the male palp with large RTA, long filamentous embolus and long conductor, as well as the presence of a tutalculum and cymbial diverticulum (Zhang & Zhang 2019: fig. 4E).
Distribution. Central China (Shaanxi) (Fig. 12B).
description: Published as part of Zamani, Alireza & Marusik, Yuri M., 2022, New taxonomic considerations in Zodariellum Andreeva & Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera, pp. 161-177 in Zootaxa 5178 (2) on page 174, DOI: 10.11646/zootaxa.5178.2.3, http://zenodo.org/record/7022237
Zodariellum spasskyi Zamani & Marusik 2022, comb. n.
Creators: name: Zamani, Alireza
name: Marusik, Yuri M.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Zodariidae
Zodariellum
Zodariellum spasskyi

description: Zodariellum spasskyi (Charitonov, 1946) comb. n.
Fig. 12A
Zodarion spasskyi Charitonov, 1946: 22, fig. 11 (♀).
Zodarion spasskyi: Charitonov 1969: 72.
Comments. This species is transferred to Zodariellum due to its screw-like copulatory ducts, a character unknown in other Zodariinae.
Distribution. According to WSC (2022), this species is known from Uzbekistan and Kazakhstan. Accounting that the only figure provided with the original description is not very informative, it is very much possible that the record from Kazakhstan belongs to another species. We have only marked the type locality on the map (Fig. 12A).
description: Published as part of Zamani, Alireza & Marusik, Yuri M., 2022, New taxonomic considerations in Zodariellum Andreeva & Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera, pp. 161-177 in Zootaxa 5178 (2) on page 174, DOI: 10.11646/zootaxa.5178.2.3, http://zenodo.org/record/7022237
Zodariellum tadzhikum Zamani & Marusik 2022, comb. n.
Creators: name: Zamani, Alireza
name: Marusik, Yuri M.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Zodariidae
Zodariellum
Zodariellum tadzhikum

description: Zodariellum tadzhikum (Andreeva & Tyshchenko, 1968) comb. n.
Fig. 12A
Zodarion tadzhikum Andreeva & Tyshchenko, 1968: 686, fig. 3 (♀).
Zodarion tadzhikum: Andreeva 1976: 39, fig. 47 (♀).
Comments. There are no proper figures of the epigyne available for this species.We have transferred it to Zodariellum considering that Andreeva (1976) compared it with Z. bactrianum and Z. zebra. The holotype is in the Zoological Institute of the Polish Academy of Sciences (W. Wawer, pers. comm.).
Distribution. Tajikistan (Fig. 12A).
description: Published as part of Zamani, Alireza & Marusik, Yuri M., 2022, New taxonomic considerations in Zodariellum Andreeva & Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera, pp. 161-177 in Zootaxa 5178 (2) on page 174, DOI: 10.11646/zootaxa.5178.2.3, http://zenodo.org/record/7022237
Zodariellum testaceofasciatum Zamani & Marusik 2022, comb. n.
Creators: name: Zamani, Alireza
name: Marusik, Yuri M.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Zodariidae
Zodariellum
Zodariellum testaceofasciatum

description: Zodariellum testaceofasciatum (Spassky, 1941) comb. n.
Fig. 12A
Zodarion testaceo-fasciatum Spassky, 1941: 14, fig. 3 (♀).
Zodarion testaceo-fasciatum Spassky & Luppova, 1945: 45 (♀).
Zodarion testaceofasciatum: Brignoli 1983: 213.
Comments. This species was described twice, in two different publications and under the same name. Apparently, the original manuscript was sent to the publisher in Riga prior to World War II, and Spassky was unaware that his paper was published during the war (in 1941) and resubmitted the manuscript again, this time in coauthorship with E.P. Luppova to the Soviet journal Entomologicheskoe Obozrenie. The second manuscript was published in 1945, without any figures.
The species is known only from the type locality, and only one figure showing the abdominal pattern has been published for it. It is very likely that Z. zebra, a species with similar abdominal pattern and type locality not far from that of Z. testaceofasciatum, is a junior synonym of it. Although the epigyne of this species is not described, we decided to transfer it to Zodariellum considering its similarity to Z. zebra and the fact that only Zodariellum and “ Zodarion ” denisi Spassky, 1938 are known to occur in the region.
Distribution. Tajikistan (Fig. 12A).
description: Published as part of Zamani, Alireza & Marusik, Yuri M., 2022, New taxonomic considerations in Zodariellum Andreeva & Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera, pp. 161-177 in Zootaxa 5178 (2) on page 174, DOI: 10.11646/zootaxa.5178.2.3, http://zenodo.org/record/7022237
Zodariellum turanicum Zamani & Marusik 2022, sp. n.
Creators: name: Zamani, Alireza
name: Marusik, Yuri M.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Zodariidae
Zodariellum
Zodariellum turanicum

description: Zodariellum turanicum sp. n.
Figs 1C, 2D, 3D, 4D, 5D, 6D, 9D, 12A
Type material. Holotype ♂ (ZMMU), TURKMENISTAN: Lebap Region: Kugitang-Tau Mnt. range, environs of Khodzhapil, [37°56'N 66°37'E], ca 1100 m, on the ground, 20.05.1985 (A.V. Tanasevitch).
Etymology. The specific epithet refers to Turan, a historical region in Central Asia.
Diagnosis. The new species has abdominal pattern similar to those of Z. zebra and Z. testaceofasciatum, two species described not far from each other (eastern Uzbekistan and western Tajikistan) and currently known only from females. It differs from the males of its congeners by having two transverse light stripes on the abdomen (vs. lacking in other species), and a V-shaped dark marking on a light carapace (vs. carapace uniformly coloured or with small longitudinal spots along the fovea). The male palp of Z. turanicum sp. n. is most similar to that of Z. bactrianum by having two processes in the distal half of RTA, but can be differentiated by the angled anterodorsal part of RTA (Ap) (vs. rounded), and branches located at the same level (vs. mesal one [=closer to cymbium] located more posteriorly).
Description. Male. Habitus as in Fig. 1C. Total length 3.55. Carapace 1.60 long, 1.50 wide. Eye sizes: AME: 0.18, ALE: 0.10, PME: 0.10, PLE: 0.08. Carapace, sternum, chelicerae, labium and maxillae yellowish. Carapace light coloured with irregular V-shaped marking. Legs slightly lighter than carapace.Abdomen dorsally dark maroon, with 2 pale transverse stripes in anterior half and 3 obtuse triangular spots in anterior half; venter light beige. Measurements of legs: I: 6.78 (1.82, 0.62, 1.59, 1.77, 0.98), II: 6.23 (1.67, 0.55, 1.37, 1.78, 0.86), III: 6.57 (1.77, 0.61, 1.38, 1.91, 0.90), IV: 8.22 (2.34, 0.63, 1.88, 2.44, 0.93).
Palp as in Figs 2D, 3D, 4D, 5D, 6D, 9D. Tibial apophysis longer than tibia, 2.3 times longer than wide in proximal part, tip inclined, with anterodorsal part (Ap) angled; dorsal part with 2 processes originated at anterior 1/3, processes of subequal length and shape and located at the same level (one behind the other), ectal margin smoothly rounded in ventral view; cymbium length/width ratio 1.8; embolus originating at ca. 4:30 o’clock position.
Female. Unknown.
Distribution. Known only from the type locality in Lebap Region, northeastern Turkmenistan (Fig. 12A)
description: Published as part of Zamani, Alireza & Marusik, Yuri M., 2022, New taxonomic considerations in Zodariellum Andreeva & Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera, pp. 161-177 in Zootaxa 5178 (2) on page 175, DOI: 10.11646/zootaxa.5178.2.3, http://zenodo.org/record/7022237