Stenus trifidus L. BENICK 1938
Creators: name: Puthz, Volker
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Stenus
Stenus trifidus

description: Stenus trifidus L. BENICK, 1938
(Abb. 61, 62, 72, 73, 159, 160, 235-237, 248-251, 302)
Stenus trifidus L. BENICK, 1938: 261; PUTHZ, 1969: 57.
Material: COSTA RICA, San José Prov.: ♂ - Syntypus:, La Caja, 8 km W San José, 1934, H. Schmidt (FMCh); ♀ -ST: Vara Blanca- Poás, 2000m (DEI); 7♂♂, 5 ♀♀: ibidem, VII., VIII.1938, 8.XII.1940, A. Bierig (FMCh, cP); 1♀: La Estrella, 24.X.1941, idem (FMCh); 2♂♂, 1 ♀: La Palma, 1550m, Carribean Escarpment, 10 o 3’N, 83 o 58’W, concentrated leaf humus on floor, 5.IV.1973, Wagner & Kethley (FMCh, cP); 4♂♂, 14♀♀: Zurqui de Moravia, 1600m, Malaise, III.- VI.1992, 30.III.- 30.V.1995, P. Hanson (FMCh, cP); 4♀♀: San Juan, 14 km NE, Finca Zurqui, 10 o 28’57’’N, 84 o 0‘22’’W, 6.VII.2000, Ashe & al. (KSEM); 6♂♂, 4♀♀: 2,4 km ENE San Gerardo de Rivas, Cloudbridge Reserve, River trail, 1750m, 9 o 28.47’N, 83 o 34.20’W, 9.VI.2004, fungus covered logs, 10., 11.VI.2004, litter under treefall, Ashe, Falin & Hinojosa (KSEM, cP); Puntarenas Prov.: 2♂♂, 3 ♀♀: Reserva de Monteverde, 5000 feet, 27.-29.V.1979, J.M. & B.A. Campbell (CNC, cP); 2♀♀: ibidem, 1400m, 1700m, 1.VI., 4.VI.1979, H. & A. Howden (CNC); 1♂, 2♀♀: ibidem, ca. 4 km E Monteverde, 5000-5500 feet, II, III.1985, L. Herman (AMNH, cP); 1♂, 2♀♀: Monteverde 1520 (1550)m, under bark; light interception trap / coll. on Auricularia, 15., 21., 23.V.1990, Ashe & al. (KSEM); 1♂, 1♀: Monteverde Reserve (trail near lab), FIT, 1.- 2.VI.1993, C. Michalski (KSEM, cP); 2♂♂: Monteverde area, 1400-1700m, 6.-14.VI.1973, Erwin & Hevel Centr. Am. Exp. (USNM, cP); 12♂♂, 13 ♀♀: Monteverde, Est. Biologica Monteverde, 1750-1800m, cloud forest litter, 10.-15.VI.2001, R. Anderson (KSEM, cP); Cartago Prov.: 1♂, 3♀♀: Tapanti, 1100m, VIII.1938, VII., IX.1939, III.1940, A. Bierig (FMCh, USNM); 1♂, 1♀: San Isidoro-La Estrella, 1600m, 28.IX.1941, idem (FMCh); 1♂: 10 km S Tapanti, Rio Grande de Orosi, 1500m, 9 o 42’N, 83 o 47’W, mixed floor litter, 14.IV.1973, Wagner & Kethley (FMCh); 2♀♀: Embalse El Llano on Rio Macho, 5 km S Orosi, 1500m, lights, 25.V.1992, Ashe & al. (KSEM); 1♂: Embalse el Llano, Rio Macho, 1500m, 8 km S. Orosi, lights, 25.V.1992, J. Jameson (KSEM); 2♂♂, 2♀♀: P. N. Tapanti, 1500m, 9 o 45’N, 83 o 49’W, montane tropical forest, leaf litter, 4.VII.1997, R. Anderson (KSEM, cP); 3♀♀: ibidem 1650m, 9 o 43’2’’N, 83 o 46’22’’W, cloud forest litter (dry), 18.II.1998, idem (KSEM); 3♂♂, 7♀♀: ibidem 1150-1480m, 17.-19.VII.2000, Ashe & al. (KSEM, cP); 1♂, 1♀: Refugio Nacional de Fauna Silvestre Tapanti, 4.- 12 km E Station, 1430-1820m, 9 o 44.2’’N, 83 o 46.8’’W, tree and shrub slash, 30.X.2001, R. Brooks (KSEM); 1♀: 1 km W Santa Cruz de Turrialba, Route 230, 5500 feet, litter near stream, 13.III.1991, L. Herman (KSEM); 1♀: La Cangreja, 1950m, VI.-VII.1992, P. Hanson (KSEM); 23♂♂, 12♀♀: Cartago / Heredia, P.N. Tapanti, 1500m, 9 o 45’0’’N, 83 o 49’0’’W, 4.VI.1977, R. Anderson (KSEM, cP); 4♂♂, 17 ♀♀: San José / Cartago, km 45 International American Highway, 6 km NE El Empalpe, 1975m, 9 o 45’0’’N, 83 o 58’30’’W, 8.VI.1997, idem (KSEM, cP); Heredia: 15♂♂, 12♀♀: 3,2 km N Junction Routes 126 (9) and 120, 0,3 km E on Road to San Rafael, 5600 feet, litter on shore of stream, 18.III.1991, L. Herman (AMNH, cP); 5♂♂, 4♀♀: Porrosati, 6 km N San José de la Montana, 1900m, 10 o 5’30’’N, 84 o 7’0’’W, 27.VI.1997, R. Anderson (KSEM, cP); 2♂♂, 1 ♀: Cerro Chompipe ca. 2 km N Monte de la Cruz, 1950m, 10 o 5’13’’N, 84 o 4’45’’W, 25. VII.2000, Ashe & al. (KSEM); 8♂♂, 6♀♀: 6 km ENE Vara Blanca, 2000m, 10 o 11’N, 84 o 07’W, montane forest leaf litter, 15.-22. IV.2002, R. Anderson (KSEM, cP); 1♀: 6 km ENE Vara Blanca, 1950-2050m, 10 o 11’N, 84 o 07’W, INBio-OET-ALAS transect, 21.-25.IV.2004, J. Prena (coll. Kleeberg); 6♂♂, 1♀: Finca Zurqui, Tajo, 1575m, 10 o 2.3’N, 84 o 1.26’W, wet cloud forest litter, 27.VI.2000, idem (KSEM, cP); Alajuela: 1♀: Poasito, Rio Poasito, 10km below Volcan Pass, 2000m, 10 o 9’N, 84 o 13’W, concentrated floor mold under tree ferns, on slope, 16.IV.1973, Wagner & Kethley; 2♂♂, 1 ♀: 7,7 km N Junction Road 126 (9) & 120 Road to Puerto Viejo, 4600 feet, litter near stream, 17.III.1991, L. Herman (KSEM, cP); Guanacaste: 3♂♂ 1♀: Santa Elena, SE. Cloud Forest Reserve, 1650m, 10 o 20.42’N, 84 o 47.53’W, cloud forest litter, 11.VI.2001, R. Anderson (KSEM, cP). PANAMA: Chriqui Prov.: 1♂, 3♀♀: 5,6 km N Boquete, La Culebra trail, 1450m, 8 o 49’23’’N, 82 o 25’18’’W, fungusy log, 15.VI.1996, Ashe & Brooks (KSEM, cP); 6♀♀: ibidem, 5,7 km NE Boquete, 1500m, 8 o 48’0’’N, 82 o 26’0’’W, oak forest litter, 19.VI.1995, R. Anderson (KSEM, cP); 1♀: ibidem 5,6 km N Boquete, 1400m, Xantosoma robustum rotting leaves, 19.VI.1996, idem (KSEM); 1♂, 1♀: 5,4 km NE Boquete, 8 o 48’N, 82 o 26’W, o 1520m, treefall slash/fogging fungusy log, Ashe & Brooks (KSEM); 1♀: Cerro Pando, 1875m, 8 54’42’’N, fungusy log, 17.VI.1996, Ashe & Brooks (KSEM); 1♂, 1♀: near Bajo Boquete, Pate de Machu Mt., 11.VII.1983, leaf litter, N. Sorkin (AMNH); 1♂: Boquete / Cerro Punta Volcan Baru, Sendero Los Quetzales, Nebelwald, 21.II.2004, I. & H. Schatz (cP); 1♀: 27,7 km W Volcan, Hartmann’s Finca, 1800m, 8 o 45’0’’N, 82 o 48’0’’W, oak forest litter, 16.VI.1996, A. Anderson (KSEM); 1♀: La Fortuna, Quebrada Al. Trail, 8 o 42’N, 82 o 14’W, 1250m, flower fall litter, 11.VI.1995, Ashe & Brooks (KSEM); 1♂: Embalse el Llano, Rio Macho, 1500m, 8 km S. Orosi, lights, 25.V.1992, J. Jameson (KSEM); 1♀: Hornito, Finca La Suiza, 1220m, 8 o 39’N, 82 o 12’W, FIT, 3.VI.2000, H. & A. Howden (KSEM); Bocas del Toro Prov.: 1♂: Cerro Colorado, Qbda 3w, 1490m, along patch on slope, 24.I.1981, W. Suter (FMCh); 1♀ ibidem 1460m, moss on rocks with sphagnum, 18.I.1981, idem (FMCh); 1♂: ibidem off highway 17, 1490m, grass litter, path side clearing, 11.I.1981, idem (FMCh); 2♀♀: ibidem 1460m, litter under tree ferns exposed slope, 18.I.1981, idem (FMCh, cP); 1♂, 1♀: ibidem 1490m, conc. of wood/slash, small clearing, 11.I.1981, idem (FMCh, cP); 1♀: Qbda Gato, 1375m, tree fern trunk core on roadside, 21.I.1981, idem (FMCh); 2♂♂, 1 ♀: ibidem 2w, 1475m, litter on bulldozed slope along highway 17, 18.I.1981, idem (FMCh, cP); 2♂♂, 2♀♀: ibidem, 1475m, litter under roadside vegetation along highway 17, 15.I.1981, idem (FMCh, cP); 2♀♀: ibidem, trail to peak, 1475m, litter at burn tree fern, 28.I.1981, idem (FMCh); 3♂♂, 1♀: ibidem, 1290m, semidry hillside litter, 24.I.1981, idem (FMCH, cP); 1♀: ibidem, 1290m, ecotonal slash and bamboo litter, 17.I.1981, idem (FMCh); 1♀: Cerro Bollo (La Torre), 1800m, under bark, mossy stage II log, 20.I.1981, idem (FMCh); 3♂♂: ibidem, litter under blooming bromeliad, 20.I.1981, idem (FMCh, cP); 1♂: ibidem, litter under broad leaved epiphyte, 20.I.1981, idem (FMCh); 1♂: Fortuna to Soldad Trail, Soledad side, 20.V.1978, O’Brien & Marshall (FMCh).
Bemerkungen: Diese Art gehört zu den in Costa Rica und in Panama am häufigsten gesammelten Arten der Gattung. Sie ist, was das Exosklelett angeht, bemerkenswert variabel (Abb. 61, 62, 72, 73). ♂: Metasternum (Abb. 159, 160), der Aedoeagus (Abb. 237) zeigt einen schlanken Medianlobus, dessen Apikalpartie kräftig dorsad gebogen ist und an seiner Spitze eine rundliche, aussen schwach sklerotisierte, knopfförmige Erweiterung besitzt (Abb. 248-251); im Innern werden 10-14 kräftige Zähne und in einem häkchenbesetzten Innensack erkennbar (Abb. 235, 236). Der apikale Ausschnitt des 8. Sternits ist tiefer als bei der ähnlichen Art S. mucronatus SHARP (Sternitlänge: Tiefe des Ausschnitts = 2.15- 2.61 (Ø 2,45, N= 13); S. mucronatus: 3,28- 3,46). Das 8. Sternit des Weibchens besitzt einen starken apikomedianen Vorsprung (Abb. 302).
description: Published as part of Puthz, Volker, 2016, Übersicht über die neotropischen Arten der Gattung Stenus LATREILLE mit seitlich gerandetem Abdomen und ungelappten Tarsen (Coleoptera, Staphylinidae) 350. Beitrag zur Kenntnis der Steninen, pp. 1551-1634 in Linzer biologische Beiträge 48 (2) on pages 1564-1565, DOI: 10.5281/zenodo.5355748