Leptophlebiidae
Creators: name: Kluge, Nikita J.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Ephemeroptera
Leptophlebiidae

description: Key to imagines of Asian Leptophlebiidae with both claws of each leg pointed
1(2) Hind wings absent............................................................................ Nathanella
2(1) Hind wings present (Kimminsula -complex)................................................................ 3
3(4) Paracercus vestigial, about as long as 10th abdominal segment (Fig. 271)......................... Ceylonula femoralis
4(3) Paracercus as long as cerci.
5(8) Vein MP of hind wing not furcated (Fig. 52). Tibiae of middle and hind legs with groove crossing inner margin and corresponding to patella-tibial suture (Figs 54–55) (Ghatula gen. n.)..................................................... 6
6(7) Abdominal terga VIII and IX each with pair of dark brown maculae (as in Fig. 69) (presumably, based on larval hypodermal coloration)................................................................... Ghatula quadrimaculata sp. n.
7(6) Abdominal terga VIII and IX without such maculae (Figs 57–58)................................ Ghatula rufa sp. n.
8(5) Vein MP of hind wing furcated: either with intercalary forming triad (Fig. 153, 157), or without intercalary (Fig. 217), or with both branches fused distally (Fig. 158). All tibiae (including those of middle and hind legs) without groove crossing inner margin (Figs 159–164, 213–214, 461–462, 680–681)............................................................ 9
9(12) Hind wing with anterior-proximal margin bordered with brown (Figs 157–158, 671)............................... 10
10(11) Hind wing with posterior-distal margin bordered with brown (Figs 157–159). ♂: eyes meet on meson (Fig. 168); penis with pair of apical spines directed ventrally (Figs 172–173, 179, 183)............................ Petersula heptagenoides sp. n.
11(10) Hind wing with posterior-distal margin not colored (Fig. 671). ♂: eyes widely separated (Fig. 674); penis without spines directed ventrally (Figs 688, 690, 697)......................................... Hubbardula heterolepida gen. sp. n.
12(9) Hind wing with brown coloration at base only (Figs 217, 365, 412, 454, 540)..................................... 13
13(16) ♂: dorsal eyes meet on meson (as in Fig. 168); ventral spines on apices of penis grooved (as in Figs 172–173, 179; Sivaramakrishnan 1984: fig. 6, 9) (Petersula)...................................................................... 14
14(15) ♂ imago: abdominal terga III–V without paired lines (Sivaramakrishnan 1984: figs 10, 12); terga I–IX light, tergum X dark................................................................................... Petersula courtallensis
15(14) ♂ imago: abdominal terga III–V with submedian pair of longitudinal lines (Sivaramakrishnan 1984: fig. 11); terga I–VII light, terga VIII–X dark....................................................................... Petersula nathani
16(13) ♂: dorsal eyes widely separated (Figs 369, 531); ventral spines on apices of penis cone-shaped, without groves (Figs 374–375, 377, 380,417, 418, 421, 469–470, 534, 545) (Kimminsula)................................................... 17
17(18) Each abdominal tergum V–VIII or V–IX with unpaired triangular dark spot adjacent to anterior margin of tergum (as in Fig. 580) (presumably, based on larval hypodermal coloration)................................ Kimminsula latifolia sp. n.
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TABLE 1. (Continued)
18(17) Abdominal terga either without smaller dark spots (Figs 369–370, 373, 463, 532), or with large spot adjacent to posterior margin of tergum (Figs 414, 416)........................................................................... 19
19(20) ♂: penis with unpaired projection between pair of lobes (Figs 534, 545); tibiae black-brow, contrastingly darker than femora (Figs 535–536); abdominal coloration as in Fig. 532 ........................................ Kimminsula podi sp. n.
20(19) ♂: penis lobes contiguous basally, without projection between them (Figs 374–375, 377, 380); tibiae brown or ocher, not darker than femora (Figs 370, 414); abdominal coloration different (Figs 369–370, 373, 414, 416, 463)...................... 21
21(22) Abdominal terga mostly brown, with antero-lateral areas lighter ocher (Figs 413–416)............ Kimminsula taprobanes
22(21) Abdominal terga mostly light ocher, with brown markings (Figs 369–370, 373, 463).............................................................. Kimminsula fasciata (and the form intermediate between K. taprobanes and K. fasciata)
description: Published as part of Kluge, Nikita J., 2022, Review of the Kimminsula-complex (Ephemeroptera, Leptophlebiidae), pp. 1-140 in Zootaxa 5212 (1) on pages 10-13, DOI: 10.11646/zootaxa.5212.1.1, http://zenodo.org/record/7345276