Cheumatopsyche hasalaka Olah & Johanson 2008, new species
Creators: name: Oláh, J.
name: Johanson, K. A.
Publisher: name: Zenodo
Year: 2008
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Hydropsychidae
Cheumatopsyche
Cheumatopsyche hasalaka

description: Cheumatopsyche hasalaka Oláh & Johanson, new species   
Fig. 320–324  
 This species is similar to Cheumatopsyche afra Mosely due to the presence of a rectangular segment X in lateral view; bifid segment X in dorsal view formed by the short apicoventral setose lobes; and the sinuate and filiform harpagones. The new species is also easily distinguished by the presence of light pattern on the forewing membranes, and having laterad producing apicoventral setose lobe, not mesally as in C. afra.   
Male. Body brown with dense lighter pubescence, especially on legs. Maxillary palp formula I-(III, IV)- II-V, segment V as long as sum of segments I–IV. Head dorsum dark brown, with 7 brown setal warts; anteromedial and anterolateral warts fused. Swollen setal wart absent on proepisternum. Setal surface present on each precoxale. Pretarsal claws asymmetrical, laterally flanked by setal bundle; claws on mid- and hind-legs asymmetrical.  
Wings. Forewing membrane (Fig. 320) evenly brown, with slightly darker veins and pale patches of various size and shape. Forewing crossveins m-cu and cu almost tangential in oblique line. Sc and R separate before costa. Cu2 and A1 running separately before costa. Hind wing Sc and R meeting at crossvein r. Crossvein r obligue, almost meeting crossvein s. Fork 1 absent. Forewing length 4.9 mm.  
Male genitalia (Fig. 321–324). Abdominal segment IX fused annularly, tergum half as long as sternum (Fig. 321). Anterior margins of segment IX regularly bow-shaped, more sinuate in lateral aspect (Fig. 321). Apical lobe on posterolateral margins blunt obtusely angled, located well above less sclerotized articulation cavity of gonocoxites. Spine row on posterior margins of segment IX intermittent, heterogeneous. Spines on dorsolateral lobes 3 times longer than spines on apical lobes; 3–4 spines forming short row between apical lobes and dorsolateral spiny lobes about half as long as spines on apical lobes. Dorsolateral spiny lobes meeting mesally into small triangular (Fig. 321). Intersegmental step between segments IX and X sloping. Segment X medium sized, quadrangular, with almost right angled apical corners in lateral view (Fig. 321). Segment X bilobed in dorsal aspect, broadening slightly apicad due to slightly laterally curving apicoventral setose lobes (Fig. 322). Dorsal interlobular gap wide, medium sized (Fig. 322). Sutures on each side well developed, transverse, crossing segment X obliquely, reaching middle of dorsum. Longitudinal sutures broad, forming continuation of apicoventral setose lobes (Fig. 321), meeting transverse suture in Y. Smooth mesocaudal plate reduced, ending into straight margin. Apicoventral setose lobes short, curving dorsad, apex blunt in lateral view (Fig. 321); obliquely cut; slightly producing laterad in dorsal view (Fig. 322). Lateral setose areas (superior or preanal appendages) forming small, elevated, elongated wart, located behind transverse and above longitudinal sutures. Coxopodites exceeding apex of segment X, forming straight rod, dilating gradually apically, with small dorsal hump at mid-length; apex capitate dorsally (Fig. 321); apex slightly curving mesad in ventral view (Fig. 323). Harpagones arising from ventrum of coxopodite apices; slender almost filiform, curving dorsad, sinuate in lateral view (Fig. 321), S-shaped in ventral view (Fig. 323); with 2 minute apical setae in ventral view (Fig. 323). Phallotheca with long and slightly broadening, ventrad curving basal part (Fig. 324); horizontal section with straight dorsum and weakly concave ventrum. Phallothecal apex slightly broadening ventrad in lateral view (Fig. 324). Endophallus long, running through entire phallotheca, ending in narrow tube at gonopore. Chitinized endothecal process circular. Phallotremal sclerites rounded, and vertically elongating in lateral view. Vestigial membranous ventral endothecal lobes visible.  
 Holotype male: SRI LANKA: Kan Dist. Hasalaka, 500’, 22–25.xi.1970 [O.S. Flint, Jr.] - (NMNH).   
Paratypes: same data as holotype - 2 males, 1 female (NMNH), 1 male (OPC); Kan Dist. Hasalaka Irrigation Bungalow, 5 mi. NW Mahiyangana, 30.iii.–9.iv.1971, black light [P. Spangler] - 2 males (NMNH); Col. Dist. Tunmodera, 200’, 17.xi.1970 [O.S. Flint, Jr.] - 5 females (NMNH); Anu. Dist. Irrigation Bungalow, Padaviya 180 ft. 27.ii.–9.iii.1970 [Davis & Rowe] - 1 female (NMNH); Kan.Dist. Garulupota nr. Hasalaka, 800’, 22.xi.1970 [O.S. Flint, Jr.] - 1 male, 2 female (NMNH); Rat. Dist. Uggalkaltota, 500’, 10–14.x.1970 [O.S. Flint, Jr.] - 1 female (NMNH); Rat. Dist. Uggalkaltota 350 ft., Irrigation Bungalow, 31.i.–8ii.1970 [Davis & Rowe] - 1 female (NMNH); Ham. Dist. Palatupana 10’, 22–25.x.1970 [O.S. Flint, Jr.] - 1 female (NMNH).  
 Distribution. Sri Lanka.   
 Etymology. hasalaka, derived from the type locality, Hasalaka.
description: Published as part of Oláh, J. & Johanson, K. A., 2008, Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae), pp. 1-248 in Zootaxa 1802 on pages 193-194