Nyctiophylax (Nyctiophylax) tapan Oláh & Johanson 2010, new species
Creators: name: Oláh, János
name: Johanson, Kjell Arne
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Polycentropodidae
Nyctiophylax
Nyctiophylax tapan

description: Nyctiophylax (Nyctiophylax) tapan, new species   
Figs 27–30  
 The genitalia of this species of subgenus Nyctiophylax resemble those of N. khaoyaiensis and N. salma from Thailand. Nyctiophylax tapan, new species is distinguished from the 2 similar species by the shape of the ventroapical keels of the gonopods and the ventral paraproctal plate of sternite IX; the shape of sternite IX in lateral view; and the presence of a pronounced antecostal suture running around the entire sternite IX except at the posterior margin.   
 Male (in alcohol). Body uniformly brown. Maxillary palp formula II-I-IV-III-V, with segment III originating from mesal face before apex of segment II. Anterior tentorial pits located laterally, near eyes. Frontal and clypeal areas wide; frontogenal and clypeogenal areas narrow. Pair of large, triangular, frontal lateral compact setose warts filling dark frontal area, with white alveoli; warts almost tangential mesally; clypeal sclerites light brown, covered by large, single, clypeal diffuse setose wart; few anterior diffuse setae present on labrum. Forewing length 4.5 mm; discoidal cell and median cell closed; apical forks 2, 3, 4 and 5 present; anal veins A1, A2, and A3 without loop. Hind wing discoidal cell closed, median cell open; apical forks 2 and 5 present.   
Spur formula 3,4,4.  
Male genitalia. Sternite IX robust, sclerotized, quadrangular in lateral view; delimited by pronounced antecostal suture except on posterior margin; ventroapical keel developed into slightly S-shaped, tapering process in lateral view. Tergum IX membranous, with semisclerotized integument above sternite IX and behind paraproctal complex visible, confluent with paraprocts. Segment X membranous, elongate, nearly quadrangular in lateral view. Cerci elongate, club-shaped, originating from paraproctal bodies. Paraproctal complex with 2 pairs of paraproctal processes: dorsal paraproctal processes forming arching spines originating anteriorly before running dorsad and turning posterad; ventral paraproctal processes forming pair of large, horizontally elongated plates, each with capitate apex and short, dorsal process with setae on terminal area. Plates of paraproctal complex meeting ventrally, forming sclerous subphallic bridge and phallic guide. Gonopods elongate, almost parallel-sided, each with bifid apex in lateral view. Phallic apparatus located dorsally in genitalia, fixed, guided by paraproctal processes; phallotheca short, sclerotized, with apical rim and basidorsal plate. Phallobase with wide anterior foramen; membranous endotheca (or phallicata) with lesssclerotic lobe apicoventrally, spine-shaped in lateral view; pair of obscure phallotremal sclerites present inside.  
 Holotype male: INDONESIA (Sumatra): Kerinci National Park, Tapan Road, 1000 m, 10.vii.2008, light trap [J. Oláh, Jr.]—(OPC, in alcohol).   
 Distribution: Indonesia (Sumatra).   
 Etymology: Tapan, named after the type locality of the species, Tapan Road in Kerinci National Park.
description: Published as part of Oláh, János & Johanson, Kjell Arne, 2010, Fifteen new Trichoptera (Insecta) species from Sumatra, Indonesia, pp. 1-35 in Zootaxa 2618 (1) on pages 17-18, DOI: 10.11646/zootaxa.2618.1.1, http://zenodo.org/record/5304087