Pseudoundacypha yananensis Huang 2023, sp. nov.
Creators: name: Huang, Di-Ying
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lophioneurida
Westphalothripidesidae
Pseudoundacypha
Pseudoundacypha yananensis

description: Pseudoundacypha yananensis sp. nov.   
 Holotype. NIGP203301, an almost complete forewing, basal-upper section slightly broken.   
 Etymology. Yan’an, a City of Shaanxi Province, where the specimen was collected.   
 Diagnosis. As for genus.   
 Type locality and horizon. Fossil locality near the Peizhuang Village, Yan’an City, Shaanxi Province; Middle Jurassic Yan’an Formation.   
 Description. Forewing small-sized, 2.4 mm long, 0.9 mm wide, widest at circa 3/4 of wing length; wing width distinctly increase backwards; ScP present, elongate, nearly parallel wing margin; RA slightly curved apically, reaching 3/5 of wing length; common stem of RP straight, RP 1 almost straight, RP2 curved, a little longer than RP 1; M stem straight, M 1 almost straight, reaching wing apex, M 2 straight, distinct shorter than M 1; CuA slightly curved posteriorly; RP branched earlier than M branched; CuP nearly straight, rather elongate, reaching more than 1/3 of wing length; a short crossvein cua-cup present; A 1 nearly straight, A 2 small and near wing base.   
 Remarks. Pseudoundacypha gen. nov. can be assigned into the clade Lophioneurida by the position of cua-cup (Nel et al., 2012). Pseudoundacypha gen. nov. generally resembles Undacypha Vishniakova, 1981, but differs from the latter by presence of ScP and RA making an angle with R stem (Vishniakova, 1981; Ansorge, 1996; Nel et al., 2014). The new genus is also similar to Jantardachus reductus Vishniakova, 1981, but they can be also distinguished by the above differences, as well as a distinct longer stem of RP in Pseudoundacypha gen. nov. (Vishniakova, 1981; Nel et al., 2014).   
 The presence of a long ScP is a common character for Panthysanoptera, but it also occurs in some species of Lophioneurida such as Lophiocypha permiana Tillyard, 1935, Lophioneura ustulata Tillyard, 1921, and Lophioneurodes sarbalensis Becker-Migdisova, 1953. Among them, the new species resembles the Permian L. permiana. Pseudoundacypha yananensis differs from the latter by RP and M forked with larger angles and presence of a relatively straight CuA. Pseudoundacypha yananensis also differs from the Late Jurassic Karatavocypha oculata Vishniakova, 1981 by a longer ScP, RP, M forked with smaller angles, and a longer common stem of RP.   
 Abundant fossils of Thripida from the Middle-Late Jurassic Daohugou biota were assigned within the two genera Undacypha and Lophiosina (Nel et al., 2014; Huang et al., 2018; Lian et al., 2021). These genera are clearly different from the new genus by absence of ScP. Another species, Mesopsocus divaricatus Hong, 1983, was initially classified within Psocoptera. In the original description and line drawing, a weak ScP is evident. It distinguishes itself from the new species by RP and M forked with much smaller angles and a distinct longer CuA (Hong, 1983).
description: Published as part of Huang, Di-Ying, 2023, Discovery of Thripida from the Middle Jurassic Yan'an Formation, pp. 22-25 in Zootaxa 5396 (1) on page 24, DOI: 10.11646/zootaxa.5396.1.6, http://zenodo.org/record/10441081