Quadrimaera quadrimana
Creators: name: Krapp-Schickel, Traudl
Publisher: name: Zenodo
Year: 2009
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Amphipoda
Maeridae
Quadrimaera
Quadrimaera quadrimana

description: <i>Quadrimaera quadrimana</i> (Dana, 1853) (Fig. 21) <i>Gammarus quadrimanus</i> Dana, 1853: 955, pl. 65, fig. 9. <i>Maera quadrimana</i>. — Schellenberg, 1938: 45, figs 21–22. —J.L. Barnard, 1965: 511, fig. 17. —J.L. Barnard, 1970: 152, figs 94–95. — Ledoyer, 1972: 229, pl. 45. — Berents, 1983: 128, fig. 22. — Myers, 1985: 116, fig. 91. <i>Quadrimaera quadrimana</i>. — Lowry &amp; Stoddart, 2003: 186. <b>Material examined.</b> 1 female "intersex", 3.5 mm (slide), AM P77724 (JDT / LIZ 2); 1 male, AM P77726 (JDT / LIZ 3); 1 male, AM P77725 (JDT / OPH 2); 1 female, 5 juveniles, AM P70601 (QLD 1622); 1 female incomplete, AM P70963 (QLD 1687); 1 female incomplete, AM P71030 (QLD 1711); 1 male, 5.0 mm (2 slides), AM P71041 (QLD 1715); 2 males, 9 females, AM P71270 (QLD 1767); 1 female, AM P71280 (QLD 1770); 1 male, 1 female, AM P71474 (QLD 1792); 1 male, 8 females, AM P77727 (SEL /LZI-2-7). <b>Type locality.</b> Fiji (~ 17°40'S 178°31'E). <b>Description.</b> Based on male, 5.0 mm, AM P71041. <b>Head.</b> <i>Head</i> anteroventral corner produced into a sharp cusp, lacking anteroventral slit. <i>Antenna 1</i> peduncular article 2 longer than article 1; accessory flagellum with 5–6 articles. <i>Antenna 2</i> gland cone reaching end of short article 3. <i>Mandible</i> palp article 3 longer than article 2. <b>Pereon.</b> <i>Gnathopod 1</i> coxa quadrangular, anteroventral corner rounded; carpus triangular, with notch on anterodistal margin; propodus ovoid. <i>Gnathopod 2</i> propodus rectangular with parallel margins, palm defined by V-shaped incision next to prominent acute spine; medially two smaller narrow incisions defining two molar-shaped elevations; dactylus outer margin with 1 seta, inner margin smooth. <i>Pereopods 3–7</i> with bifid dactylus. <i>Pereopods 5–7</i> basis narrow with sub-parallel margins and a posteroventral corner. <i>Pereopod 7</i> propodus posterior margin with one group of long setae. <b>Pleon.</b> <i>Epimera</i> smooth. <i>Uropod 3</i> rami subequal, longer than peduncle, outer ramus somewhat longer, truncate and with many robust setae that are shorter than ramus. <i>Telson</i> deeply cleft, lobes distally evenly truncate and with 4 robust setae, some longer, some shorter than telson length. <b>Female</b> (sexually dimorphic characters). Based on female, 3.5 mm, AM P77724. <i>Gnathopod 2</i> propodus palm with regularly serrate palm. <b>Variation.</b> Ovigerous females may have regularly serrate palmar margin or in larger individuals, a palm similar to that of males with two deeper incisions (see also Schellenberg 1938 and Myers 1985). <b>Habitat.</b> Tunicates, coral rubble, mixed coralline algae with epiphytes, brown algal tufts, bommies and sand patches, seagrass, 1–17 m depth. <b>Remarks.</b> This species is found in the same group with <i>Q. pacifica</i> (see description above). Many authors described problems with highly variable material: juveniles have very narrow rectangular pereopod bases with ratio of length to width of about 2 (see Schellenberg, 1938: 47 fig. 22), while a male of 6.4 mm is described by J.L. Barnard (1970:153 fig. 94) with a pereopod 7 basis ratio of length to width of 1.4. J.L. Barnard (1965: 511 fig. 17) shows that the middle spine on the palmar margin can be worn. <b>Distribution.</b> <i>Australia.</i> Queensland, Lizard Island (Berents 1983, current study). <i>Madagascar</i> (Ledoyer 1972, 1983). <i>India</i> (Sivaprakasam 1968). <i>Marshall Islands</i> and <i>Kiribati</i> (Schellenberg 1938; J.L. Barnard 1965). <i>Fiji</i> (Dana 1853, Schellenberg 1938, Myers 1985). <i>Hawaii</i> (Schellenberg 1938, J.L. Barnard 1970).
description: Published as part of <i>Krapp-Schickel, Traudl, 2009, Maeridae, the Ceradocus group *, pp. 598-642 in Zootaxa 2260 (1)</i> on pages 629-630, DOI: 10.11646/zootaxa.2260.1.32, http://zenodo.org/record/5309853
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