cometadata/datacite-rtg-text-other-reclassification-records huggingface dataset:
Hemidactylus Murrayi Gleadow 1887
Creators: name: Mahony, Stephen
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Reptilia
Squamata
Gekkonidae
Hemidactylus

description: Hemidactylus Murrayi Gleadow, 1887
Hemidactylus murrayi was described from a series of 24 specimens (8 males, 16 females) collected from two adjacent localities “Pimpri and Garvi, in the “Dangs”, …., at the northwest extremity of the Syhadri or Ghat range, between Khandesh and Surat ” (Gleadow 1887). Pimpri (ca. 20°47’55”N, 73°35’00”E) and Garvi (ca. 20°50’12”N, 73°40’40”E) are two towns currently situated in the Dangs District of southeast Gujarat. Unfortunately, the repository of the syntypes was not mentioned in the description. Representative specimens are not currently present in BMNH or ZSI. Gleadow also did not deposit specimens in BNHM (Das & Chaturvedi 1998), despite his later plea on behalf of the society for lizard specimens (Gleadow 1905). The whereabouts of these type specimens have not been reported and are herein considered lost. In order to clarify the status of this species in the future, a topotypical specimen should be designated as a neotype. Based on the original description, this species appears morphologically to be a representative of the “rock-dwelling morph”, and thus very unlikely to be conspecific with H. brookii s.s.
description: Published as part of Mahony, Stephen, 2011, Taxonomic revision of Hemidactylus brookii Gray: a re-examination of the type series and some Asian synonyms, and a discussion of the obscure species Hemidactylus subtriedrus Jerdon (Reptilia: Gekkonidae), pp. 37-67 in Zootaxa 3042 on page 44, DOI: 10.5281/zenodo.278832
Hemidactylus Tenkatei Lidth
Creators: name: Mahony, Stephen
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Reptilia
Squamata
Gekkonidae
Hemidactylus

description: Hemidactylus Tenkatei Lidth de Jeude, 1895
Hemidactylus tenkatei was described based on two unsexed adults and one juvenile specimen presented to the Leyden Museum, The Netherlands (currently RMNH) by Dr. H. ten Kate, from “Rotti” (Lidth de Jeude 1895). The description, though detailed, is inadequate to identify the species affinities within H. brookii s.l. The type specimens are still present in the collection of RMNH (RMNH 4353––three specimens; Ronald de Ruiter in litt. 2010). Rösler and Glaw (2010) provide a description of two topotypes, which they used to elevate this nomen as a valid species. Photographs provided to me of the two adult syntypes of H. tenkatei reveal several important characters that diagnose this species from all but one other H. brookii group taxa redefined herein. Subcaudal scales completely transverse the tail width at approx. 50% of the original tail length and enlarged femoral scales of one of the female specimens indicate seven enlarged scales on each thigh separated medially by six smaller scales. These enlarged scales likely represent the enlarged pore-bearing scales of males, separated by a diastema of non-porebearing scales. These diagnosing characters are thus far only confirmed on H. subtriedroides. One H. “ brookii ” specimen (BMNH 1926.10.30.46) from West Timor, nearby Palau Roti, was examined in this study and is indistinguishable from H. subtriedroides. Additionally, I can find no characters from the Rösler and Glaw (2010) description of H. tenkatei topotypes that distinguish this species from Hemidactylus subtriedroides, thus based on morphological evidence I here consider the two as one and the same species. The revalidation of H. tenkatei by Rösler & Glaw (2010), disqualifies the possible retention of the more commonly used name H. subtriedroides as a nomen protectum (Article 23.9.1.2, 23.9.2). Chronologically H. tenkatei has nomenclatural priority over H. subtriedroides, and thus Hemidactylus subtriedroides Annandale, 1905 is here considered a junior subjective synonym of Hemidactylus Tenkatei Lidth de Jeude, 1895. The subsequent taxonomic section for this species is based on the morphological examination of H. subtriedroides types and other material as the syntypes of H. tenkatei were not directly examined in this study.
description: Published as part of Mahony, Stephen, 2011, Taxonomic revision of Hemidactylus brookii Gray: a re-examination of the type series and some Asian synonyms, and a discussion of the obscure species Hemidactylus subtriedrus Jerdon (Reptilia: Gekkonidae), pp. 37-67 in Zootaxa 3042 on page 44, DOI: 10.5281/zenodo.278832
Gecko Tytleri Tytler 1865, s.l.
Creators: name: Mahony, Stephen
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Squamata
Gekkonidae
Gecko

description: Gecko Tytleri Tytler, 1865
The name Gecko Tytleri was coined based on living animals in the private collection of the original author, collected from “dark cellars at Moulmein [= Mawlamyaing, Mon State, Myanmar] where the species is common”. The author stated that he had no means of comparing his animals with those already described by other taxonomists, but intended to provide each species a new name regardless of this fact. Unusually despite clearly indicating one of his species was conspecific with Phelsuma andamanense Blyth, Tytler still unjustifiably created the nomen Gecko chameleon indicating a lack of knowledge or respect for the basic fundamentals of nomenclatural classification followed by others of that period. Indeed all nine “new” species described in this paper are currently considered synonyms. Furthermore, he disregarded the generic classifications followed by others, which he was clearly aware of, and simply placed all “new” species of geckos in his collection in the genus Gecko.
The species Gecko tytleri was very briefly described and included only characters distinguishing it from other geckos in his collection. Due to the lack of diagnostic characters in the original description, several problems are associated with the designation of a topotype as a neotype nearly 150 years later. Because the specimen was collected in the cellars of buildings, thus associated with an anthropogenically modified habitat, the species to which the nomen Gecko tytleri was originally intended may not be a native species, but a representative of an introduced population from literally any part of the H. brookii s.l. range. Furthermore this population/species may or may not still be extant at this locality. It may even have been displaced by another more competitive introduced species of the H. brookii species group. I have not been able to locate any specimens deposited by Tytler from the type locality of this species in the collections of ZSI or BMNH, therefore it is safe to assume that type material does not exist for this nomen. Tytler’s name remains available, however, since this species has not been considered valid by any authors after 1899, a future neotype designation would not threaten the status of H. tenkatei (the only H. brookii group taxa observed in this study from Myanmar) as it would fail to comply with Article 23.9.2 of the Code.
description: Published as part of Mahony, Stephen, 2011, Taxonomic revision of Hemidactylus brookii Gray: a re-examination of the type series and some Asian synonyms, and a discussion of the obscure species Hemidactylus subtriedrus Jerdon (Reptilia: Gekkonidae), pp. 37-67 in Zootaxa 3042 on page 44, DOI: 10.5281/zenodo.278832
Hemidactylus brooki subsp. parvimaculatus Deraniyagala 1953
Creators: name: Mahony, Stephen
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Reptilia
Squamata
Gekkonidae
Hemidactylus
Hemidactylus brooki
Hemidactylus brooki parvimaculatus deraniyagala, 1953

description: Hemidactylus brooki parvimaculatus Deraniyagala, 1953
Hemidactylus brooki [sic] parvimaculatus was described based on a male type specimen (RG. 15) collected from “Colombo [Ceylon]” (= Sri Lanka), deposited at the Colombo National Museum (NMSL). It is not clear from the brief description the basis for which this population was considered a subspecies of H. brookii as all of the characters stated fell within the then accepted morphological variation of H. brookii from elsewhere in its range. Amarasinghe et al. (2009) provide a figure of the holotype. Wickramsinghe & Somaweera (2005) provide a photograph of a live individual and referred all populations of H. brookii from Sri Lanka to this subspecies. In this study, no specimens of the H. brookii complex from Sri Lanka were examined. In recent molecular phylogenetic and morphological analysis of Asian Hemidactylus, this subspecies was found to be distinct from other populations of H. “ brookii ”, and thus elevated to specific status (Bauer et al. 2010a, 2010b; Rösler & Glaw 2010). The statistical analysis of Rösler & Glaw (2010) found that the significant characters for differentiating their samples of H. parvimaculatus from Nepal H. “ brookii ” were number of lamellae on digit IV of pes and precloacal-femoral pores. Based on the redescription of H. brookii s.s. provided here, these characters completely overlap, but the two species differ slightly in other characters (see “Comparisons” section below).
description: Published as part of Mahony, Stephen, 2011, Taxonomic revision of Hemidactylus brookii Gray: a re-examination of the type series and some Asian synonyms, and a discussion of the obscure species Hemidactylus subtriedrus Jerdon (Reptilia: Gekkonidae), pp. 37-67 in Zootaxa 3042 on page 45, DOI: 10.5281/zenodo.278832
Hemidactylus mahendrai Shukla 1983, s.l.
Creators: name: Mahony, Stephen
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Reptilia
Squamata
Gekkonidae
Hemidactylus
Hemidactylus mahendrai

description: Hemidactylus mahendrai Shukla, 1983
This species was described based on four adult females and one juvenile collected from the “suberbs [sic] of Kanpur of Uttar Pradesh” (Shukla 1983). The diagnosis of the new species from congeners was based on the general descriptions of Smith (1935). The comparison poorly diagnosed this species from H. brookii s.l., and it has been regarded by recent authorities to be a synonym pending a review of the species based on examination of specimens (Zug et al. 2007; Bauer et al. 2010b; Rösler & Glaw 2010). Regarding this present review, it would not be possible to assign the name to the synonymy of any of the species dealt with here without examining topotype material. If considering the defining characters used in that description, e.g., high lamellae counts and geographical proximity, this species may be a synonym of H. kushmorensis, but I here retain it provisionally within the synonymy of H. brookii s.l..
description: Published as part of Mahony, Stephen, 2011, Taxonomic revision of Hemidactylus brookii Gray: a re-examination of the type series and some Asian synonyms, and a discussion of the obscure species Hemidactylus subtriedrus Jerdon (Reptilia: Gekkonidae), pp. 37-67 in Zootaxa 3042 on page 45, DOI: 10.5281/zenodo.278832
Hemidactylus luzonensis Taylor 1915
Creators: name: Mahony, Stephen
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Reptilia
Squamata
Gekkonidae
Hemidactylus
Hemidactylus luzonensis

description: Hemidactylus luzonensis Taylor, 1915
Hemidactylus luzonensis was described based on a single female specimen (No. 1774) collected from Manila, Luzon Island, Philippines and deposited at the Bureau of Science, Manila (Taylor 1915). This specimen has not since been described in literature and was reported as destroyed during World War II (Brown & Alcala 1978). Taylor (1915) provided a detailed description, however it is not possible to verify its affinities amongst the synonyms of H. brookii s.l., and thus would require the designation of a neotype to resolve its taxonomic status. No specimens of the H. brookii complex from the Philippines were available in this study. Brown and Alcala (1978) consider this species to be identical to the Bornean types of H. brookii, which at the time incorrectly included the “ Australia ” syntype, referred here to H. tenkatei. Rösler and Glaw (2010) suspect that this synonym may represent a junior subjective synonym of H. tenkatei. Herein I provisionally retain it in the synonymy of H. brookii pending a neotype designation.
description: Published as part of Mahony, Stephen, 2011, Taxonomic revision of Hemidactylus brookii Gray: a re-examination of the type series and some Asian synonyms, and a discussion of the obscure species Hemidactylus subtriedrus Jerdon (Reptilia: Gekkonidae), pp. 37-67 in Zootaxa 3042 on page 45, DOI: 10.5281/zenodo.278832
Hydrellia Robineau-Desvoidy
Creators: name: Júnior, Francisco De Assis Rodrigues
name: Mathis, Wayne Nielsen
name: Couri, Márcia Souto
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Ephydridae
Hydrellia

description: Genus Hydrellia Robineau-Desvoidy
Hydrellia Robineau-Desvoidy 1830: 790. Type species: Hydrellia communis Robineau-Desvoidy 1830 (= Notiphila griseola Fallén), subsequent designation, Duponchel in d'Orbigny 1845: 743. Cresson 1947: 37 –39 (review, Neotropical species). Wirth 1968: 13 (Neotropical catalog). Zatwarnicki 1988 (world catalog). Mathis & Zatwarnicki 1995: 61 –96 (world catalog). Deonier 1998: 1 –354 (revision of North American species).
Diagnosis. Hydrellia is distinguished from other genera of Hydrelliini by the following combination of characters: body frequently microtomentose, especially face. Head: ocellar setae smaller than pseudopostocellar setae; frontoorbital setae 2, anterior seta proclinate, posterior seta reclinate to lateroreclinate, sometimes with a third, much smaller proclinate seta. Eye bearing numerous interfacetal setulae. Face planate to shallowly carinate. Thorax: dorsocentral setae 2–3 (0+2, 1+2); acrostichal setulae usually in rows; postsutural supra-alar seta usually short, not longer than posterior notopleural seta; scutellar setae 3, middle pair much shorter; notopleural setae 2, these inserted at more or less the same level. Wings: hyaline; costa extended to vein M. Midtibia lacking dorsal, spinelike setae. Abdomen: dorsum comprising tergites 1–5, with tergites 1–2 somewhat fused; sternite 1 greatly reduced or lacking, 2–5 present; male 5th sternite variable in shape but usually U-shaped with posteriorly extended lobes from lateral arms modified and called “copulobi” (Deonier 1998: 15).
description: Published as part of Júnior, Francisco De Assis Rodrigues, Mathis, Wayne Nielsen & Couri, Márcia Souto, 2014, Hydrellia Robineau-Desvoidy (Diptera: Ephydridae) from Brazil with an emphasis on the faunas from the states of Parana and Rio de Janeiro, pp. 501-541 in Zootaxa 3753 (6) on page 503, DOI: 10.11646/zootaxa.3753.6.1, http://zenodo.org/record/226733
Stygiiulus seewaldi Vagalinski & Borissov & Bobeva & Canciani & Antić 2022, comb. nov.
Creators: name: Vagalinski, Boyan
name: Borissov, Simeon
name: Bobeva, Aneliya
name: Canciani, Giacomo
name: Antić, Dragan Ž.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Julida
Julidae
Stygiiulus
Stygiiulus seewaldi

description: Stygiiulus seewaldi (Strasser, 1967) comb. nov.
Figs 10E, 13
Alpityphlus seewaldi Strasser, 1967: 146–150, figs 4–9.
Typhloiulus seewaldi – Fritsch 1998: 149–150, 16. — Vagalinski et al. 2015: 346.
Diagnosis
A species of Stygiiulus stat. nov. with normal mouthparts. Easily distinguishable from congeners by its specific gonopod conformation (Fig. 10E) including a narrow, pointed velum considerably exceeding the solenomere and bearing a small, pointed, distal outgrowth (do) (the latter structure present also in S. insularis comb. nov. and in (some specimens of) S. tobias comb. nov.), in combination with the complete absence of a posterior hump on the opisthomere. Also outstanding by the small and rounded internal lobe of the promere, which is positioned close to promeral base.
Distribution
Germany, Upper Bavaria, Berchtesgadener Land, Untersberg Massif, Cave Hollerloch (the type locality); Austria, Dachstein Mts, tunnel Warmwasserstollen near Lake Hallstatt and Obere Brangrabenhöhle Cave (Fritsch 1998) (Fig. 13, orange squares).
Remark
Fritsch (1998) listed this species under the genus Typhloiulus, relying on J.-P. Mauriès’ opinion expressed in a letter to him. According to Mauriès who studied material of seewaldi, the erection of the genus Alpityphlus by Strasser (1967) was unfounded, the species being a member of Stygiiulus, at that time a subgenus of Typhloiulus. A similar view was expressed by Antić et al. (2017, 2018), but without a formal transfer to Stygiiulus. Based on the detailed original drawings we fully agree on that view, hence the new name combination is formalized here.
description: Published as part of Vagalinski, Boyan, Borissov, Simeon, Bobeva, Aneliya, Canciani, Giacomo & Antić, Dragan Ž., 2022, The mostly cavernicolous millipede genus Stygiiulus Verhoeff, 1929, stat. nov.: taxonomy, distribution and phylogenetic relationships (Diplopoda, Julida, Julidae), pp. 30-69 in European Journal of Taxonomy 798 on pages 53-55, DOI: 10.5852/ejt.2022.798.1669, http://zenodo.org/record/6323002
Stygiiulus maximus Vagalinski & Borissov & Bobeva & Canciani & Antić 2022, comb. nov.
Creators: name: Vagalinski, Boyan
name: Borissov, Simeon
name: Bobeva, Aneliya
name: Canciani, Giacomo
name: Antić, Dragan Ž.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Julida
Julidae
Stygiiulus
Stygiiulus maximus

description: Stygiiulus maximus (Verhoeff, 1929) comb. nov.
Fig. 10D
Mesoporoiulus maximu s Verhoeff, 1929: 19–20, fig. 2.
Typhloiulus (Stygiiulus) maximus – Verhoeff 1930: 9, 12–13, figs 1–2. — Manfredi 1932: 81.
Typhloiulus (Mesoporoiulus) maximus – Pretner & Strasser 1931: 87.
Typhloiulus maximus var. longicauda Strasser, 1962: 59–60, fig. 71.
Typhloiulus maximus var. maximus – Strasser 1962: 59, fig. 72.
Typhloiulus maximus – Attems 1949: 145. — Strasser 1971a: 14. — Vagalinski et al. 2015: 342–343.
not Typhloiulus tobias – Attems 1927: 250–251, figs 352–354.
Diagnosis
A species of Stygiiulus stat. nov. with normal mouthparts. Distinguishable from congeners by the combination of certain gonopodal characters (Fig. 10D), viz., a mostly straight pro- and mesomere, and an opisthomere with a faint and blunt posterior hump, a marginally broad (not tapering) and deeply serrated velum, and a solenomere with both the anterior and the posterior solenomeral branch being well developed and clearly discernible.
Distribution
This species has the widest distribution of all representatives of the genus Stygiiulus stat. nov. Known from numerous caves, as well as epigean habitats, ranging from the Julian Alps in the east, across the southern parts of the Carnic Alps Range, through the Venetian Prealps, all the way to the Piave River in the west (Fig. 13, pink squares).
Remark
Attems (1927: 250, 251, figs 352–354) gave a short description and drawings of what he thought was already described as S. tobias comb. nov. from Monte Cavallo (Lombardy). Just two/three years later, Verhoeff (1930) described another blind julid, S. maximus comb. nov., from a cave in the same area. What is evident from the gonopod drawings of both Attems and Verhoeff alone – that is that Attems’ (1927) record actually refers to S. maximus comb. nov. – was already confirmed by Strasser (1962: 60) based on re-examination of the specimens from Monte Cavallo.
description: Published as part of Vagalinski, Boyan, Borissov, Simeon, Bobeva, Aneliya, Canciani, Giacomo & Antić, Dragan Ž., 2022, The mostly cavernicolous millipede genus Stygiiulus Verhoeff, 1929, stat. nov.: taxonomy, distribution and phylogenetic relationships (Diplopoda, Julida, Julidae), pp. 30-69 in European Journal of Taxonomy 798 on page 48, DOI: 10.5852/ejt.2022.798.1669, http://zenodo.org/record/6323002
Leptoiulus trilineatus
Creators: name: Vagalinski, Boyan
name: Borissov, Simeon
name: Bobeva, Aneliya
name: Canciani, Giacomo
name: Antić, Dragan Ž.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Julida
Julidae
Leptoiulus
Leptoiulus trilineatus

description: Leptoiulus cf. trilineatus (C.L. Koch, 1847)
Fig. 11E
Material examined
BULGARIA • 1 ♀; Belasitsa Mtn, Castanea sativa forest; 3–10 May 2010; B. Georgiev and H. Delchev leg.; pitfall trap; IBER.
Descriptive notes
Vulva (Fig. 11E). Vulva of nearly equal width in both the sagittal and the transverse planes, mostly symmetric; bursa with a deep and rather broad median cleft; evenly setose throughout, without distinct rows; operculum (op) with markedly concave apical margin, exceeding bursa by ca ¼ of total height of vulva, distally with several long setae. Receptaculum seminis consisting of a very long and thin, somewhat bent, mesal tube (mt) ending in a medium-sized, piriform ampula (ma), and a somewhat ticker and mostly straight lateral tube (lt) forming a large, strongly elongate ampulla (la) at bottom.
description: Published as part of Vagalinski, Boyan, Borissov, Simeon, Bobeva, Aneliya, Canciani, Giacomo & Antić, Dragan Ž., 2022, The mostly cavernicolous millipede genus Stygiiulus Verhoeff, 1929, stat. nov.: taxonomy, distribution and phylogenetic relationships (Diplopoda, Julida, Julidae), pp. 30-69 in European Journal of Taxonomy 798 on pages 58-59, DOI: 10.5852/ejt.2022.798.1669, http://zenodo.org/record/6323002
Serboiulus spelaeophilus Gulicka 1967
Creators: name: Vagalinski, Boyan
name: Borissov, Simeon
name: Bobeva, Aneliya
name: Canciani, Giacomo
name: Antić, Dragan Ž.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Julida
Julidae
Serboiulus
Serboiulus spelaeophilus

description: Serboiulus spelaeophilus Gulička, 1967
Fig. 11F
Material examined
BULGARIA • 1 ♀ topotype; Belogradchik District, Dolni Lom, Vodni Pech Cave; 25 Feb. 2000; B. Petrov leg.; guano-clay; NMNHS.
Descriptive notes
Vulva (Fig. 11F). Strongly compressed in the sagittal plane, mostly symmetric; bursa with a shallow and very narrow median cleft; each valve distolaterally with a few setae; operculum (op) with a deeply concave apical margin, just slightly higher than bursa, distally bearing long setae. Receptaculum seminis consisting of two similarly narrow and twisted tubes – a lateral (lt) and a mesal one (mt), each forming an ovoid ampulla at bottom, the lateral one (la) being somewhat larger than the mesal one (ma).
description: Published as part of Vagalinski, Boyan, Borissov, Simeon, Bobeva, Aneliya, Canciani, Giacomo & Antić, Dragan Ž., 2022, The mostly cavernicolous millipede genus Stygiiulus Verhoeff, 1929, stat. nov.: taxonomy, distribution and phylogenetic relationships (Diplopoda, Julida, Julidae), pp. 30-69 in European Journal of Taxonomy 798 on page 59, DOI: 10.5852/ejt.2022.798.1669, http://zenodo.org/record/6323002
Typhloiulus (Inversotyphlus) lobifer Attems 1951
Creators: name: Vagalinski, Boyan
name: Borissov, Simeon
name: Bobeva, Aneliya
name: Canciani, Giacomo
name: Antić, Dragan Ž.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Julida
Julidae
Typhloiulus
Typhloiulus lobifer

description: Typhloiulus (Inversotyphlus) lobifer Attems, 1951
Fig. 11G
Material examined
CROATIA • 1 ♀; island of Brač, Škrip, Minjera-Bauxite Mine; 11 Sep. 2015; D. Antić leg.; D. Antić ded. 2021; IBER.
Descriptive notes
VULVA (Fig. 11G). Slender, of nearly equal width in both the sagittal and the transverse planes, slightly asymmetric: lateral valve being somewhat higher than mesal one, as well as more densely setose; bursa with a broad and relatively deep median cleft; several, mostly vertically arranged setae on each valve; operculum (op) with narrowly rounded apical margin, exceeding bursa by ca 2 ⁄ 7 of total height of vulva, distally with a dense submarginal row of setae each side. Receptaculum seminis consisting of a long, thin and twisted mesal tube (mt) ending in a lemon-shaped ampula (ma), and a somewhat ticker, nearly straight, lateral tube (lt), not forming an ampulla at bottom.
description: Published as part of Vagalinski, Boyan, Borissov, Simeon, Bobeva, Aneliya, Canciani, Giacomo & Antić, Dragan Ž., 2022, The mostly cavernicolous millipede genus Stygiiulus Verhoeff, 1929, stat. nov.: taxonomy, distribution and phylogenetic relationships (Diplopoda, Julida, Julidae), pp. 30-69 in European Journal of Taxonomy 798 on page 59, DOI: 10.5852/ejt.2022.798.1669, http://zenodo.org/record/6323002
Ectatomminae Emery 1895
Creators: name: Camacho, G P
name: Franco, W
name: Branstetter, M G
name: Pie, M R
name: Longino, J T
name: Schultz, T R
name: Feitosa, R M
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae

description: Ectatomminae Emery
= Heteroponerinae Bolton new synonym
Diagnosis: Presenting the characters of ‘poneromorph’ subfamilies described by Bolton (2003: p.40). Clypeus broadly inserted between frontal lobes (Bolton 2003); anterior clypeal margin with a narrow lamellar apron (Bolton 2003). Torulus not completely fused to frontal lobe (Bolton 2003). Antenna with 12 segments (Bolton 2003). Pronotum with the humeral corners angled, forming a distinct delimitation between the anterior and lateral margins (Lattke 2004). Antero-ventral angle of pronotum triangular (Feitosa 2015). Pretarsus without arolium (Lattke 2004). Petiole pedunculate (Keller 2011). Petiolar node as wide or wider than long (Feitosa 2015). Subpetiolar process very well developed, occupying more than onethird of the ventral portion of the petiolar sternite (Feitosa 2015). Helcium projecting from about midheight of the anterior face of abdominal segment III. Abdominal segment IV presclerites separated from the rest of segment by a constriction or slight thickening (Lattke 1994). Fourth abdominal tergite arched and larger than the sternite, giving the segment a curved appearance (Keller 2011).
Tribes: Ectatommini and Heteroponerini
Incertae sedis: † Canapone, † Electroponera, † Pseudectatomma.
Notes: In spite of the reciprocal monophyly of the subfamilies Ectatomminae and Heteroponerinae, the morphological evidence strongly suggests that all ectaheteromorph genera could be combined into a single subfamily Ectatomminae, which is the oldest available name. Ectatomminae, as defined here, presents a combination of 10 diagnostic characters that can be used to differentiate those ants from any other ant subfamily, making the identification of those groups more accessible.
description: Published as part of Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R & Feitosa, R M, 2022, UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus, pp. 6 in Insect Systematics and Diversity 6 (1) on pages 9-10, DOI: 10.1093/isd/ixab026, http://zenodo.org/record/5907609
Ectatommini Emery 1895
Creators: name: Camacho, G P
name: Franco, W
name: Branstetter, M G
name: Pie, M R
name: Longino, J T
name: Schultz, T R
name: Feitosa, R M
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae

description: Tribe Ectatommini Emery
= Stictoponerini Arnol’di
= Typhlomyrmecini Emery new synonym
Diagnosis (Females): Ectatommine ants of small to large size (head width 0.44-2.84mm, head length 0.56-3.8mm). Antennal scrobe usually absent. Eye absent to well-developed (Bolton 2003). Acetabulum of antennal socket apparatus spherical (Keller 2011); accessory chamber of antennal socket present (Keller 2011). Labial palp with two palpomeres (Keller 2011). Promesonotal suture fused and immobile to complete and flexible (Bolton 2003). Ventral flap on metapleural gland opening present (Keller 2011). Metacoxal cavity open (Bolton 2003). Petiolar sternite fused with tergite over its entire length (except in Rhytidoponera) (Bolton 2003, Keller 2011); laterotergites of petiole indistinct to absent.
Genera: Alfaria status revived, Ectatomma, Gnamptogenys, Holcoponera status revived, Poneracantha status revived, Rhytidoponera, Stictoponera status revived, and Typhlomyrmex.
description: Published as part of Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R & Feitosa, R M, 2022, UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus, pp. 6 in Insect Systematics and Diversity 6 (1) on page 10, DOI: 10.1093/isd/ixab026, http://zenodo.org/record/5907609
Alfaria Emery, status revived
Creators: name: Camacho, G P
name: Franco, W
name: Branstetter, M G
name: Pie, M R
name: Longino, J T
name: Schultz, T R
name: Feitosa, R M
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Alfaria

description: Alfaria Emery status revived
= Opisthoscyphus Mann new combination
Type Species: Alfaria simulans Emery
Diagnosis (Females): Head subquadrate; occipital lobe usually present; frontal carina broadly expanded laterad; row of stout setae on base of foretarsus opposite to strigil present; promesonotal suture absent to lightly impressed, never interrupting dorsal mesosomal sculpture; petiolar spiracle facing directly ventrad and sunken within a pit; second gastral (IV abdominal) sternite usually strongly reduced, so that the gaster is directed ventrally and anterad.
Species: caelata new combination, falcifera new combination, fieldi new combination, minuta revived combination, petiscapa new combination, piei new combination, simulans revived combination, striolata revived combination, and vriesi new combination (and the junior synonyms soror new combination, carinata revived combination, emeryi revived combination, mus revived combination, panamensis revived combination, pneodonax new combination, scabrosus new combination, and bufonis revived combination).
Distribution: Exclusively Neotropical, from southern Mexico to northern Argentina.
Notes: Alfaria is a very morphologically distinct lineage among the Ectatommini, given the extreme anterior curvature of the gaster in profile. In fact, these ants are usually mistakenly identified as Proceratium Roger, 1863, due to the impressive convergence in this character. We here resurrect the name Alfaria, firstly proposed by Emery (1896) and synonymized under Gnamptogenys by Brown (1958), to comprise the species previously included in the minuta group of Gnamptogenys sensu Brandão and Lattke (1990). All Alfaria species can be identified using the work of Camacho et. al. (2020) under the previous combination in Gnamptogenys.
description: Published as part of Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R & Feitosa, R M, 2022, UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus, pp. 6 in Insect Systematics and Diversity 6 (1) on page 10, DOI: 10.1093/isd/ixab026, http://zenodo.org/record/5907609
Ectatomma Smith
Creators: name: Camacho, G P
name: Franco, W
name: Branstetter, M G
name: Pie, M R
name: Longino, J T
name: Schultz, T R
name: Feitosa, R M
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Ectatomma

description: Ectatomma Smith
Type Species: Ectatomma tuberculatum (Olivier)
Diagnosis (Females): Occipital lobe absent. Antennal club absent. Palp formula 2,2. Pronotum usually with two or three tubercles. Mesonotum prominent and clearly differentiated from propodeum, separated by a deep transverse suture. Promesonotal suture well marked, interrupting or not the dorsal mesosomal sculpture. Propodeal spiracle elliptical or slit-shaped and separated from the declivous face of propodeum by a distance longer than its diameter. Apex of protibia with a stout seta close to the strigil base; dorsum of posterior coxa without projections.
Species: brunneum, confine, edentatum, gibbum, goninion, † gracile, lugens, muticum, opaciventre, parasiticum, permagnum, planidens, ruidum, suzanae, tuberculatum, and vizottoi.
Distribution: Exclusively found in the New World, from USA (Texas) to Argentina (Buenos Aires).
Notes: Ectatomma are among the most conspicuous elements of the ant fauna in Neotropical ecosystems. Currently, the most comprehensive work including an identification key for the species in the genus is the revision by Kugler and Brown (1982). However, this work does not include the species Ectatomma parasiticum Feitosa and Fresneau, in Feitosa et al. (2008), E. suzannae Almeida (1986) and E. vizottoi Almeida (1987).
description: Published as part of Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R & Feitosa, R M, 2022, UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus, pp. 6 in Insect Systematics and Diversity 6 (1) on page 10, DOI: 10.1093/isd/ixab026, http://zenodo.org/record/5907609
Gnamptogenys Roger 1863
Creators: name: Camacho, G P
name: Franco, W
name: Branstetter, M G
name: Pie, M R
name: Longino, J T
name: Schultz, T R
name: Feitosa, R M
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Gnamptogenys

description: Gnamptogenys Roger
= Commateta Santschi
= Emeryella Forel
= Tammoteca Santschi
Type Species: Gnamptogenys sulcata (Smith)
Diagnosis (Females): Head subquadrate to elongate. Mandible subtriangular to subfalcate. Occipital lobe absent. Antennal club absent. Palp formula 2,2 to 3,2. Pronotum unarmed and without tubercles. Promesonotal suture feebly impressed to absent, never interrupting dorsal mesosomal sculpture, sometimes with a small pit frequently situated medially on a weakly impressed promesonotal suture. Mesonotum not prominent, forming a continuous line with the propodeum, separated by a transverse suture. Propodeal spiracle oval or rounded, separated from the declivous face of propodeum by a distance longer or shorter than its diameter. Apex of protibia without a stout seta close to the strigil base; dorsum of posterior coxae frequently with a lobe or spine.
Species: acuminata, alfaroi, andersoni, annulata, biquetra, boliviensis, bruchi, † casca, concinna, continua, curvoclypeata, ericae, † europaea, falcaria, fernandezi, flava, haenschi, hartmani, horni, interrupta, kempfi, † levinates, lucaris, mordax, nana, † pristina, regularis, rimulosa, † rohdendorfi, rugimala, rumba, schmitti, siapensis, stellae, sulcata, tortuolosa, transversa, and volcano.
Distribution: Exclusively found in the New World, from USA (Texas) to Argentina (Buenos Aires), with one species occurring in Cuba.
Notes: In the new concept proposed here, Gnamptogenys is now restricted to the species from the previous sulcata, concinna, and mordax groups (sensu Lattke 1995), considering that G. sulcata is the type-species of the genus. All except one of the species of Gnamptogenys can be identified using the work of Camacho et. al. (2020). Gnamptogenys rugimala, a newly described species, can be identified using the paper by Marcineiro and Lattke (2020).
description: Published as part of Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R & Feitosa, R M, 2022, UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus, pp. 6 in Insect Systematics and Diversity 6 (1) on pages 10-11, DOI: 10.1093/isd/ixab026, http://zenodo.org/record/5907609
Rhytidoponera Mayr 1862
Creators: name: Camacho, G P
name: Franco, W
name: Branstetter, M G
name: Pie, M R
name: Longino, J T
name: Schultz, T R
name: Feitosa, R M
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Rhytidoponera

description: Rhytidoponera Mayr
= Chalcoponera Emery
Type Species: Rhytidoponera araneoides (Le Guillou)
Diagnosis (Females): Occipital lobe frequently present. Antennal club absent. Palp formula 2,2 to 3,2. Pronotum unarmed. Mesonotum not prominent, forming a continuous line with the propodeum, separated by a transverse suture. Promesonotal suture well marked, totally interrupting dorsal mesosomal sculpture. Propodeal spiracle oval or rounded, separated from the declivous face of propodeum by a distance longer than its diameter. Apex of protibia with a stout seta close to the strigil base; dorsum of posterior coxa without projections.
Species: abdominalis, acanthoponeroides, aciculata, aenescens, anceps, aquila, araneoides, arborea, aspera, atropurpurea, aurata, barnardi, barretti, borealis, carinata, celtinodis, cerastes, chalybaea, chnoopyx, clarki, confusa, convexa, cornuta, crassinodis, cristata, croesus, depilis, dubia, enigmatica, eremita, ferruginea, flavicornis, flavipes, flindersi, foreli, foveolata, fulgens, fuliginosa, † gibsoni, greavesi, gregoryi, haeckeli, hanieli, hilli, impressa, incisa, inops, inornata, insularis, † kirghizorum, koumensis, kurandensis, laciniosa, lamellinodis, laticeps, levior, litoralis, luteipes, maledicta, maniae, mayri, metallica, micans, mimica, mirabilis, nexa, nitida, nitidiventris, nodifera, nudata, numeensis, opaciventris, peninsularis, pilosula, pulchella, punctata, punctigera, punctiventris, purpurea, reflexa, reticulata, rotundiceps, rufescens, rufithorax, rufiventris, rufonigra, scaberrima, scabra, scabrior, socrus, spoliata, strigosa, subcyanea, tasmaniensis, taurus, tenuis, terrestris, trachypyx, turneri, tyloxys, versicolor, victoriae, violacea, viridis, † waipiata, wilsoni, and yorkensis.
Distribution: Exclusively Australasian.
Notes: This speciose ectatommine genus could be considered an ecological equivalent of Ectatomma in the Australian region. The most recent taxonomic tools for the identification of Rhytidoponera species include the papers by Ward (1980, 1984) and Heterick (2009).
description: Published as part of Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R & Feitosa, R M, 2022, UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus, pp. 6 in Insect Systematics and Diversity 6 (1) on page 12, DOI: 10.1093/isd/ixab026, http://zenodo.org/record/5907609
Poneracantha Emery, status revived
Creators: name: Camacho, G P
name: Franco, W
name: Branstetter, M G
name: Pie, M R
name: Longino, J T
name: Schultz, T R
name: Feitosa, R M
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Poneracantha

description: Poneracantha Emery status revived
= Barbourella Wheeler
= Parectatomma Emery
Type Species: Poneracantha bispinosa (Emery)
Diagnosis (Females): Head subquadrate or wider anterad than posterad in frontal view; anterior clypeal margin usually straight; frontal surface of mandible usually striate or rugulose; scape usually surpassing vertex; promesonotal suture feebly impressed to absent, never interrupting dorsal mesosomal sculpture; metanotal suture well impressed; propodeum usually armed with denticles or spines; petiolar node low; subpetiolar process shape variable, usually projecting anterad but sometimes subquadrate; metacoxal teeth generally present, usually acicular; second gastric segment slightly arched ventrally.
Species: banksi new combination, bispinosa revived combination, † brunoi, cuneiforma new combination, enodis new combination, ingeborgae new combination, insularis new combination, lanei new combination, laticephala new combination, lineolata new combination, lucaris new combination, mecotyle new combination, mediatrix new combination, menozzii revived combination, perspicax new combination, rastrata new combination, semiferox new combination, triangularis new combination, and wilsoni new combination (and the junior synonyms schubarti new combination, trigona new combination, aculeaticoxae new combination, and triangularis richteri new combination).
Distribution: Exclusively Neotropical, occurring from Guatemala to Uruguay, and in the Caribbean islands of Hispaniola and Lesser Antilles.
Notes: Here we revive the name Poneracantha from synonymy under Gnamptogenys to include all the species representing the previous rastrata group sensu Lattke (1995). All Poneracantha species can be identified using the work of Camacho et. al. (2020) under the previous combination in Gnamptogenys.
description: Published as part of Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R & Feitosa, R M, 2022, UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus, pp. 6 in Insect Systematics and Diversity 6 (1) on pages 11-12, DOI: 10.1093/isd/ixab026, http://zenodo.org/record/5907609
Typhlomyrmex Mayr 1862
Creators: name: Camacho, G P
name: Franco, W
name: Branstetter, M G
name: Pie, M R
name: Longino, J T
name: Schultz, T R
name: Feitosa, R M
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Typhlomyrmex

description: Typhlomyrmex Mayr
Type Species: Typhlomyrmex rogenhoferi Mayr
Diagnosis (Females): Head subquadrate; antennal club sometimes well-defined and formed by 3 or 4 segments; cephalic vertex mostly smooth and shining, sometimes presenting faded striae or rugulae; eye absent or reduced, with less than 15 ommatidia; promesonotal suture well marked, totally interrupting dorsal mesosomal sculpture; propodeal spiracle separated from declivity margin by a distance longer than its diameter; metacoxal dorsum unarmed or at most with a small lobe or denticle; petiole pedunculate, sometimes with a prominent anteroventral process.
Species: clavicornis, foreli, lavra new combination, lenis new combination, major, meire, prolatus, pusillus, reichenspergeri new combination, and rogenhoferi.
Distribution: Exclusively Neotropical, occurring from Mexico to Argentina (Buenos Aires).
Notes: Mostspecies of Typhlomyrmex can be identified usingthe key of Lacau et al. (2008), while T. lavra, T. lenis, and T. reichenspergeri (formerly included in the striatula group of Gnamptogenys) can be identified using Camacho et al. (2020).
description: Published as part of Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R & Feitosa, R M, 2022, UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus, pp. 6 in Insect Systematics and Diversity 6 (1) on pages 12-13, DOI: 10.1093/isd/ixab026, http://zenodo.org/record/5907609
Acanthoponera Mayr 1862
Creators: name: Camacho, G P
name: Franco, W
name: Branstetter, M G
name: Pie, M R
name: Longino, J T
name: Schultz, T R
name: Feitosa, R M
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Acanthoponera

description: Acanthoponera Mayr
Type Species Acanthoponera mucronata (Roger)
Diagnosis (Females): Ants of comparatively medium size (head width 0.90–1.61, head length 1.00–1.75). Mandible triangular. Palp formula 6,4. Frontal lobe reduced, only partially covering antennal insertions. Antennal club with four antennomeres. Antennal scrobe deeply impressed. Eye well-developed, with clear limits between ommatidia. Propodeum with a pair of welldeveloped spines. Tarsal claw with conspicuous preapical teeth and a basal lobe. Petiole with a long posterodorsal projection. Anterior face of abdominal segment III with an arched carina above the helcium.
Tribe Heteroponerini Bolton new combination
Diagnosis: Ectatommine ants of small to medium size (head width 0.42–1.61 mm, head length 0.53–1.75 mm); cephalic dorsum with a longitudinal carina extending from anterior margin of clypeus to posterior margin of head (Bolton 2003); antennal scrobe present (Bolton 2003); eye present; acetabulum of antennal socket apparatus hemispherical (Keller 2011); accessory chamber of antennal socket absent (Keller 2011); labial palp with three or four palpomeres (Bolton 2003, Keller 2011); promesonotal suture complete and flexible (Bolton 2003); ventral flap on metapleural gland opening absent (Keller 2011); metacoxal cavity closed (Bolton 2003); petiolar sternite articulated with tergite over its entire length (Bolton 2003, Keller 2011); laterotergites of petiole present.Genera: Acanthoponera,Aulacopone,Boltonia
new genus,Heteroponera.
Species: goeldii, minor, mucronata, and peruviana.
Distribution: Exclusively Neotropical, from southern Mexico to northern Argentina.
Notes: Acanthoponera represents a lineage of arboreal and nocturnal Neotropical ants. Most species can be identified using the work of Feitosa and Prada-Achiardi (2019) for the Colombian fauna.
description: Published as part of Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R & Feitosa, R M, 2022, UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus, pp. 6 in Insect Systematics and Diversity 6 (1) on pages 13-14, DOI: 10.1093/isd/ixab026, http://zenodo.org/record/5907609
Aulacopone Arnol'di
Creators: name: Camacho, G P
name: Franco, W
name: Branstetter, M G
name: Pie, M R
name: Longino, J T
name: Schultz, T R
name: Feitosa, R M
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Aulacopone

description: Aulacopone Arnol’di
Type Species: Aulacopone relicta Arnol'di
Diagnosis (Queens): Ants of comparatively medium size. Mandibles subfalcate. Median portion of clypeus modified, raised as a short, blunt triangular point projecting from the antennal insertions to the mandible. Frontal lobe expanded, extending from the clypeal posterior margin to the vertex. Antennal scrobe wide and deep.
Species: relicta.
Distribution: The only known specimens were collected in Azerbaijan in mountainous forests.
Notes: The genus is only known from two queens collected in Azerbaijan. The first specimen was collected in 1929 in Alazapin on the border with Iran, and later designated as the holotype by Arnol’di and deposited at the Zoological Institute of the Russian Academy of Sciences. The second specimen was collected in 1936, also by Arnol’di in the same country, in the region of Khachmaz, and later deposited in his personal collection at the Institute of Evolutionary Animal Morphology in Moscow. However, the holotype has been missing since 1979 and has not been examined for any study other than the original description. The second specimen was coated in gold-palladium for the study of its external morphology using scanning electron microscopy by Taylor (1980).
description: Published as part of Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R & Feitosa, R M, 2022, UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus, pp. 6 in Insect Systematics and Diversity 6 (1) on page 14, DOI: 10.1093/isd/ixab026, http://zenodo.org/record/5907609