Dendrocellus nigripennis Liang and Kavanaugh 2007, new species
Creators: name: Liang, Hongbin
name: Kavanaugh, David H.
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Dendrocellus
Dendrocellus nigripennis

description: Dendrocellus nigripennis Liang and Kavanaugh, new species   
(Figs. 17, 46, 63, 85–86)  
 Types. Holotype: a male (ISNB), ‘‘ Kina Balu, Borneo’ ’/‘‘ Collection P. Dupuis’ ’/‘‘ Coll. R. I. Sc. N. B.’’/‘‘ HOLOTYPE, Dendrocellus nigripennis Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006’’ [red label]. Paratypes (total 2 specimens): 1 male and 1 female (ISNB), ‘‘ Bornéo’ ’/ ‘‘ Collection P. Dupuis’ ’/‘‘ Coll. R. I. Sc. N. B.’’/‘‘ PARATYPE, Dendrocellus nigripennis Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006’’ [yellow label].   
 Type locality. Mount Kinabalu, Pulau Kalimantan.   
 Diagnosis. Elytra black with very slight blue luster, intervals sparsely punctate (Fig. 46); femora with apical half black, basal half brown; tibiae and tarsi black; protarsomere 3 symmetrical in male; tarsal claws with 4–5 pectinations, length of the longest pectination nearly half width of claw base (Fig. 63); aedeagus with lamella wide, apex truncate (Figs. 85–86).   
 Description. Length 9.2–11.0 mm, width 2.8–3.4 mm. Black, head and pronotum with blue luster; elytra with very slight blue luster, restricted to outer intervals in many specimens; tibiae, tarsi, apical half of femora, and apical fourfifths of Ant 1 black, Ant 2, Ant 3, and basal fifth of Ant 1 dark brown; Ant 4–11 and palpi brown. Head wider than pronotum, HW/PW 5 1.22 (1.21–1.23); vertex convex, coarsely punctate; labrum with medial lobe slightly protruded anteriorly; Ant 1/Ant 3 5 3.48 (3.40–3.56); genae slightly tumid; eyes large, about 2.4 times as long as genae. Pronotum coarsely punctate, PL/PW 5 1.65 (1.63–1.70); lateral margins strongly sinuate before basal angles; basal angles protruded laterally; basal foveae deep. Elytra short, EL/EW 5 1.66 (1.62–1.68), EW/PW 5 2.41 (2.33–2.58); lateral margins slightly emarginate in the basal fourth, widened posteriad to apical third, then narrowed to apex; outer angles slightly dentate, pointed; intervals convex, sparsely punctate, punctures small, mixed with some larger ones, interspace between two adjacent punctures 1.5–2.0 times diameter of punctures (Fig. 46). Protarsomere 3 symmetrical in male and female; tarsal claws with 4–5 pectinations, length of the longest pectination subequal to the width of claw base (Fig. 63). Aedeagus with apical lamella wide, apex truncate (Figs. 85– 86).   
 Distribution. Known only from the type locality.   
 Etymology. The new species is named for its black elytra.   
 Remarks. Initially, we assumed this to be just a black variety of D. crassus (Andrewes). However, the color of the femora and shape of the apex of the aedeagus (Fig. 85) are both very different from those of D. crassus members, so we treat this as a distinct species.
description: Published as part of Liang, Hongbin & Kavanaugh, David H., 2007, Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species, pp. 1-39 in The Coleopterists Bulletin (Oxford, England) (Oxford, England) 61 (1) on pages 1-39, DOI: 10.1649/934.1, http://zenodo.org/record/5461668