Landouria nusakambangensis Nurinsiyah & Neiber & Hausdorf 2019, sp. nov.
Creators: name: Nurinsiyah, Ayu Savitri
name: Neiber, Marco T.
name: Hausdorf, Bernhard
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Camaenidae
Landouria
Landouria nusakambangensis

description: Landouria nusakambangensis sp. nov.
urn:lsid:zoobank.org:act: 3F1A3273-1ECA-4D55-B350-AC9FE593D6AD
Figs 18, 32, 49, 52; Tables 3–4
Landouria rotatoria – van Benthem Jutting 1941: 310 (in part). (not Pfeiffer, 1842)
Diagnosis
Landouria nusakambangensis sp. nov. is characterized by a brownish-corneous, keeled shell, a penis with a small globular bulge at the proximal end and a flagellum consisting of a crenated distal part and a long tapering proximal part.
Etymology
Landouria nusakambangensis sp. nov. is named after its occurrence on Nusa Kambangan Island.
Material examined
Holotype
INDONESIA • holotype (det. anat.); Central Java, Nusa Kambangan Island, Nature Monument Gligir; 7°44′ S, 108°53′ E; 15–16 Feb. 1927; P.F. Franck leg.; D = 14.9 mm, H = 9.1 mm; ZMA 412298 a.
Paratypes
INDONESIA • 7 spec. (vouchers for van Benthem Jutting 1941); same data as for holotype; ZMA 412298 • 16 spec. (vouchers for van Benthem Jutting 1941); same data as for holotype; ZMA 410726 • 6 spec. (vouchers for van Benthem Jutting 1941); Nusa Kambangan Island; 7°44′ S, 108°53′ E; ZMA 407622.
Description
SHELL (Fig. 49; Tables 3–4). Depressed conical, with 5.25–6 hardly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines and a granular sculpture; scaly processes sparse, mainly at periphery; tuberculate around umbilicus; brownish-corneous; body whorl keeled; aperture rounded rhombic; upper insertion of peristome descending; peristome expanded, reflexed and thickened; umbilicus slightly eccentric, wide, comprising 32–40% of shell diameter, hardly obscured by columellar edge.
GENITALIA (Figs 18, 32; Table 4). Atrium short; penis long, becoming broader proximally; with a small globular bulge at proximal end; without penis coecum; membranaceous sheath around distal part of penis connected with proximal part of epiphallus by tissue; penial retractor runs from diaphragm to proximal part of cylindrical epiphallus; flagellum moderately long, broader distal part usually slightly crenated, proximal part tapering towards its proximal end, without node; vas deferens narrow and long; vagina long; oviduct short, broad, proximally tapering; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.
Remarks
Landouria nusakambangensis sp. nov. resembles L. winteriana, from which it differs in the slightly more distinct keel, the smaller globular bulge at the proximal end of the penis and a longer tapering proximal part of the flagellum. In shell shape and the keeled body whorl, L. nusakambangensis sp. nov. also resembles L. rotatoria, from which it differs in the smaller shell (D 11.9–15.0 mm vs 15.6–19.1 mm in L. rotatoria) with a wider umbilicus (U/D 0.32–0.40 vs 0.24–0.32 in L. rotatoria).
Landouria nusakambangensis sp. nov. occurs sympatrically with L. pakidulan sp. nov. (Fig. 85), from which it can easily be distinguished by the brownish, keeled shell.
Distribution
Landouria nusakambangensis sp. nov. is known only from Nusa Kambangan Island (Fig. 52).
description: Published as part of Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, pp. 1-73 in European Journal of Taxonomy 526 on pages 34-36, DOI: 10.5852/ejt.2019.526, http://zenodo.org/record/3152217