Chimarra devoensis Johanson & Oláh, 2012, new species
Creators: name: Johanson, Kjell Arne
name: Oláh, János
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Philopotamidae
Chimarra
Chimarra devoensis

description: Chimarra devoensis, new species
Figs. 26, 139–144
The male genitalia of this species resemble those of C. lavensis and C. vanuensis, particularly in the presence of a strongly dorsad-oriented dorsal branch of tergum X. In C. devoensis the dorsal process of tergum X is accompanied by a triangular, posterad oriented process between on the dorsal branches of tergum X. Segment IX has very long, anterolateral plates that are shorter in C. lavensis and C. vanuensis. In addition, the base of each gonopod is more strongly expanded ventrad compared to the other species. The species is easily distinguished from C. vanuensis by the absence of a large pale spot between Cu1 and end of Cu2 (arculus) in each forewing.
Male. Head and thorax greyish brown, dorsal part of meso- and metathorax dark brown. Foreleg anterior claw as long as foreleg spur.
Wings (Fig. 26). Forewings 5.1 mm (n=1), brown. Forewings broad, ratio of length to breadth 3.2; R1 nearly straight before crossvein r; radial sector slightly produced anterad immediately before discoidal cell; discoidal cell originating at mid-length of wing, nearly 3x longer than wide; median cell as long as discoidal cell; crossvein r originating from basis of fork I; fork I originating before crossvein s at distance equal to length of crossvein s; nygma located near base of fork II; fork III 1 /6th as long as wing; fork V shorter than fork II; Cu2 ending in wing margin well separated from A. Hind wings 4.2 mm (n=1), brown; ratio of length to breadth 3.0; margin weakly incurved where Cu1 and Cu2 fused with margin; fork I originating slightly before anterodistal corner of discoidal cell; fork III as long as discoidal cell and1/10th as long as wing; fork V slightly shorter than fork I; 1A+2Ab 3x as long as 1A.
Male genitalia (Figs. 139–144). Segment IX about 1.5x taller than long in lateral view; anterodorsally produced into long, narrow, rounded, anterodorsad oriented plate; anteroventral plate large, triangular, pointing posterad in lateral view; anterodorsal margin widely and deeply concave, U-shaped; ventral margin irregularly convex, incised at vertical apodeme; each posterior margin forming uniformly convex plate originating immediately below cercus; segment IX ventrally with setae restricted to narrow band before posterior margin (Fig. 141). Dorsal margin of segment IX undulating in lateral view; anterior margin straight, membranous in dorsal view. In ventral view segment IX with almost parallel lateral margins, slightly narrower at transverse apodeme; anterior margin concave; posterior margin straight, without central projection. Tergum X nearly as long as segment IX, divided into pair of lateral branches at basis; in lateral view each lateral branch with large, straight, dorsad oriented dorsal branch, short, widely triangular posterad orienting process below dorsal branch, and long tube-shaped, nearly straight ventral branch with 2 apical sensillae. In dorsal and ventral view, ventral processes oriented posterad, slightly diverging, tapering along their length. Cerci small, wart-like, located at posterior margin of segment IX at mid-height of tergum X and covered by long setae. Gonopods longer than length of segment IX; divided into very broad basal third and narrow distal 2/3rds in lateral view; distal 2/3rds nearly parallel-sided; each gonopod unbranched in lateral view; with well-developed dorsomesal lobe and ventromesal lobe of mesal process in ventral view. Anterodorsal margin of each gonopod basally convex, distally straight; posteroventral margin concave, with strongly expanded setal bases; ventral margin of basis strongly expanded ventrad; distal apex without megasetae. In ventral view, gonopods slightly broader at base than rest of gonopods, with undulating margins; gonopods weakly curved mesad along their length. Mesal branch of each gonopod divided into well-developed ventromesal and dorsomesal lobe; ventromesal lobe strongly produced dorsad, darkly pigmented, covered by minute setae; dorsomesal lobe curved ventrad, with pointed apex. Phallic apparatus slightly longer than rest of genitalia, straight along its length; phallotheca, in lateral view nearly 4x thicker than posterior part; in ventral view nearly 2x thicker than posterior part; apicoventral spine absent; phallotremal sclerite forming simple, 3-rayed structure in lateral view; in ventral view forming single posterad directed arrow head situated on basis with 4 anterad pointing spines; 4 nearly black, short, variously directed endothecal spines present, about as long as diameter of narrowest part of phallotheca; endothecal spicules pointing posterodorsad, present in group below phallotremal sclerite.
Female. Unknown.
Holotype male: TAVEUNI: Cakadrove Prov., Devo Peak, Malaise trap, 10–16.i.2003, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#37] [FNIC].
Etymology: Devoensis, after Devo Peak, the type locality of the species.
Distribution: Taveuni.
description: Published as part of Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, pp. 1-58 in Zootaxa 3354 on pages 46-47, DOI: 10.5281/zenodo.210736