Tomarus pumilus
Creators: name: López-García, Margarita M.
name: Deloya, Cuauhtémoc
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Dynastidae
Tomarus
Tomarus pumilus

description: Tomarus pumilus (Prell, 1937)
(Figs. 1K, 16N, 20E, 28E; 47)
Ligyrus pumilus Prell, 1937: 90. Original combination.
Male lectotype designated by Endrödi (1969: 71) (NHMB) “Corralitos // Ligyrus / pumilus / ♂ -Type // Lectotypus / Ligyrus / pumilus / Prell / Endrody // TOMARUS / PUMILUS / (PRELL) / det. B.C. Ratcliffe 2011”. Type locality: Corralito, Córdoba, Argentina.
Ligyrus rosettae Endrödi, 1968: 68. Synonym.
Male holotype (HNHM) “ Holotypus / Ligyrus / rosettae / Endr. // Proc. San Clemente del Tuyu Arg. / Prov. Bs. Aires, xii.49 // Museum Frey / München // KEYENCE 105”. Male paratype (HNHM) “Proc. San Clemente / del Tuyu Arg. / Prov. Bs. Aires. xii.49 // Paratypus / Ligyrus / rosettae / Endr.”. Female paratype (HNHM) “Mendoza / Rep. Arg. / v.26 // Chili / Faz ded. // Paratypus / Ligyrus / rosettae / Endr.” Male paratype (HNHM) “Mendoza / Rep. Arg. / v.26 // Chili / Faz ded. // Ligyrus / scarabaeinus Pertÿ // Paratypus / Ligyrus / rosettae / Endr.” Type locality: San Clemente del Tuyu, Buenos Aires, Argentina.
Description. Habitus as in Fig. 28E. Length 19.2–23.6 mm; humeral width 9.1–12.3 mm. Color dark reddish brown to black. Head: Frons coarsely rugose. Frontoclypeal region with 2 transverse tubercles (Fig. 1K), separated by about 3 tubercle diameters. Clypeus narrowed towards apex, base 3.8 wider than apex. Clypeal teeth triangular, separated by less than a tooth diameter. Mandible with 2 apical teeth and a lateral, narrowly rounded tooth. Mentum abruptly constricted at apical 3rd. Galea of maxilla with teeth 5 and 6 well developed. Interocular distance equal to 4.1 times an eye width. Pronotum: Surface with dense, large punctures. Apical tubercle small, rounded. Fovea shallow, striate, narrow (1/5 as wide as interocular distance). Scutellum: Surface with minute, sparse punctures. Elytra: First interval punctate, punctures similar in size to the other intervals; sutural stria complete. Inner surface of apex with rounded, small tubercles forming 38 parallel lines. Abdomen: Apex of tergite IV with a triangular wide area with large, elongate tubercles forming about 16 diagonal parallel lines. Pygidium with dense rugosity on basal 3rd; apex regularly rounded. Venter: Apex of prosternal process flat, longitudinally oval. Metasternum with short, sparse setae on anterior angles. Legs: Protibia tridentate with an additional inconspicuous, basal convexity. Metatibia triangular; apex slightly crenulate, with 21–27 spinules. Male genitalia: Spiculum gastrale without basal plates. Phallobase 1.1 times longer than parameres. Parameres with lateral teeth short, acute; apical half narrowed, apices outwards (Figs. 16N, 20E). Internal sac with copulatory lamella and short lamellar spiny belt; with a complex of 1–3 spine-like, accessory lamellae, with dense granules at base.
Diagnosis. Tomarus pumilus can be recognized by the following character combination: frontal tubercles transverse (Fig. 1K); pronotal fovea and tubercle inconspicuous (Fig. 28E); pronotum deeply and densely punctate; metasternum with sparse, minute setae on anterior angles; internal sac with a complex of 1–3 spine-like, accessory lamellae; and parameres with lateral teeth short, triangular, acute; apical half narrowed, apices outwards (Figs. 16N, 20E).
Distribution. Argentina.
Locality records (Fig. 47). 20 examined specimens from HNHM, NHUMB, and USNM. Some records from Neita-Moreno & Ratcliffe (2017). ARGENTINA (33). Buenos Aires (4): Buenos Aires; San Clemente del Tuyu; Villa Gesell. Catamarca (1): Catamarca. Córdoba (4): Corralito; Tanti. Mendoza (16): Mendoza; Pedregal; Potrerillos; Puente del Inca-Las Heras; San Rafael; Telteca-La Valle. Río Negro (5): Allen; General Roca. Salta (1): Urundel. San Juan (2): Jachal; San Juan.
Natural history. Adults are attracted to lights at night (Neita-Moreno & Ratcliffe 2017) and are mainly found between November and March.
description: Published as part of López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, pp. 1-119 in Zootaxa 5211 (1) on pages 68-69, DOI: 10.11646/zootaxa.5211.1.1, http://zenodo.org/record/7345007