Hemiteles yamatonis Watanabe, 2021, sp. nov.
Creators: name: Watanabe, Kyohei
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Hemiteles
Hemiteles yamatonis

description: Hemiteles yamatonis sp. nov.
(New SJN: Haraaka-mame-togari-himebachi)
(Figs 43A–E, 65O, 66M)
Etymology. The specific name is from the old name of Japan “Yamato”.
Type series. Holotype: KPM-NK 55032, F, JAPAN, Toyama Pref., Toyama City, Kamegai, 4–11. VIII. 2009, M. Watanabe et al. leg. (MsT). Paratypes: JAPAN, KPM-NK 55033, F, Hokkaido, Sapporo City, Maruyama, 29. VII. 2009, K. Watanabe leg.; KPM-NK 55034, F, Kanagawa Pref., Yamakita Vil., Nakagawa, 3. VII. 2013, K. Watanabe leg.; KPM-NK 55035, F, Yamanashi Pref., Koushu City, Yanagisawa-toge, 5. VIII. 2008, K. Watanabe leg.; KPM-NK 55036, F, Toyama Pref., Nanto City, Togamura, Kamimomose, 11–18. VIII. 2009, M. Watanabe et al. leg. (MsT).
Description. Female (n=5). Body length 4.4–4.7 (HT: 4.7) mm. Body densely coriaceous, covered with silver setae.
Head matt. Clypeus 2.0 times as wide as maximum length, its anterior margin rounded. MSL 1.0–1.1 (HT: 1.0) times as long as BWM. OOL almost as long as POL. OOL and POL longer than OD. Occipital carina complete, its lower end joined with hypostomal carina distant from mandibular base. Upper tooth of mandible slightly longer than lower tooth. Base of mandible flat. Antenna with 22–24 (HT: 24) flagellomeres. FL III 3.3 times as long as maximum depth in lateral view.
Mesosoma. Lateral part of pronotum covered with longitudinal striae ventrally, the interspace of striae polished. Epomia indistinct. Mesoscutum matt. Notaulus short, fading out in front of the center of mesoscutum. Scutellum polished and punctate, without a lateral longitudinal carina except for posterior half. Mesopleuron covered with longitudinal striae ventrally except for smooth speculum (Fig. 43D). Posterior transverse carina of mesosternum largely absent in front of mid coxae. Metapleuron with a complete juxtacoxal carina. Propodeal carinae complete (Fig. 65O). Area basalis distinct (Fig. 65O). Area superomedia distinct, transversely elongate, its length almost as long as area basalis (Fig. 65O). Postero-lateral corner of area dentipara weakly projected. Area petiolaris strongly elongated. Propodeum polished, partly irregularly, obliquely, and transversely rugulose, with a smooth area in the median part of area petiolaris. Length of fore wing 2.8–3.6 (HT: 3.6) mm. Areolet absent (Fig. 43E). Vein 2m-cu of fore wing with a single bulla. Nervellus inclivous, intercepted behind the middle (Fig. 43E). Hind femur 4.2–4.3 (HT: 4.2) times as long as maximum depth in lateral view. Hind TS I: II: III: IV: V = 2.0: 0.8–0.9 (HT: 0.9): 0.6: 0.3: 0.4. Tarsal claws simple.
Metasoma. T I 2.1–2.2 (HT: 2.2) times as long as maximum width, longitudinally striate. Median dorsal carina of T I absent. Dorsolateral carina of T I indistinct. T II to T VI wider than long, finely and densely punctate. Ovipositor sheath 0.5–0.56 (HT: 0.5) times as long as hind tibia. Ovipositor straight, with or without an obtuse nodus and with weak ventral teeth (Fig. 66M).
Coloration (Figs 43A–E). Body (excluding wings and legs) black to blackish brown. Scape, pedicel, and base of FL I reddish brown to brown. Clypeus more or less tinged with reddish brown. Mandible yellowish brown except for darkened apex and base. Palpi and tegula whitish yellow. Collar and antero-dorsal corner of mesopleuron tinged with reddish brown. Apex of T I, T II, and base of T III reddish yellow. Posterior margins of T III to T VI narrowly tinged with reddish brown. T VII whitish yellow. T III to T VII more or less tinged with brown. Membranous part of metasomal sternites whitish yellow. Ovipositor yellowish brown. Wings hyaline. Veins and pterostigma brown to dark brown. Legs yellowish brown to reddish brown. Fore and mid coxae, basal part of fore tibia, and all tibial spurs, trochanters and trochantelli whitish yellow to yellow. Hind femur, tibia except for base, and tarsus more or less darkened.
Male. Unknown.
Distribution. Japan (Hokkaido and Honshu).
Bionomics. Unknown.
Remarks. This species resembles H. similis (Gmelin, 1790), but it can be distinguished by the following combination of character states: hind coxa reddish brown (at least partly black in H. similis); mesosoma without red area (usually with red area in H. similis); ovipositor sheath 0.5–0.56 times as long as hind tibia (ca. 0.75 in H. similis). This species also resembles H. maculipterus sp. nov. but it can be distinguished by the above key.
description: Published as part of Watanabe, Kyohei, 2021, Taxonomic and Zoogeographic Study of the Japanese Phygadeuontinae (Hymenoptera, Ichneumonidae), with Descriptions of 17 New Species, pp. 55-136 in Bulletin of the Kanagawa Prefectural Museum (Natural Science) 50 on pages 104-106, DOI: 10.32225/bkpmnh.2021.50_55, http://zenodo.org/record/12774639