Caprobolus andringitra Wesener &amp; Enghoff &amp; Sierwald 2009, sp. n.
Creators: name: Wesener, Thomas
name: Enghoff, Henrik
name: Sierwald, Petra
Publisher: name: Zenodo
Year: 2009
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Spirobolida
Pachybolidae
Caprobolus

description: <i>Caprobolus andringitra</i> Wesener, sp. n. urn:lsid:zoobank.org:act: 331040CB-492C-483D-8D12-41B8265EB54D <b>Material examined:</b> 6 &amp;male;, 1 imm. <i>Holotype:</i> 1 &amp;male; (41 mm long), FMNH-INS 13601, Madagascar, Province Fianarantsoa, Andringitra reserve, camp 3, 40 km S Ambalavao, along Volotsangana River, rainforest, 1210 m, 22°13’22’’ S, 46°58’18’’ E, leg. S. Goodman, 5–6.XII.1993. <i>Paratypes:</i> 3 &amp;male;, 1 imm., FMNH-INS 13600, same data as holotype; 1 &amp;male;, FMNH-INS 13611, same data as holotype; 2 &amp;male;, FMMC 8219, RS Ivohibe, 900 m, rainforest, 22°25’ S, 46°56 ‘E, leg. S. Goodman, 7–12.X.1997. <b>Other material examined</b> : 1 &amp;male;, CASENT 9032814, Province Fianarantsoa, Forêt de Vevembe, 66.6 km 293°WNW Farafangana, rainforest in transition to montane rainforest, 600 m, 22°47’28” S, 47°10’55” E, leg. B. L. Fisher et al., yellow pan trap, 23.IV.2006 <b>Description.</b> <i>Measurements</i>: male holotype with 44 rings, circa 41 mm long, 3.2 mm wide. Male paratypes with 43 or 44 rings (specimen from Vevembe with 46), up to 44 mm long, up to 3.5 mm wide. <i>Coloration</i> faded in alcohol. Head, legs and antennae orange-red (Figs 39 A–D). Anal valves and subanal scale reddish, preanal process greyish (Fig. 39C). Metazonites red, remaining parts of body rings up to ozopore openings orange-reddish, more dorsally greyish. Specimens dorsally with a slender, orange line. <i>Head</i>: each eye with circa 24–28 ocelli arranged in 6 or 7 vertical rows (Fig. 39D). Labrum with standard three irregular teeth and a single row of 10–12 stout marginal setae. Clypeus with two setiferous foveolae on each side. Antennae long, protruding back to ring 6. Relative lengths of antennomeres: 1&lt;&lt;2=3=4=5=6, first antennomere very short (Fig. 40A). Terminal antennomere with four large sensory cones located together inside a membranous area (Fig. 40B). Antennomere 5 latero-apically with five rows (Fig. 40C), antennomere 6 with two rows (Fig. 40B) of sensilla basiconica. <i>Gnathochilarium</i> lamellae linguales each with two standard setae located behind one another (Fig. 39E). Stipites each with three apical setae. Palpi of gnathochilarium with numerous sensilla. Hypopharyngeal crest with a field of spine-like structures (Fig. 40D). Central pads of endochilarium separated into two areas, a group of 6–8 sensilla located apically (Fig. 40E). Close to endochilarium with a large group of sensilla, located towards hypopharyngeal crest. Mentum basally subdivided by a membranous suture (Fig. 39E). Stipites each with a sclerotized ledge (Fig. 39E) <i>Mandible</i> external tooth simple, rounded; mesal tooth with three cusps (Fig. 40F). Seven to nine dense pectinate lamellae (Fig. 40G). Intermediate area between pectinate lamellae and molar plate almost completely absent. Molar plate short, only slightly longer than pectinate lamellae, with only six or seven transverse furrows (Fig. 40F). <i>Collum:</i> smooth, not protruding as far as body ring 2 (Fig. 39A). <i>Body rings:</i> mostly smooth, with some irregular punctation and some transverse impressions on mesozona. Ozopores starting at ring 6, touching suture between mesozona and metazona. Ozopore on ring 6 located more ventrally than subsequent ozopores. <i>Telson</i>: anal valves with well-developed lips and micropunctation. Preanal ring and subanal process well-rounded, not extending beyond anal valves (Fig. 39C). <i>Legs</i>: coxae 1 and 2 elongated and fused with sternum, podomeres from prefemur to tarsus in both sexes each with 4–10 ventral/mesal setae. Length of midbody legs circa 1.0 times body diameter in males (Fig. 39F). Each podomere with a pair of setae ventrally. Coxae 3 and beyond of cylindrical shape. Tarsus with a stout dorso-apical seta and three pairs of setae, apical pair stout, others successively smaller (Fig. 39G). <i>Male sexual characters</i>: tarsal pads absent (Fig. 39G). Coxae 3 and 4 each with a short, conical coxal process. Process protruding posteriorly (Fig. 39B). <i>Anterior gonopod</i> sternite elongated into a wide, almost rectangular lobe. Mesal process of coxite prominent, longer than telopodite, but shorter than telopodite process (Fig. 41A). Telopodite on anterior side with a circular opening for the posterior gonopods (Fig. 41A). Telopodite process very long, slender, slightly curved inwards. Shape resembling a sable (Fig. 41B), basally projecting to lateral side into a long, slen- der, well-rounded, retrorse process. Retrorse process laterally extending beyond telopodite (Fig. 41B). <i>Posterior gonopods</i> unique (Figs 41 C–E). Gonopod pair connected via a wellsclerotized sternite. Coxites and telopodites positioned parallel to those of opposite gonopod. Coxite wide, well-sclerotized, with a single groove (Fig. 41C). Coxite and telopodite separated by a suture. Telopodite massive, longer than and as wide as coxite (Fig. 41C). Telopodite basally divided into two branches (Figs 41D, E). Both branches of similar width and almost similar length, resembling a U (Figs 41 C–E). Sperm canal protruding at base of U, running free but somehow attached to lateral main branch (Fig. 41E), tip prolonged into flagellum (Fig. 41D). Mesal branch of telopodite on anterior side medially with a finger-shaped process (Fig. 41D). <i>Female unknown.</i> <b>Etymology:</b> <i>andringitra</i>, noun in apposition, after the type locality, the Andringitra mountain massive.
description: Published as part of <i>Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra, 2009, Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda), pp. 1-128 in ZooKeys 19 (19)</i> on pages 76-77, DOI: 10.3897/zookeys.19.221, http://zenodo.org/record/576503