Hemisyntexis lepida Li, Wang, Rasnitsyn & Shih 2024, sp. nov.
Creators: name: Li, Yi
name: Rasnitsyn, Alexandr P.
name: Shih, Chungkun
name: Ren, Dong
name: Wang, Mei
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Anaxyelidae
Hemisyntexis
Hemisyntexis lepida

description: Hemisyntexis lepida Li, Wang, Rasnitsyn & Shih, sp. nov.
urn:lsid:zoobank.org:act: CE39453F-60D6-4BDD-9850-3B70BC411AD2
Material. Only the holotype, CNU-HYM-LB2024501.
Etymology. The specific name is from Latin “lepidus”, meaning exceptional and referring to the well-preserved fossil specimen.
Horizon and locality. Huangbanjigou, Chaomidian Village, Shangyuan Township, Beipiao City, Liaoning Province, China; Lower Cretaceous (Barremiane–Aptian), Yixian Formation.
Diagnosis. As for the genus.
Description. Female. Head entirely dark, slightly darker than thorax and abdomen, with ovipositor slightly darker; legs lightly colored; wing veins dark (Fig. 1).
Head: Moderately large, narrower than thorax, nearly round shaped. Eye elongate ovoid, occupying all head side; ocelli small, in wide triangle at level before eye midlength (Fig. 1A). Antenna thin, with almost uniform width, flagellomeres partly preserved and almost equal in lengths, maximum 0.39 mm in length and 0.25 mm in width (Fig. 1E).
Thorax: Pronotum short and trapezoid, at least four times as wide as long; pronotum posterior border slightly curved in dorsal view. Mesoscutum large, with anterior margin nearly straight, with longitudinal sulcus and notauli strongly impressed (Fig. 1B); mesoscutellum tapering to acutely sharp apex; ratio of lengths of prescutum, longitudinal sulcus between notauli and scuto-scutellar sulcus and scutellum 3.3/2.1/4.0. Mesopostnotum trapezoid (Fig. 1C), slightly narrower than metascutellum; metascutum with metascutellum small, rounded quadrangularly, transverse; cenchri present and small.
Leg: Legs spindly. Tarsi pentamerous, basitarsi long but shorter than remaining tarsomeres combined; tarsomeres (Fig. 1D) about twice as long as wide; pretarsal claws more than half length of fifth tarsomere, with curved apices.
Wing: Forewing three times as long as wide (Fig. 1G); vein C and R slightly bent, 1r-rs 1.2 times as long as 2r-rs; 1-Rs long and proclival, about twice as long as 1-M; 2-M distinct, 0.2 times as long as Rs+M; M+Cu slightly bent basally; 3+4-M 2.4 times as long as 5-M; 5-M 1.2 times as long as 3r-m; 3r-m and 2m-cu vertical; 2-Cu twice as long as 1-Cu; 1cu-a straight, lying basally of cell a, as long as 1-Cu; 2+3A with distinct subbasal loop; cell 2+3rm long and pentagonal, about 3.4 times as long as wide. Cell 1r about 2.1 times as long as wide; cell 2r short, as long as wide; cell 3r 3.4 times as long as wide; cell 1mcu about 1.7 times as long as wide.
Abdomen: Abdomen only slightly narrower than mesothorax, first abdominal tergite split medially (Figs. 1A, E). Ovipositor straight, prominent significantly behind abdomen, incompletely preserved (Fig. 1E). First valvifer small, subhemicircular as visible (Fig. 1F); second valvifer large narrowing up to single thickening before meeting with valvula 3 (saw sheath), six times as long as the first valvifer; valvula 2 (dorsal stylet) visible only basally at its thickened base; ovipositor sheaths almost straight, prominent significantly behind abdomen.
Measurements (in mm). Body length at least 19.21, excluding ovipositor 16.90; head length 2.54, width 2.71; antenna 4.50 long as preserved; thorax length 4.06; abdomen length 10.51; forewing 10.24 long as preserved, maximum width 2.69; ovipositor length 8.94 as preserved.
description: Published as part of Li, Yi, Rasnitsyn, Alexandr P., Shih, Chungkun, Ren, Dong & Wang, Mei, 2024, New fossil Anaxyelidae (Insecta: Hymenoptera: Siricoidea) from the Lower Cretaceous Yixian Formation of northeastern China, pp. 107-121 in Zootaxa 5562 (1) on page 112, DOI: 10.11646/zootaxa.5562.1.13, http://zenodo.org/record/14595696