Ctenodontina baleta Walker 1849, comb. nov.
Creators: name: Vieira, Rodrigo
name: Landa, José Manuel Ayala
name: Rafael, José Albertino
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Asilidae
Ctenodontina
Ctenodontina baleta

description: Ctenodontina baleta (Walker, 1849) comb. nov.
(Figs. 6, 24–26, 34–47)
Asilus baletus Walker, 1849: 427; Schiner, 1866: 721; Williston, 1891: 90 (catalogue); Kertész, 1909: 323 (catalogue); Hull, 1962 (2): 545; Martin & Papavero, 1970: 88 (catalogue).
Lecania baleta; Artigas & Papavero, 1995: 37; Papavero, 2009: 31 (catalogue).
Pachychaeta? caracasae Martin, 1975: 85; Artigas & Papavero, 1995: 37; Papavero, 2009: 31 (catalogue).
Pachychoeta caracasae Martin, 1975: 85 syn. nov. (Figs. 27 –29)
Pachychoeta inca Martin, 1975: 86 syn. nov. (Figs. 30–33)
Addenda to the holotype original description based on specimens from Venezuela.
Male terminalia (Figs. 38–44): Black. Medioapical region of hypandrium with concavity (Fig. 39). Epandrium wide basally with finger-like lobe placed dorsally. Apex of finger-like lobe rounded in lateral view (Fig. 40); subepandrial sclerite without lateral projections, apical margin rounded (Fig. 41); gonocoxite with apical 1/3 digitiform (Fig. 42); gonostyle subrectangular, covered with small setae anteriorly and with long setae posteriorly (Figs. 42, 43); aedeagus with two long prongs (Fig. 44); ejaculatory apodeme wide in lateral view (Fig. 44). Female terminalia (Fig. 46, 47): Tergite VIII expanded laterally (Figs. 46, 47); tergite IX+X slightly membranous medially (Fig. 46); apex of sternite VIII truncate with mediaoapical slit (Fig. 47); hypoproct with spiniform setae; median sclerite at posterior end of furca (remnant of tergite IX) absent; two oval sclerotized spermathecal capsules.
Comments. Ctenodontina baleta (Walker) comb. nov., Pachychoeta caracasae Martin syn. nov. and Pachychoeta inca Martin syn. nov. were allocated in Lecania Macquart. However, these species have diagnostic characters of Ctenodontina. Lecania is a paraphyletic group and has no diagnostic characters defined. According to Artigas & Papavero (1995), Lecania “is an extremely varied assembly of species. This group needs a thorough revision to decide whether there are several genera involved or only one. The variation of the several characters is amazing, and there is an incredible combination of them”.
Ctenodontina baleta (Walker) comb. nov., P. caracasae Martin syn. nov., P. inca Martin syn. nov. as well as C. nairae and C. martini, do not have a spinous swelling ventrally on male hind femora (Figs. 6, 7). However, as discussed by Fisher (1985) and Vieira (2012b), Ctenodontina is better defined by its very distinctive male and female terminalia and C. baleta (Walker) comb. nov., P. caracasae Martin syn. nov. and P. inca Martin syn. nov. by the following characters: female ovipositor with an apical comb of spines, and tergite VIII broadly expanded laterally, densely setose and cupped beneath, completely obscuring sternite VIII in profile and male specimens with an epandrium narrowed apically (usually terminating in a finger-like lobe), distinctly shorter than sternite VIII (Fisher & Hespenheide 1992; Fisher 2009; Vieira 2012b).
Material examined. Holotype of Asilus baletus: Venezuela / One of Walkers series so named. Eaw / Type Pachychaeta baleta (Wlk) [Walker], N. Papavero det. / Holotype Asilus baletus Walker, det. J.E. Chainey, 1985 (♂ NHM); Holotype of Pachychaeta caracasae: Venezuela, 5.of Caracas, Quebrada Catia 18.xi.1968 / Chas. H. Martin Collector / California Academy of Sciences, Type No. 12602 / Pachychaeta caracasae Martin, Det. C.H. Martin / Holotype Pachychaeta caracasae Chas. H. Martin (♂ CAS); Holotype of Pachychaeta inca: San Esteban, Venezuela, 1–15.Dec.1939 / P. Anduze Collector / California Academy of Sciences, Type No. 12603 / Holotype Pachychaeta inca Chas. H. Martin / Pachychoeta inca Martin, Det. C.H. Martin 19 (♂ CAS).
Other material examined. [Venezuela], Distrito Federal, Rio Oritapo, 200m, col.: M. Ayala- 11.6.[19]67 (1 ♂, 1 ♀ INPA); [VENEZUELA], 7-x-1967, Edo. Guárico, La Puerta, [San Francisco de Tiznados-Qda. Coronado 09°31’05"N 67°37’29"W], col. Adolfo Ayala (2 ♂, 2 ♀ INPA); [Venezuela], rio Oritapo, Edo. Vargas: (10.621446°, -66.480845°) / 22-VIII- [19]65 (2 ♂ JMAC); same data, except: 26-XI- [19]65 (1 ♂ JMAC); same data, except: 11- VI- [19]67 (2 ♂, 3 ♀ JMAC); [Venezuela] rio Camurí Grande, Edo. Vargas: (10.613400°, -66.719940°) / 23-IX- [19]72 (1 ♀ JMAC); [Venezuela] rio Cojo Macuto- Edo. Vargas: (10.599056°, -66.881881°) / 9-VIII- [19]69 (1 ♂ JMAC); [Venezuela] rio Curupao, Edo. Miranda: (10.497547°, -66.644678°) / 10-VI- [19]67 (1 ♂, 1 ♀ JMAC); [Venezuela] rio Suapire, El Manguito, Edo. Miranda: (10.294581°, -66.737645°) / 6-I- [19]73 (1 ♀ JMAC); [Venezuela] rio Periquito, La trilla, Edo. Aragua: (10.388835°, -67.745727°) / 15-XII- [19]73 (1 ♂, 1 ♀ JMAC); same data, except: 12-X- [19]68 (3 ♂, 2 ♀ JMAC); [Venezuela] rio Maletero, La Victoria, Edo. Aragua: (10.236862°, -67.315002°) / 2-IX- [19]77 (3 ♀ JMAC); [Venezuela] rio Guárico - San Sebastian - La Puerta, Edo. Aragua: (9.916317°, -67.349211°) / 13-VII- [19]67 (1 ♀ JMAC); same data, except: 7-X- [19]67 (10 ♂, 6 ♀ JMAC); [Venezuela] rio Aragua, La Victoria, Pié de Cerro: (10.256486°, -67.348120°) / 18-V- [19]74 (1 ♂, ♀ JMAC); same data, except: 29-VI- [19]74 (1 ♂, ♀ JMAC); [Venezuela] Hacienda Carrizalito - Palambrota, Edo. Guárico: (9.651826°, -67.552502°) / 20-XII- [19]65 (1 ♀ JMAC); [Venezuela] rio Orituco, Edo. Guárico: (9.910627°, -66.394536°) / 11-XI- [19]72 (1 ♂, 1 ♀ JMAC); [Venezuela] rio Borburata, Edo. Carabobo: (10.425188°, -67.977241°) / 9-VIII- [19]86 (1 ♂, 1 ♀ JMAC).
description: Published as part of Vieira, Rodrigo, Landa, José Manuel Ayala & Rafael, José Albertino, 2017, Ctenodontina Enderlein, 1914 (Diptera, Asilidae, Asilinae): new combinations, synonyms, new species and new records for Argentina, pp. 279-291 in Zootaxa 4216 (3) on pages 285-289, DOI: 10.5281/zenodo.231832