Dendropaemon (Glaphyropaemon) inemarginatus Genier & Arnaud, new species
Creators: name: François Génier
name: Patrick Arnaud
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Dendropaemon
Dendropaemon inemarginatus

description: 25. Dendropaemon (Glaphyropaemon) inemarginatus Génier & Arnaud, new species
(Fig. 25, 158)
Dendropaemon (D.) refulgens: Martínez & Clavijo 1990, Bol. Ent. Ven. N.S. 5: 155 (biology)
Type locality. Coromoto, Departamento Atures, Territorio Federal Amazonas, Venezuela.
Diagnosis. The only species in the genus with a vestigial posterior pronotal margin. In some species the margin is reduced but always visible on a distance laterally and medially.
Description. Female holotype (Fig. 25). Body. Body moderately large, length 13.0 mm, maximum width 6.5 mm; body subrectangular in dorsal view; dorsum slightly convex. Color. Dorsal surface dark brown to black, glossy, with green metallic sheen; head with green metallic sheen on posterior portion of clypeus, genae and frons; pronotum with green metallic sheen except for anteromedian carina, on anterior portion of disc and surface adjacent to lateral fossae; elytra with uniform green metallic sheen; ventrum reddish brown to dark brown; pygidium with green metallic sheen; legs reddish brown to dark brown. Head. Clypeus broadly arcuate, anterior portion upturned; clypeal teeth ogival; clypeal median emargination narrowly v-shaped, clypeal edge acutely notched on external side of each clypeal tooth, clypeal teeth ventral surface lacking carina, clypeal margin illdefined, lacking sharp carina posteriorly, clypeal surface with transverse blunt rugulae; clypeogenal suture welldefined, bluntly carinate internally; genal surface irregularly punctate, transversally and shallowly sulcate on posterior half; clypeofrontal carina rather low, approximately 4 times wider than high, slightly arcuate in dorsal view, simply carinate, clypeofrontal carina apical edge straight in frontal view; eyes small in dorsal view, interocular ratio 5.8. Pronotum. Pronotum transverse in dorsal view, pronotal width/length ratio 1.4; disc of pronotum minutely punctate basally, puncture becoming fine anteriorly, with an ill-defined shallow longitudinal depression on posterior half; pronotal anterior margin wider and flat lateral to eyes; anterior portion with a trisinuous carina, carina produced into a tubercle medially; anterior angles surface simply punctate, slightly but distinctly sulcate along posterior edge of anterior margin; lateral fossae simply rounded, concave; lateral portions unmodified; pronotal basal fossae ill-defined, slightly concave; posterior margin vestigial, reduced to a minute interrupted carina throughout. Elytra. Elytra approximately as long as wide in dorsal view, elytral combined width/ length ratio 1.1; elytral base lacking distinct margin, simply convex; elytral striae 1–4 moderately wide basally and fine apically, evenly impressed throughout, elytral striae 5 atrophied, lacking fine carina on each side on disc, strial punctures fine and well-defined, adjacent strial edge encroaching on interval, stria 1 well-defined apically, connecting to marginal stria; interstriae slightly convex, minutely punctate throughout, surface glossy. Thoracic sterna. Proepisternal carina complete, extending laterally; metasternal median lobe angularly produced anteromedially, ventral ridge well-defined, v-shaped. Legs. Profemur posterior surface slightly but distinctly convex, rather coarsely punctate and glabrous internally, posterointernal margin rather thin, evenly developed, internal edge rather wide, with a contiguous row of setae along anterointernal edge and few scattered long setae on anterior half, remaining surface smooth. Protibia with four teeth on lateral edge; internal basal angle lobate; anterior surface with long aligned row of setae internally, surface glossy or feebly microsculptured between punctures; posterior surface with rugose irregular punctures externally to median carina, surface finely and irregularly microsculptured between punctures, with a single interrupted setal row along lateral teeth. Mesofemur angularly produced on anterointernal edge apically. Mesotibia rather short, gradually widening toward apex in anterior view; anteroapical edge slightly sinuate in anterior view, anteroapical row of setae complete; apicoanterior edge circularly indented internally; external edge more or less rounded, with several large elongate setiferous punctures. Mesotarsus similar in shape to metatarsus, 3-segmented, first segment short, approximately as long as wide at apex. Metafemur broadly oval in anterior view, lacking distinct depressed area anterointernally before apex, apicoposterior edge unmodified, anterior surface with a well-defined sulcus on more than half the length. Metatibia moderately slender, slightly widening toward apex in anterior view, anterior surface with distinct row of setae, with fine irregular punctures on a glossy surface, metatibial posterior surface flat between longitudinal row of setae and lateral edge, glossy between punctures. Metatarsus 3-segmented, first segment short, approximately as long as wide at apex, with anterointernal carina well defined and almost reaching apical edge. Abdominal sternites. Sternites 3–6 longitudinally flat; sternites 4–6 with 1–3 unaligned rows of setae laterally, glabrous medially; sternite 7 approximately longitudinally flat medially, shorter than segment 6 along midline; pygidium finely punctate on disc.
Measurements (1 female). Length: 13.0 mm.
Primary type data. Female holotype (CMNC): [Marz-979/ VENEZUELA / T.F.Amazonas/ D o Atures/ Coromoto/ G.y H.Martínez-leg./ Coll. A. Martínez] handwritten; [H. & A. HOWDEN/ COLLECTION/ ex. A. Martínez coll.]; [WORLD/ SCARAB./ DATABASE/ WSD00016731]; [HOLOTYPE ♀/ Dendropaemon / inemarginatus n.sp. / Génier & Arnaud, 2014] red card.
Material examined. Primary type only.
Etymology. Inemarginatus, without + margin, an adjective referring to the configuration of the posterior pronotal margin.
Natural history. The holotype was collected in a zone of loose and sandy soil in the early hours of the morning (Martínez & Clavijo, 1990).
Remarks. Male and variation unknown.
Superficially resembling species of Coprophanaeoides, however, the presence of a complete proepisternal carina place this species within Dendropaemon s. str. where it seems to occupy an isolated position.
description: Published as part of François Génier & Patrick Arnaud, 2016, Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini) in Zootaxa 4099 (1) on pages 52-53, DOI: 10.11646/zootaxa.4099.1.1, http://zenodo.org/record/268274