Dichotomius (Cephagonus) marcoscarvalhoi Nunes & Vaz-de-Mello 2019, new species
Creators: name: Nunes, Rafael V.
name: Vaz-de-Mello, Fernando Z.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Dichotomius
Dichotomius marcoscarvalhoi

description: 3.18.: Dichotomius (Cephagonus) marcoscarvalhoi new species
(Figure 39 (e-h))
Diagnosis. separated from other species in the quadraticeps group by the following combined characters: larger males pronotum with two central-rounded lobes on the anterior portion of the disc; posterior margin of pronotum having 1 – 2 rows of ocellate or/and coarse punctures; 1st ventrite with groups of 8 – 12 long hair at each side and paramera with rounded and convergent apexes.
● HOLOTYPE (male): BRASIL: Mato Grosso. Curvelândia. Fazenda Aparecido. 15³33 ʹ 43 ʹ ’ Sı 58³00 ʹ 20 ” W. ii-2004. M Santos-Filho [at CEMT].
● PARATYPES: same data of Holotype [8 ƋƋı 2 ♀♀ at CEMT]; BRASIL. Mato Grosso. Araputanga. Fazenda Sudam. 15³15 ʹ 22 ” Sı 58³27 ʹ 12 ” Wı 352 m. Floresta semidecídua. Pitfall fezes humanas. 28-i-2013. RJ Silva [2 ƋƋ at CEMT]; Campos de Júlio. 30 km N de Uirapuru. 14³17 ’ Sı 59 ³15 ʹ W. xii-2002. A Foucart [1 Ƌı 3 ♀♀ at CEMT]; Mirassol do Oeste. Fazenda Santa Helena. 15³36 ʹ 34 ” Sı 57 ³58 ʹ 12 ” W. 26-i-2013. RJ Silva [1 ♀ at CEMT].
Description (Holotypeı male). BL: 20 mmı PW: 12 mm. Colour: blackı shiny. Head: dorsal surface smooth. Interocular space with fine punctures. Clypeo-genal angle obtuse (110³). Cephalic carina longer than wide with rounded apex. Pronotum: margins with 1 – 2 rows of ocellate punctures. Posterior border groovedı almost faced; posterior angles excavated. Anterior portion of disc having a pair of central lobes on anterior portion. Region adjacent to eyes with ocellate punctures. Metasternum: sides and margins of anterior lobe with ocellate setigerous punctures. Anterior portion of anterior lobe smooth and glabrous. Setigerous punctures equally spaced at sides. Elytra: striae deeply impressed having fine punctures spaced by twice their diameter. Interstriae disc smoothı with fine punctures (viewed under 20x magnification)ı flat; 1st interstriae strongly convexı also lacking microsculpture with very fine punctures (20 x). Abdomen: 1st ventrite with groups of 8 – 12 long hair at each side. Puncture of each ventrite interrupted medially as well groove along the ventrite posterior margin of sixth ventrite. Aedeagus (Figure 39 (f-h)): dorsallyı paramere apexes acuteı parallel; longitudinal excavation present taking whole paramera length; sides sinuous; basal rounded excavation present; anterior delimitation of basal excavations forming a pair of knobs. Laterallyı subtle declivity from basis towards apex; wide posterior rounded excavation (1/5). Ventrallyı each paramera with a lateral groove; subgenital plate widely emarginated at apex and less emarginated at basis.
Morphological variation. males BL: 17 – 20 mmı PW: 9 – 12 mm. Females BL: 17 – 19 mmı PW 9 – 11 mm. Distinguished from males as follows: Head: dorsal surface striated. Cephalic carina lowı not bifurcated apically. Pronotum: simply convexı lacking lobes. Abdomen: sixth ventrite three times larger than fifth and bearing a central pair of tubercles. Some individuals (males or females) mesepimeron may have dense setaeı concealing the structure.
Distribution. Brazilı state of Mato Grosso (Figure 36).
Etymology. named after our friend Dr. Marcos André de Carvalhoı director of Instituto de Biociências (Universidade Federal de Mato Grossoı Brasil) for many years.
description: Published as part of Nunes, Rafael V. & Vaz-de-Mello, Fernando Z., 2019, Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini), pp. 2231-2351 in Journal of Natural History 53 (37) on pages 2324-2325, DOI: 10.1080/00222933.2019.1692088, http://zenodo.org/record/3665519