Cybaeus devius Chamberlin and Ivie
Creators: name: Bennett, Robb
name: Copley, Claudia
name: Copley, Darren
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Cybaeidae
Cybaeus
Cybaeus devius

description: <i>Cybaeus devius</i> Chamberlin and Ivie Figs 14–16, 32 <i>Cybaeus devius</i> Chamberlin and Ivie 1942: 18, fig. 48. Roewer 1954: 90. Roth and Brown 1986: 3. Bennett in Copley <i>et al.</i> 2009: 373, fig. 9. World Spider Catalog 2021. <b>Type material examined.</b> <b>U.S.A.: <i>California</i>: Holotype female.</b> Madera County, North Fork, 37°15'N / 119°30'W, no date, no collector (AMNH). <b>Diagnosis.</b> The male of <i>C. devius</i> is unknown. The female of <i>C. devius</i> is only likely to be confused with the females of <i>C. echo</i> <b>spec. nov.</b> or <i>C. wilsonia</i> <b>spec. nov.</b> with which it shares very short spermathecal stalks and the unique copulatory duct morphology of a heavily sclerotized apical section of each copulatory duct enveloped by a very lightly sclerotized, almost diaphanous, basal section (Figs 15–16, 18–19, 21–22). In the female of <i>C. devius</i> (Figs 15–16) the basal section of each copulatory duct is voluminous, makes three loops around the more heavily sclerotized apical section, and occupies a greater area than (and, in ventral view, partially obscures) each relatively small spermathecal base. In the female of <i>C. wilsonia</i> <b>spec. nov.</b> (Figs 18–19) the basal section of each copulatory duct is also voluminous but is looped only twice around the apical section. In the female of <i>C. echo</i> <b>spec. nov.</b> (Figs 21–22) the basal section of each copulatory duct loops three times around the apical section but is not voluminous (the basal and apical sections are of similar diameter) and occupies a much smaller area. Furthermore, the spermathecal bases are large and unobscured in females of <i>C. echo</i> <b>spec. nov.</b> (Figs 21–22) and <i>C. wilsonia</i> <b>spec. nov.</b> (Figs 18–19) bases. Finally, the basal section of each copulatory duct in <i>C. devius</i> travels from the atrium to the anterior margin of the vulva before beginning to loop around the apical section (Fig. 15) (<i>versus</i> travelling only partway to the anterior margin before looping in <i>C echo</i> <b>spec. nov.</b> [Fig. 21] and <i>C. wilsonia</i> <b>spec. nov.</b> [Fig. 18]). <b>Description.</b> Ventral tibia I macrosetae 2-1p-2-1p-0. <b> <i>Female</i>:</b> (n=1). Atrium (Figs 14–15) very small, about 1/4 width of vulva. Spermathecal heads apparently protruding dorsally from vulval ducts from within region enclosed by copulatory ducts (Fig. 16). Measurements (n=1). CL 2.11, CW 1.44, SL 1.04, SW 1.01. <b>Note:</b> <i>Cybaeus echo</i> <b>spec. nov.</b> and perhaps <i>C. wilsonia</i> <b>spec. nov.</b> may be synonyms of <i>C. devius</i>. Insufficient specimens are available to make a strongly supported conclusion of conspecificity. Although its coloration indicates the single specimen of <i>C. echo</i> is fully matured the vulva at least appears to be teneral with thin-walled ducts and weakly developed Bennett’s glands. Likewise the three known females of <i>C. wilsonia</i> all have apparently teneral vulvae. Clearly more specimens (especially males) from across the entire Sierra Nevada range of these three putative species need to be examined to analyze variation. At this time we consider the specimens examined here to be sufficiently distinct to warrant the recognition of three separate species. <b>Distribution</b>. (Fig. 32). Known only from the type locality in Madera County on the western slopes of the central Sierra Nevada.
description: Published as part of <i>Bennett, Robb, Copley, Claudia &amp; Copley, Darren, 2021, Cybaeus (Araneae: Cybaeidae) in the Nearctic: the devius and tardatus species groups of the Californian clade, pp. 451-479 in Zootaxa 5026 (4)</i> on page 458, DOI: 10.11646/zootaxa.5026.4.1, http://zenodo.org/record/5300630
description: {"references": ["Roewer, C. F. (1954) Katalog der Araneae 2 a. Institut Royal des Sciences Naturelles de Belgique, Bruxelles, 923 pp.", "Copley, C. R., Bennett, R. &amp; Perlman, S. J. (2009) Systematics of Nearctic Cybaeus (Araneae: Cybaeidae). Invertebrate Systematics, 23, 367 - 401. https: // doi. org / 10.1071 / IS 09001", "World Spider Catalog (2021) World Spider Catalog. Version 22.0. Natural History Museum, Bern. Available from: https: // wsc. nmbe. ch / (accessed 15 June 2021) https: // doi. org / 10.24436 / 2"]}