Daedalma Pyrcz, Greeney, Willmott & Wojtusiak, 2011, n. ssp.
Creators: name: Pyrcz, Tomasz W.
name: Greeney, Harold F.
name: Willmott, Keith R.
name: Wojtusiak, Janusz
Publisher: name: Zenodo
Year: 2011
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Nymphalidae
Daedalma

description: Daedalma vertex milleri Pyrcz, n. ssp.   
(Figs. 3G, 3H, 10B)  
 Material examined: PERU: HOLOTYPE male: Huánuco, Carpish, 2700–3000 m, 08.VI.1995, 0943/7606, G. Lamas leg., red, rectangular label saying: Holotype, MUSM; PARATYPES (9 males and 1 female): 3 males: Huánuco, Carpish, 2800–3000 m, 26.X.2004, P. Boyer leg., 2 PBF, 1 TWP; 1 male: same locality but 19.II.2003, TWP; 1 male: same data but 19.II.2003, PBF; 3 males: Huánuco, Carpish, 2700–2800 m, 08.VI.1995, G. Lamas leg., MUSM; 1 male: Huánuco, au dessus de Huanacaure, km 43 de Pachachupan, est de Acomayo, 3000–3100 m, P. Boyer leg., PBF; 1 female: Huánuco, 2 km au dessus de Huanacaure, route Pachachupan-Huanacaure km 42, 3000 m, 22.X.2009, P. Boyer leg., PBF.   
 Description: MALE (Fig. 3G): Head, thorax and abdomen: not differing from nominate subspecies. Wings: FW length: 26–27 mm (mean: 26.2 mm, n=7), shorter than in nominate subspecies (mean: 28,6 mm); FWD ground colour blackish brown; orange patch a shade darker and smaller than in nominate subspecies, only slightly shorter distally but nearly half as wide; blackish postmedian patch in cell Cu1-Cu2 situated at edge of orange area, rather than enclosed by orange as in nominate subspecies. HWD uniform blackish brown, same as in nominate subspecies. New subspecies differs from nominate subspecies on FWV in shape and size of patch, and brick red colour, markedly darker in milleri. HWV pattern similar to nominate subspecies but ground colour chocolate brown instead of blackish brown. Genitalia (Fig. 10B): As compared to nominate subspecies, uncus same length but stouter, especially in basal part; gnathos similar; valvae narrowing gradually towards apex; saccus and aedeagus not differing noticeably from nominate subspecies.   
 FEMALE (Fig. 3H): FWD patch wider than in the male; a shade lighter orange. HWD with orange reddish patches along costal margin to apical area, gradually darkening basally, some faint reddish markings in the submarginal and marginal area of M3-Cu1 and Cu1-Cu2. FWV similarly patterned as in the male except for more prominent orange markings. HWV patterned as in the male but with more prominent sandy yellow basal, postbasal and postdiscal markings, a dark brown median band, and olive scaling denser in postmedian and marginal areas. Genitalia: not examined.   
 Etymology: This subspecies in dedicated to Lee D. Miller, eminent American entomologist, who contributed significantly to the knowledge of Pronophilina.   
 Remarks: Daedalma vertex milleri is apparently confined to central Peru, where it occurs in Huánuco, and most probably in Pasco and Junín. Daedalma vertex milleri apparently occurs at slightly lower elevations than the nominate subspecies (Pyrcz, 2004), with available data indicating an altitudinal range of 2600–3100 m.
description: Published as part of Pyrcz, Tomasz W., Greeney, Harold F., Willmott, Keith R. & Wojtusiak, Janusz, 2011, 2898, pp. 1-68 in Zootaxa 2898 on page 24