Puto Signoret
Creators: name: Zheng, Xinyi
name: Li, Yuang
name: Tan, Zhixiang
name: Wu, San'An
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Pseudococcidae
Puto

description: Key to adult females of Puto from the Old World (updated from Danzig & Gavrilov 2014)
Notes: The key does not include P. brunnitarsis (Signoret 1875) because the available description is inadequate. The data of P. barberi (Cockerell), P. marsicanus and P. thailandicus are derived from Williams & Granara de Willink (1992), Marotta & Tranfaglia (1993) and Williams (2004) respectively.
1(0) Claw with denticle well developed........................................................................ 2
- Claw without denticle............................................................... P. borealis (Borchsenius)
2(1) Dorsal tubular ducts present............................................................................. 9
- Dorsal tubular ducts absent............................................................................. 3
3(2) Ventral tubular ducts of 2 or 3 sizes....................................................................... 4
- Ventral tubular ducts all same size........................................................................ 6
4(3) Ventral tubular ducts of 2 sizes........................................................................... 5
- Ventral tubular ducts of 3 sizes............................................................ P. orientalis Danzig
5(4) Large ducts present along margins of thorax and posterior abdominal sternites, absent from median areas of posterior abdominal sternites........................................................................... P. antennatus (Signoret)
- Large ducts absent from margins of thorax and posterior abdominal sternites, but present on median areas of posterior abdominal sternites............................................................................. P. barberi (Cockerell)
6(3) Ventral tubular ducts scattered throughout posterior part of body, but absent from anterior part of body... P. konoi Takahashi
- Ventral tubular ducts present in frontal group only, or (rarely) totally absent....................................... 7
7(6) Multilocular disc-pores numerous, scattered on all medial zone of venter.................. P. superbus (Leonardi), partly
- Multilocular disc-pores few, present on cephalothoracic sternites only or totally absent.............................. 8
8(7) Multilocular disc-pores present on cephalothoracic sternites............................... P. peyerimhoffi (Vayssiere)
- Multilocular disc-pores absent.......................................................... P. israelensis Ben-Dov
9(2) Dorsal tubular ducts of 2 sizes; smaller ducts present along margins of abdominal tergites.......... P. caucasicus Hadzibejli
- Dorsal tubular ducts all same size, large.................................................................. 10
10(9) Dorsal tubular ducts wide; diameter of their openings more than 2 times larger than diameter of a trilocular pore......... 11
- Dorsal tubular ducts narrow; diameter of their openings about the same size as diameter of a trilocular pore............. 13
11(10) Dorsal tubular ducts located only in cerarii. Ventral tubular ducts all same size, present only in frontal group................................................................................................ subericola (Vayssiere)
- Dorsal tubular ducts located between cerarii. Ventral tubular ducts of 2 or 3 sizes, numerous on all abdominal sternites.... 12
12(11) Quinquelocular disc-pores present....................................................... P. huangshanensis Wu
- Quinquelocular disc-pores absent........................................................... P. tubulifer Danzig
13(10) Dorsal tubular ducts present along body margin only........................................................ 14
- Dorsal tubular ducts present in medial zones of tergites as well as along margins.................................. 15
14(13) Dorsum with additional cerarii, forming 6 longitudinal rows. Ventral margins without small oral collar tubular ducts.................................................................................... P. superbus (Leonardi), partly
- Dorsum without additional cerarii. Ventral margins with groups of small oral collar tubular ducts.................................................................................................. P. sinensis Zheng & Wu, sp. n.
15(13) Ventral tubular ducts present on thoracic and abdominal sternites as well as in frontal group......................... 16
- Ventral tubular ducts absent from thoracic and abdominal sternites, present only in frontal group..................... 17
16(15) Dorsum with additional cerarii forming 2 longitudinal rows, plus several additional cerarii on abdominal segment VI..................................................................................... P. megriensis (Borchsenius)
- Dorsum without additional cerarii...................................................... P. thailandicus Williams
17(15) Dorsum with additional cerarii forming 6 longitudinal rows........................ P. marsicanus Marotta & Tranfaglia
- Dorsum with additional cerarii forming 2 longitudinal rows.............................. P. kondarensis (Borchsenius)
description: Published as part of Zheng, Xinyi, Li, Yuang, Tan, Zhixiang & Wu, San'An, 2025, A new species of the genus Puto Signoret (Hemiptera: Coccomorpha: Putoidae) from China and redescription of Puto huangshanensis Wu, pp. 119-137 in Zootaxa 5570 (1) on pages 133-134, DOI: 10.11646/zootaxa.5570.1.5, http://zenodo.org/record/14734630