Nebria (Epinebriola) tuberculata Huber & Schmidt, 2017, sp. nov.
Creators: name: Huber, Charles
name: Schmidt, Joachim
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Nebria
Nebria tuberculata

description: Nebria (Epinebriola) tuberculata sp. nov. (Fig. 24)
Holotype ♀: Nepal, Helambu, S-slope Ganja La, 4400 –4800 m, 5. 8. 1998, leg. C. Berndt (cSCHM).
Body length 12 mm. Colour dark brown, appendages of the head brown. Antennae brown. Legs brown, femur darkened with brown knees. Pronotum brownish lightened, slightly lighter than the black head and the elytra. 1 st interval of the elytra brownish lightened.
Head with two shallow oblique impressions between the eyes. Labrum with anterior margin straight, bearing six setae. Apical margin of clypeus straight. Vertex impunctate, transversally wrinkled. Eyes weakly prominent. Antennae long and slender extending to one third of the elytra. Antennal scape elongate, a little longer than the eye's diameter, basally narrowed, subcylindrical apically, with one dorsal seta. 2 nd antennomere with one seta ventroapically. Maxillary stipes normal. Penultimate labial palpomere trisetose. Ligula spiny, with two lateral setae. Mentum with bifid medial tooth. Submentum with a row of 14 setae. Microreticulation of the head isodiametric.
Pronotum faintly transverse, ratio width/length of the pronotum = 1.26. Lateral margin distinctly convex, narrowed basally, widely rounded to the anterior angles, convexly narrowed to the posterior angles, concavely sinuate just in front of the small posterior angle (Fig. 31E). Lateral explanation narrow with a conspicuous groove also apically, broadened basally. Lateral margin basally blade-like, obliquely upturned. Posterior angles acute but very small, slightly protruding as a tubercle (Fig. 25). Anterior angles widely rounded, not protruding. Basal margin bisinuate, with a faint incision between the basal fovea and the small posterior angles. Pronotal disc weakly convex. Basal fovea deep, anterior and posterior transverse impression distinct; median line shallow, faintly transversally wrinkled. Lateral groove, basal fovea, anterior and posterior transverse sulci impunctate, only the area between the apical margin and the anterior transverse impression faintly punctate. Apical margination of the pronotum restricted to lateral one-fourths, merges with a faint ridge at the anterior angle; the ridge continues along the lateral groove to the lateral seta. Base of the pronotum not margined. Basolateral seta present. 1–2 midlateral setae in the apical half of the pronotum, inserted in distinct grooves at the inside of the slightly upturned margin. Microreticulation of the pronotum isodiametric, impunctate on disc. Proepisternum smooth. Prosternal process laterally margined, apex bulging and unmargined.
Elytral silhouette convex, regularly ovoid-elongate, widest slightly behind the middle. Ratio length/width of the elytra = 1.60. 1.7 times wider than the pronotum. Elytral apex rounded, apical carina absent. Basal margination straight, smoothly merged with the lateral margination. Humeral carina faintly developed. Striae distinct on disc, coarsely and tightly punctate. Striae obliterate towards the apex. Intervals rather convex, more convex on disc. Interval 3 with 7–8 setae adjoining stria 3. Scutellar seta absent. Microreticulation oblong. Mesepisterna smooth. Metepisterna 1.8 times as long as wide, smooth. Metacoxa with 4–5 basal and 1–2 apical setae. Hindwings present as a strap-like vestige.
Sternum 3 with one seta, faintly transversally wrinkled. Sterna 4–6 each with 5–7 posterior paramedial setae. Anal sternum with three paramedial setae in the female.
Legs long and slender. All tarsi dorsally glabrous. Metatarsomere 4 ventroapically markedly projected. Meso- and metatarsomeres 2–4 ventroapically with long setae. All tarsomeres 5 ventrally with two rows of 3–4 short setae. Metatarsomere 5 long, as long or longer than the metatarsomeres 3+4.
Female genitalia: Gonocoxa (Fig. 10E): Gonocoxite 2 elongate-triangular, broad at base, half the length of gonocoxite 1. Apex dorsally arcuate, dorsally distinctly grooved. Ventral preapical insertion furrow elongate-oval, with two nematiforme setae. Gonocoxites ventrally unjointed and continuously sclerotized, dorsally separated by a membranous area.
Male unknown.
Body ratios: hea.w/fro.w = 1.43; prm.w/hea.w = 1.34; prm.w/prp.w = 1.54; pra.w/prp.w = 1.07; ely.w/prm.w = 1.74; prm.w/prm.l = 1.27; ely.l/ely.w = 1.59.
Etymology: The name tuberculata (tuberculum, latin = small protuberance) refers to the acutely protruding tip of the posterior angles of the pronotum.
Diagnosis: N. tuberculata sp. nov. is recognizable within the subgenus Epinebriola by its plurisetose metacoxa with 4–5 setae, its plurisetose elytral interval 3 with 7–8 setae and its tuberculous posterior angles of the pronotum.
Distribution (Fig. 41): Known only from the type locality, the pass Ganja La (= Kanja La) in the Helambu mountain range, Northwestern Central Nepal.
Habitat: Not noted.
description: Published as part of Huber, Charles & Schmidt, Joachim, 2017, Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini), pp. 1-85 in Contributions to Natural History 36 on pages 48-51, DOI: 10.5169/seals-787049, http://zenodo.org/record/6285683