Clivina apexopaca Balkenohl 2021, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina apexopaca

description: <i>Clivina apexopaca</i> sp. nov. urn:lsid:zoobank.org:act: 250646A2-6AEC-4656-A989-44F652EA13EC (Figs 29, 49, 69, 96, 113) ETYMOLOGY. The name refers to the reticulated apex of the elytron. TYPE MATERIAL. Holotype: • &amp;male;, with labels and data: white, black framed, black printed and handwritten “Tjilatjap Java Drescher 15 XI. 1926 ” / “ <i>castanea</i> West, H.E. Andrewes det.” / “ <i>C. castanea</i> Westw. 57 det. K. Kult” (ETHZ). Paratypes: JAVA. • 1 spec., Java. Drescher Tjideres 100 M Res. Cheribon iii.1924 / <i>Clivina castanea</i> West. (Andrewes det.) (ETHZ); • 1 &amp;male;, 1 &amp;female;, 4 specs., same data but 28.v.–8.vi.1934 and 7–25.v.1934 / <i>C. castanea</i> Westw. 57 det. K. Kult (ETHZ); • 4 specs., same data but 19.vi.– 6.vii.1934 (ETHZ); • 1 spec., Batoerraden G. Slamet. Java F.C. DRESCHER., ix.1932 / <i>castanea</i> Westd. H.E. Andrewes det. / <i>C. castanea</i> Westw. 57 det. K. Kult (ETHZ); • 1 &amp;female;, Djember E. Java / <i>Clivina castanea</i> Westw. / COLLECTIO KAREL KULT COLL. A. DOSTAL (CADW); • 2 &amp;female;&amp;female;, INDONESIA, E JAWA Baluran Nat. P., 50 m Wonorejo 24– 25.ii.1994, BOLM leg. (SMNS); • 1 spec., F.C. Drescher Praeanger Bandoeng Dago lamp 13.xi.1929 / <i>castanea</i> Westd H.E. Andrewes det. / <i>C. castanea</i> Westw. 57 det. K. Kult (ETHZ). BALI. • 1 &amp;male;, 1 &amp;female;, Bali, 8 km e. Cekik W Bali NP, 8°11’S 114°30’E, 13–16.xii.2000, leg. Gerstmeier / <i>Clivina castanea</i> WESTW. det. M. Baehr’ 05 (CMBM / CMBB). DIAGNOSIS. Amedium sized species with smooth lateral margin of the pronotum and elytron, with isodiametric reticulation on interval eight of the elytron, and nearly right angled lobe of the mentum. Asimilar combination of characters show <i>C. agona</i> and <i>C. apexplana</i> sp. nov. <i>C. agona</i> differs mainly by the size, the distinct lateral group of punctures in the basal half of the pronotum, and the acute angled lobe of the mentum. <i>C. apexplana</i> sp. nov. differs mainly in the flattened and glossy intervals at the apex of the elytron. Moreover, in <i>C. apexopaca</i> sp. nov. interval seven is reticulated at the humerus and behind the anterior transverse line of the pronotum, and the male aedeagus shows a very different shape and inner structure. DESCRIPTION. <i>Measurements</i>. Holotype: Body length 7.33 mm, width 2.28 mm; ratio length/width of pronotum 0.88; ratio length/width of elytra 1.84. Other material: Body length 7.13–8.22 mm (: = 7.55 mm *), width 2.18–2.54 mm (: = 2.34 mm *), ratiolength/width of pronotum 0.84–0.89 (: = 0.88*), ratiolength/widthofelytra 1.79–1.85 (: = 1.83*); (*n = 10). <i>Colour</i>: Glossy. Piceous. Intermediate and hind legs dark fuscous, antennae and tarsomeres fuscous. <i>Head</i>: Athird narrower than pronotum. Clypeus slightly excised anteriorly; wing prominent, as protruding as clypeus, separated from clypeus by distinct notch; supraantennal plate convex; clypeus antero-laterally with carina, wing amargined, hollowed out, supraantennal plate reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by distinct notch. Clypeus slightly transverse, convex and glossy in posterior four fifths, smooth, reflexed margined anteriorly, separated from frons by distinct furrow. Frons moderately convex, with distinct pore at middle, with blunt lateral carina at posterior eye-level. Clypeus and frons smooth, with minute scattered punctures, separated from wing and supraantennal plate by distinctly deep furrow continuing posteriorly as wide supraorbital furrow with two supraorbital setigerous punctures at posterior eye level. Neck constriction developed as small step, with row of irregular punctures. Eye projecting, elongated in dorsal view. Gena covering eye by about a quarter in ventral view. Antenna reaching up to posterior angle of pronotum, antennomeres not flattened, four to ten of moderate length (ratio around L/ W 1.9). Labrum convex, with isodiametric reticulation, seven setose. Mentum (Fig. 49) with isodiametrically to longitudinally reticulated surface, with lobe somewhat concavely hollowed out, shape of lobe like a skewed trapezium, with slightly sinuate lateral margin, with straight and askew anterior margin and nearly rectangular lateral anterior angle, carinate at margins, with medial angle obtuse, with medium sized carina at middle, median tooth wide, somewhat less protruding anteriorly as lobe, obtuse angled anteriorly. <i>Pronotum</i> (Fig. 69): Disk moderately and regularly convex in lateral view, in frontal view moderately convex on disc but stepped laterally to channel. Wider than long. Reflexed lateral margin smooth, attenuate and slightly convex in anterior two thirds, widest in posterior third; anterior angle not projecting; posterior angle marked by distinct sharp tooth, tooth not protruding laterally over convexity of lateral margin. Lateral channel moderately wide, slightly widened between setigerous punctures, reticulated. Median line narrow, line-like, very narrow anteriorly, joining and distinctly surpassing anterior transverse line; anterior transverse line with irregular and joining punctures, just joining basal constriction. Surface smooth, with some transverse wrinkles and longitudinal wrinkles at middle, with distinct isodiametric reticulation in basal quarter, with small band of isodiametric reticulation behind anterior transverse line. <i>Elytron</i>: Disk indistinctly convex in anterior thirdin lateral view, convex in frontal view. Nearly twice as long as wide, nearly regularly convex with maximum width at middle. Reflexed lateral margin smooth. Scutellar striole distinct; setigerous tubercle at base of first stria, with distinct tubercle at base of third interval. Striae moderately deep, punctuate-striate, one to three free at base, four to seven joining at humerus, one running up to apex, just not joining with seven, all striae ending free at apex. Intervals slightly to moderately convex apically, eights with carina at humerus and in apical quarter. Third interval with four setigerous punctures, first situated at second stria near base, the other three approaching third stria. Surface of intervals glossy, interval eight with distinct isodiametric reticulation, all intervals with isodiametric reticulation in apical quarter, interval seven with reticulation at humerus. <i>Hind wings</i>: Fully developed. <i>Lower surface</i>: Proepisternum covered with fine but distinct isodiametric reticulation, with some transverse wrinkles. Terminal sternite of abdomen developed apically as deep narrow transverse fold. <i>Legs</i>: Legs covered with irregular to longitudinal reticulation. Protibia with three spines of moderate length, not sulcate dorsally, movable spur regularly arcuate. <i>Male genitalia</i> (Fig. 96): Median lobe moderately broad, regularly arcuate, distinctly arcuate at base, apex slender, with distinctly elongated spatula, ventral rim moderately developed, base with small apophysis. Endophallus apically with small band of short bristles. Ventral paramere wide, distinctly arcuate, lateral apophysis long, slender, apex slender, with two long and one small setae at apex. Dorsal paramere slender, sinuate, at apex with two long and and one small setae. <i>Female coxostylus and laterotergite</i> (Fig. 113): Coxostylus moderately slender, regularly acuminated to apex, dorsally with nine long to medium sized and one distinctly elongated and robust nematiform setae in basal half, ventrally with numerous smaller setae, with one SSO. Laterotergite with four setae of moderate length and two smaller setae basally and one isolated seta towards coxostylus. VARIATION. In specimens from Bali the median line surpasses the anterior transverse line not as distinct as in the specimens from Java. DISTRIBUTION. Known from Java and Bali.
description: Published as part of <i>Balkenohl, Michael, 2021, Belgian Journal of Entomology, pp. 1-83 in Belgian Journal of Entomology 115</i> on pages 42-44, DOI: 10.5281/zenodo.5645953