Mecynostomella caledonia Johanson 2003, sp.n.
Creators: name: Johanson, Kjell Arne
Publisher: name: Zenodo
Year: 2003
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Kokiriidae
Mecynostomella
Mecynostomella caledonia

description: Mecynostomella caledonia sp.n.
(Figs 28­35, 53)
Material examined: Male holotype: Trib. of Karionan River, 5 km NW Païta, 122 m (N54), 14.xi.1972 [W. L. & J. G. Peters] (ROM, alcohol). 1 male paratype: bank Grand Lakes, 20.viii.1965 [F. Starmühlner] (NRM, alcohol). 3 male paratypes: Riviére Bleue, at bridge on Eaux et Forêts road, 21 km NW Ouénarou Forest Station, 183 m (N51), 6­7.xi.1972 [W. L. & J. G. Peters] (NRM, alcohol). 1 female paratype: Province Sud, stream, Koghi Mt., 22°10.327'S, 166°30.524'E, 522 m, Malaise trap, 12­16.xi.2001 [Johanson, Pape, Viklund] (NRM, alcohol). 3 male paratypes: Grand Lac, 3.i.1967 [F. Starmühlner] (ROM, alcohol). 5 male paratypes: Grand Lac W Cofimpac Camp, 250 m, 9.viii.1971 (#2) [J.D. Holloway] (ROM, alcohol). 1 male, 1 female paratype: Mt. Dzumac, 760 m, 22°02'S, 166°28'E, 27­ 28.ii.1984 [M. Pogue & M. Epstein] (NMNH, pinned). 1 male, 5 female paratypes: Riviére Bleue, 22°6'S, 166°39'E, 18.x.1986 [R. Brown & O. Pellmyr] (NMNH, pinned). 1 female paratype: Mt. Koghi, 400 m, 12­14.xi.1986 [R. Brown & O. Pellmyr] (NMNH, pinned). 9 male, 1 female paratypes: Mt. Koghi, 400 m, 12­14.xi.1986 [R. Brown & O. Pellmyr] (NMNH, alcohol). 2 male, 1 female paratypes: Col de Mouirange, 225 m, 27.x.1986 [R. Brown & O. Pellmyr] (NMNH, alcohol). 2 male paratypes: Riviére de Pirogue, 22°10'S, 166°42'E, 500 m, 7­9.ii.1984 [M. Pogue & M. Epstein] (NMNH, alcohol). 2 male paratypes: same, except (NRM, pinned from NMNH). 1 male, 4 female paratypes: Riviére Bleue, 166°40'E, 22°6'S, 8.xi.1986 [R. Brown & O. Pellmyr] (NMNH, alcohol). 2 male paratypes: Riviére Bleue, 166°37.53'E, 22°5'S, 8.xi.1986 [R. Brown & O. Pellmyr] (NMNH, alcohol). 1 male, 4 female paratypes: Riviére Bleue, 166°39.55'E, 22°6'S, 18.xi.1986 [R. Brown & O. Pellmyr] (NMNH, alcohol).
Etymology: caledonia, derived from the type locality, New Caledonia. To be treated as a noun in genitive case.
Diagnosis: M. caledonia sp.n. lacks modified sensillae in the forewings as in brevis and spinosa. The species has a slender gonocoxite apex and a shallow, ellipsoidal incision on the posteroventral margin of segment IX.
Male (Fig. 28­32).
Head: Frontal surface grey­brownish, frons covered by short, dark brownish setae; dorsal part divided into dark anterior and pale posterior half; few, long, golden brown setae present along eye margin and posterior margin of head. Flagellum with 44 flagellomeres.
Thorax: Pronotum dark yellowish­grey, with few dorsolateral, stout, dark brownish setae in distinct warts. Mesonotum dark brownish, with golden setae, setal warts absent.
Wings: Forewing length 6.9­9.5 mm (n=10), hind wing length 5.9­8.5 mm (n=10). Forewings greyish­black, hind wings grey. Venation as described for genus.
Genitalia (Figs 28­32): Segment IX, lateral view (Fig. 28), about 2.3x higher than long; anterior margin shallowly produced, slightly undulating; rounded dorsally; posterior margin bi­lobed, dorsal lobe small; ventral margin widely incised (Fig. 30). Segment X with simple, central branch about parallel­sided at distal half in dorsal view (Fig. 29); in lateral view (Fig. 28), slender, about parallel­sided proximally, tapers distally; dorsal margin slightly bent ventrally at mid­length; with few setae on dorsolateral margin at distal half; more than 2x longer than high (Fig. 28); apex narrowly rounded in lateral view, slightly incised in dorsal view (Fig. 29). Lateral branch nearly as long as central branch, tubular, slightly curving ventrally along its length (Fig. 28), apex slightly bent laterad in dorsal view (Fig. 29), with short setae confined to apex, setae on proximal part form continuous band with setae on segment IX. Median lobe, lateral view (Fig. 28), large, slightly longer than high, longer than lateral branches; with setae on ventral margin; in dorsal view (Fig. 29) oriented posteriorly, visible between the central and lateral branches, pointed. Gonocoxite, lateral view (Fig. 28), sub­straight along its length; dorsal margin sigmoid, ventral margin sub­straight, apex produced dorsally, slender sub­apical part about 0.6x as high as proximal part; in ventral view (Fig. 30) oriented posteriad and with setae along its length. Phallus simple; phallobase, lateral view (Fig. 31), length about three­quarts the height of segment IX; endothecae produced dorsally in dorsal opening, without spines; in ventral view (Fig. 32) lateral incisions at proximal part located opposite to anterior margin of phallus opening.
Female (Figs 33­35).
Wings: forewing length 9.6­10.9 mm (n=5), hind wing length 7.8­9.0 mm (n=5). Venation as in male, except fork 1 in forewing present, sessile.
Genitalia: Vagina without spines. Segments VIII and IX separate. Segment IX apparently without setae; in lateral view (Fig. 33), dorsal process absent, ventral part produced posteriorly into lobe being slightly incised posteriorly and covered by small, marginal microtrichiae; in ventral view (Fig. 35), dorsal margin of lobe with central V­shaped incision surrounded by microtrichiae; in lateral view, dorsal and ventral margins of lobe subparallel. Pair of dark microtrichiae­filled, sickle­shaped, multi­lobed cavity on ventral margin of transition between segments IX and X; in dorsal view (Fig. 34), cavity visible through the segment, circular, with apices of outgrowths being distinctly darker. Segment X oriented ventrally; in lateral view (Fig. 33) divided into wide proximal half, and narrow distal half, with sub­straight dorsal margin, slightly undulating ventral margin; in dorsal view (Fig. 34), median processes curve medially, tapers apically, bears marginal microtrichiae; lateral processes oriented posteriorly.
description: Published as part of Johanson, Kjell Arne, 2003, Revision of the New Caledonian genus Mecynostomella (Trichoptera, Kokiriidae), pp. 1-24 in Zootaxa 270 (1) on pages 12-14, DOI: 10.11646/zootaxa.270.1.1, http://zenodo.org/record/5014321