Protohermes van der Weele 1907
Creators: name: Martins, Caleb C.
name: Ardila-Camacho, Adrian
name: Rivera-Gasperín, Sara Lariza
name: Oswald, John D.
name: Liu, Xingyue
name: Contreras-Ramos, Atilano
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Megaloptera
Corydalidae
Protohermes

description: Genus Protohermes van der Weele, 1907
Protohermes van der Weele, 1907: 243. Type species. Hermes costalis Walker, 1853: 207 (by present designation; see Nomenclatural notes below).
Hermes Gray in Cuvier, 1832: 332. Type species. Chauliodes maculipennis Gray in Cuvier, 1832: 332 (by original designation).
Neurhermes Navás, 1915b: 391. Type species. Chauliodes maculipennis Gray in Cuvier, 1832: 332 (by monotypy [of Hermes Gray in Cuvier, 1832]).
Allohermes Lestage, 1927: 100. Type species. Protohermes davidi van der Weele, 1909: 254 (by original designation).
Nomenclatural notes
Oswald & Penny (1991) identified Protohermes as a genus name based on a misidentified type species and a name whose type species had yet to be validly fixed. They noted that the case (at that time) required referral to the Commission for type species fixation under Art. 70b of the Code (3 rd edition, ICZN 1985). We are not aware of any nomenclatural acts published since 1991 that have validly fixed the type species of Protohermes under either the 3 rd or 4 th editions of the Code. To resolve this long-standing issue, we act here to formally fix the type species of Protohermes under Art. 70.3 (4 th edition, ICZN 1999) in a manner that is in accordance with long-accepted usage. Van der Weele (1907) originally included five valid nominal species in Protohermes – Hemerobius grandis Thunberg, 1781; Hermes anticus Walker, 1853; Hermes albipennis Walker, 1853; Nevromus montanus McLachlan, 1869; and Nevromus dichrous Brauer, 1878 – and explicitly designated Hermes anticus Walker, 1853 (as “ N. [sic] anticus Wlk. ”) as its type species. Kimmins (1949) presented evidence that the Protohermes anticus of van der Weele (1907) was not conspecific with Hermes anticus Walker, but was, rather, a misidentification of the taxonomic species Hermes costalis Walker, 1853. Based on this situation Kimmins recommended that “ Hermes anticus Weele 1907 (nec Hermes anticus Walker, 1853) (= Hermes costalis Walker, 1853)” (a nominal species not originally included in Protohermes) be treated as the de facto type species of Protohermes. Kimmins’ proposal has been accepted and adopted by all subsequent workers on the Corydalidae. Accordingly, we here fix (under Art. 70.3.2 of the Code) as the type species of Protohermes the nominal species Hermes costalis Walker, 1853 [the original combination of the current taxonomic species Protohermes costalis (Walker, 1853)], which was misidentified as Hermes anticus Walker, 1853 in the original designation of van der Weele (1907).
Included species
Protohermes acutatus Liu, Hayashi & D. Yang, 2007 a
Oriental (China).
Protohermes albipennis (Walker, 1853)
Hermes albipennis Walker, 1853
Corydalis albipennis (Walker, 1853)
Corydalus albipennis (Walker, 1853)
Nevromus albipennis (Walker, 1853)
Oriental (India, Nepal).
Protohermes arunachalensis Ghosh, 1991
Oriental (China, India, Myanmar).
Protohermes assamensis Kimmins, 1949
Oriental (India, Myanmar).
Protohermes axillatus Navás, 1932
Oriental (Vietnam).
Protohermes basimaculatus Liu, Hayashi & D. Yang 2007 a
Oriental (China).
Protohermes bellulus Banks, 1931
Oriental (Malaysia).
Protohermes burmanus Liu & Dvořák, 2017
Oriental (Myanmar).
Protohermes cangyuanensis C.-K. Yang & D. Yang, 1988
Oriental (China).
Protohermes cavaleriei Navás, 1925c
Oriental (China).
Protohermes changninganus C.-K. Yang & D. Yang, 1988
Oriental (China).
Protohermes chebalingensis Liu & D. Yang, 2006 a
Oriental (China).
Protohermes concolorus C.-K. Yang & D. Yang, 1988
Oriental (China).
Protohermes congruens Liu, Hayashi & D. Yang, 2009 a
Oriental (Nepal).
Protohermes costalis (Walker, 1853)
Hermes costalis Walker, 1853
Corydalis costalis (Walker, 1853)
Corydalus costalis (Walker, 1853)
Protohermes griseus Stitz, 1914
Protohermes costalis (Walker, 1853)
Oriental (China).
Protohermes costatostriatus (van der Weele, 1907)
Hermes costatostriatus van der Weele, 1907 Neurhermes costatostriatus (van der Weele, 1907)
Oriental (India).
Protohermes curvicornis Liu, Hayashi & D. Yang, 2013 a
Oriental (China, India, Myanmar).
Protohermes davidi van der Weele, 1909 Allohermes davidi (van der Weele, 1909)
Oriental (China).
Protohermes decemmaculatus (Walker, 1860)
Hermes decemmaculatus Walker, 1860
Nevromus decemmaculatus (Walker, 1860)
Oriental (India).
Protohermes decolor Navás, 1931
Oriental (Malaysia).
Protohermes dichrous (Brauer, 1878)
Nevromus dichrous Brauer, 1878
Hermes dichrous (Brauer, 1878)
Oriental (Indonesia, Malaysia).
Protohermes differentialis (C.-K. Yang & D. Yang, 1986)
Neurhermes differentialis C.-K. Yang & D. Yang, 1986
Neurhermes guangxiensis C.-K. Yang & D. Yang, 1992a
Neurhermes fangchengensis D. Yang in D. Yang et al., 2004
Oriental (China, Vietnam).
Protohermes dimaculatus C.-K. Yang & D. Yang, 1988
Oriental (China).
Protohermes disjunctus Liu, Hayashi & D. Yang, 2007 a
Oriental (Japan).
Protohermes dulongjiangensis Liu, Hayashi & D. Yang, 2010b
Oriental (China).
Protohermes festivus Navás, 1932
Oriental (China).
Protohermes flavinervus Liu, Hayashi & D. Yang, 2009 a
Oriental (China).
Protohermes flavipennis Navás, 1929b
Oriental (China).
Protohermes flinti Liu, Hayashi & D. Yang, 2007b
Oriental (Laos).
Protohermes fruhstorferi (van der Weele, 1907) Hermes fruhstorferi van der Weele, 1907
Allohermes fruhstorferi (van der Weele, 1907)
Oriental (China, Vietnam).
Protohermes fujianensis C.-K. Yang & D. Yang, 1999
Oriental (China).
Protohermes furcatus Liu, Hayashi & D. Yang, 2008b
Oriental (Thailand).
Protohermes goodgeri Liu, Hayashi & D. Yang, 2013b
Oriental (Malaysia).
Protohermes grandis (Thunberg, 1781) Hemerobius grandis Thunberg, 1781 Corydalis grandis (Thunberg, 1781) Corydalus grandis (Thunberg, 1781) Hermes grandis (Thunberg, 1781) Chauliodes grandis (Thunberg, 1781) Nevromus grandis (Thunberg, 1781)
Palaearctic (Japan).
Protohermes guangxiensis C.-K. Yang & D. Yang, 1986
Oriental (China, Vietnam).
Protohermes gutianensis D. Yang & C.-K. Yang, 1995
Oriental (China).
Protohermes hainanensis C.-K. Yang & D. Yang, 1990
Oriental (China).
Protohermes horni Navás, 1932
Oriental (China).
Protohermes hubeiensis D. Yang & C.-K. Yang, 1992b
Oriental (China).
Protohermes hunanensis D. Yang & C.-K. Yang, 1992a
Oriental (China).
Protohermes immaculatus Kuwayama, 1964
Oriental (Japan).
Protohermes impunctatus Liu, Hayashi & D. Yang, 2008b
Oriental (Myanmar).
Protohermes infectus (McLachlan, 1869) Nevromus infectus McLachlan, 1869
Protohermes anticus var. striolatus Stitz, 1914
Protohermes anticus striolatus Stitz, 1914
Protohermes tibetanus C.-K. Yang & D. Yang in Huang et al., 1988
Neochauliodes anticus striolatus (Stitz, 1914)
Oriental (China, India, Myanmar).
Protohermes ishizukai Liu, Hayashi & D. Yang, 2009b
Oriental (Laos, Vietnam).
Protohermes karubei Liu, Hayashi & D. Yang, 2013b
Oriental (Malaysia).
Protohermes latus Liu & D. Yang, 2006b
Oriental (China).
Protohermes lii Liu, Hayashi & D. Yang, 2007 a
Oriental (China, Vietnam).
Protohermes maculiferus (Walker, 1853)
Hermes maculiferus maculiferus Walker, 1853
Hermes maculiferus Walker, 1853
Neurhermes maculiferus (Walker, 1853)
Oriental (India).
Protohermes maculipennis (Gray in Cuvier, 1832)
Chauliodes (Hermes) maculipennis Gray in Cuvier, 1832
Chauliodes maculipennis Gray in Cuvier, 1832
Nevromus ruficollis Rambur, 1842
Hermes ruficollis (Rambur, 1842)
Hermes maculipennis (Gray in Cuvier, 1832)
Nevromus maculipennis (Gray in Cuvier, 1832)
Neurhermes maculipennis (Gray in Cuvier, 1832)
Oriental (India (?), Indonesia, Malaysia (?)).
Protohermes montanus (McLachlan, 1869) Nevromus montanus McLachlan, 1869
Oriental (India).
Protohermes motuoensis Liu & D. Yang, 2006b
Oriental (Bhutan, China, India, Nepal, Pakistan).
Protohermes niger C.-K. Yang & D. Yang, 1988
Oriental (China).
Protohermes nigerescens (Liu, Hayashi & D. Yang, 2015f) Neurhermes nigerescens Liu, Hayashi & D. Yang, 2015f
Oriental (India).
Protohermes ohli Liu, Hayashi & D. Yang, 2013 a
Oriental (India, Myanmar).
Protohermes orientalis Liu, Hayashi & D. Yang, 2007 a
Oriental (China).
Protohermes owadai Liu, Hayashi & D. Yang, 2007c
Oriental (Vietnam).
Protohermes parcus C.-K. Yang & D. Yang, 1988
Oriental (China).
Protohermes pennyi Liu, Hayashi & D. Yang, 2009 a
Oriental (Nepal).
Protohermes piaoacanus Liu, Hayashi & D. Yang, 2008b
Oriental (Vietnam).
Protohermes sabahensis Liu, Hayashi & D. Yang, 2008c
Oriental (Malaysia).
Protohermes selysi (van der Weele, 1909)
Hermes selysi van der Weele, 1909
Neurhermes bipunctatus C.-K. Yang & D. Yang, 1988 Neurhermes selysi (van der Weele, 1909)
Oriental (Bangladesh, China, India, Myanmar).
Protohermes similis C.-K. Yang & D. Yang, 1988 Protohermes sichuanensis D. Yang & C.-K. Yang, 1997
Oriental (China).
Protohermes sinensis D. Yang & C.-K. Yang, 1992a
Oriental (China).
Protohermes sinuolatus Liu, Hayashi & D. Yang, 2009b
Oriental (Vietnam).
Protohermes sonus Liu, Hayashi & D. Yang, 2013c
Oriental (Myanmar).
Protohermes spectabilis Liu, Hayashi & D. Yang, 2008c
Oriental (Malaysia).
Protohermes stangei Liu & Dobosz, 2019
Oriental (Vietnam).
Protohermes stigmosus Liu, Hayashi & D. Yang, 2007 a
Oriental (China).
Protohermes striatulus Navás, 1926 Protohermes basiflavus D. Yang in D. Yang et al., 2004
Oriental (China, Myanmar, Vietnam).
Protohermes sublunatus Liu, Hayashi & D. Yang, 2013c
Oriental (India).
Protohermes subnubilus Kimmins, 1949 Protohermes tengchongensis C.-K. Yang & D. Yang, 1988
Oriental (China, Myanmar).
Protohermes subparcus Liu & D. Yang, 2006c
Oriental (China).
Protohermes sumatrensis (van der Weele, 1909)
Hermes sumatrensis van der Weele, 1909
Neurhermes sumatrensis (van der Weele, 1909)
Oriental (Indonesia, Malaysia).
Protohermes tenellus Liu, Hayashi & D. Yang, 2007b
Oriental (Thailand).
Protohermes tonkinensis (van der Weele, 1909)
Hermes maculiferus tonkinensis van der Weele, 1909
Hermes tonkinensis van der Weele, 1909
Hermes maculiferus var. tonkinensis van der Weele, 1909
Neurhermes tonkinensis (van der Weele, 1909)
Oriental (China, Laos, Thailand, Vietnam).
Protohermes tortuosus Liu, Hayashi & D. Yang, 2008c
Oriental (Malaysia).
Protohermes trapezius Li & Liu, 2021
Oriental (China).
Protohermes triangulatus Liu, Hayashi & D. Yang, 2007 a
Oriental (China, Laos, Thailand, Vietnam).
Protohermes uniformis Banks, 1931
Oriental (Malaysia).
Protohermes vitalisi Navás, 1919a
Oriental (Vietnam).
Protohermes walkeri Navás, 1929b
Oriental (India, Nepal, Pakistan).
Protohermes weelei Navás, 1925c
Oriental (China, Laos).
Protohermes wuyishanicus Li & Liu, 2021
Oriental (China).
Protohermes xanthodes Navás, 1914
Protohermes rubidus Stitz, 1914
Protohermes martynovae Vshivkova, 1995
Palaearctic (China, North Korea, Russia, South Korea); Oriental (China).
Protohermes xingshanensis Liu & D. Yang, 2005 a
Oriental (China).
Protohermes yangi Liu, Hayashi & D. Yang, 2007 a
Oriental (China, Laos, Vietnam).
Protohermes yunnanensis C.-K. Yang & D. Yang, 1988
Oriental (China, Vietnam).
Protohermes zhuae Liu, Hayashi & D. Yang, 2008b
Oriental (China).
description: Published as part of Martins, Caleb C., Ardila-Camacho, Adrian, Rivera-Gasperín, Sara Lariza, Oswald, John D., Liu, Xingyue & Contreras-Ramos, Atilano, 2022, A world checklist of extant and extinct species of Megaloptera (Insecta: Neuropterida), pp. 1-93 in European Journal of Taxonomy 812 (1) on pages 23-30, DOI: 10.5852/ejt.2022.812.1727, http://zenodo.org/record/6448732