Tomoglossa Kraatz 1856
Creators: name: Gusarov, Vladimir I.
Publisher: name: Zenodo
Year: 2002
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Tomoglossa

description: <i>Tomoglossa</i> Kraatz, 1856 (Figs. 1­46) <i>Tomoglossa</i> Kraatz, 1856: 342. <i>Tomoglossa</i>; Bernhauer, 1907 b: 401. <i>Noverota</i> Casey, 1910: 90, <b>syn. nov.</b> <i>Tomoglossa</i>; Fenyes, 1920: 255. <i>Atheta</i> (<i>Noverota</i>); Fenyes, 1920: 199. <i>Tomoglossa</i>; Bernhauer &amp; Scheerpeltz, 1926: 598. <i>Atheta</i> (<i>Noverota</i>); Bernhauer &amp; Scheerpeltz, 1926: 606. <i>Tomoglossa</i>; Scheerpeltz, 1963: 123. <i>Tomoglossa</i>; Benick &amp; Lohse, 1974: 107. <i>Tomoglossa</i>; Sawada, 1977: 192. <i>Noverota</i>; Seevers, 1978: 122. <i>Tomoglossa</i>; Lohse, 1989: 208. <i>Noverota</i>; Newton, Thayer, Ashe &amp; Chandler, 2000: 369. (Other references for Palaearctic <i>Tomoglossa</i> are omitted) <b>Type Species.</b> <i>Homalota luteicornis</i> Erichson, 1837, by monotypy. <b>Diagnosis.</b> <i>Tomoglossa</i> can be distinguished from other athetine genera by the combination of the following characters: parallel­sided body; slender sickle­shaped mandibles (Figs. 1­3); ligula with two completely separate lobes (Fig. 10); mentum fused with submentum (Fig. 12); pronotum with microsetae directed posteriorly throughout the disc (Type VI, Benick and Lohse 1974) (Fig. 13); pronotal macrosetae inconspicuous; pronotal hypomera fully visible in lateral aspect; metasternal process extremely short and wide, virtually non­existent, perhaps better described as convexity of the anterior margin of metasternum (Fig. 15); mesotibia with short median macroseta (shorter than tibial width); metatarsal segment 1 longer than segment 2 (Fig. 14); without empodial setae; with numerous erect macrosetae and long semi­erect microsetae on abdominal terga VII­VIII. <i>Tomoglossa</i> shares with <i>Hydrosmecta</i> Thomson, 1858 slender mandibles, inconspicuous process of the metasternum, and the lack of empodial setae. <i>Tomoglossa</i> can be easily distinguished from <i>Hydrosmecta</i> by its less slender body, more widely separated lobes of the ligula, less transverse mentum fused with submentum, pubescence directed posteriorly everywhere on the pronotal disc (Type VI), shorter metasternum, and long erect setation of the apical abdominal segments. <i>Tomoglossa</i> is similar to <i>Geostiba</i> Thomson, 1858 in having pronotal pubescence of Type VI; however, <i>Tomoglossa</i> differs in having slender mandibles, more widely separated lobes of ligula, less transverse mentum fused with submentum, inconspicuous metasternal process, long and erect setation of the apical abdominal segments and in lacking empodial setae. <b>Description.</b> Length 1.6­2.1 mm. Head reddish brown to dark brown; pronotum yellow to reddish yellow, lighter than head; elytra from brownish yellow to brown, with yellow apical (1 / 3 to 1 / 5) portion; abdominal segments III­V and VII­VIII yellow to reddish yellow; abdominal tergum VI darker, reddish brown to dark brown; antennae uniformly yellow or with darker articles 2­11; maxillar palpi uniformly yellow or with darker article 3; legs yellow to reddish yellow. Body form parallel­sided. Head transverse, eyes as long as temples (seen from above); infraorbital carina complete. Antennal articles 2 and 3 of equal length, 4­5 subquadrate, 6­10 slightly transverse, apical article as long as 9 and 10 combined, without coeloconic sensilla. Labrum (Fig. 8­9) transverse and narrow, with straight anterior margin. Mandibles (Figs. 1­3) symmetrical, slender, sickle­shaped, with a small medial tooth; ventral molar area without patches of denticles, dorsal molar region without visible “velvety patch” (400 x) (Figs. 227.22­231.22 in Newton <i>et al.</i> 2000: 309). Maxilla (Figs. 4­7) with galea as long as lacinia, apical lobe of galea covered with numerous fine and short setae; interior margin of galea with row of setae subapically; apical third of lacinia with row of closely spaced spines, middle third produced medially and covered with numerous setae. Labial palpi with three articles; ligula divided into two completely separate lobes; medial pseudopore field of prementum with 4­5 pseudopores (Sawada 1970, 1972), lateral areas with pair of twin pores, single spinose pore and 4­6 irregular pseudopores (Fig. 10). Mentum (Fig. 12) with anterior margin concave, posteriorly fused with submentum. Pronotum transverse, broadest slightly in front of middle, sides broadly rounded, anterior margin straight, anterior and posterior angles rounded; posterior margin convex; surface covered with short microsetae directed posteriorly in both midline and lateral areas (Type VI, Benick &amp; Lohse 1974); macrosetae very short, limited to one macroseta at each of anterior and posterior angles and one on each lateral margin; hypomera fully visible in lateral aspect. Posterior margin of elytra straight. Wings present, fully developed. Mesosternal process short and wide, extending 1 / 4 length of mesocoxal cavities, metasternal process almost non­existent, anterior margin of metasternum only slightly convex medially (Fig. 15); mesosternum and mesosternal process not carinate medially; relative lengths of mesosternal process: isthmus: metasternal process in ratio of about 6: 19: 1; mesocoxal cavities margined posteriorly; mesocoxae narrowly separated. Mesotibia with very short median macroseta (shorter than tibia width). Tarsal segmentation 4­5 ­ 5; metatarsal segment 1 longer than segment 2 (Fig. 14). No empodial setae present. Abdominal terga III­V with moderate transverse basal impressions. Tergum VII 1.6 times longer than VI. Puncturation of terga III­V equally sparse, puncturation of terga VI­ VII sparser. Terga VII­VIII with numerous erect macrosetae and long semi­erect microsetae. Tergum VIII (Figs. 17, 19, 34, 38, 43) in both sexes with basal row of long microsetae. Female sternum VIII (Figs. 16, 18, 35, 44) with row of apical microsetae (as in other Athetini). Median lobe of aedeagus ventrally with very characteristic structure in the area of basal pore (Figs. 21 ­22, 40­ 41). Internal sac with a deeply concave sclerite (Figs. 23 ­24, 42). Copulatory piece (Sawada 1972) without apical process (Figs. 25, 42). Spermatheca with wide umbilicus (Muona 1990) (Figs. 26, 36, 45). <b>Discussion.</b> The type species of <i>Noverota</i> (<i>N. ornatella</i> Casey, 1910, by original designation) and <i>Tomoglossa</i> (<i>T. luteicornis</i> (Erichson, 1837)) differ only in body coloration (<i>T. luteicornis</i> being darker) and in minor details of genitalia (Figs. 21­26 in this paper and Fig. 123 in Lohse 1989). As a result, <i>Noverota</i> is placed in synonymy with <i>Tomoglossa</i>. Apparently Casey was not familiar with <i>Tomoglossa</i> and when describing <i>Noverota</i> he compared it only with <i>Hydrosmecta</i>. Casey (1910) included seven species in the genus <i>Noverota</i>. Three of these species belong to <i>Tomoglossa</i>. They are redescribed below. <i>Noverota clemens</i> Casey, 1910, <i>N. finitima</i> Casey, 1910, and <i>N. scenica</i> Casey, 1910 belong to <i>Philhygra</i> Mulsant &amp; Rey, 1873 and will be redescribed elsewhere. The taxonomic position of <i>N. personata</i> Casey, 1910 is not clear, but it does not belong to <i>Tomoglossa</i>. The species is known from a single female type and a detailed study of its mouthparts is not possible. When additional specimens of this species are found the position of the species can be clarified. Two new species of <i>Tomoglossa</i> are described below. Thus, five valid Nearctic species of <i>Tomoglossa</i> are currently known. In the key by Ashe (Newton <i>et al.</i> 2000) the specimens belonging to <i>Tomoglossa</i> run to the second half of the couplet 58. In the key by Benick and Lohse (1974) the same specimens easily reach <i>Tomoglossa</i> at couplet 8.
description: Published as part of <i>Gusarov, Vladimir I., 2002, A revision of Nearctic species of the genus Tomoglossa Kraatz, 1856 (Coleoptera: Staphylinidae: Aleocharinae), pp. 1-19 in Zootaxa 30</i> on pages 3-5, DOI: 10.5281/zenodo.4619813
description: {"references": ["Kraatz, G. (1856 - 1858) Naturgeschichte der Insecten Deutschlands. Abt. 1. Coleoptera. Band 2. Verlag der Nicolaischen Buchhandlung, Berlin, 1080 pp.", "Bernhauer, M. (1907 b) Neue Aleocharini aus Nordamerika. (Col.) (3. Stuck.). Deutsche Entomologische Zeitschrift, 1907 (4), 381 - 405.", "Casey, T. L. (1910) New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera, 1, 1 - 183.", "Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.) Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Brussels, pp. 111 - 414.", "Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.) Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, 499 - 988.", "Scheerpeltz, O. (1963) Neue Arten der Gattung Tomoglossa Kraatz, mit einer Bestimmingstabelle der bis jetzt bekannt gewordenen Arten dieser Gattung (Col. Staph.). Entomologische Berichten, 23 (7), 123 - 132.", "Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.) Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220.", "Sawada, K. (1977) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). III: Japanese Species described by the previous Authors. Contributions from the Biological Laboratory, Kyoto University, 25 (2), 171 - 222.", "Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, i-vi + 1 - 275.", "Lohse, G. A. (1989) Erganzungen und Berichtigungen zu Freude-Harde-Lohse \" Die Kafer Mitteleuropas \" Band 5 (1974). In: Lohse, G. A. &amp; Lucht, W. H. (Eds.) Die Kafer Mitteleuropas. 1. Supplementband mit Katalogteil. Goecke &amp; Evers Verlag, Krefeld, pp. 185 - 243.", "Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Eds.) American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, pp. 272 - 418.", "Erichson, W. F. (1837 - 1839) Die Kafer der Mark Brandenburg. Band 1. F. H. Morin, Berlin, viii + 740 pp.", "Sawada, K. (1970) Aleocharinae (Staphylinidae, Coleoptera) of the IBP-Station in the Shiga Heights, Central Japan (I). Bulletin of the National Science Museum, 13 (1), 23 - 64.", "Sawada, K. (1972) Methodological Research in the Taxonomy of Aleocharinae. Contributions from the Biological Laboratory, Kyoto University, 24 (1), 31 - 59.", "Muona, J. (1990) The Fennoscandian and Danish species of the genus Amischa Thomson (Coleoptera, Staphylinidae). Entomologisk Tidskrift, 111, 17 - 24."]}