Leucopholis Dejean 1833
Creators: name: Calcetas, Orlando A.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Leucopholis

description: Key to the species of Leucopholis of the Philippines
1. Metaventral process distinctly long, extending in front of prosternal process (Fig. 2); large to very large sized species, length 35.0– 42.5 mm; prosternal process mound-shaped, triangulate; pronotal scales variable in shape and size; body scales elongate ovoid, not flattened, slanted at 45° angle, base of scale only attached to surface ....................................................................................................... 2
– Metaventral process distinctly short not extending in front of prosternal process; moderate to small sized species, length 22.5–29.5 mm; prosternal process ovoid to spindle-shaped anteriorly; pronotal scales nearly uniform in shape and size; body scales distinctly ovoid, flattened, not slanted at 45° angle, entire scale strongly attached to surface ................................................................................ 5
2. Metaventral process lanceolate, constricted subapically (Fig. 2); abdomen covered mostly with elongate ovoid scales; length 35.0 mm ................................................................. L. stainesi sp. nov.
– Metaventral process not lanceolate, not constricted subapically (Figs 8, 15, 18); abdomen covered mostly with short ovoid scales .......................................................................................................... 3
3. Paramere apical process of male genitalia not distinctly elongate; bird bill-like; apical process of male genitalia tapered or rounded apically (Figs 5–6, 13–14) .................................................................. 4
– Paramere apical process of male genitalia distinctly elongate; not bird bill-like; apical process of male genitalia spatulate apically (Figs 19–20); length 42.5 mm .................................. L. ratcliffei sp. nov.
4. Paramere apical process of male genitalia tapered apically; paramere basal margin of male genitalia sinuate; paramere lateral margin of male genitalia surface rugose (Figs 9–10); medial posterior margin of pronotum slightly extended posteriorly; posterolateral margin of elytron near posterior angle distinctly infleXed laterally; pygidium with vertical flattened margin subposteriorly; length 38.2 mm ................................................................................................................. L. bezdeki sp. nov.
– Paramere apical process of male genitalia rounded apically; paramere basal margin of male genitalia bisinuate; paramere lateral margin of male genitalia surface rugose (Figs 13–14); medial posterior margin of pronotum distinctly extended posteriorly; posterolateral margin of elytron near posterior angle vertically flattened laterally; pygidium without vertical flattened margin subposteriorly, with sloping downward margin. length 36.0 mm .............................................. L. semperi Brenske, 1896
5. Scales on elytra nearly uniform in size ............................................................................................. 6
– Scales on elytra variable in size; length 23.5–29.5 mm ................ L. pulverulenta Burmeister, 1855
6. Clypeus without medial cleft ............................................................................................................ 7
– Clypeus with medial cleft ................................................................................................................. 8
7. Clypeus strongly convex in males, weakly convex in females, wider than labrum in males, narrow in females; length 28.5–32.5 mm ................................................................... L. reflexa Moser, 1924
– Clypeus weakly convex in males, strongly convex in females, nearly as wide as labrum in both sex; length 22.5–25.1 mm ....................................................................................... L. bakeri Moser, 1924
8. Protibial spur long, extending beyond anterior angle of protibia; length 25.5–30 mm ...................... ................................................................................................................. L. irrorata Chevrolat, 1841
– Protibial spur distinctly shorter, not extending beyond anterior angle of protibia; length 21– 22.5 mm ................................................................................ L. guevarai Calcetas & Adorada, 2017
description: Published as part of Calcetas, Orlando A., 2023, Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896, pp. 184-203 in European Journal of Taxonomy 890 on pages 199-200, DOI: 10.5852/ejt.2023.890.2261, http://zenodo.org/record/8305633