Closterocerus petiolatus Hansson 2022, sp.nov.
Creators: name: Hansson, Christer
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Eulophidae
Closterocerus
Closterocerus petiolatus

description: Closterocerus petiolatus sp.nov.   
(Figs 137–140, 213, 224, 234)  
 Diagnosis. Antenna flattened, scape long with one-third of its length reaching above level of vertex and widest at apex, all flagellomeres distinctly separated (Figs 224); pronotal collar with a carina along anterior margin (Figs 137, 139); mesoscutum with one pair of weak setae in posterior part (Fig. 137); fore wing infuscate with two elongate hyaline parts in apical part of wing, separated by infuscation (Figs 137, 138, 140, 213); hind tarsus with T1–3 white and T4 dark brown (Fig. 138); petiole as long as wide with strong sculpture on dorsal part; length of body female 1.7–2.3mm, male 1.4–2.0mm.   
Female holotype: length of body 2.0mm.  
 Antenna dark brown. Frons metallic bluish-green. Vertex metallic purple. Mesoscutum metallic bluish-purple with posterior one-half of midlobe golden-green. Mesoscutellum golden-green with sides and posterior part metallic bluish-purple. Dorsellum metallic bluish-purple. Propodeum golden-green. Coxae black, femora, mid and hind tibiae dark brown, fore tibia infuscate with ventral part dark brown; fore tarsus infuscate, mid and hind tarsi with T1–3 white and T4 dark brown. Fore wing infuscate with two elongate hyaline parts in apical part of wing, separated by infuscation; hind wing hyaline. Petiole black. Gaster with Gt 1-2 metallic bluish-purple, remaining tergites dark brown with metallic tinges.   
Antenna flattened, scape long with one-third of its length reaching above level of vertex and widest at apex. Frons and vertex with strong reticulation. Frontofacial suture V-shaped. Subtorular sutures present. Occipital margin rounded.  
Pronotal collar with a carina along anterior margin. Mesoscutum with strong reticulation; notauli narrow and complete; midlobe of mesoscutum with one pair of weak setae in posterior part. Mesoscutellum with strong, but weaker than mesoscutum, reticulation. Dorsellum convex and smooth. Propodeum reticulate in posterior two-thirds, otherwise smooth, without a median carina; callus with two setae. Fore wing speculum small and closed; without stigmal hairline and with radial cell hairy.  
 Petiole about as long as wide with strong sculpture on dorsal surface. Gaster ovate; Gt 1 smooth, remaining tergites with weak reticulation.   
Relative measurements: head length dorsal 21; head length frontal 27; head width 46; POL 9; OOL 6.5; lateral ocellus maximum width 3; eye length 18.5; malar space 10.5; mouth width 11; mesosoma length 69; mesosoma width 43; mesoscutellum length 25; mesoscutellum width 25; fore wing length 132; fore wing width 80; marginal vein length 70; postmarginal vein length 3; stigmal vein length 9; fore wing marginal fringe length 8; gaster length 67; gaster width 47.  
Variation in paratype material. Length of body: 1.7–2.3mm. Vertex metallic purple or bluish-green. Fore wing wing infuscate with two elongate hyaline parts in apical part of wing, separated by infuscation, sometimes with a hyaline part below basal one-half of marginal vein.  
Male (Figs 139, 140). Length of body 1.4–2.0mm. Vertex golden-green or golden-red; mesoscutum golden-red with anterior one-third and sidelobes golden-green; mesoscutellum completely golden-red; gaster elongate. Otherwise as in female.  
 Hosts. Unknown.   
 Distribution. Costa Rica.   
 Material examined.   
Type material. Holotype ♀ “ COSTA RICA: San José, Cerro de la Muerte, 20km S Empalme, 2800m, iiiiv.1989, P. Hanson ” (NHMUK). Paratypes (19♀ 6♂, CNC, MZLU, MZUCR, NHMUK): 6♀ with same label data as holotype; 1♀ “ COSTA RICA: San José, 2km SE Empalme, xi.1988, P. Hanson ”; 1♀ “ COSTA RICA: San José, Cerro de la Muerte, 19km S, 3km W Empalme, 2600m, v-vi.1989, Hanson & Godoy”; 1♀ “ COSTA RICA: San José, E.B. Cerro de la Muerte, 3100m, vii.2000, P. Hanson ”; 1♀ “ COSTA RICA: San José, Zurqui de Moravia, 1600m, 10˚3’N, 84˚0’W, v.1992, P. Hanson ”; 2♀ “ COSTA RICA: San José, San Gerardo de Dota, 20-21.ii.2013, J.S. Noyes, NHM (Ent) 2012-91”; 2♀ “ COSTA RICA: Puntarenas, Potrero Grande, Cerro Kasir, 2959m, 19.vii-19.viii.2000, M.Alfaro, LS 349700/566850 #59114”; following from same locality as previous but collected 20.viii.2000 (1♀), 19.ix-19.x.2000 (1♀); 1♀ “ COSTA RICA: Puntarenas, Buenos Aires, Estación Altamira, Sendero a Casa Coca, 1700m, 16.vii-16.viii.2000, D. Rubí, MT, LS 574400 /331750 #84976”; 1♂ “ COSTA RICA: Puntarenas, E.B. Monteverde, 10º20′N, 84º49′W, 1540m, 26.ii.2007, J.S. Noyes, BMNH(Ent) 2010-21”; 2♂ “ COSTA RICA: Highway #2, Km 72, 9˚38’N, 83˚50’W, 2850m, 31.iii.1985, Goulet & Masner”; 1♀ 1♂ “ COSTA RICA: Highway #2, Km 93, 9˚36’N, 83˚45’W, 3200m, 1.iv.1985, Goulet & Masner”; 2♂ “ COSTA RICA: Highway #2, Km 66, 9˚45’N, 84˚10’W, 2400m, 4.iv.1985, Goulet & Masner”.  
 Etymology. Name referring to large petiole, as long as wide, which is unusual but not unique within this genus. Most species has petiole very short, just a narrow band.
description: Published as part of Hansson, Christer, 2022, Eulophidae Of Costa Rica (Hymenoptera: Chalcidoidea), 4 The Genus Closterocerus Westwood S. Str., pp. 1-137 in Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) (Oxford, England) 1 (1) on pages 66-67, DOI: 10.5281/zenodo.6464279