Pterostichus (Wraseiellus) comatus Shi & Sciaky, new species
Creators: name: Shi, Hongliang
name: Sciaky, Riccardo
name: Liang, Hongbin
name: Zhou, Hongzhang
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Pterostichus
Pterostichus comatus

description: Pterostichus (Wraseiellus) comatus Shi & Sciaky, new species
(Figures 6, 22, 32, 60, 70, 97, 101)
Type locality. Guangxi: Maoershan Mt. (N25.87º, E110.42º). The label of holotype does not indicate its altitude or exact locality. But from the altitude information on the two paratypes (1900m), we suppose that this species only lives in the very high altitude of that mountain, which has its highest peak at 2142m.
Type materials. Holotype (MNHN): male, body length = 12.8 mm, board mounted, genitalia dissected and glued on plastic film pinned under specimen, " CHINE (GUANGXI) / MAO'ER SHAN / VII-1995 T.DEUVE"; "Museum Paris"; " HOLOTYPE 3/ Pterostichus (Wraseiellus) / comatus new species / Des. SHI H.L. 2012" [red label] [Figs. 6, 22, 32, 60, 70, 101]. Paratypes: 2 teneral males (IZAS and SCAU), "Guangxi, Maoershan, Sanjiangyuan, 1900m, 2003.8.25, TIAN Mingyi leg." [Fig. 97].
Diagnosis. This new species can be easily distinguished from all the other species of this subgenus by their lateral margins of pronotum with three to five setae on each side, contrasting other species only with a single midlateral seta.
Description. Body length 12.3–12.8 mm; dorsal side black or with elytra slightly brownish; legs dark brown, tarsi light brown; ventral side black or dark brown. Males with elytral microsculpture transverse on the basal half, gradually turning to isodiametric after middle. Head. Frons finely and sparsely punctate, densely in the frontal furrows; antennae reaching elytra basal fifth; 3rd antennomere glabrous except apical setae; males with the terminal labial palpomere tubular. Pronotum cordiform, widest before middle, PW/PL = 1.26–1.32; lateral margins fully rounded near the middle, and strongly sinuate just before hind angles; three to five mid-lateral setae present before middle, hind angles distinct and sharp, rectangular; basal fovea moderately deep, faintly defined; inner groove of basal fovea subparallel to median line, slightly curved to the outside, ended at a distance from the posterior margin of pronotum; outer groove of basal fovea about half length as the inner one, deep and reaching the basal margin of pronotum; ridge between outer groove and lateral margin distinct; area between inner and outer basal foveal grooves rugose and hardly convex; basal fovea area, as well as area between two basal foveae finely and densely punctate; some very fine and sparse punctures also present in the lateral depression and, sometimes, on the lateral area of disc. Elytra with base slightly depressed between 3rd and 5th intervals; humeral teeth very small; intervals slightly convex; striae moderately deep, without punctures; scutellar striae incomplete, forming short grooves, located between 1st and 2nd striae; umbilical pore series on 9th interval sparse in the middle, composed of 14–15 pores (6, 2, 6–7). Ventral side. Proepisterna and metaepisterna finely punctate, posterior half of proepisterna with punctures sparser; terminal sternum of males with a very faint transverse ridge at about anterior fourth, and slightly depressed after the ridge (Fig. 97). Legs. 5th tarsomere with two or three pairs of setae beneath. Male genitalia. Median lobe of male genitalia bent less than 90 degree, apex slightly bent to the ventral side and then the turned to the dorsal side (Fig. 32 A); ventral margin almost straight in the middle, dorsal margin gradually curved; apical orifice large, slightly turned to the left side; apical lamella thick, apex slightly thickened and turned to the dorsal side (Figs. 32 B, 60B); when in dorsal view, apical lamella narrow and long, length about 2.5 times as basal width, twisted to the left (the widest surface of apical lamella orientated to the dorsal-left side of median lobe), strongly bent to the left side (Fig. 32 B); apex of apical lamella oblique to the left (Fig. 60 A). Right paramere fine and long, sickle-form, apex sharp (Fig. 70 B). Female unknown.
Distribution. Only known from the type locality, Maoershan Mt. (Guangxi) (Fig. 110).
Etymology. The name " comatus " comes from Latin, meaning hairy, referring to the additional setae on pronotum lateral margin.
Variation. The three specimens we studied are different in the number of lateral setae of pronotum: the holotype with five lateral setae, but the two paratypes with only three. We suppose that the number of lateral setae of pronotum may vary from individual to individual. Such case is also found in other members of Pterostichini with three or more lateral setae of pronotum.
description: Published as part of Shi, Hongliang, Sciaky, Riccardo, Liang, Hongbin & Zhou, Hongzhang, 2013, A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions, pp. 101-135 in Zootaxa 3664 (2) on pages 112-113, DOI: 10.11646/zootaxa.3664.2.1, http://zenodo.org/record/220503