Mecodema argentum Seldon & Buckley 2019, sp. n.
Creators: name: Seldon, David S.
name: Buckley, Thomas R.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Mecodema
Mecodema argentum

description: Mecodema argentum sp. n.
Figure 19.
Diagnosis: Distinguishable from other North Island Mecodema species by having: 1, a very broad and flat head; 2, elytral interval 7 flat; 3, the apical shape of the penis lobe and form of left paramere distinctive (Figs 19 PL and 19LP).
Description: Length 29–32 mm, pronotal width 7.4–8.9 mm, elytral width 8.5–10 mm. Colour of entire body matte black, except coxae bright reddish-brown, legs matte dark reddish-brown.
Head: Very broad and flat. Vertex smooth, except very fine wrinkles sparsely distributed laterally (Fig. 19); vertexal groove broad and shallow, more impressed laterally (micropunctate and small shallow grooves occassionally present); large supraorbital puncture bearing 4–5 setae; 4–6 poorly defined supraorbital grooves above and anterad eyes, extended onto lateral area of frons; frons (Fig. 9) with a large and shallow depression each side of midline; frontoclypeal suture more defined laterally, obscured by raised hump medially of frons-clypeus junction, tentorial pits indistinct; anterior area of clypeus with a setose puncture each side bearing 2 setae, central groove with a large setose puncture bearing 2–3 setae (central puncture may be absent). Labrum rounded laterally, anterior edge straight, slightly emarginated with 2 proximate central setae, 2 setae each side evenly spaced. Mentum lobes rounded (Fig. 12F), median process very broad and short, slightly angled upward (15°), moderately indentate (Fig. 12B); mentum setae absent. Submentum sclerite constriction narrow, strongly folded with 6 evenly distributed setae. Stipes with 2 basal setae. Gula pits small, suture defined, gula flat and smooth. Gena (Fig. 8) with rugose, transverse wrinkles forming an isodiametric pattern laterally.
Prothorax: Prothoracic carina narrow the entire length, slightly crenulated with 8–12 setae each side (Fig. 19), extended beyond anterior angle; posterior lateral sinuation evidently carinate, parallel; pronotum very broad and flattened, overall shape broadly cordate; midline narrowly defined with anterior medial triangular impression and posterior medial diamond-shaped impression present, disc with transverse wrinkles laterally; pronotal foveae deep and broad; anterior and posterior edges straight. Prosternum (Fig. 2) flat with very fine transverse lines; proepisternum without microsculpture. Procoxal setae absent; protibia distally expanded and shovel-like (Fig. 10B).
Elytra: Broad and flattened, slightly deflected laterally; humeral angle evenly convex (Fig. 6B); basal margin slightly curved and gently sloped to base, interval 1 extended to scutellum, other intervals and striae extended to margin (Fig. 19); lateral carina narrow the entire length, extended beyond humeral angle; humerus with 3–4 closely distributed anterior setose punctures, 4 th or 5 th puncture double-spaced posterad; suture less impressed than striae; striae with small, regularly distributed, asetose punctures, increasing in size laterally, all equally impressed (visible to naked eye); intervals relatively flat but convexity slightly increased laterally; interval microsculpture absent; 7 th strial setal pattern (Fig. 1) with 3 setae in anterior ½, 4–5 setae in posterior ½, setose punctures small.
Ventral surface: Mesepisternum and metepisternum without microsculpture; setose punctures present on mesocoxae (2) and metacoxae (2). Abdominal ventrite 1 lineate (sparsely); ventrite 2 (Fig. 2) finely lineate, may have fine pitting interspersed close to metacoxae; ventrites 3–5 with 1 setose puncture each side of midline; ventrite 6 setae present: ♂ with 1 setose puncture each side of midline, ♀ with 2–3 setose punctures each side located at junction of bluntly-rounded apical edge and curve anterad; ventrites 4 and 5 may be foveate laterally. Anterior metaventrite process (Fig. 2) a triangle with a very broad carina the entire length.
Male genitalia: Apical portion of penis lobe asymmetrically hooked, short apical portion with slight deflection to right of vertical axis (VV) (Fig. 15C); apex shape narrow the entire length and perpendicular (Figs 3 and 19 PL), ventral process curved downward to form a rounded point, then curved upward to form the dorsal process that is a very short, rounded hook (almost hammer-like); penis lobe shaft of equally narrow the entire length (Fig. 19 PL), ventral edge straight (VV), overall length of the penis lobe distinctly curved ventrally (LV) (Fig. 16C). Structures of the endophallus (Fig. 4): lateral form of the apex of the central spicule pointed; dorsal form of the apex of the central spicule greatly expanded with projection; setal (scales) coverage of the apical plate is extensive (76–100%); left setose flange is large; right setose flange is absent. Left paramere basal lobe a large rectangle-shape, with very pronounced anterior hump forming a steep angle (almost perpendicular) to the very short and narrow arm (Fig. 19 LP); narrow terminal lobe; a few short setae extended from apex, then setae increased in length to mid-arm; longer setae extended from ½ way along arm to anterior region of basal lobe (Fig. 19 LP); ventral edge of basal lobe relatively straight. Right paramere narrow and triangular in shape, slightly longer with very long setae extended along apical ½ of ventral edge (Fig. 19 RP); apical ⅓ slightly inwardly curved.
Female genitalia: Basal gonocoxite 1 long and broad at base, but very broad toward gonocoxite 2 with 3–4 well-defined grooves (wrinkles); internal dorso-lateral carina with 3 short robust setae at ventral apex dorsad gonocoxite 2. Gonocoxite 2 triangular, broad basally, narrowed and rounded apically. Ramus (Fig. 5) long and narrow.
Comments: The southern distributional limit along the the Coromandel Peninsula of Mecodema argentum remains unclear, but there are other large broscines present in this area. Mecodema validum is present in the Kauaeranga Valley and Mt Te Aroha, i.e. the southern areas of the Coromandel Peninsula. The widespread species, M. crenaticolle, is present as far north as the Tapu-Coroglen Rd, and M. atrox (a coastal forest dweller) has been collected at Kennedy Bay. However, M. argentum has more shared characters with north Auckland and Northland species, like that of the curvidens species group (see Seldon & Leschen 2011), than the Mecodema species of the Waikato and Bay of Plenty.
Distribution: New Zealand, North Island, northern Coromandel Range, Coromandel town, Trig Tk.
Holotype: NZAC male labelled. Coromandel Ra. 3½ km N. of Coromandel 13.1.93 G.Shirley / HOLOTYPE Mecodema argentum n. sp. design. DS Seldon, TR Buckley 2018.
Paratypes: 2♂, 1♀ NEW ZEALAND, CL, Coromandel Ra., Coromandel town, Trig T Tk, PTs 8–28 Feb 2014, DS Seldon / CAR 159 ♂; CAR 205 ♂; CAR 206 ♀ (AMNZ); 1♂, Kauri Grove Coromandel-Coroglen Rd 22- 1-60 J.I. Townsend [hw] (NZAC); 1♂, Mainridge Pitoki Stream 3 km ESE of Coromandel 13.1.93 G. Sherley [hw] (NZAC).
Etymology: Argentum is Latin for silver, and therefore, Mecodema argentum is named for my good friend Silver Bishop. Silver reminded me a few years ago that I promised to name a species after her when I got the chance, so I’m fulfilling that promise. The name is to be treated as a noun in apposition.
description: Published as part of Seldon, David S. & Buckley, Thomas R., 2019, The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand, pp. 1-148 in Zootaxa 4598 (1) on pages 27-29, DOI: 10.11646/zootaxa.4598.1.1, http://zenodo.org/record/2668063