Clivina quadrataconvexa Balkenohl 2023, nov.sp.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina quadrataconvexa

description: Clivina quadrataconvexa nov.sp. (Figs 7-9)   
T y p e m a t e r i a l: Holotype: ♁, with labels and data: white, handwritten in black ink and black framed, "N-India, UP Rishikesh, 450 m leg. Werner 8/88" (CMBB). Paratypes: 1♁, 2♀♀, same data as holotype (CMBB, OLML); 1♀, same data as holotype but with printed label; 1♁, white, black printed "KISTNA DT. Uppaluru V-51 Biffi" (CMBB).  
E t y m o l o g y: The name refers to the square pronotum which is in frontal view distinctly convex, and is expressed by the Latin adjective quadratus (= sqare) and convexus (= arched) in the feminine genus.  
 D i a g n o s i s: The new species belongs into the Clivina helferi species-group and is externally closer to C. monticola ANDREWES, 1931, and C. mekongensis LESNE, 1896. From both of the species, it differs mainly by the pronotum which is distinctly wider than long. In addition, C. mekongensis differs by the anteriorly sharply advanced anterior angles of the pronotum and the completely dull surface, C. monticola by the laterally sharply projecting posterior angles of the pronotum and its much wider lateral marginal channel.   
D e s c r i p t i o n: Measurements: Length 6.07-7.67 mm (x = 6.35 mm *), width 1.84- 1.98 mm (x = 1.93 mm *), ratio length/width of pronotum 0.84-0.88 (x = 0.86*), ratio length/width of elytra 1.77-1.89 (x = 1.83*); (*n = 6).  
Colour: Glossy. Piceous. Antenna, palpomeres and legs fuscous.  
Head: A quarter narrower than pronotum. Clypeus straight anteriorly, with its lateral teeth slightly projecting; wing of moderate size, not more protruding anteriorly than clypeus, distinctly separated from clypeus by notch; supraantennal plate more or less convex laterally; clypeus, wing, and supraantennal plate finely reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by distinct notch, separated from supraorbital carina by an oblique sulcus. Clypeus transverse, with transverse elevation, separated from frons by a rugose furrow (in some specimens indistinct). Frons moderately convex, with central pore at middle, with scattered small punctures. Supraantennal plate separated from clypeus and frons by deep moderately wide furrow continuing posteriorly up to mid-eye level as wide supraorbital furrow. With blunt distinct supraorbital carina at posterior half of eye. With two supraorbital setigerous punctures situated at mid-eye level and at the posterior end of the eye. Neck constriction developed by punctures and distinct step, step pointed posteriorly at middle. Eye hemispherical, projecting laterally. Gena small. Antenna long, reaching over base of pronotum, antennomeres four to ten elongate (ratio length/width around 1.82). Labrum nearly straight, isodiametrically reticulated, seven setose. Mandible of moderate size. Mentum with lateral lobe oval anteriorly, flattened, isodiametrically reticulated, median tooth longer than lateral lobes, hollowed out.  
Pronotum: Disk moderately convex in lateral view, distinctly convex in frontal view. Quadrate, wider than long. Anterior margin slightly excised. Reflexed lateral margin subcrenulated (lateral view), slightly convex at middle, slightly attenuating in anterior half, widest in posterior third; anterior angle rounded-off; posterior angle marked by distinct tooth, laterally indistinctly projecting. Lateral channel with irregular reticulation at sides, indistinctly foveolate (lateral view), margin from posterior angle to base flattened-convex, joining with basal carina by indistinct knob, basal channel narrow. Median line narrow, sharp, joining anterior transverse line and base; anterior transverse line slightly wider than median line, reaching lateral channel. Surface with microscopic punctures, with group of line-like punctures laterally in basal half, with two clouds of punctures anterior to the line-like punctures (arrangement resembles to an indistinct widened letter Y), few transverse wrinkles laterally, more distinct at base.  
Elytron: Disk flattened in anterior half in lateral view, distinctly and regularly convex in frontal view. Outline long oval, distinctly less than twice as long as wide, with maximum width behind middle. Humerus rounded, formed by lateral margin which joins interval seven. Reflexed lateral margin smooth. Lateral channel moderately wide. Scutellar striole of moderate length; with distinct setigerous puncture at base of first stria, with distinct tubercle at base of third interval. Striae moderately deep, with indistinct punctures, one to four free at base, five and six joining at humerus, seven shortened behind humerus, striae one and two running up to apex, three and four, and five and six joining apically. Intervals moderately convex, convexity more evident laterally, eight carinate apically. Third interval with four setigerous punctures adjoining third stria. Surface of intervals smooth and glossy on disk, with distinct isodiametric reticulation at base, at apical quarter, and on intervals six, seven and eight.  
Hind wings: Fully developed.  
Lower surface: Proepisternum covered with distinct isodiametric reticulation, with some fine transverse wrinkles. Sternites of abdomen finely isodiametrically reticulated. Abdominal sternum VII with the two anal setigerous punctures widely separated, at middle slightly projecting posteriorly where it is flattened, with narrow marginal furrow at middle (in both sexes).  
Legs: Protibia with three spines of moderate length and a small but distinct one basally, sulcate dorsally, movable spur regularly arcuate, surface with longitudinal reticulation. Mesotibia with distinct and moderately long protuberance preapically with acuminate apex and with its seta inserted laterally, with eight strong setae arranged in two rows, with few fine setae at the inner side.  
Male genitalia (Fig. 8): Median lobe regularly curved, with straight apex, apex with flattened and regularly acuminated lamella. Endophallus with group of bristles. Parameres moderately wide, acuminated and distorted in apical third, at apex with one short seta.  
Female genitalia (Fig. 9): Gonocoxites of the falciform type, dimer, gonocoxite-2 nearly straight, scapiform-like, with two long apical setae, with four lateral setae, gonocoxite-1 with two nematiform long setae. Coxites not fused, movable to each other. Epipleurite triangle shaped, with two groups of setae.  
Variation: The six specimens differ slightly in the development of the clypeal-furrow and rugae on the frons. In two of the paratypes, the furrow is developed laterally and indistinct, on others it appears distinct and doubled. In the specimen from Uppaluru, flat furrows extend to the frons. The supraantennal plates vary slightly in its lateral convexity.  
D i s t r i b u t i o n: Known from two localities in India: Rishikesh (Uttar Pradesh), and Uppaluru (Andhra Pradesh).
description: Published as part of Balkenohl, Michael, 2023, Notes on Clivinini species from Asia with the description of a new genus and a new species from Myanmar and the North of India (Coleoptera, Carabidae, Scaritinae), pp. 405-415 in Linzer biologische Beiträge 55 (2) on pages 409-410, DOI: 10.5281/zenodo.10787819