Homalopoda baunos Noyes 2023, sp.nov.
Creators: name: Noyes, John Stuart
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Encyrtidae
Homalopoda
Homalopoda baunos

description: Homalopoda baunos sp.nov.   
(Figs 119-125; Hab. E 20)  
 DIAGNOSIS. Female (length about 1.6-1.9mm): scape pale orange; funicle generally pale orange, mixed brown, clava dark brown; fore leg pale orange, mid leg white mixed pale orange; fore wing (Fig. 119) with apical area generally lightly infuscate; paired hyaline spots at apex of venation separated by about length of marginal vein; no hyaline spot present in middle of wing below marginal vein; head (Fig. 120) about 4.9X as wide as frontovertex; occipital margin with a bristle-like seta behind posterior ocellus; mandible with 4 teeth; an inconspicuous line of silvery setae adjacent to eye above scrobe; scape about 4.9X as long as broad; funicle segments subcylindrical, all at least 1.5X as long as broad (Fig. 122), linear sensilla present only on F3-F4; flagellum about as long as head width; apex of scutellum with two pairs of scale-like setae (Fig. 121); costal cell of fore wing ventrally with 3 lines of setae proximally, a single line of setae in distal half; postmarginal vein very nearly as long as stigmal vein (Fig. 123); linea calva not interrupted (Fig. 119); hind tibia without a subbasal constriction. Male: unknown.   
 Female ( holotype ): length, including ovipositor, 1.89mm; excluding ovipositor, 1.78mm (CPD).   
Head dark brown to black with a distinct metallic lustre; occiput mainly metallic dark green with some blue-green reflections; frontovertex metallic blue-green mixed with some violet; area above scrobes deep violet; scrobes and interantennal prominence metallic dark green with some violet and brassy reflections; mouth margin orange-brown; temple mixed metallic blue-green, dark blue, purple and coppery purple, posteriorly relatively dull; area immediately below lowest part of eye metallic blue-green, gena relatively dull coppery purple; frontovertex with a few, fairly inconspicuous, scattered, translucent, pale brown setae; an inconspicuous line of sparse, translucent silvery setae between eye and scrobe extending to position of malar sulcus; gena with scattered, similarly inconspicuous, translucent, silvery setae; interantennal prominence with two submedian lines of similar setae; maxillary palpus white; radicle pale orange, with a dark brown apical ring; scape pale orange, a little dusky at extreme apex; pedicel dark brown, pale orange at apex and ventrally; funicle generally pale orange, but segments dusky dorsally and proximally so that F1 is mostly dusky pale orange and F6 is mostly pale orange; clava dark brown; thorax dark brown to black with a metallic lustre; pronotum mainly with blue-green and violet sheen, posterior margin medially coppery purple; mesoscutum metallic blue-green with violet reflections, narrowly margined coppery purple laterally and posteriorly; tegula dark brown with a weak coppery purple sheen; axilla dark brown with a very dull coppery purple sheen; scutellum dorsally relatively dull, with very weak coppery purple and dark green reflections, side and apex strongly shining blue-green and green, with some brassy and violet reflections; mesopleuron anteriorly with a moderate violet, brassy and dark blue sheen, medially dark green and brassy, posteriorly coppery purple; all coxae dark brown; fore leg pale orange, tibia with a small, brown, subbasal spot externally and tarsus slightly dusky; mid femur with proximal 0.5X white, apex pale orange, tibia white to very pale orange with a diffuse, subbasal brown ring, spur and tarsus white to very pale orange; hind femur pale orange at extreme base otherwise dark brown, tibia dark brown with apical 0.3X white, tarsus white, apical tarsomere pale orange; fore wing with a pattern of hyaline and infuscate areas as in Fig. 110, with only one pair of hyaline spots; propodeum dark brown with a weak brassy and purple sheen, side with a metallic dark green and dark blue sheen mixed brassy; gaster dark purple-brown, dorsally with a coppery purple sheen, Gt1 with a metallic, dark blue sheen, side and venter with coppery, blue, brassy and green reflections; gonostylus dark brown.  
Head (Fig. 120) about 4.9X as wide as frontovertex, in profile about 1.7X as high as deep, tangent to frontovertex forming an angle of about 55° with that of face occipital margin rounded with a conspicuous, elongate seta present behind each posterior ocellus, seta about as long a width of frontovertex across anterior ocellus; a slender, oval depression adjacent to eye near occipital margin and a small, shallow median depression medially in front of occipital margin; ocelli forming an angle of about 60°; frontovertex clearly narrowing anteriorly; ocellar area with relatively deep, fine, polygonally reticulate sculpture of mesh size slightly less than diameter of eye facet (Fig. 120), area between anterior ocellus and top of scrobes with slightly shallower, finer, slightly transversely elongate polygonally reticulate to imbricate-reticulate sculpture; temple completely smooth and shiny adjacent to eye, otherwise temple and gena with relatively shallow, longitudinally elongate, fine polygonally reticulate sculpture; top of scrobes with shallow, regular, polygonally reticulate sculpture, above torulus almost completely smooth and shiny, elsewhere with shallow sculpture; interantennal prominence and mouth margin with shallow, relatively fine, polygonally reticulate sculpture; antenna as in Fig. 122; scape hardly broadened and flattened, about 4.9X as long as broad; all funicle segments at least about 1.5X as long as broad, F2 shortest, segments slightly broader distad, linear sensilla present only on F3-F4; clava slightly longer than F2-F4 combined, sutures parallel, sensory area small, at apex only giving it a rounded appearance; eye slightly overreaching occipital margin, naked; clypeal margin medially virtually straight; mandible with four teeth, two uppermost very short. Relative measurements: HW 82.5, HH 75, FV 17, POL 11, OOL 0, OCL 14 (shortest line from posterior ocellus to elongate seta on occipital margin), AOL 12, EL 43, EW 43, MS 40, SL 42, SW 8.5.  
Thorax (Fig. 121) with visible part of mesoscutum about 1.5X as broad as long; scutellum about 1.3X as broad as long, apex with 2 pairs of elongate, scale-like setae, longer (inner) pair each about 0.9X as long as scutellum, shorter (outer) pair each about 0.8X as long as scutellum; hind femur about 3.6X as long as broad; fore wing with venation and setation as in Figs 119, 123; costal cell distally with about 10 setae extending dorsally in apical 0.3X; linea calva entire; propodeum medially about 0.07X as long as scutellum. Relative measurements: FWL 116, FWW 48; HWL 84, HWW 22.  
Gaster with hypopygium (Fig. 125) reaching about 0.6X towards apex; syntergum about 0.9X as long as mid tibia, with apex acute; ovipositor (Fig. 124) slightly exserted, the exserted part about 0.4X length of mid tibial spur or about 0.2X mid tibia; second valvifer without any subapical setae. Relative measurements: OL 70.5, GL 20 [MT 48].  
Variation. Females vary in overall length from 1.60-1.89mm, otherwise little variation in material examined.  
Male. Unknown.  
DISTRIBUTION. Costa Rica.  
HOSTS. Unknown.  
MATERIAL EXAMINED.  
 Type material. Holotype E: COSTA RICA, Guanacaste, Cacao (ACG), 1100 m, MT/YPT, 26.i-24. ii.1996 (J.S. Noyes). Paratypes: COSTA RICA, 1E, Puntarenas, Res. Priv. Karen Mogensen, 9°52’N 85°03’W, 305m, 11-12.2018 (J.S. Noyes). Holotype and paratype in NHMUK.
description: Published as part of Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera), pp. 1-921 in Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) (Oxford, England) 2 (11) on pages 69-70, DOI: 10.5281/zenodo.8074943