Typhoctes jamaicensis Fernández & Brothers 2025, sp. nov.
Creators: name: Fernández, Fernando
name: Brothers, Denis J.
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Mutillidae
Typhoctes
Typhoctes jamaicensis

description: Typhoctes jamaicensis Brothers, sp. nov.
(Fig. 3)
Type material. Holotype, ♂, JAMAICA, Kingston Parish, The Palisadoes [approx. 17°56’40”N 76°47’10”W], 14 November 1986, W.J. Pulawski collector (CAS 2051222); Paratypes, 2♂, data as for holotype (CAS 2051223 & CAS 2051224).
Diagnosis. Male. Frons with fairly dense white simple decumbent setae forming a distinct lateral whorl; clypeus densely covered with long thickened semi-decumbent subplumose white setae mostly concealing the cuticle and fewer semi-decumbent thin finely subplumose fuscous setae. Lateral face of pronotum moderately punctate on anterior half, with distinct longitudinal costulation on posterior half but smooth along posterior margin; disk of propodeum densely longitudinally costulate medially but mainly densely coarsely punctate, declivity finely and sparsely punctate; lateral face of fused metepisternum-propodeum densely longitudinally costulate but densely punctate on dorsal quarter. Mid and hind tibial spurs white with extreme tip blackened. Female. Unknown.
Description. Male. Length 5.7 (5.6–5.7) mm. Body shining black; appendages and wing veins very dark reddish brown; mandible and tegula dark reddish brown; mid and hind tibial spurs white with extreme tip blackened; wings infumated throughout but very slightly darker on apical third. Vestiture of decumbent and erect, silvery white and fuscous to black hairs, forming the following notable patterns: frons with fairly dense white simple decumbent setae forming a distinct whorl on each side, numerous scattered long thin erect dark sparsely subplumose setae and a few erect subplumose white and black setae between antennal bases, clypeus densely covered with long thickened semi-decumbent subplumose white setae mostly concealing the cuticle and fewer semi-decumbent thin finely subplumose fuscous setae, gena with simple decumbent and erect finely subplumose white setae, mandible with erect apparently simple to finely subplumose fuscous setae; mesosoma generally with fairly dense decumbent apparently simple to finely subplumose white setae (some pale fuscous on scutum) and sparse apparently simple to finely subplumose dark fuscous erect setae antero-dorsally; legs with white setae except tarsal setae pale fuscous; T1–T6 fairly sparsely covered with erect finely subplumose white setae; T2–T6 with scattered erect finely subplumose fuscous setae near posterior margin becoming denser posteriorly; T7 (pygidium) with dense short erect subplumose fuscous setae; T1 and T2 with conspicuous posterior fringe of semi-erect thickened white setae; T3 with sparse fringe of fine whitish setae; T4–T6 with very inconspicuous sparse fringe of fine fuscous setae; metasomal sterna with sparse erect finely subplumose white setae, a very few scattered long erect finely subplumose fuscous setae on S4–S6; S8 (hypopygium) with scattered erect finely subplumose and denser shorter semi-decumbent fuscous setae.
Head. 1.27 (1.21–1.27) × as wide as high, 1.02 (0.97–1.02) × width of mesosoma; narrowest part of frons 0.80 (0.76–0.80) × eye height. Clypeus fairly densely and moderately punctate, almost flat but weakly convex medio-dorsally. Frons slightly convex, with very fine well separated punctures; ventral width between eyes 0.78 (0.78– 0.81) × dorsal width between eyes. Median ocellus 1.38 (1.22–1.38) × as wide as lateral ocellus, 0.22 (0.22–0.23) × narrowest width of frons; POL 0.75 (0.73–0.76) × OOL, POL 1.50 (1.45–1.60) × LOL. Eye large, shallowly reniform, eye height 0.73 (0.69–0.73) × head height; malar space short, 0.07 (0.06–0.08) × eye height. Antennal sockets large, oval; ratios of lengths of antennal segments 1–4 about 20:7:12:19 (paratypes 18:7:11:17), scape 2.22 (2.22–2.25) × as long as wide.
Mesosoma. 1.67 (1.61–1.67) × as long as wide. Pronotum 1.62 (1.62–1.77) × as wide as long, slightly widening posteriad, 2.75 (2.65–2.75) × as wide as mid-length, posterior margin evenly shallowly concave; epaulet an inconspicuous diagonal tuft of short pale setae mainly concealed beneath patch of converging decumbent white setae on weakly convex humeral tubercle; dorsum with well-marked spaced moderate punctures irregularly interspersed with fine punctures, punctation sparse medially but dense along posterior margin; lateral face scarcely concave dorsally, moderately punctate on anterior half, with distinct longitudinal costulation on posterior half but smooth along posterior margin. Mesoscutum densely moderately punctate anteriorly, punctures becoming larger and sparser posterad and surface smooth postero-medially, without notauli, shallow well defined parapsidal groove extending two-thirds scutum length from postero-lateral angle. Scutellum finely longitudinally costate but costae interspersed with coarse punctures on lateral third, narrow anterior margin and broader posterior margin smooth. Metanotum smooth with moderate ill-defined irregular punctures medially and shallow coarse to distinct dense posteriorly diverging costae laterally. Disk of propodeum mainly densely coarsely rugose-punctate but densely longitudinally costulate on narrow baso-median band, declivity very finely and sparsely punctate with few transverse costulae at postero-ventral extremity. Mesopleuron evenly gibbose, coarsely and densely punctate but less so postero-ventrally. Metepimeron weakly convex, clearly distinguishable, densely and finely punctate on posterior two-thirds, smooth along anterior margin. Lateral face of fused metepisternum-propodeum densely longitudinally costulate but densely punctate on dorsal quarter.
Wings. Fore wing 2.11 (1.99–2.11) × as long as mesosoma, 3.35 (3.26–3.35) × as long as broad; marginal cell slightly wider basally than apically, with straight posteriorly rounded apex, 2.79 (2.74–2.79) × as long as high, 0.69 (0.68–0.69) × as long as first submarginal cell; second submarginal cell 2.43 (2.13–2.43) × as long as high, 0.84 (0.84–0.86) × as long as marginal cell, 0.58 (0.58–0.59) × as long as first submarginal cell; third submarginal cell 1.15 (1.15–1.20) × as long as high, 0.58 (0.58–0.61) × as long as marginal cell, 0.40 (0.40–0.42) × as long as first submarginal cell. Hind wing with 2 (or 2) basal hamuli, and 7 (or 7 and 8) apical hamuli.
Legs. Dorsal tooth on hind coxa a low convex elongate lamina postero-dorsally. Mid and hind tibiae each with 2 longitudinal rows of 2 or 3 black bristles dorsally and 2 or 3 similar bristles apically. Longer spur of hind tibia 0.75 (0.73–0.77) × as long as hind basitarsus. Claws slender, with small tooth near middle of ventral margin.
Metasoma. 2.79 (2.79–2.95) × as long as wide. T1 strongly constricted into anterior narrow neck, bulging dorsally on posterior third, constricted again at junction with T2, 1.31 (2.31–2.39) × as long as wide and 0.68 (0.68– 0.70) × as wide as T2; indistinguishably fused with S1 on anterior third; entirely sparsely moderately punctate, smooth interspaces at least the width of punctures. T2 similarly punctate as T1 on anterior half but punctures becoming denser and finer posteriorly with dense mixture of moderate and fine punctures; anterolaterally with short sparse and pale felt line 0.23 (0.17–0.23) × lateral length of T2, its anterior extremity at 0.09 (0.04–0.10) × lateral length of T2. T3–T6 with very dense mixture of small and very fine punctures. Pygidium (T7) almost square, dorsally flattened and posteriorly slightly retuse, with dense small punctures, posterior margin straight medially and abruptly convex laterally. S1 mainly flattened, smooth and shining with very few widely scattered small punctures at about anterior third. S2–S6 punctation similar to corresponding terga but much sparser. S7 very short, smooth. Hypopygium (S8) almost square, flattened, posterior margin weakly convex, with fairly dense small punctures.
Variation. The median costulate region on the propodeal disk varies in extent and distinctness, from an area about one-sixth of the width of the propodeum and half of its length with fine regular longitudinal costulae (in the holotype), to a poorly defined area about one-quarter the width of the propodeum but extending only about a fifth of its length with irregular weak costulae interrupted by several punctures (Fig. 3C). The paratypes have the pygidium extremely dark reddish brown rather than black.
Remarks. All three specimens are glued on the right side to narrow card points.
Distribution. Jamaica.
Etymology. The name refers to the country of origin of the type series. It is an adjective.
description: Published as part of Fernández, Fernando & Brothers, Denis J., 2025, New species of Typhoctes (Hymenoptera, Thynnoidea, Chyphotidae) from Colombia and Jamaica, pp. 123-138 in Zootaxa 5633 (1) on pages 128-130, DOI: 10.11646/zootaxa.5633.1.6, http://zenodo.org/record/15372929