Perrona rosaliae Harzhauser & Landau & Janssen 2022, nov. comb.
Creators: name: Harzhauser, Mathias
name: Landau, Bernard
name: Janssen, Ronald
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Neogastropoda
Clavatulidae
Perrona
Perrona rosaliae
Clavatula

description: Perrona rosaliae (Hoernes & Auinger, 1891) nov. comb.   
 Figs 38D 1 –D 3, E 1 –E 3, 1F 1 –F 3, 4T, 5, 7    Clavatula Rosaliae —Hoernes 1891: 130 [nomen nudum].   
 Pleurotoma asperulata Lam. — Hörnes 1854: 341 (partim), pl. 37, figs 2a–b (only) [non Megaclavatula asperulata   
(Lamarck, 1822)].  
 * Pleurotoma (Clavatula) Rosaliae nov. form.— Hoernes & Auinger 1891: 347, pl. 47, figs 2–3.   
 C [lavatula]. (C [lavatula]) rosaliae (R. Hörn. et Au.) — Sieber 1958a: 157.   
 Clavatula (C.) cf. rosaliae (R. Hörn. Auing.) — Sieber 1958b: 150.   
 Clavatula (Clavatula) asperulata (Lamarck) — Schultz 1998: 74, pl. 30, fig. 1 [non Megaclavatula asperulata (Lamarck, 1822)].   
 non Clavatula rosaliae Hö. Au. — Boettger 1906: 62. non Pleurotoma rosaliae R. Hoern. und Auing. — Moisescu 1955b: 262. non Clavatula (Clavatula) cf. rosaliae R. Hörn. et Auing. — Pavlovsky 1957: 53, pl. 2, figs 2a–b [? = Megaclavatula laevigata (Eichwald, 1830)]. non Clavatula (Clavatula) rosaliae (Hoernes, Auinger, 1891) — Nikolov 1994: 57, pl. 6, figs 16–17 [? = Megaclavatula laevigata (Eichwald, 1830)]. non Clavatula suturalis rosaliae (Hoernes R. & Auinger M. 1890) — Katona et al. 2011: 9, pl. 1, fig. 8 [= Megaclavatula polonica (Pusch, 1837)].   
 Type material. Lectotype: NHMW 1999z0098/0002a, Vienna /Pötzleinsdorf (Austria), SL: 45.9 mm, MD: 18.5 mm, illustrated in Hoernes & Auinger (1891, pl. 47, fig. 2), designated by Nikolov (1994: 57), figs 38E 1 –E 3, 4 T. Paralectotype: NHMW 1999z0098/0002b, Vienna /Pötzleinsdorf (Austria), SL: 35.2 mm, MD: 12.2 mm, illustrated in Hoernes & Auinger (1891, pl. 47, fig. 3), figs 38F 1 –F 3.   
 Additional studied material. NHMW A1382, Enzesfeld (Austria), SL: 68.8 mm, MD: 23.4 mm, illustrated in Schultz (1998, pl. 30, fig. 1), figs 38D 1 –D 3; NHMW 1846 /0037/0314, 21 spec., Vienna / Pötzleinsdorf (Austria); NHMW 2015 /0389/0478, 2 spec., Enzesfeld (Austria).   
 Revised description. Shell large, solid, moderately broad fusiform with weakly coronate spire; apical angle 31– 38°. Protoconch not preserved. Teleoconch of at least nine whorls. Early teleoconch whorls with narrow indistinctly beaded subsutural cord, weakly concave mid-portion with strongly opisthocline axial riblets, with swollen abapical ends forming suprasutural row of beads. On fifth whorl sub- and suprasutural cords loose beads becoming smooth, rounded cords, separated by concave mid-portion. Faint spiral threads on entire shell surface, most prominent on concave area. On sixth to seventh whorl profile subcylindrical, subsutural spiral cord becoming broad, flattened subsutural collar with small, pointed spines, on weakly concave mid-portion spirals weaken. Suprasutural cord largely covered by succeeding whorl, fully on penultimate whorl. Suture narrowly incised, linear. Last whorl 60% of total height; broad flattened subsutural collar, weakly carinate mid-collar, with about 14 small spines placed on carina, narrow groove delimiting collar. Subsutural ramp broad. Weakly concave, shoulder placed at level of outer lip insertion, sharply angled by narrow cord bearing small tubercles. Base angled by similar narrow tubercular cord, moderately constricted. Siphonal fasciole weakly swollen, slightly twisted. Weak spirals over entire surface, stronger and of alternate strength over fasciole. Aperture moderately wide, pyriform. Outer lip not thickened, smooth within. Anal sinus deep, moderately wide, asymmetrically U-shaped, with apex mid-ramp. Siphonal canal moderately long and narrow, slightly deflected, shallowly notched at tip. Columella strongly excavated, weakly twisted at fasciole, smooth. Columellar and parietal callus thin, adherent, poorly delimited.   
 Discussion. This species is reminiscent of Megaclavatula nov. gen. species in shape and sculpture, but placement in that genus is unlikely due to the Perrona - type sculpture of the early teleoconch. The early spire of Perrona rosaliae (Hoernes & Auinger, 1891) is identical to that of Perrona styriaca (Hilber, 1879), but P. rosaliae has a deeper anal sinus and more prominent spiral cords. Both species are examples of weakly coronate Perrona species with Megaclavatula -convergent shell shapes. An additional example is Perrona francisci (Toula, 1901) from the Serravallian of Turkey (see Landau et al. 2013, pl. 47, figs 5–6, pl. 74, figs 8–9).   
 The type specimens from Vienna /Pötzleinsdorf (Austria) represent two subadult specimens. Therefore, we illustrate a fully grown specimen from Enzesfeld (Austria), which shows that Perrona rosaliae ranges in the same size-class as M. laevigata, M. grunerti and M. polonica. There is a general misconception of this species in the literature and none of the literature records except from the original descriptions from Vienna /Pötzleinsdorf (Austria) are conspecific with P. rosaliae. Specimens referred to M. rosaliae in the literature lack spiral sculpture and represent other species, such as Megaclavatula laevigata (Eichwald, 1830) (e.g., Pavlovsky 1957) or M. polonica (Pusch, 1837) (e.g., Katona et al. 2011).   
 Paleoenvironment. Inner neritic sandy bottoms based on the assemblage from Vienna /Pötzleinsdorf (own data).   
 Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: Vienna /Pötzleinsdorf, Enzesfeld (Austria) (Hoernes & Auinger 1891; Sieber 1958b; hoc opus).
description: Published as part of Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, pp. 1-172 in Zootaxa 5123 (1) on pages 130-131, DOI: 10.11646/zootaxa.5123.1.1, http://zenodo.org/record/6399827