Sweltsa coloradensis
Creators: name: Stark, Bill P.
name: Baumann, Richard W.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Plecoptera
Chloroperlidae
Sweltsa
Sweltsa coloradensis

description: <i>Sweltsa coloradensis</i> (Banks) (Figs. 1-15, 36-37, 42, 45) http://lsid.speciesfile.org/urn:lsid: Plecoptera.speciesfile.org: TaxonName:3555 <i>Chloroperla coloradensis</i> Banks, 1898 -1899:199. Holotype &amp;female; (Museum of Comparative Zoology), Colorado <i>Alloperla coloradensis</i>: Needham &amp; Claassen, 1925:113 (In part) <i>Alloperla</i> (<i>Sweltsa</i>) <i>coloradensis</i>: Ricker, 1943:135 <i>Sweltsa coloradensis</i>: Illies, 1966: 451 Material examined. UNITED STATES: California: <i>Modoc Co</i>., Rush Creek, Hwy 299, Upper Rush Creek Campground, 24 May 2007, R.W. Baumann, B.C. Kondratieff, 1&amp;male; (BYU). <b>Colorado:</b> <i>Eagle Co</i>., Deep Creek, 17 Rd, 27 June 1997, B.C. Kondratieff, 11&amp;male;, 1&amp;female; (CSUIC). <i>Gilpin Co</i>., South Boulder Creek, 2 miles W Tolland, 7 July 1991, B.C. Kondratieff, R. Durfee, 16&amp;male;, 10&amp;female; (CSUIC). <i>Grand Co</i>., Colorado River, Kawuneeche Valley, Rocky Mountain National Park, 18 June 1980, B.C. Kondratieff, 3&amp;male;, 6&amp;female; (CSUIC). <i>Gunnison Co</i>., Quartz Creek, 76 Rd, N of Pitkin, 7 July 2008, K.W. Stewart, 25&amp;male;, 38&amp;female; (BPSC). <b>Idaho:</b> <i>Bannock Co</i>., Mill Creek, Summit Campground, 5 June 2010, B. Stark, R.W. Baumann, 2&amp;male; (BPSC). <b>Nevada:</b> <i>Elko Co.,</i> Lamoille Creek, Thomas Canyon Campground, 40° 39’ N, - 115° 24.4’ W. 23 June 2006, S.M. Clark, 7&amp;male;, 10&amp;female; (BYU). <i>Humboldt Co</i>., Indian Creek, Santa Rosa Range, 29 June 1999, A.L. Sheldon, 10&amp;male;, 19&amp;female; (BYU). Mahogany Creek, Summit Lake Mountain, 1 July 1999, A.L. Sheldon, 16&amp;male;, 14&amp;female; (BYU). <i>Nye Co</i>., South Fork Pine Creek, Toquima Range, 7 July 1999, A.L. Sheldon, 14&amp;male;, 5&amp;female; (BYU). <i>Washoe Co</i>., Rock Creek, Granite Range, 5 June 1998, A.L. Sheldon, 1&amp;male; (BYU). <i>White Pine Co</i>., Berry Creek, Schell Creek Range, Berry Creek Campground, 24 June 2006, R.W. Baumann, B.C. Kondratieff, 15&amp;male;, 9&amp;female; (BYU). Timber Creek, Timber Creek Campground, Schell Creek Range, 39° 24’ N, - 114° 38’ W, R.W. Baumann, B.C. Kondratieff, 35&amp;male;, 14&amp;female; (BYU). <b>New Mexico:</b> <i>Rio Arriba Co.,</i> Rio de las Vallecitos, 4 miles N Cañón Plaza, 11 June 1985, R.L. Hassage, 1&amp;male; (BYU). <i>San Miguel Co</i>., Terrero, Pecos River, 10 June 1974, B. Stark, T. Wolfe, 3&amp;male;, 6&amp;female; (BPSC). <b>Oregon:</b> <i>Harney Co</i>., McCoy Creek, 1 mile above Fish Lake, Steens Mountains, 20 July 1969, S.G. Jewett, Jr. 3&amp;male;, 10&amp;female; (BYU). <i>Umatilla Co</i>., Camas Creek, Hwy 395, Ukiah Dale State Park, 16 May 2001, B. Stark, K.W. Stewart, J.B. Sandberg, 3&amp;male;, 2&amp;female; (BPSC). North Fork John Day River, Hwy 395, 12 miles S Ukiah, 16 May 2001, B. Stark, K.W. Stewart, J.B. Sandberg, 5&amp;male;, 1&amp;female; (BPSC). <i>Union Co</i>., Grande Ronde River, Bird Track Springs Campground, Hwy 244, 45° 18.027’ N, - 118° 18.427’ W, 15 May 2014, B. Stark, A. Harrison, 1&amp;male; (BPSC). <b>South Dakota:</b> <i>Lawrence Co</i>., Spearfish Creek, Hwy 85, Cheyenne Crossing, 7 June 1995, R.W. Baumann, B. Huntsman, 10&amp;male;, 14&amp;female; (BYU). <b>Utah:</b> <i>Box Elder Co</i>., Clear Creek, Raft River Mountains, 18 June 1979, R.W. Baumann, G.M. Webb, 23&amp;male;, 20&amp;female; (BYU). <i>Juab Co</i>., Granite Creek, Deep Creek Mountains, 14 April 1984, R.W. Baumann, 18&amp;male;, 5&amp;female; (BYU). <i>Salt Lake Co</i>., Big Cottonwood Creek, 22 June 1974, B. Stark, 10&amp;male;, 5&amp;female; (BPSC). <i>Sanpete Co.,</i> Willow Creek, Ephraim Canyon, 39.29707 N, - 111.52377 W, 28 June 2005, Johnson, Anderson, 14&amp;male;, 16&amp;female; (BYU). <i>Tooele Co</i>., Harker Creek, Harker Canyon, Sheep Rock Mountains, 39° 59’ N, - 112° 39’ W, 30 May 2012, R.W. Baumann, G.M. Webb, 8&amp;male;, 5&amp;female; (BYU). South Willow Creek, Boy Scout Campground, Stansbury Mountains, 11 June 2007, R.W. Baumann, K. Anderson, 20&amp;male;, 21&amp;female; (BYU). <i>Wasatch Co</i>., Cascade Springs, Uintah National Forest, 27 May 2000, B. Stark, 8&amp;male;, 4&amp;female; (BPSC). Daniels Canyon, Whiskey Springs Picnic Area, 40° 24.677’ N, - 111° 19.278’ W, 13 June 2015, B. Stark. 7&amp;male;, 12&amp;female; (BPSC). <b>Wyoming:</b> <i>Big Horn Co</i>., West Tensleep Creek, N of West Tensleep Lake, 19 July 2005, D.E. Martin, 2&amp;male; (BYU). <i>Fremont Co</i>., Jakeys Creek, Hwy 26, 5 miles E of Dubois, 20 July 1967, R.W. Baumann, 2&amp;male;, 2&amp;female; (BYU). <i>Washakie Co</i>., 12.5 km ESE Tensleep, 25 June 1996, K.B. Miller, 9&amp;male;, 10&amp;female; (CSUIC). <b>Adult habitus.</b> Described by Needham &amp; Claassen (1925). Male forewing length 7.0- 7.5 mm, female 8.5-9.0 mm. Head and pronotum pale yellow with dark markings; ocelli connected by a V-shaped dark brown mark, M-line and anteromedian area of head forward of M-line dark brown. Pronotum pale yellow with almost complete dark brown margins and rugosities (Fig. 42). Abdomen brown with subtle, slightly darker median brown stripe. Cerci pale, antennae pale basally but brown in apical two thirds. Wing membrane pale, veins pale brown. <b>Male epiproct</b> (n=12). Maximum width 320-370 μm, length 360-450 μm. Base relatively narrow, but gradually expanded beyond cowl, and reaching maximum width across lateral projections of epiproct body. Shoulders project conspicuously beyond hook base (Figs. 1-13, 45). <b>Aedeagus</b> (n=3). Length 500-524 μm, width 590-760 μm. Entirely membranous; apex with a pair of small finger-shaped lobes in a parallel or V-pattern, apicolateral margins, each bearing a large, projecting, rounded lobe, and mesoventral area with a pair of eye-like lobes (Figs. 14-15); Fingershaped lobes bear scattered setae of variable length, apicolateral lobes bare; mesoventral lobes bare, but each with a transverse groove. Median ventral field below mesoventral eye-shaped lobes with a low nose-like tubercle. <b>Female subgenital plate.</b> Posterior margin of plate projecting over basal third to half of sternum 9; plate truncate, to slightly concave along posterior margin. Plate usually darkly pigmented over most of surface, contrasting with the pale sternum 9 and the pale basal and lateral areas of sternum 8 (Figs. 36-37). <b>Diagnosis and distribution.</b> This species, in which the epiproct shoulders conspicuously project beyond the base of the epiproct hook (Fig. 45), is the common form in the central Rockies and Great Basin, and the probable “true” <i>S. coloradensis.</i> Published records exist for <i>S. coloradensis</i> from Alberta, British Columbia and Yukon Territory in Canada, and from Arizona, California, Colorado, Idaho, Montana, Nevada, New Mexico, Oregon, South Dakota, Utah, Washington and Wyoming in the United States (DeWalt et al. 2017; Surdick 1985). Many of the published records will apply to one of the two newly described species below. The Montana, Washington, Alberta, British Columbia and Yukon specimens examined by us have all been determined as <i>S. lyrata</i> sp.n., and the Arizona and western New Mexico specimens are determined as <i>S. mogollonica</i> sp. n. Specimens from the Central Rocky Mountains of Colorado and Utah are identified as <i>S. coloradensis</i>, Some states (e.g. Idaho and New Mexico) have populations of at least two members of the complex. We have recognized populations of <i>S. coloradensis</i> from the sites listed in the “Material examined” section above. Material examined. UNITED STATES: California: <i>Modoc Co</i>., Rush Creek, Hwy 299, Upper Rush Creek Campground, 24 May 2007, R.W. Baumann, B.C. Kondratieff, 1&amp;male; (BYU). <b>Colorado:</b> <i>Eagle Co</i>., Deep Creek, 17 Rd, 27 June 1997, B.C. Kondratieff, 11&amp;male;, 1&amp;female; (CSUIC). <i>Gilpin Co</i>., South Boulder Creek, 2 miles W Tolland, 7 July 1991, B.C. Kondratieff, R. Durfee, 16&amp;male;, 10&amp;female; (CSUIC). <i>Grand Co</i>., Colorado River, Kawuneeche Valley, Rocky Mountain National Park, 18 June 1980, B.C. Kondratieff, 3&amp;male;, 6&amp;female; (CSUIC). <i>Gunnison Co</i>., Quartz Creek, 76 Rd, N of Pitkin, 7 July 2008, K.W. Stewart, 25&amp;male;, 38&amp;female; (BPSC). <b>Idaho:</b> <i>Bannock Co</i>., Mill Creek, Summit Campground, 5 June 2010, B. Stark, R.W. Baumann, 2&amp;male; (BPSC). <b>Nevada:</b> <i>Elko Co.,</i> Lamoille Creek, Thomas Canyon Campground, 40° 39’ N, - 115° 24.4’ W. 23 June 2006, S.M. Clark, 7&amp;male;, 10&amp;female; (BYU). <i>Humboldt Co</i>., Indian Creek, Santa Rosa Range, 29 June 1999, A.L. Sheldon, 10&amp;male;, 19&amp;female; (BYU). Mahogany Creek, Summit Lake Mountain, 1 July 1999, A.L. Sheldon, 16&amp;male;, 14&amp;female; (BYU). <i>Nye Co</i>., South Fork Pine Creek, Toquima Range, 7 July 1999, A.L. Sheldon, 14&amp;male;, 5&amp;female; (BYU). <i>Washoe Co</i>., Rock Creek, Granite Range, 5 June 1998, A.L. Sheldon, 1&amp;male; (BYU). <i>White Pine Co</i>., Berry Creek, Schell Creek Range, Berry Creek Campground, 24 June 2006, R.W. Baumann, B.C. Kondratieff, 15&amp;male;, 9&amp;female; (BYU). Timber Creek, Timber Creek Campground, Schell Creek Range, 39° 24’ N, - 114° 38’ W, R.W. Baumann, B.C. Kondratieff, 35&amp;male;, 14&amp;female; (BYU). <b>New Mexico:</b> <i>Rio Arriba Co.,</i> Rio de las Vallecitos, 4 miles N Cañón Plaza, 11 June 1985, R.L. Hassage, 1&amp;male; (BYU). <i>San Miguel Co</i>., Terrero, Pecos River, 10 June 1974, B. Stark, T. Wolfe, 3&amp;male;, 6&amp;female; (BPSC). <b>Oregon:</b> <i>Harney Co</i>., McCoy Creek, 1 mile above Fish Lake, Steens Mountains, 20 July 1969, S.G. Jewett, Jr. 3&amp;male;, 10&amp;female; (BYU). <i>Umatilla Co</i>., Camas Creek, Hwy 395, Ukiah Dale State Park, 16 May 2001, B. Stark, K.W. Stewart, J.B. Sandberg, 3&amp;male;, 2&amp;female; (BPSC). North Fork John Day River, Hwy 395, 12 miles S Ukiah, 16 May 2001, B. Stark, K.W. Stewart, J.B. Sandberg, 5&amp;male;, 1&amp;female; (BPSC). <i>Union Co</i>., Grande Ronde River, Bird Track Springs Campground, Hwy 244, 45° 18.027’ N, - 118° 18.427’ W, 15 May 2014, B. Stark, A. Harrison, 1&amp;male; (BPSC). <b>South Dakota:</b> <i>Lawrence Co</i>., Spearfish Creek, Hwy 85, Cheyenne Crossing, 7 June 1995, R.W. Baumann, B. Huntsman, 10&amp;male;, 14&amp;female; (BYU). <b>Utah:</b> <i>Box Elder Co</i>., Clear Creek, Raft River Mountains, 18 June 1979, R.W. Baumann, G.M. Webb, 23&amp;male;, 20&amp;female; (BYU). <i>Juab Co</i>., Granite Creek, Deep Creek Mountains, 14 April 1984, R.W. Baumann, 18&amp;male;, 5&amp;female; (BYU). <i>Salt Lake Co</i>., Big Cottonwood Creek, 22 June 1974, B. Stark, 10&amp;male;, 5&amp;female; (BPSC). <i>Sanpete Co.,</i> Willow Creek, Ephraim Canyon, 39.29707 N, - 111.52377 W, 28 June 2005, Johnson, Anderson, 14&amp;male;, 16&amp;female; (BYU). <i>Tooele Co</i>., Harker Creek, Harker Canyon, Sheep Rock Mountains, 39° 59’ N, - 112° 39’ W, 30 May 2012, R.W. Baumann, G.M. Webb, 8&amp;male;, 5&amp;female; (BYU). South Willow Creek, Boy Scout Campground, Stansbury Mountains, 11 June 2007, R.W. Baumann, K. Anderson, 20&amp;male;, 21&amp;female; (BYU). <i>Wasatch Co</i>., Cascade Springs, Uintah National Forest, 27 May 2000, B. Stark, 8&amp;male;, 4&amp;female; (BPSC). Daniels Canyon, Whiskey Springs Picnic Area, 40° 24.677’ N, - 111° 19.278’ W, 13 June 2015, B. Stark. 7&amp;male;, 12&amp;female; (BPSC). <b>Wyoming:</b> <i>Big Horn Co</i>., West Tensleep Creek, N of West Tensleep Lake, 19 July 2005, D.E. Martin, 2&amp;male; (BYU). <i>Fremont Co</i>., Jakeys Creek, Hwy 26, 5 miles E of Dubois, 20 July 1967, R.W. Baumann, 2&amp;male;, 2&amp;female; (BYU). <i>Washakie Co</i>., 12.5 km ESE Tensleep, 25 June 1996, K.B. Miller, 9&amp;male;, 10&amp;female; (CSUIC). <b>Adult habitus.</b> Described by Needham &amp; Claassen (1925). Male forewing length 7.0- 7.5 mm, female 8.5-9.0 mm. Head and pronotum pale yellow with dark markings; ocelli connected by a V-shaped dark brown mark, M-line and anteromedian area of head forward of M-line dark brown. Pronotum pale yellow with almost complete dark brown margins and rugosities (Fig. 42). Abdomen brown with subtle, slightly darker median brown stripe. Cerci pale, antennae pale basally but brown in apical two thirds. Wing membrane pale, veins pale brown. <b>Male epiproct</b> (n=12). Maximum width 320-370 μm, length 360-450 μm. Base relatively narrow, but gradually expanded beyond cowl, and reaching maximum width across lateral projections of epiproct body. Shoulders project conspicuously beyond hook base (Figs. 1-13, 45). <b>Aedeagus</b> (n=3). Length 500-524 μm, width 590-760 μm. Entirely membranous; apex with a pair of small finger-shaped lobes in a parallel or V-pattern, apicolateral margins, each bearing a large, projecting, rounded lobe, and mesoventral area with a pair of eye-like lobes (Figs. 14-15); Fingershaped lobes bear scattered setae of variable length, apicolateral lobes bare; mesoventral lobes bare, but each with a transverse groove. Median ventral field below mesoventral eye-shaped lobes with a low nose-like tubercle. <b>Female subgenital plate.</b> Posterior margin of plate projecting over basal third to half of sternum 9; plate truncate, to slightly concave along posterior margin. Plate usually darkly pigmented over most of surface, contrasting with the pale sternum 9 and the pale basal and lateral areas of sternum 8 (Figs. 36-37). <b>Diagnosis and distribution.</b> This species, in which the epiproct shoulders conspicuously project beyond the base of the epiproct hook (Fig. 45), is the common form in the central Rockies and Great Basin, and the probable “true” <i>S. coloradensis.</i> Published records exist for <i>S. coloradensis</i> from Alberta, British Columbia and Yukon Territory in Canada, and from Arizona, California, Colorado, Idaho, Montana, Nevada, New Mexico, Oregon, South Dakota, Utah, Washington and Wyoming in the United States (DeWalt et al. 2017; Surdick 1985). Many of the published records will apply to one of the two newly described species below. The Montana, Washington, Alberta, British Columbia and Yukon specimens examined by us have all been determined as <i>S. lyrata</i> sp.n., and the Arizona and western New Mexico specimens are determined as <i>S. mogollonica</i> sp. n. Specimens from the Central Rocky Mountains of Colorado and Utah are identified as <i>S. coloradensis</i>, Some states (e.g. Idaho and New Mexico) have populations of at least two members of the complex. We have recognized populations of <i>S. coloradensis</i> from the sites listed in the “Material examined” section above.
description: Published as part of <i>Stark, Bill P. &amp; Baumann, Richard W., 2018, Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae), pp. 30-43 in Illiesia 14 (2)</i> on pages 34-36, DOI: 10.5281/zenodo.4761184
description: {"references": ["Banks, N. 1898 - 1899. Descriptions of new North American neuropteroid insects. Transactions of the American Entomological Society, 25: 199 - 218. https: // biodiversitylibrary. org / page / 7522027", "Needham, J. G. &amp; P. W. Claassen. 1925. A Monograph of the Plecoptera or Stoneflies of America North of Mexico. The Thomas Say Foundation, Volume 2, Entomological Society of America, Lafayette, Indiana. 397 pp. https: // archive. org / details / monographofpleco 00", "Ricker, W. E. 1943. Stoneflies of Southwestern British Columbia. Indiana University Publications, Science Series No. 12. Bloomington, Indiana. 145 pp.", "Illies, J. 1966. Katalog der rezenten Plecoptera. Das Tierreich, Lieferung 82. Walter de Gruyter &amp; Co., Berlin. XXX + 632 pp.", "Surdick, R. F. 1985. Nearctic Genera of Chloroperlinae (Plecoptera: Chloroperlidae). Illinois Biological Monographs, Number 54. University of Illinois Press, Urbana and Chicago, Illinois. 146 pp."]}