Clivina crenulata Balkenohl 2021, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina crenulata

description: <i>Clivina crenulata</i> sp. nov. urn:lsid:zoobank.org:act: 87F44F48-46CC-4FA4-A5FB-4F1FAFAFB87C (Figs 26, 46, 66, 81, 93) ETYMOLOGY. The name refers to the crenulated lateral margin of the pronotum and elytron (Latin <i>crenulatus</i>, -a, -um = weakly crenate). TYPE MATERIAL. Holotype: • &amp;male;, with labels and data: beige, handwritten with black ink “ Borneo ” / white, handwritten with black ink and black printed “Kauf 8. 1936 von Staudinger” / white, black printed “F. van Emden Bequest. B.M. 1960-129” / beige, handwritten with black ink “ <i>Saundersi</i> Andr.”, (BMNH). REMARKS. The following parts are missing: Left front tarsomeres three to five, right intermediate leg, and right hind tarsomeres. The specimen is very different from <i>C. saundersi</i>. The label “ <i>Saundersi</i> Andr.” is obviously a labelling error. DIAGNOSIS. Asmall sized species with crenulated lateral margin of the pronotum and elytron, irregular reticulation on all intervals of the elytron, and with five setigerous punctures on interval three. It is the only species of the group showing these characters. Moreover, it is the only species with five setigerous punctures on the third interval of the elytron. DESCRIPTION. <i>Measurements</i>. Body length 7.62 mm, width 2.33 mm; ratio length/width of pronotum 0.88; ratiolength/widthofelytra 1.78. <i>Colour</i>: Opaque. Piceous. Intermediate and hind leg dark fuscous, antenna and tarsomeres fuscous. <i>Head</i>: Athird narrower than pronotum. Clypeus distinctly excised anteriorly, with lateral angle rounded; wing less protruding as clypeus, flattened, separated from clypeus by distinct notch; supraantennal plate convex; clypeus and supraantennal plate reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by sharp notch. Clypeus slightly transverse, moderately convex, separated from frons by distinct furrow. Frons moderately convex, with flat pore at middle, with converging distinct lateral carina at posterior eye-level. Clypeus and frons with some fine irregular reticulation, separated from wing and supraantennal plate by deep and wide furrow continuing posteriorly as wide supraorbital furrow with two supraorbital setigerous punctures at posterior eye level. Neck constriction developed as distinct step, with band of punctures. Eye laterally projecting. Gena indistinct. Antenna reaching just over posterior angle of pronotum, antennomeres four to ten elongate (double as long as wide). Labrum convex anteriorly, with indistinct reticulation, seven setose. Mentum (Fig. 46) with distinct isodiametrically reticulated surface, reticulation slightly elongated on lobe, with surface undulated, shape of lobe anteriorly like a skewed trapezium with distinct antero-lateral and obtuse medial angle, reflexed margined, anterior margin sinuate, with sharp carina at middle, median tooth wide, distinctly more protruding anteriorly as lobe, obtusely angled anteriorly. <i>Pronotum</i> (Fig. 66): Disk in lateral view moderately convex in anterior half, flattened in posterior half up to basal declivity, convex in frontal view. Outline square, wider than long. Reflexed lateral margin subcrenulated, nearly straight but slightly attenuated in anterior half, widest at posterior third; anterior angle distinct, projecting; posterior angle developed as distinct rounded tooth. Lateral channel with series of small pits between the two setigerous punctures, distinctly widened between anterior and posterior setigerous puncture. Median line moderately wide; anterior transverse line as wide as median line, joining and surpassing median line, narrow before joining basal constriction. Surface with transverse wrinkles, with irregular reticulation, base with few deeper transverse rugae at declivity. <i>Elytron</i>: Disk flattened in anterior half in lateral view, convex on frontal view. Less than twice as long as wide, slightly dilated in apical half, maximum width behind middle. Reflexed lateral margin with some distinct notches posterior humerus, with indistinct notches in anterior half, smooth posteriorly. Scutellar striole distinct. Base of elytron with setigerous tubercle at base of first stria, with a distinct tubercle at base of third and fourth interval, each. Striae less deep, punctuate-striate, one to three free at base, four to seven joining at humerus, one joining with seven at apex, two to six ending free at apex. Intervals moderately convex, fourth forming short transverse carina at humerus, eights with carina at humerus and apex, Third interval with five setigerous punctures, first distinctly removed from base and situated in the middle of the interval, the other four approaching third stria. Surface of intervals opaque, covered with irregular reticulation, becoming indistinctly isodiametric on interval four and slightly more distinct towards interval eight. <i>Hind wings</i>: Fully developed. <i>Lower surface</i>: Proepisternum covered with distinct isodiametric reticulation and laterally with transverse wrinkles and short longitudinal rough structures (Fig. 81). Apex of terminal sternite of abdomen developed as narrow flat transverse sulcus. <i>Legs</i>: Legs covered with longitudinal reticulation. Protibia with three slender spines of moderate length, distinctly sulcate dorsally, movable spur regularly arcuate. <i>Male genitalia</i> (Fig. 93): Median lobe relatively short, gently arcuate at middle, straight apically, wider at base, ventral rim indistinct, apex wide, spatulate, straight in ventral and lateral view. Endophallus with no clearly visible bristles or spines. Ventral paramere moderately wide, lateral apophysis distinct, the other one indistinct, apical third slender, with two long setae at apex, with one short seta preapically. Dorsal paramere slender, regularly arcuate in basal half, straight in apical half, with slender apophyses, at apex with two long setae and one short seta preapically. <i>Female coxostylus and laterotergite</i>: Unknown. VARIATION. Unknown. DISTRIBUTION. Known from Borneo.
description: Published as part of <i>Balkenohl, Michael, 2021, Belgian Journal of Entomology, pp. 1-83 in Belgian Journal of Entomology 115</i> on pages 36-37, DOI: 10.5281/zenodo.5645953