Acatopygia olivacea Winterton, 2007, sp. nov.
Creators: name: Winterton, Shaun L.
Publisher: name: Zenodo
Year: 2007
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Therevidae
Acatopygia
Acatopygia olivacea

description: Acatopygia olivacea sp. nov.
(Figure 1)
Holotype female, AUSTRALIA: QUEENSLAND: Mt. Coot[­t]ha, 20.iii.1948 (MEI 25318) (ANIC). Condition: Good.
Paratypes, AUSTRALIA: QUEENSLAND: male, Pistol Gap, nr. Byfield, 23 10’S, 150 40’E, 10.i.1970, mv lamp, G.A. Holloway (MEI 25319) (AMSA); female, 5 km N Leyburn, 27 58’S, 151 38’E, 3.iv.1993, G.&A. Daniels, C.J. Burwell (MEI 33661) (GDCB / AMSA); male, Brisbane Forest Park: Scrub Road, 10.x.1997, 27 25’05”S, 152 50’13”E, open eucalypt forest, S. Winterton, N. Power, D. White, Malaise trap 3 (ANIC 29 21105) (QDPI). NEW SOUTH WALES: male, Mosman, 20.ii. [19]24, Mackerras [J. Mann Coll.] (MEI 25317) (QMBA); female, Warrumbungle National Park, Brownes Creek nr. Wombelong Creek, M.E. Irwin, 12–16.xii.1995, nr. cliff overhang at water hole, 31 16’24”S, 148 57’38”E (MEIC / CASC).
Diagnosis
Scutum covered with olive­green pubescence; male frons wider than ocellar tubercle, occiput with multiple, poorly defined rows of postocular setae; legs brown; male abdomen without silver velutum; male sternites 4–7 with black, scale­like setae.
Description
Male: Body length: 7.5–8.0 mm.
Head. Frons flat, width equal to ocellar tubercle at narrowest point; frons and face glossy brown; frons overlain with sparse, brown pubescence dorsally, silver pubescence laterally between eye and base of antenna, admixed with short, dark setae; eye facets uniform size throughout eye, transverse furrow absent; ocellar tubercle flattened, black, overlain with grey pubescence admixed with several dark setae; occiput convex, dark, overlain with dense green­grey pubescence dorsally, changing to silver on lower portion and on gena; 2– 3 poorly­defined rows of relatively long, black, postocular setae; gena overlain with silver­white pubescence admixed with white setae; palp and labellum brown, palp covered with dark setae; antennal length shorter than head; antennna orange­brown, overlain with brown­grey pubescence, admixed with scattered, dark setae on scape, pedicel, and base of flagellum; flagellum tapered gradually with two apical segments; style pale.
Thorax. Glossy dark brown; scutum overlain with olive­green pubescence, densely admixed with short setae; scutal macrosetae black; scutellum overlain with matte, black pubescence anteriorly, grey pubescence posteriorly; pleuron with sparse, grey pubescence anteriorly on postpronotum and proepisternum, also on meron, metanepisternum and posterior margin of katepisternum; coxae glossy dark­brown, overlain with grey pubescence, admixed with setae on anterolateral surfaces of mid and fore coxae; setae dark on fore coxa and posterolateral surface of hind coxa; setae pale on mid and hind coxae; legs glossy brown with sparse, pale setae, longer on fore femur; wing hyaline with brown or black banding, two bands across basal medial (bm) and discal (d) cells respectively, third band across the subcostal and radial fields, basal bands connected approximately midway; venation brown; haltere stem brown, knob white; scutal chaetotaxy (pairs): np, 3; sa, 1; pa, 1; dc, 3–4; sc, 1.
Abdomen. Glossy brown with scattered pale setae, setae on posterior segments a mixture of pale and dark setae, setae on terminalia pale; tergite 1 with grey pubescence laterally, brown pubescence medially on tergites 1–5; intersegmental membrane of segments 2–3 white; sternites 4–7 with covering of black, scale­like setae; terminalia dark.
Male genitalia. Epandrium quadrangular, length much shorter along midline than greatest width, posterolateral corners rounded (Fig. 1 A); cercus rounded in shape; tergite 8 broad, slightly emarginate medially (Fig. 1 B); sternite 8 ovate, anterior margin straight, posterior margin rounded with medial patch of elongate setae (Fig. 1 E); gonocoxite (Figs 1 C–D) slightly rounded in profile; gonocoxal process narrow, length equal to inner gonocoxal process, elongate setae on posterior surface; medial atrium present, relatively small; hypandrium large, fused laterally to gonocoxites, large patch of posteriorly projecting setae medially; ventral lobe broad, projecting dorsally, relatively short, velutum on apically posterior surface; gonostylus broad basally, curved medially along length, erect setae on medial surface; inner gonocoxal process spatulate, equal in length to gonostylus; gonocoxal apodeme approximately equal to length of gonocoxite; distiphallus curved ventrally (Figs 1 F–G), ventral apodeme of parameral sheath relatively broad, much shorter than ejaculatory apodeme; dorsal apodeme recurved dorsally, ‘Y’­shaped, fused to ejaculatory apodeme laterally; lateral ejaculatory apodeme small and tapered; ejaculatory apodeme acuminate anteriorly.
Female: Body length: 7.0–9.0 mm.
Similar to male except: Frons slightly wider than ocellar tubercle, covered with greenish­grey pubescence dorsally; shorter setae on head, thorax and abdomen; scutum with short dark setae; abdominal sternites without black scale­like setae.
Female genitalia. Spermathecal sac present, spherical in shape; spermathecal sac duct thickened at common junction with spermathecal ducts; furca quadrangular in shape; spermathecae spherical.
Comments
Acatopygia olivacea sp. nov. is easily recognised from other species of Acatopygia based on the scutal colouration and male abdominal characters such as the presence of black, scale­like setae on the sternites and the lack of silver velutum on the tergites. This species is known from a disparate series of specimens from northern New South Wales and Queensland.
description: Published as part of Winterton, Shaun L., 2007, Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae), pp. 51-62 in Zootaxa 1405 on pages 54-56, DOI: 10.5281/zenodo.175484