Copris Geoffroy 1762
Creators: name: Zidek, Jiri
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Copris

description: Copris Geoffroy
Geoffroy 1762: 87; type sp. Scarabaeus lunaris Linnaeus, 1758.
– Insulicopris Tsukamoto 1974: 1; type sp. Copris brachypterus Nomura, 1964.
– Pilularius Schrank 1798: 394; type sp. Pilularius lunus Schrank, 1798 (= Scarabaeus lunaris Linnaeus, 1758).
– Palaeocopris Pierce 1946: 130; type sp. Palaeocopris labreae Pierce, 1946 = Phanaeus labreae (Pierce) [late Pleistocene of California, see Miller et al. 1981: 627, Edmonds 1994: 91].
Species-group taxa
– achates Péringuey 1901: 346, 365; = cornifrons
– acuticeps Felsche; see Coptodactyla
– acuticollis (Motsch.) of Kabakov 2006: 87 [key]; = acutidens acutidens Motschulsky 1860: 13; China (Fujian, Hubei, Sichuan), Japan. afgoi Cambefort 1992b: 181; Somalia. agnus Sharp 1875: 47; Borneo, Java, Malaysia, Singapore. algol Nguyen-Phung 1989: 202; Kenya, Tanzania.
– amabile Ferreira 1958a: 3; = amabilis Kolbe amabilis Kolbe 1914: 311; DRC, Uganda.
– ammon (Fabricius 1781: 24), as Scarabaeus; = minutus amyntor Klug 1855: 655; Kenya, Mozambique, RSA, Zimbabwe.
– amyntor Harold 1869a: 61; = harrisi
– anaglypticus Say 1823: 204; = fricator fricator anceus (Olivier 1789: 116), as Scarabaeus; RSA (Cape). andrewesi (Waterhouse); see Paracopris
* angolensis Ferreira 1962b: 144; ssp. of cassius angusticornis Arrow 1933: 427; China (Fujian, Hubei, Sichuan, Yunnan), Thailand, Vietnam.
* angusticornis Nguyen-Phung and Cambefort; ssp. of laius angustus Nguyen-phung and Cambefort 1986: 407; Benin, Ivory Coast, Mali, Senegal. anomiopseoides (Boucomont); see Paracopris antares Ferreira 1958a: 5; RSA (Free State). apicepunctatus Balthasar; see Microcopris arcturus Gillet 1907: 179; DRC, Gabon. ares Zelenka 1993: 242; RSA. arimotoi Ochi and Araya 1992: 83; Indonesia (Sumatra). arizonensis Schaeffer 1906: 254, 256; Mexico (Chihuahua), USA (Arizona, New Mexico, Texas). armatus Harold 1869c: 498; Mexico (DF, Guerrero, Hidalgo, Jalisco, México, Michoacán). armeniacus Faldermann 1835: 238; Armenia, Azerbaijan, Georgia, Iran, Turkey. [sp. inquirenda in Ziani 2017] armiger Gillet 1910: 1; Burundi, DRC, Kenya, Mozambique, RSA (Transvaal), Tanzania, Zambia, Zimbabwe. arrowi Felsche 1910: 350; “South China ”, Thailand. aspericollis Gillet 1910: 3; Costa Rica, El Salvador, Guatemala, Mexico (Chiapas). atropolitus Gillet 1910: 7; DRC, Kenya, Tanzania.
– baileyi Blackburn; see Coptodactyla barclayi Ochi, Kon and Bai; see Sinocopris bartolozzii (Ochi, Kon and Pham); see Paracopris basilewskyi Ferreira 1962a: 101; DRC. basipunctatus (Balthasar); see Sinocopris
– belisama (Schrank 1798: 394), as Pilularius; = lunaris bellator Chevrolat in Guérin-Méneville 1844: 81; Indonesia (Java), Malaysia. bengalensis Gillet 1911a: 289; China (Guizhou, Xizang, Yunnan), Myanmar, Thailand. bidens Kolbe 1893: 200; Cameroon.
– bifidus (Poda von Neuhaus 1761: 18), as Scarabaeus; = lunaris
– bihamatus Balthasar 1965: 20, as Paracopris; = fidius
bootes Klug 1855: 655; Botswana, Kenya, Mozambique, RSA (Transvaal), Tanzania.
* borneensis Ochi and Kon; ssp. of gibbulus
– bornemisszai Ferreira 1972: 167, 783; = bootes
boucardi Harold 1869c: 497; El Salvador, Guatemala, Mexico (Guerrero, Oaxaca).
bovinus Gillet 1908 a: 68; Angola, DRC.
brachypterus brachypterus Nomura 1964: 50; Japan (Okinawa: Amami-Ôshima).
brachypterus toguchii Ochi and Kawahara 2012: i; Japan (Tokunoshima).
brittoni Ferreira 1962a: 103; DRC, Tanzania.
brooksi Matthews; see Coptodactyla
caelatus Fabricius 1794: 435; Lesotho, Mozambique, RSA.
* caffer Gillet; ssp. of orion
caliginosus Kohlmann, Cano and Delgado 2003: 2; Guatemala.
– calvus Sharp 1875: 46; = saundersi
cambeforti Nguyen-Phung 1988c: 12; RSA (Transvaal).
cambodiensis Ochi, Kon and Kawahara 2008: 243; Cambodia.
camerunus Felsche 1904: 402; Cameroon.
capensis Waterhouse 1891a: 513; Lesotho, RSA (Cape).
cariniceps (Felsche); see Paracopris
carinicus Gillet 1910: 8; China (Yunnan), India (Assam), Laos, Myanmar, Thailand.
carmelita Fabricius 1801: 39; Benin, Burkina Faso, DRC, Gambia, Guinea, Sierra Leone, Senegal, Uganda.
* cartwrighti Robinson 1941: 131; = ssp. of fricator
cassius Péringuey 1901: 344, 351; Angola, Botswana, Mozambique, RSA (Cape), Zimbabwe.
– castaneus Mulsant 1842: 73, as var. of C. lunaris; = lunaris
* cavolini (Petagna); ssp. of hispanus [Prevailing usage of the subsequent spelling cavolinii has not been officially demonstrated; S. Ziani and T. Branco, pers. comm.]
celebensis Lansberge 1886: 9; Indonesia (Sulawesi).
* centralis Gillet; ssp. of orion
cheni (Ochi, Kon and Bai); see Sinocopris
chinmontanus (Ochi, Kon and Kawahara); see Paracopris
* chuduraziensis Ochi, Kon and Kawahara; ssp. of pseudochus; see Sinocopris
– ciprianii Balthasar 1939a: 3, as Paracopris; = coriarius
– claudius (Harold 1877a: 48), as Catharsius; = repertus
– clavicornis Matthews and Halffter 1959: 191; = moechus
colmanti Gillet 1908 b: 56; Cameroon, DRC, Gabon.
complexus Nguyen-Phung 1989: 206; Burundi, Cameroon, DRC, Kenya, Malawi, Rwanda, Tanzania, Uganda.
compressipennis (Gillet); see Paracopris
confucius Harold 1877a: 48; Cambodia, China (Fujian, Guangdong, Guizhou, Hongkong, Sichuan, Yunnan), Laos, Malaysia, Myanmar, Taiwan, Thailand, Vietnam.
– confusus Boheman 1857: 240; = amyntor
– consobrinus (Lansberge); see Microcopris
* continentalis Ochi and Kon; ssp. of pecuarius; see Sinocopris
– contractus Boheman 1857: 236; = caelatus
– contractus Péringuey 1901: 344, 352; = obesus
convexiusculus Shipp 1897: 450; Somaliland.
coriarius Gillet 1907: 178; DRC, Eritrea, Ethiopia, Guinea, Ivory Coast, Malawi, Nigeria, RSA, Tanzania, Uganda, Zimbabwe [var. of mesacanthus in Ferreira 1961: 156].
– corniculatus Mulsant 1842: 73, as ab. of lunaris; = lunaris
cornifrons Boheman 1860: 110; Botswana, Mozambique, RSA (Cape).
corniger Sahlberg 1823: 6; Ethiopia, Kenya, Lesotho, RSA.
– corniger Harold 1869c: 1014; = bellator
– coronatus (Harold 1877b: 98), as Catharsius; = signatus
corpulentus Gillet 1910: 13; China (Fujian, Sichuan, Yunnan), India (Arunachal Pradesh), Laos, Myanmar, Thailand, Vietnam.
costaricensis costaricensis Gahan 1894: 116; Costa Rica, Mexico (Chiapas), Panama.
costaricensis dolichocerus Matthews 1961: 97; Guatemala, Mexico (Chiapas).
cribratus (Gillet); see Paracopris
cribricollis Gillet 1910: 6; Tanzania.
– curvicornis Boheman 1860: 110; = cornifrons ♀
– davidis (Deyrolle 1878: 95), as Catharsius; = Catharsius molossus (syn. Cheung et al. 2018)
davisi Nguyen-Phung and Cambefort 1986b: 104; Angola.
davisoni (Waterhouse); see Paracopris
decellei Cambefort 1992a: 29; DRC, Guinea, Rwanda, Uganda.
– deletus Mulsant 1842: 73, as var. of lunaris; = lunaris
delicatus Arrow 1931: 117; China (Yunnan), India (Arunachal Pradesh, Assam, Sikkim).
denticulatus Nguyen-Phung 1988b: 216; Mozambique, RSA, Zimbabwe.
depressa Paulian; see Coptodactyla
* dicyrtus Matthews and Halffter; ssp. of remotus
diversus Waterhouse 1891a: 518; Kenya, Somalia, Sudan, Tanzania, Madagascar; introduced into Australia (Queensland, NT, WA).
* dolichocerus Matthews; ssp. of costaricensis
doriae (Harold); see Microcopris
draco Arrow 1906: 127; Angola.
dracunculus Ferreira 1959: 13; DRC.
ducalis Blackburn; see Coptodactyla
dudleyi Cambefort 1992a: 30; Malawi.
eburneus Cambefort 1992a: 28; Ivory Coast.
elpheneroides Felsche 1910: 345; Tanzania.
elphenor Klug 1855: 655; Angola, Burkina Faso, Botswana, Cameroon, DRC, Ethiopia, Kenya, Mozambique, Namibia, RSA, Tanzania, Zimbabwe; introduced into Australia (Queensland).
– emarginatus (Olivier 1789: 115), as Scarabaeus; = lunaris
erratus erratus Lansberge 1886: 13; Sulawesi.
erratus masayukii Ochi and Kon 2004a: 28; Indonesia (Sulawesi: Peleng I.).
evanidus Klug 1855: 655; Botswana, Ethiopia, Kenya, Mozambique, RSA, Tanzania, Uganda, Zimbabwe.
– excavatus Klug 1855: 655; = bootes
excisus (Waterhouse); see Paracopris
fallaciosus Gillet 1907: 180; Botswana, Ethiopia, Kenya, Mozambique, Namibia, RSA, Somalia, Uganda, Zimbabwe; introduced into Australia (Queensland).
fallax Felsche 1910: 349; Namibia.
felschei Reitter 1892: 216 [= 1893: 95]; Armenia, Azerbaijan, Georgia, Iran, e. Turkey.
– ferreirae Frey 1974: 273; = megaceratoides
fidius (Olivier 1789: 184), as Scarabaeus; DRC, Mozambique, Namibia, RSA.
* formosanus Gillet; ssp. of punctatus
– frankenbergeri Balthasar 1934: 149; = acutidens
frankenbergerianus Balthasar 1958: 477; China (Sichuan).
freyi Ferreira 1961a: 138; Gabon.
fricator fricator (Fabricius 1787: 15), as Scarabaeus; India, Sri Lanka; introduced into much of e.+c. USA and se. Canada (Ontario).
fricator cartwrighti Robinson 1941: 131, as sp.; USA (South Carolina).
– fricator Harold 1877a: 46; = erratus
fujiokai fujiokai Ochi and Kon; see Microcopris
fujiokai poringensis Ochi and Kon; see Microcopris
fukiensis Balthasar 1953: 225; China (Fujian, Zhejiang), Taiwan.
furciceps (Felsche); see Paracopris
– furcillatus Felsche 1910: 345; = costaricensis
* gabonicus Cambefort; ssp. of phungae
garambae Cambefort 1992a: 27; DRC
gazellarum Gillet 1918: 148; Burkina Faso, DRC, Sudan, Uganda.
* genopunctatus Nguyen-Phung; ssp. of vankhaii
gibbulus gibbulus Lansberge 1886: 17; Malaysia, Borneo, Java, Sumatra.
gibbulus borneensis Ochi and Kon 2005: 71; Malaysia (Sabah).
gilleti Kolbe 1907: 25; Tanzania.
– gistelianus Gistel 1857: 587; = lunaris
–glabriceps Felsche 1910: 346; = acutidens
glabricollis (Hope); see Coptodactyla
gladiator Arrow 1933: 425; Myanmar.
– globulipennis Waterhouse 1891a: 512; = sexdentatus Thunberg
gopheri Hubbard 1894: 310; USA (Florida).
gracilis Waterhouse 1891a: 518; Namibia.
* grandis Nguyen-Phung and Cambefort; ssp. of inhalatus
graueri Kolbe 1914: 312; Tanzania.
grossa (Thomson); see Pseudopedaria
halffteri Matthews 1959: 133; Mexico (Guerrero, Morelos, Puebla, San Luis Potosí).
– haroldi Lansberge 1886: 15; = spinator
harrisi Waterhouse 1891a: 515; Benin, CAR, DRC, Eritrea, Ethiopia, Kenya, Sierra Leone, Somalia, Sudan, Tanzania, Uganda, Zimbabwe.
hartli Frey 1975: 294; Tanzania.
hatayamai Ochi and Araya 1992: 79; Malaysia.
hidakai Ochi and Kon; see Microcopris
– hintoni Saylor 1933: 238; = klugi
– hispanicus (Poiret 1789: 297), as Scarabaeus; = hispanus hispanus
hispanus hispanus (Linnaeus 1764: 12), as Scarabaeus; western ssp.: s. France incl. Corsica, Sardinia (Italy), Iberian peninsula, Morocco, Tunisia; introduced into Australia (WA).
hispanus cavolini (Petagna 1792: 140), as sp. of Scarabaeus; eastern ssp.: continental Italy + Sicily, Balkans, Turkey, Israel, n. Iran, c. Asia. [For C. hispanus and C. cavolini as separate species see Falahee and Angus 2010.]
– hispanus persianus Mikšić 1958: 1; = hispanus cavolini
hosakai Ochi and Kon; see Microcopris
howdeni Matthews and Halffter 1959: 200; USA (Florida, Georgia?, South Carolina).
humilis Gillet 1908 a: 71; Tanzania.
hybridus Gillet 1910: 11; Tanzania.
igualensis Warner 1990: 236; Mexico (Guerrero).
illotus Balthasar 1942b: 189; China (Fujian).
imitans (Felsche); see Paracopris
– imitatrix Balthasar; see Coptodactyla
inaequabilis Zhang 1997: 758; China (Hubei).
incertus Say 1835: 175; Colombia, Ecuador, Mexico (Hidalgo, Puebla, San Luis Potosí, Tamaulipas, Yucatán), Panama; introduced into Hawaii, New Zealand, Vanuatu.
– indicus Gillet 1910: 5; = fricator fricator
inemarginatus Blatchley 1918: 54; USA (Florida).
inhalatus inhalatus Quedenfeldt 1884: 272; Angola, DRC, Mozambique, Namibia, RSA (Cape), Zimbabwe.
inhalatus grandis Nguyen-Phung and Cambefort 1986b: 102; Zimbabwe.
inhalatus perturbator Péringuey 1901: 352, as sp.; Botswana, Namibia, RSA, Zimbabwe.
inhalatus sanctaeluciae Nguyen-Phung and Cambefort 1986b:102; RSA (Natal).
insidiosus Péringuey 1901: 346, 366; Burundi, DRC, Rwanda, Tanzania, Uganda, Zimbabwe.
integer Reiche 1850: 312; Burkina Faso, Burundi, DRC, Ethiopia, Kenya, Rwanda, Tanzania, Zimbabwe.
– integer Harold 1869a: 61; = fallaciosus
interioris Kolbe 1897: 143; Burundi, Cameroon, DRC, Gambia, Guinea, Nigeria, Senegal, Uganda, Tanzania.
iris Sharp 1875: 45; Laos, Malaysia, Myanmar, Thailand, Vietnam.
* isthmiensis Matthews; ssp. of lecontei
jacchoides Nguyen-Phung and Cambefort 1987: 70; Lesotho, RSA.
jacchus (Fabricius 1775: 20), as Scarabaeus; Lesotho, Namibia, RSA, Zimbabwe.
jahi Nguyen-Phung and Cambefort 1986a: 413; Cameroon.
javanensis (Bui, Ziegler and Bonkowski); see Paracopris
– jenisonianus Gistel 1857: 586; = lunaris
jucundus Gillet 1933: 326; Cameroon, Guinea.
kachinensis Ochi, Kon and Kawahara 2008: 245; Myanmar.
kasagii (Ochi and Kon); see Paracopris
katangae Cambefort and Nguyen-Phung 1996: 286; DRC.
keralensis Gill 1986: 632; India (Kerala).
kershawi Blackburn; see Thyregis
kiuchii (Masumoto); see Sinocopris
klapperichi Balthasar 1942a: 116; China (Fujian).
klugi klugi Harold 1869c: 498; Mexico (most states), Central America.
klugi sierrensis Matthews 1961: 72; Mexico.
kusakabei (Ochi, Kon and Kawahara); see Sinocopris
laeviceps Harold 1869c: 498; Belize, Colombia, Costa Rica, Guatemala, Honduras, Mexico (Chiapas, DF, Oaxaca, Veracruz).
– laevicollis Mulsant 1842: 69, as var. of hispanus; = hispanus hispanus
laevigatus (Gillet); see Sinocopris
laiiformis Nguyen-Phung and Cambefort 1987: 72; Zimbabwe.
laioides Boucomont 1932: 46; DRC, Namibia, Zambia, Zimbabwe.
laius laius Harold 1868a: 81; Burkina Faso, DRC, Gambia, Guinea Bissau, Senegal, Sudan.
laius angusticornis Nguyen-phung and Cambefort 1987: 73; Cameroon, Chad, Sudan, Uganda.
lannathai Hanboonsong, Masumoto and Ochi 2003: 108; Thailand.
– lar (Fabricius 1787: 13), as Scarabaeus; = minutus
– laticornis Boheman 1857: 226; = elphenor
– laticornis Wallengren 1881: 20; = jacchus
– laticornis Péringuey 1901: 343, 347; = corniger, fallaciosus
lecontei lecontei Matthews 1961: 98; USA (Arizona, New Mexico), Mexico (Chihuahua, Colima, Guerrero, Jalisco, Nayarit, Oaxaca, Sinaloa, Sonora).
lecontei isthmiensis Matthews 1961: 102; Mexico.
lesnei Paulian; see Coptodactyla
– lineatus Wiedemann 1823: 9; = sexdentatus
lugubris Boheman 1858: 42; Belize, Costa Rica, El Salvador, Guatemala, Honduras, Mexico (most states), Nicaragua, Panama.
– lunarioides Waterhouse 1891a: 513; = corniger
lunaris (Linnaeus 1758: 346), as Scarabaeus; UK, s. Sweden, c.+s.+e. Europe, w. China, w. Siberia, Turkey, Syria, Iran, c. Asia, Kazakhstan; introduced to Australia.
lunaticeps (Gillet); see Pseudocopris
– lunus (Schrank 1798: 393), as Pilularius; = lunaris
macacus Lansberge 1886: 14; Indonesia (Sulawesi).
macclevei Warner 1990: 232; Mexico (Sonora), USA (Arizona).
macer Péringuey 1901: 343, 346; Angola, Mozambique, Namibia, RSA (Transvaal), Swaziland, Tanzania, Zimbabwe.
maesi Ratcliffe 1998: 93; Nicaragua.
magicus Harold 1881: 88; China (Guizhou, Hubei, Hunan, Sichuan, Xinjiang, Yunnan), India (Sikkim), Laos, Myanmar, Thailand, Vietnam.
manni (Ochi, Kon and Bai); see Sinocopris marcoberrai Ochi, Kon and Pham 2019: 435; c. Vietnam (Quang Nam).
– marcus (Gillet); see Microcopris
– marianus (Petiver 1764: 27), as Scarabaeus; = hispanus hispanus martinae Nguyen-Phung 1988a: 257; Tanzania. martinezi Matthews and Halffter 1968: 154; Mexico (Durango).
* masayukii Ochi and Kon; ssp. of erratus
matthewsi Reid; see Coptodactyla
matthewsi matthewsi Delgado and Kohlmann 2001: 344; Guatemala, Mexico (Chiapas).
matthewsi pacificus Delgado and Kohlmann 2001: 348; El Salvador, Guatemala, Mexico.
medogensis Zhang 1988: 228; China (Xizang).
megaceratoides Waterhouse 1891a: 511; DRC, Gambia, Guinea, Nigeria, Senegal.
megasoma Matthews and Halffter 1959: 194; Mexico (Durango, Michoacán).
merdeka Reid; see Coptodactyla
meridionalis Matthews; see Coptodactyla
mesacanthus mesacanthus Harold 1878: 45; Angola, DRC, Malawi, Mozambique, Namibia, RSA, Zambia, Zimbabwe.
mesacanthus transvaalensis Nguyen-Phung 1988c: 10; RSA.
mexicanus Matthews and Halffter 1959: 194; Mexico (Michoacán).
– minator Harold 1880: 152; = anceus [valid in Gillet 1910] mindorensis (Ochi); see Microcopris
– minutissimus (Kryzhanovskii and Medvedev); see Microcopris minutus (Drury 1770: 78), as Scarabaeus; e.+c. USA. misellus Péringuey 1901: 344, 349; Botswana, DRC, Mozambique, Namibia, RSA, Zambia, Zimbabwe.
– modestus Boheman 1857: 242; = evanidus moechus LeConte 1854: 222; Mexico (Chihuahua, Durango), USA (Texas). moffartsi Gillet 1907: 177; Burundi, Cameroon, DRC, Kenya, Rwanda, Uganda. mongkhoni Hanboonsong, Masumoto and Ochi 2003: 114; Thailand. monstrosa Felsche; see Coptodactyla monteithi Matthews; see Thyregis
* montivagus Gillet 1908 a: 80, as var. of harrisi [valid sp. in Gillet 1910, var. of harrisi in Ferreira 1961: 209]. montreuili (Ochi, Kon and Pham); see Paracopris
– morgani Waterhouse 1891a: 514; = carmelita morphaeus Gillet 1932: 325; Kenya. mourgliai Zelenka 1993: 239; Somalia.
– muelleri Balthasar 1939b: 42; = acutidens muticus (Boheman); see Litocopris
– muticus (Péringuey); see Litocopris neglectus Moxey 1963: 25; India (Tamil Nadu). nepos Gillet 1908 a: 76; Burundi, DRC, Kenya, Rwanda, Tanzania, Uganda, Zimbabwe.
– nepticulus Kolbe 1897: 144; = diversus
– neptis Reiche 1850: 314; = orphanus Guérin-Méneville nevinsoni Waterhouse 1891a: 519; Cambodia, Laos, Malaysia, Thailand, Vietnam. nitida Paulian; see Coptodactyla novaki Zelenka 1992: 167; Malaysia. nubilosus Kohlmann, Cano and Delgado 2003: 4; Guatemala, Honduras. numa Lansberge 1886: 19; India (Assam), Indonesia (Borneo, Sumatra), Malaysia, Myanmar. obenbergeri (Balthasar); see Sinocopris obesus Boheman 1857: 237; Lesotho, Mozambique, RSA, Tanzania, Zimbabwe.
– obliteratus Mulsant 1842: 73, as var. of lunaris; = lunaris ochus (Motschulsky); see Sinocopris
– oedipus (Fabricius 1775: 20), as Scarabaeus; = jacchus
olivieri (Ochi, Kon and Kawahara); see Paracopris
onitoides Gillet; see Coptodactyla
– orientalis Fabricius 1792: 52; = repertus
orion orion Klug 1835: 34; Angola, Burkina Faso, Burundi, Cameroon, DRC, Gambia, Kenya, Mozambique, Namibia, RSA, Rwanda, Senegal, Tanzania.
orion caffer Gillet 1908 a: 70, as var.; RSA.
orion centralis Gillet 1908 a: 70, as var.; DRC.
orion overlaeti (Balthasar 1961: 98), as sp.; DRC, Gabon, Rwanda (?).
orphanus Guérin-Méneville 1847: 308; Angola, Burundi, DRC, Ethiopia, Guinea, Kenya, Rwanda, Tanzania, Uganda.
– orphanus Péringuey 1901: 344, 350; = evanidus
– orphanus Ferreira 1958b: 501; = gracilis
– oryctes (Herbst); see Microcopris
* overlaeti (Balthasar); ssp. of orion
* pacificus Delgado and Kohlmann; ssp. of matthewsi
– paniscus (Fabricius 1775: 123), as Scarabaeus; = hispanus hispanus
papua Lansberge; see Coptodactyla
parapecuarius (Ochi, Kon and Kawahara); see Sinocopris
parasonensis Ochi, Kon and Pham 2019: 437; Laos (Hua Phan).
pauliani Nguyen-Phung and Cambefort 1986a: 413; Cameroon.
– pauliani (Frey); see Thyregis
pecuarius pecuarius (Lewis); see Sinocopris
pecuarius continentalis Ochi and Kon; see Sinocopris
pedarioides (Lansberge); see Paracopris
– persianus Mikšić 1958:1; = hispanus cavolini
– persimilis Felsche 1908: 273; = fallaciosus
* perturbator Péringuey; ssp. of inhalatus
phungae phungae Cambefort 1992b: 181; Gabon, Ivory Coast.
phungae gabonicus Cambefort 1992b: 182; Gabon.
phylax Gillet 1908 b: 59; Cameroon, DRC, Ivory Coast.
– pluridens Gillet 1908 a: 70; = interioris
– plutus (Fabricius 1794: 436), as Scarabaeus; = fidius
poggii Ochi and Kon 2005: 69; Malaysia (Sabah).
– pompilius Waterhouse 1875: 75; = sarpedon
popaensis (Ochi, Kon and Kawahara); see Paracopris
* poringensis Ochi and Kon; ssp. of fujiokai; see Microcopris
potanini Semenov 1891: 310; China (Gansu).
– prociduus Say 1835: 176; = incertus [var. of incertus in Gillet 1910]
– pronus Gerstaecker 1884: 50; = integer
propinquus (Felsche); see Microcopris
– pseudobootes Ferreira 1962a: 104; = typhoeus
pseudochus pseudochus (Ochi and Kon); see Sinocopris
pseudochus chuduraziensis Ochi, Kon and Kawahara; see Sinocopris
– pseudolunaris Ferreira 1969: 3; = hispanus hispanus
pseudosinicus Balthasar 1958: 475; “ China ”.
pueli Mollandin de Boissy 1905: 110; Algeria, Tunisia, Armenia?, Iran?, Turkey?
pumiliaris Wallengren 1881: 20; RSA (Transvaal).
punctatus punctatus Gillet 1910: 4; China (Fujian, Sichuan), Myanmar, Thailand.
punctatus formosanus Gillet 1921: 126; Taiwan.
puncticollis Boheman 1857: 243; Mozambique, Zimbabwe.
punctipennis Boucomont 1914: 333, 335; Indonesia (Sumatra).
punctiventris Waterhouse; see Litocopris
punctulatus punctulatus (Wiedemann); see Paracopris
punctulatus septentrionalis (Ochi, Kon and Pham); see Paracopris
punjabensis (Gillet); see Paracopris
– quadridentatus (DeGeer 1778: 638), as Scarabaeus; = lunaris
quasilaevigatus (Ochi, Kon and Bai); see Sinocopris
ramosiceps (Gillet); see Paracopris
rebouchei Harold 1869c: 497; Mexico (Guerrero, Jalisco, México, Michoacán, Morelos, Nuevo León, Puebla, Veracruz).
reflexus (Fabricius); see Microcopris
– reflexus (Panzer 1794: 7), as Scarabaeus; = minutus
relictus Matthews; see Thyregis
remotus remotus LeConte 1866: 381; Mexico (Coahuila, Nuevo León, Tamaulipas), USA (Oklahoma, Texas).
remotus dicyrtus Matthews and Halffter 1959: 198; Mexico.
renwarti Nguyen-Phung and Cambefort 1986a: 407; Ivory Coast.
repertus Walker 1858: 208; China (Xizang, Yunnan), India (Himachal Pradesh, Sikkim, Uttar Pradesh), Nepal, Sri Lanka, Thailand.
– retusus Mulsant 1842: 68, as var. of C. paniscus; = hispanus hispanus
ribbei Lansberge 1886 a: 12; Indonesia (Sulawesi).
riekoae Ochi and Kon 2003: 51; Laos.
ritsemae Harold 1875: 137; Lesotho, RSA.
ruandanus (Balthasar); see Litocopris
rugosus Gillet 1908 a: 77: Uganda.
ruricola Balthasar 1933: 266; China (Sichuan).
sabinus (Gillet); see Sinocopris
sacontala (Kollar and Redtenbacher) [in Redtenbacher]; see Sinocopris
sallei Harold 1869c: 496; Mexico (Veracruz to Chiapas).
* sanctaeluciae Nguyen-Phung and Cambefort; ssp. of inhalatus
sarpedon Harold 1869b: 104 [replacement name for sexdentatus Kollar and Redtenbacher]; India (Himachal Pradesh, Uttar Pradesh), Nepal, Thailand, Vietnam–Laos (Annam).
saundersi Harold 1868b: 99; Sulawesi.
schoolmeestersi Ochi and Kon 2010: 127; Nepal.
– schoutedeni Boucomont 1928: 419; = wiesei
scorpio Zelenka 1993: 241; Tanzania.
senegalensis Gillet 1910: 15; Senegal.
* septentrionalis (Ochi, Kon and Pham); ssp. of punctulatus; see Paracopris
serius Nguyen-Phung 1987: 271; Tanzania.
– servius Harold 1877a: 46; = agnus
sexdentatus Thunberg 1818: 404; RSA (Cape), Zimbabwe.
– sexdentatus Kollar and Redtenbacher 1844: 520 [in Redtenbacher]; = sarpedon
siamensis Gillet 1910: 12; Thailand.
siangensis Biswas 1979: 339; India (Arunachal Pradesh).
* sierrensis Matthews; ssp. of klugi
signatus (Walker); see Paracopris
– silenus (Fabricius 1775: 21), as Scarabaeus; = minutus
– similis Balthasar 1939a: 4, as Paracopris; = mesacanthus
– simillimus Felsche 1910: 346 [erroneously as from “ Sierra Leone ”]; = minutus
simonettai Cambefort 1992b: 179; Somalia.
simplex (Harold); see Litocopris
simulator Nguyen-Phung and Cambefort 1986a: 409; Chad.
singularis Nguyen-Phung and Cambefort 1986a: 406; Ethiopia.
sinicus Hope 1842a: 60; China (Fujian, Hubei, Jiangxi, Sichuan, Shanghai, Yunnan), Cambodia, Indonesia (Java), Malaysia, Thailand, Vietnam–Laos (Annam).
sinon (Olivier 1789: 123), as Scarabaeus; Gambia, Mauritania, Senegal.
– sinuatus Mulsant 1842: 68, as var. of C. paniscus; = hispanus cavolini sodalis Walker 1858: 208; Sri Lanka. sorex Balthasar 1942a: 117; China (Fujian). sphaeropterus Harold 1877b: 98; RSA (Cape), Zimbabwe. spinator Harold 1881: 89; India (Nicobar Is.), Indonesia (Borneo, Sumatra), Malaysia. stereocera Gillet; see Coptodactyla storeyi Reid; see Coptodactyla subaenea Harold; see Coptodactyla subdolus Balthasar 1958: 476; China (Fujian). subpunctatus Gillet 1910: 3; Costa Rica, Panama. subsidens Péringuey 1901: 344, 355; Botswana, Namibia.
– sulcicollis Lansberge 1886: 10; = sinicus [valid in Gillet 1910] sumatrensis Gillet 1921: 123; Sumatra. surdus (Arrow); see Paracopris szechouanicus Balthasar 1958: 479; China (Fujian, Guizhou, Hunan, Sichuan, Zhejiang).
– taprobanus Gillet 1910: 9; = sodalis tarsatus (Paulian); see Thyregis tetraodon Gillet 1910: 11; Philippines.
– tiberius Schaufuss 1885: 188; = saundersi
* toguchii Ochi and Kawahara; ssp. of brachypterus torresica Matthews; see Coptodactyla
* transvaalensis Nguyen-Phung; ssp. of mesacanthus tridens Felsche 1901: 150; Cameroon, DRC.
– tridens Mulsant 1842: 68, as var. of paniscus; = hispanus cavolini tridentatus Solís and Kohlmann 2003: 2; Costa Rica. tripartitus Waterhouse 1875: 74; China (Fujian, Sichuan, Yunnan), Taiwan, Japan, Korean peninsula.
– troglodytarum Roth 1851: 124; = integer truncatus Felsche 1901: 150; Ivory Coast. tsukamotoi (Ochi, Kon and Kawahara); see Sinocopris tuberculata Gillet; see Coptodactyla tubericollis (Felsche); see Pseudocopris
– tullius (Olivier 1789: 118), as Scarabaeus; = fricator fricator
– tullius Harold 1877a: 45; = spinator typhoeus Gerstaecker 1884: 50; Kenya. uenoi (Hanboonsong, Masumoto and Ochi); see Sinocopris umbilicatus Abeille de Perrin 1901: 68; Albania, Bulgaria, Croatia, France (incl. Corsica), Greece, Hungary, Italy, Romania, Slovakia, Turkey. urus Boheman 1857: 229; Mozambique, RSA, Swaziland. usambaricus Gillet 1908 a: 80; Kenya, Tanzania. vankhaii vankhaii Nguyen-Phung 1988c: 12; Ethiopia, Somalia. vankhaii genopunctatus Nguyen-Phung 1988c: 15; Kenya, Tanzania. victorini Boheman 1857: 234; RSA (Cape, Natal).
– victorini Péringuey 1901: 344, 354; = ritsemae vietnamicus Kabakov 1994: 300; Vietnam. vilhenai Ferreira 1962b: 154; Angola. villiersi Walter; see Pseudopedaria vitalisi (Gillet); see Microcopris vrydaghi Ferreira 1962a: 105; DRC, RSA (Natal). warneri McCleve and Kohlmann 2005: 18; Mexico (Chihuahua, Sonora). wiesei Kolbe 1914: 310; Burundi, DRC, RSA (Cape), Rwanda, Tanzania, Uganda.
– yamamotoi (Tesař); see Sinocopris yangi (Ochi, Kon and Bai); see Sinocopris yokoae Ochi, Kusui and Pham; see Microcopris youngai Balthasar 1967: 49, 50; PRC (Brazzaville).
zhangi (Ochi, Kon and Bai); see Sinocopris
description: Published as part of Zidek, Jiri, 2020, Catalogue of species-group names assigned to Copris Geoffroy, Coptodactyla Burmeister, Litocopris Waterhouse, Microcopris Balthasar, Paracopris Balthasar, Pseudocopris Ferreira, Pseudopedaria Felsche, Sinocopris Ochi, Kon and Bai, and Thyregis Blackburn (Coleoptera: Scarabaeidae: Scarabaeinae: Coprini), pp. 1-24 in Insecta Mundi 756 on pages 3-12, DOI: 10.5281/zenodo.3746274