Orphnebius undetermined MOTSCHULSKY 1858
Creators: name: Assing, Volker
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Orphnebius
Orphnebius undetermined

description: 3.1.2. Key to the named Orphnebius species of Laos
Since as many as 20 Orphnebius species are described from Laos, a country from where the genus had not been reported before, a key to species is provided.
1. Abdominal tergites IX and X strongly modified, with conspicuously dense and long pubescence; hemi-tergites IX strongly elongated in female, ventrally connected by thin sclerotized clasp-shaped structure, antero-ventrally with thin sclerotized projections in male. Posterior margin of tergite VIII mostly smooth, rarely coarsely serrate. Median lobe of aedeagus large in relation to body and with bulbous capsule (exception: O. reductus); paramere of more or less distinctly derived shape. Body in nearly all species distinctly bicoloured, with the forebody dark-brown to black (in one species only head blackish-brown and the remainder of the body reddish) and the abdomen reddish (occasionally with segment VIII infuscate). Antennae often more or less distinctly asymmetric. Sternites and paratergites III–V without distinct postero-lateral processes. Orphnebius hauseri group. ....................................... 2
– Abdominal tergites IX and X not distinctly modified, short and without conspicuously dense pubescence, not distinctly elongated in female. Posterior margin of tergite VIII finely serrate or with two to three more or less pronounced tooth-like projections (in one species with additional short teeth). Median lobe of aedeagus small in relation to body, usually more or less dorso-ventrally depressed, with blade-shaped ventral process, and without sclerotized internal structures; paramere not distinctly modified. Body in most species of different coloration. Antennae symmetric. Sternites and paratergites III–V often with conspicuous postero-lateral processes. Deroleptus. .................................................................................................................................................................. 15
2. Head blackish-brown, distinctly contrasting with the reddish pronotum, elytra, and abdomen (Fig. 45). Antennae uniformly reddish (Fig. 12). Eyes large, occupying all of lateral margins of head (Fig. 45). Median lobe of aedeagus with very acute ventral process (Figs 279–280); paramere as in Fig. 281. Orphnebius retunsus subgroup. ............... ................................................................................................................................................................ retunsus spec. nov.
– Pronotum (nearly) as dark as head. Antennae with at least antennomeres VI–X dark-brown to blackish. Eyes usually smaller. Aedeagus of different morphology. .................................................................................................................... 3
3. Head conspicuously transversely wedge-shaped; eyes strongly convex or somewhat wedge-shaped, situated at posterior margin of head (Figs 28, 31). Antennomere I strongly elongate, longer than the combined length of antennomeres II–IV (Figs 6, 8). Orphnebius dilatatus subgroup. ................................................................................. 4
– Head not transversely wedge-shaped; eyes less strongly convex, not wedge-shaped. Antennomere I much shorter than the combined length of antennomeres II–IV. ......................................................................................................... 5
4. Eyes somewhat wedge-shaped (Fig. 31). Labrum with median tubercle and truncate anterior margin. Antennomeres V–X more transverse and more distinctly asymmetric (Fig. 6). Tibiae reddish. Median lobe of aedeagus and paramere as in Figs 233–235. .............................................................................................................. extensus spec. nov.
– Eyes strongly convex, but not wedge-shaped (Fig. 28). Labrum without tubercle, anterior margin convexly produced in the middle. Antennomeres V–X weakly transverse and not distinctly asymmetric (Fig. 15). Tibiae brown. Median lobe of aedeagus and paramere as in Figs 229–231. Spermatheca as in Fig. 232. ........................ dilatatus spec. nov.
5. Head of transversely quadrangular shape. Antennomeres V–X distinctly transverse and distinctly asymmetric. Clypeus largely membranous. Tergite VIII laterally with a cluster of thin setae on either side, posterior margin smooth or distinctly serrate. Pro- or mesotibiae sometimes modified (distinctly dilated). Median lobe of aedeagus distinctly sclerotized, dark, with short and mostly bifid ventral process. Paramere with slender condylite. Spermatheca with modified proximal portion. Orphnebius serratus subgroup. ................................................................ 6
– Head of more or less orbicular shape. Clypeus with larger sclerotized portion. Tergite VIII without lateral clusters of thin setae, posterior margin never serrate. Pro- and mesotibiae unmodified. Median lobe of aedeagus less strongly sclerotized, ventral process never bifid. Spermatheca of different shape. ................................................................... 9
6. Posterior margin of tergite VIII smooth (Fig. 260). Median lobe of aedeagus and paramere as in Figs 253–256, 258–259. Spermatheca as in Fig. 257. ................................................................................................. bicuspis spec. nov.
– Posterior margin of tergite VIII serrate (Figs 240, 249, 252). ........................................................................................ 7
7. Protibia dilated (Fig. 39); mesotibia unmodified. Median lobe of aedeagus with bifid ventral process (Figs 236–238). Paramere as in Fig. 239. Spermatheca as in Figs 241–243. .............................................................. serratus spec. nov.
– Protibiae unmodified. Mesotibia smoothly dilated or strongly dilated.
8. Mesotibia smoothly dilated. Ventral process of aedagus not bifid (Figs 244–246). Paramere as in Fig. 247. ........... .................................................................................................................................................................... integer spec. nov.
– Mesotibia conspicuously modified, strongly dilated and posterior surface with pronounced longitudinal excavation (Fig. 38). Spermatheca with very long and thin proximal portion (Fig. 251). .......................... latitibialis spec. nov.
9. Posterior margins of tergite and sternite VIII truncate in the middle (Fig. 272). Eyes very large, occupying nearly all of lateral margins of head (Fig. 49). Antenna with pronounced sexual dimorphism, longer in male (1.3–1.4 mm) than in female (1.1–1.2 mm), male antenna with longer antennomere XI and with less transverse antennomeres VI–X. Median lobe of aedeagus with relatively small capsule and with ventral process of distinctive shape (Figs 269–270). Condylite of paramere strongly reduced (Fig. 274). Spermatheca as in Fig. 273. Orphnebius reductus subgroup. ........................................................................................................................ reductus spec. nov.
– Posterior margins of tergite and sternite VIII convex. Eyes in most species smaller. Antenna without pronounced sexual dimorphism. Median lobe of aedeagus with large capsule and a ventral process of completely different shape. Spermatheca of different shape. ...................................................................................................................................... 10
10. Antennomeres V–X conspicuously coniform, brown to dark-brown (Fig. 11). Pronotum strongly transverse, approximately 1.45 times as broad as long, and 1.3 times as broad as head (Fig. 46). Median lobe of aedeagus, paramere, and spermatheca as in Figs 263–265, 268. Orphnebius conicornis subgroup. ............ fusicollis spec. nov.
– Antennomeres V–X not coniform and more or less distinctly flattened, dark-brown to black. Pronotum less transverse and smaller in relation to head (exception: O. grandicollis). Male and female primary sexual characters different. O. hauseri subgroup. ........................................................................................................................................ 11
11. Abdominal segment VIII infuscate (Fig. 35). Antennomere XI conspicuously elongate (Fig. 16). Sculpture of tergite VII composed primarily of dense longitudinal striae. ..................................................................................... 12
– Abdomen uniformly reddish. Antennomere XI less strongly elongate. Sculpture of tergite VII composed primarily of non-setiferous punctation (exception: O. grandicollis). ........................................................................................... 13
12. Antennomeres I–III reddish. Tibiae dark-reddish. Ventral process of aedeagus longer, more slender in ventral view (Figs 216–217). Paramere as in Figs 218–219. ............................................................................ fuscapicalis spec. nov.
– Antennomeres I–III blackish-brown. Tibiae dark-brown. Ventral process of aedeagus shorter, broader in ventral view (Figs 220–221). Paramere as in Fig. 222. ............................................................................. nigrapicalis spec. nov.
13. Pronotum large in relation to head, 1.4 times as broad as long and approximately 1.25 times as broad as head (Fig. 22). Antenna 1.4 mm long (Fig. 17). Sculpture of tergite VII predominantly composed of dense longitudinal striae. Median lobe of aedeagus and paramere as in Figs 212–215. ......................................... grandicollis spec. nov.
– Pronotum distinctly smaller in relation to head, 1.3 times as broad as long and 1.2 times as broad as head at most. Antenna much shorter, <1.2 mm long. Sculpture of tergite VII predominantly composed of non-setiferous punctation. Male primary sexual characters different. ............................................................................................................ 14
14. Eyes much larger, occupying all of lateral margins of head, approximately three times as long as distance from posterior margin of eye to posterior constriction of head (Fig. 24). Median lobe of aedeagus and paramere as in Figs 202–204. Spermatheca as in Fig. 206. .......................................................................................... cernens spec. nov.
– Eyes distinctly smaller, approximately as long as distance from posterior margin of eye to posterior constriction of head (Fig. 25). Median lobe of aedeagus and paramere as in Figs 208–210. ................................. lunatus spec. nov.
15. Sternites III–V without, or with inconspicuous, postero-lateral processes. .............................................................. 16
– At least sternite III, IV, or V with long postero-lateral processes. .............................................................................. 17
16. Head and pronotum weakly transverse; elytra without tubercle near suture (Fig. 85). Sternites III–V unmodified (Fig. 91). Antenna very slender, 1.9–2.2 mm long; antennomeres V–VII oblong, IX–X weakly transverse (Fig. 58). Metafemur distinctly bicoloured, with the basal half yellowish and the apical half blackish. Tergite VIII oblong and with two large tooth-like projections posteriorly (Fig. 320). Median lobe of aedeagus, paramere, and spermatheca as in Figs 317–318, 324. ....................................................................................................................... spoliatus spec. nov.
– Head and pronotum more transverse; elytra with a small tubercle near suture (Fig. 102). Sternite IV and paratergite IV with a short postero-lateral process on either side (Fig. 117). Antenna much shorter, 1.2 mm long, distinctly incrassate apically, and with distinctly transverse antennomeres V–X (Fig. 64). Metafemur blackishbrown. Tergite VIII transverse, posterior margin distinctly concave and finely serrate (Fig. 351). Spermatheca as in Fig. 353. ............................................................................................................................................ carinatus spec. nov.
17. Head truncate or even weakly concave posteriorly; posterior angles marked (Figs 89–90). Eyes enormous, nearly reaching posterior margin of head and more than twice as long as distance from posterior margin of eye to posterior constriction of head in dorsal view (Fig. 89–90). ......................................................................................................... 18
– Head convex posteriorly, lateral and posterior margin practically forming a semi-circle, posterior angles obsolete. Eyes much smaller, approximately as long as, or slightly longer than distance from posterior margin of eye to posterior constriction of head in dorsal view (Figs 86–87). ........................................................................................ 19
18. Sternite V with long lateral process; sternites III–IV weakly modified (Fig. 97). Antenna shorter and finer, 1.2 mm long (Fig. 61). Posterior margin of tergite VIII with a long lateral tooth and two shorter sublateral teeth on either side (Fig. 343). Small species, length of forebody 1.6 mm. Median lobe of aedeagus and paramere as in Figs 340–342. ....................................................................................................................................................................... vates spec. nov.
– Sternite V unmodified; paratergite IV and sternite IV each with a pronounced postero-lateral process on either side, these processes contiguous (Fig. 101). Antenna longer and more massive, approximately 1.8 mm long (Fig. 63). Posterior margin of tergite VIII concave and serrate (Fig. 348). Larger species, length of forebody 2.5 mm. Spermatheca as in Fig. 350. ..................................................................................................................... baccillatus spec. nov.
19. Sternites III and IV each with a long postero-lateral process on either side (Figs 92–94). Elytral punctation very fine and sparse (Fig. 86). Antenna shorter, approximately 1.5 mm long; antennomeres VI–VII not flattened and VI–X not oblong (Fig. 59). Metafemur not distinctly bicoloured, dark-brown. Tergite VIII transverse, with a posterior excision, in postero-median portion with a tubercle (Fig. 328). Sternite VIII conspicuously pointed posteriorly (Figs 329–331). Median lobe of aedeagus of distinctive shape, ventral process knife-shaped (Figs 325–326). Spermatheca as in Fig. 333–334. ............................................................................................................. cultellatus spec. nov.
– Only sternite III with a long postero-lateral process on either side (Fig. 95). Elytral punctation conspicuously coarsely granulose and rather dense (Fig. 87). Antenna much longer, approximately 2.0 mm long; antennomere I conspicuously club-shaped, strongly dilated apically; antennomeres III–VII distinctly flattened and VI–IX oblong (Fig. 60). Metafemur distinctly bicoloured, blackish with the basal third pale-yellowish. Tergite VIII oblong, posteriorly with a long median, two even longer lateral, and four smaller teeth (Fig. 338). Sternite VIII convex posteriorly (Fig. 339). Median lobe of aedeagus and paramere as in Figs 335–337. .................................................. septemcuspis spec. nov.
description: Published as part of Assing, Volker, 2016, On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae), pp. 13-111 in Beiträge Zur Entomologie = Contributions to Entomology 66 (1) on pages 19-21, DOI: 10.21248/contrib.entomol.66.1.13-111, http://zenodo.org/record/6421171