Amplirhagada uwinsensis Köhler 2010, n.sp.
Creators: name: Köhler, Frank
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Camaenidae
Amplirhagada
Amplirhagada uwinsensis

description: <i>Amplirhagada uwinsensis</i> n.sp. <b>Type locality</b> (Fig. 1). Western Australia, Kimberley, Bonaparte Archipelago, Hanover Bay, Uwins Island, 15°15'32"S 124°46'08"E; KIS-3–106. Vine thicket and large fig trees on upper slopes, piles of sandstone rocks, under large slabs. (leg. V. Kessner, 09 June 2006). <b>Type material</b>. Holotype WAM S34621 (Pl. 2.7). Paratypes AMS C463715 (15 preserved specimens, as holotype), WAM S36991 (35 preserved specimens, as holotype), AMS C463716 (15 shells, 15°15'25"S 124°48'04"E), WAM S37441 (30 shells, 15°15'25"S 124°48'04"E). <b>Additional, non-type material</b>. WAM S36987 –90, WAM S37440, WAM S37442 –4, AMS C463717 (Uwins Island). <b>Etymology</b>. In reference to Uwins Island, where this species occurs. <b>Shell</b> (Fig. 69A–C, Pl. 2.7–9). Semi-globose with medium high spire; solid. Periphery angulate; upper and basal sectors of whorls rounded. Umbilicus completely concealed by columellar reflection. Background colour light reddish brown; sub-sutural band diffuse to well defined, light brown to chestnut brown, varies greatly in thickness and contrast; mid-whorl band diffuse, brownish to chestnut brown, thin, varies greatly in contrast and thickness, visible on last whorl(s) only; bands may blend into each other across whorl surface; ventral colour yellowish-greenish brown to ochre; outer lip colour blending into chestnut brown; inner lip blending into dark ochre to chestnut brown. Protoconch c. 2.7 mm in diameter, comprising 1.7 whorls, with fine, indistinct axial lirae. Teleoconch with marked growth lines, becoming oblique towards base of shell. Angle of aperture 45°; outer lip rounded, slightly expanded, not reflected, basal node of lip weak, palatal node absent. Parietal wall of inner lip absent. <b>Pallial morphology</b>. Pallial cavity short, extending ½ to <b> 3</b> ⁄ <b>4</b> whorls; mantle pigmentation consists of indistinctive, sparsely distributed brown spots. Kidney extending about half or more than half of pallial cavity. <b>Genital morphology</b> (Figs. 67–68). Penis rather straight, more or less of same length as anterior part of oviduct. Vas deferens coils once before entering penis. Penial retractor muscle very short to stubby. Penial verge very short (barely visible), slender to spatulate, with pointed tip. Penial wall pustules comparatively large, elongated, arranged in rows on median and basal portion of penial chamber, forming densely packed, narrow longitudinal pilasters at apical end of penial chamber. Main stimulatory pilaster welldifferentiated, large, cone-shaped, covered with flattened pustules, comprising anterior to median portion of penial chamber. Two additional pilasters are formed that are covered by flattened pustules or smooth narrow ridges. Vas deferens entering penial sheath in upper third. Vagina moderately wide, tubular to posteriorly inflated; inner vaginal wall with weakly developed longitudinal pilasters. Spermathecal duct wide, internally with smooth longitudinal pilasters. Spermathecal head globular, connected with oviduct by connective tissue, internally smooth, with delicate wall. Free oviduct comprising less than half to about half of anterior part of oviduct, more or less straight. Spermoviduct of same length as anterior part of oviduct or longer. Talon embedded in albumen gland close to anterior end of albumen gland. <b>Radular morphology</b> (Fig. 69E–F). Rectangular. Tooth formula C+14–16+4+22. In average 149±16.0 rows of teeth, 29.5±1.0 rows per mm (n = 2). Central teeth with sharply pointed, triangular mesocones, shorter than base of teeth; ectocones vestigial. Lateral teeth with sharply pointed, triangular mesocones, length equal to base of teeth; ectocones and endocones vestigial. Marginal teeth with elongate mesocones; ectocones smaller and narrower than mesocones; endocones shorter than ectocones. <b>Comparative remarks</b>. One of the few species with large, dome-shaped shells. Differs from <i>A. tricenaria</i> and <i>A. regia</i> by conspicuously dark brown outer lip colour and by huge, cone-shaped main pilaster, presence of additional pilasters and characteristically elongated pustules of inner penial wall.
description: Published as part of <i>Köhler, Frank, 2010, Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae), pp. 217-284 in Records of the Australian Museum 62 (2)</i> on pages 261-263, DOI: 10.3853/j.0067-1975.62.2010.1554, http://zenodo.org/record/5239086