Toechorychus marcapatus Tedesco & Aguiar 2013, sp. nov.
Creators: name: Tedesco, Anazélia M.
name: Aguiar, Alexandre P.
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Toechorychus
Toechorychus marcapatus

description: Toechorychus marcapatus Tedesco, sp. nov.   
(Figs 36, 42, 91, 117, 131, 180)  
Description. Holotype FEMALE. Fore wing length 5.78 mm.  
 Head. Mandible 1.38 as long as basal width, moderately pilose; ventral margin projected as crest; dorsal tooth distinctly longer than ventral tooth. Clypeus 1.70 as wide as high, subtriangular, minutely strigulate, apically smooth; apex 1.45 as long as base, truncate; apical margin sharp, medially slightly convex. Supraclypeal area entirely strigate, with scarce punctures, densely pilose, medially slightly prominent; between antennal foramens with V-shaped carina; radicle foveolate. Antenna with 25 flagellomeres; white band starting at flagellomere 4, reaching flagellomere 12; flagellum slender; subapical flagellomeres slightly flattened. Supra-antennal area with very faint but distinct median longitudinal carina, medially without longitudinal elevation, near antennal sockets striate; dorsal half medially rugulose. Paraocular area smooth. Vertex rugulose around ocelli, without sulcus; gena and vertex behind ocelli smooth, sparsely pilose; gena in lateral view uniformly wide; occipital carina stout, dorsally uniformly arched, ventrally slightly projected as crest, reaching hypostomal carina far from mandible base; hypostomal carina projected as crest; malar space 0.79 as long as basal width of mandible.   
 Mesosoma. Mesosoma in dorsal view 2.08 as long as wide; in lateral view, mesosoma middle width 0.17 mm. Pronotum centrally smooth, latero-ventrally rugose behind collar, margin near mesopleuron ventrally corrugated; pronotal swelling strigulate; collar dorso-laterally rounded, distinctly swollen, anteriorly punctulate; epomia absent. Mesoscutum densely pilose, lobes dorsally punctulate, laterally markedly corrugated or covered with sparse coarse punctures, central lobe without longitudinal sulcus or carina; notauli deeply impressed, posteriorly parallel throughout. Scutoscutellar groove smooth, polished; scutellum 1.04 as long as wide, smooth; scutellar carina incomplete, restricted approximately to anterior 0.6; postscutellum 0.27 as long as wide; hind margin of metanotum with two short lateral carinae extending towards transverse furrow at base of propodeum. Subalar ridge somewhat elongate; mesopleuron dorsally medially markedly strigate, ventrally strigulate; mesopleural groove corrugated; epicnemial carina faint, dorsally straight; dorsal end of epicnemial carina reaching 0.2 of posterior margin of pronotum, behind epicnemial carina without rounded confused-rugulose area; sternaulus medially shallow, interrupted, reaching base of mid coxa, markedly sinuate, posteriorly almost faint, faint from anterior 0.6 to 0.8, not corrugated; scrobe deeply impressed, forming pit. Mesothoracic venter punctate and strigulate; median portion of posterior transverse carina of mesothoracic venter short and straight, regularly shaped, without projections. Transverse furrow at base of propodeum laterally 0.05 as long as anterior portion of propodeum, shallow and narrow, very faintly corrugated, almost smooth, medially some carinae stouter; propodeum 1.21 as long as wide medially; anterior margin of propodeum without teeth; anterior portion of propodeum finely strigulate, medially without longitudinal carinae; lateral longitudinal carina of propodeum indistinct; posterior portion of propodeum punctate and strigate, medially without distinctly stout longitudinal carina; behind posterior transverse carina with two divergent longitudinal carinae; anterior transverse carina of propodeum complete, stout, straight; posterior transverse carina absent, only with lateral slight swellings; propodeal spiracle 1.33 as long as wide; pleural carina of propodeum absent; lower division of metapleuron strigulate, densely pilose, juxtacoxal carina absent.   
 Wings. Fore wing: vein 2+3Rs slightly concave; ramellus absent; vein 1M+Rs uniformly curved; vein 1cu-a posteriorly slightly curved, arising far from vein 1M+Rs base, angle with vein M+Cu distinctly obtuse; vein 2Cu 0.63 as long as vein 2cu-a; vein 4Rs 0.91 as long as vein 4M, straight; cell 1+2Rs 0.81 as high as pterostigma, rectangular; vein 2M distinctly longer than vein 3M; vein 3M distinct, 1.00 as long as vein 2M. Hind wing: vein 1Cu 1.30 as long as vein cu-a; vein 2-1A reaching 0.80 of distance to posterior margin.   
 Legs. Tibia with very sparse short bristles; hind coxa globose.   
 Metasoma. T1 0.53 as long as hind femur, without basolateral tooth; anteriorly smooth, posteriorly coriarious; spiracle at anterior 0.54, slightly prominent; postpetiole dorsally, at level of spiracles, faintly but distinctly concave; T2–8 minutely coriarious; centrally almost glabrous, laterally pilose; T2 1.14 as long as wide at apex; apex of T2 2.06 as wide as base. Dorsal valve of ovipositor with nodus indistinct; apex of lower valve with series of teeth; notch absent.   
 Color. Head and metasoma mostly black and white, mesosoma mostly orange (154,060,018). Head: pale yellow (226,200,132); mandible teeth, supra-antennal area medially, interocellar area, vertex and gena, except complete large orbital band, and occiput, black; scape ventrally yellow, dorsally brown; flagellum ventrally entirely brown or orange, dorsally entirely whitish at f5–10, partially at f4 and f11. Mesosoma: propleuron pale yellow; pronotum black except entire collar and pronotal swelling, pale yellow; mesoscutum black, except longitudinal lanceolate mark at entire length of central lobe; mesopleuron ventrally orange, dorsally pale yellow, except medially, around hypoepimeron, dark brown (062,029,012); remaining of mesosoma orange, except for the following pale yellow marks: axillary carina, scutellum, postscutellum, tegula basally, upper division of metapleuron centrally. Legs: orange (178,064,013), yellowish towards apex; all t5 dark brown.. Metasoma: T1–2 orange, brownish towards apex, with posterior 0.2 white; T3–8 anteriorly dark brown, laterally and at posterior 0.3, white; S1 orange; S2–5 white with lateral brown spot; bright yellow, laterally black; S6–8 white.   
 Variation. Flagellomeres dorsally entirely whitish at f5–11, partially at f4 and f12. T1 medially and t2 basally, black. Scutoscutellar groove faintly corrugated. Postscutellum sometimes orange. Dark brown mark around hypoepimeron ventrally lighter, or faded. Black portions of mesoscutum, pronotum and mesopleuron dark brown in one specimen. T 2 in one specimen basally orange (same tone of body, except blackish over thyridium). Differences in tonality include variations from whitish (246,238,222) to bright yellow (240,214,097), including pale yellow (226,200,132).   
 Comments. Very similar to T. melgassus sp. nov., from which it can be readily differentiated by having pronotum, mesoscutum and mesopleuron marked with black areas (vs. completely orange in T. melgassus sp. nov.; Fig. 37); collar dorso-laterally rounded, distinctly swollen, as in Fig. 47 (vs. carinated, not swollen, as in Fig. 46); pleural carina of propodeum absent (vs. present only posteriorly). Color pattern very similar to T. paramaribus sp. nov., from which it can be distinguished by having supraclypeal area entirely pale yellow (vs. dorsal half of supraclypeal area with median longitudinal stripe; Fig. 74); collar laterally pale yellow (vs. dark brown; Fig. 38); gena and malar space pale yellow (vs. gena ventrally and malar space, black; Fig. 74); mesopleuron with black mark anteriorly around hypoepimeron (vs. orange, except for subalar ridge and epicnecium, pale yellow; Fig. 38); posterior transverse carina of propodeum absent (vs. complete, laterally with crest); pleural carina of propodeum absent (vs. present only posteriorly).   
 Male. Unknown.   
 Distribution. Suriname, Brazil, and Peru. Recorded from five localities from Suriname to Peru (Fig. 216). The type locality is the Floresta Nacional de Caxiuanã, Pará (2°4’54”S 51°51’5”W). These records comprise a range of 26°09’ in latitude.   
 Biology. Unknown.   
 Material examined. 8 females. Holotype ♀, BRAZIL: Pará, Melgaço, Floresta Nacional de Caxiuanã, Estação Científica Ferreira Pena, 22–25.XI.2003, Malaise trap M5, A.P.Aguiar & J.Dias leg. P05144 [field point] (UFES). Right hind leg beyond femur missing; otherwise in good shape. Paratypes: BRAZIL: 1 ♀ from Amazonas, Manaus, INPA, Reserva Florestal Adolpho Ducke, 08–14.X.2005, Malaise trap, pt. 10, A.P.Aguiar et al. leg. 1 ♀ from Pará, Melgaço, Floresta Nacional de Caxiuanã, Trilha Igarapé Tijucaquara, 18–24.XI.2003, Malaise trap M11, A.P.Aguiar & J.Dias leg., P05128 [field point]. 1 ♀ from Espírito Santo, Cariacica, Reserva Biológica de Duas Bocas, 27.V.1997, sweeping, C.O.Azevedo leg. (UFES). Other specimens: SURINAME: 1 ♀ from 45 km S Paramaribo, 29.IX.1963, D.C.Geijskes leg.; 1 ♀, same data, except 29.IX–03.X.1963; 1 ♀, same data, except 26.X–01.XI.1963 (AEIC). PERU: 1 ♀ from Avispa, nr. Marcapata, 01–15.X.1962, 30 m, L.Peña leg. (AEIC).
description: Published as part of Tedesco, Anazélia M. & Aguiar, Alexandre P., 2013,  Phylogeny and Revision of  Toechorychus  Townes (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of thirty-five new species , pp. 1-138 in Zootaxa 3633 (1) on pages 47-48, DOI: 10.11646/zootaxa.3633.1.1, http://zenodo.org/record/5261931