Cormocephalus (Cormocephalus) ungulatus
Creators: name: Schileyko, Arkady A.
name: Cupul-Magaña, Fabio G.
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Chilopoda
Scolopendromorpha
Scolopendridae
Cormocephalus
Cormocephalus ungulatus

description: Cormocephalus (C.) ungulatus (Meinert, 1886)
Figs 33–38
Cupipes ungulatus Meinert, 1886: 187;
Cupipes graecus Meinert, 1887: 123;
Cupipes ungulatus: Pocock, 1893: 460;
Cupipes ungulatus: Kraepelin, 1903: 177;
Cupipes ungulatus: Chamberlin, 1914: 184;
Cupipes ungulatus: Chamberlin, 1921: 17;
Cupipes ungulatus: Chamberlin, 1922: 7;
Cupipes ungulatus: Chamberlin, 1925: 37;
Cormocephalus (C.) ungulatus: Attems, 1930: 101;
Cormocephalus (C.) ungulatus: Bücherl, 1941: 300;
Cormocephalus mundus Chamberlin, 1955: 46;
Cormocephalus (Cupipes) tingonus Chamberlin, 1957: 31;
Cormocephalus mundus: Kraus, 1957: 383;
Cormocephalus tingonus: Bücherl, 1974: 103;
Cormocephalus ungulatus: Schileyko 2018: 72;
Cormocephalus ungulatus: Martínez-Muñoz & Perez-Gelabert 2018: 83, 92.
Terra typica: Haiti (= Hispaniola) Island and Pernambuco (Eastern Brazil).
Studied material. Panama, IBISCA project, Colón Province, San Lorenzo Forest, tree canopies, 1 sad. (Rc 7155), 2004, col. I. Tuf. Brazil, Amazonas, Reserva Florestal A. Ducke, near [about 25 km N of] Manaus, 59 59’W, 2 55’S, primary tropical rainforest forest on Terra firme, 1 ad. (Rс 6483), 06-011. III.1998, col. S.I. Golovatsch.
Description of ad. Rc 6483 [data on sad. Rc 7155 in square brackets where it differs].
Length ca. 30 [15] mm. Coloration in alcohol uniformly yellow [greyish] with head visibly darker [head, body (including sternites) and ultimate legs with small aggregations of dark pigment, Figs 33, 34]; numerous small setae at all body surface.
Head (Fig. 35) with much shortened anteriorly (as long as ½ [1/3] of head), but well-developed [indefinite] paramedian sutures; very short antennae (left 15, right 17 articles [both 17]) hardly extending to the posterior margin of tergite 1 [2] when reflexed.
Forcipular coxosternite (Figs 36, 33) with two much shortened posteriorly (as long as 1/2 [1/3] of coxosternite) and converging anteriorly typical longitudinal sutures; transverse suture absent. Tooth-plate with 4 teeth, of these the most lateral and the most median ones are approximately of the same length, being much shorter than 2 remaining teeth; long and pointed trochanteroprefemoral process with one lateral tubercle.
Tergites 1–20 with complete paramedian sutures; tergite 21 (Fig. 38), with complete median suture [plus a median depression in the posterior half], only this tergite is marginated laterally.
Sternites 2–20 with complete paramedian sutures disposed in paramedian sulci; sternite 21 (Fig. 38) trapeziform, slightly [noticeably] longer than wide, its posterior margin rounded [straight]; sternite 21 with a short, shallow median sulcus / depression in the posterior third.
Legs with 2 short [minute] but clearly recognizable [hardly recognizable] pretarsal accessory spines.
Ultimate LBS (Fig. 37): coxal pore field as long as [very slightly longer than] sternite 21, coxopleuron slightly longer than sternite 21, with rounded posterior margin, lacking either any process or spines [the only minute spine at the very inner corner of the coxopleuron].
Ultimate legs (Figs 37, 38): prefemur, femur and tibia much flattened dorsally. These articles are each with characteristic dorso-medial longitudinal depression apically; tibia and tarsus 1 somewhat swollen / bulbous ventrally. Prefemur with the only spine at the position of the corner spine (Fig. 38) [prefemur (Fig. 34) with 3 unusually large dorso-medial spines—2 of them disposed at a remarkably enlarged corner spine—plus 1 ventro-medial and 1 medial one, both of the latter are disposed at the distal margin of the prefemur, forming a transverse row together with the corner spine; no any ventral spines].
Remarks. Adult spm Rc 6483 has been mentioned by Schileyko (2002) as C. brasiliensis Humbert & Saussure, 1870 but the present re-investigation shows that it is C. ungulatus, because it shares with the latter such important peculiarities of the forcipular coxosternite as very short paramedian sutures plus a total absence of a transverse one. This specimen also has no paired dorso-distal “bifid spines” on the ultimate femur (!), which should be diagnostic for C. brasiliensis (see Chamberlin 1914: 183) and being very unusual (if not unique) within Cormocephalus; Lewis (1989: 1006) wrote that C. brasiliensis “was described on the basis of a defective specimen”.
description: Published as part of Schileyko, Arkady A. & Cupul-Magaña, Fabio G., 2021, Cormocephalus (Cormocephalus) guildingii Newport, 1845 (Chilopoda: Scolopendromorpha): a composite description, new samples from Western Mexico and a new species subgroup of Neotropical Cormocephalus (Cormocephalus), pp. 301-325 in Zootaxa 5071 (3) on pages 314-316, DOI: 10.11646/zootaxa.5071.3.1, http://zenodo.org/record/5723692