Aname warialda Raven 1985
Creators: name: Wilson, Jeremy D.
name: Harvey, Mark S.
name: Simmons, Leigh W.
name: Rix, Michael G.
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Anamidae
Aname
Aname warialda

description: Aname warialda Raven, 1985
Figs 1, 11, 81–82
Aname warialda Raven, 1985: 407, figs 7, 18, 22, 35, 46, 56, 58.
Aname “MYG687” – Rix et al. 2021: figs 3, 6–7.
Diagnosis
Males of A. warialda can be distinguished from all species for which males are known except A. bifaceta sp. nov., A. occivillosa sp. nov., A. scutitheca sp. nov., and A. villosa by a moderate to large body size (carapace length>4.0 mm), the presence of a long embolus (embolus length / bulb length>1.5) that is relatively straight, a short tibial megaspine (megaspine length/ tibia length <0.2), and three or more spines on the prolateral patellae of the pedipalp and leg I (Fig. 81A–Q). Males of A. warialda can be distinguished from those of A. occivillosa and A. villosa by the presence of a longer embolus (embolus length / bulb length>2.5) and a longer asetose depression on the palp tibia (depression length / palp tibia length> 0.6) (Fig. 81J–M; cf. Figs 74, 78). Males of A. warialda can be distinguished from those of A. scutitheca by the presence of a tibia I that stays about the same width from the proximal end to the base of the tibial spur when in lateral view and a less spiny palp tibia (Fig. 81J–K, N–P; cf. Fig. 76). Males of A. warialda can be distinguished from those of A. bifaceta by the presence of a longer asetose depression on palp tibia (depression length / palp tibia length ~0.71), and a knuckle at the base of the tibial spur (Fig. 81J–K, N–P; cf. Fig. 71).
Females of A. warialda can be distinguished from all species for which females are known except A. bifaceta sp. nov. and A. scutitheca sp. nov. by the presence of spermathecae with a single elongate, undulating vesicle (lateral vesicle length/ genitalia width>0.5) (Fig. 82L). Females of A. warialda can be distinguished from those of A. scutitheca by the absence of a large rounded extension of the posterior genital plate (“scute”) over the epigastric furrow (Fig. 82D, L; cf. Fig. 77). Females of A. warialda can be distinguished from those of A. bifaceta by the presence of larger, longer spermathecae (lateral vesicle length / genitalia width ~ 0.8; cf. ~ 0.5 in A. bifaceta) (Fig. 82L; cf. Fig. 72).
Type material
Holotype
AUSTRALIA – New South Wales • ♂; Warialda; 29°32′ S, 150°35′ E; 7 Jan. 1980; R. Hobbs leg.; QMB S1291.
Paratypes
AUSTRALIA – Queensland • 1 ♂; Girraween National Park; 28°50′ S, 151°56′ E; 5 Feb. 1979; H. Todd leg.; QMB S1296. – New South Wales • 1 ♀; Toowoomba; 27°34′ S, 151°57′ E; 10 Feb. 1965; T. Passlow leg.; QMB S1299 • 2 ♂♂; Tenterfield; 29°06′ S, 151°54′ E; Apr. 1983; G. Colvin leg.; QMB S1294 • 1 ♂; Texas Caves [Ashford Caves]; 29°12′ S, 150°59′ E; 4 Apr. 1979; L.R. Jeffrey leg.; QMB S1298 • 1 ♀; Inverell; 29°46′ S, 151°07′ E; May 1979; QMB S1314 • 1 ♂; Bingara; 29°49′ S, 150°25′ E; Jan. 1980; R. Hobbs leg.; QMB S1292.
Other material examined
AUSTRALIA – Queensland • 1 ♂; Carnarvon National Park, Mount Moffatt Section, near Marlong Arch; 24°59′ S, 147°54′ E; 760 m a.s.l.; 12 Dec. 2012 – 16 Jan. 2013; G.B. Monteith and G. Keith leg.; gutter trap, eucalypt forest; QMB S52397 • 2 ♂♂; Carnarvon National Park, Mount Moffatt Section; 25°04′ S, 148°02′ E; 13–15 Dec. 1987; G.B. Monteith, J. Thompson and D. Yeates leg.; QMB S11261 • 1 ♀; Mount Tabor Station, ca 147.2 km NNE of Morven; 25°10′ S, 147°36′ E; 8 Dec. 2020; C. Eddie and E. Amsters leg.; excavated, eucalypt forest; QMB S118236 • 1 ♀; Mount Tabor Station, ca 145.4 km NNE of Morven; 25°11′ S, 147°37′ E; 22 Oct. 2021; E. Amsters and R. Aisthorpe leg.; excavated, eucalypt forest, near watercourse; QMB S118245 • 1 ♀; Mount Tabor Station, ca 145.0 km NNE of Morven; 25°11′ S, 147°36′ E; 21 Oct. 2021; E. Amsters and R. Aisthorpe leg.; excavated, eucalypt forest; QMB S118242 • 1 ♀; 51.5 km NNW of Injune; 25°25′ S, 148°21′ E; 21 Sep. 2021; E. Amsters leg.; excavated, Callitris forest; QMB S118241 • 1 ♀; 51.3 km NNW of Injune; 25°26′ S, 148°21′ E; 21 Sep. 2021; E. Amsters leg.; excavated, Callitris forest; QMB S118239 • 1 ♀; 51.3 km NNW of Injune; 25°26′ S, 148°21′ E; 21 Sep. 2021; E. Amsters leg.; excavated, Callitris forest; QMB S118240 • 1 ♂; Hutton Creek, “Oak Wells”; 25°46′ S, 148°17′ E; 600 m a.s.l.; 15 Dec. 2001 – 7 Mar. 2002; G.B. Monteith and D.J. Cook leg.; pitfall trap, vine scrub; QMB S59308 • 1 ♂; Kingaroy; 26°31′ S, 151°48′ E; 1 Dec. 1995; D. Kamholtz leg.; QMB S29591 • 1 ♂; Roma; 26°34′ S, 148°47′ E; 24 Feb. 1986; R. Kunde leg.; QMB S100534 • 1 ♂; 40 km S of Charleville, “ Wallal ”; 26°44′ S, 146°07′ E; 23 Jan. 1996; P. McRae leg.; QMB S29635 • 1 ♂; Tara; 27°16′ S, 150°27′ E; 20 Nov. 2001; Queensland Department of Primary Industries leg.; QMB S118360 • 1 juv.; Lake Broadwater Conservation Park, N of lake; 27°20′ S, 150°33′ E; 335 m a.s.l.; 7 Dec. 2019; M.G. Rix and M.S. Harvey leg.; excavated, open burrow on ground, open woodland; QMB S111487 • 1 ♀; Lake Broadwater Conservation Park, Wilga Bush Camping Area; 27°20′ S, 151°06′ E; 345 m a.s.l.; 28 Oct. 2022; J.D. Wilson, E. Briggs, M.G. Rix and A.G. Rix leg.; excavated; QMB S118219 • 1 ♀; Lake Broadwater Conservation Park, Wilga Bush Camping Area; 27°20′ S, 151°06′ E; 345 m a.s.l.; 28 Oct. 2022; J.D. Wilson, E. Briggs, M.G. Rix and A.G. Rix leg.; excavated; QMB S118220 • 1 ♂; Lake Broadwater Conservation Park; 27°21′ S, 151°05′ E; 3 Jan.–25 Feb. 1986; Queensland Museum Party leg.; pitfall trap; QMB S3153 • 1 ♀; Moonbah, campsite [Alton National Park]; 27°59′ S, 149°19′ E; 9–11 Jan. 1979; R.J. Raven leg.; QMB S1295 • 1 ♂; Ula Ula National Park; 27°59′ S, 149°28′ E; 11 Nov. 2016; R.C. Santana leg.; QMB S110191 • 1 ♀; Durikai State Forest, Durikai Road; 28°12′ S, 151°37′ E; 531 m a.s.l.; 19 Feb. 2023; M.G. Rix, J.D. Wilson and M.S. Harvey leg.; excavated, open burrow on ground; QMB S118230 • 1 ♂; 18 km N of Inglewood, Robert Wicks Research Centre; 28°18′ S, 151°09′ E; 10 Jan. 1999; G. Rettke leg.; hand collected, on path at night, Callitris forest; QMB S42819 • 1 juv.; Oman Ama, E of Inglewood, Donovans Road; 28°23′ S, 151°20′ E; 342 m a.s.l.; 19 Feb. 2023; M.G. Rix, J.D. Wilson and M.S. Harvey leg.; excavated, open burrow on ground next to road; QMB S118229 • 1 juv.; Warroo, SE of Inglewood, Cement Mills Road; 28°31′ S, 151°23′ E; 409 m a.s.l.; 14 Aug. 2022; E.J. Briggs and M. S. Woolley leg.; excavated, y-shaped burrow, Callitris forest; QMB S118218 • 1 ♀; Warroo, SE of Inglewood, Cement Mills Road; 28°32′ S, 151°21′ E; 374 m a.s.l.; 19 Feb. 2023; M.G. Rix, J.D. Wilson and M.S. Harvey leg.; excavated, open burrow on ground next to road; QMB S118219 • 1 juv.; Warroo, SE of Inglewood, Cement Mills Road; 28°32′ S, 151°21′ E; 374 m a.s.l.; 19 Feb. 2023; M.G. Rix, J.D. Wilson and M.S. Harvey leg.; excavated, open burrow on ground next to road; QMB S118228 • 1 ♂; The Summit; 28°34′ S, 151°57′ E; 16 Jan. 1984; Queensland Department of Primary Industries leg.; QMB S100536 • 1 ♂; Stanthorpe; 28°37′ S, 151°30′ E; 21 Dec. 1994; Mr and Mrs Kuenstner leg.; QMB S26087 • 1 ♂; Stanthorpe; 28°37′ S, 151°30′ E; Apr. 2016; M. Hodgetts leg.; QMB S118359 • 1 ♀; Nundubbermere Falls, 25 km SW of Stanthorpe; 28°47′ S, 151°41′ E; 1–4 Apr. 1988; G.B. Monteith leg.; QMB S4192 • 1 ♂; Tenterfield, Wallangarra; 28°55′ S, 151°56′ E; Apr. 1983; M. Wanstall leg.; QMB S10000. – New South Wales • 1 ♂; Texas; 28°51′ S, 151°09′ E; 4 Feb. 1992; E.J. Greentree leg.; QMB S19135 • 1 ♂; Texas Caves [Ashford Caves]; 29°12′ S, 150°59′ E; 1–2 Feb. 1975; G.V. Czechura leg.; QMB S1297 • 1 ♂; Deepwater; 29°27′ S, 151°49′ E; 10 Dec. 1985; R. Hobbs leg.; QMB S9804 • 1 ♂; Emmaville; 29°30′ S, 150°19′ E; 13 Jun. 1996; F. Ginns leg.; QMB S34590 • 1 ♂; The Brigalows, 25 km N of Inverell; 29°37′ S, 151°06′ E; 1 Jan. 1991; F. Ginns leg.; QMB S34586 • 1 ♂; 15 km W of Armidale; 30°31′ S, 151°31′ E; 7 Dec. 1980; R. Hobbs leg.; QMB S9747 • 1 ♀; Armidale, near cemetary; 30°31′ S, 151°40′ E; AMS KS31709 • 1 ♂; Moore Creek, 8–10 km N of Tamworth; 31°02′ S, 150°58′ E; Jan. 1984; D. McCalee leg.; AMS KS13803.
Description
Male (holotype, QMB S1291)
GENERAL (Fig. 81A–Q). Body length 22.77, in good condition, colour presumably faded due to preservation.
DORSAL PROSOMA (Fig. 81A, E–F). Carapace length 8.81, width 7.71, length/width 1.14, clypeus to fovea length/carapace length 0.71, caput width/carapace width 0.66, carapace red, caput slightly darker than thorax, reflective setae present, heavy on caput, moderate on thorax, fovea procurved, fovea width/ carapace length 0.16 (Fig. 81A, F); chelicerae dark red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.46 (Fig. 81A); eye group rectangular, width/length 1.84, eye tubercle present (Fig. 81E).
ABDOMEN (Fig. 81B, D). Abdomen length 9.63, brown, dorsal pattern absent, with consistent cover of short setae.
VENTRAL PROSOMA (Fig. 81C, G–I). Labium cuspules absent (Fig. 81H); maxillae heel distinct, cuspules present, count =about 150, extending posteriorly onto heel, extending laterally about 40% of maxillae length (Fig. 81C, I); coxae cuspules absent, thorn-like setae on prolateral face present (Fig. 81C, I); sternum length/width 1.09, central sternum with consistent covering of short setae, row of longer setae around posterior edges (Fig. 81G–H); posterior sigilla elongate, central sternum to posterior sigilla length/sternum length 0.25, posterior sigilla length/sternum length 0.18 (Fig. 81G–H); other sigilla small and lateral, medial sigilla semi-elongate, anterior sigilla round (Fig. 81G–H).
LEG I (Fig. 81N–Q). Leg I orange-brown, femur length 6.88, patella length 4.41, tibia length 4.86, metatarsus length 4.63, tarsus length 3.24, total length 24.02, leg I length/carapace length 2.73 (Fig. 81N– O); scopulae on distal metatarsus and tarsus (Fig. 81N–O); spine count Fe D 4, Fe PL 1, Pa PL 3, Ti PL 0, Ti RL 0, Me PL 0, Me RL 0, Ta 0 (Fig. 81N–O); tibia length/width [TIL/TID] 3.21, even width along length, spur present, digitiform, knuckle present, megaspine angled at 3 degrees, length to distal face of spur/tibia length [TIS/TIL] 0.51, spur height/tibia width [TISH/TID] 0.61, megaspine length/tibia length 0.18 (Fig. 81N–P); metatarsus slightly sinuous, proximal excavation present, excavation concave with pronounced heel, heel rounded, excavation length/metatarsus length [MIPEL/MIL] 0.39, metatarsus length/width [MIL/MID] 3.78 (Fig. 81N–O, Q).
PEDIPALP (Fig. 81J–M). Tibia length 3.62, width 1.48, length/width [PTL/PTD] 2.45, asetose depression present, depression length/palp tibia length [PDL/PTL] 0.71, retrolateral face with consistent covering of light setae, ventral face with two elongate bristle-like setae below depression, prolateral face with patch of spines on distal half, disto-medial spine present (Fig. 81J–K); patella prolateral face with 3 spines (Fig. 81J–K); cymbium with scopulae present distally (Fig. 81J–K); copulatory organ total length 2.78, length/palp tibia length 0.77 (Fig. 81L–M); bulb length/width 0.94 (Fig. 81L–M); embolus demarcated and roughly perpendicular to bulb, attenuate, long and straight with slightly swollen base, width at base/ bulb width 0.24, embolus length/bulb length 2.56 (Fig. 81L–M).
Female (QMB S118220)
GENERAL (Fig. 82A–L). Body length 25.16, in good condition.
DORSAL PROSOMA (Fig. 82A, E–F). Carapace length 8.96, width 8.65, length/width 1.04, clypeus to fovea length/carapace length 0.78, caput width/carapace width 0.74, carapace red-brown, reflective setae present, light on caput, light on thorax, fovea procurved, fovea width/carapace length 0.21 (Fig. 82A, F); chelicerae dark red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.62 (Fig. 82A); eye group rectangular, width/length 1.82, eye tubercle present (Fig. 82E).
ABDOMEN (Fig. 82B, D). Abdomen length 10.77, dark brown, dorsal pattern absent, with consistent cover of short setae.
VENTRAL PROSOMA (Fig. 82C, G–I). Labium cuspules absent (Fig. 82H); maxillae heel distinct, cuspules present, count =about 216, extending posteriorly onto heel, extending laterally about 40% of maxillae length (Fig. 82C, I); coxae cuspules absent, thorn-like setae on prolateral face present (Fig. 82C, I); sternum length/width 1.06, central sternum with consistent covering of short setae, row of longer setae around posterior edges, setae at higher density around anterior edges (Fig. 82G–H); posterior sigilla elongate, central sternum to posterior sigilla length/sternum length 0.23, posterior sigilla length/sternum length 0.20 (Fig. 82G–H); other sigilla small, round and lateral (Fig. 82G–H).
LEG I (Fig. 82J–K). Leg I dark coffee-brown, darker on patella and tibia, femur length 6.85, patella length 4.64, tibia length 4.58, metatarsus length 4.29, tarsus length 2.76, total length 23.12, leg I length/ carapace length 2.58; scopulae on distal metatarsus and tarsus; spine count Fe D 0, Fe PL 1, Pa PL 1, Ti PL 0, Ti RL 4, Me PL 1, Me RL 4, Ta 0; tibia length/width [TIL/TID] 3.01.
GENITALIA (Fig. 82D, L). Epigastric furrow extending slightly, posterior edge with rounded shape (Fig. 82D); spermathecae with one vesicle each (Fig. 82L); lateral vesicle undulating, with several distinct bends, length 1.47, lateral vesicle length/genitalia width 0.81, length/width at base 4.04, crown slightly wider than stem (Fig. 82L).
Distribution and natural history
Aname warialda has a wide distribution in northern New South Wales and south-eastern Queensland, in the Brigalow Belt South, Nandewar and New England Tablelands bioregions, from Tamworth north to Carnarvon National Park, and east to Toowoomba (Fig. 11). It constructs an open, silk-lined burrow with silk spilling out from the entrance, sometimes with a built-up mound of soil around the entrance. The burrow has a short, hidden ‘wishbone’ entrance, which, when excavated, is revealed to terminate in a distinctly thick, white sheath of silk (Fig. 11).
description: Published as part of Wilson, Jeremy D., Harvey, Mark S., Simmons, Leigh W. & Rix, Michael G., 2025, An integrative systematic revision of the wishbone spiders (Araneae: Anamidae: Aname L. Koch, 1873) of subtropical and tropical eastern Australia, with the description of 55 new species, pp. 1-298 in European Journal of Taxonomy 985 on pages 126-129, DOI: 10.5852/ejt.2025.985.2845, http://zenodo.org/record/15169471