cometadata/datacite-rtg-text-other-reclassification-records huggingface dataset:
(Homobezzia)
Creators: name: Huerta, Herón
name: Spinelli, Gustavo R.
name: Grogan Jr, William L.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia

description: Subgenus Homobezzia glabra group
Diagnosis. (Modified from Spinelli & Wirth 1989a). Moderately large species, pollinose gray with brownish scutal vittae and yellowish scutellum. Interocular space without stout median seta. Male flagellum with distal flagellomeres elongated; segment 13 without stout black basal seta. Scutum usually pollinose, pale pinkish to dark grayish, usually with median brown vitta anteriorly and a pair of lateral vittae posteriorly; setae of scutellum and above wing bases poorly developed. Wing translucent, usually unmarked. Legs extensively yellowish, with narrow dark bands usually present; femora and tibiae at most with broad brown bands on 1 pair of legs; vestiture not strong or bristly; fore femur unarmed ventrally; female claws small; with 1 or 2 pairs of gland rods. Male gonocoxite short, gonostyles about as long as gonocoxite, slightly curved and usually with blunt tip; aedeagus various, usually triangular; height of basal arch, breadth of distal process, and extent of ventral spicules vary with species; parameres with long rod-like distal process, sometimes slightly narrowed in midportion and or slightly broader apically, with tip rounded.
This group includes six species in the Neotropical region (Spinelli & Wirth 1989a), one species ranges north into the Nearctic region (B. glabra).
description: Published as part of Huerta, Herón, Spinelli, Gustavo R. & Grogan Jr, William L., 2023, New records of predaceous midges in Bezzia Kieffer and Phaenobezzia Haeselbarth from Mexico with description of two new species of Bezzia (Diptera: Ceratopogonidae), pp. 535-552 in Zootaxa 5323 (4) on page 543, DOI: 10.11646/zootaxa.5323.4.5, http://zenodo.org/record/8212446
Bezzia (Homobezzia) pulchripes Kieffer 1917
Creators: name: Huerta, Herón
name: Spinelli, Gustavo R.
name: Grogan Jr, William L.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Ceratopogonidae
Bezzia
Bezzia pulchripes

description: Bezzia (Homobezzia) pulchripes Kieffer
(Fig. 4A–D)
Bezzia pulchripes Kieffer, 1917: 330 (female; Paraguay). Lane 1958: 27 (key); Mayer 1959: 232 (pupa; Brazil); Wirth 1959: 234 (male; Brazil); Wirth 1974: 14.53 (catalogue of the Americas south of USA); Spinelli & Wirth 1989a: 768 (female, male, pupa; redescription; figures; distribution); Dippolito & Spinelli 1995: 54 (record Rondonia, Brazil); Borkent & Wirth 1997: 128 (world catalog); Borkent & Spinelli 2000: 61 (catalog of new world south of the United States); Borkent & Spinelli, 2007: 94 (Neotropical catalog; distribution); Anjos et al. 2018: 152 (diagnostic characters of pupae); Borkent & Dominiak 2020: 203 (world catalog); Santarém & Felippe˗Bauer 2021: 34; Spinelli et al. 2023: 63 (catalog of biting midges from Argentina).
Diagnosis. Only species in the glabra group with the following combination of characters: palpal segment 5 reduced, shorter than segment 4; thorax dark brown with small anteromedian vertical spine; legs dark brown with narrow subapical pale bands on femora and tibiae, sub-basal on tibiae; bases of fore and mid femora broadly pale (Fig. 4A); tarsomeres 1–4 yellowish brown with apices infuscated, tarsomeres 5 darker; female abdomen with a pair of long sclerotized gland rods; two unequal-size spermathecae with long slender necks; male sternite 9 with shallow caudomedian excavation, tergite 9 with two apical lobes; aedeagus triangular, slightly longer than basal breadth; parameres with bilobed anterolateral plates.
Material examined. MEXICO, Yucatan, Reserva Ría Lagartos, Peten “Tucha”, 30-nov-1995, light trap, GPS [21° 35′ 46″ N, 88° 08′ 47″ W], Huerta, H., 1 female CAIM [CAIMCrt/lam-00266]. New record from Yucatan.
Distribution. Widespread, from Mexico (Tabasco, Yucatan) to northeastern Argentina.
Remarks. Spinelli & Wirth (1989a) indicated that the subapical pale band on the hind tibiae is not present in some specimens from Belize and Jamaica, and it is absent in the above specimen from Yucatan.
description: Published as part of Huerta, Herón, Spinelli, Gustavo R. & Grogan Jr, William L., 2023, New records of predaceous midges in Bezzia Kieffer and Phaenobezzia Haeselbarth from Mexico with description of two new species of Bezzia (Diptera: Ceratopogonidae), pp. 535-552 in Zootaxa 5323 (4) on pages 545-546, DOI: 10.11646/zootaxa.5323.4.5, http://zenodo.org/record/8212446
Bezzia (Homobezzia) snowi Lane 1958
Creators: name: Huerta, Herón
name: Spinelli, Gustavo R.
name: Grogan Jr, William L.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Ceratopogonidae
Bezzia
Bezzia snowi

description: Bezzia (Homobezzia) snowi Lane
(Fig. 1E)
Bezzia snowi Lane, 1958: 34 (female, male; Guatemala; figures). Wirth 1974: 53 (world catalog); Spinelli & Wirth 1991: 10 (redescription, female, male, pupa; key, figures); Borkent & Wirth 1997: 129 (world catalog); Borkent & Spinelli 2000: 62 (catalog of new world south of the United States); Borkent & Spinelli 2007: 94 (Neotropical catalog); Spinelli & Wolff 2016: 126 (Ceratopogonidae of Colombia; distribution); Ronderos et al. 2018: 645 (list immature stages); Anjos et al. 2018: 152 (diagnostic characters of pupae); Borkent & Dominiak 2020: 204 (world catalog); Santarém & Felippe˗Bauer 2021: 34 (Brazilian distribution); Spinelli et al. 2023: 64 (catalog of biting midges from Argentina).
Diagnosis. Only Neotropical species in the venustula group with the following combination of characters: female with legs yellowish, distal 1/3 of hind femur and distal 1/5 of hind tibia brown; femora armed ventrally with 5–8 spines on fore leg, 3–6 on mid leg, and 1–4 on hind leg; tarsi yellowish brown except 5th tarsomeres dark; halter dark brown; three pairs of long slender gland rods; male sternite 9 with shallow caudomedian excavation, tergite 9 with elongated apicolateral lobes; gonocoxite with blunt anteromesal protuberance; aedeagus triangular; parameres long with bilobed well-developed anterolateral plates, distal process with rounded tip.
New country record. MEXICO, Guerrero, Ayutla de los Libres, La Unión, “Rancho Camalote”, jun-2009, Malaise trap, Bibiano-Marín, W., 1 female CAIM. Oaxaca, San Juan Guichicovi, El Zacatal, potrero, 28-jul-2009, CDC trap, Salceda-Sanchez, B., Rodríguez, A. A., 1 female CAIM.
Distribution. Mexico (Guerrero, Oaxaca), Guatemala, Panama, Colombia, Trinidad, Brazil.
Remarks. This species differs from other related species of this group mainly by their femora with ventral spines (5–8 fore, 3–6 middle and 1–4 hind), legs yellowish with distal 1/3 of hind femur and distal 1/5 of hind tibia brown, two subspherical spermathecae with long necks, slightly unequal and parameres twice as long as length of aedeagus.
description: Published as part of Huerta, Herón, Spinelli, Gustavo R. & Grogan Jr, William L., 2023, New records of predaceous midges in Bezzia Kieffer and Phaenobezzia Haeselbarth from Mexico with description of two new species of Bezzia (Diptera: Ceratopogonidae), pp. 535-552 in Zootaxa 5323 (4) on page 546, DOI: 10.11646/zootaxa.5323.4.5, http://zenodo.org/record/8212446
Phaenobezzia Haeselbarth 1965
Creators: name: Huerta, Herón
name: Spinelli, Gustavo R.
name: Grogan Jr, William L.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Ceratopogonidae
Phaenobezzia

description: Phaenobezzia Haeselbarth, 1965
Type species: Probezzia pistiae Ingram & Macfie, by original designation.
description: Published as part of Huerta, Herón, Spinelli, Gustavo R. & Grogan Jr, William L., 2023, New records of predaceous midges in Bezzia Kieffer and Phaenobezzia Haeselbarth from Mexico with description of two new species of Bezzia (Diptera: Ceratopogonidae), pp. 535-552 in Zootaxa 5323 (4) on page 548, DOI: 10.11646/zootaxa.5323.4.5, http://zenodo.org/record/8212446
Phaenobezzia maya Spinelli & Wirth 1986
Creators: name: Huerta, Herón
name: Spinelli, Gustavo R.
name: Grogan Jr, William L.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Ceratopogonidae
Phaenobezzia
Phaenobezzia maya

description: Phaenobezzia maya Spinelli & Wirth
Phaenobezzia maya Spinelli & Wirth, 1986: 234 (male, female; Belize; figures; distribution). Dippolito & Spinelli 1995: 56 (record Rondonia, Brazil); Borkent & Wirth 1997: 135 (world catalog); Borkent & Spinelli 2000: 65 (new world catalog south of USA); Borkent & Spinelli 2007: 97 (Neotropical catalog; distribution); Borkent & Dominiak 2020: 213 (world catalog).
Diagnosis. The only species of Phaenobezzia with the following combination of characters: eyes separated by the diameter of 2–3 ommatidial facets; antenna with narrow bases of flagellar segments pale; legs yellowish brown; ventral palisade setae sparse on tarsomeres 1–3 of mid leg, dense on tarsomeres 1–3 of hind leg; tarsomeres on all legs with 4–5 pairs of stout, sharp-pointed ventral spines. Male genitalia with short gonostylus.
Material examined. MEXICO, Veracruz, Juchique de Ferrer, Dos Arroyos, LC15, cafetal, 7-sep-2008, CDC trap, 1 female CAIM. Yucatan, Reserva Ría Lagartos, Peten “Tucha”, 14-oct-1996, light trap, GPS [21° 35′ 46″ N, 88° 08′ 47″ W], Huerta, H., 1 male CAIM [CAIMCrt/lam-97-02897]; same date except, Tizimin, entrada Zac-Boo, selva baja caducifolia, light trap, 19-mar-1996, Ibáñez-Bernal, S., 1 female CAIM [CAIMCrt/lam-98-00731]. New Yucatan record.
Distribution. USA (Texas), Mexico (Morelos, Veracruz, Yucatan), Belize, El Salvador, Honduras, Costa Rica, Panama, Brazil.
Remarks. This species is widely distributed in the Neotropical region, but it is only known in the Nearctic region from Texas. We provide the first record from Yucatan.
description: Published as part of Huerta, Herón, Spinelli, Gustavo R. & Grogan Jr, William L., 2023, New records of predaceous midges in Bezzia Kieffer and Phaenobezzia Haeselbarth from Mexico with description of two new species of Bezzia (Diptera: Ceratopogonidae), pp. 535-552 in Zootaxa 5323 (4) on page 549, DOI: 10.11646/zootaxa.5323.4.5, http://zenodo.org/record/8212446
Phaenobezzia fulvithorax
Creators: name: Huerta, Herón
name: Spinelli, Gustavo R.
name: Grogan Jr, William L.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Ceratopogonidae
Phaenobezzia
Phaenobezzia fulvithorax

description: Phaenobezzia fulvithorax (Malloch)
Probezzia fulvithorax Malloch, 1915: 354 (male, female; USA, Illinois).
Phaenobezzia fulvithorax (Malloch): Wirth & Grogan 1982: 185 (redescription; combination; distribution); Spinelli & Wirth 1986: 234 (Mexico record); Borkent & Wirth 1997: 135 (world catalog); Borkent & Spinelli 2000: 65 (new world catalog south of USA); Borkent & Dominiak 2020: 213 (world catalog).
Diagnosis. The only species of Phaenobezzia with the following combination of characters: eyes broadly separated by the diameter of 5–6 ommatidia; antenna dark brown, flagellomere 13 bluntly rounded, not tapered distally; legs yellowish brown, hind femur and tibia with dark apices; ventral palisade setae sparse on tarsomere 1 of mid leg, dense on tarsomeres 1–2 on hind leg, tarsomeres 5 on all legs with 4 pairs of stout, sharp-pointed ventral spines. Male gonostylus about half as long as the gonocoxite.
Material examined. MEXICO, Guerrero, Cocula, Apipilulco, sept-1995, light trap, 1 female CAIM [CAIM / Crt/lam-00265]. Morelos, San Rafael Vicente Aranda, 6-may-1983, light trap, 10 females, 7 males CAIM [CAIM / Crt/lam−00267−00284]. New records from Guerrero and Morelos.
Distribution. British Columbia and Quebec (Canada), in USA from Washington, east to Michigan, south to California, east to Texas and Arkansas, Mexico (Baja California, Nuevo Leon, Morelos, Guerrero).
Remarks. This species has a wide Nearctic distribution, included in Baja California, Mexico (Wirth & Grogan 1982). Spinelli & Wirth (1986) recorded this species from Nuevo Leon. We herein include new records from the Neotropical region of Mexico (Guerrero, Morelos).
description: Published as part of Huerta, Herón, Spinelli, Gustavo R. & Grogan Jr, William L., 2023, New records of predaceous midges in Bezzia Kieffer and Phaenobezzia Haeselbarth from Mexico with description of two new species of Bezzia (Diptera: Ceratopogonidae), pp. 535-552 in Zootaxa 5323 (4) on page 549, DOI: 10.11646/zootaxa.5323.4.5, http://zenodo.org/record/8212446
Calappa Weber 1795
Creators: name: Charbonnier, Sylvain
name: Garassino, Alessandro
name: Pasini, Giovanni
name: Chény, Cédric
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Calappidae
Calappa

description: Genus Calappa Weber, 1795
TYPE SPECIES. — Cancer granulatus Linnaeus, 1758, subsequent designation by Latreille (1810).
INCLUDED FOSSIL SPECIES. — See Schweitzer et al. (2010) and Pasini & Garassino (2017).
description: Published as part of Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni & Chény, Cédric, 2024, Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria, pp. 13-29 in Geodiversitas 46 (2) on page 16, DOI: 10.5252/geodiversitas2024v46a2, http://zenodo.org/record/10566276
Epialtidae MacLeay 1838
Creators: name: Charbonnier, Sylvain
name: Garassino, Alessandro
name: Pasini, Giovanni
name: Chény, Cédric
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Epialtidae

description: Family EPIALTIDAE MacLeay, 1838
DIAGNOSIS. — Carapace elongate, triangular or cuneate, sometimes with unusually projected antero-lateral margins; rostrum bifid or singular, long or short; eyes without true orbits, supraorbital eave weak, without intercalated spine, eyestalks short or absent, eyes protected by very long rostrum or preorbital spine, sometimes a postorbital spine present but not cupped to protect eye; merus and ischium of maxilliped equally wide; pereiopods 3-5 often very short compared to 1 and 2 (after Schweitzer et al. 2020).
INCLUDED SUBFAMILIES. — Epialtinae MacLeay,1838; Periacanthinae Lőrenthey in Lőrenthey & Beurlen, 1929; Pisinae Dana, 1851; Pliosomatinae Števčič, 1994; Tychinae Dana, 1851; Wahraniinae n. subfam. (this study).
description: Published as part of Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni & Chény, Cédric, 2024, Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria, pp. 13-29 in Geodiversitas 46 (2) on page 18, DOI: 10.5252/geodiversitas2024v46a2, http://zenodo.org/record/10566276
Wahraniinae Charbonnier & Garassino & Pasini & Chény 2024, n. subfam.
Creators: name: Charbonnier, Sylvain
name: Garassino, Alessandro
name: Pasini, Giovanni
name: Chény, Cédric
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Epialtidae

description: WAHRANIINAE n. subfam.
urn:lsid:zoobank.org:act: 176801F1-BE77-451A-B53A-7DF9308518FF
TYPE GENUS. — Wahrania n. gen.
DIAGNOSIS. — Carapace pyriform; long, diverging pseudorostral spines (= rostrum); rimmed supraorbital eave with sharp postorbital spine; one very long, sharp hepatic spine with a pair of basal accessorial spines; one long, sharp epibranchial spine; one long, sharp mesobranchial spine; two long, sharp metabranchial spines; dorsal surface of carapace and spines strongly tuberculate.
DISCUSSION
Following Schweitzer et al. (2020), Wahraniinae n. subfam. is placed within Epialtidae based on the weak supraorbital eave without intercalated spine, the very long rostrum bifid, protecting the eye, and one postorbital spine present but not cupped to protect eye. Wahraniinae n. subfam. differs from the Epialtinae, Periacanthinae, Pisinae, Pliosomatinae, and Tychinae by its very elongate antero- and posterolateral spines developed only on the margin and not on the dorsal surface, and by its hepatic spine flanked by basal accessorial spines. The marginal position of antero- and posterolateral spines is closed to Periacanthinae but Periacanthus Bittner, 1875, besides a short bifid rostrum (versus very long bifid rostrum in Wahrania n. gen.), shows a trifurcate spine on branchial region (versus single and simple spine in Wahrania n. gen.) and a preorbital spine to protect the eye (versus absent in Waharnia n. gen.)
description: Published as part of Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni & Chény, Cédric, 2024, Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria, pp. 13-29 in Geodiversitas 46 (2) on page 18, DOI: 10.5252/geodiversitas2024v46a2, http://zenodo.org/record/10566276
Wahrania arambourgi Charbonnier & Garassino & Pasini & Chény 2024, n. comb.
Creators: name: Charbonnier, Sylvain
name: Garassino, Alessandro
name: Pasini, Giovanni
name: Chény, Cédric
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Goneplacidae
Wahrania
Wahrania arambourgi

description: Wahrania arambourgi (Van Straelen, 1937), n. comb.
(Fig. 2E, F)
Maia [sic] arambourgi Van Straelen, 1937: 475, 476, pl. 33, fig. 3. — Glaessner 1969: R502, fig. 314.9.
Maja arambourgi – Saint Martin & Müller 1988: 251. — Moissette & Müller 1990: 738, 739. — Schweitzer et al. 2010: 97.
TYPE MATERIAL. — Holotype by monotypy (MNHN.F.R03773, Arambourg coll.).
TYPE LOCALITY. — Les Planteurs quarry, Sanaouber district of Sidi El Houari, Oran.
TYPE AGE. — Late Miocene (late Messinian).
DESCRIPTION
Carapace
Carapace pyriform;pseudorostral spines (= rostrum)longer than the half of carapace length; pseudorostral spines sharp, straight, diverging,slightly deflected;supraorbital eave rimmed with sharp postorbital spine; one very long, sharp hepatic spine, as long as the pseudorostral spines, slightly turned downward; a pair of short,sharp, straight accessorial spines at the base of hepatic spine; one long, sharp epibranchial spine, slightly turned upward; one long, sharp, straight mesobranchial spine, as long as epibranchial spine, directed outward; two long, sharp, straight metabranchial spines, as long as mesobranchial spine, directed posteriorly; narrow posterior margin poorly preserved; meso- and metagastric regions slightly inflated; urogastric and cardiac regions slightly inflated and marked by a deep branchiocardiac groove; dorsal surface of carapace and spines strongly tuberculate(based onVan Straelen 1937: pl. 33, fig. 3, we note that the surface of carapace was altered and eroded, inducing the loose of ornamentation).
DISCUSSION
The fossil record of the representatives of the Epialtidae from the Miocene is very scarce. Indeed, only eight genera of the Epialtinae and Pisinae are known to date from the Miocene of Europe (Hungary and Poland), North America (USA), South America (Argentina, Peru, and Venezuela), Taiwan, and Japan (Schweitzer et al. 2020). However, only two fossil species are known to date from the Miocene of the Mediterranean Basin: Herbstia melitense Gatt & De Angeli, 2010 (Pisinae Dana, 1851) from Malta and Wahrania arambourgi (Van Straelen, 1937), n. comb. (Wahraniinae n. subfam.) from Algeria.
description: Published as part of Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni & Chény, Cédric, 2024, Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria, pp. 13-29 in Geodiversitas 46 (2) on page 19, DOI: 10.5252/geodiversitas2024v46a2, http://zenodo.org/record/10566276
Oregoniidae Garth 1958
Creators: name: Charbonnier, Sylvain
name: Garassino, Alessandro
name: Pasini, Giovanni
name: Chény, Cédric
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Oregoniidae

description: Family OREGONIIDAE Garth, 1958
INCLUDED GENERA. — Chionoecetes KrØyer, 1838; Macroregonia Sakai, 1978; Oregonia, Dana, 1851; Hyas Leach, 1814 (after Ng et al. 2008).
description: Published as part of Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni & Chény, Cédric, 2024, Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria, pp. 13-29 in Geodiversitas 46 (2) on page 19, DOI: 10.5252/geodiversitas2024v46a2, http://zenodo.org/record/10566276
Parahyas Charbonnier & Garassino & Pasini & Chény 2024, n. gen.
Creators: name: Charbonnier, Sylvain
name: Garassino, Alessandro
name: Pasini, Giovanni
name: Chény, Cédric
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Epialtidae
Parahyas

description: Parahyas n. gen.
urn:lsid:zoobank.org:act: B38E2FCA-0FC0-43E8-99A4-3F94395BEA5E
ETYMOLOGY. — A combination between Para (Latin: similar to) and the extant genus Hyas Leach, 1814. The gender of the genus is masculine.
TYPE SPECIES. — Hyas oranensis Van Straelen, 1937.
DIAGNOSIS. — Carapace pyriform; orbit composed of supraorbital rim and widely cup-shaped postorbital spine bearing marginal spines; inflated branchial regions with posterolateral margin bearing row of tubercles; intestinal region protruding distally beyond and under branchial regions; urogastric and cardiac regions limited by deep branchiocardiac groove.
DISCUSSION
Parahyas n. gen. is assigned to the Oregoniidae based upon the widely cup-shaped postorbital spine, which is well visible in several extant representatives such as Hyas coarctatus Leach, 1815 and H. ursinus Rathbun, 1924. Parahyas n. gen. differs from Oregonia, Chionoecetes, Macroregonia by the presence of widely cup-shaped postorbital spine (absent in all these genera). Parahyas n. gen. differs from Hyas by the presence of a row of large tubercles extending along the entire length of the posterolateral margin (versus sometimes scarce small tubercles along the margin in Hyas), dorsal surface of carapace covered by small tubercles arranged uniformly (versus scattered granules and tubercles of varying size in Hyas), and gastric, urogastric, and cardiac regions without median raised tubercles (versus above-mentioned regions with raised median tubercles in Hyas).
description: Published as part of Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni & Chény, Cédric, 2024, Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria, pp. 13-29 in Geodiversitas 46 (2) on page 19, DOI: 10.5252/geodiversitas2024v46a2, http://zenodo.org/record/10566276
Parahyas oranensis Charbonnier & Garassino & Pasini & Chény 2024, n. comb.
Creators: name: Charbonnier, Sylvain
name: Garassino, Alessandro
name: Pasini, Giovanni
name: Chény, Cédric
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Epialtidae
Parahyas
Parahyas oranensis

description: Parahyas oranensis (Van Straelen, 1937), n. comb.
(Fig. 3)
Hyas oranensis Van Straelen, 1937: 474, 475, pl. 33, fig. 2. — Saint Martin & Müller 1988: 251. — Moissette & Müller 1990: 738. — Schweitzer et al. 2010: 97. — Kato et al. 2013: 270, 275-277. — Guinot et al. 2019: 307.
TYPE MATERIAL. — Holotype MNHN.F.R03767, one paratype MNHN.F.A28903 (Arambourg coll.).
TYPE LOCALITY. — Les Planteurs quarry, Sanaouber district of Sidi El Houari, Oran.
TYPE AGE. — Late Miocene (late Messinian).
DESCRIPTION
Carapace
Carapace pyriform, convex both longitudinally and transversely, length slightly exceeding width; widest posteriorly; rostrum not preserved; orbit composed of supraorbital rim and widely cup-shaped postorbital spine bearing marginal spines; anterolateral margins poorly preserved; posterolateral margins rounded, bearing row of 6-7 wide tubercles extending along entire length of margin; posterior margin partially preserved; hepatic region depressed; proto-, meso-, and metagastric regions raised; diamond-shaped cardiac region strongly elevated; branchial regions inflated; urogastric and cardiac regions raised and limited laterally by deep branchiocardiac groove; intestinal region protruding distally beyond and under branchial regions; dorsal surface of carapace covered with small tubercles arranged uniformly.
DISCUSSION
Van Straelen (1937) figured only the holotype but the paratype is better preserved and shows some additional characters such as the wide cup-shaped postorbital spine, the well-developed intestinal region beyond and under the inflated branchial regions, and the posterolateral margins with a row of large aligned tubercles.
Extant members of the Oregoniidae are relatively rare in the Mediterranean Basin and only Ergasticus A. Milne-Edwards, 1882 has been reported (GBIF 2022:https://www.gbif.org/species/4375129). In the fossil record, Hyas meridionalis Glaessner, 1928 was described from the Miocene of Austria, but the type specimen is poorly preserved and, even if it is surely a member of Majoidea, the generic assignment seems to be doubtful. In conclusion, Parahyas n. gen. is most likely the only Mediterranean representative of the Oregoniidae in the fossil record.
description: Published as part of Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni & Chény, Cédric, 2024, Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria, pp. 13-29 in Geodiversitas 46 (2) on pages 19-22, DOI: 10.5252/geodiversitas2024v46a2, http://zenodo.org/record/10566276
Cancer Linnaeus 1758
Creators: name: Charbonnier, Sylvain
name: Garassino, Alessandro
name: Pasini, Giovanni
name: Chény, Cédric
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Cancridae
Cancer

description: Genus Cancer Linnaeus, 1758
TYPE SPECIES. — Cancer pagurus Linnaeus, 1758, subsequent designation by Latreille (1810).
INCLUDED FOSSIL SPECIES. — See Schweitzer et al. (2010).
description: Published as part of Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni & Chény, Cédric, 2024, Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria, pp. 13-29 in Geodiversitas 46 (2) on page 22, DOI: 10.5252/geodiversitas2024v46a2, http://zenodo.org/record/10566276
Cancer sismondai Meyer 1843
Creators: name: Charbonnier, Sylvain
name: Garassino, Alessandro
name: Pasini, Giovanni
name: Chény, Cédric
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Cancridae
Cancer
Cancer sismondai

description: Cancer sismondai Meyer, 1843
(Fig. 4 A-E)
Cancer sismondae Meyer, 1843: 590.
Cancer deshayesii A. Milne-Edwards, 1864a: 74-76, pl. 4, fig. 1, 2, pl. 5, fig. 2.
Lobocarcinus sismondai – Ossó et al. 2022: 213, 214, fig. 4a-c (cum syn.).
TYPE MATERIAL. — Cancer sismondai: holotype from the Pliocene of Santo Stefano Roero, Piedmont, Italy, figured by A. Sismonda (1839: figs A, B), but lost during Second World War after Bonfiglio & Donadeo (1982).
Cancer deshayesii: syntype MNHN.F.B39264 (Marès coll.) from the Pliocene of Alger, Algeria; other syntypes (Deshayes coll.) from the Messinian of Oran are housed in the collection of the Ecole des Mines de Paris deposited at the University Claude Bernard Lyon 1, France.
EXAMINED MATERIAL. — 23 specimens from Les Planteurs quarry, Oran (MNHN.F.A51668, A51669, A51670, A51671, A51672, A51673, A51674, A51675, A51676, A51677, A51738, A51739, A51740, A51741, A41743, A41744, A41745, A41746, A41747, A51748, A51750, A51761, A51946, MNHN.F.R03507, R03537, Arambourg coll.).
DISCUSSION
A. Milne-Edwards(1864a) first described Cancer deshayesii based on a spectacular specimen from the Pliocene marls of Alger and other specimens (Deshayes coll.) from the Miocene of Oran. This species was later synonymized with Cancer sismondai Meyer, 1843 by Ristori (1886). This synonymy and the placement in Lobocarcinus Reuss, 1857, was discussed by several authors through the 20th century (e.g., Van Straelen 1937; Bonfiglio & Donadeo 1982) until Ossó et al. (2022). All the Algerian specimens assigned to deshayesii or sismondai are currently in review by the authors and will be the subject of a future publication.
description: Published as part of Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni & Chény, Cédric, 2024, Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria, pp. 13-29 in Geodiversitas 46 (2) on page 22, DOI: 10.5252/geodiversitas2024v46a2, http://zenodo.org/record/10566276
Cancer undetermined
Creators: name: Charbonnier, Sylvain
name: Garassino, Alessandro
name: Pasini, Giovanni
name: Chény, Cédric
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Cancridae
Cancer
Cancer undetermined

description: Cancer sp.
(Fig. 4F)
Cancer sp. – Van Straelen 1937: 447.
EXAMINED MATERIAL. — MNHN.F.A51742 (Arambourg coll.) from the Messinian of Les Planteurs quarry, Sanaouber district of Sidi El Houari, Oran.
DESCRIPTION
Giant left P1 chela (length: ca 9 cm from index apex to carpopropodial articulation), dorsal margin strongly convex, bearing strong tubercules; ventral margin almost straight and smooth; index slightly curved with smooth occlusal margin; dactylus strongly curved with smooth occlusal margin; occlusal opening wide; palm without carinae but with outer surface uniformly covered by very small scaly pits; index and dactylus bearing brownish coloured pattern contrasting with whitish general colour of chela.
DISCUSSION
Van Straelen (1937) proposed to place this isolated P1 chela in Cancer sp. based on its similarity with the robust chelae of Cancer pagurus. He added that the curvature of the dactylus was different from this extant species. Our examination leads to highlight some new characters: occlusal margin of index strongly incurved and smooth, smooth occlusal margin of dactylus, and strongly convex dorsal margin of propodus with at least three big tubercles. These characters do not fit those of C. pagurus, which shows the P1 chela with occlusal margins of fingers bearing molariform teeth and straight smooth dorsal margin of propodus. Based on Nations (1975), some species of Cancer show the dorsal margin of propodus with spines or tubercles, but the margin is always straight or slightly convex and the occlusal margins of both finger always bearing big teeth. Moreover, all species of Cancer show the outer surface of palm with a variable number of raised tuberculate carinae. The subquared outline of the P1 palm from Oran is also reminiscent of the P1 palm of some extant genera (Anatolikos Schweitzer& Feldman, 2000, Glebocarcinus Nations, 1975, and Platepistoma Rathbun, 1906). However, all the P1 chela of these living genera bear occlusal margin with molariform teeth and longitudinal carinae on palm (see Schram & Ng 2012: fig. 2). In conclusion, according to Van Straelen (1937), we leave this giant P1 chela in open nomenclature within Cancer, though it could represent a new species of cancrinid crab. We point out the orange brown colour pattern of the fingers, which constitutes a case of exceptional preservation
description: Published as part of Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni & Chény, Cédric, 2024, Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria, pp. 13-29 in Geodiversitas 46 (2) on page 22, DOI: 10.5252/geodiversitas2024v46a2, http://zenodo.org/record/10566276
Goneplax Leach 1814
Creators: name: Charbonnier, Sylvain
name: Garassino, Alessandro
name: Pasini, Giovanni
name: Chény, Cédric
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Goneplacidae
Goneplax

description: Genus Goneplax Leach, 1814
TYPE SPECIES. — Ocypoda bispinosa Lamarck, 1801 by original designation (junior subjective synonym of Cancer rhomboides Linnaeus, 1758).
INCLUDED FOSSIL SPECIES. — Goneplax gulderi Bachmayer, 1953, G. rhomboides (Linnaeus, 1758).
description: Published as part of Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni & Chény, Cédric, 2024, Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria, pp. 13-29 in Geodiversitas 46 (2) on page 25, DOI: 10.5252/geodiversitas2024v46a2, http://zenodo.org/record/10566276
Chlinocephalus Ristori 1886
Creators: name: Charbonnier, Sylvain
name: Garassino, Alessandro
name: Pasini, Giovanni
name: Chény, Cédric
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Malacostraca
Decapoda
Euryplacidae
Chlinocephalus

description: Genus Chlinocephalus Ristori, 1886
TYPE SPECIES. — Chlinocephalus demissifrons Ristori, 1886, by monotypy.
INCLUDED FOSSIL SPECIES. — The sole type species.
description: Published as part of Charbonnier, Sylvain, Garassino, Alessandro, Pasini, Giovanni & Chény, Cédric, 2024, Review of brachyuran crabs from the late Miocene (Messinian) of Oran, Algeria, pp. 13-29 in Geodiversitas 46 (2) on page 22, DOI: 10.5252/geodiversitas2024v46a2, http://zenodo.org/record/10566276
Limonia
Creators: name: Podenas, Sigitas
name: Podeniene, Virginija
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Limoniidae
Limonia

description: Key to pupae of Korean species of Limonia
1. Pronotal horns approximaly as long as wide (Fig. 52).............................. Limonia nubeculosa Meigen, 1804
- Pronotal horns almost twice as long as wide (Figs. 34, 69).....................................................2
2. The inner margin of pronotal horns straight (Fig. 34)......................... Limonia macrostigma (Schummel, 1829)
- The inner margin of pronotal horns with outgrowth posteriorly (Fig. 69)............ Limonia parvipennis Alexander, 1940
description: Published as part of Podenas, Sigitas & Podeniene, Virginija, 2017, Limonia crane flies (Diptera: Limoniidae) of Korea, pp. 1-37 in Zootaxa 4231 (1) on page 6, DOI: 10.11646/zootaxa.4231.1.1, http://zenodo.org/record/293646
Limonia
Creators: name: Podenas, Sigitas
name: Podeniene, Virginija
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Limoniidae
Limonia

description: Key to last instar larvae of Korean species of Limonia
1. Dorsal and ventral sclerites of spiracular field heavily sclerotised (Fig. 33)....... Limonia macrostigma (Schummel, 1829)
- Dorsal and ventral sclerites of spiracular field pale (Figs. 51, 68)................................................2
2. Caudal end of head capsule (frons) tridentate (Fig. 48)............................. Limonia nubeculosa Meigen, 1804
- Caudal end of head capsule (frons) without remarkable teeth (Fig. 61).............. Limonia parvipennis Alexander, 1940
description: Published as part of Podenas, Sigitas & Podeniene, Virginija, 2017, Limonia crane flies (Diptera: Limoniidae) of Korea, pp. 1-37 in Zootaxa 4231 (1) on page 6, DOI: 10.11646/zootaxa.4231.1.1, http://zenodo.org/record/293646
Japanagromyza cercariae Sasakawa
Creators: name: Sasakawa, Mitsuhiro
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Agromyzidae
Japanagromyza
Japanagromyza cercariae

description: Japanagromyza cercariae Sasakawa
Japanagromyza cercariae Sasakawa, 1963a: 26.
Remarks. This species is similar to Japanagromyza cupreata in coloration, but is recognizable by its less shining mesoscutum, presence of spine on the antero-ventral apex of the cercus, and two apical spines on the claw-like surstylus, short membranous hypophallus, and two long tubules of the distiphallus (Sasakawa 1963a: Fig. 2). The external character of a female found for the first time was similar to that of the holotype male, except for the abdomen was densely dusted with brownish gray as well as the mesoscutum..
Material examined. VIETNAM – Ƥ, 18 km NW of Dalat, 1300m, 4–5. V. 1960, S. & L. Quate (BPBM). Distribution. Vietnam, China (Hong Kong).
description: Published as part of Sasakawa, Mitsuhiro, 2010, A review of the Oriental Japanagromyza Sasakawa (Diptera: Agromyzidae), with descriptions of four new species, pp. 16-32 in Zootaxa 2485 on page 21, DOI: 10.5281/zenodo.195513
Japanagromyza cupreata Sasakawa
Creators: name: Sasakawa, Mitsuhiro
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Agromyzidae
Japanagromyza
Japanagromyza cupreata

description: Japanagromyza cupreata Sasakawa
(Figs. 8–10)
Japanagromyza cupreata Sasakawa, 1963a: 27.
Remarks. This species was described by a single female from Vietnam in having greenish- or bluish-coppery mesoscutum and abdomen. The males examined agreed with the original description except for the presence of only one fore-tibial lateral bristle, and r-m situated at basal 1/3 of discal cell. Therefore, the possession of two fore-tibial bristles in the female holotype is not a diagnostic character for the species.
The male genitalia are described herein for the first time: surstylus narrower than epandrium, with apical spine; cercus with two strong setae on apex and a group of 10 or 11 setulae near anterior apex; proctiger with lateral processes extremely long, each with six or seven apical setulae. Hypandrium 660 µm long, sidepiece narrow, basal apodeme 1/5 as long as sidepiece; postgonite 170 µm long, as wide as hypandrial sidepiece, hairy along outer margin. Phallapodeme 800 µm long, broadened on basal 1/5; phallus 670 µm long; epiphallus short; basiphallic sclerites united distally; right hypophallus long and broadened distally, left hypophallus with narrow annulation; right paraphallus longer than left claw-like one; distiphallus long tubulous, narrowing distally. Ejaculatory apodeme 340 µm long, blade broadened asymmetrically. The hypophallus and paraphallus of this species are distinctive.
Material examined. THAILAND – 33, Phetchabun Prov., Khao Kho National Park, savanna at nursery, 16º52.568΄N, 520 m, 12–19. X. 2006, S. Chachumnan and S. Singtong, Malaise trap (NHMLA).
Distribution. Vietnam, Laos, Thailand (new record).
description: Published as part of Sasakawa, Mitsuhiro, 2010, A review of the Oriental Japanagromyza Sasakawa (Diptera: Agromyzidae), with descriptions of four new species, pp. 16-32 in Zootaxa 2485 on page 22, DOI: 10.5281/zenodo.195513
Japanagromyza duchesneae Sasakawa
Creators: name: Sasakawa, Mitsuhiro
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Agromyzidae
Japanagromyza
Japanagromyza duchesneae

description: Japanagromyza duchesneae (Sasakawa)
Agromyza duchesneae Sasakawa, 1954b: 106.
Japanagromyza duchesneae: Sasakawa, 1958: 139.
Remarks. This is a leafminer of Duchesnea indica Focke (Rosaceae) and also of Potentilla sp. The male and female genitalia of this species are very similar to those of Japanagromyza tristella (Sasakawa 1961: Figs. 17 and 20). But, these two species belong to the different species-group by color of the halter and shape of the larval posterior spiracle as noted in the key.
Material examined. TAIWAN – Ƥ, Chiayi Prov., Alishan Mts., 2400 m, 12–16. VI. 1965, Maa and Lin (BPBM). THAILAND – Ƥ, Chaiyaphum Prov., Tat Tone National Park, near stream, 15º58.538'N, 280 m, 26. VII–7. VIII. 2006, T. Jaruphan and O. Budsawong, Malaise trap; Ƥ, Sakon Prov., Nakhon Phu Pha Yon National Park, Channel, 16º55.639΄N, 295 m, 5–11. VII. 2006, M. Ngojansri and C. Cheaukamjan, Malaise trap (NHMLA). VIETNAM – Ƥ, Lamdon Prov., Dalat, 1500 m, 29. IV.–4. V. 1960, S. & L. Quate (BPBM).
Distribution. Japan; Taiwan, Thailand (new record), Vietnam (new record), Vanuatu, Papua New Guinea.
description: Published as part of Sasakawa, Mitsuhiro, 2010, A review of the Oriental Japanagromyza Sasakawa (Diptera: Agromyzidae), with descriptions of four new species, pp. 16-32 in Zootaxa 2485 on page 24, DOI: 10.5281/zenodo.195513
Japanagromyza elaeagni Sasakawa
Creators: name: Sasakawa, Mitsuhiro
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Agromyzidae
Japanagromyza
Japanagromyza elaeagni

description: Japanagromyza elaeagni (Sasakawa)
Agromyza elaeagni Sasakawa, 1954a: 38.
Japanagromyza elaeagni: Sasakawa, 1958: 140.
Remarks. This is a leafminer of Elaeagnus umbellata Thunb. (Elaeagnaceae) in Japan. The male genitalia are characteristic in having a pair of three black finger-shaped processes projected laterally in postero-lateral corners of the phallic hood, and one tubule of the distiphallus (Sasakawa 1954a: Fig. 5b). The larval posterior spiracle is trifurcate as that of Japanagromyza tristella, but the pores are only 27 or 28 in total (in tristella, 37- 48 pores) (Sasakawa 1961: Figs. 18 and 20).
Material examined. TAIWAN – Ƥ, Kao Feag, 1260 m, 3.2 km from Wu Sheh, 17. I. 1962, C.M. Yoshimoto (BPBM). BONIN ISLANDS– 3, Chichi-jima I., Mt. Mikazuki-yama, 150 m, 14.VIII. 1980, J.L. Gressitt; 3,Ƥ, Haha-jima I., South Ridge of Mt. Chibusa-yama, 300 m, 20–22. VIII. 1980, Gressitt (BPBM).
Distribution. Japan, Bonin Islands; Taiwan (new record).
description: Published as part of Sasakawa, Mitsuhiro, 2010, A review of the Oriental Japanagromyza Sasakawa (Diptera: Agromyzidae), with descriptions of four new species, pp. 16-32 in Zootaxa 2485 on page 24, DOI: 10.5281/zenodo.195513
Japanagromyza eucalypti Spencer
Creators: name: Sasakawa, Mitsuhiro
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Agromyzidae
Japanagromyza
Japanagromyza eucalypti

description: Japanagromyza eucalypti Spencer
(Figs. 11–13)
Japanagromyza eucalypti Spencer, 1963a: 309.
Remarks. This is a leafminer of Eucalyptus camaldulensis Dehnhardt (Myrtaceae) in Australia. This species is readily recognizable by a pale knob of the halter, the yellowish brown tarsi, the elongate surstylus with an apical spine, the proctiger with a pair of long lateral processes, and the distiphallus with long tubule.
Spencer (1963a, 1977a) described briefly and illustrated roughly the long phallus (1963a: Fig. 1A), and the elongated epandrium and cercus (1977a: Fig. 174). His ‘distinctive epandrium’ means accurately both the elongate surstylus and process of the proctiger, and also his ‘distinctive ventral appendage’ consists of the hypophallus and paraphallus. Diagnostic character of the male genitalia is described here in detail: the surstylus has an apical spine; proctiger is well sclerotized, setulose on apices of long lateral processes; hypandrium is narrow U-shaped, 620 µm long, with short basal apodeme (140 µm long); phallapodeme is 740 µm long; phallus is 630 µm long; basiphallus with left sclerite distinctly longer than right one; hypophallus is narrow, right paraphallus is distinctly broader than the left; distiphallic tubule is approximately twice as long as basiphallus; ejaculatory apodeme is broadened asymmetrically, 255 µm long (not narrow as shown in Fig. 1B by Spencer, probably in lateral view).
Female genitalia: see Sasakawa 1996.
Material examined. THAILAND – 3, Chaiyaphum Prov., Pa Hin Ngam National Park, Ecotone between mix deciduous and dry dipterocarp, 15º38.100΄N, 700 m, 18–24. VIII. 2006, K. Sa-nog and B. Adnafai, Malaise trap (NHMLA).
Distribution. Australia, Papua New Guinea, Belau, Northern Marianas, Indonesia (Java), Thailand (new record).
description: Published as part of Sasakawa, Mitsuhiro, 2010, A review of the Oriental Japanagromyza Sasakawa (Diptera: Agromyzidae), with descriptions of four new species, pp. 16-32 in Zootaxa 2485 on pages 24-25, DOI: 10.5281/zenodo.195513
Japanagromyza hymenoedemia Sasakawa, 2010, sp. nov.
Creators: name: Sasakawa, Mitsuhiro
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Agromyzidae
Japanagromyza
Japanagromyza hymenoedemia

description: Japanagromyza hymenoedemia sp. nov.
(Figs. 14–16)
Diagnosis. This species is characterized by a yellow halter, densely spinose surstylus, short hypophallus and two tubules of the distiphallus surrounded with large membranous lobes.
Description. MALE. Black; parafrontalia weakly shiny; antenna brownish black; thorax very slightly grayish pollinose, mesoscutum shiny; abdomen strongly shiny. Wing with calypter brownish gray, black along margin, and fringe brown; halter yellow, stalk slightly brown tinged.
Frons only a little wider than eye; basal swellings of ors sockets projecting above eye in profile; lunule lower than semicircle; eye 1.3 times as high as broad; gena 1/10 as high as eye. First antennal flagellomere as long as wide (arista missing).
Mesoscutum with eight rows of acr, prsc weaker than anterior dc. Wing 1.9–2.0 (2.0 in holotype) mm long, costa with three sections in proportion of 3.4: 1: 0.7, r-m at basal 1/3 of discal cell (5: 9–10), ultimate section of M1 2.5–2.7 times as long as penultimate, ultimate section of CuA1 3/4 as long as penultimate. Fore tibia with lateral bristle.
S6 about 1.5 times as wide as long, four times as long as S5. Genitalia: Epandrium slightly projected on median dorso-caudal apex; surstylus long, broadly lobate in lateral view, bearing 18–20 spines; cercus almost as high as epandrium; hypandrium very long (800 µm), Y-shaped, with basal apodeme slightly shorter than sidepiece; phallapodeme 750 µm long; phallus 440 µm long, basiphallus bifurcate distally, hypophallus short, narrow, and weakly chitinized, distiphallus with distal tubules curved dorsally at ends and surrounded by membranous lobes which are covered with transparent spinules on dorsal side; ejaculatory apodeme 260 µm long, blade narrow (60 µm wide).
Body length 2.0– 2.2 mm.
FEMALE and HOSTPLANT. Unknown.
Type Material. Holotype 3 (BPBM 17133), Thailand, Chiangdao, 5–11. IV. 1958, T.C. Maa. Paratype: 3, Thailand, Chiangmai Prov., Fang, 12–19. IV. 1958, Maa (BPBM).
Distribution. Thailand.
Remarks. This species is similar to Japanagromyza stylata Sasakawa in the coloration, wing venation, and phallic structure, but differs distinctly from it in the shape and spinosity of the surstylus, and the distiphallic structure: in stylata, the surstylus is long cylindrical, bearing only two apical spines, and the distiphallic tubules are naked (Sasakawa 1963a: Fig. 5).
Etymology. The specific name refers to the membranous tumour (Greek: hymenion + oidema) of the distiphallus.
description: Published as part of Sasakawa, Mitsuhiro, 2010, A review of the Oriental Japanagromyza Sasakawa (Diptera: Agromyzidae), with descriptions of four new species, pp. 16-32 in Zootaxa 2485 on page 25, DOI: 10.5281/zenodo.195513
Japanagromyza involuta Spencer
Creators: name: Sasakawa, Mitsuhiro
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Agromyzidae
Japanagromyza
Japanagromyza involuta

description: Japanagromyza involuta Spencer
Japanagromyza involuta Spencer, 1977b: 357.
Remarks. This species is notable for its all reclinate or, hairy anterior margin of the frontalia, linear gena (less then 1/30 as high as eye), and two-bulbiform distiphallus. A male examined was easily identical with involuta by the original description, especially in structure of the male genitalia. However, some different points from the original description were found in the external and internal structures: knob of the halter is narrowly yellow in inner side (not entirely black in the original description), oh are minute, reclinate, and arranged in row (not lacking), acr are arranged in eight rows (not six rows), fore-tibial lateral bristle is present (not absent); ejaculatory apodeme is slightly longer than the phallus (not twice as long as phallus).
The male genitalia are described in detail: epandrium with a pair of strongly chitinized sclerites in inner side just above incurved surstylus; cercus about 1.5 times as high as epandrium, with five strong setae in antero-ventral part; hypandrium 600 µm long, with basal apodeme only 1/6 as long as sidepiece; phallapodeme 950 µm long; phallus 520 µm long; basiphallus weakly sclerotized, left sclerite of which with spinulose membranous process; hypophallus short; distiphallus bulbous, ventrally with spinulose lobes; ejaculatory apodeme 550 µm long, blade expanded asymmetrically, 340 µm wide (phallus: see Spencer 1977b: Figs. 30, 31).
Material examined. 3, Malaysia, Sabah, 19 km N of Kalabakan, Forest Camp, 60 m, 14. X. 1962, K.J. Kuncheria (BPBM).
Distribution. Papua New Guinea, Malaysia (new record).
description: Published as part of Sasakawa, Mitsuhiro, 2010, A review of the Oriental Japanagromyza Sasakawa (Diptera: Agromyzidae), with descriptions of four new species, pp. 16-32 in Zootaxa 2485 on pages 26-28, DOI: 10.5281/zenodo.195513