Cheumatopsyche rantsoa Olah & Johanson 2008, new species
Creators: name: Oláh, J.
name: Johanson, K. A.
Publisher: name: Zenodo
Year: 2008
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Hydropsychidae
Cheumatopsyche
Cheumatopsyche rantsoa

description: Cheumatopsyche rantsoa Oláh & Johanson, new species   
Fig. 375–379  
 This species is similar to C. masiposa, new species from which it is distinguished by the more drawn bowshaped anterior margins of segment IX; segment IX being more strongly shifted posterad, especially on dorsum; the more deeply excised anterodorsally in lateral view; the perfectly rounded apex of segment X; the basal half of the coxopodites being thicker; and the harpagones with narrower basis. The most pronounced distinguishing character is the darker forewing pattern, compared to the uniformly coloured forewings of C. masiposa.   
Male. Body brown, forewing membrane pale brown with dark pubescence. Maxilary palp formula I-(III, IV)-II-V, segment V as long as sum of segments I–IV. Head dorsum brown with brown setal warts. Swollen setal wart absent on proepisternum. Pretarsal claws asymmetrical, laterally flanked by setal bundle. Protibiae with 2 apical spurs, posteroapical spur small.  
Wings. Forewing (Fig. 375) crossveins m-cu and cu tangential. Cu2 and A1 running close before meeting at wing margin. Fork 1 present. Wing membrane pale brown with pronounced pattern; apical area dark, dark patches on crossveins s, s-m, m, and along tangential area of crossveins m-cu and cu. Forewing length 9.0 mm.  
Male genitalia (376–379). Abdominal segment IX fused annularly, tergum shorter than sternum (Fig. 376). Anterior margins of segment IX almost regular bow-shaped with long straight middle section (Fig. 376); dorsal section strongly drawn in lateral aspect. Apical lobe on posterolateral margins short, nearly triangular (Fig. 376); located above articulation cavity of gonocoxites. Antecosta well developed, widening before margin, with well visible external groove of antecostal suture; vestigial acrotergite visible. Spine row on posterior margins of segment IX complete (Fig. 376); spines heterogenous; irregularly bending on laterocaudal lobes of dorsum; spines on border between segments IX and X about half as long as spines on posterior margins; spines on apical lobes about one-third as long as spines on posterior margins; stiff, straight. Intersegmental step between segments IX and X filled and occupied by pair of setose protuberances; forming setose, laterocaudal lobes. Body of segment X, long, nearly triangular, with dorsad curving, blunt, apex in lateral aspect (Fig. 376); simple, rounded, long lobe-shaped in dorsal aspect (Fig. 377); ventrum not bellied. Apicoventral setal lobes slightly curving dorsad (Fig. 376), mesally fused with segment X, dominating distal part of segment X; delineated by setae; rounded; dorsum flat in dorsal view (Fig. 377). Smooth, mesal lobes reduced; dorsal interlobular gap small, narrow, shallowly half-circular (Fig. 377). Apicoventral lobes separated by dorsal, interlobular gap; lobes forming flaps with straight-cut dorsum in dorsal view (Fig. 377); shape highly dependent on observation angle. Unique setose structures present at basis of dorsolateral areas of segment X forming pair of well-developed protuberances representing laterocaudal lobes derived from dorsal apex of spine row on posterior margin of segment IX. Transverse sutures not visible. Longitudinal sutures running along ventral margin of segment (Fig. 376); forming continuation of apicoventral setose lobes. Setose areas (superior or preanal appendages) forming pair of elevated, circular, flap-like warts located centrally on segment X. Coxopodites exceeding apex of segment X, basally slender; distally broadening and slightly bending downwards from mid-length. Harpagones with short, broad triangular basis (Fig. 376); continuing into long, dorsad curving, thin, digitate process; cickle-shaped in ventral view (Fig. 378). Phallothecal phallobase enlarged (Fig. 379), straight, horizontal shaft parallel-sided, slightly constricting at mid-length. Endophallus long, not filling entire phallotheca. Chitinezed endothecal process circular, almost as wide as shaft. Phallotremmal sclerites broad, rounded and sinuate in lateral aspect.  
 Holotype male: MADAGASCAR: Fianarantsoa Province, 7 km W Ranomafana, 900 m, black light, mountain rainforest near river and stream, 18.x.1990 [W.E. Steiner, C. Kremen & P. Daniels] - 2 males (NMNH, pinned).   
Paratypes: same data as holotype — 1 male (NMNH), 1male (OPC); same data as holotype, except 8– 13.iii.1990 [W.E. Steiner] - 1 male (NMNH); ditto, except 20–24.iii.1990 - 1 male (NMNH); ditto, except - 1 male (NMNH); ditto, except - 1 male (NMNH).  
 Distribution. Madagascar.   
 Etymology. rantsoa, name derived from the type locality, Fianarantsoa Province.
description: Published as part of Oláh, J. & Johanson, K. A., 2008, Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae), pp. 1-248 in Zootaxa 1802 on pages 223-225