Clivina passimpunctata Balkenohl, 2025, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina passimpunctata

description: Clivina passimpunctata sp. nov.
(Figs 20, 55, 89)
Type material. Holotype: ♂, with labels and data: white, black printed “ Lope de Vega, Samar, Central Visayas JANUARY 2020 ” / backside handwritten in black ink “reg. coll. leg.” / white, black printed “ Coll. Balkenohl ” (CMBB).
Remark. The right front leg is missing as well as the left tibia onwards.
Description. Measurements. Body length 5.35 mm, width 1.51 mm, ratio L/W of pronotum 0.94, ratio L/W of elytra 1.91. Colour glossy, piceous. Supraantennal plate at margin and legs fuscous. Antennae and mouthparts somewhat more pale.
Head about a sixth narrower than pronotum. Clypeus with central part straight, with indistinctly projecting rounded teeth laterally, separated from clypeal-wing by obtuse-angled notch; wing slightly wider than teeth of clypeus, rounded, not as projecting as clypeal-teeth, separated from supraantennal plate by nearly rectangular notch, supraantennal plate with margin laterally convex, clypeus, wing, and anterior part of supraantennal plate distinctly reflexed margined. Supraantennal plate extended posteriorly into blunt keel, separated from wing by narrow furrow. Clypeus continuing into frons without clypeal-furrow. Clypeus slightly convex, frons moderately convex, with small indistinct flat pore-like depression at middle, separated from supraantennal plate by moderately wide furrow, with clypeal seta in front; supraorbital carinae blunt, with second smaller supraantennal carina, with two supraorbital setigerous punctures at middle of eye level and at posterior gena-level. Supraantennal plates, clypeus, and frons densely covered with scattered small punctures. Neck constriction closed at middle, developed as band of medium sized punctures and slight step. Eye prominent, gena small but distinct, enclosing eye posteriorly by about one quarter. Antenna elongate, reaching up to base of elytron. Labrum excised anteriorly, seemingly bilobed, finely isodiametrically reticulated, seven-setose. Mentum with lobes flat, convex laterally, with apex acute, isodiametrically reticulated, distinctly margined, median tooth flat, as high as lobe.
Pronotum (Fig. 55) with disk flat in lateral view, moderately and regularly convex in frontal view. Shape sub-cordate to square, indistinctly wider than long. Reflexed lateral margin sub-crenulate (120-fold), slightly convex at middle, widest at middle; anterior angle distinct, posterior angle distinct, indicated by distinct anteriorly projecting tooth; lateral channel moderately narrow, becoming wider posteriorly, reticulated, continuing up to base as sharp, convex margin. Median line complete, engraved. Anterior transverse line narrow, joining median line. Disk densely covered with small scattered punctures, with flat transverse wrinkles laterally, with isodiametric reticulation at base, with posterior setigerous puncture removed from margin by two diameters of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 30°. Basal channel on flange distinctly widened, rugose, with sharp carina.
Elytron (Fig. 20) with disk flattened in anterior half in lateral view, distinctly convex in frontal view. Shape of elytra elongate, nearly two times longer than wide, with lateral margin almost straight in anterior two thirds but slightly diverging, maximum width posterior middle. Humerus distinct, rounded, base somewhat retracted at middle. Reflexed lateral margin crenulate in anterior quarter. Row of setigerous punctures in the lateral channel of equally distance, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity falling perpendicularly to pedunculus, with isodiametric reticulation. Striae not very deep, impunctate, irregularly reticulated, one to four free at base, five and six joining at humerus, one to four ending free at apex, five and six joining apically. Intervals slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven and eight carinate at apex. Surface of intervals six to eight with isodiametric reticulation, interval one to five densely covered with small scattered punctures.
Hind wing fully developed.
Lower surface. Proepisternum isodiametrically reticulated, with small scattered punctures, with few transverse wrinkles laterally. Sternite of abdomen with isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with moderately projection posteriorly at middle, with narrow marginal furrow at middle.
Legs. Front tibia and front tarsomeres unknown. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of intermediate and hind legs of same width.
Male genitalia (Fig. 89). Median lobe, distinctly arcuate in basal half, indistinctly arcuate in apical half, apex regularly arcuate to tip, in lateral view apically with duckbill-like spatula. Parameres distinctly arcuate, with distinct apophyses, with small seta at apex, each.
Female gonocoxites and epipleurite unknown.
Variation unknown.
Distribution. Known from Lope de Vega on the island of Samar, Philippines.
Diagnosis. A medium sized piceous species with sub-cordate to square pronotum, intervals six to eight with isodiametric reticulation. In addition, interval one to five, head, pronotum, and proepisternum is densely covered with scattered small punctures. It is the only species of the group with this puncture pattern on the whole dorsal surface of the body.
Etymology. The name refers to the intense scattered punctures on the head, pronotum and elytron (Latin: passim = scattered; punctatus = with punctures).
description: Published as part of Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), pp. 1-70 in Zootaxa 5592 (1) on pages 34-35, DOI: 10.11646/zootaxa.5592.1.1, http://zenodo.org/record/14953674