Mada hirsuta Szawaryn & González, 2017, sp. nov.
Creators: name: Szawaryn, Karol
name: González, Guillermo
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Coccinellidae
Mada
Mada hirsuta

description: Mada hirsuta sp. nov.
(Figs. 1– 3)
Etymology. The species name is derived from its distinctive elytral pubescence. Diagnosis. This species can be quite easily distiguished from other Mada species by the surface of elytra covered with long setae irregularly arranged in whirling pattern (Fig. 1 A). Moreover it has protibiae distinctively broadened (Fig. 2 G), male genitalia with penis guide narrow, pointed, shorter than parameres, and female genitalia with coxites much longer than wide, narrow at base and broadened at posterior part.
Description. Length 3.9–4.3 mm; width 3.4–3.7 mm; TL/EW = 1.14–1.16; PL/PW = 0.50–0.51; PL/EL = 0.30– 0.31; PW/EW = 0.57–0.61; EL/EW = 0.95–1.00.
Body oval, convex (Figs. 1 A, 1B); pronotum and elytra covered with long setae. Head and mouthparts yellowishbrown (Fig. 1 C); pronotum yellowish-brown with anterior angles more pale, disc with five dark spots, one at base and four in a curved row across middle (Fig. 1 C); elytra yellowish-brown with lateral margin pale yellow (Fig. 1 B). Ventral surface light-brown. Head and pronotum punctate; elytron dually punctate.
Head exposed, transverse; ventral antennal grooves present. Eye finely faceted, with inner orbits emarginate anteromedially (Fig. 1 C). Antenna shorter than width of head, 11-segmented (Fig. 2 D); scape large, swollen, about twice as long as pedicel; pedicel longer than wide, swollen; antennomere 3 elongate, shorter than antennomeres 4–6 combined; antennomeres 4–7 elongate, antennomere 8 transverse; antennal club relatively compact, 3-segmented, asymmetrical. Clypeus transverse; labrum transverse with anterior part membranous, covered with long setae (Fig. 2 F). Mandibles multidentate, with large teeth on inner edge, with several small denticles on dorsal surface (Fig. 2 E).
Pronotum transverse, widest at base and gradually narrowing anteriorly; anterior angles and base not bordered; disc convex, finely punctate (Fig. 1 C). Prothoracic hypomeron smooth; notosternal suture distinct; prosternal process with weak grooves along its lateral margins. Mesoventrite with anterior edge emarginate with raised border for entire length; mesoventral process smooth, as broad as coxal diameter; meso-metaventral articulation with suture straight. Scutellar shield small, triangular. Elytra at base broader than pronotum; with dual punctation; humeral angles well developed; lateral margins narrow, visible from above throughout (Fig. 1 A); elytral epipleuron complete, with distinct depressions for receiving apices of mid and hind femora, with submarginal carina complete; metaventrite with intercoxal process broadly bordered; metaventral postcoxal lines forming straight line with two rounded, setose projections, laterally complete, recurved; metaventrite with discrimen complete.
Legs short and stout, not protruding from outer margin of elytral epipleuron. Trochanters roundly produced, with cavities on their inner surfaces for receiving tip of tibiae in repose. Femora cylindrical with weak grooves throughout for receiving tibiae. Tibiae with apical spurs of formula 1-2-2, protibiae expanded with weak grooves throughout for receiving tarsi (Fig. 2 G), mid and hind tibiae on outer edge near apex with oblique carina; tarsal claw double with additional large, subtriangular basal tooth (Fig. 2 N).
Abdomen with five ventrites in males with tip of 6th ventrite visible (Fig. 2 B) and five ventrites in females (Fig. 2 A). Abdominal postcoxal lines recurved roundly and incomplete, almost reaching posterior margin of ventrite 1 (Figs. 2 A, B). In males apical margin of ventrite 5 truncate (Fig. 2 B); ventrite 6 emarginate medially; tergite VIII rounded. Apodeme of male sternum IX simple, thin, rod-like. Tergite X rounded at apex (Fig. 2 M). In females apical margin of ventrite 5 rounded (Fig. 2 A); sternite VIII rounded, entire, with membranous part in middle appearing divided into two parts (Fig. 2 C), tergite VIII rounded. Proctiger (TX) membranous at base, elongate, rounded apically (Fig. 2 L).
Male terminalia and genitalia (Figs. 2 H–J). Penis guide symmetrical, thin, with sides almost parallel, spindle shaped, widest at basal part in lateral view, with pointed apex; shorter than parameres. Parameres well developed, simple, setose apically. Penis base with capsule reduced. Penis simple, rod-like.
Female genitalia (Figs. 2 K, L). Coxites elongate, narrow at base, broadly widened at posterior part, styli not observed. Bursa copulatrix ending with oviduct, sperm duct diverges slightly below oviduct; sperm duct short, lightly sclerotized, simple; spermatheca small, membranous, vermiform, without clear nodulus and ramus; accessory gland thin, a bit longer than spermatheca.
Type material. Holotype, male, Ecuador (Orellana), Payamino Research Station 0°29'36.01''S 77°17'29.15''W 300m Tropical Rainforest Malaise trap, 30.VII-12.VII.2007 / coll. CPDT Gillet BMNH (E) 2007-65 (BMNH). Paratypes, ECUADOR: 1 female, Ecuador (Orellana), Payamino Research Station 0° 29' 36.01''S 77° 17' 29.15''W 300m Tropical Rainforest, malaise trap, 30.VII–12.VII.2007 / coll. CPDT Gillet BMNH (E) 2007-65 (BMNH); PERU: 1 female, Peru, Madre de Dios, CICRA Field Stn, garden. 12.56940°S 70.10100°W 260m, 25.X–1.XI.2010, MJ Endara, malaise trap, PER10-10-MAT-021/ SEMC1097096 KUNHM-ENT (MUSM).
Distribution. Ecuador, Peru (Fig. 3).
description: Published as part of Szawaryn, Karol & González, Guillermo, 2017, A new species of Mada Gordon from the Andes (Coleoptera: Coccinellidae: Epilachnini), pp. 442-446 in Zootaxa 4226 (3) on pages 442-445, DOI: 10.11646/zootaxa.4226.3.7, http://zenodo.org/record/264098