Leucogeorgia borealis Antić & Reip 2020, sp. nov.
Creators: name: Antić, Dragan Ž.
name: Reip, Hans S.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Julida
Julidae
Leucogeorgia
Leucogeorgia borealis

description: Leucogeorgia borealis sp. nov.
urn:lsid:zoobank.org:act: D8CFFCD0-8D33-42A0-B7FD-E8FDE92E99E2
Figs 16–18, 56–57
Diagnosis
This species belongs to the group of Leucogeorgia spp. without modified mouthparts and with teeth on the mesomeral claw (vs absent in L. golovatchi sp. nov., L. lobata sp. nov. and L. prometheus sp. nov.). Leucogeorgia borealis sp. nov. differs from L. abchasica, L. satunini and L. gioi sp. nov. by having a stout mesomeral claw (vs elongate and slender in L. abchasica, L. satunini and L. gioi sp. nov). Leucogeorgia borealis sp. nov. differs from L. oculata sp. nov. by having a strongly serrate mesomeral lamella without a posterior rise (vs lamella non-serrate and with a posterior rise in L. oculata sp. nov.), and in the absence both of pigmentation and ommatidia, and the presence of metazonal setae (vs a pigmented body, ommatidia present and metazonal setae absent in L. oculata sp. nov.).
Etymology
From the Latin ʻ boreas ʼ (= ʻnorthʼ), reflecting its having the northernmost species in the genus.Adjective.
Type material
Holotype
RUSSIA • ♂; Krasnodar Province, near Goryachiy Klyuch, NW karst area of Greater Caucasus, Fanagoriyskaya (= Bolshaya Fanagoriyskaya, = Psekupskaya) Cave; 44.47° N, 38.98° E; 3 Nov. 1943; J. Birstein leg.; ZMUM.
Paratypes
RUSSIA • 1 ♂, 1 ♀, 2 juvs; same collection data as for holotype but 8 Jul. 1961; S.I. Ljovuschkin leg.; ZMUM • 1 juv.; same collection data as for holotype but 19 May 1983; S.I. Golovatch leg.; ZMUM • 1 ♀; same collection data as for holotype but 20 Sep. 1968; ZMUM.
Description
SIZE AND NUMBER OF BODY RINGS. Holotype male 24 mm long, vertical diameter of largest body ring 1.5 mm, body with 43 podous rings + 1 apodous ring + telson. Paratype male 28 mm long, vertical diameter of largest body ring 1.55 mm, body with 49 podous rings + 2 apodous rings + telson. Paratype females 21 and 24 mm long, vertical diameter of largest body ring 1.4–1.5 mm, body with 37 or 44 podous rings + 1 apodous ring + telson.
COLOUR (Fig. 16). Yellowish white in alcohol.
HEAD (Figs 16B, 17C). Without ommatidia. Frontal setae absent. Labrum with three labral teeth, four supralabral setae and 17 labral setae in paratype male. Gnathochilarium with a rhomboid promentum; lamellae linguales with 4+4 setae, stipites with 3+3 distolateral and 5+6 medial setae. Antennae 1.9 mm long in smaller paratype female, their length ca 140% of vertical diameter of largest body ring. Lengths of antennomeres I–VIII (in mm): 0.13 (I), 0.45 (II), 0.36 (III), 0.30 (IV), 0.40 (V), 0.18 (VI), 0.06 (VII) and 0.02 (VIII). Length/width ratio of antennomeres I–VII: 0.8 (I), 3 (II), 2.4 (III), 2 (IV), 2.7 (V), 1 (VI) and 0.5 (VII). Antennomeres V and VI each with a terminal corolla of large sensilla basiconica bacilliformia; antennomere VII with a terminal corolla of small sensilla basiconica bacilliformia.
BODY RINGS (Fig. 16E). Entire metazonal area with longitudinal striations. Length of midbody setae ca 5% of vertical diameter of rings.
TELSON (Fig. 16C). Epiproct with a short and blunt preanal process, covered with dorsal and lateral setae. Paraprocts rounded, setose. Hypoproct without any modifications.
LEGS IN MALES. First pair of legs modified, hook-shaped (Figs 17 A–B, 18D–E), with three complete podomeres; coxa with two setae; prefemur with four setae; femur, postfemur and tibiotarsus coalesced; femur with two setae; postfemur with one seta; tibiotarsus with a prominent distal lobe (tarsal remnant). Tip slightly tuberculated. Postfemoral and tibial ventral pads well developed on anterior legs, then gradually disappearing towards posterior legs.
VENTRAL MARGIN OF MALE BODY RING 7 (Fig. 16D). Poorly developed, low, rounded in lateral view.
PENES (Fig. 18C). In form of a short trapezoid, apically with two small subtriangular lobes.
GONOPODS (Figs 17D, 18 A–B). Promere (p) long and slender, with a flagellum (f); apical part spatulate, with denticulate margins; basal half with two developed ridges. Mesomere (m) with a well-developed, robust and denticulate mesomeral claw (mc); mesomeral lamella (ml) convex in central part, distal margin serrate, posterior part finely fimbriate. Opisthomere (o) bipartite. Anterior branch of o with a solenomere (s), with a medium-sized tip, and a well-developed and fimbriate velum (v). Posterior branch
of o in form of a shield-like protective lamella (pl). Mesomere and opisthomere connected basally through an accessory membrane (am).
Distribution
Known only from its type locality (Fig. 57, blue triangle).
Remarks
This is the northwesternmost record of the entire genus.
description: Published as part of Antić, Dragan Ž. & Reip, Hans S., 2020, The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae), pp. 1-106 in European Journal of Taxonomy 713 on pages 28-32, DOI: 10.5852/ejt.2020.713, http://zenodo.org/record/4020752