Passalus (Pertinax) umbriensis Hincks, 1950: 1044
Creators: name: Bevilaqua, Marcus
name: Fonseca, Claudio Ruy Vasconcelos da
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Passallidae
Passalus
Passalus umbriensis

description: Passalus (Pertinax) umbriensis Hincks, 1950 (Figures 7 – 8)   
Reyes-Castillo and Amat-Garcia 1991: 504; Boucher 2015: 118; Miles 2017: 124, fig. 25; Bevilaqua and Fonseca 2019: 7, fig. 3e  
 Passalus (Passalus) umbriensis Hincks, 1950: 1044, fig. 3; Hincks and Dibb 1958: 17; Reyes-Castillo 1970: 204; Amat-García et al. 2004: 177; Fonseca and Reyes-Castillo, 2004: 7 Diagnosis. Small size species; convex body; anterior edge of the front with a notch in the middle whose edges rise reminiscent of secondary mediofrontal tubercles; inconspicuous inner tubercles; mentum with mediobasal area with punctuations and setae; pronotum with protruded and acute anterior angles; abundant pronotal punctuation laterally; almost transverse and deep mesosternal scars; non-delimited metasternal disc.   
 Redescription. Body (Figure 7): habitus: convex; size: small (> 20 mm in length). Head (Figure 8 (a,c)): Labrum: anterior border straight. Clypeus: hidden under the frons; with anterior angles under the lateral + mediofrontal tubercles. Anterior frontal edge: straight, with a notch in the median region whose edges are high resembling secondary mediofrontal tubercles. Secondary mediofrontal tubercles: absent. Laterofrontal + mediofrontal tubercles: conspicuous, acute, and projected forward. Mediofrontal area: caliciform, as long as wide (1.1x), flat and shallow; with punctuations scattered throughout the anterior region. Cephalic nodule: small, not well marked and rounded. Inner tubercles: small, inconspicuous, smaller than the laterofrontal + mediofrontal tubercles, of which they are separated, located midway between them and the apex of the central tubercle. Anterofrontal ridges: straight, smaller, and weaker than the posterofrontal ridges, practically absent. Posterofrontal ridges: low, weak, and slightly sinuous, beginning at the apex of the central tubercle. Laterofrontal areas: flat and smooth surface. Central tubercle: conical, low, with non-free apex. Lateroposterior tubercles: inconspicuous, separated from the central tubercle. Postfrontal area: flat, with smooth surface. Postfrontal groove: well marked, sinuous with a slight notch located medially. Lateropostfrontal areas: shallow, smooth, and glabrous surface. Epicranial sutures: poorly marked. Epicranial pits: shallow but very visible. Anterior angles of the head: well developed, with acute apexes, slightly smaller than the laterofrontal + mediofrontal tubercles. Canthus ocular: apexes rounded not reaching half eye. Antennas: trilamellate, with narrow lamellae, the distal being wider than the other two. Mouthparts (Figures 7 (b), 8(b)): Ligula: tridentate with middle tooth larger than the lateral teeth. Hypostomal process: wide, glabrous and separated from mentum. Mentum mediobasal area: with coarse punctuations and bristles scattered throughout the area, with a protruding anterior region with no notch in the middle. Mentum lateral lobes: rounded apexes. Mentum lateral scars: large, shallow, and rounded. Mandibles: incisor lobe with three well-formed teeth at apex; robust suprainternal teeth; inconspicuous infrabasal pits. Maxilla: lacinia bidentate at the apex. Prothorax. Pronotum (Figures 7 (a,c), 8(a – c)): Anterior edge: straight. Anterior angles: acute and fairly protracted. Marginal groove: well marked, deep and with coarse punctures throughout its length, without dilation at the apex; reaching four-fifths of pronotal width. Lateral fossae: small, well marked and softly deep; slightly rounded shape. Pronotal punctuations: coarse and fairly dense, diffused throughout pronotum, concentrated mainly above the lateral fossae. Prosternum (Figures 7 (b), 8(d)): Prepisternum: not dilated in the anterior region; finely pubescent in lateroposterior region. Prepimerum: glabrous. Prosternelum: rhomboid with narrow apex. Mesosternum (Figures 7 (b), 8(d)): smooth, shiny, and glabrous. Mesosternal scars: long, narrow, and deep; practically transversal; without punctuations or pubescence. Metasternum (Figures 7 (b), 8(e)): Metasternal disc: poorly delimited, but more dilated than the lateral region of the metasternum. Metasternal punctuations: group of punctures only in the lateroposterior region; setigerous punctuations in the anterior region. Metasternal pubescence: only in the anterior region quite sparsely. Metasternal lateral groove: narrow, thinner than mesotibiae, deep, without punctures or bristles. Elytra (Figure 7 (a)): Approximately 2.6x longer and 0.9x wider than pronotum. Striae: narrower than the interstriae, marked with round and fine punctures, well defined in both the dorsal and lateral striae. Epipleura: glabrous. Humeri: with small and sparse bristles. Legs (Figures 7 (a,b), 8(e)): Profemur: ventral anterior border with well-marked groove, not reaching the apex of the profemur; ventral posterior border with few setae only near the apex. Protibiae: not dilated. Mesotibiae: one small spine on the outer face. Metatibiae: without spine. Abdomen (Figure 7 (b)): sternite VII with full groove and well marked; rough sides. Aedeagus (Figure 8 (f)): Median lobe wider than the parameres and the basal piece together and almost as long as these. In the ventral view with two plaques completely sclerotised with a recess in the basal region; basal piece and parameres fused forming a tegmen; parameres projections reaching less than one-third of the length of the median lobe; arched parameres anterior edge. In lateral view, the apex of the parameres is narrow and rounded. In dorsal view with widely separated parameres.   
 Dimensions. Total length: 21 mm; cephalic length: 2.1 mm; cephalic width: 4.8 mm; mediofrontal area length: 0.7 mm; mediofrontal area width: 1.9 mm; canthus ocular length: 0.6 mm; canthus ocular width: 0.3 mm; area of the mediofrontal area: 1 mm 2; mandibles external angle: 139º; antennal club length: 0.7 mm; antennal club width: 1.4 mm; distal lamella length: 0.4 mm; medial lamella length: 0.1 mm; width of the mentum at the lateral scars: 2.2 mm; mentum mediobasal area width: 1 mm; diameter of the mentum lateral scars: 0.4 mm; pronotal length: 4.7 mm; pronotal width: 6.4 mm; length of the pronotal anterior groove: 2 mm; elytral length: 12.4 mm; elytral width: 7.4 mm; humeral width: 6.1 mm; profemur length: 3.1 mm; length of the anterior ventral groove of the profemur: 2.4 mm; protibiae width: 0.8 mm; protarsal length: 2.4 mm; length of the last protarsomer: 0.9 mm; mesotibiae width: 0.5 mm; metasternal lateral groove width: 0.3 mm; metasternal disc length: 4.8 mm; abdomen length: 5.7 mm; aedeagus length: 2 mm; aedeagus width: 1.2 mm; median lobe length: 1.7 mm; paremeres projection lenght: 0.4 mm.   
 Material examined. Holotype (♂) F2439.49. COLOMBIA: Umbria/Guines Fluss/ Columbien//Ex Staudinger &/Bang Haas, Dresden.// Passalus Type/(Passalus)/ umbriensis Hincks /det. W.D. Hincks//Manchester Museum/ HOLOTYPE // ♂ dissect/S. Boucher det. 2012. F2439.49 (MMUE).   
 Remarks. Described in P. (Passalus) in the session ‘ Phoroneus ’, this species had its combination reviewed by Bevilaqua and Fonseca (2019) who allocated it within the subgenus P. (Pertinax).   
 Distribution. Colombia (Umbria) (locality type) and Brazil (Acre) Bevilaqua and Fonseca 2019.
description: Published as part of Bevilaqua, Marcus & Fonseca, Claudio Ruy Vasconcelos da, 2020, Redescription of the species of Passalus Fabricius, 1792 (Coleoptera: Passalidae) described by Walter Douglas Hincks (1906 - 1961) deposited in the Museum of the University of Manchester, pp. 321-350 in Journal of Natural History 54 (5 - 6) on pages 337-339, DOI: 10.1080/00222933.2020.1759721, http://zenodo.org/record/4609053