Manampoka atsimo Griswold & Wood & Carmichael 2012, SP. NOV.
Creators: name: Griswold, Charles E.
name: Wood, Hannah Marie
name: Carmichael, Anthea D.
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Phyxelididae
Manampoka
Manampoka atsimo

description: MANAMPOKA ATSIMO SP. NOV.  (FIGS 8A–G, 17A–C, 26A–I, 44A–C, 59)  
 Types: Holotype male and two paratype females (CASENT9031164) from primary montane rainforest at 670 m elevation in Parc National Andohahela, Parcelle I, Manangotry, Toliara Province, Madagascar, collected between 23 December 2008 and 3 January 2009 by Fernando Álvarez-Padilla and Hannah Wood, deposited in CASC.   
 Etymology: The specific name is from the Malagasy word for south; a noun in apposition.   
 Diagnosis: By the characters of the genus.   
 Male (Holotype): Total length 5.60. Markings as in Figure 8F, G. Carapace light orange-brown, margin of pars thoracica dusky, with broad, dusky, V-shaped mark anteriad of thoracic fovea and dusky longitudinal band on each side extending anteriorly along pars cephalica, with dusky reticulations laterad of eyes; black pigment surrounding each eye and extending between AM and between lateral eyes; chelicerae, sternum, labium, coxae, legs, and palpi pale yellowbrown; abdomen dark grey, dorsum with median light band and four pairs of light spots. Carapace 2.60 long, 2.00 wide; thoracic fovea 0.50 long, length 0.19 times that of carapace; clypeus 0.20 high; ocular area 0.35 long, 0.80 wide; ratio of eyes AM/AL/PM/PL, 1.46:2.61:1.07:1.00, diameter of PM 0.14. Chelicerae 1.00 long, slender, smooth, promargin of fang furrow with four small and three large teeth, retromargin with seven small teeth. Sternum 1.50 long, 1.10 wide; labium 0.50 long, 0.44 wide; palpal coxa 0.84 long, 0.40 wide. Legs elongate, leg I absent from specimen, femur II length 2.49 times carapace length; metatarsi II with prolateral concavity and clasping spine (Fig. 44A–C). Note: As all other Malagasy phyxelidids have at least leg I with a prolateral clasping concavity and spine, we assume that such a clasper occurs on metatarsus I of this species as well. Leg measurements (femur + patella + tibia + metatarsus + tarsus = [total]): I: [missing]; II: 6.25 + 1.10 + 6.00 + 5.70 + 2.40 = [21.45]; III: 5.10 + 1.00 + 4.35 + 4.70 + 1.90 = [17.05]; IV: 6.50 + 1.05 + 6.20 + 6.50 + 2.35 = [22.60]; palp: 1.65 + 0.55 + 0.70 + (absent) + 1.55 = [4.45]. Palpus as in Figures 17A–C and 26A–F; femur with five probasal thorn-like setae; tibia with dorsoapical blade (DTA) and retroapical lobe (RL); palpal bulb length 2 times width, embolus base retromedian, reservoir course in embolus base straight except initial loop, space between embolus base and proximal embolic curve 0.58 embolus base width; conductor one half bulb length, apex pointing retrolaterally; median apophysis a small, pointed cuspule, arising retromedially near embolus base.   
 Female (paratype): Total length 6.00. Markings as in Figure 8A–E, carapace orange-brown, shading to dusky laterally, pars cephalica shading anteriorly to red-brown; black pigment surrounding each eye and extending between lateral eyes; sternum, labium, and palpal coxa dusky red-brown; legs and palpi orangebrown; abdomen dark grey, dorsum with median longitudinal light band and four pair of light spots, venter with pair of faint longitudinal light bands. Carapace 2.85 long, 2.15 wide; thoracic fovea length 0.06 times that of carapace; clypeus 0.26 high; ocular area 0.34 long, 0.93 wide; ratio of eyes AM/AL/PM/PL, 1.25:3.16:1.00:1.00, diameter of PM 0.12. Chelicerae 1.20 long, robust, smooth, promargin of fang furrow with four small and two large teeth, retromargin with seven small teeth. Sternum 1.20 long, 1.18 wide; labium 0.56 long, 0.50 wide; palpal coxa 0.94 long, 0.40 wide. Palpal femur with three anterobasal thorns.   
Calamistrum origin at 0.45 from metatarsus base, length 0.55 times that of segment. Leg measurements (femur + patella + tibia + metatarsus + tarsus = [total]): I: 4.75 + 1.15 + 4.65 + 4.10 + 1.95 = [16.60]; II: 3.70 + 1.00 + 3.20 + 3.00 + 1.50 = [12.40]; III: 3.00 + 0.95 + 2.50 + 2.45 + 1.30 = [10.20]; IV: 4.10 + 1.00 + 3.65 + 3.50 + 1.15 = [13.40]; palp: 1.40 + 0.50 + 0.50 + (absent) + 1.40 = [4.05]. Epigynum as in Figure 26G, anterior margins of median lobe transverse, external copulatory openings large; vulva as in Figure 26H, I, with anterodorsally projecting capsule containing internal spiral chamber.  
 Variation (N = 6): Total length 5.40–7.00; carapace length 1.32–1.38 times width, height 0.38–0.49 times width; PER width 2.53–3.10 times OAL; OQP 1.11– 1.27 times OQA; clypeal height 1.67–2.17 times AM diameter; cheliceral length 4.58–5.40 times clypeal height, promargin of fang furrow with 6–7 teeth, retromargin of fang furrow with 5–7 teeth; sternum length 1.02–1.25 times width; femur I length 1.67– 1.91 times carapace length; metatarsus I length 1.44– 1.70 times carapace length.   
 Natural history: Recorded from primary montane rainforest.   
 Distribution: Known only from the type locality at Manangotry in Parc National Andohahela, far southern Madagascar (Fig. 59).   
 Material examined: MADAGASCAR: Toliara Prov., Parc National Andohahela, Parcelle I, Manangotry, off route 118, 34 km N Tolagnaro, 24°44′35.0″S, 46°51′23.3″E, elev. 670 m, 23 December 2008 – 3 January 2009, primary montane rainforest, general collecting day and night, F. Álvarez-Padilla and H. Wood, holotype ♂ and 2 ♀ paratypes, CASC (CASENT9031164), 10 ♀, CASC (CASENT9031168, 9031167, 9031165, 9031166, 9031167, 9031168).
description: Published as part of Griswold, Charles E., Wood, Hannah Marie & Carmichael, Anthea D., 2012, The lace web spiders (Araneae, Phyxelididae) of Madagascar: phylogeny, biogeography and taxonomy, pp. 728-810 in Zoological Journal of the Linnean Society 164 (4) on pages 797-798, DOI: 10.1111/j.1096-3642.2011.00779.x, http://zenodo.org/record/5407022