Myrovernix intricata Koekemoer 2019, comb. nov.
Creators: name: Koekemoer, Marinda
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Magnoliopsida
Asterales
Asteraceae
Myrovernix
Myrovernix intricata

description: Myrovernix intricata (Levyns) Koekemoer comb. nov.   
 Basionym:— Stoebe intricata Levyns (1937: 21).   
 Type:— SOUTH AFRICA. Western Cape. Prince Albert, Levyns 5034 (BOL!, holotype).   
 Taxonomic note:— Levyns (1937) described this species under Stoebe, because of the predominantly single-flowered capitula. All other characters (glandular leaves, shape and colour of disc florets, cypsela morphology and short-spined pollen grains), clearly places it in Myrovernix.   
 Intricately branched, bushy shrubs, up to ± 1 m tall and about the same in diam., occasionally wider when scrambling over rocks. Main branches up to 15 mm diam.; secondary and tertiary branches often perpendicular to main branches, stems densely or sparsely tomentose with a dense covering of stalked glands rising above the tomentum. Leaves glandular, mostly adpressed, oblong-lanceolate, up to 3 mm long, slightly gibbous near the apex, somewhat cobwebby or almost glabrous; leaf glands very different from those on stems, up to 0.5 mm in diam., widely spaced and wart-like, glands towards the leaf base much smaller, slender and stalked, very much like glands on stems; gland stalks very short, longer towards leaf bases. Capitula homogamous, discoid, 1-flowered (rarely 2-flowered), in small groups of 5–12 capitula, both lateral and terminal. Receptacle small, conical, bare. Involucral bracts in several rows, the outer bracts short, leafy and glandular, woolly or almost glabrous; the inner bracts brown, scarious, acute, equal to or slightly shorter than the florets. Disc florets ± 3 mm long, deep purple; lobes small, erect, triangular, reflexed at maturity. Cypselas smooth, with ± 10 inconspicuous ribs. Pappus setae ± 20, slightly joined at the base, as long as the corolla. Flowering March to May.   
 Diagnostic features: —Leaf glands wart-like, widely spaced; innermost involucre bract tips acuminate, capitula 1-flowered.   
 Conservation status: —Least Concern (http://Redlist.sanbi.org).   
 Distribution and habitat: — Western Cape. Rocky mountain slopes. Elevations 700–1 100 m. Figure 13.   
 Specimens examined:— SOUTH AFRICA, Western Cape. Ceres Div., Schurweberg, N of Bokkeveld Tafelberg, 19 April 1954, Esterhuysen 22884 (PRE); Waaihoek Mts., Chavonnes Mt, 31 March 1934, Galpin 12744 (PRE); Nieuwoudtville Plateau (3119AC), 19 March 1948, Acocks 14165 (PRE); Nieuwoudville plateau (3119AC), 4 April 1974, Goldblatt 1327 (K, NBG, PRE); Calvinia, W of Nieuwoudtville (3119AC), 3 June 1980, Goldblatt 5518 (PRE); Viewpoint above Van Rhyns Pass (3119AC), 11 September 1990, Koekemoer 423 (PRE); Clanwilliam, Pakhuis Pass, below Kleinkliphuis (3218BB), 7 March 1991, Koekemoer 582 (PRE); Clanwilliam, Amon jeeptrack, N Cederberg, TR048 (3218BB), 8 May 1989, Taylor 12042 (NBG); Clanwilliam, Amon jeeptrack, N Cederberg, TR005 (3218BB), 8 May 1989, Taylor 12045 (NBG); Clanwilliam, Amon jeeptrack, N Cederberg, TR004 (3218BB), 8 May 1989, Taylor 12047 (NBG); Clanwilliam, Amon jeeptrack, N Cederberg, TR006 (3218BB), 8 May 1989, Taylor 12051 (NBG); N Cederberg, path Pakhuisvlakte to Dwarsrivier, between Taaiboskraal River and Kliprivier (3218BB), 9 May 1989, Taylor 12061 (NBG, PRE); Clanwilliam, Grey’s Pass (3218DB), 19April 1937, Compton 6766 (NBG); Dist. Piquetberg, Greys Pass (Piquenierskloof) (3218DB), 10 May 1934, Galpin 12917 (PRE); Grey’s Pass, Clanwilliam (3218DB), May 1934, Galpin s.n. (BOL); Clanwilliam, Oupoort, NW of Pakhuis Pass on rd. to Botterkloof (3219AA), 24 April 2004, Bergh 1339 (NBG); Cederberg, E side of Pakhuis Pass (3219AA), 21 March 1996, Koekemoer 1223 (PRE); Cederberg, Uitkyk Pass, Algeria Forest Station (3219AA), 22 March 1996, Koekemoer 1232 (PRE); Lower slopes of Pakhuis Pass (3219AA), 11 September 1990, Koekemoer 428 (PRE); Cederberg, between Dwars River & Uitkyk Pass (3219AA), 4 April 1952, Levyns 9823 (BOL); Cederberg, Pakhuisberge, from Clanwilliam on rd. to Pakhuis (3219AA), 25 August 1995, Rodriguez-Oubina 2077 (PRE); Clanwilliam, Amon jeeptrack, N Cederberg, near Crossroads Camp at TR053 (3219AA), 9 May 1989, Taylor 12058 (NBG); Wupperthal, Pakhuis-Heuningvlei jeeptrack, N Cederberg above ‘Groot Rondevlak’ at TR008 (3219AA), 10 May 1989, Taylor 12067 (NBG); Wupperthal, Groot Rondevlak at Fortyn-se-kloof at TR064 (3219AA), 10 May 1989, Taylor 12069 (NBG); Wupperthal, Pakhuis-Heuningvlei jeeptrack, N Cederberg (3219AA), 10 May 1989, Taylor 12071 (NBG); Cederberg, Algeria, Rooikop, W of Algeria Forestry Station (3219AC), February 2004, Bergh 1182 (NBG); Cederberg, path up to Helshoogte (Helskloof?) avove Algeria campsite (3219AC), 26 February 2000, Bergh 141 (NBG); Cederberg, Algeria, N of Algeria Forestry Station, Helskloof (3219AC), 26 February 2000, Bergh 145 (NBG); Cederberg, Cederberg Pass, on the way to Kromrivier (3219AC), 26 February 2000, Bergh 173 (NBG); Clanwilliam Div., S Cedarberg, Sneeuberg (3219AC), 12 April 1952, Esterhuysen 20058 (PRE); Cederberg, near jeep track to Sneeukop hut, above Crystal Pools (3219AC), 12 September 2002, Koekemoer 2415 (PRE); Algeria Forest Station, near camping site along footpath to waterfall (3219AC), 7 March 1991, Koekemoer 592 (PRE); Gonnafontein, Cederberg, Plot 2, Pond’s (3219AC), 30 March 2000, Pond 38 (NBG); Clanwilliam Div., S Cederberg (3219AC), April 1947, Stokoe SAM63073 (SAM); Clanwilliam, Bath (3219CA), 6 July 1935, Compton 5338 (NBG); Clanwilliam Div. Olifantsrivier Mt., below Olifantsrivier Dome (3219CA), 16 April 1949, Esterhuysen 15305 (BOL, PRE); Elandskloof, Cold Bokkeveld (3219CA), 20 April 1935, Levyns 4888 & 4900 & 4906 (BOL); Cederberg, Matjiesrivier Nature Reserve, Stadsaal (3219CB), 4 October 1997, Lechmere-Oertel 752 (NBG); Wupperthal, Gideonskop (3219CB), 22 April 1957, Stokoe PRE42412 (NBG, PRE); Gideonskop (3219CB), 22 April 1957, Stokoe s.n. (NBG); Ceres Div., rocky ridge above Winkelhaaks River, E of Bokkeveld Sneeuwkop (3219CC), 20 April 1946, Esterhuysen 12687 (BOL, PRE); Clanwilliam near Thee River Kloof (3219CC), 16 April 1949, Esterhuysen 15332 (BOL); Cold Bokkeveld Mount, Ceres Div., Schoongezicht Pk. (3219CC), 4 April 1953, Esterhuysen 21290 (PRE); Ceres, Schurweberg (N of Bokkeveld Tafelberg) (3219CC), 19 April 1954, Esterhuysen 22885 (BOL, K, PRE); Ceres Div., Schurweberg (3219CC), 19 April 1946, Esterhuysen 23334 (BOL); Citrusdal, Platkloof (3219CC), 18 April 1998, Hanekom 3020 (NBG); Ceres, Cold Bokkeveld, Skoongezicht (3219CC), 8 April 1967, Hanekom 877 (PRE); Ceres, rd. NE from Op-Die-Berg, past Boplaas (3219CD), 24 April 2004, Bergh 1313 (NBG); Ceres Div., Winkelhaaks River, E of Bokkeveld Sneeukop (3219DC), 20 April 1946, Esterhuysen 12688 (BOL); rd. between R303 and Witzenberg Valley (3319AB), 21 December 2001, Bayer 01139 (PRE); Gydo Pass (3319AB), March 1951, Stokoe SAM 65885 (PRE); Tulbagh, above bridge in Nuwekloof, from Tulbagh on rd. to Gouda (3319AC), 23 March 1996, Koekemoer 1239 a & b (PRE); Ceres Div., Swartruggens, NE of Onverwacht (3319BA), 16 June 1965, Acocks 23662 (PRE); Ceres, Karoopoort (3319BA), 10 December 2003, Bergh 1067 (NBG); Ceres Karoo, Karoopoort, (3319BA), 10 December 2003, Bergh 1068 (NBG); Ceres, along R46/ R355 Kareekloof Private Nature Reserve, Kleinfontein farm (3319BA), 16 July 2003, Murray HMM 117 (NBG); Worcester Div., Keeromsberg (3319DA), 18 May 1944, Esterhuysen 10158 (BOL); Keeromsberg, Robertson-Worcester Div. (3319DA), 18 April 1954, Esterhuysen 22855 (PRE); Poesjenels River, Robertson (3319DC), 12 April 1936, Levyns 5512 (BOL); Montagu, S end of Kogmanskloof (3320CC), 14 March 1959, Acocks 20348 (K, PRE) and 27 June 2000, Bergh 529 (NBG); Montagu, between town & Baths (3320CC), 5 July 1938, Levyns 6456 (BOL); Montagu (3320CC), 1 May 1951, Levyns 9577 (BOL); S of Montagu Bath (3320CC), October 1922, Michell 124 (PRE); Calitzdorp, Gamka Reserve (3321BB), 24 May 1975, Esterhuysen 33832 (BOL); Gamkaskloof, Swartberg, (3321BD), 8 September 2001, van Wyk 359 (PRE); Riversdale, Langeberg, Garcia’s Pass (3321CC), 8 April 2004, Bergh 1277 (NBG); Garcias Pass (3321CC), 20 May 1950, Esterhuysen 17033 (BOL); Ladismith, Langeberg, Garcias Pass (3321CC), 13 May 1971, Kruger 1273 (NBG, PRE); Muiskraal, Riversdale (3321CC), 16 July 1957, Levyns 10742 (BOL); Muiskraal, Riversdale (3321CC), July 1927, Levyns 2122 (BOL); Little Karoo, Gamka Mountain Nature Reserve, upper reaches of Suiferwaterkloof (3321DB), 13 April 1988, Erasmus 156 (NBG, PRE); Oudtshoorn, Boomplaas, Cango Valley (3322AC), 21 January 1976, Hugo 26 (NBG, PRE); Swartberg Pass (3322AC), July 1935, Levyns 5034 (BOL); Prince Albert (3322AC), May 1926, Pocock 5180 (S. 180 in BOL) (BOL, PRE); Prins Albert Dist., Swartberg Pass (3322AC), April 1972, Reyneke & studente 143 (PRE); R.P.N.R. Suikerbosrug (3418AB), 8 May 2004, Duckitt 606 (NBG); R320 from Hermanus to Caledon (3419AD), 4 April 2004, Bester 5036 & 5054 (PRE); Robertson Div., Riviersonderendberge (3419BB), 7 April 1974, Theron & students 3020 (PRE); Clanwilliam Div., Uitkyk Pass (3219AA), 11 March 1938, Acocks 8597 (PRE); Cold Bokkeveld; Ceres Div.; Schurweberg (3219CC), 19 April 1946, Esterhuysen 12671 (PRE).
description: Published as part of Koekemoer, Marinda, 2019, Taxonomy and reclassification of South African Asteraceae genus Elytropappus (Gnaphalieae, Asteraceae), the description of two new genera and two new species, pp. 248-284 in Phytotaxa 403 (4) on page 276, DOI: 10.11646/phytotaxa.403.4.1, http://zenodo.org/record/13715872