Sclerocoelus tantus Kuwahara, Marshall & Paiero, 2025, sp. nov.
Creators: name: Kuwahara, Gregory K.
name: Marshall, Stephen A.
name: Paiero, Steven M.
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Sphaeroceridae
Sclerocoelus
Sclerocoelus tantus

description: Sclerocoelus tantus sp. nov.
urn:lsid:zoobank.org:act: A3F1AA4B-F179-484A-BF53-4B9206DEFE7D
Figs 9E, 136–138
Etymology
The species name (from the Latin ‘ tantus ’, meaning ‘so much, so many’) reflects the abundance of S. tantus sp. nov., specimens of which constitute almost 19% of the Neotropical Sclerocoelus specimens in the University of Guelph Insect Collection.
Material examined
Holotype
VENEZUELA • ♂; Mérida, Mérida, Los Chorros; 2100 m a.s.l.; 5 May 1988; S.A. Marshall leg.; sweep at trail head; DEBU.
Paratypes
ARGENTINA – Salta • 6 ♂♂; 22 km N of La Caldera, El Ucumar; 1150 m a.s.l.; 2–30 Dec. 1987; S. and J. Peck leg.; subtropical humid forest, Malaise flight-intercept trap; CNCI • 10 ♂♂, 4 ♀♀; La Caldera; 1500 m a.s.l.; 27 Feb. 1992; S.A. Marshall leg.; roadside forest sweep; DEBU.
BRAZIL – Rio de Janeiro • 15 ♂♂, 20 ♀♀; Teresopolis, trail to Pedra do Sino; 1300–1700 m a.s.l.; 25 Jan. 1990; S.A. Marshall leg.; DEBU • 15 ♂♂, 20 ♀♀; same data as for preceding; MNRJ.
ECUADOR – Napo • 2 ♀♀; Baeza, west along road; 1500 m a.s.l.; 16–19 May 1987; Coote and Brown leg.; montane rainforest/pasture, Malaise trap; ROME. – Pichincha • 8 ♂♂, 7 km SE of Nanegalito, trout farm ‘San Jose’; 1500 m a.s.l.; 27–30 Oct. 1999; S.A. Marshall leg.; riverine forest, treefall sweep; DEBU • 8 ♂♂, 5 ♀♀; same data as for preceding; QCAZ • 12 ♂♂, 5 ♀♀; Maquipucuna Biological Reserve; 0°07′34″ N, 78°27′57″ W; 1200 m a.s.l.; 27–28 Apr. 2002; M. Buck leg.; sweep; QCAZ • 7 ♂♂, 12 ♀♀; Maquipucuna Biological Reserve; 1200 m a.s.l.; 29 Oct. 1999; S.A. Marshall leg.; river trail, sweep; DEBU;
PARAGUAY – Caazapá • 1 ♂, 6 ♀♀; Hermosa, San Rafael Reserve, Lopez family property; 26°19′15″ S, 55°44′55″ W; 90 m a.s.l.; 3–6 Dec. 2000; Z.H. Falin leg.; flight-intercept trap; DEBU.
PERU – Cusco • 1 ♂; Cock-of-the-Rock Lodge; 13°03′21″ S, 71°32′46″ W; 1380 m a.s.l.; 18–20 Oct. 2006; J. Skevington leg.; Malaise trap; MUSM.
VENEZUELA – Aragua • 1 ♂; Colonia Tower; 2300 m a.s.l.; 10 Mar. 1995; S.A. Marshall leg.; sweep steep forest trail; DEBU. – Mérida • 12 ♂♂, 18 ♀♀; same data as for holotype; DEBU. – Trujillo • 7 ♂♂, 11 ♀♀; 10 km E of Bocono, Laguna de Lucerdo; 1760 m a.s.l.; 3 Mar. 1995; S.A. Marshall leg.; slash/compost; DEBU • 3 ♀♀; same data as for preceding; sweep; DEBU • 4 ♂♂, 1 ♀; Bocono, road to Guaramacal; 2130 m a.s.l.; 2 Mar. 1995; S.A. Marshall leg.; mud sweep; DEBU • 1 ♀; same data as for preceding; 2200 m a.s.l.; under dung; DEBU • 2 ♀♀; Bocono-Guaramacal Road; 2130 m a.s.l.; S.A. Marshall leg.; wet litter sweep; DEBU • 1 ♀; Bocono-Guaramacal Road; 3000 m a.s.l.; 3 Mar. 1995, S.A. Marshall leg.; wet debris along stream; DEBU.
Other material examined
ARGENTINA – Jujuy • 1 ♀; Calilegua National Park, El Cortadeval km 6; 800 m a.s.l.; 18–28 Dec. 1987; S. and J. Peck leg.; forest, Malaise trap; DEBU • 2 ♀♀; Calilegua National Park, Estación El Cero; 900 m a.s.l.; 18–28 Dec. 1987; S. and J. Peck leg.; forest, Malaise/flight-intercept trap; DEBU • 1 ♀; Calilegua National Park, Mirador; 600 m a.s.l.; 18–28 Dec. 1987; S. and J. Peck leg.; forest, Malaise/flight-intercept trap; DEBU. – Misiones • 1 ♀; 5 km E of Puerto Iguazú; 2–7 Feb. 1992; S.A. Marshall leg.; flight-intercept trap/pan/dung pans; DEBU • 2 ♂♂, 2 ♀♀; 5 km E of Puerto Iguazú; 3 Feb. 1992; S.A. Marshall leg.; sweep by small stream; DEBU • 1 ♂, 3 ♀♀; 5 km E of Puerto Iguazú; 1–6 Feb. 1992; S.A. Marshall leg.; sweep Malaise behind Hotel Orquidaes; DEBU. – Salta • 1 ♂, 6 ♀♀; 17 km N of La Caldera, Alto de la Sierra; 1550 m a.s.l.; 2–30 Dec. 1987; S. and J. Peck leg.; subtropical humid forest, Malaise/flight-intercept trap; DEBU • 131 ♂♂, 398 ♀♀; 22 km N of La Caldera, El Ucumar; 1550 m a.s.l.; 2–30 Dec. 1987; S. and J. Peck leg.; subtropical humid forest, Malaise/flight-intercept trap; DEBU • 1 ♀; 30 km E of Salta, Campo Quijano, El Alisal; 18–20 Feb. 1992; S.A. Marshall leg.; forest remnant, flight-intercept trap; DEBU • 4 ♂♂, 4 ♀♀; 30 km E of Salta, Campo Quijano, El Alisal; 18 Feb. 1992; S.A. Marshall leg.; forest vestige; DEBU • 21 ♂♂, 10 ♀♀; 30 km E of Salta, Campo Quijano, 20 Feb. 1992; S.A. Marshall leg.; forest vestige, sweep; DEBU • 55 ♂♂, 15 ♀♀; same data as for preceding; 24 Feb. 1992; wet debris along stream; DEBU • 17 ♂♂, 4 ♀♀; same data as for preceding; 20 Feb. 1992; stream debris, leaf litter; DEBU • 3 ♂♂, 1 ♀; same data as for preceding; sweep, dung; DEBU • 125 ♂♂, 81 ♀♀; 40 km N of Salta, Camino La Cornisa; 27 Feb. 1992; S.A. Marshall leg.; roadside, forest sweep; DEBU • 3 ♂♂; 50 km W of Chicoana, El Maray; 28 Feb. 1992; S.A. Marshall leg.; dry area, sweep pampas grass along creek; DEBU • 1 ♂, 1 ♀; Campo Quijano; 28 Feb. 1992; S.A. Marshall leg.; Malaise/flight-intercept trap; DEBU • 17 ♂♂, 8 ♀♀; Cañada La Gotera, 15 km W of Chicoana; 16–28 Feb. 1992; S.A. Marshall leg.; forest remnant; DEBU • 19 ♂♂, 19 ♀♀; same data as for preceding; 19 Feb. 1992; S.A. Marshall leg.; sweeping wet litter; DEBU • 1 ♀; Cañada La Gotera; 16–28 Feb. 1992; S.A. Marshall leg.; pan traps; DEBU • 1 ♂, 1 ♀; El Rey National Park, Pozo Verde Trail, km 5; 950 m a.s.l.; 5–15 Dec. 1987; S. and J. Peck leg.; Yungas forest, Malaise/flight-intercept trap; DEBU • 16 ♂♂, 39 ♀♀; El Rey National Park, Pozo Verde Trail, km 7; 1000 m a.s.l.; 5–15 Dec. 1987; S. and J. Peck leg.; Yungas forest, Malaise/flight-intercept trap; DEBU • 2 ♀♀; El Rey National Park, Rio La Sala; 900 m a.s.l.; 5–10 Dec. 1987; S. and J. Peck leg.; open stream side in forest, Malaise/flight-intercept trap; DEBU • 127 ♂♂, 87 ♀♀; La Caldera; 1500 m a.s.l.; 27 Feb. 1992; S.A. Marshall leg.; roadside, forest sweep; DEBU • 14 ♂♂, 20 ♀♀; La Caldera, Camino La Cornisa; 27 Feb. 1992; S.A. Marshall leg.; roadside sweep; DEBU • 3 ♀♀; Rosario De Lerma, INESALT yard; 16–28 Feb. 1992; S.A. Marshall leg.; Malaise trap; DEBU • 1 ♂, 1 ♀; Rosario De Lerma; 19 Feb. 1992; S.A. Marshall leg.; swept in hedgerow ditch; DEBU • 3 ♀♀; same data as for preceding; 24 Feb. 1992; swept over bulldozed road; DEBU • 4 ♀♀; same data as for preceding; swept over foul ditch; DEBU.
BOLIVIA – La Paz • 13 ♂♂, 9 ♀♀; 0.5 km SE of Coroico; 16°11′ S, 67°44′ W; 15 Apr. 2001; S.A. Marshall leg.; DEBU • 2 ♂♂, 2 ♀♀; 10 km NW of Caranavi, road to ENTEL tower; 15°46′35″ S, 67°35′48″ W; 1400 m a.s.l.; 13 Apr. 2001; S.A. Marshall leg.; DEBU • 2 ♂♂; 23 km NW of Caranavi; 14 Apr. 2001; S.A. Marshall leg.; roadside/stream, sweep; DEBU • 1 ♂; 38 km S of Guanay; 7–12 Apr. 2001; S.A. Marshall leg.; roadside stream, pan traps; DEBU • 26 ♂♂, 17 ♀♀; Chulumani, Apa-Apa Reserve; 16°21′15″ S, 67°30′21″ W; 2000 m a.s.l.; 1 Apr. 2001; S.A. Marshall leg.; sweep; DEBU • 1 ♀; San Antonio, 8 km S of Mapiri; 15°20′56″ S, 68°13′31″ W; 11 Apr. 2001 S.A. Marshall leg.; dung baits; DEBU. – Santa Cruz • 2 ♂♂, 2 ♀♀; Siringalito, near Pampa Grande; 2 Oct. 1996; Bettella and Rossi leg.; DEBU.
BRAZIL – Minas Gerais • 1 ♀; Prado; 21 Feb. 1990; S.A. Marshall leg.; sweep along river; MZSP. – Paraná • 1 ♂; Curitiba, 30 km SE of SEBR 277; 6 Feb. 1990; S.A. Marshall leg.; MZSP • 11 ♂♂, 7 ♀♀; Curitiba; 5 Feb. 1990; S.A. Marshall leg.; under fallen epiphyte in Mata near National History Museum; DEBU • 1 ♀; Curitiba, University campus; 2–5 Feb. 1990; S.A. Marshall leg.; wet areas, flight-intercept trap/pans; MZSP • 12 ♂♂, 9 ♀♀; Curitiba campus; 10 Feb. 1990; S.A. Marshall leg.; sweep in forest with Araucaria; MZSP • 3 ♂♂, 4 ♀♀; east of Curitiba, near Ponta Grossa; 800 m a.s.l.; 8 Feb. 1990; S.A. Marshall leg.; swept along road; MZSP • 28 ♂♂, 24 ♀♀; east of Curitiba; 8 Feb. 1990; S.A. Marshall leg.; swept near creek debris; DEBU • 3 ♀♀; Londrina, Parque Municipal Arthur Thomas; 1 Feb. 1990; S.A. Marshall leg.; sweep; MZSP • 4 ♂♂, 2 ♀♀; Londrina; 29 Jan. 1990; S.A. Marshall leg.; sweep; DEBU. – Rio de Janeiro • 14 ♂♂, 26 ♀♀; Muri, near Hotel Garlipp; 2 Mar. 1990; S.A. Marshall leg.; sweep over trail; MZSP • 25 ♂♂, 16 ♀♀; Nova Friburgo, Sitio Edelweiss; 26–27 Jan. 1990; S.A. Marshall leg.; Malaise trap; MZSP • 16 ♂♂, 15 ♀♀; Nova Friburgo, 10 km S of Sitio Edelweiss; 27 Jan. 1990; S.A. Marshall leg.; DEBU • 1 ♂, 2 ♀♀; Nova Friburgo, 10 km S of Sitio Edelweiss, Muri; 1–28 Feb. 1990; S.A. Marshall leg.; Malaise head; MZSP • 10 ♂♂, 7 ♀♀; Nova Friburgo; 27 Jan. 1990; S.A. Marshall leg.; along creek; MZSP • 13 ♂♂, 8 ♀♀; Teresópolis; 1200 m a.s.l.; 13 Mar. 1990; S.A. Marshall leg.; swept near river; MZSP • 47 ♂♂, 128 ♀♀; Teresópolis, trail to Pedra do Sino; 1300–1700 m a.s.l.; 25 Jan. 1990; S.A. Marshall leg.; sweep; DEBU • 47 ♂♂, 128 ♀♀; same data as for preceding; MZSP • 3 ♀♀; Teresópolis, trail to Pedra do Sino; 1500–2000 m a.s.l.; 14 Mar. 1998; S.A. Marshall leg.; DEBU. – São Paulo • 20 ♂♂, 10 ♀♀; São Paulo, Jaraguá; 8 Feb. 1990; S.A. Marshall leg.; MZSP.
ECUADOR – Napo • 5 ♀♀; Baeza; 1500 m a.s.l.; 16–19 May 1987; L.D. Coote and B.V. Brown leg.; wet montane rainforest, Malaise trap; DEBU • 1 ♀; Baeza; 1500–1700 m a.s.l.; 14 May 1987; L.D. Coote leg.; wet montane forest/pasture, sweeping; DEBU • 5 ♂♂, 4 ♀♀; Baeza; 1550 m a.s.l.; 15–19 May 1987; L.D. Coote and B.V. Brown leg.; wet montane rainforest/pasture, Malaise trap off stone trail; ROME • 3 ♂♂, 3 ♀♀; Baeza; 1700 m a.s.l.; 16–19 May 1987; L.D. Coote and B.V. Brown leg.; wet montane rainforest/pasture, near small creek, Malaise head; ROME • 1 ♂, 4 ♀♀; Baeza, W along road; 1500 m a.s.l.; 16–19 May 1987; L.D. Coote and B.V. Brown leg.; montane rainforest/pasture, Malaise trap; DEBU. – Pichincha • 8 ♂♂, 14 ♀♀; 3.5 km SE of Tandayapa; 28 Oct. 1999; S.A. Marshall leg.; in green leaf litter (treefall); DEBU • 1 ♀; 7 km SE of Nanegalito, trout farm ‘San Jose’; 1500 m a.s.l.; 30 Oct. 1999; S.A. Marshall leg.; DEBU • 2 ♀♀; same data as for preceding; 30–31 Oct. 1999; river edge, pan traps; DEBU • 46 ♂♂, 17 ♀♀; same data as for preceding; 27–30 Oct. 1999; riverine forest, sweep treefalls; DEBU • 3 ♂♂, 1 ♀; 7 km SE of Nanegalito, trout farm ‘San Jose’; 27 Oct. 1999; S.A. Marshall leg.; on logs/sweep; DEBU • 1 ♂; 8 km N of Nono, road to Mindo; 25 Oct. 1999; S.A. Marshall leg.; roadside, sweeping; DEBU • 1 ♂; 11.7 km SE of Tandayapa, road to Nono; 28 Oct. 1999; S.A. Marshall leg.; near stream, litter/wood; DEBU • 1 ♂; Bellavista Could Forest Reserve; 0°01′13″ S, 78°40′30″ W; 2200 m a.s.l.; 9–13 May 2009; S.A. Marshall leg.; DEBU • 2 ♀♀; Bellavista Reserve; 2200 m a.s.l.; 30 Oct. 1999; S.A. Marshall leg.; DEBU • 3 ♀♀; Bellavista Reserve, ridge trail; 28 Oct. 1999; S.A. Marshall leg.; sweeping/aspirating; DEBU • 4 ♂♂, 1 ♀; Bellavista Reserve, trail B; 30 Oct. 1999; S.A. Marshall leg.; sweeping; DEBU • 27 ♂♂, 8 ♀♀; Maquipucuna Biological Reserve; 0°07′34″ N, 78°37′57″ W; 1200 m a.s.l.; 27–28 Apr. 2002; M. Buck leg.; sweep; DEBU • 1 ♀; Maquipucuna Biological Reserve; 1200 m a.s.l.; 27 Oct. 1999; S.A. Marshall leg.; on leaves or small dung bait; DEBU • 29 ♂♂, 26 ♀♀; Maquipucuna Biological Reserve, main trail; 1300 m a.s.l.; 27 Oct. 1999; S.A. Marshall leg.; DEBU • 2 ♂♂, 1 ♀; Maquipucuna Biological Reserve, river trail; 1200 m a.s.l.; 27 Oct. 1999; S.A. Marshall leg.; DEBU • 54 ♂♂, 36 ♀♀; same data as for preceding; 29 Oct. 1999; sweeping; DEBU. – Zamora-Chinchipe • 4 ♂♂, 1 ♀; 8 km NW of Zamora, mouth of Río Sabanilla; 1420 m a.s.l.; 1 Nov. 1987; C. Young, R. Davidson and J. Rawlins leg.; CMNH • 1 ♂; San Francisco Biological Reserve, Río San Francisco trail; 3°58′30″ S, 79°04′25″ W; 2000 m a.s.l.; 25 Feb.–3 Mar. 2009; M. Pollet and A. De Braekeleer leg.; yellow pan traps; RBINS.
PARAGUAY – Caazapá • 5 ♂♂, 21 ♀♀; Hermosa, San Rafael Reserve, Lopez family property; 26°19′15″ S, 55°44′55″ W; 90 m a.s.l.; 3–6 Dec. 2000; Z.H. Falin leg.; flight-intercept trap; DEBU.
PERU – Cusco • 3 ♂♂; Cock-of-the-Rock Lodge; 13°03′21″ S, 71°32′46″ W; 1380 m a.s.l.; 18–20 Oct. 2006; J. Skevington leg.; Malaise trap; DEBU.
VENEZUELA – Aragua • 1 ♂; Colonia Tovar; 2300 m a.s.l.; 10 Mar. 1995; S.A. Marshall leg.; sweep steep forest trail; DEBU • 1 ♀; Henri Pittier National Park, Rancho Grande, Portachuelo Pass; 9 Apr. 1994; L. Masner leg.; DEBU. – Lara • 1 ♂, 27 ♀♀; Yacambú; 1200 m a.s.l.; 7 May 1981; H. Townes leg.; cloud forest; CNCI. – Trujillo • 8 ♂♂, 12 ♀♀; 10 km E of Bocono, Laguna de Lucerdo; 1760 m a.s.l.; 3 Feb. 1995; S.A. Marshall leg.; slash/compost; DEBU • 1 ♂, 4 ♀♀; same data as for preceding; sweep; DEBU • 5 ♂♂, 2 ♀♀; Bocono, road to Guaramacal; 2130 m a.s.l.; 2 Mar. 1995; S.A. Marshall leg.; sweep mud; DEBU • 2 ♀♀; same data as for preceding; 2200 m a.s.l.; under dung; DEBU • 1 ♂, 2 ♀♀; Bocono–Guaramacal Road; 2130 m a.s.l.; S.A. Marshall leg.; sweep wet litter; DEBU • 1 ♂; BoconoGuaramacal Road; 3000 m a.s.l.; 3 Mar. 1995; S.A. Marshall leg.; wet debris along stream; DEBU • 2 ♂♂; Guaramacal National Park; 2000–3000 m a.s.l.; 26 Aug.–1 Sep. 1992; L. Masner leg.; car net; DEBU • 14 ♂♂, 27 ♀♀; Guaramacal National Park, 14 km NE of Bocono; 2000 m a.s.l.; 25 Aug.–1 Sep. 1992; L. Masner leg.; DEBU • 8 ♂♂, 1 ♀; Mosquey near Bocono; 1500 m a.s.l.; 24 Aug. 1992; L. Masner leg.; coffee plantation, maxinet; DEBU.
Description
BODY (Fig. 136A). Length 2.3–3.8 mm. Head brown, lower quarter of frons orange; face, gena, and antennae orange-brown. Frontal width 2.3–2.4× frontal height. Three pairs of strong interfrontal bristles surmounting a very fine fourth pair; anterior orbital 0.6–0.7 × length of posterior. Palpus yellow. Eye large, greatest height about 3.5 × shortest genal height. Thorax brown, scutum with paler lateral edges. Two pairs of dorsocentral bristles (anterior pair only slightly larger than surrounding acrostichal setulae) separated by 7–8 rows of acrostichal setulae. Membrane around prosternum sometimes with a small, setula-bearing sclerite. Legs brown, foreleg slightly paler. Fore femur with three large ventral preapical setae. Ventral surface of male mid tibia with two rows of stout setae in apical third. Wing (Fig. 9E) slightly infuscate. CS2 0.7–0.8× CS3. Halter pale brown.
MALE ABDOMEN (Figs 136B–C, 137). Dark brown, posterior edges of tergites sometimes slightly desclerotized. T2–5 and S2–4 uniformly long-setose with large posterolateral setae. S5 almost quadrate, 2.3× length of S4 with anterior third underneath S4; posterolaterally sparsely long-setose, left side strongly reduced posterolaterally, with a dark, deflexed, trapezoidal medial patch of very dense setulae flanked by desclerotized areas. Anterior flange of S6+7 2.0 × as long as wide. Sclerite A small with short setae; sclerite B and sclerite C absent; sclerite D triangular, blade-like and attached to posteromedial lobe of S5; sclerite E long, tapered, and sinuate; sclerite F long and tapered, with a short anterobasal lobe; sclerite G very large, U-shaped; ring sclerite indistinct and slightly desclerotized. Epandrium large, 1.1× length of S8, height 1.2× maximum length and 0.6 × maximum width, uniformly setose with larger setae posteroventrally; posterior third of epandrium separated from anterior two-thirds by a deep lateral groove extending halfway up epandrium; anal fissure triangular, widest at base; perianal pads bulging slightly. Pseudocercus small, separate from epandrium and bearing 3 setae; halves of subepandrial sclerite reduced to a pair of arches fused medially forming an M-shape and articulating basally with subcercus. Subcercus with large inner part posteroventrally extended into a prominent process and a smaller process which forms a U-shaped connection between subcerci; outer part with a broadly rounded lobe articulating with surstylus and a hooked lobe contacting pseudocercus. Hypandrium with long, thin anteromedial apodeme. Surstylus stout, wider than deep, densely setose with a prominent anteroventral lobe and a bare inner lobe. Postgonite short, strongly curved anteriorly, tapered with apex blunt and notched and a long posteroventral lobe at about midlength. Phallapodeme very large with a tall dorsal ‘fin’; basiphallus stout with two large lateral plates, divergent basally and distally fused with trunk-like epiphallus; distiphallus largely membranous, reduced to a broad, distally bifurcate dorsal sclerite and a pair of small ventral sclerites.
FEMALE ABDOMEN (Fig. 138). T7 broad, posteromedially shortened; T8 divided into a small, pale dorsal sclerite and two dark lateral sclerites with elongate, acute posteroventral corner. Epiproct small, pale, and setulose. Cercus short and blunt with large apical and dorsal setae. S7 broad, rounded posteriorly with four large posterior setae; S8 reduced to two dark, ovoid lateral sclerites. Two spermathecae, bulb spherical and finely striate with a basal invagination.
Distribution
Neotropical: Argentina, Bolivia, Brazil, Ecuador, Paraguay, Peru, Venezuela.
Remarks
Sclerocoelus tantus sp. nov. is closely related to S. copiosus sp. nov. and S. elephas sp. nov., which also have a distinctive trunk-like epiphallus, but differs by the trapezoidal posteromedial setulose patch on the male S5, stout surstylus with a prominent anterodorsal lobe, rectangular female S7, and spherical spermatheca with a basal invagination. Sclerocoelus tantus is a frequently collected and widespread South American species often found with other species of the S. galapagensis group.
description: Published as part of Kuwahara, Gregory K., Marshall, Stephen A. & Paiero, Steven M., 2025, A revision of the genus Sclerocoelus Marshall (Diptera: Sphaeroceridae), pp. 1-277 in European Journal of Taxonomy 979 on pages 112-116, DOI: 10.5852/ejt.2025.979.2803, http://zenodo.org/record/14980852