Hemiteles japonicus Watanabe, 2021, sp. nov.
Creators: name: Watanabe, Kyohei
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Hemiteles
Hemiteles japonicus

description: Hemiteles japonicus sp. nov.
(New SJN: Nihon-mame-togari-himebachi)
(Figs 40A–E, 63H, 64C, 66J)
Etymology. The specific name is from Japan.
Type series. Holotype: KPM-NK 55021, F, JAPAN, Kagoshima Pref., Kimotsuke Town, Minamikata, Kanaedake, 18–20. VII. 2019, K. Watanabe leg. (YPT). Paratypes: JAPAN, KPM-NK 55022, F, Toyama Pref., Nanto City, Togamura, Kamimomose, 21–28. VII. 2009, M. Watanabe et al. leg. (MsT); KPM-NK 55023, F, Kochi Pref., Umaji Vil., 6. VIII. 2010, Y. Matsubara & K. Fukuda leg. (MsT).
Description. Female (n=3). Body length 4.6–5.6 (HT: 4.7) mm. Body matt and densely punctate and coriaceous, covered with silver setae.
Head. Clypeus 1.9 times as wide as maximum length, its anterior margin rounded. MSL 0.9–0.95 (HT: 0.9) times as long as BWM. Gena punctate and coriaceous. OOL shorter than POL. OOL and POL longer than OD. Occipital carina complete, its lower end joined with hypostomal carina distant from mandibular base. Upper tooth of mandible as long as lower tooth. Base of mandible flat. Antenna with 23–24 (HT: 24) flagellomeres. FL III 6.7 times as long as maximum depth in lateral view.
Mesosoma. Epomia distinct. Mesoscutum with some longitudinal striae near posterior end of notaulus. Notaulus long, fading our just behind of the center of mesoscutum. Scutellum without a lateral longitudinal carina except for distinct basal part. Mesopleuron longitudinally and obliquely striate just anterior to and below speculum, with coriaceous speculum. Posterior transverse carina of mesosternum largely absent in front of mid coxae. Metapleuron obliquely striate medially, without a juxtacoxal carina. Anterior transverse carina and posterior transverse carina of propodeum complete (Figs 40E, 64C). Lateral longitudinal carina and lateromedian longitudinal carina of propodeum absent (Fig. 64C). Area basalis indistinct (Fig. 64C). Area superomedia indistinct laterally (Fig. 64C). Area petiolaris not strongly elongated (Fig. 64C). Median parts of propodeum partly longitudinally striate (Fig. 64C). Length of fore wing 3.7–4.0 (HT: 3.7) mm. Areolet absent (Fig. 40D). Vein 2m-cu of fore wing with a single bulla (Fig. 40D). Nervellus inclivous, intercepted behind the middle (Fig. 40D). Hind femur 5.5– 5.7 (HT: 5.5) times as long as maximum depth in lateral view. Hind TS I: II: III: IV: V = 2.0: 0.6: 0.35: 0.15: 0.35–0.4 (HT: 0.4). Tarsal claws simple.
Metasoma. T I 1.55–1.7 (HT: 1.7) times as long as maximum width, longitudinally striate (Fig. 40E). Median dorsal carina of T I absent. Dorsolateral carina of T I complete except for indistinct area near spiracle. T II to T VI wider than long. Ovipositor sheath 0.9–0.93 (HT: 0.93) times as long as hind tibia. Ovipositor straight, with a distinct nodus and ventral teeth (Fig. 66J).
Coloration (Figs 40A–E). Body (excluding wings and legs) black to blackish brown. Ventral surface of scape, pedicel, and FL I reddish brown. Dorsal surfaces of scape, FL V to VII, and base of FL VIII white. Mandible reddish brown except for darkened apex and base. Palpi whitish yellow. T III sometimes tinged with reddish brown anteriorly. Apices of T VII and T VIII white. Membranous part of metasomal sternites whitish yellow. Ovipositor reddish brown. Wings hyaline. Veins and pterostigma brown to dark brown. Fore and mid legs brown to yellowish brown except for white trochanters, trochantelli, and base of tibiae. Hind coxa blackish brown, its apex tinged with white. Hind trochanter and base of tibia white. Hind trochantellus, femur and tibia except for base brown to dark brown. Median part of hind tibia more or less tinged with whitish yellow dorsally. Hind tibial spurs and tarsus yellow to yellowish brown.
Male. Unknown.
Distribution. Japan (Honshu, Shikoku, and Kyushu).
Bionomics. Unknown.
Remarks. This species resembles H. hirashimai, but it can be distinguished by the above key. This species and H. hirashimai are differed from other Japanese species by the short area petiolaris, the short hind TS II, and the ovipositor with a distinct nodus. They may be classified into a separate genus, while its taxonomic treatment requires the additional specimens and comparison with world species.
description: Published as part of Watanabe, Kyohei, 2021, Taxonomic and Zoogeographic Study of the Japanese Phygadeuontinae (Hymenoptera, Ichneumonidae), with Descriptions of 17 New Species, pp. 55-136 in Bulletin of the Kanagawa Prefectural Museum (Natural Science) 50 on pages 101-102, DOI: 10.32225/bkpmnh.2021.50_55, http://zenodo.org/record/12774639