Mymaridae
Creators: name: Guzmán-Larralde, Adriana J.
name: Huber, John T.
name: Martínez, Humberto Quiroz
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Mymaridae

description: Key to Mexican genera of Mymaridae. Females
1 Female: flagellum with a distinct apical clava of 1–3 segments (Guzmán-Larralde et al. 2001, fig. 11)..................2
- Male: flagellum filiform, at most with apical two segments more widely joined together (Gibson et al. 2007, fig. 84) but these similar in width to preceding segments (Huber 1997, figs 83, 84).......................................NOT keyed (males of most genera can be identified in Huber 1997)
2(1) Wings extremely short (appearing absent) and ocelli absent.......................... Chrysoctonus Mathot (Fig. 1 A)
- Wings well developed, extending beyond apex of gaster..................................................... 3
3(2) Tarsi 5-segmented.................................................................................... 4
- Tarsi 4-segmented................................................................................... 16
4(3) Funicle 8-segmented................................................................................. 5
- Funicle with 7 or fewer segments....................................................................... 9
5(4) Face with subantenal grooves (Huber 1997, fig. 1); pronotum divided medially into two lobes by a longitudinal groove; propodeum smooth, or with 2 longitudinal submedian lines or carinae dividing its surface into a median and two lateral areas (Huber 1997, figs 37, 38; Guzmán-Larralde et al. 2001, fig. 6)....................................................... 6
- Face without subantenal grooves; pronotum entire, not divided medially by a longitudinal groove; propodeum with a diamondlike pattern of several carinae (Guzmán-Larralde et al. 2001, fig. 4) dividing its surface into 5 large areoles............................................................................................ Ooctonus Haliday (Fig. 1 B)
6(5) Dorsellum strap shaped, narrow, at least 5× as wide as long, with anterior and posterior margins parallel and posterior margin straight or almost so (Huber 2015, fig. 7); pronotum longitudinally divided, with the two lobes widely separated by membranous or sclerotized trapezoidal median area defined laterally by longitudinal sutures, carinae or pale lines (Huber 2015, fig. 8); ocellar triangle usually with 3 setae (Huber 2015, fig. 180), occasionally with 4 setae; fore wing with apex of stigmal vein truncate (Huber 2015, figs 175, 176).................................................. Lymaenon Walker (Fig. 1 C)
- Dorsellum rhomboidal, triangular, or biconvex (Huber 2015, fig. 5), usually much less than 5× as wide as long and anterior margin not at all parallel with posterior margin (Huber 2015, fig. 5); pronotum with the two lobes abutting medially; ocellar triangle usually with 2 setae; fore wing with apex of stigmal vein usually oblique (Huber 2015, fig. 3)................... 7
7(6) Propodeum medially usually with 1–3 longitudinal carinae between submedian carinae (Huber 2015, fig. 102); propodeal spiracle enormous, clearly larger than pronotal spiracle (Huber 2015, fig. 102); gaster extending anteriorly under mesosoma (Huber 2015, figs 95, 105–107) though sometimes only slightly so; occiput divided into dorsal and ventral sections by a curved transverse suture extending above foramen to ventral margin of each eye (Huber 2015, fig. 99); fore wing with microtrichia absent behind and slightly beyond apex of venation, and relatively sparse beyond venation (Huber 2015, fig. 95)............................................................................... Gastrogonatocerus Ogloblin (Fig. 1 D)
- Propodeum without longitudinal carina between submedian carinae; propodeal spiracle normal, the same size or smaller than pronotal spiracle; gaster not extending anteriorly ventral to mesosoma; occiput entire (Huber 2015, figs 11, 45), rarely with a sulcus that constricts the occiput but the sulcus not extending to eyes (Huber 2015, fig. 46); fore wing either with microtrichia behind venation or, if not, then microtrichia beginning at apex of venation, and much denser than above (Huber 2015, fig. 19)................................................................................................... 8
8(7) Fore wing with microtrichia uniformly distributed to base of parastigma (Huber 2015, fig. 108) and usually fairly narrow, at least 3.8× as long as wide, with rounded apex; dorsellum triangular to rhomboidal and sometimes margined with lighter colour (Huber 2015, fig. 6); propodeum smooth medially, rarely with two submedian carinae (Matthews 1986, fig. 3; Triapitsyn 2013, fig. 22); funicle often with funicle segment 1 obliquely truncate dorsoapically, funicle segments 2 and 3 almost always longer than following funicle segments................................................... Gonatocerus Nees (Fig. 1 E)
- Fore wing almost always bare behind venation (Huber 2015, figs 3, 19) or, if setose, then microtrichia usually not uniformly distributed and less densely spaced than those beyond venation (Huber 2015, fig. 41) and usually wider than above and often with a somewhat truncate (less rounded) apex; propodeum with 2 submedian carinae; funicle with funicle segment 1 not obliquely truncate, and funicle segments 2 and 3 not longer than following funicle segments................................................................................................... Cosmocomoidea Howard (Fig. 1 F)
9(4) Fore wing with venation much shorter than half length of wing; mandible with 1 or 2 teeth......................... 10
- Forewing with venation longer than half wing length (Huber 1987, fig. 88; Triapitsyn & Berezovskiy 2004, figs 15, 17, 20); mandible with 4 teeth............................................................. Arescon Walker (Fig. 2 A)
10(9) Metasoma widely joined to mesosoma; petiole short, scarcely or not visible and almost as wide as mesosoma or metasoma; mesophragma extending through petiole into mesosoma (slide mounts, preferably cleared, needed to observe this)...... 11
- Metasoma separated from mesosoma by a visible petiole narrower than width of mesosoma or metasoma width and not
extending into mesosoma............................................................................. 15 11(10) Metasoma compressed, oval and much higher than wide in cross section (resembling that of Figitidae [Hymenoptera: Cynipoidea]); gastral tergum 1 greatly enlarged, almost 0.75 length of gaster; funicle 6-segmented (Guzmán-Larralde et al. 2001, fig. 11).......................................................................... Litus Haliday (Fig. 2 B)
- Metasoma somewhat depressed, widely oval or somewhat triangular in cross section, usually wider than high; gastral tergum 1 usually short, at most 0.5 length of gaster but usually much shorter and similar to following terga; funicle 5- or 7-segmented.................................................................................................. 12
12(11) Funicle 5-segmented; fore wing basally with posterior margin more or less distinctly excised (Guzmán-Larralde et al. 2001, fig. 3)........................................................................ Alaptus Westwood (Fig. 2 C)
- Funicle 7-segmented (segment 2 sometimes very short, ring-like); fore wing with posterior margin at level of venation not excised........................................................................................... 13
13(12) Head in anterior view narrowing ventrally, somewhat pointed; mandibles directed ventrally (not crossed when closed); mandible with 1 long and 1 short tooth; fore wing near midpoint extremely narrow then distinctly widening towards apex......................................................................................... Dicopus Enock (Fig. 2 D)
- Head in anterior view scarcely narrowing ventrally, somewhat square; mandibles not pointing ventrally (usually crossing each other when closed); mandible with 2 equal sized teeth; fore wing near midpoint wider, about the same as width near apex......................................................................................................14
14(13) Occiput with a distinct transverse suture extending above foramen from eye to eye........ Callodicopus Ogloblin (Fig. 2 E)
- Occiput without a transverse suture extending above foramen from eye to eye.......... Dicopomorpha Ogloblin (Fig. 2 F)
15(10) Fore wing fairly wide, with posterior margin straight at apex, the wing surface with dark areas medially and apically separated by a clear area and uniformly covered with several rows of microtrichia; propodeum completely and uniformly covered with distinct, isodiametric reticulate sculpture.............. Camptopteroides Viggiani (subgenus Alalinda Huber) (Fig. 3 A)
- Fore wing narrow, with posterior margin concave at apex, the wing surface uniformly clear and with only one or two longitudinal rows of microtrichia medially; propodeum less uniformly covered by faint, more elongate reticulate sculpture..................................................................................... Camptoptera Foerster (Fig. 3 B)
16(3) Funicle 8-segmented; propodeum, petiole and gastral tergum 1 each with large, translucent, reticulate laminae.............................................................................. Ptilomymar Annecke and Doutt (Fig. 3 C)
- Funicle 6-segmented; propodeum, petiole and gastral tergum 1 without laminae.................................. 17
17(16) Metasoma with petiole much shorter than wide, ring-like and scarcely visible (Guzmán-Larralde et al. 2001, fig. 8)..... 18
- Metasoma with petiole distinctly longer than wide, rarely only about as long as wide.............................. 28
18(17) Clava 2- or 3-segmented............................................................................. 19
- Clava entire....................................................................................... 24
19(18) Clava 2-segmented.................................................................................. 20
- Clava 3-segmented.................................................................................. 21
20(19) Ovipositor strongly exserted beyond apex of gaster, the exserted part usually about as long as the gaster............................................................................................ Omyomymar Schauff (Fig. 3 D)
- Ovipositor at most barely exserted beyond apex of gaster.................................................... 22
21(19) Clava compact, with sutures between segments strongly oblique; scutellum longitudinally divided by wide, shallow furrow............................................................................... Stethynium Enock (Fig. 3 E)
- Clava loose, with sutures between segments perpendicular to long axis of clava; scutellum entire................................................................................................ Neostethynium Ogloblin (Fig. 3 F)
22(20) Fore wing beyond and behind apex of venation with two longitudinal bare areas separated by a row of microtrichia extending apically from just behind apex of venation; frenum entire, without median longitudinal groove; mesophragma with apex truncate or at most very slightly indented medially; propodeum strongly sloping relative to scutellum and with a median carina................................................................................ Anaphes Haliday (Fig. 4 A)
- Fore wing not as described above; instead, either uniformly setose beyond the venation or with only a few rows of setae in a row; frenum apparently divided by a median longitudinal groove; propodeum the same level as scutellum and without a median carina.......................................................................................23
23(22) Fore wing wide, with anterior and posterior margins diverging; funicle segments longer than wide; mesophragma with apex more or less strongly indented medially....................................... Schizophragma Ogloblin (Fig. 4 B)
- Fore wing narrow, knife-like, with anterior and posterior margins parallel; funicle segments short, the apical ones quadrate or wider than long; mesophragma with apex truncate.............................. Platystethynium Ogloblin (Fig. 4 C)
24(18) Ovipositor strongly exserted beyond apex of gaster and with several setae along the exserted portion of each sheath; fore wing venation extending to about 0.5× fore length of wing.............................. Australomymar Girault (Fig. 4 D)
- Ovipositor not or scarcely exserted beyond apex of gaster and with 1 seta near apex of each sheath; fore wing venation shorter, extending to at most about 0.4× fore wing length............................................................25
25(24) Frenum entire, without a median longitudinal groove....................................................... 26
- Frenum divided by a median longitudinal groove...................................... Anagrus Haliday (Fig. 4 E)
26(25) Metasoma with hypopygium conspicuous and extending to apex of gaster................. Erythmelus Enock (Fig. 4 F)
- Metasoma with hypopygium inconspicuous and not extending to apex of gaster.................................. 27
27(26) Fore wing mostly uniformly covered with microtrichia, except for two longitudinal bare areas beyond and behind apex of venation separated by a row of microtrichia extending apically from just below apex of venation; funicle segments mostly longer than wide; mandible with 3 teeth; head in lateral view slightly rounded, with face only slightly bulging anteriorly ventral to torulus; propodeum with a median longitudinal carina................................... Anaphes Haliday (Fig. 4 A)
- Fore wing mostly bare, with one longitudinal row of microtrichia; funicle segments almost all as wide as long, bead-like; mandible with 2 teeth; head in lateral view somewhat triangular, with face projecting anteriorly ventral to torulus; propodeum without a median longitudinal carina................................................... Cleruchus Enock (Fig. 5 A)
28(17) Mandibles pointing ventrally, not crossing when closed, with 1 large tooth serrated ventrally and 1 small tooth; fore wing with uniform brown tinge except for a clear spot at wing apex; petiole short, about as wide as long..................................................................................................... Anagroidea Girault (Fig. 5 B)
- Mandibles crossing when closed, with 3 smooth teeth; fore wing either uniformly clear or with 1 or more brown bands or spots.................................................................................................. 29
29(28) Fore wing oar-like, with basal half to two-thirds extremely narrow and apex suddenly widened with a dark spot at apex and sometimes also at base of widened area; torulus almost touching transverse trabecula...............................30
- Fore wing gradually widening from base to apex and almost always without a dark apical spot; torulus separated from transverse trabecula by at least half diameter of torulus...........................................................31
30(29) Hind wing without membrane, without or almost without marginal setae except for 1 long apical seta; fore wing narrow and without membrane for about 0.75 its length, and widened apical membranous area with 1 or 2 rows of microtrichia and a brown spot in apical half; scape about 7× as long as wide................................... Mymar Curtis (Fig. 5 C)
- Hind wing normal, with membrane and numerous marginal setae; scape about 4× as long as wide................................................................................................. Neomymar Crawford (Fig. 5 D)
31(29) Metatibia with setae conspicuous, distinctly longer than maximum width of tibia in lateral view.................................................................................................. Cnecomymar Ogloblin (Fig. 5 E)
- Metatibia with setae inconspicuous, not or barely longer than width of tibia..................................... 32
32(31) Scutellum divided medially by a longitudinal groove.............................. Tetrapolynema Ogloblin (Fig. 5 F)
- Scutellum entire......................................................................................33
33(32) Scape with inner surface with strong imbricate sculpture; pronotal spiracle midway between tegula and anterior apex of notaulus.......................................................................... Stephanodes Enock (Fig. 6 A)
- Scape with inner surface smooth or with inconspicuous reticulate sculpture; pronotal spiracle much nearer tegula than anterior apex of notaulus.................................................................................... 34
34(33) Propodeum with V–shaped or somewhat H–shaped carinae submedially........................................ 35
- Propodeum without carinae or with a single, median carina...................................................36
35(34) Propodeum with V–shaped carina; fore wing usually with some basally thickened setae and often with 1 or 2 distinct dark transverse bands with microtrichia thicker than those on hyaline areas of wing......... Acmopolynema Ogloblin (Fig. 6 B)
- Propodeum with somewhat H-shaped carinae; fore wing without thickened setae and with a rather indistinct dark band just beyond venation but with microtrichia uniformly thin............................... Platyfrons Yoshimoto (Fig. 6 C)
36(34) Fore wing with 2 wide, distinct brown bands and with a curved longitudinal row of long erect microtrichia contrasting strongly in length with remaining more or less appressed microtrichia...................... Entrichopteris Yoshimoto (Fig. 6 D)
- Fore wing hyaline or with a single brown band and with uniform microtrichia all short and more or less appressed...... 37
37(36) Prosternum open, not separated anteriorly from head by propleura (propleura not abutting medially)................................................................................ Polynema (subgenus Polynema Haliday) (Fig. 6 E)
- Prosternum closed, separated anteriorly from head by propleura (the propleura abutting medially or almost so)......... 38
38(37) Face with a pit next to each torulus...................................... Polynema (subgenus Doriclytus Foerster)
- Face without a pit next to each torulus......................................... Palaeoneura Waterhouse (Fig. 6 F)
description: Published as part of Guzmán-Larralde, Adriana J., Huber, John T. & Martínez, Humberto Quiroz, 2017, Generic key and catalogue of Mymaridae (Hymenoptera) of Mexico, pp. 1-38 in Zootaxa 4254 (1) on pages 3-5, DOI: 10.11646/zootaxa.4254.1.1, http://zenodo.org/record/545688