Sycophila (Tineomyza) wiebesi Lotfalizadeh & Delvare & Cruaud & Rasplus 2024, sp. n.
Creators: name: Lotfalizadeh, Hossein
name: Delvare, Gérard
name: Cruaud, Astrid
name: Rasplus, Jean-Yves
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Eurytomidae
Sycophila
Sycophila wiebesi

description: Sycophila (Tineomyza) wiebesi Lotfalizadeh & Rasplus sp. n.   
(Figs 121–123)  
  Type material examined. Holotype  ♀: Guinea, Monts Nimba, Mifergui, ex syconia of Ficus artocarpoides, 1.iv.1993, Rasplus J.-Y. leg. Deposited in CBGP. Paratypes: same data as holotype, 11♀. Monts Nimba, Mifergui, ex syconia of Ficus laurifolia, 19.iv.1993, Rasplus J.-Y. leg., 2♀ & 1♂. Gabon, Makokou, ex syconia of F. artocarpoides, 2.ix.1988, Maldès J.M. leg., 1♀.   
 Etymology. This species is named after J.T. Wiebes for his great contribution to our knowledge of fig wasps.   
 Diagnosis. Female body mainly dark (Fig. 121A). POL less than 2× OOL (Fig. 121D). All funiculars distinctly longer than wide, fu1 much longer than pedicel, clava short (less than 2× as long as wide) (Fig. 121B). Petiole long (longer than wide), without ventral transverse carina between petiole and St1 (Fig. 122B). Basal carina of propodeum inverted V-shaped, medially surrounded by a large areola (Fig. 122D). Fore wing with a large dark brown spot, basal cell setose, costal cell densely setose ventrally and a row of setae dorsally (Fig. 122C).   
Male body mostly dark or dark brown (Fig. 123A). Fu1 longer than wide, fu2–fu4 at most as long as wide (Fig. 123B). Fore wing entirely setose, without speculum, basal cell setose (Fig. 123D).  
 Description of holotype ♀ (Figs 121–122). Length 4.03 mm. Body generally dark or dark brown (Fig. 121A), except the following parts, all yellowish-brown: entire orbital margin, torular margin, dorsal margin of dorsal ocelli, antenna, pronotum dorsolaterally and posteriorly, tegula, mesopleuron, legs (except fore coxae and femur, mid coxae, hind femur and tibia), gaster ventrally, Gt4 anterolaterally; antenna dark–brown; pronotum anterolaterally testaceous; fore wing with a dark brown spot below mv, reaching base of mv, lower margin of mv not obscured by dark brown spot (Fig. 122C); gaster not uniformly coloured (Fig. 122B); body setation whitish, setation of fore wing entirely dark.   
 Head 1.32× as broad as high [HW 188 & HH 142] (Fig. 121C); in dorsal view transverse, 1.85× as broad as long [HW 240 & HL 130]; face and vertex coriaceous with sparse punctures; malar space 0.53× length of eye [ms 45 & EH 85] (Fig. 121D); malar sulcus distinct, sinuous (Fig. 121D); genal margin regularly curved, edged but not carinate; antennal toruli widely separated; interantennal space not raised, not carinate laterally; scrobal depression smooth, reaching the median ocellus, not carinate on margin; POL 1.86× as long as OOL [POL 41 & OOL 22] (Fig. 121D); temple rounded, 0.27× the eye length (in dorsal view) [TL 28 & EL 105]; setation oriented downwards on face, outwards on the punctuate frons.   
 Antenna. Scape reaching lower margin of median ocellus, 3.91× as long as broad [SL 90 & SW 23]; fu1 2.25× as long as broad [fu1L 45 & fu1W 20], distinctly longer than pedicel, not bulging dorsally; funiculars (fu2–fu4) with three rows of mps and 2–3 whorls of adpressed short setae; fu2–fu5 distinctly longer than broad (Fig. 121B).   
 Mesosoma weakly arched dorsally, 1.55× as long as wide (170:110), with deep sculpture (Fig. 122A); pronotum with coarse umbilicate punctures; mesepisternum flattened, without any process or raised sculpture on epicnemium; mesoscutum 0.6× as long as wide (65:110), 0.75× longer than length of pronotal collar (65:85), with dense umbilicate punctures, transversely strigose anteriorly; notauli conspicuous, slightly obliterated by mesoscutum sculpture; prepectus bare, with raised sculpture; mesoscutellum umbilicate laterally and on mesoscutellar disc; mesepimeron transversally strigose; metepimeron subtriangular and areolate with long white hairs; propodeum (Fig. 122B) with deep median depression, with a smooth median band, without any median carina; lateral rows of areolae absent; basal carina inverted V, truncate mesally; lateral branches of basal carina slightly convex; distance between lateral carinae less than 0.5× width of nucha.   
 Fore wing with short marginal fringe, apically subtruncate (pointed anteriorly); dark brown spot below mv; basal cell more setose; mv slightly enlarged, normally broadened, in lower margin straight, longer than pmv; stv not curved; pmv distinct, longer than stv; ventral side of costal cell densely setose, dorsally with row of setae; speculum present; parastigmal bristles absent (Fig. 122C).   
 Legs. Fore coxa without depression on frontal surface, hind coxa bare basodorsally; fore (38:12) and hind femora (40:15); hind tibia dorsally with more than 10 setae, setae shorter than basal tibial width.   
 Metasoma. Petiole longer than broad dorsally, ventral transverse carina between petiole and St1 absent. Metasoma 1.6× as long as wide (170:105) (Fig. 122B), longer than mesosoma, strongly compressed laterally and dorsally angulate (Fig. 122E); ventral transverse carina between petiole and St1 absent; petiole in dorsal view longer than wide, shorter than half-length of hind coxa; tergites finely sculptured; Gt4 (in lateral view) slightly as long as Gt3, not sinuous posteriorly; Gt5 partly covering Gt6, not sinuous posteriorly; Gt5 with few setae dorsally, Gt6 sparsely setose ventrally; ovipositor upturned (Fig. 122B).   
 Male (Fig. 123). Length 2.74–6.22 mm. Similar to female including the colouration with longer petiole (Fig. 123A); antennal formula 11143, with much shorter segments, at most as long as broad; each funicular with one row of mps and several whorls of short erected setae (Fig. 123B); fore wing more setose than in female, without speculum (Fig. 123D).   
 Variation. Length 3.33–5.15 mm. In pale specimens, tergites with a yellowish band anteriorly, legs partly yellowish, mesopleuron yellowish-brown.   
 Host plants. Ficus artocarpoides and F. laurifolia.   
 Geographical distribution. Gabon and Guinea.
description: Published as part of Lotfalizadeh, Hossein, Delvare, Gérard, Cruaud, Astrid & Rasplus, Jean-Yves, 2024, Morphological phylogeny and revision of Sycophila and Ficomila (Hymenoptera: Chalcidoidea, Eurytomidae) associated with Afrotropical fig trees (Moraceae, Ficus), pp. 1-190 in Zootaxa 5401 (1) on pages 167-171, DOI: 10.11646/zootaxa.5401.1.1, http://zenodo.org/record/10524258