cometadata/datacite-rtg-text-other-reclassification-records huggingface dataset:
Crematogaster treubi subsp. apilis Forel
Creators: name: Hosoishi, Shingo
name: Ogata, Kazuo
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Crematogaster
Crematogaster treubi
C. treubi apilis forel

description: Key to species based on the worker caste (except for C. treubi apilis Forel, which was not examined)
1 Clypeus with distinct rugulae. Petiole rectangular, with angular corners in dorsal view (Fig. 3 a). Occipital carina developed as distinct ridge (Fig. 2 a).................................................................................. 2
- Clypeus relatively smooth and shining without distinct rugulae. Petiole broader anteriorly than posteriroly, but without angular corners (Fig. 3 b). Occipital carina undeveloped, without distinct ridge (Fig. 2 b).................................... 3
2 Clypeus with one longitudinal rugula on the median portion and two distinct pairs of longitudinal rugulae laterally. Dorsal promesonotum with dense reticulate sculpture (Fig. 4 a)................................................... brevis
- Clypeus with two distinct pairs of longitudinal rugulae, but lacking median rugula. Dorsal promesonotum with more open reticulate sculpture with shiny interspaces (Fig. 4 b).................................................... overbecki
3 Propodeal spines undeveloped (Fig. 12)................................................................ yappi
- Propodeal spines developed (Fig. 6)....................................................................... 4
4 Postpetiole strongly bilobed behind (Fig. 3 c)........................................................... walshi
- Postpetiole weakly bilobed (Fig. 3 b)....................................................................... 5
5 Metanotal groove distinct and deep (Fig. 5 a).......................................................... bouvardi
- Metanotal groove less distinct and deep (Fig. 5 b)............................................................ 6
6 Propodeal spines short and stout, their length shorter than the diameter of propodeal spiracles (Fig. 5 b)............. kojimai
- Propodeal spines long and slender, their length longer than the diameter of propodeal spiracles (Fig. 5 c)............. treubi
description: Published as part of Hosoishi, Shingo & Ogata, Kazuo, 2012, Revision of the Crematogaster brevis complex in Asia (Hymenoptera: Formicidae), pp. 18-30 in Zootaxa 3349 on page 20, DOI: 10.11646/zootaxa.3349.1.2, http://zenodo.org/record/212508
Crematogaster brevis Emery
Creators: name: Hosoishi, Shingo
name: Ogata, Kazuo
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Crematogaster
Crematogaster brevis

description: Crematogaster brevis Emery
(Figs 3 a, 4a, 7)
Crematogaster brevis Emery, 1887: 467 (footnote), pl. 2, fig. 19.
LECTOTYPE worker (top specimen of two on one pin) (by present designation) and one paralectotype worker, one paralectotype queen, four paralectotype males from INDONESIA: Buitenzorg [Bogor, Java] (Conte Solms Laubach) (MCSN) [examined].
Combination in C. (Orthocrema) by Emery, 1922: 132; in C. (Mesocrema) by Santschi, 1928: 33.
Measurements and indices. HW 0.7-0.71; HL 0.68-0.72; CI 99-103; SL 0.49-0.53; SI 70-75; EL 0.14; PW 0.4- 0.44; WL 0.62-0.63; PSL 0.1-0.11; PtL 0.23; PtW 0.19-0.21; PtH 0.15; PpL 0.12-0.13; PpW 0.19-0.2; PtHI 65; PtWI 91; PpWI 154-158; WI 95-100 (Lectotype and one paralectotype workers measured).
General description of worker. Compound eyes slightly projecting from lateral margin of head. Scape reaching posterior corner of head, the scape with standing and appressed setae.
Pronotal shoulders developed as ridges. Mesonotal dorsum convex in lateral view. Metanotal groove concave. Propodeal spines longer than the diameter of propodeal spiracles, curved upward. Anterior margin of metapleural gland bulla not reaching anterior margin of propodeal spiracle.
Petiole in dorsal view rectangular with angular corners. Subpetiolar process not visible in the specimens examined. Postpetiole bilobed behind, but without distinct longitudinal median sulcus.
Dorsal surface of head generally smooth, but the malar region with longitudinal rugulae. Clypeus with one median and two distinct pairs of lateral rugulae. The rugulae connecting the anterior and posteror clypeal margins. Dorsal face of promesonotum with dense reticulate sculpture. Lateral surface of pronotum generally smooth and shining, but some longitudinal rugulae extending from the anterior portion of the pronotum. Mesopleuron smooth, but lower portion of mesopleuron weakly sculptured. Longitudinal rugulae developed on the front and behind the metanotal groove in lateral view.
Fourth abdominal tergite with abundant appressed setae and sparse standing setae.
Distribution. This species is known only from the type locality in Indonesia (Java).
Remarks. This species is similar to C. overbecki. It differs in having a single median rugula in addition to two pairs of lateral rugulae on the clypeus, and in the pronounced sculpture on the dorsum of the mesonotum.
The type specimens were collected from the vine Dischidia major (Vahl) (= Dischidia rafflesiana Wallich) with Dolichoderus bituberculatus Mayr (Emery, 1887).
description: Published as part of Hosoishi, Shingo & Ogata, Kazuo, 2012, Revision of the Crematogaster brevis complex in Asia (Hymenoptera: Formicidae), pp. 18-30 in Zootaxa 3349 on pages 22-23, DOI: 10.11646/zootaxa.3349.1.2, http://zenodo.org/record/212508
Crematogaster overbecki Viehmeyer
Creators: name: Hosoishi, Shingo
name: Ogata, Kazuo
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Crematogaster
Crematogaster overbecki

description: Crematogaster overbecki Viehmeyer
(Figs 2 a, 4b, 9)
Crematogaster overbecki Viehmeyer, 1916: 123, fig. 3.
Syntype worker from SINGAPORE (H. Overbeck). (NHMB) [examined].
Combination in C. (Orthocrema) by Emery, 1922: 132; in C. (Mesocrema) by Santschi, 1928: 33.
Measurements and indices. HW 0.63; HL 0.63; CI 100; SL 0.44; SI 70; EL 0.12; PW 0.4; WL 0.64; PSL 0.11; PtL 0.18; PtW 0.19; PtH 0.13; PpL 0.12; PpW 0.18; PtHI 72; PtWI 106; PpWI 150; WI 95 (One syntype worker measured).
General description of worker. Compound eyes slightly projecting from lateral margin of head. Scape reaching posterior corner of head, the scape with standing and appressed setae.
Pronotal shoulders developed as ridges. Mesonotal dorsum highly convex in lateral view. Metanotal groove concave. Propodeal spines longer than the diameter of propodeal spiracles, curved upward. Anterior margin of metapleural gland bulla not reaching anterior margin of propodeal spiracle.
Petiole in dorsal view rectangular with angular corners. Subpetiolar process weakly developed, angulate. Postpetiole bilobed behind, but without distinct longitudinal median sulcus.
Dorsal surface of head generally smooth, but the malar region with longitudinal rugulae. Clypeus with two distinct pairs of rugulae laterally. The rugulae connecting the anterior and posteror clypeal margins. Dorsal face of promesonotum with relatively open, reticulate sculpture with shiny interspaces. Lateral surface of pronotum generally smooth and shining, but some longitudinal rugulae extending from the anterior portion of pronotum.
Mesopleuron smooth, but lower portion of mesopleuron weakly sculptured. Longitudinal rugulae developed on the front and behind the metanotal groove in lateral view. Some rugulae on the mesonotum extending to the anterior portion of propodeal dorsum.
Fourth abdominal tergite with abundant appressed setae, and sparse standing setae.
Distribution. This species is known only from the type locality in Singapore.
Remarks. This species is similar to C. brevis, but can be distinguished by lacking median rugula on the clypeus, and by the distinctive sculpture of the mesonotal dorsum.
description: Published as part of Hosoishi, Shingo & Ogata, Kazuo, 2012, Revision of the Crematogaster brevis complex in Asia (Hymenoptera: Formicidae), pp. 18-30 in Zootaxa 3349 on pages 24-25, DOI: 10.11646/zootaxa.3349.1.2, http://zenodo.org/record/212508
Crematogaster kojimai, sp. nov.
Creators: name: Hosoishi, Shingo
name: Ogata, Kazuo
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Crematogaster
Crematogaster kojimai

description: Crematogaster kojimai sp. nov.
(Figs 5 b, 8)
HOLOTYPE worker from MALAYSIA: path 3, Cameron Highlands, 13. iii. 2010 (SH10-Mal-53) (S. Hosoishi) (FRIM). Eight paratype workers, same data as holotype (BMNH, KUM, MCZC, MHNG, NHMB, MCSN, MZB, THNHM).
Measurements and indices. HW 0.6-0.71; HL 0.61-0.69; CI 95-104; SL 0.45-0.51; SI 69-77; EL 0.1-0.13; PW 0.34-0.4; WL 0.66-0.77; PSL 0.05-0.09; PtL 0.16-0.2; PtW 0.18-0.22; PtH 0.11-0.14; PpL 0.1-0.14; PpW 0.14- 0.19; PtHI 60-82; PtWI 100-129; PpWI 114-158; WI 78-94 (Twelve workers measured).
General description of worker. Workers monomorphic.
Compound eyes slightly projecting from lateral margin of head. Scape usually just reaching posterior corner of head in smaller workers, not reaching it in larger workers, the scape with standing setae.
Anterior margin of pronotal collar almost straight in dorsal view. Metanotal groove concave in lateral view. Propodeal spines short and stout, their length shorter than the diameter of propodeal spiracles. Anterior margin of metapleural gland bulla exceeding anterior margin of propodeal spiracle.
Petiole in dorsal view broader anteriorly than posteriorly. Postpetiole weakly bilobed, without distinct longitudinal median sulcus. Subpetiolar and subpostpetiolar processes undeveloped.
Clypeus relatively smooth and shining, but with feeble rugulae. Dorsal face of mesosoma almost smooth and shining. Dorsolateral face of pronotum smooth in dorsal view. Lateral surface of pronotum smooth and shining. Lateral surface of mesonotum with rugulae. Mesopleuron relatively smooth and shining, but the anterodorsal and lower portions weakly sculptured.
Standing setae abundant. Dorsal face of head, clypeus and mesosoma with standing setae. Fourth abdominal tergite with sparse standing setae.
Distribution. This species is known from Malaysia (Peninsula).
Remarks. This species is similar to C. treubi, but can be distinguished by the short and stout propodeal spines.
The nest series in Cameron Highlands (e. g. SH10-Mal-83) was collected from Dischidia astephana (c.f. Rintz, 1980).
Specimens examined. MALAYSIA: 2 workers, path 3, Cameron Highlands, 11. iii. 2005 (2005c9) (S. Hosoishi); 3 workers, path 3, Cameron Highlands, 11. iii. 2005 (2005c14) (S. Hosoishi); 5 workers, path 3, Cameron Highlands, 13. iii. 2010 (SH10-Mal-83) (S. Hosoishi); 13 workers, St. 8, Gunung Berembun, Cameron Highlands, 15. vii. 2003 (Fogging) (H. Kojima et al).
description: Published as part of Hosoishi, Shingo & Ogata, Kazuo, 2012, Revision of the Crematogaster brevis complex in Asia (Hymenoptera: Formicidae), pp. 18-30 in Zootaxa 3349 on page 24, DOI: 10.11646/zootaxa.3349.1.2, http://zenodo.org/record/212508
Crematogaster treubi subsp. apilis Forel
Creators: name: Hosoishi, Shingo
name: Ogata, Kazuo
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Crematogaster
Crematogaster treubi
Crematogaster treubi apilis forel

description: Crematogaster treubi apilis Forel
Crematogaster treubi r. apilis Forel, 1913: 195. worker from TAIWAN: Pilam (Stück) (types not found in MHNG) [not seen]. Combination in C. (Orthocrema) by Emery, 1922: 132.
Remarks. We have not been able to examine type-material of C. treubi apilis. From the original description, this subspecies belongs to the C. treubi group. Forel (1913) described the following features: (1) scape exceeding posterior corner of head by two times of the width of the scape; (2) the metanotal groove deep. The first feature is not found in the other species of the C. treubi group. It is impossible to judge the depth of the metanotal groove from the original description. The taxonomic status of this subspecies will remain uncertain until the type-material can be examined.
description: Published as part of Hosoishi, Shingo & Ogata, Kazuo, 2012, Revision of the Crematogaster brevis complex in Asia (Hymenoptera: Formicidae), pp. 18-30 in Zootaxa 3349 on page 26, DOI: 10.11646/zootaxa.3349.1.2, http://zenodo.org/record/212508
Crematogaster walshi Forel
Creators: name: Hosoishi, Shingo
name: Ogata, Kazuo
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Crematogaster
Crematogaster walshi

description: Crematogaster walshi Forel
(Figs 3 c, 11)
Crematogaster walshi Forel, 1902: 205.
LECTOTYPE worker (top specimen of three on one pin, MHNG worker) (by present designation) and five paralectotype workers from INDIA: Pooree, Bengale [Puri, Bengal] (Walsh) (MHNG, NHMB) [examined].
Combination in C. (Orthocrema) by Emery, 1922: 132; in C. (Mesocrema) by Santschi, 1928: 33.
Measurements and indices. HW 0.72-0.93; HL 0.76-0.92; CI 93-103; SL 0.49-0.58; SI 62-69; EL 0.13-0.17; PW 0.42-0.53; WL 0.83-1.03; PSL 0.04-0.08; PtL 0.21-0.27; PtW 0.21-0.27; PtH 0.13-0.17; PpL 0.13-0.18; PpW 0.2- 0.26; PtHI 59-68; PtWI 93-105; PpWI 139-162; WI 87-100 (Lectotype and five paralectotype workers measured).
General description of worker. Workers monomorphic.
Compound eyes not projecting from lateral margin of head, the outer margin of eye almost flush with head capsule in full face view. Scape not reaching posterior corner of head, the scape with standing setae.
Anterior margin of pronotal collar almost straight or slightly convex in dorsal view. Metanotal groove concave in lateral view. Propodeal spines developed, but their length variable. Anterior margin of metapleural gland bulla exceeding anterior margin of propodeal spiracle.
Petiole in dorsal view broader anteriorly than posteriorly in smaller workers, broader at middle portion in larger workers. Postpetiole strongly bilobed behind without a distinct longitudinal median sulcus. Subpetiolar and subpostpetiolar processes undeveloped.
Clypeus relatively smooth, but with feeble rugulae. Dorsal face of mesosoma almost smooth and shining. Dorsolateral face of pronotum smooth in dorsal view. Lateral surface of pronotum smooth and shining. Mesopleuron weakly sculptured, but the higher portion with longitudinal rugulae in lateral view.
Dorsal face of head, clypeus and mesosoma with abundant standing setae. Fourth abdominal tergite with sparse standing setae.
Distribution. This species is known only from the type locality in India.
Remarks. This species is distinct in having the strongly bilobed postpetiole.
description: Published as part of Hosoishi, Shingo & Ogata, Kazuo, 2012, Revision of the Crematogaster brevis complex in Asia (Hymenoptera: Formicidae), pp. 18-30 in Zootaxa 3349 on pages 26-28, DOI: 10.11646/zootaxa.3349.1.2, http://zenodo.org/record/212508
Crematogaster yappi Forel
Creators: name: Hosoishi, Shingo
name: Ogata, Kazuo
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Crematogaster
Crematogaster yappi

description: Crematogaster yappi Forel
(Fig 12)
Crematogaster yappi Forel, 1901: 374.
LECTOTYPE worker (bottom specimen of three on one pin, MHNG worker) (by present designation) and five paralectotype workers from MALAYSIA: Gunung Inas, Kedah (M. R. H. Yapp) (MHNG, NHMB) [examined].
Combination in C. (Acrocoelia) by Emery, 1922: 152.
Measurements and indices. HW 0.65-0.74; HL 0.66-0.75; CI 94-105; SL 0.48-0.54; SI 70-77; EL 0.11-0.15; PW 0.34-0.4; WL 0.71-0.81; PSL 0; PtL 0.18-0.22; PtW 0.21-0.24; PtH 0.12-0.15; PpL 0.12-0.14; PpW 0.17-0.2; PtHI 67-74; PtWI 105-111; PpWI 129-167; WI 77-90 (Lectotype and five paralectotype workers measured).
General description of worker. Compound eyes slightly projecting from lateral margin of head. Scape reaching posterior corner of head, the scape with standing setae.
Anterior margin of pronotal collar almost straight in dorsal view. Metanotal groove concave in lateral view. Propodeal spines undeveloped. Anterior margin of metapleural gland bulla almost reaching anterior margin of propodeal spiracle.
Petiole in dorsal view broader anteriorly than posteriorly. Postpetiole weakly bilobed, but without a distinct longitudinal median sulcus. Subpetiolar and subpostpetiolar processes undeveloped.
Dorsal surface of head generally smooth and shining, but the lower portion near clypeal margin with longitudinal rugulae. Clypeus relatively smooth, but with feeble rugulae. Dorsal face of mesosoma almost smooth and shining. Dorsolateral face of pronotum smooth in dorsal view. Lateral surface of pronotum smooth and shining. Mesopleuron relatively smooth, but with feeble rugulae on the higher portion.
Fourth abdominal tergite with sparse standing setae.
Distribution. This species is known only from the type locality in Malaysia (Peninsula).
Remarks. This species can be easily distinguished from other species of the C. brevis complex found in Asia by its undeveloped propodeal spines.
description: Published as part of Hosoishi, Shingo & Ogata, Kazuo, 2012, Revision of the Crematogaster brevis complex in Asia (Hymenoptera: Formicidae), pp. 18-30 in Zootaxa 3349 on page 28, DOI: 10.11646/zootaxa.3349.1.2, http://zenodo.org/record/212508
Crematogaster (Crematogaster) jacobsoni Forel
Creators: name: Hosoishi, Shingo
name: Ogata, Kazuo
Publisher: name: Zenodo
Year: 2012
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Formicidae
Crematogaster
Crematogaster jacobsoni

description: Crematogaster (Crematogaster) jacobsoni Forel
Crematogaster jacobsoni Forel, 1911b: 201.
Three syntype workers from INDONESIA: Semarang, Java (MHNG) and one syntype worker (NHMB) [examined].
Combination in C. (Orthocrema) by Emery, 1922: 132; in C. (Acrocoelia) by Santschi, 1928: 34; in C. (Mesocrema) by Santschi, 1937: 310.
Remarks. Examination of syntype workers in MHNG and NHMB reveals that C. jacobsoni is not similar to the C. brevis group or C. treubi group in Asia. Characteristic features include: occipital carinae developed; petiole weakly broader anteriorly, but without angular corners; postpetiole weakly bilobed with longitudinal median sulcus.
This species is similar to C. (C.) semperi Emery, 1893 or C. (C.) simoni Emery, 1893. We hereby transfer C. jacobsoni to the subgenus Crematogaster, to compare it with closely related species in this group in the future.
description: Published as part of Hosoishi, Shingo & Ogata, Kazuo, 2012, Revision of the Crematogaster brevis complex in Asia (Hymenoptera: Formicidae), pp. 18-30 in Zootaxa 3349 on pages 28-29, DOI: 10.11646/zootaxa.3349.1.2, http://zenodo.org/record/212508
Copelatus glyphicus NA
Creators: name: Alarie, Yves
name: Michat, Mariano C.
name: Watanabe, Kohei
name: Shaverdo, Helena
name: Wang, Liang-Jong
name: Watts, Chris H. S.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Dytiscidae
Copelatus
Copelatus glyphicus

description: Copelatus glyphicus (Say, 1823)
Source of material. The five instar III larvae studied were associated with adults collected at the following locality: Canada. Québec, Trois-Rivières. Ephemeral pond in red maple forest at Chemin des Oblats, 06.VIII.1986, 08.VIII.1986, 12.VIII.1986; Y. Alarie leg. The identification is firm as C. glyphicus is the only Copelatus species known from that region.
Diagnosis (instar III). The third instar of Copelatus glyphicus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80–0.90 mm; L3 <2.00 mm; U <0.40 mm; head capsule rounded, strongly constricted at level of occipital region (Fig. 19), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus broadly convex, extending mesally beyond level of adnasalia (Fig. 19); adnasalia margined with short bluntly rounded and truncated teeth (Fig. 63); MP2/MP1> 1.50; GA/MP1> 1.50; 1.30 <LP2/LP1 <1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U/HW <0.40; U/LAS <0.50; Nearctic.
Description, instar III (Figs 19–20, 63)
Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.
Head (Figs 19, 63): Head capsule rounded, strongly constricted at level of occiput; anterior margin of frontoclypeus broadly convex, extending beyond level of adnasalia; scale-like sculpticels well-developed both on frontoclypeus and parietale; adnasalia margined with short bluntly rounded and truncated teeth (Fig. 63); HL = 0.81–0.94 mm; A/MP = 1.34–1.38; MP/LP = 2.12–2.17; MP2/MP1 = 1.50–1.73; GA/MP1 = 1.79–1.88; LP2/LP1 = 1.40–1.56; MNL/MNW = 2.85–3.04.
Thorax: L3 = 1.60–1.74 mm.
Abdomen (Fig. 20): LAS = 0.65–0.74 mm; LAS subcylindrical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.28–0.32 mm, composed of one urogomphomere; U/HW = 0.31–0.33; U/LAS = 0.42–0.44.
Chaetotaxy: Parietale with 6–9 temporal spines; position and number of secondary setae on legs are shown in Table 7.
description: Published as part of Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong & Watts, Chris H. S., 2022, An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae), pp. 151-205 in Zootaxa 5175 (2) on page 168, DOI: 10.11646/zootaxa.5175.2.1, http://zenodo.org/record/7003344
Copelatus japonicus Sharp 1884
Creators: name: Alarie, Yves
name: Michat, Mariano C.
name: Watanabe, Kohei
name: Shaverdo, Helena
name: Wang, Liang-Jong
name: Watts, Chris H. S.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Dytiscidae
Copelatus
Copelatus japonicus

description: Copelatus japonicus Sharp, 1884
Source of material. The four instar III larvae studied were associated with adults collected at the following locality: Taiwan. Taipei, Beitou, 24.VIII.1994; L.J. Wang leg. The identification is firm as C. japonicus is the only Copelatus species collected at this locality.
Diagnosis (instar III). The third instar of Copelatus japonicus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.90 mm; L3 <2.00 mm; U <0.40 mm; head capsule rounded, strongly constricted at level of occipital region (Fig. 21), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally at about level of adnasalia (Fig. 21); adnasalia margined with bluntly rounded and truncated teeth (Fig. 63); MP2/MP1> 1.50; GA/ MP1> 1.50; 1.30 <LP2/LP1 <1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of two urogomphomeres; U/HW <0.40; U/LAS <0.50; Palaearctic.
Description, instar III (Figs 21–24, 63)
Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.
Head (Figs 21, 63): Head capsule rounded, strongly constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending mesally at about level of adnasalia; scale-like sculpticels well-developed over frontoclypeus and parietale; adnasalia margined with bluntly rounded and truncated teeth (Fig. 63); HL = 0.86–0.90 mm; A/MP = 1.51–1.60; MP/LP = 1.77–1.88; MP2/MP1 = 1.57–1.82; GA/MP1 = 1.67–1.85; LP2/LP1 = 1.28–1.61; MNL/MNW = 2.50–2.72.
Thorax (Figs 23–24): L3 = 1.84–1.92 mm.
Abdomen (Fig. 22): LAS = 0.66–0.72 mm; LAS subconical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus, U = 0.28–0.30 mm, composed of two urogomphomeres; U/HW = 0.28–0.30; U/LAS = 0.42–0.43.
Chaetotaxy: Parietale with 4–6 temporal spines; position and number of secondary setae on legs are shown in Figs 23–24 and Table 7.
description: Published as part of Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong & Watts, Chris H. S., 2022, An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae), pp. 151-205 in Zootaxa 5175 (2) on pages 168-169, DOI: 10.11646/zootaxa.5175.2.1, http://zenodo.org/record/7003344
Copelatus tenebrosus Regimbart 1880
Creators: name: Alarie, Yves
name: Michat, Mariano C.
name: Watanabe, Kohei
name: Shaverdo, Helena
name: Wang, Liang-Jong
name: Watts, Chris H. S.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Dytiscidae
Copelatus
Copelatus tenebrosus

description: Copelatus tenebrosus Régimbart, 1880
Source of material. The single instar III larvae studied was reared ex ovo from were adults collected at the following locality: Japan. Mantaburu, Yonaguni-cho, Yonaguni-jima Island, Okinawa Prefecture, 24°27’04.1”N 122°57’36.8”E, 13.VI.2019; E. K. Watanabe leg.
Diagnosis (instar III). The third instar of Copelatus tenebrosus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80 mm; L3 <2.00 mm; U <0.40 mm; head capsule rounded, slightly constricted at level of occipital region (Fig. 52), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally well beyond level of adnasalia (Fig. 52); adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); MP2/MP1> 1.50; GA/MP1> 1.50; 1.30 <LP2/LP1 <1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of two urogomphomeres; U/HW <0.40; U/LAS <0.50; Australia, Oriental, Palaearctic.
Description, instar III (Figs 52–53, 65)
Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.
Head (Figs 52, 65): Head capsule rounded, slightly constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending mesally well beyond level of adnasalia; scale-like sculpticels present over frontoclypeus and parietale; adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); HL = 0.84 mm; A/MP = 1.40; MP/LP = 2.03; MP2/MP1 = 1.64; GA/MP1 = 1.85; LP2/LP1 = 1.43; MNL/MNW = 2.87.
Thorax: L3 = 1.73 mm.
Abdomen (Fig. 53): LAS = 0.63 mm; LAS subconical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus, U = 0.30 mm, composed of two urogomphomeres; U/HW = 0.33; U/LAS = 0.47.
Chaetotaxy: Parietale with 5–6 temporal spines; position and number of secondary setae on legs are shown in Table 7.
description: Published as part of Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong & Watts, Chris H. S., 2022, An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae), pp. 151-205 in Zootaxa 5175 (2) on page 187, DOI: 10.11646/zootaxa.5175.2.1, http://zenodo.org/record/7003344
Exocelina australiae AU
Creators: name: Alarie, Yves
name: Michat, Mariano C.
name: Watanabe, Kohei
name: Shaverdo, Helena
name: Wang, Liang-Jong
name: Watts, Chris H. S.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Dytiscidae
Exocelina
Exocelina australiae

description: Exocelina australiae (Clark, 1863)
Source of material. The three instar III larvae studied were collected in association with adults at the following locality: South Australia. Ten kilometers North of Forreston, 9.VIII.1999; C. H. S. Watts leg.
Diagnosis (instar III). The third instar of Exocelina australiae can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 1.85–1.90 mm; L3> 2.90 mm; U> 0.50 mm; head capsule subquadrate, strongly constricted at level of occipital region (Fig. 54), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus broadly convex, extending mesally at level of adnasalia (Fig. 54); adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); MP2/MP1> 1.50; GA/MP1> 1.50; LP2/LP1> 2.00; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with more than 5 PV and 5 AV secondary setae; metafemur with more than 4 AV secondary setae; urogomphus composed of one urogomphomere; U/HW <0.40; U/LAS <0.50; Australian.
Description, instar III (Figs 54–55, 65)
Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.
Head (Figs 54, 65): Head capsule subquadrate, strongly constricted at level of occiput; anterior margin of frontoclypeus broadly convex, extending mesally at about level of adnasalia; scale-like sculpticels present over frontoclypeus and parietale; adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); HL = 1.85–1.90 mm; A/MP = 1.37–1.41; MP/LP = 2.36–2.41; MP2/MP1 = 1.56–1.67; GA/MP1 = 1.53–1.73; LP2/LP1 = 2.00–2.67; MNL/MNW = 2.72–2.96.
Thorax: L3 = 3.71–3.87 mm.
Abdomen (Fig. 55): LAS = 1.33–1.63 mm; LAS subcylindrical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus, U = 0.56–0.58 mm, composed of one urogomphomere; U/HW = 0.25–0.26; U/LAS = 0.35–0.44.
Chaetotaxy: Parietale with 9–10 temporal spines; position and number of secondary setae on legs are shown in Table 7.
description: Published as part of Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong & Watts, Chris H. S., 2022, An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae), pp. 151-205 in Zootaxa 5175 (2) on page 187, DOI: 10.11646/zootaxa.5175.2.1, http://zenodo.org/record/7003344
Liopterus haemorrhoidalis
Creators: name: Alarie, Yves
name: Michat, Mariano C.
name: Watanabe, Kohei
name: Shaverdo, Helena
name: Wang, Liang-Jong
name: Watts, Chris H. S.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Dytiscidae
Liopterus
Liopterus haemorrhoidalis

description: Liopterus haemorrhoidalis (Fabricius, 1787)
Source of material. The four instar III larvae were collected in association with adults at the following locality: Belarus, Vicebsk Region, area near upy Village, wetland of evinka River, Carex, moss. 12.VIII.1998;H.Shaverdo leg.
Diagnosis (instar III). The third instar of Liopterus haemorrhoidalis can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 1.40 mm; L3> 2.90 mm; U> 0.50 mm; head capsule subquadrate, strongly constricted at level of occipital region (Fig. 58), scale-like sculpticels lacking over frontoclypeus and parietale; anterior margin of frontoclypeus broadly convex, extending mesally at level of adnasalia (Fig. 58); adnasalia margined with short bluntly rounded teeth (Fig. 65); MP2/MP1 <1.30; GA/MP1 <1.50; 1.30 <LP2/LP1 <1.90; inner margin of stipes with a dorsal linear row of spinulae (Fig. 60); profemur with less than 3 PV and more than 5 AV secondary setae; metafemur with more than 4 AV secondary setae (Fig. 61); urogomphus composed of one urogomphomere; U/HW = 0.40; U/LAS> 0.50; Palaearctic.
Description, instar III (Figs 58–62, 65)
Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.
Head (Figs 58, 60, 65): Head capsule subquadrate, strongly constricted at level of occiput; anterior margin of frontoclypeus broadly convex, extending mesally at about level of adnasalia; scale-like sculpticels lacking over frontoclypeus and parietale; adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); HL = 1.36–1.39 mm; A/MP = 1.16–1.27; MP/LP = 1.67–1.74; MP2/MP1 = 1.15–1.28; GA/MP1 = 1.27–1.49; LP2/LP1 = 1.54–1.71; MNL/MNW = 2.63–2.71.
Thorax (Figs 61–62): L3 = 2.86–3.06 mm.
Abdomen (Fig. 59): LAS = 1.09–1.17 mm; LAS subconical, constricted posteriorly at point of insertion of urogomphi. Urogomphus, U = 0.60–0.63 mm, composed of one urogomphomere; U/HW = 0.38–0.40; U/LAS = 0.54–0.57.
Chaetotaxy: Parietale with 7–10 temporal spines; position and number of secondary setae on legs are shown in Figs 61–62 and Table 7.
description: Published as part of Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong & Watts, Chris H. S., 2022, An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae), pp. 151-205 in Zootaxa 5175 (2) on pages 193-197, DOI: 10.11646/zootaxa.5175.2.1, http://zenodo.org/record/7003344
Trochosa C. L. Koch 1847
Creators: name: Marusik, Yuri M.
name: Nadolny, Anton A.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Lycosidae
Trochosa

description: Trochosa C.L. Koch, 1847
Trochosa C.L. Koch, 1847: 95; Engelhardt 1964: 224; Fuhn & Niculescu-Burlacu 1971: 221; Dondale & Redner 1990: 22; Zyuzin 1990: 422; Almquist 2005: 246; Tanaka 1988: 94.
Type species: Aranea lupus ruricola De Geer, 1778.
Diagnosis. Trochosa resembles Pirata Sundevall, 1833 and the related genera Piratula Roewer, 1960 and Trebacosa Dondale & Redner, 1981 by the carapace pattern: a pair of longitudinal stripes in the anterior part of a light median band. However, the genus can be easily separated from the others due to its large size, and the parallel longitudinal stripes in Trochosa (Fig. 1) which are converging posteriorly in Pirata and related genera (see Omelko et al. 2011). Trochosa can be differentiated from the genera mentioned before by the copulatory organs, which are similar to those of Hogna Simon, 1885. Members of Trochosa and Hogna differ from other genera by having a triangular tegular apophysis (ventral view), an elliptically curved embolus with a basal transparent membrane, a pointed synembolus, and the epigyne has an anchor-shaped septum and a pair of deep anterior hoods. Trochosa can be differentiated from Hogna by: 1) the lack of strong macrosetae on the cymbial apex (vs. present); 2) the epigyne having a small fovea, and the septal stem covering more than half of the fovea (in Hogna, the fovea is larger, equal to three times the width of the septal stem); 3) the width of the septal base is equal to the length of the septum (in the West Palaearctic, Hogna ’s septum length is 1.2–1.4 times longer than the width of the septal base); 4) the septal stem widens anteriorly (in Hogna, the septum edges are parallel).
Comments. A search for published figures and types of species described by Roewer (1955a) reveals that some arachnologists are not aware of the key characters of the genus. For example, Roewer (1955a) described T. hispanica under three different names in three different genera but none of them in Trochosa. Simultaneously, he placed a species, Hippasa loeffleri (Roewer, 1955a), belonging to a different subfamily, as well as Arctosa tbilisiensis Mcheidze, 1946 (= Trochosa impercussa Roewer, 1955a) in Trochosa. In the revision of Australian Trochosa, McKay (1979) treated 12 species, although 11 of them are now transferred to other genera. The remaining Trochosa glarea McKay, 1979, known only from the female, has an epigyne that differs from those of the generotype.
description: Published as part of Marusik, Yuri M. & Nadolny, Anton A., 2020, On the identity of Trochosa hispanica (Araneae, Lycosidae), with notes on the synonymy of West Palaearctic " Trochosa " species, pp. 56-80 in Zootaxa 4859 (1) on page 58, DOI: 10.11646/zootaxa.4859.1.2, http://zenodo.org/record/4412755
Arctosa tangerana Marusik & Nadolny 2020, comb. n.
Creators: name: Marusik, Yuri M.
name: Nadolny, Anton A.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Lycosidae
Arctosa
Arctosa tangerana

description: Arctosa tangerana (Roewer, 1960) comb. n.
Fig. 17 F–G
Trochosina tangerana Roewer, 1960a: 909, figs 504a–b (♀).
Trochosa tangerana: World Spider Catalog 2020.
Comments. This species, 11 mm long, was described based on the holotype female from Tangier (Morocco) (Roew- er 1960a). Habitus and epigyne figures are present with the description. The carapace is uniformly coloured (a character unknown in Trochosa), lacking a pair of longitudinal stripes (present in all Trochosa) and the epigyne has a large fovea and lacks the distinct, deep anterior hoods characteristic of Trochosa. The shape of the epigyne is similar to that of Arctosa villica (Lucas, 1846) as illustrated in Nentwig et al. (2020) and Oger (2020), and A. villica is known from Morocco. Given the patternless carapace, the shape of the epigyne and the occurrence in Morocco, it is most likely that the two species names could be synonymous. Because of the similarity of Arctosa villica and Trochosa tangerana, we transfer the latter species to Arctosa.
description: Published as part of Marusik, Yuri M. & Nadolny, Anton A., 2020, On the identity of Trochosa hispanica (Araneae, Lycosidae), with notes on the synonymy of West Palaearctic " Trochosa " species, pp. 56-80 in Zootaxa 4859 (1) on page 70, DOI: 10.11646/zootaxa.4859.1.2, http://zenodo.org/record/4412755
Alopecosa garamantica Marusik & Nadolny 2020, comb. n.
Creators: name: Marusik, Yuri M.
name: Nadolny, Anton A.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Lycosidae
Alopecosa
Alopecosa garamantica

description: Alopecosa garamantica (Caporiacco, 1936) comb. n.
Fig. 16 A–C
Pirata garamantica Caporiacco, 1936: 89, fig. 6 (♀). Arkalosula garamantica: Roewer 1955b: 231.
Trochosomma garamantica: Roewer 1960a: 853, figs 474a–b (♀ transferred from Arkalosula = Arctosa).
Trochosa garamantica: World Spider Catalog 2020.
Comments. This species was described based on several females collected in Fezzan (Southwestern Libya). Caporiacco indicated its size as 7 mm (3 mm carapace and 4.1 mm abdomen) in length. Judging from the shape of the epigyne given in Caporiacco (1936) and Roewer (1960a), it is neither Pirata nor Trochosa but is most similar to Alopecosa simoni (Thorell, 1872) and A. pentheri (Nosek, 1905), members of the A. cursor -group (see Marusik et al. 2018; Nentwig et al. 2020) and because of this similarity, we transfer this species to Alopecosa.
description: Published as part of Marusik, Yuri M. & Nadolny, Anton A., 2020, On the identity of Trochosa hispanica (Araneae, Lycosidae), with notes on the synonymy of West Palaearctic " Trochosa " species, pp. 56-80 in Zootaxa 4859 (1) on pages 69-70, DOI: 10.11646/zootaxa.4859.1.2, http://zenodo.org/record/4412755
Hogna ferox
Creators: name: Marusik, Yuri M.
name: Nadolny, Anton A.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Lycosidae
Hogna
Hogna ferox

description: Hogna ferox (Lucas, 1838)
Fig. 14
Geolycosa altera Roewer, 1955a: 764, fig. 12 (♀). Syn. n.
Geolycosa atroscopulata Roewer, 1955a: 765, figs 14–16 (♂ ♀). Syn. n.
Lycorma nigrichelis Roewer, 1955a: 767, figs 17–19 (♂ ♀). Syn. n.
Hogna radiata: Mozaffarian & Marusik 2001: 69 (synonymized with G. altera without examination of the type).
Hogna ferox: Wunderlich 1992: 458, figs 716–719 (♂ ♀); Hepner & Paulus 2009: 341, figs 14–16 (♂ ♀).
Trochosa altera: Logunov 2012: 355, figs 20–21 (examined type, transferred from Geolycosa).
Hogna ferox: Lecigne 2016b: 104, figs 7L–O, 11J–K (♂).
For the complete list of taxonomic references see World Spider Catalog (2020).
Types. Geolycosa atroscopulata Roewer, 1955 (Fig. 14 A–G): lectotype ♂ and paralectotypes 2♂ 1♀ designated here (SMF), “Arachn. Coll. Rwr. -Lfd. No. 11441, No. 703, Typ., Iran: Sabzawaran”, examined (in the text Roewer mentioned this material as: 1♂ 1♀, Sabzawaran, Typus, H. Löffler leg., 2♂, Sabzawaran, Paratipoide, F. Starmuhl- ner leg.); 1♀ (SMF), “Sabzawaran (nördl. Flussbett), Paratypoid”, H. Löffler leg., not examined.
Geolycosa altera Roewer, 1955: 1 ♀ (SMF), “bei Tschalus (Kaspi-Küste), Typus ”, H. Löffler leg., not exam- ined.
Lycorma nigrichelis Roewer, 1955 (Fig. 14 H–I): holotype ♀ (SMF), “Arachn. Coll. Rwr. -Lfd. No. 11444, No. 706, Typus, Kurusch-Gol (Tabris)”, H. Loffler leg., examined; paratype 1♂ (SMF), “ Allotypus, bei Tschalus (Kaspi- Kuste)”, F. Starmuhlner leg., not examined.
Comments. This species is considered here because Geolycosa altera was recently (Logunov 2012; World Spi- der Catalog 2020) placed in Trochosa. Although we have not studied the type, this species is adequately illustrated by Logunov (2012). Its peculiarly shaped vulva is exactly the same as that of H. ferox. The female allotype of G. atroscopulata is a subadult with a well-developed ‘pre-epigyne’. The structure of the epigyne and habitus in the type of Lycorma nigrichelis are identical to those of H. ferox.
description: Published as part of Marusik, Yuri M. & Nadolny, Anton A., 2020, On the identity of Trochosa hispanica (Araneae, Lycosidae), with notes on the synonymy of West Palaearctic " Trochosa " species, pp. 56-80 in Zootaxa 4859 (1) on page 72, DOI: 10.11646/zootaxa.4859.1.2, http://zenodo.org/record/4412755
Arctosa annulipes Marusik & Nadolny 2020, comb. n.
Creators: name: Marusik, Yuri M.
name: Nadolny, Anton A.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Lycosidae
Arctosa
Arctosa annulipes

description: Arctosa annulipes (L. Koch, 1875) comb. n.
Fig. 17 A–D
Trochosa annulipes L. Koch, 1875: 77, pl. 7, fig. 4 (♀); World Spider Catalog 2020.
Tarentula annulipes: Strand 1908: 319.
Arctosa annulipes: Caporiacco 1941: 44; Denis 1966: 128, fig. 45 (♀).
Trochosomma annulipes: Roewer 1955b: 304; Roewer 1960a: 852, fig. 473a–b (♀).
Comments. This species was described based on a female from Cairo (Egypt). Its length is reported as 7 mm (3.5 mm carapace and 3.5 mm abdomen). The habitus figures in L. Koch (1875) and Roewer (1960a) show a uniformly colored carapace lacking the pair of parallel stripes within the median light band, which is present in all Trochosa species occurring in Europe. The figures of the epigyne provided by Roewer (1960a) and Denis (1966) are very different. Denis (1966) mentioned that Roewer’s figures are too schematic and poorly reflect the actual shape of certain structures. We faced the same problems studying the types of Roewer’s species described from Iran, the figures of the epigyne provided by Roewer are rather different from studied types (for instance, Figs 12 C–C’, E–E’, F–F’). Judging from the carapace pattern and figures of the epigyne given by Denis (1966), this species belongs to Arctosa; therefore, we transfer this species from Trochosa to Arctosa. Trochosa annulipes L. Koch, 1875 is the type species of Trochosomma Roewer, 1960 syn. n. which is currently considered a synonym of Trochosa (World Spider Catalog 2020) but should be considered a synonym of Arctosa C.L. Koch, 1847.
description: Published as part of Marusik, Yuri M. & Nadolny, Anton A., 2020, On the identity of Trochosa hispanica (Araneae, Lycosidae), with notes on the synonymy of West Palaearctic " Trochosa " species, pp. 56-80 in Zootaxa 4859 (1) on page 70, DOI: 10.11646/zootaxa.4859.1.2, http://zenodo.org/record/4412755
Pardosa persica Marusik & Nadolny 2020, comb. n.
Creators: name: Marusik, Yuri M.
name: Nadolny, Anton A.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Lycosidae
Pardosa
Pardosa persica

description: Pardosa persica (Roewer, 1955) comb. n.
Fig. 13 A–C
Dingosa persica Roewer, 1955a: 762, fig. 11 (♀).
Trochosa persica: Mozaffarian & Marusik 2001: 70; Framenau & Baehr 2007: 1626.
Type: holotype ♀ (SMF), “Arachn. Coll. Rwr. -Lfd. No.10467, No.469, Typ., Iran: Lahidschan”, F. Starmuhlner leg., examined.
Other material examined: IRAN: 2♀ (ZMUT), Khuzestan, #2251, Abadam-B-Emam, 12 KE Abadam Rd., 29.03.1999 (K. Elmi); 1♀ (ZMUT), #475, collecting locality and collector unknown.
Comment. This species was transferred from Dingosa Roewer, 1955, a genus known from Australia, to Trochosa (Mozaffarian & Marusik, 2001) because the epigyne drawn by Roewer (1955a: fig. 11) corresponds well to Trochosa. Examination of the type revealed that the epigyne was inaccurately drawn (cf. Figs. 13B and 13C) and that the holotype female has a pattern typical to Pardosa s.l. Judging from the epigyne, it may belong to the Pardosa nebulosa -group. This group was once considered a separate genus, Acroniops Simon, 1898, currently a junior synonym of Pardosa C.L. Koch, 1847. The new combination provided for Roewer’s species led to secondary homonymy with Pardosa persica Marusik, Ballarin & Omelko, 2012, the species belonging to the Pardosa monticola - group that was also once considered a separate genus, Pardosops Roewer, 1955. In any case, we herein replace the secondary junior homonym with P. persiana nom. n. (pro Pardosa persica Marusik, Ballarin & Omelko, 2012).
description: Published as part of Marusik, Yuri M. & Nadolny, Anton A., 2020, On the identity of Trochosa hispanica (Araneae, Lycosidae), with notes on the synonymy of West Palaearctic " Trochosa " species, pp. 56-80 in Zootaxa 4859 (1) on page 72, DOI: 10.11646/zootaxa.4859.1.2, http://zenodo.org/record/4412755
Trochosa adjacens O. Pickard-Cambridge 1885
Creators: name: Marusik, Yuri M.
name: Nadolny, Anton A.
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Lycosidae
Trochosa
Trochosa adjacens

description: Trochosa adjacens O. Pickard-Cambridge, 1885
Fig. 10C
Trochosa adjacens O. Pickard-Cambridge, 1885: 83 (♀).
Type: holotype ♀ in OUMNH, vial “## faded label, 1680”. Information in the text “Yangihissar, April 1874 ” (= China, Xinjiang).
Note. This species was recognized among the “Yarkand spiders”, a collection lacking species or geographical labels, due to its size, “over 5 lines” (~ 10.5 mm), and pale coloration. In the description, O. Pickard-Cambridge (1885) compared this species with T. terricola Thorell, 1856 and T. ruricola, the generotype.
Description. See O. Pickard-Cambridge (1885). Holotype faded. Measurements of the holotype ♀: total length 11.0; carapace 5.0 long, 4.0 wide; Leg I 11.7 (3.3, 1.9, 2.6, 2.3, 1.6).
Comments. The shape of the epigyne indicates that this species belongs to Trochosa, unlike other species placed by O. Pickard-Cambridge (1885) in this genus. The relationship of T. adjacens with other congeners remains unclear. Most, if not all, females in the genus Trochosa cannot be distinguished from each other, especially singletons. To correctly classify this species, males are required from the type locality.
Distribution. This species is known from the type locality only, Yangihissar Town in SW Xinjiang.
description: Published as part of Marusik, Yuri M. & Nadolny, Anton A., 2020, On the identity of Trochosa hispanica (Araneae, Lycosidae), with notes on the synonymy of West Palaearctic " Trochosa " species, pp. 56-80 in Zootaxa 4859 (1) on page 73, DOI: 10.11646/zootaxa.4859.1.2, http://zenodo.org/record/4412755