Erymanthus pustulosus Quedenfeldt 1888
Creators: name: Gerstmeier, Roland
name: Salvamoser, Max
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Cleridae
Erymanthus
Erymanthus pustulosus

description: Erymanthus pustulosus Quedenfeldt, 1888
(Figs 26–27, 40 D)
Quedenfeldt 1888: 182.
Erymanthus variegatus (Pic 1950), n. syn. Pic 1950: 160.
Specimens examined: Erymanthus pustulosus: Lectotype (designated here): Kassai, 4/9, Eurymanthus pustulosus Af. c. Qdf, Ex Musaeo Quedenfeldt, Muséum Paris 1952, Coll. R. Oberthúr, Holotype, pustulosus H. Type (MNHN). Erymanthus variegatus: Lectotype (designated here): Holotypus variegatus Pic, Coll. Mus. Congo, Ruanda: Kibungu, X-XII.1937, R. Verhulst, R. Det. FF. 5621, désiré, Erymanthus variegatus M. Pic (MRAC).
Comment on type specimens: Neither Quedenfeldt (1888: 182) nor Pic (1950: 160) mentioned number of examined specimens, therefore all type material are syntypes (ICZN 1999: Article 73.2) and available for lectotype designation (ICZN 1999: Article 74.1) as it cannot be assumed that these taxa are based on a single specimen. The specimens from MNHN and MRAC are designated lectotypes.
Others: Dahomey, Muséum Paris, Coll. M. Pic (MNHN). Percy Fyfe Nat. Res., Potgietersrus Dist., 18.– 20.1.1971, R. Jones, Eurymanthus pustulosus (Quedenfeldt), det. G. Ekis 1985 (MNHN). Côte d'Ivoire, Lamto, 10.V.1982, Leroux Rec., Muséum Paris, lumière (MNHN). Côte d'Ivoire, Dimbokro, Museum Paris, Le Moult. 1916 (MNHN). Senegal, Ex Musaeo Maiszech, Muséum Paris 1952, Coll R. Oberthur (MNHN). Dahomey, Muséum Paris, Coll. M. Pic (MNHN). Beira Port. E. A., 28.9.1899, G. A. K. Marshall. 1908-212, Erymanthus horridus Westw., Erymanthus pustulosus Qued. ♂, J. J. Menier det. 1981 (BMNH). van Someren, Garissa Bura, Tana Riv: 11 48., V. G. L. van Someren Collection, Brit. Mus. 1959-468. (BMNH). N. W. Rhodesia, Kashitu, N of Broken Hill, Nov. 1914, H. C. Dollman, H. C. Dollman Coll. 1919-79, J. B. Corporaal det. 1928, Erymanthus pustulosus Qued., Erymanthus pustulosus Qued. ♀, J. J. Menier det. 1981, tree trunk (BMNH). Br. Somaliland: W. A. Macfadyen. B. M. 1929-398, ex BMNH, Erymanthus pustulosus, J. J. Menier det. 1981, ♂ (BMNH). Rhodesia, J. A. W. Lucas, St. Luke's Hopital, ca 30km S E Lupane, 17-12-1975 (RMNH). N. W. Rhodesia, Kashitu, N of Broken Hill, Jan. 1915, H. C. Dollman, H. C. Dollman Coll. 1919-79, J. B. Corporaal det. 1928, Erymanthus pustulosus Qued., ex Brit. Museum 1928, tree trunk (BMNH). Natal, Popp, 17814 (ZMB). Kamerun, Sina, L. Colin. IV.12 (ZMB). Zimbabwe, 3.xii.1993, 20°13'S / 31°00'E, Kyle Rec. Park at Lake Mutirikwe, lux, leg. J. Deckert (ZMB). Namibia: b. Omaruru, Farm Otjua, 5.–7.X.1991, leg. U. Göllner (ZMB). Südafrika: Transvaal, 24°05' S / 30°15' E, Lekgalameetse Nat. Res., 21.xii.1995, leg. F. Koch (ZMB). Central Malawi. Lilongwe Dist. Dzalanyama forest 1300m, 14-29 S–33-39 E, XI-XII. 2005/I. G. 31.593, Don C. Joly, Coll. I. R. Sc. N. B. (ISNB). Kenya, Eastern, E of Thika 29.12., Mwingi, Nguni, Lgt. Snížek 2007, Coll. I. R. Sc. N. B. (ISNB). Kenya SE 12.12. Voi 2007, S Foot Sagala, Lgt. Snížek Mts., Coll. I. R. Sc. N. B. (ISNB). Kenya SE 12.12. Voi 2007, S Foot Sagala, Lgt. Snížek Mts., Coll. I. R. Sc. N. B. (ISNB). Kenya, 9/ 10.V.2000, Hola, Werner & Lizler leg. (SMNS). Kenya XII.93, Sagala Hills, Werner leg. (SMNS). Kenya, 12.V.2001, E. Prov., Hola, Werner & SMRZ leg.
(SMNS). Namibia, Popa Falls, Kavango, 24.xi.1993, 18°07' S / 21°35' E, leg.: F. Koch (SMNS). RSA: Limpopo Prov., Lekgalameetse Nature Reserve, 800-1000 m, 27.XI.–1.XII.2008, leg. W. Schawaller (SMNS). RSA, KwaZulu-Natal, Ndumo NR, Campside, 90 m, 10.II.–13.II.2010, leg. D. Bartsch & Berg (SMNS). S. Afr.; Mpumalanga, Ezemvelo, 1374m, 25.42 S – 29.01 E, 27.1.2004; E-Y: 3608, vegetation, highveld, leg. TMSA Staff (TMSA). Zimbabwe, MasvingoPro, LakeMutirikweLake Kyle, 20.13 S – 31.00 E, 27.11.2006; E-Y: 3736, light trap, leg. Gussmann, Müller (TMSA). Zimbabwe, MasvingoPro, LakeMutirikweLake Kyle, 20.13 S – 31.00 E, 27.11.2006; E-Y: 3736, light trap, leg. Gussmann, Müller (TMSA). S. Afr.; Mpumalanga, Skukuza, 25.00 S – 31.35 E, 26.11.– 5.12.1993, leg. Krüger, Dunning (TMSA). S. Afr.; Krüger Nat. Pk, Skukuza, 25.00 S – 31.35 E, 16.– 24.12.1993, light trap, Krüger, Dunning (TMSA). S. Afr.; Gauteng, Cullinan, Windy Brow, 25.41 S – 28.30 E, 8.1.2002; E-Y: 3466, groundtraps, leg. TMSA staff (TMSA). S. Afr.; North Wes Prov., Tswaing, 25.24 S – 28.06 E, December; M. Krüger (TMSA). S. Afr.; North West Prov., Tswaing, 25.24 S – 28.06 E, December; M. Krüger (TMSA). S. Afr.; Gauteng Prov, Tswaing, 25.24 S – 28.06 E, 24.11.2003; E-Y: 3614, light trap, leg. TMSA staff (TMSA). S. Afr.; Waterberg, Geelhoutbush farm, 24.22 S – 27.33 E, 7.12.1998: E-Y: 3251, at light, leg. CL Bellamy (TMSA). S. Afr.; Waterberg, Dorndraai Lodge, c. 24.38 S – 27.48 E, 18.– 22.12.2004, leg. M. Krüger, (TMSA). S. Afr.; Limpopo Prov., Soutpan, near Vivo, 10.1.2009; leg. T. Beyers (TMSA). S. Afr.; Mpumalanga Prov, Der Brochen, 25.04 S – 30.07 E, 7.12.2005, at light, leg. James Harrison (TMSA). Elisabethville, Novbre 1948, Ch. Seydel, C19866, Coll. Mus. Congo, R. Det. H. 5763, Erymanthus horridus West. (MRAC). South Africa, KZN Tembe Elephant Res., 27.03S 32.25E, 100m, 28.x.–02.xi.1995, M. W. Mansell, light trap, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, TVL, D'Nyala Nat. Res., Ellisras District, 23.45S 27.49E, 18.–20.xii.1987, B. Grobbelaar, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, TVL, D'Nyala Nat. Res., Ellisras District, 23.45S 27.49E, 17.–20.xii.1987, M. W. Mansell, Light, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, TVL, D'Nyala Nat. Res., Ellisras District, 23.45S 27.49E, 850m, 24.–26.ix.1990, R. G. Oberprieler, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, TVL, D'Nyala Nat. Res., Ellisras District, 23.45S 27.49E, 18.–20.xii.1987, B. Grobbelaar, Erymanthus det. J. R. Mawdsley 2008, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, TVL, D'Nyala Nat. Res., Ellisras District, 23.45S 27.49E, 18.xii.1987, B. Grobbelaar, E. v. d. Linde, Collected at light, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, MPU, Kampersrus, near Hoedspruit, 24°30'S 30°53'E, 10.xi. 2005, 547m, W. Breytenbach, Collected using a light trap, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, TVL, Kruger Nat. Park, Renosterkoppies, 25°07'S 31°36'E, Collected at light, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, TVL, Loskopdam Nat. Res., 25°25'S 29°20'E, 12.–13.xii.1985, C. G. E. Woolman, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, TVL, Loskopdam Nat. Res., 25°25'S 29°20'E, 12.–13.xii.1985, I. M. Millar, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, TVL, Pretoria, Garden of Union Buildings, 25°45'S 28°12'E, 25.II.1982, NC Grobbelaar, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, Natal, Mkuze, 27°37'S 32°03'E, XII.1981, P. Reavell, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, TVL, Soutpan, Pretoria Dist., 25°24'S 28°06'E, 09.xii.1987, M. W. Mansell, Collected at light, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, TVL, Soutpan, Pretoria Dist., 25°24'S 28°06'E, ii.1985, M. W. Mansell, Collected at light, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa, TVL, Swadini, Blydepoort Nat. Res., 24°32'S 30°54'E, 26.–29.i.1987, B. Grobbelaar, Collected at light, National Coll. of Insects, Pretoria, S. Afr. (SANC). South Africa: Natal, Umhlanga Rocks, 3106E 2944S, XI.83, P. E. Reavell, 10m, light, National Coll. of Insects, Pretoria, S. Afr. (SANC). Swa/ Namibia, Caprivi, Katima Mulilo, SE17 24 Ad 10–11.I.1985, Univ. Pretoria, Dept. Entomol. Tour '85, National Coll. of Insects, Pretoria, S. Afr. (SANC). N Rhodesia, 16.XI.1941, Dr. W. Eichler, 15, ex coll. Witold Eichler, Inst. Zool. P. A. N. Warszawa 64/63 (ZMPA). Kenya: central-south, Kibwezi Forest, Spring Camp, 28.– 29.11.2004, leg. Puchner (CRSW). Kenya: central-south, Kibwezi Forest, Spring Camp, 28.– 29.11.2004, leg. Puchner (CRSW). Ethiopia, Oromia, 17 km SEE Yabelo, N04°36', N04°51' 1580m, 17.5.12, leg. Egger M. (CMEW). St. Lucia, Natal 1/87, Werner leg., coll. R. Gerstmeier CRG Munich (RGCM). Somalia, Afgoi, IV.1978, A. Simonetta! Museo ''La Specola'' ex coll. J. Simonetta Num. Mag. 2812 (MZUF). Kenya, 12.1993, Sagala Hills, leg. K. Werner, coll. R. Gerstmeier CRG Munich (RGCM). RSA, KwaZulu-Natal, SW Magudu, 27°34'S –31°35W, 4.– 5.1.2009, P. Schüle leg., coll. R. Gerstmeier CRG Munich (RGCM). Coll. I. R. Sc. N. B., Congo Belge, Mwadingusha, XI– XII–1957, leg. N. Magis (RGCM). Tanzania, 16.xii.1999, near Mitundo, Dodoma prov., Werner + Lizler lgt., Jakub Rolčík Collection Prague (JRCP). Tanzania, 16.xii.1999, near Mitundo, Dodoma prov., Werner + Lizler lgt., Jakub Rolčík Collection Prague (JRCP). Tanzania, 16.xii.1999, near Mitundo, Dodoma prov., Werner + Lizler lgt., Jakub Rolčík Collection Prague (JRCP).
Tanz.- Dodoma pr., 60km N Kondoa, S 04°36' E 35°46', 1570m, 19.12.2006, J. Halada lgt., Jakub Rolčík Collection Prague (JRCP). Zambie, Liwingstone, 24.12.2002, Smrž P. lgt., Erymanthus horridus Westw., J. Rolčík det. 2004, Jakub Rolčík Collection Prague (JRCP). Zambie, Liwingstone, 24.12.2002, Smrž P. lgt., Erymanthus horridus Westw., J. Rolčík det. 2004, Jakub Rolčík Collection Prague (JRCP). Zambia - SW, 30km NW Kazangula, 23.XII.2002, J. Halada leg., Jakub Rolčík Collection Prague (JRCP). RSA, Maphutoland, SW of Emangusi, 29.1.2003, M. Snižek lgt., Jakub Rolčík Collection Prague (JRCP). RSA, Noth. Transvaal, Kruger NP, Punda Maria, xii.2005, J. Hlásek lgt., Jakub Rolčík Collection Prague (JRCP). RSA, Noth. Transvaal, Kruger NP, Punda Maria, xii.2005, J. Hlásek lgt., Jakub Rolčík Collection Prague (JRCP). S. Africa, S 25.46 E 27.54, 25km W Pretoria, Saartjiesnek, S. Bílý leg. 16.– 18.12.1997, Jakub Rolčík Collection Prague (JRCP). Swaziland, 20.– 21.xii.2003, Mlawula Game Res., Werner & Lizler lgt., Jakub Rolčík Collection Prague (JRCP). Zimbabwe c.9.– 11.12., Mushandike Sanct (Masvingo env.) 1998, S. Bečvář lgt., (RGCM). Zimbabwe, centr, 20kmW Ngezi Park, Kwekwe, 30.11.1998, leg. M. Halada, Jakub Rolčík Collection Prague (JRCP). Kenya, Hola, 9.– 10.5.2000, Werner lgt., Jakub Rolčík Collection Prague (JRCP). Kenya, Sagala Hills, XI.1993, Werner leg., Jakub Rolčík Collection Prague (JRCP). Kenya, XI.93, Sagala Hills, Werner leg., Jakub Rolčík Collection Prague (JRCP). Zambia NW, 90 km Solwezi E of Chisasa, 2005 18.XI., Snížek lgt, Collection Vladimír Beneš (JRCP). Kenya, Sagala Hills, XI.1993, Werner leg., Collection Vladimír Beneš (JRCP). Kenya SE, 12.12. Voi 2007, s Foot Sagala, lgt. Snížek Mts., Collection Vladimír Beneš (RGCM). Kenya, Eastern, E of Thika 29.12., Mwingi, Nguni, Lgt. Snížek 2007, Collection Vladimír Beneš (JRCP). Kenya, Eastern, E of Thika 29.12., Mwingi, Nguni, Lgt. Snížek 2007, Collection Vladimír Beneš (JRCP). Kenya, Coast, N of Garsen, 6.12.2007, lgt. Snížek, Collection Vladimír Beneš (JRCP). Zambia NW, E Mutumbwe (NW Kasempa), 15.XI.2005, Snížek, Collection Vladimír Beneš (JRCP). Zambia NW, 80 km SSW of Solwezi, 13.XI.2005, Snížek lgt., Collection Vladimír Beneš (JRCP). Zambia, o. Northern prov., Chipona fals, 5.–8.xi.2002, 30km of Chinaséi, F. a. L. Kantner lgt., Collection Vladimír Beneš (JRCP).
Diagnosis: Erymanthus pustulosus can be confused with E. revoili, E. ater and E. horridus. Erymanthus revoili has the pronotal gibbosities conical and a conspicuous rasp-like coarse punctation. In Erymanthus ater the antennomeres 9–11 are longer than the antennomeres 1–8. Erymanthus horridus has a longitudinal ridge in the pronotal groove, a conical phallic tip and strongly pointed tips of ventrite VI. Erymanthus pustulosus is recognizable by the characteristic maculation of the pronotum and the elytra. Erymanthus variegatus is treated as a synonym because we observed no differences in external morphology of genitalia.
Length: (59 specimens measured) 11–20mm (average 14.69mm).
Head: Black, antennae brown to black; frons with two gibbosities; with anteriorly directed flavotestaceous and black pilosity; posterior gibbosity with fine punctation; antennomere 11 elongate.
Pronotum: Pronotal length to width ratio 0.86:1 to 1.16:1 (average 1.07:1); black with a characteristic brown maculation (Fig. 40 D); pronotal groove shallow, gibbosities conspicuously elevated; with isolated flavotestaceous and black pilosity; with isolated fine punctation.
Scutellum: Brown; vested with flavotestaceous setae.
Elytra: Elytral length to width ratio 2.07:1 to 2.67:1 (average 2.25:1); dark brown to black, with light brown and brown maculation; punctation widely distributed, irregular, brown; in the middle with fine punctation, lateral punctation coarse, punctation circular, width of interstices variable; with isolated brown pilosity; gibbosities yellow to brown, all gibbosities with black setal tuft.
Legs: Coxae, femora and tibiae black.
Lower surface: Black; dense fine punctation; with flavous pilosity.
Male genitalia, pygidium, ventrite VI: Tips of tegminal lobes blunt, tegminal lobes one-fourth length of tegmen; tegminal struts broadened after one-eighth of length and tapering after three-sevenths, broad furcate tip; phallic tip blunt; posterior side of pygidium rounded; processes of ventrite VI rather blunt.
Distribution: Senegal, Sierra Leone, Ivory Coast, Benin, Cameroon, Ethiopia, Somalia, Kenya, DR Congo, Rwanda, Tanzania, Zambia, Mozambique, Zimbabwe, Namibia, Swaziland, South Africa.
Seasonal occurrence: Collected from January to December.
description: Published as part of Gerstmeier, Roland & Salvamoser, Max, 2014, Revision of the checkered beetle genus Erymanthus Spinola, 1841 (Coleoptera, Cleridae, Clerinae), pp. 501-548 in Zootaxa 3755 (6) on pages 527-531, DOI: 10.11646/zootaxa.3755.6.1, http://zenodo.org/record/251514