Clivina tuberculifer Balkenohl, 2025, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina tuberculifer

description: Clivina tuberculifer sp. nov.
(Figs 8, 43, 78, 109)
Type material. Holotype: ♂, with labels and data: white, black printed “ NEPAL East, Kosi Zone, Dhankutta distr., Arun valley, Dharan env., 500 m, 22-23.v.1996, P. Čechovský lgt.” / “ Clivina sp. ad. wallacei DET. DR.A. DOSTAL 2011 ” (CADW).
Paratype: 1 ♀, same data as holotype (CADW).
Description. Measurements. Body length 4.85 / 4.96 mm, width 1.45 / 1.53 mm, ratio L/W of pronotum 0.90 / 0.91, ratio L/W of elytra 1.78 / 1.79. Colour glossy, fuscous. Supraantennal plate slightly brightened. Antennae, mouthparts, intermediate and hind legs yellowish.
Head less than a quarter narrower than pronotum. Clypeus straight anteriorly, with its lateral teeth slightly projecting; wing of moderate size, not more protruding anteriorly than clypeus, distinctly separated from clypeus by notch; supraantennal plate convex laterally, slightly convex in anterior half; clypeus, wing, and supraantennal plate finely reflexed margined. Supraantennal plate extended up to mid-eye level, ending at mid-eye level as blunt knob, separated from wing by distinct notch, separated from supraorbital carina by deep moderately wide furrow. Clypeus transverse, slightly convex, separated from frons by flat furrow. Frons moderately convex, with central longitudinal pore at middle, with scattered small punctures. Supraantennal plate separated from clypeus and frons by moderately wide furrow continuing posteriorly up to mid-eye level as moderately wide supraorbital furrow. With blunt distinct supraorbital carina at posterior half of eye. With two supraorbital setigerous punctures situated at mid-eye level and posterior gena-level. Neck constriction developed by punctures and distinct step. Eye convex, projecting laterally but not hemispherical. Gena distinct, enclosing eye posteriorly. Antenna moderately long, not reaching basal carina of pronotum, antennomeres four to ten moniliform (L/W around 1.06). Labrum nearly straight, isodiametrically reticulated, seven-setose. Mandible of moderate size. Mentum with lateral lobe pentagonal with rounded angles, with irregular and oblique wrinkles, median tooth spatulate, slightly higher as lateral lobes.
Pronotum (43) with disk somewhat flattened but indistinctly convex in lateral view, in frontal view flattened at middle and moderately convex laterally. Quadrate, wider than long. Anterior margin nearly straight. Reflexed lateral margin indistinctly sub-crenulate, regularly moderately convex, widest at middle; anterior angle rounded-off; posterior angle rounded, marked by obtuse tooth. Width of lateral channel medium sized, reticulated, basal part of lateral margin from posterior angle to flange of base slanted with angle of around 20°, convex, flange short but distinct, basal channel moderately developed, basal carina fine but sharp. Median line narrow, crenulated, joining anterior transverse line and base; anterior transverse line narrow, complete. Surface with microscopic punctures, with scattered medium sized punctures, base isodiametrically reticulated.
Elytron (Fig. 8) with disk flattened in anterior half in lateral view, regularly convex in frontal view. Outline regularly long oval, distinctly less than twice as long as wide, with maximum width about at middle. Humerus rounded, formed by lateral margin which joins stria five laterally by a short carina. Reflexed lateral margin nearly smooth posterior humerus. Lateral channel moderately wide. Scutellar striole of moderate length, not deep; with distinct setigerous puncture at base of first stria, with conspicuous tubercle at base of third interval. Striae moderately deep, punctate-striate, one to four free at base, five and six joining at humerus, seven shortened behind humerus, striae one and two running up to apex, three and four, and five and six just not joining apically. Intervals slightly convex, seven and eight carinate apically. Third interval with four setigerous punctures adjoining third stria. Surface of intervals smooth and glossy on disk, with distinct isodiametric reticulation at base, at apical tip, and on interval seven and eight.
Hind wing fully developed.
Lower surface. Proepisternum with distinct isodiametric reticulation, with some fine transverse wrinkles laterally. Sternites of abdomen with shingle-like reticulation, transverse at middle. Abdominal sternum VII with the two anal setigerous punctures widely separated, at middle slightly projecting posteriorly where it is flattened, with narrow marginal furrow at middle (in both sexes).
Legs. Protibia with three spines of moderate length and a small knob basally, distinctly sulcate dorsally, movable spur moderately arcuate, surface with intricate reticulation. Mesotibia with distinct and moderately long protuberance preapically with acuminate apex and with its seta inserted laterally, with eight setae furnished tubercles above protuberance, with few fine setae at the inner side. Front leg with tarsomeres slightly widened, first tarsomere elongated, slender and with carina baso-laterally, laterally with acute tubercle furnished with four short setae.
Male genitalia (Fig. 78). Median lobe slender, moderately convex, more curved at root of apical lamella, apex with flattened wide lamella. Parameres moderately slender, with three setae at apex.
Female gonocoxites and epipleurite (Fig. 109). Gonocoxites scapiform, extraordinary slender, less sclerotized, not fused, completely movable; basal gonocoxite 1 elongated, with four setae, apical gonocoxite 2 slender, distinctly elongated, indistinctly curved in its whole length, cylindrical in apical half, with two long setae at apex, with one long setae at lateral margin at middle, with one minute seta. Epipleurite with three setae, two close together and one isolated.
Variation. The separation by a notch between the clypeus and the clypeal-wings varies between the two specimens, as well as the rugosity of the clypeal-furrow.
Distribution. Known from the East of Nepal.
Diagnosis. A small sized fuscous species with square pronotum and interval seven and eight of the elytron reticulated. It differs mainly from the most similar species C. quadrataconvexa Balkenohl and convexaquadrata sp. nov. by the conspicuous tubercle at the base of interval three of the elytron, the different reticulation pattern of the lateral intervals, the smaller eyes, and its small size.
Etymology. The name refers to the base of the elytron which carries a large tubercle, and is expressed by a combination of the Latin substantive tuberculum (= tubercle) and the Latin verb ferre (= to carry).
description: Published as part of Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), pp. 1-70 in Zootaxa 5592 (1) on pages 18-19, DOI: 10.11646/zootaxa.5592.1.1, http://zenodo.org/record/14953674