Inella harryleei Rolan & Fernandez-Garces 2008
Creators: name: Fernandes, Maurício Romulo
name: Pimenta, Alexandre Dias
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Triphoridae
Inella
Inella harryleei

description: “Inella” harryleei Rolán & Fernández-Garcés, 2008
Figs 2L, 30–32
Material examined
USA – Florida State • [1, d] spec.; off Saint Petersburg; 28°26´56″ N, 84°40´37″ W; 63 m depth; 24 May 2012; G. Paulay leg.; FLMNH 450495.
Description of basic anatomy
OPERCULUM. Yellowish, ovate, flat, moderately thin but rigid, semi-transparent, 3.25 distinct whorls, nucleus slightly eccentric, dislocated ~37% from center toward margin; diameter of operculum does not exceed diameter of opercular pouch.
JAW. Wing-shaped; outer side with ~30 rows of scales, each scale often rhombus-shaped/squared, occasionally puzzle-shaped, rectangular or irregular; micro-pores in the posterior region, up to 380 nm in diameter; inner side with scales gem-like or leaf-shaped, surface smooth; scales of outer side 10.9– 16.5 µm long, 9.1–11.0 µm wide, ratio length/width 1.0–1.6 (rhombus-shaped/squared), 7.9–11.8 µm long, 4.5–6.1 µm wide, ratio length/width 1.3–2.6 (puzzle-shaped), 8.0– 9.9 µm long, 2.9–4.3 µm wide, ratio length/width 2.1–3.3 (rectangular); scales of inner side 11.8–19.4 µm long, 4.5–7.4 µm wide, ratio length/width 2.3–2.9 (gem-like), 15.1–17.7 µm long, 4.2–6.6 µm wide, ratio length/width 2.3–4.2 (leafshaped).
RADULA. Up to 37 overcrowded and undifferentiated teeth per row, comb-like and usually with five elongated cusps, but occasionally four or six/seven cusps, outermost cusps 39–77% of length of inner cusps; last marginal teeth with four to five triangular and pointed cusps, outer ones and median cusp (cusps 1, 3 and 5, when five cusps present) usually 53–74% of length of larger cusps; teeth with four cusps 4.4–4.7 µm wide, teeth with five cusps 3.7–5.5 µm wide (teeth in the center of row slightly wider than marginal teeth), teeth with six/seven cusps 6.0– 6.6 µm wide, last marginal teeth 3.0– 4.2 µm wide.
Remarks
“ Inella ” harryleei has a different radular morphology when compared to Inella obliqua (May, 1915), which is the single species with illustrated soft parts of Inella s.s. (Marshall 1983). The operculum of “ I ”. harryleei is slightly eccentric (Fig. 30C), similarly to I. obliqua and S. gaesona.
The undifferentiated and overcrowded teeth of “ I ”. harryleei resemble those of Monophorus olivaceus (Figs 17–18) and M. erythrosoma, with a typical tooth bearing five comb-like cusps, in spite of slight variations in the number of cusps. Just like M. olivaceus, the last marginal teeth of “ I ”. harryleei are considerably reduced in size. Species of Monophorus are usually sampled at mid-low depths of the continental shelf (Fernandes & Pimenta 2017a), considering that “ I ”. harryleei similarly inhabits depths of 63–90 m (Rolán & Fernández-Garcés 2008; present study). However, “ I ”. harryleei shows significant differences in the shell (e.g., abapical spiral cord of teleoconch much developed) and operculum (with an ovoid shape, almost elliptical, and nucleus more eccentric) compared to Monophorus. Although it seems premature to reallocate “ I ”. harryleei in Monophorus, “ I ”. harryleei is certainly related to the informal “ Inella group” (Marshall 1983).
description: Published as part of Fernandes, Maurício Romulo & Pimenta, Alexandre Dias, 2019, Basic anatomy of species of Triphoridae (Gastropoda, Triphoroidea) from Brazil, pp. 1-60 in European Journal of Taxonomy 517 on page 46, DOI: 10.5852/ejt.2019.517, http://zenodo.org/record/2647891