Multidentula squalina subsp. acutior Lindholm 1922
Creators: name: Ruud, A. Bank
name: Menkhorst, Henk P. M. G
name: Neubert, Eike
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Enidae
Multidentula
Multidentula squalina
Multidentula squalina acutior (lindholm, 1922)

description: <i>Multidentula squalina acutior</i> (Lindholm, 1922) (Fig. 18) <i>Chondrula acutior</i> Lindholm, 1922b: 358 -359. Type locality: "Von mir im Genist des Fl. Tschoroch bei Kapandidi (unweit Batum).... in einem tadellos erhaltenen leeren Stücke gefunden (coll. m.)". Holotype in ZIN (Sysoev &amp; Schileyko, 2009: 85, 259, fig. 38E). Description. – Shell dextral, oval in outline, with a rather wide, open, slit-like umbilicus. The 6.5-7.7 whorls are rather convex with a rather deep suture. Teleoconch with irregular, oblique striae which are mostly rather fine; there are no spiral striae. Shell solid, not or hardly translucent, rather dark brownish coloured, with a whitish band behind the peristome. The last whorl has an impression at the position of the infrapalatalis; the infrapalatalis and palatalis superior are visible outside as blurred white stripes. Peristome somewhat reflected, thickened by a prominent labial callus, the columellar and palatal insertion connected by a clearly visible callus (but the callus is mostly not extra thickened near the insertions). The subangularis is well developed and fused with the palatal peristome. The subangularis is connected to the parietalis by a more or less thickened callus. The very prominent parietalis is curved and deeply recessed. A spiralis is missing. Columellaris perpendicular to the columellar peristome, deeply recessed. Basalis present, well developed. Infrapalatalis and palatalis superior strongly developed; the infrapalatalis is the most prominent one. A suprapalatalis is also present, but is less prominent than the palatalis superior. A weak suturalis is available. Measurements (n = 13). – H = 5.0-6.3 (mean 5.5); LWH = 2.6-3.1 (mean 2.9); MH = 1.7-2.1 (mean 1.8); LWD = 2.0-2.6 (mean 2.3); LWM = 2.3-2.9 (mean 2.6); MD = 1.5-1.8 (mean 1.6); NW = 6.5-7.7 (mean 7.0). Localities. – Vilayet Artvin: crossing Artvin/&amp;Scedil;av&amp;scedil;at/Demirkent, 17 km towards Yusufeli, suspension bridge Çoruh Nehri (41.0925°N 41.8146°E) (NMBE/2); 5 km N. Erkinis (= castle N. Demirkent) (40.9258°N 41.7619°E) (HMT/1); N. K&amp;imath;nal&amp;imath;çam (= NNW. Ca&amp;gbreve;liyan) (40.7567°N 41.6449°E) (HMT/41; NMBE 544678/1); Yusufeli (40.8215°N 41.5434°E) (HMT/3); 16 km SE. Yusufeli (40.7826°N 41.6967°E) (HMT/14); 17 km SE. Yusufeli, 5 km N. province boundary Erzurum/Artvin (40.7361°N 41.6651°E) (HMT/47). Vilayet Erzurum: 6 km SE. Çaml&amp;imath;yamaç (40.5854°N 41.5952°E) (HMT/81); Ispir, 38 km towards Yusufeli (40.6517°N 41.2299°E) (NMBE/4); deposition Tortum Gölü (40.6361°N 41.6312°E) (HMT/59); 9 km SW. Uzundere, 2 km S. exit to Dikmen (40.4698°N 41.5019°E) (HMT/117). Vilayet Gümü&amp;scedil;hane: 6 km N. Arpal&amp;imath; (= 22 km NW. Bayburt) (40.5792°N 39.1334°E) (HMT/14); Salyaz&amp;imath;, deposition Sulama Gölet&amp;imath; (40.2446°N 39.8070°E) (HMT/29). Differentiation. – <i>Multidentula s. squalina</i> (Fig. 16) differs from <i>M. s. acutior</i> by the more densily packed and more pronounced oblique striae, the more pointed apex, the wider umbilicus, the mostly absent suturalis, the more reflected peristome, the less slen- der suprapalatalis, and the palatalis superior and infrapalatalis are generally more deeply situated in the aperture (i.e. they are situated further away from the peristome). <i>Multidentula s. eudoxina</i> (Fig. 17) has a different dentition compared to <i>M. s. acutior</i>. The former lacks a suprapalatalis, the infrapalatalis is rather broad and seems slightly bifid, the palatalis superior is more prominent and the angularis is not fused with the palatal peristome, but instead connected to it by a thin callus. Remarks. – The type locality "Kapandidi“ of <i>acutior</i> could not be traced. It is likely that Maradidi is meant, which is located in Georgia along the river Chorochi (= Çoruh Nehri) a few km north of the Turkish border (41.5030N 41.7123E). An isolated locality from <i>M. squalina</i> has been reported by Fehér &amp; Er&amp;odblac;ss (2009: 12, pl. 1 figs 8-10) from Albania. All other localities of <i>M. squalina</i> are restricted to Turkey (except one: the type locality of <i>acutior</i>). The shells of Albania (as interpreted from the mentioned figures by Fehér &amp; Er&amp;odblac;ss as well as those published by Welter-Schultes [2012: 194]) do not belong to the nominotypical subspecies, as the subangularis is not fused with the palatal peristome, the presence of a suturalis, the oblique striae are less pronounced, the whorls are more convex, and the apex is less pointed. Furthermore, in <i>M. s. squalina</i> the basalis is shifted to a more subcolumellar position, and the infrapalatalis and palatalis superior are more deeply situated in the aperture. The Albanian subspecies differs from <i>M. s. acutior</i> by the less pronounced subangularis which is non-fused. Another isolated locality of <i>M. squalina</i> is within Turkey itself, namely Çivril and Senirkent (both in the Göller Bölgesi area) (Y&amp;imath;ld&amp;imath;r&amp;imath;m &amp; Schütt, 1997: 52, fig. 16; Schütt, 2001: 148, fig. c; Schütt, 2010: 146, fig. c). Unfortunately, the figures are too vague to determine its subspecific state.
description: Published as part of <i>Ruud, A. Bank, Henk P. M. G Menkhorst &amp; Eike Neubert, 2016, Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae), pp. 5-30 in Basteria 80 (1)</i> on pages 22-23, DOI: 10.5281/zenodo.439745
description: {"references": ["LINDHOLM, W. A., 1922 a. Description of two Bulimini (Gastropoda Pulmonata) from Russian Central Asia. - Annuaire de Musee zoologique de l'Academie des Sciences de Russie 23 (2): 273 - 275.", "LINDHOLM, W. A., 1922 b. Beschreibung einiger kaukasischer Landschnecken (Gastropoda Pulmonata). - Annuaire du Musee Zoologique de l'Academie des Sciences de Russie 23 (3 / 4): 355 - 361.", "SYSOEV, A. &amp; SCHILEYKO, A. A., 2009. Land snails and slugs of Russia and adjacent countries: 1 - 312, 142 plates. Pensoft, Sofia / Moscow. [Pensoft Series Faunistica No 87].", "FEHER, Z. &amp; EROSS, Z. P., 2009. Contribution to the Mollusca fauna of Albania. Results of the field trips of the Hungarian Natural History Museum between 1992 and 2007. - Schriften zur Malakozoologie 25: 3 - 21.", "YILDIRIM, M. Z. &amp; SCHUTT, H., 1997. Verbreitung der Enidae im Seengebiet SW-Anatoliens (Gastropoda Pulmonata: Buliminoidea). - Club Conchylia Informationen 29 (1 / 2): 47 - 55.", "SCHUTT, H. &amp; SESEN, R., 2001. Rediscovery of Sesteria gallandi Bourguignat, 1884 (Mollusca, Pulmonata, Buliminoidea, Enidae). - Turkish Journal of Zoology 25 (4): 427 - 429.", "SCHUTT, H., 2010. Turkish land snails 1758 - 2005. 5 th, revised edition with colour photos: 1 - 559. Verlag Natur &amp; Wissenschaft, Solingen."]}