Scapanoclypeus bicoloratus Sehnal, new species
Creators: name: Sehnal, Richard
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Melolonthidae
Scapanoclypeus
Scapanoclypeus bicoloratus

description: Scapanoclypeus bicoloratus Sehnal, new species (Figs. 1A–E, 3 J)
Type locality. Namibia, Hardap Region, Strampiert.
Type material. Holotype and 3 paratypes (all males): “ Namibia / 10km E Stampiert [= Strampiert, Hardap prov.] / 6.II.1995; leg. K. Werner [p]”. Type depository: holotype and 2 paratypes in TMSA, 1 paratype in RSCV. Description of holotype (Ƌ). Body length 7.6 mm. Body elongate. Head and pronotum black with blackish-brown margins, scutellum black; antennae dark brown (Fig. 1B); elytra bicolored with disc yellowish brown and brownishblack margins (Fig. 1A). Protibiae and metabiae brownish-black, mesotibiae reddish brown, protarsi and mesotarsi testaceous, metatarsi reddish brown. Abdomen brown.
Head. Clypeus broadly rounded; deeply concave in spoon-like fashion; without a large, central, round disc; completely covered by evenly spaced, deep punctures with each bearing 1 long, columnar macroseta (often broken off); sides slightly prolonged backward. Frontoclypeal suture weakly indicated; medially with a definite edge; with broadbased, tricuspid process toward frons (Fig. 1C); anteromedial termination markedly punctate, non-setose. Labrum reduced, triangular; lobes rounded. Frons deeply rugose; edge of clypeus strongly punctate; each puncture bearing a long, semierect, posteriorly inclined, yellow macroseta; macrosetae longest at edge of clypeus and around eye canthus. Eyes large, exceeding genae externally in dorsal aspect; distance between eyes in ventral aspect shorter than diameter of eye. Genae rugopunctate, with group of long macrosetae. Antennae with nine antennomeres; antennomeres 7–9 forming long, strongly curved club at least 6.4x times longer than antennal shaft; antennomeres 1–4 with sparse, long macrosetae; antennal club completely densely punctate, without smooth areas. Antennomere 2 bulbous and as long as antennomeres 3–6 combined. Terminal maxillary palpomere elongate, longer than palpomeres 2 and 3 combined; sub-basally with apically rounded, flat, oval alutaceous area tapering toward apex.
Pronotum weakly convex, approximately tetragonal, 1.32x times wider than long, broadest approximately at middle, with finely impressed medial line; nearly completely bordered, anterior border widely interrupted in the middle. Anterior and posterior angles strongly rounded, poorly defined. Surface with long, yellow macrosetae; deeply punctate, punctures behind head each with a macroseta adhering to disc.
Scutellum shiny, impunctate, slightly longer than wide; margins broadly arcuate, lateral margins without macrosetae. Elytra moderately convex, only slightly dilated posteriorly. Disc strongly wrinkled and punctate; punctures evenly distributed, separated approximately by more than 6x their diameter; macrosetose as on pronotum, inclined posteriorly, absent from distinct humeri. Sides punctate, with lateral margins and apices diffusely darker than disc.
Macropterous.
Legs. All femora shiny and irregularly, coarsely punctate; macrosetae relatively long. Protibia tridentate [appearing bidentate], with basal tooth obscure, broadly angulate, without inner subapical spur. Claws simple, each with small, blunt bulge at base. Mesofemora dark brown. Metafemora with dark brown margins and yellow medially. Mesotibiae moderately expanded apically. Metatibiae strongly expanded apically, with 1 oblique carina externally; apical edge with row of long, stout, macrosetae of equal length; terminal calcars stout, long, lower calcar slightly shorter than upper calcar. Protarsomeres, mesotarsomeres, metatarsomeres without patches of short, dense macrosetae; metatarsomeres ventrally covered with long, isolated, sparse macrosetae.
Abdominal sternites reddish brown with yellow, recumbent setae. Pygidium yellow, flat, finely punctate.
Male genitalia (Figs 1D – E). Aedeagus symmetrical, parameres slender, relatively long.
Variability in males. Paratypes somewhat variable in body length (6.3–8.2 mm), slightly variable in dorsal punctation density and length and distribution of macrosetae. Color as in holotype.
Female. Unknown.
Diagnosis. Based on characters stated in Evans (1987), the new species differs from its congeners by the combination of following characters: tridentate protibia [appearing bidentate], inner subapical spur missing; clypeus punctate, without a large, central, round disc; elytra bicolored - disc yellowish brown, with brownish-black margins; protarsi and mesotarsi testaceous, metatarsi reddish brown. Generally, the new species is similar to S. triapicalis, from which it differs by having bicolored elytra, protibiae and metabiae brownish black, mesotibia reddish brown, antennal club long, curved club at least 6.4x times longer than antennal shaft, and clypeus without a large, central, round disc. In contrast, S. triapicalis has monochromatic elytra, tibiae reddish brown, antennal club long, strongly curved club at least 5.1x times longer than antennal shaft, and clypeus with a large, central, round disc.
Etymology. The specific epithet is given for the two colors of the elytra. The name is an adjective in the nominative singular.
Distribution. Namibia, Hardap Region (Fig. 3 J).
description: Published as part of Sehnal, Richard, 2017, Scapanoclypeus bicoloratus new species from Hardap, Namibia (Coleoptera: Scarabaeidae: Melolonthinae: Tanyproctini), pp. 593-598 in Zootaxa 4247 (5) on pages 593-594, DOI: 10.11646/zootaxa.4247.5.6, http://zenodo.org/record/439188