Lannapsyche kamba Oláh & Johanson 2010, new species
Creators: name: Oláh, János
name: Johanson, Kjell Arne
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Odontoceridae
Lannapsyche
Lannapsyche kamba

description: Lannapsyche kamba, new species   
Figs 249–252  
 In the genitalia, the shape of the harpagones of L. kamba are intermediate between the slender ones in L. suksma and the wide ones in L. birathena. The coxopodites of L. kamba are different from those in the 2 other species in that they have an apicomesal, lobe-like, broadening at the base of the bifurcation, as seen in ventral view,. The phallic apparatus is much reduced in size, but with the same V-shaped, configuration in lateral view.   
 Male (in alcohol). Body medium sized; light brown; legs, antennae and palps lighter than body; forewings pale brown, with pale setae. Head rectangular in dorsal view, about 2 times broader than long. Ocelli absent. Tentorium slender, without dorsal arm; posterior arms short, robust in dorsal view, ending in pair of large posterior tentorial pits; strong tentorial bridge without anteromesal protuberance, but with posteromesal hump; anterior tentorial arms slender, without median lamellate process; slightly broadening immediately before anterior apex. Facial groove pattern simple. Frontogenal vertical grooves forming long, dorsad orienting continuation from anterior tentorial pits. Clypeogenal vertical grooves located ventrally of anterior tentorial pits; short, running obliquely, almost horizontally laterad; subantennal grooves running vertically, with slightly mesad turning anterior end, apparently not confluent to clypeogenal groove. Vertexal grooves with thick rim of compact warts; stem of epicranial groove complete, dominating vertex; occipito-postgenal grooves partly merging anterad with skeletal ring of occipital setal warts. Labrum semi-circular, without setal warts; narrow, weakly pigmented anterior part movable, freely hanging. Frontal setal warts absent; frontal interantennal warts absent, or represented by rounded compact, elevated, posterior setal warts. Pair of round frontogenal, posterior, compact, setal warts present dorsally on pregenae, between vertical frontogenal and subantennal grooves. Pair of large, transverse, elongate, setose warts present below clypeogenal grooves, representing main setal warts on facial area of head. Vertexal, lateroantennal compact warts rounded, located below antennal grooves; pair of vertexal medioantennal compact setose warts rounded, elevated on anterior ending of coronal groove, separated by deep cleft. Paired frontal grooves nearly invisible. Pair of large, occipital compact warts covering posterior half of head dorsum, almost fusing with vertexal ocellar compact setal warts. Vertically elongate, postgenal compact warts curving along posterior section of ocular grooves. Maxillary palps broken, first and second segments intact; each with basal segment with subapicomesal nodule with setose apex; base of nodule somewhat membranous, indicating flexibility to eversion. Each scapes slightly longer than head. Each pedicel shorter than first segment of each flagellum. Pronotum with 2 pairs setal warts. Mesal warts compact, slightly elongated obliquely, occupying entire top of dorsally elevated area; elevated area separated by wide gap. Lateral warts small, rounded. One pair mesoscutal, fragmented warts arranged in longitudinal lines composed of 4 to 5 fragments. One pair mesoscutellar diffuse warts forming longitudinal patch. Each proepisternum with large, ovoid setose wart. Precoxal warts small. Pair of large, compact setal warts present on membranous part of cervix, tangential with anterior arm of each cervical sclerite. Each lateral cervical sclerite forming narrow anterior arm articulating anteriorly with occipital condyle above posterior tentorial pits of head; fusing with posterior cervical sclerite; each posterior cervical sclerite forming large, quadrangular plate reaching prothoracic episternum, articulating to weakly sclerotized anteromedian band of prothoracic eusternum by ventral intercervical sclerites. Leg claws symmetrical; spur formula 2, 4, 4; foreleg posterior spur half as long as anterior spur; midleg and hind leg anterior spurs half as long as posterior spurs. Forewings: length 8.9 mm; membrane light brown without visible pattern, with pale setae; R1 confluent with, or recurrent into, R2 short before C; base of discoidal cell located proximally of mid-length of wing; forks I, II, III, V present; forks I, II, and V sessile, fork III petiolate, crossveins ic, r, s, r-m and m-cu present; crossvein m absent; crossveins arranged in an almost regular vertical line of anastomis; postanal vein present. Hind wings: R1 confluent with R2 well before C.   
Male genitalia. Abdominal segment IX fused annularly, with longitudinal groove separating dorsal and ventral parts; groove straight oblique, ventrad curving at posterior margins, continuing further along ventral margin of segment X as dark rim; tergum straight, flat, descending posterad, in lateral view, as long as venter. Anterior margins of segment IX straight, concave at venter; posterior margins concave, with pronounced triangular above end of longitudinal groove. Antecostae forming weak marginal rims, being thick ventrally of longitudinal grooves; external groove of antecostal sutures weak; each tergum with central line only in dorsal view, separating concave surface of reduced preanal appendages; spine rows on posterior margins of segment IX reduced, merged with, or replaced by, long setae of abbreviated preanal appendages. Intersegmental depression between segment IX and segment X visible as rounded step in lateral view. Segment X forming broad lobe in lateral view, with slightly excised apex; in dorsal view with broad base and with bifid apex. Apicoventral setose lobes and apicodorsal setose lobes fusing into integrated setose area on dorsum; with very short, stout setae. Dorsal interlobular gap shallow, and narrowly triangular. Superior appendages reduced into large, circular, concave surface dominating on dorsum of genitalia, with long setae. Each coxopodites much longer than apex of segment X, parallel-sided, stout, straight, slightly ventrad curving, slightly broadening apically; harpagones as broad as apical process of coxopodites, capitate, emerging from deep apical gap of each coxopodite; each with capitate apex covered by black, stout, short triangular setae. Phallic apparatus small, forming inverted V in lateral view; about half as long as each coxopodite; most sclerotized basal section tuboid, curving ventrad, continuing into obscurely membranous section, possibly representing part of endotheca; ventrally followed by more sclerotized part; dorsally ending in membranous phallicata. Ejaculatory duct running into circular, heavily striped endophallus and phallotremal sclerites. Most pigmented part of phallus elongate in lateral view.  
 Holotype male: MYANMAR: North East: Kambaiti, 2000 m, 26.v.1934, Malaise trap (Malaise B.M. 1938-258) ─ (NHML).   
 Distribution: Myanmar.   
 Etymology: Kamba, name derived from Kambaiti, the type locality of the species.
description: Published as part of Oláh, János & Johanson, Kjell Arne, 2010, Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457, pp. 1-128 in Zootaxa 2457 (1) on pages 111-113, DOI: 10.11646/zootaxa.2457.1.1, http://zenodo.org/record/5313709