Maymena Gertsch 1960
Creators: name: Lopardo, Lara
name: Hormiga, Gustavo
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Mysmenidae
Maymena

description: MAYMENA GERTSCH, 1960
(FIGS 10–16, 128B, D, E, 131I, J, 140M–O, 141A–I, 147D,E: CLADE C168)
Maymena Gertsch, 1960a: 30–38. Gertsch, 1971: 94–95. Brignoli, 1974: 224. Baert, 1990: 17. Platnick, in Eberhard, Platnick & Schuh, 1993: 11. Griswold et al., 1998: 63.
Type species
Maymena mayana (Chamberlin & Ivie, 1938) by original designation, type material in MCZ, not examined.
Familial placement and composition
Transferred to Mysmenidae from Symphytognathidae by Forster & Platnick (1977). Maymena is closely related to Mysmenopsinae in our working phylogenetic hypothesis (Fig. 161B). Currently, Maymena comprises 13 described species (Platnick, 2014), and it is represented here by three described plus three undescribed species (the latter species scored only for molecular data): M. mayana, M. ambita, M. rica, Maymena -MYSM-003- ARG, Maymena -MYSM-004- MEX, and Maymena - MYSM-016-ARG).
Monophyly
Morphological synapomorphies of Maymena include: posterior lateral spinnerets with an anterior flat spatulate modified seta (Fig. 11H); aciniform gland spigots of the posterior spinnerets of two different shapes (Fig. 13D); seta on major ampullate field with one row of long ‘branches’ (Figs 11E, 13C, 16B); leg spination (i.e. macrosetae occurring on tibiae, femora, and metatarsi; Figs 140M, 141C); and males with cylindrical palpal tibia (Fig. 10A). Ambiguously optimized synapomorphies for Maymena include: males with a femoral spot at least on femur I; metatarsal clasping spine particularly proximal (Fig. 16G); embolus rim deeply grooved (Fig. 10H); the primary cymbial conductor apically bent over the ventral side (Fig. 10D, G); epiandrous fusules dispersed in a row (Figs 12B, 16A); and macrosetae on female palpal tarsus (Figs 13A, 15A). Maymena is one of the few clades in this data set that is strongly supported and stable, and its monophyly is supported by most data partitions and parameter combinations. The genus is also supported by 125 molecular synapomorphies.
Diagnosis
Maymena shares with Mysmeninae the anterior flat spatulate modified seta on PLS and the aciniform gland spigots of two different shapes; with Trogloneta the presence of femoral spot on males; and with Mysmenopsinae the dispersed male epiandrous fusules, the macrosetae on female palpal tarsus, the leg spination, and the absence of tegular conductor (see also Gertsch, 1960a). Maymena differs from all other mysmenid genera by the combination of the aforementioned features, and by the presence of one row of long ‘branches’ on the major ampullate field seta, males with cylindrical palpal tibia, metatarsal clasping spine particularly proximal, seemingly interacting with the particularly distal tibial clasping spine in most species, and the male palpal morphology, with primary cymbial conductor apically bent over the ventral side. In addition, the respiratory arrangement distinguishes Maymena from other mysmenids. It consists of anterior booklungs and two long lateral tracheal tubes and two shorter median apodemes arising from a narrow posterior single spiracle (L. Lopardo, P. Michalik & G. Hormiga, unpubl. data; also Gertsch, 1960a).
description: Published as part of Lopardo, Lara & Hormiga, Gustavo, 2015, Out of the twilight zone: phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea), pp. 527-786 in Zoological Journal of the Linnean Society 173 (3) on page 778, DOI: 10.1111/zoj.12199, http://zenodo.org/record/5331625