Pterocyclos frednaggsi Sutcharit &amp; Panha 2014, new species
Creators: name: Sutcharit, Chirasak
name: Tongkerd, Piyoros
name: Panha, Somsak
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Architaenioglossa
Cyclophoridae
Pterocyclos
Pterocyclos frednaggsi

description: <i>Pterocyclos frednaggsi</i> Sutcharit &amp; Panha, new species (Figs. 1, 2A <b>–</b> C, E, 3Q <b>–</b> S; Table 1) <b>Type specimens.</b> Holotype CUMZ 4594 (height 12.1 × diameter 27.8 mm; Fig. 3Q). Type locality: Bukit Chintamanis, Pahang, Malaysia (03°26.798' N, 102°00.987' E). Paratypes CUMZ 4581 (18 specimens in ethanol, Fig. 3R), 4571 (29 shells), NHMUK (2 shells), NMW (2 shells), SMF (2 shells) are from the type locality. <b>Other material examined.</b> Gua Pulai, Gua Musang, Kelanta, Malaysia CUMZ 4597 (9 shells, Fig. 3S), 4944 (4 specimens in ethanol). <b>Etymology.</b> The specific epithet “ <i>frednaggsi</i> ” comes from Fred Naggs, malacologist at the Natural History Museum, London, who has enthusiastically encouraged and continuously supported land snail research in Thailand and other Southeast Asian countries. <b>Diagnosis.</b> <i>Pterocyclos frednaggsi</i> Sutcharit &amp; Panha, new species, differs from <i>P. blandi</i> by having a thick and yellowish periostracum, a broad, dark brown peripheral band, a channel shaped accessory respiratory structure, and a slight expansion of the upper part of the apertural lip. It differs from <i>P. spaleotes</i> by having a larger shell, a thicker periostracum, and a broad, dark brown peripheral band. The differences from <i>P. diluvium</i> Sutcharit &amp; Panha, new species, include the broad, dark brown peripheral band and the channel shaped accessory respiratory structure. This new species is superficially similar to <i>P. umbraticus,</i> but differs from this latter species by its relatively smooth shell surface, the broad dark brown peripheral band, and the fact that its distribution is limited to the lowland limestone area (altitude less than 200 m amsl). In contrast, <i>P. umbraticus</i> has a chevron-shaped, zigzag shell sculpture and tends to live in non-limestone forests of highland areas (altitude about 200 <i>–</i> 1600 m amsl) at Maxwell’s Hill and Gunung Brinchang, Pahang (Benthem Jutting, 1949). <b>Description.</b> Shell medium sized, depressed, thickened, widely umbilicated. Apex acute; spire flattened to slightly elevated; suture wide; 4 <i>–</i> 5 convex and regularly increasing whorls. Shell surface with thin growth lines; periostracum brown and corneous. Last whorl rounded, stout, with a broad dark brown peripheral band. Shell colour brownish or with a variegated, dark brown zigzag pattern on early whorl. Aperture rounded. Apertural lip white, upper part slightly expanded near suture. Accessory respiratory structure channel-like to short tubular shaped. Operculum calcareous, slightly concave inside, and multilamellae outside. <b>Radula.</b> Central tooth with well developed central cusp and two smaller lateral cusps on each side; central cusp small with pointed tip; four smaller lateral cusps on both sides with dull to pointed head. Lateral teeth have three cusps; outer cusp large, elongate shape, and two smaller inner lateral cusps with curved tips. Inner marginal teeth have three cusps; central cusp large and convex head, and flanked with smaller and pointed head of one inner and one outer lateral cusps. Outer marginal cusp bicuspid, each cusp with pointed head (Fig. 2E). <b>Distribution.</b> This limestone dwelling species is known from the type locality and from Gua Musang, Kelantan, Malaysia. <b>Remarks.</b> Specimens of this species from Gua Musang, Kelantan (Fig. 3S) have slightly larger and darker shells with a narrower peripheral band than typical shells. However, the unique shape of the short tubular accessory respiratory structure and the multilamellae operculum suggest that these atypical forms represent intraspecific geographic variation.
description: Published as part of <i>Sutcharit, Chirasak, Tongkerd, Piyoros &amp; Panha, Somsak, 2014, The land snail genus Pterocyclos Benson, 1832 (Caenogastropoda: Cyclophoridae) from Thailand and Peninsular Malaysia, with descriptions of two new species, pp. 330-338 in Raffles Bulletin of Zoology 62</i> on pages 336-337, DOI: 10.5281/zenodo.5458831
description: {"references": ["van Benthem Jutting WSS (1949) On collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells. Bulletin of the Raffles Museum, 19: 50 - 77."]}