Geomyphilus ungulatus
Creators: name: Dellacasa, Marco
name: Dellacasa, Giovanni
name: Skelley, Paul E.
name: Gordon, Robert D.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Aphodiidae
Geomyphilus
Geomyphilus ungulatus

description: Geomyphilus ungulatus (Fall, 1901)   
(Fig. 51–55)  
 Aphodius ungulatus Fall, 1901: 254; Schmidt 1922: 336 (subgenere incerto).   
 Aphodius (Chilothorax) ungulatus; Dellacasa 1988: 212.   
 Geomyphilus ungulatus; Gordon and Skelley 2007: 411 (lectotype designation).   
 Type locality. Pasadena, California [U.S.A.].   
 Type repository. Museum of Comparative Zoology. Harvard University, Cambridge. MA (U.S.A.) (type not examined).   
 Redescription. Length 5.0–6.0 mm; oblong, moderately convex, moderately shiny, glabrous. Head and pronotum piceous; clypeal margin and pronotal sides shadowy reddish; elytra dirty yellow with a lateral fuscous shade widened from shoulder to beyond the middle; legs pale brown; antennal club fuscous. Head with epistome feebly convex, rather evenly, moderately punctured; punctation distally almost granulo-rugose, finer and sparser on disc; clypeus broadly sinuate at middle, distinctly angulate at sides, thinly bordered, edge slightly reflexed, finely and sparsely fimbriate laterally; genae obtusely round, elongately ciliate, protruding from the eyes; frontal suture obsolete; front moderately evenly punctured. Pronotum transverse, feebly depressed near hind angles, dually punctured; large punctures, three times larger than small ones, close and coarse on sides, lacking on disc; small punctures dense and deeper on sides, sparse and superficial on disc; lateral margins slightly arcuate, thinly bordered, edge sparsely shortly fimbriate; hind angles obtuse; base almost regularly arcuate, thickly bordered toward hind angles, thinly bordered medially. Scutellum flat, coarsely densely punctured on basal half. Elytra oval, scarcely broader posteriorly, finely striate; striae superficially not closely punctured, subcrenulate; interstriae almost flat, strongly alutaceous on preapical declivity, finely irregularly punctured. Hind tibiae upper spur shorter than first tarsal segment; latter as long as following two segments combined. All claws very slender and exceptionally elongate. Male: head and pronotum relatively less convex and less coarsely punctured; aedeagus Fig. 54–55. Female: head and pronotum relatively more convex and more coarsely punctured.   
 Material examined. U.S.A.: California: Cypress Co., 22.XI.1930, leg. A. C. Davis, hole of Citellus (1 ex., DCGI); Rt. 71 16.22 mi. fr. San Bernardino County line, Riverside Co., 03.X.1972, leg. J. Saulnier, refuge chamber of Thomomys bottae (8 exx., DCGI).   
 Distribution. U.S.A. (California).   
 Biology. Apparently a rodent associate species, collected from November to February.
description: Published as part of Dellacasa, Marco, Dellacasa, Giovanni, Skelley, Paul E. & Gordon, Robert D., 2017, Systematic revision of the species of Geomyphilus (Coleoptera: Scarabaeidae: Aphodiinae) of Mexico and neighboring countries with description of a new Mexican species, pp. 1-19 in Insecta Mundi 2017 (590) on pages 11-12, DOI: 10.5281/zenodo.5169501