Calyxochaetus tohono Runyon, 2024, sp. nov.
Creators: name: Runyon, Justin B.
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Dolichopodidae
Calyxochaetus
Calyxochaetus tohono

description: Calyxochaetus tohono sp. nov.
(Figs 111, 127, 143, 148)
Type material. HOLOTYPE ♂, labeled: “ ARIZONA: Santa Cruz Co. / Sonoita Crk nr Patagonia / Lake inlet 3775 ft / N31°29.95′ W110°50.69′ / 03APRIL2016, J.B. Runyon ”; “HOLOTYPE/ Calyxochaetus / tohono / J.B. Runyon [red label]” (USNM) (Fig. 143). PARATYPES: USA: Arizona: Same data as holotype (2♂, CAS; 2♂, CNC; 10♂, MTEC; 2♂, USNM); Cochise County: Southwestern Research Station, Portal, 5–9.vi.1972, Malaise, W.W. Wirth (1♂, USNM); South Fork Cave Creek, Chiricahua Mts, 5235 ft, 17.iv.2003, JBR & RLH (1♂, MTEC); San Pedro River, 4 mi. E Sierra Vista, 4025 ft, 23.iv.2002, RLH & JBR (2♂, 2♀, MTEC); Garden Canyon, Huachuca Mts, 5300 ft, 16.iv.2003, JBR & RLH (4♂, MTEC); Pima County: Madera Canyon, 23.vi.1982, RLH (1♂, MTEC); Santa Cruz County: Tubac, 29.ix.1972, L.J. Ogden & FCH (1♂, FSCA); Sycamore Canyon, 14 mi. NW Nogales, 4000 ft, 14.iv.2003, JBR & RLH (3♂, 12♀, MTEC); Patagonia Lake, 3750 ft, 15.iv.2003, JBR & RLH (9♂, 2♀, MTEC); Ruby Road W Peña Blanca Lake, ca. 4200 ft, 31°23.4′N 111°07.4′W, 21–22.viii.2007, yellow pan trap along creek, J.M. Cumming & S.E. Brooks (2♂, 1♀, CNC); same data except, sweep creek margin, S.E. Brooks (2♂, CNC); same data except, J.M. Cumming (1♂, 1♀, CNC).
Diagnosis. Males can be recognized by the medium-sized lanceolate lamellae on the arista-like stylus of males (Fig. 111), yellow metepimeron, and hind tibia with posteroventral row of slender setae (as in Fig. 134). This species is similar to C. ornatus but is smaller and males have an arista-like stylus with smaller lanceolate lamella.
Description. Male. Wing length 2.8–3.3 mm. Similar to C. ornatus except as noted: Head: Face silver with golden yellow tinge. Postpedicel (Fig. 111) slightly longer then wide, often with slight subapical ventral indentation (a few specimens have more rounded postpedicel); arista-like stylus (Fig. 111) with smaller lanceolate lamella (width less than half width of postpedicel). Scutellum without distinct blue-violet reflections medially (at most weak reflections). Legs: Foreleg: Tarsomere 3 two-thirds length of tarsomere 4. Hindleg: Tibia with posteroventral row of slender setae on basal half slightly smaller, more uniform in length. Abdomen: Hypopygium: Postgonites with apical lobes bluntly rounded with dorsal edge slightly curved, shiny light to dark brown.
Female. Wing length 2.9–3.3 mm. Similar to C. ornatus except postpedicel usually as long as wide; tergite 4 usually wholly brown occasionally with small faint yellow ventrolateral spot.
Distribution and seasonal occurrence. Known from southeastern Arizona (Fig. 148). Adults have been collected from April through September.
Remarks. Adults were observed at several sites sitting on, and flying around, low vegetation and algae-covered stagnant water of side-pools of creeks in the shade of willows (Fig. 3) and large rocks.
Etymology. Named in honor of the Tohono Oʼodham, Native American people whose traditional territory included southern Arizona. The species name is considered a noun in apposition.
description: Published as part of Runyon, Justin B., 2024, Revision of the genus Calyxochaetus Bigot (Diptera: Dolichopodidae: Sympycninae), pp. 1-74 in Zootaxa 5539 (1) on pages 70-71, DOI: 10.11646/zootaxa.5539.1.1, http://zenodo.org/record/14240426