Cheiloneurus danessa Noyes 2023, sp.nov.
Creators: name: Noyes, John Stuart
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Encyrtidae
Cheiloneurus
Cheiloneurus danessa

description: Cheiloneurus danessa sp.nov.   
(Figs 547-553; Hab. E 85)  
 DIAGNOSIS. Female (length about 0.9 mm): body generally dark brown with a weak to moderate metallic sheen; mesoscutum with uniform brown setae; fore wing (Fig. 553) hyaline; fore coxa white; head about 2.7X as wide as frontovertex; antenna (Fig. 547) with pedicel about as long as F1-F2 combined; all funicle segments very slightly longer than broad and with linear sensilla; clava slightly longer than F4-F6 combined, with apex rounded; mandible with 1 acute tooth and a broad, upper truncation; mesoscutum (Fig. 552) with longitudinally elongate, lineolate sculpture, some elongate, polygonally reticulate sculpture medially and rounded cells near posterior margin; scutellum with longitudinally elongate, lineolate sculpture, a few polygonally reticulate cells in middle; scutellum without tuft of setae medially in apical half; fore wing (Fig. 553) about 2.3X as long as broad; parastigma slightly swollen, not downcurved; stigmal vein (Fig. 550) with 3 campaniform sensilla at apex; gaster (Fig. 551) with “glands” on Gt1 and Gt5, those on Gt1 continuous, not separated, those on Gt5 separated by less than 0.2X their own lengths; syntergum broader than long. Male: unknown.   
 Female ( holotype ): length, including ovipositor, 0.90mm; excluding ovipositor, 0.86mm (CPD).   
Head dark brown to black with a metallic sheen; occiput with a weak coppery sheen, some metallic blue-green reflections below occipital margin; frontovertex generally slightly shiny with a weak coppery purple sheen along occipital margin and above scrobes, some dull coppery purple elsewhere, with some hardly discernible metallic dark blue-green; scrobal area with a moderate coppery purple sheen, area between eye and scrobes dull coppery purple with a hint of metallic dark blue-green; temple and gena with a dull coppery purple sheen; frontovertex, gena and interantennal prominence with a few scattered, inconspicuous, pale brown setae; palpi white; radicle orange-brown; scape mostly dark brown on external surface with apical 0.4X ventrally pale orange, internal surface similar but with most of apical half pale orange; pedicel mainly dark brown, apex very pale orange; flagellum slightly dusky, yellow, clava dusky pale orange; thorax dark brown to black with a weak to moderate metallic lustre; pronotum with a moderate coppery sheen; mesoscutum quite dull with a slight brassy and weak purple sheen, posteriorly a little more shiny and coppery along posterior margin; tegula dark brown with a slight coppery sheen; axilla with a weak to moderate coppery purple sheen; scutellum dorsally about as shiny as mesoscutum, medially with a weak coppery purple sheen, laterally mixed with weak to moderate metallic dark blue-green, extreme side and apex with a moderate metallic dark blue-green sheen mixed coppery; mesopleuron fairly shiny with a slight brassy and coppery purple sheen with a hint of metallic dark blue-green; fore coxa white, mid and hind coxae dark brown; fore femur white to very pale yellow with a very thin dark brown line along dorsal margin in middle half and a very faint, shorter one opposite, tibia white at extreme base, otherwise pale orange with a very weak dusky spot subbasally on outer surface, tarsus pale orange with apical tarsomere slightly dusky; mid femur white with a narrow, incomplete, dark brown subapical band, tibia white with knee very narrowly dark brown and with a narrow, dark brown subbasal band, spur very pale yellow, tarsus white to very pale yellow; hind femur dark brown with extreme base and apex white, tibia dark brown with extreme base white and apical two-fifths white, tarsus white to pale yellow with apical tarsomere dusky; wings hyaline; propodeum dark brown with a weak purple sheen, side with a moderate coppery sheen; gaster dark brown with coppery purple, brassy, green and blue reflections; gonostylus brown.  
Head about 2.7X as wide as frontovertex, in profile about 1.4X as high as deep, tangent to frontovertex forming an angle of about 95° with that of face; occipital margin sharp, not carinate; no distinct oval depression adjacent to eye near occipital margin; ocelli forming an angle of about 80°; inner eye margins subparallel; frontovertex with moderately deep, polygonally reticulate sculpture of mesh size generally subequal to eye facet, not shallower above top of scrobes; temple and gena with shallower, fine, elongate, polygonally reticulate sculpture; top of scrobes with shallow, regular, polygonally reticulate sculpture, scrobes smooth; interantennal prominence and mouth margin with slightly shallower, polygonally reticulate sculpture; scrobes shallow, indistinctly margined, broadly ∩-shaped; interantennal prominence dorsally narrowly rounded; antenna as in Fig. 547; scape relatively short and stout, slightly broadened and flattened, about 3.4X as long as broad; funicle with segments all slightly longer than broad and becoming progressively larger distad so that F6 is largest; linear sensilla present on all funicle segments; clava about 3.2X as long as broad and about 1.1X as wide as F6, not curved downwards towards apex, slightly longer than F4-F6 combined, sutures parallel, sensory area small, at apex only giving it a rounded appearance; eye slightly overreaching occipital margin, appearing naked; malar sulcus absent; clypeal margin weakly concave; mandible with one tooth and a broad, slightly oblique, truncation. Relative measurements: HW 79, HH 61, FV 29, POL 14.5, OOL 2, OCL 4, AOL 9, EL 42, EW 38.5, MS 24, SL 27.5, SW 8.  
Thorax (Fig. 552) with pronotum hardly exposed behind head, its posterior margin broadly concave, with a shallow median invagination; pronotum with shallower, finer polygonally reticulate sculpture than ocellar area; mesoscutum with conspicuously longitudinally lineolate striate-reticulate to striate sculpture that is finer than that in ocellar area and about as deep, a few rounded cells near posterior margin; axilla with slightly transversely elongate, polygonally reticulate sculpture that is slightly shallower than that on mesoscutum; scutellum dorsally with striate to striate-reticulate sculpture that is slightly deeper and coarser than that on mesoscutum, cellular sculpture along mid-line more rounded, extreme side and apex quite smooth and shiny; visible part of mesoscutum about 1.8X as broad as long; scutellum about 1.2X as broad as long, apex with a pair of slender, elongate setae; hind femur about 3.4X as long as broad; fore wing with venation and setation as in Figs 550, 553; linea calva entire and open; propodeum (Fig. 552) medially less than 0.1X as long as scutellum. Relative measurements: FWL 67.5, FWW 29.5; HWL 47.5, HWW 11.  
Gaster with hypopygium reaching about 0.5X to apex; last tergite about 0.9X as long as mid tibia, with apex acute; “glands” (Fig. 551) present on Gt1 and Gt5, those on Gt1 continuous, not separated, those on Gt5 separated by less than 0.2X their own lengths; hypopygium Fig. 551; ovipositor (Fig. 548) slightly exserted, the exserted part about 0.5X as long as mid tibial spur or about 0.15X as long as mid tibia. Relative measurements: OL 75, GL 20 [MT 48].  
Variation. Only holotype examined.  
Male. Unknown.  
DISTRIBUTION. Costa Rica.  
HOSTS. Unknown.  
MATERIAL EXAMINED.  
 Type material. Holotype E: COSTA RICA, Guanacaste, 18km N. Liberia, Buena Vista Lodge, 10°48’N 85°24’W, 770m, 15-17.ii.2016 (J.S. Noyes). Holotype in NHMUK.   
COMMENTS. The holotype has the right flagellum missing but is otherwise in good condition.
description: Published as part of Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera), pp. 1-921 in Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) (Oxford, England) 2 (11) on pages 223-225, DOI: 10.5281/zenodo.8074943