Pterostichus (Wraseiellus) stictopleurus subsp. cangshanensis Shi & Sciaky, new subspecies
Creators: name: Shi, Hongliang
name: Sciaky, Riccardo
name: Liang, Hongbin
name: Zhou, Hongzhang
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Pterostichus
Pterostichus stictopleurus
Pterostichus (wraseiellus) stictopleurus cangshanensis shi &amp; sciaky

description: Pterostichus (Wraseiellus) stictopleurus cangshanensis Shi & Sciaky, new subspecies
(Figures 12, 25, 36, 44, 65, 66, 73, 88, 91, 105)
Type locality. Yunnan: Dali, E. slope of Cangshan Mt. (N25.63º, E100.14º)
Type materials. Holotype (IZAS), male, body length = 12.8 mm, pinned, genitalia dissected and glued on plastic film pinned under specimen, " CHINA, Yunnan prov., / Dali, E. slope of Cang- / shan Mt., 2008. X.18, / leg. HUANG Rong etc., / NO.2008101801"; " / 2008. X.18, / NO.2008101801"; "IOZ(E)1891916"; " HOLOTYPE 3 / Pterostichus (Wraseiellus) / stictopleurus cangshanensis / new subspecies / Des. SHI H.L. 2012" [red label] [Figs. 12, 25, 36, 44, 65, 73, 105]. Paratypes (total 18 specimens all from Dali): 3 males and 2 females (IZAS), the same label data as holotype, 2 of them (1 male, 1 female) with different IOZ database label (1891914, 1891915), others without database label. 1 male (IZAS), " CHINA, Yunnan prov., Dali, S. slope of Cangshan Mt., 2008. IX.6, leg. LI Yunchun etc., NO.2008090608". 1 male (IZAS), " CHINA, Yunnan prov., Dali, E. slope of Cangshan Mt., 2008. IX.6, leg. LI Yunchun etc., NO.2008090608". 1 male (IZAS), " CHINA, Yunnan prov., Dali, E. slope of Cangshan Mt., 2008. IX.13, leg. ZHANG Youliang etc., NO.2008091301" [Fig. 66]. 2 males (CRS), " CHINA, Yunnan prov. Dali, 6.7.1993, lgt.S.Becvar". 3 males, 3 females (SCAU), " CHINA, Yunnan prov., Dali, Cangshan Mt., 2009. VI.25, 2600 m, leg. YANG Zhenxing, ZHANG Youliang etc., S1986WDm" [Fig. 91]. 1 male, 1 female (SCAU), " CHINA, Yunnan prov., Dali, Cangshan Mt., 2009. VII.8, 2500 m, leg. Ma Yanhong, WANG Mingtao etc., S1991EDo" [Fig. 88].
Diagnosis. The new subspecies can be distinguished from all the other members of this subgenus by the combination of the following characters: (1) 5th tarsomere setose beneath; (2) males with elytral microsculpture isodiametric; (3) pronotum hind angles obtuse, not pointed.
Comparing with the nominotypical subspecies, the new subspecies is different in: (1) 5th tarsomere setose beneath (glabrous in the nominotypical subspecies); (2) ridge between outer groove of basal fovea and lateral margin of pronotum stronger than in the nominotypical subspecies (Figs. 104, 105); (3) punctures on proepisterna much finer and sparser than in the nominotypical subspecies; (4) outer margin of right paramere slightly curved (straight in the nominotypical subspecies) (Figs. 72 B, 73B); (5) median lobe of male genitalia straight before apical lamella (in lateral view), and slightly bent to the dorsal side at apex (Figs 65, 66), but in the nominotypical subspecies, median lobe slightly sinuate and then strongly bent to the dorsal side (Figs. 63, 64).
Description. Body length 11.6–13.4 mm; dorsal side black, shining in males; tibiae dark brown, tarsi brown; ventral side black or dark brown. Males with elytral microsculpture isodiametric. Head. Frons and vertex impunctate, few punctures present in frontal furrows; antennae reaching elytra basal eighth; 3rd antennomere glabrous except apical setae; males with the terminal labial palpomere similar as in females, slightly expanded and truncate. Pronotum rounded, widest at about middle, PW/PL = 1.32–1.39; lateral margins fully rounded near the middle and then gradually curved posteriorly, almost straight before hind angles; hind angles obtuse, not pointed; one mid-lateral seta present at about anterior third; basal fovea shallow, faintly defined; inner groove of basal fovea weak, subparallel to median line, gradually disappeared posteriorly, ended at a distance from the posterior margin of pronotum; outer groove of basal fovea about two-third length as inner one, somewhat more distinct than the inner one, reaching the basal margin of pronotum; ridge between outer groove and lateral margin distinct, weakly convex; area between inner and outer basal foveal grooves slightly convex and rugose; basal foveal area densely punctate, some sparse punctures usually present in the area between two basal foveae. Elytra with base slightly depressed between 3rd and 5th intervals; elytral shoulders slightly narrowed, humeral teeth small but distinct; intervals feebly convex; striae moderately deep, with very faint punctures; scutellar striae complete or incomplete, between 1st stria and elytral suture or between 1st and 2nd striae; umbilical pore series on 9th interval sparse in the middle, composed of 15–17 pores (5–6, 2–3, 7–8). Ventral side. Proepisterna finely rugose, usually with sparse and fine punctures at anterior half; metaepisterna finely punctate; terminal sternum of males without special structure. Legs. 5th tarsomere with one to three pairs of setae beneath. Male genitalia. Median lobe of male genitalia bent at a 90 degrees angle at about basal two-fifth, ventral margin almost straight in the middle (Fig. 36 A); apical part straight and then the apical lamella slightly bent to the dorsal side (Figs. 65, 66); apical orifice large, slightly turned to the left side, apical lamella twisted to the left (Fig. 36 B); apical lamella laminar, length about twice as its basal width, apex slightly capitate and rounded, hardly bent to the left (Fig. 44). Right paramere dirk-form, elongate and curved, right margin slightly curved, narrowed apically, apex sharp (Fig. 73 B). Female genitalia (Fig. 88) very similar to those of Pt. diversus Fairmaire. Spermatheca with the seminal canal about ten times long as the receptaculum; spermathecal gland strongly expanded; the seminal canal inserted at the base of common oviduct, base of seminal canal sclerotized (as in Fig. 85). Stylomere 2 with two or three ensiform setae at the outer margin and three or four at inner margin (Fig. 91).
Distribution. This subspecies is only known from Cangshan Mt., near Dali (Xiaguan Township) (Figs. 110, 111).
Etymology. The Latinlized name " cangshanensis " refers to the type locality of the new subspecies, Cangshan Mt., a mountain range located on the west of Dali, Yunnan.
Remarks. We propose this new subspecies mainly for the presence of setae on 5th tarsomeres, which is an important character in Pterostichus, even on subgenera level. Besides this, the external and genital differences (see diagnosis section) are constant though slight, and its geographical range is isolated from the nominotypical subspecies (Cangshan and Dapingdi belong to mountain ranges isolated by Erhai basin), which support the validity of the new subspecies. But the differences between their male genitalia are so small that we prefer to regard it as a subspecies rather than a different species.
description: Published as part of Shi, Hongliang, Sciaky, Riccardo, Liang, Hongbin & Zhou, Hongzhang, 2013, A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions, pp. 101-135 in Zootaxa 3664 (2) on pages 118-119, DOI: 10.11646/zootaxa.3664.2.1, http://zenodo.org/record/220503