Leptochirus (Tropiochirus) pachoensis Bernhauer 1917
Creators: name: Asenjo, Angelico
name: Ribeiro-Costa, Cibele S.
Publisher: name: Zenodo
Year: 2013
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Leptochirus
Leptochirus pachoensis

description: Leptochirus (Tropiochirus) pachoensis Bernhauer, 1917   
(Figures 2E, 3E, 5, 6G–L)  
 Leptochirus (Tropiochirus) pachoensis Bernhauer, 1917: 47 [original description, comparison with L. (T.) proteus Fauvel, 1864. Type locality: “Westkolumbien: Pacho in den Ostkordilleren, 2000 m ”]; Scheerpeltz, 1933: 1000 [world catalog].   
 Leptochirus pachoensis Blackwelder, 1944: 110 [checklist from Neotropical region]; Herman, 2001: 1105 [world catalog]; Newton, Gutiérrez and Chandler, 2005: 30 [checklist from Colombia].   
 Type material. Leptochirus (T.) pachoensis. Holotype (female): “Pacho Col. [Colombia] / dh.es will / 2000 m oklis” “Columbia occ.[western] / cali, Fassl” “pachoensis / Bernh. Typus unic” “Chicago NHMus / M.Bernhauer / Collection” (FMNH). Note: In the original description, Bernhauer (1917) specified that he studied one specimen. We received from the FMNH a female with the label “Typus”, therefore the specimen is the holotype for the species.    Additional specimens   
  ECUADOR. Napo:  2 km south from Oritoyacu and 22km south from Baeza, 1500 m, 4-5-iii.1971, J. M. Campbell (1 male CNC); idem 4-5-iii.1976 (1 female ZMHB);  COLOMBIA. Distrito Capital:  Bogotá (1 female ZMHB); Nueva Granada (1 male DZPU, 1 female ZMHB).    Diagnosis   
 Leptochirus (T.) pachoensis and L. (T.) helleri have the accessory tooth of the left mandible far from the lateral longitudinal ridge (Figure 3A, E), but L. (T.) pachoensis can be distinguished by the first dorsal tooth of the left mandible bifurcate (Figure 3E).    Description   
BL: 14.73; BW: 2.46  
Body strongly depressed. Body all black except tarsi red-brown.  
 Head. Transverse (Figure 2E). Inner lateral teeth separated by a depression extending to base. Inner and outer lateral teeth fused and resembling a septum; median sulcus deep, sides of median sulcus parallel; laterofrontal depressions oval. Labrum with outer edge strongly angled and lateral-apical edge concave. Mandibles (Figures 2E, 3E) with lateral longitudinal ridge occupying less than two-thirds of apical region; external margin strongly developed, attenuated anteriorly. Left mandible (Figure 3E) with first dorsal tooth developed, straight and bifurcate, accessory tooth far from lateral longitudinal ridge; second dorsal tooth little developed and resembling a small hill; first ventral tooth with tooth accessory pointed. Submentum with setae arranged in row on anterolateral margin.   
 Thorax. Pronotum (Figure 2E) a little wider than long (PL: 2.54; PW: 2.98); superior marginal line of hypomeron not interrupted, not bent to dorsal region at anterior margin and bent to slightly dorsal region at posterior margin; without pair of small depressions near basal constriction. Inferior marginal line of hypomeron reaching anterior margin. Apex of prosternal process strongly expanded and resembling a circular plate, extending considerably beyond the lateral process of hypomeron. Metaventrite with microsculpture longitudinally undulate in median region and with small longitudinal median depression near posterior edge. Mesoscutellum with apex rounded. Elytra longer than broad and without pair of small depressions on posterior-lateral edge (EL: 3.27, EW: 2.95).   
 Abdomen. Cylindrical. Sternite III with keel of apex rounded. Tergite IX (Figure 6J) sclerotized with left and right plates separated, each with six or seven setae of various lengths. Tergite X (Figure 6J) membranous except for sclerotized apical part, with two pairs of long apical setae.   
 Male. Sternite IX (Figure 6K) composed of two plates, distal plate sclerotized except for membranous apex, anterior margin rounded, posterior margin sinuated and with two long setae; proximal plate less sclerotized, greater than apical, broadened in middle and with a projecting angle at posterior border. Aedeagus (Figure 6G–I) with median lobe slightly bulbous at base, pointed at apex, weakly sclerotized on dorsal side and a small plate sclerotized on ventral side; parameres short, fused dorsally.   
 Female. Similar to male, except for the gonocoxites divided longitudinally (Figure 6L). The distal plate with apex membranous and long seta; proximal plate with the posterior region pointed.    Distribution   
 Leptochirus (T.) pachoensis is currently known from Colombia. In the current study this species is listed from Ecuador (Figure 5).
description: Published as part of Asenjo, Angelico & Ribeiro-Costa, Cibele S., 2013, Revision of the neotropical subgenus Tropiochirus of the genus Leptochirus Germar 1824 (Coleoptera: Staphylinidae: Osoriinae), pp. 1257-1285 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 47 (19 - 20) on pages 1272-1274, DOI: 10.1080/00222933.2012.763053, http://zenodo.org/record/10091554