Eutrichodesmus armatocaudatus Golovatch, Geoffroy, Mauries &amp; VandenSpiegel 2009, sp. n.
Creators: name: Golovatch, Sergei
name: Geoffroy, Jean-Jacques
name: Mauriès, Jean-Paul
name: VandenSpiegel, Didier
Publisher: name: Zenodo
Year: 2009
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Polydesmida
Haplodesmidae
Eutrichodesmus
Eutrichodesmus armatocaudatus

description: <i>Eutrichodesmus armatocaudatus</i> Golovatch, Geoffroy, Mauriès &amp; VandenSpiegel, sp. n. urn:lsid:zoobank.org:act: B2718568-A85A-4FDE-80E9-574F67AB90E9 Figs 5-8. <b>Type material.</b> Vietnam, Thanh Hoa Prov., Pu Luong, Lung Cao, Hang Lang Lua Cave, 11.XII.2003, leg. L. Deharveng &amp; team (Vn0312-05), holotype &amp;male; (MNHN JC 310), paratypes: 1 &amp;male;, 1 juv. (MNHN JC 310), 1 &amp;male; (ZMUM); Ha Nam Ninh Prov., Cuc Phuong National Park, Cave of Prehistoric Man, 10.X.1998, leg. L. Deharveng (VIET-537), 1 &amp;male; (MNHN JC 310); same locality, Water Fairy Cave, 11.X.1998, leg. L. Deharveng (VIET-541), 1 &amp;male;, 2 juv. (MNHN JC 310); same locality, cave 4 (Son Cung Cave?), 21.VI.2003, leg. P. Trontelj &amp; B. Sket, 1 &amp;male;, 1 &amp;female;, 3 juv. (MNHN JC 310), 1 &amp;male;, 1 juv. (ZMUC), 1 &amp;male;, 1 juv. (NMNHS), 1 &amp;male; (MCSNV), 1 &amp;male;, 1 juv. (OBBFUL), 1 &amp;female; (SEM); Ha Nam Ninh Prov., outside Cuc Phuong National Park, cave 3, 20.VI.2003, leg. P. Trontelj &amp; B. Sket, 1 &amp;male;, 1 &amp;female; (ZMUM). <b>Name.</b> To emphasize the obvious dorsal projections on body segments 17-19. <b>Diagnosis.</b> Most similar to <i>Parapauroplus monodentus</i> Zhang <i>in</i> Zhang &amp; Wang, 1993, which also shows evident mid-dorsal projections only on body segments (16)17- 19, but differs by the presence of porosteles, the slightly lower metatergal tubercles devoid of microsetae near each of the main setae, and the slightly stouter gonopod telopodite showing a shorter midway process. <b>Description.</b> Length of adults of both sexes <i>ca</i> 10-14 mm, width 2.3-2.6 mm, body broadest at segment 3 or 4; &amp;male;&amp;male; usually a little smaller than &amp;female;&amp;female;. Holotype <i>ca</i> 11 mm long and 2.5 mm wide. Coloration uniformly pallid or light yellowish. Adults with 20 segments, pattern of conglobation typical of “ Doratodesmidae ” (Figs 5A). Head (Fig. 6A) and tegument (Figs 5B, C; 6F, G) as in preceding species, but antennae somewhat longer (Fig. 6A), collum slightly flattened mid-dorsally, not covering the head from above (Fig. 6A). Metaterga behind collum with three transverse, rather irregular rows of tubercles (Figs 5 A-C), most of which still retain short, bisegmented setae (Fig. 6G), metaterga 17-19 each with an evident dorsal outgrowth (Fig. 5C). Paraterga mostly directed ventrolaterad, rather broad, slightly surpassing level of venter, clearly trisinuate caudolaterally at base, due to two lobulations (Figs 5B, D; 6B); paraterga 2 strongly enlarged, subvertical, margin nearly entire, with only very faint traces of a series of small lobulations anterolaterally (Fig. 6A), both schism and hyposchism very small; paraterga 3 and 4 slightly shorter than others, overlap of following paraterga typical, latter broadly rounded and slightly 3- or 4-lobate. Limbus spiculate (Fig. 6F). Pore formula normal, ozopores located on top of a small porostele, slightly above ventrocaudal lobulation (Figs 6 C-E). Pleurotergal carinae wanting. Epiproct elongated, also with tubercles, directed ventrocaudad, with the usual four cones just below tip; para- and hypoprocts as in Figs 5E, F. Sterna usually with a deep, narrow depression between coxae (Figs 5D; 6B), only sterna between coxae 6, 7 and 9 much wider (Fig. 7A). Gonopod aperture transverse-oval, relatively small, far from reaching lateral sides of segment 7. Legs relatively long; femoral and tarsal segments longest and equal; claw simple, very slightly curved ventrad; some setae very sparsely microdenticulate (Figs 5D; 6B). Gonopods (Figs 7; 8) very simple. Coxae subquadrate, large, microtuberculate and abundantly setose ventrolaterally, with a conspicuous rounded lobe frontolaterally. Telopodite considerably longer than coxite, slightly stouter, setose in its basal half, with an inconspicuous, digitiform, simple, lateral, distofemoral process (dp) at about midway, more distally with a slightly folded and enlarged shaft (= solenomere) with dense pilosity and a hairpad; seminal groove terminating subapically, acropodite short. <b>Remarks.</b> This species seems to be close to some of the “doratodesmids” showing evident mid-dorsal projections on some of the metaterga, e.g. <i>Ascetophacus macclurei</i> Hoffman, 1977, <i>Cerastelachys cavernicola</i> (Sinclair, 1901), <i>Pocillidorsus dorsiangulatus</i> Zhang <i>in</i> Zhang &amp; Wang, 1993 and <i>Parapauroplus monodentus</i> Zhang <i>in</i> Zhang &amp; Wang, 1993, etc. (Hoffman 1977a, 1977b; Zhang and Wang 1993). Cave 4 also supports another millipede, <i>Glyphiulus mediobliteratus</i> Golovatch, Geoffroy, Mauriès &amp; VandenSpiegel, 2007 (Golovatch et al. 2007b).
description: Published as part of <i>Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; VandenSpiegel, Didier, 2009, Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida), pp. 1-53 in ZooKeys 7 (7)</i> on pages 13-16, DOI: 10.3897/zookeys.7.117, http://zenodo.org/record/576436
description: {"references": ["Zhang Chunzhou, Wang Daqing (1993) Diplopoda in caves of Yunnan - 1. A study of new genera and species of the millipede family Doratodesmidae (Diplopoda: Polydesmida). In: Song Linhua, Ting Huaiyuan (eds) Karst Landscape and Cave Tourism. China Environmental Science Press, Beijing: 205 - 220.", "Hoffman RL (1977 a) The systematic position of the diplopod family Doratodesmidae, and description of a new genus from Malaya (Polydesmida). Pacific Insects 17 (2 - 3): 699 - 719.", "Sinclair FG (1901) On the myriapods collected during the \" Skeat Expedition \" to the Malay Peninsula, 1899 - 1900. Proceedings of the Zoological Society of London 2: 505 - 533.", "Hoffman RL (1977 b) Diplopoda from Malayan caves collected by M. Pierre Strinati. Revue suisse de Zoologie 84 (3): 699 - 719.", "Golovatch SI, Geoffroy J-J, Mauries J-P, VandenSpiegel D (2007 b) Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda: Spirostreptida: Cambalopsidae). Part 2. The javanicus - group. Zoosystema 29 (3): 417 - 456."]}