Canthidium (Neocanthidium) kelleri
Creators: name: Carvalho De Santana, Edrielly C.
name: Pacheco, Thaynara L.
name: Vaz-De, Fernando Z.
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Canthidium
Canthidium kelleri

description: Canthidium (Neocanthidium) kelleri (Martínez et al., 1964)
Figs 1B, 5, 8
Neocanthidium kelleri Martínez et al., 1964: 172–177 (original description).
Dichotomius (Luederwaldtinia) paraguayanus Gandini & Aguilar, 2009: 139, figs 1–8 (original description) (synonymized by Nunes & Vaz-de-Mello 2013: 416).
Canthidium (Canthidium) kelleri – Martínez & Halffter 1986: 28 (new combination). — Vaz-de-Mello 2000: 191 (checklist). — Nunes & Vaz-de-Mello 2013: 416 (senior synonymy).
Canthidium (Neocanthidium) kelleri – Cupello 2018: 470 (new subgeneric assignment).
Diagnosis
Canthidium kelleri differs from other species of the gigas group by the presence of a conical projection on the head which varies in development according to the size of the specimen.
Type specimens examined
Neocanthidium kelleri
Holotype ARGENTINA • ♂; handwritten label; “ARGENTINA/ MISSIONES / LORETO / En mongas him./ Coll. Martínez // Enc: 958”; “HOLOTYPUS”; “ Neocanthidium / kelleri sp. n. ♂ / M., H y P./ A. MARTÍNEZ DET.1963”; “ MACN-En 1223 ”; MACN.
Dichotomius paraguayanus
Holotype
PARAGUAY • ♂; “PARAGUAY: Concepcion, Zanja Moroti. 2.xi.2004 ”; originally in Gandini’s collection, now deposited in CEMT; CEMT.
Paratypes
PARAGUAY • 1 ♀; Cordillera, Cerro Naranjo; 25°30 ′ S, 57°04 ′ 59 ″ W; 15 May 2005; J. Rivas leg.; CEMT.
Other specimens examined
BRAZIL • 1 ♀; Distrito Federal, Brasília; 15°47 ′ 39.22 ″ S, 47°52 ′ 55.81 ″ W; R.F. IBGE; Dec.1997; I. Diniz leg.; CEMT • 1 ♀; Brasília, Fazenda Água Limpa; 15°57 ′ 11.05 ″ S, 47°58 ′ 23.76 ″ W; 16 Sep. 1979; Amadur leg.; CEMT • 1 ♂ [paratype of N. kelleri]; Goiás, Rio Verde; 17°47 ′ 33.79 ″ S, 50°55 ′ 10.77 ″ E; Dr. Nick leg.; CEMT • 1 ♀; Mato Grosso, Cuiabá, Fazenda [“Faz.”] Mutuca, Eucalipto-5; 15°18 ′ 51 ″ S, 55°58 ′ 18 ″ W; 6 Mar. 2009; CEMT • 1 ♀; same locality as preceding, Cerrado-1; 3 Feb. 2009; CEMT • 1 ♂; Chapada dos Guimarães, Cachoeira da Geladeira; FIT; 15°25 ′ 25 ″ S, 55°42 ′ 58 ″ W; 6,200 m a.s.l.; 9–30 Jan. 2013; D. Gimo leg.; CEMT • 1 ♀; same locality as for preceding, Comunidade João Carro; 15°02 ′ 01 ″ S, 55°43 ′ 00 ″ W; 300 m a.s.l., 3 May 2016; M. Serrano leg.; CEMT • 1 ♂; Mato Grosso do Sul; Selvíria, UNESP farm, Brachiaria decumbens pasture; black light trap; 20°25 ′ 6.82 ″ S, 51°20 ′ 51.87 ″ W; 21 Sep. 2000; C.A.H. Flechtmann leg.; CEMT • 1 ♀; Minas Gerais, Cordisburgo, Fazenda [“Faz”] Pontinha; 19°5 ′ 42.30 ″ S, 44°17 ′ 3.58 ″ W; Jan. 1994; F.Z. Vaz-de-Mello leg.; CEMT • 1 ♂; same locality as for preceding; Jan. 1999; Falqueto and Vaz-de-Mello leg.; BMNH • 1 ♀; same data as for preceding; CEMT • 1 ♀; Uberaba; 19°44 ′ 50.52 ″ S, 47°56 ′ 20.95 ″ W; CEMT • 1 ♀; Lavras; 21°18 ′ 50.15 ″ S, 44°59 ′ 36.95 ″ W; 20 Jan. 2008; M.R. Rocha and D.H.T. Takahashi leg.; CEMT • 1 ♀; Arinos, Rio Urucuia; 16°14 ′ 34.15 ″ S, 45°32 ′ 44.46 ″ W; 20 Apr. 1994; N. Degallier leg.; BMNH • 1 ♂; Buritis, Fazenda [“Faz”] Querência; FIT; 15°10 ′ 23,5 ″ S, 46°32 ′ 43,1 ″ W; 978 m a.s.l.; 10 Jul. 2013 – 4 Feb. 2014; C.M. Oliveira leg.; CEMT.
Redescription
HEAD. Surface densely and distinctly punctated, especially on the clypeus. Frontoclypeal suture with conical projection.
PRONOTUM. Regularly convex, without projections, sometimes excavated anteriorly. Anterior edge slightly curved, sides arched.
HYPOMERON. Hypomeral carina incomplete.
METAVENTRITE. With simple and shallow punctation, laterobasal region with stronger punctation, lateral region with microsculpture obscurely variolate, almost semicircular.
MESEPIMERON AND MESEPISTERNUM. With shallow punctation, stronger microsculpture obscurely variolate, almost semicircular.
HIND LEG. Metafemora with a carina along posterior edge, anterior edge with setal fringe, surface with fine punctuation. Metatibiae elongated and gradually dilated towards the apex; external margin serrated.
AEDEAGUS. Parameres, in lateral view, with dorsal angulation at basal two-fifths.
Variation and sexual dimorphism
Length ranging from 8.95 to 11.54 mm. Smaller specimens have the frontal horn reduced in size, appearing as a transverse gibbosity. Pronotum with the anteromedial depression less visible and shallower in some specimens. Pygidium slightly less convex if compared with C. bokermanni. Males present a small tooth in the upper ventral part of their metatibiae. Males with ventrite VI wider medially, female ventrite VI with the same length throughout. Males also have protibial spur flattened, while in females the protibial spur is narrowed and pointed.
Geographical distribution
The specimens examined were collected in Paraguay (Cordillera) and Brazil (Mato Grosso, Mato Grosso do Sul, Goiás, Distrito Federal and Minas Gerais). It is present in the Cerrado and the Chaco biomes, which lie in the Chacoan dominion.
Comments
The holotype and one paratype originally identified as male of D. paraguayanus were examined by us and were shown to be C. kelleri, thus confirming the name’s synonymy with C. kelleri as firstly proposed by Nunes & Vaz-de-Mello (2013). However, two paratypes originally said to be females by Gandini & Aguilar (2009) are actually males, and belong to C. bokermanni. See comments below.
description: Published as part of Carvalho De Santana, Edrielly C., Pacheco, Thaynara L. & Vaz-De, Fernando Z., 2019, Taxonomic revision of the Canthidium Erichson, 1847 species of the gigas group (Coleoptera, Scarabaeidae, Scarabaeinae), pp. 1-24 in European Journal of Taxonomy 530 on pages 9-11, DOI: 10.5852/ejt.2019.530, http://zenodo.org/record/3243046