Landouria petrukensis Nurinsiyah & Neiber & Hausdorf 2019, sp. nov.
Creators: name: Nurinsiyah, Ayu Savitri
name: Neiber, Marco T.
name: Hausdorf, Bernhard
Publisher: name: Zenodo
Year: 2019
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Camaenidae
Landouria
Landouria petrukensis

description: Landouria petrukensis sp. nov.
urn:lsid:zoobank.org:act: 32D7E6E9-561F-4E05-95D2-66295BFD6C5E
Figs 19, 33, 50, 52; Tables 3–4
Diagnosis
Landouria petrukensis sp. nov. is characterized by a rather high, sharply keeled shell with small scales all over the teleoconch, a penis that is divided into a long, broad, cylindrical distal part and a slightly dilated proximal part by a narrow point and a flagellum, which is broadest near its proximal end.
Etymology
Landouria petrukensis sp. nov. is named after its occurrence in the surroundings of Petruk Cave in Karangbolong.
Material examined
Holotype
INDONESIA • holotype (det. anat.); Karangbolong, way to Petruk Cave; 7°42′ S, 109°24′ E; 78 m a.s.l.; 5 Aug. 2014; A.S. Nurinsiyah, F.L.H. Irsyad, F.J. Sari and E. Nurlaela leg.; D = 8.7 mm, H = 5.2 mm; MZB 19203.
Paratypes
INDONESIA – Central Java • 6 spec.; Karangbolong, near Petruk Cave, agroforestry; 7°42′17″ S, 109°23′59″ E; 78 m a.s.l.; MZB 19273 • 2 spec.; same data as for preceding; ZMH 133409 • 1 spec.; same data as for preceding; MZB 19204 • 5 spec.; Karangbolong, near Petruk Cave, agroforestry; 7°42′19″ S, 109°24′02″ E; 78 m a.sl.; MZB 19265 • 4 spec.; same data as for preceding; ZMH 133417 • 4 spec.; Karangbolong, wall and surroundings of Petruk Cave; 7°42′21″ S, 109°24′03″ E; 104 m a.s.l.; MZB 19266 • 3 spec.; same data as for preceding; MZB 19267 • 5 spec.; Karangbolong, near Petruk Cave, agroforestry; 7°42′18″ S, 109°24′05″ E; 138 m a.s.l.; MZB 19274 • 1 spec.; same data as for preceding; ZMH 133407 • 23 spec.; Karangbolong, Adiraja Hill, agroforestry; 7°42′48″ S, 109°24′05″ E; 168 m a.s.l.; MZB 19271 • 4 spec.; Karangbolong, Adiraja Hill, teak plantation; 7°42′50″ S, 109°24′05″ E; 166 m a.s.l.; MZB 19275 • 3 spec.; same data as for preceding; ZMH 133414.
Description
SHELL (Fig. 50; Tables 3–4). Depressed conical, with 4.75–5.5 hardly convex whorls; protoconch almost smooth; teleoconch with irregular wrinkles, apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes all over teleoconch; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture oval; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 24–37% of shell diameter, hardly obscured by columellar edge.
GENITALIA (Figs 19, 33; Table 4). Atrium short; penis divided by a narrowing into a long, broad cylindrical distal part and a slightly dilated proximal part, without penis coecum; membranaceous sheath around distal part of penis connected with proximal part of epiphallus by tissue; flagellum moderately long, broadest near its proximal end, where it rapidly tapers, slightly crenated, without node; vas deferens narrow and long; vagina short, distally dilated; oviduct longer, proximally hardly tapering; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part. Right ommatophoral retractor runs between penis and vagina.
Remarks
With regard to the presence of scales all over the teleoconch, L. petrukensis sp. nov. resembles L. tholiformis sp. nov. (Fig. 51) and L. madurensis sp. nov. (Fig. 57). It differs from both species in the smaller (D 8.1–9.8 mm vs 13.0– 14.4 mm in L. tholiformis sp. nov. and 11.4–12.0 mm in L. madurensis sp. nov.), more depressed shell (D/H 1.49–1.79 vs 1.77–1.88 in L. tholiformis sp. nov. and 1.75–2.11 in L. madurensis sp. nov.) with fewer whorls (4.75–5.5 vs 6–6.25 in L. tholiformis sp. nov. and 5.5–5.75 in L. madurensis sp. nov.), an on average narrower umbilicus (U/D 0.24–0.37 vs 0.32–0.40 in L. tholiformis sp. nov. and 0.38–0.40 in L. madurensis sp. nov.) and in the penis that is divided by a constriction into a long, broad, cylindrical distal part and a slightly dilated proximal part.
We could only investigate the anatomy of the holotype, from Karangbolong in southern Central Java (MZB 19203). There are several samples from other regions of Central Java, West Java and Banten with similar shells. Thus, it is possible that this species is more widely distributed, but the identity of the samples from other regions must be confirmed anatomically.
Distribution
Landouria petrukensis sp. nov. is known from Karangbolong in southern Central Java (Fig. 52).
description: Published as part of Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, pp. 1-73 in European Journal of Taxonomy 526 on pages 36-37, DOI: 10.5852/ejt.2019.526, http://zenodo.org/record/3152217