Empis (Enoplempis) submanca Sinclair, Brooks & Cumming, 2025, sp. nov.
Creators: name: Sinclair, Bradley J.
name: Brooks, Scott E.
name: Cumming, Jeffrey M.
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Empididae
Empis
Empis submanca

description: Empis (Enoplempis) submanca sp. nov.
urn:lsid:zoobank.org:act: 71232F6D-45A8-4716-8DF3-3ACBA583C94E
(Figs 199–204, 208)
Type material. HOLOTYPE ♂, labelled: “USA: CA: Tulare Co. / Ash Mtn. Power Stn. 3 [36°29′N, 118°50′W] 3/ 8.iv.1993 ”; “ CNC/ 1078373 ”; “ HOLOTYPE / Empis (Enoplempis) / submanca Sinclair,/ Brooks & Cumming [red label]” (CNC). PARATYPES: USA. California: El Dorado County: El Dorado Hills, New York Creek, 680 ft, 38°41′43″N 121°04′38″W, 17.iii.2004, S.D. Gaimari (7♂, 2♀, CSCA); same data except, P.H. Kerr (1♂, CSCA). Kern County: Havilah [35°31′04″N 118°31′07″W], 3000 ft, 28.iv.1964, JAP (1♂, EMEC); same data except, 2 mi. NE, 3900 ft, C.A. Toschi (1♂, EMEC); Miracle Springs [35°34′N 118°32′W], 29.iv.1964, C.A. Toschi (1♂, EMEC); 8 mi. W Wolford Hts. [35°42′N 118°27′W], 28.iv.1964, Coreopsis bigelovii, P. Rude (1♂, EMEC). Mendocino County: NCCRP, 3 mi. N Branscomb [39°42′N 123°39′W], 5–6.v.1989, MT, S. O’Keefe (1♂, EMEC). Stanislaus County: Del Puerto Cyn, 1200 ft, Frank Raines Pk [37°25ʹN 121°22ʹW], 9.iv.1971, P. Loggins (1♂, EMEC). Tulare County: Same data as holotype (1♂, 1♀, CNC); same data as holotype except, Ash Mtn Kwh Power Sta. 3, 23.iii.1993, J.F. MacDonald (1♀, CNC); same data except, 11.iv.1993 (1♂, CNC); same data except, 27.iv.1985 (3♂, 3♀, CNC); Ash Mt. HQ [36°29′N 118°49′W], 1700 ft, 27.iv.1979, R. Coville (2♂, EMEC); same data except, 28.iv.1979, JAP (1♂, EMEC); same data except, 30.iv.1979, J. T. Doyen (1♂, EMEC); same data except, 1–2.v.1979, J.A. Chemsak (1♂, 1♀, EMEC); 0.5 mi. SE Ash Mtn Seq. NP HQ, 7.iv.1984, R.D. Haines, flume debris trap (2♂, 4♀, CSCA); same data except, 4.ii.1984 (1♀, CSCA); same data except, 25.iv.1984 (1♂, 2♀, CSCA); same data except, 3.v.1984 (2♂, 1♀, CSCA); Coffee Camp [36.1524°N 118.7464°W], 1.v.1979, J. T. Doyen, beating Quercus wislizenii (1♂, 1♀, EMEC); Kern River, Campgrd No. 3, 2900 ft, 11.v.1971, PHA & M. Arnaud (1♂, USNM).
Diagnosis. This species is distinguished from other species of the E. (En.) manca species group by the male hind femur with row of strong posteroventral setae opposite sickle-shaped process (Fig. 204), sickle-shaped process greatly tapered and narrowly prolonged apically, male hind tibia with low, short anteroventral lobe on basal quarter, and phallus with short, subapical projection.
Description. Wing length 5.6–6.8 mm. Male. Similar to E. (En.) manca, except as follows: head with postpedicel long and tapered, 4.25× longer than basal width.
Thorax with postpronotal lobe and postalar ridge pale brown.Scutum with dc longer, stronger than acr, increasing in length posteriorly with prescutellar dc long, strong. Scutellum with 1 pair of long sctl, with outer marginal setae, more than half length of apical pair; sometimes pair of setulae between apical pair.
Legs short, stocky; yellow to yellowish brown, including coxae, slightly darker on apical tarsomeres; coxae often with thin greyish pruinescence (Fig. 201). Hind femur with strongly arched, anteroventral sickle-shaped process, drawn out and narrowly prolonged apically; row of slender anteroventral setae proximal to sickle-shaped process; short knob-like process distal to process; row of strong posterior setae opposite sickle-shaped process; short posteroventral ridge distal to sickle-shaped process bearing row of strong, arched setae, appressed distally; small semi-circular preapical posteroventral ridge with appressed crown of setae; apex without preapical row of posterior setae; with strong preapical anterior and anterodorsal setae (Figs 202–204). Hind tibia slightly twisted at base, not greatly narrowed; low, short anteroventral lobe with tuft of apical setae on basal quarter; row of strong anteroventral and posteroventral setae proximal to lobe, posteroventral setae longer proximally; posteroventral row curved medially to link with anteroventral row; seta in posteroapical comb; anterodorsal and posterodorsal setae slightly longer than width of tibia (Figs 201–204).
Terminalia (Figs 199, 200): yellowish brown. Cercus slender, rounded apically; dorsal margin straight; setae slender, shorter than width of cercus; base weakly fused to epandrial lamella. Subepandrial process extending on either side of phallus. Epandrial lamella narrow, linear, longer than height; posterior margin strongly upturned; bearing long setae ventrally, subequal to width of lamella, including 1–2 stronger setae. Hypandrium short, apical margin rounded, not prolonged laterally; without setae. Phallus (Fig. 200) with basal third expanded, tapered, gently arched anteriorly; apex emerging beyond cercus; apex expanded with broad opening, with short subapical projection; roughened, spiculate apically; ejaculatory apodeme elongate, half-length of epandrium, T-shaped, with lateral apodemes on lower third.
Female. Similar to female of E. manca.
Geographical distribution and seasonal occurrence (Fig. 208). Empis (En.) submanca sp. nov. is known from northern and central California. Adults have been collected from February to May.
Etymology. The species name is in reference to the similarity of this species with E. manca.
Nuptial gift presentation. Form unknown.
description: Published as part of Sinclair, Bradley J., Brooks, Scott E. & Cumming, Jeffrey M., 2025, Revision of the western Nearctic species of Empis subgenus Enoplempis (Diptera: Empididae), pp. 1-200 in Zootaxa 5615 (1) on pages 107-109, DOI: 10.11646/zootaxa.5615.1.1, http://zenodo.org/record/15217957