Achilia fiura Kurbatov & Cuccodoro & Sabella 2018, n. sp.
Creators: name: Kurbatov, Sergey A.
name: Cuccodoro, Giulio
name: Sabella, Giorgio
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Achilia
Achilia fiura

description: Achilia fiura n. sp.
Figs 3, 15, 18, 23, 54, 56, 58, 83
Type material (13 ex.): CENTRAL CHILE: Región Araucanía: Malleco prov.: MHNG; 1 ♂ (holotype); 45 km W Angol; 1400 m; 09.XII.1984 / 16.II.1985; S. & J. Peck; Nothofagus forest litter, car trap. – MHNG; 5 ♂ (paratypes); same data; S. & J. Peck; Nothofagus forest litter, car trap. – MNHS; 1 ♂ (paratype); same data; S & J. Peck; Nothofagus forest litter; car trap. – MHNG; 2 ♂ (paratypes); Nahuelbuta National Park, 40 km W Angol; 1200-1500 m; 19.XII.1984 / 17.II.1985; S. & J. Peck; Nothofagus-Araucaria; for FIT. – MHNG; 3 ♂ (paratypes); 17 km W Angol; 800 m; 08.XII.1984 / 16. II.1985; S. & J. Peck; FIT mixed Nothofagus. – FMNH (FMHD #2002-95); 1 ♂ (paratype); Nahuelbuta National Park, 2.3 km W Los Portones entrance; 37° 49.41’S 72° 58.95’W; 1150 m; 25.XII.2002; Solodovnikov 1057; Nothofagus dombeyi + antarctica, mostly open understory, berlese, leaf & log litter, forest floor.
Description: Body 1.45-1.50 mm long, usually entirely reddish with darker head and abdomen, and palpi yellowish. Head with eyes longer than temples. Pronotum wider than head, with maximal width on anterior half; median antebasal fovea as large as lateral ones. First abdominal tergite with basal striae slightly diverging, extending to less than one-third of paratergal length, and separated at base by more than one-third of tergal width.
Male: Head as in Figs 54 and 56, flanked on entire length by two very deep pits with outer edge toothed at level of eyes (Fig. 58). Narrow frontal protuberance slightly convex at base and anteriorly flattened to vertexal sulcus. Antennae (Fig. 15) with scape and pedicel longer than wide; antennomere III wider than long; antennomere IV transverse, with protruding mesal margin pointed in middle and bearing a tuft of long setae; antennomere V transverse, with protruding mesal margin pointed in middle and bearing a long seta; antennomeres VI-VII distinctly wider than long; antennomere VIII transverse, with protruding mesal margin; antennomeres IX and X strongly transverse, with protruding mesal margin bearing long setae; antennomere X wider and longer than IX; antennomere XI elongate and longer than VIII-X combined, its medial margin enlarged at base and bearing a tuft of long setae. Metasternum with large median impression. Legs with ventral margin of mesotrochanters (Fig. 23) forming short median spine; profemora and mesofemora slightly thickened; mesotibiae (Fig. 18) forming subbasal spur slightly recurved externally and with short stout apical seta; metatibiae slightly sinuate on distal half.Abdominal tergites unmodified; first abdominal ventrites with faint median impression, all other ventrites flattened at middle; ventrite I with short median carina extending from posterior margin the posterior edge of median impression. Aedeagus (Fig. 3) 0.28-0.30 mm long; dorsal plate ovoid with dorsal longitudinal struts divergent; copulatory pieces consisting of pair of long medial sclerites recurved and strongly sclerotised at base, and apically pointed, associated on each side with sclerite ending with two large spines. Parameres very wide with long seta on well-developed outer lobe; tips strongly recurved posteriorly and bearing wide subapical seta.
Female: Unknown.
Collecting data: Collected from December to February in Nothofagus and Araucaria forests, where it was found from 1150 m to 1500 m. Most specimens were taken by flight intercept traps and by car netting; one specimen was collected in sifted samples of leaf and log litter.
Distribution: Achilia fiura is known only from Malleco Province (Región Araucanía) (Fig. 83: green diamonds).
Comments: Achilia fiura is very similar to A. excisa and A. pachycera, from which it is easily distinguished by the male features of the antennae (compare Figs 13 and 15), mesotrochanters (compare Figs 20, 21 and 23), mesotibiae (compare Figs 16, 17 and 18), and the copulatory pieces of the aedeagus (compare Figs 1 and 4-5; 6-8, and 3).
description: Published as part of Kurbatov, Sergey A., Cuccodoro, Giulio & Sabella, Giorgio, 2018, A revision of the Chilean Brachyglutini – Part 3. Revision of Achilia Reitter, 1890: A. frontalis species group (Coleoptera: Staphylinidae: Pselaphinae), pp. 165-188 in Revue suisse de Zoologie 125 (1) on pages 172-174, DOI: 10.5281/zenodo.1196048