Olegia mandici Harzhauser & Landau & Janssen 2022, nov. sp.
Creators: name: Harzhauser, Mathias
name: Landau, Bernard
name: Janssen, Ronald
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Phylloxeridae
Olegia
Olegia mandici

description: Olegia mandici nov. sp.   
 Figs 26A 1 –A 2, B 1 –B 2, C 1 –C 2, D 1 –D 2, 3 Ac, 5, 7   
 Pleurotoma doderleini Hörn. — Pilar 1874: 216, pl. 2, figs 20–21 [non Olegia doderleini (Hörnes, 1854)].   
 Clavatula doderleini brigittae Hoernes & Auinger, 1892 — Strausz 1966: 410, pl. 16, fig. 15, pl. 17, fig. 1 [non Olegia brigittae Hörnes & Auinger, 1891]   
 Type material. Holotype: NHMW 2021 /0140/0001, SL: 27.8 mm, MD: 11.2 mm, Bekića vrelo, Čremušnica at Donji Kirin (Croatia), figs 26A 1 –A 2. Paratypes: NHMW 2021 /0140/0002, SL: 25.3 mm, MD: 10.5 mm, Bekića vrelo, Čremušnica at Donji Kirin (Croatia), figs 26B 1 –B 2; NHMW 1872 /0031/0008a, SL: 23.4 mm, MD: 9.4 mm, Bekića vrelo, Čremušnica at Donji Kirin (Croatia), figs 26C 1 –C 3; NHMW 1872 /0031/0008b, SL: 19.4 mm, MD: 8.4 mm, Bekića vrelo, Čremušnica at Donji Kirin (Croatia), figs 26D 1 –D 3; NHMW 1872 /0031/0008c, SL: 27.9 mm, MD: 13.5 mm, Bekića vrelo, Čremušnica at Donji Kirin (Croatia), fig. 3Ac; NHMW 1872 /0031/0008, 4 spec., Bekića vrelo, Čremušnica at Donji Kirin (Croatia).   
 Additional studied material. NHMW 1858 /0015/0148, 1 spec., Totterfeld at Hartberg (Austria); GBA 1875/006/0011, Hafnertal at Lichtenwald (Austria), illustrated in Hoernes (1875b, pl. 2, fig. 4).   
 Type locality. Bekića vrelo, Čremušnica at Donji Kirin (Croatia), Glina Basin.   
 Type stratum. Badenian siliciclastic deposits (no formation name available).   
 Age. Middle Miocene, Sarmatian (late Serravallian).   
 Etymology. In honor of Oleg Mandic (paleontologist at the NHMW and friend of the first author) in recognition of his contributions on Paratethyan stratigraphy.   
 Diagnosis. Olegia species of medium size, solid moderately slender fusiform shell, with strong sculpture of pointed tubercles on subsutural collar and rounded tubercles on suprasutural cord separated by wide, smooth concave mid-whorl portion. Four tubercular spiral cords on last whorl with secondary and tertiary spiral threads intercalated.   
 Description. Shell small, solid, moderately slender fusiform with conical spire; apical angle ~35°. Protoconch not preserved. Teleoconch of at least eight whorls. Early teleoconch whorls flat-sided with bipartite sculpture; welldefined, smooth subsutural cord, smooth mid-whorl portion. Suprasutural cord bearing large, rounded, slightly opisthocline tubercles. Later whorls with moderate width subsutural collar bearing pointed tubercles, concave midwhorl with comma-shaped riblets, and suprasutural row of larger orthocline tubercles. Entire surface covered by secondary and tertiary cords. Suture narrowly but deeply incised, weakly undulating, running just below abapical row of tubercles. Last whorl ~ 65% of total height; subsutural collar well-defined bearing about 14 small, sharp tubercles, subsutural ramp concave, delimited by tubercular shoulder cord, convex below, weakly to moderately constricted at base. Siphonal fasciole swollen, strongly twisted. Sculpture of four tubercular primary spiral cords. Adapical cord delimiting shoulder strongest, second cord at mid-whorl delimiting periphery, third peribasal, fourth perifasciolar, one further slightly weaker primary cord intercalated between shoulder and peripheral primaries and one or two on base between peribasal and perifasciolar cords. Threads of tertiary strength cover entire surface. Aperture moderately wide, ovate. Outer lip not thickened, smooth within. Anal sinus deep, moderately wide, symmetrically U-shaped, with apex just below collar. Siphonal canal moderately long and narrow, deflected to the left, shallowly notched at tip. Columella weakly excavated, twisted at fasciole, smooth. Columellar and parietal callus strongly thickened, sharply delimited, forming very broad callus rim and narrow pseudumbilical chink.   
 Discussion. Olegia mandici nov. sp. is reminiscent of Olegia rumana (Simionescu & Barbu, 1940) and might represent an endemic offshoot of that species. It differs from O. rumana in its larger size, the less numerous but larger beads at the abapical suture of early teleoconch whorls, the regular sculpture and more pointed tubercles, the wider spiral cords on the last whorl bearing larger tubercles, and the presence of secondary and tertiary spiral sculpture. The early Miocene Olegia agathae (Hoernes & Auinger, 1891) might be another closely related species, which differs especially in its smaller size and the higher, weakly constricted base. Olegia szokolyensis (Strausz, 1960) differs in its lower spire, the subcylindrical last whorl and the much weaker spiral sculpture on the base. This specimen is either a misplaced Sarmatian specimen or the only known Badenian representative.   
 Distribution in Central Paratethys. Sarmatian (middle Miocene): Styrian Basin: Totterfeld at Hartberg (Austria) (hoc opus), Hafnertal at Lichtenwald (Austria) (Hoernes 1875b); Glina Basin: Bekića vrelo, Čremušnica at Donji Kirin (Croatia) (hoc opus). Strausz (1966) described a specimen from the Badenian of Budapest as Clavatula doderleini brigittae, which seems to represent O. mandici based on its spiny sculpture and spiral sculpture.
description: Published as part of Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, pp. 1-172 in Zootaxa 5123 (1) on pages 85-86, DOI: 10.11646/zootaxa.5123.1.1, http://zenodo.org/record/6399827