Rhinolophus francisi subsp. thailandicus Soisook & Struebig & Noerfahmy & Bernard & Maryanto 2015, subsp. nov.
Creators: name: Soisook, Pipat
name: Struebig, Matthew J.
name: Noerfahmy, Sephy
name: Bernard, Henry
name: Maryanto, Ibnu
name: Chen, Shiang- Fan
name: Rossiter, Stephen J.
name: Kuo, Hao- Chih
name: Deshpande, Kadambari
name: Bates, Paul J. J.
name: Sykes, Dan
name: Miguez, Roberto Portela
Publisher: name: Zenodo
Year: 2015
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Mammalia
Chiroptera
Rhinolophidae
Rhinolophus
Rhinolophus francisi
Rhinolophus francisi thailandicus soisook and bates

description: Rhinolophus francisi thailandicus Soisook and Bates subsp. nov.
(Figs. 1, 3–6; Tables 1–2)
Holotype
PSUZC-MM.2008.51 (field number PS080420.6), adult male, body in alcohol, skull and baculum extracted, collected by Pipat Soisook, Tuanjit Srithongchuay, Piyawan Niyomwan and Priwan Srisom, on 20 April, 2008.
Type locality
Pu Nam Ron Stream, Mae Nam Pha Chi WS., Ratchaburi Province, Western Thailand (13º15’N, 99º2’E, 431 m a.s.l.).
Diagnosis
This subspecies is very similar to the nominate subspecies from Borneo with a FA of 52.97 mm and a SL of 24.37 mm. The rostral swelling is relatively narrower than in the specimens from Borneo, with an AMSW and ALSW of 4.76 and 6.53 mm, respectively. The 3D1P is relatively short; it is 66.74% of the length of the 3D2P. The sagittal crest is less
developed than in the nominate subspecies. Genetically, it differs from the nominate subspecies by 10.03% at a section of the cytochrome oxidase-I gene.
Etymology
The subspecific name, thailandicus, means ‘of or from Thailand’ indicating where this subspecies is found. The proposed English name of the subspecies is ‘ Thailand Woolly Horseshoe Bat’.
Description and taxonomic notes
As in the nominate subspecies from Borneo, the Thai subspecies, R. f. thailandicus is a mediumlarge Rhinolophus. The general appearance is similar to francisi from Borneo and overlaps in size, with a FA of 52.97 mm (Table 1). The body mass (MASS) is 16.0 g. The height of the ear is of 23.81 mm. The noseleaf width (NL) is 12.21 mm (Table 1). The wing measurements are larger than those of R. f. francisi; the third metacarpal (3MET) is 37.47 mm; the fourth (4MET) is 42.47 mm and the fifth metacarpal (5MET) is 43.99 mm (Table 1).
In the skull, the SL and CCL are 24.37 mm and 21.72 mm, respectively (Table 2). The AMSW and ALSW are of 4.76 mm and 6.53 mm, respectively. The supraorbital ridge is very well defined and connected to a well-developed sagittal crest (Fig. 3a). The zygomatic breadth (ZB) is 12.16 mm which exceeds the mastoid width (MW), 11.31 mm (Fig. 3a; Table 2). Each zygoma has a distinct triangular-shaped dorsal arch (Fig. 3a). Both the upper and lower dentition are as in R. f. francisi but are slightly smaller in size; C–M3 is 9.30 mm; C1–C1 6.07 mm; M3–M3 8.86 mm. The palatal length (PL) is 3.21 mm (Table 2).
The morphological comparison with other species is as in the nominate form of R. francisi. The size of Thai specimen is the smallest of the species. It is intermediate in size between the slightly larger R. beddomei and the slightly smaller R. trifoliatus (Tables 1–2; Fig. 5). As noted above, a future study with more specimens from Thailand may prove that the subspecies from Thailand is specifically distinct.
description: Published as part of Soisook, Pipat, Struebig, Matthew J., Noerfahmy, Sephy, Bernard, Henry, Maryanto, Ibnu, Chen, Shiang- Fan, Rossiter, Stephen J., Kuo, Hao- Chih, Deshpande, Kadambari, Bates, Paul J. J., Sykes, Dan & Miguez, Roberto Portela, 2015, Description of a new species of the Rhinolophus trifoliatus-group (Chiroptera: Rhinolophidae) from Southeast Asia, pp. 21-36 in Acta Chiropterologica 17 (1) on pages 33-34, DOI: 10.3161/15081109ACC2015.17.1.002, http://zenodo.org/record/3942447