Coneuplecta (Durgellina) calculosa
Creators: name: Hyman, Isabel T.
name: Ponder, Winston F.
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Euconulidae
Coneuplecta
Coneuplecta calculosa

description: Co <i>neuplecta</i> (<i>Durgellina</i>) <i>calculosa</i> (Gould, 1852) Figures 16D, H, 26G, 27G, 28G, 39D, 40D–F <i>Helix calculosa</i> Gould, 1852: 48. <i>Helix umbraculorum</i> Cox, 1864: Addendum, species No. 136. <i>Helix dendrobia</i> Crosse, 1868: 96–97, pl. i, fig. 5. <i>Helix</i> (<i>Conulus</i>) <i>russelli</i> Brazier, 1875b: 668. <i>Sitala?</i> <i>rimicola</i> (Benson) Blanford and Godwin-Austen, 1908: 236–237. <i>Coneuplecta</i> (<i>Durgellina</i>) <i>calculosa</i>: Baker, 1941a: 234–236, pl. 44, figs 8–9, pl. 55, figs 4–5, pl. 62, fig. 13; Solem, 1961: 471; Solem, 1988: 544, figs 173, 177–178; Smith, 1992: 227. <b>Material examined.</b> <b>New South Wales</b>: AM C374873 (one shell measured, photographed), Coffs Harbour, behind N end of Park Beach (30º17' S, 153º8' E), on leaves of sapling trees in littoral rainforest, Feb. 1983, M. Shea; AM C377743 (one specimen dissected), Zeehan Rd. at Cooks Creek Trail (30º31' S, 152º45' E), arboreal, rainforest regrowth, 13 Nov. 1999, J. Waterhouse; AM C377891 (one specimen dissected), Nymboi–Binderay National Park N of Bosterbrick, Mills Road 200–300 m from Moonpar Rd. (30º13' S, 152º39' E), arboreal, tall wet sclerophyll, vine thicket understorey, 9 Nov. 1999, M. Shea. <b>Description.</b> <i>External morphology</i>: Shell (Figures 16D, H, 26G, 27G, 28G) 4.7–5.1 whorls, pale golden brown, not glossy; protoconch sculptured with spiral and radial ribs (spirals stronger); teleoconch with spiral and radial lirae (radials stronger); shell shape as for genus. Animal white. Sole of foot and caudal apparatus as for genus; caudal horn very small. <i>Mantle cavity</i>: As for genus. Mantle with pigmentation of black and white spots, mantle gland present. <i>Digestive system</i>: As for genus. <i>Genital system</i> (Figure 39D): As for genus. Free oviduct long; capsular gland present. Bursa copulatrix very short, shorter than free oviduct; duct of bursa copulatrix wide, not distinguishable from bursa copulatrix. Vagina short. Penis moderately long, epiphallus enters penis through simple pore; penis internally smooth; basal diverticulum on penis present. Penial sheath present, enclosing only penis. Epiphallus shorter than penis. <i>Radula</i> (Figure 40D–F): As for genus. Central tooth with large ectocones and smaller accessory cusps. Two lateral teeth present, distinguished from marginal teeth by presence of endocone; marginal teeth with endocones absent; ectocones approximately same length and breadth as mesocone, subdivided into two to five extra teeth (increasing towards the outer edge). Radular formula (113.2.1.2.113) × 102 rows. <b>Range and habitat.</b> Within Australia, this species is found from Narooma, southern New South Wales to islands of Torres Strait in far northern Queensland and in the northwestern part of Northern Territory. Outside Australia, <i>Coneuplecta calculosa</i> has been recorded from India, New Caledonia, Western Samoa and the Society Islands (Solem 1988). <b>Remarks.</b> The anatomy and radula figured here for <i>Coneuplecta calculosa</i> is identical to that figured by Baker (1941a) for this species. No anatomical details are available for the type species of <i>Durgellina</i>, <i>Coneuplecta vitrina</i> (Thiele, 1928), which Schileyko (2002a) gives as a synonym of <i>C. calculosa</i>, although we were unable to confirm this. Solem (1988) is followed in including <i>Helix dendrobia</i> Crosse, 1868 and <i>Sitala rimicola</i> (Benson) Blanford and Godwin-Austen, 1908 as synonyms of <i>Coneuplecta calculosa</i>. The species figured by Odhner (1917) as <i>Sitala wilcoxi</i> (Cox, 1865) may also be <i>C. calculosa</i>, based on the radula and reproductive system. It is possible that <i>wilcoxi</i>, which has since been synonymised with <i>Sodaleta umbraculorum</i> (Iredale 1937c), is also a synonym of <i>C. calculosa</i>. Alternatively, Odhner (1917) may have misidentified the species he dissected. He did not figure a shell. In the phylogenetic analysis, <i>Coneuplecta calculosa</i> grouped with the other species of <i>Coneuplecta</i> and diverged from the base of the tree (Figures 1–3). In the molecular phylogenetic analysis of Hyman <i>et al.</i> (2007), <i>C. calculosa</i> also occupied a relatively basal position as the sister taxon to (Trochomorphidae + Microcystinae) (Figure 44A).
description: Published as part of <i>Hyman, Isabel T. &amp; Ponder, Winston F., 2010, A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462, pp. 1-148 in Zootaxa 2462 (1)</i> on pages 109-110, DOI: 10.11646/zootaxa.2462.1.1, http://zenodo.org/record/5312710
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