Scopaeus sulawesianus Frisch 2022, spec. nov.
Creators: name: Frisch, Johannes
name: Narakusumo, Raden Pramesa
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Scopaeus
Scopaeus sulawesianus

description: Scopaeus sulawesianus Frisch, spec. nov.
(Figs 24, 49, 116–118, 145, 178)
Type specimens: Indonesia: Holotype ♂, Sulawesi Selatan, Engrekang (Sadang River), (03°34’S, 119°46’E), 46 m, 27.7.2015, leg. Puchner (MZB). Paratypes (9 specimens): 2 ♂, 1 ♀, same data as holotype (MFNB, MZMB). 3 ♂, 3 ♀, Sulawesi Tengah, 6 km E Tambarana (01°11’15’’S, 120°28’06’’E), 50 m, 9. – 11.7.1999, leg. Bolm (MFNB, SMNS).
Description: Habitus and coloring as in Fig. 24. Head subquadrate. Penultimate antennal segment elongate. Mesotibia moderately thickened. Body surface dull with extremely fine, dense, setose punctation; microreticulation absent. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body including appendages light brown; disc of abdominal segments darker brown. Total body length 3.1 – 3.3 mm; forebody length 1.8 – 1.9 mm.
Male: Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII in almost posterior half with deep, triangular emargination with somewhat concave margins and acute anterior end; disc of sternite VIII with large depression wider than half sternite width stretching from emarginate posterior margin close to subbasal ridge; depression of sternite VIII surrounded laterally by long, medioposteriorly pointing macrosetae neighboured by sublateral, lateroposteriorly pointing macrosetae situated within depression (Fig. 145). Aedeagus (Figs 49, 116 – 118) about 0.5 mm long; phallobase stout, strongly sclerotized, thus dark reddish brown, shorter than distal lobes; distal portion of aedeagus seemingly attached to phallobase ventromedially owing to remarkable, dorsodistal extension of phallobase evenly narrowed towards wide, membranous, dorsodistal window (Figs 49, 116); apical lobes in lateral view very narrow basally but convexly widened distad (Figs 49, 116), in ventral and dorsal view with pronounced proximal waist, widened distolaterally and extended in remarkable, wing-like lobes with triangularly enlarged, mediodistad curved ends (Figs 117, 118); apical lobes ventrally separated in lateral halves with straight, parallel inner margins; ends of apical lobes membranous, in lateral view narrow and curved dorsad (Figs 49, 116), in ventral and dorsal view widened laterally, subflabellate apically (Figs 117, 118); dorsal lobe curved apicoventrad (Figs 49, 116), with membranous apex triangularly projecting between membranous ends of apical lobes (Figs 117, 118); flagellum inconspicuous (Fig. 117); small convex lobe, presumably ventral lobe, projecting from distal portion of aedeagus ventroproximally; median foramen oblong, with ends of lateral ridges curved proximolaterad; transverse ridge slender, curved distad (Fig. 117).
Female: Sperm pump with long, slender process segment; chamber without apophysis; bursal duct long, spirally tortuous (Fig. 178); bursa membranous.
Distribution: Scopaeus sulawesianus is known from Sulawesi only.
Etymology: With the epithet sulawesianus (adjective, Latin, composed of the geographic name Sulawesi and the suffix – anus, which indicates the geographical affiliation), reference is made to the distribution of this presumed Sulawesian endemic.
Comment: Scopaeus bipectenatus, S. bipenicillatus, S. cuspidatus and S. sulawesianus share the dark, strongly sclerotized aedeagus with small phallobase, which has a dorsodistal, membranous window of unknown function (Figs 107 – 118), and the missing apophysis of the chamber segment of the sperm pump (Figs 173 – 175, 178). They constitute a distinctive, speciose phylogenetic lineage of Scopaeus in the Oriental, Australasian, Nearctic and Neotropical regions with a large number of undescribed species.
description: Published as part of Frisch, Johannes & Narakusumo, Raden Pramesa, 2022, Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species, pp. 23-73 in Soil Organisms 95 (1) on pages 59-60, DOI: 10.25674/so95iss1id311, http://zenodo.org/record/10724236