Calobata guttata Walker, n. sp.
Creators: name: Ferro, Gustavo Borges
name: Marshall, Stephen A.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy

description: 1 Wing dark with three round hyaline spots on apical third (Fig. 40); discal band sometimes dark with up to four hyaline spots (Fig. 84)............................................................................................ 2
- Wing apex either entirely dark (Fig. 86) or clear (Fig. 103) or clear with a narrow preapical pigmentation (Fig. 62) or three preapical black spots (Figs. 35, 43, 74); discal band entirely dark................................................ 3
2 Paracephalon swollen and conical (Fig. 40). Vertex flattened. Wing with discal band black, without hyaline spots............................ C. guttata (Walker). Brazil, Colombia, French Guiana, Guiana, Surinam, Panama, Trinidad, Venezuela
- Paracephalon without swelling (Fig. 82). Vertex rounded. Wing with discal band dissected by four hyaline rounded spots................................................................................. C. secunda Hennig. Ecuador
3 Wing apex entirely clear (Fig. 103)..................................................... C. vitrata n. sp. Ecuador
- Wing apex not as above, either entirely dark (Fig. 86) or clear with a narrow preapical pigmentation (Fig. 62) or three preapical black spots (Figs. 35, 43, 74)............................................................................ 4
4 Clypeus long, length more than half of width in frontal view (Fig. 90). Fore tarsomeres 1–3 white (tarsomere one with brown pubescence basally), 4–5 covered with brown pubescence (Figs. 28, 92).......................................... 5
- Clypeus short, length less than half of width in frontal view. Fore tarsus not as above (either tarsomere one dark brown with gold/whitish ventral pubescence basally (Fig. 9) OR fore tarsomere one dark brown and 2–5 white (Fig. 30) OR fore tarsomere one white and 2–5 dark brown (Fig. 24)).................................................................. 6
5 Thorax brown. Paired spermathecal duct splitting into two convoluted stems marked by several long finger-like process (Fig. 91).............................................................................. C. spinosa n. sp. Brazil
- Thorax black. Paired spermathecal duct splitting into two convoluted and broadened stems, without finger-like process (Fig. 29)......................................................................... C. camptomera (Bigot). Brazil
6 Three frontal setae (Fig. 4). Fore tarsomere one dark brown, tarsomeres 2–5 white.................................. 7
- Two frontal setae (Fig. 14). Fore tarsomeres 2–5 dark brown................................................... 9
7 Scutellum upturned and peaked, slightly elongated at apex (Fig. 4)............ C. acuminata n. sp. Colombia, Costa Rica
- Scutellum upturned and broadly triangular (Figs. 30, 86)..................................................... 8
8 Paracephalon swollen, swelling rounded (Fig. 30). Abdominal pleural membrane dark brown, sometimes with rusty red regions. Yellowish species...................... C. cinnameus (Hennig). Bolivia, Brazil, Colombia, French Guiana, Peru
- Paracephalon smooth, without swelling. Abdominal pleural membrane black. Blackish species.............................................................................. C. setosa (Hennig). Bolivia, Brazil, Colombia, Ecuador, Peru
9 Mid and hind tibiae flattened, sulcus on outer face strongly pronounced and curved (Figs. 18, 48, 55, 97). Blackish species...................................................................................................... 10
- Mid and hind tibiae slightly flattened, sulcus on outer face weak. Colour variable.................................. 13
10 Inner vertical setae insertions slightly protuberant (Fig. 16, 46). Wing apex clear with a dark and narrow preapical pigmentation................................................................................................ 11
- Inner vertical setae insertions flat. Wing apex darkened....................................................... 12
11 Paracephalon swollen (Fig. 16). Paired spermathecae flat, disc-shaped...... C. aspera n. sp. Bolivia, Ecuador, Guiana, Peru
- Paracephalon without swelling (Fig. 46). Paired spermathecae spherical................................................................................................ C. lobulosa n. sp. Bolivia, Brazil, Colombia, Ecuador, Peru
12 Epicephalon black (Fig. 55). Ejaculatory apodeme large, blade wider than sperm pump.......................................... C. nigra Schiner. Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Honduras, Mexico, Peru, Trinidad, Venezuela
- Epicephalon velvet brown (Fig. 97). Ejaculatory apodeme small, sperm pump wider than blade..... C. velutinosa n. sp. Peru
13 Mid and hind tibia bicoloured, basal half dark brown and apical half yellowish (Fig. 59). Crossvein r-m clear, without dark spot........................................................................ C. peruana (Hennig). Bolivia, Peru
- Mid and hind tibia uniformly black, brown or yellowish. Crossvein r-m with small dark spot......................... 14
14 Paracephalon swollen (Figs. 10, 24, 33, 72) or slightly swollen (Fig. 38), extending conically beyond the vertex........ 15
- Paracephalon without swelling, not extending conically beyond the vertex...................................... 19
15 Limits between paracephalon, epicephalon and vertex not distinguishable (Fig. 38). Ocellar triangle displaced anteriorly (closer to anterior frons margin), vertex occupying more than half the length of the head.... C. erugata n. sp. Ecuador, Peru
- Limits between paracephalon, epicephalon and vertex clearly distinguishable. Ocellar triangle centralized, vertex occupying no more than half the length of the head................................................................... 16
16 Inner vertical setae inserted in strong tubercles (Fig. 33). A1+CuA2 less than half as long as CuA2............................................................................... C. conifera Hendel. Brazil, Colombia, Guiana, Paraguay
- Inner vertical seta insertion flat, without tubercle. A1+CuA2 as long as CuA2..................................... 17
17 Female cervical sclerite with a small nipple-like swelling in posterior half. Wing apex clear with three preapical black spots (Fig. 72). Black species...................................... C. podagrica Rondani. Bolivia, Brazil, Ecuador, Peru
- Female cervical sclerite smooth, without nipple-like swelling in posterior half. Wing apex clear with a dark and narrow preapical pigmentation. Yellowish to brown species, sometimes slightly rusty red....................................... 18
18 Swelling on paracephalon pointing dorsally (Fig. 9). Fore tarsomere one dark brown with gold/whitish ventral pubescence basally. Yellowish species............................................................ C. angularis n. sp. Peru
- Swelling on paracephalon posteriorly pointed (Figs. 24, 26). Fore tarsomere one white, fore tarsomeres 2–5 dark brown. Brown, sometimes slightly rusty red species................... C. bulla n. sp. Bolivia, Brazil, Colombia, Ecuador, Peru
19 Vertex flattened, head depressed posteriorly (Figs. 7, 68, 74, 93)................................................ 20
- Vertex rounded, head globose (Figs. 14, 43, 50, 53, 64, 78, 80, 99)............................................. 24
20 Mid femur white in basal third. Black/dark brown species.................................................... 21
- Mid femur uniformly brown in basal third. Brown/yellowish species........................................... 22
21 Fore tarsomere one white, fore tarsomeres 2–5 dark brown. Wing apex clear with three preapical black spots. Phallic bulb subequal in length and width, almost spherical (Fig. 96).......... C. umbra n. sp. French Guiana, Tobago, Trinidad, Venezuela
- Fore tarsomere one dark brown with gold/whitish ventral pubescence basally. Wing apex clear with a dark and narrow preapical pigmentation. Phallic bulb cylindrical, about three times as long as wide (Fig. 23).......................................................... C. brevipennis (Walker). Bolivia, Brazil, Colombia, Ecuador, French Guiana, Peru, Venezuela
22 Epicephalon smooth, without wrinkles posterior to ocellar triangle (Fig. 74). Fore tarsomere one white, fore tarsomeres 2–5 dark brown. Wing apex clear with three preapical black spots....................... C. punctipennis (Macquart). Brazil
- Epicephalon wrinkled posterior to ocellar triangle (Figs. 7, 68). Fore tarsomere one dark brown with gold/whitish ventral pubescence basally. Wing apex clear with a dark and narrow preapical pigmentation............................... 23
23 Yellowish species. Paracephalon and epicephalon yellow (Fig. 68). Paired spermathecae cylindrical and thin (Fig. 67). Phallic bulb cylindrical, about three times as long as wide (Fig. 69).................................................................................................. C. planivertex n. sp. Brazil, Colombia, Ecuador, French Guiana, Peru
- Brownish species, head and abdomen usually rusty red. Paracephalon and epicephalon darkened (Figs. 5, 7). Paired spermathecae sac-like, tapered basally and apically swollen (Fig. 6). Phallic bulb subequal in length and width, almost spherical (Fig. 8)............................................................................. C. aeruginosa n. sp. Ecuador
24 Inner vertical setae insertions with strong tubercles (Fig. 50)................ C. modesta (Cresson). Colombia, Guatemala
- Inner vertical setae insertions flat, without tubercles........................................................ 25
25 Fore tarsomere one white, fore tarsomeres 2–5 dark brown................................................... 26
- Fore tarsomere one dark brown with gold/whitish ventral pubescence basally..................................... 27
26 Vertex brownish anterior to postocellar setae. Mid femur uniformly brown in basal third. Thorax brown....................................................................................... C. rotunda n. sp. Colombia, Costa Rica
- Vertex with a rounded black spot anterior to postocellar setae (Fig. 53). Mid femur white in basal third. Thorax dark brown with blue metallic shine...................................................... C. nicaraguensis n. sp. Nicaragua
27 Female cervical sclerite slightly swollen posteriorly. Mid and hind femur white basally, apical third with an angled whitish yellow ring (Figs. 62, 64).......................................... C. picina (Cresson). Costa Rica, Ecuador, Panama
- Female cervical sclerite flat, without posterior swelling. Mid and hind femur with basal and apical third brown or yellow. 28
28 R4+5 and M1+2 ending separately in wing margin. Paired spermathecae cylindrical and strongly curved in middle (Fig. 100)............................................................................. C. venezuelensis n. sp. Venezuela
- R4+5 and M1+2 meeting at wing margin. Paired spermathecae spherical or sac-like (tapered basally and distally swollen).... 29
29 Epicephalon smooth and shiny, not depressed anterior to postocellar setae (Fig. 14). Yellowish species, sometimes with abdomen darkened at least on T4–T6.......................................................................................... C. arthritica (Wiedemann). Belize, Colombia, Costa Rica, Ecuador, Honduras, Mexico, Panama, Venezuela
- Epicephalon somewhat wrinkled and depressed before postocellar setae (Figs. 43, 80). Brownish/ferruginous species with darkened regions on mesonotum and katepisternum........................................................ 30
30 Wing apex clear with three preapical black spots. Paired spermathecal duct stems with a preapical rounded swelling (Fig. 44).................................... C. harenosa (Cresson). Colombia, Costa Rica, Ecuador, Guiana, Mexico, Panama
- Wing apex clear with a dark and narrow preapical band. Paired spermathecal duct stems very narrow in basal third, then expanded and convoluted (Fig. 81)............................................... C. schildi (Cresson). Costa Rica
description: Published as part of Ferro, Gustavo Borges & Marshall, Stephen A., 2018, A revision of the Neotropical ant-like genus Cardiacephala Macquart, including Plocoscelus Enderlein syn. nov. (Diptera: Micropezidae, Taeniapterinae), pp. 401-458 in Zootaxa 4429 (3) on pages 407-409, DOI: 10.11646/zootaxa.4429.3.1, http://zenodo.org/record/1285294