Gomphostemma nutans Hook. f.
Creators: name: Bongcheewin, Bhanubong
name: Ingrouille, Martin J.
name: Paton, Alan J.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Magnoliopsida
Lamiales
Lamiaceae
Gomphostemma
Gomphostemma nutans

description: <b> 28. Gomphostemma nutans <i>Hook.f.</i></b> (Hooker 1883: 695); Prain (1891: 228); Mukercee (1920: 206). Type: India, Khasia, Oct. 1829, <i>Simons</i> s.n. (K [Herb. Hookerianum, K000826323], lectotype selected here). Climbing herb. <i>Stems</i> slender, scrambling, obtusely quadrangular with longitudinal grooves, with stellate hairs and sometimes dendroid hairs. <i>Leaoes</i> petiolate or subsessile, chartaceous; blades ovate or elliptic-ovate, 2 – 12 × 2 – 6 cm, apex acute, margin serrate or obtusely serrate, base attenuate, cuneate or obtuse, upper side with stellate hairs and sometimes simple hairs, lower side with dense stellate hairs; petiole 3 – 20 mm long. <i>Inflorescence</i> terminal or terminal and axillary, 20 – 110 mm long, congested; verticils few-flowered; subtending bracts present, spathulate, obovate, lanceolate or ellipticlanceolate, 10 – 20 × 2 – 10 mm, apex acute, margin entire or serrate, base attenuate, with dense stellate hairs on both sides; bracteoles linear, 3 – 10 mm long. <i>Flowering calyx</i> infundibular, 10 – 13 mm long, 10-ridged, outside with dense stellate hairs, inside glabrous below and densely tomentose above; tube 3 – 5 mm long; lobes spreading, lanceolate-ovate, 5 – 8 mm long, apex acute or slightly acuminate. <i>Fruiting calyx</i> infundibular, 11 – 13 mm long; tube 2 – 5 mm long; lobes lanceolate-ovate, 5 – 8 mm long, apex acute, margin sometimes slightly revolute. <i>Corolla</i> yellow, 20 – 33 mm long, throat inflated, abruptly dilated near the throat; tube slender, incurved, 13 – 30 mm long, outside tomentose, inside with an annulus of simple hairs; posterior lip ovate, 3 – 6 mm long; anterior lip 3-lobed, spreading, 8 – 10 mm long, margin undulate. <i>Staminal filaments</i> fleshy, tomentose. <i>Style</i> slightly fleshy, glabrous, shorter than corolla with apex subequally lobed. <i>Nutlets</i> 1 – 2, obovate or ovate, 3 – 2 mm long, wrinkled. <b>DISTRIBUTION.</b> India (Assam, Nagaland, Meghalaya, Manipur), Burma. Map 5. <b> SPECIMENS EXAMINED. INDIA. Assam. Bamanigaon [Bamungaon, 26.9165°, 92.1665°], Sal Forest, near sea level, 3?ec. 1929, <i>Koelz</i> 22266 (E);</b> Nagaland. Naga Hills, Narum, 1300 m [3000 ft],?ec. 1905, <i>Meebold</i> 5203 (K); Meghalaya. Khasia, Oct. 1829, <i>Simons</i> s.n. (lectotype K [Herb. Hookerianum)]; ibid., <i>Lobb</i> s.n. (K [Herb. Hookerianum]); Khasia, Nunklow, 1030 m [3300 ft], 2 Nov. 1851, <i>Clarke</i> 16302 (lectoparatype K [Herb. Hookerianum]); Khasia, Nya Bungalhi, 530 m [2300 ft], 3 Oct. 1883, <i>Clarke</i> 20623A (K); Manipur. Mairing [22.53°, 92.3665°], Naga Hills, 1300 m [3000 ft],?ec. 1905, <i>Meebold</i> 5185 (E); Kanglatongbi [Kanglatongi, 22.9833°, 93.9°], 900 m [3000 ft], 3 Oct. 1923, <i>3ullock</i> 633 (K); On the way to Nungba [22.53°, 93.2333°], 900 m [3000 ft], 2 Jan. 1882, <i>Watt</i> 6662 (K). <b>MYANMAR.</b> Sagaing. Tamu, Upper Chindwin [22.6°, 93.1°],?ec. 1905, <i>Meebold</i> 5398 (E, K); near Kendat [23.5333°, 92.2333°], Sept. 1890, <i>Prazer</i> 352 (K); Suigon, Katha [22.1833°, 96.33°], 20 Nov. 1908, <i>Lace</i> 2211 (E 2 sheets, K). <b>HABITAT.</b> In hill evergreen forest, 530 – 1300 m. <b>CONSERVATION STATUS.</b> <i>Gomphostemma nutans</i> has AOO of 20 km 2. Google Earth imagery shows the evidence of habitat loss due to agricultural activity but the habitats in India remains intact. Hence, the Least Concern (LC) category is applied. <b>PHENOLOGY</b>. Flowering and fruiting September – January. <b>NOTES.</b> <i>Gomphostemma nutans</i> is easy to recognise by its climbing habit with slender stem. The specific epithet ‘ <i>nutans</i> ’ which means nodding may derive from the specimen mounted in an artificial position, rendering the inflorescence in a nodding state. It is a similar case for the species name of <i>Clinacanthus nutans</i> (Burm.f.) Lindau (Bongcheewin <i>et al.</i> 2019). The inflorescence may be upright in nature.
description: Published as part of <i>Bongcheewin, Bhanubong, Ingrouille, Martin J. &amp; Paton, Alan J., 2022, A revision of Gomphostemma (Lamiaceae), pp. 27-92 in Kew Bulletin 77 (1)</i> on page 75, DOI: 10.1007/s12225-021-09991-y, http://zenodo.org/record/7615581
description: {"references": ["Hooker, J.?. (1883). Labiatae, Flora of 3 ritish India 2: 602 - 503. Reeve &amp; Co., London.", "____ (1891). An account of the genus Gomphostemma Wall. Ann. Roy. 3 ot. Gard., Calcutta 3: 225 - 251.", "____ (1920). A revision of the Labiatae of the Indian Empire. Rec. 3 ot. Suro. India 12: 1 - 228."]}