Nebria (Sadonebria) hakusana Sasakawa 2022, sp.n.
Creators: name: Sasakawa, K.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Nebria
Nebria hakusana

description: Nebria (Sadonebria) hakusana Sasakawa, sp.n.
Figs 2–7.
TYPE MATERIALS: Holotype, ♂ (KS), Shinmata Pass, Nomuki, Katsuyama, Fukui Prefecture, Japan, 1.vi.2020, S. Inoue leg.; paratypes, 1♂ (KS), Mt. Sanpôkuzure, ca 1,200 m, Shirakawa City, Gifu Prefecture, Japan, 23–24.vi.2005, H.Ikeda leg.; 1♂ 2♀♀ (EUM), “KAGA-Hakusan Nakahanba ~ Bettonozoki (1,500~ 1,750 m) 1 VIII. 1973 M. Miyatake ” [Nakahanba (1,500 m) ~ Bettonozoki (1,500 m), Mt. Hakusan, Shiramine, Hakusan, Ishikawa Prefecture, Japan].
DESCRIPTION. External characters (Figs 2–3). Body length (including mandibles): ♂, 12.5–13.6 mm; ♀, 12.6– 12.8 mm. PL / PW: 0.73–0.76; PAW/ PW: 0.73–0.76; PPW/ PW: 0.64–0.68. EL/EW: 1.59–1.71. Habitus slender. Hind wings atrophied. Dorsal surface shiny and almost black; head with a dark-brown patch between eyes; pronotal and elytral margins dark-brown; mouth appendages and antennae reddish to dark brown; legs dark brown except for femora, which are blackish brown. Pronotum cordate and convex; lateral margins reflexed throughout; hind angle acute; laterobasal impression large and deep; median line distinct in the middle, absent near the anterior margin, rudimentary near the posterior margin; surface of central part and near the anterior margin almost smooth; surface near the posterior margins very sparsely punctate; surface of the lateral margins very sparsely punctate and/or shallowly, transversely wrinkled; two marginal setae on each lateral side, anterior setae near the widest pronotal point, and posterior setae near hind angles. Elytra oblong, widest behind the middle; 7–10 setigerous pores on interval 3. Abdominal sterna 4–7 usually with a pair of setae, but in the male from Mt. Sanpôkuzure, additional setae on left side of sternum 4 and on both sides of sternum 7.
Male genitalia (Figs 4–6). Aedeagus slender and strongly bent at apical 2/5; ventral contour of subapical part from lateral view arcuate ventrally; apex simple and widely rounded.Endophallus straight, elongate, directed posterodorsally; laterobasal lobes large, hemispherical; lateroapical lobes absent; dorsobasal lobe bifid from lateral view, with the subapex of basal one smaller than that of apical one; dorsoapical lobe widely swollen, and smaller than the laterobasal and dorsobasal lobes; ventroapical lobe barely swollen. Both right and left parameres spatulate, with the former larger than the latter.
DIAGNOSIS. Distinguished from N. quinquelobata Sasakawa, 2016, N. kiso, and N. mikawa, which are relatively closely distributed (Fig. 1), by the following characters: from N. quinquelobata by dark brown legs [versus yellowish brown legs in N. quinquelobata; Sasakawa, 2016], and from N. kiso and N. mikawa by less cordate and less punctate pronotum (Figs 7–9).
ETYMOLOGY. Named after the Hakusan Mountains, where the type materials were collected.
description: Published as part of Sasakawa, K., 2022, A new species of the East Asian endemic subgenus Sadonebria Ledoux et Roux (Coleoptera: Carabidae: Nebria) ground beetles from Japan, pp. 10-14 in Russian Entomological Journal 31 (1) on pages 11-13, DOI: 10.15298/rusentj.31.1.02, http://zenodo.org/record/10920153