Groehnizomus oculiferans DE FRANCESCO MAGNUSSEN & S.P. MÜLLER 2022, SP. NOV.
Creators: name: Magnussen, Ilian De Francesco
name: Müller, Sandro P.
name: Hammel, Jörg U.
name: Kotthoff, Ulrich
name: Harms, Danilo
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Schizomida
Protoschizomidae
Groehnizomus
Groehnizomus oculiferans

description: † GROEHNIZOMUS OCULIFERANS DE FRANCESCO
MAGNUSSEN & S.P. MÜLLER SP. NOV.
(FIGS 12–19)
Zoobank registration: urn:lsid:zoobank.org:act: 4291F40C-1BA1-4A9C-A668-0E1AA2E65B38
Type material: Holotype (Figs 12A–D, 13A–D), adult homeomorphic ♂, collection no. GPIH05049, ex collection Carsten Gröhn (no. 11266); paratype (Fig. 17A–D), adult heteromorphic ♂, collection no. GPIH05055, ex collection Carsten Gröhn (no. 11210).
Diagnosis
† Groehnizomus oculiferans can be characterized by the following features: slender pedipalps; flagellum shape flat and subrhomboidal with a rounded tip.
Etymology
The specific name is derived from Latin oculus, eye and feree, to bear, referring to the presence of eyes in this specimen, which is a rare occurence in Recent schizomids. It is masculine in gender.
Description
Total length from the anterior process to base of the flagellum 2.56 mm.
Colour (in amber): Legs I–III, end of the propeltidium and metapeltidium yellowish brown, nearly transparent; anterior propeltidium, pedipalps, opisthosoma, legs IV and flagellum brownish red, original colouring equivocal.
Prosoma: Length 1.13 mm; propeltidium length 0.91 mm, width 0.35 mm; anterior process broad at the base with apical tip, length 0.08 mm; setation: one seta near the tip (visible only from directly above) followed by two setae, one next to the other. Propeltidium setation: only one more single median seta on the propeltidium between the anterior process and the eyes (Fig. 14A, B). No other setae observed owing to taphonomic processes. Corneate eyes present, clearly elevated above the surface of the propeltidium. Mesopeltidia width 0.15 mm. Metapeltidium divided (Fig. 14C, D), length 0.24 mm, width (from outer margin of left plate to outer margin of right plate) 0.52 mm. Anterior sternum with 10 + 2 sternapophysial setae, length 0.36 mm, width 0.24 mm. Posterior sternum with six setae, length 0.14 mm, width 0.15 mm.
Pedipalps: Without armature, except for mesal spur on trochater; trochanter produced distally (Fig. 15A–F), setation: six large and strong ventral setae, four ventroectal, two mesal; femur: marginal longer than trochanter, setation: nine dorsal, a single seta, eight in a row, three ventroectal, four ventromesal; patella: nearly as long as the femur, setation: four dorsal, three mesal, two dorsoectal, four ectal, two ventroectal; tibia: shorter than patella, setation: five ventroectal, five ventromesal, four dorsal, five dorsoectal, three dorsomesal; tarsus: with asymmetrical tarsal spurs, spur length: 0.02 mm, setation: + three dorsal, three ventromesal, four dorsomesal, four dorsoectal, three ventroectal, two mesal; claw: bent downwards, about one-third of the length of tarsus.
Chelicerae: Setation of the mesal side of chelicerae not visible, except for G6-1.
Legs: Left leg I aborted at the trochanter, right one well preserved and complete, tarsus with seven segments, leg I ~1.54 times as long as the body; anterodorsal margin of femur of leg IV produced at an angle of ~90°, femur of leg IV is 3.4 times longer than deep, leg IV 1.18 times longer than the body, leg formula: 1423.
Opisthosoma: Length 1.43 mm, width 0.48 mm; tergites X–XII not elongated, tergite XII with a tiny posterodorsal process. Abdominal tergites setation: I: Dm, right one missing; II–VII: Dm; VIII: Dm, Dl1; IX: Dl1, Dl2; X: not visible; XI: Dl1; XII: Dm, Dl1, Dl2;
setation pattern: 2:2:2:2:2:2:2:4:4:?:2:6. Flagellum: dorsoventrally flattened, subrhomboidal in shape but with a rounded tip (holotype Fig. 16A–G; paratype Fig. 19A–D), length 0.26 mm, width 0.26 mm; stalk length 0.08 mm; setation: Dm1, Dl2 paired, Dm4, Dl3 paired, Vm1, Vm2 paired, Vm3 paired, Vl1 paired, Vm5, Vl2 paired.
Holotype further measurements: Pedipalp: trochanter 0.34mm; femur 0.36 mm; patella 0.37 mm; tibia 0.29 mm; tarsus 0.22 mm; claw 0.07 mm; total length 1.65 mm. Leg I: trochanter 0.18 mm; femur 1.01 mm; patella 1.24 mm; tibia 0.80 mm; metatarsus 0.31 mm; tarsus (tarsus segments) 0.41 mm (0.04 mm; 0.03 mm; 0.04 mm; 0.05 mm; 0.06 mm; 0.06 mm; 0.13 mm); total length 3.95 mm. Leg II: trochanter 0.11 mm; femur 0.69 mm; patella 0.31 mm; tibia 0.39 mm; metatarsus 0.40 mm; tarsus (tarsus segments) 0.30 mm (0.14 mm; 0.07 mm; 0.09 mm); claw 0.05 mm; total length 2.25 mm. Leg III: trochanter 0.15 mm; femur 0.58 mm; patella 0.22 mm; tibia 0.35 mm; metatarsus 0.38 mm; tarsus (tarsus segments) 0.30 mm (0.14 mm; 0.07 mm; 0.09 mm); claw 0.05 mm; total length 2.03 mm. Leg IV: trochanter 0.18 mm; femur 1.06 mm; patella 0.27 mm; tibia 0.61 mm; metatarsus 0.57 mm; tarsus (tarsus segments) 0.39 mm (0.16 mm; 0.08 mm; 0.15 mm); claw 0.07 mm; total length 3.01 mm.
Paratype further measurements: Total length from the anterior process to base of the flagellum: 2.56 mm. Prosoma length 1.18 mm; propeltidium length 0.77mm, width 0.35 mm. Pedipalp: trochanter 0.42 mm; femur 0.69 mm; patella 0.85 mm; tibia 0.41 mm; tarsus 0.25 mm; total length 2.62 mm. Leg I: trochanter 0.25 mm; femur> 1.27 mm; patella missing; tibia missing; metatarsus missing; tarsus missing; total length> 1.52 mm. Leg II: trochanter 0.13 mm; femur 0.65 mm; patella 0.38 mm; tibia 0.45 mm; metatarsus 0.31 mm; tarsus 0.34 mm; total length 2.26 mm. Leg III: trochanter 0.15 mm; femur 0.66 mm; patella 0.29 mm; tibia 0.37 mm; metatarsus 0.51 mm; tarsus 0.27 mm; total length 2.25 mm. Leg IV: trochanter 0.25 mm; femur 1.09 mm; patella 0.41 mm; tibia 0.68 mm; metatarsus 0.61 mm; tarsus 0.42 mm; total length 3.46 mm. Leg IV is 1.35 times longer than the body. Opistosoma: length 1.38 mm, width 0.51 mm. Flagellum: length 0.29 mm, width 0.23 mm; stalk length 0.08 mm.
Variations: Paratype with heteromorphic pedipalps, nearly as long as the body (Fig. 18A, B), 0.62 times the body length in the holotype; femur of leg IV is 2.75
818 I. DE FRANCESCO MAGNUSSEN ET AL.
times longer than deep, 3.4 times longer than deep in the holotype.
description: Published as part of Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich & Harms, Danilo, 2022, Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane, pp. 792-844 in Zoological Journal of the Linnean Society 196 on pages 810-818, DOI: 10.1093/zoolinnean/zlac034, http://zenodo.org/record/7187082