Calciphilopteris ludens Yesilyurt & H. Schneider, comb. nov.
Creators: name: Yesilyurt, Jovita C.
name: Schneider, Harald
Publisher: name: Zenodo
Year: 2010
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Polypodiopsida
Polypodiales
Pteridaceae
Calciphilopteris
Calciphilopteris ludens

description: 2. Calciphilopteris ludens (Wall. ex Hook.) Yesilyurt & H.Schneider, comb. nov. — Fig. 1A.
Basionym:— Pteris ludens Wallich (1828: 88), nomen nudum, Pteris ludens Wall. ex. Hooker (1858: 210), Lithobrochia ludens (Wall. ex Hook.) Beddome (1865: t. 27), Pellaea ludens (Wall. ex Hook.) Prantl (1882: 419), Doryopteris ludens (Wall. ex Hook.) Smith (1875: 289). Holotype: INDIA: Wallich 88 (K).
Rhizome predominantly long-creeping, scales up to 3 mm, dark brown, lanceolate, margin entire. Petiole terete to sulcate adaxially towards the base of the lamina, dark brown to atropurpureous, indumentum usually with hairs (whitish), non-clathrate scales and fibrils. Sterile and fertile fronds glabrescent to glabrous, indumentum on the abaxial side with fibrils and non-clathrate microscales. Sterile fronds up to 35 cm; lamina up to 20 cm, sagittate to pedate; ovate to pentagonal; pinnae/segments1–2(–3) pairs; hydathodes generally present (sometimes obscure). Fertile fronds up to 45 cm; lamina up to 25 cm, pentagonal to broad-cordate, pedate, pedate-pinnatifid (to pedate-2-pinnatifid); pinnae/segments in 2–4(–5) pairs; basal pinna/segment falcate, with 1(–2) acroscopic lobes and 2–3 basiscopic segments/lobes, which are ascendant, falcate to lanceolate, rarely the first basiscopic segment/lobe is divided; other pinnae/segments al equally ascendant, lanceolate to long-lanceolate. Indusium with margin entire to slightly eroded. Sporangium with pedicel 1–2(– 2½) × the capsule length; capsule with annulus with 17–34 indurated cells. Spores dark castaneous, cristate.
Distribution:— India, Indochina, southern China, western parts of Malesia (Flores, Java, Malay Peninsula, Simbawa, Sumatra), Thailand, and Australia (where it was once recorded from north-eastern Queensland by D.L. Jones s.n. (BRI), see: Bostock 1998).
Note:—some specimens (Congdon 617, Molesworth-Allen 2785, E. Smith 250A, Kerr 250, 11365) show unique leaf and spore characters that are similar but not identical with characters observed in other species of Calciphilopteris. It was decided to assign these specimens here to C. ludens, because available collections are insufficient to make unequivocal taxonomic assignments. These specimens may represent either a unique form of C. ludens or an undiscovered species.
Selected specimens:— MALAYSIA: Perlis: Molesworth-Allen 2816 (US, S). Kedah: Alphonso & Samsuri 123 (BM); Henderson 29066 (BM); Littke 260 (L). INDONESIA: Java: Horsfield s.n. 110 (BM); Mosset 188 (C, GH, L, PE). THAILAND: Geesink et al. 5949 (BM, L); Hennipman 3029 (BM, L, NY, U, US); Kerr 5476 (BM); Kostermanns 762 (L); Larsen 5790 (BM, GH); Rock 503 (US); Shimizu et al. 13325 (C, L); Tagawa et al. 1903 (L, US);. MYANMAR: Dickason 7762 (GH); Kress et al. 97-6017 (US); J. F. Smith 89 (GH). VIETNAM (Cochinchina): Larsen 8964 (GH); Pierre s.n. (GH). SINGAPORE: Corner & Henderson 22815 (C). CHINA: Yunan: Henry 13325 (MO, NY). INDIA: Assam: Mann s.n. (UC).
description: Published as part of Yesilyurt, Jovita C. & Schneider, Harald, 2010, The new fern genus Calciphilopteris (Pteridaceae), pp. 52-59 in Phytotaxa 7 on page 56, DOI: 10.11646/phytotaxa.7.1.7, http://zenodo.org/record/4893512