Syleter sinepunctatus subsp. sinepunctatus sinepunctatus 2021, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Syleter
Syleter sinepunctatus
Syleter sinepunctatus sinepunctatus

description: Syleter sinepunctatus sinepunctatus sp. nov.
Figs 11, 18, 25
Holotype: BMNH; ♂, with labels and data: white, handwritten in black ink “32 BRUNEI Bukit Sulang Nn hamunin, 28.VIII.-5.IX.82, B.E. Stork B.M. 1982-388”, printed “at light”.
Paratypes: BMNH; 1 ♀, with labels and data: orange, handwritten in black ink “ Kuching SARAWAK 28.7.1961 ”, “ at light ”, white, black printed “R.O.S.Clarke. B.M. 1970-557.”, green, handwritten with pencil “ Syleter n.sp. nr. paradoxus PUTZ. det. Balkenohl, 2000”. – MFNB; 1 ♀, with labels and data: beige, black printed “S.O. Borneo Grabowsky.”, “Zool. Mus. Berlin”. – ADCW; 3 ♂♂, 2 ♀♀, 7 specimens., with labels and data: white, black printed “ INDONESIEN: Kalimantan Timur, Tanjung Isuy 20.1.1995 leg. Seyfert & Greindl (34)”. – ADCW; 1 ♂, 3 ♀♀, 5 specimens, same data but “(33)” and “(34)” and orange, handwritten: “Coll. Zettel Wien ”.
Diagnosis: A medium sized species with seven-setose labrum, and the clypeus and frons isodiametrically reticulated. Differentiated from the similar species S. paradoxus and S. gradus sp. nov. by the missing punctures on the disk of the pronotum and the missing clypeo-frontal furrow. S. paradoxus differs in addition by the smooth clypeus. S. gradus sp. nov. differs in addition by the complete and step-like developed neck constriction, and by the larger size.
Etymology: The name refers to the completely missing punctures on the disc of the pronotum (Latin: sine – without).
Description
Measurements: Body length 3.37-3.8 mm (x = 3.64 mm *), width 1.16-1.27 mm (x = 1.22 mm *), ratio length/width of pronotum 1.01-1.05 (x = 1.03*), ratio length/width of elytra 1.58-1.66 (x = 1.62*); (*n = 10).
Colour: Piceous to fuscous. Legs fuscous. Antenna, tarsomeres and palpomeres testaceous.
Head: A third narrower than pronotum, frons and clypeus anterior eye-level conspicuously elongated. Clypeus straight anteriorly, with lateral angle obtuse rounded; completely fused with clypeal wing and supraantennal plate, all reflexed margined, clypeus convex, separated from elongated convex supraantennal plate by parallel running deep and conspicuously wide clypeal furrows, extending up to anterior eye-level; supraantennal plate extended up to mid-eye level; clypeus separated from frons by indistinct transverse nearly not traceable change of the direction of the convexity. Frons moderately convex, with wide frontal furrows, with distinct lateral carina separating frontal furrow from supraorbital furrow, frontal furrow and lateral carina wide, slightly diverging and extending nearly up to neck constriction. Clypeal, frontal, and supraorbital furrow not connected. Clypeus and frons with distinct isodiametric reticulation, clypeal wing and supraantennal plate with longitudinal reticulation. Neck at middle at same level as frons, constriction developed laterally as row of punctures. Eye moderately projecting laterally. Gena indistinct. Antenna reaching over posterior angle of pronotum, antennomeres four to ten elongate (around 1.8 times longer than wide). Labrum straight anteriorly, reticulated, seven setose.
Pronotum: Disk in lateral view moderately convex, convex in frontal view. Outline trapezoid. Reflexed lateral margin smooth, nearly straight attenuated in anterior half, widest at posterior third; anterior angle slightly projecting; posterior angle developed as distinct rounded tooth, tooth laterally much less projecting than widest part of lateral margin. Lateral channel as wide as reflexed lateral margin, anterior setigerous punctures removed from channel by diameter of pore, posterior setigerous puncture situated in the channel close to the tooth of the posterior angle. Median line moderately narrow; anterior transverse line slightly wider than median line, joining with median line. Surface with isodiametric to transverse reticulation; basal constriction distinct, flange 2.5 times wider than constriction.
Elytron: Disk slightly convex in anterior half in lateral view, convex on frontal view. Around 1.6 times as long as wide, slightly dilated in apical half, maximum width slightly behind middle. Reflexed lateral margin smooth. Scutellar striole short but visible. Base of elytron with setigerous tubercle at base of second interval, with a small tubercle at base of third interval. Striae moderately deep, crenulate-striate, four to seven reaching base, the seventh at level of the humerus, humerus marked by distinct tooth. Stria one running to apex, six and seven joining apically, the others partly joining at apex (variable). Intervals moderately convex. Third interval with three setigerous punctures, first distinctly removed from base and approaching third stria, the second situated at middle of the interval, the third approaching second stria. Interval one to seven smooth, glossy, interval eight with indistinct longitudinal reticulation.
Hind wings: Fully developed.
Lower surface: Proepisternum covered with isodiametric reticulation and few transverse wrinkles. Sternites of abdomen transversely sulcate, with transverse reticulation.
Legs: Profemur dorsally sulcate. Protibia covered with longitudinal reticulation, with two slender spines of moderate length, the third one obtuse, movable spur regularly arcuate.
Male genitalia (Fig. 18): Median lobe regularly curved in basal half, gently curved towards apex, apically distinctly acute in ventral view, in lateral view widened at middle, developed apically as asymmetric distorted spatula, broadly rounded at tip of apex; endophallus with two longitudinal groups of bristles; parameres asetose.
Female genitalia (Fig. 25): Coxostylus one and two partly fused; coxostylus two gently curved at apex, ventromedially conspicuously narrowed and with conspicuously wide and long nematiform seta, apically with moderately large cone-like seta, with twelve long and one short nematiform setae, coxostylus one with two long nematiform apical setae, ventromedially with four moderately long setae, with two shorter setae at middle; laterotergite with five moderately long nematiform setae, one of them situated basally.
Variability: The reticulation on interval eight is more or less distinct. At the apex of the elytron the striae two to five are more or less joining.
Distribution: Known from two locations in Borneo (Brunei and South East Borneo).
description: Published as part of Balkenohl, Michael, 2021, Revisiting the Oriental genus Syleter Andrewes, 1941 (Coleoptera: Carabidae: Clivinini), pp. 357-374 in Revue suisse de Zoologie 128 (2) on pages 365-367, DOI: 10.35929/RSZ.0050, http://zenodo.org/record/5640135