Lomanellidae Mendes &amp; Derkarabetian 2021, fam. nov.
Creators: name: Derkarabetian, Shahan
name: Baker, Caitlin M.
name: Hedin, Marshal
name: Prieto, Carlos E.
name: Giribet, Gonzalo
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Araneae
Lomanellidae

description: Family LOMANELLIDAE Mendes &amp; Derkarabetian, fam. nov. ZooBank LSID: urn:lsid:zoobank.org:act: B2825A44-1616-4103-91F1-A2BFF07BBC3F <i>Included genera</i>: <i>Lomanella</i> Pocock, 1903, <i>Abaddon</i> Derkarabetian &amp; Baker, gen. nov. <i>Type genus</i>: <i>Lomanella</i> Pocock, 1903. <i>Type species</i>: <i>Lomanella raniceps</i> Pocock, 1903, by original designation <i>Diagnosis</i> This Australian family (Fig. 5) is most closely related to the morphologically unique Synthetonychiidae from New Zealand, but differs from the family-level diagnosis of Forster (1954) by the presence of an obvious eye mound and shorter pedipalps bearing tubercles, which are sometimes more robust. Synthetonychiidae are diagnosed by their lack of an eye mound and elongate pedipalps that are slender and without tubercles (e.g. Forster 1954, fig. 708). Lomanellidae can be distinguished from Triaenonychidae by their relatively simple palps with weak spination (Fig. 6 <i>E</i>) (Hunt and Hickman 1993, fig. 1), a process on the dorsal surface of coxae II, and lateral projections on the carapace above coxae II associated with the ozopore. Additionally, compared to male triaenonychid genitalia, which typically possess the full complement of plates (dorsal, dorsolateral, and ventral), the male genitalia of lomanellids are extremely attenuated and show a complete loss of the dorsal plate (Fig. 7 <i>A</i>) (Hunt and Hickman 1993, fig. 3). <i>Distribution</i> Lomanellids are restricted to Australia (Fig. 5). The genus <i>Lomanella</i> is widespread in the temperate forests of southern New South Wales, Victoria and Tasmania, with a single species that is restricted to a small region of south-western Western Australia. The new genus <i>Abaddon</i>, gen. nov. is extremely restricted in distribution, known only from two localities in south-western Western Australia, an area where other arthropod taxa inhabiting these temperate rainforests show similar species distribution patterns (e.g. Rix <i>et al</i>. 2015; Sato <i>et al</i>. 2018; Schwentner and Giribet 2018). <i>Remarks</i> Amanda Mendes is included as a taxon author for this family to acknowledge her previous work, being the first to identify, recognise, and name this family in her unpublished dissertation. Although the morphological analyses of Mendes (2009) recovered <i>Pyenganella</i> in Lomanellidae, she expressed caution about the recovered relationship of <i>Pyenganella</i> with <i>Lomanella</i> and <i>Synthetonychia</i>. Simultaneously, she was certain that <i>Lomanella</i> represented a new family-level lineage as it was the only new name proposed based on her extensive morphological phylogenetic analyses. She approved the family diagnosis and agreed to authorship.
description: Published as part of <i>Derkarabetian, Shahan, Baker, Caitlin M., Hedin, Marshal, Prieto, Carlos E. &amp; Giribet, Gonzalo, 2021, Phylogenomic re-evaluation of Triaenonychoidea (Opiliones: Laniatores), and systematics of Triaenonychidae, including new families, genera and species, pp. 277-288 in Invertebrate Systematics 167 (1)</i> on page 140, DOI: 10.1071/is20047, http://zenodo.org/record/4531309
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