Empis (Enoplempis) shoshona Sinclair, Brooks & Cumming, 2025, sp. nov.
Creators: name: Sinclair, Bradley J.
name: Brooks, Scott E.
name: Cumming, Jeffrey M.
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Empididae
Empis
Empis shoshona

description: Empis (Enoplempis) shoshona sp. nov.
urn:lsid:zoobank.org:act: 4E7B127B-D438-493F-850C-FC0D52153F5A
(Figs 32–34, 38–40)
Type material. HOLOTYPE ♂, labelled: “USA: ID: Shoshone Co. / Rd 50; E Calder; St. Joe NF/ 20.vi. 2014, 680 m/ N 47°14.725′ W 116°02.473′/ ex. roadside str. B.J. Sinclair ”; “ CNC/ 1078372 ”; “ HOLOTYPE / Empis (Enoplempis) / shoshona Sinclair,/ Brooks & Cumming [red label]” [dissected] (CNC). PARATYPE: USA. Idaho: same data as holotype except, 47°15.820′N 116°08.803′W, roadside streams/ seeps (1♀, CNC).
Diagnosis. This species is distinguished from other species of the E. (En.) arrecta species group by a pair of narrow processes on the male hind tibia, hind tibia not broadened apically (Fig. 33), scutum without three brownish vittae and male cercus with broad, prolonged posterior process along epandrial margin with short ventroapical lobe.
Description. Wing length 4.0 mm. Male. Similar to E. (En.) diokra sp. nov., except as follows: head with eyes closely approximated, only very narrowly separated.
Thorax with thin whitish vittae along dc rows and either side of acr row when viewed at certain angles (Fig. 32). Anterior and posterior spiracles whitish (dried from alcohol).
Legs with apical fourth of hind femur and apical half of hind tibia darker. Hind femur slightly expanded preapically, with short, knob-like anteroventral process, shorter than quarter width of femur, with tuft of black setae directed posteriorly; anteroventral process flanked by broader inner lobe with ring of short apical setae and posterior face with apically projected spine; short, conical lobe posterior to inner lobe bearing hook-like seta; shorter, pointed ventral lobe posterior to hooked-tipped lobe; without posteroventral preapical lobe; apex with preapical row of 3–4 flattened, black, short blunt posterior setae; preapical anterior seta strong, shorter than width of femur (Figs 33, 34). Hind tibia narrowed basally and slightly twisted; with narrow, straight anteroventral digitiform process on basal quarter, slightly proximal to process of femur (when legs folded), with row of lateral setae; posteroventrally with digitiform process similar to anteroventral process, tapered with several lateral setae; with seta in posteroapical comb; apical half of tibia with long anterodorsal and posterodorsal setae, twice as long as width of tibia (Fig. 33).
Abdomen with pregenital segments unmodified; tergite 6 without posteroventral corner prolonged. Terminalia (Figs 38, 39): cerci separated; erect, projecting dorsally with crown of dense, black thickened setae; posterior margin with several setose projections; base broad, prolonged posteriorly along margin of epandrium, with ventrally directed apical extension. Subepandrial process strongly hooked, directed ventrally towards hypandrium. Epandrial lamella subrectangular, longer than high; broadly fused with cercus anteriorly; posterior margin without knob-like projection; setae similar to cercal setae. Hypandrium elongate, truncate, with shallow broad notch medially; bare, without setae. Phallus with expanded base, tuft of papilla-like projections at apex of expanded base near apex of hypandrium; strongly attenuated beyond base, arched apically with cylindrical, papilla-like projections along posterior margin; anterior margin with roughened surface (Fig. 39); apex emerging slightly beyond epandrium; ejaculatory apodeme about half-length of epandrium, T-shaped, with lateral apodemes medially.
Female. Similar to male, except as follows: frons subequal in width to width of anterior ocellus; hindleg without modified setae and processes; hind femur with strong anteroventral seta on apical fourth.
Geographical distribution and seasonal occurrence (Fig. 40). Empis (En.) shoshona sp. nov. is only known from Shoshone County in Idaho, collected in June.
Etymology. The species name is in reference to the type locality, Shoshone County, Idaho and is a noun in apposition.
Nuptial gift presentation. Form unknown.
description: Published as part of Sinclair, Bradley J., Brooks, Scott E. & Cumming, Jeffrey M., 2025, Revision of the western Nearctic species of Empis subgenus Enoplempis (Diptera: Empididae), pp. 1-200 in Zootaxa 5615 (1) on pages 17-20, DOI: 10.11646/zootaxa.5615.1.1, http://zenodo.org/record/15217957