Heraeus chamamecinus Dellapé & Melo & Henry 2016, SP. NOV.
Creators: name: Dellapé, Pablo M.
name: Melo, María C.
name: Henry, Thomas J.
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Rhyparochromidae
Heraeus
Heraeus chamamecinus

description: HERAEUS CHAMAMECINUS SP. NOV.  (FIGS 8F–G, 15C, 16C, 17I–L, 19)  
 Diagnosis   
Strongly setose. Ocelli located posteriorly. Distiflagellomere without or with only a weakly developed pale band. Profemur irregularly pigmented. Metafemur with a conspicuous dark apical band. Posterior margin of pygophore in dorsal view weakly bilobed; aedeagus with spines laterally to the ejaculatory reservoir and on the sclerotized vesical lobes forming an arc.  
 Heraeus chamamecinus sp. nov. shares with H. illitus and H. similis sp. nov. a highly setose body, and the presence of a slightly bilobed posterior margin of pygophore, but it can be distinguished by the absence of a well-defined pale band on distiflagellomere, and the short labium not surpassing the mesocoxae.   
 Description (Fig. 15C)   
Total length 6.18. Strongly setose.  
 Head: Convex dorsally, dark brown, shiny, coriaceus, with short recumbent and long erect setae. Head length 1.25, width 1.01, postocular length 0.46. Eyes protruding, not surpassing dorsal margin of head in lateral view. Ocelli placed posterior to an imaginary line passing posterior border of eyes. Interocular width 0.51, interocellar width 0.34. Labium pale brown, apex of article IV darker; with erect setae, extending to mesocoxae. Labial segment lengths: I 0.80, II 0.88, III 0.78, and IV 0.32. Antennal tubercles parallel. Antennae pale brown, except distal third of basiflagellomere and distiflagellomere darker; distiflagellomere without pale band; all segments with short recumbent and long erect setae, pedicel with abundant erect setae, equal to or longer than diameter of segment. Antennal lengths: scape 0.48, pedicel 1.18, basiflagellomere 0.96, distiflagellomere 1.02.   
 Thorax: Pronotum pruinose; dark brown, posterior lobe with irregular pale areas, and humeral angles dark brown with a small pale spot. Surface punctate, more conspicuous on posterior lobe. Anterior lobe with long erect and short recumbent setae, posterior lobe with erect setae shorter than setae of anterior lobe. Collar length 0.08, anterior lobe length 0.62, posteri- or lobe length 0.54; anterior lobe width 0.99, posteri- or lobe width 1.57. Pleurae brown. Evaporative area extended. Scutellum brown, pruinose, punctate, setae similar to setae on anterior lobe. Hemelytra pruinose, with erect and semi-erect setae. Colour pale brown, costal margin pale on proximal three-quarters, with a subapical corial spot, dark pattern as in Figure 15 (C); membrane with a pale, ovoid, apical spot, veins pale. Legs: Pale brown, coxae, profemur, and pretarsus darker; profemur irregularly pigmented; metafemur with a conspicuous dark band apically, less evident on mesofemur. Meso- and metatibia with a subproximal dark band, apex of tibiae darker (Fig. 16C). Legs with abundant erect setae, longest on profemur. Profemur with spines restricted to apical two-thirds. Protibia slightly curved.   
 Abdomen: Brown with abundant short recumbent setae. Male genitalia: Pygophore (Fig. 17I, J) subquadrangular, with posterior margin slightly bilobed and depressed towards aperture, anterior margin of dorsal aperture rounded, inner projections subquadrangular. Parameres: Figure 17 (K, L). Aedeagus (Fig. 8F, G): conjunctiva with an anterior lobe basal to ejaculatory reservoir; vesica with spines lateral to ejaculatory reservoir, forming an arc on the sclerotized vesical lobes; processus gonopori long and slender.   
 Variability observed in paratypes   
Some specimens show a weakly developed pale band on distiflagellomere.  
 Distribution   
Argentina and Brazil (Fig. 19).  
 Etymology   
The specific epithet is a noun in aposition and refers to the typical music style (Chamamé) from the Argentinian province of Corrientes.  
 Type material   
 Holotype: ♂, ARGENTINA, Corrientes: Ituzaingo, Reserva Santa María, 30-X-2003, T. luz, M.C. Melo (MLP).   
 Paratypes: 22♂, 14♀, same data as for holotype (MLP); 6♂, 8♀, same data as for holotype (USNM); 1♀, same data as for holotype, P.M. Dellapé (MLP); 2♂, 4♀, same data as for holotype, M. Chayle (MLP); 1♂, 31-X-2003, M.C. Coscarón col. (MLP); 2♂, 3♀, 29- X-2003, P.M. Dellapé col. (AMNH); 2♂, 1♀, 29-X- 2003, P. Dellapé col. (USNM); 6♂, 7♀, 29-X-2003, P.M. Dellapé col. (MLP); 1♂, 1♀, 27-IV-2003, M.C. Melo col. (MLP); 3♂, P.M. Dellapé col. (MLP); Tucumán: 1♂, Ciudad Universitaria, 800 m a.s.l., 18- II-[19]59, J.F.G. Clarke (USNM); 1♀, Depto Burruyacú, Rio Salas, 12-II-1982, R. Golbach (IFML); 1♀, Horco Molle, 1/ 18-IX-1966, L. Stange (IFML); Entre Rios: 5♂, X-1994 (USNM); 1♀, X-[19]93, E. Foerster (USNM); 1♂, 1♀, X-[19]93, Peña & Ugarte (USNM). BRAZIL: 1♀, Banhado Purane, 14- II-[19]02, E.G. & E.A. Monroe (CNC); Paraná: 1♂, 58 km S Curitiba, Sao Jose dos Pinhais, 24-XI- 1985, T.J. Henry (USNM).
description: Published as part of Dellapé, Pablo M., Melo, María C. & Henry, Thomas J., 2016, A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species, pp. 29-134 in Zoological Journal of the Linnean Society 177 (1) on pages 65-67, DOI: 10.1111/zoj.12362, http://zenodo.org/record/10109299