Hydromanicus naraik Olah & Barnard 2008, new species
Creators: name: Oláh, J.
name: Johanson, K. A.
Publisher: name: Zenodo
Year: 2008
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Trichoptera
Hydropsychidae
Hydromanicus
Hydromanicus naraik

description: Hydromanicus naraik Oláh & Barnard, new species   
Fig. 63–66  
 This species is morphologically similar to H. topali, new species. Hydromanicus naraik is distinguished in the ventral view by the presence of a clearly falciform cornute sclerotized endothecal process; the harpagones do not broaden in lateral view, not in ventral view; the foliose branches of the preanal appendages are one-third the length of the filiform branches, which have a sub-apical, lateral hump seen in dorsal view, and being capitate in lateral view. A striking difference is also the smaller endophallus located exclusively in the horizontal section of the phallotheca.   
Male. Body and wings brown. Forewings membrane uniformly coloured. Process and protuberance not seen on the cleared sternum V. Maxillary palp formula I-IV-III-II-V; segment V very thin, slender, slightly longer than segments I–IV together. Maxillary palps short, reaching end of mesonotum. Head dorsum brown with 7 brown setal warts. Swollen setal wart absent on proepisternum. Small setal wart, or setal area, present on precoxale (proepisternum). Pretarsal claws symmetrical, without laterally flanked setal bundle. Spur formula 244. Protibial posteroapical spur smaller than anteroapical spur.  
Wings. Venation typical for the genus. Forewing crossveins m-cu and cu separate. Sc and R separate before costa. Cu2 and A separate before wing margin. Hind wing Sc and R meet before costa at crossveins r and s. Crossveins r and s tangent. Stem of M and Cu1 located distantly of each other. False vein of convolution runs near and along stem of Cu1. Median cell open. Fork 1 present. Forewing length 9.2 mm, hind wing length 6.9 mm.  
Male genitalia. Abdominal segment IX fused annularly, short; dorsum one-fifth to one-sixth as long as ventrum (Fig. 63); median keel very short and broad (Fig. 64), with granulose dorsal and lateral surfaces; keel with slopes separated into 2 narrow lobes in dorsal view (Fig. 64); anterior margin shallowly arciform (Fig. 63). Antecosta forming single line narrowing gradually dorsally and ventrally before margin. External groove of antecostal suture well-developed. Apical lobe at posterolateral margin small, obtusely triangular, located immediately above basis of gonocoxites. Longitudinal sutures dividing segment IX ventrally of apical lobe. Posterior spine row intermittent, present from apical lobe to segment X; 2 or 3 short, stout setae present on slope of median keel (Fig. 63). Setal patches located ventrally of longitudinal sutures, composed of same type and length of stout setae. Depression between segments IX and X stepwise, with obtusely angled step corner. Segment X long, triangular, with rounded, blunt apex, slightly curving ventrad (Fig. 63), narrow in dorsal view (Fig. 64). Two pairs preanal appendages present, forming long, filiform, and short filiform plates covered by setae nested in elevated alveoli. Long filiform preanal appendages capitate in lateral view (Fig. 63), with subapical, lateral hump in dorsal view (Fig. 64). Apex of segment X with apicoventral setose lobes modified into pair of short, digitiform processes in dorsal view (Fig. 64); bluntly broad in lateral view (Fig. 63); apical halves covered by setae. Dorsal, interlobular gap narrow, deep, demarcated by apicoventral setose processes. Apicodorsal setose lobe reduced to setose surface merged with setose surface of apicoventral lobe. Segment X without smooth cavity. Traverse and longitudinal sutures absent. Coxopodites basally wide; straight along their lengths; strongly exceeding apex of segment X; in ventral view (Fig. 65) stout, slightly narrowing at mid-length before broadening apically. Harpagones broad in lateral view (Fig. 63), parallel-sided and less broad in ventral view; apex slightly excised (Fig. 65), with mesal lobe longer than lateral lobe. Phallic apparatus (Fig. 66) uniformly bent; anteriorly obtusely angled ventrad; broad tube of horizontal phallotheca with nearly straight dorsum, slightly concave ventrally and at mid-length. Distal part of phallotheca ending in 2 pairs sclerous processes; pair of smaller processes located dorsally, forming beak-shaped phallotremal sclerites surrounding phallotrema; pair of large cornute processes originating from middle of apex. Sclerous endothecal process curving mesodorsad, slender; forming long corniform arch. Cornute, sclerotized endothecal process falciform in ventral view. Pair of medium-sized, membranous, ventral, endothecal lobes orienting apicoventrad covering basal part of endothecal process. Endophallus located in horizontally in phallotheca, anteriorly ending in narrow, short tube with gonopore.  
 Holotype male: INDIA: Tamil Nadu, Naraikkadu, 2500–3000 feet, Tinnevelly [Tirunelveli], 11– 13.iii.1936 (Brit. Mus. 1936-512, NHML, alcohol).   
 Distribution. India.   
 Etymology. naraik, derived from the name of the type locality, Naraikkadu.
description: Published as part of Oláh, J. & Johanson, K. A., 2008, Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae), pp. 1-248 in Zootaxa 1802 on pages 43-46