Phygopoda chaquensis Clarke 2017, sp. nov.
Creators: name: Clarke, Robin O. S.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Cerambycidae
Phygopoda
Phygopoda chaquensis

description: Phygopoda chaquensis sp. nov.   
Fig. 5, 6  
 Holotype. Male: 6.20 mm. Deposited at MNKM.   
 Description of holotype (Fig. 5). Small. Forebody and abdomen of equal length. Prothorax subcylindrical, 1.13 wider than head with eyes.   
 Surface features. Color: forebody and sides of abdomen black, rest of abdomen dark chestnut. Labial and maxillary palpomeres chestnut. Elytra chestnut, with all margins darker (and elytral apex chestnut); but humeri paler and large translucent panels dark testaceous. Antennae chestnut basally, rest browner. Legs generally chestnut (including all tarsi); front and middle legs, and basal half of metafemoral peduncle paler; rest of metafemoral peduncle and lateral surface of mesofemoral clave darker; metatibiae (and its setae) almost black; metatarsal setae chestnut. Pubescence: away from midline especially dense on abdomen; dense on underside of pro- and mesofemora; metatibial setae short, moderately sparse and only denser for apical fifth (but not thick enough to be called a brush). Puncturation: punctures on translucent panels small, and separated by slightly more, to three times their diameter; on center of metasternum shallow and not rugose; on abdomen dense at base of each segment and towards sides of each one (in places deeper and strongly beveled), towards midline sparse.   
 Structure. Head: rostrum moderately long and wide (width/length 3.00). Clypeus narrow, flat, on lower plane than labrum and separated from it by raised border. Frons moderately short, transversely depressed; frontal suture eliminated by dense punctures. Inferior lobes 1.18 longer than wide; width of one lobe/interocular distance about 5.70. Superior lobes of eyes narrow (ca. 0.10 mm wide medially, slightly narrowed laterally); with about 7 rows of ommatidia medially, 6 laterally; and separated by 3.00 times their own width. Antennal tubercles separated by 2.33 width of scape. Antennae: apex reaching middle of urosternite II and passing apex of elytra towards apex of antennomere VII. Scape nearly cylindrical with weakly bent base (when viewed laterally). Lengths (mm) antennal segments as follows: scape 0.45, pedicel 0.20, III 0.55, IV 0.45, V-VI 0.55, VII 0.50, VIII 0.40, IX 0.35, X 0.30, XI 0.40.   
 Prothorax: length 1.25 mm, 1.09 longer than wide; sides slightly sinuate for apical half, widest behind middle (prothoracic quotient 1.67). Prosternum declivous across apical fifth, nearly flat to base of prosternal process. Prosternal process: flat; base sub-laminate (ca. 0.10 mm), about 3.0 times narrower than procoxal cavities; apex of process a moderately small, flat, equilateral triangle (but details partially hidden).   
 Mesothorax: elytral length 1.70 mm, 1.55 longer than wide (1.10 mm); humeri wide enough to hide sides of mesothorax; towards apex surface with slight transverse declivity. Mesosternum: width of mesocoxal cavity 2.00 times wider than base of mesosternal process. Length of mesosternum/length of metasternum 0.70.   
 Metathorax: width 1.2 mm, body length/width metathorax about 5.0; moderately rounded at sides, oblique to middle of metasternal apex. Metasternum convex, moderately tumid at center; longitudinal suture represented by shallow stria basally, ending in small oval depression towards apex.   
 Abdomen: somewhat more fusiform; relatively short; convex; urosternites I-IV widest (0.85 mm) at middle of III, narrowest (0.65 mm) at apex of IV. Urosternite I slightly elongate (0.95 mm), sides straight; II and III (0.60 mm), II sides almost straight, III sides weakly rounded, IV (0.50 mm) sides weakly rounded. Urosternite V (0.35 mm) trapezoidal, transverse; sides sharply pointed (and the same when viewed laterally). Abdominal process a large, wide isosceles triangle (with broad apex) coplanar with abdomen. Apical tergite convex.   
 Legs: ratio length front/middle/hind leg 1.0:1.1:3.4; and body length/length of leg 2.6, 2.4, 0.8 respectively. Front and middle legs: moderately slender, front leg rather long; their claves somewhat tumid mesally and widest at middle (when viewed from above). Front leg: tibia gradually widening to apex; apico-lateral margin not toothed. Middle leg: femur 1.60 longer than tibia; length of femur/lateral width of clave 4.00. Hind leg: length 8.25 mm; clave comparatively long and weakly abrupt; lengths clave/ peduncle 0.49; femora (3.65 mm) 1.07 longer than tibiae (3.40 mm); tibia bisinuate (when viewed from above), almost straight (when viewed from the side); tarsus short (1.20 mm), about one third length of tibia, and narrower than apex of latter; metatarsomere I gradually widened to apex, 1.25 longer than lengths of II+III.   
 Male variation. Among the paratypes two have paler legs, about half have paler chestnut abdomens and antennae; and in two specimens elytral apices are pale chestnut; in one pronotum slightly chestnut; abdomen paler chestnut in two paratypes. Antennal variation: minor differences include V-VII may be equal in length; VI nearer filiform; VIII narrower at base and slightly more serrate; IX and X slightly more serrate. In two paratypes prothorax widest well behind middle (prothoracic quotient 1.38); in all paratypes sides of pronotum lack noticeable smooth areas (as in other species), and one lacks any smooth interstices (as described for the holotype). Elytra: dehiscent for apical half; in half of the paratypes elytral punctures slightly larger and denser; in some elytra slightly narrower towards apex; in one apical margin more rounded. Small differences in proportions of abdomen are common; in one paratype it is narrow and cylindrical; lengths forebody/abdomen 0.95-1.09.   
 Description of female (Fig. 6). Female allotype differs from male holotype as follows: total length 7.60 mm. More robust than male. Lengths forebody/abdomen 0.78.   
 Surface features. Color: underside of prothorax and abdomen clouded chestnut; basal segments of antenna orange, apical ones duskier; apical half of metafemora, metatibiae (and its setae) and metatarsi blacker. Surface ornamentation: similar to holotype on dorsad; but generally less pubescent, noticeably so on underside (with longer pubescence much reduced on forebody); and less densely punctured on metathorax and abdomen. Pubescence on abdomen dense but shorter, fascia on sides of abdominal segments much broader; metatibiae more strongly setose, almost brush-like towards apex.   
 Structure. Head: narrower (widths head with eyes/prothorax 1.28). Rostrum longer (width/length 2.60). Inferior lobes of eyes considerably shorter and narrower (length 0.40 mm, width 0.33 mm); less convex (lying slightly below level of interocular); well separated (width interocular 0.25 mm, widths lobe/interocular 1.30). Superior lobes slightly wider, separated by about 2.67 width of lobes. Interocular flat, densely punctured; frontal suture reaching frons and antennal tubercles. Antennae more robust and slightly shorter, reaching basal third of urosternite III; only VII and VIII serrate (but weakly so); XI less elongate.   
 Prothorax: more elongate, 1.13 longer than wide; prothoracic quotient 1.63; and lacking smooth areas towards sides of pronotum. Prosternum almost completely flat (including base of process); widths procoxal cavity/base of process 3.50; apex of process triangular and transverse.   
 Meso- and metathorax: Elytra slightly shorter (length/width 1.50) only reaching middle of metacoxae); less dehiscent; humeri hardly hide mesepimerum. Meso- and metathorax very similar to those of holotype, except: mesosternal declivity less abrupt; widths mesosternal process/coxal cavity 2.33; metathorax almost parallel-sided (body length/width metathorax 5.6).   
 Abdomen: more fusiform and convex, wider than holotype; length urosternites 1-V: 2.3, 1.6, 1.5, 1.2, 1.6), V conical and much longer. Last visible tergite conical, elongate, weakly convex, rounded at apex.   
 Legs: slightly more robust and longer than in male, ratio of length from front to hind leg 1.0:1.2:4.1; body length/length of leg 3.8, 3.1, 0.8 respectively. Middle leg: femur 1.42 longer than tibia; length of femur/lateral width of clave 4.50; clave 1.70 longer than peduncle. Hind leg: longer (9.55 mm) than in male, clave more abrupt and shorter (with longer abdomen base of clave passes middle of urosternite V); peduncle longer (lengths clave/peduncle 0.43); femur distinctly longer than tibia (lengths femur/tibia 1.20); tibia straight (when viewed from the side); tarsus 3.17 shorter than tibia; metatarsomere I narrower and more cylindrical than in male, but also 1.25 longer than lengths of II+III.   
 Color variation in females. The single paratype varies from the allotype by the following: antennae as in holotype; disc of pronotum clouded chestnut; apices of elytra black; abdomen pale chestnut; hind leg with femoral claves and tarsi chestnut.   
 Diagnosis. Lack of distinct brush on metatibia (especially in males); inferior lobes of eyes in males less contiguous; abdomen densely pubescent (in males long and erect, in female shorter and less erect). Chaco forest species.   
 Species sample data.   
 Measurements (mm): 13 males, 2 females: total length 5.4-6.9/7.4-7.6; length of pronotum 0.90- 1.25/1.25-1.40; width of pronotum 0.80-1.05/1.10-1.15; length of elytra 1.40-1.70/1.75-1.80; width at humeri 0.85-1.05/1.10-1.20.   
 Type material: Holotype male, BOLIVIA, Tarija, 4 km W Villamontes, 2 km S Puente Usterez, west bank of Rio Pilcomayo, 21°16’S / 63°27’W, 366 m, flying to/on flowers of Croton sp. A, 12.XII.2011, Clarke and Zamalloa col. (MNKM).   
 Paratypes with same data as holotype: male, 12.XII.2011 (RCSZ); same data (MZSP); same data (EMEC); same data (CMNH); same data (BMNH).   
 Paratypes with different data from holotype: Santa Cruz, Prov. Cordillera, road to Itai, 83 km N Camiri, 19°20.547S / 63°28.870W, 890 m, male 17.XII.2011, S. Lingafelter col. (SLPC); 24 km S Camiri, 20°12.0’S / 63°28.4’W, 885 m, on Croton flowers, male, 18.XII.2011, S. Lingafelter col. (SLPC); same data, 2 males, Wappes and Lingafelter col. (ACMT).   
 Tarija, G. Chaco Rd to Tarija, 22 Km W Villamontes, 21°15’S / 63°34’W, 3 males, 13-15.XII.2011, Wappes, Bonasso and Morris (ACMT); same data, male (RFMC); same data, female (RCSZ).   
 Etymology. The specific epithet, “chaquensis”, simply means “from the Chaco ” in Latin.
description: Published as part of Clarke, Robin O. S., 2017, Bolivian Rhinotragini XI. New species of Phygopoda Thomson, 1864 and Phygopoides Peñaherrera-Leiva and Tavakilian, 2007 (Coleoptera, Cerambycidae), pp. 1-18 in Insecta Mundi 2017 (568) on pages 9-11, DOI: 10.5281/zenodo.5170467