cometadata/datacite-rtg-text-other-reclassification-records huggingface dataset:
Ectobius punctatissimus Ramme 1922
Creators: name: Hristov, Georgi H.
name: Chobanov, Dragan P.
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Blattodea
Ectobiidae
Ectobius
Ectobius punctatissimus

description: Ectobius punctatissimus Ramme, 1922
Figure 60
Ectobius punctatissimus was recorded from Strandzha Mountain (Gramatikovo village, 5.VI–6.VI.1959, leg. R. Bańkowska) by Bazyluk (1961: 432). All subsequent publications mentioning this species from Bulgaria (Princis 1963: 68; Princis 1965: 17, 26; Princis 1971: 1072; Harz & Kaltenbach 1976: 249; Hubenov et al. 1998: 341) rely on the latter record.
This species occurs in Italy and the western part of the Balkan Peninsula as south as Albania. No confirmed data for the occurrence of this species in Bulgaria exists. Though we were unable to study the material recorded by Bazyluk (1961), based on the remote range of the species and own unpublished data (Hristov, unp.) we consider the records for this species in Bulgaria erroneous.
description: Published as part of Hristov, Georgi H. & Chobanov, Dragan P., 2016, An annotated checklist and key to the Bulgarian cockroaches (Dictyoptera: Blattodea), pp. 351-388 in Zootaxa 4154 (4) on page 360, DOI: 10.11646/zootaxa.4154.4.1, http://zenodo.org/record/264825
Megachile (Rhyssomegachile) Mitchell
Creators: name: Gonzalez, Victor H.
name: Griswold, Terry
name: Engel, Michael S.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Megachilidae
Megachile

description: Subgenus Rhyssomegachile Mitchell
Figures 1, 4A, C, D, E, 14–20
Cressoniella (Rhyssomegachile) Mitchell, 1980: 63.
description: Published as part of Gonzalez, Victor H., Griswold, Terry & Engel, Michael S., 2018, South American Leaf-Cutter Bees (Genus Megachile) Of The Subgenera Rhyssomegachile And Zonomegachile, With Two New Subgenera (Hymenoptera: Megachilidae), pp. 1-73 in Bulletin of the American Museum of Natural History 2018 (425) on pages 1-73, DOI: 10.1206/00030090-425.1.1, http://zenodo.org/record/4613364
Megachile (Rhyssomegachile)
Creators: name: Gonzalez, Victor H.
name: Griswold, Terry
name: Engel, Michael S.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Megachilidae
Megachile

description: Key to Species of Rhyssomegachile
FEMALES
1. Carina behind gena continuing onto vertex (fig. 15C, D); dorsum of mesosoma with long (≥2.0× OD), light reddish-brown setae (fig. 15B, C); forewing yellowish with yellow to light brown veins (fig. 15B) ....................... ............................. M. guayaqui Schrottky
— Carina behind gena not continuing onto vertex; dorsum of mesosoma with shorter (≥1.0× OD), yellowish or grayish setae; forewing brownish with dark brown veins........2
2(1).Disc of mesoscutum with punctures distinctly spaced, not contiguous, separated by 1–2× a puncture width (fig. 20D) ................. ............................................... M. urbana Smith
— Disc of mesoscutum with punctures contiguous or nearly so (fig. 4A) ..................... 3
3(2).Ventral surface of mesepisternum and fore- and middle legs with coxae, trochanters, base of femora, and outer surfaces of tibiae and basitarsi densely covered by corkscrewlike setae (fig. 4D, E); clypeus and supraclypeal area finely and densely punctate (fig. 16B); T2–T5 without apical fasciae (fig. 16D); T6 with short, appressed brownishgray setae, without longer, erect setae .......... ......................... M. kartaboensis Mitchell — Ventral surface of mesepisternum and all legs distinctly covered by thick, minutely branched, and apically curved setae (fig. 17C); clypeus and supraclypeal area coarsely punctate (fig. 17A); T2–T5 each with apical fasciae at least laterally (fig. 17D, E); T6 covered mostly by short, appressed yellowish setae from which scattered, longer, erect setae arise ................... M. simillima Smith
description: Published as part of Gonzalez, Victor H., Griswold, Terry & Engel, Michael S., 2018, South American Leaf-Cutter Bees (Genus Megachile) Of The Subgenera Rhyssomegachile And Zonomegachile, With Two New Subgenera (Hymenoptera: Megachilidae), pp. 1-73 in Bulletin of the American Museum of Natural History 2018 (425) on pages 1-73, DOI: 10.1206/00030090-425.1.1, http://zenodo.org/record/4613364
Chrysosarus (Zonomegachile) Mitchell
Creators: name: Gonzalez, Victor H.
name: Griswold, Terry
name: Engel, Michael S.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Megachilidae
Chrysosarus

description: Subgenus Zonomegachile Mitchell
Figures 2, 4F, 5F, H, 21–34
Chrysosarus (Zonomegachile) Mitchell, 1980: 72.
description: Published as part of Gonzalez, Victor H., Griswold, Terry & Engel, Michael S., 2018, South American Leaf-Cutter Bees (Genus Megachile) Of The Subgenera Rhyssomegachile And Zonomegachile, With Two New Subgenera (Hymenoptera: Megachilidae), pp. 1-73 in Bulletin of the American Museum of Natural History 2018 (425) on pages 1-73, DOI: 10.1206/00030090-425.1.1, http://zenodo.org/record/4613364
Megachile (Zonomegachile) paisa Gonzalez & Griswold & Engel 2018, new species
Creators: name: Gonzalez, Victor H.
name: Griswold, Terry
name: Engel, Michael S.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Megachilidae
Megachile
Megachile paisa

description: Megachile (Zonomegachile) paisa, new species
Figures 21, 31
DIAGNOSIS: This species can be recognized by the following combination of features: head and mesosoma mainly with light reddish-brown pubescence (fig. 31A–C); mesoscutum microalveolate to imbricate among setiferous punctures; probasitarsus robust, about 2.5× longer than broad (fig. 31D); S2 and S3 basally with white scopal setae, remaining of S3 and S4 with light reddish-brown setae, sides of S3, S4, and entire S5 and S6 with black setae. This species is most similar to M. durantae from northwestern Brazil. In addition to the geographic separation, it can be distinguished by the pubescence of the head and mesosoma, which is light reddish brown in M. paisa and whitish in M. durantae. The wings in M. paisa are slightly more yellow than those of M. durantae.
DESCRIPTION: Holotype, Female: As described for M. durantae except as follows: total body length 11.9 mm; forewing length 10.3 mm; head width 4.4 mm. Interocellar distance 2.5× OD, about as long as ocellocular distance. Wings slightly more yellow than those of M. durantae. Pubescence predominantly light reddish brown on head and mesosoma except whitish on outer surfaces of tibiae. S2 and S3 basally with white scopal setae, remaining of S3 and S4 with light reddish-brown setae, sides of S3, S4, and entire S5 and S6 with black setae.
HOLOTYPE: ♀, Colombia: Porce. Ant. [Antioquia], (Col.), San Ignacio B, 2–4 PM, Pescado, 16/04/1997 [16 April 1997], Allan Smith P (BBSL).
ETYMOLOGY: The specific epithet is a selfreferential nickname for inhabitants of the department of Antioquia, the area where this species is currently known.
COMMENTS: The type specimen is not in very good condition. The apical margins of the wings are strongly notched and the setae are plastered against the integument, including the white fasciae on the apical margins of the terga and sterna. Megachile paisa is exceptionally similar morphologically to M. durantae, differing in the color of the wings and body pubescence. Both species are known only from the female type and they could be conspecifics. However, M. paisa is known from northwestern Colombia, on the eastern slope of the Cordillera Occidental. Thus, it is on the other side of the Andes, more than 2000 km distant from the type locality of M. durantae in Rondônia, Brazil, localities that represent rather different habitats.
description: Published as part of Gonzalez, Victor H., Griswold, Terry & Engel, Michael S., 2018, South American Leaf-Cutter Bees (Genus Megachile) Of The Subgenera Rhyssomegachile And Zonomegachile, With Two New Subgenera (Hymenoptera: Megachilidae), pp. 1-73 in Bulletin of the American Museum of Natural History 2018 (425) on pages 1-73, DOI: 10.1206/00030090-425.1.1, http://zenodo.org/record/4613364
Megachile (Zonomegachile) nigribarbis Vachal
Creators: name: Gonzalez, Victor H.
name: Griswold, Terry
name: Engel, Michael S.
Publisher: name: Zenodo
Year: 2018
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Megachilidae
Megachile
Megachile nigribarbis

description: Megachile (Zonomegachile) nigribarbis Vachal
Megachile nigribarbis Vachal, 1909: 6 (holotype ♂, MNHN: Mato Grosso, Brazil). Raw, 2002: 37 (placement in Zonomegachile
description: Published as part of Gonzalez, Victor H., Griswold, Terry & Engel, Michael S., 2018, South American Leaf-Cutter Bees (Genus Megachile) Of The Subgenera Rhyssomegachile And Zonomegachile, With Two New Subgenera (Hymenoptera: Megachilidae), pp. 1-73 in Bulletin of the American Museum of Natural History 2018 (425) on pages 1-73, DOI: 10.1206/00030090-425.1.1, http://zenodo.org/record/4613364
Yezoceryx albimaculatus Sheng & Sun 2010
Creators: name: Achterberg, Cornelis Van
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Yezoceryx
Yezoceryx albimaculatus

description: Yezoceryx albimaculatus Sheng & Sun, 2010
(Figure 1)
Yezoceryx albimaculatus Sheng & Sun, 2010: 38. Holotype: ♀, China: Guangdong (GSFPM).
Diagnosis. Occipital carina narrowly dipped dorsally; vertex behind lateral ocelli strongly prominent above level of ocelli; first tergite about 2.1× as long as its posterior width; ovipositor length about 3.1× length of hind tibia, first sternite sharply hooked anteriorly; body black with many yellowish/whitish marks.
Material examined. Vinh Phuc Province, Tam Dao NP: 1♂ (OMNH), 1.vi.1997, 500– 1000 m a.s.l.; 1♂ (OMNH), 1.v.1998, 900– 1200 m a.s.l., R. Matsumoto leg.
Distribution. Previously known only from China (Guangdong Province), these are the first records of this species from Vietnam.
Remarks. Vietnamese specimens have the subalar prominence entirely black and in males the middle part of the median lobe of the mesoscutum is not so deeply impressed as in males from China.
description: Published as part of Achterberg, Cornelis Van, 2017, First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species, pp. 345-372 in Zootaxa 4311 (3) on page 347, DOI: 10.11646/zootaxa.4311.3.2, http://zenodo.org/record/847854
Yezoceryx Uchida 1928
Creators: name: Achterberg, Cornelis Van
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Yezoceryx

description: Yezoceryx Uchida, 1928
Yezoceryx Uchida, 1928: 36.
Type species: Yezoceryx scutellaris Uchida, Original designation.
Diagnosis. Upper tooth as long as to shorter than lower tooth; clypeus somewhat flattened, with a semi-circular or straight transverse subapical ridge, median tubercle usually present on transverse ridge; frons always with strong median longitudinal carina; notauli deep and posteriorly convergent; epicnemial carina usually complete, sometimes erased dorsally; fore wing without areolet, vein 2 rs-m opposite or basad of 2 m-cu; fore and mid tarsal claws with acute accessory tooth near apex; hind tarsal claw simple; posterior transverse carina of propodeum absent medially so that area superomedia and area petiolaris confluent; first sternite usually with a sharp tubercle; female hypopygium large and triangular, folded on midline, and reaching or surpassing metasomal apex; ovipositor tip with faint transverse ridges.
description: Published as part of Achterberg, Cornelis Van, 2017, First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species, pp. 345-372 in Zootaxa 4311 (3) on page 346, DOI: 10.11646/zootaxa.4311.3.2, http://zenodo.org/record/847854
Yezoceryx lii Sheng & Sun 2013
Creators: name: Achterberg, Cornelis Van
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Yezoceryx
Yezoceryx lii

description: Yezoceryx lii Sheng & Sun, 2013
(Figure 5)
Yezoceryx lii Sheng & Sun, 2013: 35. Holotype: ♀, China: Jiangxi (GSFPM).
Diagnosis. Mesoscutum with a triangular black mark on posterior half of median lobe; scutellum with lateral carina extending beyond middle; first tergite 1.8× as long as its posterior width; first sternite with pointed hook; ovipositor about 2.3× length of hind tibia.
Material examined. Dak Lak, Chu Yang Sin NP, Krong K’Mar: 1♀ (RMNH), 2–10.vi.2007, 740– 900 m a.s.l., Malaise trap, C. v. Achterberg & R. de Vries leg.
Distribution. Previously known from China (Jiangxi Province) (Sheng et al., 2013). This is the first record of this species from Vietnam.
Remarks. The female specimen from Vietnam has a broader black mark on the ocellar area laterally. Metasomal tergites have black stripes rather than the brown stripes of the holotype.
description: Published as part of Achterberg, Cornelis Van, 2017, First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species, pp. 345-372 in Zootaxa 4311 (3) on page 354, DOI: 10.11646/zootaxa.4311.3.2, http://zenodo.org/record/847854
Yezoceryx maculatus Chao 1981
Creators: name: Achterberg, Cornelis Van
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Yezoceryx
Yezoceryx maculatus

description: Yezoceryx maculatus Chao, 1981
(Figure 7)
Yezoceryx maculatus Chao, 1981: 203. Holotype: ♀, China: Jianyang xian (FACS).
Diagnosis. Brown with black marks on mesosoma and metasomal tergites; basal area of propodeum inverted trapezoid; first tergite about 2.0–2.4× as long as posterior width; first sternite with short, anteriorly projecting hook; ovipositor 2.6–3.1× length of hind tibia.
Material examined. Vinh Phuc, Tam Dao NP: 1♀ (OMNH), 6.v.1998, 900– 1200 m a.s. l., hand net, R. Matsumoto leg.; Ninh Binh, Cuc Phuong NP: 1♂ (RMNH) near entrance, 14.iv–1.v.2000, Malaise trap, Q.P. Mai leg; 1♀ (RMNH) near entrance, 15–27.v.2000, Malaise trap, Q.P. Mai leg.; Thua Thien-Hue, Phong Dien NR: 1♀ (RMNH), nr base camp, 15 km W Phong My, 23.iii–6.iv.2001, 80– 210 m a.s.l., Malaise trap, C. v. Achterberg & R. de Vries leg.; Nghe An, Pu Mat NP: 1♂ (IEBR), 17.iv.2006, hand net, N.T. Pham leg.; Dong Nai, Cat Tien NP: 2♀ (RMNH, IEBR), 9.iv–13.v. 2007, 100 m a.s.l., Malaise trap, Q.P. Mai, M.T. Nguyen & C. v. Achterberg leg.; Son La, Thuan Chau, Co Ma: 1♀ (IEBR), 5.vi.2008, 1200 m a.s.l., hand net, H.X. Le leg.; Hoa Binh, Mai Chau, Tan Son: 1♀ (IEBR), 10–15.vi.2010, 800– 900 m a.s.l., Malaise trap, L.D. Khuat leg.; Cao Bang, Phia Oac NR: 1♀ 1♂ (IEBR), 8–18.v.2013, Malaise trap, T.H. Pham leg.
Distribution. Previously known only from China (Chao, 1981). These are the first records of this species from Vietnam.
Remarks. Vietnamese specimens have a slightly more elongate first tergite (2.1–2.4×) than the holotype (2.0×), according to its original description (Chao, 1981).
description: Published as part of Achterberg, Cornelis Van, 2017, First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species, pp. 345-372 in Zootaxa 4311 (3) on pages 355-356, DOI: 10.11646/zootaxa.4311.3.2, http://zenodo.org/record/847854
Yezoceryx nebulosus Wang 1982
Creators: name: Achterberg, Cornelis Van
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Yezoceryx
Yezoceryx nebulosus

description: Yezoceryx nebulosus Wang, 1982
(Figure 8)
Yezoceryx nebulosus Wang, 1982a: 97. Holotype: ♀, China: Sichuan (IZCAS) Diagnosis. Dark brown to black; face largely black; pronotum and mesopleuron smooth, without striae; first tergite elongate, 2.6–3.0× as long as its posterior width; first sternite with blunt, short projecting hook; ovipositor 4.0× length of hind tibia.
Material examined. Cao Bang, Phia Oac NR: 1♀ (IEBR), 8–18.v.2013, Malaise trap, T.H. Pham leg.; Son La, Copia NR: 2♂ (IEBR), Malaise trap, 27.iv-2.v.2016, T.V. Hoang leg.
Distribution. Previously known only from China (Sichuan) (Wang, 1982a), these are the first records of this species from Vietnam.
Remarks. The first tergite of the Vietnamese specimens is about 3.0–3.3× longer than its posterior width, a little more elongate than that of the holotype (2.6×).
description: Published as part of Achterberg, Cornelis Van, 2017, First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species, pp. 345-372 in Zootaxa 4311 (3) on pages 356-357, DOI: 10.11646/zootaxa.4311.3.2, http://zenodo.org/record/847854
Yezoceryx maculataexcels Wang 1993
Creators: name: Achterberg, Cornelis Van
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Yezoceryx
Yezoceryx maculataexcels

description: Yezoceryx maculataexcels Wang, 1993
(Figure 6)
Yezoceryx maculataexcels Wang, 1993: 206. Holotype: ♂, China: Yunnan (IZCAS).
Diagnosis. Yellow with black marks; median carina of frons forming broad tubercle between antennal sockets; mesoscutum with U-shaped black mark; first tergite about 1.7–1.9× as long as its posterior width; first sternite with blunt tubercle or weak anteriorly projecting hook; ovipositor 2.5× length of hind tibia.
Material examined. Hoa Binh, Yen Thuy, Lac Thinh: 1♀ (IEBR), 1–10.x.2002, Malaise trap, L.D. Khuat leg.; 1♂ (IEBR), 20.iv.2003, hand net, L.D. Khuat leg.; Hoa Binh, Yen Thuy, Da Phuc: 1♂ (IEBR), 10–20.x.2003, Malaise trap, L.D. Khuat leg.; Nghe An, Chau Khe, Khe Bu: 2♂ (IEBR), 17.iv.2006, hand net, N.T. Pham leg.; Bac Can, Ba Be NP: 1♂ (OMNH), 2.v.2006, 20.23 0 N 105.37 0E, 200 m a.s.l., R. Matsumoto leg.; Son La, Co Ma, Thuan Chau: 1♀ (IEBR), 4.vi.2008, 1200 m a.s.l., hand net, H.X. Le leg.; Kon Tum, Chu Mon Ray NP: 1♂ (IEBR), 21.v.2009, hand net, T.V. Hoang leg.; Kon Tum, Ngoc Linh NR: 1♂ (IEBR), 11.iv.2015, 1204 m a.s.l., 15 0 13.78N 107 0 43.7E, hand net, T.P.L. Nguyen & D.D. Nguyen leg.; Hoa Binh, Ngoc Son-Ngo Luong NR: 5♂ (IEBR), 17– 22.ix. 2016, 690 m a.s.l., Malaise trap, P.V. Pham leg.
Distribution. Previously known from China (Wang, 1993). These are the first records of this species from Vietnam.
description: Published as part of Achterberg, Cornelis Van, 2017, First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species, pp. 345-372 in Zootaxa 4311 (3) on page 354, DOI: 10.11646/zootaxa.4311.3.2, http://zenodo.org/record/847854
Yezoceryx purpuratus Sonan 1936
Creators: name: Achterberg, Cornelis Van
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Yezoceryx
Yezoceryx purpuratus

description: Yezoceryx purpuratus Sonan, 1936
(Figure 11)
Yezoceryx purpuratus Sonan, 1936: 250. Holotype: ♀, Formosa: Mekaran in Chikuto-gun in Shinchiku Prefecture (TARI).
Diagnosis. Blue purple; occipital carina down-curved dorsally; clypeus without median tooth, apical margin distinctly up-curved; propodeum with dense, irregular wrinkles; metasomal first tergite about 2.0× as long as its apical width; first sternite with anteriorly projecting hook; ovipositor 3.6–4.0× as long as hind tibia.
Material examined. Vinh Phuc, Tam Dao NP: 1♀ (OMNH), 3.v.1998; 1♀ (OMNH), 4.v.1998, 900– 1000 m a.s.l., hand net, R. Matsumoto leg.; 1♀ (IEBR), Hoa Binh, Mai Chau, Tan Son, 20–25, v.2009, 850– 900 m a.s.l., Malaise trap, L. D. Khuat leg.
Distribution. Previously known from Taiwan (Sonan, 1936), these are the first records of this species from Vietnam.
Remarks. The Vietnamese specimens have partly yellow inner orbits (at the level of the upper face).
description: Published as part of Achterberg, Cornelis Van, 2017, First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species, pp. 345-372 in Zootaxa 4311 (3) on page 360, DOI: 10.11646/zootaxa.4311.3.2, http://zenodo.org/record/847854
Yezoceryx qinlingensis Wang 1993
Creators: name: Achterberg, Cornelis Van
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Yezoceryx
Yezoceryx qinlingensis

description: Yezoceryx qinlingensis Wang, 1993
(Figure 12)
Yezoceryx qinlingensis Wang, 1993: 208. Holotype: ♂, China: Shaanxi Province, Taibai Mt. (IZCAS).
Diagnosis. Body brown with black marks; antenna of female unicolourous, without white band; first tergite 2.3× as long as posterior width; first sternite with anteriorly projecting hook; ovipositor about 3.6× as long as hind tibia.
Material examined. Cao Bang, Phia Oac NR: 1♀ (IEBR), 8–18.v.2013, Malaise trap, T.H. Pham leg. Distribution. Previously known from China (Shaanxi Province) (Wang, 1993), this is the first record of this species from Vietnam.
Remarks. The Vietnamese specimen has the mesopleuron and metapleuron darker than those of the holotype.
description: Published as part of Achterberg, Cornelis Van, 2017, First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species, pp. 345-372 in Zootaxa 4311 (3) on pages 361-362, DOI: 10.11646/zootaxa.4311.3.2, http://zenodo.org/record/847854
Yezoceryx umaculosus Wang 1982
Creators: name: Achterberg, Cornelis Van
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Yezoceryx
Yezoceryx umaculosus

description: Yezoceryx umaculosus Wang, 1982
(Figure 17)
Yezoceryx umaculosus Wang, 1982b: 204. Holotype: ♀, China: Yunnan (IZCAS).
Diagnosis. Yellow with black marks; mesoscutum with U-shaped black mark; scutellum usually with black spot posteriorly; first tergite about 1.4–1.6× as long as its apical width; first sternite with anteriorly projecting hook; ovipositor about 1.8× length of hind tibia.
Material examined. Vinh Phuc, Tam Dao NP: 1♀ (IEBR), 2.vi.1997, 900– 1000 m a.s.l.; 1♂ (OMNH), 7.v.1998, 3♂ (OMNH), 8.v.1998, hand net, R. Matsumoto leg.; Ninh Binh, Cuc Phuong NP: 1♂ (OMNH), 26.iv.1998; 1♀1♂ (OMNH), 27.iv.1998, 200– 300 m a.s.l., hand net, R. Matsumoto leg.; Thua Thien Hue, Phong Dien NR: 5♀ 8♂ (RMNH, IEBR), 15 km W Phong My, 23.iii–6.iv.2001, 80– 210 m a.s.l.; 1♀ 2♂ (RMNH), 12 km W Phong My, 24.iii-6.iv.2001, 60 m; 1♀ (RMNH), 15 km W Phong My, 23.iii–6.iv.2001, 50– 60 m a.s.l.; 1♀ (RMNH), 19 km W Phong My, 23.iii-6.iv.2001, 50- 100m; 1♀ (RMNH), 15 km W Phong My, 23.iii–6.iv. 2001, 150m, Malaise trap, C. v. Achterberg & R. de Vries leg.; Thua Thien-Hue, Nam Dong, Thuong Lo: 1♂ (IEBR), 1-4.ix.2005, Malaise trap, T.Q. Nguyen leg.; Hoa Binh, Xa Linh, Pa Co: 1♂ (IEBR), 23.iv.2002, 1100 m a.s.l., hand net, T.V. Hoang leg.; Bac Can, Ba Be NP: 1♂ (OMNH), 1.v.2006, 20.23 0 N 105.37 0E, 200 m a.s.l., R. Matsumoto leg.
Distribution. Previously known from China (Yunnan Province) (Wang, 1982b), these are the first records of this species from Vietnam.
description: Published as part of Achterberg, Cornelis Van, 2017, First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species, pp. 345-372 in Zootaxa 4311 (3) on pages 368-369, DOI: 10.11646/zootaxa.4311.3.2, http://zenodo.org/record/847854
Yezoceryx walkleyae Momoi 1968
Creators: name: Achterberg, Cornelis Van
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Yezoceryx
Yezoceryx walkleyae

description: Yezoceryx walkleyae Momoi, 1968
(Figure 18)
Yezoceryx walkleyae Momoi, 1968. Kontyû, 36(3): 217. Holotype: ♀, China: Sichuan (USNM).
Diagnosis. Body entirely reddish; first tergite 2.8× its posterior width; first sternite with anteriorly projecting hook; ovipositor about 4.5× as long as hind tibia.
Material examined. Hoa Binh, Mai Chau, Tan Son: 1♀ (IEBR), 1–5.vi.2000, 850– 900 m a.s.l., Malaise trap, L.D. Khuat leg.
Distribution. Previouly known only from China (Sichuan) (Momoi, 1968; Yu et al., 2012), this is the first record of this species from Vietnam.
description: Published as part of Achterberg, Cornelis Van, 2017, First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species, pp. 345-372 in Zootaxa 4311 (3) on page 369, DOI: 10.11646/zootaxa.4311.3.2, http://zenodo.org/record/847854
Palaeoproteus miocenicus
Creators: name: Villa, Andrea
name: Macaluso, Loredana
name: Mörs, Thomas
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Amphibia
Caudata
Batrachosauroididae
Palaeoproteus
Palaeoproteus miocenicus

description: ? Palaeoproteus cf. miocenicus
Figure 5
Material. Hambach 6C: one humerus (IPB-HaH 2390).
Description. This humerus is rather large and robust. It misses only most of the distal epiphysis. The length of the preserved portion is about 16 mm. The bone expands strongly towards the epiphyses. Based on the preserved portion, it appears rather short. The proximal epiphysis is poorly ossified. It has an asymmetrical shape, more expanded toward the ventral side. The posterior surface displays a distinct fossa.
Remarks. The size of this humerus suggests that it belongs to one of the two largest caudate taxa recognised in Hambach (i.e., either cryptobranchids or batrachosauroidids). The bone appears rather short in general appearance, which hints against Chelotriton, cryptobranchids, and possibly also proteids. Its shortness rather recalls the shortened limbs of P. klatti (see Vasilyan and Yanenko, 2020). Furthermore, the general morphology of the bone differs from the fragment of humerus attributed to Mioproteus gardneri Venczel and Codrea, 2018, whereas the poorly ossified proximal end hints against Chelotriton. Therefore, IPB-HaH 2390 is here attributed with doubt to the batrachosauroidid taxon found in the Hambach assemblage, Palaeoproteus cf. miocenicus.
description: Published as part of Villa, Andrea, Macaluso, Loredana & Mörs, Thomas, 2024, Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe, pp. 1-56 in Palaeontologia Electronica (a 3) (a 3) 27 (1) on pages 8-9, DOI: 10.26879/1323, http://zenodo.org/record/10962296
Euronecturus grogu Macaluso, Villa, and Mors 2022
Creators: name: Villa, Andrea
name: Macaluso, Loredana
name: Mörs, Thomas
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Amphibia
Caudata
Proteidae
Euronecturus
Euronecturus grogu

description: Euronecturus grogu Macaluso, Villa, and Mörs, 2022
Material. Hambach 6C: five atlases (IPB-HaH 2119/2121, IPB-HaH 2150, IPB-HaH 2165).
Remarks. These atlases were recently described as types of a new proteid taxon, E. grogu. The reader is referred to Macaluso et al. (2022b) for further information.
description: Published as part of Villa, Andrea, Macaluso, Loredana & Mörs, Thomas, 2024, Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe, pp. 1-56 in Palaeontologia Electronica (a 3) (a 3) 27 (1) on page 9, DOI: 10.26879/1323, http://zenodo.org/record/10962296
Chelotriton undetermined
Creators: name: Villa, Andrea
name: Macaluso, Loredana
name: Mörs, Thomas
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Amphibia
Caudata
Salamandridae
Chelotriton
Chelotriton undetermined

description: ? Chelotriton sp.
Figure 9
Material. Hambach 6C: two humeri (IPB-HaH 2389, IPB-HaH 2393).
Description. These humeri are moderately large and well ossified. IPB-HaH 2389 (Figure 9 A-B) preserves only the proximal half, which is about 9.5 mm long. The humerus head is rather flat proximally. On the ventral side, the ventral crest of the humerus is well developed and robust. It has an almost horizontal distal margin and a rather steep proximal margin. The crest does not bear a distinct sulcus ventrally. On the other side of the bone, the dorsal crest of the humerus is present as a short and pointed process. IPB-HaH 2393 (Figure 9 C-D), on the other hand, is represented only by the distal epiphysis, which is about 6.5 mm in length. It has deep cubital ventral and olecranon fossae. The distal portion of the epiphysis was not coossified with the rest of it.
Remarks. These humeri are tentatively assigned to Chelotriton because of the size and the strong ossification. Proteids and likely batrachosauroidids have poorly ossified long bones, and the two specimens are larger and more ossified than bones of other salamandrids identified in Hambach and not as large as those of adult cryptobranchids.
description: Published as part of Villa, Andrea, Macaluso, Loredana & Mörs, Thomas, 2024, Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe, pp. 1-56 in Palaeontologia Electronica (a 3) (a 3) 27 (1) on page 13, DOI: 10.26879/1323, http://zenodo.org/record/10962296
Palaeobatrachidae Cope 1865
Creators: name: Villa, Andrea
name: Macaluso, Loredana
name: Mörs, Thomas
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Amphibia
Anura
Palaeobatrachidae

description: ? Palaeobatrachidae indet.
Figure 18
Material. Hambach 6C: one maxilla (IPB-HaH 2279).
Description. IPB-HaH 2279 is a fragmentary and robust maxilla, lacking the anterior and posterior portions. The lamina horizontalis is robust and moderately developed in medial direction. On the ventral surface of the lamina, a low longitudinal ridge separates a toothless medial half from the crista dentalis. Teeth were pleurodont, but no one is preserved; they were separated by small processes, which are now eroded. Nine tooth positions are recognizable. In the middle of the dorsal surface of the lamina, a deep and circular recess is visible. By the recess, the lateral wall of the maxilla bends medially. The processus palatinus should be located by this bending, but it is not clearly recognizable in this specimen; this could be either due to a real absence of the processus or, more likely, to preservational reasons. The lateral surface of the bone is smooth.
Remarks. This specimen from Hambach 6C is rather similar to the maxillae of P. eurydices in (Villa et al., 2016): overall robustness; presence of the circular recess (recessus vaginiformis for the processus maxillaris anterior, which is the anterior termination of the subocular bar; Roček, 2003: 1941) on the lamina horizontalis; presence of processes separating the teeth (even though it is not possible to say if they were knob-like in origin). On the ventral surface of the lamina horizontalis, the low ridge delimitating medially the toothed area recalls the maxilla RGM 632039, possibly referred to a young P. eurydices by Villa et al. (2016). However, the Hambach 6C specimen has more teeth than the latter. IPB-HaH 2279 may thus represent a palaeobatrachid, maybe even related to P. eurydices, but in absence of further, better preserved and more taxonomically significant, material from the Miocene level of Hambach, it appears more confident to propose only a cautious identification.
description: Published as part of Villa, Andrea, Macaluso, Loredana & Mörs, Thomas, 2024, Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe, pp. 1-56 in Palaeontologia Electronica (a 3) (a 3) 27 (1) on page 27, DOI: 10.26879/1323, http://zenodo.org/record/10962296
Eopelobates undetermined
Creators: name: Villa, Andrea
name: Macaluso, Loredana
name: Mörs, Thomas
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Amphibia
Anura
Pelobatidae
Eopelobates
Eopelobates undetermined

description: ? Eopelobates sp.
Figure 19 K-O
Material. Hambach 11C: two humeri (IPB-HaR 2414/2415); one ilium (IPB-HaR 2421).
Description. The two humeri (Figure 19 K-M) are represented only by their distal epiphyses. They reach a moderately large size. A spherical eminentia capitata is present, as well as a fossa cubitalis ventralis. The latter is open on the lateral side. Both humeri have a well-developed epicondylus ulnaris and a less developed epicondylus radialis. On the dorsal surface, a wide and elongated olecranon scar is present. Based on the position of the scar, it seems that the eminentia capitata was laterally shifted, but this cannot be confidently evaluated given the absence of the diaphysis.
The ilium (Figure 19 N-O) is very poorly preserved, with only the base of the shaft and the anterior portion of the body remaining. It clearly has no dorsal crest and no dorsal tubercle. The acetabular rim is slightly higher in its anteroventral part. There are no supraacetabular and preacetabular fossae and the spiral groove is not distinct. The dorsal acetabular expansion appears rather short, whereas the development of the ventral one cannot be evaluated due to breakage. The medial surface of the ilial body is flat in its preserved portion. The absence of its posterior part hinders an evaluation of the presence or absence of striae.
Remarks. These elements are here tentatively attributed to Eopelobates because of the overall morphological similarity with those from Hambach 11 referred to this genus, in particular in the indistinct spiral groove on the ilium. However, poor preservation hinders confidence in the attribution.
description: Published as part of Villa, Andrea, Macaluso, Loredana & Mörs, Thomas, 2024, Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe, pp. 1-56 in Palaeontologia Electronica (a 3) (a 3) 27 (1) on page 30, DOI: 10.26879/1323, http://zenodo.org/record/10962296
Pelobatidae Bonaparte 1850
Creators: name: Villa, Andrea
name: Macaluso, Loredana
name: Mörs, Thomas
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Amphibia
Anura
Pelobatidae

description: Pelobatidae indet.
Figure 19 P-U
Material. Hambach 6C: one trunk vertebra (IPB-HaH 2220); one humerus (IPB-HaH 2399).
Discussion. IPB-HaH 2220 (Figure 19 P-S) is a moderately large-sized and procoelous trunk vertebra. It has a cylindrical centrum and a long neural arch, with a low carina neuralis and a well-developed posterior point. Transverse processes are broken off, but they were located under the prezygapophyses.
The humerus (Figure 19 T-U) is small, but preserves part of the diaphysis and part of the epiphysis. The eminentia capitata is shifted laterally. A shallow fossa cubitalis ventralis, which is open on the lateral side, is present. The epicondyles are missing. The cristae medialis and lateralis are not developed. On the dorsal side, the visible part of the olecranon scar is poorly marked.
Remarks. Following Bailon (1999), these two fossils can be assigned to pelobatids. This attribution is based on: vertebra procoelous, with a long neural arch, transverse processes located ventral to the prezygapophyses, a cylindrical centrum, and a well-developed posterior point; humerus with a laterally-open fossa cubitalis ventralis and maybe also lateral shift of the eminentia capitata. The absence of more taxonomically significant elements in the Miocene level of Hambach, however, hinders a discrimination of either Eopelobates or Pelobates.
description: Published as part of Villa, Andrea, Macaluso, Loredana & Mörs, Thomas, 2024, Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe, pp. 1-56 in Palaeontologia Electronica (a 3) (a 3) 27 (1) on page 30, DOI: 10.26879/1323, http://zenodo.org/record/10962296
Hyla undetermined
Creators: name: Villa, Andrea
name: Macaluso, Loredana
name: Mörs, Thomas
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Amphibia
Anura
Hylidae
Hyla
Hyla undetermined

description: Hyla sp.
Figure 20
Material. Hambach 6C: one sacral vertebra (IPB-HaH 2195). Hambach 11: one ilium (IPB-HaR 2088).
Description. The sacral vertebra (Figure 20 A-D) is moderately small and has an anterior cotyle and two posterior condyles. The posterior condyles are well separated. The neural canal is elliptical and the dorsal surface of the neural arch does not present a distinct carina neuralis. The prezygapophyses are suboval and slightly tilted dorsally (about 30°). The transverse processes are broken off, but, judging from the preserved portion of the left one, they seem to have been anteroposteriorly extended.
IPB-HaR 2088 (Figure 20E) is a small ilium with a globular and laterally-bending dorsal tubercle. The dorsal crest is absent and the preacetabular zone is strongly expanded anteroventrally. The posterior portion of the ilial body is broken.
Remarks. Following the diagnostic criteria reported by Bailon (1999), these two fossils can be attributed to a tree frog. The combination of features that is diagnostic for the sacral vertebra is: anterior cotyle; two well-differentiated posterior condyles; extended transverse processes; and no carina neuralis. The diagnostic combination for the ilium is, on the other hand: no dorsal crest; expanded preacetabular zone; and globular and laterally-bending dorsal tubercle. A specific identification of European Hyla species based on isolated bones is considered impossible by most authors (e.g., Bailon, 1999; Sanchiz, 1981; Sanchiz and Młynarski, 1979a; Sanchiz and Sanz, 1980). Nevertheless, it is possible that at least the ilium coming from the most recent level in Hambach pertains to the Hyla arborea (Linnaeus, 1758) species complex, including the extant European species of the genus (Faivovich et al., 2005; Stöck et al., 2008).
description: Published as part of Villa, Andrea, Macaluso, Loredana & Mörs, Thomas, 2024, Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe, pp. 1-56 in Palaeontologia Electronica (a 3) (a 3) 27 (1) on page 30, DOI: 10.26879/1323, http://zenodo.org/record/10962296
Bufo bufo
Creators: name: Villa, Andrea
name: Macaluso, Loredana
name: Mörs, Thomas
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Amphibia
Anura
Bufonidae
Bufo
Bufo bufo

description: BUFO BUFO (Linnaeus, 1758)
Bufo gr. bufo
Figure 21
Material. Hambach 11: one sacral vertebra (IPB-HaR 2020).
Description. IPB-HaR 2020 is a medium-sized sacral vertebra provided with an anterior cotyle and two posterior condyles. The neural canal is subelliptical. A low carina neuralis is present on the dorsal surface of the neural arch, whereas there are no fossettes. Transverse processes are broken and so it is not easy to evaluate their complete anteroposterior extension. Nevertheless, the preserved bases suggest that they were not cylindrical and also not strongly expanded.
Remarks. This sacral vertebra is here attributed to a representative of the common toad species group due to (Bailon, 1999): presence of an anterior cotyle; transverse processes not strongly extended; presence of a carina neuralis; and absence of fossettes on the dorsal surface of the neural arch.
description: Published as part of Villa, Andrea, Macaluso, Loredana & Mörs, Thomas, 2024, Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe, pp. 1-56 in Palaeontologia Electronica (a 3) (a 3) 27 (1) on page 31, DOI: 10.26879/1323, http://zenodo.org/record/10962296
Pelophylax undetermined
Creators: name: Villa, Andrea
name: Macaluso, Loredana
name: Mörs, Thomas
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Amphibia
Anura
Ranidae
Pelophylax
Pelophylax undetermined

description: Pelophylax sp.
Figure 22 A-D
Material. Hambach 6C: two ilia (IPB-HaH 2307, IPB-HaH 2387).
Description. These rather large and moderately preserved ilia have a well-developed and almost vertical dorsal tubercle, which is continued anteriorly by a high dorsal crest. The dorsal acetabular expansion is moderately short. The angle comprised between the latter and the tubercle approaches 90°. A shallow supraacetabular fossa is present. The medial surface of the ilial body is flat, without interiliac groove. The shape of the ilioischiatic juncture cannot be clearly described due to preservational reasons.
Remarks. Identification of these ilia as pertaining to green frogs is supported by (Bailon, 1999): high dorsal crest; well-developed dorsal tubercle; and angle comprised between the tubercle and the dorsal acetabular expansion equals 90°. Specific identification of green frogs is possible only with a population-sized amount of ilia (Blain et al., 2015).
description: Published as part of Villa, Andrea, Macaluso, Loredana & Mörs, Thomas, 2024, Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe, pp. 1-56 in Palaeontologia Electronica (a 3) (a 3) 27 (1) on page 32, DOI: 10.26879/1323, http://zenodo.org/record/10962296
Ranidae Batsch 1796
Creators: name: Villa, Andrea
name: Macaluso, Loredana
name: Mörs, Thomas
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Amphibia
Anura
Ranidae

description: Ranidae indet.
Figure 22 G-N
Material. Hambach 6C: nine sacral vertebrae (IPB-HaH 2193/2194, IPB-HaH 2209/2215); one scapula (IPB-HaH 2323). Hambach 11: one trunk vertebra (IPB-HaR 2184); three sacral vertebrae (IPB-HaR 2018/2019; IPB-HaR 2031); one ilium (IPB-HaR 2087).
Description. IPB-HaR 2184 is a small and amphicoelous trunk vertebra, with a short neural arch and laterally directed transverse processes.
Sacral vertebrae (Figure 22 G-J) are small and provided with one anterior and two posterior condyles. They have cylindrical transverse processes.
IPB-HaH 2323 (Figure 22 K-L) is an elongated scapula. It has no ridges on the anterior margin, but it is provided with a ridge on the inner surface. The cavitas glenoidalis opens in posterior direction and is partially hidden by the pars acromialis in ventral view.
The ilium (Figure 22 M-N) is small and poorly preserved, missing completely the shaft. It has a large and subcircular acetabular fossa. Dorsal to the latter, a deep supraacetabular fossa is present. The dorsal tubercle is also present, even though mostly broken away. Nevertheless, it appears moderately distinct. Anteriorly, it is in continuation with a dorsal crest. Both the dorsal and ventral acetabular expansions are broken, but the former appears well developed based on what is preserved. Medially, there is no interiliac groove or tubercle. The ilioischiatic juncture seems rather low and large, but it is not completely preserved.
Remarks. All these specimens present a combination of features supporting attribution to indeterminate ranids according to the criteria presented by Bailon (1999). For the trunk vertebra, this combination includes amphicoely, the short neural arch, and the laterally directed processes; this further suggests that it represents the eighth vertebra in the column (Bailon, 1999). In the sacral vertebra, the diagnostic combination consists of the anterior condyle, the two posterior condyles, and the cylindrical processes. For the scapula, significant are the elongation, the absence of crista anterior, the cavitas glenoidalis hidden in ventral view, as well as the presence of an inner ridge on scapula. The attribution of the ilium is supported by the presence of the dorsal crest and the absence of interiliac tubercles and grooves. The low and large juncture could hint at green frogs for the ilium (Gleed-Owen, 1998), but it is not clear how much this apparent morphology may be influenced by the preservation.
description: Published as part of Villa, Andrea, Macaluso, Loredana & Mörs, Thomas, 2024, Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe, pp. 1-56 in Palaeontologia Electronica (a 3) (a 3) 27 (1) on page 33, DOI: 10.26879/1323, http://zenodo.org/record/10962296
Rana undetermined
Creators: name: Villa, Andrea
name: Macaluso, Loredana
name: Mörs, Thomas
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Amphibia
Anura
Ranidae
Rana
Rana undetermined

description: Rana sp.
Figure 22 E-F
Material. Hambach 6C: one ilium (IPB-HaH 2321).
Description. IPB-HaH 2321 is moderately preserved. Laterally, a wide and subcircular acetabular fossa is visible. Dorsal to it, there is a shallow supraacetabular fossa. The dorsal tubercle is moderately distinct, low, and anteriorly inclined. Its lateral surface is smooth. The dorsal crest is present, but almost completely broken off. Both dorsal and ventral acetabular expansions are preserved and well distinct, with the latter that is more developed than the former. The dorsal acetabular expansion and the dorsal tubercle define an obtuse angle. The flat medial surface has neither an interiliac tubercle nor a groove. The ilioischiatic juncture is high and narrow.
Remarks. The following combination of features allows referral of this ilium to a brown frog (Gleed-Owen, 1998; Bailon, 1999): presence of a dorsal crest; poorly distinct dorsal tubercle; angle between tubercle and dorsal acetabular expansion obtuse; and ilioischiatic juncture high and narrow.
description: Published as part of Villa, Andrea, Macaluso, Loredana & Mörs, Thomas, 2024, Miocene and Pliocene amphibians from Hambach (Germany): New evidence for a late Neogene refuge in northwestern Europe, pp. 1-56 in Palaeontologia Electronica (a 3) (a 3) 27 (1) on pages 32-33, DOI: 10.26879/1323, http://zenodo.org/record/10962296
Deuterocopus socotranus Rebel 1907
Creators: name: Ustjuzhanin, Petr
name: Kovtunovich, Vasily
name: Anikin, Vasily
name: Aarvik, Leif
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Pterophoridae
Deuterocopus
Deuterocopus socotranus

description: Deuterocopus socotranus Rebel, 1907
Deuterocopus socotranus Rebel, 1907: 115. (Type locality: Yemen, Sokotra). Deuterocopus viticola Meyrick, 1911: 104. (Type locality: Sri Lanka).
Deuterocopus triannulatus Meyrick, 1913: 107. (Type locality: Australia).
Deuterocopus deltoptilus Meyrick, 1930: 565. (Type locality: Uganda).
Deuterocopus henrioti Bigot & Boireau, 2006: 16 –17. (Type locality: Cote d'Ivoire).
Material examined. 1 Ƌ1 ♀, Western Region, Kibale Distr., Kibale Biol. Field. Station of Maker SU N 0033', W 03021', 1511 m, 19–24.x.2014, V. Anikin.
Distribution. Yemen, Australia, Ivory Coast, Malawi, Mozambique, Republic of South Africa, Somalia, Uganda, Zambia, Zimbabwe (De Prins & De Prins 2015); Oman, Japan, India, Sri Lanka, Thailand, Taiwan, Burma, Indonesia, New Guinea, Australia.
description: Published as part of Ustjuzhanin, Petr, Kovtunovich, Vasily, Anikin, Vasily & Aarvik, Leif, 2016, Additions to the fauna of plume moths (Lepidoptera, Pterophoridae) of Uganda, pp. 393-402 in Zootaxa 4132 (3) on page 394, DOI: 10.11646/zootaxa.4132.3.7, http://zenodo.org/record/265451
Walsinghamiella niniella Gielis 2011
Creators: name: Ustjuzhanin, Petr
name: Kovtunovich, Vasily
name: Anikin, Vasily
name: Aarvik, Leif
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Pterophoridae
Walsinghamiella
Walsinghamiella niniella

description: Walsinghamiella niniella Gielis, 2011
Walsinghamiella niniella Gielis, 2011: 41. (Type locality: Tanzania).
Material examined. 1 ♀, Western Region, Kibale Distr., Kibale Biol. Field. Station of Maker SU N 0033', W 03021', 1511 m, 19–24.x.2014, V. Anikin.
Distribution. Tanzania, Malawi, Uganda (De Prins & De Prins 2015). Notes. New to Uganda.
description: Published as part of Ustjuzhanin, Petr, Kovtunovich, Vasily, Anikin, Vasily & Aarvik, Leif, 2016, Additions to the fauna of plume moths (Lepidoptera, Pterophoridae) of Uganda, pp. 393-402 in Zootaxa 4132 (3) on page 394, DOI: 10.11646/zootaxa.4132.3.7, http://zenodo.org/record/265451