Ulivellia pseudinsolita Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev 2024, sp. nov.
Creators: name: Kovac, Damir
name: Kameneva, Elena P.
name: Korneyev, Severyn V.
name: Araújo, Alexandre Santos
name: Savaris, Marcoandre
name: Smit, John T.
name: Schneider, Alexander
name: Schreiber, Robert
name: Korneyev, Valery A.
Publisher: name: Zenodo
Year: 2024
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Ulidiidae
Ulivellia
Ulivellia pseudinsolita

description: Ulivellia pseudinsolita Kameneva & V. Korneyev sp. nov.
urn:lsid:zoobank.org:act: F2BE9A18-8FAA-4FB2-9D3C-37489BFC0066
Figs 4 G, 55–56
Material examined. Type. Holotype ♂: Brazil: “ Corupa [Corupá], S. Cath.[Santa Catarina] / (Hansa Humbolt) / II 1945 ”, “ A. Maler, Coll. / Frank Johnson / Donor” (abdomen dissected and attached in a plastic tube with glycerin under specimen) (AMNH).
Diagnosis. Ulivellia pseudinsolita differs from other species of the genus Ulivellia in having a wing pattern with oblique discal crossband, prescutellar acrostichal setae present, halter and legs entirely black; it most closely resembles U. tenoris in having wing pattern with four narrow brown crossbands, of which discal and preapical are oblique and clearly diverging posteriorly, differing from it by the wider wing and the hyaline interspace between discal and preapical crossbands narrower, colouration of legs and halters and presence / absence of acrostichal setae (see the key). Both species have a wing pattern similar to that of Euxesta insolita Hendel 1909 (Fig. 4 I), clearly differing by the at least partly hyaline or yellow wing base (in E. insolita completely black); as well as by the almost twice larger size (in E. insolita wing length = 3.0– 3.5 mm), and the epistome / clypeus sizes.
Description. Male. Head (Figs 55 C–D) ratio (length: height: width) = 1: 1.4: 1.7, frons, parafacial, and gena yellowish brown, face and occiput black. Frons shiny brown, narrowed posteriorly (Fig. 55 C), 1.2× as long (from the lunule to inner vertical seta) as wide at lunule and 1.5× longer than wide at vertex or 0.85× as long (from the lunule to anterior ocellus) as wide at lunule and 1.1× longer than wide at vertex, with metallic shining, brown or black ocellar triangle and vertical plates; orbits, vertex, gena and occiput silver-white microtrichose; parafacial narrow, 0.3× broader than postpedicel, sparsely white microtrichose. Frontal plates with 6–8 proclinate parafrontal setulae and 6–7 moderately strong, frontal setae and the frontal vitta with 3 interfrontal setae on each side, anteriormost crossing, longer than other setulae and half as long as outer vertical seta; 1 very long orbital seta 1.2× longer than ocellar seta, 0.8× longer than inner vertical seta and twice as long as outer vertical seta and 1 reclinate anterior orbital seta as long as longest frontal setae; postocellar seta long, 0.7× longer than inner vertical seta; outer vertical seta short, 0.4× longer than inner vertical seta and twice as long as postocular setae forming one row, allied by 15–20 dorsal and 18–25 ventral occipital setae. Lunule indistinguishable. Eye 1.6× higher than long. Face above transverse fold 0.6× higher than wide at fold, brown, white microtrichose; epistome entirely black or dark brown, sparsely white microtrichose, with metallic cyan sheen, strongly produced anteriorly. Clypeus shiny brown, sparsely white microtrichose, 0.3× higher than epistome. Gena brownish yellow, with long genal seta and 4–5 peristomal setae anterior of it, 0.4–0.5× longer than genal seta. Occiput black, except postgena partly brown, with cyan metallic sheen, greyish microtrichose; ventral part of occiput (postgena) with 5–7 setae almost as long as genal seta.
Antenna brown; scape and pedicel with black setulae; postpedicel brown, whitish microtrichose, twice as long as wide, apically rounded; arista black to brown at base, almost bare. Mouthparts black, prementum black, with steel sheen. Palp brown, moderately wide crescent, apically rounded, with 9–12 long black setulae, of them medial setulae longer than others.
Thorax (Fig. 55 E) brown to black, with bluish sheen and sparse white microtrichia not obscuring underlying cuticle. Mesonotal scutum 1.1×as long as wide; with brown microtrichia posterolateral to transverse suture, black setulose, with 12–16 rows of setulae between rows of dorsocentral setulae; acrostichal prescutellar setae present; prescutellar area setulose behind anterior dorsocentral setae to acrostichal seta. Scutellum dorsally very slightly convex, black, almost matt, sparsely microtrichose, without setulae, laterally with bluish sheen. Subscutellum and mediotergite black, grey microtrichose. Other setae as described for the genus. All the setae and setulae black.
Wing (Figs 4 G, 55 A, B, F) 5.5 mm (♂) long, 3× longer than wide; basicostal cell hyaline or yellowish; costal cell 5× longer than wide, slightly widened apically, brown in basal half, with costa very slightly curved posteriorly, but forming neither lobe nor cleft before apex of vein Sc; pterostigma entirely brown, narrow triangular, 4.4× longer than wide, vein R 1 with 15–17 setulae dorsal to Sc apex, terminating at level of crossvein dm-m; vein R 2+3 weakly arcuate, subparallel to costa, turning towards it before very apex. Crossvein r-m distal to center of cell dm. Cell r 4+5 5.8× longer than wide, widened distally of crossvein dm-m, narrowed apically. Cell m 1 narrowly triangular; the ultimate section of M 1 3.7× longer than crossvein dm-m and 2.8× longer than penultimate section, markedly deepened posteriorly at middle. Vein CuA Z-shaped, forming moderately long posteroapical lobe of cell cua along vein CuP, 1.3× longer than its anterior shoulder. Wing pattern with subbasal crossband from basal half of costal cell through cell cua into alula, wide oblique brown discal crossband from pterostigma to base of cell m 4 except its very base and apex; narrow and oblique preapical crossband from cell r 1 through crossvein dm-m merging with apical crossband in cell r 1 reaching apex of cell m 1; hyaline interspace between discal and preapical crossband 1.3–1.6 as wide as discal crossband in cell dm (Fig. 4 G, 55 A). Cell cup and anal lobe pale grey, narrower than cell cua. Alula 3× longer than wide, grey in apical 0.5–0.6. Calypters white, with white cilia. Halter dark brown to black.
Legs (Fig. 55 A) with yellow or brownish yellow coxae and trochanters, femora and tibiae entirely black, except coxae and knees narrowly brown, with bluish sheen; tarsi entirely brownish yellow, black setose and setulose. Fore femur with 2 rows of posterodorsal and posterior setae slightly shorter than femur width, and 4–5 strong posteroventral setae in apical half. Mid femur anteriorly with row of short suberect setulae; mid tibia with one strong medioventral seta 2.1× longer than tibia width.
Abdomen moderately elongate (Figs 55 A, B, 56 F), black, sparsely whitish microtrichose, with cyan metallic sheen. Tergite 5 not microtrichose, somewhat shagreened, dull.
Cerci wide oval, moderately developed, partly fused at bases elongate, sack-like; setulose (Figs 56 A–D). Epandrium ovoid, dorsally short, moderately setulose; surstyli basally wide, outer surstylus flattened, posteroventrally directed, apically oval with single trichoid sensillum; inner surstylus with four very strong claw-like mesally directed prensisetae (Figs 56 A, C, D). Pregonites symmetrical, almost flat, with 5 moderately developed setulae ventro-mesally, conspicuously anterior to postgonites (Figs 56 C, E). Postgonites on laterobasal part of the phallic guide lobes, with 4–5 small trichoid sensilla (Figs 56 C, E). Phallus moderately short and narrow, about 2.5–3× longer than epandrium high, thin and bare (Figs 56 A–E).
Female unknown.
Etymology. The species name is an adjective and is composed of the Greek “pseud-” (“false”) and the species name “insolita ”, reflecting the superficial similarity of the wing pattern to Euxesta insolita Hendel, 1909.
description: Published as part of Kovac, Damir, Kameneva, Elena P., Korneyev, Severyn V., Araújo, Alexandre Santos, Savaris, Marcoandre, Smit, John T., Schneider, Alexander, Schreiber, Robert & Korneyev, Valery A., 2024, Revision of the Aspistomella group of genera (Diptera: Ulidiidae: Pterocallinae: Lipsanini), pp. 1-117 in Zootaxa 5530 (1) on pages 98-102, DOI: 10.11646/zootaxa.5530.1.1, http://zenodo.org/record/14023149