Tylopus punctus Likhitrakarn & Golovatch & Panha 2016, sp. nov.
Creators: name: Likhitrakarn, Natdanai
name: Golovatch, Sergei I.
name: Panha, Somsak
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Diplopoda
Polydesmida
Paradoxosomatidae
Tylopus
Tylopus punctus

description: Tylopus punctus sp. nov.
urn:lsid:zoobank.org:act: 35585ACA-8490-49EE-8DA5-73EB3EB04A5C
Figs 21–22, 26
Diagnosis
Very similar to T. silvestris (Pocock, 1895), but differs in the smaller size, the width of midbody being 2.1–2.5 mm (versus 3.0 mm); the caudal corner of midbody paraterga very narrowly rounded and not drawn behind the rear tergal margin (versus nearly pointed and produced behind the rear tergal margin), and gonopod process h small and pointed (versus absent).
Etymology
To emphasize gonopod process h being small and pointed; adjective.
Material examined
Holotype
MYANMAR: ♂, Mintaingbin Forest Camp, ca 35 km north of Aungban, Chan State, 20°55'20" N, 96°33'60" E, ca 1300 m a.s.l., 11–23 Jun. 2004, leg. H. Schillhammer & H.V. Shaverdo (NHMW- 8248).
Paratype
MYANMAR: 1 ♀, same data as for holotype (NHMW- 8249).
Description
MEASUREMENTS AND COLOUR. Length 27.5 (♂) or 29 mm (♀), width of midbody pro- and metazonae 1.56 and 2.1 mm (♂) or 1.98 and 2.5 mm (♀), respectively. Coloration of alcohol material upon long-term preservation rather uniformly light brown to brown, with light yellow paraterga, antennae, venter and legs (Fig. 21 A–I).
HEAD. Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae rather short (Fig. 21B), reaching only in front of body segment 3 (♂, ♀) when stretched dorsally. In width, segment 3 <4 <head <collum <segment 2 <5–15; thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 4+4 anterior, 2+2 intermediate and 3+3 posterior; an evident lateral incision at about anterior 1/3; caudal corner of paraterga very broadly rounded, declined ventrad, not drawn behind rear tergal margin.
BODY. Tegument smooth and shining, prozonae very finely shagreened, metaterga rather smooth and leathery, with posterior halves faintly rugulose, surface below paraterga finely microgranulate (Fig. 21 A–F). Postcollum metaterga with two transverse rows of setae: 2+ 2 in anterior (pre-sulcus) row, mostly abraded, and 2+ 2 in posterior (post-sulcus) row, mostly traceable as insertion points, except for segments 18 and 19, each latter with 4+4 setae in posterior row. Tergal setae simple, slender, about 1/3 of metatergal length (Fig. 21A, F). Axial line visible both on pro- and metazonae. Paraterga well-developed (Fig. 21 A–G), especially so in ♂, set high (at upper ¼ of body height), anterior edge rounded and narrowly bordered, fused to callus; caudal corner very narrowly rounded, segments 2 and 3 slightly extending behind rear tergal margin, thereafter not surpassing rear tergal margin until segment 15 (Fig. 21 A–D). Paratega 2 broad, anterior edge angular, lateral edge with two small, but evident incisions in anterior half. Following paraterga with an evident incision in anterior 1/3 of lateral edge (Fig. 21A, C, F). Calluses on paraterga delimited by a sulcus only dorsally, rather narrow, a little wider on pore-bearing segments. Ozopores distinct, lying at about ¼ of metatergal length in front of posterior edge of metaterga. Transverse sulcus usually distinct (Fig. 21A, D, F), slightly incomplete on segment 19, complete on segments 5–18, narrow, line-shaped, rather deep, reaching bases of paraterga, beaded at bottom. Stricture between pro- and metazonae evident, rather wide and deep, beaded at bottom down to base of paraterga (Fig. 21A, C–F). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2 and 3, thereafter increasingly reduced and remaining a sharp caudal tooth until segment 9 (♂) or 6 (♀), retained as a small caudal tooth and increasingly reduced until segment 17, absent from segment 18 (♂) or 7 (♀). Epiproct (Fig. 21 E–G) conical, flattened dorsoventrally, with two evident, rather large, rounded, apical papillae; tip subtruncate; lateral pre-apical papillae clear, lying close to tip. Hypoproct roundly subtriangular, setigerous knobs at caudal edge small and well-separated (Fig. 21G).
STERNA. Sparsely setose, without modifications, but with a pair of small, blunt, fully separated cones between ♂ coxae 4 (Fig. 21H, I). Legs rather short and slender, midbody ones ca 1.1–1.2 (♂) or 0.8–0.9 (♀) as long as body height, ♂ prefemora not swollen, ♂ tarsal brushes present only on legs 1 and 2.
GONOPOD. Rather simple (Fig. 22); coxa slightly curved caudad, sparsely setose distoventrally. Prefemur sparsely setose, about 1/3 as long as femorite + postfemoral part. Femorite curved and stout, with an evident mesal groove and a strong distolateral sulcus demarcating a postfemoral part; process h small, slender and pointed; solenophore (sph) clearly coiled, long and flattened.
Remark
Pocock (1895) described only T. doriae (Pocock, 1895) and T. silvestris from northern Myanmar, the above being a third species of this genus also occurring in northern Myanmar.
New faunistic record
description: Published as part of Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak, 2016, The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina, pp. 1-47 in European Journal of Taxonomy 195 on pages 29-32, DOI: 10.5852/ejt.2016.195, http://zenodo.org/record/3837846