Promalactis denticuligera Wang & Liu 2021, sp. nov.
Creators: name: Wang, Shuxia
name: Liu, Chen
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Lepidoptera
Oecophoridae
Promalactis
Promalactis denticuligera

description: Promalactis denticuligera Wang, sp. nov.
(Figs 10, 32, 49)
Type material. CHINA, Tibet: Holotype ♂, Tongmai Town (30.10°N, 95.08°E), 2029 m, 12.VIII.2018, leg. MJ Qi, slide No. LC 19310. Paratypes (23♂, 55♀): 6♂, 39♀, same data as holotype, except 12‒16.VIII.2019, slide No. LC 19311 ♀; 14♂, 5♀, same data as holotype, except 23‒26. VI.2009, leg. MJ Qi & JQ Deng, slide Nos. LC 19312 ♂, LC19313 ♀; 2♀, Yadong Village, Motuo County, 833 m, 16.VIII.2017, leg. XF Yang & MJ Qi, slide No. JYY17327; 1♂, 9♀, Pailong Town, Linzhi County, 2031 m, 17‒18.VIII.2018, leg. MJ Qi, slide Nos. LC 19326 ♂, LC19331 ♀; 1♂, Pailong Town, Linzhi County, 2010 m, 6.VII.2013, leg. JW Li, slide No. LC 19353; 1♂, Linzhi Town, Linzhi City, 2985 m, 19. VI.2019, leg. MJ Qi & JQ Deng, slide No. LC 19316.
Diagnosis. The new species can be distinguished from other members of the group by two features of the male genitalia: the asymmetrically bilobed valvae, and the costa of the left valva produced into a free dorsodistal lobe bearing a cluster of spines along the costal margin.
Description. Adult (Fig. 10). Forewing length 7.0‒9.0 mm.
Head: Frons greyish brown, vertex cream white, occiput rust brown. Labial palpus with second palpomere rust brown on outer surface, ochreous yellow on inner surface; third palpomere black on outer surface, rust brown on inner surface except white apically. Antenna with scape white dorsally, pale grey ventrally; flagellum black annulated with white on dorsal surface, black on ventral surface.
Thorax: Dorsum of mesothorax and tegula rust brown. Forewing ochreous brown on basal 2/3, ochreous yellow on distal 1/3; markings white edged with black scales: costal spot triangular, from distal 1/3 narrowed to above posterior angle of cell; basal streak slender, straight, oblique inward to base of fold; antemedian streak from dorsal 1/3 to anterior margin of cell, parallel with basal streak; dorsal streak from basal 2/3 extending obliquely outward to distal 1/4 of posterior margin of cell; tornal spot black, narrowly diffused inward to end of dorsal streak; distal patch black, extending from apex along termen to beyond tornal spot, gradually narrowed; apical spot sub-oval, small, set in black distal patch; fringe ochreous yellow except grey before tornus. Hindwing and fringe grey. Fore- and midlegs black, except fore femur white and midleg yellowish brown on ventral surface, tibiae with white spot at base and middle, with a tuft of white scales at apex, basal tarsomere white at base and apex, second tarsomere white apically; hindleg yellowish brown, tibia with long grey hairs, basal tarsomere white apically, second and third tarsomeres black except white distally, fourth and fifth tarsomeres white.
Abdomen: Male genitalia (Fig. 32). Uncus wide parallel-sided from base to about basal 2/5, then slightly narrowed to rounded apex. Gnathos longer than uncus; mesial plate triangular, wide at base, narrowed to pointed apex, granulate distally; basal arms wide basally, narrowed to mesial plate. Tegumen with height more than three times length of uncus; lateral arms widened and arched on inner margin near apex. Valvae narrow at base, widened toward apex, asymmetrically bilobed distally; left valva: dorsal lobe slender, slightly narrowed to apex, ventral lobe wide at base, narrowed to basal 2/3, then slender to rounded apex, setose, shorter than dorsal lobe; costa very narrow, with clustered spines along costal margin in distal half; sacculus broadly banded, almost uniform in width, with a large semicircular concavity dorsally before free distal process; right valva: costa narrow, produced to a free digitate dorsal lobe distally, ventral lobe much larger and longer than dorsa lobe, elongate sub-triangular, with a flat apex; sacculus narrowly banded, wide at base, narrowed to basal 2/3, fused with valva and setose in distal 1/3. Saccus longer than uncus, elongate triangular, wide at base, narrowed to rounded apex. Juxta a large sclerotized plate, reaching posterior 1/4 of tegumen, narrow basally, widened distally, rounded apically; basal lobe clubbed, reaching middle of saccus. Aedeagus straight, almost as long as left valva, tubular, distal 1/3 spatulate, denticulate laterally and apically. Cornutus absent.
Female genitalia (Fig. 49). Apophyses posteriores about 1/3 length of apophyses anteriores. Eighth sternum sub-rectangular, with a notch at middle on posterior margin.Antrum heavily sclerotized, slightly narrowed to ductus bursae. Ductus bursae membranous, slightly inflated, less than half length of antrum, with a coiled belt posteriorly. Corpus bursae slightly longer than ductus bursae, membranous, ovate; signum sub-ovate, with dense, large and small denticles.
Distribution. China (Tibet).
Etymology. The specific epithet is derived from the Latin denticuliger, referring to the distally and apically toothed aedeagus.
description: Published as part of Wang, Shuxia & Liu, Chen, 2021, Taxonomic study of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) X. The commotica species-group, with descriptions of twenty-two new species, pp. 293-330 in Zootaxa 4980 (2) on pages 311-312, DOI: 10.11646/zootaxa.4980.2.4, http://zenodo.org/record/4889068