Belisana zham Zhu & Yao & Zheng & Li 2020, sp. nov.
Creators: name: Zhu, Wenhui
name: Yao, Zhiyuan
name: Zheng, Guo
name: Li, Shuqiang
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Pholcidae
Belisana
Belisana zham

description: Belisana zham Yao & Li sp. nov.   
Figs 9–10, 11 G–H  
 Type material. Holotype: Male (IZCAS Ar39813), Zham Town (27°59 ꞌ 01.05 ꞌꞌ N, 85°58 ꞌ 58.32 ꞌꞌ E, elevation 2450 ± 3 m), Nyalam County, Xigaze, Tibet, China, 29 July 2014, Y. Li leg. Paratypes: 2 males (IZCAS Ar39814–39815) and 4 females (IZCAS Ar39816–39819), same data as holotype.   
 Etymology. The specific name refers to the type locality and is a noun in apposition.   
 Diagnosis. This species resembles B. xigaze sp. nov. (Figs 7–8, 11 E–F) and B. yadongensis (Hu, 1985) (see Zhang et al. 2006: 203, figs 41–50) with similar male chelicerae (Fig. 10D), but can be distinguished by procursus that is not protruding dorso-subdistally (Figs 9 A–B), by absence of retrolateral membranous flap on procursus (Fig. 9D), by presence of large, triangular, disto-retrolateral membranous lamella on procursus (arrow in Fig. 9D) and by having slender bulbal apophysis (Fig. 10C). It can also be distinguished from B. xigaze sp. nov. by absence of black lateral margins but presence of pair of large brown marks on carapace (Figs 10 E–F). It differs from B. yadongensis by having pore plates that are nearly elliptic in shape (Figs 10B, 11H).   
 Description. Male (holotype, IZCAS Ar39813): Total length 2.25 (2.38 with clypeus), carapace 1.02 long, 0.80 wide, opisthosoma 1.23 long, 1.01 wide. Leg I: 11.90 (2.75 + 0.36 + 2.78 + 4.38 + 1.63), leg II: 8.02 (2.19 + 0.29 + 2.13 + 2.97 + 0.44), leg III: 4.98 (1.47 + 0.13 + 1.25 + 1.72 + 0.41), leg IV: 6.85 (2.03 + 0.25 + 1.41 + 2.75 + 0.41); tibia I L/d: 56. Habitus as in Figs 10 E–F. Carapace pale, with pair of large brown marks; ocular area with brown stripe medially; sternum pale, without marks. Legs pale, without darker rings. Opisthosoma pale, with black spots dorsally and laterally. Distance PME-PME 0.11, diameter PME 0.06, distance PME-ALE 0.04, AME absent. Ocular area not elevated. Thoracic furrow absent. Clypeus unmodified. Sternum slightly wider than long (0.75/0.68). Chelicerae (Fig. 10D) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses each with hardened and pointed tip (distance between tips: 0.46). Pedipalps as in Figs 9 A–B; coxa unmodified; trochanter with short retrolatero-ventral apophysis; femur without proximo-dorsal apophysis; procursus (Figs 9 A–D) simple proximally and complex distally, with sclerites, spine, and large, triangular, disto-retrolateral membranous lamella (arrow in Fig. 9D); bulb (Fig. 10C) with hooked apophysis and simple embolus. Retrolateral trichobothria of tibia I at 8% proximally; legs with short vertical setae on metatarsi, without spines or curved setae; tarsus I with 24 distinct pseudosegments.   
 Female (IZCAS Ar39816): Similar to male, habitus as in Figs 10 G–H. Total length 2.31 (2.38 with clypeus), carapace 0.82 long, 0.69 wide, opisthosoma 1.49 long, 1.41 wide. Tibia I: 4.45; tibia I L/d: 31. Distance PME-PME 0.10, diameter PME 0.05, distance PME-ALE 0.05, AME absent. Sternum slightly wider than long (0.58/0.45). Carapace only with dark radiated lines; ocular area without median stripe. External female genitalia (Figs 10A, 11G) simple and flat, with pair of pockets 0.38 apart (ep in Figs 10 A–B, 11G–H). Vulva (Figs 10B, 11H) with roof-shaped anterior arch and pair of small, nearly elliptic pore plates.   
 Variation: Leg I missing in male paratypes (IZCAS Ar39814–39815). Tibia I in two female paratypes (IZCAS Ar39817–39818; leg I missing in the other female paratype IZCAS Ar39819): 4.17, 4.55.   
 Natural history. The species was found on the underside of rocks.   
 Distribution. China (Tibet, type locality only; Fig. 12).
description: Published as part of Zhu, Wenhui, Yao, Zhiyuan, Zheng, Guo & Li, Shuqiang, 2020, The Belisana spiders (Araneae: Pholcidae) from Tibet, China, pp. 111-128 in Zootaxa 4802 (1) on pages 121-127, DOI: 10.11646/zootaxa.4802.1.7, http://zenodo.org/record/3904926