Sanaungulus ypogaeum Fanti & Müller, 2022, sp. nov.
Creators: name: Fanti, Fabrizio
name: Müller, Patrick
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Cantharidae
Sanaungulus
Sanaungulus ypogaeum

description: Sanaungulus ypogaeum sp. nov.   
(Fig. 22)  
 Holotype. Female, adult specimen in a Burmese (Kachin) amber piece: Catalog number SNSB­BSPG 2021 XII 67 in the Bayerische Staatssammlung für Paläontologie und Geologie (ex Müller’s collection number: BUB4165).   
 Type locality. Myanmar: Kachin state, Myitkyina District, Hukawng Valley.   
 Type horizon. Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.   
 Differential diagnosis. Sanaungulus ypogaeum sp. nov. differs from other species of Sanaungulus with filiform antennae: Sanaungulus ruicheni (Hsiao & Huang, 2018), Sanaungulus morellii Fanti & Damgaard, 2020, Sanaungulus rosenzweigi Fanti & Damgaard, 2020, and the species described here based on its pronotum that is strongly narrow anteriorly and with a rounded anterior margin (Hsiao & Huang 2018; Fanti & Damgaard 2020).   
 Description. Adult, winged, slender. Female, defined on the shape of the last urites and on the basis of the short antennae. Body length 4.9 mm. Head and pronotum testaceous­brown, elytra and legs brown.   
Head elongated, very narrow, strongly constricted (triangular­shaped) behind the eyes, fitted with rugosity and impressed punctuation. Eyes prominent, very large, roundish, inserted in the upper and dorsolateral part of the head, inter­ocular dorsal distance about 0.6 times greater than the eye diameter. Mandibles elongated, robust, falciform, without teeth. Maxillary palps 4­ segmented with the last palpomere strongly securiform. Labial palps 3­ segmented, with the last palpomere slightly securiform.Antennae 11­ segmented, filiform, robust, short and slightly surpassing the humeral zone of the elytra, antennal insertion in the eyes proximity; scape elongated, robust, club­shaped; antennomere II very short, globular; antennomere III short, thinner and longer than second; antennomere IV elongated, the longest; antennomeres V–VII about 0.2 times shorter than antennomere IV; antennomeres VIII–X shorter than previous ones; antennomere XI filiform, longer than previous ones, rounded at apex; all the antennomeres with short pubescence. Pronotum transverse, wider than head, narrow anteriorly, anterior margin rounded and very slightly bordered, posterior margin almost straight, sides rounded, posterior corners obtuse, pronotum disc smooth. Scutellum triangular­shaped. Elytra short which reveals various abdominal segments, at humeri wider than pronotum and as wide as pronotum at the apex, humeral zone enlarged, posteriorly parallel­sided, rounded apexes, surface sligthly pubescent and with a very small rugosity. Metathoracic wings almost completely covered by elytra. Sternum elongated with posterior margin rounded, sternites narrow and transverse, last tergite deeply incised apically and divided in two lobes, last sternite small and rounded. Legs thin, relatively long, pubescent; coxae massive; trochanters elongated, with rounded apex; femora nearly straight, more robust than tibiae, cylindrical; tibiae flattened, thin, without spurs near the apex, protibiae slightly longer than profemora, metatibiae very longer than metafemora. Tarsal formula 5­ 5­ 5; first tarsomere long, about 2.0 times longer than second; third tarsomere short and bilobed; fourth strongly bilobed; fifth tarsomere elongated, thin, slightly flat and curved; claws simple, without denticle at the base.  
 Etymology. Derived from the Latin noun “ ypogaeum (hypog Á um)” = hypogeum, tomb. In reference to the fact that the amber that has incorporated the specimen is its tomb. Specific epithet is to be treated as noun in apposition.   
 Syninclusions. Air bubbles, detritus.   
 Remarks. Piece of clear, golden amber in Cabochon cut 18x14x 2.8 mm. The inclusion is complete.
description: Published as part of Fanti, Fabrizio & Müller, Patrick, 2022, Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information, pp. 331-380 in Baltic Journal of Coleopterology 22 (2) on pages 370-372, DOI: 10.5281/zenodo.13203160