Braunus ocellatus Thomas 2001, comb. nov.
Creators: name: Barão, Kim Ribeiro
name: Garbelotto, Thereza De Almeida
name: Campos, Luiz Alexandre
name: Grazia, Jocelia
Publisher: name: Zenodo
Year: 2016
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Pentatomidae
Braunus
Braunus ocellatus

description: Braunus ocellatus (Thomas, 2001) comb. nov.
(Figures 11, 54–79, 81–82)
Lojus ocellatus Thomas, 2001: 854 –857, figs 1–4.
Material examined. Holotype ♂, ECUADOR, Napo, Cabañas San Isidro, 2 km NW Cosanga, 0°40’S 77°55’W, 22–30.I.1996, T.J. Henry leg. (USNM). Paratype ♀, same label data as holotype (DBTC). Paratype ♂, ECUADOR, Napo, CabañasSan Isidro, 2 km NW Cosanga, 0°33’S 77°55’W, 20–23.VII.1998, 2150 m, premontaine rain forest, Ratcliffe, Jameson, Smith and Villatoro leg. (UNSM). Additional material examined. 3 ♂ 3 ♀, ECUADOR, Napo, Yanayacu Biological Station, 2144 m, 0.60°S 77.89°W, 24–30.XI.2009, D. Forero leg. [EC09_L1] (1♂ 1♀ UCRC; 1♂ 1♀ USNM; 1♂ 1♀ MNRJ); 1♀, ECUADOR, Napo, 1.3 km SW of Baeza, 2050 m, 00°28’34”S 77°53’47”W, 11.XI.2006, J. Skuhrovec leg. (NMPC).
Diagnosis. Body color light brown (Figs 54–55). Anterior angles of pronotum each with obtuse projection; each anterolateral margin with three small spines, not evenly distributed; humeral angles quadrate, each with bilobed apex, anterior lobe with 1–2 spines and posterior lobe with a digitoid process; posterior margin weakly concave (Fig. 11). Apex of scutellum reaching abdominal segment V. Apex of hemelytral radial vein calloused; corium reaching abdominal segment VI; membrane reaching anterior margin of abdominal segment VII on females (Fig. 57) and surpassing abdominal apex on males (Fig. 56). Male genitalia: posterolateral angles obtuse; inferior layer of ventral rim emarginate with one projection medially (Figs 60–65). Female genitalia: posterior margin of each gonocoxite 8 concave; posterior margin of each laterotergite 8 acute; lateral margin of each laterotergite 9 straight (Figs 74–75).
Male genitalia. Dorsal rim sinuous (Figs 60–61; dr). Posterolateral angles obtuse (Figs 60–61; pa). Inferior layer of ventral rim with median projection (Figs 64–65; vr); superior layer of ventral rim depressed medially (Figs 62–63; sl). Segment X rectangular (Figs 62–63, 67; X). Parameres F-shaped, apex acute dorsally and ventrally; basal process rounded bearing setal tuft (Figs 60–63, 66; par). Phallus: limit of sclerotized and membranous area of phallotheca straight (Figs 70–71; ph). Ventral conjunctival processes each curved toward vesica apically, dorsal surface concave (Figs 68–73; vcp). Each laterodorsal conjunctival process digitiform, sclerotized, longer than vesica (Figs 68–73; lcp). Vesica cup-like, strongly projected dorsally, with dorsal margin uniformly concave (Figs 70–73; v). Ductus seminis distalis ventrally curved, originating in central part of vesica (Figs 70–73; ds). Process of gonopore absent.
Female genitalia. Gonocoxites 8 longer than wide; sutural margins juxtaposed at 1/3 basal portion, each widely separated and sinuous on the remaining 2/3; posterior margins concave; each sutural margin with fringe of setae covering gonapophyses 8 (Figs 74–75; gc8). Gonapophyses 8 visible, carinated medially (Figs 74–75; g8). Laterotergites 8 triangular, longer than wide; each posterior margin acute surpassing tergite 8; each spiracle placed posterior to lateral angle of each gonocoxite 8 (Figs 74–75; la8). Gonocoxites 9 trapezoidal, each with 1+1 tufts of long setae on disc (Figs 74–77; gc9). Each laterotergite 9 digitiform, depressed basally; apex rounded and surpassing tergite 8 (Figs 74–75; la9). Segment X rectangular (Figs 74–75; X). Gonapophyses 9 each 1+1 sclerotized areas surrounding chitinellipsen (Figs 76–77; ch) and 1+1 lateral folds reaching posterior margin of chitinellipsen (Figs 76–77; ch). Chitinellipsen circular (Figs 76–77; ch). Thickening of vaginal intima tube-like (Figs 76–77; t vi). Median duct of vesicular area with a sclerotized ring at base (Figs 76–77; md). Capsula seminalis with three processes (Figs 76–77; cs).
Measurements. Total length, 8.5 ± 0.37 (7.7–9.5); head length, 2.2 ± 0.11 (1.9–2.5); head width, 1.4 ± 0.04 (1.3–1.5); length of pronotum, 1.7 ± 0.05 (1.6–1.9); width of pronotum at base, 3.9 ± 0.16 (3.5–4.3); maximum width of pronotum at humeral angle, 5.4 ± 0.20 (4.9–6.0); length of scutellum, 2.6 ± 0.15 (2.3–3.0); width of scutellum, 2.5 ± 0.10 (2.5–2.8); width of abdomen across third segment, 4.6 ± 0.21 (4.2–5.4).
Distribution. Ecuador (Napo) (Figs 78–79, 81–82).
description: Published as part of Barão, Kim Ribeiro, Garbelotto, Thereza De Almeida, Campos, Luiz Alexandre & Grazia, Jocelia, 2016, Unusual looking pentatomids: reassessing the taxonomy of Braunus Distant and Lojus McDonald (Hemiptera: Heteroptera: Pentatomidae), pp. 168-186 in Zootaxa 4078 (1) on pages 181-182, DOI: 10.11646/zootaxa.4078.1.16, http://zenodo.org/record/261493