Engistanoxia sagala Sehnal 2014, sp. nov.
Creators: name: Sehnal, Richard
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Melolonthidae
Engistanoxia
Engistanoxia sagala

description: Engistanoxia sagala sp. nov.   
(Figs 1–7)  
 Type locality. Kenya, Taita Taveta county, Voi env., Sagala Hills.   
 Type material. HOLOTYPE: J, ދ KENYA, Coast prov. [the earlier division of the country] / Voi env. & Sagala Hills / 23. XI. - 4. XII. 2000 / M. BednaĜík leg. [p]ތ (NMPC). PARATYPES: No. 1, ♀ and Nos 2–6 and No. 12, JJ, same data as holotype; Nos 7–11, JJ, ދ KENYA, Coast prov. / Voi env. & Sagala Hills / 23. XI. - 4. XII. 2000 / Ivo MartinĤ leg. [p]ތ; Nos 13–14, JJand No. 15, ♀, ދ KENYA XI-92 / near Voi, Sagala reg. / Werner leg. // Engistanoxia [bold] / dessertineae / M. Lacroix, 2002 / Coll. M. Lacroix [p]ތ. Paratypes Nos 1–10 in RSCV, No. 11 in MBCO, Nos 12–15 in MLCR.   
  Description. Male  (holotype). Body length 23.2 mm (without pygidium). Body elongate, posteriorly slightly dilated. Dorsal surface and abdomen reddish brown, legs partly lighter, antennae pale brown, elytra pruinose (Fig. 1).   
 Head. Clypeus 2.19 times wider than long, with anterior angles broadly round and anterior margin strongly upturned and bisinuate; surface punctate, with smooth facets between punctures; posterolaterally oriented, pointed, semierect scale issues from centre of each puncture; some medial scales narrower than others, fronto-clypeal suture weakly indicated. Frons densely and coarsely punctate, with a scale issuing from each puncture; scales near midline point laterally toward a longitudinal bulge in each half of frons, forming a bisinuate pattern (Fig. 2). Labrum bilobed, anterior angles obtuse but distinct, macrosetae longest at edge of clypeus and around eye canthus. Genae rugo-punctate, with group of short macrosetae. Occiput deeply but sparsely punctate, with pointed scales oriented posteriorly. Eye canthus long, with macrosetae in two regular rows; anterior conical macrosetae long, posterior conical macrosetae ¿ve times shorter than clypeal scales. Antennae decamerous. Antennal club hexamerous, straight and 1.5 times longer than antennal shaft (antennomeres 1–4 combined). Antennal club on apex with irregular, ¿ne, short macrosetae with sensilla pits. Antennomere 1 flat, ventrally with long setae, as long as antennomers II and III combined; antennomere 4 short, anteriorly pointed (Fig. 3). Terminal maxillary palpomere widest at midlength, apically truncate, with a terminal tubercle on inner, more angular side, and on dorsal surface with a large oval alutaceous area widening toward apex (Fig. 4).   
 Pronotum weekly convex, approximately heptagonal (Fig. 5), 1.7 times wider than long, widest in posterior third, with distinct impressed medial line.Anterior and posterior angles round but discernible. Lateral margins crenulate, with a yellow hook-shaped macroseta issuing from each crenulation. Surface densely punctate; punctures unevenly distributed, spaces between them smooth and matte; each puncture bearing pointed scale only about half as long as scales on clypeus and frons. Scales forming groups oriented mostly toward midline (Fig. 5).   
 Scutellar shield wider than long, triangular, evenly punctate in anterior half, with scales similar to those on pronotum (Fig. 5) and elytra, along midline with a strip of long, ¿ne hairs.   
 Elytra convex, elytron 1.4 times longer than wide. Each elytron quadricostate (including sutural costa), costae reaching from humeral to terminal bulge, costae 2 and 4 somewhat shorter. Part of elytron between terminal bulge and apex short, steeply inclined. Crests of costae glabrous, their sides and intervals unevenly covered by punctures and scales similar to those on pronotum. Spaces between punctures and scales ¿nely rugate and matte. Lateral margins with hairs twice as long as those on pronotal margins.   
Macropterous.  
 Legs. All femora shiny, irregularly punctate, macrosetae long. Protibia tridentate, claws strongly curved, microrugate, each with a minute basal tooth and a much larger tooth at midlength. Meso- and metatibiae expanded apically, densely covered by broad macrosetae, spaces between individual macrosetae larger than diameter of macrosetae, with one oblique interrupted carina externally. Apical edge with a row of short, stout macrosetae of equal length, terminal calcars stout, long, lower calcar little shorter than upper; pro-, meso- and metatarsomeres without patches of short, dense macrosetae; metatarsomeres covered by isolated, sparse macrosetae ventrally; meso- and metatibiae shallowly puctate, ¿ne long macrosetation recumbent.   
 Abdominal ventrites reddish brown, matte, ¿nely punctate, covered by oval white scales and a row of semierect yellowish-white macrosetae. Pygidium slightly concave, shallowly punctate, with yellowish-white caudally oriented scales becoming toward posterior margin ¿ne, short, yellowish-white macrosetae.   
 Male genitalia (Figs 6–7). Aedeagus symmetrical. Phallobase in lateral view with three extensions, ventrolaterally deeply excised. Parameres forming a symmetrical lobe with medial oval summit and an oval concavity in its centre. Dorsal part of lobe steeply elevated and then reclined at right angle ventral to base.   
 Female. Similar to male but has a shorter antennal club and wider and shorter protibia.   
 Variability. Males. Paratypes somewhat variable in body length (18.4–23.2 mm). Colour as in holotype.   
 Differential diagnosis. The habitus of Engistanoxia sagala sp. nov. is close to E. dessertineae, from which it differs in the pointed but not prolonged antennomere IV, size of the accessory tooth on tarsal claws, crenulate lateral margins of the pronotum, shape and pattern of scales on the head and pronotum, shape of the terminal maxillary palpomere and its pit, frontal margin of clypeus, bisinuate phallobase and shape of the parameres (Figs 6–7).   
 Etymology. The species epithet refers to the type locality, Sagala Hills in Kenya (Taita Taveta county), and the name is a noun in apposition.   
 Collection circumstaces. Adults were collected at light (M. BednaĜík and I. MartinĤ, pers. comm.).
description: Published as part of Sehnal, Richard, 2014, Engistanoxia sagala sp. nov. from Kenya (Coleoptera: Scarabaeidae: Melolonthinae: Melolonthini), pp. 563-569 in Acta Entomologica Musei Nationalis Pragae 54 (2) on pages 564-568, DOI: 10.5281/zenodo.5301245