Clivina margineserrata Balkenohl, 2025, sp. nov.
Creators: name: Balkenohl, Michael
Publisher: name: Zenodo
Year: 2025
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Clivina
Clivina margineserrata

description: Clivina margineserrata sp. nov.
(Figs 21, 56, 115)
Type material. Holotype: ♀, with labels and data: white, black printed “ SARAWAK (Borneo), ca 40 km SE KAPIT 03.1994, J. Kodada leg.” (NHMW). Paratype: 1 ♀, same data as holotype but “ca 25 km E KAPIT” (CMBB).
Remark. In the paratype, the right hind leg and some tarsomeres of the left hind leg are missing. In addition, the elytra seem to be somewhat more slender, possibly due to a damage.
Description. Measurements. Holotype / paratype: Body length 6.16 mm / 5.92 mm, width 1.86 mm / 1.63 mm, ratio L/W of pronotum 0.90 / 0.92, ratio L/W of elytra 1.76 / 1.94. Colour glossy, piceous. Supraantennal plate indistinctly translucent. Antennae, mouthparts, intermediate and hind legs rufous-fuscous.
Head a quarter narrower than pronotum. Clypeus with central part straight, with slightly but distinctly projecting teeth laterally, separated from clypeal-wing by distinct notch; wing distinct, rounded, wider but not as projecting as clypeal-teeth, separated from supraantennal plate by distinct notch, supraantennal plate with margin laterally obtuse-angular, clypeus, wing, and supraantennal plate reflexed margined. Supraantennal plate extended posteriorly into a blunt keel, separated from wing by furrow. Clypeus continuing into frons without furrow, slightly convex, with scattered small punctures laterally. Frons with indistinct indentation at middle, separated from supraantennal plate and supraorbital carina by furrow, with clypeal seta in front; supraorbital carinae blunt, with second shorter carina laterally, with two supraorbital setigerous punctures at middle of eye and posterior to gena-level. Neck constriction closed, formed by a straight step from frons. Eye moderately large, gena distinct but small. Antenna elongate, reaching up to base of elytron. Labrum slightly excised anteriorly, isodiametrically reticulated, seven-setose. Mentum isodiametrically reticulated, with oblique wrinkles, lobe with lateral margin convex, with acute apical tip, medially concave to cleft, median tooth much lower than lobes, acute, not carinate.
Pronotum (Fig. 56) with disk slightly convex in lateral view, in frontal view flattened at middle and moderately convex laterally. Shape square to cordate, wider than long. Reflexed lateral margin distinctly serrate, nearly parallel, widest at middle; anterior angle somewhat rounded but well-marked, posterior angle distinct, with robust tooth; lateral channel distinctly widened, rugose and reticulated, continuing up to base as narrow, straight margin. Median line engraved, complete, joining and surpassing anterior transverse line. Anterior transverse line distinct, complete. Disk with pattern of smaller punctures resembling with some phantasy to the letter Y, with some transverse wrinkles, with isodiametric reticulation at lateral margin and at base, with posterior setigerous puncture removed from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 30°. Basal channel on flange wide, rugose, with sharp carina.
Elytron (Fig. 21) with disk flattened in anterior half in lateral view, distinctly convex in frontal view. Shape of elytra elongate, about two-fifth longer than wide, with lateral margin indistinctly convex and diverging, maximum width behind middle. Humerus distinct, rounded. Reflexed lateral margin slightly crenulate behind humerus. Row of setigerous punctures in the lateral channel regular in size and distance. Scutellar striole distinct; setigerous tubercle at base of first stria distinct, with small tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, indistinctly punctate, one to four free at base, five and six joining at humerus, one and two ending free at apex, three and four and five and six indistinctly joining at apex. Intervals slightly convex at middle, moderately convex laterally, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as very short carina, interval seven with distinct carina at apex, eight more sharp carinate at apex. Surface of all intervals isodiametrically reticulated, reticulation more distinct laterally.
Hind wing fully developed.
Lower surface. Proepisternum with fine isodiametrical reticulation, with extremely fine wrinkles laterally. Sternite of abdomen with isodiametric reticulation, weaker medially. Last visible sternite with the two marginal setigerous punctures widely separated, with slight projection posteriorly at middle where it is slightly hollowed out and smooth, with narrow marginal furrow at middle.
Legs. Protibia slender, with longitudinal reticulation, with narrow sulcus dorsally, with three spines, movable spur moderately arcuate, basal tarsomere slender at base, carinate, with acute tubercle and two wide setae at middle of tarsomere. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg widened.
Male genitalia unknown.
Female gonocoxites and epipleurite (Fig. 115). Basal gonocoxite with two long and two minute setae, apical gonocoxite elongated, with one long seta at base and one long seta at beginning of apical third, with two very long setae apically. Epipleurite trapezoid-shaped, with one long seta arising from tubercle-like lappet.
Variation. In the paratype, the elytra are slenderer, and the lateral margin of the pronotum is not as conspicuously serrate as in the holotype.
Distribution. Known from Sarawak on the island of Borneo.
Diagnosis. A medium sized piceous species with square to cordate shaped pronotum and all intervals of the elytron reticulated. The most similar species C. punctaticollis sp. nov., differs mainly by the elytra with intervals one to six glossy, the pronotum with the lateral margin sub-serrate, narrow lateral channel, much smaller tooth at the posterior angle, and the different puncture pattern on the disk. In addition, it is much smaller.
Etymology. The name refers to the pronotum with its lateral border (Latin: margo, marginis) which is distinctly serrated (Latin: serrate).
description: Published as part of Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), pp. 1-70 in Zootaxa 5592 (1) on pages 35-36, DOI: 10.11646/zootaxa.5592.1.1, http://zenodo.org/record/14953674