Novispathodus pingdingshanensis
Creators: name: Leu, Marc
name: Bucher, Hugo
name: Vennemann, Torsten
name: Bagherpour, Borhan
name: Ji, Cheng
name: Brosse, Morgane
name: Goudemand, Nicolas
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Chordata
Ozarkodinida
Gondolellidae
Novispathodus
Novispathodus pingdingshanensis

description: Novispathodus pingdingshanensis (Zhao & Orchard, 2007)
Fig. 17D–F; M–P, R, S, U, X, AA, AB, AD, AF
*2007 Neospathodus pingdingshanensis n. sp.; Zhao & Orchard, Zhao et al., p.36, pl. 1, fig. 4A–C.
2012a Novispathodus pingdingshanensis (Zhao & Orchard); Goudemand & Orchard in Goudemand et al., p. 1030–1031, figs. 2B, F, G, I–J, M, P, Q, AD, 3T-U, 6.
2013 Neospathodus pingdingshanensis Zhao & Orchard; Chen et al., p 825, fig. 3.10, 3.12.
2013 Neospathodus waageni subsp. nov. A; Metcalfe et al., p. 1144, figs. 9.1–9.5, 9.7, 9.8, 9.10.
2014 Novispathodus pingdingshanensis (Zhao & Orchard); Maekawa & Igo in Shigeta, p. 239–240, figs. 171.13–171.31.
2015 Novispathodus pingdingshanensis (Zhao & Orchard); Chen et al., p. 111, 112, figs. 7.1–7.4, 8.5, 8.6.
2016 Novispathodus ex. gr. pingdingshanensis (Zhao & Orchard); Komatsu et al., p. 69, figs. 5.4a–5.5c.
2016 Neospathodus robustus Koike; Chen & Kolar-Jurkovšek in Chen et al., p. 93, fig. 9.5 (only).
2018 Novispathodus pingdingshanensis (Zhao & Orchard); Maekawa in Maekawa et al., p. 36–37, figs. 20.2–20.18, 21.1–21.13.
2019 Novispathodus pingdingshanensis (Zhao & Orchard); Chen et al., fig. 3, nr. 8.
2019 Novispathodus pingdingshanensis (Zhao & Orchard); Liu et al., p. 13, pl. 3 fig. 5 (only).
Material.> 100 specimens.
Diagnosis (Zhao and Orchard, in Zhao et al., 2007; emended by Goudemand, in Goudemand et al., 2012b). Small segminate P 1 elements characterized by a length/ height ratio in the range of 1.32–2.34, and about 4–9 robust, wide, mostly fused, and distinctively posteriorly recurved denticles. In lateral view, the basal margin is usually straight. A large, broadly expanded oval to sub-rounded basal cavity is upturned on the inner margin and flat to downturned on the outer margin.
Remarks. Goudemand (in Goudemand et al., 2012b) revised the original diagnosis by noticing that the basal margin is not necessarily straight and therefore, the most strikingly difference to Nv. waageni is the denticulation: the denticles axes are distinctively recurved posteriorly whereas in Nv. waageni and Nv. abruptus, the denticles are straight, inclined or radiating. Te two or three denticles anterior of the cusp are often clearly asymmetrical and the posterior edge of the element is much shorter than the anterior one. In most sections worldwide, Nv. pingdingshanensis first occurs within the positive δ 13 Ccarb excursion of the latest Smithian and may extend to the earliest Spathian (Goudemand et al., 2019; Leu et al., 2019; Zhang et al., 2019). Some elements within our rich material resemble Ns. Pingdingshanensis, but appear to have a relatively small basal cavity, more posteriorly recurved denticles or more numerous denticles that what was previously described for this species. Tese elements (Figs. 17C, J, K, T; 18E, I–K; 19O, Q; 20H, N–O, S; here assigned to N. ex gr. pingdingshanensis) may deserve differentiation in the future. Two elements illustrated by Metcalfe et al., (2013, figs. 9.6, 9.9, p. 1144) as ‘ Neospathodus waageni subsp. nov. A’ display a higher anterior end and more recurved denticles than the other elements they included in ‘ Neospathodus waageni subsp. nov. A’, which we synonymized with Ns. pingdingshanensis: elements like these two elements do not seem to fit in Ns. pingdingshanensis and may deserve assignment to a new species. Some further elements resembling Ns. pingdingshanensis but distinctively shorter than Ns. pingdingshanensis are herein assigned to a new species (Nv. gryphus n. sp. see below).
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Occurrence. Worldwide occurrence. China: Jinya/ Waili area, Guangxi (Goudemand et al., 2012b) Jiarong, southern Guizhou (Chen et al., 2013, 2015). Anshun Fm., Qingyan section, Guizhou (Ji et al., 2011). Chaohu (Zhao et al., 2007, 2008). Daxiakou, Hubei (Zhao et al., 2013). Vietnam: Bac Tuy Fm., Xenoceltites variocostatus and Tirolites beds, (Komatsu et al., 2016; Shigeta et al., 2014). Canada: Scythogondolella mosheri zone, Wapiti Lake (Orchard & Zonneveld, 2009). Australia: Hovea Member of Kockatea Shale, upper part of Neospathodus waageni Zone, Smithian substage (Metcalfe et al., 2013).
description: Published as part of Leu, Marc, Bucher, Hugo, Vennemann, Torsten, Bagherpour, Borhan, Ji, Cheng, Brosse, Morgane & Goudemand, Nicolas, 2022, A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China, pp. 1-61 in Swiss Journal of Palaeontology (19) (19) 141 (1) on pages 27-29, DOI: 10.1186/s13358-022-00259-x, http://zenodo.org/record/12000082