Rhagovelia angustipes Uhler 1894
Creators: name: Galindo-Malagón, Ximena Alejandra
name: Morales, Irina
name: Moreira, Felipe Ferraz Figueiredo
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hemiptera
Veliidae
Rhagovelia
Rhagovelia angustipes

description: Key to species of the Rhagovelia angustipes complex from Colombia
1. Monomorphic apterous; tarsal formula 2-2-2 (articulation more easily viewed on middle leg); found mainly on brackish or salt water (e.g., mangroves, estuaries).......................................................... [salina group] ... 2
-. Dimorphic apterous or macropterous; tarsal formula 3-3-3 (articulation more easily viewed on middle leg); found mainly on freshwater (e.g., streams, rivers)........................................... [bisignata and hambletoni groups] ... 7
2. Eyes covered with abundant setae........................................................................ 3
-. Eyes covered with one or two setae only.................................................................. 4
3. Body length 2.20–2.40 mm in the male and 3.30–3.50 mm in the female; posterior surface of male hind femur with a row of 3–7 spines; paramere and proctiger as in Figs. 19D, 20D ............................ Rhagovelia plumbea Uhler, 1894
-. Body length ~ 1.75 mm in the male and ~ 2.90 mm in the female; posterior surface of male hind femur with a row of 6–10 spines; paramere and proctiger as in Figs. 19E, 20E ..................... Rhagovelia tintipan Molano, Morales & Moreira, 2018
4. Male fore tibia straight or slightly curved (Fig. 21B); male abdominal segment VIII large, longer dorsally than abdominal tergum VII (Fig. 1B); paramere strongly sculptured (Fig. 19B); proctiger as in Fig. 20B; dorsum of female abdominal terga VI–VIII (sometimes also V) shiny black centrally (Fig. 3B)........ Rhagovelia colombiana (Polhemus & Manzano, 1992)
-. Male fore tibia strongly curved (Fig. 21A); male abdominal segment VIII small, approximately equal in length dorsally to abdominal terga VII (Figs. 1A, 1C, 1D, 1E); paramere shape simpler (Figs. 19A, 19C, 19D, 19E); at most dorsum of female abdominal terga VII–VIII shiny black or brown centrally (Fig. 1C, 1E, 3A, 3C, 3E)................................ 5
5. Body length ~3.00 mm in the male and ~ 4.20 mm in the female; paramere and proctiger as in Fig. 19A, 20A ........................................................................ Rhagovelia arcuata (Polhemus & Manzano, 1992)
-. Body length 2.20–2.60 mm in the male and 3.00– 3.20 mm in the female; paramere and proctiger not as above........... 6
6. Posterior surface of male hind femur with about 6 spines beyond middle; male abdominal sterna without median carina; female abdominal laterotergites slightly elevated (Fig. 3E)........................... Rhagovelia rosarensis Padilla-Gil, 2010
-. Posterior surface of male hind femur with about 5 spines basally, followed by one long spine before middle, then about 13 smaller spines towards apex; male abdominal sterna VII–VIII with weak median carina; female abdominal laterotergites vertical or nearly so (Fig. 3C)................................. Rhagovelia nuqui Molano, Morales & Moreira, 2018
7. Hind femur without spines in both sexes (Fig. 21D).......................................................... 8
-. Hind femur with spines in both sexes (Figs. 21E, 21F)...................................................... 10
8. Dorsum of abdominal terga V–VIII shiny black centrally; paramere and proctiger as in Figs. 19V, 20V ................................................................................. Rhagovelia guachiconoense Padilla-Gil, 2019
-. At most dorsum of abdominal segments VII–VIII shiny black centrally; paramere and proctiger not as above............ 9
9. Antennomere II shorter than III; fore and hind coxae and trochanters dark brown to black (Figs. 10F, 12F), rarely brown to yellow; male fore tibia very thin for about 3/4 of length, then expanding to apex (Fig. 21D).................................................................................................... Rhagovelia longipes Gould, 1931
-. Antennomere II longer than III; fore and hind coxae and trochanters yellow (Figs. 10A); male fore tibia thicker and more uniform throughout length (Fig. 10A)....................................... Rhagovelia caunapi Padilla-Gil, 2015
10. Male metasternum and abdominal sterna II–IV strongly swollen, with brushes of long golden setae medially (Fig. 10C); lateral margins of male abdominal segment VIII clearly divergent posteriorly (Fig. 9C); paramere and proctiger as in Figs. 19L, 20L. ................................................................. Rhagovelia gastrotricha Padilla-Gil, 2011
-. Male metasternum and abdominal sterna II–IV not strongly swollen, covered only with short setae in most species; male abdominal segment VIII subcylindrical, with lateral margins parallel or bowed, but not clearly divergent posteriorly (Figs. 5B, 5C, 5D, 5E, 5F, 9B, 9D, 9E, 13A, 13B, 13C, 13D, 17A, 17E); paramere not as above.............................. 11
11. Male hind trochanter with spines; male hind femur much thicker than middle femur, enormously incrassate in most specimens (Fig. 21F); paramere and proctiger as in Figs. 19I, 20I ........................ Rhagovelia calopa Drake & Harris, 1927
-. Male hind trochanter without spines; male hind femur at most slightly thicker than middle femur, never enormously incrassate (Fig. 21E); paramere and proctiger not as above........................................................... 12
12. Male abdominal sternum VII anteriorly with a stout, slightly curved spine; paramere and proctiger as in Figs. 19S, 20S .............................................................................. Rhagovelia spinosa Gould, 1931
-. Male abdominal sternum VII without spine; paramere and proctiger not as above................................. 13
13. Body length 2.60–2.90 in the male and 2.80–3.10 in the female............................................... 14
-. Body length 3.00– 4.20 in the male and 3.20–4.70 in the female............................................... 18
14. Male hind tibia without spines or apical spur (Figs. 13A, 17C) (apex of tibia can sometimes bear a tuft of setae, but never a spur).............................................................................................. 15
-. Male hind tibia with short spines and an apical spur (Fig. 9E, 13C, 13D)........................................ 16
15. Lateral margins of male abdomen more evenly tapering to apex (Fig. 17C); paramere and proctiger as in Figs. 19T, 20T; female hind femur thinner (Fig. 18A); female abdomen elongated (Fig. 18C).......... Rhagovelia tantilla Drake & Harris, 1933
-. Lateral margins of male abdomen more strongly bowed on segments IV–V (Fig. 13A); paramere and proctiger as in Fig. 19P, 20P; female hind femur thicker (Fig. 15A); female abdomen short and robust (Fig. 15A)...... Rhagovelia molanoi sp. nov.
16. All coxae and trochanters yellowish (Fig. 10E, 12E); male abdominal sterna II–V with median comb of long golden setae (Fig. 10E); male abdominal segment VIII large and robust (Fig. 9E); paramere and proctiger as in Figs. 19N, 20N; lateral margins of female abdomen sinuous (Fig. 11E); posterior margin of female abdominal tergum VIII with long setae (Fig. 11E)...................................................................................... Rhagovelia graziae sp. nov.
-. Middle trochanter dark brown to black; male abdominal sterna only with short setae (Figs. 14C, 14D); male abdominal segment VIII small (Figs. 13C, 13D); paramere and proctiger not as above; lateral margins of female abdomen more evenly tapering or bowed (Figs. 16C, 16D); posterior margin of female abdominal tergum VIII without long setae (Figs. 15C, 15D)........ 17
17. Male hind femur with 9–11 spines; male hind tibia with 11 short spines and an apical spur; lateral margins of male abdomen more evenly tapering to apex (Fig. 13C); paramere and proctiger as in Figs. 19Q, 20Q; female hind femur with 6–7 spines; female abdomen more elongated, with laterotergites vertical or nearly so (Fig. 15C)...................................................................................................... Rhagovelia rosensis Padilla-Gil, 2011
-. Male hind femur with 4–7 spines; male hind tibia with 2 short spines and an apical spur; lateral margins of male abdomen more strongly bowed on segments IV–V (Fig. 13D); paramere and proctiger as in Figs. 19R, 20R; female hind femur with 4–5 spines; female abdomen more robust, with laterotergites slightly elevated (Fig. 15D)..... Rhagovelia santanderi Padilla-Gil, 2015
18. Body length 3.00– 3.40 in the male and 3.20–3.75 in the female............................................... 19
-. Body length 3.50–4.20 in the male and 4.00– 4.70 in the female............................................... 23
19. Male hind tibia only with an apical spur, without small spines throughout length; paramere and proctiger as in Figs. 19F, 20F. ...................................................................... Rhagovelia angustipes Uhler, 1894
-. Male hind tibia with small spines throughout length and an apical spur; paramere and proctiger not as above........... 20
20. Male hind femur thicker, shorter than hind tibia (ratio ~0.90/1.00) (Fig. 13B); male hind femur with 4–5 spines; only abdominal tergum VIII shiny black.................................................... Rhagovelia penta Padilla-Gil, 2015
-. Male hind femur thinner, longer than hind tibia (ratio ~1.05/1.00) (Figs. 5B, 5D, 17E); male hind femur with at least 6 spines; central shiny black areas at least on dorsum of abdominal segments VII–VIII.................................... 21
21. Male hind femur slightly surpassing apex of abdomen, with 6 spines (Fig. 5D); paramere and proctiger as in Fig. 19H, 20H; female hind femur with 4 spines........................................... Rhagovelia calceola Padilla-Gil, 2011
-. Male hind femur distinctly surpassing apex of abdomen, with at least 8 spines (Fig. 5B, 17E); paramere and proctiger not as above; female hind femur with at least 7 spines............................................................ 22
22. Middle coxa black (Figs. 17F, 18D); fore and hind trochanters black (Figs. 17F, 18D); male hind femur with at least 10 spines; paramere and proctiger as in Figs. 19U, 20U; female abdominal mediotergites approximately in the same horizontal plane (Fig. 18C); female abdominal laterotergites horizontal or slightly elevated (Fig. 18C)...... Rhagovelia tenuipes Champion, 1898
-. All coxae yellow (Figs. 6B, 8B); fore and hind trochanters yellow (Figs. 6B, 8B); male hind femur with 8 spines; paramere not as above; female abdominal mediotergites I–III raised, IV–VI depressed, VII horizontal, tergum VIII declining posteriorly (Fig. 7B); female abdominal laterotergites elevated, more strongly on last two segments (Fig. 7B)........................................................................................... Rhagovelia barbacoensis Padilla-Gil, 2015
23. Fore and hind trochanters yellow (Figs. 10D, 12D); middle coxa yellow (Figs. 10D, 12D); male hind femur shorter and thicker, with 6–7 spines (Fig. 9D); paramere and proctiger as in Figs. 19M, 20M. ........... Rhagovelia grandis Padilla-Gil, 2011
-. Fore and hind trochanters black; middle coxa black; male hind femur longer and thinner, with at least 10 spines (Figs. 5C, 5F, 9B); paramere and proctiger not as above................................................................. 24
24. Legs with abundant long black setae (Figs. 5C, 7C); paramere and proctiger as in Fig. 19G, 20G; female abdominal laterotergites strongly elevated, almost touching at apex of last segment, which bears abundant brown setae posteriorly (Fig. 7C).................................................................................. Rhagovelia boyacensis sp. nov.
-. Legs without abundant long black setae (Figs. 5F, 7F, 9B, 11B); paramere and proctiger not as above; female abdominal laterotergites not as above (Fig. 11B).................................................................... 25
25. Central shiny black areas on dorsum of abdominal segments VII–VIII in the male and VI–VIII in the female; male hind femur with 14–16 spines; paramere and proctiger as in Figs. 19J, 20J ..................... Rhagovelia cardia Padilla-Gil, 2011
-. Central shiny black areas on dorsum of abdominal segments V–VIII in both sexes; male hind femur with 10 spines; paramere and proctiger as in Figs. 19K, 20K ....................................... Rhagovelia cimarrona Padilla-Gil, 2011
description: Published as part of Galindo-Malagón, Ximena Alejandra, Morales, Irina & Moreira, Felipe Ferraz Figueiredo, 2021, Revision of the Rhagovelia angustipes complex (Insecta: Hemiptera: Veliidae from Colombia, pp. 167-225 in Zootaxa 4958 (1) on pages 169-171, DOI: 10.11646/zootaxa.4958.1.11, http://zenodo.org/record/4691515