Pupinidius obrutschewi subsp. obrutschewi Moellendorff 1901
Creators: name: Wu, Min
name: Zheng, Wei
Publisher: name: Zenodo
Year: 2009
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Stylommatophora
Enidae
Pupinidius
Pupinidius obrutschewi
Pupinidius obrutschewi obrutschewi (moellendorff, 1901)

description: 9. Pupinidius obrutschewi obrutschewi (Moellendorff, 1901)
Figs 1, 4 E–G, 5A–B, 8E, 13
Buliminus obrutschewi Sturany, 1900: 29, pl. 3, figs 26–28.
Buliminus (Pupinidius) obrutschewi Moellendorff, 1901: 342.
Buliminus (Pupinidius) obrutschewi eurystoma Moellendorff, 1901: 342, pl. 8, figs 23, 24; Wiegmann, 1901: 246, pl. 10, figs 40–44; Kobelt, 1902: 846, pl. 108, figs 1, 2, pl. 120, figs 23–27.
Pupinidius obrutschewi obrutschewi Yen, 1939: 83, pl. 7, fig. 46.
Material examined. HBUMM-05672 (20 out of 36 mature shells were measured; specimen-1 for dissection): 1233 m a.s.l., 33°41΄47.3΄ N, 104°29΄10.3΄ E, eastern bank of Lianghekou, Gansu, 2006-X-2, limestone, coll. J.Liu & W.Zheng; HBUMM-05678 (30 fmss): 1286 m a.s.l., 33°41΄38.5΄ N, 104°28΄48.6΄ E, western bank of Lianghekou, Gansu, 2006-X-2, other stone replacing limestone, coll. J.Liu & W.Zheng (measurement: ewh 1.500–1.669–1.875, wh 6.500–7.044–7.500, h 18.7–21.0–23.3, diam. maj. 10.1–11.1–11.8, aph 9.0–9.7–10.9, apw 6.4–7.5–8.0, h/diam. maj. 1.62–1.89–2.12, h/aph 1.99–2.16–2.33); ZI RAS-N512: type material, 1885- VI-22, coll. Potanin. SMF-41982: Lectotype v. B. eurystoma Mlldff. Liang-dshia-pa, Kansu, China. Slg. O.v.Moellendorff. SMF-41983: Paratypes v. B. eurystoma Mlldff. 2 fmss. Liang-dshia-pa, Kansu, China. Slg. O.v.Moellendorff (measurement of types: ewh 1.500–1.563–1.625, wh 6.875–7.129–7.375, h 21.1–22.2–23.8, diam. maj. 12.9–13.4–13.9, aph 9.9–10.7–11.2, apw 8.5–8.8–9.0, h/diam. maj. 1.58–1.66–1.72, h/aph 1.97–2.06–2.13). SMF-41984: One juvenile. Wen-hsien, Kansu, China. Slg. O.v.Moellendorff.
Distribution. Gansu (type locality, Liang-dshia-pa, Lianghekou, Wenxian).
Type locality. Schy-pa (S Gansu).
Diagnosis. Shell bandless; striate. Aperture faintly with or without angular tubercle. Most swollen part at body whorl. With 5.750–7.375 whorls; height 15.9–23.8, diam. maj. 10.4–13.9. Penis without penial caecum; internally without fine longitudinal pilaster. Papilla developed by fused proximal pilasters forming V-shaped structure present. Epiphallus straight. Epiphallic caecum present.
Re-description of shell. Pointed-ovate; apex protruded. Most swollen part at body whorl. Shell dextral; thin; solid; opaque; glossy; speckled or not; not spirally grooved;with 5.750–6.250–6.625 whorls. Whorls flattened; not shouldered. Growth lines usually not very clear. Embryonic shell smooth; polished; with 1.500–1.551–1.750 whorls. Postnuclear whorls smooth. Suture normal, without narrow defined zone on beneath whorl. Last whorl slightly gradually ascending towards aperture; slightly angular at periphery (in lectotype indistinct); without smoothed spiral peripheral depression nor rugate region. Shell 15.9–19.5–21.5 high; 10.4–11.3–12.0 in diam. maj. Height/width ratio 1.44–1.73–2.06. Aperture in a plane; completely attached to body whorl; truncate-ovate; not adnate; oblique; not armed; faintly with angular tubercle, or without angular tubercle; 8.2–9.6–10.2 high; 6.7–7.4–8.0 wide. Ratio of shell height to aperture height 1.77–2.03–2.28. Peristome narrowly reflexed with distinct cuff; expanded; sharp. Parietal callus indistinct. Columellar margin reflexed. Columella vertical. Outer edge of columellar lip vertical. Umbilicus very narrowly open. Apical whorls normally coloured. Shell multicoloured; in darker or lighter brown. With narrow or thick white stripes below protoconch. Aperture white; not brightly banded; spirally without colour band.
Re-description of genitalia. Vas deferens slightly swollen proximally; entering epiphallus apically with distinct demarcation. Epiphallus more or less short; slightly narrowed towards proximal end; normally straight; externally smooth. Epiphallic caecum present; blunt apically; located near vas deferens entrance. Flagellum short; tubular; proximally normal; with tip pointed. Penis with terminal entrance of epiphallus; clavate; without penial caecum; distally enlarged; thin-walled; internally without fine pilasters, with with 2–3 circular ridges and 1 V-shaped pilaster. V-shaped pilaster with free proximal end approaching at penial retractor insertion, distally developing a small nipple-like papilla. Penis without process. Penial verge absent. Penial appendix moderately long; branched off from proximal portion of penis; divided into sections including A-1+A-2, A-3 and A-4+A-5. A-1 and A-2 fused, interiorly smooth. A-2 and A-3 not fused. Boundary between A-4 and A-5 indistinct. A-5 long; almost straight. Penial retractor biramous; attaching to penis in middle part and to A-1+ A-2; with penial retractor arms arising from diaphragm closed to each other.
Additional retractor other than penial or appendicular absent. Muscular band connecting vagina and epiphallus absent. Atrium short; with weak retractor. Free oviduct longer than vagina. Vagina short; not swollen; straight; not lined with loose, spongy tissue; unpigmented. Bursa copulatrix duct moderately long; proximally straight. Bursa copulatrix with stalk; without apical ligament; normal in size; with short neck; well defined. Diverticle normally present; longer than reservoir; unexpanded. Bursa copulatrix and diverticle distinguishable; forked more distally from their base.
Measurement of genitalia. P: 3; Ep: 7.8; Fl: 1.4; VD: 15.0; Va: 2.3; FO: 4.3; BCD: 5.9; BC: 1.1; D: 4.5; A- 1+A-2: 3.8; A3: 1.8; A-4+A-5: 12 (in HBUMM-05672-specimen 1).
Remarks. The new collected specimens are more or less smaller than the type specimens in shell size. The population (HBUMM-05678), habited in the opposite bank of Bailongjiang River, differs slightly from the individuals represented by HBUMM-05672 by larger shell size, darker colouration, and more dense striation on shell.
description: Published as part of Wu, Min & Zheng, Wei, 2009, A review of Chinese Pupinidius Moellendorff (Gastropoda, Stylommatophora: Enidae), with the description of a new species, pp. 1-31 in Zootaxa 2053 on pages 14-16, DOI: 10.5281/zenodo.186646