Mortoniella (Mortoniella) chalalan Blahnik & Holzenthal 2017, new species
Creators: name: Blahnik, Roger J.
name: Holzenthal, Ralph W.
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Magnoliopsida
Gentianales
Apocynaceae
Mortoniella
Mortoniella chalalan

description: Mortoniella (Mortoniella) chalalan, new species   
Fig. 57  
This diminutive species has a relatively simple overall morphology, but is reasonably distinctive. In addition to the spine-like dorsal processes of the phallicata, which character was used to place it in the punensis subgroup, the species has two straight, narrow endophallic spines, 1 ventromesal spine and 1 apicodorsal spine. This combined configuration is not known for any other species. The dorsal phallic spine is symmetrical in shape and has its apex depressed (dorsoventrally flattened), as is typical of the subgroup. The shape of tergum X, which is short, with relatively broad apicolateral processes and a concave mesal invagination, is also distinctive.  
 Adult —Length of forewing: male 2.6-2.7 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0:3:4. Overall color medium brown, apices of tarsal segments whitish or pale brown. Tibial spurs darker than legs, contrasting in color. Wing bar at anastamosis indistinct, marked with light brown setae, most evident at arculus.   
 Male genitalia —Ventral process of segment VI relatively small, laterally compressed, posteriorly directed, length about 1½ times width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short, lateral margins subparallel, apicomesal margin somewhat concave between apicolateral lobes, apicolateral lobes relatively short and broad, subtruncate; ventrolateral lobes rounded, weakly projecting. Inferior appendages short, without dorsolateral projections, apicoventral projections short, acute apically, ventromesal margin with very short subtruncate projection. Mesal pockets of inferior appendage with spine-like apical processes short, posterodorsally curved. Paramere appendage moderately elongate (shorter than dorsal phallic spine), nearly uniformly narrow, apex acute, opposite appendages converging mesally. Dorsal phallic spine stout, nearly uniform in width, without distinct ventral projection, depressed apically, apex gradually upturned, acute in both lateral and dorsal views. Phallicata with dorsal margin sclerotized and produced anteriorly into anvil-like projection, laterally with pair of curved spine-like projections, projections hooked over paramere appendages; ventral margin of phallicata simple, without lobes or projections. Endophallic membrane elongate, with small rounded membranous preapical and apicomesal lobes; ventromesal spine nearly straight, very narrow and nail-like; endophallic membrane with additional apical dorsomesal spine, spine very narrow, attenuate, slightly longer than ventromesal spine; phallotremal spines absent.   
 Holotype male (pinned)— BOLIVIA: LaPaz: ANMI Madidi, Chalalan Ecolodge, Raya Mayo river at Anta Trail, 14.43557° S, 67.92935° W, el 264 m, 26.vii.2003, Robertson and Blahnik (UMSP000094778) (UASC).   
  Paratypes — BOLIVIA: Santa Cruz:  PN and ANMI Amboró, Guardia Parque Mataracú, Quebrada Verde Uno, 17.55389° S, 63.86917° W, el 374 m, 19-23.xi.2004, Robertson, Garcia and Vidaurre– 2 males (alcohol) (UMSP); PN & ANMI Amboró, Guardia Parque Mataracú, Río Yapacani, 17.52072° S, 63.86795° W, 329 m, 26.xi.2004, Roberson, Garcia, Vidaurre– 1 male (UMSP).   
 Etymology —This species is named M. chalalan, used as a noun in apposition, for the name of the Ecolodge in Madidi National Park where the type specimen was collected.
description: Published as part of Blahnik, Roger J. & Holzenthal, Ralph W., 2017, Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *, pp. 1-251 in Insecta Mundi 2017 (602) on pages 72-73, DOI: 10.5281/zenodo.5170203