Forcipomyia Szadziewski, Dominiak &amp; Withers, 2020, sp. nov.
Creators: name: Szadziewski, Ryszard
name: Dominiak, Patrycja
name: Withers, Phil
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Ceratopogonidae
Forcipomyia

description: <i>Forcipomyia</i> (<i>F</i>.) <i>pyrenaica</i> sp. nov. (Figs 1–3) <b> <i>Diagnosis</i>.</b> The new species can be easily distinguished by having very low tarsal ratio of hind leg TR(3) 0.45, palpomere 3 enlarged on proximal 1 / 3, parameres separated, stout on proximal 2 / 3 and filamentous on distal 1 / 3, aedeagus with triangular median projection and distinct lateral shoulders. Female unknown. <b> <i>Description</i>.</b> Male. Body dark brown with pale parts of legs and abdomen (Fig. 2a). Head dark brown. Eyes bare, touching at distance of 6 ommatidia. Frons with distinct tubercle bearing stout, black spine-like seta (with a barb) in submedian position (Fig. 1c). Flagellum with 13 flagellomeres (Fig. 1a), length 1.10 mm, AR 1.11, distal flagellomeres 10–13 elongated, plume on flagellomeres 2–10 well developed, flagellomere 10 about 2.45 times longer than flagellomere 11. Palpus 5-segmented (Fig. 1b); palpomere 3 with distinct sensory pit on enlarged basal 1 / 3, length 0.14 mm, PR 4.0; palpomere 4 about 1.4 times longer than palpomere 5. Thorax including scutellum dark brown (Fig. 2a). Wing with two darker patches at anterior margin, on first radial cells and on radial fork (Fig. 2b); length 1.55 mm, CR 0.45; first radial cell slit-like, second one small; wing membrane with scale-like slender macrotrichia. Legs pale brownish, femur of hind leg brown (Fig. 2a), tarsomeres 4 distinctly longer than tarsomere 5, claws long, empodium well developed, TR (1) 0.63, TR(2) 0.36, TR(3) 0.45. Abdomen dark brown with pale caudal margins of tergites (Fig. 2a). Genitalia dark brown with pale gonostyli (Fig. 3a). Caudal margin of sternite 9 slightly convex with broad and shallow median incision. Gonocoxite without modified setae, gonostylus almost straight. Aedeagus shield-shaped, well sclerotized with well developed long triangular median projection and distinct lateral shoulders (Fig. 3b). Parameres separat- ed, stout, rod-like on proximal 2 / 3 and narrow, filamentous on distal 1 / 3 (Fig. 3c). Female unknown. <i>Material examined</i>. <i>Forcipomyia</i> (<i>F</i>.) <i>pyrenaica</i> sp. nov., holotype male, France, Pyrenees-Orientales, RNN foret de la Massane, Malaise trap, about 600 m a.s.l., 9.09.2008, P. Withers. The holotype is deposited in the Museum of Amber Inclusions, University of Gda&amp;nacute;sk, in the Collection of Extant Invertebrates. <b> <i>Etymology</i>.</b> The specific name refers to the Pyrenees mountains (Latin Pyrenaei) where the species was collected. <b> <i>Discussion</i>.</b> The new species is a typical member of the subgenus <i>Forcipomyia</i> s. str. (Alwin and Szadziewski 2013). The subgenus including almost 350 extant species is distributed worldwide (Borkent 2016). The low hind tarsal ratio (less than 0.5) is known among a few species of <i>Forcipomyia</i> s. str. outside of Europe. European species of the subgenus have higher hind TR, usually about 0.8–1.2 (Goetghebuer 1934, Remm 1962, 1980, Szadziewski 1983, Navai and Szadziewski 2016). In the holotype the tubercle of the frons in sub- median position has stout black spine-like seta with a distinct barb (Fig. 1c). In all, known for us, adults of biting midges this tubercle is devoid of setae and we treat this character as a developmental aberration. Similar setae (forked, with serrations, barbs or spurs) are known in immature stages (larvae and pupae) of <i>Forcipomyia</i> (Lewa&amp;nacute;czyk <i>et al</i>. 2009).
description: Published as part of <i>Szadziewski, Ryszard, Dominiak, Patrycja &amp; Withers, Phil, 2020, Two New Species Of Biting Midges From France And Algeria (Diptera: Ceratopogonidae), pp. 113-120 in Annales Zoologici 70 (1)</i> on pages 113-116, DOI: 10.3161/00034541ANZ2020.70.1.006, http://zenodo.org/record/3776615
description: {"references": ["Alwin, A. and R. Szadziewski. 2013. Biting midges of the subgenus Trichohelea of Forcipomyia in Poland, with keys for the determination of Polish subgenera (Diptera: Ceratopogonidae). Polish Journal of Entomology, 82: 113 - 126.", "Borkent, A. 2016. World species of biting midges (Diptera: Ceratopogonidae). Available from: http: // www. inhs. uiuc. edu / research / FLYTREE / CeratopogonidaeCatalog. pdf.", "Goetghebuer, M. 1934. 13 a. Heleidae (Ceratopogonidae), In: E. Lindner (ed.), Die Fliegen der palaearctishen Region 3, 1 - 48, pls 1 - 6, Lfg 77, Stuttgart.", "Remm, H. 1962. A survey of species of the genus Forcipomyia Meigen (Diptera, Heleidae) from Estonia. [In Russian, Estonian and English summary]. Loodusuurijata Seltsi Aastaraamat, 54: 165 - 195.", "Remm, H. 1980. New species of the family Ceratopogonidae (Diptera) from the Middle Asia [in Russian]. Tartu Riikliku Ulikooli Toimetised, 516: 85 - 128.", "Szadziewski, R. 1983. Ceratopogonidae (Diptera) from Algeria. II. New species, new records and new synonymy in the genus Forcipomyia Meig. Polskie Pismo Entomologiczne, 53: 363 - 384.", "Navai, S. and R. Szadziewski. 2016. Diptera Stelviana 2. 3.3. 05 Ceratopogonidae. Biting midges of the genus Forcipomyia Meigen (Diptera, Ceratopogonidae). Studia Dipterologica, Supplement 21: 65 - 95.", "Lewanczyk A., Szadziewski, R. and P. Dominiak. 2009. Diagnosis of Forcipomyia sahariensis Kieffer (Diptera: Ceratopogonidae) with the first description of immature stages. Fragmenta Faunistica, 52: 149 - 155."]}