Cochlostoma (Clessiniella) waldemari
Creators: name: Zallot, Enrico
name: Mattia, Willy De
name: Fehér, Zoltán
name: Gittenberger, Edmund
Publisher: name: Zenodo
Year: 2021
Subjects: Biodiversity
Taxonomy
Animalia
Mollusca
Gastropoda
Architaenioglossa
Cochlostomatidae
Cochlostoma
Cochlostoma waldemari

description: Cochlostoma (Clessiniella) waldemari (Wagner, 1897)
Figs 1, 3D, 14E, 15E, 16E, 31–33, 34 (top), 35 (top), 36 (bottom)
Pomatias (Auritus) waldemari A.J. Wagner, 1897: 616, pl. 9, 90 (“Ogulin”).
Cochlostoma waldemari – Welter-Schultes 2012: 103.
Diagnosis
Spotless shell. Moderately strong, more or less widely spaced ribs. Columellar lobe gently curved backwards. External lobe of type “II” in almost all the populations. The body spermiduct is huge and twisted instead of straight as in all the other species of Clessiniella.
Type material
Syntypes CROATIA • ♀; Ogulin; A.J. Wagner leg.; MIZPAS-W-6979a (Fig. 33) • 2 ♀♀, 3 ♂♂; same collection data as for preceding; MIZPAS-W-6979/5.
Other material
CROATIA • Kapela pass, Ogulin; 45.07534° N, 15.20837° E; 2009; De Mattia leg.; EZ0909 • Klek Mt, close to the top; 45.25904° N, 15.14044° E; 2009; Zallot leg.; EZ0904 • Klek Mt, cliffs halfway; 45.25438° N, 15.14561° E; 1996; Zallot leg.; EZ0905 • Mt Risnjak; 45.43095° N, 14.6188° E; 1400 m a.s.l.; 2009; De Mattia leg.; EZ0845 • Mt Risnjak; 45.39072° N, 14.61748° E; 1000 m a.s.l.; 2010; Zallot leg.; EZ0907 • Ročko Polje; 45.3607° N, 14.09534° E; 2009; De Mattia leg.; WdM6697 • Rudnica; 45.23402° N, 15.33827° E; 2009; De Mattia leg.; EZ0906 • Spilja Vrelo; 45.31777° N, 14.71065° E; 2009; Zallot leg.; EZ0899 • Vitunj-Ogulin; 45.29114° N, 15.14041° E; 2005; Zallot leg.; EZ0903.
SLOVENIA • Benete, 45.76192°N, 14.56752°E;2009; De Mattia leg.; WdM6805 • Bresovica; 45.95607°N, 14.43533° E; 2006; De Mattia leg.; EZ0897 • Bresovica pri Predgradu; 45.55164° N, 15.0437° E; 2009; De Mattia leg.; WdM6612 • Divje Jezero; 45.9827° N, 14.0277° E; 2009; De Mattia leg.; WdM6840 • Dol-E of Čepovan; 46.06445° N, 13.81189° E; 2009; De Mattia leg.; WdM6837 • Jelenja Vas; 45.5115° N, 15.05346° E; 2009; De Mattia leg.; WdM6804 • Krempa Kolpa; 45.53417° N, 15.09528° E; 1996; Mildner leg.; EZ0902 • Marija Ceplje Kolpa; 45.53444° N, 15.09389° E; 2006; Mildner leg.; EZ0901 • Mt Snežnik 1150; 45.56913° N, 14.321° E; Jun. 2010; De Mattia leg.; WdM7014 • Mt Snežnik 1620; 45.58845° N, 14.44209° E; Jun. 2010; De Mattia leg.; EZ0963 • Mt Snežnik car parking; 45.58235° N, 14.43117° E; Jun. 2010; De Mattia leg.; WdM7006 • Polog-E of Čepovan; 46.06387° N, 13.81275° E; 2009; De Mattia leg.; WdM6842 • SW Pescena Glava; 46.05156° N, 13.73844° E; 843 m a.s.l.; 2009; De Mattia leg.; WdM6829 • Tisovec; 45.81809° N, 14.76859° E; 2009; De Mattia leg.; WdM6749 • same collection data as for preceding; WdM6750 • Vrsic Pass; 46.44278° N, 13.76417° E; 1985; Zallot leg.; EZ0896 • Zagradec- Polyane; 45.86028° N, 14.83417° E; Bodon leg.; EZ0900.
Description
SHELL. The shell is more or less light corneous brown and spotless. The size of the aperture varies considerably among the populations; shells with a large aperture and a correspondingly large lip occur next to shells with a poorly developed lip and a relatively small aperture. The lip is gently curved backwards on the columellar side.
FEMALE GENITALIA. The female genitalia are hardly distinguishable from those of Co. (Cl.) tergestinum if not for the more relaxed loops.
MALE GENITALIA. There is a long, inflated penis. The penial spermiduct is slightly twisted and occupies only the central part of the penis. The body spermiduct is twisted and conspicuously protruding (see Figs 34–35).
Distribution
The species inhabits the north-western and south-eastern part of Slovenia and an adjacent area in Croatia. It is reported from Austria for the Karawanken and the Gailtaler Alpen (Edlinger & Mildner 1979). Its range largely surrounds that of Co. (Cl.) tergestinum (see Fig. 1).
Habitat preference
The snails are found on rocky cliffs as well as on boulders in woody areas.
Remarks
To assign a given population to Co. (Cl.) waldemari rather than to Co. (Cl.) tergestinum on the basis of the morphology of the shell and the female genitalia alone could be challenging if not impossible (see Figs 34–35). Only the structure of the male genitalia allows a convincing identification of the species. The H3 analysis (Fig. 7) confirms that these sibling species belong to separate clades within Clessiniella.
Boeckel (1939) considered Co. (Cl.) waldemari a subspecies of Co. (Cl.) tergestinum. This view was contradicted by Bole (1994). In some areas, like on Mount Snežnik in southern Slovenia, the two species live not far from each other and here they can be distinguished also conchologically while differing in the presence or absence of the spots on the teleoconch (with Co. (Cl.) waldemari having a spotless shell) and in the shape of the body whorl approaching the aperture, in lateral view (Fig. 36).
description: Published as part of Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán & Gittenberger, Edmund, 2021, Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae), pp. 49-95 in European Journal of Taxonomy 762 (1) on pages 87-89, DOI: 10.5852/ejt.2021.762.1453, http://zenodo.org/record/5176909