Coryphomimus levigatus Yin 2022, sp. nov.
Creators: name: Yin, Zi-Wei
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Staphylinidae
Coryphomimus
Coryphomimus levigatus

description: Coryphomimus levigatus sp. nov.
(Figs 42, 81B, 98C, D, 99F, 100A)
Chinese common name: 光ẚṈỀȁà甲
Type material (14 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xiang, Mêdog County, nr. 80K, 29°41’09”N, 95°30’10”E, alt. 2330 m, mixed leaf litter, sifted, 09.vii.2018, Cheng, Peng & Shen leg. (中ǞƱẪẸṘ县 80K ṉ ữ)’ (SNUC). PARATYPES: CHINA: 2 ♂♂, 7 ♀♀, same data as that of holotype; 1 ♂, 2 ♀♀, ‘ China: Mêdog County, pass from Bari Village to Renqingbeng Temple, 29°18’45”N, 95°21’27”E, ca. 1830 m, 19.iii.2017, X.-B. Song leg. [ƱẪẸṘ县巴U村 -仁ů崩寺]’; 1 ♀ ‘ China: Xizang A. R., Motuo Co., Hanmi, alt. 2200 m, 19.viii.2006, Tang Liang leg.’ (all paratypes in SNUC).
Diagnosis. Male. Body length 2.16–2.20 mm; dorsal surface of body with scattered exceptionally long setae and normal pubescence. Head subtruncate at base, narrower than pronotum, tempus moderately short, rounded at posterolateral angle, vertex lacking sulcus, with relatively small, setose foveae, with long mediobasal carina extending from head base anteriorly to frons, with thin lateral carinae from subbase to posterior margin of strongly raised antennal tubercles; antenna elongate, with modified antennomeres 10–11, antennomere 10 with moderately large basal cavity on ventral surface, 11 with short basal projection. Pronotum with laterally carinate median longitudinal sulcus, with pairs of discal and short lateral carinae, lacking antebasal or marginal spines. Discal stria of elytron long, extending posteriorly to more than apical 1/5 of elytral length; disc finely punctate. Pro- and mesotrochanter and mesofemur with small, blunt ventral spine; metatrochanter with short, apically curved projection on ventral margin. Aedeagus strongly asymmetric, stout; median lobe with large basal capsule and slightly oval foramen, ventral stalk broad at middle and then abruptly narrowing apically on left side, dorsal lobe embracing median lobe, non-articulated, with elongate sclerite forked at apex. Female. Body length 1.89–2.04 mm; antenna lacking modifications, legs lacking spine or projection, genitalia as in Fig. 42J.
Description. Male. Body (Fig. 42A) length 2.16–2.20 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence, with scattered exceptionally long setae present on vertex, lateral margins and discs of pronotum and elytra, meso- and metatibia and tergites 1–4 (IV–VII).
Head (Fig. 42B) subtruncate at base, much wider than long, length 0.43 mm, width across eyes 0.51–0.52 mm; vertex smooth, markedly raised at middle, lacking sulcus, vertexal foveae (dorsal tentorial pits) setose, relatively small, mediobasal carina distinct and long, extending from head base anteriorly to level of bases of antennal tubercles, lateral carina thin, extending from subbase anteriorly to base of antennal tubercle; posterolateral angle rounded; frons anteriorly demarcated from clypeus by frontal-clypeal ridge, weakly impressed between large, strongly raised antennal tubercles; clypeus with smooth surface, its entire anterior margin strongly carinate and moderately raised; ocular-mandibular carina complete. Venter with smooth surface; small gular foveae (posterior tentorial pits) originating from shared oval opening, with median carina faint from opening and more distinct near mouthparts. Compound eyes prominent, composed of approximately 40 ommatidia. Antenna elongate, length 1.25 mm, distinct club (Fig. 42C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each slightly elongate, 8 smallest, moniliform, 9 much wider and longer than 8, sub-moniliform, 10 (Fig. 42D) much broader and longer than 9, asymmetric, ventral surface with moderately large cavity at base, 11 largest, much longer than 9 and 10 combined, sub-fusiform, with short and broad projection at base of mesal margin.
Pronotum (Fig. 42B) wider than long, length 0.49–0.50 mm, width 0.58 mm, widest anterior middle; lateral margins rounded, strongly incised posterior to lateral humps, convergent basally and sub-parallel at basal 1/4; disc slightly convex, finely punctate, broad median longitudinal sulcus with carinate sides, posteriorly confluent with oval antebasal impression and short mediobasal carina, with pair of discal and short lateral longitudinal carinae, lacking antebasal, discal, or marginal spines/tubercles; with large, setose anterolateral and lateral antebasal foveae; with small outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit and large, setose antero-hypomeral fovea; margin of coxal cavity broadly carinate.
Elytra much wider than long, length 0.63 mm, width 0.75 mm; each elytron with three moderately large, asetose basal foveae, inner two close; long discal stria extending posteriorly from outer basal fovea to more than apical 1/5 of elytral length; humerus moderately prominent, subhumeral fovea present, carinate marginal stria extending from fovea to posterior margin of elytron.
Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, originating from shared opening, large lateral mesoventral foveae not forked internally, with short mesoventral process, with complete marginal stria. Metaventrite prominent admesally, inclined towards middle, with well-developed lateral mesocoxal foveae, two lateral metaventral foveae moving medially into large, setose shared impression; posterior margin weakly emarginate at middle, with thin split.
Legs elongate, femora; procoxa with exceptionally long seta at base, protrochanter with blunt ventral spine; mesotrochanter (Fig. 42E) with distinct blunt spine on ventral margin, mesofemur (Fig. 42E) with small ventral denticle near base; metatrochanter (Fig. 42F) with broad, apically curved projection on ventral margin, metatibia with short pencil-like apical tuft of setae.
Abdomen widest at lateral margins of tergite 1 (IV), length 0.64–0.69 mm, width 0.66–0.68 mm. Tergite 1 (IV) more than twice as long as 2 (V), with large mediobasal foveae at lateral margins of short median sulcus, with one pair of large basolateral foveae, lacking discal carinae, marginal carinae complete, oblique inner carina thinner than outer one; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of small basolateral foveae and pair of thin lateral carinae, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lacking lateral carina; midlength of sternites 2–4 (IV–VI) gradually shorter, 5 (VII) slightly longer than 4, sternites 3–4 with tiny tubercle at middle, 3–5 each with two pairs of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 42G) weakly sclerotized, slightly oval.
Aedeagus (Fig. 42H, I) 0.41 mm long, in dorsal view asymmetric, stout; median lobe with broad basal capsule and slightly oval foramen, with long basoventral projection, ventral stalk in dorsal view broad at middle and abruptly narrowing apically at left side, with rounded, membranous apex; dorsal lobe broad, embracing median lobe, with elongate sclerite apically forked; parameres membranous, apically split into two lobes.
Female. Similar to male in external morphology; antenna slightly shorter; antenna lacking modifications, legs lacking spines or projections; each compound eye composed of approximately 13 ommatidia; sternite 2 (IV) with one pair of mediobasal and two pairs of basolateral foveae. Measurements (as for male): body length 1.89–2.04 mm; length/width of head 0.41/ 0.49 mm, pronotum 0.44–0.46/ 0.54–0.56 mm, elytra 0.49–0.52/ 0.67–0.71 mm; abdomen 0.61–0.66/ 0.67–0.70 mm; length of antenna 1.00– 1.10 mm; genitalia (Fig. 42J) weakly sclerotized, transverse, maximum width 0.36 mm.
Distribution. Mêdog County, Tibet, SW China (Figs 81B, 98C, D, 99F, 100A).
Etymology. The specific epithet ‘ levigatus (- a, - um)’ is a Latin adjective meaning ‘smoothed, polished’, referring to the smooth, aspinose lateral margins of the pronotum of this species.
description: Published as part of Yin, Zi-Wei, 2022, The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth's " Third Pole " (Coleoptera, Staphylinidae, Pselaphinae), pp. 1-211 in Zootaxa 5111 (1) on pages 98-99, DOI: 10.11646/zootaxa.5111.1.1, http://zenodo.org/record/6340729