Caracladus daguanshanensis Muhammad & Wang & Zhang 2023, sp. n.
Creators: name: Muhammad, Irfan
name: Wang, Lu-Yu
name: Zhang, Zhi-Sheng
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Arachnida
Araneae
Linyphiidae
Caracladus
Caracladus daguanshanensis

description: Caracladus daguanshanensis sp. n. (kḂXĿĦ)
Figures 1–4
Type material. Holotype male: CHINA, Chongqing, Wuxi County, YNR, Daguanshan, 31°29′43.91″N, 109°43′44.17″E, elev. 2404 m, 2.IX.2020, Z.S. Zhang et al. leg. (SWUC-T-LIN-25-01). Paratypes (6 females): 2 females, same data as holotype (SWUC-T-LIN-25-02~03); 1 female, YNR, Guanshan Forest farm, Huajiaya, 31°31′33.83″N, 109°42′17.11″E, elev. 2192 m, 1.IX.2020, Z.S. Zhang et al. leg. (SWUC-T-LIN-25-04); 3 females, YNR, Guanshan Forest farm, Huajiaya, Yezhuping, 31°29′43.30″N, 109°43′44.55″E, elev. 2411 m, 9.IV.2022, Z.S. Zhang et al. leg. (SWUC-T-LIN-25-05~07).
Etymology. This epithet is derived from the type locality.
Diagnosis. The new species resembles Caracladus montanus Sha & Zhu, 1994 and C. tsurusakii Saito, 1988 in having similar ventral plate of epigyne with broad atrium and dorsal plate extending anteriorly inside the atrium of epigyne (Fig. 2A, B; Sha & Zhu 1994, fig. 6; Frick & Muff 2009, fig. 40) and male palp with similar embolic division (Fig. 1A–D; Sha & Zhu 1994, fig. 3; Frick & Muff 2009, fig. 41), but can be distinguished by the distal end of cephalic lobe straight in C. daguanshanensis sp. n. (Fig. 3B), vs., distally curved both in C. montanus (Sha & Zhu 1994, fig. 2) and C. tsurusakii (Frick & Muff 2009, figs 44, 45). Moreover, in the male palp of C. daguanshanensis sp. nov., the dorsal tibial apophysis with two arms (Fig. 1C), vs., with two apophyses, prolateral tibial apophysis and retrolateral tibial apophysis in C. montanus (Sha & Zhu 1994, fig. 5) and C. tsurusakii (Frick & Muff 2009, fig. 42); ventral surface of dorsal tibial apophysis with several teeth in C. daguanshanensis sp. n. (Fig. 1B), vs., teeth absent in C. montanus (Sha & Zhu 1994, fig. 3) and C. tsurusakii (Saito 1988, fig. 14). The inner margin of ventral plate of epigyne trapezoid in C. daguanshanensis sp. n. (Fig. 2A, B), vs., almost rectangular in C. montanus (Sha & Zhu 1994, fig. 6) and oval in C. tsurusakii (Frick & Muff 2009, fig. 40).
Description. Male (holotype, Figs 3A, B, D, 4A, B): Total length: 2.38. Carapace 1.37 long, 0.72 wide, yellowish-brown, cephalic lobe 0.49 long, projecting forward, narrowing towards tip with sparse stout setae; cephalic pits absent; thoracic region slightly elevated, fovea, cervical and radial grooves distinct. Clypeus 0.24 high, directed obliquely anteriorly. Chelicerae with 5 promarginal and 5 retromarginal teeth. AER straight, present at the base of cephalic lobe, PMEs present on the dorsal surface of the cephalic lobe. Eye sizes and interdistances: AME 0.04, ALE 0.05, PME 0.04, PLE 0.05, AME–AME 0.01, PME–PME 0.04, AME–ALE, 0.02, PME–PLE 0.19, ALE–ALE 0.17, PLE–PLE 0.18, ALE–PLE contiguous. Length of legs: I 2.44 (0.74, 0.82, 0.47, 0.41), II 2.19 (0.66, 0.76, 0.43, 0.34), III 1.91 (0.57, 0.63, 0.40, 0.31), IV 2.46 (0.76, 0.85, 0.48, 0.37). TmI 0.57. Tibial spine formula: 1-1-1-1. Abdomen 1.01 long, 0.76 wide, oval, grey, dorsally with two pairs of sigillae, ventral side grey.
Palp (holotype, Fig. 1 A−D): Patella two times longer than wide, ventrally slightly curved; tibia with two dorsal trichobothria, dorsal tibial apophysis with two arms; paracymbium simple, J-shaped; tegulum with short and long protegular papillae on protegulum; suprategulum with median tooth, distal suprategular apophysis sclerotized, distal end curved with pointed end; embolic membrane slender, running along the embolus; radix simple, tailpiece footshaped in prolateral view; embolus whip-like, gradually narrowing towards tip..
Female (Figs 3C, 4C): Total length: 2.01 mm. Carapace 0.97 long, 0.69 wide, yellowish-brown, cephalic region slightly elevated, fovea, cervical and radial grooves distinct. Clypeus 0.14 high. Chelicerae with 5 promarginal and 5 retromarginal teeth. AER straight, PER procurved, slightly wider than AER. Eye sizes and interdistances: AME 0.04, ALE 0.07, PME 0.06, PLE 0.06, AME–AME 0.02, PME–PME 0.04, AME–ALE, 0.02, PME–PLE 0.05, ALE–ALE 0.24, PLE–PLE 0.28, ALE–PLE contiguous. Length of legs: I 2.44 (0.71, 0.80, 0.47, 0.39), II 2.18 (0.66, 0.73, 0.44, 0.35), III 1.98 (0.61, 0.65, 0.39, 0.33), IV 2.46 (0.74, 0.87, 0.50, 0.35). TmI 0.41. Tibial spine formula: 1-1-1-1. Abdomen 1.01 long, 0.76 wide, oval, grey, dorsally with two pairs of sigillae, ventral side grey.
Epigyne (Fig. 2 A−C): Ventral plate wider than long, with broad trapezoid opening at the center; dorsal plate extending anteriorly inside the atrium; copulatory opening at anterior end of trapezoid dorsal plate.Vulva: Copulatory pouch almost round, posteriorly connected with copulatory ducts; copulatory ducts short, relatively sclerotized; spermathecae globular, pointing dorso-laterally, separated by distance one and half times longer than their diameter; fertilization ducts extending mesally.
Distribution. Known only from the type locality.
description: Published as part of Muhammad, Irfan, Wang, Lu-Yu & Zhang, Zhi-Sheng, 2023, One new genus and nine new species of Linyphiidae spiders from Yintiaoling Nature Reserve, Chongqing of China, pp. 82-114 in Zootaxa 5257 (1) on pages 83-87, DOI: 10.11646/zootaxa.5257.1.7, http://zenodo.org/record/7766060