Tomarus rosettae
Creators: name: Neita-Moreno, Jhon César
name: Alexander, Recursos Biológicos
name: Agustín, San
name: de, Villa
name: Leyva
name: Boyacá
name: Ratcliffe, Brett C.
name: Collections, Systematics Research
name: of, University
name: Museum, Nebraska State
name: Hall, Nebraska
name: of, University
name: Nebraska
name: Lincoln
name: Ne
name: Usa
Publisher: name: Zenodo
Year: 2017
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Coleoptera
Dynastidae
Tomarus
Tomarus rosettae

description: Tomarus rosettae (Endrödi, 1968)   
(Fig. 1d, 2a, 3i, 4f, 5f, 6k–l, 7f, 8f, 10k–l, 11f, 12f, 13f, 14f, 15f, 17b)  
 Ligyrus rosettae Endrödi 1968: 168 (original combination).   
 Redescription. Male. Length 19.4–21.5 mm; width 9.4–10.8 mm. Color dark reddish brown to black. Head: Frons and clypeus coarsely rugose. Frontoclypeal region with 2 distinctly transverse tubercles; tubercles relatively narrowly separated, usually about 2.5 tubercle diameters apart (Fig. 4f). Clypeus strongly rugopunctate to transversely rugose, tapering to broad, slightly emarginate apex with 2 reflexed teeth. Mandibles with 2 teeth, apical tooth rounded, subapical notch shallow (Fig. 6k–l); labium densely setose on sides, paraglossa developed, apex with small tooth at center (Fig. 7f); maxilla with galea developed and with 6 teeth (Fig. 5f). Pronotum: Surface with disc moderately punctate, sides and anterior angles densely punctate to rugopunctate; punctures moderate in size on disc, becoming larger laterally; fovea rugose within. Anterior margin with strong, median tubercle; fovea small, as wide as width between eyes (Fig. 8f). Elytra: Surface densely punctate; punctures moderate in size with sparser, minute punctures between larger punctures; larger punctures usually ocellate, and with 3 pairs of double rows. Pygidium: Surface densely punctate, punctures becoming denser in basal and lateral angles; punctures small to moderate in size, weakly ocellate. In lateral view, male with convex surface, surface weakly convex in females. Legs: Protibia tridentate and with weak, basal convexity suggestive of 4 th tooth, teeth subequally spaced from each other. Protarsus in male simple, not enlarged. Mesotibia relatively broad, transverse carinae arcuate, spinules long, stout (Fig. 10k–l). Metatibia with 1 basal, transverses carina interrupted at middle (carina with spine-like setae) and 1 medial, transverse carina; apex truncate with about 21 spinules (Fig. 10l). Venter: Prosternal process long, thick, apex longitudinally oval. Mesosternal surface glabrous, slightly concave at middle. Metasternal surface with sparse, ocellate punctures. Male Genitalia: Parameres (Fig. 11f, 12f, 13f) subtriangular, acute apices curving outwards. Internal sac with 3 copulatory lamellae (Fig. 15f).   
 Female. Length 20.0– 21.6 mm; width 9.8–10.0 mm. Similar to male, but pronotal surface with punctures slightly less dense. Gonocoxite larger than gonocoxal sternite, with 2 semicircular furrows (Fig. 14f).
description: Published as part of Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, pp. 1-36 in Insecta Mundi 2017 (547) on page 10, DOI: 10.5281/zenodo.5172580