Enicospilus centraliscleritiger Pham & Pham & Matsumoto & Shimizu & Broad 2023, sp. nov.
Creators: name: Pham, Nhi Thi
name: Pham, Phu Van
name: Matsumoto, Rikio
name: Shimizu, So
name: Broad, Gavin R.
Publisher: name: Zenodo
Year: 2023
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Enicospilus
Enicospilus centraliscleritiger

description: Enicospilus centraliscleritiger sp. nov.
urn:lsid:zoobank.org:act: EC337820-8FBE-4ED4-B8C9-F0E9494E83C1
Fig. 16
Diagnosis
Interocellar area reddish brown; clypeus convex, ventral margin blunt; mandible twisted ca 20°, outer surface without a diagonal setose groove; fore wing fenestra with only central sclerite present, small, circular to oval, vein 1m-cu&M sinuate, angled medially.
Differential diagnosis
The new species has only one circular to oval central sclerite similar to E. abessyniensis (Szépligeti, 1907) from the Afrotropical region and Sri Lanka. It differs from the latter by the position of the central sclerite (centrodistal vs central), vein 2r&RS less sinuate and vein 1m-cu&M strongly sinuate, angled medially.
Etymology
The adjective belonging to the Latin neuter noun ‘ centrum ʼ is ‘ centralis, - is, - e ʼ. Combined with the sclerite, the specific epither ‘ centraliscleritiger ʼ meaning ‘bearing a central scleriteʼ, referring to the appearance of the fore wing, which has only a central sclerite.
Material examined
Holotype
VIETNAM • ♀; Tuyen Quang Province, Na Hang, Son Phu; 10 Jun. 2017; Khuat D.L. leg.; Malaise trap; IEBR.
Paratype
VIETNAM • 1 ♂; Gia Lai Province, Kon Chu Rang NP; 27 Apr. 2016; Nguyen T.P.L. leg.; light trap; IEBR.
Description
Female (holotype)
MEASUREMENTS. Body length 26.1 mm, fore wing length 17.5 mm.
HEAD with FI = 0.7, GOI = 4.0 (Fig. 16d). Lower face 0.7 × as wide as high, shiny, finely punctate (Fig. 16b). Clypeus 1.4× as wide as high, moderately convex in profile, sparsely punctate, ventral margin blunt (Fig. 16b). Malar space 0.3× as long as basal mandibular width (Fig. 16b). Mandible twisted by ca 20°, evenly narrowed, outer surface without diagonal setose groove. Upper tooth of mandible about 1.3× as long as lower tooth (Fig. 16b). Frons, vertex and gena shiny with fine setae (Fig. 16c). Posterior ocellus close to eye (Fig. 16c). Occipital carina complete, ventral end meeting hypostomal carina at about 1.0× basal mandible width from base of mandible. Antenna with 65 flagellomeres; F1 1.9× as long as F2; F20 2.3 × as long as wide.
MESOSOMA. Moderately shiny (Fig. 16e). Pronotum finely striate (Fig. 16e). Mesoscutum 1.4× as long as maximum width, evenly rounded in profile. Notaulus absent. Scutellum moderately convex, with irregular rugae, lateral longitudinal carinae along entire length of scutellum. Epicnemium matt, densely punctate (Fig. 16e). Epicnemial carina moderately strong, dorsal end bent to reach anterior margin of mesopleuron (Fig. 16e). Mesopleuron entirely transversely striate (Fig. 16e). Submetapleural carina broadened anteriorly (Fig. 16e). Metapleuron strigose (Fig. 16e). Propodeum evenly rounded in profile; anterior transverse carina complete; anterior area longitudinally striate; spiracular area smooth; posterior area coarsely rugose; propodeal spiracle elliptical, joined by ridge to pleural carina (Fig. 16e).
WINGS (Fig. 16f). Fore wing with AI = 0.83, CI = 0.40, DI = 0.39, ICI = 0.83, SDI = 1.43, SI = 0.24, SRI = 0.34; vein 1m-cu&M strongly sinuate, angled medially; vein 2r&RS slightly sinuate; vein RS rather evenly curved; fenestra and sclerites of discosubmarginal cell as in Fig. 16f; proximal and distal sclerites absent; central sclerite nearly oval in shape, pigmented, positioned in centrodistal part of fenestra; proximal corner of marginal cell evenly setose; posterodistal corner of second discal cell ca 100° and of subbasal cell ca 95°; vein 1cu-a antefurcal to M&RS by 0.2× length of 1cu-a. Hind wing with NI = 2.0; vein RS relatively straight; vein RA with 9 uniform hamuli.
LEGS. Hind leg with coxa in profile 1.8× as long as high; basitarsus 2.0× as long as second tarsomere; fourth tarsomere 2.6× as long as wide; tarsal claws simply pectinate.
METASOMA. With DMI = 1.4, PI = 2.8, THI = 2.7; thyridium oval; ovipositor sheath not longer than posterior height of metasoma.
COLOUR (Fig. 16a). Reddish brown except for apex of mandible and posterior 0.6 of T3 onwards black. Wings hyaline; central sclerite pigmented and amber; veins and pterostigma dark brown.
Variation in female
Unknown.
Male
Similar to female, except for following characters: F1 2.0× as long as F2, F20 1.8 × as long as wide, FI = 0.7; fore wing length 15.8 mm, AI = 0.90, CI = 0.44, DI = 0.40, ICI = 0.90, SDI = 1.60, SI = 0.23, SRI = 0.27, vein 1m-cu&M with short ramellus, central sclerite circular; metasoma with DMI = 1.3, PI = 2.6, THI = 3.0.
Distribution
Currently known from Tuyen Quang Province in the northeast and Gia Lai Province, the Central Highlands of Vietnam.
description: Published as part of Pham, Nhi Thi, Pham, Phu Van, Matsumoto, Rikio, Shimizu, So & Broad, Gavin R., 2023, A review of the genus Enicospilus Stephens (Ichneumonidae: Ophioninae) from Vietnam, with descriptions of ten new species, pp. 1-151 in European Journal of Taxonomy 873 (1) on pages 37-39, DOI: 10.5852/ejt.2023.873.2133, http://zenodo.org/record/8038022