Trieces confusus Mazón & Bordera 2020, sp. nov.
Creators: name: Mazón, Marina
name: Bordera, Santiago
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Hymenoptera
Ichneumonidae
Trieces
Trieces confusus

description: Trieces confusus sp. nov.
(Figs 1, 2, 3)
Material Examined: Holotype. Spain: ♀, P.N. Cabañeros (Ciudad Real), Abierto, Raña del Pocico, TM 1, 18.III/12. IV.2005, #2095, leg. Mazón & Bordera (CEUA). Paratypes: Spain: 2♀, Alicante, Alcoy, Font Roja Menejador, 1352 m, TM, 23. V/5. VI.2001 (CEUA); 1♀, P.N. Cabañeros (Ciudad Real), Abierto, Raña del Pocico, TM 1, 12.IX/8. X.2004, #1515; 1♂, same data, #1516, leg. Mazón & Bordera (CEUA).
Diagnosis: Trieces confusus can be placed within the texanus species group sensu Tolkanitz (2010) because of metapleuron lacking a vertical slit-shaped depression posteriorly and having dorsal hairs in the posterodorsal area of metapleuron (Fig. 2E). It can be distinguished from all other Palaearctic species by the combination of the following characters: lateral longitudinal carinae conspicuous and complete only on tergite I (Figs 1D, 1F, 2A, 2C, 3D, 3F); eyes strongly converging towards mandibles in female (Fig. 1B), with short and sparse setae (Fig. 2F); propodeal spiracle not joining pleural carina (Figs 1G, 2B, 2E, 3G); metapleuron smooth and shiny, with a few and sparse setiferous punctures on the upper edge, under the pleural carina, and on the hind lower corner, short wrinkles are present only on its hind part, in front of hind coxa insertion (Fig. 2E); legs dark brown to black, with yellow marks on tibiae and tarsus (Figs 1A, 3A).
Description: Female: Body length (without ovipositor) 3.8–4.0 mm. Fore wing 2.5-2.6 mm long.
Head (Figs 1B, 1C, 1G, 2F).—Transverse, 2.3–2.6 × as wide as long, barely constricted behind the eyes, slightly rounded (Fig. 1C). Gena 0.8–1.1 × as long as eye (viewed from above). Vertex high, the back of the head strongly declivous behind the posterior ocelli. Frons barely convex (Fig. 1G). Vertex, face and frons granulate but shiny. Gena sparsely and finely punctured on a smooth and shiny background. Malar space short, strongly granulate, about 0.45–0.6 × as long as basal width of mandible. Face and clypeus convex (in lateral view), face strongly protruding (Fig. 1G), clypeal suture absent, clypeus with apical margin truncate. Interantennal process short, ending in a right angle (Fig. 1B). Eyes strongly converging towards mandibles (Fig. 1B) with short and sparse hairs (Fig. 2F). Mandible barely narrowed towards apex, teeth of apparently equal length. Maxillary palp 5-segmented, the palpomeres slender; labial palp 4-segmented, the second palpomere slightly inflated, the distal two quite slender. Antenna with 21–22 flagellomeres, first flagellomere 3.0–3.1 × as long as wide, second one 1.5–1.8 ×. Hind ocelli separated from eye about 1.3–1.7 × their diameter. Space between hind ocelli 2.3–2.7 × as long as their diameter. Occipital carina present, medially situated far behind ocelli (Fig. 1C). Head covered by soft hairs.
Mesosoma (Figs 1E, 1G, 2B, 2D, 2E).—Pronotum smooth and polished in its lower part, strongly concave, convex and finely punctate on upper part (Fig. 2B). Epomia absent (Fig. 2B). Mesoscutum slightly ovoid from above, clearly convex, very finely and densely punctate on a smooth and shiny background. Notauli absent (Fig. 1E). Prescutellar groove smooth, without longitudinal wrinkles. Scutellum convex, same sculpture than mesoscutum but punctures smaller, completely delimited by strong carinae. Mesopleuron strongly convex, shiny and punctate, hind part polished (Figs 1G, 2B). Epicnemial carina weak, reaching a little above centre of pronotum (Fig. 2D). Mesopleural fovea moderately deep, mesopleural suture absent (Fig. 2B). Metapleuron smooth and shiny, with a few and sparse setiferous punctures on the upper edge, under the pleural carina, and on the hind upper and lower corner; short wrinkles are present only on its hind part, in front of the hind coxa. Metapleural fovea weak and small (Figs 2B, 2E). Propodeum with lateromedian, lateral longitudinal and posterior transverse carinae present, punctate on shiny and strongly granulate background (Figs 1E, 2B); propodeal spiracle elliptic, slightly protruding, not joining pleural carina (Fig. 2B). Hind femur 2.6–2.7 × as long as high. Hind tibia short, 3.9–4.1 × longer than wide; tibial spurs strongly asymmetric, inner spur about 4.0 × longer than outer spur. Tarsal claws not pectinate.
Wings (Fig. 1A).—Fore wing with ramulus absent; vein 3 rs-m absent, thus areolet open; vein 2 m-cu vertical, reaching areolet at a distance from 2 rs-m approximately equal to its length, with a single bulla situated slightly above the middle; vein Cu-a distal to Rs&M; abscissa of Cu 1 between 1 m-cu and Cu 1 a as long as Cu 1 b; Cu 1 b reclivous, slightly angular to clearly curved. Hind wing with vein cu-a + first abscissa of Cu 1 between M and cu-a not intercepted by distal abscissa of Cu 1, which is absent.
Metasoma (Figs 1D, 1F, 2A, 2C).—Tergite I 0.8–1.0 × as long as wide, spiracle situated just below lateral longitudinal carina, joining it; lateromedian longitudinal carinae conspicuous, parallel, slightly divergent on its basal part and reaching the posterior margin of tergite (Figs 1D, 2C); lateral longitudinal carinae conspicuous and complete. Tergites II–III without lateromedian and lateral longitudinal carinae, with traces of a weak single median carina (Fig. 1D), sometimes absent (Fig. 2C). Tergites II–VIII clearly transverse (Fig. 1D), tergite II 0.67–0.8 × as long as wide. All tergites strongly and very densely punctate on smooth and shiny background (Figs 1D, 2C). Ovipositor short and straight, nodus present but not clearly distinct, without teeth; apex gradually tapered. Ovipositor sheaths nearly as long as the tergite I.
Colouration (Fig. 1). — Body black. Face from interantennal process, clypeus, malar space and mandibles (except teeth), palpi, tegulae, and mostly all tibiae and tarsi, yellow. Scape, pedicel and flagellomeres beneath, light brown. Coxae and femora dark brown to black.
Male: Differs from female as follows:
Body length 4.0 mm. Fore wing 2.5 mm long.
Head (Figs 3B, 3C, 3G).—Malar space long, about twice as wide as basal width of mandible. Eyes not converging towards mandibles (Fig. 3B). Antenna with 23 flagellomeres (Fig. 1A), without tyloids; first flagellomere about 2.6 × as long as its maximum width, second one 2.0 ×. Hind ocelli separated from eye by about 1.9 × their diameter. Space between hind ocelli 2.7 × their diameter (Fig. 3C).
Mesosoma (Figs 3A, 3E, 3G).—Hind femur 2.4 × as long as high. Inner spur of tibiae a little longer than in female, reaching about basal third of outer one.
Colouration (Fig. 3).—Yellow coloration of head only represented at mandibles (except teeth). Upper face reddish in the center. Clypeus and malar space black.
Etymology: The species name refers to the possession of characters from both Trieces and Chorinaeus.
Hosts: Unknown.
description: Published as part of Mazón, Marina & Bordera, Santiago, 2020, Discovery of a new species of Trieces (Hymenoptera: Ichneumonidae: Metopiinae) questioning the traditional delimitation between Trieces and Chorinaeus, pp. 594-600 in Zootaxa 4821 (3) on pages 596-599, DOI: 10.11646/zootaxa.4821.3.10, http://zenodo.org/record/4400925