Cardiodactylus halmahera Gorochov & Robillard 2014, n. sp.
Creators: name: Robillard, Tony
name: Gorochov, Andrej V.
name: Poulain, Simon
name: Suhardjono, Yayuk R.
Publisher: name: Zenodo
Year: 2014
Subjects: Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Orthoptera
Haglotettigoniidae
Cardiodactylus
Cardiodactylus halmahera

description: Cardiodactylus halmahera Gorochov & Robillard, n. sp.
(Figs 4, 5H, 13F, 14F, 27)
Type material. Male holotype: Indonesia. Maluku Utara Province, Halmahera I., environs of Subaim Vill. (to S from Lolobata Vill. but not far from it) near coast of Wasile Bay, low hills, disturbed forest, on leaf of bush, at night, 27.I–1.II.2011, A. Gorochov (ZIN). Paratypes (1♂, 3♀): same information as holotype, 1♂, 1♀ (ZIN); 1♀ (MNHN-EO-ENSIF3655). Halmahera [I.] Noord-Oost [North-West], Bernstein, 1♀ (RMNH).
Type locality. Indonesia, Halmahera Island, environs of Subaim Village (to S from Lolobata Village but not far from it) near coast of Wasile Bay.
Etymology. Named after type locality.
Distribution. Indonesia, Halmahera Island.
Diagnosis. The new species is characterized by the male epiphallus with assymetrical dorsal ridges folded laterally, close to that of C. kotandora and with a rather deep notch at middle in profile view.
Description. Male (holotype): General appearance similar to that of C. empagatao but with following differences: face of head with a pair of larger dark brown spots under antennal cavities connected with each other by reversed U-shaped brown narrow stripe almost reaching rostral apex, and with rather long dark stripe along clypeal suture; dorsum of head almost completely brown (including dorsal surface of rostrum) with narrow light brown stripe outlined anterior part of this dorsum, and with longitudinal yellowish stripe behind each eye (Fig. 27A–C); pronotum with dark brown most part of lateral lobes and of disk (only 2 pairs of longitudinal light stripes along lateral edges of disk and along ventral edges of lateral lobes developed; Fig. 27B–C); hind tibiae with somewhat more numerous darkish spots; tegmina with slightly darker most part of distal half of dorsal field and more contrast (white) light marks, but cell membranes in region of large light spot at base of apical area slightly darker, with grayish tinge; Fig. 27D–E); hind wings with more distinct (light brown) venation; abdominal tergites somewhat darker (mainly grayish brown), and almost all sternites of almost same color (however with lateral parts light brown); wings somewhat longer, with narrower mirror (ratio length/width about 2) and 11–12 branches on Sc (Fig. 27E). Genitalia distinguished by assymetrical epiphallic dorsal ridges, they dorsal edge folded laterally, and by presence of rather deep notch at middle in profile view (Fig. 27H–K).
Female: General appearance similar to that of holotype, but sometimes hind part of head dorsum slightly lighter and face without dark stripe along clypeal suture, tegmina in all females brown with less contrast light marks similar to those of female of C. empagatao and with additional yellowish spot between basal and lateral light spots on dorsal field but not along lateral edge of this field (area between this additional spot and basal spot almost dark brown) (Fig. 27F–G). Sc comb with 8–9 branches. Ovipositor short (hind femur 1.2–1.3 times as long as ovipositor) (Fig. 13F).
Female genitalia (Fig. 14F): Copulatory papilla more or less triangular, with small baso-lateral sclerites, its apex elongate, slightly sclerotized, folded ventrally.
Juvenile: unknown.
Variations. Second male somewhat lighter and with less contrasted coloration of head, pronotum and tegmina; its genitalia with some parts less sclerotized, and with notch on right epiphallic ridge less deep (this male possibly collected very shortly after moulting).
Measurements. Length in mm. Body: male 20–22, female 22–23; body with wings: male 27–31, female 31–33; pronotum: male 3–3.3, female 3.5–3.7; tegmina: male 15.5–18, female 18.5–19.5; hind femora: male 18.5–20.5, female 20–21; ovipositor 16–17.
Habitat and life history traits. Unknown.
Behavior. Unknown.
description: Published as part of Robillard, Tony, Gorochov, Andrej V., Poulain, Simon & Suhardjono, Yayuk R., 2014, Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species, pp. 1-104 in Zootaxa 3854 (1) on pages 45-47, DOI: 10.11646/zootaxa.3854.1.1, http://zenodo.org/record/4929336