New remarkable hell ants (Formicidae: Haidomyrmecinae stat. nov.) from mid-Cretaceous amber of northern Myanmar
Creators: name: Perrichot, Vincent
name: Wang, Bo
name: Barden, Phillip
Publisher: name: Zenodo
Year: 2020
Subjects: Biodiversity
Taxonomy

description: <i>Key to genera and species of Haidomyrmecinae</i> 1. Clypeus posteriorly with a slightly elevated setose lobe positioned just ventrad or between antennal insertion <i>………………………………………………………………</i>.. <i>………</i>... 2 Clypeus posteriorly projecting forward into a cephalic horn arising ventrad, between, or dorsad antennal insertion <i>………………………………………………………………</i>.. <i>………</i>... 5 2. Antenna compact, with flagellomeres short (&lt;2 as long as broad); first flagellomere as long as or longer than second one <i>…………………………………………………………………………</i> 3 Antenna elongate, with flagellomeres long (&gt;2 as long as broad); first flagellomere shorter than second one <i>………… ……………………………………………… Haidomyrmex</i> species: 4 3. All flagellomeres nearly of same length; clypeal setose pad positioned anterior to antennal insertion; ocelli present; two metatibial spurs <i> <i>…………………………………………………………</i> …………………</i> <i>Haidomyrmodes mammuthus</i> Perrichot et al., 2008. Flagellomere I longest of basal four flagellar articles; clypeal setose pad positioned slightly posterior to antennal insertion; ocelli absent; one metatibial spur <i>……………………… …</i>.. <i>…</i>.. <i>…… Haidoterminus cippus McKellar, Glasier &amp; Engel, 2013</i>. 4. Flagellomere II longest of basal three flagellar articles; frontal triangle moderately elevated; vertex, ventral margin of mandibles, and antennomeres densely setose; mandibles with medioventral blade bearing at least one large apical tooth, with tip of apical portion tapered to acute point, smooth; ocelli reduced to pit-like traces; subpetiolar process present <i>……………………………</i>. <i>… Haidomyrmex cerberus Dlussky,1996</i> Flagellomere II longest; frontal triangle strongly elevated; vertex, ventral margin of mandibles, and antennomeres mostly glabrous; mandibles with medioventral blades asymmetrical, left one bearing three small teeth, right one bearing two larger teeth; tip of apical portion tapered to sharp point, smooth; ocelli and subpetiolar process absent <i>………………………………… ……………… Haidomyrmex scimitarus Barden &amp; Grimaldi, 2012</i> Apical flagellomere longest; frontal triangle feebly elevated; vertex, ventral margin of mandibles, and antennomeres mostly glabrous; mandibles with medioventral blade bearing a single apical tooth, with tip of apical portion tapered to blunt point and minutely serrate; ocelli absent; subpetiolar process a minute tooth <i>……………………………………………………………………</i>. <i>…………………</i>. <i>Haidomyrmex zigrasi</i>.. Barden &amp; Grimaldi, 2012. 5. Posterior clypeal projection not furrowed ventrally (<b>¼</b> without lateral ridges), forming a pointed or paddle-shaped horn; long, paired trigger setae present, arising from base of clypeal process or ventral surface of apical setose pad; peg-like denticles absent or present only on ventral setose pad; labrum not obviously exposed <i>……………………………………………………………</i>. <i>…</i> 6 Posterior clypeal projection distinctly furrowed ventrally, with lateral ridges originating above mandible insertions, and converging posteriorly to form the lateral margins of a horn; paired trigger setae absent; peg-like denticles present along lateral clypeal ridges, or on labrum, or both; labrum exposed ventrally of horn <i>……………………………………………………</i> 9 6. Clypeal horn a small triangulate cuticular elevation; one pair of trigger setae positioned at base of horn; compound eyes small (OI 24); ocelli absent; flagellomeres IIeX compact, barely longer than wide <i>……………………………………………………………… ……………………… Protoceratomyrmex revelatus</i> gen. et. sp. nov. Clypeal horn paddle-shaped, comprising basal stalk and apical setose pad; compound eyes small to large (OI 26e39); ocelli present; flagellomeres IIeX elongate, distinctly longer than wide <i>………………………………………………………………</i>.... <i>………</i> 7 7. Clypeal horn and curved apical portion of mandibles extremely long, reaching high above vertex (HoI 140; MDI 130); peg-like denticles present on ventral setose pad; 2 pairs of trigger setae positioned at base of horn's stalk; frontal triangle fused with horn, not distinguishable; gastral constriction between AIII and AIV at most faintly impressed <i>…………………………………………………………………………</i>... <i>……… Ceratomyrmex ellenbergeri Perrichot, Wang &amp; Engel, 2016</i> Clypeal horn and curved apical portion of mandibles distinctly shorter, not reaching above vertex (MDI 70); peg-like denticles absent; 1 pair of trigger setae positioned on ventral surface of apical setose pad; frontal triangle connecting with horn by cleared cuticular membrane; gastral constriction between AIII and AIV deeply impressed <i>………………………………………… ……………………………………………</i>. <i>Linguamyrmex</i> species: 8 8 Clypeal horn reduced (HoI 25e35), with stalk almost indistinct and apical setose pad hexagonal, as broad as (worker) or smaller than (gyne) compound eye's length (EL); mandibles with medioventral blade bearing two teeth and, in worker, surpassing midlength of apical curved portion; compound eyes small in worker (OI 25); propodeum higher than long <i>…</i>.. <i>………………</i>. <i>…… Linguamyrmex brevicornis</i> gen. et. sp. nov. Clypeal horn conspicuous (HoI 56), with stalk short, not surpassing frontal triangle in dorsal view, and apical setose pad circular, broader than EL in worker; mandibles with medioventral blade smooth and not reaching to midlength of apical curved portion; compound eyes large in worker (OI 39); propodeum longer than high <i>……………………………… …………………</i>. <i>… Linguamyrmex vladi Barden &amp; Grimaldi, 2017</i>. Clypeal horn enormous (HoI 95e105), with apical setose pad circular, nearly as broad as head in dorsal view, and stalk about as long as pad; mandibles with medioventral blade smooth and, in worker, not reaching to midlength of apical curved portion; compound eyes large in worker (OI 44); propodeum longer than high <i>…………………………………………………………………… …………………</i>.. <i>Linguamyrmex rhinocerus</i> Miao &amp; Wang, 2019. 9 Body and legs robust; mandibles scythe-shaped, with medioventral triangular blade present; in frontal view, the outline of apical portion of mandibles aligned with clypeal lateral ridges; clypeal horn gradually tapering to rounded tip; genae elongate, the mandibles inserted far from compound eyes <i>……………………………………………………………………</i>.. <i>…</i> 10 Body and legs gracile; mandibles sickle-shaped, with medioventral triangular blade absent; clypeal horn and labrum protruding horizontally, their tip with a deep median notch; apical lateral margins of horn bilobed; genae strongly reduced, the mandibles inserted close to compound eyes <i>……………………</i>. <i>………</i>. <i>Aquilomyrmex huangi</i> gen. et. sp. nov. 10 Mandibles with apical curved portion much longer than basal portion and entirely serrate along dorsal margin; with medioventral blade an isosceles tooth not projecting beyond mandibular elbow (labrum and clypeal surface below horn entirely exposed when mandibles closed); ocelli large, antennomeres and compound eyes distinctly elongate <i>…………………………… Dhagnathos autokrator</i> gen. et. sp. nov. Mandibles with apical curved portion not distinctly longer than basal portion; with medioventral blade a large triangle gradually tapering to tip of apical portion (labrum and clypeal surface below horn entirely concealed when mandibles closed); dorsal margin of mandibles serrate along distal half of medioventral projection, bidentate apically; ocelli, antennomeres, and compound eyes more reduced <i>………… ……………………………… …………………………………… Chonidris insolita</i> gen. et. sp. nov.
description: Published as part of <i>Perrichot, Vincent, Wang, Bo &amp; Barden, Phillip, 2020, New remarkable hell ants (Formicidae: Haidomyrmecinae stat. nov.) from mid-Cretaceous amber of northern Myanmar, pp. 1-18 in Cretaceous Research 109</i> on pages 16-17, DOI: 10.1016/j.cretres.2020.104381, http://zenodo.org/record/3653072
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