Gomphostemma dolichobotrys Merr.
Creators: name: Bongcheewin, Bhanubong
name: Ingrouille, Martin J.
name: Paton, Alan J.
Publisher: name: Zenodo
Year: 2022
Subjects: Biodiversity
Taxonomy
Plantae
Tracheophyta
Magnoliopsida
Lamiales
Lamiaceae
Gomphostemma
Gomphostemma dolichobotrys

description: <b> 3. Gomphostemma dolichobotrys <i>Merr</i>.</b> (Merrill 1932a: 128); Keng (1958: 319); Walsingham (2019: 195). Type: Indonesia, Atceh, Takengon, in primary forest, 1100 m, 5 Jan. 1932, <i>3angham &amp; 3angham-Masters</i> 528 (holotype GH [GH00001123]; isotype NY [NY00000333]). <i>Gomphostemma racemosum</i> Steenis ex H. Keng (1969: 80). Type: Indonesia, Northern Sumatra, Atceh, Gacolanden, 1000 m, 5 Feb. 1935, <i>oan Steenis</i> 8682 (holotype L [L0003692]; isotypes BO n.v., K [K000822953], L [L0003691]). Perennial herb up to 2 m tall. <i>Stems</i> erect, obtusely quadrangular with longitudinal grooves, with a mixture of coppery-coloured dense dendroid hairs with a 3 – many-celled stalk and stellate hairs. <i>Leaoes</i> petiolate, chartaceous; blades broadly elliptic-ovate, 13 – 23 × 9 – 13 cm, apex acute, margin shallowly dentate, base cuneate; upper side dull green with 2 – 3-celled simple hairs, lower side pale green tomentose with dense stellate hairs and dendroid hairs confined on midrib and veins and with sparse pale yellow punctate glands; petioles 30 – 50 mm long. <i>Inflorescence</i> axillary, raceme-like with opposite cymes unbranched, lax with axis visible between flowers, 20 – 80 (– 100) mm long, inserted at the upper nodes at which leaves are usually present; verticils 3 – 6 (– 8)-flowered; peduncles (13 –) 30 – 60 mm long; bracts deciduous, not seen; bracteoles linear, narrowly elliptic or slightly falcate, 3 – 10 mm long. <i>Flowering calyx</i> tubular or narrowly infundibular, 6 – 10 mm long, 10-ridged, outside with a mixture of dense hairs, inside with adpressed simple hairs; tube 2 – 5 mm long; lobes 2 – 3 mm long, apex acute. <i>Fruiting calyx</i> tubular-infundibular, 5 – 11 mm long; tube 3 – 8 mm long; lobes 2 – 3 mm long. <i>Corolla</i> white, 13 – 20 mm long, throat narrow, gradually dilated towards the throat, outside tomentose with dense stellate hairs, inside glabrous; tube slender, straight, 8 – 13 mm long, hardly exserted from calyx; posterior lip hood-like, ovate, 2 – 3 mm long; anterior lip slightly longer than posterior lip, 3-lobed, subequal, ovate. <i>Staminal filaments</i> fleshy, tomentose. <i>Style</i> slightly fleshy, glabrous with apex subequally lobed. <i>Nutlets</i> solitary, 3 – 6 mm long. <b>DISTRIBUTION.</b> Indonesia (Sumatra). Map 1. <b>SPECIMENS EXAMINED. INDONESIA. SUMATRA</b>. Aceh: Gunung Leuser Nature Reserve, Climbing Gunung Leuser West top, from Penosan via Putcuk Angasan, 23 km SW of Blang Kedceren [3.9956°, 95.2069°], 1300 – 1300 m, 31 March 1953, <i>de Wilde &amp; de Wilde-Duyfjes</i> 13892 (L); Takengon, 1100 m, 5 Jan. 1932, <i>3angham &amp; 3angham-Masters</i> 528 (holotype GH; isotype NY); Gacolanden, 1000 m, 5 Feb. 1935, <i>oan Steenis</i> 8682 (holotype of <i>G. racemosum</i>, L; isotypes K, L). <b>HABITAT.</b> In primary forest, riverine forest and scrub edge; alt. 1000 – 1300 m. <b>CONSERVATION STATUS.</b> There are only three collections of <i>Gomphostemma dolichobotrys</i> collected from Gunung Leuser Nature Reserve in North Sumatra giving an EOO of 563 km 2 suggesting Endangered status. However, the localities are well-protected as a nature reserve and no evidence of habitat loss, hence, the Least Concern (LC) category is applied. <b>PHENOLOGY</b>. Flowering and fruiting January – March. <b>NOTES.</b> <i>Gomphostemma dolichobotrys</i> is recognised by its erect stems, upright inflorescence with deciduous bracts, and corolla tube hardly exserted from the calyx. It is morphologically similar to <i>G. microdon</i> in having an unbranched, raceme-like inflorescence.
description: Published as part of <i>Bongcheewin, Bhanubong, Ingrouille, Martin J. &amp; Paton, Alan J., 2022, A revision of Gomphostemma (Lamiaceae), pp. 27-92 in Kew Bulletin 77 (1)</i> on page 43, DOI: 10.1007/s12225-021-09991-y, http://zenodo.org/record/7615581
description: {"references": ["Merrill, E.?. (1932 a). Plants collected in Sumatra. Contr. Arnold Arbor. 8: 128.", "____ (1958). Labiatae. In: C. G. G. J. van Steenis (ed.), Flora Malesiana ser. 1, 8: 301 - 392. Sicthoff &amp; Noordhoff, Alpen Aan? en Ricn.", "Walsingham, L. (2019). Gomphostemma. In: G. L. C. Bramley (ed.), Flora Malesiana ser. 1, 23: 1 - 222. Foundation Flora Malesiana, The Netherlands.", "Keng, H. (1969). Flora Malesianae precursors 28: A revision of Malesian Labiatae. Gard. 3 ull. Singapore 22: 13 - 180."]}