Dinosauroid footprints of French Upper Triassic. Discrimination, interpretation and comparison
Authors: Gand Georges , Demathieu Georges , Grancier Max , Sciau Jacques 
Published: June 2005

Abstract
Dinosauroid tracks are common in France. They are mainly tridactyl II–IV feet tracks, sometimes with those of hands. A lot of these latter Coelurosaurichnus grancieri Courel & Demathieu, 2000 was discovered by MG near the village of Payzac (Ardèche department) in beds belonging to the “Ensemble gréso-dolomitique gris” (= EGDG) (fig. 1. 2). They are dated Carnian from palynological data [Fauconnier et al., 1996]. C. grancieri are tracks of feet and hands respectively 10 cm and 2.5 cm long (fig. 1. 3–5, 7–21; tabl. I-1, tabl. II). From the trackways characters, it was inferred that trackmakers were long legged, bipedal-quadrupedal reptiles, 1–2 m long.
New discoveries made by MG in Payzac site showed the existence of a new ichnopopulation Grallator sp. (fig. 1. 25–33 ; tabl. I. II) They are only feet tracks whose trackmakers were also long-legged but only bipedal animals, a little larger that the preceding ones. Statistical comparisons with the use of Student and Snédecor tests, discriminant analysis between C. grancieri and Grallator sp. show that these two ichnopopulations are different (CG, GP, fig. 3). Also they are well distinguished with other Carnian ichnospecies of Coelurosaurichnus of Franconie = Grallator or Atreipus (Haubold & Klein, 2000) (Aal, fig. 3) and Hettangian G. variabilis of Causses (Gv, fig. 4).
Coelurosaurichnus grancieri and Grallator sp. trackmakers were carnivorous reptiles and probably belonging to different taxa. Maybe they were only Dinosauromorpha or already theropod Dinosauria. Anyway, their presence in the lower-medium Carnian means a biodiversity for these bipedal reptiles which wwas also underlined by palichnological data from South Africa [Ellenberger, 1972] and osteological discoveries in Argentina [Bonaparte, 1993; Cuny, 1993].
Other beds bearing dinosauroid footprints were found in Ucel Formation dated Norian [Razin et al., 1996] well exposed near Corbès [Ellenberger, 1965] and the town of la Grand-Combe [Gand et al., 2000]. All these traces are Grallator, therefore II–IV feet, about 20 cm long, probably made by Ceratosauria. The ichnopopulation G. andeolensis are clearly discriminative of Carnian Coelurosaurichnus grancieri/ Grallator sp. and Rhaetian Eubrontes (fig. 4). It was found with large Sauropoda and Prosauropoda footprints. Then, in France, frequent dinosauroid footprints have been found in Grès d’Olemps F. (Causse de Rodez) in which are Eubrontes giganteus, Grallator cf. minusculus and Dilophosauripus sp. The first two are probably due to Theropods and the latter to Ornithopodia.
French palichnological data and those coming from West USA [Lockley and Hunt, 1995] suggest that the carnivorous bipedal reptiles size increases slowly but surely during Upper Triassic with a speeding up at the end of Rhaetian. But the presence of large Eubrontes, 43 cm long in Carnian beds of Australia means this is not true [Lucas, 2003]. Of course, it would be very interesting to find outside of Australia such large early Eubrontes. In France, also we have not met Brachychirotherium and Chirotherium : ichnites of Thécodonts which are still present in the Carnian-Norian of Germany and America [Haubold, 1984, 1986].