bluuebunny/crossref_metadata_2025_split huggingface dataset:
Macro- and microevolutionary aspects of the early Paleogene recovery of the planktonic foraminifera
Authors: Arnold Anthony J. , Kelly Daniel C. , Parker William C. 
Published: January 2018

To date, most macroevolutionary studies have focused on taxonomic data; there are very few data sets that provide a large enough statistical population for extensive macroevolutionary studies involving direct morphometric data. The Cenozoic planktonic foraminifera are exceptional in this regard. Digitally captured shape descriptions of 342 species of Cenozoic planktonic foraminifera have been combined with the available data on their phylogenetic relationships to examine the interplay between speciation rates, size change, and various geometric characters. The results tend, at least for the planktonic foraminifera, to support Stanley's (1973) hypothesis regarding the causal mechanisms behind Cope's Rule. The tendency toward size increase may result from the disadvantages of large size during times of mass extinction rather than from the (conventionally assumed) advantages of larger size.Additional results derived from morphotypic longevities and morphotypic turnover rates suggest an enhanced probability of speciation early in the Cenozoic, and enhanced longevity in the later Cenozoic.Microevolutionary studies of patterns of changing variance within the Paleocene and Eocene genus Morozovella suggest that conventional foraminiferal taxonomic practice may not accurately reflect biological realities within the group, thus implying that our macroevolutionary results might be interpreted in other ways.Paedomorphosis has now been seen in several foraminiferal lineages, including Morozovella angulata, a focus of the present study. Isotopic data (Shackleton, Corfield, and Hall, 1985) suggests that evolution of this group is accompanied by the invasion of a stratified water column. These observations suggest that one might look for systematic macroevolutionary morphologic tendencies in chamber expansion rate and size as a guide to understanding paleoecological conditions. It also seems reasonable to suggest that the complex morphological changes seen in the morozovellids may not represent morphological adaptation, but resource-related heterochronic shifts with ancillary morphological consequences.
Replacement events among tetrapods: expansion or competition?
Authors: Benton Michael J. , Storrs Glenn W. 
Published: January 2018

Many macroevolutionary patterns have been explained in the past on the basis of expectations and prejudice rather than quantitative testing. Large-scale ecological replacements have been a prime example, where expectations of progress and competitive advantage have coloured interpretations. Equally, no doubt, others have looked for, and found, evidence for entirely noncompetitive processes to explain major replacements (reviewed, Benton 1987).A partially quantitative approach is presented, based on a study of tetrapod families, marine and nonmarine, through the interval from Devonian to the present day. New data bases of the history of tetrapod diversification have the advantage over older attempts of (1) a fully cladistic phylogeny, and hence monophyletic families; (2) a detailed overview of stratigraphy and improved range data; (3) palaeobiogeographic control; and, (4) definition of ecological categories in terms of mean body size categories, diet, main habitat types (fully marine; partially marine/ coastal; freshwater; aquatic/terrestrial; lowland; upland; arboreal, aerial).The bulk of the diversification of tetrapods is correlated with increases in the ranges both of habitats occupied and of diets (Benton 1990). However, tetrapod families came and went, and a large proportion of the diversification could have been driven by competition and progressive replacement of archaic families by competitively superior ones in ecological relays. In order to test this, the ranges of all tetrapod families were plotted against time and sorted into a variety of habitat, dietary, and palaeogeographic categories.The results do not support a generalised view of competitive ecological relays. The majority of families appear to have arisen within new cells of the matrix, and did not apparently overlap in time with, or follow closely after, any potential competitors. This would be the minimum requirement for a postulate of competition. Only some 5% of cases could be open to a competitive explanation, and these include certain well known examples, such as multituberculates/ rodents, plesiadapiforms/ rodents, ichthyosaurs/ mosasaurs. The present study has narrowed the odds againt large-scale competition as a significant motor of large-scale biotic replacements. Most new taxa of tetrapods appear to have arisen in response to the availability of empty adaptive space.Environmental shifts may also be detected during the course of tetrapod evolution, with expansions offshore to ever more fully aquatic lifestyles among marine tetrapods (because they all arose from terrestrial ancestors!), and with expansions from lowland tropical zones to temperate belts, uplands, and marginal lifestyles (burrowing, climbing, flying) among nonmarine tetrapods.
Biogeographic and evolutionary patterns of change in the terrestrial biota across the Cretaceous\Tertiary boundary
Authors: Clemens William A. 
Published: January 2018

The currently available fossil record suggests, 1) biogeographic differentiation of the Late Cretaceous terrestrial biota and, 2) distinctly different patterns of evolution of terrestrial faunas and floras across the Cretaceous\Tertiary boundary.Discovery in Alaska of dinosaurs and mammals that lived at Late Cretaceous northern high latitudes provides evidence that many groups of terrestrial vertebrates had extensive geographic ranges and faunas were biogeographically differentiated. The Alaskan dinosaurs, represented by individuals that range in size from hatchlings to adults, might have been migratory forms living at high latitudes only during the summer months. In contrast, the small mammals probably were not migratory.Although recent discoveries are expanding our knowledge of the evolution of the terrestrial biota, the fossil record of terrestrial vertebrates during the Cretaceous\Tertiary transition is still heavily biased in favor of the northern Western Interior of North America. Here evolutionary change of the terrestrial fauna did not just involve extinction of lineages, some already decreasing in taxonomic diversity, and survival of many others. Shifts in biogeographic range and immigration of new groups played a significant role in remodeling the terrestrial fauna. The paleobotanical record is more extensive but also is biased with the most detailed record coming from the Western Interior where floral change is characterized as “massive” or “catastrophic” in scope.These conflicting evolutionary patterns are well founded being based on analysis of substantial fossil records. A new program of research directed toward resolution of the apparent paradox tests the hypothesis that at the end of the Cretaceous the terrestrial biota was biogeographically heterogeneous and evolutionary patterns of faunal and floral change in the Western Interior cannot be taken as globally representative.
Environmental patterns in the origin and diversification of deep-water scleractinian and rugose corals
Authors: Droser Mary L. , Hampt Gretchen , Clements Steven 
Published: January 2018

Scleractinian and rugose corals demonstrate a broad environmental distribution and have been documented in strata reflecting deposition from high-energy, nearshore reefal to low-energy basinal settings. Both clades have a relatively high diversity of representatives which live(d) at great depths. While deep-water scleractinian coral banks are well-documented, commonly the deep-water representatives of both clades are the small, solitary, and often delicate forms. In order to determine the nature of diversification of these deep-water forms the paleoenvironments of the earliest Rugosa and representatives of extant deep-water scleractinian families were determined. Environmental patterns of the subsequentdiversification of the Rugosa and scleractinian sub-order Caryophylliina were also documented. Environmental information was determined from independent sedimentological data.The earliest reported Rugosa is Lambeophyllum from the Chazyan Crown Point Limestone (VT, U.S.A.), which was deposited in an nearshore/inner shelf setting. Subsequentto this occurrence, rugosans appear nearly simultaneously in North America, Australia, Europe and China. However, common Lambeophyllum, within the Blackriverian Orwell Limestone (NY, U.S.A), appearsto be the earliest of these occurrences. The Orwell is, likewise interpreted to be shallow water in origin. After their initial appearance in an onshore environment, the Rugosa rapidly expand offshore. While Middle Ordovician Rugosaare of low diversity (1 genus at a given locality), by the end of the Blackriverian they are found in middle and outer shelf settings.Nearly all representatives of families of extant scleractinians now living in deep-water (>200’) also first occur in an onshore environment - primarily the inner shelf. The caryophylliids, the most speciousdeep-water coral clade, first appear in Toarcian age inner shelf strata and soon after occur in outer shelf deposits. However, occurrences of high diversities of caryophylliids (generic diversity greater than 4 at a given locality) are confined to inner shelf settings until the Cretaceous when generic diversities of 2 or more are common in the outer shelf. In the Tertiary, high diversities of caryophylliids are generally restricted to slope and deep basin settings, a pattern which characterises their modern distribution.Both rugose and scleractinian corals expand offshore at low diversities within 5 million years of their first appearance and thus, demonstrate environmental diversification before taxonomic diversification.
Dinosaur behavior and growth
Authors: Horner John R. 
Published: January 2018

Geological and paleontological data derived from the Upper Cretaceous Two Medicine Formation of Montana indicates that at least some dinosaur species exhibited complex social behaviors comparable to many living birds. Two species of duck-billed dinosaurs, a hypsilophodontid and a troodontid, nested in colonies and attended their respective young. Duckbilled dinosaurs had altricial young, whereas the hypsilophodontid and troodontid had precocial young. Morphological evidence indicates that several of the cranial elements of the nestling duckbills experienced retarded development, and a retention of juvenile features. Following their respective nesting periods duck-billed and horned dinosaurs aggregated into large herds, and apparently migrated seasonally. Cranial ornamentations possessed by duck-billed and horned dinosaurs were most likely used for sexual display and mate recognition.Osteohistological data indicates that the duck-billed dinosaurs hatched from their eggs at .50 to .75 meters in length, and exited their nests at 1.5 to 2 meters in length, at an age of about one month. These dinosaurs reached 3 to 4 meters in length their first year, and 7 to 8 meters in length after about 5 years. Growth and metabolic rates decreased substantially upon reaching adult size.
Spine function in the odontopleurid trilobites

Leonaspis

and

Dicranurus

from the Devonian of Oklahoma
Authors: Kloc Gerald J. 
Published: January 2018

The Lower Devonian Haragan Formation of Oklahoma has well preserved odontopleurid trilobites found in repeated calcarous mudstones. These layers (5–9cm. thick) represent distal tempestites that rapidly buried and preserved completely articulated odontopleurids which provide important paleobiological information. The genal and pleural spines of Leonaspis williamsi and Dicranurus hamatus elegantus of the Haragan Formation are examined and compared.In Leonaspis the ventrally directed genal and pleural spines are straight and lay flat. The first pleural spine is the longest followed by progressively shorter pleural spines. Several collected and prepared enrolled specimens have the spines radiating outward. In an enrolled position the spines would protect Leonaspis from a large predator.The pleural spines of Dicranurus have a very different pattern from Leonaspis. The first three spines are short and laterally directed, the fourth, sixth and seventh are very long, curved and directed ventrally, the eighth, ninth and pygidial spines are short and decrease in length toward the pygidium. There is no pleural spine on the fifth thoracic segment and the genal spine is very long and curved. On these spines at many growth stages there is direct and indirect evidence of being encrusted based on the following: (1) Epibionts such as bryozoans, brachiopods and a crinoid; 2) Endoliths (25–150 microns in diameter) of unknown origin are found all over Dicranurus but are predominantly on the spines; 3) Pyrite (now limonite) coating the spines apparently formed from the decay of organic matter. If the source of the organic matter came from the trilobite, then pyrite would be found all over Dicranurus. However, it is only found randomly on the spines. This pattern is like some of the encrusting bryozoans suggesting that the source of the organic matter may be from an encrusting organism.It is doubtful that the pleural spines of Dicranurus would be adapted for resting on the bottom when bryozoans are found encrusting the entire surface of the spine. However, the very long pleural and genal spines with attched organisms would break up the outline of Dicraurus making it less conspicuous to a predator.
Competitive overgrowth and post-Paleozoic macroevolution of marine bryozoa
Authors: McKinney Frank K. , Lidgard Scott 
Published: January 2018

The Lower Ordovician through Upper Jurassic record of the phylum Bryozoa consists almost entirely of the class Stenolaemata, excepting a very few Gymnolaemata (ichnofossils of the order Ctenostomata and, in the uppermost Upper Jurassic, Pyriporopsis, which is the oldest known Cheilostomata). Along with the decline to eventual extinction of most Paleozoic stenolaemates beginning with the end-Permian crisis, the stenolaemate order Cyclostomata radiated in Late Triassic and Jurassic to include the diversity of growth forms and locally the abundance common for Paleozoic bryozoan faunas. By late Cretaceous, however, cheilostomes had diversified and had come to dominate in bryozoan faunas; they have continued to dominate to the present.In the abundant and diverse bryozoan fauna of the northern Adriatic Sea, cheilostomes are four to five times as diverse and are orders of magnitude more abundant than are cyclostomes. Within this fauna, encrusting sheet-like cheilostomes consistently overgrow adjacent encrusting cyclostomes. Different ontogenetic patterns of skeleton and organs, as well as larger zooid sizes at colony margins, produce patterns of feeding currents that allow cheilostomes to deliver their filtered water to and to overtop competing cyclostomes at interacting margins. These Bauplan differences apparently constitute a key evolutionary innovation characterizing the Cheilostomata. Along with higher growth rates in most cheilostomes, this key evolutionary innovation should have given cheilostomes a competitive edge since the origin of multiserial colonies within the clade in Early Cretaceous. It is hypothesized that superior overgrowth capability of cheilostomes has been an important factor in their success and the relative decline of cyclostomes.
Vendian body fossils (?) and isotope stratigraphy from the Caborca area, Sonora, Mexico
Authors: McMenamin Mark A.S. , Rowland Stephen M. , Corsetti Frank , Dix Anne M. , Nance Ronald P. 
Published: January 2018

In the Proterozoic Clemente Formation at Cerro Rajon, south of Caborca, Sonora, we have collected sandstone beds that contain many small, sediment-filled structures. Each structure is a thin-walled, elongate sack, about 1.5 cm tall, oriented perpendicular to bedding. The sacklike structures are laterally flattened and roughly elliptical in transverse section, with the major axis of the ellipse measuring several mm and the minor axis 2-3 mm. The walls are not mineralized and consist of a thin layer of cryptocrystalline clay. One or both walls of each specimen is undulatory with a wavelength of 3-4 mm. The sacks are closed at the bottom, but appear to be open at the top, like an uninflated toy balloon. On the bottom surface of the sandstone bed, beneath each vertical sack structure, is a prolate ellipsoid-shaped sole mark. The long axis of each ellipsoid sole mark is parallel to the long axis of the associated sack structure, and the sole marks are oriented sub-parallel to one another on the surface of the bed. We tentatively interpret these structures to be the holdfasts of gregarious, attached, benthic organisms oriented sub-parallel to one another on the sea floor.A preliminary study of carbon and oxygen isotope stratigraphy was carried out on the 2 km-thick Proterozoic through Lower Cambrian section at Cerro Rajon. Samples were collected every 50 m, and analyses were done in the laboratory of J.M. Hayes at Indiana University, whose assistance is gratefully acknowledged. The isotope record shows a primary signature for most of the section. Although the wide spacing of sample collection does not permit an unequivocal intercontinental correlation, a prominent negative δ13C excursion near the base of the Papalote Formation may correlate with similar excursions at the Vendian-Cambrian boundary on several continents. This would place the base of the Cambrian of the Caborca area several hundred meters lower than previously thought, but still several hundred meters above the possible body fossils of the Clemente Formation.
From environmental fluctuations and energy availability to evolutionary change and speciation
Authors: Parsons Peter A. 
Published: January 2018

Evolutionary patterns can be considered in terms of two interacting continuums (a) the magnitude of environmental fluctuations of physical variables especially climatic, and (b) the availability of metabolic energy beyond basic needs for maintenance and survival (Parsons, 1991) so giving four extreme combinations:(1) widely fluctuating environments, low metabolic energy availability. This situation is a feature of species borders where the energetic costs of environmental perturbations restrict normal physiological processes so that ecological range expansions are precluded. Provided extinctions do not occur, species would alter their ranges to track longer term climatic changes as described in the historical and fossil records.(2) widely fluctuating environments, high metabolic energy availability. This combination favors opportunists and colonists. Furthermore, following a relaxation of stress intensity after a mass extinction there could be a burst of speciation in such generalist species.(3) stable environments, low metabolic energy availabilty. This combination represents deep sea and cave environments where low metabolic rate is an adaptation to accomodate stresses such as anoxia and low resource availability. Consequently the capacity for evolutionary change is low in these habitats which tend to contain relict species.(4) stable environments, high metabolic energy availability. On energetic grounds speciation is theoretically possible, but high vulnerability to any stressful perturbation would be restrictive in practice since some stress is the norm in natural populations.The potential for evolutionary change and speciation is low in these four extremes, however in intermediate habitats the potential is higher as implied in (2) when stress intensity is relaxed. The intermediate habitats would be characterized by moderately fluctuating environments where the metabolic cost of stress would not preclude adaptive change. Furthermore, the stress levels in such habitats should lead to sufficient underlying genetic variability to be permissive of adaptive responses to environmental change (Parsons, 1991). This paradigm may apply to speciation in tropical rain forests. A fossil analogy would be marine invertebrates in frequently disturbed onshore habitats.
Apparatus architecture and allometry: the keys to conodont element function?
Authors: Purnell Mark A. 
Published: January 2018

Much effort has been expended in determining the geological utility of conodont elements; little has been spent on determining their usefulness to the conodont animal. Most conodonts had an apparatus made up of a number of morphologically distinct, complex elements. Such apparatuses were borne by members of the Ozarkodinida, Prioniodinida, and Prioniodontida, together accommodating ~75 percent of known conodont genera.The ozarkodinid apparatus consisted of an anterior battery of S and M elements behind which lay opposed pairs of Pb and Pa elements. Debate over its function has centered on two main hypotheses. These have considered the apparatus either as a filter feeding system, or as teeth. Ontogenetic analysis reveals that ozarkodinid apparatus growth was incompatible with a filtering function. The rate of increase in size of the S and M elements (the postulated filtering system) was insufficient to have met the increasing food requirements of the growing animal.Like the apparatus of ozarkodinids, that of prioniodinids was differentiated into an anterior S and M element series and posterior, paired Pa and Pb elements. Despite these similarities in element arrangement, the prioniodinid apparatus does not exhibit the same degree of morphological differentiation as that of ozarkodinids. The Pa and Pb elements were probably involved in swallowing rather than cutting and grinding food.The prioniodontid apparatus is poorly known but it is clear that although the morphology of elements is comparable to ozarkodinids, their arrangement in the apparatus was markedly different. Until the architecture of the apparatus has been determined, it's mode of function can only be guessed at. Preliminary knowledge of the apparatus, however, suggests that it operated differently to those of ozarkodinids and prioniodinids.Understanding of the function of conodont elements requires that their morphology is studied within the context of an integrated multielement apparatus. There is substantial variation in apparatus composition and differentiation, and element morphology between clades. This variation undoubtedly reflects, to a large extent, differences in apparatus adaptability and versatility, functional flexibility, and food specificity; all factors that in other groups have had a major influence on evolutionary patterns.