bluuebunny/crossref_metadata_2025 huggingface dataset:
Developing a Scots pine breeding objective: a case study involving a Swedish sawmill
Authors: Berlin Mats , Lönnstedt Lars , Jansson Gunnar , Danell Öje , Ericsson Tore
Published: March 2014

The aim of this study was to develop a Scots pine (Pinus sylvestris L.) breeding objective for a vertically integrated sawmill in northern Sweden. The production system was defined as comprising the sawmill and the forests supplying it. Volume per hectare, wood density, survival and straightness were used as objective traits and the related selection criteria were measurements, collected at young tree age, of height, diameter, pilodyn penetration, vitality and straightness. A bio-economic model was used to calculate economic weights for the objective traits identified. We also investigated the efficiency of different selection indices based on these economic weights, in combination with available data on genetic parameters. Furthermore, we studied the effect of different discount rates on the calculated economic weights. The results showed that, compared to the full index (which included all selection criteria), omitting either vitality or straightness had a negligible effect, reducing predicted profit gain per hectare by less than one per cent. Height or diameter each had a greater effect, with a loss of predicted profit gain per hectare of up to 6%. Excluding pilodyn penetration from the selection index caused the largest reduction in predicted profit gain per hectare, amounting to over 10%. However, when both height and diameter were removed the predicted profit gain per hectare dropped to one-third of that based on the full index. Finally, ranking and genetic selection for the developed breeding objective was insensitive to changes in the discount rate.
Wind and snow damage in the Pyrenees pine forests: effect of stand attributes and location
Authors: Martín-Alcón Santiago , González-Olabarría José , Coll Lluís
Published: March 2014

Wind and snow-induced damage have been analyzed at stand level for three pine forests in the Central-Eastern Pyrenees (Pinus nigra Arn. salzmanii, Pinus sylvestris L. and Pinus uncinata Ram.). Stand-level models have been then developed for the most affected two species, Pinus sylvestris L. and Pinus uncinata Ram., to describe damage severity. The models were based on data from national forest inventory plots. They included variables related to the spatial location and structure of the stands, being validated using a sub-set of the database (25% of the plots randomly selected). Mountain pine forests (Pinus uncinata Ram.) were the most heavily affected by wind and snow disturbances. For both mountain and Scots pine species, topographic exposure and the severity of the local storm regime had an important effect on the degree of damage. Standâs resistance to wind and snow was found to be dependent on the combined effect of basal area and mean slenderness of the dominant trees. For a given slenderness ratio, damage increased strongly in lower-density stands, particularly in stands with basal areas below 15 m/ha. Stand structure was particularly important to define the resistance of Scots pine stands, which presented a higher vulnerability to wind and snow under higher degree of even-agedness. The models presented in this study provide empirically-based information that can be used to implement silvicultural practices to minimize the risk of those forests to suffer wind and snow-related damages.2
Growth of Pinus sylvestris after the application of wood ash or P and K fertilizer to a peatland in southern Sweden
Authors: Sikström Ulf , Almqvist Curt , Jansson Gunnar
Published: March 2014

The effects of the application of wood ash and of fertilizer regimes including phosphorus (P) and potassium (K), with and without simultaneous addition of nitrogen (N), were investigated on a stand of Scots pine (Pinus sylvestris L.) saplings growing on a drained oligotrophic peatland site in southern Sweden. A randomized block design was used. Tree growth and concentrations of various elements in the needles were measured. The addition of similar doses of P (approx. 40 kg P ha) from different sources resulted in similar growth responses, amounting to 1.6â1.9 m ha yr of stem wood over the 26-year study. The P source was either wood ash (2500 kg d.w. ha) or PK-fertilizer (raw phosphate and potassium chloride). In response to several treatments there were both increased numbers of trees and increased growth of individual trees. The high PK-dose (40 kg P ha and 80 kg K ha) appeared to result in a larger growth increase than the low dose (20 kg P ha and 40 kg K ha). The N treatment had no additional effect on growth. In the control plots, tree growth was more or less negligible (0.04 m ha yr). After almost 26 years, concentrations of P and K in the needles of treated plants were still higher than in the untreated control plants. Nevertheless, in spite of the elevated P concentration, P appears to limit the growth of Scots pine. In conclusion, after sufficient drainage of this type of peatland site, it is possible for a forest stand to develop to the pole stage if wood ash or PK-fertilizer is applied.â13â1â1â1â1â1â1â13â1â1
Comparison of two working methods for small tree harvesting with a multi tree felling head mounted on farm tractor
Authors: Belbo Helmer
Published: March 2014

In this study, the efficiency of a small multi-tree felling head, mounted on a farm tractor with a timber trailer was studied, when harvesting small trees for energy in thinnings. Both separate loading and direct loading of the felled trees was studied. Time studies were carried out in a mixed stand of Norway spruce (Picea abies (L.) Karst) and birch (Betula pubescens Ehrh.). The time consumption of the work elements in the different work methods was formulated by regression analysis, where the independent variables were tree size and degree of accumulation. The average size of the harvested trees was 0.035 m3. The time consumption for the harvesting and loading were similar for the two studied methods, 20 minutes per m3 at a tree size of 0.035 m3, but the two methods showed different characteristics for different tree sizes and level of accumulation. The direct loading method had the highest productivity when more than 0.1 m3 were collected in the felling cycle, whereas the separate loading method had the highest productivity when less than 0.05 m3 were collected in the felling cycle. The total effective time consumption for harvesting and forwarding the biomass 300 meters to roadside landing was 27 minutes per m3. The efficiency of the initial felling and collecting of the small trees was the main challenge. Both the harvesting technique and harvesting technology needs further development to provide a feasible production chain for woodfuel from energy thinning.
Coupling greenhouse gas credits with biofuel production cost in determining conversion plant size
Authors: Gan Jianbang , Smith C.
Published: March 2014

Biofuel plant size is one of the key variables in biofuel supply chain analysis as it plays a pivotal role in controlling the efficacy of both feedstock supply and feedstock-to-biofuel conversion. The unit production cost and greenhouse gas (GHG) balance of biofuels vary with plant size. We develop an analytical framework for integrating biofuel production costs and GHG balance derived from life-cycle analysis into supply chain optimization, followed by its application to ethanol production using forest biomass in the southern United States. We derive formulas for determining the optimal biofuel plant size and the corresponding feedstock supply radius based on the minimization of biofuel production costs less GHG benefits. Our results indicate that though biofuel plant size and feedstock supply radius should be augmented by considering GHG benefits, the GHG price will have a more significant impact on net biofuel production costs than on conversion plant size or feedstock supply radius. With a rise in the GHG price the net biofuel production cost tends to increase while the directions of change in plant size and feedstock supply radius are uncertain, depending upon the costs and GHG emissions of biomass transport and feedstock-to-fuel conversion. Combining GHG offset values with biofuel production costs enables us to more holistically examine the biofuel supply chain.
Capercaillie (Tetrao urogallus) habitat characteristics in north-boreal Finland
Authors: Miettinen Janne , Helle Pekka , Nikula Ari , Niemelä Pekka
Published: March 2014

This study aimed to identify tools for taking capercaillie habitats into consideration in forest management. This would provide new alternatives for ecologically more sustainable forest management. Capercaillie summer and winter locations, from wildlife monitoring counts (1998â2004) in northern Finland, and reference, non-capercaillie locations were combined with forest planning data, and the area proportions of different landscape classes in an 800-m radius circle surrounding capercaillie and reference locations were compared. Thinning stands (in summer and winter) and spruce mires (in summer) were more abundant in capercaillie habitats than in reference landscapes, whereas e.g. seedling stands, mature stands and waste land areas were less abundant. The relative habitat use was highest in mean tree diameter (DBH) classes from 10.5 to 14.5 cm in summer habitats of adult capercaillie in heath forests, whereas in peatland forests, in brood habitats and in winter habitats it peaked in diameter classes 14.5 to 18.5 cm. The tree layer density was positively associated with the relative habitat use. A trend of lower habitat use was detected in the largest diameters (17â40 cm) in comparison to middle-sized diameters (10â16 cm) in heath forests, but not in peatland forests. Relatively young managed forests (age 30â40 years or more) can form suitable capercaillie habitats in north-boreal forests. However, this suitability is not necessarily permanent. Understorey management, longer rotations and multicohort forest management are suitable tools for capercaillie habitat management, because they can increase the available cover close to the ground, canopy cover, overall forest cover at the landscape scale and bilberry cover.
Relationship between autumn cold hardiness and field performance in northern Pinus sylvestris
Authors: Persson Torgny , Andersson Bengt , Ericsson Tore
Published: March 2014

Results from 3 artificial freezing tests (one-year-old seedlings) and 15 field trials (9- to 21-year old trees) of half-sib offspring from first generation Scots pine (Pinus sylvestris L.) plus-trees were used to estimate the amount of additive genetic variance for autumn cold hardiness and traits assessed in the field, and the genetic correlations between them. Cold hardiness of individual seedlings was scored visually, based on the discoloration of their needles after freezing in a climate chamber. The field traits analyzed were tree vitality, tree height, spike knot frequency, branch diameter, branch angle, stem straightness, and susceptibility to infection by the pathogenic fungi Phacidium infestans L., Gremmeniella abietina (Lagerb.) Morelet, Melampsora pinitorqua (Braun) Rostr. and Lophodermella sulcigena (Rostr.) HÃ¶hn. Narrow sense individual heritabilities varied between 0.30 and 0.54 for autumn cold hardiness, 0 and 0.18 for tree vitality, 0.07 and 0.41 for tree height, and 0.01 and 0.26 for the remaining traits. Based on the results of the artificial freeze tests, our estimates of additive genetic correlations indicate that while early selection for cold hardiness can improve seedling survival rates in the field, it may also reduce growth in mild environments. It also has minor effects on quality traits and attack by common fungal diseases. The results indicate that artificial freeze testing is an appropriate method for identifying suitable clones for establishing seed orchards to supply stock for the reforestation of regions with harsh environments.
Tree species classification using airborne LiDAR – effects of stand and tree parameters, downsizing of training set, intensity normalization, and sensor type
Authors: Korpela Ilkka , Ørka Hans , Maltamo Matti , Tokola Timo , Hyyppä Juha
Published: March 2014

Tree species identification constitutes a bottleneck in remote sensing-based forest inventory. In passive images the differentiating features overlap and bidirectional reflectance hampers analysis. Airborne LiDAR provides radiometric and geometric information. We examined the single-trees-level response of two LiDAR sensors in over 13 000 forest trees in southern Finland. We focused on the commercially important species. Our aims were to 1) explore the relevant LiDAR features and study their dependencies on stand and tree variables, 2) examine two sensors and their fusion, 3) quantify the gain from intensity normalizations, 4) examine the importance of the size of the training set, and 5) determine the effects of stand age and site fertility. A set of 570 semiurban broad-leaved trees and exotic conifers was analyzed to 6) examine the LiDAR signal in the economically less important species. An accuracy of 88 90% was achieved in the classification of Scots pine, Norway spruce, and birch, using intensity variables. Spruce and birch showed the highest levels of confusion. Downsizing the training set from 30% to 2.5% of all trees had only a marginal effect on the performance of classifiers. The intensity features were dependent on the absolute and relative sizes of trees, especially for birch. The results suggest that leaf size, orientation, and foliage density affect the intensity, which is thus not affected by reflectance only. Some of the ecologically important species in Finland may be separable, since they gave rise to high intensity values. Comparison of the sensors implies that performance of the intensity data for species classification varies between sensors for reasons that remained uncertain. Both range and gain receiver normalization improved species classification. Weighting of the intensity values improved the fusion of two LiDAR datasets.
Soil organic carbon in Swedish spruce and pine forests – differences in stock levels and regional patterns
Authors: Stendahl Johan , Johansson Maj-Britt , Eriksson Erik , Nilsson Åke , Langvall Ola
Published: March 2014

The selection of tree species is one factor to consider if we want to mitigate carbon dioxide emissions to the atmosphere through forest management. The objectives of this study were to estimate the differences in soil organic carbon (SOC) stocks under Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) forests and to examine causes of differences in the accumulation of carbon in the forest soil. Large-scale inventory data was used to quantify variations in SOC stock in relation to stand type and the accumulation of carbon for spruce and pine stands was analysed by simulation. Based on field data, the national mean SOC stock was 9.2 kg m in spruce dominated stands and 5.7Â kg m in pine dominated stands. For both species, the SOC stock, measured in the field inventory, increased significantly with increasing temperature, although at different rates. The SOC stock was larger for spruce under all temperature conditions, but the difference between species diminished with increasing temperature. The simulations indicated that the build-up of SOC over several rotations was 22% higher in spruce stands than in pine stands under similar environmental conditions. The main difference was found to be the greater input of harvest residues for spruce. Further, the simulations showed that ground vegetation contributed considerably more to the litter production under pine than under spruce. On sites where both Scots pine and Norway spruce are considered suitable, the latter should be selected if the aim of the forest management policy is to maximize the accumulation of SOC in the forest. Further, spruce is more favourable for SOC accumulation in areas with cold temperatures and on sites with low productivity.â2â2
Export of suspended solids and dissolved elements from peatland areas after ditch network maintenance in south-central Finland
Authors: Nieminen Mika , Ahti Erkki , Koivusalo Harri , Mattsson Tuija , Sarkkola Sakari , Laurén Ari
Published: March 2014

In Finland nearly 6 million hectares of peatlands are drained for forestry purposes. Ditch network maintenance in the drained peatlands, i.e. cleaning old ditches or digging complementary ditches, deteriorates surface water quality by increasing the export of dissolved elements and suspended solids (SS). Effect of ditch network maintenance on the export of SS, dissolved organic carbon (DOC), and dissolved nitrogen (N), phosphorous (P), iron (Fe), aluminum (Al) and manganese (Mn) was studied in nine pairs of treated and control (no maintenance) catchments located in southern and central Finland. In this study we extended the paired catchment approach by combining data from several catchments and identifying the treatment effect on SS and element loads from the entire dataset. Following the method of LaurÃ©n et al. (2009) we identified how uncertainty in correlation between treatment and control catchments during pre-treatment period is reflected in the estimated treatment effect on SS and element loads. In the experiment, the export of SS increased significantly for the four year study period following the ditch network maintenance and Al export increased for one year. The export of N, P and Fe was not significantly changed and DOC and Mn export decreased after the ditch maintenance operation.
Immediate effects of logging, mounding and removal of logging residues and stumps on coarse woody debris in managed boreal Norway spruce stands
Authors: Rabinowitsch-Jokinen Ruut , Vanha-Majamaa Ilkka
Published: March 2014

Wood fuel production has increased remarkably, but its environmental effects within the forest ecosystem have not yet been studied much. We investigated the immediate effects of two series of forest management treatments, which produce timber and forest chips, on the volume and decay classes of coarse woody debris (CWD). One of the treatment series included logging and residue harvesting (LRH) and mounding (M), while the other series included LRH and mounding combined with stump harvesting (MSH). We hypothesized that, i) LRH reduces CWD, excluding stumps; ii) the more intense the soil preparation treatment is, M vs. MSH, the more CWD is destroyed; iii) both LRH and soil preparation treatments (M and MSH) reduce the occurence of snags, highly decayed CWD and deciduous CWD in particular. Ten sample plots in mature managed Norway spruce (Picea abies (L.) H. Karst.) dominated forests were located in Southern Finland. The total volume of CWD on the sample plots was measured three times: before and after LRH, and after M or MSH. LRH significantly decreased the volume of snags and the combined volume of snags and logs. MSH significantly decreased the total volume of CWD, while M had no significant effect on the volume of CWD. The middle and highly decayed CWD were destroyed most easily in the treatments.
Regional potential yields of short rotation willow plantations on agricultural land in Northern Europe
Authors: Mola-Yudego Blas
Published: March 2014

The development of short rotation forestry for bioenergy requires accurate and reliable yield estimates. This paper analyses the current, expected and potential regional productivity of short rotation willow plantations for six countries in Northern Europe. The estimations for present productivity are based on empirical models, using data regarding management, and local productivity based on the regional cereal yields. The estimates of expected yield rely on the current trends of yield increase from commercial willow plantations in the region. The estimates for potential yield are based on climatic restrictions. The results show potential average yields of 9.5, 6.8, 7.9, 9.0, 9.3, and 8.0 odt haâ1 yrâ1 for Denmark, Finland, Estonia, Latvia, Lithuania and Sweden, respectively. The results of the study also show that there is a wide regional variation between the different countries. In Denmark, Finland and Sweden there is a convergence between the future forecasts and the climatic potential yields in the areas of high productivity. The Baltic countries seem to present lower estimates of present productivity, reflecting possible socio-economic restrictions, although they show a high biomass potential. The methods presented in this study can be further developed in other areas where willow cultivation is considered, and can serve as a basis for future economic considerations.
Using mixed estimation for combining airborne laser scanning data in two different forest areas
Authors: Suvanto Aki , Maltamo Matti
Published: March 2014

Airborne laser scanning (ALS) data have become the most accurate remote sensing technology for forest inventories. When planning new inventories the costs of fieldwork could be reduced if datasets of old inventory areas are effectively reused in the new area. The aim of this study was to apply mixed estimation using a combination of existing and new field datasets in area-based approach. Additionally, combining datasets with mixed estimation was compared with constructing new local models with smaller datasets. The two forest study areas were in Juuka and Matalansalo, which are located about 120 km apart in eastern Finland. ALS-based regression models were constructed using datasets of Matalansalo (472 reference plots) and Juuka (10â212 reference plots). Models were developed for the basal area median tree diameter and height, mean tree height, stem number, basal area and volume. The work was based on a simulation approach which involved five methods for approximating the regression coefficients. The first method merged the datasets using ordinary least squares (OLS) regression models, whereas the second and third methods combined datasets using mixed estimation on different weighting principles, and the final two estimated local models with predetermined and new independent variables. The results indicate that mixed estimation can improve the accuracy of derived stand variables compared with basic OLS models. Additionally, a sample of 40â50 plots was enough to build local models for basal area and volume and produce at least the equal accuracy of results than any other methods in this study.
Operational efficiency and damage to sawlogs by feed rollers of the harvester head
Authors: Nuutinen Yrjö , Väätäinen Kari , Asikainen Antti , Prinz Robert , Heinonen Jaakko
Published: March 2014

In mechanical cutting, deep damage caused by feed rollers can reduce the yield of good quality timber for the sawmill and plywood industries. Additionally the feeding and energy efficiency of feed rollers are important for the profitability of harvester cutting. The objectives of this study were to compare the damages to sawlogs, as well as the time and fuel consumption of stem feeding with six different steel feed rollers during the processing of stems using a single grip harvester. This study tested two rollers with big spikes, two rollers with small spikes, one roller with studs in v-angle and one roller with adaptable steel plates in the ring of the roller. A highly detailed, and accurate processing and fuel consumption projection was recorded using the harvesterâs automated data collector on a log and stem level. The roller adaptable plate averaged, for unbarked sawlogs, the lowest damages of 3.7 mm. While the damages of the roller with big spikes were the deepest with an average of 7.8 mm. For medium stems, volume of 0.35 m3, the range of differences between the maximum and minimum effective feeding time per roller was 6â19%, which would increase the effective time consumption of cutting by 1â3%. Corresponding differences in fuel consumption during total stem processing were in the range of 7â15%. According to this study it can be concluded that the traditional rollers with spikes were most effective in processing and fuel consumption, but at the same time they caused the deepest damages to the sawlogs. The roller type with adaptable steel plates was the most effective for small stems, additionally it also caused the least damage to the sawlogs.
Leaf number indicates salt tolerance of young seedling families of European aspen (Populus tremula L.) growing in different soils
Authors: Vaario Lu-Min , Yrjälä Kim , Rousi Matti , Sipilä Timo , Pulkkinen Pertti
Published: March 2014

Soil salinity limits plant productivity and quality. We evaluated the response of 12 aspen (Populus tremula) families to salt stress in two different soils irrigated for 4-weeks with 0, 80 or 160 mM saline solution. Easily measurable characteristics such as shoot height, leaf number, dry mass as well as the distribution of sodium (Na) ions were measured in 5-month-old aspen seedlings raised in controlled greenhouse conditions on two different soils. Growth among families varied significantly, and the interaction between family and soil type was significant. From 2â5 months, leaf number correlated with that of the first month and salinity tolerance. Sodium ions varied significantly within plants and among families; seedlings that accumulated higher Na concentrations in root had more leaves and lower Na in shoot. These results suggest that leaf number indicates salt tolerance in young seedlings. Seedling performance was also affected by soil type, especially the root/shoot ratio, suggesting an interaction between salt tolerance and growth medium. This study has identified significant intra-specific variation in salt tolerance of aspen in 160 mM saline and highlighted the potential to select and develop a method for efficient pre-screening of trees to be used in the reclamation of salt-affected land.+++
B-SAP markers derived from the bacterial KatG gene differentiate populations of Pinus sylvestris and provide new insights into their postglacial history
Authors: Androsiuk Piotr , Zielinski Roman , Polok Kornelia
Published: March 2014

The aim of the studies was to evaluate the efficiency of the KatG gene based B-SAP markers as a tool to distinguish morphologically diversified and geographically distant Scots pine populations and to track the routes of migrations. The 19 populations growing in the IUFRO 1982 provenance experiment and representing the natural distribution of the species in Europe were scored using 103 B-SAP loci. Among them 26% loci were polymorphic. The level of polymorphism was associated with the location of primers on the KatG template. The diversity was low, H = 0.086, and deposited mostly among populations. Seven unique markers were found that identified populations and likely they were associated with morphology. The overall genetic identity was relatively low, I = 0.933 (D = 0.069). The block of six B-SAP markers discriminated populations into two groups in agreement with their geographic origin and thereby further described as the North and the South. The North group was uniform with genetic diversity, H = 0.026 and the overall genetic distance D = 0.022. Presumably, it migrated from refugia in the Alps via France, northern Germany and Denmark, to Scandinavia and Russia. The South group was heterogeneous with H = 0.063 and D = 0.047. This group migrated from the Carpathians via Slovakia to Germany and Poland. The Balkans and Asian refugia did not take part in recolonization of Europe. The block of six B-SAP/KatG markers can be recommended for tracking postglacial history of Scots pine.eee
Snag characteristics and cavity-nesting birds in the unmanaged post-fire northeastern Canadian boreal forest
Authors: Lowe Jeovanna , Pothier David , Savard Jean-Pierre , Rompré Ghislain , Bouchard Mathieu
Published: March 2014

We studied the availability and characteristics of snags and their use by cavity-nesting birds in the northeastern part of the Canadian boreal forest. We built up two long-term (> 200 years) chronosequences following time since the last fire in the unmanaged boreal forest of northeastern QuÃ©bec, one in the balsam fir-white birch domain (southern region) and one in the spruce-mosses domain (northern region). We then sampled and characterized snags and live trees in 30 stands from each of these two chronosequences. We also looked for nest cavities on all sampled snags, performed bird inventories by point counts, and calculated tree mortality rate from permanent sample plots. Results show that mortality rates follow a U-shaped pattern, with more snags of large diameter (> 20 cm DBH) in young (< 50 years) and in old (> 200 years) forests. In the latter, we also found more nest cavities than in any other age classes. Although abundance of primary cavity nesters (excavating species) did not vary among age classes, secondary cavity nesters (using cavities already available) tend to be more numerous in older forests. Our results highlight the capacity for young and old-growth forests to provide quality habitat for species that are dependent on large snags. Proper forest management should maintain a mosaic of different age forest stands, including snags, to promote biodiversity and provide important resources for resident bird species.
Examining the effect of environmental certification, wood source, and price on architects’ preferences of hardwood flooring
Authors: Macias Natalie , Knowles Chris
Published: March 2014

This article examines the importance architects place on three factors, environmental certification, wood source, and price, when specifying hardwood flooring. Architects were presented with nine flooring scenarios, in which the three factors were present in varying levels. They were asked to rank the scenarios from the least preferred to the most preferred. Data were obtained from a mail survey of architects in Oregon and Washington, U.S.A. (n = 402). Conjoint analyses determined that architects consider price and wood source to be the most important factors when specifying hardwood flooring. Interestingly, environmental certification was considered the least important factor. The respondents were then separated into three groups for further analysis based on whether they identified themselves as more influenced by environmental factors (biocentric) or human needs (anthropocentric). This analysis showed that the biocentric group favored wood source over price and environmental certification, while the anthropocentric group favored price.
Estimating tree biomass of sub-Saharan African forests: a review of available allometric equations
Authors: Henry Matieu , Picard Nicolas , Trotta Carlo , Manlay Raphaël , Valentini Riccardo , Bernoux Martial , Saint-André Laurent
Published: March 2014

In response to the growing interest in estimating carbon stocks in forests, available allometric equations have been compiled for sub-Saharan Africa. Tree, sprout and stand volume and biomass equations were reviewed. The 850 equations and 125 related references were incorporated into an open-access database on the Carboafrica website (http://www.carboafrica.net). The collected information provides a basic tool for the estimation of biomass and carbon stocks and other purposes, such as bioenergy and fodder supply assessment. A Tier-method approach was developed to illustrate the possible use of the equations. Current available biomass expansion factors that are used to convert a volume to the total aboveground biomass appear to be limited; incomplete species-specific allometric equations are preferred to generalised equations. The analysis of the database highlighted important gaps in available tools to assess forest carbon stocks and changes in these stocks. A quality control assessment revealed that 22% of the equations were misreported and recommendations were proposed to guide further research. Further statistical analyses, such as the Bayesian approach, would help to produce more accurate biomass estimates.
Factors influencing endophytic communities in poplar plantations
Authors: Martín-García Jorge , Espiga Elba , Pando Valentín , Diez Julio
Published: March 2014

The fungal species associated with leaves and twigs from stands of Populus x euramericana in northern Spain were studied with the aim of evaluating the effects of several factors on endophytic communities in these plantations. Endophyte assemblages were analysed in 12 poplar plantations (clone I-214), chosen according to a factorial scheme with two factors: age and site quality. Crown condition, dendrometric variables and foliar nutrients were recorded in each sampled tree to evaluate their effects on endophytic communities. Fungal species richness and relative isolation frequency (RIF) were higher in young stands than in adult stands. Moreover, the age-related differences depended on site quality, with the lowest richness levels observed in adult stands located in poor sites. At stand level, endophyte assemblages varied among stands according to site quality and, to a lesser extent, stand age. On the other hand, crown discoloration, total height and foliar concentrations of iron and zinc may be key indicators of endophytic communities in poplar plantations, at tree level.
Morphological mechanism of growth response in treeline species Minjiang fir to elevated CO2 and temperature
Authors: Hou Ying , Qu Jintao , Luo Zukui , Zhang Chao , Wang Kaiyun
Published: March 2014

To test whether and how morphological traits are linked with growth responses of plants to temperature and CO is important for understanding the mechanism underlying how plant growth will respond to global warming. In this study, using closed-top chambers to mimic future elevated CO and temperature, the growth response, morphological traits of Minjiang fir (Abies faxoniana Rehd.et Wils.) and the relationship of the two were investigated after two years of exposure to the single and combined elevation of CO and temperature. The results showed that biomass of Minjiang fir was 21%, 31%, and 35% greater than the control in elevated CO, elevated temperature and the combination of elevated CO and temperature treatments, respectively. Elevated CO and temperature significantly affected the morphology of Minjiang fir, and a few morphological traits were highly correlated with growth responses. Larger branch angles at the upper layer, crown volume, and relative crown length contributed to positive growth responses to elevated CO, while decreased specific leaf area (SLA) constricted any further growth response. Leaf morphological traits were more closely correlated with the response ratio than crown did in the elevated temperature, while in the combination of elevated CO and temperature, crown was more correlated with the response ratio than the leaf morphological traits. Thus, our results indicate that morphological traits may contribute differently to growth responses under different experimental conditions.22222222
Green-tree retention and life after the beetle: stand structure and small mammals 30 years after salvage harvesting
Authors: Sullivan Thomas , Sullivan Druscilla , Lindgren Pontus , Ransome Douglas
Published: March 2014

We report on a retrospective investigation of the impacts of salvage harvesting of lodgepole pine (Pinus contorta var. latifolia Engelm. ex S. Wats.), killed by an outbreak of mountain pine beetle (Dendroctonus ponderosae Hopk.) in the 1970s, with variable retention of Douglas-fir (Pseudotsuga menziesii (Mirbel) Franco). Our inference to biodiversity was coniferous stand structure and four mammal species: the southern red-backed vole (Myodes gapperi Vigors), common shrew (Sorex cinereus Kerr), red squirrel (Tamiasciurus hudsonicus Erxleben) and northern flying squirrel (Glaucomys sabrinus Shaw). We tested hypotheses that, at 30 years after salvage harvest of beetle-killed lodgepole pine trees, (1) abundance and diversity of stand structure, and (2) abundance of mammal species, will increase with higher levels of green-tree retention (GTR). Stand structure attributes and small mammals were sampled during 2005â2008 in young pine stands, with a range of GTR seed-trees (none, dispersed, and aggregated Douglas-fir), and uncut forest in south-central British Columbia, Canada. Diameters and heights of Douglas-fir and lodgepole pine and basal area of total conifers supported hypothesis (1). Mean abundance of the red-backed vole was consistently higher (2.3 to 6.4 times) in the uncut forest than other stands. Overall mean patterns of abundance for common shrews, red squirrels, and northern flying squirrels were similar among treatment stands. Mean abundance of the red-backed vole supported hypothesis (2), but numbers of the other three species did not. There is âlife after the beetleâ at 30 years after salvage harvesting, and this was enhanced by GTR.
Improvement of low level bark beetle damage estimates with adaptive cluster sampling
Authors: Coggins Sam , Coops Nicholas , Wulder Michael
Published: March 2014

Detection of low level infestation in forest stands is of principle importance to determine effective control strategies before the attack spread to large areas. Of particular concern is the ongoing mountain pine beetle, Dendroctonus ponderosae (Hopkins) epidemic, which has caused approximately 14 million hectares of damage to lodgepole pine (Pinus contorta Dougl. ex. Loud var. latifolia Engl.) forests in western Canada. At the stand level attacked trees are often difficult to locate and can remain undetected until the infestation has become established beyond a small number of trees. As such, methods are required to detect and characterise low levels of attack prior to infestation expansion, to inform management, and to aid mitigation activities. In this paper, an adaptive cluster sampling approach was applied to very fine-scale (20 cm) digital aerial imagery to locate mountain pine beetle damaged trees at the leading edge of the current infestation. Results indicated a mean number of 7.36 infested trees per hectare with a variance of 18.34. In contrast a non-adaptive approach estimated the mean number of infested trees in the same area to be 61.56 infested trees per hectare with a variance of 41.43. Using a relative efficiency estimator the adaptive cluster sampling approach was found to be over two times more efficient when compared to the non-adaptive approach.
Aggregating harvest activities in long term forest planning by minimizing harvest area perimeters
Authors: Öhman Karin , Eriksson Ljusk
Published: March 2014

The study presents a new approach for aggregating stands for harvest in strategic forest planning. In fragmented landscapes this could benefit nature conservation as well as reduce costs. The approach is built on the idea of minimizing the outside perimeter of contiguous harvest areas. The formulation allows for the use of exact solution methods such as mixed integer programming. The method was tested in a landscape consisting of 2821 stands. The application showed that large and compact harvest areas were created with limited sacrifice of financial value. The mixed integer programs were in most cases solved within a couple of hours. The method needs to be tested on different landscapes with different degrees of fragmentation. It is also necessary to evaluate the long term consequences of the large clear cuts that appear to be a consequence of this problem formulation.
Microdissection to isolate vascular cambium cells in poplar
Authors: Goué Nadia , Noël-Boizot Nathalie , Vallance Michel , Magel Elisabeth , Label Philippe
Published: March 2014

Vascular cambium is the lateral meristem producing xylem cells inwards and phloem cells outwards in plant stem. Thus, in trees, the quality and quantity of wood is a result of highly regulated developmental process depending initially on the vascular cambium cell production. The availability of accurate transcriptomics technologies based on high coverage sequencing raises the level of expectations on tissue sampling to a very high degree. What is the benefit of top-level transcriptomics in wood formation studies if we are using these technologies on raw tissues, mixing cells at the organ level or even higher scale? The presented work describes a nine-step procedure, from standing tree to isolated ray and fusiform cells from cryolyophilized tangential sections of the poplar cambial zone. The aim of this paper is to present a step by step procedure including advices on how to select the optimal tree, how to fell the tree while securing its physiological parameters, how to cryolyophilize and microdissect under binocular, presenting the time schedule of the whole process and RNA analysis.
Soil preparation reduces pine weevil (Hylobius abietis L.) damage on both peatland and mineral soil sites one year after planting
Authors: Luoranen Jaana , Viiri Heli
Published: March 2014

We studied pine weevil (Hylobius abietis (L.)) feeding damage to Norway spruce and Scots pine seedlings planted in regeneration areas located on peatlands or on mineral soil sites in Southern and Central Finland. The survey included two planting years and a total of 60 regeneration areas (40 areas on peatlands and 20 on mineral soil sites). Some sites classified as peatland were as transformed or transforming drained peatlands that also contained mineral soil on a prepared surface. The soil preparation method, type of surface material around a seedling, pine weevil, vole-induced or other damage and the health of each seedling were observed in systematically selected circular sample plots. There was slightly more pine weevil damage on peatland than on mineral soil sites. More seedlings were damaged on unprepared peat and humus than on a prepared surface. Seedlings surrounded by a prepared surface had a slightly greater risk of being gnawed by pine weevil when planted on prepared peat compared to planting on prepared mineral soil. Vole damage was observed only in one region during one year. Mounded areas had slightly less vole damage than patched areas. In order to reduce damage caused by pine weevils and voles, it is important to scarify the regeneration area properly before insecticide-treated seedlings are planted. Mounding and patching are recommended so that seedlings can be planted in mineral soil whenever possible, even in the case of peatlands.
Spatial structure in four Norway spruce stands with different management history in the Alps and Carpathians
Authors: Lamedica Silvia , Lingua Emanuele , Popa Ionel , Motta Renzo , Carrer Marco
Published: March 2014

In Europe most Norway spruce (Picea abies (L.) Karst.) mountain forests have been altered by human activities, leading to a lack of reference condition concerning their original structure. Nonetheless, remnants of Norway spruce primeval forests still exist in the Carpathians. Our objective was to assess the differences in structure between managed and unmanaged stands, concerning diameter distributions, amount of standing deadwood, spatial distributions and spatial structure of trees. We established four permanent plots: one in a virgin forest in the Eastern Carpathians and three in a previously managed forest in the Alps. In each plot, species, DBH, and position of the live and dead standing trees were collected. Spatial distribution and structure of all the trees was analysed through several indices. In the Carpathians forest there are clear signs of natural density-dependent mortality processes whereas in the Alpine plots such dynamics are less evident. In these latter plots, the lower snags volume and the random trees spatial distribution can be considered the legacies of past management. Nonetheless, despite the different history of the four stands, they all seem to converge towards a similar spatial structure with the presence of groups (30â40 m) of trees of similar size.
Reappearance of old-growth elements in lowland woodlands in northern Belgium: Do the associated species follow?
Authors: Vandekerkhove Kris , De Keersmaeker Luc , Walleyn Ruben , Köhler Frank , Crevecoeur Luc , Govaere Leen , Thomaes Arno , Verheyen Kris
Published: March 2014

The forest cover of the western European lowland plain has been very low for centuries. Remaining forests were intensively managed, and old-growth elements like veteran trees and coarse woody debris became virtually absent. Only over the last decades have these old-growth elements progressively redeveloped in parks, lanes and forests, and have now reached their highest level over the last 500â1000 years. Biodiversity associated with these old-growth elements makes up an important part of overall forest biodiversity. The ability of species to recolonise the newly available habitat is strongly determined by limitations in their dispersal and establishment. We analyse the current status and development of old-growth elements in Flanders (northern Belgium) and the process of recolonisation by means of specific cases, focussing on saproxylic fungi and saproxylic beetles. Our results show that âhotspotsâ of secondary old growth, even isolated small patches, may have more potential for specialised biodiversity than expected, and may provide important new strongholds for recovery and recolonisation of an important share of old-growth related species.
Dynamic old-growth forests? A case study of boreal black spruce forest bryophytes
Authors: Fenton Nicole , Bergeron Yves
Published: March 2014

Old-growth forests have sparked significant interest over the last twenty years and definitions have evolved from structure based to process based, acknowledging the diversity of forests that could be considered old growth. However studies frequently group all forests over a certain age into a single type, negating the dynamic processes that create old growth. In this study we examine a 2350-year chronosequence in boreal black spruce forests in northwestern Quebec to determine whether continued community change can be observed in the bryophyte layer. Bryophytes dominate the understory of boreal forests and influence ecosystem functioning, particularly in paludified forests where production exceeds decomposition in the organic layer. Community composition and richness changed throughout the chronosequence with no evidence of a steady state associated with an old-growth phase. In contrast the bryophyte community continued to evolve with multiple phases being evident. These results suggest that old-growth forests on the Clay Belt of northwestern Quebec and northeastern Ontario, Canada, should be regarded as part of the continuous gradient in forest development rather than a single state. This complicates conservation of these forests as multiple phases should be considered when planning forest reserves.
A comparison of avian diversity in spruce monocultures and spruce-birch polycultures in southern Sweden
Authors: Felton Adam , Andersson Erik , Ventorp David , Lindbladh Matts
Published: March 2014

The replacement of some spruce monocultures with stands composed of planted Norway spruce (Picea abies) and naturally regenerated birch (Betula spp.) has a range of potential benefits, but the implications for biodiversity are generally unknown. Here we conduct a paired replicated study in southern Sweden of the avian biodiversity found within Norway spruce monocultures, and within Norway spruce stands possessing approximately 20% birch. Our research leads us to three findings. First, avian diversity was significantly higher in the spruceâbirch polycultures. Second, spruceâbirch polycultures exclusively attracted broadleaf-associated bird species and retained the majority of conifer-associated bird species found in the spruce monocultures. Third, avian biodiversity within the spruceâbirch polycultures did not incorporate threatened taxa. We suggest that in addition to the apparent benefits for stand level diversity, widespread use of spruceâbirch polycultures could provide a means of softening the matrix for broadleaved-associated species, while concurrently providing an increased broadleaf base from which future conservation actions could be implemented. Our results are relevant to multi-use forestry, and recent policy initiatives by forest certification agencies which aim to increase broadleaf-associated biodiversity within conifer-dominated production forest landscapes.