bluuebunny/crossref_metadata_2025 huggingface dataset:
Redefinition and revision of Lydda Westwood, 1840 with taxonomic notes on Diostrombus Uhler, 1896 (Homoptera: Fulgoroidea, Derbidae)
Authors: Van Stalle Jan
Published: July 2010

A redefinition and taxonomic revision is given of the genus Lydda. Twenty-one species are attributed to this genus occurring in New Guinea and northern Australia. The zoraidine genera with short antennae are keyed and all Lydda species are keyed and redescribed with emphasis on the male and female genitalia. Four species are described as new: Lydda perlucida, L. guttata, L. cornuta and L. fusca; 16 species are transferred from Proutista to Lydda given 16 new combinations. P. awarensis Van Stalle, 1986 is synonymized with L. gemina Muir, 1913. Arfaka Distant, 1907 and Afakia Kirkaldy, 1909, former synonyms of Proutista are now considered as synonyms of Lydda. The Afrotropical species attributed to Lydda are transferred to Diostrombus, new subgenus Lyddastrombus, giving seven more new combinations. Proutista jezeki Dlabola, 1979 and P. jezeki Dlabola, 1981 are transferred to Diostrombus s. str., giving D. jezeki (Dlabola) comb. n. and D. adila (Dlabola) comb. n. A lectotype is selected for Lydda elongata, Lydda perkinsi, L. lutea, L. lurida, L. decisa, L. straminea, L. lumholtzi, L. australis, L. gemina, L. koebelei, L. calypso and Philadelpheia pandani. Diostrombus muiri nom. n. is proposed for D. cocos Muir, 1928, a secondary junior homonym of D. cocos (Muir, 1926) comb. n.
The fruit flies of the Canary Islands (Diptera: Tephritidae)
Authors: Merz Bernhard
Published: July 2010

This study updates our knowledge of the Tephritidae of the Canary Islands. The 30 recorded species are keyed and briefly discussed. Tephritis luteipes sp. n. is described and Trupanea tubulata Munro is recorded for the first time from the Canary Islands. Three records are regarded as erroneous: Aciura coryli (Rossi), Dectodesis augur (Frauenfeld) and Paroxyna absinthii (Fabricius). Tephritis reticulata Becker, T. duplex Becker and Paroxyna multimaculosa Dirlbek & Dirlbek are transferred to Campiglossa (combs. n.). The following new synonymies are proposed (junior synonyms in parantheses): Capitites ramulosa (Loew) (= Urellia perfecta Becker), Campiglossa duplex (Becker) (=Paroxyna aequalis Hering), Campiglossa reticulata (Becker) (=Tephritis occulta Becker), Oedaspis quinquefasciata Becker (= Oedaspis heringi Hendel), Tephritis praecox (Loew) (= Tephritis poecilura Loew). Lectotypes are designated for Paroxyna aequalis Hering, Tephritis reticulata Becker, Tetanocera walkeri Wollaston and Trypeta praecox Loew. Seven fruit fly species were reared for the first time, and 12 new fruit fly - plant relationships are mentioned. Furthermore Paroxyna martii (Becker) and Urelliosoma guimari (Becker) are recorded for the first time from Southern Spain and Tephritis praecox from Austria.
A new fossil tortricid (Lepidoptera: Tortricidae) from Dominican amber
Authors: Poinar George O. , Brown John W.
Published: July 2010

Poinar Jr, G.O. & Brown, J.W.: A new fossil tortricid (Lepidoptera: Tortricidae) from Dominican amber. Ent. scand. 23: 25-29. Copenhagen, Denmark. April 1993. ISSN 0013-8711. A tortricid, presumably of the tribe Polyorthini of the subfamily Chlidanotinae (Tortricidae: Lepidoptera), Polyvena horatis gen. n., sp. n., is described in amber from the Dominican Republic. The present find represents the first tortricid to be described from Dominican amber and the first possible fossil of the subfamily Chlidanotinae. Diagnostic subfamily characters include a generalized wing venation with widely distant M2 and M3 of the hindwing, raised scale tufts on the forewing and large valvae without a corresponding large tegumen.
The first Palaearctic record of the mycetophilid genus Drepanocercus Vockeroth (Diptera: Mycetophilidae)
Authors: Söli Geir E.E.
Published: July 2010

Söli, G. E. E.: The first Palaearctic record of the mycetophilid genus Drepanocercus Vockeroth (Diptera: Mycetophilidae). Ent. scand. 24: 73-77. Copenhagen, Denmark. April 1993. ISSN 0013-8711. Drepanocercus spinistylus sp. n. is described, based on one male captured in a Malaise trap in Jostedalen, West Norway. The genus was previously monotypic, represented by the Nearctic species, Drepanocercus ensifer Vockeroth, 1980. The male genitalia of both species are figured, and a revised diagnosis of the genus is given. Drepanocercus is suggested to be most closely related to Paratinia Mik, 1874.
Review of the Incurvaria vetulella species-group in the Alps (Lepidoptera: Incurvariidae)
Authors: Huemer Peter
Published: July 2010

Huemer, P.: Review of the Incurvaria vetulella species-group in the Alps (Lepidoptera: Incurvariidae). Ent. scand. 24: 109-120. Copenhagen, Denmark. April 1993. ISSN 0013-8711. The Incurvaria vetulella species-group in the Alps is defined and reviewed. The following species are recognized: vetulella (Zetterstedt, 1839) (eastern Alps), triglavensis Hauder, 1912 stat. n. (southeastern Alps) and ploessli sp. n. (southwestern Alps). All species are described and/or redefined and adults, male and female genitalia are illustrated. Furthermore, Scandinavian specimens of vetulella and circulella (Zetterstedt, 1839) are illustrated for comparison. The considerable external variation between various alpine populations of vetulella is discussed and regarded as infraspecific. A lectotype of triglavensis is designated.
The Erythrothrips complex of tropical Aeolothripidae (Thysanoptera) with new taxa from Australia and South Africa
Authors: Marullo Rita , Mound Laurence A.
Published: July 2010

The New World genus Erythrothrips is redefined and a key to the species provided. A new genus and species, Erythridothrips cubilis, is described from Australian tropical rain-forest. The South African genus Audiothrips is withdrawn from synonymy with Erythrothrips and a new species, A. senckenbergiana, is described. The sole African species in the American genus Stomatothrips is transferred to the African genus Allelothrips. The relationships of these taxa are discussed, together with reasons for questioning the validity of the tribe Orothripini.
Osrhoes coronta Druce, the New World palaeosetid moth: A reappraisal, with description of a new type of female genital apparatus (Lepidoptera, Exoporia)
Authors: Nielsen Ebbe S. , Kristensen Niels P.
Published: July 2010

The monobasic Colombian genus Osrhoes is redescribed on the basis of the male holotype and a female paratype of O. coronta Druce, 1900. Osrhoes is unique among the exoporian moths so far known in having a long internal duct from the ductus bursae to a chamber adjacent to the ovipore, a functional analogy of the 'ductus seminalis' of ditrysian Lepidoptera; the subgenital plates are very large and completely fused in the midline, i.e., there is no 'intergenital cleft'. Strong reasons for retaining Osrhoes in the non-ditrysian grade are the homoneurous venation and large forewing jugal lobe, while apomorphies supporting its assignment to the Exoporia-Hepialoidea include, e.g. its elongate intercalary sclerite, postapical Rs3, male genitalia with typical hepialoid hinged juxta/trulleum complex and lack of sclerotized phallus. The absence of inter-M crossveins and the extreme reduction of the maxillae are currently considered diagnostic traits of the family Palaeosetidae, which is otherwise represented by three small Australasian genera. The wing proportions and absence of a forewing 'anal loop' in Osrhoes probably also indicate relationships to palaeosetids. Although the status of the crossvein character as an apomorphy is debatable, it is preferred at present to uphold the tentative àssignment of Osrhoes to the Palaeosetidae.
A phylogenetic study of the species of the genus Vespa (Hymenoptera: Vespinae)
Authors: Archer Michael E.
Published: July 2010

A phylogenetic analysis with the program Hennig86 is performed on the 23 species of the genus Vespa, using the Polistinae and the remaining species of the Vespinae as outgroups. Two species, V. basalis and V. binghami, are unplaced while the remaining 21 species are resolved as a separate group. Within this group the following species groups are resolved as being of uncertain relationships: V. affinis group with 13 species, V. tropica group with five species, V. crabro group with two species and V. orientalis.
Cladistic relationships among higher groups of Heteroptera: congruence between morphological and molecular data sets
Authors: Wheeler Ward C. , Bang Ranhy , Schuh Randall T.
Published: July 2010

The monophyly of the 7 infraorders of Heteroptera and history of higher group concepts and interrelationships within the Heteroptera are briefly reviewed. Data from 31 morphological characters are combined with 669 bases of 18S nuclear rDNA for 29 taxa, including several outgroups to the Heteroptera, to produce a phylogeny based on the total available evidence. The molecular data alone and in conjunction with morphological data indicate that: the Homoptera are probably not monophyletic; the Auchenorrhyncha are the sister group of Coleorrhyncha + Heteroptera ; the Enicocephalomorpha are the sister group of remaining Heteroptera; the Dipsocoromorpha are the sister group of remaining Heteroptera; the Gerromorpha are the sister group of remaining Heteroptera; the Nepomorpha are the sister group of remaining Heteroptera; the Leptopodomorpha are the sister group of the Cimicomorpha + Pentatomomorpha. The molecular evidence corroborates the morphologically based theory of a sister group relationship between Aradoidea and trichophoran Pentatomomorpha. This scheme deviates from that previously published by Schuh, in which the Leptopodomorpha were treated as the sister group of the Nepomorpha.
Apicrenus fossilis gen. & sp. n. (Heteroptera: Reduviidae: Apiomerinae) from Dominican amber (lower Oligocene - upper Eocene)
Authors: Maldonado Capriles Jenaro , Santiago-Blay Jorge A. , Poinar George O.
Published: July 2010

Apicrenus fossilis, a new neotropical apiomerine fossil genus, is described and compared with annectant genera. This new genus is distinguished from other consubfamilials by the following combination of characters: anteriorly located eyes, with their margins flushing with the lateral margins of the head, anterior acetabula not visible from above, absence of collar, and presence of two triangular discal corial cells in the fore wing.
Libanorhinus succinus gen. & sp. n. (Coleoptera: Nemonychidae) from Lebanese amber
Authors: Kuschel G. , Poinar G.O.
Published: July 2010

The nemonychid, Libanorhinus succinus gen. & sp. n. represents the first weevil to be reported from Lebanese amber and the first formal description of a representative of the family Nemonychidae from any amber source. The specimen is placed in the extinct subfamily Eobelinae on the basis of its elytral punctures lined up to form striae, the presence of scutellar strioles and the possession of simple claws. The vertex of the head, antennal insertions at about the apical quarter of the rostrum and abdominal ventrites distinguish it from previously described fossil species in the Eobelinae. Since many extant nemonychids feed and develop in the male cones of representatives of the Araucariaceae, the present fossil could have developed in the cones of this resin-producing tree family.
Revision of Cornutrypeta Han & Wang, a new tephritid genus proposed for Oriental and Palaearctic species (Diptera: Tephritidae)
Authors: Ke Chung Kim  , Han Ho-Yeon , Wang Xing-Jian
Published: July 2010

A new genus, Cornutrypeta Han & Wang, is proposed for nine Palaearctic and Oriental tephritid species with enlarged male frontal setae: C. gigantocornuta Han & Wang, sp. n., C. melanonotum (Brunetti), comb. n., C. nigrifemur Han & Wang, sp. n., C. nigritata (Wang), comb. n., C. omeishana Han & Wang, sp. n., C. spinifrons (Schroeder), comb. n., C. superciliata (Frey), comb. n., C. triceratops (Bezzi), comb. n., and C. yushunia Han & Wang, sp. n. All the previously recorded species are transferred from Vidalia Robineau-Desvoidy. A lectotype is designated for C. superciliata. Cornutrypeta is placed in the newly recognized Trypeta group of the tribe Trypetini. A key to the species, descriptions, illustrations, and a cladogram are presented. We also resurrect Stemonocera Rondani, which was considered a synonym of Vidalia.
Xiaomyia, Shangomyia and Zhouomyia, three new and unusual genera of Chironomini from Oriental China (Diptera: Chironomidae)
Authors: Xinhua Wang  , Sæther Ole A.
Published: July 2010

Xiaomyia aequipedes gen. & sp. n. from Guandong in Oriental China, Shangomyia impectinata gen. & sp. n. from Hainan in Oriental China and Burma, and Zhouomyia plauta gen. & sp. n. from Hainan are described as male imagines. Xiaomyia plus Shangomyia in most features appear to form a separate tribe within the subfamily Chironominae, while a few characters indicate a placement in the Harnischia group of genera. They are preliminary placed at the base of the Chironomini. They are unique in having apical scales with 1-2 spines on all tibiae, but no combs; no dorsomedian eye extension; extended costa; leg ratios of all legs higher than 1.5; and inferior volsella fused to the gonocoxite. Zhouomyia is a typical plesiomorphic Chironomini either related to Apedilum Townes, Paralauterborniella Lenz, Lauterborniella Thienemann & Bause, and Pagastiella Brundin, or to the Endochironomus group of genera.
Specific status of Aquarius cinereus (Puton) and A. najas (De Geer) (Hemiptera: Gerridae) and the extent of hybridization in the Mediterranean region
Authors: Zimmermann Manfred , Scholl Adolf
Published: July 2010

Results of field observations, cross-breeding experiments, morphometry, and starch gel electrophoresis suggest that A. cinereus and A. najas are specifically distinct. Cross-breeding experiments between the two taxa in the laboratory were only successful when using males of A. cinereus and females of A. najas. Field observations on mixed populations in spring did not reveal any pre- or postcopulae involving partners of the two taxa (assigned by body length). Instead all pairs observed consisted of conspecific males and females. Morphometry of field caught adults of A. najas and A. cinereus yielded a clearly bimodal distribution of body length with only very few individuals of intermediate size. Therefore, body length alone allows reasonable separation of the two taxa in natural populations. Although F1-hybrids from laboratory rearings (photoperiod 18L:6D, temperature ∼ 23°C) differed only marginally in body length from A. najas reared under the same conditions, backcrosses and also F2-hybrids were of intermediate size. In the field individuals of both taxa are predominantly wingless, but both hybrids and backcrosses from the laboratory showed a net shift in morph ratio with many longwinged specimens. Vertical starch gel electrophoresis of three monomorphic (Apk, Got-2, Mdh-2), six polymorphic loci with low levels of polymorphism (Idh-1, Fum, Got-1, Mdh-1, Me, 6-Pgd) and three highly polymorphic loci (Es-4, Idh-2, Pgm) suggests that the two taxa do not share a common gene pool. Gene differentiation among all A. cinereus populations as measured by Nei's GST values is very low. Between Tyrrhenian and continental A. najas populations, however, gene differentiation is very high, and in the same order of magnitude as gene differentiation between the two taxa A. cinereus and A. najas in sympatric populations. Our field observations, together with morphometric data, laboratory rearings, and information on wing polymorphism, suggest that hybridization in natural populations is rare.
A new species of the ' marine' genus Thalassosmittia Strenzke & Remmert from Xizang (Tibet), China (Diptera: Chironomidae)
Authors: Xinhua Wang  , Sæther Ole A.
Published: July 2010

Thalassosmittia montana sp. n. from Xizang (Tibet) is described as male imago. A key to male imagines of the 6 known species of Thalassosmittia Strenzke & Remmert is presented. T. montana differs from the other species of Thalassosmittia, which are all marine, in having no anal lobe, retracted R4+5 of the wing, R2+3 running in the middle between R, and R4+5, very strong virga and heavily sclerotized, apically toothed phallapodeme.
New species and records of Tachiona Sharp (Coleoptera: Staphylinidae, Aleocharinae) with a revised key to known species
Authors: Ashe James S.
Published: July 2010

Four new species of Tachiona are described: T. costaricensis (type locality 8 km S Orosi, Cartago Prov., Costa Rica), T. elegans (type locality 15.4 km NW Filo de Caballo, Guerrero, Mexico) T. mexicanus Ashe (type locality 23.1 km E Vicente Guerrero [Santa Maria del Monte, V.G.], Puebla, Mexico), T. oaxacaensis Ashe (type locality 27.2 km S Miahuatlan, Oaxaca, Mexico). Illustrations of distinctive structural features are provided. New records of Tachiona monteverdensis Ashe from the states of Chiapas and Veracruz in Mexico, and Tachiona deplanata Sharp from near Jalapa, State of Veracruz, in Mexico, are listed. A key is provided for identification of the eight known species of Tachiona.
Re-evaluation of the genus Closterocerus Westwood (Hymenoptera: Eulophidae), with a revision of the Nearctic species
Authors: Hansson Christer
Published: July 2010

The taxonomic status and delimitation of the genus Closterocerus Westwood is discussed, partly on the basis of new characters. The following genera are synonymized with Closterocerus: Achrysocharis Girault (maintained as subgenus), Achrysocharella Girault, Cecidiophaga Erdös, Chrysocharidia Erdös, Halochariessa Erdös, Halocharis Erdös, Moserina Delucchi, Pseudochrysocharis Girault, Wolffiella Krausse. The genus Chrysonotomyia Ashmead is regarded as a valid genus, but most species previously recognized in Chrysonotomyia are transferred to Closterocerus. The Nearctic species of Closterocerus are revised, and a key to the species is given. Twenty-one species are recognized, of which 8 are described as new: brachyphagus, carolinae, cyaneus, longus, nitidus, nius, texanus, violaceus. Four new synonyms are suggested: Chrysonotomyia baliosa Yoshimoto and C. cunicularia Yoshimoto = Closterocerus cinctipennis Ashmead; Pleurotropis tricinctus Ashmead and Closterocerus winnemanae Crawford = Closterocerus trifasciatus Westwood. Seven Nearctic species are newly transferred to Closterocerus: Achrysocharis partifuscipennis Ashmead and Moserina maculata Delucchi from Omphale, Achrysocharis phenacaspia Yoshimoto, A. spiniformis Moser, A. vesiculis Moser, Chrysonotomyia solidaginis Yoshimoto and Wolffiella ruforum Krausse all from Chrysonotomyia. The species Closterocerus damastes Walker is reassigned to Closterocerus from Chrysonotomyia. The Palearctic species Achrysocharis lyonetiae Ferrière, Nesomyia cinctiventris Ashmead, Halochariessa germanica Erdös, Entedon lanassa Walker, Achrysocharis smaragdula Graham, Chrysonotomyia yamagishii Kamijo, Halochariessa pannonica Erdös and Halocharis transsylvanica Erdös are newly transferred to Closterocerus from Chrysonotomyia. A neotype is designated for Closterocerus cinctipennis. Lectotypes are designated for Closterocerus utahensis Crawford, C. winnemanae and Pleurotropis tricinctus.
Revision of the genera Marmottania Puton and Phasmosomus Kiritshenko (Heteroptera: Lygaeidae, Rhyparochrominae)
Authors: Linnavuori Rauno E.
Published: July 2010

The genera Marmottania Puton and Phasmosomus Kiritshenko are revised. The former is included in the tribe Ozophorini, the latter in the Phasmosomini. New combination: Marmottania priesneri Wagner = Phasmosomus priesneri (Wagner). New synonymies: Marmottania rungsi Vidal, 1951 = M. simonis Puton, 1887. Phasmosomus hoberlandti Kiritshenko, 1963 = P.priesneri (Wagner, 1958).
Systematics, phylogeny and zoogeography of genus Yoraperla (Plecoptera: Peltoperlidae)
Authors: Stark Bill P. , Riley Nelson C.
Published: July 2010

Yoraperla Ricker, 1952 is revised and 4 nearctic and 3 oriental species are recognized. Three new species, X siletz, Y. han and Y. uchidai, are described and Y. nigrisoma (Banks, 1948) is removed from the synonymy of Y. brevis (Banks, 1907). Phylogenetic and zoogeographic analyses support the hypothesis of two disjunct clades isolated by an increase in sea level in the Bering Straits. The east Asian clade is consistently resolved with the two Korean species Y. han and Y. uchidai forming a sister group relationship apart from the Japanese species Y. uenoi, while resolution of the western North American clade is less secure. Yoraperla probably arose in the Sierra Nevada of North America and migrated to east Asia along an inland route including the Rocky Mts. Future collection and conservation suggestions are given.
Revision of the genus Aspidiotes Schonherr (Coleoptera: Curculionidae, Tanymecini)
Authors: Sanchez-Ruiz M. , Alonso-Zarazaga M.A.
Published: July 2010

The genus Aspidiotes Schönherr, 1847 is revised; 6 species are recognized as valid. Amomphus Schonherr, 1848 is considered to be an unjustified replacement name for Aspidiotes Schönherr, 1847. Phaenognathus Schönherr, 1847 is retained as a subgenus of Aspidiotes although the cladistic analysis failed to find synapomorphies supporting this opinion. One new species, A. (Aspidiotes) gonzalezi, is described from NE Spain (Lerida, Thrragona and Zaragoza). A seventh species currently included in the genus, Amomphus reichei Desbrochers, 1872, is placed incertae sedis; the type specimen has not been traced. Distributional as well as biological data are provided. A cladistic analysis and a biogeographical discussion are provided.
A new phylogenetically basal subfamily of brown lacewings from Chile (Neuroptera: Hemerobiidae)
Authors: Oswald John D.
Published: July 2010

A new genus and species, Adelphohemerobius enigmaramus, is described from Chile and assigned to a new subfamily, Adelphohemerobiinae, of the neuropterous family Hemerobiidae (brown lacewings). A cladistic analysis of morphological data derived from the unique female holotype of A. enigmaramus supports the conclusion that it represents the sister-group of the family Hemerobiidae sensu Oswald (1993a). The holotype of A. enigmaramus is interpreted to be the first non-teratological hemerobiid specimen known to possess a single 'radial sector', i.e., only the true ancestral Rs.
A revision of the Braconidae (Hymenoptera) species described by J. W. Zetterstedt
Authors: Papp Jenö
Published: July 2010

Twenty-five species of Braconidae described in the genera Alysia Latreille, Bracon Fabricius, Lepton Zetterstedt, Mícrogaster Latreílle and Sigalphus Latreille by J. W. Zetterstedt in 1838 are revised. Lectotypes and paralectotypes are designated and the present status of Zetterstedt's species is established. Ten names of Zetterstedt are regarded as valid; sixten new synonyms and six new combinations are proposed. A lectotype of Bracon konovii Marshall is designated and the name is resurrected from synonymy with Cyanopterus nigrator (Zetterstedt).
The Trepobatinae (Heteroptera: Gerridae) of New Guinea and surrounding regions, with a review of the world fauna. Part 2. Tribe Naboandelini
Authors: Polhemus Dan A. , Polhemus John T.
Published: July 2010

The small waterstriders of the subfamily Trepobatinae have radiated extensively on New Guinea and surrounding archipelagos. The present contribution is the second in a series of reports dealing with this endemic fauna, and presents a revision of the tribe Naboandelini, proposed in Part 1 of this series to hold the genera Naboandelus Distant (type-genus), Hynesionella Poisson, and Calyptobates gen. n. A key to these genera is provided, followed by a taxonomic treatment of their constituent species occuring in the Australasian region. The following new taxa are proposed within Naboandelini: Calyptobates gen. n., including type-species Calyptobatesjourama sp. n. from Australia, Calyptobates amboina sp. n. from Ambon, Calyptobates andaman sp. n. from the Andaman Islands, Calyptobates minimus sp. n. from Australia, Calyptobates rubidus sp. n. from Australia, Calyptobates samarinda sp. n. from Borneo, and Calyptobates simplex sp. n. from New Guinea. The genus Naboandelus Distant is redescribed, and the following new species are added: Naboandelus borneensis sp. n. from Borneo, Naboandelus johorensis sp. n. from Malaysia, and Naboandelus taprobanicus sp. n. from Sri Lanka. The genus Hynesionella Poisson is redefined, Naboandelus capensis Poisson 1955 is transferred to the genus (comb. n.), and Hynesionella omercooperi Hungerford & Matsuda, 1959 is shown to be a synonym of capensis (syn. n.). Habitat and distributional data are given for these taxa, accompanied by figures of key characters and distribution maps.
Review of the family Ibaliidae (Hymenoptera: Cynipoidea) with keys to genera and species of the World
Authors: Zhiwei Liu  , Nordlander Göran
Published: July 2010

The archaic cynipoid family Ibaliidae is revised. All described taxa are included in a checklist, and keys to valid taxa are presented. Host records include larvae of several woodwasp species (Siricidae: Siricinae and Tremicinae). Ibaliidae is considered to include the East Asian genus Heteribalia Sakagami, with five valid species, and the mainly Holarctic genus Ibalia Latreille, with 13 valid species, seven in the subgenus Ibalia and six in the subgenus Tremibalia. Heteribalia aureopilosa Maa, 1949 and H. subtilis Maa, 1949 are raised from subspecies to species level. Ibalia (T.) hunanica sp. n., is described from Oriental China. Three morphs of I. (T.) anceps, differing in wing colour pattern, are recognized, and distribution records of these morphs in North America are mapped. I. scalpellator Westwood, 1837 is synonymized with I. anceps Say, 1824; I. takaehihoi Yasumatsu, 1937 with I. jakowlewi Jacobson, 1899; I.fulviceras Yang, 1991 with I. ornata Belizin, 1968; and I. yunshae Yang et Liu,1992 with I. rufipes drewseni Borries, 1891. Lectotypes are designated for I. picea Matsumura, 1912 (syn. of I. leucospoides (Hochenwarth, 1785)) and for I. japonica Matsumura, 1912.
A revision of the genus Epichlorops Becker (Diptera: Chloropidae)
Authors: Wheeler Terry A.
Published: July 2010

The primarily Nearctic genus Epichlorops Becker is revised. The genus comprises ten species in the Nearctic region: E. aquilonius sp. n.; E. californicus sp. n.; E. ectyphus sp. n.; E. elongatus sp. n.; E. exilis (Coquillett); E. glaphyrus sp. n.; E, mexieanus sp. n.; E. punctieollis (Zetterstedt); E. scaber (Coquillett); and E. stenostylus sp. n. Epichlorops punetieollis is the only species of the genus known from the Palaearctic region. A key and illustrations of all species are provided. The monophyly of the genus and phylogenetic relationships between the species are analysed and discussed.
Revision of the genus Paralopus Wagner (Heteroptera: Miridae, Phylinae)
Authors: Linnavuori Rauno E.
Published: July 2010

The genus Paralopus Wagner, 1957, is revised. New species: P. edithae sp.n. (Palestine).
The Isotomiella of Thailand (Collembola: Isotomidae), with description of five new species
Authors: Bedos Anne , Deharveng Louis
Published: July 2010

Eight species of Isotomiella Bagnall, five of which are new, are recorded from Thailand: I. symetrimucronata Najt & Thibaud, 1987, I. nummulifer Deharveng & Oliveira, 1990, I. cribrata Deharveng & Suhardjono, 1994, L brevidens sp. n., I. edaphica sp. n., I, hirsuta sp. n., I. inthanonensis sp. n., and I. leksawasdii sp. n. Isotomiella minor (Schaeffer, 1896), the only species previously recorded from Thailand, is probably misidentified. A key to the species of Thai Isotomiella is given.
The classification of the Timandra griseata group (Lepidoptera: Geometridae, Sterrhinae)
Authors: Albrecht Anders , Kaila Lauri
Published: July 2010

The Timandra griseata group is revised with special emphasis on the European taxa. It is concluded that two species with slightly overlapping distribution areas occur in Europe: T. griseata Petersen, 1902 which is confined to northern Europe and T. comai Schmidt, 1931 (= Calothysanis amataria brykaria Nordström, 1943, syn. n.) which is widespread in South and Central Europe. The differences in morphology, yearly occurrence and phenology between T. griseata and T. comai were studied in southern Finland. The following new combinations are proposed: Timandra Paralias (Prout), Timandra recompta ovidius (Bryk) and T. recompta prouti (Inoue). A lectotype for T. griseata Petersen is designated. T. paralias, although not a member of the T. griseata group, is redescribed because of its external resemblance.
Rostromyrmex, a new genus of myrmicine ants from Peninsular Malaysia (Hymenoptera: Formicidae)
Authors: Rościszewski Krzysztof
Published: July 2010

Rostromyrmex, a new genus of myrmicine ants from Peninsular Malaysia is described on the basis of workers, dealate females, and a single male. Analysis of characters suggests a closest relationship with either Lordomyrma or the Solenopsis genus group.
Revision of the milliped family Paeromopodidae, and elevation of the Aprosphylosomatinae to family status (Julida: Paeromopodoidea)
Authors: Shelley Rowland M.
Published: July 2010

The family Paeromopodidae is comprised of large, agile millipeds that include the longest diplopods in the Nearctic. It is endemic to three regions in the western United States - a large, irregular area extending from the southern Sierra Nevada Mountains and the Monterey Peninsula of California to the northern Puget Sound region of Washington, a small, ovoid area in the Warner Mountains of northeastern California and the adjacent fringes of Oregon and Nevada, and an area in the northwestern interior extending from the Wallowa Mountains of northeastern Oregon to Flathead Lake, Montana. Representatives have flat, plate-like gonopods that represent the coxites; the flagellum is absent from the anterior gonopods and the telopodite is lost from both gonopod pairs. The family includes two genera: Paeromopus Karsch, with five species - angusticeps (Wood), eldoradus, cavicolens, and buttensis, all by Chamberlin, and ocellatus Loomis - and Californiulus Verhoeff, with five species - chamberlini (Brolemann), comb. n., dorsovittatus Verhoeff, yosemitensis Chamberlin, euphanus (Chamberlin), and parvior (Chamberlin). The principal taxonomic features involve the configurations of the distal and midlength projections of the anterior gonopods and the terminal or subterminal projections of the posterior genitalia. Californiulus chamberlini, dorsovittatus, and yosemitensis display broad, yellow, middorsal stripes, the last also with a median black line, and the other species exhibit light or dark transverse bands. The following new synonymies are proposed: P. lysiopetalinus Karsch and pistus Chamberlin, and Paeromopellus sphinx Verhoeff, under P. angusticeps; Klansolus zantus Chamberlin under C. chamberlini; C. vicinus Chamberlin under C. dorsovittatus; K. mononus and obscurans, both by Chamberlin, under C. yosemitensis; and K. socius and Aigon rodocki, both by Chamberlin, under C. parvior. The monobasic subfamily Aprosphylosomatinae, accommodating Aprosphylosoma Hoffman, is elevated to family status in the superfamily Paeromopodoidea. Its sole species, A. darceneae Hoffman, known only from Oregon, is smallbodied, dark mottled brown in color, and has a separate coxite and telopodite on the anterior gonopod.
Eight new species and a revised key to Australian Helicopsyche (Trichoptera: Helicopsychidae)
Authors: Johanson Kjell Arne
Published: July 2010

Eight new species of Helicopsyche, viz., H. neboissi sp. n.; H. tribulationa sp. n.; H. curva sp. n.; H. bellangrensis sp. n.; H. braueri sp. n.; H. alicae sp. n.; H. albidela sp. n.; and H. kakadu sp. n., are described. The females of H. tillyardi Mosely, 1953 and H. heacota Mosely, 1953 are described for the first time. A revised description of the male and female H. cochleaetesta Korboot, 1964, female H. murrumba Mosely, 1953 and male H. tillyardi Mosely, 1953 are given. Revised keys to the males and females of Australian Helicopsyche species are presented.
Description of Dineura pullior sp. n. (Hymenoptera: Tenthredinidae), with quantified observations on saw wear
Authors: Schmidt S. , Walter G.H.
Published: July 2010

In sawfly taxonomy the structure of the lancet or the saw is an important character for identification of females. Because adult females use the saw to place eggs into the host plant its teeth may wear and their shape may thus be altered. We investigated the pattern of wear in a species of nematine sawfly that we also describe for the first time. Saw wear of females that deposited known numbers of eggs was examined. Saw wear was not strongly related to the number of eggs laid, despite some saws being considerably worn. The range of variation in saw profile illustrated by Lindqvist (1956) in the closely-related species D. virididorsata (Retzius) was not observed in our study. Despite the shape of the saw being altered to some extent with use, other structures on the saw do not change and provide good taxonomic characters in the genus Dineura, as evidenced by our detection of the cryptic (= sibling) species on the basis of differences in ctenidial structure. We describe this species under the name Dineura pullior sp. n.
Systematic revision of Ericydeus Pascoe (Coleoptera: Curculionidae)
Authors: Lanteri Analia A.
Published: July 2010

The genus Ericydeus Pascoe (Polydrosinae: Naupactini) distributed throughout the United States of America up to Argentina is revised and a cladistic analysis including 16 species is conducted. Three new species are described: E. bahiensis, E. argentinensis and E. cupreolus. Ericydeus humeralis Hustache is synonymized under E. nigropunctatus (Chevrolat); and E. modestus viridans (Boheman) and E. modestus duodecimpunctatus (Dalla-Torre, Emden & Emden) are elevated to species rank. Other valid species are: E. hancocki (Kirby), E. schoenherri (Perty), E. sedecimpunctatus (Linnaeus), E. yucatanus (Champion), E. roseiventris (Champion), E. quadripunctatus (Champion), E. modestus (Gyllenhal), E. forreri (Champion), E. lautus (LeConte) and E. placidus (Horn). In the cladogram obtained the species from South America gather in a clade (E. argentinensis, E. sedecimpunctatus, E. nigropunctatus, E. schoenherri, E. hancocki, and E. bahiensis) and the species from Central and North America form a separate clade (E. yucatanus, E. roseiventris - E. quadripunctatus, E. cupreolus, E. viridans - E. modestus, E. duodecimpunctatus, E. forreri, E. lautus - E. placidus). The character evolution follows a southern-northern direction. The paper includes a redescription of the genus, redescriptions or descriptions of its 16 species, a dichotomous key, habitus photographs, drawings of diagnostic structures, distribution maps, a cladogram, and a discussion of the phylogeny and distribution of the genus.
A comparative SEM study of the eggs of the Sepsidae (Diptera) with a cladistic analysis based on egg, larval and adult characters
Authors: Meier Rudolf
Published: July 2010

The eggs of 21 species in 12 genera of Sepsidae were studied using scanning electron microscopy. All but the eggs of Ortalischema have respiratory filaments which aid in gas exchange. Judged by positional differences and the lack of a central canal, the two filaments of Orygma are not homologous to the single filament of most remaining sepsids. Hinton's hypothesis that egg filaments are plesiomorphic for the Sepsidae is therefore rejected. Three egg characters are added to an existing data set comprising 58 taxa and 85 larval and adult characters. The egg characters allow the unambiguous placement of Lasionemopoda, for which the larval and adult data set had suggested two different positions on equally parsimonious trees. Based on the position of Lasionemopoda, the evolution of the fore femora of the Sepsidae is briefly discussed. The egg morphology of the Sepsidae is also compared to the morphology of drosophiline eggs which also have egg filaments.
A study of Zaleinae, a taxon transitional between Canacidae and Tethinidae (Diptera), with the description of a new genus and species
Authors: Freidberg Amnon
Published: July 2010

Suffomyia scutellaris gen. n. & sp. n. is described from specimens collected at the intertidal zone along the western coast of the Gulf of Elat (northern Red Sea), Sinai. This is the first representative of the subfamily Zaleinae in the Palaearctic Region. The phylogenetic relationships of the Zaleinae are discussed, and the subfamily is shown to be an intermediate group between typical Canacidae and Tethinidae, indicating that only one family is involved.
Revision of the genus Boopidocoris Reuter (Heteroptera: Miridae, Phylinae)
Authors: Linnavuori Rauno E.
Published: July 2010

The genus Boopidocoris Reuter, 1879, is revised. New species: B. nahidae sp. n. (Iran).
Phylogeny and classification of aquatic bugs (Heteroptera, Nepomorpha). An essay review of Mahner's 'Systema Cryptoceratum Phylogeneticum'
Authors: Andersen Nils Møller
Published: July 2010

This essay is essentially a review of the monographic work by the German zoologist Martin Mahner : 'Systema Cryptoceratum Phylogeneticum (Insecta, Heteroptera)' (Zoologica, Heft 143, Stuttgart 1993). The monograph is the most comprehensive systematic account of the aquatic bugs to date and the first major work on this group where the principles of phylogenetic (cladistic) systematics are consistently applied. Mahner follows the principles of the 'konsequentphylogenetische oder cladistischen Systematik', being Willi Hennig's phylogenetic systematics as interpreted and modified by Peter Ax. The methodological procedures recommended by this school of systematics is controversial, however, and call for a broader discussion of current trends in systematics as exemplified by the phylogeny and classification of the aquatic bugs.
The encyrtid genus Allocerellus Silvestri (Hymenoptera: Chalcidoidea), parasitoids of mealybugs (Homoptera: Pseudococcidae)
Authors: Prinsloo G.L.
Published: July 2010

The Encyrtidae genus Allocerellus Silvestri is reviewed and keys for the separation of males and females of the 12 known species are provided. The males of A. vittatus Annecke & Mynhardt and A. hortensis Annecke & Mynhardt are described for the first time and the following four species are described from South Africa as new: ater, curtus, mimus and notatus. New information on hosts and distribution is provided for all but two of the previously known species.
Systematics, cladistics, and biogeography of the Peirates collarti and P. lepturoides species groups (Heteroptera: Reduviidae, Peiratinae)
Authors: Morrone Juan J. , Coscaron MARIA DEL CARMEN
Published: July 2010

Cleptocoris StÅl is synonymized with Peirates Serville. Its species are assigned to the P. collarti and P. lepturoides species groups. The former group includes P. collarti Schouteden and P. monodi Villiers; the latter group is formed by P. amieti Villiers, P. areatus Miller, P. atromaculatus (StÅl), P. aurigans Distant, P. balteatus Germar, P. cinctiventris Horváth, P. diola (Villiers) comb. n., P. lepturoides (Wolff), P. macilentus Miller, P. maurus StÅl (= Cleptocoris leyei Villiers, syn. n.), P. niger Signoret, P. nitidicollis Reuter (= P. conspurcatus Distant, syn. n.), P. ochripennis Jeannel, P. perinetensis Villiers, P. strepitans Rambur (= P. rufescens Villiers, syn. n.), P. tellini Schouteden and P. turpis Walker (= P. brachypterus Horváth, syn. rev.). These species are redescribed, illustrated, their geographical distribution mapped, and keys for separating them are given. A phylogenetic analysis of the species is presented. In the cladogram, the P. collarti and P. lepturoides species groups are monophyletic and sister taxa. Within the latter, P. amieti is the sister species to the trichotomy formed by P. perinetensis, P. strepitans and the clade including the remaining species which comprises, in phylogenetic order, the pair P. niger - P. turpis; the group including P. cinctiventris, P. lepturoides, P. areatus, and P. ochripennis; and the group including P. atromaculatus-P. aurigans, P. balteatus, P. tellini, P. macilentus-P. nitidicollis, P. diola - P. maurus. The geographical distribution of the species shows an African - Indopacific generalized track, with an Indian Ocean baseline. The group is hypothesized to be of tropical Gondwanan origin, with subsequent spread into the Palearctic.
The status of the genera Asecodes Förster, Ionympha Graham and Teleopterus Silvestri (Hymenoptera: Eulophidae), with a review of Nearctic species
Authors: Hansson Christer
Published: July 2010

Ionympha Graham is removed from synonymy with Asecodes Förster. Teleopterus Silvestri, Metasecodes Erdös and Desmatocharis Graham are placed in synonymy with Asecodes. These transpositions are supported by new characters. Five species previously classified in Teleopterus are newly transferred to Asecodes: Desmatocharis reticulatus Kamijo, Entedon erxias Walker, Eulophus turcicus Nees, Teleopterus delucchii Boucek, T. politus Hansson. The Nearctic species of Asecodes are presented in a key, including one new species: A. orilatum, two newly recorded species: A. mento (Walker), A. coronis (Walker), hitherto known only from Europe, and two species previously in Teleopterus: A. erxias, A. politum. Ionympha is represented by two species in the Nearctic: I. carne (Walker), I. ochus (Walker).
Phylogenetic analysis of larval and adult characters of Adephaga (Coleoptera) using cladistic computer programs
Authors: Haas A. , Beutel R.G.
Published: July 2010

Eighty characters of larvae and adults of 35 adephagan genera, 2 polyphagan genera, and 1 cupedid genus were analyzed using the cladistic computer programs PAUP and MacClade. The analysis resulted in 1961 equally parsimonious trees of 131 steps (minimum length cladogram). The monophyly of Adephaga and of all adephagan families is confirmed. A sistergroup relationship between Gyrinidae and the remaining Adephaga, a sistergroup relationship between Spanglerogyrus and Gyrininae, the monophyly of Dytiscoidea, a sistergroup relationship between Noteridae and the remaining Dytiscoidea, a sistergroup relationship between Hygrobiidae and Dytiscidae, the monophyly of Caraboidea (Rhysodidae and Carabidae), and the monophyly of Harpalinae sensu Crowson (1955) are in agreement with earlier phylogenetic hypotheses by Beutel & Roughley (1988) and Beutel (1992a, 1993, 1995). The sistergroup relationship between Haliplidae and Dytiscoidea and the monophyly of Geadephaga (Trachypachidae + Caraboidea) are in contrast to Beutel (1992a, 1993, 1995). The high number of equally parsimonius trees is largely due to an inconsistent distribution of character states in basal carabid taxa, i.e. reversals and convergencies. Further evidence is needed for a clarification of these systematic problems.
The Trepobatinae (Heteroptera: Gerridae) of New Guinea and surrounding regions, with a review of the World fauna. Part 4. The marine tribe Stenobatini
Authors: Polhemus Dan A. , Polhemus John T.
Published: July 2010

The small waterstriders of the subfamily Trepobatinae have radiated extensively on New Guinea and in surrounding archipelagos. All of the marine forms of the subfamily are found in this region, with the exception of one monotypic genus occurring on brackish water in the eastern tropical Pacific. The present study, the fourth in a series of reports dealing with Melanesian Trepobatinae, covers these regional marine taxa. The tribe Stenobatini, proposed in Part 1 of this series to hold the genera Stenobates Esaki, type-genus, plus Rheumatometroides Hungerford and Matsuda, and Stenobatopsis gen. n. (as undescribed genus 2), is revised, two new genera are proposed, and keys to genera and species are provided, followed by synonymies, diagnoses and discussion of the constituent genera. The following new taxa are proposed within the Stenobatini: Pseudohalobates gen. n., monobasic, type-species S. inobonto sp. n. from Indonesia (Celebes, Moluccas, Talaud Archipelago, Biak and Yapen islands, and Vogelkop Peninsula of Irian Jaya), and the Philippines (southern Mindanao); Stenobatopsis gen. n., monobasic, type-species S. stygius sp. n. from Halmahera; Thetibates gen. n., type-species Rheumatometroides serena Lansbury, from northern Papua New Guinea and the Solomon Islands; Rheumatometroides kikori sp. n. from the Kikori delta of southern Papua New Guinea; R. insularis papar n. ssp. from Sabah, north Borneo; R. sele sp. n. from the Vogelkop Peninsula of Irian Jaya; R. wabon sp. n. from Biak Island; Stenobates fakfak sp. n. from the Vogelkop Peninsula of Irian Jaya; S. kamojo sp. n. from Biak Island, Salawati Island, Yapen Island, and the Vogelkop Peninsula of Irian Jaya; S. kasim sp. n. from Salawati Island; S. labuha sp. n. from Bacan and Halmahera; S. langoban sp. n. from Palawan; S. sangihe sp. n. from the Sangihe Archipelago; and S. zamboanga sp. n. from Mindanao. Other nomenclatural changes: Rheumatometroides carpentaria (Polhemus & Polhemus), comb. n.; Rheumatometroides insularis insularis (J. Polhemus & Cheng), comb. n.; Stenobates makraitos (Chen & Nieser) comb. n.; Thetibates matawa (Lansbury), comb. n. [ = Rheumatometroides aqaaqa (Lansbury), syn. n.]; Thetibates serena (Lansbury), comb. n. Habitat and distributional data are given for these taxa, accompanied by keys, figures of key characters and distribution maps.
The genus Bombylisoma Rondani (Diptera: Bombyliidae): phylogenetics and review of species from the Afrotropical Region
Authors: Greathead D.J.
Published: July 2010

The allocation of species to species groups within the genus Bombylisoma Rondani by previous authors is discussed and examined by means of phylogenetic analysis. Most characters previously used to define species groups are shown to exhibit a high degree of homoplasy but characters of the male genitalia provide synapomorphies uniting groups of species. As a result of the analysis a new genus Eremyia gen. n. is proposed for B. argentatum (Greathead) and allied species and species groups are defined: a B. minimum (Scopoli) group; a B. senegalense (Macquart) group; a B. tripunctatum (Macquart) group; and a B. argyropygum (Wiedemann) - B. nucale (Bezzi) group. A key is provided for the identification of the Afrotropical species. B. pectorale (Loew) and B. gibbicorne (Bezzi) are shown to be synonyms of B. senegalense (Macquart), and B. twiga sp. n. is described from Kenya.
The higher classification of butterflies (Lepidoptera): problems and prospects
Authors: Vane-Wright R.I. , De Jong R. , Ackery P.R.
Published: July 2010

Progress in understanding the higher classification of butterflies has not kept pace with increase in the number of described species. Important points of uncertainty or contention include, apart from ranking problems, monophyly of Papilionoidea plus Hesperioidea, their relationship with other Lepidoptera in general and the Hedyloidea in particular, the question of the sister group of the Pieridae (either Papilionidae, or Lycaenidae + Nymphalidae), and the division of families into subfamilies. Traditional groupings are discussed and compared with the results of a cladistic analysis using 103 characters and 74 species (59 butterflies and 15 moths). The cladistic analysis supports a number of currently held views about butterfly classification, such as monophyly of five major family groupings (Hesperiidae, Papilionidae, Pieridae, Lycaenidae and Nymphalidae) and suggests sister group relationships between Papilionoidea and Hesperioidea, and Pieridae and (Lycaenidae + Nymphalidae). Most traditional subfamilies, however, are not supported on the basis of the data set used but the Riodininae, which always appeared as a monophyletic, subordinate group within the Lycaenidae, are a notable exception. Further, the analysis suggests that, contrary to traditional ideas, the Parnassiinae, not Baroniinae, are sister to the remainder of the Papilionidae, Pseudopontiinae are internal to (Pierinae + Coliadinae), Dismorphiinae are sister to all other Pieridae, and that Liptena, Poritia and Miletus represent the closest relatives of the Riodininae. The data set is not well suited for an assessment of the position of the butterflies amongst other Lepidoptera. Nevertheless, of the moths used, Macrosoma (Hedylidae, Hedyloidea) and Urania (Uraniidae, Geometroidea) appear to be the closest relatives of the butterflies. With regard to the higher classification of the butterflies many problems thus remain, and several ways to tackle these are discussed. The need for some form of international co-operation between fieldworkers, comparative morphologists and molecular systematists is stressed.