bluuebunny/crossref_metadata_2025 huggingface dataset:
Ammophila nancy Menke, a new species in the pruinosa complex (Hymenoptera: Sphecidae: Ammophilinae)
Authors: MENKE ARNOLD S.
Published: February 2019

Ammophila nancy new species (Hymenoptera, Sphecidae), is described from Arizona, California, Nevada, New Mexico, Texas and Mexico (Sonora and Baja California). The new species is a member of the pruinosa complex which also contains californica Menke, 1964, and pruinosa Cresson, 1865. A key is provided for the identification of the three species.
Review of the leafhopper genus Pythamus Melichar (Hemiptera: Cicadellidae: Evacanthinae) in the Indian subcontinent
Authors: VIRAKTAMATH C. A. , WEBB M. D.
Published: February 2019

The genus Pythamus Melichar (type species: P. dealbatus Melichar) is redefined. The genus Dussana Distant (type species: D. quaerenda Distant) is treated as a junior synonym of Pythamus. Three new species, Pythamus biramosus sp. nov. (India: Karnataka), P. bispinosus sp. nov. (India: Tamil Nadu, Kerala) and P. montanus sp. nov. (India: Tamil Nadu) in addition to P. dealbatus are described and illustrated. Dussana quaerenda is treated as a junior synonym of P. dealbatus. Dussana assamensis Ramakrishnan is transferred to the genus Onukia. A key to species of Pythamus is also provided.
Biogeographic area relationships in Venezuela: A Parsimony analysis of Culicidae—Phytotelmata distribution in National Parks
Authors: NAVARRO JUAN-CARLOS , LIRIA JONATHAN , PIÑANGO HERMES , BARRERA ROBERTO
Published: February 2019

This study represents the first attempt to understand the current distribution and evolutionary history of Culicidae in Venezuela and adjacent areas using cladistic methods. We studied the association between immature mosquitoes and their aquatic habitats in plants (phytotelmata) in 16 protected natural areas of Venezuela. A total of 68 mosquito species was collected from 47 host-plant species. Parsimony Analysis of Endemicity using localities and mosquito species was used to find the most parsimonious cladograms depicting the biogeographic relationships. Varying degrees of mosquito-plant associations were observed, with the greatest specificity existing at the genus and subgenus levels. Implicit enumeration algorithms obtained one parsimonious tree (L=101 steps) with four well-supported groups named "Amazonas" (southern Venezuela), “Bolivar” (southeastern Venezuela), "Los Andes Cordillera” and "Coastal Cordillera" (northern Caribbean mountain range). These results support the hypothesis that the Guayana Shield was an ancestral center of speciation, followed by the Andean Region, whereas the mosquito fauna from the Central Cordillera was derived from the latter.
Three new species of Euglossa Latreille (Hymenoptera: Apidae) from Brazil
Authors: NEMÉSIO ANDRÉ
Published: February 2019

Euglossa (Euglossella) jacquelynae sp. n., from Central Brazil, Euglossa solangeae sp. n., from coastal southeastern Brazil, and Euglossa (Euglossa) sovietica sp. n., from the western Brazilian Amazon, are described as new species. Euglossa solangeae sp. n. is believed to belong to the Euglossa stellfeldi Moure species group—a discussion on the placement of this group is also presented – and Euglossa sovietica sp. n. is treated as a member of Euglossa purpurea Friese species group.
Rediagnosis of Ichthyotrema Caballero & Bravo-Hollis, 1952 (Digenea: Gyliauchenidae Fukui, 1929), including the redescription of I. vogelsangi Caballero & Bravo-Hollis, 1952
Authors: HALL KATHRYN A. , CRIBB THOMAS H.
Published: February 2019

Recent studies on the Gyliauchenidae Fukui, 1929 have lead to our reinterpretation of the morphology of the group. Here we emend the generic diagnosis of Ichthyotrema Caballero & Bravo-Hollis, 1952 to reflect current understanding of the morphology of the digestive system and male terminal genitalia. We redescribe the type-, and only, species, Ichthyotrema vogelsangi Caballero & Bravo-Hollis, 1952, based on our examinations of paratype material from the intestine of Prionurus punctatus Gill, 1862, caught in Pacific waters off Mexico. Specimens of I. vogelsangi are distinctive because of their massive bodies, and Ichthyotrema is distinguished among the Gyliauchenidae by the position of the cirrus-sac entirely posterior to the caeca, the possession of a post-testicular ovary and the coiling of the uterus between the diagonal testes. The ventral sucker of Ichthyotrema is close to the posterior extremity of the body, supporting placement of the genus within the Gyliaucheninae Fukui, 1929. Here we recognise Ichthyotrema as a monotypic genus within the Gyliauchenidae, which is restricted to acanthurid fishes in the Eastern Pacific Ocean.
The genus Scirtothrips Shull (Insecta, Thysanoptera, Thripidae) and three related genera in Japan
Authors: MASUMOTO MASAMI , OKAJIMA SHÛJI
Published: February 2019

The genus Scirtothrips and related genera in Japan were studied, and the following four genera and eight species recorded: Anascirtothrips okinawanus n. sp., Drepanothrips reuteri Uzel, Scirtothrips albidus n. sp., S. dorsalis Hood, S. flavus n. sp., S. katsura n. sp. and S. machili n. sp., Parascirtothrips fagaceae n. gen. et sp. The male of D. reuteri is recorded from Japan for the first time. Ten genera are recognized in the Scirtothrips genus group, and relationships between these are discussed.
Reticulitermes malletei (Isoptera: Rhinotermitidae): a valid Nearctic subterranean termite from Eastern North America
Authors: AUSTIN J. W. , BAGNÈRES A. -G. , SZALANSKI A. L. , SCHEFFRAHN R. H. , HEINTSCHEL B. P. , MESSENGER M. T. , CLÉMENT J. -L. , GOLD R. E.
Published: February 2019

The taxonomic status of Reticulitermes Holmgren in North America has been in need of revision for years, but because of morphological ambiguity, traditional morphological identification of Reticulitermes species has always been difficult and unreliable. Analysis of termites, applying non-morphological genetic comparisons of mitochondrial DNA from numerous populations across North America, has implicated new species which are presently under investigation. Applying the 16S rRNA gene with biometric, cuticular hydrocarbons, and ethological data, a cryptic species of Nearctic Reticulitermes has been identified from Delaware, Georgia, Maryland, North Carolina, and South Carolina and determined to constitute a valid species with an apparently Atlantic distribution. Epicuticular hydrocarbon analysis showing the presence of rare triunsaturated alkenes, and a total absence of methyl branched alkanes also support this cryptic group as a distinct species. The name for this species is Reticulitermes malletei, previously described, but not generally accepted by termite experts in the United States. Comparisons from a 403 bp region of the mtDNA 16S rRNA gene was used to discriminate known Reticulitermes species from North America: The eastern subterranean termite R. flavipes (Kollar), dark southern subterranean termite R. virginicus (Banks), light southern subterranean termite R. hageni Banks, arid subterranean termite R. tibialis Banks, and western subterranean termite R. hesperus Banks. When compared to this new species, both maximum parsimony and maximum likelihood support their genetic isolation from sympatric populations of known species and eliminate either exotic Palearctic introductions or western Nearctic involvement.
Morphology and surface structure of eggs and first instar larvae of Croce schmidti (Navás, 1927) (Neuroptera: Nemopteridae)
Authors: SATAR ALİ , SULUDERE ZEKİYE , CANDAN SELAMİ , CANBULAT SAVAŞ
Published: February 2019

The morphology of eggs and first instar larvae of Croce schmidti (Navás, 1927) is described and figured for the first time using light and scanning electron microscopy. The females were collected from Derik, Mardin Province, Turkey and maintained in the laboratory. The eggs are elongate and cylindrical with rounded ends and bear a small terminal micropylar knob. Except for the adhesive materials, the chorion surface is smooth. The newly emerged larva of C. schmidti is characterized by a slightly quadrangular head and short prothorax and has prominent pale dark brown color markings on the metatergites. The body is covered with short brown setae, which include macrotrichia and dolichasters.
A new, toxic species of Colostethus (Anura: Dendrobatidae: Colostethinae) from the Cordillera Central of Colombia
Authors: GRANT TARAN
Published: February 2019

A new species of Colostethus is described from cloud forest in the Cordillera Central of Colombia at approximately 2100–2500 m above sea level. It is a riparian species not known to occur more than a few meters from the water’s edge. Specimens were collected in grassy vegetation and on and beneath rocks along streams and pools, in both primary forest and in or at the edge of forest clearings. Most specimens were collected during the day, but several were observed to be active at night. The new species differs from other species of Colostethus in lacking pale oblique lateral and ventrolateral stripes, possessing a pale dorsolateral stripe, lacking or having at most rudimentary toe webbing, and possessing a marbled or reticulated ventral color pattern. Adult males of the new species possess a swollen third finger and a gray or black throat. In life, the new species possesses bright lemonor golden-yellow flash marks on the axilla, groin, belly, and concealed surfaces of the thigh and shank, and bluish-white ventral coloration, and it exudes a milky secretion from the skin. The aqueous extract of the skin is toxic, but additional samples are required to determine the nature of the toxic compound(s).
Rediscovery of Bathypsammis Huys & Gee, 1993 (Copepoda, Harpacticoida, Pseudotachidiidae) with description of a new species from the Antarctic deep sea
Authors: WILLEN ELKE , SCHULZ MAXIMILIAN
Published: February 2019

The species diversity of Copepoda Harpacticoida in the deep sea of the Angola Basin (DIVA I expedition with RV Meteor) and in the Weddell Sea (ANDEEP II expedition with RV Polarstern) have been investigated. From the obtained multicorer samples two female specimens of Bathypsammis Huys & Gee, 1993 were collected. The only specimen known before was described as Psammis longifurca by Bodin in 1968 from the northern Atlantic. In the present paper a new species Bathypsammis polaris sp. n. from the Antarctic Weddell Sea is described. The monophyly of Bathypsammis within the Paranannopinae Por, 1986 can be confirmed by several new autapomorphies whereas the relation to other paranannopid taxa must remain open for the moment. The new findings implicate a widespread distribution of Bathypsammis from the northern to the southern Atlantic and even to the southern polar regions. Up to now all individuals that have been found occur exclusively at deep-sea sites and in very low abundances.
Establishment of the Vultocinidae, a new family for an unusual new genus and new species of Indo-West Pacific crab (Crustacea: Decapoda: Brachyura: Goneplacoidea), with comments on the taxonomy of the Goneplacidae
Authors: NG PETER K.L. , MANUEL-SANTOS MARIVENE R.
Published: February 2019

A new family, Vultocinidae, is established for an unusual new genus and new species of crab (Crustacea: Brachyura: Goneplacoidea) here described from relatively deep waters in the central Philippines and Vanuatu. Although Vultocinus anfractus, new genus, new species, superficially resembles pilumnid genera like Lophoplax and pseudoziid genera like Planopilumnus in its deeply sculptured carapace and leg surfaces, its male abdomen and gonopods appear to affiiliate it with the Goneplacidae sensu lato instead. However, its suite of many unusual characters requires the recognition of a new family for Vultocinus. Comparisons of the Vultocinidae with the Goneplacidae sensu stricto suggests that the subfamily Mathildellinae, presently regarded as a subfamily, should be recognised as a full family; and two other problematic constituent genera, Progeryon and Conleyus, are also referred to their respective families in the Goneplacoidea. The genus Paragalene is also referred to the Progeryonidae. The extent of the sterno-abdominal cavity along the thoracic sternites, structure of the posterior thoracic sternites, position of the press button of the male abdominal locking mechanism and form of the first four male abdominal segment as characters in the taxonomy of the Goneplacoidea is also discussed.
Some New Zealand Polyclads (Platyhelminthes, Polycladida)
Authors: HOLLEMAN JOHN J.
Published: February 2019

Eight species of poyclads from New Zealand are presented. Five species were already known to science, and are redescribed. Four of these species have been previously reported from New Zealand, the fifth, Echinoplana celerrima, is new to New Zealand. Two are new species of the genus Chromoplana, Chromoplana sirena n. sp. and Chromoplana kaikouris n. sp. One species belongs to a new genus, Aotearoa ballantinensis n. gen. n. sp. All new species and the new genus are described and discussed. The taxonomic status of Postenterogonia orbicularis, Notoplana suteri with Notoplana australis, and Leptostylochus elongatus is reviewed.
Redescription of Tagulis mystacinus (Araneae: Thomisidae) from Sri Lanka
Authors: BENJAMIN SURESH P. , JALEE ZIYARD
Published: February 2019

Tagulis mystacinus Simon, 1895 from Sri Lanka is redescribed based on a pair of newly collected male and female specimens. T. mystacinus may be separated from other thomisids by the presence of very peculiar leaf-shaped macrosetae immediately posterior to anterior and posterior lateral eyes. They also may be separated by details of their genitalia.
Eupholidoptera karatolosi sp.n. and E. mirzayani sp.n. (Orthoptera, Tettigoniidae), two new bushcrickets from Greece and Iran
Authors: MOFIDI-NEYESTANAK MOHSEN , QUICKE DONALD L.J.
Published: February 2019

Eupholidoptera karatolosi Mofidi-Neyestanak & Quicke from Greece and E. mirzayani Mofidi-Neyestanak & Quicke from Iran, two new species of bushcrickets (Orthoptera, Tettigoniidae: Platycleidini), are described and distinguished from closely related species based on morphology. Species relationships within Eupholidoptera Maran are discussed and some species are given new assignments. A simplified illustrated identification key to the species of Eupholidoptera is provided to accommodate the new species. They are being described since they have been used to generate DNA sequence data that will be published elsewhere as part of a phylogenetic study of the tribe Platycleidini.
Identifying species of Bythinella (Caenogastropoda: Rissooidea): A plea for an integrative approach
Authors: HAASE MARTIN , WILKE THOMAS , MILDNER PAUL
Published: February 2019

The genus Bythinella comprises many species throughout Europe, but species delimitation, traditionally based on shell morphology and genital anatomy, is often a matter of debate. Out of an ongoing large-scale project on the phylogeny of the genus, we analyzed the relationships of species occurring in the south Austrian province Carinthia and in neighboring Slovenia as a model for similar cases of systematic and taxonomic ambiguity. Our analyses based on sequence data of a fragment of COI comprising 638 bp, morphological and anatomical investigations confirmed the presence of three species, B. opaca (Gallenstein, 1848), B. robiciana (Clessin, 1890) and B. angelitae nom. nov. for B. opaca (Frauenfeld, 1857). The latter, while genetically distinct, is morphologically and anatomically cryptic in that it can only be distinguished from B. opaca by the denticulation of the radular marginal teeth. B. robiciana, on the other hand, is morphologically well defined, but genetically not separable from B. opaca, its stem species. Thus, taxonomy in Bythinella has to be based on the integration of morphology, anatomy and genetics. Our phylogenetic analyses suggest that B. opaca has colonized Carinthia, which has largely been covered by glaciers during the last ice age, along two routes, one from the south and a second one from the southeast.
Annotated checklist of the rotifers (Phylum Rotifera), with notes on nomenclature, taxonomy and distribution
Authors: SEGERS HENDRIK
Published: February 2019

Phylum Rotifera comprises about 2030 known species classified in three main groups, the marine Seisonida (3 species), the Monogononta (1570 species) and the unique, exclusively parthenogenetic Bdelloidea with 461 clonal species. Here I present an annotated checklist of the taxon, giving the global distribution of the valid species. This checklist contains over 3,000 available names at the genusand species-group level, including valid names and a non-comprehensive list of synonyms. A number of taxonomic and nomenclatural suggestions following from the review is presented. Suggestions include several new synonyms and two nomina nova: Dissotrocha kostei for Dissotrocha aculeata Koste, 1996 non Ehrenberg, 1838 (stat. nov., ex. Dissotrocha hertzogi aculeata) and Lepadella zigzag for Lepadella bractea Myers, 1934 non (Ehrenberg, 1838). This contribution also describes features of an on-line, global names database of the Rotifera, on which this checklist is based.
New species of the amphiliid catfish genera Amphilius, Doumea and Phractura and the taxonomy of Paramphilius from West Central Africa (Siluriformes, Amphiliidae)
Authors: SKELTON PAUL H.
Published: February 2019

A recent review of the amphiliid fishes of West Central Africa (WCA) indicates the presence of several described and undescribed species in different genera. Known amphiliines include three species of Amphilius Günther, one species of Paramphilius Pellegrin and three doumeine species in two genera, Doumea Sauvage and Phractura Boulenger. The known Amphilius from the WCA include A. longirostris (Boulenger, 1901), A. pulcher Pellegrin, 1929, and A. nigricaudatus Pellegrin, 1909. Seven species new to science include Amphilius dimonikensis, a small species with a banded colour pattern, described from 6 specimens from the Kouilou River system in Congo; Amphilius caudosignatus, a very small species with a unique caudal fin coloration, described from 3 specimens from the Ogowe River system in Gabon; Amphilius korupi, a dark brown species with a short head, eyes advanced and long oral barbels, described from 43 specimens taken from the Cross, Ndian and Akpa-Yafe River systems in Cameroon; Amphilius mamonekenensis, distinguished by its light spotted appearance, described from 18 specimens taken from the Kouilou River system in Congo; Doumea gracila, a large robust species but with a relatively slender caudal peduncle, described from 44 specimens from the Lobé, and Lokoundjé rivers, Cameroon; Doumea sanaga, with the dorsal fin relatively far back and a long slender caudal peduncle, described from 32 specimens taken from the Sanaga River system, Cameroon; and Phractura stiassny, described from a single specimen with a unique pair of neural spines before the dorsal fin from the Nyanga River in Gabon. In addition Amphilius baudoni Pellegrin is assigned to Paramphilius, with Paramphilius goodi Harry, 1953, considered as a junior synonym. Distribution maps for the new species are given.
Achillesaurus manazzonei, a new alvarezsaurid theropod (Dinosauria) from the Late Cretaceous Bajo de la Carpa Formation, Río Negro Province, Argentina
Authors: MARTINELLI AGUSTÍN G. , VERA EZEQUIEL I.
Published: February 2019

A new genus and species, Achillesaurus manazzonei gen. et sp. nov., of the enigmatic clade Alvarezsauridae (Theropoda, Coelurosauria), recovered from the Santonian Bajo de la Carpa Formation (Río Negro Province, Argentina), is here described. A. manazzonei is a relatively large alvarezsaurid different from Alvarezsaurus calvoi (from the same Age and Formation) in having a lateral fossa in the proximal caudal centra, a less developed supraacetabular crest, the brevis shelf not reaching the base of the ischial pedicel, and the lateral malleolus of the tibia at the same level of the medial one. Achillesaurus differs from Patagonykus puertai, from the Portezuelo Formation (Neuquén Province, Argentina), by the presence of an almost undeveloped supracetabular crest of the ilium and the unfused condition of the astragalus and the calcaneum. The new species is excluded from the Asian Mononykinae due to the unreduced fibula distally and a non-arctometatarsalian pes. The autapomorphies of Achillesaurus are the presence of a biconcave caudal vertebra (possibly the fourth) with the cranial surface 30% larger in diameter than the caudal one. The inclusion of Achillesaurus in a phylogenetic framework resulted in an unresolved polytomy among the new taxon, Alvarezsaurus, and Patagonykus plus Mononykinae, the latter clade being weakly supported. The result here presented shows a basal stem radiation of South American alvarezsaurids. New material of the Patagonian alvarezsaurids is necessary to evaluate relevant traits to test further the phylogenetic relationships of the basal alvarezsaurids.
Three new ground living pholcid species (Araneae: Pholcidae) from Parque Estadual da Cantareira, São Paulo, São Paulo, Brazil
Authors: MACHADO ÉWERTON O. , YAMAMOTO FLÁVIO U. , BRESCOVIT ANTONIO D. , HUBER BERNHARD A.
Published: February 2019

Three pholcid species (Araneae: Pholcidae) from Parque Estadual da Cantareira, city of São Paulo, Brazil are newly described: Mesabolivar cantharus n. sp., Mesabolivar camussi n. sp., and Tupigea cantareira n. sp. Collection data suggest a preference of the species for ground level habitats. Preliminary data on Tupigea cantareira n. sp. suggest a continuous occurrence of adult individuals throughout the year, with a peak in the rainy season. It also seems that T. cantareira n. sp. prefers the natural (“Mata”) environment over the reforested Pinus area.
Trichodinids (Ciliophora: Peritrichida) parasitic on gills of freshwater fishes, Carassius auratus and Aristichthys nobilis from China, with the description of Trichodina subtilihamata sp. nov.
Authors: TANG FAHUI , ZHAO YUANJUN , TAO YANFEI
Published: February 2019

Five trichodinid ectoparasites (Ciliophora: Peritrichida) were isolated from two freshwater fishes, Carassius auratus and Aristichthys nobilis collected from Chongqing, China. Trichodina subtilihamata sp. nov. is described from the gills of Carassius auratus and characterized by a slender hooked blade. The other four species of trichodinids are: Trichodina mutabilis Kazubski & Migala, 1968, Trichodina uniforma Van As & Basson, 1989, Trichodina nigra Lom, 1960 and Trichodina kazubski Van As & Basson, 1989. Comparative descriptions, with photomicrographs and morphometric data, are presented of these trichodinids to establish locality records. Descriptions presented here were obtained by examinations of specimens prepared using the dry silver nitrate impregation and the methyl green-pyronin staining. Comparisons with closely related species are provided for the new species.
First record of a living ditaxiporine catenicellid in the Atlantic, with a description of Vasignyella ovicellata n. sp. (Bryozoa)
Authors: VIEIRA LEANDRO M. , GORDON DENNIS P. , CORREIA MONICA D.
Published: February 2019

The genus Vasignyella Gordon, 1989 is recorded for the first time from the Atlantic Ocean, in the littoral zone of Alagoas, northeastern Brazil. It is only the second species attributed to the genus, the type species of which has never been seen in a reproductive state, and which has a colony form of uniserial chains of unizooidal internodes. The new species from Brazil, herein described as V. ovicellata, has a similar colony form in the infertile state, but has ovicells in multizooidal internodes. This character allows a reconsideration of the phylogenetic relationships of Vasignyella, which was previously the sole included genus in the catenicellid bryozoan subfamily Vasignyellinae. The nearest generic relative of Vasignyella was, however, considered to be the extinct genus Ditaxiporina Stach, 1935 (subfamily Ditaxiporinae), with near-identical autozooidal morphology, but with most internodes multizooidal and also having ovicells when reproductive. The discovery of ovicells in Vasignyella allows the subfamily Vasignyellinae to be discarded and the genus transferred to the Ditaxiporinae, a subfamily with an Atlanto-American record previously known to range from the Late Paleocene to the Early Oligocene.
Contribution to the knowledge of the Neotropical genus Plagiolabra von Schulthess (Hymenoptera, Vespidae, Eumeninae)
Authors: HERMES MARCEL G.
Published: February 2019

Plagiolabra von Schulthess is a Neotropical eumenine wasp genus that has not received much attention in the last century. Brief redescriptions of the two included species are provided, along with an identification key, illustrations and a distributional map. Male genitalia are illustrated and described for the first time. A discussion of the monophyly of the genus and phylogenetically relevant characters is presented.
Three new species of Ciulfina Giglio-Tos (Mantodea: Liturgusidae) from north-eastern Australia
Authors: HOLWELL GREGORY I. , GINN SCOTT G. , HERBERSTEIN MARIE E.
Published: February 2019

The genus Ciulfina Giglio-Tos includes a number of small tree-trunk dwelling species of praying mantids that are found through eastern Queensland and northern Australia. Three new species of Ciulfina: C. baldersoni, C. klassi and C. rentzi and one existing species C. biseriata (Westwood) are formally described on the basis of male genital morphology. A key to the identification of Ciulfina based on genital morphology is also provided.
A review of the genus Leptoteratura Yamasaki, 1982 (Orthoptera: Tettigoniidae: Meconematinae) from China
Authors: MAO SHAO-LI , SHI FU-MING
Published: February 2019

Keys to the subgenera of Leptoteratura, to the species of subgenus Leptoteratura from China and to all species of subgenus Rhinoteratura are presented. One new species Leptoteratura (Rhinoteratura) lamellatus Mao et Shi, sp. nov. and generic diagnosis are presented. Type specimens are deposited in Museum of Hebei University, Baoding.
A redescription of Leptodactylus jolyi Sazima and Bokermann (Anura, Leptodactylidae) and the recognition of a new closely related species
Authors: GIARETTA ARIOVALDO A. , COSTA HUGO C.M.
Published: February 2019

Based on freshly collected and recorded topotypes, we re-describe Leptodactylus jolyi and describe a closely related new species from the municipality of Uberlândia (MG, Brazil). As some other species of the L. fuscus group, the new species present longitudinal skin folds on dorsal surface of shank. Leptodactylus marambaiae is a shorte-legged species, in L. jolyi the shank reaches 61–65% of the SVL, and 62–68 % in the new species. Males L. jolyi are larger (45 mm SVL) than those of L. gracilis (38.5 mm), than those of L. plaumanni (31–35 mm), and are smaller and more slender than those of the new species (48.3–53.9). In L. jolyi and in the new species the vomerine teeth are in two straight transverse series, in L. gracilis each is a distinct arch. Leptodactylus plaumanni has a fast call rate (13–23/s), in L. gracilis it is 2.5–4/s, in L. jolyi it is just about 0.1–0.3/s and in the new species calls are emitted irregularly (0.02–0.3/s). Leptodactylus jolyi call is composed by 1–3 pulses, while the new species has a single or double-pulsed call; the call also is 25-100% longer in L. jolyi (mode 0.04 s, our sample) than in the new species (mode 0.02 s). We predict that populations in the Cerrado now identified as L. jolyi will turn out to be conspecific with the new species described herein. In the original description the call of L. jolyi is two times longer than that we described here based on topotypes. A possible mistake in time scale in the original description may account for this difference.
Fishes (Elasmobranchii and Actinopterygii) of Picãozinho reef, Northeastern Brazil, with notes on their conservation status
Authors: DE SOUZA ALLAN T. , ILARRI MARTINA DI IULIO , DE MEDEIROS PAULO ROBERTO , GREMPEL RENATO GROTTA , ROSA RICARDO S. , SAMPAIO CLÁUDIO L.S.
Published: February 2019

Shallow reefs on the northeastern coast of Brazil are rich, productive and diverse environments, with great ecological and economical importance. Picãozinho is located 1,500 m off the coast of João Pessoa city, Paraíba state, NE Brazil. The aim of this work is to provide an updated list of the reef fishes of Picãozinho, based on a survey of approximately 350 hours of direct observation using free dive techniques, and complemented by collection data and other unpublished records. A total of 102 species of 43 families were recorded. Of these, 36 are new occurrences for the reef and seven are species that have been recently resurrected from synonymy, mainly with Caribbean species previously cited for Brazil.
Review of Cermatobius Haase, 1885 (Chilopoda: Henicopidae) of China and Neotype Designation for Cermatobius longicornis (Takakuwa, 1939)
Authors: MA HUI-QIN , SONG DA-XIANG , ZHU MING-SHENG
Published: February 2019

A review of Chinese occurences of the genus Cermatobius Haase, 1885 (Lithobiomorpha: Henicopidae) is based on both male and female specimens. Diagnoses, descriptions, distributions, and figures are presented for the two Chinese species, C. longicornis (Takakuwa, 1939) and C. longitarsis (Verhoeff, 1934a); at the same time, we designated a neotype for C. longicornis.
A new Satsuma species (Pulmonata: Camaenidae) endemic to Taiwan
Authors: WU SHU-PING , LIN YAO-SUNG , HWANG CHUNG-CHI
Published: February 2019

A new species of camaenid land snail, Satsuma longkiauwensis sp. nov. from southern Taiwan is established. This large terrestrial and herbivorous snail inhabits the lowland forests with a narrow geographical distribution. The species is characterized by having a large shell, roundly angulated peripheries adjacent to the peristome, an open umbilicus, a robust flagellum, a weak expansion on male genitalia instead of a penial caecum externally and a hemispherical verge instead of an elongated pilaster internally. A key is provided for the first time to identify camaenids from Taiwan.
A reassessment of apheloriine millipede phylogeny: additional taxa, Bayesian inference, and direct optimization (Polydesmida: Xystodesmidae)
Authors: MAREK PAUL E. , BOND JASON E.
Published: February 2019

Millipedes in the tribe Apheloriini occur throughout the eastern United States, predominately in the deciduous forests of the Appalachian Mountains. Herein we present a reassessment of apheloriine millipede phylogeny using mitochondrial DNA sequences and an additional 29 exemplar taxa (including 15 undescribed species and all of the species in the genus Brachoria, except one). In this study, first we check the results of the previous phylogeny of the tribe (Marek and Bond, 2006) with different alignment and phylogenetic techniques (direct optimization and maximum likelihood), and second reconstruct a new phylogeny evaluating it in the same way with Bayesian, maximum likelihood, and direct optimization. Using this updated and expanded phylogeny, we tested historical classifications with Bayes factor and Shimodaira-Hasegawa hypothesis testing, consistently finding very strong evidence against their implied phylogenetic hypotheses. Lastly, using the new phylogeny as a foundation, we make taxonomic modifications and provide an updated species list of Apheloriini (106 species/17 genera).
A new species of Hemigrammus Gill, 1858 (Characiformes: Characidae) from Río Caura, Venezuela
Authors: BENINE RICARDO C. , LOPES GUILHERME A.M.
Published: February 2019

A new characid species, Hemigrammus taphorni, is described from the Río Caura drainage, Venezuela. The new species is distinguished from its congeners by characters related to its color pattern, number of scale rows below the lateral line, teeth morphology, and dorsal-fin length. A brief discussion about the putative relationships of H. taphorni is provided.
A new species of deep-water spider crab of the genus Rochinia A. Milne-Edwards, 1875, from Guam (Crustacea: Brachyura: Majidae)
Authors: NG PETER K.L. , DE FORGES BERTRAND RICHER
Published: February 2019

A new species of deep-water spider crab (Majidae) of the genus Rochinia A. Milne-Edwards, 1875, is described from Guam. Rochinia paulayi, new species, is distinguished from the allied R. pulchra (Miers, 1886), R. riversandersoni (Alcock, 1895) and R. griffini Davie & Short, 1989, in the form and position of the carapace spines.
A new species and first stage associations in Crinitella (Ephemeroptera: Ephemerellidae: Ephemerellinae)
Authors: JACOBUS LUKE M. , McCAFFERTY W. P. , SITES ROBERT W.
Published: February 2019

The alates of Crinitella coheri (Allen and Edmunds) are described for the first time, and morphological variability of the species is discussed. Male adults have penes with dorsal projections at the base of the gonopores and forceps with a relatively straight lateral profile. Crinitella coheri female alates and associated eggs have been misidentified as those of a Hyrtanella species. Crinitella coheri is reported from Vietnam for the first time. We describe C. lacuna, new species, based on male and female subimagoes and larvae, and the species is included in Crinitella on the basis of genitalia morphology. Alates of C. lacuna are distinguished from C. coheri by the presence of a prominent dorsal longitudinal stripe, and larvae are distinguished by having paired median spines on abdominal terga, mouthparts that are not greatly reduced, and gills 3 that are relatively larger than those of C. coheri. Crinitella lacuna is reported from India, Thailand and Vietnam. The structures of Crinitella and Hyrtanella gill cavities are different and do not represent a synapomorphy for the two genera.
Taxonomic status and ecology of Oriental Pheretima darnleiensis (Fletcher, 1886) and other earthworms (Oligochaeta : Megascolecidae) from Mt Kinabalu, Borneo
Authors: BLAKEMORE ROBERT J. , CSUZDI CSABA , ITO MASAMICHI T. , KANEKO NOBUHIRO , KAWAGUCHI TATSUYA , SCHILTHUIZEN MENNO
Published: February 2019

The synonymy of Perichaeta darnleiensis Fletcher, 1886, first described from Australian Darnley Island in the Torres Straits, was confused since Beddard (1900) and Michaelsen (1900) wrongly applied the names Perichaeta cingulata Schmarda, 1861 (= Megascolex cingulatus) and Megascolex indicus Horst, 1883 [= Amynthas corticis (Kinberg, 1828)], respectively. Taxonomic revision of this species now merges several parthenogenetic morphs, including Pheretima decipiens Beddard, 1912 syn. nov. from Luzon, Philippines. Final resolution depends on location of missing types or, alternatively, on neotypification, nevertheless new material from Mt. Kinabalu, north Sabah (Borneo) allows augmentation of the description of Pheretima darnleiensis from that given by Sims & Easton (1972) who, for its lobate/serrate intestinal caeca, remarked that these “cannot be regarded as taxonomic characters as they are more fully formed in the larger specimens”. Body size range is now 45–700 mm, although this suggests either high plasticity or too wide synonymy. Its distribution in the tropical Indo-Australasian Archipelago and islands of Malaysia, Indonesia and Fiji is attributed to human-mediated introductions in recent and pre-historic times that mask its true provenance; reports from Hawaii, Sri Lanka, South Africa and South America are disputed. Sympatric Amynthas omeimontis kinabalu Sims & Easton, 1972 was elevated to species level by Blakemore (2005). Descriptions of Pheretima (Parapheretima) saba Sims & Easton, 1972 and Polypheretima everetti (Beddard & Fedarb, 1895: 69) – somewhat similar to Polypheretima kinabaluensis (Beddard & Fedarb, 1895: 71) – are augmented and specimens are figured on newly collected material. A new Mt Kinabalu earthworm, Metaphire paka Blakemore sp. nov. is proposed that compares with Metaphire cai (Michaelsen, 1916) from Java [originally “Pheretima inflata (Horst) var. cai”], bringing the total known earthworms from Mt Kinabalu to six species. Ecological associations are briefly discussed.
First records of water mites (Acari: Hydrachnidia) from Bhutan, with description of two new species
Authors: PESIC VLADIMIR , SMIT HARRY
Published: February 2019

New records of water mites (Acari: Hydrachnidia) from streams in Bhutan are presented. Two species new to science are described: Sperchon himalayus sp. nov. and Torrenticola bhutana sp. nov. Furthermore, the first records from Bhutan are given for Monatractides tristis (Lundblad), Testudacarus tripeltatus Walter and Sperchon cf. indicus Kumar et al.
Nomenclatural Acts: homonymy in the Ascidiacea (Tunicata) and proposed nomina nova
Authors: BRUNETTI RICCARDO
Published: February 2019

In a revision of 4356 species and sub species of the class Ascidiacea, 15 were found to be homonyms for which nomina nova are proposed in this paper. The paper refers only to the nomenclature of these entities; and their taxonomy is not discussed here.
Two new species of Aclis (Mollusca: Gastropoda: Aclididae) from the continental slope of northeast Brazil
Authors: DE BARROS JOSÉ CARLOS N. , DE LIMA SILVIO FELIPE B. , DE A. FRANCISCO JONATA
Published: February 2019

Two new species of Aclis Lovén, 1846 are described from the continental slope of northeast Brazil. Aclis macrostoma n. sp. is recorded off the state of Rio Grande do Norte at depths ranging from 375 to 510 meters. Aclis watsoni n. sp. was dredged off the state of Sergipe at depths ranging from 365 to 510 meters. There are no similar species along the Brazilian coast. Aclis watsoni is similar to Aclis angulata Smith, 1890, Aclis trilineata Watson, 1897, Aclis supranitida (Wood, 1842) and Aclis ascaris (Turton, 1819) due to the presence of spiral keels and/or indication of a subsutural ramp. Aclis macrostoma is similar to Discaclis canariensis Moolenbeek & Warén, 1987, Aclis attenuans Jeffreys, 1883 and Aclis gulsonae (Clark, 1850) regarding its circular aperture and dilatation of the peristome, with a non-thickened outer lip. The latter two species are much more slender.
A new species of Huntemannia (Copepoda: Harpacticoida: Huntemanniidae) from the Yellow Sea, Korea
Authors: SONG SUNG JOON , RHO HYUN SOO , KIM WON
Published: February 2019

Huntemannia doheoni sp. nov. is described from a muddy sand flat off the coast of the Yellow Sea, Korea. The new species can be distinguished from its congeners by a combination of characters that include: a strong, hook-like outer process on the P1 coxa; incompletely fused exopod segments and reduced setal formulae of the P2–P4 in the female; and modified outermost spine on the distal segment of P3–P4 in the male. In addition, a key to the species of Huntemannia is provided.
Larval morphology in four species of Madagascan frogs ofthe subgenus Brygoomantis (Mantellidae: Mantidactylus)
Authors: KNOLL ANGELIKA , KÖHLER JÖRN , GLAW FRANK , TESCHKE MEIKE , VENCES MIGUEL
Published: February 2019

We describe the tadpole morphology of four species of frogs classified in the endemic Madagascan subgenus Brygoomantis of the genus Mantidactylus, based on larval specimens identified by DNA barcoding: Mantidactylus betsileanus and M. biporus, and two so far undescribed species that are here named M. sp. aff. betsileanus "very slow calls" and "Vohidrazana" referring to their bioacoustic features or collecting locality. The tadpoles of these four species are brown to yellowish benthic forms with a depressed body shape, dorsally directed eyes and relatively low fins. The oral discs are generalised and all species exhibit a wide gap in dorsal papillae. There is variation among different developmental stages with regard to the keratodont row formulae. In Mantidactylus betsileanus, younger stages (25 and 26) exhibit the formula 1:3+3/1+1:2, whereas it is 1:4+4/1+1:2 in older stages (36 and 38). In M. sp. aff. betsileanus "very slow calls", keratodont row formulae vary within one developmental stage (25–26) from 1:3+3/3 to 1:4+4/3, whereas it seems to be constantly 1:4+4/3 in all examined developmental stages of M. sp. aff. betsileanus "Vohidrazana". In general, the larvae of the different Brygoomantis species studied are morphologically similar to each other. Differences between species mainly concern colouration, shape of body and the number of keratodonts per millimetre, which is approximately 38 in M. betsileanus, 50 in M. biporus, and 60 in the two undescribed species.
A new species of Phrynopus from Departamento Cusco, southern Peru(Anura: Brachycephalidae)
Authors: CHAPARRO JUAN C. , DE LA RIVA IGNACIO , PADIAL JOSÉ M. , OCHOA JOSÉ A. , LEHR EDGAR
Published: February 2019

We describe a new species of Phrynopus (Anura: Brachycephalidae) from two close localities at the upper limits of cloud forest in the southern Peruvian Departamento Cusco, between 3555–3950 m a.s.l. The new species is characterized by having medium size (maximum snout-vent length 23.4 mm), dentigerous processes of vomers absent, tympanic membrane inconspicuous, dorsal skin coarsely shagreen in life, dorsolateral folds, ventral skin areolate, dorsum tan, venter bold black with conspicuous bluish-gray spots, and a bluish-white iris.
Hydromastodon sallesi, new genus and new species of Atalophlebiinae (Insecta: Ephemeroptera: Leptophlebiidae) from West and North of Brazil, and notes on systematics of Hermanella group
Authors: POLEGATTO C. M. , BATISTA J. D.
Published: February 2019

The genus Hydromastodon is described here with a new species H. sallesi, and a new combination: Hydromastodon (=Hydrosmilodon) mikei (Thomas & Boutonnet 2004). H. sallesi was described based on nymphs from Pindaíba Stream, near Nova Xavantina, State of Mato Grosso, and from Bem Querer Stream, State of Roraima, Brazil. Hydromastodon can be distinguished from other genera of the Hermanella group by presence of posterolateral spines on abdominal segments 6 or 7 to 9, tarsal claws with subequal denticles in two separate rows, without large subapical denticle, and clavate setae on legs. Hydromastodon seems to be closely related to Hydrosmilodon and Leentvaaria.
A new species of Gonatocerus (Hymenoptera: Mymaridae) from Argentina, an egg parasitoid of Tapajosa rubromarginata (Hemiptera: Cicadellidae)
Authors: TRIAPITSYN SERGUEI V. , LOGARZO GUILLERMO A. , VIRLA EDUARDO G. , DE LEÓN JESSE H.
Published: February 2019

Specimens of Gonatocerus virlai S. Triapitsyn, Logarzo & de León sp. n., which belongs to the ater species group of Gonatocerus Nees (Mymaridae), were reared in Argentina mostly from wild-collected and sentinel eggs of the sharpshooter Tapajosa rubromarginata (Signoret) (Cicadellidae: Cicadellinae: Proconiini) on citrus and corn plants. The distribution and host associations (both natural and laboratory) of G. virlai are given. Under quarantine laboratory conditions in the USA, it was successfully reared for many generations on an unnatural host, the glassy-winged sharpshooter, Homalodisca vitripennis (Germar).
A morphological and molecular description of a new Teleopsis species (Diptera: Diopsidae) from Thailand
Authors: FÖLDVÁRI MIHÁLY , POMIANKOWSKI ANDREW , COTTON SAMUEL , CARR MARTIN
Published: February 2019

A new species of Teleopsis (Diptera, Diopsidae) from Chiang Mai, Thailand is described and illustrated. Teleopsis thaii Földvári & Carr is shown to be a member of a species group, termed the dalmanni species group, along with three previously described species. Presented here are a morphological description of T. thaii and an allometric comparison of the species with other members of the Teleopsis genus. We also present multi-gene phylogenetic analyses to highlight the possible position of T. thaii within the dalmanni species group.
On the taxonomy of the genus Hymenicoides Kemp, 1917 (Crustacea: Decapoda: Brachyura: Hymenosomatidae), with resurrection of Limnopilos Chuang & Ng, 1991, and descriptions of two new species
Authors: NARUSE TOHRU , NG PETER K.L.
Published: February 2019

The systematics of the false spider crab genus Hymenicoides Kemp, 1917 (Hymenosomatidae), is revised. Hymenicoides carteri, type species of the genus, has a distinct locking structure of male abdomen as well as possession of a strong tubercle on the palm of the chela, features which are quite different from H. naiyanetri (Chuang & Ng, 1991), and H. microrhynchus Ng, 1995. The present study restricts Hymenicoides Kemp, 1917, for H. carteri and resurrects Limnopilos Chuang & Ng, 1991, for the other two species. Two new species, one of Hymenicoides from Myanmar, and one of Limnopilos from Sumatra, are also described. A key to the species of both genera is provided.
Nomenclatural notes on the names Gaeolaelaps and Geolaelaps (Acari: Laelapidae)
Authors: HALLIDAY R. B. , LINDQUIST E. E.
Published: February 2019

The genus-group names Geolaelaps and Gaeolaelaps (Acari: Laelapidae) are in widespread use in the formal taxonomic literature of acarology, for a genus or subgenus of predatory soil mites. They are also in use in the literature of ecology and agriculture, especially in discussions of the biological control of western flower thrips, Frankliniella occidentalis (Pergande), and sciarid larvae and gnats of the genus Bradysia, in greenhouse vegetables and ornamentals (Zhang, 2003). Examples of usage in mite taxonomy are Karg (1971), Evans & Till (1979) and Casanueva (1993) (Gaeolaelaps), and Karg (1979, 1993) and Xu & Liang (1996) (Geolaelaps). Examples of usage in ecology and agriculture include House et al. (1987) (Gaeolaelaps) and Gillespie & Quiring (1990) (Geolaelaps). There are many other examples of usage of both of these names in both print and internet documents, and there is considerable confusion surrounding their spelling and authorship. The purpose of this note is to resolve that confusion.
Kotumsaridae, a new family of subterranean amphipod crustaceans from India, with description of Kotumsaria bastarensis, new genus, new species. Zootaxa, 1589, 33–46. (ERRATUM)
Authors: MESSOULI M. , HOLSINGER J. R. , REDDY Y. R.
Published: February 2019

ERRATUM
New species and records of Keilbachia Mohrig (Diptera, Sciaridae) from the Oriental region
Authors: HIPPA HEIKKI , VILKAMAA PEKKA
Published: February 2019

The following new species are described: Keilbachia adunca (Brunei), K. ancylonema (Malaysia, Selangor and Myanmar, Kambaiti), K. camptonema (Malaysia, Selangor), K. clinata (Malaysia, Sabah), K. mirabilis (Malaysia, Selangor and Pahang), and K. orthonema (Malaysia, Sabah). K. curvispina (Edwards), comb. nov., from Sumatra, is redescribed and a lectotype is designated. K. acerspina Mohrig, previously known from Nepal, is recorded from Myanmar, Kambaiti.
First fossil Micropholcommatidae (Araneae), imaged in Eocene Paris amber using X-Ray Computed Tomography
Authors: PENNEY DAVID , DIERICK MANUEL , CNUDDE VEERLE , MASSCHAELE BERT , VLASSENBROECK JELLE , VAN HOOREBEKE LUC , JACOBS PATRIC
Published: February 2019

We apply Very-High-Resolution X-Ray Computed Tomography (VHR-CT) to a minute fossil spider (~1 mm long) from Eocene amber of the Paris Basin, France. We demonstrate that the newly described genus and species of Micropholcommatidae, Cenotextricella simoni, retains excellent details of the somatic and male pedipalpal morphology that allows unqualified comparison with extant species. Thus, in addition to calibrating the tree of life, such fossils can now be incorporated into cladistic matrices and their resultant phylogenies. This is the first fossil record of the family Micropholcommatidae, extending the known geological range of the family by 53 million years (lowermost Eocene) and that of the symphytognathoid clade by approximately 5–9 million years.
New taxa and records of Phaneropterinae (Orthoptera: Tettigoniidae) from China
Authors: LIU CHUN-XIANG , KANG LE
Published: February 2019

Two new genera, Semicarinata gen. nov., and Qinlingea gen. nov., are described from China, with description of one new species, S. colorata sp. nov., and one new combination, Q. brachystylata (Liu and Wang, 1998). Two species, i.e., Tamdaopteron major Gorochov, 2005, and Indogneta lata Ingrisch and Shishodia, 2000, as well as these two genera, are first records for China. We also gave additional descriptions about male stridulatory area of these two genera and the first female description of the species I. lata. A key to the genera with open tympana in China is provided.