bluuebunny/crossref_metadata_2025 huggingface dataset:
Genetic Variation Within and Among Mats of the Reindeer Lichen, Cladina Subtenuis
Authors: Beard Karen H. , Depriest Paula T.
Published: September 2002

Cladina subtenuis is the most commonly found ‘ reindeer lichen ’ in the southeastern United States. In the present study C. subtenuis was examined for polymorphism in the small subunit (SSU) ribosomal DNA (rDNA) within and among small clumps of podetia, here referred to as mats, from five geographical locations. Polymorphism in the SSU rDNA was not detected among podetia within mats, but polymorphism was detected among mats from a single location. Therefore, mats of C. subtenuis may represent a single genetic individual, although genetically distinct mats may occur in close proximity. Their proximity couldencourage sexual reproduction between genetically distinct individuals, which would maintain variation in these populations. The SSU rDNA types represented three different lengths and restriction-site patterns, reflecting the presence of sequence insertions of at least 200 nucleotides in the SSU rDNA coding region. Thei nsertions, differing in position, number and sequence, are group I introns similar to those previously reported in the Cladonia chlorophaea complex. The intron variation among the SSU rDNA types indicates that molecular evolution has occurred within this species, perhaps since its divergence from related taxa and dispersal across the southeastern United States.
Foliicolous Lichens From Madeira, with the Description of a New Genus and two New species and a World-Wide key of Foliicolous Fellhanera
Authors: Sérusiaux Emmanuël
Published: September 2002

A preliminary annotated list of the foliicolous lichens and of their lichenicolous fungi occurring in the laurisilva in Madeira is presented. Fellhaneropsis Séerus. & Coppins gen. nov. is introduced to accommodate Bacidia myrtillicola Erichsen and Bacidia vezdae Coppins & James, both being present in Madeira on living leaves. Two new species are described: Byssoloma kalbii Sérus. sp. nov. and Fellhanera seroexspectata Sérus. sp. nov. The following combinations are introduced: Fellhanera lambinonii (Sérus.) Lücking & Sérus.,Fellhaneropsis myrtillicola (Erichsen) Sérus. & Coppins, F. vezdae (Coppins & James) Sérus. & Coppins, Woessia apiahica (Müll. Arg.) Sérus., W. canariensis (Lumbsch & Vězda) Sérus. and W. vasakii (Vězda) Sérus. A key to all foliicolous species of Fellhanera is presented. Bacidia buxi Vězda & Vivant and Bacidia gorgonea. Vězda & Poelt are reduced into synonymy with Fellhaneropsis myrtillicola, Tapellaria similes Kalb with Byssoloma marginatum(Arnold) Sérus., and Bacidia michaeliana Sérus. with .Bacidia lambinonii Sérus.
Phenolic levels in Umbilicaria Americana in Relation to Enzyme Polymorphism, Altitude and Sampling Date
Authors: Swanson Andrew , Fahselt Dianne , Smith Dale
Published: September 2002

A significant negative correlation was found between the quantity of acetone-extractable phenolic compounds and the degree of enzyme polymorphism in eight eastern North American populations of Umbilicaria americana. In sites higher elevations, where UV irradiance was more intense, phenolic levels were significantly lower and within-stand enzyme variability significantly greater than in low elevation sites. Similarly, the phenolic content of thalli collected in Ontario, Canada, over a 16-month sampling period was negatively correlated with mean monthly UV-B irradiance.
Coenogonium: A Green Algal Lichen Without Photosynthetic Depression at High Water Contents
Authors: Thomas Michael A. , Nash Thomas H. , Tucker Shirley C.
Published: September 2002

For experiments run at 350 ppm CO2, the subtropical lichen Coenogonium does not exhibit net photosynthetic depression at water contents above those at which initial maximal values are reached. This is in contrast to most lichens for which such a depression occurs. We infer that the results reflect the unusual morphology of Coenogomum, in which the alga is the dominant symbiont, occurring as filaments at the lichen surface. This arrangement should minimize CO2 diffusion resistance to the algal surface. With light saturation occurring by 70 μmol photons m−2 s−1, Coenogoniium has one of the lowest light saturation values reported among lichens.
Lichens Growing on Glass in Antarctica
Authors: Schroeter B. , Sancho L. G.
Published: September 2002

Early stages of lichen colonization on glass are reported for the first time from Antarctica. In the initial establishment of the lichen thallus rhizomorphs play an importantrole for the attachment to the substratum and the exploitation of nutrient and photobiont resources. The micrographs presented indicate that in Antarctica colonization of bare substrata bylichens is not necessarily an extremely slow process. Furthermore, our findings demonstrate that the use of artificial substrata such as glass could provide a simple method for studies in lichencolonization and development in extreme environments.
Comparison of Lipids and Fatty Acids of the Lichen Rhizoplaca Peltata Collected from Foothill and High Mountain Habitats in the Same Region
Authors: Bychek Irene A. , Bychek Eugene A.
Published: September 2002

Lipids and fatty acids from the lichen Rhizoplaca peltata, growing either high in the mountains or in nearby foothills, were analysed and compared. It was shown that the same lichen species had considerable differences in the content of triacylglycerols, free sterols, wax esters, steryl esters, glycolipids, betaine and phospholipids, as well as in fatty acid content, depending on altitude.
Lichens of Skomer Island, West Wales
Authors: Wolseley P. A. , James P. W. , Coppins B. J. , Purvis O. W.
Published: September 2002

Skomer Island, a National Nature Reserve, lies off the West coast of Wales. This small island holds lichen communities of international importance associated with large populations of seabirds. Saxicolous lichen communities are described, 248 species listed and details given of a base-line monitoring programme, which will provide data on changes in the lichen flora over time and distinguish cyclical from permanent deleterious changes. Threats to island populations are discussed, such as the likely effects of oil pollution following the Sea Empress accident in February 1996.
A Review of Lichens in Metal-Enriched Environments
Authors: Purvis O. W. , Halls C.
Published: September 2002

The importance of mineralogy in determining lichen assemblages occurring on metalliferous rocks and derived soils is discussed with reference to environments enriched in metals from mine and smelter wastes and other sites involving human activities. A study of lichens growing on Cu and Fe sulphide-rich rocks in Scandinavia is presented. This data suggests that the composition of the lichen communities is governed by mineralogy, which influences the mobility and availability of Cu2+ and Fe2+ in the environment, rather than the total concentrations of either metals within the substratum. A new lichen community, the Lecideion inopis Purvis, is described characterized by the faithful species Lecidea inops, Psilolechia leprosa and several Cu-rich ecotypes. This community occurs alkaline environments where Cu is fixed predominantly as a secondary hydroxide or carbonate. Conversely, low pH, Fe sulphide-rich environments, where Cu may also be present, are characterized by the Acarosporion sinopicae, which includes several species with a rusty colouration. Contrasting assemblages rich in terricolous ephemeral species, especially Vezdaea spp. occurring in Pb/Zn-rich environments, and the comparatively species-poor assemblages on Cr and Ni-rich ultrabasic rocks, are also reviewed. Although mine spoil heaps and other mineralized environments are perceived in human terms as toxic, they may support a diversity of lichens, including rare taxa restricted to these sites. The systematic description of these lichen communities in relation to rock mineralogy and chemical processes are necessary if physiological tolerance and speciation are to be understood.
Australiaena Streimannii, a New Genus and Species in the Physciaceae From Tropical Australasia
Authors: Matzer M. , Mayrhofer H. , Elix J. A.
Published: September 2002

The genus Australiaena (lichenized Ascomycetes, Physciaceae) with the single species A. streimannii sp. nov. is described as new from tropical Australasia. Morphologically it is characterized by crustose thalli with an often radiate-plicate margin, ascospores with a type-B ontogeny, and filiform spermatia. The chemistry of this taxon is diverse (lobaric acid, conlobaric acid, xanthones, gyrophoric acid, lecanoric acid, 5-O- ethylhiascic acid, perlatolic acid), and three chemotypes are distinguished.
Effects of Thallus Damage on Interactions of Lichens with Non-Lichenized Fungi Under Natural and Laboratory Conditions
Authors: Glenn Marian G. , Gomez-Bolea Antonio , Orsi Ernest V.
Published: September 2002

The response of corticolous lichens to artificially induced chemical damage was compared to symptoms found on damaged lichens in the field. Experimental thalli were damaged and observed (1) in situ in a relatively pristine area, and (2) in a growth chamber either hydrated continuously or periodically. Lichenicolous fungi encountered included Cornutispora lichenicola, Lichenoconium erodens, Phoma cytospora, Vouauxiomyces truncatus, Hobsonia christiansenii, Pronectria oligospora, Nectria rubefaciens and its anamorph, Acremonium rhabdosporum, and a variety of hyphomycetes. The results support the idea that both thallus necrosis and moist conditions are necessary to produce the full suite of symptoms observed on ailing lichens in the field.
Photosynthetic Performance of Two Closely Related Umbilicaria Species in Central Spain: Temperature as a Key Factor
Authors: Sancho L. G. , Schroeter B. , Valladares F.
Published: September 2002

Net photosynthesis (NP) and dark respiration (DR) of thalli of the lichen species Umbilicaria grisea and U. freyi growing together in the same habitat the Sierra de Guadarrama, central Spain, were measured under controlled conditions in the laboratory and under natural conditions in the field over a range of photosynthetic photon flux densities (PPFD), thallus temperatures and thallus water contents. Laboratory experiments revealed that the photosynthetic response to PPFD at optimum thallus water content is very similar in both species. The light compensation points of NP increased from PPFD of c. 20 µmol m−2 s−1 at 0°C up to c. 100 µmol m−2 s−1 PPFD at 25°C. In both species light saturation was not reached up to 700 µmol m−2 s−1 PPFD except at 0°C. By contrast, the temperature dependence of CO2 gas exchange differed substantially between U. grisea and U. freyi. Both species gave significant rates at 0°C. Optimal temperatures of NP were always higher in U. grisea at various PPFD levels if the samples were kept at optimal thallus water content. NP showed maximal rates at 95% dw in U. grisea and 110% dw in U. freyi respectively. In U. grisea a much stronger depression of NP was observed with only 5% of maximal NP reached at 180% dw. At all PPFD and temperature combinations U. freyi showed higher rates of NP and more negative rates of DR if calculated on a dry weight basis. This was also true under natural conditions at the same site, when U. freyi was always more productive than U. grisea. The differences in the photosynthetic response to temperature between both species correlated well with the different distribution patterns of both species. The possibility of genetic control of the physiological performance of these species and its influence on their distribution patterns and autecology is discussed.
Ft-Raman Spectroscopy of Dirina Massiliensis f. Sorediata Encrustations Growing on Diverse Substrata
Authors: Edwards H. G. M. , Farwell D. W. , Seaward M. R. D.
Published: September 2002

The FT-Raman spectra of encrustations of Dirina massiliensis f. sorediata on a variety of substrata collected from several locations have been recorded and analysed. This is the first time that such a comprehensive spectroscopic study has been made of lichen encrustations of the same species on different substrata. This lichen is able to bring about deterioration in all six different lichen substrata studied. Calcium oxalate dihydrate is produced in significant amounts by the reaction of lichen-generated oxalic acid with various calcareous rock substrata, as well as particles of the particular substratum itself. However, for specimens growing on a non-calcareous substratum in an exposed maritime location only traces of calcium oxalate are produced and more noticeable quartz particles are incorporated into the thalli. The results indicate that this lichen can degrade its substratum highly effectively and there is evidence of different strategies being exhibited by lichens in different habitats and geographical locations.
A New Saxicolous Species of Arthothelium
Authors: Makhija Urmila , Patwardhan P. G.
Published: September 2002

Arhothelium saxicolum Makhija & Patwardhan sp. nov. is described from India. It occurs abundantly only on basaltic rock and is restricted to the northern parts of Western Ghats, South-West India.
Frequency, Diversity and Ecological Strategies of Epiphytic Lichens in the Swiss Central Plateau and the Pre-Alps
Authors: Dietrich Michael , Scheidegger Christoph
Published: September 2002

To identify representative quantitative criteria for the creation of a future Red List of epiphytic lichens, 849 trees in 132 long-term ecological observation plots in the Swiss Central Plateau and the Pre-Alps were surveyed by standard sampling. Based on the trees, frequency data of the lichen taxa observed are described by the log series model, indicating the controlling effect of few ecological factors. Based on the plots, four classes of scarcity, each comprising 25% of the species, were established. As a contribution to the development of a national, representative survey of lichens, α-diversity (species richness, species density) and β-diversity (dissimilarity) were calculated in terms of region, vegetation formation, vegetation belt and for their combinations. Differences in lichen diversity between the Central Plateau and the Pre-Alps were caused by the bigger elevational range in the Pre-Alps, which resulted in a higher species richness. α-Diversity of forest and non-forest were similar, whereas each vegetation formation showed one third of its species restricted to it. The contributions to the total lichen diversity of crustose, foliose and fruticose as well as of generative and vegetative species was calculated. Specific features along the altitudinal gradient of vegetation belts emerged: the percentage of crustose and generative lichens declined with every altitudinal step, increased in fruticose and vegetative lichens, and was the same in foliose species.
Continuous Monitoring of CO2 Exchange of Lichens in the Field: Short-Term Enclosure with an Automatically Operating Cuvette
Authors: Lange O. L. , Reichenberger H. , Walz H.
Published: September 2002

Field measurements of CO2 exchange by lichens are difficult because these poikilohydric organisms rely on direct hydration by rain, dew or fog. Continuous enclosure of thalli in conditioned, measurement cuvettes is, therefore, not experimentally reasonable. Porometric instruments with brief enclosure times, have proved useful for studies of lichen gas exchange but allow only spot measurements and recording is not easily possible for extended time periods. We describe a newly developed, automatically operating cuvette that allows long-term monitoring of lichen photosynthetic and respiratory CO2 exchange in the field. Samples, e.g. flat stone slabs covered with epilithic lichens, are positioned on the base of the cuvette where, when the cuvette is open, they are exposed to the same environmental conditions as naturally growing thalli. At regular intervals (typically 30 min), an upper lid automatically encloses the lichen for c. 3 min in a stirred cuvette; CO2 exchange is measured using IRGA techniques and microclimate data arc recorded. The successful operation of the cuvette is illustrated by means of diel time courses of CO2 exchange for selected, very different weather conditions.
Three New Species of Porina (Trichotheliaceae) from Lord Howe Island, Australia
Authors: McCarthy P. M.
Published: September 2002

Porina howeana sp. nov., P. Mariae sp. nov. and P. ocellatoides sp. nov. are described from shaded basalt on Lord Howe Island, New South Wales, Australia.
New Species or Interesting Records of Foliicolous Lichens. I. Trichothelium Argenteum (Lichenized Ascomycetes: Trichotheliaceae)
Authors: LÜcking Robert , Ferraro Lidia Itati
Published: September 2002

Trichothelium argenteum R. Lücking & L. I. Ferraro sp. nov. is distinguished by its very large perithecia with a well-developed, silvery white, basally black crown of setae, and by a non-radiate phycobiont. In young perithecia, the setae are confluent and form a characteristic, two-coloured disc-like expansion. Ascospores are 7-septate and bacillar. Trichothelium argenteum is known from a few populations in Costa Rica, Ecuador, Brazil, Paraguay, and Argentina.
New Species or Interesting Records Of Foliicolous Lichens. II. Flavobathelium Epiphyllum (Lichenized Ascomycetes: Melanommatales)
Authors: LÜcking Robert , Aptroot André , Thor Göran
Published: September 2002

Flavobathelium epiphyllum gen. et sp. nov. is described and illustrated. The new taxon is characterized by perithecia and pyenidia embedded in pulverulaceous, uni- to plurilocular pseudostromata, small, transversely septate ascospores, and the presence of bacillar to filiform macroconidia with gelatinous appendages and simple microconidia. Possible relationships are seen with the genus Phyllobathclium. Flavobathelium epiphyllum is a common species in Central and South American lowland to submontane rain forests with undisturbed vegetation. It is mostly found along moderately illuminated parts of the understory, such as small, natural light gaps.
Growth from the Centre in an Umbilicate Lichen
Authors: Hestmark Geir
Published: September 2002

The pustules on the surface of the umbilicate lichen Lasallia pustulata are moved away from the centre and towards the thallus margins with an average speed of 1.8 mm per year, revealing a growth pattern organized radially and flowing from the centre of the thallus. Pustules close to the margin move substantially faster than those close to the centre; for each 10 mm added to the initial distance from the centre, the speed of outward movement is about doubled. This suggests that intercalary growth over the entire thallus is displaced outwards in an accumulating manner, accelerating the pustules. Moving away from the centre the pustules expand, often collapse in the middle, and mature by developing isidia. In the less active (senescent ?) margins the pustules are eroded away and the supplementary intercalary growth seems insufficient to keep the outflowing thallus intact. Thus the margins are disrupted into irregular lobes.
The Phylogeny of Lecanactis (Opegraphaceae)
Authors: Tehler Anders , Egea José M.
Published: September 2002

The genus Lecanactis, with 24 species, has been phylogenetically analysed using cladistic parsimony methods and support tests. Morphological, anatomical and chemical data were used, comprising 38 characters. Twelve equally most parsimonious trees were obtained. The successive approximations character weighting method gave one most parsimonious tree. The ingroup, Lecanactis, is supported as monophyletic. Although parsimony jackknifing and Bremer support indicate that the trees are poorly supported, some groups are wholly or partly distinguished in both the strict consensus tree, the successive weighting tree and the Jac tree.
Protoparmelia Hypotremella, a New Sterile Corticolous Species from Europe, and its Lichenicolous Fungi
Authors: Aptroot A. , Diederich P. , van Herk C. M. , Spiers L. , Wirth V.
Published: September 2002

Protoparmelia hypotremella van Herk, Spier & Wirth, a sterile corticolous lichen, is described as a species new to science, so far known from Austria, SW Germany and the Netherlands. It is rapidly spreading in Germany and the Netherlands. It is apparently close to P. oleagina, and is often found growing together with that species on oak trees along roads in the Netherlands. Both species are host to the lichenicolous fungi Sphinctrina anglica and Tremella wirthii.
Intraspecific Interactions and Parasitism in an Association of Rhizocarpon lecanorinum and R. Geographicum
Authors: Clayden Stephen R.
Published: September 2002

Intraspecific thallus interactions in mixed populations of Rhizocarpon lecanorinum and R. geographicum in North Wales are described. In contrast to the most commonly occurring morphotype of R. geographicum, which forms mosaics comprised of sharply delimited individual thalli, R. lecanorinum thalli consistently merge with one another at intraspecific contacts, leaving no visible trace of their initial boundaries. On the analogy of mycelial interactions in populations of wood-rotting and other non-lichen-forming fungi, it is postulated that these differences are the outcome, in R. geographicum, of somatic incompatibility between distinct genotypes, and in R. lecanorinum of somatic compatibility among genetically identical or very closely related thalli. Possible mechanisms for the evolution of clonal population structure in R. lecanorinum and links between somatic interactions and reproductive output in these species are discussed. Facultative parasitism of R. lecanorinum by Schaereria fuscocinerea, and resistance to such parasitism by cooccurring R. geographicum, are also documented.
Population Structure of the Epiphytic Lichen Usnea Longissima in a Boreal Picea Abies Canopy
Authors: Gauslaa Yngvar
Published: September 2002

Usnea longissima was sampled in SE Norway on six main branches of one Picea abies, comprising its full vertical canopy range. Nearly all specimens (n = 781) were unattached and fragmented, in contrast with associated lichens. A combination of weak axes and lack of holdfasts resulted in a population of repeatedly displaced thalli, a similar situation as in ground-dwelling vagant lichens. Comparisons with a litterfall population indicated that thalli, regardless of size, had a similar probability to end as litter on the ground. Specimens were frequently sorediate, but only 12 small thalli with a basal holdfast had been directly recruited through soredial establishment. Its patchy distribution within apparently homogenous forest stands is probably a consequence of a rare successful long-range dispersal through soredia coupled with an abundant local dispersal of coarse thallus fragments. Old forests with ample, diffuse skylight at lower and humid canopy levels are probably essential to maintain a vagant epiphytic life form with a predominantly downward dispersal of thallus fragments.
Visualization of Extracellular Deposits in Recent and Subfossil Umbilicaria Hyperborea
Authors: Fahselt Dianne , Alstrup Vagn
Published: September 2002

Field emission scanning electron microscopy was used to characterize mycobiont wall surfaces in Umbilicaria hyperborean from Greenland. To determine the precise intrathalline distribution of phenolics, comparisons were made of hyphal surface features and dimensions before and after acetone extraction. Stratification was evident within the medulla, as extracellular phenolics were observed only on hyphae near or within the algal zone. The outside diameter of hyphae in this region was thus significantly greater than in the remainder of the medulla. Surface deposits were also examined in 1350-year-old subfossil thalli and hyphal diameters were compared statistically to those in extant thalli. The mean hyphal diameter in the upper medulla was not significantly less in subfossil specimens than in recent thalli, suggesting that phenolic cover was maintained in spite of glaciation. However, after ice burial phenolic masses tended to be flatter than in recent specimens and finely tubcrculate. The appearance of mycobiont hyphae in the cortex of subfossil thalli seemed to be the same as in extant thalli, except that there tended to be more compressible, smaller and less conglutinated filaments near the algal layer.
Lichenicolous Fungi from the Western Pyrenees, France and Spain. IV. Ascomycetes
Authors: Etayo Javier , Diederich Paul
Published: September 2002

Twenty species of lichenicolous ascomycetes are recorded for the western Pyrenees. Capronia hypotrachynae Etayo & Diederich (on Hypotrachyna species, also known from the Canary Islands and from Papua New Guinea), Llimoniella pubescens Etayo & Diederich (on Lepraria species, also from Scotland and Papua New Guinea), Polycoccum microcarpum Diederich & Etayo (on Cladonia species, also from Scotland), Skyttea megalosporae Etayo & Diederich (on Megalospora tuberculosa), Sphaerellothecium cinerascens Etayo & Diederich (on Cladonia parasitica) and S. parmeliae Diederich & Etayo (on Parmelia s. str., also from Finland) are described as new.
A New Species of Catillaria from Coastal Mediterranean Regions
Authors: Tretiach M. , Hafellner J.
Published: September 2002

A revision of several specimens of Catillaria mediterranea Hafellner revealed that two species can be recognized: C. mediterranea s. str., which has 8–16 spores per ascus, grows on foliose and fruticose lichens, and has a Mediterranean- Macaronesian, montane distribution, and C. praedicta Tretiach & Hafellner sp. nov., which has (16-) 24–32 (-48) spores per ascus, occurs on shrubs in coastal Mediterranean maquis, and has a Mediterranean, maritime distribution. The systematic affinities of the two species with the 8-spored C. nigrodavata (Nyl.) Schuler are briefly discussed.
The Effect of Agronomic Photosystem-II Herbicides on Lichens
Authors: Jensen Manfred , Linke Kerstin , Dickhäuser Andreas , Feige G. Benno
Published: September 2002

The sensitivity of Hypogymnia physodes, Lobaria pulmonaria and Peltigera aphthosa to six photosystem II herbicides and to DBMIB was tested in the laboratory by chlorophyll fluorescence and oxygen-exchange measurements. In addition, experiments with freshly isolated photobiont cells from H. physodes and L. pulmonaria were performed. Generally, the lichens were most sensitive to the urea herbicides diuron and isoproturon, whereas the triazines atrazine, terbuthylazine, and simazine and the triazinone metamitron were less inhibitory. Among the three lichen species investigated, H. physodes was the most sensitive to the urea herbicides. For the other agents, no significant differences between lichen species could be found. The highest pI50 values obtained from dose response curves were around 6.5 for isolated photobionts, but most values for lichen thalli were in the range 5–6. Thus, there is no particular sensitivity of green algal lichen photobionts to photosystem II herbicides as compared to other algae, higher plant chloroplasts or protoplasts. In nature, we observed recovery from (damaging) treatment with 10−5 mol diuron 1−1 for H. physodes within weeks. Therefore, damage to lichens from the use of photosystem-II herbicides in agriculture is probably only of very local occurrence.
Lecanora Barkmaneana, a new Nitrophilous Sorediate Corticolous Lichen from the Netherlands
Authors: Aptroot A. , van Herk C. M.
Published: September 2002

Lecanora barkmaneana, a sorediate corticolous lichen belonging to the Lecanora subfusca group, is described as a species new to science on the basis of numerous sterile collections and two richly fertile collections. It is known so far only from the Netherlands, where it often grows on roadside trees. It seems to be spreading rapidly at nutrient-rich sites.
How Agriculture Affects Lichen Vegetation in Central Switzerland
Authors: Ruoss Engelbert
Published: September 2002

The results from several floristic and bioindication projects in Central Switzerland (1986–1998), focusing on the effects of agriculture, particularly from nitrogen and agrochemical emissions, are summarized. The abundance of nutrienttolerant lichen species and the decreased occurrence of nitrophytic lichens are both correlated with agricultural land use and high atmospheric deposition. The impact could be demonstrated not only by the distribution patterns of nutrient-tolerant species, but also by the high N content of lichens such as Physcia caesia. Positive growth response of Cetraria islandica to application of mineral nutrients and increased CO2 in open-top chambers at high altitudes was also observed. The methods and a survey of the results from studies conducted within the framework of the lichen research programme of the Natural History Museum of Lucerne are presented here. A discussion of the issues shows the need for further studies concerning the influence of introduced nutrients on the biodiversity of lichens.
The Response of Different Cladonia Species after Treatment with Fertilizer or Lime in Heathland
Authors: Vagts Irene , Kinder Michael
Published: September 2002

The effect of deposition of different fertilizers and of lime on lichens was investigated under field conditions. Three terricolous lichens were tested: Cladonia floerkeana, C. furcata and C. portentosa. The lichens were exposed to three types of powdered fertilizer (NPK-, N-, P-fertilizer) and lime. The applied nutrients were recorded in lichen thalli and soil. The influence on lichen growth rate and vegetation cover was estimated over 2 years. The response was a clear change in vegetation cover with an exceptional stimulating effect of NPK on the moss Polytrichum formosum. A lethal effect was observed on Cladonia floerkeana following exposure to lime. NPK and P had the most significant stimulatory effect on C. furcata. Only NPK promoted the growth of C. portentosa and P, N or Ca had a negative effect.
Bacidia Neosquamulosa, a new and Rapidly Spreading Corticolous Lichen Species from Western Europe
Authors: Aptroot A. , van Herk C. M.
Published: September 2002

Bacidia neosquamulosa Aptroot & van Herk, a new corticolous lichen species, is described from the Netherlands, where it seems to be rapidly spreading and where it is considered to be a neophytic species. It is also reported from Belgium and Great Britain. It resembles corticolous morphs of the rather variable B. arnoldiana, but differs from this and other species by the longer, conspicuously septate, filiform, curved, macroconidia, the longer ascospores and the squamuloseisidiate thallus.
Ecology of Foliicolous Lichens at the ‘Botarrama’ Trail (Costa Rica), a Neotropical Rainforest. IV. Species Associations, their Salient Features and Their Dependence on Environmental Variables
Authors: Lücking Robert
Published: September 2002

A community ecology approach using an association of species test and a principal component analysis resulted in the recognition of two major and two minor foliicolous lichen communities in a neotropical rainforest. The major communities arc governed by microclimatic factors, one characteristic of the shady understorcy and the other confined to light gaps. The minor communities are due to subtle phorophyte preferences towards palm and dicot leaves. The shady understorey community is dominated by the families Anhoniaceae, Opegraphaceae, Tnchoihdiaceae and Pilocarpaceae, which predominantly have Phycepeltis as their phycobiont, thin thalli, abundant sexual reproduction, small ascospores produced in high numbers, and pyenidial conidiomata. The light-gap community is mainly composed of the families Gomphillaceae and Ectolechiaceae, with Trebouxia as their phycobiont, thickly crystalline or whitish, dispersed thalli, frequent asexual reproduction, large ascospores produced in low numbers, and specialized campylidia and hyphophores as conidiomata. Phycobiont, thallus structure, and the mode of reproduction are considered as adaptations to different microsites, whereas the shape and size of ascospores and conidia seem to be of less importance. The foliicolous lichen communities reflect the spatial and temporal structure of the forest, in particular the continuity of the shady understorey in comparison to the discontinuity of the light gaps. While the formation of the shady understorey community follows more deterministic patterns, the light-gap community exhibits high stochasticity.
Post-Fire Colonization of a Mediterranean Forest Stand by Epiphytic Lichens
Authors: Longán Angels , Gaya Ester , Gómez-Bolea Antonio
Published: September 2002

The post-fire colonization of a Quercus ilex forest by epiphytic lichens has been studied in Catalonia (NE Spain), eleven years after a fire. Specific richness and lichen biomass have been studied separately on Quercus ilex, Erica arborea, Rosmarinus officinalis. and Cistus albidus, to reveal possible differences among phorophytes in facilitating lichen establishment. Shrubs play an important role in colonization by common species whereas the stools of Quercus ilex offer a suitable substratum for rarer species. Cistus albidus, with the highest lichen diversity and 98·2% of the total biomass of macrolichens, is the most suitable phorophyte for lichen establishment.
A New Species of Schaereria from Tasmania
Authors: Kantvilas Gintaras
Published: September 2002

The new species, Schaereria bullata Kantvilas from alpine Tasmania, is described and illustrated, with notes on its characterization, distribution and habitat. A key to the seven known species of the genus is provided.
Steinia Australis, A New Species in the Lichen Family Aphanopsidaceae
Authors: Kantvilas G. , McCarthy P. M.
Published: September 2002

Steinia australis Kantvilas & P. M. McCarthy is described, based on a specimen from Tasmania. The new species, the second in the genus and only the third in the family Aphanopsidaceae, differs from S. geophana by its larger ascospores, well-developed exciple and scurfy, leprose thallus.
The Lichens of Rarotonga, Cook Islands, South Pacific Ocean I: Pyrenocarpous Taxa
Authors: McCarthy P. M.
Published: September 2002

Four new pyrenocarpous lichens are described from Rarotonga, Cook Islands, South Pacific Ocean: Porina deminuta (Trichotheliaceae), Pyrenula ochraceoflava var. pacifica (Pyrenulaceae), Strigula decipiens var. divisa (Strigulaceae) and Verrucaria fortuita (Verrucariaceae). The new combinations Porina eminentior (Nyl.) P. M. McCarthy and P. exocha (Nyl.) P. M. McCarthy are made for Thelenella eminentior Nyl. (Clathroporina eminentior (Nyl.) Müll. Arg.) and Verrucaria exocha Nyl. (Clathroporina exocha (Nyl.) Müll. Arg.), respectively. Brief descriptions and additional notes are provided for 26 other taxa. The phytogeographical affinities and possible origins of the pyrenocarpous lichens of Rarotonga and Lord Howe Island are discussed.
Changes in Water Economy in Relation to Anatomical and Morphological Characteristics During Thallus Development in Parmelia Acetabulum
Authors: Fos Simón , Deltoro Vicente I. , Calatayud Ángeles , Barreno Eva
Published: September 2002

The influence on uptake and water loss of the structural changes experienced by Parmelia acetabulum during thallus development were investigated. Small specimens were characterized by flat lobes and a thin thallus and cortex. Large specimens appeared strongly rugose, imbricate and irregularly folded, and had a significantly thicker cortex and medulla than small thalli. Maximum water storage capacity did not differ between large and small thalli, although water retention was much higher in large thalli, presumably due to the interaction of structural characteristics and a higher boundary layer resistance. This translated into a longer duration of the period of thallus hydration in large thalli compared to small thalli. Incubation of thalli in water-vapour-saturated atmospheres induced full recovery of photosynthetic electron transport to the values before desiccation in small thalli but only induced a partial recovery in large thalli. The close correlation between photosynthetic electron transport and net photosynthesis during desiccation found in this species suggested that carbon-fixation activity could be regained to a larger extent by incubation of thalli in water vapour in small compared to large thalli. The higher ability for water vapour uptake of small thalli might allow them to efficiently use small amounts of intermittently available water or periods of high relative humidity. The significance of this differential ability to utilize water is discussed with regard to the known ecological preferences of the species.
Dispersal Patterns of Terricolous Lichens by Thallus Fragments
Authors: Heinken Thilo
Published: September 2002

Dispersal patterns from seven terricolous lichen species, with a high capacity for asexual reproduction by fragmentation (Cetraria muricata, Cladonia species), were studied in differing vegetation types in north-eastern Germany. Marked lichen thalli were fragmented by trampling. After 15 days the spread of thallus fragments was monitored. Whereas most of the fragments that were dispersed by wind remained within a 20-cm radius from the source, the maximal dispersal distance was 57 cm in a dry sand grassland and 68 cm in an open pioneer pine forest. Dispersal was negligible in a closed old-growth pine forest. Several fragmented lichen cushions were disturbed and removed by animals, and led to a maximal dispersal distance of 9·70 m. These results suggest that: (a) thallus fragments provide good short-distance dispersal in open vegetation, but are inefficient for long-distance dispersal, and (b) wind and animals are important factors for the dispersal of thallus fragments. For restoration management of man-made substrata, artificial introduction of lichen thalli is proposed.
Pcr Primers for the Amplification of Mitochondrial Small Subunit Ribosomal DNA of Lichen-forming Ascomycetes
Authors: Zoller Stefan , Scheidegger Christoph , Sperisen Christoph
Published: September 2002

Four primers for the amplification of mitochondrial DNA of lichenforming ascomycetes are presented. The primers match the conserved regions U2, U4, and U6, respectively, of mitochondrial small subunit (SSU) ribosomal DNA (rDNA). Polymerase chain reaction using different combinations of the primers produced single amplification products from DNA of eight lichen-forming fungal species but did not amplify DNA of two axenic cultured algal species. The amplification product obtained from Lobaria pulmonaria was sequenced and the 894-bp sequence was compared with the mitochondrial SSU rDNA sequence of Podospora anserine. The two sequences revealed more than 76% identity in the conserved regions U3 to U5 demonstrating that we amplified mitochondrial DNA. The primers matching U2 and U6 yielded amplification products of 800–1000 bp depending on the species examined. The variation observed suggests that mitochondrial SSU rDNA may be useful for phylogenetic analyses of lichen-forming ascomycetes.
A Comparison of Morphological, Chemical and Molecular Characters in Some Parmelioid Genera
Authors: Crespo Ana , Gavilán Rosario , Elixt John A. , Gutiérrez Gabriel
Published: September 2002

Morphological and chemical data from 64 genera of Parmeliaceae, including the most common genera in Europe, were analysed by means of neighbour–joining and parsimony analysis. Two presence/absence matrices were constructed, one for the 64 genera and a smaller one for the most common European genera. Probabilities were too low to allow grouping of the genera and only a few pairs of genera had a probability of >50% for both matrices. A molecular matrix (nuclear rDNA-ITS) of the most common genera in Europe was constructed and analysed by the method of maximum likelihood. These results were much more informative than the non-molecular matrix for establishing phylogenetic relationships among the genera.
Variability of Ribonuclease Activity in Lichen Thalli
Authors: de los Ríos Asunción , Ramírez Rosalía , Estévez Pilar
Published: September 2002

RNase activity in Lasallia hispanica, Parmelia omphalodes and Cornicularia normoerica collected at different times and in different habitats, have shown variability that cannot be solely attributable to variation in habitat or season. Electromorph pattern of RNase activity in L. hispanica and C. normoerica, showed variations among different collection times; some bands are constant but other showed high variability, which could be related to physiological changes. It is also noteworthy that in both species one band is exclusively present in thalli from one of the sampling localities. This could be the result of an adaptation to this habitat and/or due to genetic differences.
Photobiont Inventory of a Lichen Community Growing on Heavy-Metal-Rich Rock
Authors: Beck Andreas
Published: September 2002

The photobiont inventory of a stand of the Acarosporetum sinopicae, lichen community comprising saxicolous, chalcophilous lichens, has been analysed. Investigated lichen species were Acarospora rugulosa, A. sinopica, Bellemerea diamartha, Lecanora polytropa, L. subaurea, Lecidea silacea, L. lapicida, Rhizocarpon geographicum, and Umbilicaria cylindrical. For all these lichen species this is the record of the photobionts, except for L. lapicida. The photobionts were cultured axenically and investigated using light microscopical and molecular methods (ITS-sequence analyses). Every lichen species contained only one photobiont species. All photobionts belong to Trebouxia jamesii, but two different subspecies were found with the morphological differences corresponding to molecular differences. The new subspecies T. jamesii subsp. angustilobata is described, differing from the typical T. jamesii by a crenulate chloroplast but identical to the latter taxon in respect to the pyrenoid structure in the light microscope. These results are discussed with respect to the photobiont inventory of the Physcietum adscendentis, analysed in an earlier study.
Two New Species of Micarea From South Africa
Authors: Coppins B.J.
Published: September 2002

Two new species of Micarea are described from South Africa: M. almbornii Coppins, on loose sandstone from Stellenbosch (Western Cape) and M. endoviolascens Coppins, on damp soil from Transvaal. A note is given on Lecidea geïna Stizenb.
The Lichens Of Rarotonga, Cook Islands, South Pacific Ocean II: Parmeliaceae
Authors: Simone Louwhoff H. J. J. , Elix John A.
Published: September 2002

Ten parmelioid taxa in five genera were identified from Rarotonga, Cook Islands, South Pacific Ocean. These comprise: three species of Bulbothrix, one species of Canoparmelia, four species of Parmotrema, one species of Rimelia and one species of Xanthoparmelia. Canoparmelia rarotongensis Louwhoff & Elix is described as new, being distinguished from other species of this genus by its unique secondary chemistry (lecanoric acid) and conspicuous, inflated isidia. The majority of parmelioid lichens on Rarotonga have cosmopolitan and pantropical distributions. Bulboihrix is reported for the first time from the Polynesian region.
Sulcopyrenula, A New Pyrenocarpous Lichen Genus (Pyrenulaceae, Lichenized Ascomycota)
Authors: Harada H.
Published: September 2002

Sulcopyrenida H. Harada gen. nov. is described. It differs from other genera of Pyrenulaceae by having longitudinally sulcate spores. The genus is represented by two species: S. canellae-albae (Fée) H. Harada comb. nov. and S. staurospora (Tuck.) H. Harada comb. nov. Both species have been previously reported from N. and S. America, and the latter species is now reported from Asia.
Proposal for a Standardized Nomenclature and Characterization Of Insoluble Lichen Pigments
Authors: Meyer Barbara , Printzen Christian
Published: September 2002

A method for the characterization of insoluble lichen pigments is presented. The method relies on pH–dependent colour reactions before and after pretreatment with HNO3. Diagnostic reactions for 22 different pigments and a key for their identification are given. A standardized nomenclature and a procedure similar to the typification of species in systematic botany is proposed in order to avoid ambiguous naming.
Bark pH and susceptibility to toxic air pollutants as independent causes of changes in epiphytic lichen composition in space and time
Authors: van Herk C. M.
Published: September 2002

The lichen composition on wayside Quercus robur in the Netherlands was related tobark properties (pH, EC, NH4+, SO42-, NO3-)and levels of air pollution (SO2 and NH3). The pH of the bark and the susceptibility to toxic substances appear to be the two major primary factorsaffecting epiphytic lichen composition. These factors have independent effects on the lichen composition.Most of the so-called nitrophytic species appear to have a low sensitivity to toxiceffects of SO2; their only requirement being a high bark pH. An increased bark pH appears to be the primary cause of the enormous increase in nitrophytic species and the disappearance of acidophytic species over the last decade in the Netherlands. Measurements of ambient NH3 concentrations in air show that there is a nearly linear relationship between the NH3concentration andthe abundance of nitrophytes on Quercus. The abundance of nitrophytes was not correlated with SO2 concentrations. Most of the acidophytic species appear very sensitive to NH3 since in areas with concentrations of 35 µg m-3 or more, all acidophytic species have disappeared. Current methods using species diversity to estimate or monitor SO2 air pollution need some modification, otherwise the air quality may be erroneously considered to be relatively good in areas with high NH3 levels.
A new method for the accurate in situ monitoring of chlorophyll a fluorescence in lichens and bryophytes
Authors: Schlensog Mark , Schroeter Burkhard
Published: September 2002

A new method has been developed to measure chlorophyll a fluorescence of bryophytes and lichens in situ in the field. Specially designed aluminium stands fixed to rocks or other suitable substrata allow several cryptogam thalli to be measured sequentially with only one pulse amplitude modulated (PAM) chlorophyll a fluorescence analyser. The stands guarantee a fixed position for the fluorescence probe in relation to the thallus surface and also provide the possibility to darken the thalli so that both the apparent and the potential quantumefficiency of PSII (▵F/Fm′ and Fv/Fm) can be measured. A detailed analysis of chlorophyll a quenching parameters (qP, qN, NPQ) can also be performed with only minimal changes in the environmental conditions. The method allows easy detection of periods of metabolic activity and inactivity of the poikilohydric organisms. In combination with gas–exchange and microclimatic measurements the carbon balance also can be calculated. The method is described in detail and examples of its application in the field are presented
Rhizocarpon Timdalii, a New Lichen Species from North-West Europe and North-east North America
Authors: Ihlen Per G. , Fryday Alan M.
Published: September 2002

The new species Rhizocarpon timdalii, is described from north-west Europe and north-east North America. The ecology and distribution of the new species are also discussed.