bluuebunny/crossref_metadata_2025 huggingface dataset:
Rhombognathus auster, a new rhombognathine mite (Acari, Halacaridae) from southern South America
Authors: BARTSCH ILSE
Published: July 2005

A new species of rhombognathine mites. Rhomhognathus auster, is described. The species lives intcrtidally on the shores of eastern Tierra del Fuego. Judging by the shape of idiosoma. legs and claws, R. auster is presumed to live amongst slender, tubular algae in semi‐exposcd and sheltered habitats.
Myxinidocotyle gen.n. and Lophocotyle Braun (Platyhelminthes, Monogenea, Acanthocotylidae) with descriptions of three new species from hagfishes (Chordata, Myxinidae)
Authors: MALMBERG GÖRAN , FERNHOLM BO
Published: July 2005

Three new species of monogencans parasitizing hagfish are described: Myxinidocotyle californica gen. et sp.n., M. japonica sp.n. and Lophocotyle novaezeelandica sp.n. The pseudohaptor of Myxinidocotyle has 8+8 transverse ridges that never attain the radial arrangement characterizing Lophocotyle and Acanthocotyle. There are no sclerites in the pseudohaptoral ridges of Myxinidocotyle and Lophocotyle. The male reproductive system of Myxinidocotyle and Lophocotyle differs from that of Acanthocotyle in having a bilateral accessory gland apparatus and an armed copulatory organ. The femrle reproductive system has a large vaginal receptaculum scminis; in Myxinidocotyle with two vagino‐intestinal canals. The uterine pore and the male genital pore are close together and the vaginal pore is situated posterior to them, on the left ventral side of the body. We conclude that Myxinidocotyle, Lophocotyle and Acunrkocotyle represent different levels of acanthocotylid evolution. Myxinidocotyle being the most primitive. The family Acanthocotylidac is divided into the subfamilies Myxinidoeutylinae subfam.n., Lophocotylinae and Acanthoeutylinae.
Studies on the development of the oncosphere and procercoid of Cyathocephalus truncatus (Cestoda) in the intermediate host, Gammarus pulex
Authors: OKAKA CHRISTOPHER E.
Published: July 2005

Cultured eggs of Cyathocephalus truncatus were found to develop optimally at 20°C, but they did not develop at 6°C and above. The oncosphere of the tapeworm has been seen to be a true hexacanth embryo. Specimens of the intermediate host. Gammaurs pulex, were infected by being forced to feed on pieces of lettuce leaves on which infective eggs of the tapeworm were placed. The young drveloping porocercoid was first recovered from the haemocoel of Gammarus pulex 5 days after infection. The procercoid was found to reach the infective stage within the amphipod after about ten weeks.
Echinoderes higginsi sp.n. (Kinorhyncha, Cyclorhagida) from the southern North Sea with a key to the genus Echinoderes Claparède
Authors: HUYS RONY , COOMANS AUGUST
Published: July 2005

Echinderes higginsi sp.n. is described from shallow subtidal sand north of the mouth of the Westerscheldt estuary, The Netherlands. The species is compared with other species having the same spine formulac. It appears to be most closely related to E. kristenseni Higgins. 1983 and, to a lesscr extent. to E. riedli Higgins, 1966. Condyldres multispinosus (McIntyre. 1962) is reported from the same locality, the first report of this centroderid kinorhynch since the original description. Some additional information on E. levunderi Karling, 1954 is presented. The presence of lateroventral adhesive tubes on the fourth segment, lateral spines on segment 7 and small additional setae on segment 12 in the male is noted. Echinoderes canariensis Greeff, 1869 and E. agigens Bǎcescu, 1968 are considered species inquirendae. The distribution of adult North Sea Kinorhyncha is reviewed. Finally, an up‐dated key. covering 43 valid species of Echhinderes, is presented.
Peculiar new deep‐sea desmoscolecid species of the genus Protricoma (Nematoda) from Corsica
Authors: DECRAEMER WILFRIDA , SOETAERT KARLINE
Published: July 2005

Three new species of Protricoma were found among the material collected during a deep‐sea survey near Corsica: P. Maramphis sp.n., P. mocrocephala sp.n. and P. corsicana sp.n. The new species are characterized by their small body size. a slender cylindrican body with finely annulated body cuticle and a very small head.
Namibnema papillata gen. et sp.n. and Axonolaimus deconincki sp.n. (Nematoda, Axonolaimoidea) from marine and estuarine beaches of southern Africa
Authors: VINCX MAGDA , FURSTENBERG JOHAN P.
Published: July 2005

Namibrema papillata gen. et sp.n. is closely related to Nicascolaimus punctatus Ricmann, 1986.hnema papillutu gen. ct sp.n. is closcly related to Nicuwolaimus punctutur Ricmann. 1986. The presence of twelve stomatal rugae, the punctated cuticle and the shape of the pre‐anal supplements indicate a relationship with species of the Chromadorina. Axonolaimus deconincki sp.n. is chjaracterized by the complex nature of the gubernaculum.
Sabellaria wilsoni sp.n. (Polychaeta, Sabellariidae) from the southeast coast of Brazil
Authors: DA CUNHA LANA PAULO , GRUET YVES
Published: July 2005

A new species of Sabellaria is described from the southeast coast of Brazil and compared with related species known from South or North America.
The phylogenetic position of the heteropodAtlanta gaudichaudiSouleyet (Mollusca, Gastropoda), a spermatological investigation
Authors: JAMIESON BARRIE G. M. , NEWMAN LESLIE J.
Published: July 2005

The male adult, the spermatophore and the spermatozoa ofAtlanta gaudichaudiare described. No parasperm are demonstrable. The ultrastructure of the spermatozoa is shown to differ fundamentally from that of the Archaeogastropoda and Neritopsina and to conform broadly with that of the Mesogastropoda (excepting Cyclophoracea, Viviparacea and Cerithacea), the Neogastropoda and the epitonacean heterogastropods. In having a nuclear fossa, containing the axoneme, which penetrates the entire length of the nucleus as the intranuclear canal, the spermatozoon especially resembles that of some members of the Rissoacea, Littorinacea, Eulimacea, Tonnacea, Volutacea and Conacea. Absence, inAtlantaof type 7 parasperm (sensuKoike 1985), which characterize the Calyptraeacea, Cypraeacea, Tonnacea, Buccinacea, Volutacea (including the Mitracea) and most Conacea, or of type 8 parasperm of the Conidae, contraindicates heteropod relationship with these taxa. Relationship to the Rissoacea cannot be ruled out, though presence of oligopyrene, cusperm‐like sperm in these (Bythiniidae) and pterotracheid heteropods is a questionable indication of affinity. Relationship of heteropods with the Littorinacea is suggested by similarity of the sperm, presence in carinariid heteropods and littorinids of nurse cells, and apparent homology of the male reproductive apparatus in both groups. The latter two similarities require confirmation, however. It is suggested that deposition of spermatophores on the exterior of the female in atlantid heteropods is a primitive mode of sperm transfer compatible with relationship with the Littorinacea.
Chromosomes of two species of Tisbe (Copepoda, Harpacticoida) belonging to different groups of sibling species
Authors: LAZZARETTO IVANA , LIBERTINI ANGELO
Published: July 2005

Karyological investigations curried out on Tisbe marmorata and T. dobzhanskii confirmed the diploid value 2n= 24 for the former species; on the basis of more than 200 observations it was ascertained that the diploid number of the latter species corresponds to 22, in contrast with a previous report. The karyotypes of these two species differ both in morphology and in location of satellites, not only from each other, but also from all other species so far examined. A close comparison of the two species with their respective siblings is made.
A re‐investigation and re‐interpretation of the cumacean photoreceptor
Authors: MEYER‐ROCHOW V. BENNO
Published: July 2005

Evidence for and against the view that the singular eye in Diastylis rathkei (Cumacea) represents a regressed compound eye is summarised and supplemented with new ultrastructural observations. Neither in the adult nor in the larval manca stage are even traces of a facetation found. The eye consists of two closely apposed eye halves and the four lenticular complexes in each appear to increase in size with age along an isometric growth curve. Each lenticular complex consists of a lens rich in glycogen‐like particles and a rhabdom made up of regularly aligned microvilli 0.075 μm in diameter. No more than three retinula cells contribute to each lenticular compkx. Retinula cells contain presumed carotenoid bodies 0.4–0.5 μm in diameter and clusters of electron‐opaque glycogen matcrial near the proximally‐located nuclei. The bulk of the eye is occupied by cells crowded with reflecting vesicles of approximately 0.8 μm diameter. Though it appears too early to offer a final and decisive conclusion as to the nature and origin of the eye of D. rathkei, comparisons with compound eyes of other peracaridan crustaceans and anatomical parallels to isopod and other malacostracan ocelli make clear that the long‐held view that the cumacean photoreceuptor represents a regressed compound eye is not necessarily the correct one.
External ultrastructure of sensory organs in the subfamily lrumuinae (Arachnida, Opiliones, Assamiidae)
Authors: KAURI HANS
Published: July 2005

The external ultrastructure of sensory organs in edaphic reprerentatives of the subfamily Irumuinae is described. The types of receptor are either bristle‐shaped or bulbiform. The bristle‐shaped receptors occur on the ventral side of the coxae and displace sensory hairs there. The bulbiform ones are distributed over the scute and on proximal joints (femur, trochanter) of legs. These structures are considered as adaptations to the soil environment.
Three new genera of Braconini from Australasia and Malaysia (Insecta, Hymenoptera, Braconidae)
Authors: QUICKE DONALD L. J.
Published: July 2005

Ancilibracon gen.n. (type species: Ancilinracon townesi sp.n.). Ancilibracon bakeri sp.n., Gelasinibracon gen.n. (type species: Gelasinibracon sedlaceki sp.n.), Gelasinibracon simplicicaudatus sp.n., Esengoides gen.n. (type species: Esengoides crentdatus sp.n.) and Esengoides fulvus sp.n. are each described and illustrated. Ancilibracon towensi is from Malaysia. A. bakeri is from Borneo, both Gelasinibracon species are from Guinéa, E. crenulatus is from the solomon Islands and E. fulvus is from Australia. These three new genera belong to the Plesiobrscon Cameron group of the Braconina. Their relationship are discussed.
Acregoliathidae (Osteichthyes, Teleostei), a new family of fishes from the Cenozoic of Acre State, Brazil
Authors: RICHTER MARTHA
Published: July 2005

A new fossil fish family (Acregoliathidac) with only one known species (Acregoliathidac rancii gen. et sp.n.) is described. The material consists of large Scales coming from the Plio‐Pleistocene of Acre State, Northern Brazil. Studies involving X‐ray radiograph, SEM and optical microscopic methods demonstrated the peculiarity of the new materia1. whose affinities with the classical fish scale models are hardly evident. The scales of A. rancii are round, with large exposed and emberddted areas. They bear concentric circuli and the posterior fields is covered with many layers of a glassy and birefringent tissue. Two bony layers are clearly distinguishable: the basal isopedin and, above, an important cell‐containing spongiosa (protohaversian bone tissue). The affinities of the fossil with both sarcopterygian and actinoplerygian fishes are discussed and the Acregoliathidac are considered to be teleostean fishes of uncertain systematic relationships with other families or oders of this subdivision.
Fine structure of the nematocytes of Polypodium hydriforme Ussov (Cnidaria)
Authors: RAIKOVA EKATERINA V.
Published: July 2005

The fine structure of the stinging cells (nematocytes) and stinging capsules (nematocysts) is described for Polypodium hydriforme. a freshwater coclenterate with a prominent endoparasitie stage in its life cycle. All the nematocysts belong to the type of lesser glutinants (atrichous isorhiza) and fall into three size classes. The internal structure of the capsules is similar in the three classes. A novel type of organization of the cnidocil apparatus of the nematocysts is described. The cnidocil lacks a root fibre and its kinctosome sits directly on the operculum of the nematocyst, so that the entire cnidocil apparatus has a radial rather than bilateral symmetry. It is compared with that of other types of nematocytes and its similarity with the mechanoreccptors of the coelentcratcs is noted. The possible place of the Polypodium nematocytes in the evolution of the collar receptors of the Metazoa is discussed.
A monograph of the Dimarcusidae (Platyhelminthes, Seriata, Tricladida)
Authors: SLUYS RONALD
Published: July 2005

The triclad family Dimarcusidae contains the species Rhodax evelinae, Opisthobursa mexicana, O. josephinae, Balliania thetisae and Mitchellia sarawakana. A comprehensive description is given of each of these species, covering external features, alimentary system, reproductive system, life cycle, ecology and distribution. Postulated apomorphies of the Dimarcusidae are (1) a penis bulb with gland cells, (2) horizontal orientation of female genital duct or bursal canal, combined with a dorsal opening of the common oviduct or diverticulum and (3) location of the ovaries at some distance posterior to the brain. The genera Rhodax, Opisthobursa and Mitchellia form a monophyletic group, the sistergroup being B. thetisae. The genera Rhodax and Opisthobursa share a sistergroup relationship. The Dimarcusidae should not be classified among the Maricola, Terricola or Paludicola. The family represents a fourth lineage within the Tricladida and is more closely related to the Paludicola and Terricola than to the Maricola.
Chromadorida (Nematoda) from Guadeloupe and Polynesia with evidence of intersexuality
Authors: GOURBAULT NICOLE , VINCX MAGDA
Published: July 2005

Spirobolbolaimus boucherorum sp.n. (Microlaimidac) and Sabatieria maboyae sp.n. (Comesomatidae) arc described, and Dorylaimopsis pellucidum (Cobb, 1920) is discussed. Intersex specimens have been found in all three species. Spirobolbolaimus boucherorum sp.n. is characterized by its multispiral amphidcal fovea, number and localization of the cervical setae arranged in six rows posterior to the amphids, and large banana‐shaped sperm. Sabatieria maboyae sp.n., a member of the S. pulchra group, possesses long posteriorly located cervical setae and long spicules.
Cladistic analysis of the subfamilies within the Tubificidae (Oligochaeta)
Authors: ERSÉUS CHRISTER
Published: July 2005

A hypothesis of phylogenetic relationships within the family Tubificidae is presented, based on a Wagner analysis of morphological characters in the different subfamilies. Two major lincages are recognized. One, including the subfamilies Tubificinae, Telmatodrilinae and Limnodriloidinae, is supported by a synapomorphic ability to form slender spermatozeugmata in the spermathecae; the other, including Rhyacodrilinae (paraphyletic), Phallodrilinae, and the (present family) Naididae, is supported by two synapomorphics, the possession of modified penial setae and numerous coelomocytes (the latter secondarily lost in the Phallodrilinae). Some implications for the classification of the Tubificidae are discussed.
Identity between Ramulus libanicus (Uvarov) and Ramulus turcus (Karabaḡ) (Insecta, Phasmatodea): body, egg and chromosome analysis
Authors: SCALI VALERIO , MANTOVANI BARBARA , MARESCALCHI OMBRETTA
Published: July 2005

A detailed study of bisexual populations of stick insects collected near Samandaḡ (southern Turkey) showed that the brown females were Ramulus turcus (Karabaḡ), while males and greenish females morphologically corresponded to R. libanicus (Uvarov) from Mount Carmel. The identity of these two taxa was further supported by egg morphology and structure, which agree with earlier descriptions of R. libanicus. Karyological analysis furthermore showed a chromosome set of 2n = 30 with an XX‐XY sex‐chromosome mechanism which is unusual for stick insects and has never been described before among Ramulini. This supports the identity of R. turcus and R. libanicus and we conclude that these Turkish and Mount Carmel stick insects belong to the same bisexual species and that its name should be R. libanicus (Uvarov).
A contribution to the Dicyrtomidae (Collembola) of Hawaii
Authors: SNIDER RICHARD J.
Published: July 2005

Ten new species in three subgenera of Dicyrtoma are described from the Hawaiian Islands. Specimens were received from collections made on Hawaii, Maui, Kauai and Oahu. Species definitions are based on chaetotaxy of head, legs and the circumanal region. In addition, presence or absence of lateral spines (neosminthurid setae) on the parafurcular lobes may assist in grouping species within subgenera. Previous records of dicyrtomids from Hawaii include only one species, Ptenothrix (Papaairioides) daubia(Folsom). The following new species are described: Ptenothrix(Ptenothrix) hawaiicnaeis sp.n., Ptenothrix (Papirioides) kauaiensis sp.n., P. (Papirioides) serrata sp.n., Dicyrtoma (Calvatomina) sylvestratilis sp.n., D. (Calvatomina) brevifibra sp.n., D. (Calvatomina) tesselata sp.n., D. (Calvatomina) longidigita sp.n., D. (Calvatomina) bellingeri sp.n., D. (Calvatomina) madestris sp.n., and D. (Calvatomina) microdentata sp.n.
Gibson's reclassification of the enoplan nemerteans (Enopla, Nemertea): a critique and cladistic analysis
Authors: SUNDBERG PER
Published: July 2005

Gibson's (1988) reclassification of the class Enopla (phylum Nemertea), and in particular the forming of the taxon Paramonostilifera, is assessed by a cladistic analysis based on 35 characters. This analysis does not support the reclassification, and it is suggested that the previous classification is kept for the time being, although it is recognized that certain of the higher categories are non‐monophyletic and should be reclassified.
Multivariate analysis of morphometric differentiation within the Laonice cirrata‐group (Polychaeta, Spionidae)
Authors: ORRHAGE LARS , SUNDBERG PER
Published: July 2005

Principal component, and canonical variate analysis based on six characters show that there are morphological differences between three Laonice forms (L. bahusiensis, L. sarsi, and Laonice sp.). This supports Söderström's division of L. cirrata into four species; a decision often questioned by other taxonomists. We cannot exclude the possibility that the morphological differences are caused by polymorphism or seasonal variation within a single species. There is, however, no information to vindicate this and we conclude that the three forms should be regarded as separate species until this hypothesis can be rejected.
Ontogenetic changes in the trochoidean (Archaeogastropoda) radula, with some phylogenetic interpretations
Authors: WARÉN ANDERS
Published: July 2005

Radulae have been examined of juvenile specimens of 18 species belonging to the Turbinidae, the Tricoliidae. and 6 subfamilies of Trochidae. The investigations show that all species pass through profound changes, usually finishing at a size of 1.5–3 teleoconch whorls. Most species have an identical, shared morphology at a size of up to 1.5–2 teleoconch whorls, except the species of Calliostomatinac and Solariellinae. The evolutionary consequences are discussed and it is suggested that such an unspecialized starting point means a high potential for specializations and future evolution. For the taxonomist this means a reduction in the reliability of radular differences as a systematic tool.
Vipera pontica sp.n., a new viper species in the kaznakovi group (Reptilia, Viperidae) from northeastern Turkey and adjacent Transcaucasia
Authors: BILLING HERBERT , NILSON GÖRAN , SATTLER UWE
Published: July 2005

Vipera pontica sp.n. from northeastern Turkey and Transcaucasia is a member of the Vipera kaznakovi group characterized by partial fragmentation of frontal and parietals, sharp canthus rostralis, and a colour pattern on the head of the Vipera dinnicki type. It differs from all vipers in the Near and Middle East in having an upturned but hornless snout, and in a number of scalation and colour pattern characters. It is most similar to the west European V. aspis, but differs in having a yellow green tail tip, more loreals and fewer subcaudals. The affinities to the latter species as well as to V. barani are discussed.
Revision of the Antarctic genus Notaeolidia (Gastropoda, Nudibranchia), with a description of a new species
Authors: WÄGELE HEIKE
Published: July 2005

New material from the Atlantic sector of the Antarctic Ocean allows a revision of the Antarctic genus Notaeolidia Eliot, 1905. One new species (Notaeolidia schmekelae sp.n.) is described. N. depressa Eliot, 1905 and N. gigas Eliot, 1905, which arc poorly known, arc redescribed. N. rufopicta Thiele, 1912, N. robsoni Odhner, 1934, N. subgigas Odhner, 1944, N. alutacea Minichev, 1972 and N. flava Minichev, 1972 are synonymized with N. depressa. N. purpurea Eliot, 1905 is synonymized with N. gigas. The monogeneric family Notaeolidiidae Odhner, 1926. is characterized by the following autapomorphy: “nephroproct in front of the genital apertures”. Too little is known to elaborate the phylogenetic position of the family Notaeolidiidae within the cladobranch Nudibranchia.
A synopsis of the genus Atypus (Araneae, Atypidae)
Authors: SCHWENDINGER PETER J.
Published: July 2005

Twelve of sixteenAtypusspecies previously described, and one undescribed, are examined.Atypus javanusThorell is re‐diagnosed and the male is described. The vulva of this species, and ofA. coreanusKim andA. karschiDönitz, arc illustrated for the first time. The taxonomic characters given by Kraus & Baur [Verh, naturw. Ver. Hamb. 17: 85–116(1974)] and additional details of male‐and female genital organs are discussed. A granular texture on male chelicerae and front legs, presumably a stridulatory apparatus, is noted and used for distinguishing species. A deep cymbial pit of unknown function is illustrated and discussed. Specific character combinations for the species examined are presented in a table. Notes on relationships, habitat, zoogeography, web, phenology and development are given.
Loricate choanoflagellates (Acanthoecidae, Choanoflagellida) from the Weddell Sea, Antarctica
Authors: THOMSEN HELGE A. , BUCK KURT R. , COALE SUSAN L. , GARRISON DAVID L. , GOWING MARCIA M.
Published: July 2005

A study of the choanoflagellate species composition of an Antarctic ice edge zone (northern Weddeil Sea, March 1986) has resulted in the finding of 16 taxa of which one,Cosmoeca takahashiiThomsen sp.n., is described here. An emended description is given ofParvicorbicula circularis.Morphological, numerical and dimensional aspects of other species have been analysed in an attempt to improve the circumscription of certain taxa, and to increase our understanding of the nature of intraspecific variability. Data on the relative abundance of choanoflagellate species along seaward transects perpendicular to the ice edge showed that seven species account for more than 95% of all choanoflagellates identified. The two most abundant species wereParvicorbicula socialisandDiaphanoeca pedicellata.
Pipunculidae (Diptera) from Papua New Guinea: the genera Cephalosphaera, Cephalops and Beckerias
Authors: DE MEYER MARC , GROOTAERT PATRICK
Published: July 2005

Seven new pipunculid species are described from Papua New Guinea: Cephalosphaera gymne sp.n., C. immodica sp.n. C. wauensis sp.n., Cephalops bifidus sp.n., C. multidenticulatus sp.n., C. papuaensis sp.n., and C. parmatus sp.n. Cephalops argutus (Hardy) is considered as a true Beckerias sp. and thus placed in a new combination: Beckerias argutus. The phylogenetic relationship of the new species is shortly discussed.
Revision of the genus Elaphostrongylus Cameron (Nematoda, Metastrongyloidea) with particular reference to species of the genus occurring in Norwegian cervids
Authors: GIBBONS LYNDA M. , HALVORSEN O. , STUVE G.
Published: July 2005

The three species of the genus Elaphostrongylus occurring in Norway. namely, E. cervi, E. rangiferi and E. alces from Cervus elaphus, Rangijer tarandus and Alces alces respectively are redescribed and figured using light and scanning electron microscopical techniques. The taxonomic status of the three species is reviewed and their validity confirmed.
Four new species of Hydromedusae (Cnidaria, Hydrozoa) from the coast of south‐western Africa
Authors: PAGÈS FRANCESC , BOUILLON JEAN , GILI JOSEP‐MARIA
Published: July 2005

Four previously unknown species of hydromedusae are described from plankton samples collected during oceanographic cruises undertaken along the south‐western coast of Africa. The Anthomedusa Bythotiara capensis sp.n., known from a single specimen, was found in shallow waters of the Cape region (South Africa). The Leptomedusa Margalefia intermedia gen. et sp.n. collected in Namibian waters, possesses desmoneme cnidocysts. This category of cnidocysts is previously unreported in the Leptomedusae, but is common in Anthomedusae and in the limnomedusan family Proboscidactylidae. Margalefia seems to occupy a position intermediate between the families Tirannidae and Laodiceidae. Two species of Limnomedusae, Aglauropsis edwardsii sp.n. and Proboscidactyla menoni sp.n. were frequent at several stations in Namibian waters between depths of 50 m and the surface.
Gnathostomulida from Hawaii
Authors: STERRER WOLFGANG
Published: July 2005

As thc second in a series on Gnathostomulida from the Pacific Ocean this paper describes an additional eight species from Hawaii of which four are new to science. All are accommodated within existing genera: Haplognathia asymmetrica sp.n., Haplognathia ruberrima (Sterrer, 1966), Haplognathia rufa sp.n., Cosmognathia arcus Sterrer, 1991, Cosmognathia manubrium sp.n., Pterognathia ctenifera Sterrer, 1970, Pterognathia hawaiiensis sp.n., and Austrognathia novaeze landiae Sterrer, 1991.
Gnathostomulida from Tahiti
Authors: STERRER WOLFGANG
Published: July 2005

This paper deals with nine species of Gnathostomulida from the Pacific Ocean, all in established genera. Of the nine species one is cosmopolitan (Haplognathia rosacea (Sterrer, 1970)), four have been described previously from other localities in the Indo‐Pacific (Cosmognathia manubrium Sterrer, 1991; Gnathostomula maldivarum Gerlach, 1958; Austrognathia nannulifera Sterrer, 1991; Austrognatharia pecten Sterrer, 1991), and four are new to science (Pterognathia ugera sp.n.; Gnathostomula algreti sp.n.; Austrognatharia macroconifera sp.n.; Austrognatharia mooreensis sp.n.).
Pseudocatamachilis gen.n., a new genus of Machilidae (Insecta, Apterygota, Microcoryphia) from Spain
Authors: GAJU‐RICART MIQUEL , de ROCA CARMEN BACH
Published: July 2005

A new genus, Pseudocatamachilis gen.n. is established on the basis of a thorough study of many specimens male and female of Catamachilis torquata (Nav#aAs, 1905). All type material was lost during the Spanish civil war (1936‐39). A neotype is designated and the species redescribed. Genus Pseudocatamachilis is related to Catamachilis Silvestri and Promesomachilis Silvestri, but distinguished from these genera by the type of ovipositor and the presence of two pairs of parameres, among other characteristics given in the text.
Coscinoderma sporadense sp.n. from the Aegean Sea with comments on Coscinoderma confragosum (Porifera, Dictyoceratida)
Authors: VOULTSIADOU‐KOUKOURA ELENI , VAN SOEST ROB W. M. , KOUKOURAS ATHANASIOS
Published: July 2005

Coscinoderma sporadense sp.n. is described from shallow waters in the North Aegean. This is the first record of a Coscinoderma species from the Mediterranean. The new species is compared to the deep water East Atlantic Coscinoderma confragosum Poléjaeff, 1884, and clear differences between them are established. The status of the genus Coscinoderma is discussed.
Crassophyllum thessalonicae sp.n. (Octocorallia, Pennatulacea), from the Aegean Sea
Authors: VAFIDIS DIMITRIS , KOUKOURAS ATHANASIOS
Published: July 2005

A new pennatulacean species, Crassophyllum thessalonicae sp.n. is described from the Mediterranean (North Aegean Sea). This is the first record of a Crassophyllum Tixier‐Duri vault, 1961 species from the Mediterranean Sea. The differences between the new species and C. cristatum Tixier‐Durivault, 1961, the other known species of the genus, are discussed, and the genus Crassophyllum is critically compared with the other two related genera Pteroeides Herklots, 1858 and Sarcoptilus Gray, 1848.
Zeylanurotrerna spearei sp.n. (Digenea: Brachylaimidae) from the cane toad, Bufo marinus, in Australia
Authors: CRIBB THOMAS H. , BARTON DIANE P.
Published: July 2005

Zeylanurotrema spearei sp.n. is described from the urinary bladder of the introduced cane toad, Bufo marinus, from North Queensland, Australia. The other species in Zeylanurotrema, Z.lyriocephali, has been recorded only from an agamid lizard from Sri Lanka. The new species has differently arranged gut caeca, vitelline follicles and ovary as compared with Z. lyriocephali. Z. spearei has a Laurer's canal that does not open to the exterior but which forms a glandular Juel's organ. We suggest that Z. spearei is an Australian species that has been acquired by the cane toad from a native host. In juvenile specimens of Z. spearei a persistent microcercous tail is present. From this and the anatomy of the adult we conclude that Zeylanurotrema is a brachylaimid rather than a urotrcmatid as originally proposed. A new subfamily, the Zeylanurotrematinae, is proposed to accommodate Zeylanurotrema.
Two new species of Achaeta (Enchytraeidae, Oligochaeta) from meadow and pasture soils of Germany
Authors: HECK MICHAEL , ROMBKE JOERG
Published: July 2005

Two new, closely related species of the genus Ackaeta, A. urbana sp.n. and A. microcosmi sp.n., (Enchytraeidae, Oligochaeta) from meadow and pasture soils of Berlin and Frankfurt (Germany) are described. Both show the same form and length/width ratio of the seminal funnel and the same general shape of the spermathecal system. They are distinguishable by distributions of epidermal sacs: only dorsal (A. microcosmi) or dorsal and ventral (A. urbana). Also the origin of the dorsal blood vessel is different.
Two new species and a phylogenetic analysis of the genus Tectidrilus (0Iigochaeta, Tubifcidae)
Authors: ERSÉUS CHRISTER
Published: July 2005

Tectidrilus probus sp.n. and Tectidrilus profusus sp.n. are described from offshore localities in southern California. Both species appear closely related to T. diversus Ersëus, 1982, which occurs in the same area, but they differ from the latter by possessing small oesophageal diverticula; T. probus differs also by its very small spermathecal vestibules. Tectidrilus profusus is a highly apomorphic form with elaborate, partly muscular, male ducts and spermathecae. Monophyly of Tectidrilus is supported by the short clitellum and the unisetal ‘bundles’ in postclitellar segments. Patterns of apomorphic character states within the genus are analyzed under the principle of Camin‐Sokal parsimony (irreversibility of character states), giving three equally parsimonious cladograms. In the strict consensus tree of these cladograms, eight of the twelve members of Tectidrilus form an apomorphic group defined by the papillated body wall, but the individual species within this group cannot be unequivocally resolved from each other.
Composition and preservation of the Chengjiang fauna –a Lower Cambrian soft‐bodied biota
Authors: XIANGUANG HOU , RAMSKÖLD LARS , BERGSTRÖM JAN
Published: July 2005

The Lower Cambrian Chengjiang fauna is reviewed and shown to be closely comparable with the younger Burgess Shale fauna. but with various differences in detail. A diverse group of more or less annulated lobopod animals including ‘armoured lobopods’ are regarded as representatives of the phylum Onychophora. ‘Trilobitomorphs’ include several new types. Probable protaspides of the trilobitomorph Naraoia are described. No molluses or deuterostomes have been identified. The preservational orientations of the various taxa are reviewed and compared with orientations of the Burgess Shale taxa. Orientation in the sediment is found to be closely correlated to the original shape of individuals. Several new genera and species are described: the segmented. worm‐shaped Yunnanozoon lividum gen. et sp.n., the ‘armoured lobopods’Onychodictyon ferox gen. et sp.n. and Cardiodictyon catenulum gen. et sp.n. and the arthropods Saperion glumaceum gen. et sp.n., Sinoburius Iunaris gen. et sp.n., and Xandarella spectaculum gen. et sp.n.
Relationships among turtles of the genus Clemmys (Reptilia, Testudines, Emydidae) as suggested by plastron scute morphology
Authors: LOVICH JEFFREY E. , LAEMMERZAHL ARNDT F. , ERNST CARL H. , McBREEN JOHN F.
Published: July 2005

The Conservative morphology of hardshelled turtles has fostered the use of size relationships between epidermal scutes (scales) on the shell to differentiate between species and subspecies of many taxa. The size relationship of the six major pairs of plastral scutes were used to compare the four currently recognized species of the genus Clemmys with each other. as well as with the distantly related Graptemys barbouri using Jaccard Coefficients. Shannon‐Weiner diversity indices, and multivariate analysis. Results were concordant among the three techniques used and confirm our prediction that plastral morphology varies little among closely related species and widely among distantly related taxa. Clemmys muhlenbergii appears to he more different from Clemmys guttata than previously suggested. Analysis of plastral morphology shows promise as a taxonomic tool for turtle systematists.
A generic revision of the Phallodrilinae (Oligochaeta, Tubificidae)
Authors: ERSÉUS CHRISTER
Published: July 2005

The generic classification of the subfamily Phallodrilinae (with a total of 252 species) is revised according to phylogenetic principles. Morphological character patterns are assessed under the principle of maximum parsimony, but due to the poor resolution and instability in the resulting cladograms, the analysis is supplemented with intuitive weighting of synapomorphic similarities. Thirty‐one genera are recognized. The former definition of the type genus Phallodrilus Picrantoni, 1902, is revised and the members of this genus are reclassified into 20 taxa, each hypothesized to be monophyletic: Phallodrilus s.str. (revised to contain the type species only), Somalidrilus gcn.n., Milliganius gen.n., Phallodriloides gen.n., Albanidrilus gen.n., Abyssidrilus gen.n., Paraktedrilus gen.n., Uniporodrilus Erséus, 1979, Inermidrilus gen.n., Gianius gen.n., Mexidrilus gen.n., Peosidrilus Baker & Erséus, 1979, Atlantidrilus Erséus, 1983, Thalassodrilus Brinkhurst, 1963, Pirodrilus gen.n., Pirodriloides gen.n., Marionidrilus gen.n., Pectinodrilus gen.n., Pacifidrilus gen.n. and Pseudospiridion gen.n. Nine new species are described from the Northwest Atlantic: Phallodriloides pinnulatus sp.n., Abyssidrilus opulentus sp.n., A. potens sp.n., A. altoides sp.n., Mexidrilus obtusus sp.n., Peosidrilus dalei sp.n., P. aduncus sp.n., P. riseri sp.n., and Atlantidrilus hamulus sp.n. The genus Discordiprostatus Baker, 1982, is synonymized with Nootkadrilus Baker, 1982.
Caridina bruneiana, a new species of freshwater shrimp (Decapoda, Caridea, Atyidae) from Negara Brunei Darussalam, Borneo
Authors: CHOY SATISH C.
Published: July 2005

A new species of the freshwater atyid shrimp, Caridina bruneiana from Negara Brunei Darussalam, Borneo is described. Caridina bruneiana is characterized by its moderately long and deep rostrum, pereiopod shape, spination and segmental ratios, the shape and spination of its posterior telsonic margin, egg size and the presence of an appendix interna on the endopod of the first male pleopod. The distribution of atyid shrimps in forested streams and their possible use as bioindicators of stream water quality in tropical rainforests are briefly discussed.
New deep‐sea Tardigrada (Arthrotardigrada, Halechiniscidae) from a manganese nodule area of the eastern South Pacific*
Authors: BUSSAU CHRISTIAN
Published: July 2005

Three new Tardigrada species are described from a manganese nodule area of thc abyssal castern South Pacific. Moebjergarctus mangunis gen. et sp.n. is characterized by its segmcntcd ccphalic cirri, club‐shaped and forward bent primary clavae, spherical secondary clavae, spines on legs I, 11, 111, small clavoid papillae on legs IV, long cirri E, and corkscrew‐shaped digits with simple claws. Angursa capsula sp.n. is distinguished by the presence of secondary and tertiary clavac, occurring as elevated cuticular areas, club‐shaped primary clavae, short lateral cirri, short median cirrus, spines on legs I and hcmisphcrical, capsulc‐shaped papillae on legs IV. Angursa lingua sp.n. is characterized by the presence of secondary and tertiary clavae, occurring as elevated cuticular areas, balloon‐shaped primary clavae, long lateral cirri, long median cirrus, spines on legs I, clavoid papillae on legs IV and weakly broadened distal parts of cirri E, resulting in tongue‐shaped swellings. A simple key to the species of Angursa is presented.
New Pferastericola species (Platyhelminthes, Rhabdocoela) and a cladistic analysis of the Pterastericolidae
Authors: JONDELIUS ULF
Published: July 2005

Two ncw species of the family Pterastericolidae which inhabit starfish arc dcscribcd. Pterastericola bergensis sp.n. from thc oesophagus and cardiac stomach of Psilaster andromecla has posterior gonopore and lateral vitellaria. Pterastericola asamushii sp.n. from the pyloric caeca of Astropecren scopurius has unciliated epithelium postcriorly and an anterior gonopore. A cladistic analysis of thc phylogcny of the Pterastcricolidae is presented. The withdrawal of the genus Trilohorhynchus is proposed.
Identification of the Kalyptorhynchia (Plathelminthes) in Meixner's ‘Turbellaria’ 1938 with remarks on the morphology and distribution of the species in the North Sea and the Baltic Sea
Authors: KARLING TOR G.
Published: July 2005

Remarks arc given on taxonomy, structure and distribution of 38 species of Kalyptorhynchia, incompletely described (in part nomina nuda) in part I of ‘Turbellaria (Strudclwürmer)’ in Tierwelt der Nord‐und Ostsee (Meixncr 1938). The remarks are based mainly on the existing parts of the basic material for Meixner's study in the collections of the Zoological Institution in Graz, Austria, including incomplete proof sheets of the unpublished part II. Available information on the species published after 1938 has been taken into account.
Platynereis nadiae sp. n. (Polychaeta: Nereididae) from Italian coasts
Authors: ABBIATI MARCO , CASTELLI ALBERTO
Published: July 2005

A new species, Platynereis nadiae sp. n. (Polychacta: Ncrcididac), from thc Mediterranean Sea is described. It is clearly distinguished from the other Platynereis species by the distribution and the morphology of the homogomph falciger setae and by the structure of thc tcntacular cirri.
Axiid collections of the Zoological Museum, Copenhagen, with the description of one new genus and six new species (Axiidae, Thalassinidea, Crustacea)
Authors: SAKAI KATSUSHI
Published: July 2005

The collections of the family Axiidae in the Zoological Museum. Copenhagen. were examined: seven genera including one new genus, Eucalastacus gen.n. and 14 Indo‐Pacific species containing five species, Eiconaxius farreae, E. weberi, E. sibogae, Axius australiensis and Axiopsis serratifrons; six new species, Eiconaxius demani sp.n., E. mortenseni sp.n., Bouvieraxius keiensis sp.n., Eucalastacus torbeni sp.n., Axiopsis tsushimaensis sp.n., and Calocarirdes vigila sp.n.; and three uncertain species. Eutrichocheles aff. brocki, Eutrichochelts sp. and Axiopsis sp. Postorlocaris. Sakaiocaris and Ca1listocaris are referred, respectively. as junior synonyms of Bouvieraxius. Spongiaxius and Ambiaxius.
Further data on Prokennalestes (Mammalia, Eutheria inc. sed.) from the Early Cretaceous of Mongolia
Authors: SIGOGNEAU‐RUSSELL DENISE , DASHZEVEG DEMBERLYIN , RUSSELL DONALD E.
Published: July 2005

Four new incomplete lower jaws of Prokennalestes are described. bringing new data for establishing the primitive cutherian dental formula and its evolution in later placentals, as well as the possible relationships of the genus.
Pseudomastus deltaicus gen. et sp.n. (Polychaeta: Capitellidae) from a shallow water bay in the North‐ Western Mediterranean Sea
Authors: CAPACCIONI‐AZZATI ROMANA , MARTÍN DANIEL
Published: July 2005

Specimens of Pseudomastus deltaicus gen. et sp.n. (Capitellidae), were collected from muddy bottoms in Els Alfacs Bay (Ebre Delta), a semi‐enclosed shallow water area. The main diagnostic characteristics are the possession of 12 thoracic setigers, of which the first 10 have only capillary setae and the last two have capillary notosetae and neuropodial hooded hooks, and presence of branchiae in the abdominal region. Pygidium bearing three anal cirri.
Trochonerilla mobilis gen. et sp.n., a meiofaunal nerillid (Annelida, Polychaeta) from a marine aquarium in Moscow
Authors: TZETLIN ALEXANDER B. , SAPHONOV MICH AIL V.
Published: July 2005

A nerillid species new to science is described from the marine aquarium system of Moscow Zoo. Trochonerilla mobilis gen. et sp.n. is characterized by eight chaetigerous segments, three antennae, absence of palps, parapodia with two bundles of simple chaetae and pygidium with two anal cirri. In contrast to other eight‐segmented nerillids, Trochonerilla mobilis are very active and mobile animals. They are also able to swim through the water column by means of the strong ring of cilia on the first chaetigerous segment. Their geographical origin is unknown.
The pterolichoid feather mites (Acarina, Astigmata) of the Megapodiidae (Aves, Galliformes)
Authors: ATYEO WARREN T.
Published: July 2005

The pterolichoid feather mites of megapodes are reviewed. Named taxa are briefly discussed and most are figured. The Pterolichidae (Pterolichinae) are: Ascetohchus Perez & Atyeo, three species; Echinozonus Atyeo & Perez, six species; Pereziella Atyeo, two species; and Phycoferus Atyeo & Perez, two species. New pterolichine taxa, with the type species listed first, include: Botryaspis cordiforma gen. et sp.n. and B. cordata gen. et sp.n.; Cycloprotarsus lineatus gen. et sp.n., C. centralis gen. et sp.n. and C. monacrotrichus gen. et sp.n.; Eurypterolichus gen.n. for Pterolichus navicula comb.n. Trouessart & Neumann and E. coniger gen. et sp.n.; Goniodurus gen.n. for Pterolichus (Pseudalloptes) quadratus comb.n. Trouessart and G. bilobatus gen. et sp.n.; Haptepigynus gen.n. for Pterolichus (Pseudalloptes) tridentiger comb.n. Trouessart and H. holonotus gen. et sp.n.; Heliaspis ventralis gen. et sp.n.; Leipobius ocellatus gen. et sp.n.; Maleolichus maleo gen. et sp.n.; Mayracarus gen.n. for Pterolichus (P.) tritilobus comb.n. Trouessart; Megapodobius arcuatus gen. et sp.n. and M. striatus gen. et sp.n.; Oxygynurus brevissimus gen. et sp.n., O. longicaulis gen. et sp.n., O. mediocaulis gen. et sp.n. and O. parvicaulis gen. et sp. n.; Prionoturus amembranatus gen. et sp. n.; Talegallobius bidentatus gen. et sp.n.; and Tanysomacarus imperfectus gen. et sp.n. and T. brachymeles gen. et sp.n. A new taxon of the Thoracosathesidae is: Thoracosathes caudiculata sp.n. Keys are provided and host‐ commensal associations are discussed. All taxa are restricted to the Megapodiidae. Cheylabis fuscina Trouessart is assigned to Pereziella and has as a synonym P. dupilcata Atyeo.
Telmapsyllagen.n., an unusual psyllid from black mangrove in Florida and Costa Rica (Insecta, Homoptera, Psylloidea)
Authors: HODKINSON IAN D.
Published: July 2005

Telmapsylla minuta. gen. et sp.n., a diminutive species of psyllid collected from black mangrove,Avicenniaspecies (Verbenaceae), is described from Florida and Costa Rica.Telmapsyllais particularly significant in that it possesses characters currently used to define both the families Psyllidae and Calophyidae. Its taxonomic position and host‐plant relationships are discussed.Telmapsyllais most probably a primitive calophyid.
Ultrastructure of spermiogenesis and spermatozoa of four Oriopsis species (Sabellinae, Sabellidae, Polychaeta)
Authors: ROUSE GREG W.
Published: July 2005

The ultrastructure of sperm from four species of Oriopsis is described. Males of Oriopsis bicoloris produce sperm with an elongate nucleus divided into four rods, connected proximally, and four long mitochondria lying along the nucleus. The axoneme runs in the middle of the nuclear and mitochondria1 rods. Oriopsis brevicollaris males have sperm with an elongate nucleus and long midpiece comprised of four mitochondria wrapped around the axoneme. Males of O. mobilis have sperm with an elongate nucleus and long midpiece similar to that of O. brevicollaris, although the midpiece is much longer. Oriopsis dentata males havc sperm with a flattened head, similar to those of mammals, though the midpiece is simple and the axoneme free. The variability of reproductive mechanisms within the Polychaeta is discussed with reference to the elucidation of the function of various sperm morphologies. The implications for the taxonomy and systematics of Oriopsis, and the Sabellidae as a whole, are also discussed. It is concluded that Oriopsis is not monophyletic and examination of reproductive structures in other small sabellids is required. Study on reproduction in the type species, O. armandi, is needed to establish the reproductive method of Oriopsis and so allow revision of this genus.
Revision of Trachoderna Blanchard with the description of an allied genus from central Chile (Insecta, Coleoptera, Curculionidae)
Authors: MORRONE JUAN J.
Published: July 2005

Trachodema Blanchard includes the species T. tuberculosa Blanchard and T. minuta (Hustache). Lumiarhinus gen.n. is established for L. aelficus gen. ct sp.n. and L. horridus (Germain) comb. n. Both genera are endemic to central Chile. They have tuberculate elytra, but arc separated on the basis of the body vestiture, presence or absence of frontal fovea, depth of the scrobes, shape of the hind margin of the prothorax, and shape of the elytra. Diagnostic characters. dcscriptions and illustrations of the genera and species are given and keys to the species are presented. Lectotypes are designated for T. minuta and L. horridus.
Shallow water rissoiform gastropods from Tristan da Cunha, South Atlantic Ocean, with records of species from Gough Island
Authors: WORSFOLD T. M. , AVERN G. , PONDER W. F.
Published: July 2005

Shallow water rissoiform gastropods collected by the Norwegian Scientific Expedition (1937‐1938) to Tristan da Cunha are described. The fauna consists of Powellisefia cf. philomelae (Watson) (Rissoidae), three new species of Eatoniella, E. trochiformisa, E. lineuta, and E. tristanensis (Eatoniellidae). two new species of Onoba, O. crassicordara and O. tristanensis (Rissoidae) and Rissoella cf. irma (Bartsch) (Rissoellidae). A small collection from Cough Island is also described, containing three species referred to Onoha, one of which is described as new. O. merelinoides, and a species of Eutoniella. The biogeographie relationships of the fauna are briefly discussed.
The Onisimus problem (Amphipoda, Lysianassoidea, Uristidae)
Authors: LOWRY JAMES K. , STODDART HELEN E.
Published: July 2005

Redescriptions ofAnonyx litoralisKrølyer, 1845 (type species ofOnisimusBoeck, 1871 andPseudalibrotusDella Valle, 1893),Anonyx edwardsiiKrøyer, 1846 (type species ofBoeckosimusBarnard, 1969) andParonesimus barentsiStebbing, 1894 (type species ofParonesimusStebbing, 1894) show that these species are all congeneric. Consequently,Pseudalibrotusbecomes an objective junior synonym andParonesimusandBoeckosimusbecome subjective junior synonyms ofOnisimusand all species currently assigned to any of these three genera are provisionally assigned toOnisimus.